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Checklist
Checklist of the Diptera families Acartophthalmidae, Canacidae (including Tethinidae), Carnidae and Milichiidae of Finland (Insecta)
expand article infoJere Kahanpää
† University of Helsinki, Helsinki, Finland
Open Access

Abstract

A checklist of 29 species in the smaller carnoid families Acartophthalmidae, Canacidae, Carnidae and Milichiidae (Diptera) recorded from Finland is presented. Tethinidae are also included as a subfamily of Canacidae. Phyllomyza tetragona Hendel is removed from the list as no reliable records of this species within the post-1944 borders of Finland could be found.

Keywords

Checklist, Finland, Diptera, biodiversity, faunistics

Introduction

The superfamily Carnoidea (sensu Buck 2006 but excluding Cryptochetidae, see Wiegmann et al. 2011) includes seven families. The largest by far is Chloropidae, treated in a separate article in this issue. Four of the remaining six are found in Finland: Acartophthalmidae, Canacidae, Carnidae and Milichiidae. The remaining two families, Inbiomyiidae and Australimyzidae, are respectively restricted to the Neotropical and Australasian Regions.

All four families are rather poorly studied in Finland. The presence of additional, even undescribed, species is very likely in Carnidae and Milichiidae. The identification of all species of Meoneura was verified from male genitalia during the preparation of this checklist with one exception: the male of Meoneura elongella (Zetterstedt, 1838) remains undescribed. It is the only North European Meoneura with black halteres and as such easily recognised.

World catalogues have recently been published for Canacidae (Munari and Mathis 2010), Carnidae (Brake 2011), and Milichiidae (Brake 2000). The Finnish species of these families were last listed in Hackman (1980).

Number of species by family.

Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Acartophthalmidae 5 3 3 average
Canacidae 322 38 1 average
Carnidae 93 (Brake 2011) 39 13 poor
Milichiidae 278 43 12–13 average

Checklist

suborder Brachycera Macquart, 1834

clade Eremoneura Lameere, 1906

clade Cyclorrhapha Brauer, 1863

infraorder Schizophora Becher, 1882

clade Muscaria Enderlein, 1936

parvorder Acalyptratae Macquart, 1835

superfamily Carnoidea Newman, 1834

ACARTOPHTHALMIDAE Czerny, 1928

ACARTOPHTHALMUS Czerny, 1902

Acartophthalmus bicolor Oldenberg, 1910

Acartophthalmus nigrinus (Zetterstedt, 1848)

Acartophthalmus pusio Frey, 1947

CANACIDAE Jones, 1906

PELOMYIINAE Foster, 1976

PELOMYIELLA Hendel, 1934

Pelomyiella cinerella (Haliday, 1837)

CARNIDAE Newman, 1834

CARNUS Nitzsch, 1818

Carnus hemapterus Nitzsch, 1818

MEONEURA Rondani, 1856

Meoneura anceps Frey, 1935

Meoneura elongella (Zetterstedt, 1838)

Meoneura flavifacies Collin, 1930

Meoneura glaberrima Becker, 1907

= neglecta Collin, 1930

Meoneura lacteipennis (Fallén, 1823)

Meoneura lamellata Collin, 1930

Meoneura minutissima (Zetterstedt, 1860)

Meoneura neottiophila Collin, 1930

Meoneura obscurella (Fallén, 1823)

Meoneura prima (Becker, 1903)

= seducta Collin, 1937

Meoneura triangularis Collin, 1930

Meoneura vagans (Fallén, 1823)

MILICHIIDAE Schiner, 1862

MADIZINAE Czerny, 1909

DESMOMETOPA Loew, 1866

Desmometopa m-nigrum (Zetterstedt, 1848)

Desmometopa sordida (Fallén, 1820)

Desmometopa varipalpis Malloch, 1927

LEPTOMETOPA Becker, 1903

Leptometopa latipes (Meigen, 1830)

MADIZA Fallén, 1810

Madiza glabra Fallén, 1820

MILICHIINAE Schiner, 1862

MILICHIA Meigen, 1830

Milichia ludens (Wahlberg, 1847)

PHYLLOMYZINAE Curran, 1934

NEOPHYLLOMYZA Melander, 1913

Neophyllomyza acyglossa (Villeneuve, 1920)

PHYLLOMYZA Fallén, 1810

? Phyllomyza equitans (Hendel, 1919)

Phyllomyza formicae Schmitz, 1923

Phyllomyza longipalpis (Schmitz, 1924)

Phyllomyza rubricornis Schmitz, 1923

Phyllomyza securicornis Fallén, 1823

Phyllomyza silesiaca (Duda, 1935)

Excluded species

Phyllomyza tetragona Hendel, 1924 not found within present borders

Tethina grisea (Fallén, 1823) misidentification?

Tethina illota (Haliday, 1838)

Notes

Meoneura. At least two additional, possibly undescribed species occur in Finland.

Phyllomyza equitans (Hendel, 1919). Only females have been found in Finland.

Tethina grisea (Fallén, 1823) and T. illota (Haliday, 1838). Munari and Mathis (2010) list two Tethina species from Finland. Krogerus (1932) is mentioned as the source for the T. grisea record; there is no listed source for the record of T. illota. An older version of the world catalogue (Mathis and Munari 1996) mentions only T. grisea from Finland. No Finnish specimens of these species could be found in the Finnish Museum of Natural History (MZH) where most of the material collected by Krogerus is kept, or in any other Finnish insect collection. Unfortunately the identifications in Krogerus (1932) are rather unreliable. It is possible that Tethina species occur in Finland, but none are included in this checklist because the evidence for their presence is, in my opinion, insufficient.

References

  • Brake I (2000) Phylogenetic systematics of the Milichiidae (Diptera, Schizophora). Entomologica Scandinavica Supplement 57: 1–120.
  • Buck M (2006) A new family and genus of acalypterate flies from the Neotropical region, with a phylogenetic analysis of Carnoidea family relationships (Diptera, Schizophora). Systematic Entomology 31(3): 377–404. doi: 10.1111/j.1365-3113.2006.00328.x
  • Hackman W (1980) A check list of the Finnish Diptera. Notulae entomologicae 60: 17–48, 117–162.
  • Krogerus R (1932) Über die Ökologie und Verbreitung der Arthropoden der Triebsandgebiete an den Küsten Finnlands. Acta Zoologica Fennica 12: 1–308.
  • Mathis WN, Munari L (1996) World catalog of the family Tethinidae (Diptera). Smithsonian Contributions to Zoology 584: 1–27. doi: 10.5479/si.00810282.584
  • Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang Z-Q (Ed) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/zt03148p229.pdf
  • Wiegmann BM, Trautwein MD, Winkler IS, Barr NB, Kim J-W, Lambkin C, Bertone MA, Cassel BK, Bayless KM, Heimberg AM, Wheeler BM, Peterson KJ, Pape T, Sinclair BJ, Skevington JH, Blagoderov V, Caravas J, Kutty SN, Schmidt-Ott U, Kampmeier GE, Thompson FC, Grimaldi DA, Beckenbach AT, Courtney GW, Friedrich M, Meier R, Yeates DK (2011) Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the United States of America 108(14): 5690–5695. doi: 10.1073/pnas.1012675108