Monograph |
Corresponding author: Alexander Riedel ( riedel@smnk.de ) Academic editor: Miguel Alonso-Zarazaga
© 2014 Alexander Riedel, Rene Tänzler, Michael Balke, Cahyo Rahmadi, Yayuk R. Suhardjono.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Riedel A, Tänzler R, Balke M, Rahmadi C, Suhardjono YR (2014) Ninety-eight new species of Trigonopterus weevils from Sundaland and the Lesser Sunda Islands. ZooKeys 467: 1-162. https://doi.org/10.3897/zookeys.467.8206
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The genus Trigonopterus Fauvel, 1862 is highly diverse in Melanesia. Only one species, Trigonopterus amphoralis Marshall, 1925 was so far recorded West of Wallace’s Line (Eastern Sumatra). Based on focused field-work the fauna from Sundaland (Sumatra, Java, Bali, Palawan) and the Lesser Sunda Islands (Lombok, Sumbawa, Flores) is here revised. We redescribe T. amphoralis Marshall and describe an additional 98 new species: T. acuminatus sp. n., T. aeneomicans sp. n., T. alaspurwensis sp. n., T. allopatricus sp. n., T. allotopus sp. n., T. angulicollis sp. n., T. argopurensis sp. n., T. arjunensis sp. n., T. asper sp. n., T. attenboroughi sp. n., T. baliensis sp. n., T. batukarensis sp. n., T. bawangensis sp. n., T. binodulus sp. n., T. bornensis sp. n., T. cahyoi sp. n., T. costipennis sp. n., T. cuprescens sp. n., T. cupreus sp. n., T. dacrycarpi sp. n., T. delapan sp. n., T. dentipes sp. n., T. diengensis sp. n., T. dimorphus sp. n., T. disruptus sp. n., T. dua sp. n., T. duabelas sp. n., T. echinatus sp. n., T. empat sp. n., T. enam sp. n., T. fissitarsis sp. n., T. florensis sp. n., T. foveatus sp. n., T. fulgidus sp. n., T. gedensis sp. n., T. halimunensis sp. n., T. honjensis sp. n., T. ijensis sp. n., T. javensis sp. n., T. kalimantanensis sp. n., T. kintamanensis sp. n., T. klatakanensis sp. n., T. lampungensis sp. n., T. latipes sp. n., T. lima sp. n., T. lombokensis sp. n., T. merubetirensis sp. n., T. mesehensis sp. n., T. micans sp. n., T. misellus sp. n., T. palawanensis sp. n., T. pangandaranensis sp. n., T. paraflorensis sp. n., T. pararugosus sp. n., T. parasumbawensis sp. n., T. pauxillus sp. n., T. payungensis sp. n., T. porcatus sp. n., T. pseudoflorensis sp. n., T. pseudosumbawensis sp. n., T. punctatoseriatus sp. n., T. ranakensis sp. n., T. relictus sp. n., T. rinjaniensis sp. n., T. roensis sp. n., T. rugosostriatus sp. n., T. rugosus sp. n., T. rutengensis sp. n., T. saltator sp. n., T. santubongensis sp. n., T. sasak sp. n., T. satu sp. n., T. schulzi sp. n., T. sebelas sp. n., T. sembilan sp. n., T. sepuluh sp. n., T. seriatus sp. n., T. serratifemur sp. n., T. setifer sp. n., T. silvestris sp. n., T. singkawangensis sp. n., T. singularis sp. n., T. sinuatus sp. n., T. squalidus sp. n., T. sumatrensis sp. n., T. sumbawensis sp. n., T. sundaicus sp. n., T. suturalis sp. n., T. syarbis sp. n., T. telagensis sp. n., T. tepalensis sp. n., T. tiga sp. n., T. trigonopterus sp. n., T. tujuh sp. n., T. ujungkulonensis sp. n., T. variolosus sp. n., T. vulcanicus sp. n., T. wallacei sp. n.. All new species are authored by the taxonomist-in-charge, Alexander Riedel. Most species belong to the litter fauna of primary wet evergreen forests. This habitat has become highly fragmented in the study area and many of its remnants harbor endemic species. Conservation measures should be intensified, especially in smaller and less famous sites to minimize the number of species threatened by extinction.
Southeast Asia, integrative taxonomy, turbo-taxonomy, weevils, hyperdiverse, morphology, cox1 , DNA barcoding, conservation, endemism, Coleoptera , Curculionidae , Cryptorhynchinae
Trigonopterus Fauvel, a genus of flightless weevils placed in the subfamily Cryptorhynchinae of Curculionidae (
The area covered herein includes parts of Sundaland (i.e. Sumatra, Java, Borneo, and the Philippine island of Palawan) and the Lesser Sunda Islands. No species of Trigonopterus is known to us from the Malay Peninsula and the western and central parts of Sumatra. The intensive field work for this study largely focused on East Sumatra, Java and the Lesser Sunda Islands, while material available from only a few localities in Borneo and Palawan could be included. Thus, many more species can be expected from these islands. The fauna of Sulawesi will be treated in a separate publication.
We have previously established that Trigonopterus weevils are suitable for accelerated taxonomic study combining morphology and the DNA barcoding approach using mitochondrial cox1 data (
This study is based on > 4,000 specimens of Trigonopterus, collected specifically for a project on this genus. Unless otherwise stated, specimens were collected by sifting the litter of primary forests with subsequent extraction by hand or, more efficiently, using eclectors (
ARC Alexander Riedel Collection, stored in SMNK, Germany.
MZB LIPI Research Center of Biology, Division of Zoology, Museum Zoologicum Bogoriense, Widyasatwaloka, Cibinong, Indonesia.
SMNK Staatliches Museum für Naturkunde, Karlsruhe, Germany.
ZSM Zoologische Staatssammlung, München, Germany.
The methods applied for DNA sequencing and sequence analysis are described by
Morphological terminology follows
Trigonopterus insignis Fauvel, 1862, by monotypy.
Fully apterous genus of Cryptorhynchinae. Length 1.5–6.0 mm. Rostrum in repose not reaching center of mesocoxa. Scutellar shield completely absent externally. Mesothoracic receptacle deep, posteriorly closed. Metanepisternum completely absent externally. Elytra with 9 striae (sometimes superficially effaced). Tarsal claws minute. Usually body largely unclothed, without dense vestiture. For additional information, see http://species-id.net/wiki/Trigonopterus.
Holotype, male (Fig.
Holotype (MZB): ARC2482 (EMBL # LM655819), East Java Prov., Kediri, Mt. Wilis, Besuki, sample 1, S07°51.785', E111°50.188', 1379 m, 21-XI-2011. Paratypes (MZB, SMNK, ZSM): E-Java Prov.: 17 exx, ARC2483 (EMBL # LM655820), ARC2484 (EMBL # LM655821), same data as holotype; 2 exx, Kediri, Besuki, Mt. Wilis, sample 4, S07°51.852', E111°50.185', 1388 m, 22-XI-2011; 4 exx, Kediri, Besuki, Mt. Wilis, sample 5, S07°51.650', E111°50.174', 1485 m, 22-XI-2011; 1 ex, Kediri, Besuki, Mt. Wilis, sample 6, S07°51.709', E111°50.100', 1409 m, 22-XI-2011.
E-Java Prov. (Mt. Wilis). Elevation: 1379–1485 m.
This epithet is based on the Latin adjective acuminatus (pointed) and refers to the shape of the elytra.
Trigonopterus acuminatus Riedel, sp. n. was coded as “Trigonopterus sp. 357” by
Holotype, male (Fig.
Holotype (MZB): ARC1472 (EMBL # LM655570), West Nusa Tenggara Prov., Lombok, Gn. Rinjani, Tetebatu, Kokok Belimbing, Rinjani-track, sample 3, S08°28.837', E116°26.553', 1245 m, 02-IV-2010. Paratypes (MZB, SMNK, ZSM): West Nusa Tenggara Prov.: 9 exx, ARC1471 (EMBL # LM655569), same data as holotype; 19 exx, ARC0168, Tetebatu, Rinjani-trail, 1200-1450 m, 07-XII-2004; 6 exx, Tetebatu, Rinjani-trail from Orong Gerisak, sample 5, S08°30.096', E116°25.062', 1010 m, 04-IV-2010; 19 exx, ARC1462 (EMBL # LM655560), ARC1463 (EMBL # LM655561), Lombok, Gn. Rinjani, Tetebatu, Orong Gerisak, Rinjani-track, sample 6, S08°29.577', E116°24.782', 1195 m, 04-IV-2010; 10 exx, Lombok, Gn. Rinjani, Tetebatu, Orong Gerisak, Rinjani-track, sample 7, S08°29.433', E116°24.746', 1240 m, 04-IV-2010; 18 exx, ARC1479 (EMBL # LM655577), ARC1480 (EMBL # LM655578), Lombok, Gn. Rinjani, Tetebatu, Orong Gerisak, Rinjani-track, sample 8, S08°29.173', E116°24.517', 1345 m, 04-IV-2010; 3 exx, Lombok, Gn. Rinjani, Tetebatu, Orong Gerisak, Rinjani-track, sample 9, S08°28.981', E116°24.353', 1430 m, 04-IV-2010; 3 exx, ARC2259 (EMBL # LM655713), Lombok, Senaru, Rinjani-track, sample 1, S08°19.429', E116°24.082', 900 m, 21-III-2011; 30 exx, ARC2261 (EMBL # LM655715), ARC2262 (EMBL # LM655716), Lombok, Senaru, Rinjani-track, sample 2, S08°20.570', E116°23.969', 1320 m, 21-III-2011; 10 exx, Lombok, Senaru, Rinjani-track, sample 3, S08°20.780', E116°23.790', 1465 m, 21-III-2011; 8 exx, Lombok, Senaru, Rinjani-track, sample 4, S08°20.439', E116°24.047', 1240 m, 21-III-2011; 13 exx, Lombok, Senaru, Rinjani-track, sample 5, S08°19.800', E116°24.107', 1015 m, 21-III-2011; 7 exx, Lombok, Senaru, Rinjani-track, sample 6, S08°19.359', E116°24.070', 860 m, 23-III-2011; 12 exx, Senaru, Rinjani-trail, sample 7, S08°19.643', E116°24.033', 935 m, 23-III-2011; 14 exx, Senaru, Rinjani-trail, sample 8, S08°19.719', E116°24.040', 955 m, 23-III-2011; 6 exx, ARC2277 (EMBL # LM655731), ARC2278 (EMBL # LM655732), Santong, Rinjani-track, sample 1, S08°20.715', E116°19.695', 830 m, 24-III-2011; 8 exx, Santong, Rinjani-track, sample 2, S08°20.813', E116°19.778', 870 m, 24-III-2011; 4 exx, Santong, Rinjani-track, sample 3, S08°20.996', E116°20.001', 960 m, 24-III-2011; 10 exx, Santong, Rinjani-track, sample 4, S08°21.160', E116°20.067', 1005 m, 25-III-2011; 2 exx, Santong, Rinjani-track, sample 5, S08°21.351', E116°20.208', 1105 m, 25-III-2011; 6 exx, Santong, Rinjani-track, sample 6, S08°21.754', E116°20.476', 1315 m, 25-III-2011; 17 exx, ARC2293 (EMBL # LM655747), ARC2294 (EMBL # LM655748), Gn. Rinjani, Bawnau, near Sembalun, sample 1, S08°21.136', E116°29.257', 1140 m, 26-III-2011; 3 exx, Mt. Rinjani, Bawnau, near Sembalun, S08°21.136', E116°29.257', 1140 m, collected from foliage, 26-III-2011; 44 exx, ARC1522 (EMBL # LM655620), ARC1523 (EMBL # LM655621), ARC1524 (EMBL # LM655622), Sumbawa, Batu Dulang, Mt. Batu Pasak, sample 2, S08°37.028', E117°15.783', 1305 m, 12-IV-2010; 5 exx, Sumbawa, Batu Dulang, Mt. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 12-IV-2010; 12 exx, Sumbawa, Batu Dulang, Mt. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 18-IV-2010; 14 exx, Sumbawa, Batu Dulang, Mt. Batu Pasak, sample 4, S08°37.318', E117°15.339', 1280 m, 18-IV-2010; 11 exx, Sumbawa, Batu Dulang, Mt. Batu Pasak, sample 5, S08°37.005', E117°15.790', 1350 m, 18-IV-2010; 23 exx, ARC1502 (EMBL # LM655600), ARC1503 (EMBL # LM655601), ARC1504 (EMBL # LM655602), West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 2, S08°35.884', E117°08.384', 1310 m, 15-IV-2010; 7 exx, West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 3, S08°35.386', E117°08.251', 1415 m, 15-IV-2010; 1 ex, West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 4, S08°35.168', E117°08.175', 1515 m, 15-IV-2010; 8 exx, West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 5, S08°35.740', E117°08.721', 1330 m, 16-IV-2010; 17 exx, West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 6, S08°35.533', E117°08.605', 1350 m, 16-IV-2010; 30 exx, West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 7, S08°35.176', E117°08.295', 1490 m, 16-IV-2010.
West Nusa Tenggara Prov., Lombok (Santong, Sembalun, Senaru, Tetebatu), Sumbawa (Batu Dulang, Tepal). Elevation: 830–1350 m.
This epithet is a combination of the Latin adjectives aeneus (copper, bronze) and micans (shining) and refers to the metallic lustre of this species´ elytra.
Trigonopterus aeneomicans Riedel, sp. n. was coded as “Trigonopterus sp. 288” by
Holotype, male (Fig.
Holotype (MZB): ARC2326 (EMBL # LM655779), East Java Prov., Banyuwangi Reg., Alas Purwo N.P., Pancur, sample 3, S08°41.533', E114°22.593', 25 m, 09-IV-2011. Paratypes (MZB, SMNK): E-Java Prov., Banyuwangi Reg., Alas Purwo N.P.: 6 exx, ARC2323 (EMBL # LM655776), ARC2324 (EMBL # LM655777), ARC2325 (EMBL # LM655778), Triangulasi, sample 1, S08°39.744', E114°21.976', 25 m, 09-IV-2011; 1 ex, ARC2327 (EMBL # LM655780) same data as holotype; 1 ex, Triangulasi, sample 4, S08°39.335', E114°21.772', 25 m, 10-IV-2011; 3 exx, Triangulasi, sample 5, S08°39.426', E114°21.890', 25 m, 10-IV-2011; 1 ex, Triangulasi, sample 7, S08°39.373', E114°21.842', 25 m, 10-IV-2011; 1 ex, Triangulasi, sample 8, S08°39.559', E114°21.823', 25 m, 10-IV-2011.
E-Java Prov. (Alas Purwo N.P.). Elevation: 25 m.
This epithet is based on the type locality, Alas Purwo National Park.
Trigonopterus alaspurwensis Riedel, sp. n. was coded as “Trigonopterus sp. 345” by
Holotype, male (Fig.
Holotype (MZB): ARC2680 (EMBL # LM655938), West Java Prov., Sumedang, Sukajadi, Mt. Cakrabuana, sample 6, S07°01.943', E108°08.029', 1593 m, 20-IV-2012. Paratypes (MZB, SMNK, ZSM): W-Java Prov.: 2 exx, ARC3583 (EMBL # LM656016), ARC3584 (EMBL # LM656017), Ciwidey, Mt. Tikukur, sample 2, S07°07.816', E107°24.298', 1784 m, 01-XII-2011; 4 exx, ARC2512 (EMBL # LM655849), ARC2513 (EMBL # LM655850), Ciwidey, Mt. Tikukur, sample 3, S07°07.723', E107°23.898', 1929 m, 01-XII-2011; 2 exx, ARC2516 (EMBL # LM655853), Bandung, Lembang, Pangli, Mt. Bukittinggul, sample 1, S06°48.810', E107°44.216', 1753 m, 03-XII-2011; 2 exx, ARC2517 (EMBL # LM655854), ARC3585 (EMBL # LM656018), Bandung, Lembang, Pangli, Mt. Bukittinggul, sample 2, S06°48.845', E107°44.074', 1898 m, 03-XII-2011; 3 exx, ARC2724 (EMBL # LM655980), Garut, Wanaraja, Talagabodas, sample 1, S07°12.511', E108°03.554', 1741 m, 27-IV-2012; 7 exx, ARC2717 (EMBL # LM655973), ARC2718 (EMBL # LM655974), ARC2719 (EMBL # LM655975), ARC3595 (EMBL # LM656026), ARC3596 (EMBL # LM656027), ARC3597, Garut, Wanaraja, Talagabodas, sample 2, S07°11.966', E108°03.984', 1719 m, 27-IV-2012; 8 exx, ARC2715 (EMBL # LM655971), ARC2716 (EMBL # LM655972), Garut, Cikajang, Mt. Payung, sample 3, S07°25.320', E107°48.259', 1560 m, 26-IV-2012; 3 exx, ARC0186 (EMBL # LM655411), ARC0158, Garut, Cilawu, Mt. Cikuray, sample 2, S07°18.840', E107°52.512', 1800 m, 24-IX-2005; 4 exx, Garut, Cilawu, Mt. Cikuray, sample 4, S07°18.927', E107°52.460', 1930 m, 24-IX-2005; 2 exx, Garut, Cilawu, Mt. Cikuray, sample 3, S07°19.075', E107°52.338', 2050 m, 24-IX-2005; 13 exx, ARC2679 (EMBL # LM655937), same data as holotype; 4 exx, Mt. Sawal, Panjalu, Tembong, sample 3, S07°11.102', E108°16.310', 1569 m, 29-XI-2011; 9 exx, ARC2509 (EMBL # LM655846), ARC2510 (EMBL # LM655847), ARC2511 (EMBL # LM655848), Mt. Sawal, Panjalu, Tembong, sample 4, S07°11.275', E108°16.280', 1723 m, 29-XI-2011.
W-Java Prov. (Mt. Bukittinggul, Mt. Cakrabuana, Mt. Cikuray, Mt. Payung, Mt. Sawal, Talagabodas, Mt. Tikukur). Elevation: 1560–2050 m.
This epithet is based on the Greek allos (other) and the Latin patria (homeland) and refers to its fragmented distribution.
Trigonopterus allopatricus Riedel, sp. n. was coded as “Trigonopterus sp. 305”.
Holotype, male (Fig.
Holotype (MZB): ARC1513 (EMBL # LM655611), West Nusa Tenggara Prov., Sumbawa, Batu Dulang, Mt. Batu Pasak, sample 2, S08°37.028', E117°15.783', 1305 m, 12-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang). Elevation: 1305 m.
This epithet is the latinized form of a combination of the Greek adjective allos (foreign) and the noun topos (place). The species marks the Southwestern limit of the T. politus-group´s distribution which is most diverse in the Papuan region.
Trigonopterus allotopus Riedel, sp. n. was coded as “Trigonopterus sp. 331” by
Holotype, male (Fig.
Holotype (MZB): ARC2654 (EMBL # LM655912), West Java Prov., Sukabumi, Mt. Gede, Cisaat, Situ Gunung, sample 1, S06°49.377', E106°55.729', 1281 m, 15-IV-2012. Paratypes (MZB, SMNK): W-Java Prov., Sukabumi, Mt. Gede, Cisaat, Situ Gunung: 3 exx ARC2655 (EMBL # LM655913), same data as holotype; 2 exx, ARC2658 (EMBL # LM655916), ARC2659 (EMBL # LM655917), sample 2, S06°49.038', E106°55.852', 1342 m, 15-IV-2012.
W-Java Prov. (Mt. Gede). Elevation: 1281–1342 m.
This epithet is a combination of the Latin adjective angularis (angular) and the noun collum (neck, pronotum).
Trigonopterus angulicollis Riedel, sp. n. was coded as “Trigonopterus sp. 314” by
Trigonopterus amphoralis
Selected male (Fig.
Syntypes: “Pedada Bay, Lampongs, 2 males, 22.I.1922 (Dr. K.W.Dammerman)”
(MZB, SMNK, ZSM): East Sumatra, Lampung Prov.: 2 exx, ARC0261 (EMBL # LM655452), ARC0262 (EMBL # LM655453), Bawang, Pedada Bay, Mt. Tanggang, sample 2, S05°43.947', E105°06.480', 659 m, 09-VIII-2006; 1 ex, ARC0273 (EMBL # LM655464), Bawang, Pedada Bay, Mt. Tanggang, sample 4, S05°43.871', E105°06.393', 744 m, 09-VIII-2006; 2 exx, ARC0259 (EMBL # LM655451), ARC0260, Bawang, Pedada Bay, Mt. Tanggang, sample 1, S05°43.933', E105°06.598', 579 m, 09-VIII-2006; 1 ex, Bawang, Pedada Bay, Mt. Tanggang, sample 3, S05°43.938', E105°06.440', 673 m, 09-VIII-2006; 1 ex, Bawang, Pedada Bay, Mt. Tanggang, sample 4, S05°43.871', E105°06.393', 744 m, 09-VIII-2006; 6 exx, ARC2725 (EMBL # LM655981), ARC2726 (EMBL # LM655982), Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 2, S05°30.330', E104°25.838', 660 m, 01-V-2012; 4 exx, ARC2729 (EMBL # LM655985), Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 5, S05°31.044', E104°25.664', 562 m, 02-V-2012; 4 exx, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 6, S05°31.678', E104°25.508', 608 m, 02-V-2012; 4 exx, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 7, S05°33.768', E104°25.361', 371 m, 02-V-2012; 3 exx, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 9, S05°32.611', E104°26.353', 455 m, 02-V-2012; 1 ex, Bukit Barisan Selatan N.P., Liwa, sample 6, S05°04.753', E104°03.183', 809 m, 05-V-2012; 2 exx, ARC2735 (EMBL # LM655991), ARC2736 (EMBL # LM655992), Bukit Barisan Selatan N.P., Liwa, sample 7, S05°04.958', E104°03.379', 813 m, 05-V-2012.
Lampung Prov. (Bukit Barisan Selatan, Pedada Bay). Elevation: 371–813 m.
Trigonopterus amphoralis Marshall was coded as “Trigonopterus sp. 319”. The original description was based on two syntypes collected by Dammerman who was curator at the Zoological Museum Bogor. A search at MZB was without success, but a specimen bearing the syntypes´ original labels was located at BMNH. However, it is NOT the syntype: the characters (ovate body with smooth elytra) are in stark contrast to the original description and it belongs to a group never recorded west of Wallace Line, but very common to the east of it. Probably, several Trigonopterus-specimens in the same box became detached from their cards, and they were glued to the wrong cards in an attempt to restore order. The other specimens in the drawer were checked, but unfortunately none was found resembling the description of T. amphoralis. Specimens collected at the type locality match the original description perfectly and thus the situation appears resolved sufficiently.
Holotype, male (Fig.
Holotype (MZB): ARC2457 (EMBL # LM655795), East Java Prov., Krucil, Bremi, Mt. Argopuro (trail to Taman Hidup), sample 2, S07°58.478', E113°31.069', 1457 m, 10-XI-2011. Paratypes (MZB, SMNK): E-Java Prov., Krucil, Bremi, Mt. Argopuro (trail to Taman Hidup): 5 exx, ARC2458 (EMBL # LM655796), ARC2459 (EMBL # LM655797), same data as holotype; 3 exx, ARC2464 (EMBL # LM655802), sample 3, S07°58.586', E113°31.345', 1628 m, 10-XI-2011, sample 4, S07°58.680', E113°31.519', 1785 m, 10-XI-2011; 1 ex, ARC2465 (EMBL # LM655803), sample 7, S07°58.449', E113°31.002', 1457 m, 11-XI-2011.
E-Java Prov. (Mt. Argopuro). Elevation: 1457–1785 m.
This epithet is based on the type locality, Mt. Argopuro.
Trigonopterus argopurensis Riedel, sp. n. was coded as “Trigonopterus sp. 360” by
Holotype, male (Fig.
Holotype (MZB): ARC2474 (EMBL # LM655811), East Java Prov., Mt. Arjuno, road Batu - Pacet, Cangar, sample 5, S07°43.400', E112°31.768', 1419 m, 15-XI-2011. Paratypes (MZB, SMNK, ZSM): E-Java Prov.: 4 exx, ARC2473 (EMBL # LM655810), ARC2475 (EMBL # LM655812), same data as holotype; 1 ex, ARC2476 (EMBL # LM655813), Mt. Arjuno, road Batu - Pacet, Cangar, sample 4, S07°43.247', E112°31.641', 1388 m, 15-XI-2011; 4 exx, ARC2477 (EMBL # LM655814), ARC2478 (EMBL # LM655815), ARC2479 (EMBL # LM655816), Kediri, Mt. Wilis, Besuki, sample 1, S07°51.785', E111°50.188', 1379 m, 21-XI-2011; 4 exx, Kediri, Besuki, Mt. Wilis, sample 6, S07°51.709', E111°50.100', 1409 m, 22-XI-2011; 9 exx, ARC2485 (EMBL # LM655822), ARC2486 (EMBL # LM655823), ARC2487 (EMBL # LM655824), ARC2488 (EMBL # LM655825), Kediri, Mt. Wilis, Besuki, sample 7, S07°51.638', E111°50.010', 1432 m, 22-XI-2011.
E-Java Prov. (Mt. Arjuno, Mt. Wilis). Elevation: 1388–1432 m.
This epithet is based on the type locality Mt. Arjuno.
Trigonopterus arjunensis Riedel, sp. n. was coded as “Trigonopterus sp. 356”.
Holotype, male (Fig.
Holotype (MZB): ARC2520 (EMBL # LM655857), West Java Prov., Bandung, Lembang, Pangli, Mt. Bukittinggul, sample 3, S06°48.807', E107°43.897', 1983 m, 03-XII-2011. Paratypes (MZB, SMNK, ZSM): Bandung, Lembang, Pangli, Mt. Bukittinggul: 10 exx, ARC2521 (EMBL # LM655858), ARC3639 (EMBL # LM656061), same data as holotype; 4 exx, sample 2, S06°48.845', E107°44.074', 1898 m, 03-XII-2011; 10 exx, ARC2522 (EMBL # LM655859), sample 4, S06°48.660', E107°43.722', 2185 m, 03-XII-2011.
W-Java Prov. (Mt. Bukittinggul). Elevation: 1983–2185 m.
This epithet is based on the Latin adjective asper (rough, coarse) and refers to its body sculpture.
Trigonopterus asper Riedel, sp. n. was coded as “Trigonopterus sp. 359”. It is closely related to T. variolosus Riedel, sp. n. from which it can be separated by a larger penis with a longer flagellum. The minimal p-distance between both species is 9.5%.
Holotype, male (Fig.
Holotype (MZB): ARC2543 (EMBL # LM655880), W-Kalimantan Prov., Bengkayan, Suka-Bangun, Mt. Bawang, sample 5, N00°54.103', E109°22.515', 652 m, 11-XII-2011 (MZB). Paratypes (SMNK): 1 ex, ARC2542 (EMBL # LM655879), same data as holotype.
W-Kalimantan Prov. (Mt. Bawang). Elevation: 652 m.
This species is named in honour of Sir David F. Attenborough in recognition of his outstanding documentaries on natural history.
Trigonopterus attenboroughi Riedel, sp. n. was coded as “Trigonopterus sp. 366” by
Holotype, male (Fig.
Holotype (MZB): ARC0580 (EMBL # LM655516), Bali, Bedugul, Tamblingan, above the village along water pipe, sample 4, S08°15.854', E115°05.312', 1310 m, 06-XI-2007. Paratypes (ARC, MZB, SMNK, ZSM): Bali: 7 exx, ARC166, Bedugul, mountain NW of botanical garden, 1400 m, 04-XII-2004 (ARC); 1 ex, ARC0575 (EMBL # LM655511), Bedugul, Mt. Pohen, S08°16.526', E115°08.634', 1580 m, 01-XI-2007; 10 exx, ARC0576 (EMBL # LM655512), ARC0577 (EMBL # LM655513), ARC0578 (EMBL # LM655514), ARC0579 (EMBL # LM655515), Bedugul, Mt. Pohen, S08°16.643', E115°08.875', 1405 m, 01-XI-2007; 6 exx, Bedugul, Mt. Pohen, sample 6, S08°16.607', E115°08.773', 1460 m, 01-XI-07; 15 exx, ARC0581 (EMBL # LM655517), ARC0582 (EMBL # LM655518), ARC0583 (EMBL # LM655519), same data as holotype; 43 exx, Tamblingan, sample 5, S08°15.108', E115°05.573', 1350 m, 03-IV-2011; 2 exx, Tamblingan, sample 1, S08°16.029', E115°05.854', 1210 m, 06-XI-2007; 2 exx, Tamblingan, sample 2, S08°15.015', E115°06.202', 1255 m, 06-XI-2007; 13 exx, Lake Tamblingan, 1250 m, 18-19-XII-2008; 1 ex, ARC0588 (EMBL # LM655524), Bedugul, Mt. Catur, S08°15.626', E115°11.354', 1690 m, 07-XI-2007; 26 exx, ARC2307 (EMBL # LM655760), ARC2308 (EMBL # LM655761), ARC2309 (EMBL # LM655762), Mt. Batukaru, Wangayagede, sample 3, S08°21.431', E115°05.699', 1170 m, 31-III-2011; 49 exx, Mt. Batukaru, Wangayagede, sample 4, S08°21.373', E115°05.564', 1310 m, 31-III-2011; 53 exx, Mt. Batukaru, Wangayagede, sample 5, S08°21.130', E115°05.321', 1520 m, 31-III-2011; 8 exx, Angseri, Mt. Adeng, sample 1, S08°19.824', E115°08.604', 1190 m, 05-IV-2011; 8 exx, Angseri, Mt. Adeng, sample 2, S08°19.726', E115°08.583', 1280 m, 06-IV-2011; 39 exx, ARC2319 (EMBL # LM655772), ARC2320 (EMBL # LM655773), Angseri, Mt. Adeng, sample 3, S08°19.626', E115°08.554', 1385 m, 06-IV-2011; 1 ex, Angseri, Mt. Adeng, sample 4, S08°19.463', E115°08.537', 1535 m, 06-IV-2011; 4 exx, ARC0730 (EMBL # LM655547), ARC0731 (PCR failed), Pangkung village, trail to Mesehe-waterfall, S08°18.30', E114°41.68', 350-600 m, 14 &17-XII-2008; 3 exx, ARC2252 (EMBL # LM655707), ARC2253 (EMBL # LM655708), ARC2254 (PCR failed), Negara, Kampung Pasatan, Desa Pohsanten, Mt. Mesehe, sample 5, S08°16.589', E114°41.772', 745 m, 18-III-2011; 1 ex, Negara, Kampung Pasatan, Desa Pohsanten, Mt. Mesehe, sample 4, S08°17.466', E114°41.331', 490 m, 17-III-2011; 17 exx, Negara, Kampung Pasatan, Desa Pohsanten, Mt. Mesehe, sample 5, S08°16.589', E114°41.772', 745 m, 18-III-2011; 4 exx, Negara, Kampung Pasatan, Desa Pohsanten, Mt. Mesehe, sample 6, S08°16.234', E114°41.528', 935 m, 18-III-2011; 1 ex, Negara, Kampung Pasatan, Desa Pohsanten, Mt. Mesehe, sample 7, S08°16.314', E114°41.649', 890 m, 18-III-2011; 52 exx, ARC2959 (EMBL # LM656012), ARC2960 (EMBL # LM656013), ARC2961 (EMBL # LM656014), Negara, Kampung Pasatan, Desa Pohsanten, Mt. Mesehe, sample 8, S08°16.454', E114°41.725', 825 m, 18-III-2011.
Bali (Mt. Andeng, Mt. Batukaru, Bedugul, Mt. Mesehe). Elevation: 600–1690 m.
This epithet is based on the island of Bali.
Trigonopterus baliensis Riedel, sp. n. was coded as “Trigonopterus sp. 328” by
Holotype, male (Fig.
Holotype (MZB): ARC2303 (EMBL # LM655757), Bali, Mt. Batukaru, Wangayagede, sample 3, S08°21.431', E115°05.699', 1170 m, 31-III-2011. Paratypes (MZB, SMNK): Bali, Mt. Batukaru, Wangayagede: 3 exx, ARC2304 (EMBL # LM655758), ARC2305 (PCR failed), same data as holotype; 1 ex, ARC2302 (EMBL # LM655756), sample 1, S08°22.295', E115°06.048', 835 m, 31-III-2011; 1 ex, Mt. Batukaru, Wangayagede, sample 4, S08°21.373', E115°05.564', 1310 m, 31-III-2011; 3 exx, ARC2951 (EMBL # LM656004), ARC2952 (EMBL # LM656005), ARC2953 (EMBL # LM656006), sample 4, S08°21.373', E115°05.564', 31-III-2011.
Bali (Mt. Batukaru). Elevation: 835–1310 m.
This epithet is based on the type locality Mt. Batukaru.
Trigonopterus batukarensis Riedel, sp. n. was coded as “Trigonopterus sp. 340” by
Holotype, male (Fig.
Holotype (MZB): ARC2535 (EMBL # LM655872), W-Kalimantan Prov., Bengkayan, Suka-Bangun, Mt. Bawang, sample 1, N00°53.429', E109°22.230', 246 m, 10-XII-2011. Paratypes (MZB, SMNK, ZSM): W-Kalimantan Prov., Bengkayan, Suka-Bangun, Mt. Bawang: 3 exx, ARC2536 (EMBL # LM655873), same data as holotype; 3 exx, ARC2534 (EMBL # LM655871), sample 2, N00°53.514', E109°22.301', 275 m, 10-XII-2011; 3 exx, ARC2537 (EMBL # LM655874), sample 3, N00°53.621', E109°22.475', 400 m, 10-XII-2011; 3 exx, sample 6, N00°53.992', E109°22.502', 556 m, 11-XII-2011; 1 ex, ARC2546 (EMBL # LM655883), sample 8, N00°53.736', E109°22.316', 411 m, 11-XII-2011.
W-Kalimantan Prov. (Mt. Bawang). Elevation: 246–556 m.
This epithet is based on the type locality.
Trigonopterus bawangensis Riedel, sp. n. was coded as “Trigonopterus sp. 364”.
Holotype, male (Fig.
Holotype (MZB): ARC0202 (EMBL # LM655424), West Java Prov., Ciamis, Mt. Sawal, Batu Cakra, sample 1, S07°14.920', E108°15.762', 990 m, 01-X-2005 (MZB). Paratypes (MZB, SMNK, ZSM): W-Java Prov., Ciamis: 13 exx, ARC0160, ARC0171, ARC0201 (EMBL # LM655423), ARC0203 (EMBL # LM655425), ARC0218 (EMBL # LM655440), same data as holotype; 2 exx, ARC2688 (EMBL # LM655946), ARC2689 (EMBL # LM655947), Pangandaran, sample 3, S07°43.070', E108°39.634', 156 m, 22-IV-2012.
W-Java Prov. (Pangandaran, Mt. Sawal). Elevation: 156–990 m.
This epithet is a combination of the Latin prefix bi- (two) and the noun nodulus (small swelling) and refers to the structure of the elytral apex.
Trigonopterus binodulus Riedel, sp. n. was coded as “Trigonopterus sp. 315”.
Holotype, male (Fig.
Holotype (MZB): ARC0710 (EMBL # LM655535), E-Kalimantan Prov., Berau Dist., Hutan Wisata Sei Tangap, ca. 8 km W of Tanjungredeb, N02°08.07', E117°24.65', 30 m, 02-X-2008. Paratypes (MZB, SMNK): E-Kalimantan Prov., Berau Dist.: 9 exx, ARC0711 (EMBL # LM655536), ARC0712 (EMBL # LM655537), ARC0713 (EMBL # LM655538), same data as holotype; 1 ex, Hutan Mayang Mangurai, ca. 15 km SW of Tanjungredeb, N02°06.217', E117°24.003‘, 20m, 30-XI-2008.
E-Kalimantan Prov. (Tanjungredeb). Elevation: 20–30 m.
This epithet is based on the island of Borneo.
Trigonopterus bornensis Riedel, sp. n. was coded as “Trigonopterus sp. 311”.
Holotype, male (Fig.
Holotype (MZB): ARC2678 (EMBL # LM655936), West Java, Prov., Sumedang, Sukajadi, Mt. Cakrabuana, sample 6, S07°01.943', E108°08.029', 1593 m, 20-IV-2012. Paratypes (MZB, SMNK, ZSM): W-Java Prov.: 9 exx, Sumedang, Sukajadi, Mt. Cakrabuana, sample 4, S07°01.734', E108°08.059', 1530 m, 19-IV-2012; 8 exx, same data as holotype; 59 exx, ARC0155, ARC0172, ARC0193 (EMBL # LM655417), ARC0194 (EMBL # LM655418), ARC0195 (EMBL # LM655419), Ciamis, Mt. Sawal, Batu Cakra, sample 1, S07°14.920', E108°15.762', 990 m, 01-X-2005 (MZB); 13 exx, sample 3, S07°14.900', E108°15.893', 940 m, 01-X-2005; 11 exx, Ciamis, Mt. Sawal, Blok Cireong, sample 2, S07°14.127', E108°15.568', 1120 m, 01-X-2005; 3 exx, ARC2504 (EMBL # LM655841), Ciamis, Mt. Sawal, Panjalu, Tembong, sample 1, S07°10.528', E108°16.423', 1314 m, 29-XI-2011; 8 exx, ARC2508 (EMBL # LM655845), Ciamis, Mt. Sawal, Panjalu, Tembong, sample 3, S07°11.102', E108°16.310', 1569 m, 29-XI-2011.
W-Java Prov. (Mt. Cakrabuana, Mt. Sawal). Elevation: 990–1593 m.
The species is named for arachnologist Cahyo Rahmadi, who collected the first specimens of this species. The epithet is a noun in the genitive case.
Trigonopterus cahyoi Riedel, sp. n. was coded as “Trigonopterus sp. 320” by
Holotype, male (Fig.
Holotype (MZB): ARC2470 (EMBL # LM655807), E-Java, Mt. Semeru, road Senduro - Ranupani, sample 4, S08°01.801', E112°59.931', 1430 m, 13-XI-2011. Paratypes (MZB, SMNK, ZSM): E-Java Prov.: 2 exx, ARC2469 (EMBL # LM655806), ARC2472 (EMBL # LM655809), same data as holotype; 2 exx, ARC2471 (EMBL # LM655808), Mt. Semeru, road Senduro - Ranupani, sample 3, S08°02.052', E112°59.356', 1572 m, 13-XI-2011; 6 exx, Mt. Semeru, road Senduro - Ranupani, sample 4, S08°01.801', E112°59.931', 1430 m, 13-XI-2011; 1 ex, Mt. Semeru, road Senduro - Ranupani, sample 5, S08°01.952', E113°00.379', 1345 m, 13-XI-2011; 18 exx, ARC2480 (EMBL # LM655817), ARC2481 (EMBL # LM655818), Kediri, Besuki, Mt. Wilis, sample 1, S07°51.785', E111°50.188', 1379 m, 21-XI-2011; 12 exx, Kediri, Besuki, Mt. Wilis, sample 4, S07°51.852', E111°50.185', 1388 m, 22-XI-2011; 4 exx, ARC2489 (EMBL # LM655826), ARC2490 (EMBL # LM655827), Kediri, Besuki, Mt. Wilis, sample 5, S07°51.650', E111°50.174', 1485 m, 22-XI-2011; 10 exx, Kediri, Besuki, Mt. Wilis, sample 6, S07°51.709', E111°50.100', 1409 m, 22-XI-2011; 15 exx, Kediri, Besuki, Mt. Wilis, sample 7, S07°51.638', E111°50.010', 1432 m, 22-XI-2011.
E-Java Prov. (Mt. Semeru, Mt. Wilis). Elevation: 1345–1572 m.
This epithet is a combination of the Latin nouns costa (rib, ridge) and penna (elytron) refers to its elytral sculpture.
Trigonopterus costipennis Riedel, sp. n. was coded as “Trigonopterus sp. 358”.
Holotype, male (Fig.
Holotype (MZB): ARC2235 (EMBL # LM655691), East Nusa Tenggara Prov., Flores, Labuhan Bajo, Tebedo, sample 1, S08°29.848', E119°59.633', 525 m, lowland forest, 14-III-2011. Paratypes (MZB, SMNK): Flores, Labuhan Bajo: 3 exx, ARC2233 (EMBL # LM655689), ARC2234 (EMBL # LM655690), same data as holotype; 1 ex, Roe, sample 4, S08°35.442', E120°00.366', 785 m, thick litter under Ficus-tree, 13-III-2011; 1 ex, ARC2215 (EMBL # LM655671), Roe, earthworm-dominated habitat, sample 1, S08°36.540', E120°01.871', 955 m, 13-III-2011.
East Nusa Tenggara Prov., Flores (Labuhan Bajo). Elevation: 525–955 m.
This epithet is based on the Latin adjective cuprescens (coppery).
Trigonopterus cuprescens Riedel, sp. n. was coded as “Trigonopterus sp. 347”.
Holotype, male (Fig.
Holotype (MZB): ARC1506 (EMBL # LM655604), West Nusa Tenggara Prov., Sumbawa, Batu Dulang, Mt. Batu Pasak, sample 2, S08°37.028', E117°15.783', 1305 m, 12-IV-2010. Paratypes (MZB, SMNK, ZSM): West Nusa Tenggara Prov., Sumbawa: 6 exx, ARC1505 (EMBL # LM655603), ARC1507 (EMBL # LM655605), ARC1508 (EMBL # LM655606), same data as holotype; 6 exx, Batu Dulang, Mt. Batu Pasak, sample 5, S08°37.005', E117°15.790', 1350 m, 18-IV-2010; 1 ex, ARC3589 (PCR failed), Tepal, Pc. Nengas, sample 7, S08°35.176', E117°08.295', 1490 m, 16-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang, Tepal). Elevation: 1305–1350 m.
This epithet is based on the Latin adjective cupreus (copper-colored).
Trigonopterus cupreus Riedel, sp. n. was coded as “Trigonopterus sp. 332”.
Holotype, male (Fig.
Holotype (MZB): ARC1527 (EMBL # LM655625), West Nusa Tenggara Prov., Sumbawa, Batu Dulang, Mt. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 18-IV-2010. Paratypes (MZB, SMNK, ZSM): 2 exx, ARC1525 (EMBL # LM655623), ARC1526 (EMBL # LM655624), same data as holotype; 1 ex, Batu Dulang, Mt. Batu Pasak, sample 5, S08°37.005', E117°15.790', 1350 m, 18-IV-2010; 1 ex, ARC1532 (EMBL # LM655630), Sumbawa, Tepal, Pc. Nengas, sample 7, S08°35.176', E117°08.295', 1490 m, 16-IV-2010; 13 exx, ARC2200 (EMBL # LM655656), ARC2201 (EMBL # LM655657), ARC2202 (EMBL # LM655658), ARC2203 (EMBL # LM655659), Flores, Mt. Ranaka, sample 8, S08°37.781', E120°31.184', 1730 m, 10-III-2011; 1 ex, ARC2194 (EMBL # LM655650), Flores, Lake Ranamese, sample 3, S08°37.024', E120°33.400', 1270 m, 08-III-2011.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang, Tepal), East Nusa Tenggara Prov., Flores (Mt. Ranaka, L. Ranamese). Elevation: 1270–1730 m.
This epithet is based on the name of Dacrycarpus trees under which this weevil was usually sifted – it may however feed on other plants occurring in the same habitat.
Trigonopterus dacrycarpi Riedel, sp. n. was coded as “Trigonopterus sp. 329”. Cox1 sequences of the populations from Sumbawa and Flores differ 7.5–9.7% p-distance, but morphologically no differences could be found.
Holotype, male (Fig.
Holotype (MZB): ARC2242 (EMBL # LM655697), East Nusa Tenggara Prov., Flores Isl., Labuhan Bajo, Tebedo, sample 1, S08°29.848', E119°59.633', 525 m, 14-III-2011. Paratypes (MZB, SMNK): 7 exx, ARC2591 (EMBL # LM655900), ARC2592 (EMBL # LM655901), ARC3636 (EMBL # LM656058), same data as holotype.
East Nusa Tenggara Prov., Flores (Labuhan Bajo). Elevation: 525 m.
This epithet is based on the Indonesian word for “eight” and is treated as a noun in apposition.
Trigonopterus delapan Riedel, sp. n. was coded as “Trigonopterus sp. 298” by
Holotype, male (Fig.
Holotype (MZB): ARC1468 (EMBL # LM655566), West Nusa Tenggara Prov., Lombok, Tetebatu, Rinjani-trail from Orong Gerisak, sample 7, S08°29.433', E116°24.746', 1240 m, 04-IV-2010 (MZB). Paratypes (ARC, MZB, SMNK, ZSM): Lombok: 4 exx, ARC0163, N Tetebatu, 1200-1450 m, 07-XII-2004; 1 ex, ARC1461 (EMBL # LM655559), Tetebatu, Rinjani-trail from Orong Gerisak, sample 6, S08°29.577', E116°24.782', 1195 m, 04-IV-2010; 2 exx, ARC1474 (EMBL # LM655572), ARC1475 (EMBL # LM655573), Tetebatu, Rinjani-trail from Orong Gerisak, sample 5, S08°30.096', E116°25.062', 1010 m, 04-IV-2010; 1 ex, ARC2270 (EMBL # LM655724), Senaru, Rinjani-track, sample 6, S08°19.359', E116°24.070', 860 m, 23-III-2011; 4 exx, ARC2271 (EMBL # LM655725), ARC2272 (EMBL # LM655726), ARC2273 (EMBL # LM655727), Santong, Rinjani-track, sample 1, S08°20.715', E116°19.695', 830 m, 24-III-2011; 1 ex, Santong, Rinjani-track, sample 2, S08°20.813', E116°19.778', 870 m, 24-III-2011; 1 ex, ARC2283 (EMBL # LM655737), Santong, Rinjani-track, sample 3, S08°20.996', E116°20.001', 960 m, 25-III-2011; 2 exx, ARC2284 (EMBL # LM655738), Santong, Rinjani-track, sample 4, S08°21.160', E116°20.067', 1005 m, 25-III-2011.
West Nusa Tenggara Prov., Lombok (Santong, Senaru, Tetebatu). Elevation: 830–1240 m.
This epithet is based on the combination of the Latin nouns dens (tooth) and pes (foot).
Trigonopterus dentipes Riedel, sp. n. was coded as “Trigonopterus sp. 322” by
Holotype, male (Fig.
Holotype (MZB): ARC0290 (EMBL # LM655480), C-Java Prov., N slopes of Dieng figau, Petungkriyono, mountain N Tinalum, sample 1, S07°06.418', E109°44.514', 1115 m, 22-VIII-2006. Paratypes (MZB, SMNK, ZSM): 13 exx, ARC0288 (EMBL # LM655478), ARC0289 (EMBL # LM655479), ARC0291 (EMBL # LM655481), ARC0292 (EMBL # LM655482), same data as holotype.
C-Java Prov. (Dieng). Elevation: 1115 m.
This epithet is based on the type locality.
Trigonopterus diengensis Riedel, sp. n. was coded as “Trigonopterus sp. 335”.
Holotype, male (Fig.
Holotype (MZB): ARC0211 (EMBL # LM655433), West Java Prov., Mt. Halimun N.P., Citalahab, sample 2, S06°44.503', E106°31.550', 1200 m, 12-IX-2005. Paratypes (ARC, MZB, SMNK): W-Java Prov., Mt. Halimun N.P.: 3 exx, ARC0101, W-Java Prov., Mt. Halimun N.P., Citalahab, S06°44.503', E106°31.550', 1150 m, 20-21-IX-2004; 10 exx, ARC0212 (EMBL # LM655434), ARC0213 (EMBL # LM655435), same data as holotype; 1 ex, Citalahab, sample 1, S06°44.423', E106°31.738', 1100 m, 12-IX-2005; 2 exx, ARC0170, N Ciptarasa, sample 3, S06°49.867', E106°30.085', 1100 m, 19-IX-2005; 2 exx, ARC0216 (EMBL # LM655438), ARC0217 (EMBL # LM655439), Panguyangan, Mt. Talaga, ca.1000 m, 17-IX-2005.
W-Java Prov. (Mt. Halimun N.P.). Elevation: 1000–1200 m.
This epithet is a combination of the prefix di- (two, double) and the adjective derived from the Greek morphe (gestalt).
Trigonopterus dimorphus Riedel, sp. n. was coded as “Trigonopterus sp. 293”.
Holotype, male (Fig.
Holotype (MZB): ARC1490 (EMBL # LM655588), West Nusa Tenggara Prov., Lombok, road between Sembalun and Sapit, sample 5, S08°26.453', E116°31.859', 1195 m, 31-III-2010. Paratypes (MZB, SMNK, ZSM): West Nusa Tenggara Prov., Lombok: 1 ex, ARC1491 (EMBL # LM655589), same data as holotype; 1 ex, ARC1486 (EMBL # LM655584), road between Sembalun and Sapit, sample 1, S08°25.110', E116°31.947', 1450 m, 30-III-2010; 2 exx, ARC1488 (EMBL # LM655586), road between Sembalun and Sapit, sample 4, S08°25.331', E116°31.841', 1405 m, 31-III-2010; 1 ex, ARC1460 (EMBL # LM655558), Tetebatu, Rinjani-trail from Orong Gerisak, sample 6, S08°29.577', E116°24.782', 1195 m, 04-IV-2010; 5 exx, ARC0164, Tetebatu, Rinjani-trail, 1200-1450 m, 07-XII-2004; 2 exx, ARC1466 (EMBL # LM655564), ARC1467 (EMBL # LM655565), sample 7, S08°29.433', E116°24.746', 1240 m, 04-IV-2010; 1 ex, ARC1485 (EMBL # LM655583), Sesaot, Rinjani-trail, sample 3, S08°29.841', E116°13.679', 625 m, 28-III-2010; 1 ex, ARC2265 (EMBL # LM655719), Senaru, Rinjani-trail, sample 4, S08°20.439', E116°24.047', 1240 m, 21-III-2011; 1 ex, ARC2266 (EMBL # LM655720), Senaru, Rinjani-trail, sample 5, S08°19.800', E116°24.107', 1015 m, 21-III-2011; 2 exx, Lombok, Senaru, Rinjani-track, sample 6, S08°19.359', E116°24.070', 860 m, 23-III-2011; 5 exx, ARC2269 (EMBL # LM655723), Senaru, Rinjani-trail, sample 7, S08°19.643', E116°24.033', 935 m, 23-III-2011; 7 exx, Senaru, Rinjani-trail, sample 8, S08°19.719', E116°24.040', 955 m, 23-III-2011; 7 exx, ARC2275 (EMBL # LM655729), ARC2276 (EMBL # LM655730), Santong, Rinjani-trail, sample 1, S08°20.715', E116°19.695', 830 m, 24-III-2011; 2 exx, Santong, Rinjani-track, sample 2, S08°20.813', E116°19.778', 870 m, 24-III-2011; 2 exx, Santong, Rinjani-track, sample 3, S08°20.996', E116°20.001', 960 m, 24-III-2011; 2 exx, Santong, Rinjani-track, sample 4, S08°21.160', E116°20.067', 1005 m, 25-III-2011; 1 ex, Santong, Rinjani-track, sample 6, S08°21.754', E116°20.476', 1315 m, 25-III-2011; 2 exx, ARC2286 (EMBL # LM655740), ARC2287 (EMBL # LM655741), Sajang, sample 1, S08°19.370', E116°29.570', 845 m, 26-III-2011; 4 exx, Mt. Rinjani, Bawnau, near Sembalun, sample 1, S08°21.136', E116°29.257', 1140 m, 26-III-2011; 3 exx, Mt. Rinjani, Bawnau, near Sembalun, S08°21.136', E116°29.257', 1140 m, collected from foliage, 26-III-2011; 2 exx, ARC2291 (EMBL # LM655745), ARC2292 (EMBL # LM655746), Sajang, sample 1, S08°21.136', E116°29.257', 1140 m, 26-III-2011; 4 exx, Mt. Pengasingan, SE Sembalun, sample 2, S08°19.354', E116°30.530', 945 m, 27-III-2011; 1 ex, Mt. Pengasingan, SE Sembalun, sample 3, S08°19.445', E116°30.603', 1005 m, 27-III-2011; 4 exx, ARC2299 (EMBL # LM655753), ARC2300 (EMBL # LM655754), Gn. Pengasingan, SE Sembalun, sample 4, S08°19.504', E116°30.767', 1035 m, 27-III-2011.
West Nusa Tenggara Prov., Lombok (Mt. Pengasingan, Sajang, Santong, Sembalun, Senaru, Sesaot, Tetebatu). Elevation: 625–1450 m.
This epithet is based on the Latin participle disruptus (broken apart) and refers to its phylogeography which appears to reflect the division of the species in two populations.
Trigonopterus disruptus Riedel, sp. n. was coded as “Trigonopterus sp. 292” by
Holotype, male (Fig.
Holotype (MZB): ARC2232 (EMBL # LM655688), East Nusa Tenggara Prov., Flores Isl., Labuhan Bajo, Roe, sample 3, S08°36.259', E120°01.539', 975 m, 13-III-2011. Paratypes (MZB, SMNK, ZSM): Flores Isl., Labuhan Bajo: 1 ex, ARC2217 (EMBL # LM655673), Roe, sample 1, S08°36.540', E120°01.871', 955 m, earthworm-dominated habitat, 13-III-2011; 2 exx, ARC2589 (EMBL # LM655898), ARC3626 (EMBL # LM656056), Roe, sample 3, S08°36.259', E120°01.539', 975 m, 13-III-2011; 1 ex, ARC2226 (EMBL # LM655682), Roe, sample 5, S08°35.395', E120°00.383', 790 m, 13-III-2011; 1 ex, same data as holotype; 3 exx, ARC2244 (EMBL # LM655699), ARC2593 (EMBL # LM655902), ARC2594 (EMBL # LM655903), Tebedo, sample 1, S08°29.848', E119°59.633', 525 m, 14-III-2011.
East Nusa Tenggara Prov., Flores (Labuhan Bajo). Elevation: 525–975 m.
This epithet is based on the Indonesian word for “two” and is treated as a noun in apposition.
Trigonopterus dua Riedel, sp. n. was coded as “Trigonopterus sp. 284”.
Holotype, male (Fig.
Holotype (MZB): ARC0284 (EMBL # LM655474), C-Java Prov., N slopes of Dieng figau, Petungkriyono, Mt. Deles, sample 1, S07°08.221', E109°43.599', 1505 m, 24-VIII-2006. Paratypes (MZB, SMNK): C-Java Prov.: ARC0283 (EMBL # LM655473), same data as holotype; 2 exx, ARC2491 (EMBL # LM655828), ARC2492 (EMBL # LM655829), Mt. Slamet, Guci, sample 2, S07°13.255', E109°11.005', 1858 m, 26-XI-2011; 4 exx, ARC2493 (EMBL # LM655830), ARC2494 (EMBL # LM655831), ARC2495 (EMBL # LM655832), Mt. Slamet, Guci, sample 6, S07°11.983', E109°10.556', 1671 m, 27-XI-2011; 1 ex, Mt. Slamet, Guci, sample 7, S07°11.906', E109°10.486', 1653 m, 27-XI-2011.
C-Java Prov. (Dieng, Mt. Slamet). Elevation: 1505–1858 m.
This epithet is based on the Indonesian word for “twelve” and is treated as a noun in apposition.
Trigonopterus duabelas Riedel, sp. n. was coded as “Trigonopterus sp. 444”. Cox1 sequences of the populations from Dieng and Mt. Slamet differ 4.6–5.4 % p-distance, but no distinct morphological differences could be found.
Holotype, male (Fig.
Holotype (MZB): ARC0204 (EMBL # LM655426), W-Java Prov., Ciamis, Mt. Sawal, Batu Cakra, sample 1, S07°14.920', E108°15.762', 990 m, 01-X-2005. Paratypes (MZB, SMNK): W-Java Prov.: 1 ex, ARC2503 (EMBL # LM655840), Ciamis, Panjalu, Tembong, Mt. Sawal, sample 1, S07°10.528', E108°16.423', 1314 m, sifted; 8 exx, ARC2696 (EMBL # LM655952), ARC2697 (EMBL # LM655953), ARC2698 (EMBL # LM655954), Garut, Cikajang, Mt. Payung, sample 1, S07°25.345', E107°48.825', 1085 m, 26-IV-2012.
W-Java Prov. (Mt. Payung, Mt. Sawal). Elevation: 990–1314 m.
This epithet is based on the Latin adjective echinatus (prickly) and refers to the species´ erect scales that resemble spines.
Trigonopterus echinatus Riedel, sp. n. was coded as “Trigonopterus sp. 337”.
Holotype, male (Fig.
Holotype (MZB): ARC2192 (EMBL # LM655648), East Nusa Tenggara Prov., Flores Isl., Ruteng, Danau Ranamese, sample 1, S08°37.705', E120°33.702', 1320 m, 08-III-2011. Paratypes (SMNK): Flores Isl., Ruteng, Mt. Ranaka: 1 ex, ARC2583 (EMBL # LM655892), sample 9, S08°37.641', E120°31.204', 1685 m, 10-III-2011.
East Nusa Tenggara Prov., Flores (Ruteng). Elevation: 1320–1685 m.
This epithet is based on the Indonesian word for “four” and is treated as a noun in apposition.
Trigonopterus empat Riedel, sp. n. was coded as “Trigonopterus sp. 303” by
Holotype, male (Fig.
Holotype (MZB): ARC2224 (EMBL # LM655680), East Nusa Tenggara Prov., Flores Isl., Labuhan Bajo, Roe, sample 5, S08°35.395', E120°00.383', 790 m, 13-III-2011 (MZB). Paratypes (SMNK, ZSM): Flores Isl., Labuhan Bajo: 2 exx, ARC2587 (EMBL # LM655896), ARC2588 (EMBL # LM655897), Roe, sample 2, S08°36.338', E120°01.637', 965 m, 13-III-2011; 1 ex, ARC3625 (EMBL # LM656055), Roe, sample 3, S08°36.259', E120°01.539', 975 m, 13-III-2011.
East Nusa Tenggara Prov., Flores (Labuhan Bajo). Elevation: 790–965 m.
This epithet is based on the Indonesian word for “six” and is treated as a noun in apposition.
Trigonopterus enam Riedel, sp. n. was coded as “Trigonopterus sp. 295” by
Holotype, male (Fig.
Holotype (MZB): ARC2211 (EMBL # LM655667), East Nusa Tenggara Prov., Flores Isl., Ruteng, Danau Ranamese, sample 6, S08°38.370', E120°33.798', 1215 m, 11-III-2011. Paratypes (SMNK): 1 ex, ARC2212 (EMBL # LM655668), same data as holotype.
East Nusa Tenggara Prov., Flores (Lake Ranamese). Elevation: 1215 m.
This epithet is based on a combination of the Latin adjective fissus (cleft) and the Greek noun tarsos and refers to the shape of the protarsi.
Trigonopterus fissitarsis Riedel, sp. n. was coded as “Trigonopterus sp. 290” by
Holotype, male (Fig.
Holotype (MZB): ARC3618 (EMBL # LM656048), East Nusa Tenggara Prov., Flores, Ruteng, Mt. Ranaka, sample 1, S08°37.321', E120°31.463' 1535 m, 08-III-2011. Paratypes (MZB, SMNK, ZSM): Flores: 3 exx, ARC3616 (EMBL # LM656046), ARC3617 (EMBL # LM656047), ARC3619 (EMBL # LM656049), same data as holotype; 2 exx, ARC2189 (EMBL # LM655646), ARC2190 (EMBL # LM655647), Ruteng, Lake Ranamese, sample 1, S08°37.705', E120°33.702', 1320 m, 08-III-2011; 1 ex, ARC3621 (EMBL # LM656051), Ruteng, Lake Ranamese, sample 4, S08°37.705', E120°33.702', 1250 m, 08-III-2011; 2 exx, ARC2210 (EMBL # LM655666), ARC3620 (EMBL # LM656050), Ruteng, Golo Lusang, sample 1, S08°39.864', E120°27.322', 1590 m, 11-III-2011.
East Nusa Tenggara Prov., Flores (Golo Lusang, Mt. Ranaka, Lake Ranamese). Elevation: 1250–1590 m.
This epithet is based on the island of Flores.
Trigonopterus florensis Riedel, sp. n. was coded as “Trigonopterus sp. 440”. Morphologically it is very similar to T. pseudoflorensis Riedel, sp. n. and T. paraflorensis Riedel, sp. n. but its cox1 sequences differ 6.69%, respectively 7.75% smallest interspecific p-distance.
Holotype, male (Fig.
Holotype (MZB): ARC2690 (EMBL # LM655948), W-Java, Ciamis, Pangandaran, sample 1, S07°42.836', E108°39.478', 157 m, 22-IV-2012. Paratypes (MZB, SMNK, ZSM): W-Java: 4 exx, ARC2684 (EMBL # LM655942), ARC2685 (EMBL # LM655943), same data as holotype; 1 ex, Ciamis, Pangandaran, sample 3, S07°43.070', E108°39.634', 156 m, 22-IV-2012; 4 exx, ARC2693 (EMBL # LM655949), ARC2694 (EMBL # LM655950), ARC2695 (EMBL # LM655951), Garut, Pameungpeuk, Leuweung Sancang, sample 2, S07°43.807', E107°53.700', 74 m, 24-IV-2012.
W-Java Prov. (Leuweung Sancang, Pangandaran). Elevation: 74–157 m.
This epithet is based on the Latin adjective foveatus (pitted) and refers to its sculpture.
Trigonopterus foveatus Riedel, sp. n. was coded as “Trigonopterus sp. 375”.
Holotype, male (Fig.
Holotype (MZB): ARC2274 (EMBL # LM655728), West Nusa Tenggara Prov., Lombok, Santong, Rinjani-track, sample 1, S08°20.715', E116°19.695', 830 m, 24-III-2011. Paratypes (MZB, SMNK, ZSM): 3 exx, same data as holotype; 4 exx, Santong, Rinjani-track, sample 2, S08°20.813', E116°19.778', 870 m, 24-III-2011; 10 exx, ARC2281 (EMBL # LM655735), ARC2282 (EMBL # LM655736), Lombok, Santong, Rinjani-track, sample 3, S08°20.996', E116°20.001', 960 m, 25-III-2011; 1 ex, Santong, Rinjani-track, sample 4, S08°21.160', E116°20.067', 1005 m, 25-III-2011; 1 ex, Santong, Rinjani-track, sample 5, S08°21.351', E116°20.208', 1105 m, 25-III-2011; 1 ex, Santong, Rinjani-track, sample 6, S08°21.754', E116°20.476', 1315 m, 25-III-2011; 9 exx, ARC2256 (EMBL # LM655710), ARC2257 (EMBL # LM655711), ARC2258 (EMBL # LM655712), Lombok, Senaru, Rinjani-track, sample 1, S08°19.429', E116°24.082', 900 m, 21-III-2011; 2 exx, Lombok, Senaru, Rinjani-track, sample 4, S08°20.439', E116°24.047', 1240 m, 21-III-2011; 16 exx, Lombok, Senaru, Rinjani-track, sample 6, S08°19.359', E116°24.070', 860 m, 23-III-2011; 3 exx, Senaru, Rinjani-trail, sample 7, S08°19.643', E116°24.033', 935 m, 23-III-2011; 7 exx, Senaru, Rinjani-trail, sample 8, S08°19.719', E116°24.040', 955 m, 23-III-2011; 12 exx, ARC2288 (EMBL # LM655742), ARC2289 (EMBL # LM655743), ARC2290 (EMBL # LM655744), Lombok, Mt. Rinjani, Bawnau, near Sembalun, sample 1, S08°21.136', E116°29.257', 1140 m, 26-III-2011.
West Nusa Tenggara Prov., Lombok (Santong, Sembalun, Senaru). Elevation: 830–1240 m.
This epithet is based on the Latin adjective fulgidus (shining, gleaming) and refers to its vivid coloration.
Trigonopterus fulgidus Riedel, sp. n. was coded as “Trigonopterus sp. 349” by
Holotype, male (Fig.
Holotype (MZB): ARC0249 (EMBL # LM655441), West Java Prov., Cianjur, above kebun Gede, sample 2, S06°47.735', E107°00.658', 2005 m, sifted. Paratypes (MZB, SMNK, ZSM): W-Java Prov., Mt. Gede: 5 exx, ARC0250 (EMBL # LM655442), ARC0251 (EMBL # LM655443), ARC0252 (EMBL # LM655444), ARC0253 (EMBL # LM655445), same data as holotype; 22 exx, ARC0003, Cibodas, 2000 m, 07-VI-2002; 4 exx, ARC2664 (EMBL # LM655922), Sukabumi, Cisaat, Situ Gunung, sample 5, S06°47.912', E106°56.457', 1940 m, 16-IV-2012; 5 exx, ARC2665 (EMBL # LM655923), Sukabumi, Cisaat, Situ Gunung, sample 6, S06°47.706', E106°56.560', 2117 m, 16-IV-2012.
W-Java Prov., (Mt. Gede). Elevation: 1940–2117 m.
This epithet is based on the type locality.
Trigonopterus gedensis Riedel, sp. n. was coded as “Trigonopterus sp. 307” by
Holotype, male (Fig.
Holotype (MZB): ARC2647 (EMBL # LM655905), West Java Prov., Bogor, Cidahu, Mt. Salak (near Javana Spa), sample 1, S06°43.733', E106°42.711', 1429 m, 13-IV-2012. Paratypes (ARC, MZB, SMNK, ZSM): W-Java Prov.: 1 ex, ARC2646 (EMBL # LM655904), same data as holotype; 1 ex, Bogor, Cidahu, Mt. Salak (near Javana Spa), sample 2, S06°43.499', E106°42.970', 1550 m, 13-IV-2012; 5 exx, ARC2650 (EMBL # LM655908), ARC2651 (EMBL # LM655909), Bogor, Cidahu, Mt. Salak (near Javana Spa), sample 3, S06°43.425', E106°43.227', 1756 m, 13-IV-2012; 4 exx, ARC0103, Mt. Halimun N.P., Pasir Banteng, Mt. Botol, 1550 m, 17-XI-2004; 3 exx, ARC0183 (EMBL # LM655408), Mt. Halimun N.P., Citalahab, Mt. Kendeng, 1550 m, 19-XI-2004; 5 exx, ARC0185 (EMBL # LM655410), Mt. Halimun N.P., Citalahab, Mt. Kendeng, sample 2, S06°45.523', E106°31.435', 1600 m, 10-IX-2005; 1 ex, ARC0188 (PCR failed), Mt. Halimun N.P., Citalahab, Mt. Kendeng, sample 1B, S06°45.723', E106°31.473', 1695 m, 10-IX-2005.
W-Java Prov. (Mt. Halimun Salak N.P.). Elevation: 1429–1756 m.
This epithet is based on the type locality.
Trigonopterus halimunensis Riedel, sp. n. was coded as “Trigonopterus sp. 299” by
Holotype, male (Fig.
Holotype (MZB): ARC1547 (EMBL # LM655645), Java, Banten-Prov., Ujung Kulon N.P., Tama Jaya, Mt. Honje, sample 4, S06°46.141', E105°31.649', 395 m, 23-IV-2010. Paratypes (MZB, SMNK, ZSM): Java, Banten-Prov., Ujung Kulon N.P., Tama Jaya, Mt. Honje: 6 exx, ARC1542 (EMBL # LM655640), ARC1543 (EMBL # LM655641), ARC1544 (EMBL # LM655642), sample 2, S06°45.973', E105°31.915', 540 m, 23-IV-2010; 16 exx, sample 4, S06°46.141', E105°31.649', 395 m, 23-IV-2010; 2 exx, sample 3, S06°46.009', E105°31.761', 465 m, 23-IV-2010.
Banten Prov. (Ujung Kulon N.P.). Elevation: 395–540 m.
This epithet is based on the type locality, Mt. Honje.
Trigonopterus honjensis Riedel, sp. n. was coded as “Trigonopterus sp. 338”.
Holotype, male (Fig.
Holotype (MZB): ARC0305 (EMBL # LM655495), East Java Prov., Banyuwangi, Mt. Ijen, Licin, sample 4, S08°06.673', E114°14.488', 1225 m, 31-VIII-2006. Paratypes (MZB, SMNK, ZSM): E-Java Prov., Banyuwangi, Mt. Ijen, Licin: 11 exx, ARC0304 (EMBL # LM655494), ARC0306 (EMBL # LM655496), ARC0307 (EMBL # LM655497), same data as holotype; 1 ex, ARC0303 (EMBL # LM655493), sample 2, S08°05.603', E114°14.366', 1575 m, 28-VIII-2006; 2 exx, sample 3, S08°06.056', E114°14.621', 1400 m, 28-VIII-2006; 5 exx, Banyuwangi, Mt. Ijen, Licin, sample 4, S08°06.673', E114°14.488', 1255 m, 28-VIII-2006.
E-Java Prov. (Mt. Ijen). Elevation: 1225–1575 m.
This epithet is based on the type locality.
Trigonopterus ijensis Riedel, sp. n. was coded as “Trigonopterus sp. 316”.
Holotype, male (Fig.
Holotype (MZB): ARC0192 (EMBL # LM655416), West Java Prov., Mt. Halimun N.P., N Ciptarasa, sample 3, S06°49.867', E106°30.085', 1100 m, 19-IX-2005. Paratypes (MZB, SMNK, ZSM): 17 exx, ARC0156, ARC0190 (EMBL # LM655414), ARC0191 (EMBL # LM655415), same data as holotype; 4 exx, W-Java Prov., Citalahab, sample 2, S06°44.503', E106°31.550', 1200 m, 12-IX-2005; 6 exx, ARC2656 (EMBL # LM655914), ARC2657 (EMBL # LM655915), W-Java Prov., Sukabumi, Mt. Gede, Cisaat, Situ Gunung, sample 1, S06°49.377', E106°55.729', 1281 m, 15-IV-2012; 3 exx, ARC2660 (EMBL # LM655918), ARC2661 (EMBL # LM655919), W-Java Prov., Sukabumi, Mt. Gede, Cisaat, Situ Gunung, sample 2, S06°49.038', E106°55.852', 1342 m, 15-IV-2012; 4 exx, ARC0196, ARC0197 (EMBL # LM655420), ARC0198, W-Java Prov., Ciamis, Mt. Sawal, Batu Cakra, sample 1, S07°14.920', E108°15.762', 990 m, 01-X-2005; 2 exx, W-Java Prov., Ciamis, Mt. Sawal, Blok Cireong, sample 2, S07°14.127', E108°15.568', 1120 m, 01-X-2005; 1 ex, ARC3593 (EMBL # LM656024), Sumedang, Sukajadi, Mt. Cakrabuana, sample 5, S07°01.804', E108°08.044', 1563 m, 20-IV-2012; 4 exx, ARC0285 (EMBL # LM655475), ARC0286 (EMBL # LM655476), ARC0287 (EMBL # LM655477), C-Java Prov., N slopes of Dieng figau, Petungkriyono, Mt. Deles, sample 1, S07°08.221', E109°43.599', 1505 m, 24-VIII-2006; 4 exx, ARC0297 (EMBL # LM655487), C-Java Prov., N slopes of Dieng figau, Petungkriyono, Mt. Deles, sample 3, S07°08.225', E109°43.555', 1495 m, 24-VIII-2006; 4 exx, ARC0296 (EMBL # LM655486), C-Java Prov., N slopes of Dieng figau, Petungkriyono, mountain N Tinalum, sample 1, S07°06.418', E109°44.514', 1115 m, 22-VIII-2006; 1 ex, C-Java Prov., Mt. Slamet, Guci, sample 5, S07°11.953', E109°10.497', 1620 m, 27-XI-2011; 6 exx, ARC2496 (EMBL # LM655833), ARC2497 (EMBL # LM655834), ARC2498 (EMBL # LM655835), C-Java Prov., Mt. Slamet, Guci, sample 6, S07°11.983', E109°10.556', 1671 m, 27-XI-11.
W-Java Prov. (Mt. Gede, Mt. Halimun N.P., Mt. Cakrabuana, Mt. Sawal), C-Java Prov. (Dieng, Mt. Slamet). Elevation: 990–1671 m.
This epithet is based on the name of Java Island.
Trigonopterus javensis Riedel, sp. n. was coded as “Trigonopterus sp. 294”.
Holotype, male (Fig.
Holotype (MZB): ARC1421 (EMBL # LM655548), E-Kalimantan Prov., Berau Dist., 1 km off road Tanjungredeb - Tanjungselor, ca. 45 km N of Tanjungredeb, N02°29.55', E117°28.77', 190 m, 29-IX/03-X-2008 (MZB). Paratypes (SMNK): 2 exx, ARC1422 (EMBL # LM655549), ARC1423 (EMBL # LM655550), same data as holotype.
E-Kalimantan Prov. (Tanjungredeb). Elevation: 190 m.
This epithet is based on the Indonesian name for Borneo, Kalimantan.
Trigonopterus kalimantanensis Riedel, sp. n. was coded as “Trigonopterus sp. 310” by
Holotype, male (Fig.
Holotype (MZB): ARC0589 (EMBL # LM655525), Bali, Kintamani, Mt. Abang, sample 1, S08°17.118', E115°24.868', 1440 m, 09-XI-2007. Paratypes (SMNK): Bali: 1 ex, ARC0590 (EMBL # LM655526), same data as holotype.
Bali (Kintamani). Elevation: 1440 m.
This epithet is based on the type locality, Kintamani.
Trigonopterus kintamanensis Riedel, sp. n. was coded as “Trigonopterus sp. 286” by
Holotype, male (Fig.
Holotype (MZB): ARC0595 (EMBL # LM655531), Bali, National Park, Mt. Klatakan, sample 1, S08°13.220', E114°29.509', 545 m, 12-XI-2007. Paratypes (MZB, SMNK): Bali National Park, Mt. Klatakan: 3 exx, ARC0596 (EMBL # LM655532), ARC0597 (EMBL # LM655533), ARC0598 (EMBL # LM655534), same data as holotype; 2 exx, ARC2954 (EMBL # LM656007), ARC2955 (EMBL # LM656008), sample 2, S08°13.029', E114°29.466', 600 m, 12-XI-2007; 7 exx, sample 3, S08°13.325', E114°29.477', 500 m, 12-XI-2007.
Bali (Mt. Klatakan). Elevation: 500–600 m.
This epithet is based on the type locality, Mt. Klatakan.
Trigonopterus klatakanensis Riedel, sp. n. was coded as “Trigonopterus sp. 285” by
Holotype, male (Fig.
Holotype (MZB): ARC0266 (EMBL # LM655457), East Sumatra, Lampung Prov., Bawang, Pedada Bay, Mt. Tanggang, sample 2, S05°43.947', E105°06.480', 659 m, 09-VIII-2006. Paratypes (MZB, SMNK, ZSM): E-Sumatra, Lampung Prov.: 1 ex, ARC0267 (EMBL # LM655458), same data as holotype; 1 ex, ARC0274 (EMBL # LM655465), Bawang, Pedada Bay, Mt. Tanggang, sample 3, S05°43.938', E105°06.440', 673 m, 09-VIII-2006; 1 ex, Bawang, Pedada Bay, Mt. Tanggang, sample 4, S05°43.871', E105°06.393', 744 m, 09-VIII-2006; 1 ex, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 5, S05°31.044', E104°25.664', 562 m, 02-V-2012; 1 ex, ARC2731 (EMBL # LM655987), Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 8, S05°33.032', E104°25.985', 421 m, 02-V-2012; 2 exx, ARC2732 (EMBL # LM655988), ARC2733 (EMBL # LM655989), Bukit Barisan Selatan N.P., Liwa, sample 1, S05°04.409', E104°03.265', 670 m, 04-V-2012; 1 ex, Bukit Barisan Selatan N.P., Liwa, sample 6, S05°04.753', E104°03.183', 809 m, 05-V-2012; 1 ex, ARC2734 (EMBL # LM655990), Bukit Barisan Selatan N.P., Liwa, sample 7, S05°04.958', E104°03.379', 813 m, 05-V-2012; 1 ex, Bukit Barisan Selatan N.P., Liwa, sample 8, S05°05.015', E104°03.442', 801 m, 05-V-2012.
Lampung Prov. (Bukit Barisan Selatan N.P., Pedada Bay). Elevation: 421–813 m.
This epithet is based on the Indonesian province of Lampung.
Trigonopterus lampungensis Riedel, sp. n. was coded as “Trigonopterus sp. 301”.
Holotype, male (Fig.
Holotype (MZB): ARC2466 (EMBL # LM655804), East Java Prov., Mt. Semeru, road Senduro - Ranupani, sample 4, S08°01.801', E112°59.931', 1430 m, 13-XI-2011. Paratypes (SMNK, ZSM): E-Java Prov., Mt. Semeru, road Senduro - Ranupani: 2 exx, ARC2467 (EMBL # LM655805), ARC2468, same data as holotype; 1 ex, sample 3, S08°02.052', E112°59.356', 1572 m, 13-XI-2011.
E-Java Prov. (Mt. Semeru). Elevation: 1430 m.
This epithet is based on a combination of the Latin adjective latus (wide) and the noun pes (foot) and refers to the unusual size of the protarsi.
Trigonopterus latipes Riedel, sp. n. was coded as “Trigonopterus sp. 355”.
Holotype, male (Fig.
Holotype (MZB): ARC2199 (EMBL # LM655655), East Nusa Tenggara Prov., Flores Isl., Ruteng, Mt. Ranaka, sample 3, S08°38.099', E120°31.745', summit area, 2205 m, 09-III-2011. Paratypes (MZB, SMNK): Flores Isl., Ruteng, Mt. Ranaka: 2 exx, ARC2579 (EMBL # LM655888), ARC2580 (EMBL # LM655889), sample 5, S08°38.277', E120°31.616', 2010 m, 09-III-2011; 1 ex, ARC2581 (EMBL # LM655890), sample 6, S08°38.243', E120°31.569', 1995 m, 09-III-2011; 2 exx, ARC2204 (EMBL # LM655660), sample 8, S08°37.781', E120°31.184', 1730 m, 10-III-2011.
East Nusa Tenggara Prov., Flores (Ruteng). Elevation: 1730–2205 m.
This epithet is based on the Indonesian word for “five” and is treated as a noun in apposition.
Trigonopterus lima Riedel, sp. n. was coded as “Trigonopterus sp. 297”.
Holotype, male (Fig.
Holotype (MZB): ARC1458 (EMBL # LM655556), West Nusa Tenggara Prov., Lombok, Tetebatu, Rinjani-trail from Orong Gerisak, sample 6, S08°29.577', E116°24.782', 1195 m, 04-IV-2010. Paratypes (ARC, MZB, SMNK, ZSM): Lombok: 11 exx, ARC1459 (EMBL # LM655557), same data as holotype; 23 exx, Tetebatu, Rinjani-trail from Orong Gerisak, sample 5, S08°30.096', E116°25.062', 1010 m, 04-IV-2010; 13 exx, Tetebatu, Rinjani-trail from Orong Gerisak, sample 7, S08°29.433', E116°24.746', 1240 m, 04-IV-2010; 10 exx, ARC1476 (EMBL # LM655574), ARC1477 (EMBL # LM655575), ARC1478 (EMBL # LM655576), Tetebatu, Rinjani-trail from Orong Gerisak, sample 8, S08°29.173', E116°24.517', 1345 m, 04-IV-2010; 10 exx, N Tetebatu, 1200-1450 m, 07-XII-2004; 1 ex, ARC1483 (EMBL # LM655581), Sesaot, Rinjani-trail, sample 3, S08°29.841', E116°13.679', 625 m, 28-III-2010; 7 exx, ARC2260 (EMBL # LM655714), Senaru, Rinjani-track, sample 1, S08°19.429', E116°24.082', 900 m, 21-III-2011; 12 exx, Senaru, Rinjani-track, sample 2, S08°20.570', E116°23.969', 1320 m, 21-III-2011; 17 exx, Lombok, Senaru, Rinjani-track, sample 3, S08°20.780', E116°23.790', 1465 m, 21-III-2011; 48 exx, Lombok, Senaru, Rinjani-track, sample 4, S08°20.439', E116°24.047', 1240 m, 21-III-2011; 47 exx, ARC2267 (EMBL # LM655721), ARC2268 (EMBL # LM655722), Senaru, Rinjani-track, sample 5, S08°19.800', E116°24.107', 1015 m, 21-III-2011; 31 exx, Lombok, Senaru, Rinjani-track, sample 6, S08°19.359', E116°24.070', 860 m, 23-III-2011; 40 exx, Senaru, Rinjani-trail, sample 7, S08°19.643', E116°24.033', 935 m, 23-III-2011; 48 exx, Senaru, Rinjani-trail, sample 8, S08°19.719', E116°24.040', 955 m, 23-III-2011; 87 exx, ARC2279 (EMBL # LM655733), ARC2280 (EMBL # LM655734), Santong, Rinjani-track, sample 1, S08°20.715', E116°19.695', 830 m, 24-III-2011; 150 exx, Santong, Rinjani-track, sample 2, S08°20.813', E116°19.778', 870 m, 24-III-2011; 67 exx, Santong, Rinjani-track, sample 3, S08°20.996', E116°20.001', 960 m, 24-III-2011; 27 exx, Santong, Rinjani-track, sample 4, S08°21.160', E116°20.067', 1005 m, 25-III-2011; 28 exx, Santong, Rinjani-track, sample 5, S08°21.351', E116°20.208', 1105 m, 25-III-2011; 27 exx, Santong, Rinjani-track, sample 6, S08°21.754', E116°20.476', 1315 m, 25-III-2011.
West Nusa Tenggara Prov., Lombok (Santong, Senaru, Sesaot, Tetebatu). Elevation: 625–1345 m.
This epithet is based on the island of Lombok.
Trigonopterus lombokensis Riedel, sp. n. was coded as “Trigonopterus sp. 323” by
Holotype, male (Fig.
Holotype (MZB): ARC2332 (EMBL # LM655785), East Java Prov., Banyuwangi Reg., Meru Betiri N.P., Sukamade, sample 2, S08°32.221', E113°51.985', 260 m, 12/13-IV-2011. Paratypes (MZB, SMNK, ZSM): E-Java Prov., Banyuwangi Reg., Meru Betiri N.P., Sukamade: 5 exx, ARC2333 (EMBL # LM655786), same data as holotype; 2 exx, ARC2334 (EMBL # LM655787), ARC2335 (EMBL # LM655788), sample 4, S08°32.163', E113°51.904', 340 m, 13-IV-2011.
E-Java Prov. (Meru Betiri N.P.). Elevation: 260–340 m.
This epithet is based on the type locality, Meru Betiri National Park.
Trigonopterus merubetirensis Riedel, sp. n. was coded as “Trigonopterus sp. 346”.
Holotype, male (Fig.
Holotype (MZB): ARC2250 (EMBL # LM655705), Bali, Negara, Kampung Pasatan, Desa Pohsanten, Mt. Mesehe, sample 2, S08°17.299', E114°41.494', 460 m, 17-III-2011. Paratypes (MZB, SMNK, ZSM): Bali, Negara, Kampung Pasatan, Desa Pohsanten, Mt. Mesehe: 5 exx, ARC2251 (EMBL # LM655706), same data as holotype; 6 exx, ARC2248 (EMBL # LM655703), ARC2249 (EMBL # LM655704), sample 1, S08°17.369', E114°41.392', 550 m, 17-III-2011; 15 exx, sample 3, S08°17.403', E114°41.424', 515 m, 17-III-2011; 5 exx, sample 4, S08°17.466', E114°41.331', 490 m, 17-III-2011; 1 ex, sample 6, S08°16.234', E114°41.528', 935 m, 18-III-2011; 3 exx, ARC2247 (EMBL # LM655702), S08°16.454', E114°41.725', 840 m, hand-collected from foliage, 18-III-2011; 26 exx, ARC0727, ARC0728, ARC0729, ARC0732 (PCR failed), trail to Mesehe waterfall, S08°18.30', E114°41.68', 350–600 m, 14.&17.XII.2008; 8 exx, ARC2962 (EMBL # LM656015), sample 8, S08°16.454', E114°41.725', 825 m, 18-III-2011.
Bali (Mt. Mesehe). Elevation: 460–840 m.
This epithet is based on the type locality Mt. Mesehe.
Trigonopterus mesehensis Riedel, sp. n. was coded as “Trigonopterus sp. 334” by
Holotype, male (Fig.
Holotype (MZB): ARC2213 (EMBL # LM655669), East Nusa Tenggara Prov., Flores, Labuhan Bajo, Roe, earthworm-dominated habitat, sample 1, S08°36.540', E120°01.871', 955 m, 13-III-2011. Paratypes (MZB, SMNK, ZSM): 3 exx, ARC2214 (EMBL # LM655670), ARC2216 (EMBL # LM655672), same data as holotype; 3 exx, ARC2220 (EMBL # LM655676), ARC2221 (EMBL # LM655677), ARC2222 (EMBL # LM655678), Flores, Labuhan Bajo, Roe, sample 5, S08°35.395', E120°00.383', 790 m, 13-III-2011; 1 ex, ARC2230 (EMBL # LM655686), Flores, Labuhan Bajo, Roe, sample 3, S08°36.259', E120°01.539', 975 m, 13-III-2011.
East Nusa Tenggara Prov., Flores (Labuhan Bajo). Elevation: 790–975 m.
This epithet is based on the Latin adjective micans (shining) and refers to the metallic coloration of this species.
Trigonopterus micans Riedel, sp. n. was coded as “Trigonopterus sp. 348” by
Holotype, male (Fig.
Holotype (MZB): ARC0263 (EMBL # LM655454), E-Sumatra, Lampung Prov., Bawang, Pedada Bay, Mt. Tanggang, sample 1, S05°43.933', E105°06.598', 579 m, 09-VIII-2006. Paratypes (MZB, SMNK, ZSM): E-Sumatra, Lampung Prov., Bawang, Pedada Bay, Mt. Tanggang: 2 exx, ARC0264 (EMBL # LM655455), ARC0265 (EMBL # LM655456), same data as holotype; 2 exx, sample 2, S05°43.947', E105°06.480', 659 m, 09-VIII-2006.
Lampung Prov. (Pedada Bay). Elevation: 579–744 m.
This epithet is based on the Latin adjective misellus (poor).
Trigonopterus misellus Riedel, sp. n. was coded as “Trigonopterus sp. 336”.
Holotype, male (Fig.
Holotype (SMNK): ARC1440 (EMBL # LM655554), PHILIPPINES, Palawan, Puerto Princessa region, Sabang, Mt. Bloomfield, N10°11.62', E118°52.35', 500-700 m, 10-XII-2009. Paratypes (SMNK): 2 exx, ARC1441 (EMBL # LM655555), ARC1442 (PCR failed), same data as holotype.
Palawan (Mt. Bloomfield). Elevation: ca. 500–700 m.
This epithet is based on the type locality, the island of Palawan.
Trigonopterus palawanensis Riedel, sp. n. was coded as “Trigonopterus sp. 344”.
Holotype, male (Fig.
Holotype (MZB): ARC2682 (EMBL # LM655940), West Java Prov., Ciamis, Pangandaran, sample 2, S07°42.836', E108°39.478', 157 m, 22-IV-2012. Paratypes (SMNK, ZSM): W-Java Prov., Ciamis, Pangandaran: 3 exx, ARC2683 (EMBL # LM655941), same data as holotype; 3 exx, ARC2686 (EMBL # LM655944), ARC2687 (EMBL # LM655945), ARC3637 (EMBL # LM656059), sample 3, S07°43.070', E108°39.634', 156 m, 22-IV-2012.
W-Java Prov. (Pangandaran). Elevation: 156–157 m.
This epithet is based on the type locality, Pangandaran.
Trigonopterus pangandaranensis Riedel, sp. n. was coded as “Trigonopterus sp. 372”.
Holotype, male (Fig.
Holotype (MZB): ARC3622 (EMBL # LM656052), East Nusa Tenggara Prov., Flores, Ruteng, Mt. Ranaka, sample 7, S08°37.781', E120°31.184', 1730 m, 10-III-2011. Paratypes (MZB, SMNK): Flores: 4 exx, ARC2205 (EMBL # LM655661), ARC2206 (EMBL # LM655662), ARC3623 (EMBL # LM656053), ARC3624 (EMBL # LM656054), same data as holotype.
East Nusa Tenggara Prov., Flores (Mt. Ranaka). Elevation: 1730 m.
This epithet is based on a combination of the Greek prefix para- (next to; near by) and the specific epithet of T. florensis Riedel, sp. n., a sibling species.
Trigonopterus paraflorensis Riedel, sp. n. was coded as “Trigonopterus sp. 441”. Morphologically it is very similar to T. florensis Riedel, sp. n. and T. pseudoflorensis Riedel, sp. n. but its cox1 sequences differ 7.75%, respectively 7.14% smallest interspecific p-distance.
Holotype, male (Fig.
Holotype (MZB): ARC0585 (EMBL # LM655521), Bali, Bedugul, Tamblingan, above the village along water pipe, sample 4, S08°15.854', E115°05.312', 1310 m, 06-XI-2007. Paratypes (MZB, SMNK, ZSM): Bali: 2 exx, ARC0584 (EMBL # LM655520), same data as holotype; 2 exx, ARC3586 (EMBL # LM656019), ARC3587, Tamblingan, sample 2, S08°15.015', E115°06.202', 1255 m, 06-XI-2007; 2 exx, ARC2315 (EMBL # LM655768), ARC2316 (EMBL # LM655769), Tamblingan, sample 5, S08°15.108', E115°05.573', 1350 m, 03-IV-2011; 1 ex, ARC2306 (EMBL # LM655759), Mt. Batukaru, Wangayagede, sample 3, S08°21.431', E115°05.699', 1170 m, 31-III-2011; 2 exx, ARC2310 (EMBL # LM655763), ARC2311 (EMBL # LM655764), Mt. Batukaru, Wangayagede, sample 5, S08°21.130', E115°05.321', 1520 m, 31-III-2011; 1 ex, ARC2255 (EMBL # LM655709), Negara, Kampung Pasatan, Desa Pohsanten, Mt. Mesehe, sample 8, S08°16.454', E114°41.725', 825 m, 18-III-2011, sifted.
Bali (Mt. Batukaru, Mt. Mesehe, Tamblingan). Elevation: 825–1520 m.
This epithet is based on a combination of the Greek prefix para- (next to; near by) and the specific epithet of T. rugosus Riedel, sp. n., a sibling species.
Trigonopterus pararugosus Riedel, sp. n. was coded as “Trigonopterus sp. 327” by
Holotype, male (Fig.
Holotype (MZB): ARC3603 (EMBL # LM656033), West Nusa Tenggara Prov., Sumbawa, Batu Dulang, Gn. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 12-IV-2010. Paratypes (MZB, SMNK, ZSM): West Nusa Tenggara Prov., Sumbawa: 5 exx, ARC3641 (EMBL # LM656062), ARC3642 (EMBL # LM656063), ARC3644 (EMBL # LM656065), ARC3645 (EMBL # LM656066), same data as holotype; 1 ex, ARC3649 (EMBL # LM656068), Batu Dulang, Gn. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 18-IV-2010; 1 ex, ARC3656 (EMBL # LM656074), Batu Dulang, Gn. Batu Pasak, sample 5, S08°37.005', E117°15.790', 1320 m, 18-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang). Elevation: 1385 m.
This epithet is based on the Greek prefix para- (next to; near by) and the name of Trigonopterus sumbawensis Riedel, sp. n., a sibling species.
Trigonopterus parasumbawensis Riedel, sp. n. was coded as “Trigonopterus sp. 443”.
Holotype, male (Fig.
Holotype (MZB): ARC2340 (EMBL # LM655790), West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 7, S08°35.176', E117°08.295', 1490 m, 16-IV-2010. Paratypes (MZB, SMNK, ZSM): West Nusa Tenggara Prov., Sumbawa: 3 exx, ARC1534 (EMBL # LM655632), ARC2342 (EMBL # LM655792), ARC2344 (EMBL # LM655794), same data as holotype; 2 exx, ARC1514 (EMBL # LM655612), ARC1515 (EMBL # LM655613), Batu Dulang, Mt. Batu Pasak, sample 2, S08°37.028', E117°15.783', 1305 m, 12-IV-2010; 1 ex, ARC3652 (EMBL # LM656070), Batu Dulang, Mt. Batu Pasak, sample 4, S08°37.318', E117°15.339', 1280 m, 18-IV-2010; 1 ex, ARC3612 (EMBL # LM656042), Tepal, Pc. Nengas, sample 3, S08°35.386', E117°08.251', 1415 m, 15-IV-2010; 2 exx, ARC3605 (EMBL # LM656035), ARC3607 (EMBL # LM656037), Tepal, Pc. Nengas, sample 2, S08°35.884', E117°08.384', 1310 m, 15-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang, Tepal). Elevation: 1280–1490 m.
This epithet is based on the Latin adjective pauxillus (small) and refers to its body size.
Trigonopterus pauxillus Riedel, sp. n. was coded as “Trigonopterus sp. 324” by
Holotype, male (Fig.
Holotype (MZB): ARC2710 (EMBL # LM655966), West Java Prov., Garut, Cikajang, Mt. Payung, sample 2, S07°25.268', E107°48.492', 1250 m, 26-IV-2012. Paratypes (MZB, SMNK, ZSM): 7 exx, ARC3598 (EMBL # LM656028), ARC3599 (EMBL # LM656029), ARC3600 (EMBL # LM656030), ARC3601 (EMBL # LM656031), same data as holotype; 3 exx, ARC2702 (EMBL # LM655958), ARC2703 (EMBL # LM655959), ARC3590 (EMBL # LM656021), W-Java Prov., Garut, Cikajang, Mt. Payung, sample 1, S07°25.345', E107°48.825', 1085 m, 26-IV-2012.
W-Java Prov. (Mt. Payung). Elevation: 1085–1250 m.
This epithet is based on the type locality.
Trigonopterus payungensis Riedel, sp. n. was coded as “Trigonopterus sp. 312”.
Holotype, male (Fig.
Holotype (MZB): ARC2699 (EMBL # LM655955), West Java, Garut, Cikajang, Mt. Payung, sample 1, S07°25.345', E107°48.825', 1085 m, 26-IV-2012. Paratypes (MZB, SMNK, ZSM): W-Java Prov., Garut, Cikajang, Mt. Payung: 10 exx, ARC2700 (EMBL # LM655956), ARC2701 (EMBL # LM655957), ARC3638 (EMBL # LM656060), same data as holotype; 7 exx, ARC2711 (EMBL # LM655967), sample 2, S07°25.268', E107°48.492', 1250 m, 26-IV-2012; 8 exx, ARC2714 (EMBL # LM655970), sample 3, S07°25.320', E107°48.259', 1560 m, 26-IV-2012; 1 ex, ARC3594 (EMBL # LM656025), Sumedang, Sukajadi, Mt. Cakrabuana, sample 5, S07°01.804', E108°08.044', 1563 m, 20-IV-2012.
W-Java Prov. (Mt. Payung, Mt. Cakrabuana). Elevation: 1085–1563 m.
This epithet is based on the Latin porca (ridge between two furrows) and to be treated as an adjective. It refers to the elytral sculpture.
Trigonopterus porcatus Riedel, sp. n. was coded as “Trigonopterus sp. 370”.
Holotype, male (Fig.
Holotype (MZB): ARC2209 (EMBL # LM655665), East Nusa Tenggara Prov., Flores, Ruteng, Mt. Ranaka, sample 6, S08°38.243', E120°31.569', 1995 m, 09-III-2011. Paratypes (SMNK, ZSM): Flores: 3 exx, ARC3613 (EMBL # LM656043), ARC3614 (EMBL # LM656044), ARC3615 (EMBL # LM656045), Ruteng, Mt. Ranaka, sample 5, S08°38.277', E120°31.616', 2010 m, 09-III-2011.
East Nusa Tenggara Prov., Flores (Mt. Ranaka). Elevation: 1995–2010 m.
This epithet is based on the Greek prefix pseudo (false) and the name of Trigonopterus florensis Riedel, sp. n., a sibling species.
Trigonopterus pseudoflorensis Riedel, sp. n. was coded as “Trigonopterus sp. 352” by
Holotype, male (Fig.
Holotype (MZB): ARC3602 (EMBL # LM656032), West Nusa Tenggara Prov., Sumbawa, Batu Dulang, Gn. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 12-IV-2010. Paratypes (SMNK): Sumbawa, Batu Dulang, Gn. Batu Pasak: 2 exx, ARC3604 (EMBL # LM656034), ARC3643 (EMBL # LM656064), ARC3646 (EMBL # LM656067), ARC3651 (EMBL # LM656069), same data as holotype; 1 ex, ARC3657 (EMBL # LM656075), sample 5, S08°37.005', E117°15.790', 1320 m, 18-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang). Elevation: 1385 m.
This epithet is based on the Greek prefix pseudo (false) and the name of Trigonopterus sumbawensis Riedel, sp. n., a sibling species.
Trigonopterus pseudosumbawensis Riedel, sp. n. was coded as “Trigonopterus sp. 442”.
Holotype, male (Fig.
Holotype (MZB): ARC1497 (EMBL # LM655595), West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 2, S08°35.884', E117°08.384', 1310 m, 15-IV-2010. Paratypes (MZB, SMNK, ZSM): West Nusa Tenggara Prov., Sumbawa: 1 ex, ARC1498 (EMBL # LM655596), same data as holotype; 1 ex, ARC1535 (EMBL # LM655633), Tepal, Pc. Nengas, sample 5, S08°35.740', E117°08.721',1330 m, 16-IV-2010; 2 exx, Tepal, Pc. Nengas, sample 6, S08°35.533', E117°08.605',1350 m, 16-IV-2010; 1 ex, ARC1518 (EMBL # LM655616), Batu Dulang, Mt. Batu Pasak, sample 2, S08°37.028', E117°15.783', 1305 m, 12-IV-2010; 1 ex, ARC1528 (EMBL # LM655626), Batu Dulang, Mt. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 18-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang, Tepal). Elevation: 1305–1385 m.
This epithet is based on a combination of the Latin participles punctatus (punctate) and seriatus (in rows) and refers to the elytral sculpture.
Trigonopterus punctatoseriatus Riedel, sp. n. was coded as “Trigonopterus sp. 287” by
Holotype, male (Fig.
Holotype (MZB): ARC2196 (EMBL # LM655652), East Nusa Tenggara Prov., Flores Isl., Ruteng, Mt. Ranaka, sample 1, S08°37.321', E120°31.463', 1535 m, 08-III-2011 (MZB). Paratypes (MZB, SMNK): Flores Isl., Ruteng, Mt. Ranaka: 1 ex, ARC2197 (EMBL # LM655653), same data as holotype; 1 ex, ARC2198 (EMBL # LM655654), sample 3, S08°38.099', E120°31.745', summit area, 2205 m, 09-III-2011; 1 ex, ARC2582 (EMBL # LM655891), sample 7, S08°37.714', E120°31.509', 1850m, 09-III-2011.
East Nusa Tenggara Prov., Flores (Ruteng). Elevation: 1535–2205 m.
This epithet is based on the type locality.
Trigonopterus ranakensis Riedel, sp. n. was coded as “Trigonopterus sp. 342” by
Holotype, male (Fig.
Holotype (MZB): ARC0308 (EMBL # LM655498), E-Java Prov., Banyuwangi, Mt. Ijen, Licin, sample 4, S08°06.673', E114°14.488', 1255 m, 31-VIII-2006. Paratypes (MZB, SMNK, ZSM): E-Java Prov.: 19 exx, ARC0309 (EMBL # LM655499), same data as holotype; 33 exx, ARC0301 (EMBL # LM655491), ARC0302 (EMBL # LM655492), Banyuwangi, Mt. Ijen, Licin, sample 2, S08°05.603', E114°14.366', 1575 m, 28-VIII-2006; 12 exx, ARC0310 (EMBL # LM655500), ARC0311 (EMBL # LM655501), Banyuwangi, Mt. Ijen, Licin, sample 3, S08°06.056', E114°14.621', 1400 m, 28-VIII-2006; 2 exx, Banyuwangi, Mt. Ijen, Licin, sample 4, S08°06.673', E114°14.488', 1255 m, 28-VIII-2006; 2 exx, Banyuwangi, Mt. Ijen, Licin, sample 6, S08°07.115', E114°14.650', 1100 m, 31-VIII-2006; 9 exx, ARC2460 (EMBL # LM655798), ARC2461 (EMBL # LM655799), Krucil, Bremi, Mt. Argopuro (trail to Taman Hidup), sample 2, S07°58.478', E113°31.069', 1457 m, 10-XI-2011; 4 exx, ARC2462 (EMBL # LM655800), ARC2463 (EMBL # LM655801), Krucil, Bremi, Mt. Argopuro (trail to Taman Hidup), sample 3, S07°58.586', E113°31.345', 1628 m, 10-XI-2011; 2 exx, Krucil, Bremi, Mt. Argopuro (trail to Taman Hidup), sample 7, S07°58.449', E113°31.002', 1457 m, 11-XI-2011.
E-Java Prov. (Mt. Argopuro, Mt. Ijen). Elevation: 1100–1628 m.
This epithet is based on the Latin participle relictus (left behind), referring to the distribution of the species which inhabits an isolated patch of wet forest.
Trigonopterus relictus Riedel, sp. n. was coded as “Trigonopterus sp. 317” by
Holotype, male (Fig.
Holotype (MZB): ARC2263 (EMBL # LM655717), West Nusa Tenggara Prov., Lombok, Gn. Rinjani, Senaru, Rinjani-track, sample 3, S08°20.780', E116°23.790', 1465 m, 21-III-2011. Paratypes (MZB, SMNK, ZSM): West Nusa Tenggara Prov., Lombok, Gn. Rinjani: 2 exx, ARC2264 (EMBL # LM655718), same data as holotype; 2 exx, ARC1464 (EMBL # LM655562), ARC1465 (EMBL # LM655563), Tetebatu, Rinjani-trail from Orong Gerisak, sample 8, S08°29.173', E116°24.517', 1345 m, 04-IV-2010; 1 ex, ARC1469 (EMBL # LM655567), Tetebatu, Rinjani-trail from Orong Gerisak, sample 7, S08°29.433', E116°24.746', 1240 m, 04-IV-2010; 1 ex, ARC1470 (EMBL # LM655568), Tetebatu, Rinjani-trail from Kokok Belimbing, sample 3, S08°29.433', E116°24.746', 1245 m, 02-IV-2010; 1 ex, ARC1473 (EMBL # LM655571), Tetebatu, Rinjani-trail from Kokok Belimbing, sample 4, S08°28.320', E116°26.459', 1420 m, 02-IV-2010; 5 exx, ARC0167, Tetebatu, Rinjani-trail, 1200-1450 m, 07-XII-2004; ARC1487 (EMBL # LM655585), road between Sembalun and Sapit, sample 1, S08°25.110', E116°31.947', 1450 m, 30-III-2010; 1 ex, ARC1489 (EMBL # LM655587), road between Sembalun and Sapit, sample 4, S08°25.331', E116°31.841', 1405 m, 31-III-2010; 1 ex, ARC1492 (EMBL # LM655590), road between Sembalun and Sapit, sample 5, S08°26.453', E116°31.859', 1195 m, 31-III-2010.
West Nusa Tenggara Prov., Lombok (Senaru, Sembalun, Tetebatu). Elevation: 1195–1465 m.
This epithet is based on the type locality, Mt. Rinjani.
Trigonopterus rinjaniensis Riedel, sp. n. was coded as “Trigonopterus sp. 282” by
Holotype, male (Fig.
Holotype (MZB): ARC2219 (EMBL # LM655675), East Nusa Tenggara Prov., Flores, Labuhan Bajo, Roe, sample 1, S08°36.540', E120°01.871', 955 m, 13-III-2011. Paratypes (MZB, SMNK): Flores: 1 ex, ARC2218 (EMBL # LM655674), same data as holotype; 2 exx, ARC2227 (EMBL # LM655683), ARC2228 (EMBL # LM655684), Labuhan Bajo, Roe, sample 5, S08°35.395', E120°00.383', 790 m, 13-III-2011; 2 exx, ARC2245 (EMBL # LM655700), ARC2246 (EMBL # LM655701), Labuhan Bajo, Tebedo, lowland forest, sample 1, S08°29.848', E119°59.633', 525 m, 14-III-2011.
East Nusa Tenggara Prov., Flores (Labuhan Bajo). Elevation: 525–955 m.
This epithet is based on the type locality.
Trigonopterus roensis Riedel, sp. n. was coded as “Trigonopterus sp. 353” by
Holotype, male (Fig.
Holotype (MZB): ARC2672 (EMBL # LM655930), West Java Prov., Sumedang, Sukajadi, Mt. Cakrabuana, sample 4, S07°01.734', E108°08.059', 1530 m, 19-IV-2012. Paratypes (MZB, SMNK, ZSM): W-Java Prov.: 3 exx, ARC2673 (EMBL # LM655931), ARC2674 (EMBL # LM655932), ARC3591 (EMBL # LM656022), same data as holotype; 4 exx, ARC2675 (EMBL # LM655933), ARC2676 (EMBL # LM655934), ARC2677 (EMBL # LM655935), ARC3592 (EMBL # LM656023), Sumedang, Sukajadi, Mt. Cakrabuana, sample 5, S07°01.804', E108°08.044', 1563 m, 20-IV-2012; 1 ex, ARC2681 (EMBL # LM655939), Sumedang, Sukajadi, Mt. Cakrabuana, sample 6, S07°01.943', E108°08.029', 1593 m, 20-IV-2012; 1 ex, ARC2671 (EMBL # LM655929), Sumedang, Sukajadi, Mt. Cakrabuana, sample 3, S07°01.493', E108°08.065', 1394 m, 19-IV-2012; 5 exx, ARC0157, ARC0408, Ciamis, Mt. Sawal, Blok Cireong, sample 2, S07°14.127', E108°15.568', 1120 m, 01-X-2005; 3 exx, Batu Cakra, sample 1, S07°14.920', E108°15.762', 990 m, 01-X-2005; 3 exx, ARC2505 (EMBL # LM655842), ARC2506 (EMBL # LM655843), ARC2507 (EMBL # LM655844), Panjalu, Tembong, sample 3, S07°11.102', E108°16.310', 1569 m, 29-XI-2011.
W-Java Prov. (Mt. Cakrabuana, Mt. Sawal). Elevation: 990–1569 m.
This epithet is a combination of the Latin adjective rugosus (wrinkled) and the participle striatus (provided with furrows) and refers to the species´ body-sculpture.
Trigonopterus rugosostriatus Riedel, sp. n. was coded as “Trigonopterus sp. 354”.
Holotype, male (Fig.
Holotype (MZB): ARC0586 (EMBL # LM655522), Bali, Bedugul, Mt. Catur, sample 3, S08°15.626', E115°11.354', 1690 m, 07-XI-2007. Paratypes (MZB, SMNK, ZSM): Bali: 3 exx, ARC0587 (EMBL # LM655523), same data as holotype; 3 exx, Bedugul, Mt. Catur, sample 2, S08°15.147', E115°11.323', 1930 m, 07-XI-2007; 3 exx, ARC0571 (EMBL # LM655507), ARC0572 (EMBL # LM655508), Bedugul, Mt. Pohen (= Mt. Tapak = Mt.Keramat), sample 2, S08°16.328', E115°08.360', 1785 m, 01-XI-2007; 2 exx, ARC0573 (EMBL # LM655509), Bedugul, Mt. Pohen (= Mt. Tapak = Mt.Keramat), sample 3, S08°16.386', E115°08.402', 1735 m, 01-XI-2007; 1 ex, ARC0574 (EMBL # LM655510), Bedugul, Mt. Pohen (= Mt. Tapak = Mt.Keramat), sample 5, S08°16.526', E115°08.634', 1580 m, 01-XI-2007; 1 ex Bedugul, Mt. Pohen (= Mt. Tapak = Mt.Keramat), sample 6, S08°16.607', E115°08.773', 1460 m, 01-XI-07; 4 exx, ARC0591 (EMBL # LM655527), ARC0592 (EMBL # LM655528), ARC0593 (EMBL # LM655529), ARC0594 (EMBL # LM655530), Kintamani, Mt. Abang, sample 1, S08°17.118', E115°24.868', 1440 m, 09-XI-2007, 01-XI-2007; 6 exx, ARC2317 (EMBL # LM655770), ARC2318 (EMBL # LM655771), Angseri, Mt. Adeng, sample 3, S08°19.626', E115°08.554', 1385 m, 06-IV-2011; 5 exx, Angseri, Mt. Adeng, sample 4, S08°19.463', E115°08.537', 1535 m, 06-IV-2011; 9 exx, Angseri, Mt. Adeng, sample 5, S08°19.402', E115°08.545', 1590 m, 06-IV-2011; 5 exx, ARC2321 (EMBL # LM655774), ARC2322 (EMBL # LM655775), Angseri, Mt. Adeng, sample 6, S08°19.138', E115°08.416', 1760 m, 06-IV-2011; 1 female, Bedugul, mountain NW of botanical garden, 1400 m, 04-XII-2004 (ARC).
Bali (Mt. Andeng, Bedugul, Kintamani). Elevation: 1385–1785 m.
This epithet is based on the Latin adjective rugosus and refers to the wrinkled integument.
Trigonopterus rugosus Riedel, sp. n. was coded as “Trigonopterus sp. 280” by
Holotype, male (Fig.
Holotype (MZB): ARC2193 (EMBL # LM655649), East Nusa Tenggara Prov., Flores Isl., Danau Ranamese, sample 2, S08°37.235', E120°33.554', 1295 m, 08-III-2011. Paratypes (MZB, SMNK, ZSM): Flores Isl., Ruteng: 1 ex, ARC2207 (EMBL # LM655663), Mt. Ranaka, sample 2, S08°37.164', E120°31.106', 1450 m, 08-III-2011; 1 ex, ARC3627 (EMBL # LM656057), Mt. Ranaka, sample 11, S08°37.386', E120°31.156', 1575 m, 08-III-2011; 4 exx, ARC2584 (EMBL # LM655893), ARC2585 (EMBL # LM655894), Mt. Ranaka, sample 10, S08°37.481', E120°31.410', 1610 m, 10-III-2011; 3 exx, ARC2208 (EMBL # LM655664), ARC2577 (EMBL # LM655886), ARC2578 (EMBL # LM655887), Danau Ranamese, sample 4, S08°38.385', E120°33.568', 1215 m, 08-III-2011; 2 exx, ARC2586 (EMBL # LM655895), Danau Ranamese, sample 6, S08°38.370', E120°33.798', 1215 m, 11-III-2011.
East Nusa Tenggara Prov., Flores (Ruteng). Elevation: 1215–1610 m.
This epithet is based on the type locality.
Trigonopterus rutengensis Riedel, sp. n. was coded as “Trigonopterus sp. 306”.
Holotype, male (Fig.
Holotype (MZB): ARC1530 (EMBL # LM655628), West Nusa Tenggara Prov., Sumbawa, Batu Dulang, Gn. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 18-IV-2010. Paratypes (SMNK): 2 exx, ARC1529 (EMBL # LM655627), ARC1531 (EMBL # LM655629), same data as holotype.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang). Elevation: 1385 m.
This epithet is based on the Latin noun saltator (leaper) and refers to the biogeographic pattern where close relatives of this species are found on islands to the West.
Trigonopterus saltator Riedel, sp. n. was coded as “Trigonopterus sp. 281” by
Holotype, male (Fig.
Holotype (SMNK): ARC2523 (EMBL # LM655860), MALAYSIA, Sarawak, Kuching, Mt. Santubong, sample 3, N01°43.884', E110°19.727', 496 m, 06-XII-2011. Paratypes (SMNK, ZSM): MALAYSIA, Sarawak, Kuching, Mt. Santubong: 1 ex, ARC2524 (EMBL # LM655861), same data as holotype; 13 exx, ARC2527 (EMBL # LM655864), ARC2528 (EMBL # LM655865), sample 1, N01°43.760', E110°19.113', 138 m, 05-XII-2011; 6 exx, sample 2, N01°43.779', E110°19.183', 173 m, 05-XII-2011; 1 ex, sample 3, N01°43.884', E110°19.727', 496 m, 06-XII-2011; 10 exx, sample 4, N01°43.884', E110°19.715', 487 m, 06-XII-2011; 8 exx, sample 5, N01°43.843', E110°19.672', 478 m, 06-XII-2011; 10 exx, sample 6, N01°43.830', E110°19.601', 375 m, 06-XII-2011.
Sarawak (Santubong). Elevation: 138–496 m.
This epithet is based on the type locality.
Trigonopterus santubongensis Riedel, sp. n. was coded as “Trigonopterus sp. 361”.
Holotype, male (Fig.
Holotype (MZB): ARC1482 (EMBL # LM655580), West Nusa Tenggara Prov., Lombok, Sesaot, Rinjani-trail, sample 4, S08°30.047', E116°13.826', 615 m, 28-III-2010. Paratypes (MZB, SMNK, ZSM): Lombok: 2 exx, ARC1481 (EMBL # LM655579), same data as holotype; 1 ex, ARC1484 (EMBL # LM655582), Sesaot, Rinjani-trail, sample 3, S08°29.841', E116°13.679', 625 m, 28-III-2010; 1 ex, ARC2301 (EMBL # LM655755), W Sengigi, Mt. Pusuk, sample 3, S08°27.723', E116°04.160', 585 m, 29-III-2011; 1 ex, ARC2285 (EMBL # LM655739), Sajang, sample 1, S08°19.370', E116°29.570', 845 m, 26-III-2011; 10 exx, ARC2295 (EMBL # LM655749), ARC2296 (EMBL # LM655750), ARC2297 (EMBL # LM655751), ARC2298 (EMBL # LM655752), Mt. Pengasingan, SE Sembalun, sample 2, S08°19.354', E116°30.530', 945 m, 27-III-2011.
West Nusa Tenggara Prov., Lombok (Mt. Pusuk, Sajang, Sembalun, Sesaot). Elevation: 615–945 m.
This epithet is based on the Sasak tribe traditionally inhabiting the island of Lombok. It is a noun in apposition.
Trigonopterus sasak Riedel, sp. n. was coded as “Trigonopterus sp. 330” by
Holotype, male (Fig.
Holotype (MZB): ARC2331 (EMBL # LM655784), E-Java Prov., Banyuwangi Reg., Meru Betiri N.P., Sukamade, sample 2, S08°32.221', E113°51.985', 260 m, 12/13-IV-2011. Paratypes (MZB, SMNK, ZSM): E-Java Prov., Banyuwangi Reg., Meru Betiri N.P., Sukamade: 17 exx, ARC2328 (EMBL # LM655781), ARC2329 (EMBL # LM655782), ARC2330 (EMBL # LM655783), same data as holotype; 1 ex, sample 4, S08°32.163', E113°51.904', 340 m, 13-IV-2011.
E-Java Prov. (Meru Betiri N.P.). Elevation: 260–340 m.
This epithet is based on the Indonesian word for “one” and is treated as a noun in apposition.
Trigonopterus satu Riedel, sp. n. was coded as “Trigonopterus sp. 341”.
Holotype, male (Fig.
Holotype (SMNK): ARC1438 (EMBL # LM655552), PHILIPPINES, Palawan, Puerto Princessa region, Sabang, Mt. Bloomfield, N10°11.62', E118°52.35', 500–700 m, 10-XII-2009. Paratypes (SMNK): 2 exx, ARC1437 (EMBL # LM655551), ARC1439 (EMBL # LM655553), same data as holotype.
Palawan (Mt. Bloomfield). Elevation: ca. 500–700 m.
This species is named in honour of Andreas Schulz (Leverkusen, Germany), a successful collector of edaphic arthropods.
Trigonopterus schulzi Riedel, sp. n. was coded as “Trigonopterus sp. 343” by
Holotype, male (Fig.
Holotype (MZB): ARC0715 (EMBL # LM655540), E-Kalimantan Prov., Berau Dist., Hutan Wisata Sei Tangap, ca. 8 km W of Tanjungredeb, N02°08.07', E117°24.65', 30 m, 02-X-2008. Paratypes (MZB, SMNK, ZSM): E-KALIMANTAN, Berau Dist.: 26 exx, ARC0714 (EMBL # LM655539), ARC0716 (EMBL # LM655541), ARC0717 (EMBL # LM655542), ARC0718 (EMBL # LM655543), ARC0719 (EMBL # LM655544), ARC0720 (EMBL # LM655545), ARC0721 (EMBL # LM655546), same data as holotype; 3 exx, Hutan Mayang Mangurai, ca. 15 km SW of Tanjungredeb, N02°06.217', E117°24.003‘, 20 m, 30-XI-2008; 15 exx, ARC1418, ARC1419, ARC1420, ARC1426 (PCR failed), ca. 45 km N of Tanjungredeb, 1 km off road Tanjungredeb - Tanjungselor, N02°29.55', E117°28.77', 190 m, 29.IX./3.X.2008; 2 exx, near Kampung Suaran, ca 40 km S of Tanjungredeb, primary forest on limestone, N01°59.70', E117°36.05', 50 m, 01-X-2008.
E-Kalimantan Prov. (Tanjungredeb). Elevation: 20–190 m.
This epithet is based on the Indonesian word for “eleven” and is treated as a noun in apposition.
Trigonopterus sebelas Riedel, sp. n. was coded as “Trigonopterus sp. 308” by
Holotype, male (Fig.
Holotype (MZB): ARC1499 (EMBL # LM655597), West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 2, S08°35.884', E117°08.384', 1310 m, 15-IV-2010 (MZB). Paratypes (MZB, SMNK, ZSM): West Nusa Tenggara Prov., Sumbawa: 2 exx, ARC1500 (EMBL # LM655598), ARC1501 (EMBL # LM655599), same data as holotype; 2 exx, Tepal, Pc. Nengas, sample 7, S08°35.176', E117°08.295', 1490 m, 16-IV-2010; 3 exx, ARC1519 (EMBL # LM655617), ARC1520 (EMBL # LM655618), ARC1521 (EMBL # LM655619), Batu Dulang, Mt. Batu Pasak, sample 2, S08°37.028', E117°15.783', 1305 m, 12-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang, Tepal). Elevation: 1305–1490 m.
This epithet is based on the Indonesian word for “nine” and is treated as a noun in apposition.
Trigonopterus sembilan Riedel, sp. n. was coded as “Trigonopterus sp. 283”.
Holotype, male (Fig.
Holotype (MZB): ARC2538 (EMBL # LM655875), W-Kalimantan Prov., Bengkayan, Suka-Bangun, Mt. Bawang, sample 5, N00°54.103', E109°22.515', 652 m, 11-XII-2011 (MZB). Paratypes (MZB, SMNK, ZSM): 8 exx, ARC2539 (EMBL # LM655876), ARC2540 (EMBL # LM655877), ARC2541 (EMBL # LM655878), same data as holotype.
W-Kalimantan Prov. (Mt. Bawang). Elevation: 652 m.
This epithet is based on the Indonesian word for “ten” and is treated as a noun in apposition.
Trigonopterus sepuluh Riedel, sp. n. was coded as “Trigonopterus sp. 365” by
Holotype, male (Fig.
Holotype (MZB): ARC2730 (EMBL # LM655986), E-Sumatra,, Lampung Prov., Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, S05°31.678', E104°25.508', 608 m, 02-V-2012. Paratypes (MZB, SMNK, ZSM): E-Sumatra, Lampung Prov.: 5 exx, ARC0271 (EMBL # LM655462), Bawang, Pedada Bay, Mt. Tanggang, sample 1, S05°43.933', E105°06.598', 579 m, 09-VIII-2006; 5 exx, ARC0268 (EMBL # LM655459), ARC0269 (EMBL # LM655460), ARC0270 (EMBL # LM655461), Bawang, Pedada Bay, Mt. Tanggang, sample 2, S05°43.947', E105°06.480', 659 m, 09-VIII-2006; 1 ex, Bawang, Pedada Bay, Mt. Tanggang, sample 3, S05°43.938', E105°06.440', 673 m, 09-VIII-2006; 4 exx, ARC0272 (EMBL # LM655463), Bawang, Pedada Bay, Mt. Tanggang, sample 4, S05°43.871', E105°06.393', 744 m, 09-VIII-2006; 4 exx, ARC2727 (EMBL # LM655983), ARC2728 (EMBL # LM655984), Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 2, S05°30.330', E104°25.838', 660 m, 01-V-2012; 2 exx, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 3, S05°32.289', E104°26.295', 495 m, 01-V-2012; 11 exx, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 4, S05°33.406', E104°25.680', 405 m, 01-V-2012; 2 exx, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 5, S05°31.044', E104°25.664', 562 m, 02-V-2012; 1 ex, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 6, S05°31.678', E104°25.508', 608 m, 02-V-2012; 2 exx, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 7, S05°33.768', E104°25.361', 371 m, 02-V-2012; 1 ex, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 8, S05°33.032', E104°25.985', 421 m, 02-V-2012; 1 ex, Bukit Barisan Selatan N.P., Kota Agung, Sukaraja, sample 9, S05°32.611', E104°26.353', 455 m, 02-V-2012.
Lampung Prov. (Bukit Barisan Selatan N.P., Pedada Bay). Elevation: 371–744 m.
This epithet is based on the Latin adjective seriatus (arranged in rows) and refers to the elytral scales.
Trigonopterus seriatus Riedel, sp. n. was coded as “Trigonopterus sp. 339”.
Holotype, male (Fig.
Holotype (MZB): ARC2236 (EMBL # LM655692), East Nusa Tenggara Prov., Flores, Labuhan Bajo, Tebedo, sample 1: S08°29.848', E119°59.633', 525 m, 14-III-2011. Paratypes (MZB, SMNK, ZSM): 12 exx, ARC2237 (EMBL # LM655693), ARC2238 (EMBL # LM655694), ARC2239 (EMBL # LM655695), ARC2240 (EMBL # LM655696), same data as holotype; 1 ex, ARC2223 (EMBL # LM655679), Flores, Labuhan Bajo, Roe, sample 5, S08°35.395', E120°00.383', 790 m, 13-III-2011; 1 ex, ARC2229 (EMBL # LM655685), Flores, Labuhan Bajo, Roe, sample 3, S08°36.259', E120°01.539', 975 m, 13-III-2011.
East Nusa Tenggara Prov., Flores (Roe, Tebedo). Elevation: 525–975 m.
This epithet is a combination of the Latin nouns serra (saw) and femur (thigh) and is to be treated as noun in apposition.
Trigonopterus serratifemur Riedel, sp. n. was coded as “Trigonopterus sp. 351”.
Holotype, male (Fig.
Holotype (MZB): ARC2713 (EMBL # LM655969), West Java, Garut, Cikajang, Mt. Payung, sample 3, S07°25.320', E107°48.259', 1560 m, 26-IV-2012. Paratypes (MZB, SMNK): W-Java Prov., Garut, Cikajang, Mt. Payung: 4 exx, ARC2704 (EMBL # LM655960), ARC2705 (EMBL # LM655961), ARC2709 (EMBL # LM655965), sample 2, S07°25.268', E107°48.492', 1250 m, 26-IV-2012; 1 ex, same data as holotype.
W-Java Prov. (Mt. Payung). Elevation: 1250–1560 m.
This epithet is a combination of the Latin noun seta (bristle) and the suffix –fer (bear, carry) and refers to the elytral vestiture.
Trigonopterus setifer Riedel, sp. n. was coded as “Trigonopterus sp. 374”.
Holotype, male (Fig.
Holotype (MZB): ARC2668 (EMBL # LM655926), West Java Prov., Sumedang, Sukajadi, Mt. Cakrabuana, sample 3, S07°01.493', E108°08.065', 1394 m, 19-IV-2012. Paratypes (MZB, SMNK, ZSM): W-Java Prov., Sumedang, Sukajadi, Mt. Cakrabuana: 7 exx, ARC2669 (EMBL # LM655927), ARC2670 (EMBL # LM655928), same data as holotype; 1 ex, sample 2, S07°01.243', E108°08.056', 1292 m, 19-IV-2012; 2 exx, sample 4, S07°01.734', E108°08.059', 1530 m, 19-IV-2012; 4 exx, sample 5, S07°01.804', E108°08.044', 1563 m, 20-IV-2012.
W-Java Prov. (Mt. Cakrabuana). Elevation: 1292–1563 m.
This epithet is based on the Latin adjective silvestris (living in the forest) and refers to the species habitat.
Trigonopterus silvestris Riedel, sp. n. was coded as “Trigonopterus sp. 373”.
Holotype, male (Fig.
Holotype (MZB): ARC2547 (EMBL # LM655884), W-Kalimantan Prov., Singkawang, Mt. Poteng, sample 3, N00°51.211', E109°03.418', 651 m, 13-XII-2011. Paratypes (SMNK): 1 ex, ARC2548 (EMBL # LM655885), same data as holotype.
W-Kalimantan Prov. (Singkawang). Elevation: 651 m.
This epithet is based on the type locality.
Trigonopterus singkawangensis Riedel, sp. n. was coded as “Trigonopterus sp. 367”.
Holotype, male (Fig.
Holotype (MZB): ARC0281 (EMBL # LM655471), East Sumatra, Lampung Prov., Kalianda, Mt. Rajabasa, sample 1, S05°46.896', E105°37.687', 1281 m, 15-VIII-2006. Paratypes (MZB, SMNK): E-Sumatra, Lampung Prov., Kalianda, Mt. Rajabasa: 1 ex, ARC0282 (EMBL # LM655472), same data as holotype; 2 exx, ARC2746 (EMBL # LM656002), ARC2747 (EMBL # LM656003), sample 1, S05°46.896', E105°37.687', 1281 m, 07-V-2012; 1 ex, sample 6, S05°46.651', E105°37.574', 1257 m, 07-V-2012.
Lampung Prov. (Mt. Rajabasa). Elevation: 1257–1281 m.
This epithet is based on the Latin adjective singularis (alone) and refers to the fact that no other Trigonopterus species could be found on the type locality.
Trigonopterus singularis Riedel, sp. n. was coded as “Trigonopterus sp. 321”.
Holotype, male (Fig.
Holotype (MZB): ARC1510 (EMBL # LM655608), West Nusa Tenggara Prov., Sumbawa, Batu Dulang, Mt. Batu Pasak, sample 2, S08°37.028', E117°15.783', 1305 m, 12-IV-2010. Paratypes (MZB, SMNK, ZSM): West Nusa Tenggara Prov., Sumbawa: 13 exx, ARC1509 (EMBL # LM655607), ARC1511 (EMBL # LM655609), ARC1512 (EMBL # LM655610), same data as holotype; 1 ex, Batu Dulang, Mt. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 12-IV-2010; 1 ex, Batu Dulang, Mt. Batu Pasak, sample 3, S08°37.524', E117°15.423', 1385 m, 18-IV-2010; 5 exx, ARC1493 (EMBL # LM655591), ARC1494 (EMBL # LM655592), ARC1495 (EMBL # LM655593), ARC1496 (EMBL # LM655594), Tepal, Pc. Nengas, sample 2, S08°35.884', E117°08.384', 1310 m, 15-IV-2010; 2 exx, Tepal, Pc. Nengas, sample 3, S08°35.386', E117°08.251', 1415 m, 15-IV-2010; 7 exx, Tepal, Pc. Nengas, sample 5, S08°35.740', E117°08.721', 1330 m, 16-IV-2010; 1 ex, Tepal, Pc. Nengas, sample 6, S08°35.533', E117°08.605', 1350 m, 16-IV-2010; 2 exx, Tepal, Pc. Nengas, sample 7, S08°35.176', E117°08.295', 1490 m, 16-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang, Tepal). Elevation: 1305–1490 m.
This epithet is based on the Latin adjective sinuatus (bent) and refers to the elytral outline.
Trigonopterus sinuatus Riedel, sp. n. was coded as “Trigonopterus sp. 333” by
Holotype, male (Fig.
Holotype (MZB): ARC2737 (EMBL # LM655993), East Sumatra, Lampung Prov., Gisting, Sidokaton, Mt. Tanggamus, sample 1, S05°25.475', E104°41.063', 1539 m, 03-V-2012. Paratypes (MZB, SMNK, ZSM): E-Sumatra, Lampung Prov., Gisting, Sidokaton, Mt. Tanggamus: 2 exx, ARC2741 (EMBL # LM655997), ARC2742 (EMBL # LM655998), sample 2, S05°25.491', E104°41.006', 1636 m, 03-V-2012; 4 exx, ARC2744 (EMBL # LM656000), sample 3, S05°25.508', E104°40.882', 1759 m, 03-V-2012; 3 exx, sample 4, S05°25.484', E104°41.027', 1623 m, sifted.
Lampung Prov. (Mt. Tanggamus). Elevation: 1539–1759 m.
This epithet is based on the Latin adjective squalidus (rough, dirty) and refers to the rough integument often covered with dirt incrustations.
Trigonopterus squalidus Riedel, sp. n. was coded as “Trigonopterus sp. 371”.
Holotype, male (Fig.
Holotype (MZB): ARC2743 (EMBL # LM655999), East Sumatra, Lampung Prov., Gisting, Sidokaton, Mt. Tanggamus, sample 2, S05°25.491', E104°41.006', 1636 m, 03-V-2012. Paratypes (MZB, SMNK, ZSM): E-Sumatra, Lampung Prov.: 23 exx, ARC0275 (EMBL # LM655466), ARC0276 (EMBL # LM655467), ARC0277 (EMBL # LM655468), ARC0278 (EMBL # LM655469), ARC0279 (EMBL # LM655470), ARC0393, Sumberjaya, Bodongjaya, Mt. Rigis, sample 3, S05°03.140', E104°26.912', 1351 m, 12-VIII-2006; 27 exx, ARC2738 (EMBL # LM655994), ARC2739 (EMBL # LM655995), ARC2740 (EMBL # LM655996), Gisting, Sidokaton, Mt. Tanggamus, sample 1, S05°25.475', E104°41.063', 1539 m, 03-V-2012; 5 exx, Gisting, Sidokaton, Mt. Tanggamus, sample 2, S05°25.491', E104°41.006', 1636 m, 03-V-2012; 8 exx, ARC2745 (EMBL # LM656001), Gisting, Sidokaton, Mt. Tanggamus, sample 4, S05°25.484', E104°41.027', 1632 m, 03-V-2012.
Lampung Prov. (Sumberjaya, Mt. Tanggamus). Elevation: 1351–1636 m.
This epithet is based on the name of Sumatra Island.
Trigonopterus sumatrensis Riedel, sp. n. was coded as “Trigonopterus sp. 300”.
Holotype, male (Fig.
Holotype (MZB): ARC3609 (EMBL # LM656039), West Nusa Tenggara Prov., Sumbawa, Tepal, Pc. Nengas, sample 3, S08°35.386', E117°08.251', 1415 m, 15-IV-2010. Paratypes (MZB, SMNK, ZSM): Sumbawa: 2 exx, ARC3610 (EMBL # LM656040), ARC3611 (EMBL # LM656041), same data as holotype; 4 exx, ARC1533 (EMBL # LM655631), ARC2339 (EMBL # LM655789), ARC2341 (EMBL # LM655791), ARC2343 (EMBL # LM655793), Tepal, Pc. Nengas, sample 7, S08°35.176', E117°08.295', 1490 m, 16-IV-2010; 2 exx, ARC3606 (EMBL # LM656036), ARC3608 (EMBL # LM656038), Tepal, Pc. Nengas, sample 2, S08°35.884', E117°08.384', 1310 m, 15-IV-2010; 2 exx, ARC1516 (EMBL # LM655614), ARC1517(EMBL # LM655615), Batu Dulang, Mt. Batu Pasak, sample 2, S08°37.028', E117°15.783', 1305 m, 12-IV-2010; 3 exx, ARC3653 (EMBL # LM656071), ARC3654 (EMBL # LM656072), ARC3655 (EMBL # LM656073), Batu Dulang, Mt. Batu Pasak, sample 4, S08°37.318', E117°15.339', 1280 m, 18-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Batu Dulang, Tepal). Elevation: 1280–1490 m.
This epithet is based on the island of Sumbawa.
Trigonopterus sumbawensis Riedel, sp. n. was coded as “Trigonopterus sp. 326” by
Holotype, male (Fig.
Holotype (MZB): ARC1536 (EMBL # LM655634), Banten Prov., Pandegelang, Mt. Karang, sample 3, S06°17.110', E106°03.263', 1145 m, 20-III-2010. Paratypes (ARC, MZB, SMNK, ZSM): 5 exx, ARC0102, W-Java Prov., Mt. Halimun N.P., Citalahab, S06°44.503', E106°31.550', 1150 m, 20-21-IX-2004; 9 exx, W-Java Prov., Mt. Halimun N.P., Citalahab, sample 1A, S06°44.503', E106°31.55', 1200 m, 11-IX-2005; 5 exx, W-Java Prov., Mt. Halimun N.P., Citalahab, sample 1, S06°44.423', E106°31.738', 1100 m, 12-IX-2005; 29 exx, ARC0208 (EMBL # LM655430), ARC0209 (EMBL # LM655431), ARC0210 (EMBL # LM655432), ARC0392, W-Java Prov., Mt. Halimun N.P., Citalahab, sample 2, S06°44.503', E106°31.550', 1200 m, 12-IX-2005; 9 exx, W-Java Prov., N Ciptarasa, sample 2, S06°50.045', E106°30.218', 1050 m, 19-IX-2005; 6 exx, ARC0189 (EMBL # LM655413), W-Java Prov., N Ciptarasa, sample 3, S06°49.867', E106°30.085', 1100 m, 19-IX-2005; 3 exx, W-Java Prov., N Ciptarasa, sample 4, S06°49.802', E106°30.020', 1150 m, 19-IX-2005; 3 exx, ARC1537 (EMBL # LM655635), ARC1539 (EMBL # LM655637), same data as holotype; 1 ex, Banten Prov., Pandegelang, Mt. Karang, sample 2, S06°17.266', E106°03.262', 1025 m, 20-III-2010; 1 ex, Banten Prov., Pandegelang, Mt. Karang, sample 3, S06°17.110', E106°03.263', 1145 m, 26-IV-2010; 1 ex, ARC1538 (EMBL # LM655636), Banten Prov., Pandegelang, Mt. Karang, sample 8, 1100 m, 26-IV-2010.
Banten Prov. (Mt. Karang), W-Java Prov. (Mt. Halimun N.P.). Elevation: 1100–1200 m.
This epithet is based on the type locality which is situated just east of the Sunda Strait; it is also part of the Sunda region of Java.
Trigonopterus sundaicus Riedel, sp. n. was coded as “Trigonopterus sp. 296”.
Holotype, male (Fig.
Holotype (MZB): ARC2652 (EMBL # LM655910), West Java Prov., Bogor, Cidahu, Mt. Salak (near Javana Spa), sample 3, S06°43.425', E106°43.227', 1756 m, 13-IV-2012. Paratypes (ARC, MZB, SMNK, ZSM): W-Java Prov.: 6 exx, ARC0104, ARC0184 (EMBL # LM655409), Mt. Halimun N.P., Pasir Banteng, Mt. Botol, 1550 m, 17-XI-2004; 3 exx, Mt. Halimun N.P., Citalahab, Mt. Kendeng, 1550 m, 19-XI-2004; 3 exx, ARC2648 (EMBL # LM655906), ARC2649 (EMBL # LM655907), Bogor, Cidahu, Mt. Salak (near Javana Spa), sample 1, S06°43.733', E106°42.711', 1429 m, 13-IV-2012; 5 exx, same data as holotype; 1 ex, ARC2653 (EMBL # LM655911), Sukabumi, Mt. Gede, Cisaat, Situ Gunung, sample 1, S06°49.377', E106°55.729', 1281 m, 15-IV-2012; 1 ex, ARC2663 (EMBL # LM655921), Sukabumi, Mt. Gede, Cisaat, Situ Gunung, sample 5, S06°47.912', E106°56.457', 1940 m, 16-IV-2012; 1 ex, Sukabumi, Mt. Gede, Cisaat, Situ Gunung, sample 4, S06°48.250', E106°56.271', 1720 m, 16-IV-2012; 4 exx, ARC2666 (EMBL # LM655924), Sukabumi, Mt. Gede, Cisaat, Situ Gunung, sample 7, S06°48.377', E106°56.183', 1637 m, 16-IV-2012, under Lithocarpus sp.; 1 exx, ARC0159, Garut, Kawah Kamojang, sample 2, S07°09.578', E107°47.802', 1400 m, 26-IX-2005; 2 exx, ARC0182 (EMBL # LM655407), Garut, Kawah Kamojang, sample 3, S07°09.497', E107°47.828', 1430 m, 26-IX-2005; 4 exx, ARC0256 (EMBL # LM655448), ARC0257 (EMBL # LM655449), ARC0258 (EMBL # LM655450), Telaga Warna, between Puncak and Cipanas, sample 1, S06°42.253', E106°59.755', 1585 m, 06-VIII-2006; 11 exx Telaga Warna, between Puncak and Cipanas, sample 2, S06°42.097', E106°59.833', 1477 m, 06-VIII-2006.
W-Java Prov., (Mt. Gede, Mt. Halimun N.P., Kawah Kamojang, Mt. Salak). Elevation: 1400–1940 m.
This epithet is based on the Latin adjective suturalis and refers to the basally swollen elytral suture.
Trigonopterus suturalis Riedel, sp. n. was coded as “Trigonopterus sp. 304”.
Holotype, male (Fig.
Holotype (MZB): ARC2195 (EMBL # LM655651), East Nusa Tenggara Prov., Flores, Ruteng, Mt. Ranaka, sample 1, S08°37.321', E120°31.463', 1535 m, 08-III-2011.
East Nusa Tenggara Prov., Flores (Ruteng). Elevation: 1535 m.
This epithet is based on the weevil genus Syarbis to which it bears some superficial resemblance e.g. in the reduction of the onychium. It is to be treated as a noun in apposition.
Trigonopterus syarbis Riedel, sp. n. was coded as “Trigonopterus sp. 350”.
Holotype, male (Fig.
Holotype (MZB): ARC2312 (EMBL # LM655765), Bali, Busungbiu – Pupuan, Telaga, Primatani reserve, sample 2, S08°17.304', E114°56.635', 625 m, 02-IV-2011. Paratypes (MZB, SMNK, ZSM): Bali: 6 exx, ARC2313 (EMBL # LM655766), ARC2314 (EMBL # LM655767), ARC2956 (EMBL # LM656009), ARC2957 (EMBL # LM656010), ARC2958 (EMBL # LM656011), same data as holotype; 21 exx, ARC0723, ARC0724, ARC0725, ARC0726 (PCR failed), Busungbiu – Pupuan, S08°17.64', E114°56.72', 600–700 m, sifted.
Bali (Telaga). Elevation: 625 m.
This epithet is based on the type locality, Telaga.
Trigonopterus telagensis Riedel, sp. n. was coded as “Trigonopterus sp. 289” by
Holotype, male (Fig.
Holotype (MZB): ARC3588 (EMBL # LM656020), West Nusa Tenggara Prov., W-Sumbawa, Tepal, Pc. Nengas, sample 5, S08°35.740', E117°08.721', 1330 m, 16-IV-2010.
West Nusa Tenggara Prov., Sumbawa (Tepal). Elevation: 1330 m.
This epithet is based on the type locality.
Trigonopterus tepalensis Riedel, sp. n. was coded as “Trigonopterus sp. 439”.
Holotype, male (Fig.
Holotype (MZB): ARC2243 (EMBL # LM655698), East Nusa Tenggara Prov., Flores Isl., Labuhan Bajo, Tebedo, sample 1, S08°29.848', E119°59.633', 525 m, 14-III-2011. Paratypes (SMNK): Flores Isl., Labuhan Bajo: 1 ex, ARC2225 (EMBL # LM655681), Roe, sample 5, S08°35.395', E120°00.383', 790 m, 13-III-2011.
East Nusa Tenggara Prov., Flores (Labuhan Bajo). Elevation: 525–790 m.
This epithet is based on the Indonesian word for “three” and is treated as a noun in apposition.
Trigonopterus tiga Riedel, sp. n. was coded as “Trigonopterus sp. 291” by
Holotype, male (Fig.
Holotype (MZB): ARC2531 (EMBL # LM655868), W-Kalimantan Prov., Bengkayan, Suka-Bangun, Mt. Bawang, sample 2, N00°53.514', E109°22.301', 275 m, 10-XII-2011. Paratypes (MZB, SMNK, ZSM): W-Kalimantan Prov., Bengkayan, Suka-Bangun, Mt. Bawang: 7 exx, ARC2530 (EMBL # LM655867), ARC2532 (EMBL # LM655869), ARC2533 (EMBL # LM655870), same data as holotype; 3 exx, sample 3, N00°53.621', E109°22.475', 400 m, 10-XII-2011; 1 ex, sample 6, N00°53.992', E109°22.502', 556 m, 11-XII-2011; 6 exx, ARC2544 (EMBL # LM655881), sample 7, N00°53.921', E109°22.444', 515 m, 11-XII-2011; 1 ex, ARC2545 (EMBL # LM655882), sample 8, N00°53.736', E109°22.316', 411 m, 11-XII-2011.
W-Kalimantan Prov. (Mt. Bawang). Elevation: 275–556 m.
This epithet is based on the generic name, which is describing the triangular shape of the elytra very well. It is to be treated as a noun in apposition.
Trigonopterus trigonopterus Riedel, sp. n. was coded as “Trigonopterus sp. 363” by
Holotype, male (Fig.
Holotype (MZB): ARC2231 (EMBL # LM655687), East Nusa Tenggara Prov., Flores Isl., Labuhan Bajo, Roe, sample 3, S08°36.259', E120°01.539', 975 m, 13-III-2011 (MZB). Paratypes (MZB, SMNK): 4 exx, ARC2590 (EMBL # LM655899), same data as holotype.
East Nusa Tenggara Prov., Flores (Labuhan Bajo). Elevation: 975 m.
This epithet is based on the Indonesian word for “seven” and is treated as a noun in apposition.
Trigonopterus tujuh Riedel, sp. n. was coded as “Trigonopterus sp. 325” by
Holotype, male (Fig.
Holotype (MZB): ARC1540 (EMBL # LM655638), Java, Banten Prov., Ujung Kulon N.P., Tama Jaya, Mt. Honje, sample 5, S06°46.241', E105°31.542', 260 m, 24-IV-2010 (MZB). Paratypes (MZB, SMNK, ZSM): Java, Banten Prov., Ujung Kulon N.P., Tama Jaya, Mt. Honje: 35 exx, ARC1541 (EMBL # LM655639), same data as holotype; 23 exx, ARC1545 (EMBL # LM655643), ARC1546 (EMBL # LM655644), sample 4, S06°46.141', E105°31.649', 395 m, 23-IV-2010; 55 exx, sample 4, S06°46.141', E105°31.649', 395 m, 24-IV-2010; 24 exx, sample 6, S06°46.206', E105°31.592', 305 m, 24-IV-2010.
Banten Prov. (Ujung Kulon N.P.). Elevation: 260–395 m.
This epithet is based on the type locality.
Trigonopterus ujungkulonensis Riedel, sp. n. was coded as “Trigonopterus sp. 318”.
Holotype, male (Fig.
Holotype (MZB): ARC0314 (EMBL # LM655504), West Java Prov., SW Bandung, Ciwidey, Mt. Patuha, sample 2, S07°09.699', E107°24.377', 2100 m, 06-IX-2006. Paratypes (MZB, ARC, SMNK, ZSM): W-Java Prov.: 2 exx, ARC0312 (EMBL # LM655502), ARC0313 (EMBL # LM655503), same data as holotype; 5 exx, ARC0315 (EMBL # LM655505), ARC0316 (EMBL # LM655506), SW Bandung, Ciwidey, Mt. Patuha, sample 3, S07°09.336', E107°24.260', 2015 m, 06-IX-2006; 6 exx, ARC0004, Ciwidey, Gambung, Mt. Tilu, 1900 m, 12-VI-2002; 13 exx, ARC0169, ARC0214 (EMBL # LM655436), ARC0215 (EMBL # LM655437), Cilawu, Mt. Cikuray, sample 3, S07°19.075', E107°52.338', 2050 m, 24-IX-2005.
W-Java Prov., (Mt. Cikuray, Mt. Patuha, Mt. Tilu). Elevation: 1900–2100 m.
This epithet is based on the Latin adjective variolosus (scarred) and refers to the species´ rough sculpture.
Trigonopterus variolosus Riedel, sp. n. was coded as “Trigonopterus sp. 302”.
Holotype, male (Fig.
Holotype (MZB): ARC2514 (EMBL # LM655851), West Java Prov., Bandung, Lembang, Pangli, Mt. Bukittinggul, sample 1, S06°48.810', E107°44.216', 1753 m, 03-XII-2011. Paratypes (MZB, SMNK, ZSM): 2 exx, ARC0254 (EMBL # LM655446), ARC0255 (EMBL # LM655447), W-Java Prov., Telaga Warna, between Puncak and Cipanas, S06°42.253', E106°59.755', sample 1, 1584 m, 06-VIII-2006; 1 ex, ARC2662 (EMBL # LM655920), W-Java Prov., Sukabumi, Mt. Gede, Cisaat, Situ Gunung, sample 3, S06°48.628', E106°56.077', 1462 m, 16-IV-2012; 1 ex, ARC2667 (EMBL # LM655925), W-Java Prov., Sukabumi, Mt. Gede, Cisaat, Situ Gunung, sample 7, under Lithocarpus sp., S06°48.377', E106°56.183', 1637 m, 16-IV-2012; 3 exx, ARC2515 (EMBL # LM655852), same data as holotype; 4 exx, ARC2518 (EMBL # LM655855), ARC2519 (EMBL # LM655856), W-Java Prov., Bandung, Lembang, Pangli, Mt. Bukittinggul, sample 2, S06°48.845', E107°44.074', 1898 m, 03-XII-2011; 7 exx, ARC2706 (EMBL # LM655962), ARC2707 (EMBL # LM655963), ARC2708 (EMBL # LM655964), W-Java Prov., Garut, Cikajang, Mt. Payung, sample 2, S07°25.268', E107°48.492', 1250 m, 26-IV-2012; 6 exx, ARC2712 (EMBL # LM655968), W-Java Prov., Garut, Cikajang, Mt. Payung, sample 3, S07°25.320', E107°48.259', 1560 m, 26-IV-2012; 3 exx, ARC0205 (EMBL # LM655427), ARC0206 (EMBL # LM655428), ARC0207 (EMBL # LM655429), W-Java Prov., Garut, Cilawu, Mt. Cikuray, sample 1, S07°18.578', E107°52.622', 1648 m, 24-IX-2005; 3 exx, ARC0161, ARC0187 (EMBL # LM655412), W-Java Prov., Garut, Cilawu, Mt. Cikuray, sample 2, S07°18.840', E107°52.512', 1800 m, 24-IX-2005; 7 exx, ARC2720 (EMBL # LM655976), ARC2721 (EMBL # LM655977), ARC2722 (EMBL # LM655978), W-Java Prov., Garut, Wanaraja, Talagabodas, sample 2, S07°11.966', E108°03.984', 1719 m, 27-IV-2012; 1 ex, ARC2723 (EMBL # LM655979), W-Java Prov., Garut, Wanaraja, Talagabodas, sample 1, S07°12.511', E108°03.554', 1741 m, 27-IV-2012; 6 exx, ARC0162, ARC0173, ARC0199 (EMBL # LM655421), ARC0200 (EMBL # LM655422), W-Java Prov., Ciamis, Mt. Sawal, Batu Cakra, S07°14.920', E108°15.762', 990 m, 01-X-2005; 1 ex, W-Java Prov., Ciamis, Mt. Sawal, Blok Cireong, sample 2, S07°14.127', E108°15.568', 1120 m, 01-X-2005; 9 exx, ARC0293 (EMBL # LM655483), ARC0294 (EMBL # LM655484), ARC0295 (EMBL # LM655485), C-Java Prov., N slopes of Dieng figau, Petungkriyono, mountain N Tinalum, sample 1, S07°06.418', E109°44.514', 1115 m, 22-VIII-2006; 6 exx, ARC0298 (EMBL # LM655488), ARC0299 (EMBL # LM655489), ARC0300 (EMBL # LM655490), C-Java Prov., N slopes of Dieng figau, Petungkriyono, Mt. Deles, sample 3, S07°08.225', E109°43.555', 1495 m, 24-VIII-2006; 7 exx, C-Java Prov., N slopes of Dieng figau, Petungkriyono, Mt. Deles, sample 1, S07°08.221', E109°43.599', 1505 m, 24-VIII-2006; 2 exx, C-Java, Mt. Slamet, Guci, sample 5, S07°11.953', E109°10.497', 1620 m, 27-XI-2011; 6 exx, ARC2499 (EMBL # LM655836), ARC2500 (EMBL # LM655837), Central Java Prov., Mt. Slamet, Guci, sample 6, S07°11.983', E109°10.556', 1671 m, 27-XI-2011; 9 exx, ARC2501 (EMBL # LM655838), ARC2502 (EMBL # LM655839), C-Java Prov., Mt. Slamet, Guci, sample 4, S07°12.304', E109°10.450', 1406 m, 27-XI-2011.
W-Java Prov. (Mt. Gede N.P., Mt. Bukittinggul, Mt. Cikuray, Mt. Payung, Talagabodas, Mt. Sawal), C-Java Prov. (Dieng, Mt. Slamet). Elevation: 990–1898 m.
This epithet is based on the Latin adjective vulcanicus and refers to its presence on volcanic mountains.
Trigonopterus vulcanicus Riedel, sp. n. was coded as “Trigonopterus sp. 313”. This species is divided into three populations that exhibit slightly different shapes of the penis; however, they hardly justify recognition as separate species. The western population is found at Mt. Gede; the central population at Mt. Bukittinggul, Mt. Cikuray, Mt. Payung and Talagabodas; and the Eastern population at Mt. Sawal, Dieng, and Mt. Slamet.
Holotype, male (Fig.
Holotype (SMNK): ARC2525 (EMBL # LM655862), MALAYSIA, Sarawak, Kuching, Mt. Santubong, sample 3, N01°43.884', E110°19.727', 496 m, 06-XII-2011. Paratypes (ARC, SMNK): MALAYSIA, Sarawak, Kuching: 2 exx, ARC2526 (EMBL # LM655863), same data as holotype; 1 ex, ARC2529 (EMBL # LM655866), Mt. Santubong, sample 4, N01°43.884', E110°19.715', 487 m, 06-XII-2011; 2 exx, sample 5, N01°43.843', E110°19.672', 478 m, 06-XII-2011; 2 exx, sample 6, N01°43.830', E110°19.601', 375 m, 06-XII-2011; 1 ex, Mt. Santubong, 700 m, 25-III-1990; 2 exx, Bako N.P., 27-III-1990.
Sarawak (Santubong). Elevation: 487–700 m.
This species is named in honour of Alfred Russel Wallace (1823–1913) for laying the foundation of research on the Natural History of the Indo-Australian Archipelago. He spent some time at this species´ type locality where he prepared the draft of his “Sarawak Law”.
Trigonopterus wallacei Riedel, sp. n. was coded as “Trigonopterus sp. 362” by
1 | Metafemur subapically simple, without stridulatory patch. Length 1.90–2.65 mm. (subgenus Mimidotasia) | 2 |
1' | Metafemur subapically with stridulatory patch. Length 2.35–3.80 mm | 9 |
2 (1) | Elytra (Figs |
3 |
2' | Elytra (e.g. Figs |
4 |
3 (2) | Sutural interval of elytra swollen subapically, projecting in dorsal aspect. Penis (Fig. |
T. costipennis Riedel, sp. n. |
3' | Elytral apex with sutural interval costate but not distinctly projecting in dorsal aspect. Penis (Fig. |
T. misellus Riedel, sp. n. |
4 (2') | Elytral striae with rows of elongate-claviform, rather slender scales | 5 |
4' | Elytral striae with rows of claviform, wider scales | 6 |
5 (4) | Elytral intervals costate-tuberculate. Penis (Fig. |
T. diengensis Riedel, sp. n. |
5' | Elytral intervals costate. Penis (Fig. |
T. echinatus Riedel, sp. n. |
6 (4') | Elytral intervals with relatively low costae. Penis (Figs |
7 |
6' | Elytral intervals with costae more distinct. Penis (Figs |
8 |
7 (6) | Color largely ferruginous. Penis (Fig. |
T. honjensi s Riedel, sp. n. |
7' | Color largely black. Penis (Fig. |
T. seriatus Riedel, sp. n. |
8 (6') | Penis with endophallic structures as in Fig. |
T. satu Riedel, sp. n. |
8' | Penis with endophallic structures as in Fig. |
T. foveatus Riedel, sp. n. |
9 (1') | Elytra with striae deeply impressed; intervals costate to carinate. Femora each with small tooth. Penis (Fig. |
T. relictus Riedel, sp. n. |
9' | If elytral intervals costate, at least profemur edentate. Penis different, without pair of subapical blade-like processes | 10 |
10 | Anteroventral ridge of femora forming blunt tooth, especially in meso- and metafemur | 11 |
10' | All femora without tooth, or if species from Sumatra, meso- and metafemur with small denticle | 16 |
11 (10) | Sutural interval of elytra at apex markedly swollen, forming pair of distinct apical protrusions. Species from W-Java | 12 |
11' | Sutural interval of elytra at apex simple or weakly swollen. Species from E-Java | 14 |
12 (11) | Sutural interval of elytra at apex forming median, apically bifid protrusion. Penis (Fig. |
T. vulcanicus Riedel, sp. n. |
12' | Sutural interval of elytra at apex markedly swollen, forming pair of rounded apical protrusions. Penis (Figs |
13 |
13 (12') | Transfer apparatus of penis (Fig. |
T. dimorphus Riedel, sp. n. |
13' | Transfer apparatus of penis (Fig. |
T. binodulus Riedel, sp. n. |
14 (11') | Disk of pronotum (Fig. |
T. merubetirensis Riedel, sp. n. |
14' | Punctation of pronotum more coarsely, forming more or less distinct pattern (Figs |
15 |
15 (14') | Disk of pronotum (Fig. |
T. ijensis Riedel, sp. n. |
15' | Disk of pronotum (Fig. |
T. argopurensis Riedel, sp. n. |
16 (10') | Elytral intervals each with secondary rows of punctures resembling striae; at middle with transverse subglabrous band | T. alaspurwensis Riedel, sp. n. |
16' | Elytra with sculpture different; if sculpture coarse, never with transverse subglabrous band at middle | 17 |
17 (16') | Elytra with striae indistinct, marked by fine lines and rows of small punctures; intervals flat, more or less coriaceous | 18 |
17' | Elytra with striae deeply impressed and / or intervals elevated | 20 |
18 (17) | Tarsomere 3 of protarsus 2.8–3.0 × as wide as tarsomere 1, markedly larger than of mesotarsus, (Fig. |
T. latipes Riedel, sp. n. |
18' | Tarsomere 3 of protarsus 2.4–2.7 × as wide as tarsomere 1, at most slightly larger than of mesotarsus (Figs |
19 |
19 (18') | Body (Fig. |
T. arjunensis Riedel, sp. n. |
19' | Body (Fig. |
T. acuminatus Riedel, sp. n. |
20 (17') | Pronotum laterally without distinct preapical constriction. Usually larger species, length of elytra+pronotum 2.05–3.44 mm | 21 |
20' | Pronotum laterally with distinct preapical constriction. Usually smaller species, length of elytra+pronotum 1.76–3.22 mm | 26 |
21 (20) | Pronotum in apical half slightly rounded apicad. Species from East Sumatra | 22 |
21' | Pronotum anterolaterally subangularly projecting. Species from Java | 23 |
22 (21) | Elytra 2.62–2.65 × as long as pronotum. Penis (Fig. |
T. singularis Riedel, sp. n. |
22' | Elytra 2.50–2.56 × as long as pronotum. Sides of penis (Fig. |
T. amphoralis Marshall |
23 (21') | Elytral intervals carinate; striae with rows of long erect piliform scales | 24 |
23' | Elytral intervals costate; striae with rows of shorter subrecumbent scales | 25 |
24 (23) | Rostrum apically without distinct median denticle. Apex of penis (Fig. |
T. silvestris Riedel, sp. n. |
24' | Rostrum apically with distinct median denticle. Apex of penis (Fig. |
T. cahyoi Riedel, sp. n. |
25 (23') | Elytral apex (Fig. |
T. pangandaranensis Riedel, sp. n. |
25' | Elytral apex (Fig. |
T. ujungkulonensis Riedel, sp. n. |
26 (20') | Sides of pronotum behind preapical constriction rounded, without distinct angular projections | 27 |
26' | Sides of pronotum behind preapical constriction angularly projecting | 31 |
27 (26) | Sutural intervals of elytra extended into median apical tooth. Mt. Gede | T. gedensis Riedel, sp. n. |
27' | Sutural intervals of elytra at most swollen, but without distinct tooth | 28 |
28 (27') | Elytral intervals costate-tuberculate; striae each with sparse row of suberect bristles | 29 |
28' | Elytral intervals carinate; striae each with row of suberect piliform scales or without conspicuous pubescence | 30 |
29 (28) | Penis (Fig. |
T. asper Riedel, sp. n. |
29' | Penis (Fig. |
T. variolosus Riedel, sp. n. |
30 (28') | Elytral striae each with row of minute recumbent scales. E-Sumatra | T. lampungensis Riedel, sp. n. |
30' | Elytral striae with rows of suberect piliform scales. W-Java | T. setifer Riedel, sp. n. |
31 (26') | Elytral intervals irregularly costate-tuberculate; especially towards sides costae laterally corrugate | 32 |
31' | Elytral intervals costate-carinate | 36 |
32 (31) | Rostrum apically with denticle | 33 |
32' | Rostrum apically without denticle | 34 |
33 (32) | Metatibia ventrally with sparse row of long erect setae. Penis (Fig. |
T. halimunensis Riedel, sp. n. |
33' | Metatibia ventrally simple, without row of long setae. Penis (Fig. |
T. duabelas Riedel, sp. n. |
34 (32') | Elytral intervals partly flat, partly costate-tuberculate; interval 5 subbasally swollen. E-Sumatra | T. squalidus Riedel, sp. n. |
34' | Elytral intervals irregularly costate-tuberculate; interval 5 subbasally simple. W-Java | 35 |
35 (34') | Penis (Fig. |
T. allopatricus Riedel, sp. n. |
35' | Penis (Fig. |
T. variolosus Riedel, sp. n. |
36 (32') | Elytra subnude, with sparse small recumbent setae | 37 |
36' | Elytral striae each with row of suberect piliform scales | 38 |
37 (36) | Anterolateral angular projections of pronotum distinct. Elytra setae recumbent but distinct. Java | T. javensis Riedel, sp. n. |
37' | Anterolateral angular projections of pronotum indistinct. Elytral setae minute, easily overlooked. E-Sumatra | T. lampungensis Riedel, sp. n. |
38 (36') | Profemur edentate, meso- and metafemur ventrally with small denticle. E-Sumatra | T. sumatrensis Riedel, sp. n. |
38' | All femora edentate. Java | 39 |
39 (38') | Sutural interval of elytra basally swollen, widened, raised above level of other intervals | T. suturalis Riedel, sp. n. |
39' | Sutural interval of elytra surmounted by other intervals | 40 |
40 (39') | Rostrum subapically with median denticle. Abdominal ventrite 5 swollen, at middle with shallow impression | 41 |
40' | Rostrum subapically simple. Abdominal ventrite 5 flat or slightly swollen in apical half | 42 |
41 (40) | Profile of abdominal ventrite 2 projecting angularly. Penis (Fig. |
T. sundaicus Riedel, sp. n. |
41' | Profile of abdominal ventrite 2 simple. Penis (Fig. |
T. angulicollis Riedel, sp. n. |
42 (40') | Body (Fig. |
T. porcatus Riedel, sp. n. |
42' | Body (Fig. |
43 |
43 (42') | Penis (Fig. |
T. payungensis Riedel, sp. n. |
43' | Penis (Fig. |
T. rugosostriatus Riedel, sp. n. |
1 | Body in dorsal aspect with marked constriction between pronotum and elytron. Pronotum in basal half with pair of sublateral, kidney-shaped impressions (Fig. |
T. baliensis Riedel, sp. n. |
1' | Body subovate, in dorsal aspect with weak constriction between pronotum and elytron. Pronotum without pair of distinct sublateral impressions | 2 |
2 | Profile with distinct constriction between pronotum and elytron. Pronotum (Figs |
T. rugosus Riedel, sp. n. and T. pararugosus Riedel, sp. n. |
2' | Profile dorsally convex. Pronotum coarsely punctate, reticulate | 3 |
3 (2') | Elytral intervals (Figs |
4 |
3' | Elytral intervals (Figs |
5 |
4 (3) | Median rod of transfer apparatus thick, straight, 2.0 × as long as body of penis (Fig. |
T. klatakanensis Riedel, sp. n. |
4' | Median rod of transfer apparatus flagelliform, forming almost full coil, ca 2.7 × as long as body of penis (Fig. |
T. mesehensis Riedel, sp. n. |
5 (3') | Median rod of male transfer apparatus thin, shorter (1.4 × as long as body of penis), projecting basad (Fig. |
T. batukarensis Riedel, sp. n. |
5' | Median rod of male transfer apparatus thick, longer (1.6–1.7 × as long as body of penis), projecting basad further than apicad (Figs |
6 |
6 (5') | Body of penis (Fig. |
T. kintamanensis Riedel, sp. n. |
6' | Body of penis (Fig. |
T. telagensis Riedel, sp. n. |
1 | Body in dorsal aspect with marked constriction between pronotum and elytron (e.g. Figs |
2 |
1' | Body subovate, in dorsal aspect with weak constriction between pronotum and elytron (e.g. Figs |
20 |
2 (1) | Length 1.44–2.20 mm. Pronotum in basal half with pair of sublateral impressions. Coloration of body dark ferruginous to black | 3 |
2' | Length 2.20–4.04 mm. Pronotum without pair of distinct impressions. Coloration of body partly ferruginous, coppery, bronze, or black | 11 |
3 (2) | Flagelliform transfer apparatus (Fig. |
T. lombokensis Riedel, sp. n. |
3' | Flagelliform transfer apparatus (Figs |
4 |
4 (3') | Penis (Figs |
5 |
4' | Penis with weakly subangulate or rounded apex never extended medially | 6 |
5 (4) | Flagelliform transfer apparatus (Fig. |
T. sumbawensis Riedel, sp. n. |
5' | Flagelliform transfer apparatus (Fig. |
T. pseudosumbawensis Riedel, sp. n. |
6 (4') | Flagelliform transfer apparatus (Fig. |
T. roensis Riedel, sp. n. |
6' | Flagelliform transfer apparatus (Figs |
7 |
7 (6') | Flagelliform transfer apparatus ca. 3.0–3.3 × longer than body of penis; apodemes 2.4–2.5 × as long as body of penis. Sumbawa | 8 |
7' | Flagelliform transfer apparatus ca. 4.0–7.0 × longer than body of penis; apodemes 3.2–4.1 × as long as body of penis. Flores | 9 |
8 (7) | Penis (Fig. |
T. parasumbawensis Riedel, sp. n. |
8' | Penis (Fig. |
T. pauxillus Riedel, sp. n. |
9 (7') | Penis (Fig. |
T. paraflorensis Riedel, sp. n. |
9' | Penis (Figs |
10 |
10 (9') | Elytra more compact (Fig. |
T. florensis Riedel, sp. n. |
10' | Elytra more slender (Fig. |
T. pseudoflorensis Riedel, sp. n. |
11 (2') | Profemur with large subacute tooth. Color of elytra black, usually basal third ferruginous | T. dentipes Riedel, sp. n. |
11' | Profemur with blunt tooth or simple | 12 |
12 (11') | Color of elytra with bright metallic lustre, coppery | 13 |
12' | Color of elytra black, sometimes basally ferruginous; at most with bronze lustre, never bright metallic | 17 |
13 (12) | Onychium minute, drop-shaped, with median spine instead of pair of claws; weakly projecting beyond margin of tarsomere 3 | T. syarbis Riedel, sp. n. |
13' | Onychium distinctly projecting beyond margin of tarsomere 3, apically with pair of claws | 14 |
14 (13') | Elytra with humeri markedly swollen, laterally markedly projecting (Figs |
15 |
14' | Elytra with humeri simple (Fig. |
16 |
15 (14) | Elytra (Fig. |
T. fulgidus Riedel, sp. n. |
15' | Elytra (Fig. |
T. micans Riedel, sp. n. |
16 (14') | Elytra (Fig. |
T. cupreus Riedel, sp. n. |
16' | Elytra (Fig. |
T. cuprescens Riedel, sp. n. |
17 (12') | Body (Fig. |
T. serratifemur Riedel, sp. n. |
17' | Body (Figs |
18 |
18 (17') | Elytra (Fig. |
T. sinuatus Riedel, sp. n. |
18' | Elytra basally ferruginous; intervals 5 and 6 simple | 19 |
19 (18') | Elytral interval 5 in front of middle swollen, dorsally projecting. Penis (Fig. |
T. dacrycarpi Riedel, sp. n. |
19' | Elytral interval 5 simple. Penis (Fig. |
T. sasak Riedel, sp. n. |
20 (1') | species from Lombok | 21 |
20' | species from Sumbawa | 23 |
20' | species from Flores | 27 |
21 (20) | Elytral intervals flat, subglabrous. Pronotum longitudinally rugose | T. disruptus Riedel, sp. n. |
21' | Elytral intervals costate. Pronotum coarsely punctate | 22 |
22 (21') | Elytra black except sutural interval ferruginous; intervals 3 and 5 more distinctly swollen, with row of punctures | T. rinjaniensis Riedel, sp. n. |
22' | Elytra with bronze lustre; intervals 3 and 5 as other intervals, subglabrous | T. aeneomicans Riedel, sp. n. |
23 (20') | Elytra with intervals flat, striae marked by regular rows of punctures | T. punctatoseriatus Riedel, sp. n. |
23' | Elytra with striae deeply impressed, intervals costate | 24 |
24 (23') | Elytra with bronze lustre | T. aeneomicans Riedel, sp. n. |
24' | Elytra largely black, without distinct metallic lustre | 25 |
25 (24') | Sutural intervals of elytra costate, with row of punctures facing laterad. Abdominal ventrite 5 medially with shallow impression | T. sembilan Riedel, sp. n. |
25' | Sutural intervals of elytra forming pair of ridges, inner face with dense row of punctures. Abdominal ventrite 5 with median carina | 26 |
26 (25') | Elytral interval 3 with dense row of punctures. Penis as in Fig. |
T. saltator Riedel, sp. n. |
26' | Elytral interval 3 subglabrous, with few punctures. Penis as in Fig. |
T. tepalensis Riedel, sp. n. |
27 (20') | Tarsomere 3 of protarsus enlarged, medially deeply incised. Penis (Fig. |
T. fissitarsis Riedel, sp. n. |
27' | Tarsomere 3 of protarsus of similar shape as tarsomere 3 of mesotarsus. Penis without marked constriction | 28 |
28 (27') | Elytral intervals weakly costate, almost flat; striae consisting of rows of deeply impressed punctures | T. tujuh Riedel, sp. n. |
28' | Elytral intervals distinctly costate; striae incised | 29 |
29 (28') | Abdominal ventrite 5 with median carina, at least in basal third | 30 |
29' | Abdominal ventrite 5 without median carina | 32 |
30 (29) | Elytral interval black; suture broadly sunk-in. Abdominal ventrite 5 with median carina in basal third | T. delapan Riedel, sp. n. |
30' | Elytral interval ferruginous. Abdominal ventrite 5 with complete median carina | 31 |
31 (30') | Elytral intervals subglabrous, forming regular costae. Penis (Fig. |
T. tiga Riedel, sp. n. |
31' | Elytral intervals on dorsum punctate, along basal margin corrugate. Penis (Fig. |
T. ranakensis Riedel, sp. n. |
32 (29') | Metatibia apically with uncus and small premucro | 33 |
32' | Metatibia apically with uncus, without premucro | 34 |
33 (32) | Body (Fig. |
T. dua Riedel, sp. n. |
33' | Body (Fig. |
T. rutengensis Riedel, sp. n. |
34 (32') | Body (Fig. |
T. enam Riedel, sp. n. |
34' | Body wider (Figs |
35 |
35 (34') | Penis (Fig. |
T. lima Riedel, sp. n. |
35' | Penis (Fig. |
T. empat Riedel, sp. n. |
1 | Body subovate. Elytra with striae each bearing distinct row of suberect scales, intervals costate | 2 |
1' | Body subovate or subrhomboid. Elytra with striae simple, punctures containing small scale or seta; intervals flat. IF elytra with sparse rows of suberect scales and costate intervals, then body subrhomboid | 4 |
2 | Epistome with dorsoposteriad directed horn and rostrum at middle with dorsal protrusion | 3 |
2' | Epistome with denticle; rostrum with median and pair of submedian ridges, without dorsal protrusion between antennal insertions | T. palawanensis Riedel, sp. n. |
3 (2) | Length 1.52–1.79 mm. Elytral intervals narrow, weakly costate | T. wallacei Riedel, sp. n. |
3' | Length 2.32–2.45 mm. Elytral intervals subcarinate | T. schulzi Riedel, sp. n. |
4 (1') | Elytra with humeri swollen, laterally markedly projecting | 5 |
4' | Elytra with humeri simple, not markedly projecting | 6 |
5 (4) | Pronotum anteriorly simple. Elytral striae deeply impressed; intervals costate. Anteroventral ridge of meso- and metafemur simple | T. trigonopterus Riedel, sp. n. |
5' | Pronotum anteriorly with flanges projecting laterally. Elytral striae marked by small punctures; intervals flat. Anteroventral ridge of meso- and metafemur denticulate | T. sebelas Riedel, sp. n. |
6 (4') | Profemur in basal half posteroventrally with slightly bifid tooth. Male metatibia ventrally with dense fringe of long, stiff setae | T. santubongensis Riedel, sp. n. |
6' | Profemur in basal half posteroventrally without tooth. Male metatibia with sparse rows of setae | 7 |
7 (6') | Elytral apex with suture distinctly incised. Abdominal ventrite 2 in profile simple. | |
7' | Elytral apex with suture simple, not incised. Abdominal ventrite 2 in profile projecting dentiform | 9 |
8 (7) | Penis as in Fig. |
T. bornensis Riedel, sp. n. |
8' | Penis as in Fig. |
T. kalimantanensis Riedel, sp. n. |
9 (7') | In front of forehead median ridge of rostrum swollen, projecting subangularly from profile | 10 |
9' | Base of rostrum in profile simple, evenly confluent with forehead | |
10 (9) | Penis as in Fig. |
T. bawangensis Riedel, sp. n. |
10' | Penis as in Fig. |
T. singkawangensis Riedel, sp. n. |
11 (9') | Color largely black. Elytral intervals flat. Penis as in Fig. |
T. sepuluh Riedel, sp. n. |
11 | Color largely ferruginous. Elytral intervals weakly costate. Penis as in Fig. |
T. attenboroughi Riedel, sp. n. |
subgenus Mimidotasia Voss: T. costipennis Riedel, sp. n., T. diengensis Riedel, sp. n., T. echinatus Riedel, sp. n., T. foveatus Riedel, sp. n., T. honjensis Riedel, sp. n., T. misellus Riedel, sp. n., T. satu Riedel, sp. n., T. seriatus Riedel, sp. n.
T. attenboroughi-group: T. attenboroughi Riedel, sp. n., T. bawangensis Riedel, sp. n., T. santubongensis Riedel, sp. n., T. sebelas Riedel, sp. n., T. sepuluh Riedel, sp. n., T. singkawangensis Riedel, sp. n.
T. bornensis-group: T. bornensis Riedel, sp. n., T. kalimantanensis Riedel, sp. n.
T. dimorphus-group: T. acuminatus Riedel, sp. n., T. alaspurwensis Riedel, sp. n., T. allopatricus Riedel, sp. n., T. amphoralis Marshall, T. angulicollis Riedel, sp. n., T. argopurensis Riedel, sp. n., T. arjunensis Riedel, sp. n., T. asper Riedel, sp. n., T. binodulus Riedel, sp. n., T. cahyoi Riedel, sp. n., T. cuprescens Riedel, sp. n., T. cupreus Riedel, sp. n., T. dacrycarpi Riedel, sp. n., T. dentipes Riedel, sp. n., T. dimorphus Riedel, sp. n., T. duabelas Riedel, sp. n., T. fulgidus Riedel, sp. n., T. gedensis Riedel, sp. n., T. halimunensis Riedel, sp. n., T. ijensis Riedel, sp. n., T. javensis Riedel, sp. n., T. lampungensis Riedel, sp. n., T. latipes Riedel, sp. n., T. merubetirensis Riedel, sp. n., T. micans Riedel, sp. n., T. pangandaranensis Riedel, sp. n., T. payungensis Riedel, sp. n., T. porcatus Riedel, sp. n., T. rugosostriatus Riedel, sp. n., T. sasak Riedel, sp. n., T. serratifemur Riedel, sp. n., T. setifer Riedel, sp. n., T. silvestris Riedel, sp. n., T. singularis Riedel, sp. n., T. sinuatus Riedel, sp. n., T. squalidus Riedel, sp. n., T. sumatrensis Riedel, sp. n., T. sundaicus Riedel, sp. n., T. suturalis Riedel, sp. n., T. syarbis Riedel, sp. n., T. ujungkulonensis Riedel, sp. n., T. variolosus Riedel, sp. n., T. vulcanicus Riedel, sp. n.
T. politus-group: T. allotopus Riedel, sp. n.
T. relictus-group: T. baliensis Riedel, sp. n., T. florensis Riedel, sp. n., T. lombokensis Riedel, sp. n., T. paraflorensis Riedel, sp. n., T. parasumbawensis Riedel, sp. n., T. pauxillus Riedel, sp. n., T. pseudoflorensis Riedel, sp. n., T. pseudosumbawensis Riedel, sp. n., T. relictus Riedel, sp. n., T. roensis Riedel, sp. n., T. sumbawensis Riedel, sp. n.
T. saltator-group: T. aeneomicans Riedel, sp. n., T. batukarensis Riedel, sp. n., T. delapan Riedel, sp. n., T. disruptus Riedel, sp. n., T. dua Riedel, sp. n., T. empat Riedel, sp. n., T. enam Riedel, sp. n., T. fissitarsis Riedel, sp. n., T. kintamanensis Riedel, sp. n., T. klatakanensis Riedel, sp. n., T. lima Riedel, sp. n., T. mesehensis Riedel, sp. n., T. pararugosus Riedel, sp. n., T. punctatoseriatus Riedel, sp. n., T. ranakensis Riedel, sp. n., T. rinjaniensis Riedel, sp. n., T. rugosus Riedel, sp. n., T. rutengensis Riedel, sp. n., T. saltator Riedel, sp. n., T. sembilan Riedel, sp. n., T. telagensis Riedel, sp. n., T. tepalensis Riedel, sp. n., T. tiga Riedel, sp. n., T. tujuh Riedel, sp. n.
T. trigonopterus-group: T. trigonopterus Riedel, sp. n.
T. wallacei-group: T. palawanensis Riedel, sp. n., T. schulzi Riedel, sp. n., T. wallacei Riedel, sp. n.
Herein, we apply our fast track taxonomy approach (
At the start of this project and based on literature the high diversity of Trigonopterus in the Papuan region (
The field work for this study focused on eastern parts of Sumatra, Java and the Lesser Sunda Islands. Materials stored in museum collections neither contained Trigonopterus from the western parts of Sumatra, nor from the Malay Peninsula. A dedicated search in the field could possibly lead to interesting discoveries, but equally well the genus´ real area of distribution excludes this region or parts of it. More promising for future discoveries of species are Borneo and Palawan where only material from a few localities was available. Borneo obviously possesses a rich fauna of Trigonopterus and its species number will be magnitudes higher than the ten species recorded so far. Species discovery on Java (35 species recorded), Bali (8 species recorded), and Lombok (7 species recorded) may approach saturation, while additional species are likely to be found on Sumatra (7 species recorded), Sumbawa (12 species recorded), and Flores (18 species recorded).
Most localities sampled by us harbour at least one species of Trigonopterus; in some places many more were found, such as seven species each at Mount Sawal of West Java, and Mount Ranaka of Flores Island. Although these figures are dwarfed by comparable localities of New Guinea (
Wide areas of East Java and the Lesser Sunda Islands are markedly influenced by seasonal climate resulting in extensive monsoon forests, while wet evergreen forests persist on the slopes of higher mountains (
Looking at the distribution of Trigonopterus species in the Sunda Arc and the areas of remaining primary forests, it appears likely that some of these newly discovered species will become extinct in the next few decades. The blame will be largely on human activities which have reduced the once extensive areas of forest to small fragments. It is our responsibility to help preserving these remaining areas of habitat. We should intensify our efforts especially with those forests harbouring endemic species but nevertheless holding a relatively weak conservation status, as exemplified by the reserves of Mount Sawal and Mount Ranaka. In these cases, a limited investment in better protection could achieve a significant improvement for species conservation.
Herein, we provide faces to 98 hitherto unknown species by publishing their descriptions and images, in parallel also on the wiki-site Species-ID. The new names will be useful to deposit scientific data reliably and sustainably. Separate studies on the phylogeny and distribution patterns of Trigonopterus are in progress and will build on this taxonomic foundation. Most of all, it is hoped that knowledge of this previously unknown fauna will create an awareness for the conservation value of the remaining fragments of wet evergreen primary forest in the region. There may be a certain redundancy of conservation areas when we look at the distribution of many vertebrates – however, the larger portion of biodiversity as represented by arthropods exhibits a different picture. The message that a study of flightless weevils of the Sunda Arc conveys is clear: with almost every area of remaining primary forest being destroyed we will lose some more species – with or without names.
We thank LIPI (Indonesian Institute of Sciences), RISTEK (Ministry of State for Research and Technology, Indonesia) and the Indonesian Department of Forestry for providing relevant permits. The field work in Indonesia would not have been possible without the generous hospitality and help of many local people, and we thank all of them very warmly. Michi Schrödl (Munich) gave inspiration to use numbers as species epithets. We are grateful to Lyubomir Penev (Sofia) for his constant support during the publication of this study. We thank the German Research Foundation DFG (RI 1817/1-1, 3-1, 3-3) for financial support.