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Checklist of the fungus gnats of Finland: BolitophilidaeDiadocidiidae, Ditomyiidae, Keroplatidae and Mycetophilidae (Diptera)
expand article infoJevgeni Jakovlev
† Finnish Environment Institute, Helsinki, Finland
Open Access

Abstract

A checklist of the ’fungus gnat’ group of families in Sciaroidea: Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae and Mycetophilidae (Diptera) recorded from Finland. At present, the known Finnish fauna comprises 767 recognized species in 78 genera; 21 species of the family Bolitophilidae, 5 of the family Diadocidiidae, 2 of the family Ditomyiidae, 47 of the family Keroplatidae and 692 of the family Mycetophilidae. The unassigned genus Sciarosoma Chandler, 2002 is not included in this paper because it is listed with Sciaridae.

The most relevant synonyms, on which species were ever listed from Finland in Hackman (1980) and subsequent authors are indicated in the checklist. Eighteen species names of wrong records were removed.

Keywords

Species list, Finland, Diptera, biodiversity, faunistics

Introduction

Fungus gnats are a rich assemblage of nematocerous flies (Diptera, Nematocera) placed in the infraorder Bibionomorpha and superfamily Sciaroidea, currently divided into eight families (Blagoderov and Grimaldi 2004). Five of these families (Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae and Mycetophilidae) are called informally ‘fungus gnats’ by most European authors. Species in these families are morphologically closely related and the group is also ecologically uniform.

Adults of fungus gnats are generally quite easily recognized by their habitus. They are slender to moderately robust flies between 2 and 10 mm (a few up to 15 mm) in body length, with long antenna, a high and often strongly arched thorax made distinctly higher by the long, stout and tight-sitting coxae of their legs. Their wings have reduced and variable venation that is quite distinctive for the different families and much used also to classify genera. Their body and legs are often covered with strong bristles. As larvae the majority of fungus gnat species are mycetophagous, associated with fungi, either fruiting bodies or mycelia in dead wood and soil litter.

Here I cover all the Finnish fungus gnat fauna of the families Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae, and Mycetophilidae. In Europe, the superfamily Sciaroidea further includes two large families, the Sciaridae – black-winged fungus gnats and the Cecidomyiidae – gall midges (Søli et al. 2000) although some authors, e.g. Hippa and Vilkamaa (2006) consider the Cecidomyiidae as the sister group of all other Sciaroidea. They differ markedly in habitus and larval biology and are presented elsewhere. Finally, the unassigned genus Sciarosoma Chandler, 2002 is not included in this checklist since it is listed with the family Sciaridae according Hippa and Vilkamaa (2006), but see also opinion by Jaschhof et al. (2006).

Systematics and nomenclature of the enumeration follow the online checklist of Nordic fungus gnats (Kjærandsen 2012). Only the most relevant synonyms, on which species were ever listed from Finland, are included in the checklist.

Adults of the North European species can largely be identified using Hutson et al. (1980) and Zaitzev (1994)Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae, and Mycetophilidae in part (Mycomyiinae, Sciophilinae, Gnoristinae and Leiinae), and Zaitzev (2003)Mycetophilidae in part (Mycetophilinae and Manotinae). However, since many new taxa are added, and corrections, synonyms and revisions continually published, one needs a complete set of the fragmented taxonomic literature to cover all the species. A good source for obtaining most important literature on the Sciaroidea is the ViBrant/Scratchpad module Fungus Gnats Online (2014). Species identification of fungus gnats based on examination of larvae is currently impossible because the larvae seemingly lack unique morphological characteristics and search for species-specific molecular markers are yet to be implemented.

Fungus gnats are distributed all over the world with about 5,300 known species (Table 1), more than 1,450 of them are known from the Palaearctic region (Søli et al. 2000). According to the current version of the Fauna Europaea Web-database (Chandler 2004), 1,098 species of fungus gnats had then been recorded from Europe. Taking into account literature published subsequently these figures could be increased in ca 100 additional species, i.e. some 1,200 species are known in Europe at present. As opposed to many other insect groups, fungus gnats, at least Mycetophilidae, seem to display an increasing diversity towards the north in Europe, and in the European perspective, they are especially diverse in the boreal zone. The current version of the checklist of North European (Sweden, Norway, Finland, Russian Karelia, Russian Lapland, Denmark, The Faroes, and Iceland) fungus gnats (Kjærandsen 2012) includes 896 species.

In Finland, 269 species of the fungus gnats (of which ca. 40 currently recognized species were described by Carl Lundström) were first listed in the Enumeration Insectorum Fenniae (Frey et al. 1941). Hackman (1980) presented the second checklist of Finnish Diptera that incorporates 486 species of fungus gnats including five with a question mark. This list of species, with minor additions and corrections concerning Mycetophilidae after systematical revisions of the genera Trichonta (Gagné 1981) and Mycomya (Väisänen 1984) in the Holarctic region was included into Catalogue of the Palaearctic Diptera (Hackman et al. 1988). During the recent two decades, large-scale inventories (e.g. Polevoi and Jakovlev 2004, Polevoi et al. 2006, Jakovlev et al. 2006, 2014, Jakovlev and Penttinen 2007) have increased the total number of Finnish fungus gnats with ca. 300 species. The 2010 Finnish Red List assessment of fungus gnats was based on a regional species pool consisting on 734 species (Penttinen et al. 2010). At present, the Finnish fungus gnat fauna incorporates 767 recognized species (except Sciarosoma) and ca. 20 species awaiting descriptions. Altogether, 18 species names of wrong records from Finland were removed from the checklist. These species and 3 doubtful species names are listed in the end.

Number of species by family.

Family Number of species (except Sciarosoma) in Level of knowledge
World
(Pape et al. 2011)
Europe
(Chandler 2004)
Nordic Region (Kjærandsen 2012) Finland
Bolitophilidae 59 36 31 21 average
Diadocidiidae 34 5 5 5 average
Ditomyiidae 94 4 3 2 average
Keroplatidae 953 110 62 47 average
Mycetophilidae 4150 942 794 692 average

Checklist

Nematocera Dumeril, 1805

infraorder Bibionomorpha Hennig, 1954

superfamily Sciaroidea Schiner, 1864

BOLITOPHILIDAE Winnertz, 1864

BOLITOPHILA Meigen, 1818

sg. Bolitophila Meigen, 1818

Bolitophila austriaca (Mayer, 1950)

Bolitophila basicornis (Mayer, 1951)

Bolitophila caspersi Plassmann, 1956

Bolitophila cinerea Meigen, 1818

Bolitophila saundersii (Curtis, 1836)

Bolitophila tenella Winnertz, 1864

sg. Cliopisa Enderlein, 1936

Bolitophila aperta Lundström, 1915

Bolitophila bimaculata Zetterstedt, 1838

Bolitophila dubia Siebke, 1863

= disjuncta Loew, 1869

Bolitophila fumida Edwards, 1941

Bolitophila glabrata Loew, 1869

Bolitophila hybrida (Meigen, 1804)

Bolitophila ingrica Stackelberg, 1969

Bolitophila limitis Polevoi, 1996

Bolitophila maculipennis Walker, 1836

= coronata Mayer, 1951

Bolitophila modesta Lackschewitz, 1937

= tarsata Mayer, 1951

Bolitophila nigrolineata Landrock, 1912

Bolitophila obscurior Stackelberg, 1969

Bolitophila occlusa Edwards, 1913

Bolitophila pseudohybrida Landrock, 1912

Bolitophila rossica Landrock, 1912

DIADOCIDIIDAE Winnertz, 1864

DIADOCIDIA Ruthe, 1831

sg. Adidocidia Laštovka & Matile 1972

Diadocidia trispinosa Polevoi, 1996

Diadocidia valida Mik, 1874

sg. Diadocidia Ruthe, 1831

Diadocidia ferruginosa (Meigen, 1830)

Diadocidia fissa Zaitzev, 1994

Diadocidia spinosula Tollet, 1948

DITOMYIIDAE Keilin, 1919

SYMMERUS Walker, 1848

Symmerus annulatus (Meigen, 1830)

Symmerus nobilis Lackschewitz, 1937

KEROPLATIDAE Rondani, 1856

KEROPLATINAE Rondani, 1856

tribe Keroplatini Rondani, 1856

CEROTELION Rondani, 1856

Cerotelion striatum (Gmelin, 1790)

= lineatum (Fabricius, 1775) preocc.

KEROPLATUS Bosc, 1792

Keroplatus testaceus Dalman, 1818

Keroplatus tipuloides Bosc, 1792

= sesioides Wahlberg, 1839

* Keroplatus tuvensis Zaitzev, 1991 (see Notes)

ROCETELION Matile, 1988

Rocetelion humerale (Zetterstedt, 1850)

tribe Orfeliini Matile, 1990

ISONEUROMYIA Brunetti, 1912

Isoneuromyia semirufa (Meigen, 1818)

MACRORRHYNCHA Winnertz, 1846

Macrorrhyncha flava Winnertz, 1846

Macrorrhyncha rostrata (Zetterstedt, 1851)

MONOCENTROTA Edwards, 1925

Monocentrota lundstroemi Edwards, 1925

NEOPLATYURA Malloch, 1928

Neoplatyura flava (Macquart, 1826)

Neoplatyura modesta (Winnertz, 1864)

ORFELIA Costa, 1857

Orfelia discoloria (Meigen, 1818)

Orfelia falcata Zaitzev, 1994

Orfelia fasciata (Meigen, 1804)

Orfelia krivosheinae Zaitzev, 1994

Orfelia lugubris (Zetterstedt, 1851)

= tristis (Lundström, 1911)

Orfelia nemoralis (Meigen, 1818)

Orfelia nigricornis (Fabricius, 1805)

Orfelia ochracea (Meigen, 1818)

= unicolor (Staeger, 1840)

Orfelia pallida (Staeger, 1840)

PYRATULA Edwards, 1929

Pyratula perpusilla (Edwards, 1913)

Pyratula subcanariae Chandler, 2001

Pyratula zonata (Zetterstedt, 1855)

URYTALPA Edwards, 1929

Urytalpa atriceps (Edwards, 1913)

Urytalpa dorsalis (Staeger, 1840)

= ochracea (Meigen, 1818)

Urytalpa galdes Hedmark & Kjærandsen, 2009

Urytalpa macrocera (Edwards, 1913)

Urytalpa trivittata (Lundström, 1914)

MACROCERINAE Rondani, 1856

tribe Macrocerini Rondani, 1856

MACROCERA Meigen, 1803

Macrocera angulata Meigen, 1818

Macrocera centralis Meigen, 1818

Macrocera crassicornis Winnertz, 1864

Macrocera estonica Landrock, 1924

Macrocera fasciata Meigen, 1804

Macrocera fascipennis Staeger, 1840

Macrocera grandis Lundström, 1912

= magna Landrock, 1917

Macrocera lutea Meigen, 1804

Macrocera maculata Meigen, 1818

Macrocera nigricoxa Winnertz, 1864

Macrocera parva Lundström, 1914

Macrocera phalerata Meigen, 1818

Macrocera pilosa Landrock, 1917

Macrocera pumilio Loew, 1869

Macrocera pusilla Meigen, 1830

Macrocera stigma Curtis, 1837

Macrocera stigmoides Edwards, 1925

Macrocera vittata Meigen, 1830

Macrocera zetterstedti Lundström, 1914

= nana Zetterstedt, 1860

MYCETOPHILIDAE Newman, 1834

MYCOMYINAE Edwards, 1925

MYCOMYA Rondani, 1856

sg. Calomycomya Väisänen, 1984

Mycomya avala Väisänen, 1984

Mycomya pulchella (Dziedzicki, 1885)

sg. Coheromyia Väisänen, 1984

Mycomya branderi Väisänen, 1984

= tantilla auct. nec (Loew, 1869) misid.

sg. Cymomya Väisänen, 1984

Mycomya circumdata (Staeger, 1840)

sg. Lycomya Väisänen, 1984

Mycomya pectinifera Edwards, 1924

sg. Mycomya Rondani, 1856

Mycomya annulata (Meigen, 1818)

= incisurata (Zetterstedt, 1838)

Mycomya bialorussica Landrock, 1925

Mycomya bicolor (Dziedzicki, 1885)

Mycomya bisulca Lackschewitz, 1937

Mycomya britteni Kidd, 1955

Mycomya brunnea (Dziedzicki, 1885)

Mycomya cinerascens (Macquart, 1826)

= hyalinata (Meigen, 1830)

Mycomya collini Edwards, 1941

Mycomya danielae Matile, 1972

Mycomya denmax Väisänen, 1979

Mycomya digitifera Edwards, 1925

Mycomya disa Väisänen, 1984

Mycomya dziedzickii Väisänen, 1984

= fulva (Dziedzicki, 1885) preocc.

Mycomya egregia (Dziedzicki, 1885)

Mycomya festivalis Väisänen, 1984

Mycomya flavicollis (Zetterstedt, 1852)

Mycomya forestaria Plassmann, 1978

Mycomya fuscata (Winnertz, 1864)

Mycomya griseovittata (Zetterstedt, 1852)

= fasciata (Zetterstedt, 1838) preocc.

= clavigera (Lundström, 1912)

Mycomya hackmani Väisänen, 1984

Mycomya heydeni Plassmann, 1970

Mycomya hians (Lundström, 1912)

Mycomya hiisi Väisänen, 1979

Mycomya humida Garrett, 1924

= curvata Fisher, 1937

Mycomya indistincta Polevoi, 1995

Mycomya insignis (Winnertz, 1864)

= wrzesniowskii (Dziedzicki, 1885)

Mycomya islandica Väisänen, 1984

Mycomya karelica Väisänen, 1979

Mycomya kaurii Väisänen, 1979

Mycomya kuusamoensis Väisänen, 1979

Mycomya lambi Edwards, 1941

Mycomya leporina Väisänen, 1984

Mycomya levis (Dziedzicki, 1885)

Mycomya livida (Dziedzicki, 1885)

Mycomya maculata (Meigen, 1804)

Mycomya marginata (Meigen, 1818)

= limbata (Winnertz, 1864)

Mycomya mituda Väisänen, 1980

Mycomya neohyalinata Väisänen, 1984

Mycomya nigricornis (Zetterstedt, 1852)

= melanoceras Edwards, 1924

Mycomya nitida (Zetterstedt, 1852)

= exigua (Winnertz, 1864)

Mycomya norna Väisänen, 1984

Mycomya occultans (Winnertz, 1864)

Mycomya ornata (Meigen, 1818)

Mycomya parva (Dziedzicki, 1885)

Mycomya prominens (Lundström, 1913)

Mycomya pseudoapicalis Landrock, 1925

Mycomya pseudocurvata Väisänen, 1979

Mycomya punctata (Meigen, 1804)

Mycomya ruficollis (Zetterstedt, 1852)

Mycomya safena Väisänen, 1984

Mycomya shermani Garrett, 1924

= kingi Edwards, 1941

Mycomya shewelli Väisänen, 1984

Mycomya siebecki (Landrock, 1912)

Mycomya sieberti (Landrock, 1930)

Mycomya sigma Johannsen, 1910

= duplicata Edwards, 1925

Mycomya simulans Väisänen, 1984

Mycomya spinicoxa Väisänen, 1979

Mycomya storai Väisänen, 1979

Mycomya subarctica Väisänen, 1979

Mycomya tenuis (Walker, 1856)

Mycomya thula Väisänen, 1984

Mycomya trivittata (Zetterstedt, 1838)

Mycomya tumida (Winnertz, 1864)

Mycomya vittiventris (Zetterstedt, 1852)

= elegans (Lundström, 1912)

Mycomya wankowiczii (Dziedzicki, 1885)

Mycomya winnertzi (Dziedzicki, 1885)

sg. Mycomyopsis Väisänen, 1984

Mycomya affinis (Staeger, 1840)

= flava (Winnertz, 1864)

Mycomya confusa Väisänen, 1977

Mycomya fennica Väisänen, 1979

Mycomya maura (Walker, 1856)

Mycomya neolittoralis Väisänen, 1984

Mycomya paradentata Väisänen, 1984

= dentata auct. nec Fisher, 1937 misid.

Mycomya penicillata (Dziedzicki, 1885)

Mycomya permixta Väisänen, 1984

Mycomya trilineata (Zetterstedt, 1838)

= fusca (Meigen, 1818)

sg. Neomycomya Väisänen, 1984

Mycomya fimbriata (Meigen, 1818)

NEOEMPHERIA Osten Sacken, 1878

Neoempheria bimaculata (von Roser, 1840)

Neoempheria pictipennis (Haliday, 1833)

Neoempheria striata (Meigen, 1818)

Neoempheria tuomikoskii Väisänen, 1982

Neoempheria winnertzi Edwards, 1913

SCIOPHILINAE Rondani, 1840

ACNEMIA Winnertz, 1864

Acnemia amoena Winnertz, 1864

Acnemia angusta Zaitzev, 1982

Acnemia falcata Zaitzev, 1982

Acnemia longipes Winnertz, 1864

Acnemia nitidicollis (Meigen, 1818)

Acnemia trifida Zaitzev, 1982

ALLOCOTOCERA Mik, 1886

Allocotocera pulchella (Curtis, 1837)

ANACLILEIA Meunier, 1904

Anaclileia dispar (Winnertz, 1864)

Anaclileia dziedzickii (Landrock, 1911)

AZANA Walker, 1856

sg. Azana Walker, 1856

Azana anomala (Staeger, 1840)

COELOPHTHINIA Edwards, 1941

Coelophthinia thoracica (Winnertz, 1864)

EUDICRANA Loew, 1869

Eudicrana nigriceps (Lundström, 1909)

LEPTOMORPHUS Curtis, 1831

sg. Leptomorphus Curtis, 1831

Leptomorphus forcipatus Landrock, 1918

Leptomorphus subforcipatus Zaitzev & Ševčík, 2002

= quadrimaculatus auct. nec Matsumura, 1910 misid. (see Notes)

Leptomorphus walkeri Curtis, 1831

MEGALOPELMA Enderlein, 1911

Megalopelma nigroclavatum (Strobl, 1910)

MONOCLONA Mik, 1886

Monoclona braueri (Strobl, 1895)

Monoclona rufilatera (Walker, 1837)

Monoclona silvatica Zaitzev, 1983

= mikii Kertesz,1898 misid. (see Notes)

NEURATELIA Rondani, 1856

Neuratelia nemoralis (Meigen, 1818)

Neuratelia sintenisi Lackschewitz, 1937

PARATINIA Mik, 1874

Paratinia sciarina Mik, 1874

PHTHINIA Winnertz, 1864

Phthinia congenita Plassmann, 1984

Phthinia humilis Winnertz, 1864

Phthinia mira (Ostroverkhova, 1977)

= plassmanni Caspers, 1984

Phthinia setosa Zaitzev, 1994

Phthinia winnertzi Mik, 1869

POLYLEPTA Winnertz, 1864

Polylepta borealis Lundström, 1912

Polylepta guttiventris (Zetterstedt, 1852)

SCIOPHILA Meigen, 1818

Sciophila adamsi Edwards, 1925

Sciophila antiqua Chandler, 1987

= hebes auct. nec Johannsen, 1910 misid. (see Notes)

Sciophila bicuspidata Zaitzev, 1982

Sciophila buxtoni Freeman, 1956

Sciophila caesarea Chandler, 2001

Sciophila dziedzickii Edwards, 1925

Sciophila fenestella Curtis, 1837

Sciophila fuliginosa Holmgren, 1883

Sciophila geniculata Zetterstedt, 1838

Sciophila hirta Meigen, 1818

Sciophila karelica Zaitzev, 1982

Sciophila krysheni Polevoi, 2001

Sciophila limbatella Zetterstedt, 1852

Sciophila lutea Macquart, 1826

Sciophila minuta Zaitzev, 1982

Sciophila modesta Zaitzev, 1982

Sciophila nigronitida Landrock, 1925

Sciophila nonnisilva Hutson, 1979

Sciophila persubtilis Polevoi, 2001

Sciophila plurisetosa Edwards, 1921

Sciophila pomacea Chandler, 2006

= ochracea Stephens in Walker, 1856 preocc.

Sciophila pseudoflexuosa Kurina, 1991

Sciophila rufa Meigen, 1830

Sciophila salassea Matile, 1983

Sciophila setosa Garrett, 1925

Sciophila thoracica Staeger, 1840

= quadriterga (Hutson, 1979)

Sciophila varia (Winnertz, 1864)

Sciophila yakutica Blagoderov, 1992

GNORISTINAE Edwards, 1925

ACOMOPTERA Vockeroth, 1980

Acomoptera difficilis (Dziedzicki 1885)

Acomoptera spinistylus (Søli, 1993)

APOLEPHTHISA Grzegorzek, 1885

Apolephthisa subincana (Curtis, 1837)

BOLETINA Staeger, 1840

Boletina atridentata Polevoi & Hedmark, 2004

Boletina basalis (Meigen, 1818)

Boletina bidenticulata Sasakawa & Kimura, 1974

Boletina borealis Zetterstedt, 1852

Boletina brevicornis Zetterstedt, 1852

Boletina cincticornis (Walker, 1848)

Boletina cordata Polevoi & Hedmark, 2004

Boletina cornuta Zaitzev, 1994

Boletina digitata Lundström, 1914

Boletina dispecta Dziedzicki, 1885

Boletina dispectoides Jakovlev & Penttinen, 2007

Boletina dissipata Plassmann, 1986

Boletina dubia (Meigen, 1804)

Boletina edwardsi Chandler, 1992

Boletina falcata Polevoi & Hedmark, 2004

Boletina fennoscandica Polevoi & Hedmark, 2004

Boletina gripha Dziedzicki, 1885

Boletina griphoides Edwards, 1925

Boletina groenlandica Staeger, 1845

Boletina gusakovae Zaitzev, 1994

Boletina hedstroemi Polevoi & Hedmark, 2004

Boletina intermedia Lundström, 1915

Boletina jamalensis Zaitzev, 1994

= struthioides Polevoi & Hedmark, 2004

Boletina kivachiana Polevoi & Hedmark, 2004

Boletina kurilensis Zaitzev, 1994

Boletina landrocki Edwards, 1924

Boletina lapponica Polevoi & Hedmark, 2004

Boletina lundbecki Lundström, 1912

Boletina lundstroemi Landrock, 1912

Boletina maculata Holmgren, 1870

= apicalis (Walker, 1848) preocc.

Boletina minuta Polevoi in Zaitzev & Polevoi, 1995

Boletina moravica Landrock, 1912

Boletina nasuta (Haliday, 1839)

Boletina nigricans Dziedzicki, 1885

Boletina nigricoxa Staeger, 1840

Boletina nigrofusca Dziedzicki, 1885

Boletina nitida Grzegorzek, 1885

Boletina nitiduloides Zaitzev, 1994

Boletina onegensis Polevoi in Zaitzev & Polevoi, 1995

Boletina pallidula Edwards, 1925

Boletina palmata Polevoi, 2013

Boletina pectinunguis Edwards, 1932

Boletina pinusia Maximova, 2001

Boletina plana (Walker, 1856)

Boletina polaris Lundström, 1915

Boletina populina Polevoi in Zaitzev & Polevoi, 1995

Boletina pseudonitida Zaitzev, 1994

Boletina rejecta Edwards, 1941

Boletina sciarina Staeger, 1840

Boletina silvatica Dziedzicki, 1885

Boletina subtriangularis Polevoi & Hedmark, 2004

Boletina takagii Sasakawa & Kimura, 1974

Boletina tiroliensis Plassmann, 1980

Boletina triangularis Polevoi in Zaitzev & Polevoi, 1995

Boletina trispinosa Edwards, 1913

Boletina trivittata (Meigen, 1818)

Boletina verticillata Stackelberg, 1943

Boletina villosa Landrock, 1912

COELOSIA Winnertz, 1864

Coelosia bicornis Stackelberg, 1946

Coelosia flava (Staeger, 1840)

Coelosia fusca Bezzi, 1892

= silvatica Landrock, 1918

Coelosia gracilis Johannsen, 1912

Coelosia limpida Plassmann, 1986

Coelosia tenella (Zetterstedt, 1852)

Coelosia truncata Lundström, 1909

DZIEDZICKIA Johannsen, 1909

Dziedzickia marginata (Dziedzicki, 1885)

ECTREPESTHONEURA Enderlein, 1911

Ectrepesthoneura colyeri Chandler, 1980

Ectrepesthoneura hirta (Winnertz, 1846)

Ectrepesthoneura nigra Zaitzev, 1984

Ectrepesthoneura ovata Ostroverkhova, 1977

= bucera Plassmann, 1980

Ectrepesthoneura pubescens (Zetterstedt, 1860)

Ectrepesthoneura referta Plassmann, 1976

Ectrepesthoneura tori Zaitzev & Økland, 1994

GNORISTE Meigen, 1818

Gnoriste apicalis Meigen, 1818

Gnoriste bilineata Zetterstedt, 1852

GRZEGORZEKIA Edwards, 1941

Grzegorzekia collaris (Meigen, 1818)

HADRONEURA Lundström, 1906

Hadroneura palmeni Lundström, 1906

IMPLETA Plassmann, 1978

Impleta consorta Plassmann, 1978

KATATOPYGIA Martinsson & Kjærandsen, 2012

Katatopygia erythropyga (Holmgren, 1883)

Katatopygia sahlbergi (Lundström, 1906)

PALAEODOCOSIA Meunier, 1904

Palaeodocosia vittata (Coquillett, 1901)

= janickii (Dziedzicki, 1923)

= alpicola auct. nec (Strobl, 1894) misid. (see Notes)

SAIGUSAIA Vockeroth, 1980

Saigusaia flaviventris (Strobl, 1894)

SPEOLEPTA Edwards, 1925

Speolepta leptogaster (Winnertz, 1864)

SYNAPHA Meigen, 1818

Synapha fasciata Meigen, 1818

Synapha vitripennis (Meigen, 1818)

SYNTEMNA Winnertz, 1864

Syntemna daisetsuzana Okada, 1938

Syntemna elegantia Plassmann, 1978

Syntemna hungarica (Lundström, 1912)

Syntemna morosa Winnertz, 1864

Syntemna nitidula Edwards, 1925

Syntemna oulankaensis Polevoi, 2003

Syntemna penicilla Hutson, 1979

Syntemna relicta (Lundström, 1912)

Syntemna setigera (Lundström, 1914)

= haagvari Økland, 1995

Syntemna stylata Hutson, 1979

Syntemna stylatoides Zaitzev, 1994

TETRAGONEURA Winnertz, 1846

Tetragoneura ambigua Grzegorzek, 1885

Tetragoneura obirata Plassmann, 1990

Tetragoneura pudogensis Polevoi & Jakovlev, 2011

Tetragoneura ruuhijarvii Polevoi & Jakovlev, 2011

Tetragoneura sylvatica (Curtis, 1837)

LEIINAE Edwards, 1925

CLASTOBASIS Skuse, 1890

Clastobasis alternans (Winnertz, 1864)

DOCOSIA Winnertz, 1864

Docosia expectata Laštovka & Ševčík, 2006

Docosia flavicoxa Strobl, 1900

= pallipes Edwards, 1941

Docosia gilvipes (Haliday in Walker, 1856)

Docosia landrocki Laštovka & Ševčík, 2006

Docosia muelleri Plassmann, 1986

Docosia tibialis Laštovka & Ševčík, 2006

GREENOMYIA Brunetti, 1912

Greenomyia baikalica Zaitzev, 1994

Greenomyia borealis (Winnertz, 1864)

Greenomyia mongolica Laštovka & Matile, 1974

LEIA Meigen, 1818

Leia bilineata (Winnertz, 1864)

Leia bimaculata (Meigen, 1804)

Leia crucigera Zetterstedt, 1838

Leia cylindrica (Winnertz, 1864)

Leia fascipennis Meigen, 1818

Leia longiseta Barendrecht, 1938

Leia picta Meigen, 1818

Leia subfasciata (Meigen, 1818)

Leia winthemii Lehmann, 1822

RONDANIELLA Johannsen, 1909

Rondaniella dimidiata (Meigen, 1804)

MANOTINAE Edwards, 1925

MANOTA Williston, 1896

Manota unifurcata Lundström, 1913

MYCETOPHILINAE Newman, 1834

tribe Exechiini Edwards, 1925

ALLODIA Winnertz, 1864

sg. Allodia Winnertz, 1864

Allodia anglofennica Edwards, 1921

Allodia confusa Zaitzev, 2003

= simplex Zaitzev, 1983 preocc.

Allodia embla Hackman, 1971

Allodia lugens (Wiedemann, 1817)

Allodia lundstroemi Edwards, 1921

Allodia ornaticollis (Meigen, 1818)

Allodia pyxidiiformis Zaitzev, 1983

Allodia septentrionalis Hackman, 1971

Allodia truncata Edwards, 1921

Allodia tuomikoskii Hackman, 1971

Allodia zaitzevi Kurina, 1998

sg. Brachycampta Winnertz, 1864

Allodia adunca Zaitzev, 1992

Allodia alternans (Zetterstedt, 1838)

Allodia angulata (Lundström, 1913)

Allodia barbata (Lundström, 1909)

Allodia bohemica Ševčík, 2004

Allodia czernyi (Landrock, 1912)

Allodia foliifera (Strobl, 1910)

Allodia grata (Meigen, 1830)

Allodia huggerti Kjærandsen, 2007

Allodia neglecta Edwards, 1925

Allodia penicillata (Lundström, 1912)

Allodia pistillata (Lundström, 1911)

Allodia protenta Laštovka & Matile, 1974

= mendli Plassmann, 1977

Allodia silvatica (Landrock, 1912)

Allodia subpistillata Ševčík, 1999

ALLODIOPSIS Tuomikoski, 1966

Allodiopsis domestica (Meigen, 1830)

Allodiopsis gracai Ševčík & Papp 2003

Allodiopsis korolevi Zaitzev, 1982

Allodiopsis pseudodomestica (Lackschewitz, 1937)

Allodiopsis rustica (Edwards, 1941)

ANATELLA Winnertz, 1864

Anatella ankeli Plassmann, 1977

Anatella aquila Zaitzev, 1989

Anatella bremia Chandler, 1994

Anatella ciliata Winnertz, 1864

Anatella crispa Zaitzev, 1994

Anatella dampfi Landrock, 1924

Anatella dentata Zaitzev, 1989

Anatella emergens Caspers, 1987

Anatella flavicauda Winnertz, 1864

Anatella flavomaculata Edwards, 1925

Anatella gibba Winnertz, 1864

Anatella lenis Dziedzicki, 1923

Anatella maritima Ostroverkhova, 1979

Anatella minuta (Staeger, 1840)

Anatella novata Dziedzicki, 1923

Anatella setigera Edwards, 1921

Anatella simpatica Dziedzicki, 1923

Anatella turi Dziedzicki, 1923

Anatella unguigera Edwards, 1921

BRACHYPEZA Winnertz, 1864

sg. Brachypeza Winnertz, 1864

Brachypeza armata Winnertz, 1864

Brachypeza bisignata Winnertz, 1864

= hilaris Winnertz, 1864 preocc.

sg. Paracordyla Tuomikoski, 1966

Brachypeza obscura Winnertz, 1864

BREVICORNU Marshall, 1896

Brevicornu arcticoides Caspers, 1985

Brevicornu arcticum (Lundström, 1913)

Brevicornu auriculatum (Edwards, 1925)

Brevicornu beatum (Johannsen, 1912)

Brevicornu bellum (Johannsen, 1912)

Brevicornu bipartitum Laštovka & Matile, 1974

Brevicornu canescens (Zetterstedt, 1852)

= griseolum auct. nec (Zetterstedt, 1852)

Brevicornu cognatum Ostroverkhova, 1979

Brevicornu fasciculatum (Lackschewitz, 1937)

Brevicornu fennicum (Landrock, 1927)

Brevicornu fissicauda (Lundström, 1911)

Brevicornu foliatum (Edwards, 1925)

Brevicornu fuscipenne (Staeger, 1840)

Brevicornu glandis Laštovka & Matile, 1974

Brevicornu griseicolle (Staeger, 1840)

Brevicornu griseolum (Zetterstedt, 1852)

= boreale (Lundström, 1914)

Brevicornu improvisum Zaitzev, 1992

Brevicornu kingi (Edwards, 1925)

Brevicornu luteum (Landrock, 1925)

Brevicornu melanderi Zaitzev, 1988

Brevicornu nigrofuscum (Lundström, 1909)

Brevicornu occidentale Zaitzev, 1988

Brevicornu parafennicum Zaitzev in Zaitzev & Polevoi, 1995

Brevicornu proximum (Staeger, 1840)

Brevicornu rosmellitum Chandler, 2001

Brevicornu ruficorne (Meigen, 1838)

Brevicornu serenum (Winnertz, 1864)

Brevicornu sericoma (Meigen, 1830)

Brevicornu setulosum Zaitzev, 1988

Brevicornu spathulatum (Lundström, 1911)

Brevicornu verralli (Edwards, 1925)

CORDYLA Meigen, 1803

Cordyla brevicornis (Staeger, 1840)

Cordyla crassicornis Meigen, 1818

Cordyla fasciata Meigen, 1830

Cordyla fissa Edwards, 1925

Cordyla flaviceps (Staeger, 1840)

Cordyla fusca Meigen, 1804

Cordyla insons Laštovka & Matile, 1974

Cordyla murina Winnertz, 1864

Cordyla nitens Winnertz, 1864

Cordyla nitidula Edwards, 1925

= bergensis (Barendrecht, 1938)

Cordyla parvipalpis Edwards, 1925

Cordyla pusilla Edwards, 1925

= sixi (Barendrecht, 1938)

Cordyla semiflava (Staeger, 1840)

EXECHIA Winnertz, 1864

Exechia bicincta (Staeger, 1840)

Exechia borealis Lundström, 1912

Exechia cincta Winnertz, 1864

Exechia confinis Winnertz, 1864

Exechia contaminata Winnertz, 1864

Exechia cornuta Lundström, 1914

Exechia dentata Lundström, 1916

Exechia dizona Edwards, 1924

Exechia dorsalis (Staeger, 1840)

Exechia exigua Lundström, 1909

Exechia festiva Winnertz, 1864

Exechia frigida (Boheman, 1865)

Exechia fusca (Meigen, 1804)

Exechia lucidula (Zetterstedt, 1838)

Exechia lundstroemi Landrock, 1923

Exechia macula Chandler, 2001

= maculipennis (Stannius, 1831) preocc.

Exechia micans Laštovka & Matile, 1974

Exechia nigra Edwards, 1925

Exechia nigrofusca Lundström, 1909

Exechia nigroscutellata Landrock, 1912

Exechia papyracea Stackelberg, 1948

Exechia parva Lundström, 1909

Exechia parvula (Zetterstedt, 1852)

= nana Staeger, 1840 preocc.

Exechia pectinivalva Stackelberg, 1948

Exechia pseudocincta Strobl, 1910

Exechia pseudofestiva Lackschewitz, 1937

Exechia repanda Johannsen, 1912

Exechia repandoides Caspers, 1984

Exechia separata Lundström, 1912

Exechia seriata (Meigen, 1830)

= pallida Stannius, 1831

Exechia similis Laštovka & Matile, 1974

Exechia spinigera Winnertz, 1864

Exechia spinuligera Lundström, 1912

Exechia styriaca Strobl, 1898

= sororcula Lackschewitz, 1937

Exechia subfrigida Laštovka & Matile, 1974

Exechia unifasciata Lackschewitz, 1937

Exechia unimaculata (Zetterstedt, 1860)

EXECHIOPSIS Tuomikoski, 1966

sg. Exechiopsis Tuomikoski, 1966

Exechiopsis aemula Plassmann, 1984

Exechiopsis clypeata (Lundström, 1911)

Exechiopsis distendens (Lackschewitz, 1937)

Exechiopsis dumitrescae (Burghele-Balacesco, 1972)

Exechiopsis fimbriata (Lundström, 1909)

Exechiopsis forcipata Lackschewitz, 1937)

Exechiopsis grassatura (Plassmann, 1978)

Exechiopsis hammi (Edwards, 1925)

Exechiopsis indecisa (Walker, 1856)

Exechiopsis ingrica Stackelberg, 1948

Exechiopsis intersecta (Meigen, 1818)

Exechiopsis januarii (Lundström, 1913)

Exechiopsis lackschewitziana (Stackelberg, 1948)

Exechiopsis landrocki (Lundström, 1912)

Exechiopsis ligulata (Lundström, 1913)

Exechiopsis pseudindecisa Laštovka & Matile, 1974

Exechiopsis pseudopulchella (Lundström, 1912)

Exechiopsis pulchella (Winnertz, 1864)

Exechiopsis sagittata Laštovka & Matile, 1974

Exechiopsis subulata (Winnertz, 1864)

sg. Xenexechia Tuomikoski, 1966

Exechiopsis crucigera (Lundström, 1909)

Exechiopsis davatchii Matile, 1969

Exechiopsis leptura (Meigen, 1830)

Exechiopsis membranacea (Lundström, 1912)

Exechiopsis perspicua (Johannsen, 1912)

Exechiopsis pollicata (Edwards, 1925)

Exechiopsis praedita (Plassmann, 1976)

Exechiopsis seducta (Plassmann, 1976)

Exechiopsis stylata Laštovka & Matile, 1974

MYROSIA Tuomikoski, 1966

Myrosia maculosa (Meigen, 1818)

NOTOLOPHA Tuomikoski, 1966

Notolopha brachycera (Zetterstedt, 1852)

= tuomikoskii (Zaitzev & Maximova, 2000)

Notolopha cristata (Staeger, 1840)

Notolopha sibirica (Zaitzev & Maximova, 2000)

PSEUDEXECHIA Tuomikoski, 1966

Pseudexechia aurivernica Chandler, 1978

Pseudexechia canalicula (Johannsen, 1912)

Pseudexechia parallela (Edwards, 1925)

Pseudexechia pectinacea (Ostroverkhova, 1979)

Pseudexechia tristriata Stackelberg, 1969

Pseudexechia trivittata (Staeger, 1840)

PSEUDOBRACHYPEZA Tuomikoski, 1966

Pseudobrachypeza helvetica (Walker, 1856)

PSEUDORYMOSIA Tuomikoski, 1966

Pseudorymosia fovea (Dziedzicki, 1910)

= optiva auct. nec (Dziedzicki, 1910) misid. (see Notes)

RYMOSIA Winnertz, 1864

Rymosia acta Dziedzicki, 1910

Rymosia affinis Winnertz, 1864

= gracilipes Dziedzicki, 1910

Rymosia batava (Barendrecht, 1938)

Rymosia bifida Edwards, 1925

Rymosia britteni Edwards, 1925

Rymosia connexa Winnertz, 1864

Rymosia fasciata (Meigen, 1804)

Rymosia fraudatrix Dziedzicki, 1910

? = spiniforceps Matile, 1963

Rymosia guttata Lundström, 1912

Rymosia pinnata Ostroverkhova, 1979

Rymosia placida Winnertz, 1864

Rymosia setiger Dziedzicki, 1910

Rymosia signatipes (van der Wulp, 1859)

= winnertzi Barendrecht, 1938

STIGMATOMERIA Tuomikoski, 1966

Stigmatomeria crassicornis (Stannius, 1831)

SYNPLASTA Skuse, 1890

= Gymnogonia Tuomikoski, 1966

Synplasta bayardi Matile, 1971

Synplasta dulcia (Dziedzicki, 1910)

Synplasta exclusa (Dziedzicki, 1910)

= sintenisi (Lackschewitz, 1937)

Synplasta gracilis Winnertz, 1864

= excogitata auct. nec (Dziedzicki, 1910) misid. (see Notes)

Synplasta ingeniosa (Kidd 1969)

Synplasta praeformida (Dziedzicki, 1910)

Synplasta pseudingeniosa Zaitzev, 1993

Synplasta rufilatera (Edwards, 1941)

TARNANIA Tuomikoski, 1966

Tarnania fenestralis (Meigen, 1838)

Tarnania tarnanii (Dziedzicki, 1910)

tribe Mycetophilini Newman, 1834

DYNATOSOMA Winnertz, 1864

Dynatosoma cochleare Strobl, 1895

Dynatosoma dihaeta Polevoi, 1995

= schachti Plassmann, 1999

Dynatosoma fuscicorne (Meigen, 1818)

Dynatosoma majus Landrock, 1912

Dynatosoma nigromaculatum Lundström, 1913

= abdominale auct. nec Staeger, 1840

Dynatosoma nobile Loew, 1873

Dynatosoma reciprocum (Walker, 1848)

Dynatosoma rufescens (Zetterstedt, 1838)

= bukowskii Zaitzev, 1986

= lutescens (Zetterstedt, 1852)

Dynatosoma silesiacum Ševčík, 2001

Dynatosoma thoracicum (Zetterstedt, 1838)

= norwegiense Zaitzev & Økland, 1994

EPICYPTA Winnertz, 1864

Epicypta aterrima (Zetterstedt, 1852)

Epicypta fumigata (Dziedzicki, 1923)

Epicypta limnophila Chandler, 1981

Epicypta scatophora (Perris, 1849)

MYCETOPHILA Meigen, 1803

= FUNGIVORA Meigen, 1800 suppr.

Mycetophila abbreviata Landrock, 1914

Mycetophila abiecta (Laštovka, 1963)

Mycetophila adumbrata Mik, 1884

Mycetophila alea Laffoon, 1965

= guttata Dziedzicki, 1884

Mycetophila attonsa (Laffoon, 1957)

= uncta Plassmann, 1999

Mycetophila autumnalis Lundström, 1909

Mycetophila bialorussica Dziedzicki, 1884

* Mycetophila biformis Maximova, 2002 preocc. (see Notes)

Mycetophila biusta Meigen, 1818

Mycetophila blanda Winnertz, 1864

Mycetophila bohemica (Laštovka, 1963)

Mycetophila boreocruciator Ševčík, 2003

Mycetophila brevitarsata (Laštovka, 1963)

Mycetophila caudata Staeger, 1840

Mycetophila cingulum Meigen, 1830

Mycetophila confluens Dziedzicki, 1884

= fulva Winnertz, 1864

Mycetophila confusa Dziedzicki, 1884

= affluctata Edwards, 1941

Mycetophila curviseta Lundström, 1911

Mycetophila deflexa Chandler, 2001

Mycetophila dentata Lundström, 1915

Mycetophila devioides Bechev, 1988

Mycetophila distigma Meigen, 1830

= w-fuscum Dziedzicki, 1884

Mycetophila dziedzickii Chandler, 1977

= obscura Dziedzicki, 1884 nec Walker, 1848

Mycetophila edwardsi Lundström, 1913

Mycetophila estonica Kurina, 1992

Mycetophila evanida Laštovka, 1972

Mycetophila exstincta Loew, 1869

= mikii Dziedzicki, 1884

Mycetophila finlandica Edwards, 1913

Mycetophila flava Winnertz, 1864

Mycetophila forcipata Lundström, 1913

Mycetophila formosa Lundström, 1911

Mycetophila freyii Lundström, 1909

Mycetophila fungorum (De Geer, 1776)

Mycetophila gibbula Edwards, 1925

Mycetophila hetschkoi Landrock, 1918

Mycetophila ichneumonea Say, 1823

Mycetophila idonea Laštovka, 1972

Mycetophila immaculata (Dziedzicki, 1884)

Mycetophila laeta Walker, 1848

= calva Lundström, 1912

Mycetophila lapponica Lundström, 1906

Mycetophila lobulata Zaitzev, 1999

Mycetophila lubomirskii Dziedzicki, 1884

Mycetophila luctuosa Meigen, 1830

= modesta Winnertz, 1864

Mycetophila magnicauda Strobl, 1895

Mycetophila marginata Winnertz, 1864

Mycetophila mitis (Johannsen, 1912)

Mycetophila mohilevensis Dziedzicki, 1884

Mycetophila monstera Maximova, 2002

Mycetophila morata Zaitzev, 1999

Mycetophila moravica Landrock, 1925

Mycetophila morosa Winnertz, 1864

Mycetophila nigrofusca Dziedzicki, 1884

Mycetophila occultans Lundström, 1913

= tarsata Winnertz, 1864 preocc.

Mycetophila ocellus Walker, 1848

Mycetophila ornata Stephens, 1846

Mycetophila ostentanea Zaitzev, 1998

Mycetophila pecinai (Laštovka, 1963)

Mycetophila perpallida Chandler, 1993

Mycetophila pictula Meigen, 1830

= bimaculata Fabricius, 1805 nec Meigen, 1804

Mycetophila pseudoforcipata Zaitzev, 1998

Mycetophila pumila Winnertz, 1864

Mycetophila pyrenaica Matile, 1967

Mycetophila quadra Lundström, 1909

Mycetophila ruficollis Meigen, 1818

Mycetophila schnabli (Dziedzicki, 1884)

Mycetophila sigmoides Loew, 1869

Mycetophila signata Meigen, 1830

Mycetophila signatoides Dziedzicki, 1884

= assimilis Matile, 1967

Mycetophila sinuosa Plassmann & Schacht, 1999

Mycetophila sordida van der Wulp, 1874

Mycetophila stolida Walker, 1856

Mycetophila stricklandi (Laffoon, 1957)

Mycetophila strigata Staeger, 1840

Mycetophila strigatoides Landrock, 1927

Mycetophila strobli Laštovka, 1972

Mycetophila stylata (Dziedzicki, 1884)

Mycetophila sublunata Zaitzev, 1998

Mycetophila subsigillata Zaitzev, 1999

Mycetophila sumavica (Laštovka, 1963)

Mycetophila triangularis Lundström, 1912

Mycetophila trinotata Staeger, 1840

Mycetophila uliginosa Chandler, 1988

Mycetophila unguiculata Lundström, 1913

Mycetophila uninotata Zetterstedt, 1852

Mycetophila unipunctata Meigen, 1818

Mycetophila xanthopyga Winnertz, 1864

Mycetophila zetterstedtii Lundström, 1906

PHRONIA Winnertz, 1864

Phronia aviculata Lundström, 1914

Phronia avidoides Jakovlev & Polevoi, 2009

Phronia biarcuata (Becker, 1908)

= johannae Steenberg, 1924

= praecox Edwards, 1925

Phronia bicolor Dziedzicki, 1889

= fusciventris van Duzee, 1928

Phronia borealis Hackman, 1970

Phronia braueri Dziedzicki, 1889

= annulata Winnertz, 1864

Phronia caliginosa Dziedzicki, 1889

= trivittata Dziedzicki, 1889

Phronia cinerascens Winnertz, 1864

Phronia conformis (Walker, 1856)

= girschneri Dziedzicki, 1889

Phronia cordata Lundström, 1914

Phronia coritanica Chandler, 1992

Phronia cornuta Lundström, 1914

Phronia crassitarsus Hackman, 1970

Phronia digitata Hackman, 1970

Phronia disgrega Dziedzicki, 1889

Phronia distincta Hackman, 1970

Phronia dziedzickii Lundström, 1906

Phronia egregia Dziedzicki, 1889

Phronia electa Dziedzicki, 1889

Phronia elegans Dziedzicki, 1889

Phronia elegantula Hackman, 1970

Phronia exigua (Zetterstedt, 1852)

Phronia fennica Jakovlev & Polevoi, 2009

Phronia flavipes Winnertz, 1864

Phronia forcipata Winnertz, 1864

Phronia forcipula Winnertz, 1864

Phronia gagnei Chandler, 1992

Phronia gracilis Hackman, 1970

Phronia humeralis Winnertz, 1864

Phronia interstincta Dziedzicki, 1889

Phronia longelamellata Strobl, 1898

= minuta Landrock, 1928

Phronia lutescens Hackman, 1970

Phronia maculata Dziedzicki, 1889

Phronia mutabilis Dziedzicki, 1889

Phronia mutila Lundström, 1911

Phronia nigricornis (Zetterstedt, 1852)

= dubia Dziedzicki, 1889

Phronia nigripalpis Lundström, 1909

Phronia nitidiventris (van der Wulp, 1859)

= squalida Winnertz, 1864

= vitiosa Winnertz, 1864

Phronia notata Dziedzicki, 1889

Phronia obscura Dziedzicki, 1889

Phronia obtusa Winnertz, 1864

Phronia peculiaris Dziedzicki, 1889

Phronia persimilis Hackman, 1970

Phronia petulans Dziedzicki, 1889

Phronia portschinskyi Dziedzicki, 1889

Phronia siebeckii Dziedzicki, 1889

Phronia signata Winnertz, 1864

= austriaca Winnertz, 1864

Phronia spinigera Hackman, 1970

Phronia strenua Winnertz, 1864

= flavicollis Winnertz, 1864 nom. dubium

Phronia subsilvatica Hackman, 1970

Phronia sudetica Dziedzicki, 1889

Phronia sylvatica Dziedzicki, 1889

Phronia taczanowskyi Dziedzicki, 1889

Phronia tenuis Winnertz, 1864

Phronia tiefii Dziedzicki, 1889

= marginata Dziedzicki, 1889

Phronia triangularis Winnertz, 1864

Phronia unica Dziedzicki, 1889

Phronia vitrea Plassmann, 1999

= carli Chandler, 2001

Phronia willistoni Dziedzicki, 1889

PLATUROCYPTA Enderlein, 1910

Platurocypta punctum (Stannius, 1831)

Platurocypta testata (Edwards, 1925)

SCEPTONIA Winnertz, 1864

Sceptonia concolor Winnertz, 1864

Sceptonia costata (van der Wulp, 1859)

Sceptonia demeijerei Bechev, 1997

Sceptonia flavipuncta Edwards, 1925

Sceptonia fumipes Edwards, 1925

Sceptonia fuscipalpis Edwards, 1925

Sceptonia hamata Ševčík, 2004

Sceptonia longisetosa Ševčík, 2004

Sceptonia membranacea Edwards, 1925

Sceptonia nigra (Meigen, 1804)

Sceptonia pughi Chandler, 1991

Sceptonia regni Chandler, 1991

Sceptonia tenuis Edwards, 1925

Sceptonia thaya Ševčík, 2004

TRICHONTA Winnertz, 1864

Trichonta apicalis Strobl, 1898

Trichonta atricauda (Zetterstedt, 1852)

= adunca Edwards, 1925

Trichonta beata Gagné, 1981

Trichonta bicolor Landrock, 1912

Trichonta bifida Lundström, 1909

Trichonta brevicauda Lundström, 1909

Trichonta canora Gagné, 1981

Trichonta clara Gagné, 1981

Trichonta comica Gagné, 1981

Trichonta comis Gagné, 1981

Trichonta concinna Gagné, 1981

Trichonta conjungens Lundström, 1909

Trichonta delicata Gagné, 1981

Trichonta eximia Gagné, 1981

Trichonta facilis Gagné, 1981

Trichonta falcata Lundström, 1911

Trichonta fissicauda (Zetterstedt, 1852)

= claripennis Lundström, 1914

Trichonta flavicauda Lundström, 1914

Trichonta fragilis Gagné, 1981

Trichonta fusca Landrock, 1918

Trichonta generosa Gagné, 1981

Trichonta girschneri Landrock, 1912

Trichonta hamata Mik, 1880

Trichonta icenica Edwards, 1925

Trichonta melanura (Staeger, 1840)

Trichonta nigritula Edwards, 1925

Trichonta palustris Maximova, 2002

Trichonta patens Johannsen, 1912

Trichonta perspicua van der Wulp, 1881

= mediastinalis (Lundström, 1909)

Trichonta subfusca Lundström, 1909

Trichonta submaculata (Staeger, 1840)

Trichonta subterminalis Zaitzev & Menzel, 1996

Trichonta terminalis (Walker, 1856)

* Trichonta tristis (Strobl, 1898) (see Notes)

Trichonta trivittata Lundström, 1916

Trichonta venosa (Staeger, 1840)

Trichonta vitta (Meigen, 1830)

Trichonta vulcani (Dziedzicki, 1889)

= appropinquata (Strobl, 1900)

? = trifida Lundström, 1909 (see Notes)

Trichonta vulgaris Loew, 1869

= nigricauda Lundström, 1906

ZYGOMYIA Winnertz, 1864

Zygomyia angusta Plassmann, 1977

Zygomyia humeralis (Wiedemann, 1817)

Zygomyia kiddi Chandler, 1991

Zygomyia notata (Stannius, 1831)

Zygomyia pictipennis (Staeger, 1840)

Zygomyia pseudohumeralis Caspers, 1980

Zygomyia semifusca (Meigen, 1818) (see Notes)

Zygomyia valida Winnertz, 1864

Zygomyia vara (Staeger, 1840)

Zygomyia zaitzevi Chandler, 1991

Excluded species

Bolitophilidae

Bolitophila spinigera Edwards, 1925 misidentified

Keroplatidae

Macrocera nigropicea Lundström, 1906 not found within present borders

Macrocera sudetica Landrock, 1924 only females, unidentifiable (see Notes)

Orfelia bicolor (Macquart, 1826) only females, unidentifiable (see Notes)

Mycetophilidae

Allodia triangularis (Strobl, 1895) misidentified

Cordyla styliforceps (Bukowski, 1934) misidentified (see Notes)

Exechiopsis unguiculata (Lundström, 1911) not found within present borders

Leptomorphus quadrimaculatus Matsumura, 1916 misidentified (see Notes)

Monoclona mikii Kertesz, 1898 misidentified (see Notes)

Mycetophila gratiosa Winnertz, 1864 misidentified (see Notes)

Mycetophila lunata Meigen, 1804 misidentified (see Notes)

Mycetophila vittipes Zetterstedt, 1852 misidentified (see Notes)

Palaeodocosia alpicola (Strobl, 1895) misidentified (see Notes)

Pseudexechia trisignata (Edwards, 1913) not found within present borders

Pseudorymosia optiva (Dziedzicki, 1910) misidentified (see Notes)

Sciophila hebes Johannsen, 1910 misidentified (see Notes)

Synplasta excogitata (Dziedzicki, 1910) misidentified (see Notes)

Trichonta fidelis Gagné, 1981 mistake (see Notes)

Notes

Cordyla styliforceps (Bukowski, 1934) was added to the list of Finnish insects by Silfverberg (2007) with a reference to Polevoi (2001) which most probably based on misidentified species (A.Polevoi, pers. comm).

Leptomorphus quadrimaculatus Matsumura, 1916 was added to the list of Finnish insects by Silfverberg (2001) with a reference to Polevoi (1995). This record has later shown to belong to Leptomorphus subforcipatus (Zaitzev and Ševčík 2002).

Macrocera sudetica Landrock, 1924 and Orfelia bicolor (Macquart, 1826) (= basalis Winnertz, 1864). Listed in Hackman (1980) by the only female specimen located in the collection of the Finnish Museum of Natural History “Diptera Fennica” that cannot be recognised. No further records from Finland.

Monoclona mikii Kertesz, 1898. Listed in Hackman (1980) by erroneously identified male specimen of Monoclona braueri and by the female specimen (both are located in “Diptera Fennica”) that cannot be recognized. Later this species has been recorded from Finland under its valid name, Monoclona silvatica (Jakovlev et al. 2014).

Mycetophila gratiosa Winnertz, 1864. Listed in Hackman (1980) by erroneously identified male specimens of Mycetophila luctuosa located in “Diptera Fennica”.

Mycetophila lunata Meigen, 1804 was interpreted broadly in old sources, and although it was confirmed by Silfverberg (2001) with a reference to Polevoi (1995) this record has later shown to belong to another related species, M. sublunata Zaitzev, 1999. Hence, M. lunata still needs to be confirmed from Finland.

Mycetophila vittipes Zetterstedt, 1852. Listed in Hackman (1980) by erroneously identified male specimens of Mycetophila vittipes-group located in “Diptera Fennica”.

Palaeodocosia alpicola (Strobl, 1895) Listed in Hackman (1980) by erroneously identified male specimen of P. vittata located in “Diptera Fennica”.

Pseudorymosia optiva (Dziedzicki, 1910). The small genus Pseudorymosia contains only two known Palaearctic species, P. fovea (Dziedzicki, 1910) and P. optiva (Dziedzicki, 1910), that in fact might be only one species (Zaitzev 2003). Re-examination of specimens in the “Diptera Fennica” collections formerly identified as P. optiva and published by Hackman (1980) revealed that all of them are in fact the widely distributed P. fovea as interpreted in newer determination sources (Jakovlev et al. 2006).

Sciophila hebes Johannsen, 1910. Recorded by Komonen (2001) by erroneously identified specimens of Sciophila antiqua.

Synplasta excogitata (Dziedzicki, 1910). Listed in Hackman (1980) by erroneously identified male specimen of Synplasta gracilis.

Trichonta fidelis Gagné, 1981. Listed in Hackman (1980) with the note that T. fidelis Gagné was only a manuscript of Gagné (1981) at the time. Apparently Gagné changed his mind before publication and used this name for an oriental species instead (Hackman 1982).

Zygomyia semifusca (Meigen, 1818). Listed in Hackman (1980) as Mycetophila semifusca.

Doubtful and unrecognized species

* Keroplatus tuvensis Zaitzev, 1991. This species might be a junor synonym of Keroplatus testaceus (Kjærandsen et al. 2007). With this reference, it was deleted from the list of Finnish insects by Silfverberg (2012), but this synonym has not been adopted in the checklist of North European fungus gnats (Kjærandsen 2012). Hereby we keep K. tuvensis in the Finnish checklist pending revision of the K. testaceus complex of species.

* Mycetophila biformis Maximova, 2002 preocc. Added by Jakovlev et al. (2014). This is a junior primary homonym of Mycetophila biformis Duret, 1992. A replacement name will be given by the author in the near future (Yulia Maximova, pers. comm.).

* Trichonta tristis (Strobl, 1898). Judging from illustration of Dziedzicki (1889), he actually illustrated this species as Trichonta vulcani. This makes vulcani a junior synonym of tristis. In this case, Trichonta trifida Lundström, 1909 becomes the valid species that we now call vulcani (J. Kjærandsen, pers.comm.). Here we list T. vulcani and T. tristis as valid species sensu Zaitzev (2003), figures 107,1 and 109,5.

Species listed by Hackman (1980) with a question marks

Five species, Docosia flavicoxa Strobl, 1900, Cordyla bergensis (Barendrecht, 1938), Cordyla murina Winnertz, 1864, Mycetophila lunata Meigen, 1804 and Phronia petulans Dziedzicki, 1889 were listed in the former checklist of Finnish Diptera (Hackman 1980) with a question marks. Later, Finnish records were confirmed for D. flavicoxa by Jakovlev et al. (2014), for C. murina by Jakovlev et al. (2006), and for P. petulans by Jakovlev and Polevoi (2009). Cordyla bergensis was excluded as a junior synonym of Cordyla nitidula (Jakovlev et al. 2006). Mycetophila lunata – see above.

Acknowledgements

The author thanks Dr. Jukka Salmela (Finland) who has discovered tens of fungus gnat species new to Finland resulting from his studies of under recorded fauna of wetlands, mires and forests along brooks in northern boreal and oroarctic zones of Finnish Lapland. Jere Kahanpää and Dr. Jukka Salmela, the editors of the Checklist of Finnish Diptera, Dr. Jostein Kjærandsen (Norway), the editor of the the online checklist of Nordic fungus gnats (Kjærandsen 2012), Dr. Pekka Vilkamaa (Finland), and two referees: Dr. Jukka Salmela and Dr. Alexei Polevoi (Petrozavodsk, Russia) are thanked for useful comments on the manuscript.

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