An updated checklist of the Atelestidae, Brachystomatidae, Dolichopodidae, Empididae and Hybotidae (Diptera) recorded from Finland is presented. The genera with uncertain placement within superfamily Empidoidea (= the Iteaphila group) are also included in this paper.

Checklist, Finland, Diptera , Atelestidae , Brachystomatidae , Dolichopodidae , Hybotidae , Empididae

Empidoidea (dance flies and long-legged flies) is one of the most diverse lineages of Diptera with some 13,000 species currently recognized worldwide (Pape et al. 2011) and many more still undescribed. The superfamily is generally accepted as monophyletic, but its subdivision into families, subfamilies and tribes is still being debated.

The families Atelestidae, Brachystomatidae, Hybotidae and Empididae were traditionally all included in Empididaesensu lato. Chvála (1983) split the empidids into five families: Empididaesensu stricto, Hybotidae, Atelestidae, Dolichopodidae and Microphoridae. Sinclair and Cumming (2006) analyzed the whole superfamily Empidoidea, elevated Brachystomatidae to a full family and transferred Microphorinae to the long-legged flies (Dolichopodidae). Two of the Finnish genera, Anthepiscopus Becker and Iteaphila Zetterstedt, remain unassigned on family level (Sinclair and Cumming 2006, Shamshev and Sinclair 2009). The classification proposed by Sinclair and Cumming (2006) is adopted for this checklist.

The world fauna of the Empidoidea has recently been catalogued by Yang et al. (2006, 2007). The Finnish species were last listed by Kahanpää and Winqvist (2005) for Atelestidae, Brachystomatidae, Hybotidae and Empididae (plus Microphorinae) and by Kahanpää and Grichanov (2004) for Dolichopodidae. Two updates to the dolichopodid checklist have since been published (Kahanpää 2006, 2013).

suborder Brachycera Macquart, 1834

clade Eremoneura Lameere, 1906

superfamily Empidoidea Latreille, 1809

Uncertain family position within Empidoidea (incertae familiae)

ITEAPHILA Zetterstedt, 1838

Iteaphila macquarti Zetterstedt, 1838

Iteaphila nitidula Zetterstedt, 1838

= obscura Zetterstedt, 1849

ANTHEPISCOPUS Becker, 1891

Anthepiscopus oedalinus (Zetterstedt, 1838)

EMPIDIDAE Latreille, 1809

HEMERODROMIINAE Schiner, 1862

Chelipodini Hendel, 1936

CHELIPODA Macquart, 1823

Chelipoda albiseta (Zetterstedt, 1838)

Chelipoda inexspectata Tuomikoski, 1966

Chelipoda vocatoria (Fallén, 1816)

PHYLLODROMIA Zetterstedt, 1837

Phyllodromia melanocephala (Fabricius, 1794)

Hemerodromiini Schiner, 1862

CHELIFERA Macquart, 1823

Chelifera concinnicauda Collin, 1927

= lapponica Frey, 1950

= stigmatica misid.

Chelifera flavella (Zetterstedt, 1838)

Chelifera frigelii (Zetterstedt, 1838)

Chelifera precabunda Collin, 1961

Chelifera precatoria (Fallén, 1816)

Chelifera subangusta Collin, 1961

Chelifera trapezina (Zetterstedt, 1838)

HEMERODROMIA Meigen, 1822

Hemerodromia adulatoria Collin, 1927

Hemerodromia oratoria (Fallén, 1816)

Hemerodromia raptoria Meigen, 1830

EMPIDINAE Latreille, 1809

Empidini Latreille, 1809

EMPIS Linnaeus, 1758

sg. Anacrostichus Bezzi, 1909

Empis lucida Zetterstedt, 1838

sg. Coptophlebia Bezzi, 1909

Empis hyalipennis Fallén, 1816

sg. Empis Linnaeus, 1758

Empis acinerea Chvala, 1985

= cinerea Zetterstedt, 1855 preocc.

Empis bicuspidata Collin, 1927

Empis caudatula Loew, 1867

Empis chioptera Meigen, 1804

Empis laminata Collin, 1927

Empis nigripes Fabricius, 1794

= pennaria Fallén, 1816

= vernalis Meigen, 1822

Empis pennipes Linnaeus, 1758

Empis prodromus Loew, 1867

Empis staegeri Collin, 1963

= planetica misid.

Empis syrovatkai Chvála, 1985

= plumipes Zetterstedt, 1842 preocc.

sg. Euempis Frey, 1953

Empis picipes Meigen, 1804

= maculipes Zetterstedt, 1842

Empis tessellata Fabricius, 1794

sg. Kritempis Collin, 1926

Empis livida Linnaeus, 1758

sg. Leptempis Collin, 1926

Empis grisea Fallén, 1816

sg. Platyptera Meigen, 1803

Empis borealis Linnaeus, 1758

sg. Xanthempis Bezzi, 1909

Empis aemula Loew, 1873

Empis digramma Meigen in Gistl, 1835

= diagramma misspelling

Empis laetabilis Collin, 1926

Empis stercorea Linnaeus, 1761

Empis univittata Loew, 1867

RHAMPHOMYIA Meigen, 1822

sg. Aclonempis Collin, 1926

Rhamphomyia galactoptera Strobl, 1893

sg. Amydroneura Collin, 1926

Rhamphomyia gibba (Fallén, 1816)

sg. Holoclera Schiner, 1860

Rhamphomyia bohemica Barták & Kubík, 2012

= caliginosa misid.

= trigemina misid.

Rhamphomyia culicina (Fallén, 1816)

Rhamphomyia nigripennis (Fabricius, 1794)

Rhamphomyia sciarina (Fallén, 1816)

Rhamphomyia umbripennis Meigen, 1822

Rhamphomyia variabilis (Fallén, 1816)

= tenuirostris (Fallén, 1816)

sg. Lundstroemiella Frey, 1922

Rhamphomyia dudai Oldenberg, 1927

Rhamphomyia hybotina Zetterstedt, 1838

sg. Megacyttarus Bigot, 1880

Rhamphomyia anomalina Zetterstedt, 1838

Rhamphomyia anomalipennis Meigen, 1822

= anomala misid.

Rhamphomyia crassirostris (Fallén, 1816)

= nigripes auct. nec (Fabricius, 1794)

Rhamphomyia gufitar Frey, 1922

Rhamphomyia maculipennis Zetterstedt, 1842

= tephraea misid.

= poissoni misid.

Rhamphomyia nodipes (Fallén, 1816)

= spissirostris (Fallén, 1816)

Rhamphomyia paradoxa Wahlberg, 1844

sg. Pararhamphomyia Frey, 1922

Rhamphomyia albidiventris Strobl, 1898

= woldstedti Frey, 1913

Rhamphomyia albipennis (Fallén, 1816)

Rhamphomyia albitarsis Collin, 1926

Rhamphomyia albissima Frey, 1913

Rhamphomyia alpina Zetterstedt, 1838

Rhamphomyia amoena Loew, 1840

Rhamphomyia angulifera Frey, 1913

Rhamphomyia breviventris Frey, 1913

Rhamphomyia caesia Meigen, 1822

= filata Zetterstedt, 1842

Rhamphomyia caudata Zetterstedt, 1838

= aethiops Zetterstedt, 1838

Rhamphomyia chibinensis Frey, 1922

Rhamphomyia cribrata Oldenberg, 1927

Rhamphomyia curvula Frey, 1913

Rhamphomyia dentata Oldenberg, 1910

Rhamphomyia fascipennis Zetterstedt, 1838

Rhamphomyia filicaudula Frey, 1949

Rhamphomyia fuscipennis Zetterstedt, 1838

Rhamphomyia fuscula Zetterstedt, 1838

Rhamphomyia geniculata Meigen, 1830

= plumipes auct. nec (Meigen, 1804)

Rhamphomyia griseola (Zetterstedt, 1838)

= dispar Zetterstedt, 1838

= aperta Zetterstedt, 1859

Rhamphomyia helleni Frey, 1922

Rhamphomyia lividiventris Zetterstedt, 1838

Rhamphomyia longestylata Frey, 1916

= caudata auct. nec Zetterstedt, 1838

Rhamphomyia lucidula Zetterstedt, 1842

Rhamphomyia marginata (Fabricius, 1787)

Rhamphomyia modesta Wahlberg, 1844

Rhamphomyia murina Collin, 1926

Rhamphomyia niveipennis Zetterstedt, 1838

Rhamphomyia obscura Zetterstedt, 1838

Rhamphomyia obscuripennis Meigen, 1830

= nitidicollis Frey, 1913

Rhamphomyia physoprocta Frey, 1913

Rhamphomyia pilifer Meigen, 1838

= dentipes Zetterstedt, 1842

= glaucella Zetterstedt, 1842

= intermedia Frey, 1922

Rhamphomyia plumifera Zetterstedt, 1838

Rhamphomyia poplitea Wahlberg, 1844

Rhamphomyia praestans Frey, 1913

Rhamphomyia pusilla Zetterstedt, 1838

Rhamphomyia rufipes Zetterstedt, 1838

= lapponica Frey, 1955

Rhamphomyia simplex Zetterstedt, 1849

Rhamphomyia subglaucella Frey, 1922

Rhamphomyia tibiella Zetterstedt, 1842

Rhamphomyia tipularia (Fallén, 1816)

Rhamphomyia unguiculata Frey, 1913

Rhamphomyia sp. aff. albipennis

sg. Rhamphomyia Meigen, 1822

Rhamphomyia albosegmentata Zetterstedt, 1838

Rhamphomyia cinerascens (Meigen, 1804)

Rhamphomyia coracina Zetterstedt, 1849

Rhamphomyia dorsata Becker, 1915

= phanerostigma Frey, 1918

Rhamphomyia hambergi Frey, 1916

Rhamphomyia ignobilis Zetterstedt, 1859

= attenuata Frey, 1913

Rhamphomyia laevipes (Fallén, 1816)

= tephraea Meigen, 1822

Rhamphomyia latifrons Frey, 1913

Rhamphomyia morio Zetterstedt, 1838

Rhamphomyia nitidula Zetterstedt, 1842

Rhamphomyia palmeni Frey, 1913

Rhamphomyia plumipes (Meigen, 1804)

= vespertilio Zetterstedt, 1842

Rhamphomyia reflexa Zetterstedt, 1838

Rhamphomyia spinipes (Fallén, 1816)

Rhamphomyia stigmosa Macquart, 1827

= conformis auct. nec Kowarz, 1867

Rhamphomyia sulcata (Meigen, 1804)

= tibialis auct. nec Meigen, 1822

Rhamphomyia trilineata Zetterstedt, 1859

= sulcatina Collin, 1926

= tibialis auct. nec Meigen, 1822

= propinqua misid.

Rhamphomyia vesiculosa (Fallén, 1816)

Hilarini Collin, 1961

HILARA Meigen, 1822

Hilara abdominalis Zetterstedt, 1838

= obscuritarsis Zetterstedt, 1859

Hilara albitarsis von Roser, 1840

Hilara anglodanica Lundbeck, 1913

Hilara barbipes Frey, 1908

Hilara beckeri Strobl, 1892

Hilara biseta Collin, 1927

Hilara bistriata Zetterstedt, 1842

Hilara brevistyla Collin, 1927

Hilara campinosensis Niesiolowski, 1986

Hilara canescens Zetterstedt, 1849

Hilara chorica (Fallén, 1816)

Hilara clavipes (Harris, 1776)

= spinimana Zetterstedt, 1838

Hilara clypeata Meigen, 1822

Hilara coracina Oldenberg, 1916

= quadrifaria Strobl, 1892

Hilara cornicula Loew, 1873

Hilara discoidalis Lundbeck, 1910

Hilara diversipes Strobl, 1892

= germanica Engel, 1941

Hilara eviana Straka, 1976

Hilara femorella Zetterstedt, 1842

Hilara gallica (Meigen, 1804)

Hilara griseola Zetterstedt, 1838

Hilara hirta Strobl, 1892

= hirtella misid.

Hilara hybrida Collin, 1961

Hilara hyposeta Straka, 1976

Hilara implicata Collin, 1927

Hilara intermedia (Fallén, 1816)

= pubipes Loew, 1873

Hilara interstincta (Fallén, 1816)

Hilara monedula Collin, 1927

= longifurca auct. nec Strobl, 1892

Hilara lapponica Chvála, 2002

Hilara litorea (Fallén, 1816)

Hilara longivittata Zetterstedt, 1842

Hilara lurida (Fallén, 1816)

Hilara nigritarsis Zetterstedt, 1838

= infans Zetterstedt, 1842

Hilara nitidula Zetterstedt, 1838

Hilara pilipes Zetterstedt, 1838

Hilara pseudochorica Strobl, 1892

Hilara pulchripes Frey, 1913

Hilara quadrifasciata Chvála, 2002

= quadrivittata auct. nec Meigen, 1822

Hilara sturmii Meigen, 1822

= cingulata misid.

Hilara submaura Collin, 1927

Hilara tanythrix Frey, 1913

Hilara tenuinervis Zetterstedt, 1838

Hilara woodiella Chvála, 1999

CLINOCERINAE Collin, 1928

CLINOCERA Meigen, 1803

Clinocera appendiculata (Zetterstedt, 1838)

= simplicinervis Frey, 1913

Clinocera aucta (Zetterstedt, 1849)

Clinocera nivalis (Zetterstedt, 1838)

Clinocera stagnalis (Haliday, 1833)

Clinocera wesmaeli (Macquart, 1835)

DOLICHOCEPHALA Macquart, 1823

Dolichocephala guttata (Haliday, 1833)

Dolichocephala irrorata (Fallén, 1815)

Dolichocephala thomasi Wagner, 1983

= ocellata misid.

WIEDEMANNIA Zetterstedt, 1838

sg. Eucelidia Mik, 1881

Wiedemannia zetterstedti (Fallén, 1826)

sg. Philolutra Mik, 1881

Wiedemannia bohemani (Zetterstedt, 1838)

Wiedemannia simplex (Loew, 1862)

= fallaciosa auct. nec Loew, 1873

sg. Wiedemannia Zetterstedt, 1838

Wiedemannia bistigma (Curtis, 1834)

Uncertain subfamily position within Empididae (incertae sedis)

HORMOPEZA Zetterstedt, 1838

Hormopeza copulifera Melander, 1927

Hormopeza obliterata Zetterstedt, 1838

RAGAS Walker, 1837

Ragas unica Walker, 1837

ATELESTIDAE Hennig, 1970

ATELESTINAE Hennig, 1970

ATELESTUS Walker, 1837

Atelestus pulicarius (Fallén, 1816)

MEGHYPERUS Loew, 1850

Meghyperus sudeticus Loew, 1850

HYBOTIDAE Meigen, 1820

TRICHININAE Chvála, 1983

TRICHINA Meigen, 1830

Trichina bilobata Collin, 1926

Trichina clavipes Meigen, 1830

Trichina elongata Haliday, 1833

Trichina opaca Loew, 1864

= picipes Tuomikoski, 1935

Trichina pallipes (Zetterstedt, 1838)

TRICHINOMYIA Tuomikoski, 1959

Trichinomyia flavipes (Meigen, 1830)

Trichinomyia fuscipes (Zetterstedt, 1838)

OCYDROMIINAE Schiner, 1862

Ocydromiini Schiner, 1862

CHVALAEA Papp & Földvári, 2001

Chvalaea sopianae Papp & Földvári, 2001

LEPTODROMIELLA Tuomikoski, 1936

Leptodromiella crassiseta Tuomikoski, 1936

LEPTOPEZA Macquart, 1834

Leptopeza borealis Zetterstedt, 1842

Leptopeza flavipes (Meigen, 1820)

OCYDROMIA Meigen, 1820

Ocydromia glabricula (Fallén, 1816)

Ocydromia melanopleura Loew, 1840

OEDALEINAE Chvála, 1983

ALLANTHALIA Melander, 1927

Allanthalia pallida (Zetterstedt, 1838)

ANTHALIA Zetterstedt, 1838

Anthalia schoenherri Zetterstedt, 1838

EUTHYNEURA Macquart, 1836

Euthyneura albipennis (Zetterstedt, 1842)

Euthyneura gyllenhali (Zetterstedt, 1838)

Euthyneura myrtilli Macquart, 1836

= myricae Haliday, 1851 misid.

OEDALEA Meigen, 1820

Oedalea freyi Chvála, 1983

Oedalea holmgreni Zetterstedt, 1852

Oedalea hybotina (Fallén, 1816)

Oedalea stigmatella Zetterstedt, 1842

Oedalea tibialis Macquart, 1827

Oedalea zetterstedti Collin, 1926

TACHYDROMIINAE Meigen, 1822

Symballophthalmini Sinclair & Cumming, 2006

SYMBALLOPHTHALMUS Mecker, 1889

Symballophthalmus dissimilis (Fallén, 1815)

Symballophthalmus fuscitarsis (Zetterstedt, 1859)

= scapularis Collin, 1961

Tachydromiini Meigen, 1822

PLATYPALPUS Macquart, 1827

Platypalpus agilis (Meigen, 1822)

Platypalpus albicornis (Zetterstedt, 1842)

Platypalpus albiseta (Panzer, 1806)

Platypalpus albocapillatus (Fallén, 1815)

Platypalpus alpinus Chvála, 1971

Platypalpus alter (Collin, 1961)

Platypalpus annulatus (Fallén, 1815)

= fulvipes (Meigen, 1822)

Platypalpus annulipes (Meigen, 1822)

Platypalpus articulatoides (Frey, 1918)

? Platypalpus articulatus Macquart, 1827

Platypalpus ater (Wahlberg, 1844)

= atra misspelling

Platypalpus boreoalpinus Frey, 1943

Platypalpus brachystylus (Bezzi, 1892)

= brunneitibia (Strobl, 1899)

Platypalpus brevicornis (Zetterstedt, 1842)

Platypalpus calceatus (Meigen, 1822)

Platypalpus candicans (Fallén, 1815)

Platypalpus ciliaris (Fallén, 1816)

Platypalpus confiformis Chvála, 1971

Platypalpus confinis (Zetterstedt, 1842)

Platypalpus cothurnatus Macquart, 1827

Platypalpus cryptospina (Frey, 1909)

= tantulus (Collin, 1961)

Platypalpus cursitans (Fabricius, 1775)

= bicolor (Meigen, 1804)

Platypalpus ecalceatus (Zetterstedt, 1838)

Platypalpus excavatus Yang & Yao, 2007

= excisus (Becker, 1907) preocc.

Platypalpus exilis (Meigen, 1822)

Platypalpus fenestella Kovalev, 1971

Platypalpus flavicornis (Meigen, 1822)

Platypalpus fuscicornis (Zetterstedt, 1842)

Platypalpus hackmani Chvála, 1972

Platypalpus infectus (Collin, 1926)

Platypalpus interstinctus (Collin, 1926)

Platypalpus laestadianorum (Frey, 1913)

Platypalpus lapponicus Frey, 1943

Platypalpus longicornis (Meigen, 1822)

Platypalpus longiseta (Zetterstedt, 1842)

= extricatus (Collin, 1926)

Platypalpus luteicornis (Meigen, 1838)

= difficilis (Frey, 1907)

= interjectus (Lundbeck, 1910)

Platypalpus luteus (Meigen, 1804)

Platypalpus maculus (Zetterstedt, 1842)

Platypalpus maculimanus (Zetterstedt, 1842)

= articulatus auct. nec Macquart, 1827

Platypalpus maculipes (Meigen, 1822)

Platypalpus major (Zetterstedt, 1842)

Platypalpus melancholicus (Collin, 1961)

Platypalpus minutus (Meigen, 1804)

Platypalpus nigricoxa (Mik, 1884)

Platypalpus nigritarsis (Fallén, 1816)

Platypalpus nigrosetosus (Strobl, 1893)

Platypalpus nonstriatus Strobl, 1901

Platypalpus notatus (Meigen, 1822)

Platypalpus pallidicornis (Collin, 1926)

Platypalpus pallidicoxa (Frey, 1913)

Platypalpus pallidiventris (Meigen, 1822)

= flavipes (Fabricius, 1794)

Platypalpus pallipes (Fallén, 1815)

Platypalpus pectoralis (Fallén, 1815)

Platypalpus pseudofulvipes (Frey, 1909)

= coarctatus (Collin, 1926)

Platypalpus pseudorapidus Kovalev, 1971

Platypalpus pulicarius (Meigen, 1830)

Platypalpus rapidus (Meigen, 1822)

Platypalpus sahlbergi (Frey, 1909)

Platypalpus scandinavicus Chvála, 1972

Platypalpus sordidus (Zetterstedt, 1838)

Platypalpus stabilis (Collin, 1961)

Platypalpus stackelbergi Kovalev, 1971

Platypalpus stigmatellus (Zetterstedt, 1842)

Platypalpus strigifrons (Zetterstedt, 1849)

Platypalpus subbrevis (Frey, 1913)

Platypalpus subtilis (Collin, 1926)

Platypalpus sylvicola (Collin, 1926)

Platypalpus tuomikoskii Chvála, 1972

Platypalpus unguiculatus (Zetterstedt, 1838)

Platypalpus verralli (Collin, 1926)

Platypalpus vividus (Meigen, 1838)

= albisetoides Chvála, 1973

Platypalpus zetterstedti Chvála, 1971

TACHYDROMIA Meigen, 1803

Tachydromia aemula (Loew, 1864)

Tachydromia arrogans (Linnaeus, 1761)

Tachydromia connexa Meigen, 1822

Tachydromia incompleta (Becker, 1900)

Tachydromia lundstroemi (Frey, 1913)

Tachydromia morio (Zetterstedt, 1838)

Tachydromia punctifera (Becker, 1900)

Tachydromia sabulosa Meigen, 1830

Tachydromia terricola Zetterstedt, 1819

Tachydromia umbrarum Haliday, 1833

TACHYPEZA Meigen, 1830

Tachypeza fennica Tuomikoski, 1932

Tachypeza fuscipennis (Fallén, 1815)

Tachypeza heeri (Zetterstedt, 1838)

Tachypeza nubila (Meigen, 1804)

Tachypeza truncorum (Fallén, 1815)

Tachypeza winthemi (Zetterstedt, 1838)

Drapetini Collin, 1961

CHERSODROMIA Haliday, 1851

Chersodromia arenaria (Haliday, 1833)

Chersodromia cursitans (Zetterstedt, 1819)

CROSSOPALPUS Bigot, 1857

Crossopalpus curvinervis (Zetterstedt, 1842)

Crossopalpus curvipes (Meigen, 1822)

Crossopalpus humilis (Frey, 1913)

Crossopalpus nigritellus (Zetterstedt, 1842)

Crossopalpus setiger (Loew, 1859)

DRAPETIS Meigen, 1822

Drapetis arcuata Loew, 1859

Drapetis assimilis (Fallén, 1815)

Drapetis exilis Meigen, 1822

Drapetis infitialis Collin, 1961

Drapetis ingrica Kovalev, 1972

Drapetis parilis Collin, 1926

Drapetis pusilla Loew, 1859

Drapetis simulans Collin, 1961

Elaphropeza Macquart, 1827

Elaphropeza ephippiata (Fallén, 1815)

STILPON Loew, 1859

Stilpon graminum (Fallén, 1815)

HYBOTINAE Meigen, 1820

Bicellariini Sinclair & Cumming, 2006

BICELLARIA Macquart, 1823

= Cyrtoma Meigen, 1824

Bicellaria austriaca Tuomikoski, 1955

Bicellaria intermedia Lundbeck, 1910

Bicellaria nigra (Meigen, 1824)

Bicellaria pilosa Lundbeck, 1910

Bicellaria simplicipes (Zetterstedt, 1842)

Bicellaria spuria (Fallén, 1816)

Bicellaria subpilosa Collin, 1926

Bicellaria sulcata (Zetterstedt, 1842)

Bicellaria uvens Melander, 1928

= bisetosa Tuomikoski, 1936

Hybotini Meigen, 1820

HYBOS Meigen, 1803

Hybos culiciformis (Fabricius, 1775)

Hybos femoratus (Müller, 1776)

= femoralis misspelling

Hybos grossipes (Linnaeus, 1767)

SYNDYAS Loew, 1857

Syndyas nigripes (Zetterstedt, 1842)

BRACHYSTOMATIDAE Melander, 1908

TRICHOPEZINAE Vaillant, 1981

GLOMA Meigen, 1822

Gloma fuscipennis Meigen, 1822

HELEODROMIA Haliday, 1833

sg. Heleodromia Haliday, 1833

Heleodromia immaculata Haliday, 1833

TRICHOPEZA Rondani, 1856

Trichopeza albocincta (Boheman, 1863)

= albicincta Frey, 1913

Trichopeza longicornis (Meigen, 1822)

DOLICHOPODIDAE Latreille, 1809

MICROPHORINAE Collin, 1960

MICROPHOR Macquart, 1827

= Microphorus misspelling

Microphor anomalus (Meigen, 1824)

Microphor crassipes Macquart, 1827

Microphor holosericeus (Meigen, 1804)

= velutinus Macquart, 1827

PARATHALASSIINAE Chvála, 1981

MICROPHORELLA Becker, 1909

Microphorella praecox (Loew, 1864)

DOLICHOPODINAE Latreille, 1809

DOLICHOPUS Latreille, 1796

= Hygroceleuthus Loew, 1857

sg. Dolichopus Latreille, 1796

Dolichopus acuticornis Wiedemann, 1817

Dolichopus annulitarsis Ringdahl, 1920

Dolichopus apicalis Zetterstedt, 1849

Dolichopus argyrotarsis Wahlberg, 1850

Dolichopus armillatus Wahlberg, 1850

= stenhammari var. b Zetterstedt, 1843

Dolichopus atripes Meigen, 1824

Dolichopus austriacus Parent, 1927

Dolichopus bonsdorffi Frey, 1915

Dolichopus brevipennis Meigen, 1824

Dolichopus caligatus Wahlberg, 1850

= flavipes misid.

= albifrons misid.

Dolichopus calinotus Loew, 1871

Dolichopus campestris Meigen, 1824

Dolichopus cilifemoratus Macquart, 1827

= pseudocilifemoratus Stackelberg, 1930

Dolichopus cinctipes Wahlberg, 1850

Dolichopus claviger Stannius, 1831

Dolichopus clavipes Haliday, 1832

Dolichopus costalis Frey, 1915

Dolichopus discifer Stannius, 1831

?= nigricornis Meigen, 1824

Dolichopus discimanus Wahlberg, 1851

Dolichopus fraterculus Zetterstedt, 1843

Dolichopus griseipennis Stannius, 1831

Dolichopus gubernator Mik, 1878

Dolichopus hilaris Loew, 1862

Dolichopus lancearius Hedström, 1966

Dolichopus latilimbatus Macquart, 1827

Dolichopus latipennis Fallén, 1823

Dolichopus lepidus Staeger, 1842

= cruralis Wahlberg, 1850

= lapponicus Becker, 1917

Dolichopus linearis Meigen, 1824

Dolichopus lineatocornis Zetterstedt, 1843

Dolichopus longicornis Stannius, 1831

Dolichopus longitarsis Stannius, 1831

Dolichopus maculipennis Zetterstedt, 1843

Dolichopus mannerheimi Zetterstedt, 1838

Dolichopus migrans Zetterstedt, 1843

Dolichopus nigripes Fallén, 1823

Dolichopus nitidus Fallén, 1823

Dolichopus notatus Staeger, 1842

= notabilis Zetterstedt, 1843

Dolichopus nubilus Meigen, 1824

Dolichopus pennatus Meigen, 1824

Dolichopus picipes Meigen, 1824

= consimilis Wahlberg, 1850

Dolichopus planitarsis Fallén, 1823

Dolichopus plumipes (Scopoli, 1763)

= parvicaudatus Zetterstedt, 1843

= pectinitarsis Stenhammar, 1851

Dolichopus popularis Wiedemann, 1817

Dolichopus pseudomigrans Ringdahl, 1928

Dolichopus punctum Meigen, 1824

Dolichopus remipes Wahlberg, 1839

Dolichopus rupestris Haliday, 1833

Dolichopus ruthei Loew, 1847

Dolichopus sabinus Haliday, 1838

Dolichopus setiger Negrobov, 1973

Dolichopus signatus Meigen, 1824

Dolichopus signifer Haliday, 1832

Dolichopus simplex Meigen, 1824

Dolichopus stenhammari Zetterstedt, 1843

= annulipes Zetterstedt, 1838 invalidated

Dolichopus subpennatus d’Assis-Fonseca, 1976

Dolichopus trivialis Haliday, 1832

= cilifemoratus auct. nec Macquart, 1827

Dolichopus ungulatus (Linnaeus, 1758)

Dolichopus urbanus Meigen, 1824

Dolichopus vitripennis Meigen, 1824

Dolichopus wahlbergi Zetterstedt, 1843

Dolichopus zetterstedti Stenhammar, 1852

sg. Macrodolichopus Stackelberg, 1933

Dolichopus diadema Haliday, 1832

ETHIROMYIA Brooks, 2005

Ethiromyia chalybeus (Wiedemann, 1817)

GYMNOPTERNUS Loew, 1857

Gymnopternus aerosus (Fallén, 1823)

Gymnopternus angustifrons (Staeger, 1842)

Gymnopternus brevicornis (Staeger, 1842)

Gymnopternus celer (Meigen, 1824)

Gymnopternus metallicus (Stannius, 1831)

HERCOSTOMUS Loew, 1857

Hercostomus germanus (Wiedemann, 1817)

Hercostomus nigrilamellatus (Macquart, 1827)

Hercostomus nigriplantis (Stannius, 1831)

Hercostomus sahlbergi (Zetterstedt, 1838)

SYBISTROMA Meigen, 1824

Sybistroma discipes (Germar, 1817)

Sybistroma obscurellum (Fallén, 1823)

TACHYTRECHUS Haliday, 1851

= Ammobates Stannius, 18312 preocc.

Tachytrechus ammobates (Haliday, 1851)

= plumipes (Fallén, 1823) preocc.

Tachytrechus hamatus Loew, 1871

Tachytrechus notatus (Stannius, 1831)

SCIAPODINAE Becker, 1917

SCIAPUS Zeller, 1842

= Sciopus misspelling

Sciapus albifrons (Meigen, 1830)

Sciapus basilicus Meuffels & Grootaert, 1990

Sciapus lobipes (Meigen, 1824)

Sciapus longulus (Fallén, 1823)

Sciapus maritimus Becker, 1918

= flavomaculatus Ringdahl, 1949

Sciapus platypterus (Fabricius, 1805)

Sciapus wiedemanni (Fallén, 1823)

Sciapus zonatulus (Zetterstedt, 1843)

= contristans misid.

SYMPYCNINAE Aldrich, 1905

CAMPSICNEMUS Haliday, 1851

= Ectomus Mik, 1878

Campsicnemus alpinus (Haliday, 1833)

Campsicnemus armatus (Zetterstedt, 1849)

Campsicnemus articulatellus (Zetterstedt, 1843)

= pilosellus (Zetterstedt, 1843)

= dasycnemus Loew, 1857

Campsicnemus compeditus Loew, 1857

Campsicnemus curvipes (Fallén, 1823)

Campsicnemus femoratus Ringdahl, 1949

Campsicnemus loripes (Haliday, 1832)

Campsicnemus lumbatus Loew, 1857

Campsicnemus marginatus Loew, 1857

Campsicnemus paradoxus (Wahlberg, 1844)

Campsicnemus picticornis (Zetterstedt, 1843)

Campsicnemus pumilio (Zetterstedt, 1843)

= pectinulatus Loew, 1864

Campsicnemus pusillus (Meigen, 1824)

Campsicnemus scambus (Fallén, 1823)

LAMPROCHROMUS Mik, 1878

Lamprochromus strobli Parent, 1925

SYMPYCNUS Loew, 1857

Sympycnus aeneicoxa (Meigen, 1824)

Sympycnus pulicarius (Fallén, 1823)

?= annulipes (Meigen, 1824)

?= desoutteri Parent, 1925

SYNTORMON Loew, 1857

= Bathycranium Strobl, 1892

Syntormon bicolorellus (Zetterstedt, 1843)

Syntormon filiger Verrall, 1912

= rufipes misid.

= simplicipes Frey, 1915

Syntormon freymuthae Loew, 1873

= S. denticulatus misid.

Syntormon metathesis (Loew, 1850)

Syntormon pallipes (Fabricius, 1794)

Syntormon pumilus (Meigen, 1824)

Syntormon tarsatus (Fallén, 1823)

= aulicus misid.

TELMATURGUS Mik, 1874

Telmaturgus tumidulus (Raddatz, 1873)

TEUCHOPHORUS Loew, 1857

= Teucophorus misspelling

Teuchophorus monacanthus Loew, 1859

Teuchophorus nigricosta (von Roser, 1840)

= pectinifer Kowarz, 1868

= signatus Zetterstedt, 1849

Teuchophorus spinigerellus (Zetterstedt, 1843)

DIAPHORINAE Schiner, 1864

ARGYRA Macquart, 1834

= Leucostola Loew, 1857

Argyra argentina (Meigen, 1824)

Argyra argyria (Meigen, 1824)

Argyra auricollis (Meigen, 1824)

Argyra diaphana (Fabricius, 1775)

Argyra elongata (Zetterstedt, 1843)

Argyra ilonae Geoffries, 1989

= confinis (Zetterstedt, 1849) preocc.

Argyra leucocephala (Meigen, 1824)

Argyra magnicornis (Zetterstedt, 1838)

Argyra setulipes Becker, 1918

Argyra setimana Loew, 1859

= subarctica misid.

Argyra spoliata Kowarz, 1879

Argyra vestita (Wiedemann, 1817)

ASYNDETUS Loew, 1869

Asyndetus latifrons (Loew, 1857)

CHRYSOTUS Meigen, 1824

Chrysotus angulicornis Kowarz, 1874

Chrysotus cilipes Meigen, 1824

Chrysotus cupreus (Macquart, 1827)

Chrysotus femoratus Zetterstedt, 1843

Chrysotus gramineus (Fallén, 1823)

= microcerus Kowarz, 1874

= varians Kowarz, 1874

Chrysotus laesus (Wiedemann, 1817)

= amplicornis Zetterstedt, 1849

Chrysotus neglectus (Wiedemann, 1817)

Chrysotus obscuripes Zetterstedt, 1838

= kowarzi Lundbeck, 1912

Chrysotus pulchellus Kowarz, 1874

Chrysotus suavis Loew, 1857

DIAPHORUS Meigen, 1824

Diaphorus hoffmannseggi Meigen, 1830

Diaphorus nigricans Meigen, 1824

Diaphorus oculatus (Fallén, 1823)

MELANOSTOLUS Kowarz, 1884

Melanostolus melancholicus (Loew, 1869)

MEDETERINAE Lioy, 1864

DOLICHOPHORUS Lichtward, 1902

Dolichophorus kerteszi Lichtward, 1902

MEDETERA Fischer von Waldheim, 1819

Medetera abstrusa Thuneberg, 1955

Medetera acanthura Negrobov & Thuneberg, 1970

Medetera adjaniae Gosseries, 1989

= breviseta Parent, 1927 preocc.

Medetera ambigua (Zetterstedt, 1843)

Medetera apicalis (Zetterstedt, 1843)

Medetera belgica Parent, 1936 sensu Negrobov & Stackelberg, 1972

Medetera betulae Ringdahl, 1949

Medetera borealis Thuneberg, 1955

Medetera cuspidata Collin, 1941

Medetera dichrocera Kowarz, 1877

Medetera excellens Frey, 1909

Medetera fasciata Frey, 1915

Medetera freyi Thuneberg, 1955

Medetera fumida Negrobov, 1967

Medetera impigra Collin, 1941

Medetera incrassata Frey, 1909

Medetera infumata Loew, 1857

Medetera inspissata Collin, 1952

Medetera jacula (Fallén, 1823)

Medetera jugalis Collin, 1941

Medetera melancholica Lundbeck, 1912

Medetera muralis Meigen, 1824

Medetera nitida (Macquart, 1834)

= stackelbergi Parent, 1927

Medetera obscura (Zetterstedt, 1838)

Medetera pallipes (Zetterstedt, 1843)

Medetera parenti Stackelberg, 1925

= collini Thuneberg, 1955

Medetera pinicola Kowarz, 1877

= nuortevai Thuneberg, 1955

Medetera plumbella Meigen, 1824

Medetera prjachinae Negrobov & Stackelberg, 1974

Medetera protuberans Negrobov, 1967

Medetera pseudoapicalis Thuneberg, 1955

Medetera seguyi Parent, 1926

Medetera senicula Kowarz, 1877

Medetera setiventris Thuneberg, 1955

Medetera signaticornis Loew, 1857

Medetera striata Parent, 1927

Medetera tristis (Zetterstedt, 1838)

Medetera vagans Becker, 1917

= fennica Thuneberg, 1955

Medetera veles Loew, 1861

= bilineata Frey, 1915

Medetera zinovjevi Negrobov, 1967

SYSTENUS Loew, 1857

Systenus bipartitus (Loew, 1850)

Systenus pallipes (von Roser, 1840)

= adpropinquus (Loew, 1857)

Systenus scholtzi (Loew, 1850)

THRYPTICUS Gerstäcker, 1864

Thrypticus atomus Frey, 1915

Thrypticus bellus Loew, 1869

Thrypticus cuneatus (Becker, 1917)

Thrypticus divisus Strobl, 1880

= fennicus Becker, 1917

Thrypticus intercedens Negrobov, 1967

Thrypticus laetus Verrall, 1912

Thrypticus nigricauda Wood, 1913

Thrypticus pollinosus Verrall, 1912

Thrypticus pruinosus Parent, 1932

Thrypticus smaragdinus Gerstäcker, 1864

Thrypticus tarsalis Parent, 1932

Thrypticus sp. A

HYDROPHORINAE Lioy, 1864

HYDROPHORUS Fallén, 1823

Hydrophorus albiceps Frey, 1915

Hydrophorus alpinus Wahlberg, 1844

Hydrophorus altivagus Aldrich, 1911

= wahlgreni Frey, 1915

Hydrophorus bipunctatus (Lehmann, 1822)

Hydrophorus borealis Loew, 1857

Hydrophorus brunnicosus Loew, 1857

Hydrophorus callosoma Frey, 1915

= albosignatus Ringdahl, 1919

Hydrophorus freyi Storå, 1954

Hydrophorus geminus Frey, 1915

Hydrophorus litoreus Fallén, 1823

Hydrophorus nebulosus Fallén, 1823

Hydrophorus norvegicus Ringdahl, 1928

Hydrophorus pectinatus Gerstäcker, 1864

= forcipatus Frey, 1915

Hydrophorus pilipes Frey, 1915

Hydrophorus praecox (Lehmann, 1822)

Hydrophorus rufibarbis Gerstäcker, 1864

= micans Frey, 1915

Hydrophorus signiferus Coquillett, 1899

= magnicornis Frey, 1915

Hydrophorus viridis (Meigen, 1824)

LIANCALUS Loew, 1857

Liancalus virens (Scopoli, 1763)

PEODES Loew, 1857

Peodes petsamoensis Frey, 1930

SCELLUS Loew, 1857

Scellus spinimanus (Zetterstedt, 1843)

THINOPHILUS Wahlberg, 1844

sg. Thinophilus Wahlberg, 1844

Thinophilus flavipalpis (Zetterstedt, 1843)

Thinophilus ruficornis (Haliday, 1838)

NEURIGONINAE Aldrich, 1905

NEURIGONA Rondani, 1856

Neurigona abdominalis (Fallén, 1823)

Neurigona pallida (Fallén, 1823)

Neurigona quadrifasciata (Fabricius, 1781)

Neurigona suturalis (Fallén, 1823)

Neurigona sp. A

RHAPHIINAE Bigot, 1852

RHAPHIUM Meigen, 1803

= Xiphandrium Loew, 1857

= Porphyrops auctt.

Rhaphium albifrons (Zetterstedt, 1843)

Rhaphium appendiculatum (Zetterstedt, 1849)

= macrocerum auct. nec Meigen, 1824

Rhaphium basale Loew, 1850

Rhaphium caliginosum Meigen, 1824

= zetterstedti (Parent, 1925)

Rhaphium commune (Meigen, 1824)

= spinicoxa Loew, 1850

Rhaphium confine Zetterstedt, 1843

Rhaphium crassipes (Meigen, 1824)

Rhaphium discolor Zetterstedt, 1838

= consobrinum Zetterstedt, 1843

= riparium auct. nec (Meigen, 1824: 53)

Rhaphium elegantulum (Meigen, 1824)

Rhaphium fasciatum Meigen, 1824

Rhaphium fascipes (Meigen, 1824)

Rhaphium fissum Loew, 1850

Rhaphium glaciale (Ringdahl, 1920)

Rhaphium holmgreni (Mik, 1878)

= luteipenne (Frey, 1915)

Rhaphium lanceolatum Loew, 1850

= caliginosum auct. nec Meigen, 1824

Rhaphium laticorne (Fallén, 1823)

= nemorum Meigen, 1830

Rhaphium latimanum Kahanpää, 2007

Rhaphium longicorne (Fallén, 1823)

Rhaphium micans (Meigen, 1824)

Rhaphium monotrichum Loew, 1850

= auctum misid.

Rhaphium nasutum (Fallén, 1823)

Rhaphium nigribarbatum (Becker, 1900)

Rhaphium patulum (Raddatz, 1873)

= antennatum misid.

Rhaphium penicillatum Loew, 1850

Rhaphium riparium (Meigen, 1824)

= praerosum Loew, 1850

Rhaphium rivale (Loew, 1869)

Rhaphium tridactylum (Frey, 1915)

Rhaphium umbripenne (Frey, 1915)

ACHALCINAE Grootaert & Meuffels, 1997

ACHALCUS Haliday, 1857

Achalcus cinereus (Haliday, 1851)

Achalcus flavicollis (Meigen, 1824)

Achalcus nigropunctatus Pollét & Brunhues, 1996

Achalcus vaillanti Brunhues, 1987

PELOROPEODINAE Robinson, 1970

CHRYSOTIMUS Loew, 1857

Chrysotimus molliculus (Fallén, 1823)

MICROMORPHUS Mik, 1878

Micromorphus claripennis (Strobl, 1899)

XANTHOCHLORINAE Aldrich, 1905

XANTHOCHLORUS Loew, 1857

Xanthochlorus ornatus (Haliday, 1832)

Xanthochlorus tenellus (Wiedemann, 1817)

Chelifera astigma Collin, 1927 misidentified

Chrysotus arcticus Frey, 1915 not found within present borders

Chrysotus longipalpus Aldrich, 1896 imported

= pallidipalpus van Duzee, 1933

Chrysotus ringdahli Parent, 1929 not found within present borders

Clinocera nigra Meigen, 1804 misidentified

Crossopalpus abditus Kovalev, 1972 not found within present borders

Dolichopus angustipennis Kertesz, 1901 not found within present borders

= adustus Frey, 1915

Dolichopus grandicornis Wahlberg, 1850 not found within present borders

Dolichopus mediicornis Verrall, 1875 not found within present borders

Dolichopus plumitarsis Fallén, 1823 misidentified

Dolichopus propinquus Zetterstedt, 1852 misidentified

Drapetis incompleta Collin, 1926 misidentified

Empis punctata Meigen, 1804 not found within present borders

Gymnopternus assimilis (Stæger, 1842) misidentified

Hilara flavipes Meigen, 1822 misidentified

Hilara maura (Fabricius, 1776) not found within present borders

Hilara scrobiculata Loew, 1873 misidentified

Hilara tetragramma Loew, 1873 misidentified

Hydrophorus balticus (Meigen, 1824) not found within present borders

Hydrophorus ponojensis Frey, 1915 not found within present borders

Iteaphila furcata (Zetterstedt, 1842) not found within present borders

Medetera annulitarsus von Roser, 1840 misidentified

Medetera glauca Loew, 1869 misidentified

Medetera feminina Negrobov, 1967 misidentified

Medetera truncorum Meigen, 1824 imported

Oedalea flavipes Zetterstedt, 1842 misidentified

Parathalassius krogerusi Frey, 1927 nomen nudum

Rhamphomyia tibialis Meigen, 1822 misidentified

Rhaphium antennatum (Carlier, 1835) misidentified

Rhaphium obscuripes Zetterstedt, 1849 misidentified

Rhaphium suave (Loew, 1859) not found within present borders

Tachytrechus insignis (Stannius, 1831) misidentified

Trichoclinocera lapponica (Ringdahl, 1933) not found within present borders

Dolichopus caligatus Wahlberg, 1850 was synonymized with D. flavipes Stannius, 1831 by Becker (1918) but Collin (1940) reinstated D. caligatus as a valid species. True D. flavipes (if it is a valid species) has not been found in Finland but some specimens have been recorded from Northwestern European Russia.

Dolichopus stenhammari Zetterstedt, 1843. Zetterstedt (1838) first described this species as Dolichopus annulipes. Later he proposed D. stenhammari Zett., 1843 as an (unnecessary) replacement name for the taxon: Porphyrops annulipes Meigen, 1824 was recognized in Dolichopus at the time, but only as a junior synonym of what is now Sympycnus pulicarius (Fallén, 1823) (Zetterstedt 1843). Nonetheless this act means, according to article 59.3 of The Code (ICZN 1999), that D. annulipes Zett., 1838 is permanently invalid.

Hilara pseudochorica Strobl, 1892. Erroneously deleted from the Finnish checklist by Kahanpää and Winqvist (2005).

Iteaphila furcata (Zetterstedt 1842). Sinclair and Shamshev (2012) list a specimen from Kuusamo with code label 1546. According to R. Frey’s notes this specimen was collected in 1917 from eastern Kuusamo, an area ceded to Russia in 1944.

Neurigona sp. A is probably N. uralensis Becker, 1918.

Platypalpus articulatus and P. maculimanus. According to Allen (1986) these two names are not synonymous and the species identified as P. articulatus Macquart by Chvála (1975, 1989) is in fact P. maculimanus (Zett.). Upon re-examination, all Finnish material previously identified as P. articulatus belonged to P. maculimanus and P. articulatoides Frey. It remains unclear whether the real P. articulatus occurs in Finland.

Rhamphomyia sp. nr. albipennis. An apparently undescribed species near R. albipennis (Fallén, 1816) (K. Winqvist & M. Barták, priv. comm.).

Sympycnus pulicarius (Fallén, 1823). Pollet (2004) (Fauna Europaea) treats S. annulipes (Meigen, 1824) and S. desoutteri Parent, 1925 as valid species and lists S. pulicarius (Fallén, 1823) as a synonym of the latter but notes that it might actually be a synonym of S. annulipes.

Thrypticus sp. A is probably T. incanus Negrobov, 1967 described from the Leningrad Oblast, Russia (Negrobov 1967).

Trichoclinocera lapponica (Ringdahl, 1933). The type locality of this species, ”Lappland nära Tjuonajaure”, (Ringdahl 1933) is in Sweden, not Finland as stated in Chvála and Wagner (1989).