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Checklist
Checklist of the superfamilies Conopoidea, Diopsoidea and Nerioidea of Finland (Insecta, Diptera)
expand article infoJere Kahanpää, Jens-Hermann Stuke
† University of Helsinki, Helsinki, Finland
‡ Unaffiliated, Leer, Germany
Open Access

Abstract

A checklist of the Diptera superfamilies Conopoidea (Conopidae), Nerioidea (Micropezidae, Pseudopomyzidae) and Diopsoidea (Megamerinidae, Psilidae, Strongylophthalmyiidae, Tanypezidae) from Finland in presented. Myopa vicaria Walker, 1849 is formally recorded for the first time from the country.

Keywords

Checklist, Finland, Diptera, biodiversity, faunistics

Introduction

Four fly families form the superfamily Nerioidea. Two of them, Micropezidae and Pseudopomyzidae, occur in Finland. Neriids are mostly tropical and cypselomatids (sensu stricto) have only been found in the Australasian and Oriental regions. The pseudopomyzids were included in Cypselosomatidae Hendel, 1931 in the previous checklist (Hackman 1980) and are still often included in Cypselosomatidae sensu lato. The Finnish micropezid fauna is relatively well known. One or two species of Cnodacophora Czerny, 1930 and Rainieria Rondani, 1843 could be present but are so far undetected.

Diopsoidea consists of seven to ten families (depending on the source), none particularly species-rich. Only four are found in Finland: Psilidae, Tanypezidae, Strongylophthalmyiidae and Megamerinidae. Tanypezidae and Strongylophthalmyiidae are here treated as separate families following Lonsdale (2013). Megamerinidae has been assigned to several superfamilies, most often Diopsoidea following McAlpine (1989), but also Nerioidea, Sciomyzoidea, and recently Opomyzoidea (Wiegmann et al. 2011). These families have raised no particular interest among Finnish dipterists. The genus Chamaepsila Hendel, 1917 in particular needs more attention. It is likely that a few Finnish species remain unrecognised in this genus.

The systematic position of the Conopidae is unclear. It has traditionally been considered basal within Schizophora. Wiegmann et al. (2011) listed Conopidae within Sciomyzoidea while others (see Skevington et al. 2010) place the family near Tephritoidea. Within the family the recent classification of Gibson and Skevington (2013) is followed. The Finnish conopid fauna is now relatively well-understood and few additional species are expected to occur in the country.

The Finnish species of these families were last listed by Hackman (1980).

Number of species by family.

Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Conopidae ~796 83 19 good
Nerioidea:
Micropezidae 573 22 6 average
Pseudopomyzidae 18 1 1 good
Diopsoidea:
Psilidae 320 47 29 average
Tanypezidae 27 (Lonsdale 2013) 1 1 good
Strongylophthalmyiidae 47 (Lonsdale 2013) 2 2 good
Megamerinidae 15 1 1 good

Checklist part 1: Conopoidea

suborder Brachycera Macquart, 1834

clade Eremoneura Lameere, 1906

clade Cyclorrhapha Brauer, 1863

infraorder Schizophora Becher, 1882

? clade Archischiza Enderlein, 1936

superfamily Conopoidea Latreille, 1802

CONOPIDAE Latreille, 1802

CONOPINAE Latreille, 1802

CONOPS Linnaeus, 1758

sg. Conops Linnaeus, 1758

Conops quadrifasciatus De Geer, 1776

Conops strigatus Wiedemann, 1824

Conops vesicularis Linnaeus, 1761

PHYSOCEPHALA Schiner, 1861

Physocephala nigra (De Geer, 1776)

ZODIONINAE Rondani, 1856

ZODION Latreille, 1796

Zodion cinereum (Fabricius, 1794)

= notatum Meigen, 1804

SICINAE Zimina, 1960

SICUS Scopoli, 1763

Sicus ferrugineus (Linnaeus, 1761)

MYOPINAE Macquart, 1834

MYOPA Fabricius, 1775

Myopa buccata (Linnaeus, 1758)

Myopa fasciata Meigen, 1804

Myopa hirsuta Stuke & Clements, 2008

= strandi auct. nec Duda, 1940

Myopa occulta Wiedemann, 1824

= nigrifrons von Bonsdorff, 1866

Myopa tessellatipennis Motschulsky, 1859

= polystigma auct. nec Rondani, 1857

Myopa testacea (Linnaeus, 1767)

Myopa vicaria Walker, 1849

= villosa Ringdahl, 1945

= strandi Duda, 1940

MYOPOTTA Zimina, 1969

Myopotta pallipes (Wiedemann, 1824)

THECOPHORA Rondani, 1845

= Occemyia Robineau-Desvoidy, 1853

Thecophora cinerascens (Meigen, 1804)

= pusilla (Meigen, 1824)

= atra misid.

Thecophora distincta (Wiedemann, 1824)

= melanopa misid.

Thecophora fulvipes (Robineau-Desvoidy, 1830)

= sundewalli (Zetterstedt, 1844)

Thecophora jakutica Zimina, 1974

= atra misid.

DALMANNIINAE Hendel, 1916

DALMANNIA Robineau-Desvoidy, 1830

Dalmannia dorsalis (Fabricius, 1794)

= punctata misid.

Checklist part 2: Nerioidea and Diopsoidea

suborder Brachycera Macquart, 1834

clade Eremoneura Lameere, 1906

clade Cyclorrhapha Brauer, 1863

infraorder Schizophora Becher, 1882

clade Muscaria Enderlein, 1936

parvorder Acalyptratae Macquart, 1835

superfamily Nerioidea Westwood, 1840

MICROPEZIDAE Blanchard, 1840

CALOBATINAE Bigot, 1853

CALOBATA Meigen, 1803

Calobata petronella (Linnaeus, 1761)

NERIA Robineau-Desvoidy, 1830

Neria cibaria (Linnaeus, 1761)

Neria commutata (Czerny, 1930)

= nigricornis auct. nec Zetterstedt, 1838

Neria ephippium (Fabricius, 1794)

Neria nigricornis (Zetterstedt, 1838)

= helleni (Frey, 1918)

= nitidicollis (Frey, 1947)

MICROPEZINAE Blanchard, 1840

MICROPEZA Meigen, 1803

Micropeza corrigiolata (Linnaeus, 1767)

PSEUDOPOMYZIDAE McAlpine, 1966

PSEUDOPOMYZA Strobl, 1893

Pseudopomyza atrimana (Meigen, 1830)

superfamily Diopsoidea Billberg, 1820

MEGAMERINIDAE Hendel, 1913

MEGAMERINA Rondani, 1861

= Lissa Meigen, 1826 preocc.

Megamerina dolium (Fabricius, 1805)

= loxocerina (Fallén, 1820)

PSILIDAE Macquart, 1835

CHYLIZINAE Rondani, 1856

CHYLIZA Fallén, 1820

sg. Chyliza Fallén, 1820

Chyliza annulipes Macquart, 1835

= fuscipennis auct. nec Robineau-Desvoidy, 1830

Chyliza leptogaster (Panzer, 1798)

= scutellata (Fabricius, 1798)

Chyliza nova Collin, 1944

Chyliza vittata Meigen, 1826

PSILINAE Macquart, 1835

tribe Loxocerini Macquart, 1835

IMANTIMYIA Frey, 1925

Imantimyia albiseta (Schrank, 1803)

= ichneumonea (Linnaeus, 1758) nom. dubium (see Notes)

Imantimyia fulviventris (Meigen, 1826)

Imantimyia nigrifrons (Macquart, 1835)

Imantimyia sylvatica (Meigen, 1826)

LOXOCERA Meigen, 1803

sg. LOXOCERA Meigen, 1803

Loxocera aristata (Panzer, 1801)

= ichneumonea (Linnaeus, 1758) nom. dubium (see Notes)

tribe Psilini Macquart, 1835

CHAMAEPSILA Hendel, 1917

= Tetrapsila Frey, 1925

Chamaepsila atra (Meigen, 1826)

Chamaepsila bicolor (Meigen, 1826)

= nigromaculata (Strobl, 1909)

Chamaepsila buccata (Fallén, 1826)

= gracilis (Meigen, 1826)

Chamaepsila humeralis (Zetterstedt, 1847)

Chamaepsila limbatella (Zetterstedt, 1847)

Chamaepsila morio (Zetterstedt, 1835)

Chamaepsila nigra (Fallén, 1820)

Chamaepsila nigricornis (Meigen, 1826)

Chamaepsila nigrosetosa Frey, 1925

Chamaepsila obscuritarsis (Loew, 1856)

Chamaepsila pallida (Fallén, 1820)

Chamaepsila pectoralis (Meigen, 1826)

Chamaepsila rosae (Fabricius, 1794)

Chamaepsila rufa (Meigen, 1826)

Chamaepsila unilineata (Zetterstedt, 1847)

PSILA Meigen, 1803

sg. FREYOPSILA Shatalkin, 1986

Psila sibirica (Frey, 1925)

sg. PSILA Meigen, 1803

Psila fimetaria (Linnaeus, 1761)

Psila merdaria Collin, 1944

PSILOSOMA Zetterstedt, 1860

Psilosoma audouini (Zetterstedt, 1835)

Psilosoma lefebvrei (Zetterstedt, 1835)

STRONGYLOPHTHALMYIIDAE Hendel, 1917

STRONGYLOPHTHALMYIA Heller, 1902

Strongylophthalmyia pictipes Frey, 1935

Strongylophthalmyia ustulata (Zetterstedt, 1847)

TANYPEZIDAE Rondani, 1856

TANYPEZA Fallén, 1820

Tanypeza longimana Fallén, 1820

Excluded species

Dalmannia punctata (Fabricius, 1794) not found within present borders.

Notes

Chamaepsila unilineata (Zetterstedt, 1847) and C. nigrosetosa Frey, 1925 were synonymized with C. pallida but they have recently been reinstated as valid species (Shatalkin and Merz 2010).

Loxocera ichneumonea (Linnaeus, 1758). This name has been used for L. aristata (Panzer) and L. albiseta (Schrank). Unfortunately no type material is known. Linné’s original description is insufficient and may in fact belong to a hoverfly (see Buck and Marshall 2006).

Myopa vicaria Walker, 1849. One of the common Myopa species through Finland, but previously unreported due to difficulties in identification. Recorded from provinces Ab, N, Ka, St, Ta, Sa, Kl, Om, Ok, Lkoc and Li.

References

  • Buck M, Marshall SA (2006) Revision of New World Loxocera (Diptera: Psilidae), with phylogenetic redefinition of Holarctic subgenera and species groups. European Journal of Entomology 103: 193–219. doi: 10.14411/eje.2006.022
  • Gibson JF, Skevington JH (2013) Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data. Zoological Journal of the Linnean Society 167(1): 43–81 doi: 10.1111/j.1096-3642.2012.00873.x
  • Hackman W (1980) A check list of the Finnish Diptera. Notulae entomologicae 60: 17–48, 117–162.
  • McAlpine JF (1989) Phylogeny and classification of the Muscomorpha. In: McAlpine JF, Wood DM (Eds) Manual of Nearctic Diptera Volume 3. Agriculture Canada Monographs 32. Research Branch, Agriculture Canada, Ottawa, 1397–1518.
  • Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang Z-Q (Ed) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/zt03148p229.pdf
  • Shatalkin AI, Merz B (2010) The Psilidae (Diptera, Acalyptrata) of Switzerland, with description of two new species from Central Europe. Revue Suisse de Zoologie 117(4): 771–800. www.biodiversitylibrary.org/part/117602
  • Skevington JH, Thompson FC, Camras S (2010) Conopidae (Thick-headed Flies). In: Brown BV, Borkent A, Cumming JM, Wood DM, Woodley NE, Zumbado MA (Eds) Manual of Central American Diptera, Volume 2. NRC Research Press, Ottawa, 847–855.
  • Wiegmann BM, Trautwein MD, Winkler IS, Barr NB, Kim J-W, Lambkin C, Bertone MA, Cassel BK, Bayless KM, Heimberg AM, Wheeler BM, Peterson KJ, Pape T, Sinclair BJ, Skevington JH, Blagoderov V, Caravas J, Kutty SN, Schmidt-Ott U, Kampmeier GE, Thompson FC, Grimaldi DA, Beckenbach AT, Courtney GW, Friedrich M, Meier R, Yeates DK (2011) Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the United States of America 108(14): 5690–5695. doi: 10.1073/pnas.1012675108