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Checklist of the families Lonchopteridae and Phoridae of Finland (Insecta, Diptera)
expand article infoJere Kahanpää
† University of Helsinki, Helsinki, Finland
Open Access

Abstract

A checklist of the Lonchopteridae and Phoridae recorded from Finland is presented.

Keywords

Checklist, Finland, Diptera, Lonchopteridae, Phoridae

Introduction

The superfamily Phoroidea includes at least the fly families Phoridae, Lonchopteridae, and the small family Ironomyiidae known only from Australia. The flat-footed fly families Platypezidae and Opetiidae, treated in a separate paper in this volume, are placed either in their own basal superfamily Platypezoidea or included in Phoroidea (see Cumming et al. 1995, Woodley et al. 2009 and Wiegmann et al. 2011).

Lonchoptera Meigen, 1803 is the only currently recognized genus in Lonchopteridae. Five Lonchoptera species have been added to the Finnish fauna after the checklist of Hackman (1980) (Andersson 1991, Kahanpää 2013).

The scuttle flies (family Phoridae) may be the largest single family in Diptera. The true number of species in this family has been estimated to be over 30000, the vast majority still undescribed (Marshall 2012). The ‘megagenus’ Megaselia Rondani, 1856 contains about 50% of the species (Marshall 2012). Other phorid genera are relatively better known in Finland but none of them can be called well studied. It is likely that at least a hundred (or hundreds of) Finnish species remain unnoticed, most of them in Megaselia.

The checklist for Megaselia is mostly based on previously published records. The collections of Finnish Museum of Natural History (MZH) were examined by the author during the preparation of this checklist and a few obvious mistakes were taken into account. Most of the material in MZH was identified by H. Schmitz (between years 1915 and 1940) or E. Beyer (in 1958). The lists for other phorid genera are based on recently examined specimens from both MZH and private collections. Due to the difficulties encountered in identifying Megaselia, the list for this genus should definitely be considered preliminary. A revision of the Finnish material is sorely needed.

The Finnish species of these families were last listed by Hackman (1980).

Number of species (Lonchopteridae, Phoridae):

World: 63, 4105 species (Pape et al. 2011)

Europe: 13, 656 species

Finland: 8, ~224–234 species

Faunistic knowledge level in Finland: good, poor

Checklist

suborder Brachycera Macquart, 1834

clade Eremoneura Lameere, 1906

clade Cyclorrhapha Brauer, 1863

superfamily Phoroidea Curtis, 1833

LONCHOPTERIDAE Macquart, 1835

LONCHOPTERA Meigen, 1803

= Dipsa Fallén, 1810

Lonchoptera bifurcata (Fallén, 1810)

= furcata (Fallén, 1823)

Lonchoptera fallax de Meijere, 1906

Lonchoptera impicta Zetterstedt, 1848

Lonchoptera lutea Panzer, 1809

Lonchoptera meijerei Collin, 1938

Lonchoptera nigrociliata Duda, 1927

Lonchoptera nitidifrons Strobl, 1898

Lonchoptera scutellata Stein, 1890

PHORIDAE Curtis, 1833

AENIGMATIAS Meinert, 1890

Aenigmatias lubbockii (Verrall, 1877)

ANEVRINA Lioy, 1864

Anevrina thoracica (Meigen, 1804)

Anevrina unispinosa (Zetterstedt, 1860)

= fennica (Becker, 1901)

Anevrina urbana (Meigen, 1830)

BOROPHAGA Enderlein, 1924

Borophaga agilis (Meigen, 1830)

Borophaga carinifrons (Zetterstedt, 1848)

Borophaga femorata (Meigen, 1830)

Borophaga incrassata (Meigen, 1830)

Borophaga subsultans (Linnaeus, 1767)

= okellyi Schmitz, 1937

CHAETOPLEUROPHORA Schmitz, 1922

Chaetopleurophora erythronota (Strobl, 1892)

Chaetopleurophora spinosissima (Strobl, 1892)

CONICERA Meigen, 1830

Conicera dauci (Meigen, 1830)

= atra Meigen, 1830

Conicera floricola Schmitz, 1938

= similis auct. nec (Haliday, 1833)

= minuscula Schmitz, 1953

Conicera schnittmanni Schmitz, 1926

Conicera similis (Haliday, 1833)

= pauxilla Schmitz, 1920

? Conicera tarsalis Schmitz, 1920

Conicera tibialis Schmitz, 1925

= fallens Schmitz, 1948

DIPLONEVRA Lioy, 1864

= Diploneura emend.

Diplonevra florescens (Turton, 1801)

= florea (Fabricius, 1794) preocc.

= abdominalis (Fallén, 1823)

Diplonevra concinna (Meigen, 1830)

Diplonevra freyi Schmitz, 1927

Diplonevra funebris (Meigen, 1830)

= rostralis (Schmitz, 1918)

Diplonevra glabra Schmitz, 1927

= parcepilosa Schmitz, 1927

Diplonevra nitidula (Meigen, 1830)

Diplonevra oldenbergi Schmitz, 1920

DOHRNIPHORA Dahl, 1898

Dohrniphora cornuta (Bigot, 1857)

GYMNOPHORA Macquart, 1835

Gymnophora arcuata (Meigen, 1830)

Gymnophora healeyae Disney, 1980

Gymnophora nigripennis Schmitz, 1926

Gymnophora quartomollis Schmitz, 1920

HYPOCERA Lioy, 1864

Hypocera mordellaria (Fallén, 1823)

MEGASELIA Rondani, 1856

= Aphiochaeta Brues, 1903

= Plastophora Brues, 1905

Megaselia aequalis (Wood, 1909)

Megaselia affinis (Wood, 1909)

= proxima (Lundbeck, 1920)

Megaselia albicaudata (Wood, 1910)

Megaselia albiclava (Schmitz, 1926)

Megaselia altifrons (Wood, 1909)

Megaselia analis (Lundbeck, 1920)

? Megaselia angelicae (Wood, 1910)

Megaselia angularis (Schmitz, 1924)

Megaselia angusta (Wood, 1909)

= angustata emend.

= pulicaria auct. nec (Fallén, 1923)

Megaselia annulipes (Schmitz, 1921)

Megaselia armata (Wood, 1909)

Megaselia atrosericea Schmitz, 1927

Megaselia auricoma Schmitz, 1927

Megaselia baltica (Schmitz, 1924)

Megaselia barbulata (Wood, 1909)

= depilata (Lundbeck, 1921)

? Megaselia basispinata (Lundbeck, 1920)

Megaselia basiveluta Schmitz, 1935

Megaselia beckeri (Wood, 1909)

= laticrus Schmitz, 1927

Megaselia berndseni (Schmitz, 1919)

= pygmaeoides (Lundbeck, 1921)

Megaselia bovista (Gimmerthal, 1848)

Megaselia brevicostalis (Wood, 1910)

Megaselia breviseta (Wood, 1912)

Megaselia breviterga (Lundbeck, 1920)

= similata (Lundbeck, 1921)

Megaselia campestris (Wood, 1908)

Megaselia ciliata (Zetterstedt, 1848)

Megaselia cinereifrons (Strobl, 1910)

Megaselia clara (Schmitz, 1921)

Megaselia coaetanea Schmitz, 1929

Megaselia coccyx Schmitz, 1965

? Megaselia collini (Wood, 1909)

Megaselia conformis (Wood, 1909)

Megaselia costalis (von Roser, 1840)

Megaselia curvivenia Schmitz, 1928

Megaselia dahli (Becker, 1901)

Megaselia discreta (Wood, 1909)

= nudiventris (Wood, 1909)

Megaselia diversa (Wood, 1909)

Megaselia dubitalis (Wood, 1908)

Megaselia eccoptomera Schmitz, 1927

Megaselia eisfelderae Schmitz, 1948

Megaselia elongata (Wood, 1914)

Megaselia erecta (Wood, 1910)

Megaselia exarcuata Schmitz, 1927

Megaselia excavata Schmitz, 1927

Megaselia excorticata Disney, 2009

Megaselia fennicola Beyer, 1958

Megaselia flammula Schmitz, 1928

Megaselia flava (Fallén, 1823)

Megaselia flavicoxa (Zetterstedt, 1848)

Megaselia fumata (Malloch, 1909)

Megaselia funeralis Schmitz, 1928

Megaselia fungivora (Wood, 1909)

Megaselia fusca (Wood, 1909)

Megaselia fuscipalpis (Lundbeck, 1920)

Megaselia fuscoides Schmitz, 1934

Megaselia fuscovariana Schmitz, 1933

Megaselia giraudii (Egger, 1862)

Megaselia glabrifrons (Wood, 1909)

Megaselia groenlandica (Lundbeck, 1901)

Megaselia halterata (Wood, 1910)

= plurispinosa (Lundbeck, 1920)

Megaselia hectochaeta Schmitz, 1974

Megaselia hilaris Schmitz, 1927

Megaselia hirsuta (Wood, 1910)

Megaselia hirticaudata (Wood, 1910)

Megaselia hirticrus (Schmitz, 1918)

Megaselia humeralis (Zetterstedt, 1838)

= cubitalis (Becker, 1901)

Megaselia hyalipennis (Wood, 1912)

Megaselia infraposita (Wood, 1909)

Megaselia ignobilis (Schmitz, 1919)

Megaselia intonsa Schmitz, 1948

Megaselia involuta (Wood, 1910)

Megaselia latifemorata (Becker, 1901)

Megaselia limburgensis (Schmitz, 1918)

Megaselia longicostalis (Wood, 1912)

Megaselia longifurca (Lundbeck, 1921)

= spinolabella Disney, 1989

Megaselia longipalpis (Wood, 1910)

Megaselia longiseta (Wood, 1909)

Megaselia lucifrons (Schmitz, 1918)

= subnitida (Lundbeck, 1920)

Megaselia luminifrons (Schmitz, 1926)

Megaselia lutea (Meigen, 1830)

Megaselia major (Wood, 1912)

Megaselia mallochi (Wood, 1909)

Megaselia manicata (Wood, 1910)

Megaselia maura (Wood, 1910)

Megaselia meconicera (Speiser, 1925)

Megaselia meigeni (Becker, 1901)

? Megaselia miki Schmitz, 1929

Megaselia minor (Zetterstedt, 1848)

= angustifrons (Wood, 1912)

Megaselia minuta (Aldrich, 1892)

= luminosa Schmitz, 1952

Megaselia mixta (Schmitz, 1918)

Megaselia nasoni (Malloch, 1914)

= coaequalis (Schmitz, 1919)

Megaselia nigra (Meigen, 1830)

= albidihalteralis (Felt, 1896)

Megaselia nigriceps (Loew, 1866)

= projecta (Becker, 1901)

Megaselia nigripalpis (Lundbeck, 1920)

Megaselia nudipleura (Beyer, 1958)

Megaselia obscuripennis (Wood, 1909)

Megaselia offuscata (Schmitz, 1921)

Megaselia opacicornis Schmitz, 1949

Megaselia parnassia Disney, 1986

Megaselia parva (Wood, 1909)

Megaselia pectoralis (Wood, 1910)

Megaselia pectunculata Schmitz, 1927

Megaselia picta (Lehmann, 1822)

Megaselia pleuralis (Wood, 1909)

Megaselia plurispinulosa (Zetterstedt, 1860)

Megaselia posticata (Strobl, 1898)

Megaselia praeacuta (Schmitz, 1919)

Megaselia prodroma (Lundbeck, 1921)

Megaselia producta (Schmitz, 1921)

= sordescens auct. nec Schmitz, 1927

Megaselia propinqua (Wood, 1909)

Megaselia pubecula Schmitz, 1927

Megaselia pulicaria (Fallén, 1923)

= sinuata Schmitz, 1926

Megaselia pumila (Meigen, 1830)

= atripes (Brues, 1915)

Megaselia pusilla (Meigen, 1830)

Megaselia pygmaea (Zetterstedt, 1848)

Megaselia quadriseta (Schmitz, 1918)

= badia Schmitz, 1938

? Megaselia raetica Schmitz, 1934

Megaselia robertsoni Disney, 2008

Megaselia robusta Schmitz, 1928

Megaselia rubricornis (Schmitz, 1919)

Megaselia rudis (Wood, 1909)

Megaselia ruficornis (Meigen, 1830)

Megaselia rufipes (Meigen, 1804)

Megaselia scalaris (Loew, 1866)

Megaselia scutellaris (Wood, 1909)

= scutellariformis (Schmitz, 1926)

Megaselia sepulchralis (Lundbeck, 1920)

Megaselia setulipalpis Schmitz, 1938

Megaselia sordida (Zetterstedt, 1838)

= eminens Schmitz, 1953

= semiscaura Schmitz, 1927

? Megaselia specularis Schmitz, 1935

Megaselia spinicincta (Wood, 1910)

Megaselia spinigera (Wood, 1908)

Megaselia styloprocta (Schmitz, 1921)

Megaselia subcarpalis (Lundbeck, 1920)

Megaselia subpalpalis (Lundbeck, 1920)

Megaselia subpleuralis (Wood, 1909)

Megaselia subtumida (Wood, 1909)

Megaselia sulphuripes (Meigen, 1830)

Megaselia superciliata (Wood, 1910)

Megaselia superfurcata Schmitz, 1928

Megaselia sylvatica (Wood, 1910)

= impolluta (Schmitz, 1920)

Megaselia tarsalis (Wood, 1910)

Megaselia tenuiventris Schmitz, 1927

Megaselia tignorum Disney, 2009

Megaselia tumida (Wood, 1909)

= setifer Lundbeck, 1920

Megaselia unicolor (Schmitz, 1919)

Megaselia valvata Schmitz, 1935

Megaselia variana Schmitz, 1926

Megaselia vestita (Wood, 1914)

Megaselia winqvisti Disney, 2011

Megaselia woodi (Lundbeck, 1922)

Megaselia xanthozona (Strobl, 1892)

= euryprocta Schmitz, 1957

Megaselia zonata (Zetterstedt, 1838)

MENOZZIOLA Schmitz, 1927

Menozziola obscuripes (Schmitz, 1927)

METOPINA Macquart, 1835

Metopina galeata (Haliday, 1833)

= inaequalis Schmitz, 1927

Metopina oligoneura (Mik, 1867)

Metopina pileata Schmitz, 1936

MICROSELIA Schmitz, 1934

Microselia forsiusi (Schmitz, 1927)

PHALACROTOPHORA Enderlein, 1912

Phalacrotophora berolinensis Schmitz, 1920

Phalacrotophora beuki Disney in Disney & Beuk, 1997

Phalacrotophora fasciata (Fallén, 1823)

PHORA Latreille, 1796

= Trineura Meigen, 1803

Phora artifrons Schmitz, 1920

Phora atra (Meigen, 1804)

= aterrima (Fabricius, 1794) preocc.

Phora bullata Schmitz, 1927

Phora convallium Schmitz, 1928

Phora convergens Schmitz, 1920

Phora dubia (Zetterstedt, 1848)

= schineri (Becker, 1901)

Phora edentata Schmitz, 1920

Phora hamata Schmitz, 1927

Phora holosericea Schmitz, 1920

Phora hyperborea Schmitz, 1927

Phora obscura (Zetterstedt, 1848)

Phora occidentata Malloch, 1912

= zetterstedti Schmitz, 1927

Phora penicillata Schmitz, 1920

Phora praepandens Schmitz, 1927

Phora pubipes Schmitz, 1920

Phora stictica Meigen, 1830

Phora tincta Schmitz, 1920

PLECTANOCNEMA Schmitz, 1926

Plectanocnema nudipes (Becker, 1901)

PSEUDACTEON Coquillett, 1907

Pseudacteon fennicus Schmitz, 1927

Pseudacteon formicarum (Verrall, 1877)

SPINIPHORA Malloch, 1909

Spiniphora bergenstammi Mik, 1864

Spiniphora dorsalis (Becker, 1901)

Spiniphora excisa (Becker, 1901)

Spiniphora maculata (Meigen, 1830)

= helicivora (Dufour, 1841)

TRIPHLEBA Rondani, 1856

= Citrago Schmitz, 1924

Triphleba admirabilis Schmitz, 1927

? Triphleba aequalis (Schmitz, 1919)

? Triphleba autumnalis (Becker, 1901)

Triphleba bicornuta (Strobl, 1910)

= uncinata (Schmitz, 1918)

Triphleba citreiformis (Becker, 1901)

Triphleba distinguenda (Strobl, 1892)

= unicalcarata (Becker, 1901)

Triphleba excisa (Lundbeck, 1921)

Triphleba gracilis (Wood, 1907)

Triphleba hyalinata (Meigen, 1830)

? Triphleba inaequalis Schmitz, 1943

Triphleba intermedia (Malloch, 1908)

Triphleba lugubris (Meigen, 1830)

Triphleba luteifemorata (Wood, 1906)

Triphleba nudipalpis (Becker, 1901)

Triphleba opaca (Meigen, 1830)

Triphleba pachyneurella (Schmitz, 1919)

Triphleba palposa (Zetterstedt, 1848)

Triphleba renidens Schmitz, 1927

Triphleba subcompleta Schmitz, 1927

Triphleba trinervis (Becker, 1901)

VERUANUS Schmitz, 1927

Veruanus oldenbergi Schmitz, 1919

= memorabilis Schmitz, 1927

Excluded species

Megaselia cirriventris Schmitz, 1929 mistake?

Megaselia crassipes (Wood, 1909) misidentified

Megaselia digitalis Schmitz, 1957 mistake?

Megaselia indifferens (Lundbeck, 1920) not found within present borders

Megaselia lapponica Schmitz, 1928 not found within present borders

Megaselia luteipes (Schmitz, 1918) misidentified

Megaselia palmeni (Becker, 1901) not found within present borders

Megaselia septentrionalis (Schmitz, 1919) not found within present borders

Megaselia simulans (Wood, 1912) not found within present borders ?

Triphleba gilvipes Schmitz, 1943 not found within present borders

Notes

Conicera tarsalis, Megaselia collini. Records of these species are based on single females identified by H. Schmitz.

Megaselia basispinata, M. miki, M. raetica. Species recorded from Finland by Hackman (1980) and Disney (1991) but no Finnish specimens or published records with details could be located.

Megaselia cirriventris, M. digitalis. These species were recorded from ‘Fennia’ and ‘Lappland’ by Schmitz (1958), but all specimens listed are from the Abisko region in northen Sweden. It seems likely that Schmitz erroneously assigned the material from Swedish Lappland to Finland.

Megaselia luteipes. Examined specimens belong to Megaselia sp. not luteipes (females) (Espoo, leg. W. Hackman) and possibly Megaselia sp. 24 of Disney (2013) (Finland, Muonio, leg. J. Sahlberg and Russia, Kola, Kantalahti, leg. R. Frey).

Megaselia pulicaria agg. This difficult complex of species was revised by Disney (1999). The Finnish material has not been re-examined. The Finnish species involved are M. angusta, M. costalis, M. longicostalis, M. longifurca, M. pulicaria, M. subtumida and M. tumida.

Megaselia simulans. Recorded only(?) from Kuusamo, a municipality split between Finland and Russia in 1944. Most records preceding the split are from the Russian side. No material was found in MZH.

Megaselia specularis. Recorded, with some doubt, from Finland by Hackman (1963). No material could be found in MZH.

Triphleba aequalis. Schmitz (1943) mention a Finnish male stored in MZH, but no such specimen could be found.

Triphleba autumnalis, T. inaequalis. Finnish according to Disney (1991), but no specimens or detailed records could be located.

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