A Finnish checklist of the sphaeroceroid fly families Chyromyidae and Heleomyzidae is provided.

Species list, Finland, Diptera , biodiversity, faunistics

The superfamily Sphaeroceroidea is a medium-sized one, with two families of moderate diversity, Sphaeroceridae (1550 species) and Heleomyzidae (~720 species), and the small family Chyromyidae. The enigmatic afrotropical Mormotomyia hirsuta Austen, 1936 was once placed near Sphaeroceridae but it is now seen as an ephydroid fly (Kirk-Spriggs et al. 2011). McAlpine (2007) has proposed an alternative concept for Sphaeroceroidea with Sphaeroceridae and Heleomyzidae united as a single family called Heteromyzidae. This proposal has not gained significant support and for the purposes of this checklist the traditional concept of family Sphaeroceridae is retained.

There is no general agreement on the relationships of various heleomyzid tribes. Several different schemes for subfamilies have been proposed (see McAlpine 2007, McAlpine and Woodley 2010). Some taxa treated here as heleomyzids (primarily Trixoscelidinae, Chiropteromyzinae, Heteromyzinae and Borboropsini) may deserve full family status. As a conservative approach this checklist follows Marshall (2012) and keeps them as subfamilies and tribes. The heleomyzid subfamilies and tribes are listed alphabetically.

The Finnish chyromyids are small yellow flies with (at least while alive) iridescent blue or green eyes. Chyromyids are rarely collected and little is known about their ecology or the proper place of the family within Sphaeroceroidea. They may actually be a specialized lineage arising from within Heleomyzidaesensu lato.

Two of the three sphaeroceroid families are treated in this paper. The largest, Sphaeroceridae, is covered in a separate paper in this issue of ZooKeys. The Finnish species of Heleomyzidae and Chyromyidae were last listed by Hackman (1980).

suborder Brachycera Macquart, 1834

clade Eremoneura Lameere, 1906

clade Cyclorrhapha Brauer, 1863

infraorder Schizophora Becher, 1882

clade Muscaria Enderlein, 1936

parvorder Acalyptratae Macquart, 1835

superfamily Sphaeroceroidea Macquart, 1835

CHYROMYIDAE Hendel, 1916

CHYROMYINAE Hendel, 1916

CHYROMYA Robineau-Desvoidy, 1830

Chyromya flava (Linnaeus, 1758)

Chyromya oppidana (Scopoli, 1763)

GYMNOCHIROMYIA Hendel, 1933

Gymnochiromyia flavella (Zetterstedt, 1848)

= minima (Becker, 1904)

Gymnochiromyia inermis (Collin, 1933)

HELEOMYZIDAE Westwood, 1840

BORBOROPSINAE Griffiths, 1972

BORBOROPSIS Czerny, 1902

Borboropsis puberula (Zetterstedt, 1838)

= fulviceps (Strobl, 1898)

CHIROPTEROMYZINAE Frey, 1952

CHIROPTEROMYZA Frey, 1952

Chiropteromyza broersei (de Meijere, 1946)

= wegelii Frey, 1952

NEOSSOS Malloch, 1927

= Ornitholeria Frey, 1930

Neossos nidicola (Frey, 1930)

HETEROMYZINAE Fallén, 1820

HETEROMYZA Fallén, 1820

Heteromyza atricornis Meigen, 1830

Heteromyza oculata Fallén, 1820

Heteromyza rotundicornis (Zetterstedt, 1846)

TEPHROCHLAMYS Loew, 1862

Tephrochlamys flavipes (Zetterstedt, 1838)

Tephrochlamys rufiventris (Meigen, 1830)

= lapponica (Czerny, 1924)

Tephrochlamys steniusi Frey, 1930

Tephrochlamys tarsalis (Zetterstedt, 1847)

HELEOMYZINAE Westwood, 1840

tribe Heleomyzini Westwood, 1840

GYMNOMUS Loew, 1863

Gymnomus amplicornis (Czerny, 1924)

HELEOMYZA Fallén, 1810

= Helomyza Fallén, 1820 emend.

= Leria Robineau-Desvoidy, 1830

sg. Heleomyza Fallén, 1810

Heleomyza borealis Boheman, 1865

= czernyi Collart, 1933

= modesta misid.

Heleomyza hackmani Frey, 1950

Heleomyza pleuralis (Becker, 1907)

Heleomyza serrata (Linnaeus, 1758)

MORPHOLERIA Garrett, 1921

sg. Spanoparea Czerny, 1924

Morpholeria dudai (Czerny, 1924)

Morpholeria kerteszii Czerny, 1924

Morpholeria obscuriventris (Zetterstedt, 1847)

Morpholeria ruficornis (Meigen, 1830)

NEOLERIA Malloch, 1919

Neoleria inscripta (Meigen, 1830)

= minuta (Zetterstedt, 1838)

Neoleria prominens (Becker, 1897)

= tibialis misid.

Neoleria ruficauda (Zetterstedt, 1847)

Neoleria ruficeps (Zetterstedt, 1838)

SCOLIOCENTRA Loew, 1862

sg. Chaetomus Czerny, 1924

Scoliocentra confusa (Wahlgren, 1918)

Scoliocentra flavotestacea (Zetterstedt, 1838)

sg. Leriola Gorodkov, 1962

Scoliocentra brachypterna (Loew, 1873)

Scoliocentra nigrinervis (Wahlgren, 1918)

sg. Scoliocentra Loew, 1862

Scoliocentra dupliciseta (Strobl, 1894)

Scoliocentra scutellaris (Zetterstedt, 1838)

Scoliocentra villosa (Meigen, 1830)

tribe Oecotheini Gorodkov, 1972

ECCOPTOMERA Loew, 1862

Eccoptomera infuscata Wahlgren, 1918

Eccoptomera longiseta (Meigen, 1830)

Eccoptomera marginicornis Czerny, 1924

Eccoptomera microps (Meigen, 1830)

Eccoptomera obscura (Meigen, 1830)

Eccoptomera ornata Loew, 1862

Eccoptomera pallescens (Meigen, 1830)

OECOTHEA Haliday, 1837

Oecothea fenestralis (Fallén, 1820)

tribe Orbelliini Gorodkov, 1972

ORBELLIA Robineau-Desvoidy, 1830

Orbellia nivicola Frey, 1913

SUILLIINAE Wahlgren, 1917

SUILLIA Robineau-Desvoidy, 1830

= Allophyla Loew, 1862

Suillia affinis (Meigen, 1830)

Suillia apicalis (Loew, 1862)

Suillia atricornis (Meigen, 1830)

Suillia bicolor (Zetterstedt, 1838)

Suillia femoralis (Loew, 1862)

Suillia flava (Meigen, 1830)

Suillia flavifrons (Zetterstedt, 1838)

= nudipes (Czerny, 1932)

Suillia fuscicornis (Zetterstedt, 1847)

Suillia humilis (Meigen, 1830)

= inornata (Loew, 1862)

Suillia laevifrons (Loew, 1862)

Suillia lineitergum (Pandellé, 1901)

= stroblii (Czerny, 1904)

Suillia lurida (Meigen, 1830)

Suillia mikii (Pokorny, 1886)

Suillia nemorum (Meigen, 1830)

Suillia pallida (Fallén, 1820)

Suillia parva (Loew, 1862)

= collini Hackman, 1972

= flavifrons auct. nec (Zetterstedt, 1838)

Suillia quadrilineata Czerny, 1924

Suillia vaginata (Loew, 1862)

TRIXOSCELIDINAE Hendel, 1916

TRIXOSCELIS Rondani, 1856

Trixoscelis frontalis (Fallén, 1823)

? = canescens misid. (see Notes)

Trixoscelis marginella (Fallén, 1823)

Trixoscelis obscurella (Fallén, 1823)

Trixoscelis similis Hackman, 1970

Chyromya oppidana (Scopoli, 1763). Found only inside houses and farm buildings in Finland.

Orbellia nivicola Frey, 1913. This species was synonymized with O. myiopiformis R.-D. by Storå (1958), but Frey (1958) defended its validity. The status of O. nivicola as a species needs verification.

Trixoscelis canescens (Loew, 1865). This species was originally described on the basis of a single female. Soós (1979) examined the type and revived the name from synonymy with T. frontalis. Woźnica (2008) provided an illustrated diagnosis for T. canescens and synonymized T. gigans Carles-Tolrá, 2001 and T. fumipennis Papp, 2005 with it. The species was recently recorded from Finland by Flinck and Kahanpää (2013). Specimens with darkened costal veins and dorsal abdominal surfaces, both proposed diagnostic characters of T. canescens, are common among Finnish T. frontalis material (see Fig. 8 in Flinck and Kahanpää 2013). Finnish males with these features have genitalia identical with those illustrated for T. frontalis by Hackman (1970) and quite unlike the genitalia of T. gigans (= fumipennis Papp). The male specimen mentioned in Flinck and Kahanpää (2013) was later dissected and it belongs to T. frontalis. The external characters (darkened costa and dorsum of abdomen) can not be used to reliably separate T. canescens from T. frontalis. The Finnish records of T. canescens are probably all misidentifications of T. frontalis.