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Checklist of the family Sciaridae (Diptera) of Finland
expand article infoPekka Vilkamaa
† University of Helsinki, Helsinki, Finland
Open Access

Abstract

A checklist of the family Sciaridae (Diptera) recorded from Finland is provided. The genus Sciarosoma Chandler with a disputed family placement is also included in the list.

Keywords

Checklist, Finland, Diptera, Sciaridae, Sciarosoma

Introduction

Sciaridae, the black fungus gnats, is one of the large families in Sciaroidea, little studied and notoriously difficult in its taxonomy. Up-to-date keys are not available for all European species, but various publications must be consulted for identification. Our knowledge of Finnish fauna stems from two classic publications: Frey (1948) and Tuomikoski (1960) but Menzel and Mohrig (2000) made fundamental nomenclatorial changes in their study of the type materials of all Palaearctic species.

As no records of Sciaridae in Finland were published between the publication of Tuomikoski’s (1960) monograph on the family and Hackman’s (1980) check-list of Finnish nematoceran Diptera, the species lists in both works are essentially unchanged, despite some obvious lapses in the latter. Our knowledge of Finnish fauna has since grown mainly in the taxonomic treatment of various genera and species groups (see Vilkamaa et al. 2013). All species recorded in Finland in publications thus far, besides the most recent ones, appear in H. Silfverberg’s compilations of changes to the Finnish insect fauna published every five years (for the latest one, see Silfverberg 2012).

With regard to nomenclature and generic classification, the present list follows the revisions of Menzel and Mohrig (2000) and Mohrig et al. (2013), although on the basis of phylogenetic analyses Baeosciara Tuomikoski, Dolichosciara Tuomikoski, Peyerimhoffia Kieffer, Prosciara Frey and Mouffetina Frey are regarded as genera (see Vilkamaa 2000, 2003, Vilkamaa and Hippa 2005, Shin et al. 2013). For names currently understood as junior synonyms, mostly only the names listed as valid in Tuomikoski (1960) are mentioned. For complete synonym lists, one can consult Menzel and Mohrig (2000) and Mohrig et al. (2013). Although some genera can be regarded as relatively well-known for the Finnish species they contain, for some others, such as the species-rich Bradysia Winnertz, many new species including undescribed ones may be present in the native fauna. Even so, together with Germany, Sweden and the United Kingdom, Finland can be considered one of the countries best known for its sciarid fauna.

Number of species:

World: 2600 species

Europe: 700 species

Finland: 338 species

Faunistic knowledge level in Finland: average

Checklist

Nematocera Dumeril, 1805

infraorder Bibionomorpha Hennig, 1954

superfamily Sciaroidea Billberg, 1820

SCIARIDAE Billberg, 1820

BAEOSCIARA Tuomikoski, 1960

Baeosciara discolor (Lengersdorf, 1928)

= pusillima (Frey, 1948)

Baeosciara sinuata (Menzel & Mohrig, 1997)

Baeosciara scotica (Edwards, 1925)

BRADYSIA Winnertz, 1867

Bradysia affinis (Zetterstedt, 1838)

= pratincola Tuomikoski, 1960

Bradysia albanensis (Lengersdorf, 1926)

Bradysia alpicola (Winnertz, 1867)

= mutabilis (Lengersdorf, 1926)

= morio auct. nec (Fabricius, 1794)

Bradysia angustata Tuomikoski, 1960

Bradysia angustostylata Menzel, 2005

Bradysia aprica (Winnertz, 1867)

Bradysia arcana Menzel & Mohrig, 1998

= fenestralis auct. nec (Zetterstedt, 1838)

Bradysia arcula Vilkamaa, Salmela & Hippa, 2007

Bradysia ascenda Rudzinski, 1994

Bradysia bicolor (Meigen, 1818)

Bradysia bispinifera Mohrig & Krivosheina, 1983

Bradysia brevispina Tuomikoski, 1960

Bradysia browni (Shaw, 1935)

= diversiabdominalis (Lengersdorf, 1941)

= laurencei Menzel & Mohrig, 2000

Bradysia cinerascens (Grzegorzek, 1884)

= lanicauda Tuomikoski, 1960

Bradysia confinis (Winnertz, 1867)

Bradysia excelsa Menzel & Mohrig, 1998

Bradysia flavipila Tuomikoski, 1960

Bradysia forcipulata (Lundbeck, 1898)

Bradysia forficulata (Bezzi, 1914)

= nocturna Tuomikoski, 1960

Bradysia fungicola (Winnertz, 1867)

Bradysia giraudii (Egger, 1862)

Bradysia globulifera (Lengersdorf, 1934)

Bradysia hilariformis Tuomikoski, 1960

Bradysia hilaris (Winnertz, 1867)

Bradysia holsatica Heller, 2004

Bradysia hortensis Heller, 2000

Bradysia impatiens (Johannsen, 1912)

= difformis (Frey, 1948)

= paupera Tuomikoski, 1960

Bradysia inusitata Tuomikoski, 1960

Bradysia iridipennis (Zetterstedt, 1838)

Bradysia lapponica (Lengersdorf, 1926)

Bradysia latiterga Tuomikoski, 1960

Bradysia leptoptera Tuomikoski, 1960

Bradysia lilienthalae Mohrig & Menzel, 1990

Bradysia lobulifera Frey, 1948

Bradysia longicauda Mohrig & Menzel, 1990

Bradysia longicubitalis (Lengersdorf, 1924)

= cinereovittata Frey, 1948

Bradysia minima Mohrig & Mamaev, 1989

Bradysia moesta Frey, 1948

= albosetosa Frey, 1948

Bradysia moestula Tuomikoski, 1960

Bradysia nervosa (Meigen, 1818)

Bradysia nitidicollis (Meigen, 1818)

= atroparva Frey, 1948

Bradysia normalis Frey, 1948

Bradysia ocellaris (Comstock, 1882)

Bradysia pallipes (Fabricius, 1787)

= brunnipes (Meigen, 1804)

= picipes (Zetterstedt, 1838)

Bradysia pauperata (Winnertz, 1867)

Bradysia peraffinis Tuomikoski, 1960

Bradysia pilistriata Frey, 1948

Bradysia placida (Winnertz, 1867)

= fimbricauda Tuomikoski, 1960

Bradysia polonica (Lengersdorf, 1929)

Bradysia praecox (Meigen, 1818)

Bradysia rectinervis Frey, 1948

Bradysia reflexa Tuomikoski, 1960

Bradysia regularis (Lengersdorf, 1934)

= subnervosa Frey, 1948

Bradysia rufescens (Zetterstedt, 1852)

Bradysia scabricornis Tuomikoski, 1960

= subscabricornis Mohrig & Menzel, 1990

Bradysia siberica Komarova, 2001

Bradysia spinostyla Mohrig & Menzel, 1990

Bradysia strigata (Staeger, 1840)

Bradysia subalpina Frey, 1948

Bradysia subamoena Mohrig & Krivosheina, 1989

Bradysia submoesta Mohrig & Krivosheina, 1989

Bradysia tilicola (Loew, 1850)

= amoena (Winnertz, 1867)

= cellarum Frey, 1948

Bradysia trivittata (Staeger, 1840)

Bradysia vagans (Winnertz, 1868)

= callicera Frey, 1948

Bradysia vernalis (Zetterstedt, 1851)

Bradysia zonata Rudzinski, 1993

BRADYSIOPSIS Tuomikoski, 1960

Bradysiopsis vittata (Meigen, 1830)

= leucotricha (Tuomikoski, 1960)

Bradysiopsis vittigera (Zetterstedt, 1851)

CAMPTOCHAETA Hippa & Vilkamaa, 1994

Camptochaeta austriaca Heller, 2012

Camptochaeta bournei (Shaw, 1941)

Camptochaeta camptochaeta (Tuomikoski, 1960)

Camptochaeta consimilis (Holmgren, 1869)

Camptochaeta delicata (Lengersdorf, 1935)

Camptochaeta duplicata Hippa & Vilkamaa, 1994

Camptochaeta fallax Hippa & Vilkamaa, 1994

Camptochaeta hirtula (Lengersdorf, 1934)

= fulvicollis (Tuomikoski, 1960)

Camptochaeta propria Hippa & Vilkamaa, 1994

Camptochaeta scanica Hippa & Vilkamaa, 1994

Camptochaeta sicilicula Hippa & Vilkamaa, 1994

Camptochaeta simulator Hippa & Vilkamaa, 1994

Camptochaeta stammeri (Lengersdorf, 1940)

Camptochaeta tenuipalpalis (Mohrig & Antonova, 1978)

Camptochaeta uniformis (Mohrig & Menzel, 1990)

Camptochaeta vivax (Frey, 1948)

Camptochaeta xystica Hippa & Vilkamaa, 1994

CHAETOSCIARA Frey, 1942

Chaetosciara estlandica (Lengersdorf, 1929)

CLAUSTROPYGA Hippa, Vilkamaa & Mohrig, 2003

Claustropyga acanthostyla (Tuomikoski, 1960)

Claustropyga brevichaeta (Mohrig & Antonova, 1978)

Claustropyga clausa (Tuomikoski, 1960)

Claustropyga corticis (Mohrig & Antonova, 1978)

Claustropyga ctenophora Hippa, Vilkamaa & Mohrig, 2003

Claustropyga heteroclausa (Rudzinski, 1991)

Claustropyga refrigerata (Lengersdorf, 1930)

Claustropyga subcorticis (Mohrig & Krivosheina, 1985)

CORYNOPTERA Winnertz, 1867

Corynoptera barbata Tuomikoski, 1960

Corynoptera bicuspidata (Lengersdorf, 1926)

= gymnops Tuomikoski, 1960

Corynoptera bipartita Mohrig & Krivosheina, 1985

Corynoptera blanda (Winnertz, 1867)

Corynoptera boletiphaga (Lengersdorf, 1940)

= geogenia Tuomikoski, 1960

Corynoptera breviformis Mohrig & Krivosheina, 1983

Corynoptera cracentis Vilkamaa, Hippa & Heller, 2013

Corynoptera cuniculata (Lengersdorf, 1942)

= caldariorum Tuomikoski, 1960

Corynoptera defecta (Frey, 1948)

Corynoptera dentata (Bukowski & Lengersdorf, 1936)

Corynoptera deserta Heller & Menzel, 2006

= minutula (Bukowski & Lengersdorf, 1936)

Corynoptera dubitata Tuomikoski, 1960

Corynoptera fera Mohrig & Heller, 1992

Corynoptera flavicauda (Zetterstedt, 1855)

Corynoptera forcipata (Winnertz, 1867)

Corynoptera furcifera Mohrig & Mamaev, 1987

Corynoptera globiformis (Frey, 1945)

Corynoptera hypopygialis (Lengersdorf, 1926)

= piniphila (Lengersdorf, 1940)

Corynoptera inari Vilkamaa, Hippa & Heller, 2013

Corynoptera inexspectata Tuomikoski, 1960

Corynoptera irmgardis (Lengersdorf, 1930)

Corynoptera levis Tuomikoski, 1960

Corynoptera luteofusca (Bukowski & Lengersdorf, 1936)

Corynoptera marinae Mohrig & Krivosheina, 1986

Corynoptera melanochaeta Mohrig & Menzel, 1992

Corynoptera membranigera (Kieffer, 1903)

= trispina Tuomikoski, 1960

Corynoptera montana (Winnertz, 1869)

Corynoptera ninae Hippa, Vilkamaa & Heller, 2010

Corynoptera obscuripila Tuomikoski, 1960

Corynoptera parvula (Winnertz, 1867)

= uncinata (Hippa & Vilkamaa, 1994)

Corynoptera parvulaformis Mohrig, 1985

Corynoptera penna (Pettey, 1918)

= alneti Hippa, Vilkamaa & Heller, 2010

Corynoptera perochaeta (Mohrig & Menzel, 1990)

Corynoptera plusiochaeta Hippa, Vilkamaa & Heller, 2010

Corynoptera polana Rudzinski, 2009

Corynoptera postforcipata Rudzinski, 1993

Corynoptera postglobiformis Mohrig, 1993

Corynoptera praeforcipata Mohrig & Mamaev, 1987

Corynoptera quantula (Hippa & Vilkamaa, 1994)

Corynoptera saccata Tuomikoski, 1960

Corynoptera saetistyla Mohrig & Krivosheina, 1985

Corynoptera salmelai Vilkamaa, Hippa & Heller, 2013

Corynoptera sphenoptera Tuomikoski, 1960

Corynoptera spiciforceps Vilkamaa, Hippa & Heller, 2013

Corynoptera spinifera Tuomikoski, 1960

Corynoptera subblanda Tuomikoski, 1960

Corynoptera subdentata Mohrig, 1985

Corynoptera subparvula Tuomikoski, 1960

Corynoptera subsedula Mohrig & Mamaev, 1987

Corynoptera subtetrachaeta Komarova, 1995

Corynoptera subtilis (Lengersdorf, 1929)

= longicornis (Bukowski & Lengersdorf, 1936)

Corynoptera subvariegata Rudzinski, 1992

Corynoptera tetrachaeta Tuomikoski, 1960

Corynoptera trepida (Winnertz, 1867)

= clinochaeta Tuomikoski, 1960

Corynoptera triacantha Tuomikoski, 1960

Corynoptera tumidula Hippa, Vilkamaa & Heller, 2010

Corynoptera tuomikoskii Hippa, Vilkamaa & Heller, 2013

Corynoptera unidentata (Hippa & Vilkamaa, 1994)

Corynoptera vagula Tuomikoski, 1960

Corynoptera verrucifera (Lengersdorf, 1952)

Corynoptera voluptuosa Mohrig & Mamaev, 1987

Corynoptera waltraudis Mohrig & Mamaev, 1987

Corynoptera winnertzi Mohrig, 1993

COSMOSCIARA Frey, 1942

Cosmosciara perniciosa (Edwards, 1922)

CRATYNA Winnertz, 1867

= Plastosciara Berg, 1899

= Decembrina Frey, 1948

= Dendrosciara Frey, 1948

sg. Cratyna Winnertz, 1867

Cratyna ambigua (Lengersdorf, 1934)

= latiforceps (Bukowski & Lengersdorf, 1936)

Cratyna atra Winnertz, 1867

= pictiventris (Kieffer, 1898)

Cratyna betulae (Mohrig & Mamaev, 1992)

Cratyna breviflagellata (Mohrig & Mamaev, 1985)

Cratyna monumenta Rudzinski, 2009

Cratyna pernitida (Edwards, 1915)

Cratyna schineri (Winnertz, 1867)

Cratyna sicata Vilkamaa, Hippa & Heller, 2013

Cratyna symplecta (Rudzinski, 1991)

Cratyna uliginosa (Lengersdorf, 1929)

Cratyna vaporariorum (Frey, 1948)

sg. Diversicratyna Menzel & Mohrig, 1998

Cratyna spiculosa (Rudzinski, 1993)

sg. Spathobdella Frey, 1948

Cratyna colei (Freeman, 1990)

= brachialis auct. nec (Winnertz, 1871)

Cratyna falcata (Tuomikoski, 1960)

Cratyna falcifera (Lengersdorf, 1933)

Cratyna longispina (Pettey, 1918)

= tuberculata (Tuomikoski, 1960)

Cratyna nobilis (Winnertz, 1867)

= brachialis (Winnertz, 1867)

Cratyna perplexa (Winnertz, 1867)

= socialis (Winnertz, 1871)

= brevicornis (Tuomikoski, 1957)

CTENOSCIARA Tuomikoski, 1960

Ctenosciara exigua Salmela & Vilkamaa, 2005

Ctenosciara hyalipennis (Meigen, 1804)

DICHOPYGINA Vilkamaa, Hippa & Komarova, 2004

Dichopygina aculeata Vilkamaa, Hippa & Komarova, 2004

Dichopygina intermedia (Mohrig & Krivosheina, 1982)

Dichopygina nigrohalteralis (Frey, 1948)

Dichopygina ramosa Vilkamaa, Hippa & Komarova, 2004

DOLICHOSCIARA Tuomikoski, 1960

Dolichosciara flavipes (Meigen, 1804) 185

Dolichosciara hippai Komarova & Vilkamaa, 2006

Dolichosciara nigrovittata (Strobl, 1910)

Dolichosciara orcina (Tuomikoski, 1960)

Dolichosciara ornata (Winnertz, 1867)

Dolichosciara saetosa (Lengersdorf, 1929)

Dolichosciara spissispina Vilkamaa, Hippa & Heller, 2013

EPIDAPUS Haliday, 1851

sg. Epidapus Haliday, 1851

Epidapus alnicola (Tuomikoski, 1957)

Epidapus atomarius (De Geer, 1778)

Epidapus gracilis (Walker, 1848)

Epidapus ignotus (Lengersdorf, 1942)

= gracilior (Tuomikoski, 1960)

Epidapus microthorax (Börner, 1903)

= gracilicornis (Lengersdorf, 1926)

Epidapus schillei (Börner, 1903)

= titan Frey, 1948

= intermittens Tuomikoski, 1959

sg. Pseudoaptanogyna Vimmer, 1926

Epidapus abieticola Frey, 1948

Epidapus bispinulosus Mohrig & Kauschke, 1994

Epidapus echinatum Mohrig & Kozánek, 1992

Epidapus ignavus (Lengersdorf, 1941)

KEILBACHIA Mohrig, 1987

Keilbachia ferrata (Hippa & Vilkamaa, 1994)

LEPTOSCIARELLA Tuomikoski, 1960

sg. Hirtipennia Mohrig & Menzel, 1997

Leptosciarella hirtipennis (Zetterstedt, 1838)

Leptosciarella holotricha Mohrig & Menzel, 1997

sg. Leptosciarella Tuomikoski, 1960

Leptosciarella brevior (Tuomikoski, 1960)

Leptosciarella brevipalpa (Mohrig & Menzel, 1992)

Leptosciarella claviforceps (Tuomikoski, 1960)

Leptosciarella dimera (Tuomikoski, 1960)

Leptosciarella fuscipalpa (Mohrig & Mamaev, 1979)

Leptosciarella helvetica (Rudzinski, 1992)

Leptosciarella hispida (Winnertz, 1871)

Leptosciarella ignis Heller, 2012

Leptosciarella krille Heller, 2012

Leptosciarella melanoma (Mohrig & Menzel, 1990)

Leptosciarella nudinervis (Tuomikoski, 1960)

Leptosciarella pilosa (Staeger, 1840)

Leptosciarella reducta Heller & Menzel, 2013

Leptosciarella rejecta (Winnertz, 1867)

Leptosciarella scutellata (Staeger, 1840)

= elegans (Winnertz, 1867)

Leptosciarella subcoarctata Mohrig & Menzel, 1997

Leptosciarella subpilosa (Edwards, 1925)

Leptosciarella subspinulosa (Edwards, 1925)

Leptosciarella subviatica Mohrig & Menzel, 1997

Leptosciarella trochanterata (Zetterstedt, 1851)

= coarctata (Winnertz, 1867)

Leptosciarella truncata (Tuomikoski, 1960)

Leptosciarella viatica (Winnertz, 1867)

Leptosciarella viaticella Mohrig & Krivosheina, 1979

Leptosciarella yerburyi (Freeman, 1983)

sg. Leptospina Mohrig & Menzel, 1997

Leptosciarella atricha (Tuomikoski, 1960)

sg. Trichosiopsis Tuomikoski, 1960

Leptosciarella tuberculigera (Tuomikoski, 1960)

LYCORIELLA Frey, 1942

sg. Coelostylina Tuomikoski, 1960

Lycoriella eflagellata Tuomikoski, 1960

Lycoriella freyi Tuomikoski, 1960

sg. Hemineurina Tuomikoski, 1960

Lycoriella algida (Frey, 1948)

Lycoriella cochleata (Rübsaamen, 1898)

Lycoriella conspicua (Winnertz, 1867)

Lycoriella inflata (Winnertz, 1867)

= venosa auct. nec (Staeger, 1840)

Lycoriella modesta (Staeger, 1840)

Lycoriella piristylata Vilkamaa, Hippa & Heller, 2013

Lycoriella thuringiensis Menzel & Mohrig, 1991

Lycoriella vitticollis (Holmgren, 1883)

= permutata (Lundbeck, 1900)

sg. Lycoriella Frey, 1942

Lycoriella aberrans Tuomikoski, 1960

Lycoriella acutostylia Mohrig & Menzel, 1990

Lycoriella agraria (Felt, 1897)

= cellaris (Lengersdorf, 1934)

Lycoriella approximatonervis (Frey, 1948)

Lycoriella brevipila Tuomikoski, 1960

Lycoriella inconspicua Tuomikoski, 1960

Lycoriella ingenua (Dufour, 1839)

= solani (Winnertz, 1871)

Lycoriella latilobata Menzel & Mohrig, 2000

Lycoriella lundstromi (Frey, 1948)

Lycoriella micria Mohrig & Menzel, 1990

Lycoriella minutula Mohrig & Krivosheina, 1987

Lycoriella pallidior Tuomikoski, 1960

Lycoriella parva (Holmgren, 1869)

= obscuratipes (Frey, 1948)

= curvispina Tuomikoski, 1960

Lycoriella sativae (Johannsen, 1912)

= castanescens (Lengersdorf, 1940)

= fucorum (Frey, 1948)

Lycoriella subterranea (Märkel, 1844)

= vanderwieli (Schmitz, 1920)

Lycoriella tenera Vilkamaa, Hippa & Heller, 2013

Lycoriella weberi Menzel & Heller, 2013

MOUFFETINA Frey, 1942

Mouffetina expolita (Coquillett, 1900)

Mouffetina pulchricornis (Edwards, 1925)

Mouffetina silvestris (Mohrig & Antonova, 1978)

PEYERIMHOFFIA Kieffer, 1903

Peyerimhoffia crassistylata (Frey, 1948)

Peyerimhoffia infera Vilkamaa & Hippa, 2005

Peyerimhoffia menzeli Vilkamaa & Hippa, 2005

Peyerimhoffia quadrifera Vilkamaa, Hippa & Heller, 2013

Peyerimhoffia sepei Hippa & Vilkamaa, 2005

Peyerimhoffia thula Vilkamaa & Hippa, 2005

Peyerimhoffia vagabunda (Winnertz, 1867)

= brachyptera Kieffer, 1903

PHYTOSCIARA Frey, 1942

Phytosciara halterata (Lengersdorf, 1926)

Phytosciara macrotricha (Lengersdorf, 1926)

PNYXIA Johannsen, 1912

Pnyxia scabiei (Hopkins, 1895)

PNYXIOPSIS Tuomikoski, 1960

Pnyxiopsis aliger Tuomikoski, 1960

Pnyxiopsis degener (Tuomikoski, 1957)

PROSCIARA Frey, 1942

Prosciara furtiva Vilkamaa & Hippa, 1996

Prosciara plusiochaeta Hippa & Vilkamaa, 1991

Prosciara porrecta (Lengersdorf, 1929)

Prosciara producta (Tuomikoski, 1960)

Prosciara prosciaroides (Tuomikoski, 1960)

Prosciara ungulata (Winnertz, 1867)

PSEUDOLYCORIELLA Menzel & Mohrig, 1998

Pseudolycoriella brunnea (Bukowski & Lengersdorf, 1936)

Pseudolycoriella japonensis (Mohrig & Menzel, 1992)

Pseudolycoriella koreensis (Mohrig & Menzel, 1992)

Pseudolycoriella monticula (Mohrig & Menzel, 1992)

Pseudolycoriella nodulosa (Mohrig & Krivosheina, 1985)

Pseudolycoriella paludum (Frey, 1948)

Pseudolycoriella subbruckii (Mohrig & Hövemeyer, 1992)

SCATOPSCIARA Edwards, 1927

sg. Scatopsciara Edwards, 1927

Scatopsciara atomaria (Zetterstedt, 1851)

= vivida (Winnertz, 1867)

Scatopsciara bucera Rudzinski, 1994

Scatopsciara calamophila Frey, 1948

Scatopsciara curviforceps (Bukowski & Lengersdorf, 1936)

= myrmecophila Frey, 1948

Scatopsciara edwardsi Freeman, 1983

Scatopsciara fluviatilis (Lengersdorf, 1940)

Scatopsciara geophila (Tuomikoski, 1960)

Scatopsciara multispina (Bukowski & Lengersdorf, 1936)

Scatopsciara neglecta Menzel & Mohrig, 1998

Scatopsciara pusilla (Meigen, 1818)

Scatopsciara subcalamophila Menzel & Mohrig, 1991

Scatopsciara subciliata Tuomikoski, 1960

Scatopsciara tricuspidata (Winnertz, 1867)

= degenerans (Frey, 1948)

Scatopsciara vitripennis (Meigen, 1818)

sg. Xenopygina Frey, 1948

Scatopsciara gabyae (Heller, 1998)

Scatopsciara obliqua Vilkamaa, Hippa & Heller, 2013

Scatopsciara paradoxa (Frey, 1948)

Scatopsciara simillima (Tuomikoski, 1960)

SCHWENCKFELDINA Frey, 1942

Schwenckfeldina carbonaria (Meigen, 1830)

Schwenckfeldina pectinea Menzel & Mohrig, 1991

SCIARA Meigen, 1803

Sciara flavimana Zetterstedt, 1851

Sciara hebes (Loew, 1869)

= mendax Tuomikoski., 1960

= nursei Freeman, 1983

= ulrichi Menzel & Mohrig, 1998

Sciara helvola Winnertz, 1867

Sciara hemerobioides (Scopoli, 1763)

= thomae (Linnaeus, 1767)

Sciara humeralis Zetterstedt, 1851

Sciara lackschewitzi (Lengersdorf, 1934)

SCYTHROPOCHROA Enderlein, 1911

Scythropochroa quercicola (Winnertz, 1869)

Scythropochroa radialis Lengersdorf, 1926

TRICHOSIA Winnertz, 1867

= Leptosciara Frey, 1942

= Lestremioides Frey, 1942

Trichosia Winnertz, 1867

Trichosia acrotricha Tuomikoski, 1960

Trichosia borealis (Frey, 1942)

Trichosia confusa Menzel & Mohrig, 1997

Trichosia flavicoxa Tuomikoski, 1960

Trichosia diota (Garrett, 1925)

Trichosia glabra (Meigen, 1830)

Trichosia habilis (Johannsen, 1912)

= edwardsi (Lengersdorf, 1930)

Trichosia morio (Fabricius, 1794)

= caudata (Walker, 1848)

Trichosia splendens Winnertz, 1867

Trichosia ussurica Mohrig & Antonova, 1978

XYLOSCIARA Tuomikoski, 1957

sg. Protoxylosciara Tuomikoski, 1960

Xylosciara longiforceps (Bukowski & Lengersdorf, 1936)

sg. Xylosciara Tuomikoski, 1957

Xylosciara heptacantha Tuomikoski, 1957

Xylosciara lignicola (Winnertz, 1867)

Xylosciara microdon (Frey, 1948)

Xylosciara misella (Frey, 1948)

Xylosciara phryganophila (Frey, 1948)

Xylosciara senta Vilkamaa, Hippa & Heller, 2013

Xylosciara spinata (Pettey, 1918)

= betulae Tuomikoski, 1960

Xylosciara steleocera Tuomikoski, 1960

Xylosciara trimera Tuomikoski, 1960

Xylosciara validinervis Tuomikoski, 1960

ZYGONEURA Meigen, 1830

sg. Allozygoneura Menzel & Mohrig, 1998

Zygoneura calthae Tuomikoski, 1960

sg. Zygoneura Meigen, 1830

Zygoneura sciarina Meigen, 1830

UNPLACED IN SCIAROIDEA

SCIAROSOMA Chandler, 2002

Sciarosoma nigriclava (Strobl, 1898)

= borealis Chandler, 2002

Excluded species

Lycoriella auripila (Winnertz, 1867), misidentified in Tuomikoski (1960).

Bradysia betuleti (Lengersdorf, 1940), misidentified in Tuomikoski (1960).

Bradysia subbetuleti Mohrig & Krivosheina, 1989, misidentified in Salmela and Vilkamaa (2005).

Corynoptera concinna (Winnertz, 1867), misidentified in Tuomikoski (1960).

Corynoptera bistrispina (Bukowski & Lengersdorf, 1936), misidentified in Tuomikoski (1960).

Trichosia trochanterata (Zetterstedt, 1851), misidentified in Tuomikoski (1960).

Acknowledgements

I thank my colleagues Kai Heller, Heikki Hippa, Frank Menzel, Werner Mohrig and Jukka Salmela for their long-term help in identifying Sciaridae.

References

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