Lectotype ♀ (NHMB): [p] “Asia, China”, [p] “coll. Richard \ Hicker, Wien”, [h] “hickeri n. sp.”, [p] “Athemus \ hickeri (Pic) \ det. S. Kasantsev 1996”, [p] “LECTOTYPUS”, [p] “CANTHARIDAE \ CANTH00000915”.

China.

This species was described on the basis of female types, and its locality is not accurate within China. Here the abdominal sternite VIII (Fig. 34) of the female is illustrated for the first time: it is largely and roundly emarginated in middle and both sides of posterior margin, the portion between middle and lateral emarginations acute-angled at apex.

Holotype ♂ (IZAS): CHINA, Sichuan, Yajiang to 5km of Litang, 2595m, leg. Gan-Yan Yang. Paratypes: CHINA, Sichuan: 2♂♂ (IZAS): Yajiang, Hekou, Shanbeihou, 2838m, 27.V.2009, leg. Gan-Yan Yang; 1♂ (IZAS): Yajiang, 24.V.2009, leg. Feng Yuan; 1♂ (IZAS): Yajiang, Bajiaolou, 29.V.2009, leg. Zhi-Liang Wang [the above are all transliterated from Chinese labels]; 1♂ (NHMB): “CHINA, Sichuan prov., Mts W. Bamei, 3750m, 12.08.2005, leg. S. Murzin”.

China (Sichuan).

Male (Fig. 1). Body black, mandibles dark brown, pronotum red, with a large black marking in center of disc, elytra red, distinctly darkened almost along the whole length of the second elytral interstices, slightly darkened at the first elytral interstices.

Head evenly narrowed behind eyes, surface densely and finely punctate, semilustrous, covered with dense, fine, yellowish brown decumbent pubescence; eyes moderately protruding, head width across eyes moderately wider than anterior margin of pronotum; terminal maxillary palpomeres long-triangular, arcuate and sharp at apical one-third length of inner margins; antennae (Fig. 7) filiform, extending to apical one-third length of elytra, antennomeres II slightly longer than wide at apices, III–XI nearly parallel-sided, III about 2.5 times as long as wide, IV about one-third longer than III, XI slightly longer than X and pointed at apices, IV–XI each with a small round smooth impression at apical part of outer margin.

Pronotum nearly as long as wide, widest near posterior margin, anterior margin arcuate, anterior angles rounded, lateral margins slightly diverging posteriorly, posterior angles rectangular, posterior margin slightly arcuate and narrowly bordered, disc moderately convex at posterolateral parts, surface punctate and pubescent like that of head, semilustrous.

Elytra about 5.5 times longer than pronotum, 4.0 times as long as humeral width, which about one-third wider than posterior margin of pronotum, lateral margins nearly parallel, elytral venation slightly developed, not costate.

All tarsal claws simple.

Abdominal sternite IX long-triangular. Aedeagus (Figs 16–18): ventral process of each paramere normal and rounded at apex, distinctly narrowed at base and slightly bent dorsally in lateral view; dorsal fig slightly shorter than ventral process, abruptly narrowed at inner apical portion, with inner angle rounded, outer angle obtuse-angled, inner margin sinuate, apical margin rounded; laterophyse with apex pointed laterodorsally to outer angle of dorsal fig.

Female. Unknown.

Body length (males): 8.0–10.0 mm; width: 2.0–2.2 mm.

This specific name is derived from Latin gracilis (narrow) + cornu (horn), referring its antennomeres III–XI nearly parallel-sided.

This species is similar to L. hickeri Pic, but the antennae filiform, with antennomeres III–XI nearly parallel-sided; elytral venation less developed and the second elytral interstices darkened.

Holotype ♂ (MHBU): CHINA, Yunnan, Yunlong, Tianchi Nat. Res., 9.VII.2011, leg. Hao-Yu Liu. Paratype: 1♂ (MHBU): same data as holotype. [Both transliterated from Chinese labels].

China (Yunnan).

Male (Fig. 2). Body black, mandibles dark brown, pronotum red, with a large black marking in center of disc, elytra red, black almost along the whole length of the first and second elytral interstices, more or less darkened at the third and fourth elytral interstices.

Head evenly narrowed behind eyes, surface densely and finely punctate, semilustrous, covered with dense, fine, yellowish brown decumbent pubescence; eyes moderately protruding, head width across eyes moderately wider than anterior margin of pronotum; terminal maxillary palpomeres long-triangular, arcuate and sharp at apical one-third length of inner margins; antennae (Fig. 8) extending to apical one-third length of elytra, antennomeres II nearly as long as wide at apices, III–XI flattened, III–VIII obliquely widened apically, nearly long-triangular, III about twice as long as wide at apices, IV slightly longer than III, IX–XI nearly parallel-sided, XI slightly longer than X and pointed at apices, IV–XI each with an oval to round smooth impression at apical part of outer margin.

Pronotum nearly as long as wide, widest near posterior margin, anterior margin arcuate, anterior angles rounded, lateral margins slightly diverging posteriorly, posterior angles rectangular, posterior margin slightly arcuate and narrowly bordered, disc moderately convex at posterolateral parts, surface punctate and pubescent like that of head, semilustrous.

Elytra about 5.8 times longer than pronotum, 4.0 times longer than humeral width, which about one-third wider than posterior margin of pronotum, lateral margins nearly parallel, elytral venations moderately developed, slightly costate.

Pro- and meso-outer tarsal claws each with a basal projection.

Abdominal sternite IX long-triangular. Aedeagus (Figs 19–21): ventral process of each paramere normal and rounded at apex, distinctly narrowed at base and slightly bent dorsally in lateral view; dorsal fig distinctly shorter than ventral process, evenly narrowed apically, with inner angle widely rounded, outer angle obtuse-angled, inner margin nearly arcuate, apical margin rounded, around with long pubescence; laterophyse with apex pointed laterodorsally to outer angle of dorsal fig.

Female. Unknown.

Body length (males): 8.0–9.0 mm; width: 1.8–2.0 mm.

This specific name is derived from Latin longus (long) and hirtus (hairy), referring to its aedeagus: dorsal fig of each paramere covered with long pubescence along apical margin.

This species is similar to L. gracilicornis sp. n., but the antennomeres III–VIII widened apically, nearly long-triangular; elytral venation moderately developed, slightly costate; aedeagus: dorsal fig of each paramere evenly narrowed apically, inner margin arcuate, apical margin around with long pubescence.

Holotype ♂ (IZAS): CHINA, Sichuan, Mt. Emei, Xixiangchi, 1800–2000m, 12.VII.1957, leg. Zong-Yuan Wang. Paratypes: CHINA, Sichuan: 1♂ (IZAS): Mt. Emei, Jiulaodong, 1800–1900m, 26.VII.1957, leg. Ke-Ren Huang; 1♀ (IZAS): same locality, 31.VII.1957, leg. You-Cai Yu; 1♀ (IZAS): same locality, 30.VII. 1957, leg. You-Cai Yu; 1♀ (IZAS): same locality, 14.VIII.1957, leg. Zong-Yuan Wang [the above are all transliterated from Chinese labels]; 1♂ (NHMB): “CHINA, Sichuan prov., 70km West Chengdu, Qingcheng Hou Shan mts., 30°44'N, 103°08'E, 1500m, 8.–14.VI.2005, leg. S. Murzin”.

China (Sichuan).

Male (Fig. 3). Body black, mandibles dark brown, pronotum red, with a large dark brown marking, which almost extending to all margins of disc, elytra red, nearly black at the whole length of the first and second elytral interstices, more or less darkened at the third and fourth elytral interstices.

Head evenly narrowed behind eyes, surface densely and finely punctate, semilustrous, covered with dense, fine, yellowish brown decumbent pubescence; eyes strongly protruding, head width across eyes distinctly wider than anterior margin of pronotum; terminal maxillary palpomere long-triangular, nearly truncate and sharp at apical one-third length of inner margin; antennae (Fig. 10) extending to apical one-third length of elytra, antennomeres II nearly as long as wide at apices, III–XI flattened, III–VIII obliquely widened apically, nearly long-triangular, III about twice as long as wide at apices, IV slightly longer than III, IX–XI nearly parallel-sided, XI slightly longer than X and pointed at apices, IV–XI each with an oval to round smooth impression at apical part of outer margin.

Pronotum distinctly longer than wide, widest near posterior margin, anterior margin rounded, anterior angles rounded, lateral margins moderately diverging posteriorly, slightly sinuate at anterior portion, posterior angles rectangular, posterior margin slightly arcuate and narrowly bordered, disc moderately convex at posterolateral parts, surface punctate and pubescent like that of head, semilustrous.

Elytra about 4.7 times longer than pronotum, 3.7 times longer than humeral width, which about one-third wider than posterior margin of pronotum, lateral margins nearly parallel, elytral venations moderately developed, slightly costate.

Pro- and meso-outer tarsal claws each with a basal projection.

Abdominal sternite IX long-triangular. Aedeagus (Figs 22–24): ventral process of each paramere normal and rounded at apex, even and nearly straight in lateral view; dorsal fig slightly shorter than ventral process, evenly narrowed at inner apical portion and slightly widened at inner angle, with inner angle acute-angled, outer angle obtuse-angled, inner margin bisinuate, apical margin rounded; laterophyse with apex pointed laterodorsally to outer angle of dorsal fig.

Female. Similar to the male, but eyes less protruding; antennae (Fig. 9) shorter, extending to basal one-third length of elytra, antennomeres III about twice as long as wide at apices, IV–XI without impressions; pronotum nearly as long as wide, disc slightly convex on posterolateral parts; elytra with lateral margins slightly diverging posteriorly; abdominal sternite VIII (Fig. 35) roundly emarginated in middle and triangularly emarginated on both sides of posterior margin, the portion between middle and lateral emarginations obtuse-angled at apex; internal reproductive organ of genitalia see Fig. 39.

Body length (both sexes): 8.0–10.0 mm; width: 1.8–2.0 mm.

This specific name is derived from its locality, Sichuan Province, China.

This species is similar to L. hubeiensis sp. n., but differs from the latter by the antennae of female much narrower, antennomeres III about twice as long as wide at apices; aedeagus: ventral process of each paramere even and nearly straight in lateral view; abdominal sternite VIII of female with the portion between middle and lateral emarginations of posterior margin obtuse-angled at apex.

Holotype ♂ (MHBU): CHINA, Hubei, Dalaoling Nat. Res., 1200m, 9.VII.2011, leg. Xue-Song Guan. Paratypes: CHINA, Hubei: 1♀ (MHBU): Dalaoling Nat. Res., 1200m, 9.VII.2011, leg. Xiao-Long Yang; 1♀ (MHBU): same data, 10.VII.2011; 1♂ (MHBU): Badong, Lvcongpo, 1700m, 18.VII.2006, leg. Jun-Hua Wan; 1♀ (MHBU): Badong, Tiansanping, 1500m, 14.VII.2006, leg. Ping Hu; 1♀ (MHBU): Shennongjia, Bajiaomiao, 900–1300m, 17.VII.2003, leg. Yuan He; 1♀ (MHBU): same locality, 19.VII.2003, leg. Jun Ma; 1♂ (MHBU): Shennongjia, Wenshui Forestry, 1700–2000m, 20.VII.2003, leg. Hua He; 1♀ (MHBU): Wufeng, Houhe, 21.VII.2002, leg. Ying Shi; 1♀ (IZAS): Xingshan, Longmenhe, 1300m, 15.VI.1993, leg. Jian Yao; 1♀ (IZAS): same locality, 1350m, 18.VII.1993, leg. Xiao-Lin Chen; 1♀ (IZAS): same locality, 1400m, 22.VII.1993, leg. Shi-Mei Song; 1♀ (IZAS): same locality, 1670m, 23.VII.1993, leg. Xing-Ke Yang. [All are transliterated from Chinese labels].

China (Hubei).

Male (Fig. 4). Body black, mandibles dark brown, pronotum red, with a large dark brown marking, which almost extending to all margins of disc, elytra red, nearly black at the whole length of the first and second elytral interstices, more or less darkened at the third and fourth elytral interstices.

Head evenly narrowed behind eyes, surface densely and finely punctate, semilustrous, covered with dense, fine, yellowish brown decumbent pubescence; eyes strongly protruding, head width across eyes distinctly wider than anterior margin of pronotum; terminal maxillary palpomere long-triangular, nearly truncate and sharp at apical one-third length of inner margin; antennae (Fig. 12) extending to apical one-third length of elytra, antennomeres II nearly as long as wide at apices, III–XI flattened, III–IX obliquely widened apically, nearly long-triangular, III about twice as long as wide at apices, IV slightly longer than III, X–XI nearly parallel-sided, XI slightly longer than X and pointed at apices, IV–XI each with an oval to round smooth impression at apical part of outer margin.

Pronotum distinctly longer than wide, widest near posterior margin, anterior margin rounded, anterior angles rounded, lateral margins moderately diverging posteriorly, slightly sinuate at anterior portion, posterior angles rectangular, posterior margin slightly arcuate and narrowly bordered, disc moderately convex at posterolateral parts, surface punctate and pubescent like that of head, semilustrous.

Elytra about 5.3 times longer than pronotum, 3.7 times longer than humeral width, which about one-third wider than posterior margin of pronotum, lateral margins nearly parallel, elytral venations moderately developed, slightly costate.

Pro- and meso-outer tarsal claws each with a basal projection.

Abdominal sternite IX long-triangular. Aedeagus (Figs 25–27): ventral process of each paramere normal and rounded at apex, distinctly narrowed at base and slightly bent dorsally in lateral view; dorsal fig distinctly shorter than ventral process, evenly narrowed at inner apical portion and slightly widened at inner angle, with inner angle acute-angled, outer angle obtuse-angled, inner margin bisinuate, apical margin rounded; laterophyse with apex pointed laterodorsally to outer angle of dorsal fig.

Female. Similar to male, but eyes less protruding; antennae (Fig. 11) shorter, extending to elytral midlength, antennomeres III–XI distinctly widened, III about 1.6 times longer than wide at apices, IV–XI without impressions; pronotum nearly as long as wide, disc slightly convex on posterolateral parts; elytra with lateral margins slightly diverging posteriorly; abdominal sternite VIII (Fig. 36) largely and roundly emarginated in middle and both sides of posterior margin, the portion between middle and lateral emarginations rounded at apex; internal reproductive organ of genitalia see Fig. 40.

Body length (both sexes): 7.5–11.0 mm; width: 1.5–2.3 mm.

This specific name is derived from its locality, Hubei Province, China.

This species is similar to L. sichuanus sp. n. in the aedeagus, but differs from the latter by the antennae of female much wider, antennomeres III about 1.6 times longer than wide at apices; aedeagus: ventral process of each paramere narrowed at base and slightly bent dorsally in lateral view; abdominal sternite VIII of female with the portion between middle and lateral emarginations of posterior margin rounded at apex.

Holotype ♂ (ZIN): “Vietnam N, (Sa Pa), Lao Cai prov., 250km from Hanoi bearing 310#, Sa Pa vill. env., Hoang Lien Son Nat. Res., 1250-1300m, 15.-21.6.1998, leg. A. Napolov”.

Vietnam (Sa Pa).

Male (Fig. 5). Body black, mandibles dark brown, pronotum red, with a large black marking in center of disc, elytra red, nearly black at the whole length of the first and second elytral interstices.

Head evenly narrowed behind eyes, surface densely and finely punctate, semilustrous, covered with dense, fine, yellowish brown decumbent pubescence; eyes strongly protruding, head width across eyes distinctly wider than anterior margin of pronotum; terminal maxillary palpomere long-triangular, nearly truncate and sharp at apical one-third length of inner margin; antennae (Fig. 13) almost extending to apical one-fourth length of elytra, antennomeres II nearly as long as wide at apices, III–XI flattened, III–IX obliquely widened apically, nearly long-triangular, III about twice as long as wide at apices, IV slightly longer than III, X–XI nearly parallel-sided, XI slightly longer than X and pointed at apices, IV–XI each with a short narrow longitudinal smooth impression at apical part of outer margin.

Pronotum distinctly longer than wide, widest near posterior margin, anterior margin rounded, anterior angles rounded, lateral margins moderately diverging posteriorly, slightly sinuate at anterior portion, posterior angles rectangular, posterior margin slightly arcuate and narrowly bordered, disc moderately convex at posterolateral parts, surface punctate and pubescent like that of head, semilustrous.

Elytra about 5.3 times longer than pronotum, 4.0 times longer than humeral width, which about one-third wider than posterior margin of pronotum, lateral margins nearly parallel, elytral venations moderately developed, slightly costate.

All tarsal claws simple.

Abdominal sternite IX long-triangular. Aedeagus (Figs 28–30): ventral process of each paramere flattened, twist in middle and tapered at apex in ventral view, slightly narrowed at base and distinctly bent dorsally in lateral view; dorsal fig slightly shorter than ventral process, abruptly narrowed at inner apical portion, with a longitudinal ridge in middle of basal portion, membranous between inner margin and the ridge, inner angle rectangular, outer angle acute-angled and bent ventrally, inner margin distinctly protuberant in middle, apical margin nearly straight, around with long pubescence; laterophyse with apex pointed laterally to outer apical angle of dorsal fig.

Female. Unknown.

Body length (male): 9.0 mm; width: 1.5 mm.

Patronymic, dedicated to its collector, Alexander Napolov (Riga, Latvia).

This species could be easily distinguished from others by its aedeagus: ventral process of each paramere slightly flattened and twist in middle in ventral view, distinctly bent dorsally in lateral view, dorsal fig with a longitudinal ridge in middle of basal portion, membranous between inner margin and the ridge.

The left mesoleg and right metatarsomeres II–V of the holotype are missing.

Holotype ♂ (ZIN): “N. Vietnam, Sa Pa env., 1200m, 20.V.1999, leg. Ozlov”. Paratypes: 1♀ (ZIN): same data as holotype; 1♀ (ZIN): “ВЬЕТНАМ [Vietnam] горы, у ША-ПА [Sa Pa], 1600–2000m, 5.6.1963г, Каьакое [Kabakov]”.

Vietnam (Sa Pa).

Male (Fig. 6). Body black, mandibles dark brown, pronotum red, with a large dark brown marking, which almost extending to all margins of disc, elytra red, nearly black at the whole length of all elytral interstices.

Head evenly narrowed behind eyes, surface densely and finely punctate, semilustrous, covered with dense, fine, yellowish brown decumbent pubescence; eyes strongly protruding, head width across eyes distinctly wider than anterior margin of pronotum; terminal maxillary palpomere long-triangular, nearly truncate and sharp at apical one-third length of inner margin; antennae (Fig. 15) extending to apical one-fourth length of elytra, antennomeres II nearly as long as wide at apices, III–XI flattened, III–IX obliquely widened apically, nearly long-triangular, III about twice as long as wide at apices, IV slightly longer than III, X–XI nearly parallel-sided, XI slightly longer than X and pointed at apices, IV–XI each with a round smooth impression at apical part of outer margin.

Pronotum slightly longer than wide, widest near posterior margin, anterior margin rounded, anterior angles rounded, lateral margins moderately diverging posteriorly, slightly sinuate at anterior portion, posterior angles rectangular, posterior margin slightly arcuate and narrowly bordered, disc moderately convex at posterolateral parts, surface punctate and pubescent like that of head, matt.

Elytra about 4.3 times longer than pronotum, 3.5 times longer than humeral width, which about one-third wider than posterior margin of pronotum, lateral margins nearly parallel, elytral venations well-developed, moderately costate.

All tarsal claws simple.

Abdominal sternite IX long-triangular. Aedeagus (Figs 31–33): ventral process of each paramere normal and rounded at apex, even and nearly straight in lateral view; dorsal fig slightly shorter than ventral process, not narrowed apically, with inner angle rounded, outer angle obtuse-angled, inner margin nearly straight, apical margin rounded; laterophyse with apex pointed laterodorsally to outer angle of dorsal fig.

Female. Similar to the male, but eyes less protruding; antennae (Fig. 14) shorter, extending to basal one-third length of elytra, antennomeres III about 1.1 times longer than wide at apices, IV–XI without impressions; pronotum nearly as long as wide, disc slightly convex on posterolateral parts; elytra with lateral margins slightly diverging posteriorly; pro- and meso-outer and inner tarsal claws each with a basal projection; abdominal sternite VIII (Fig. 37) very slightly emarginated in middle and largely emarginated on both sides of posterior margin, the portion between middle and lateral emarginations indistinctly angled at apex; internal reproductive organ of genitalia see Fig. 41.

Body length (both sexes): 8.5–10.0 mm; width: 2.0–2.2 mm.

The specific name is derived from Latin quadrus (four) and linea (stripe), referring to its elytra darkened at all of the four elytral interstices.

This species can be easily distinguished from other species by the elytra darkened at all elytral interstices, elytral venation well-developed; all claws simple in male, pro- and meso-outer and inner tarsal claws each with a basal projection in female; aedeagus: dorsal fig of each paramere not narrowed apically.

Holotype: 1 ♂ (MNHN): [p]“MUSEUM PARIS \ SIKKIM \ DARDJILING \ HARMAND 1890”, [h]“Det. M. Pic (Digoin) \ Cantharis \ bicoloripennis \ n. sp.”, [h]“Athemus \ bicoloripennis \ (Pic) \ det. W. Wittmer”, [p]“HOLOTYPUS”.

CHINA, Xizang: 1 ♂ (IZAS): Nyalam, Zham, 2200m, 28.VI.1975, leg. Zi-Qing Wang; 1 ♂, 1 ♀ (HBUM): Zham, 5.–6.VII.2004, leg. Yi-Bin Ba & Ai-Min Shi.

China (new record: Xizang); Nepal, India (Kazantsev and Brancucci 2007).

Lectotype of Lycocerus caliginostus: 1♀ (MNHN): without locality data, [h]“Lycocerus \ caliginosus \ Gorh.”, [p] “TYPE”.

Paratype of Lycocerus vittaticollis: 1♀ (NHMB): [p]“Birmah \ Ruby-mines”, [p]“Doherty”, [p]“Frey Coll. \ 1905.100.”, [p]“♀”, [p]“Para \ type”, [p]“Lycoerus \ (?) vittaticollis, \ Champ. ”, [p]“E.M.M. 1926 \ det. G.C.C.”, [h]“L. \ vittaticollis \ Champ. \ det. W. Wittmer”, [h]“Athemus \ caliginosus (Gorh.) \det. S. Kazantsev 1996”, [p]“CANTHARIDAE \ CANTH00002784”, [p]“Naturhistorisches \ Museum Basel \ Coll. W. Wittmer”.

CHINA, Yunnan: 1♀ (IZAS): Menghai, Nannuoshan, 1100–1500m, 27.IV.1957, leg. Guang-Ji Hong; 1♀ (IZAS): Xiaomengyang, 850m, 3.V.1957, leg. Panfilov; 1♂ (IZAS): same data, 2.IV.1957, leg. Ling-Chao Zang; 1♂ (IZAS): Menghai, Nannuoshan, 1250m, 24.IV.1957, leg. Panfilov; 1♂ (IZAS): Xishuangbanna, Mengsong, 1600m, 23.IV.1958, leg. Fu-Ji Pu; 1♀ (IZAS): same data, 25.IV.1958, leg. Chun-Pei Hong; 1♀ (IZAS): same data, leg. Yi-Ran Zhang; 1♀ (IZAS): same data, 22.IV.1958, leg. Xu-Wu Meng; 1♀ (IZAS): same data, 23.IV.1958, leg. Shu-Yong Wang; 1♂ (IZAS): same data, 22.IV.1958, leg. Shu-Yong Wang; 1♂ (IZAS): Cheli, Shihuiyao, 750m, 27.IV.1958, leg. Panfilov; 1♀ (IZAS): Simao, 1300m, 31.IV.1957, leg. Bushchik.

China (new record: Yunnan); Laos, Thailand, Myanmar (Kazantsev 1999).

Holotype: 1♂ (NMPC): [p]“Laos north, 5–11.V.1997, \ 20 km NW Louang Namtha, \ N21°09.2, E 101°18.7, \ alt. 900±100 m, \ E. Jendek & O. Šauša leg.”, [p]“HOLOTYPUS \ Lycocerus \ jendeki sp. nov. \ V. Švihla det. 2005”.

CHINA, Yunnan: 1♂ (IZAS): Simao, Rd. Kunluo, 1350m, 11.V.1957, leg. Fu-Ji Pu; 1♀ (IZAS): Xishuangbanna, Meng’a, 1050–1080m, 10.V.1958, leg. Shu-Yong Wang.

China (new record: Yunnan); Laos.

Holotype: 1♂ (NHMB): [p]“N. E. Burma \ Kambaiti, 7000 ft \ 24/5 1934 \ R. Malaise”, [h]“malaisei”, [p]“HOLOTYPE”, [p]“Naturhistorisches \ Museum Basel \ coll. W. Wittmer”.

CHINA, Yunnan: 1♂ (IZAS): 60km E tengchong, 2200m, 19.–22.V2006, leg. S. Murzin & I. Shokhin; 1♂ (CAS): Longling County, Longjiang Township, Xiaoheishan Forest Reserve, Guchengshan, 2020m, N24.82888°/E098.76001°, 26.V.2005, stop# 2005-030A, D.H.Kavanaugh & C.E. Griswold collectors [CASENT 6004806]; 1♂ (CAS): same data [CASENT 1035946]; 1♂ (CAS): same data [CASENT 1035947]; 1♀ (CAS): same data [CASENT 1035948]; 1♀ (CAS): Longyang County, Bawan Township, Nankang Yakou just N of pass, N24.83178°/E098.76472°, 2180m, 25.V.2005, stop# 2005-029B, D.H. Kavanaugh & C.E. Griswold collectors [CASENT 1035849].

China (new record: Yunnan); Myanmar.

Lectotype of Lycocerus obscurus: 1♀ (MNHN): [h] “Xieng Khuang” [LAOS, Xieng Khouang]. Paralectotypes: 2♀♀(MNHN): same data as lectotype; 1♀ (MNHN): “X. K.”; 1♀ (MNHN): “Xieng Khuang \ 17-III-1919 \ R. Vitalis de Salvaza”.

Lectotype of Lycocerus obscurus var. diversus: 1♀ (MNHN): [h]“Xieng Khuang \ ex Vitalis”, [h]“L. obscurus v. \ diversus Pic ”, [h]“type”, [h]“obscurus \ v. diversus”, [p]“TYPE”, [h]“LECTOTYPUS \ L. obscurus v. \ diversus Pic \ S. Kazantsev det.”.

CHINA, Guangxi: 2♂♂, 2♀♀ (IZAS): Jingxi, 840m, 1.IV.1998, leg. Chun-Sheng Wu.

Female. Similar to male, but eyes less protruding; antennae shorter, extending to basal one-third length of elytra; pronotum wider; abdominal sternite VIII (Fig. 38) largely emarginated in middle and on both sides of posterior margin, the portion between middle and lateral emarginations distinctly protuberant and rounded at apex.

Body length (both sexes): 10.0–12.5 mm; width: 2.5–3.0 mm.

China (new record: Guangxi); Laos (Kazantsev 1999).

Based on the examination of the types of Lycocerus obscurus Pic, 1916 and some additional material at our disposal, we found no difference between it and L. rubroniger Švihla, 2011, which was presented with an adequate photo of the habitus and illustrations of aedeagus in the original publication. Therefore, L. rubroniger is synonymized with L. obscurus here, and provided with the supplementary description and illustration of abdominal sternite VIII of female.

Holotype: 1♂ (NHMB): [h]“Shillong \ Assam”, [h]“26.4.1971 \ T. Sen Gupta”, [h]“Athemus s.str. \ olivaceus \ Wittm. \ det. W. Wittmer”, [p]“HOLOTYPUS”, [p]“Naturhistorisches \ Museum Basel \ coll. W. Wittmer”, [p]“CANTHARIDAE \ CANTH00001639”.

CHINA, Yunnan: 1♂, 2♀♀ (IZAS): Gongshan, Dulong, Kongdang, 21.V.2007, leg. Yan-Lei Li.

China (new record: Yunnan); India.

Holotype: 1♂ (NHMB): [p]“Dechhi Paka 3300m \ 19.-20.5.” [p]“Nat.-Hist. Museum \ Basel-Bhutan \ Expedition 1972”, [h]“Athemus subg. \ Andrathemus \ purpurascens \ Wittm. \ det. W. Wittmer”, [p]“HOLOTYPUS”, [p]“CANTHARIDAE \ CANTH00001271”.

CHINA, Xizang: 1♂, 1♀ (IZAS): Nyingtri, 3050m, 3.VIII.1983, leg. Yin-Heng Han.

China (new record: Xizang); Bhutan (Kazantsev and Brancucci 2007).

Holotype: 1♂ (NHMB): [p]“THAI, 10.-16.V.1991 \ Chiang Dao 600m \ 15°24’N 98°55’E \ Vit Kubáň leg.”, [p]“Thailand 91 \ Thanon Thong Chai \ D. Král & V. Kubáň”, [h]“A. \ ruficornis \ Wittm. \ det. W. Wittmer”, [p]“HOLOTYPUS”, [p]“CANTHARIDAE \ CANTH00002524”.

CHINA, Yunnan: 1♂ (IZAS): Xishuangbanna, Meng’a, 1050–1080m, 2.V.1958, leg. Shu-Yong Wang; 1♀ (IZAS): Xishuangbanna, Xiaomengyang, 850m, 8.VII.1957, leg. Ling-Chao Zang. [MYANMAR]: 2♂♂ (MCSNG): Burma, Tenasserim, Thagalà, IV.1887, Fea.

China (new record: Yunnan); Myanmar (new record); Thailand.

Holotype: 1♂ (NHMB): [p]“E-Nepal \ Koshi \ M. Brancucci”, [p]“Gufa-Gorza \ 2800-2100m \ 4.VI.1985”, [h]“REM \ 94 \ 4.7”, [h]“Athemus s.str. \ semiextensus \ Wittm. \ det. W. Wittmer”, [p]“HOLOTYPUS”, [p]“CANTHARIDAE \ CANTH00001301”.

CHINA, Xizang: 1♂ (IZAS): Droma, 2800m, 8.VI.1961, leg. Lin-Yao Wang; 1 ♀ (IZAS): same data, 7.VI.1961; 1 ♂ (IZAS): same data, 5.VI.1961; 1 ♀ (IZAS): same data, 6.VI.1961.

China (new record: Xizang); India, Bhutan, Nepal (Kazantsev and Brancucci 2007).