Zoological nomenclature affects the work of all zoologists, yet only a minuscule fraction of one percent of [the craziest] zoologists deal directly with problems associated with scientific names of animals (Bock 1994).
Introduction
The accurate use of scientific names in zoology is necessary to minimize nomenclatural instability and allow the maximal retrieval of scientific information from the ever increasing body of literature. Akin to other scientific names such as species- and genus-group names, family-group names are extremely important in the exchange of information about the world we live in. Rules about how to treat family-group names, as determined by the International Commission on Zoological Nomenclature [henceforth the Commission], have been an integral part of zoological nomenclature since the early 1900s (for a historical review of family-group names refer to Bock 1994).
The first family-group names based on the stem of their type genus appeared in zoological literature in the early 19th Century (see Sabrosky 1999). The first authors inconsistently used a variety of endings (–ides, –ites, –ida, –i, –ide, etc.) for suprageneric divisions at various levels. The ending –idae was apparently first suggested by Kirby (1813) as a characteristic ending for names at the rank of family. Although many ranks have been used in the past for suprageneric names, only the following suffixes are currently regulated by the International Code of Zoological Nomenclature [henceforth the Code] (ICZN 1999a): superfamily (–oidea), family (–idae), subfamily (–inae), tribe (–ini) and subtribe (–ina).
The last twenty five years has seen an increase in the number of studies on family-group names of entire groups in zoology (e.g., Štys and Jansson 1988 for Hemiptera: Nepomorpha; Bock 1994 for Aves; McKenna and Bell 1997 for Mammalia; Ferraris and de Pinna 1999 for Pisces: Siluriformes; Sabrosky 1999 for Diptera; Engel and Krishna 2004 for Isoptera; Speidel and Naumann 2004 for Lepidoptera: Noctuoidea; Bouchet and Rocroi 2005 for Gastropoda; Engel 2005 for Hymenoptera: Apoidea; Engel and Haas 2007 for Dermaptera; Bouchet and Rocroi 2010 for Bivalvia). The contents (i.e., whether they include fossils or not) and presentation of the data (i.e., in a classification scheme, in chronological order or in alphabetic order) in those works vary greatly. The most extensive treatments of animal family-group names to date are those of Diptera (Sabrosky 1999) and Gastropoda (Bouchet and Rocroi 2005) which covered names based on approximately 2000 and 2400 distinct type genera respectively. Some of the reasons given by authors of these works for providing comprehensive lists of family-group names include avoiding the unnecessary proposal of new names, facilitating decisions on priority and promoting long-term stability of the classification. We believe that such catalogues are especially important as the number of studies dealing with the higher relationships of major clades will continue to increase as new algorithms and data sets (e.g., molecular data) become available.
Given the importance of family-group names in the scientific literature, it is rather surprising that still many authors do not cite the author or year of publication of those names correctly, or at all. According to Recommendation 51A of the Code (ICZN 1999a) “The original author and date of a name should be cited at least once in each work dealing with the taxon denoted by that name.” In addition to satisfying Recommendation 51A of the Code, citation of authorities of scientific names can also have a positive impact on modern world taxonomists (Werner 2006; Agnarson and Kuntner 2007). Adding to further confusion and errors in the literature is the fact that the Principle of Coordination (Art. 36.1) is still overlooked by some authors.
Coleoptera are currently the most species-rich group of organisms on this planet with approximately 360 000 described species (Bouchard et al. 2009). The great morphological diversity of beetles has led to the proliferation of suprageneric taxa at various ranks. Latreille (1797) was apparently the first to introduce the concept of family-level taxa (see Bock 1994: 244) but it is only a few years later (Latreille 1802) that he proposed available names for these groupings, including several in the order Coleoptera. More recent evidence suggests that available family-group names in some groups of animals other than Coleoptera (e.g., Chordata: Sauropsida) appeared in the late 18th Century (see Dubois and Bour 2010), even earlier than in Latreille (1802).
The nomenclature of family-group names in Coleoptera did not receive much attention until the treatment of Geadephaga names by Madge (1989). Similar lists, based on rules from two different editions of the Code (ICZN 1985g, 1999a), have now been published in Dytiscidae (Nilsson et al. 1989), Staphyliniformia (Newton and Thayer 1992), Cucujoidea (Pakaluk et al. 1994), Curculionoidea (Alonso-Zarazaga and Lyal 1999), Buprestoidea (Bellamy 2003, 2008a-d, 2009), Tenebrionidae (Bouchard et al. 2005), Scarabaeoidea (Smith 2006) and Cerambycidae (Bousquet et al. 2009). Another important contribution was that of Lawrence and Newton (1995) which included a review of the nomenclature of all beetle family-group names for the rank of subfamily and above.
The vast body of scientific literature dealing with beetles has been a deterrent to producing a complete review of all Coleoptera family-group names in the past. This publication was only made possible by the collaboration of several coleopterists.
The specific objectives of this study are to: 1) establish the first comprehensive list of all Coleoptera family-group names with information on type genus, author(s), year of publication and complete bibliographical references; 2) assess the availability and validity of each name using rules laid out in the most recent Code of Zoological Nomenclature (ICZN 1999a), 3) summarize Priority and Homonymy problems with currently used names and, 4) propose or implement solutions to these problems in order to promote stability. We include family-group names that were published on or before December 31, 2010.
This publication should be seen as the starting point of what could eventually serve as the basis for a submission of Part of a List of Available names in Zoology (Art. 79). With this goal in mind, we ask all coleopterists to either send us, or to publish, corrections or differences of opinions in the months and years to come. It is our intention to update our list of family-group names as new data are published and to provide a second edition of this work in approximately five years.
Methods
Criteria of availability
Decisions about the availability and validity of each Coleoptera family-group name in our catalogue were made following the process outline in Figure 1. First we established whether the name was available or unavailable based on the criteria of availability summarized below. If a name first proposed on a particular type genus was determined to be unavailable then searches were conducted to establish if a family-group name based on the same type genus was made available subsequently. If so then this available name was entered in our catalogue. From the pool of available names, we removed those that are permanently invalid (Art. 39). All the remaining available names were then separated into those that are valid based on the classification used here (at any rank from subtribe to superfamily) and those that are invalid (i.e., synonyms).
Figure 1. Overview of the process used to determine the availability and validity of family-group names in Coleoptera (modified from Bouchet and Rocroi 2005). The number of names for each category is given in parentheses.
In order to be available, a family-group name proposed before 1931 must to be a scientific name (i.e., in latinized form) in the nominative plural based on the stem of an available genus name then used as valid in the new suprageneric taxon (Art. 11.7).
In addition to the criteria of availability mentioned above, new family-group names proposed between 1931 and 1999 had to be described in words, or be associated with a bibliographic reference to such a description, in order to be considered available (Arts 13, 15). In works published after 1930 which contained more than one use of a new family-group name, we selected a page where a description was clearly associated with the new taxon for the catalogue. It should be noted that replacement names proposed in that time period are available without description (Art. 13.1.3). Additionally, “a family-group name first published after 1930 and before 1961 which does not satisfy the provisions of Article 13.1 is available from its original publication only if it was used as valid before 2000, and also was not rejected by an author who, after 1960 and before 2000, expressly applied Article 13 of the then current editions of the Code” (Art. 13.2.1). Names proposed between 1931 and 1960 can also be considered available if the description of the new family-group name and a single new genus-group name is combined (Art. 13.5). Finally, family-group names proposed since 2000 have to be explicitly indicated as new and the name of the type genus has to be clearly cited in order to be available (Art. 16). Based on our interpretation of Article 16.2, any new family-group name proposed after 1999 in a paper in which the name of the type genus is clearly cited in connection with it (although maybe not explicitly with a formula such as “Type genus = Aus Doe, 2010”) is available.
One of the most difficult tasks while working on this review was to decide on the most consistent and objective way to apply Article 11.7.2, which deals with the availability of names that were originally proposed in a vernacular form. Vernacular names are generally not treated as scientific names in zoological nomenclature (see Recommendation 11A). However, the International Commission on Zoological Nomenclature has made a single, but rather restricted, exception that applies to family-group names. According to the Code, a vernacular name “…is available with its original author and date only if it has been latinized by later authors and has been generally accepted as valid by authors interested in the group concerned and as dating from that first publication in vernacular form” (Art. 11.7.2).
We have interpreted names proposed in the following languages to be vernacular: all German names with the suffix “–en”, all Spanish names with the suffix “–os” or “–as” and all French names with the suffix “–iens”. The most important issue was to determine the correct status of names originally proposed with the suffix “–es.” We have used the principle that all family-group names proposed by non-French writers with the suffix “–es” were in fact latinized names with an ending that differs from those regulated by the Commission (these are mostly older names proposed before rules of zoological nomenclature became well-established). For each name with the suffix “–es” first proposed by French workers (e.g., Latreille, Lacordaire, Lameere) we went through the entire work containing those names and established if these authors consistently used either vernacular or latinized names in their work. French vernacular names often have accents in them (“é” or “è”) while latinized names do not. We have found that determining whether an author used vernacular or latinized names in a particular work was fairly straightforward (Bousquet et al. 2009). Latreille proposed new family-group names with the suffix “–es” in several of his works. We noticed that he most often used both the vernacular (listed first, with accents when required by French language) and latinized (listed second, always in italics) forms of each name together in the same heading. We have listed the latinized form of the name in each of those cases.
As in Bousquet et al. (2009), we have interpreted the requirements listed in Art. 11.7.2. as three separate conditions to be met. For each vernacular name we determined if it had 1) been subsequently latinized, 2) been generally accepted as valid by authors interested in the group and 3) been attributed to the author and date of original publication. Failure to fulfil any of the requirements resulted in the treatment of that name as unavailable. In those cases, we provide a comment explaining why this taxon was treated as unavailable (e.g., “original vernacular name unavailable (Art. 11.7.2): not subsequently latinized”). Many recent authors have treated family-group names first proposed in vernacular form as available if they were latinized by later authors but we do not believe that this practice is in line with the requirements of the Code. We have accepted as available all vernacular names published before 1900 that have subsequently been used in latinized form, while being used as valid, and credited to the publication in their vernacular form. For every vernacular name that meets the requirement of availability, we have added a comment in the format of the following example “original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 240, as Meracanthini), generally accepted as in Gebien (1911: 567, as Meracanthinae)”.
Formation and treatment of family-group names
Article 29.2 includes a list of suffixes to be used for groups at the superfamily (-oidea), family (-idae), subfamily (-inae), tribe (-ini) and subtribe (-ina) names. The suffix of other categories is not regulated by the Code. The rank of supertribe (-itae) was used here for a small number of groups in which recent changes in classification required an additional rank between subfamily and tribe (e.g., Staphylinoidea).
The correct spelling of family-group names depends on the stem of its type genus (Art. 29.3). As pointed out by Newton and Thayer (1992) the family-group name stems of most generic names that can be regarded as Latin nouns are determined easily by dropping the following nominative case ending –us, –um, –es and –a. Examples of such straightforward case in Coleoptera include Carabidae (type genus Carabus), Adeliini (type genus Adelium), Trechodina (type genus Trechodes) and Anthiini (type genus Anthia). A summary of other commonly encountered generic endings, along with their corresponding stems, is given in Table 1. Note that generic names ending with -gaster (Greek for stomach) can have either -gaster- or -gastr- as their correct stem and therefore we have accepted the stem formation of the first author of the family-group name based on such genera as correct. Stems were reviewed for all type genera included in our catalogue.
Table 1. List of common Coleoptera generic suffixes with their associated family-group name endings (based partly on Bouchet and Rocroi 2005, with additions by YB, AFN, MAAZ and M. K. Thayer).
Generic ending
|
Meaning
|
Derived family-group name ending
|
Type genus example
|
Family-group name example
|
---|
-apion
|
pear (Greek) |
-api- |
Aspidapion
|
Aspidapiina
|
-arthron
|
joint (Greek) |
-arthr- |
Decarthron
|
Decarthrina
|
-aspis
|
shield (Greek) |
-aspid- |
Anaspis
|
Anaspidinae
|
-baris
|
flat-bottomed boat (Greek) |
-barid- |
Baris
|
Baridini
|
-chlamys
|
mantle (Greek) |
-clamyd- |
Spodochlamys
|
Spodochlamydini
|
-celis
|
spot (Greek) |
-celid- |
Xiphoscelis
|
Xiphoscelidini
|
-ceras
|
horn (Greek) |
-cerat- |
Megaceras
|
Megaceratini
|
-cnema
|
shin or tibia (Greek) |
-cnem- |
Pachycnema
|
Pachycnemina
|
-cupes
|
dainty (Latin) |
-cuped- |
Cupes
|
Cupedidae
|
-dacne
|
bite (Greek) |
-dacn- |
Dacne
|
Dacnini
|
-deres
|
neck, throat (Greek) |
-der- |
Aglycyderes
|
Aglycyderini
|
-derma
|
skin (Greek) |
-dermat- |
Cryptoderma
|
Cryptodermatinae
|
-dytes
|
diver (Greek) |
-dyt- |
Aspidytes
|
Aspidytidae
|
-genys
|
jaw (Greek) |
-geny- |
Chaetogenys
|
Chaetogenyini
|
-hospes
|
guest (Latin) |
-hospit- |
Termitohospes
|
Termitohospitini
|
-ides
|
similar to, derived from (Greek) |
-id- |
Anaides
|
Anaidinae
|
-ifer
|
carrier of (Latin) |
-ifer- |
Undulifer
|
Unduliferinae
|
-iger
|
carrier of (Latin) |
-iger- |
Apoderiger
|
Apoderigerina
|
-ites
|
like (Latin, Greek) |
-it- |
Aegialites
|
Aegialitinae
|
-loma
|
edge or fringe (Greek) |
-lomat- |
Discoloma
|
Discolomatinae
|
-macer
|
thin (Latin) |
-macr- |
Rhynchitomacer
|
Rhynchitomacrini
|
-mycter
|
nose (Greek) |
-mycter- |
Eurymycter
|
Eurymycterini
|
-odes
|
similar to (Greek) |
-od- |
Agyrtodes
|
Agyrtodini
|
-odon
|
tooth (Greek) |
-odont- |
Pentodon
|
Pentodontini
|
-oides
|
like (Greek) |
-oid- |
Acmaeoderoides
|
Acmaeoderoidina
|
-omma
|
eye (Greek) |
-ommat- |
Omma
|
Ommatidae
|
-onyx
|
yellow gem stone (Greek) |
-onych- |
Trichonyx
|
Trichonychini
|
-ops
|
eye (Greek) |
-op- |
Achaenops
|
Achaenopina
|
-opsis
|
appearance (Greek) |
-opse- |
Brachyceropsis
|
Brachyceropseini
|
-otes
|
quality, nature (Greek) |
-ot- |
Agriotes
|
Agriotini
|
-pholis
|
horny scale (Greek) |
-pholid- |
Trachypholis
|
Trachypholidini
|
-pteryx
|
wing (Greek) |
-pteryg- |
Trichopteryx
|
Trichopterygini
|
-pus
|
foot (Greek) |
-pod- |
Baripus
|
Baripodina
|
-rhinus
|
snout (Greek) |
-rhin- |
Platyrhinus
|
Platyrhinini
|
-rhynchus
|
snout (Greek) |
-rhynch- |
Doydirhynchus
|
Doydirhynchini
|
-rhipis
|
fan (Greek) |
-rhipid- |
Xenorhipis
|
Xenorhipidini
|
-soma
|
body (Greek) |
-somat- |
Platysoma
|
Platysomatini
|
-stoma
|
mouth (Greek) |
-stomat- |
Stenostoma
|
Stenostomatini
|
-teles
|
perfect (Greek) |
-tel- |
Abroteles
|
Abrotelina
|
-termes
|
wood-worm (Latin) |
-termit- |
Philotermes
|
Philotermitini
|
-thorax
|
chest (Greek) |
-thorac- |
Mecyclothorax
|
Mecyclothoracini
|
-trox
|
gnawer (Greek) |
-trog- |
Trox
|
Trogidae
|
-trupes
|
borer (Greek) |
-trup- |
Ceratotrupes
|
Ceratotrupini
|
-typus
|
shape (Greek) |
-typ- |
Amarotypus
|
Amarotypini
|
It should be noted that if a family-group name was not formed in accordance with Art. 29.3 but its original spelling is in prevailing usage then the current spelling is to be maintained (Art. 29.5). We have conserved the spelling of several family-group names currently used as valid however we did not do so for names that are listed as synonyms. For names based on incorrect stems proposed after 1999, we have considered that prevailing usage cannot be used to conserve the original spellings because too few references using these names could be found. We have therefore corrected the stems of such names unless the name of the type genus was an arbitrary combination of letters (Art. 29.4).
As stated in Art. 35.4.1 “A family-group name based upon an unjustified emendation … or an incorrect spelling of the name of the type genus must be corrected, unless it is preserved under Article 29.5 or unless the spelling of the genus-group name used to form the family-group name is preserved under Articles 33.2.3.1 or 33.3.1.” When an unjustified emendation or an incorrect subsequent spelling of the type genus is in prevailing usage and is attributed to the original author and date (Art. 33.2.3.1; 33.3.1), the correct spelling of the type genus is that in current usage.
In the glossary of the Code, a name in prevailing usage is defined as a “name which is adopted by at least a substantial majority of the most recent authors concerned with the relevant taxon, irrespective of how long ago their work was published”. The unfortunate subjectivity in this definition, as pointed out by Ferraris (2000), left us with no choice but determine prevailing usage in an ad hoc fashion throughout.
Principle of Coordination
The Principle of Coordination (Art. 36.1) is, unfortunately, still overlooked by some authors. In some instances authors who propose new ranks for previously established suprageneric names are sometimes treated as the author of those names when in fact only a change of rank was presented. Based on the Code “A name established at any rank in the family-group is deemed to have been simultaneously established for nominal taxa at all ranks in the family-group; all these taxa have the same type genus, and their names are formed from the stem of the name of the type genus [Art. 29.3] with appropriate change of suffix [Art. 34.1]. The name has the same authorship and date at every rank.”
Principle of Priority
As for species- and genus-group names, the oldest available name for a family-group taxon should be considered as valid (Art. 23). However four important exceptions need further discussion. Firstly, when a little-known family-group name was discovered to be older than a name currently used as valid for a particular taxon, we used the Reversal of Precedence to conserve usage of the younger name if the conditions of Art. 23.9.2 could be fulfilled. Younger names conserved using Reversal of Precedence are listed in Appendix 1. In some instances we could not fulfill all conditions of Art. 23.9.2 although we considered that using the newly discovered older name as valid would threaten stability or cause confusion. In those cases we maintained usage of the younger name as valid and either submitted an application to the Commission to conserve the younger name or made a recommendation that such submission should be submitted in the near future (Art. 23.9.3).
According to Art. 35.5 “If after 1999 a name in use for a family-group taxon (e.g., for a subfamily) is found to be older than a name in prevailing usage for a taxon at higher rank in the same family-group taxon (e.g., for the family within which the older name is the name of a subfamily) the older name is not to displace the younger name.” Indeed, we encountered a small number of cases in which the replacement of a name at the higher rank (e.g., Lymexyloidea Fleming, 1821) by the discovery of an older name for a taxon at a lower rank (e.g., Hylecoetoidea Germar, 1818) would not have served the stability of well-established names. In such cases usage of the younger name at the higher rank was conserved.
In cases where a family-group name was replaced before 1961 because of the synonymy of its type genus (e.g., Lepiceridae Hinton, 1936 instead of Cyathoceridae Sharp, 1882), the substitute name is to be maintained if it is in prevailing usage (Art. 40.2). In such cases, the valid family-group name retains its own author but takes the priority of the replaced name. Based on Recommendation 40A, we have cited those names with their original author and date, followed by the date of its priority enclosed in parentheses, as determined by Art. 40.2.1 (e.g., Lepiceridae Hinton, 1936 (1882)).
Lastly, if the stability and continuity of the meaning of a family-group name is threatened by the discovery that the type genus was originally misidentified, or that the type genus was based on a misidentified type species, or that an older type species of the type genus had been overlooked, then the case is to be referred to the Commission for a ruling (Art. 65.2.1). In cases where the oldest available name for a family-group taxon is based on misidentified type genus, or an altered concept of the type genus (e.g., see Scydmaenini Reitter, 1882), we have preferred to consider this name as invalid and to use instead the family-group name which is in prevailing usage (e.g., Cyrtoscydmini L. W. Schaufuss, 1889) until an application is submitted and a ruling is rendered by the Commission.
A list of problem cases based on the Principle of Priority, with comments on implementation of solutions or necessary actions to be taken in the future, is given in Appendix 2.
Principle of Homonymy
Based on the Principle of Homonymy “when two or more names are homonyms, only the senior, as determined by the Principle of Priority…, may be used as the valid name…” (Art. 55.2). Here we report several instances in which family-group names in Coleoptera are identical to other family-group names in zoological nomenclature. These names are either based on identical type genera or on type genera that are similar but not identical. In the first instance the family-group name based on the preoccupied type genus is permanently invalid (Art. 39) but available.
When family-group names are homonyms because their type genera are similar but not identical, the case must be referred to the Commission for a ruling to remove homonymy (Art. 55.3.1). Such is the case with Adeliini Kirby, 1825 (type genus Adelium Kirby, 1819) and the hymenopteran name Adeliini Viereck, 1918 (type genus Adelius Haliday, 1833) which are both correctly formed from the stem of their type genus. More than thirty such cases were encountered during our research on Coleoptera family-group names.
Junior homonyms conserved using Reversal of Precedence are listed in Appendix 1. A list of problem cases based on the Principle of Homonymy, with comments on implementation of solutions or necessary actions to be taken in the future, is given in Appendix 3.
Submissions to the Commission
As mentioned above, we have encountered several cases which require an application to the Commission because of problems with priority and/or homonymy. Some of these cases have been submitted recently (Engel and Bouchard 2009, Bousquet et al. 2010, Bousquet and Bouchard 2010) and others will be submitted in the near future (see summary in Appendices 2 and 3). However, it is not our intention to submit applications for all cases. Since we intend to submit this work as Part of the List of Available Names in Zoology in approximately five years, we hope the coleopterist community will take this opportunity to submit applications to the Commission in order to resolve some of the remaining problems outlined here.
Bibliographic notes
Von Hayek (1973: 282, 290) and Stibick (1979: 157) reported that the important works on Elateridae classification by Candèze (1857) and Lacordaire (1857) were published in May and June of 1857 respectively. Since then, when the same new name appeared in both of these works, Candèze’s names have been given priority over Lacordaire’s. We have discovered that Lacordaire’s work was in fact published before 25 May 1857 (as recorded by the Académie des Sciences de France) which would make the names in his work the oldest based on strict adherence to the Code (Art. 21.3). In order to avoid unnecessary changes in authorship in the future and maintain prevailing usage, we have opted to continue attributing elaterid family-group names to Candèze, 1857.
Authorship of the three volumes on Coleoptera of the series Encyclopédie d’histoire naturelle is debated. The title pages on the volumes list Jean Charles Chenu as the author of the Encyclopédie d’histoire naturelle with the assistance of Eugène Desmarest for the Coleoptera section. However, the Société Entomologique de France recorded the livraisons received for 1851 in their Bulletin (p. cxxv) as “Encyclopédie d’histoire naturelle, ou traité complet de cette science, sous la direction de M. le docteur Chenu. Coléoptères, par M.E. Desmarest.” This was likely the title on the wrapper and clearly suggests that Desmarest was responsible alone for the three volume series on Coleoptera. This is also substained by Desmarest himself who in the Bulletin de la Société Entomologique de France for 1860 (p. lxiii) speaks of the three volumes series as being his work, published under the direction of Chenu. Based on the above fact, we credit the three-volume series to Desmarest alone (Desmarest 1851, 1857, 1860).
The articles presented by Mulsant (and Rey) in the “Opuscules Entomologiques” are considered here to be reprints (as a compilation) of original publications in various Annales, mostly printed at Lyon, for the following reasons: In the dedication to the first volume Mulsant wrote “ces Opuscules, publiés déjà ça et là dans nos Recueils académiques”. The dates of the dedications in the Opuscules are always later than the dates on which they were presented to the various Sociétés, frequently in the following year. Although published at Paris, the Opuscules (see verso of title pages) were printed at Lyon by F. Dumoulin and others, the same printers which published the Mémoires de l’Académie... des Sciences, Belles-Lettres et Arts de Lyon, etc. Rey himself, in later publications such as the continuation of the “Histoire Naturelle des Coléoptères de France” (e.g., Annales de la Société Linnéenne de Lyon (N. S.) 32[1885]: 1-186 + [4], pls. 1-2 [see p. 106]) cited the Annales de la Société Linnéenne de Lyon article before the Opuscules version. E. A. Fitch (1881: 46-47), in his obituary of Mulsant, indicated that both the Histoire Naturelle des Coléoptères de France and the Opuscules Entomologiques «appeared originally... in the Annals» and «later republished in separate form at Paris». These concepts were reiterated by Tottenham (1949: 352). Marseul (1882: 20) wrote, “Tous les ouvrages de Mulsant, sauf quelques-uns, ont été publiés dans les Annales des trois sociétés... et l’Académie des Sciences. Tous ont été reproduits séparément soit dans l’Histoire Naturelle des Coléoptères et des Punaises de France, soit dans les Opuscules.»
However, the first fascicles of the Histoire Naturelle series up to the “Pectinipèdes” were not reprinted in journals. It seems that most of the succeeding fascicles were submitted to the various Sociétés, were printed by Dumoulin, Barret, Pinier, etc. at Lyon, the dedications were then added and they were sent to Paris for separate publication with the same typesetting, and with changes only to the pagination and some titles. Since the issuing of the journal volumes was frequently delayed by the compilation of many papers (usually in the following year), the separate edition often appeared first, as shown by records of other journals and, in particular, the Bibliographie de la France. Therefore we have given date priority to the Histoire Naturelle versions in most instances, only those page numbers are cited with the scientific names in the catalogue unless evidence was found to the contrary, and the alternate versions are cited only in the bibliography, following each reference.
Format of the catalogue and conventions used
This paper is organized into two main parts: 1) a synoptic classification containing only the names which we consider as valid herein and 2) a catalogue of all family-group names organized in the same order as in the synoptic classification. It should be noted that our publication is first and foremost a nomenclatural treatment of family-group names. We want to emphasize that the classification used here is not based on newly generated phylogenetic data. Furthermore, we do not necessarily endorse all parts of the classification presented, particularly those which were not based on phylogenetic approaches. For the most part, we follow currently accepted concepts in recent taxon-specific catalogues as well as comprehensive syntheses such as the Handbook of Zoology (Beutel and Leschen 2005, Leschen et al. 2010), American Beetles (Arnett and Thomas 2000; Arnett et al. 2002) and the Catalogue of Palaearctic Coleoptera (Löbl and Smetana 2003, 2004, 2006, 2007, 2008, 2010). Although we recognize that the classification we use will likely become outdated in the near future, we believe that this is the best way to present the assembled data.
Only family-group names that are based on genera are included in our catalogue. For each family-group name the original spelling, author, year of publication, page number, correct stem and type genus are given. Complete data and comments regarding a particular family-group name are presented with the lowest-rank name when the same stem is used in more than one rank, since the same criteria apply in accordance with the Principle of Coordination (Art. 36.1). The correct stem to be used, which is given in square brackets (e.g., [stem: Uralocole-]), is especially important for synonyms, since the status of a name may be changed from synonym to valid in the future. Subsequent alternative spellings of family-group names (i.e., simple changes in suffix depending on change in rank) are not given here. Both available and unavailable names are listed together in the catalogue in order to enhance information retrieval. Unavailable names are preceded by an asterisk “*” and are identified as such in the “Comments” section along with the reason for this status. As is common practice in publications of the International Commission on Zoological Nomenclature, we use small capital letters for family-group names. Names in the catalogue are organized in a “phylogenetic” framework, as accepted in recent publications on the group, down to subfamily level. Valid tribes and subtribes are listed in alphabetical order within each subfamily. Synonyms are listed in chronological order under each valid taxon.
Known works of Johannes Gistel (who also published under the name Johannes Gistl and as G. Tilesius; see Evenhuis 1997a) were included in this paper. Although Gistel’s works were largely ignored by contemporaries (e.g., Gemminger and Harold 1868a) and continue to be ignored by some workers (e.g., Cate 2007) they are nevertheless broadly available and represent a significant contribution to the nomenclature of family-group and genus-group names in Coleoptera. In order to promote stability, some names proposed for the first time by Gistel (but ignored until now) that threaten names in current usage are treated here as nomina oblita (when conditions of Art. 23.9 could be met) or are subject to appeals to the Commission. According to the glossary of the Code, a nomen oblitum is a “Latin term applied after 1 January 2000 to a name, unused since 1899, which as a result of an action taken under Article 23.9.2 does not take precedence over a younger synonym or homonym in prevailing usage. The term nomen oblitum was also applied to a disused senior synonym rejected between 6 November 1961 and 1 January 1973 under Art. 23b of the Code editions then in force (see Art. 23.12.2). Nomina oblita are available names; see Articles 23.9 and 23.12 for conditions controlling their use as valid names.”
Because of the importance of the Principle of Priority, we have tried to find the most accurate date of publication (given in square brackets in the References section) for works cited in the manuscript. The date of publication was determined either from the original publication itself, from the date of “stamps” when received in libraries of natural history institutions or from secondary literature sources. A list of natural history institutions and secondary literature sources is given at the beginning of the References section. Data on dates of publication were provided by AFN, PB, AED, MAAZ, CHCL, ABTS and YB. References for type genera are not included here to conserve space and because they are for the most part available in other recent publications (e.g., Löbl and Smetana 2003, 2004, 2006, 2007, 2008, 2010).
Names on the Official List of Family-Group Names in Zoology and type genera on the Official List of Generic Names in Zoology are included in our catalogue and are summarized in Appendix 4 and 5 respectively. Cases involving family-group names and/or their type genera that are awaiting a ruling by the Commission are summarized in Appendix 6.
Results
Number of family-group names
A total of 4887 family-group names are included in our catalogue (see Figure 1). Of the names recorded, 4492 are available, of which 183 are permanently invalid because they are based on a preoccupied type genus or a genus which has been suppressed by the Commission. The majority of names were proposed in the suborder Polyphaga (4314) followed by Adephaga (531), Myxophaga (16), Archostemata (17) and Protocoleoptera (9). Within Adephaga the family Carabidae contains the highest number of names by far (441) followed by Dytiscidae (47). The five superfamilies with the highest number of names in Polyphaga are the Curculionoidea (862), Chrysomeloidea (794), Staphylinoidea (594), Tenebrionoidea (579) and Scarabaeoidea (436). Overall, the five families with the highest number of family-group names proposed are Curculionidae (555), Staphylinidae (493), Cerambycidae (468), Carabidae (441) and Tenebrionidae (323).
The number of Coleoptera family-group names that appeared in the 19th and 20th Centuries are almost identical (2331 and 2556 respectively) while the last decade saw an increase of 5% (246 names) in the total number of names in the literature (Figure 2). A large number of names proposed in the middle of the 19th Century are unavailable. Those names were generally proposed in vernacular form and were not made available subsequently (Art. 11.7.2). Unavailable names proposed after 1930 generally lacked a description or bibliographic reference to such a description (Arts 13, 15). Several names proposed in the last decade were either not explicitly introduced as new taxa or did not include necessary information about the type genus and are therefore unavailable (Art. 16).
Figure 2. Number of family-group names proposed in Coleoptera by decade. White bars = available names. Black bars = unavailable names.
Significant contributions
A summary of the most significant contributions, in terms of the total number of names proposed by author, is shown in Table 2. Eighteen authors proposed 50 or more family-group names in Coleoptera. Lacordaire was the most prolific with 353 new names. Although the majority of Lacordaire’s names were proposed in vernacular form, a large proportion of them (90%) were subsequently made available and many are used as valid today.
Table 2. Summary of number of family-group names in Coleoptera by author. Only authors that have proposed fifty or more names are included. Authors listed in decreasing order by total number of names proposed. For each author we give the total number of name proposed, the percentage of those names that are available, the range of years in which those names were proposed and the number of families in which they proposed new family-group names in.
Rank |
Author |
Country of origin |
Total family-group names |
Percent available |
Publication (year range) |
Number of families |
---|
1 |
Lacordaire, Jean Théodore |
France |
353 |
90 |
1848–1872 |
34 |
2 |
Gistel, Johannes von Nepomuk Franz Xaver |
Germany |
135 |
99 |
1848–1856 |
42 |
3 |
LeConte, John Lawrence |
USA |
131 |
100 |
1847–1883 |
45 |
4 |
Thomson, James |
USA |
124 |
99 |
1857–1877 |
6 |
5 |
Mulsant, Étienne |
France |
122 |
76 |
1839–1880 |
25 |
6 |
Chapuis, Félicien |
France |
120 |
72 |
1869–1876 |
7 |
7 |
Jeannel, René |
France |
109 |
99 |
1910–1967 |
6 |
8 |
Reitter, Edmund |
Germany |
87 |
100 |
1875–1926 |
24 |
9 |
Legalov, Andrei Aleksandrovich |
Russia |
73 |
100 |
2001–2009 |
9 |
10 |
Voss, Eduard |
Germany |
66 |
97 |
1922–1972 |
6 |
11 |
Burmeister, Hermann Carl Conrad |
Germany |
66 |
98 |
1840–1873 |
2 |
12 |
Casey, Thomas Lincoln |
USA |
64 |
100 |
1884–1922 |
14 |
13 |
Horn, George Henry |
USA |
63 |
100 |
1867–1893 |
20 |
14 |
Latreille, Pierre André |
France |
62 |
98 |
1802–1834 |
30 |
15 |
Rey, Claudius |
France |
57 |
53 |
1853–1886 |
12 |
16 |
Thomson, Carl Gustaf |
Sweden |
55 |
100 |
1857–1867 |
27 |
17 |
Blanchard, Charles Émile |
France |
55 |
71 |
1845–1853 |
20 |
18 |
Laporte, François Louis Nompar de Caumont (Comte Castelnau) |
France |
51 |
100 |
1834–1840 |
21 |
The authors with the lowest percentage of available family-group names (Mulsant, Chapuis, Rey and Blanchard) originally proposed their names in vernacular form and a significant proportion of those names were not made available subsequently. All names proposed by LeConte, Reitter, Casey, Horn, Legalov, C. G. Thomson and Laporte are considered available.
Thirteen of the authors in Table 2 were active exclusively during the 19th Century while two authors, Reitter and Casey, published their works containing new family-group names in both the 19th and 20th Centuries. Two authors published family-group names exclusively during the 20th Century (Jeannel and Voss). Legalov is our only contemporary colleague in this list; he proposed more than 50 new family-group names in the last decade. It should be noted that the recent proliferation of new scientific names proposed by Legalov is treated by some authors as “extreme splitting” (Riedel 2006) and “based on numerous spurious characters of doubtful phylogenetic value” (Oberprieler et al. 2007).
The broad taxonomic interests and expertise of Lacordaire, Gistel, LeConte, Mulsant, Latreille and C. G. Thomson are exemplified by the fact that they introduced new family-group names in 25 or more families. This was done either in catalogues (e.g., Gistel, C. G. Thomson) and/or in large scale taxonomic treatments (e.g., Lacordaire, LeConte, Latreille, Mulsant). On the other hand, the taxonomic expertise of J. Thomson, Chapuis, Jeannel, Voss, Burmeister and Legalov is much more targeted towards a small number of families (fewer than 10).
New nomenclatural acts
The following synonymies are recorded here for the first time: Agronomina Gistel, 1848 syn. nov. of Amarina Zimmermann, 1832 (Carabidae), Hylepnigalioini Gistel, 1856 syn. nov. of Melandryini Leach, 1815 (Melandryidae), Polycystophoridae Gistel, 1856 syn. nov. of Malachiinae Fleming, 1821 (Melyridae), Sclerasteinae Gistel, 1856 syn. nov. of Ptilininae Shuckard, 1839 (Ptinidae), Phloeonomini Ádám, 2001 syn. nov. of Omaliini MacLeay, 1825 (Staphylinidae), Sepedophilini Ádám, 2001 syn. nov. of Tachyporini MacLeay, 1825 (Staphylinidae), Phibalini Gistel, 1856 syn. nov. of Cteniopodini Solier, 1835 (Tenebrionidae); Agronoma Gistel 1848 (type species Carabus familiaris Duftschmid, 1812, designated herein) syn. nov. of Amara Bonelli, 1810 (Carabidae), Hylepnigalio Gistel, 1856 (type species Chrysomela caraboides Linnaeus, 1760, by monotypy) syn. nov. of Melandrya Fabricius, 1801 (Melandryidae), Polycystophorus Gistel, 1856 (type species Cantharis aeneus Linnaeus, 1758, designated herein) syn. nov. of Malachius Fabricius, 1775 (Melyridae), Sclerastes Gistel, 1856 (type species Ptilinus costatus Gyllenhal, 1827, designated herein) syn. nov. of Ptilinus Geoffroy, 1762 (Ptinidae), Paniscus Gistel, 1848 (type species Scarabaeus fasciatus Linnaeus, 1758, designated herein) syn. nov. of Trichius Fabricius, 1775 (Scarabaeidae), Phibalus Gistel, 1856 (type species Chrysomela pubescens Linnaeus, 1758, by monotypy) syn. nov. of Omophlus Dejean, 1834 (Tenebrionidae). The following replacement name is used here for the first time: Gompeliina Bouchard, 2011 nom. nov. for Olotelina Báguena Corella, 1948 (Aderidae).
Reversal of Precedence (Article 23.9) is used to conserve usage of the following names (family-group names followed by genus-group names): Perigonini Horn, 1881 nom. protectum over Trechicini Bates, 1873 nom. oblitum (Carabidae), Anisodactylina Lacordaire, 1854 nom. protectum over Eurytrichina LeConte, 1848 nom. oblitum (Carabidae), Smicronychini Seidlitz, 1891 nom. protectum over Desmorini LeConte, 1876 nom. oblitum (Curculionidae), Bagoinae Thomson, 1859 nom. protectum over Lyprinae Gistel 1848 nom. oblitum (Curculionidae), Aterpina Lacordaire, 1863 nom. protectum over Heliomenina Gistel, 1848 nom. oblitum (Curculionidae), Naupactini Gistel, 1848 nom. protectum over Iphiini Schönherr, 1823 nom. oblitum (Curculionidae), Cleonini Schönherr, 1826 nom. protectum over Geomorini Schönherr, 1823 nom. oblitum (Curculionidae), Magdalidini Pascoe, 1870 nom. protectum over Scardamyctini Gistel, 1848 nom. oblitum (Curculionidae), Agrypninae/-ini Candèze, 1857 nom. protecta over Adelocerinae/-ini Gistel, 1848 nom. oblita and Pangaurinae/-ini Gistel, 1856 nom. oblita (Elateridae), Prosternini Gistel, 1856 nom. protectum over Diacanthini Gistel, 1848 nom. oblitum (Elateridae), Calopodinae Costa, 1852 nom. protectum over Sparedrinae Gistel, 1848 nom. oblitum (Oedemeridae), Adesmiini Lacordaire, 1859 nom. protectum over Macropodini Agassiz, 1846 nom. oblitum (Tenebrionidae), Bolitophagini Kirby, 1837 nom. protectum over Eledonini Billberg, 1820 nom. oblitum (Tenebrionidae), Throscidae Laporte, 1840 nom. protectum over Stereolidae Rafinesque, 1815 nom. oblitum (Throscidae) and Lophocaterini Crowson, 1964 over Lycoptini Casey, 1890 nom. oblitum (Trogossitidae); Monotoma Herbst, 1799 nom. protectum over Monotoma Panzer, 1792 nom. oblitum (Monotomidae); Pediacus Shuckard, 1839 nom. protectum over Biophloeus Dejean, 1835 nom. oblitum (Cucujidae), Pachypus Dejean, 1821 nom. protectum over Pachypus Billberg, 1820 nom. oblitum (Scarabaeidae), Sparrmannia Laporte, 1840 nom. protectum over Leocaeta Dejean, 1833 nom. oblitum and Cephalotrichia Hope, 1837 nom. oblitum (Scarabaeidae).
Synoptic classification of the world Coleoptera
Order COLEOPTERA
†Suborder PROTOCOLEOPTERA
†Superfamily Tshekardocoleoidea Rohdendorf, 1944
†Family Tshekardocoleidae Rohdendorf, 1944
†Family Labradorocoleidae Ponomarenko, 1969
†Family Oborocoleidae Kukalová, 1969
†Superfamily Permocupedoidea Martynov, 1933
†Family Permocupedidae Martynov, 1933
†Family Taldycupedidae Rohdendorf, 1961
†Superfamily Permosynoidea Tillyard, 1924
†Family Ademosynidae Ponomarenko, 1968
†Family Permosynidae Tillyard, 1924
Suborder ARCHOSTEMATA
Family Crowsoniellidae Iablokoff-Khnzorian, 1983
Family Cupedidae Laporte, 1836
Subfamily Priacminae Crowson, 1962
†Subfamily Mesocupedinae Ponomarenko, 1969
Subfamily Cupedinae Laporte, 1836
Family Micromalthidae Barber, 1913
Family Ommatidae Sharp and Muir, 1912
†Subfamily Brochocoleinae Hong, 1982
Subfamily Tetraphalerinae Crowson, 1962
Subfamily Ommatinae Sharp and Muir, 1912
†Tribe Lithocupedini Ponomarenko, 1969
†Tribe Notocupedini Ponomarenko, 1966
Tribe Ommatini Sharp and Muir, 1912
Family Jurodidae Ponomarenko, 1985
†Family Triadocupedidae Ponomarenko, 1966
†Family Magnocoleidae Hong, 1998
†Family Obrieniidae Zherikhin and Gratshev, 1994
†Subfamily Kararhynchinae Zherikhin and Gratshev, 1994
†Tribe Kararhynchini Zherikhin and Gratshev, 1994
†Tribe Kenderlykaini Legalov, 2009
†Subfamily Obrieniinae Zherikhin and Gratshev, 1994
Suborder MYXOPHAGA
†Superfamily Asiocoleoidea Rohdendorf, 1961
†Family Asiocoleidae Rohdendorf, 1961
†Family Tricoleidae Ponomarenko, 1969
†Superfamily Rhombocoleoidea Rohdendorf, 1961
†Family Rhombocoleidae Rohdendorf, 1961
†Superfamily Schizophoroidea Ponomarenko, 1968
†Family Schizophoridae Ponomarenko, 1968
†Family Catiniidae Ponomarenko, 1968
†Family Schizocoleidae Rohdendorf, 1961
Superfamily Lepiceroidea Hinton, 1936 (1882)
Family Lepiceridae Hinton, 1936 (1882)
Superfamily Sphaeriusoidea Erichson, 1845
Family Torridincolidae Steffan, 1964
Subfamily Torridincolinae Steffan, 1964
Subfamily Deleveinae Endrödy-Younga, 1997
Family Hydroscaphidae LeConte, 1874
Family Sphaeriusidae Erichson, 1845
Suborder ADEPHAGA
†Family Tritarsidae Hong, 2002
Family Gyrinidae Latreille, 1810
Subfamily Spanglerogyrinae Folkerts, 1979
Subfamily Gyrininae Latreille, 1810
Tribe Enhydrini Régimbart, 1882
Subtribe Dineutina Desmarest, 1851
Subtribe Enhydrina Régimbart, 1882
Tribe Gyrinini Latreille, 1810
Subtribe Gyrinina Latreille, 1810
Subtribe Heterogyrina Brinck, 1956
Tribe Orectochilini Régimbart, 1882
Family Trachypachidae Thomson, 1857
†Subfamily Eodromeinae Ponomarenko, 1977
Subfamily Trachypachinae Thomson, 1857
Family Rhysodidae Laporte, 1840
Tribe Leoglymmiini Bell and Bell, 1978
Tribe Dhysorini Bell and Bell, 1978
Tribe Medisorini Bell and Bell, 1987
Tribe Rhysodini Laporte, 1840
Tribe Clinidiini Bell and Bell, 1978
Tribe Omoglymmiini Bell and Bell, 1978
Tribe Sloanoglymmiini Bell and Bell, 1991
Family Carabidae Latreille, 1802
†Subfamily Protorabinae Ponomarenko, 1977
†Subfamily Conjunctiinae Ponomarenko, 1977
Subfamily Nebriinae Laporte, 1834
Tribe Nebriini Laporte, 1834
Tribe Notiokasiini Kavanaugh and Nègre, 1983
Tribe Notiophilini Motschulsky, 1850
Tribe Opisthiini Dupuis, 1912
Tribe Pelophilini Kavanaugh, 1996
Subfamily Cicindinae Csiki, 1927
Subfamily Cicindelinae Latreille, 1802
Tribe Amblycheilini Csiki, 1903
Tribe Cicindelini Latreille, 1802
Subtribe Apteroessina Rivalier, 1971
Subtribe Cicindelina Latreille, 1802
Subtribe Dromicina Thomson, 1859
Subtribe Iresiina Rivalier, 1971
Subtribe Theratina Horn, 1893
Tribe Collyridini Brullé, 1834
Subtribe Collyridina Brullé, 1834
Subtribe Tricondylina Naviaux, 1991
Tribe Ctenostomatini Laporte, 1834
Tribe Manticorini Laporte, 1834
Tribe Megacephalini Laporte, 1834
Subfamily Carabinae Latreille, 1802
Tribe Carabini Latreille, 1802
Tribe Ceroglossini Lapouge, 1927
Tribe Cychrini Perty, 1830
Tribe Pamborini Hope, 1838
Subfamily Loricerinae Bonelli, 1810
Subfamily Omophroninae Bonelli, 1810
Subfamily Elaphrinae Latreille, 1802
Subfamily Migadopinae Chaudoir, 1861
Tribe Amarotypini Erwin, 1985
Tribe Migadopini Chaudoir, 1861
Subtribe Aquilicina Moret, 2005
Subtribe Migadopina Chaudoir, 1861
Subfamily Hiletinae Schiødte, 1848
Subfamily Scaritinae Bonelli, 1810
Tribe Carenini MacLeay, 1887
Tribe Clivinini Rafinesque, 1815
Subtribe Ardistomina Putzeys, 1867
Subtribe Clivinina Rafinesque, 1815
Subtribe Forcipatorina Bänninger, 1938
Tribe Dalyatini Mateu, 2002
Tribe Dyschiriini Kolbe, 1880
†Tribe Palaeoaxinidiini McKay, 1991
Tribe Pasimachini Putzeys, 1867
Tribe Promecognathini LeConte, 1853
Tribe Salcediini Alluaud, 1930 (1929)
Subtribe Androzelmina Bell, 1998
Subtribe Salcediina Alluaud, 1930 (1929)
Subtribe Solenogenyina Bell, 1998
Tribe Scaritini Bonelli, 1810
Subtribe Acanthoscelina Csiki, 1927
Subtribe Corintascarina Basilewsky, 1973
Subtribe Dyscherina Basilewsky, 1973
Subtribe Ochyropina Basilewsky, 1973
Subtribe Oxylobina Andrewes, 1929
Subtribe Scapterina Putzeys, 1867
Subtribe Scaritina Bonelli, 1810
Subtribe Storthodontina Jeannel, 1946
Subfamily Broscinae Hope, 1838
Tribe Broscini Hope, 1838
Subtribe Axonyina Roig-Juñent, 2000
Subtribe Baripodina Jeannel, 1941
Subtribe Broscina Hope, 1838
Subtribe Creobiina Jeannel, 1941
Subtribe Nothobroscina Roig-Juñent, 2000
Subfamily Apotominae LeConte, 1853
Subfamily Siagoninae Bonelli, 1813
Tribe Enceladini Horn, 1881
Tribe Lupercini Lecordier, 1977
Tribe Siagonini Bonelli, 1813
Subfamily Melaeninae Csiki, 1933
Subfamily Gehringiinae Darlington, 1933
Tribe Gehringiini Darlington, 1933
Subtribe Gehringiina Darlington, 1933
Subtribe Helenaeina Deuve, 2007
Subfamily Trechinae Bonelli, 1810
Tribe Bembidiini Stephens, 1827
Subtribe Anillina Jeannel, 1937
Subtribe Bembidiina Stephens, 1827
Subtribe Tachyina Motschulsky, 1862
Subtribe Xystosomina Erwin, 1994
Tribe Horologionini Jeannel, 1949
Tribe Pogonini Laporte, 1834
Tribe Trechini Bonelli, 1810
Subtribe Aepina Fowler, 1887
Subtribe Cnidina Jeannel, 1958
Subtribe Perileptina Sloane, 1903
Subtribe Plocamotrechina Jeannel, 1960
Subtribe Trechina Bonelli, 1810
Subtribe Trechodina Jeannel, 1926
Tribe Zolini Sharp, 1886
Subtribe Chalteniina Roig-Juñent and Cicchino, 2001
Subtribe Sinozolina Deuve, 1997
Subtribe Zolina Sharp, 1886
Subfamily Patrobinae Kirby, 1837
Tribe Lissopogonini Zamotajlov, 2000
Tribe Patrobini Kirby, 1837
Subtribe Deltomerina Chaudoir, 1871
Subtribe Deltomerodina Zamotajlov, 2002
Subtribe Patrobina Kirby, 1837
Subtribe Platidiolina Zamotajlov and Lafer, 2001
Subfamily Psydrinae LeConte, 1853
Tribe Amblytelini Blackburn, 1892
Tribe Mecyclothoracini Jeannel, 1940
Tribe Meonini Sloane, 1898
Tribe Moriomorphini Sloane, 1890
Tribe Psydrini LeConte, 1853
Tribe Tropopterini Sloane, 1898
Subfamily Nototylinae Bänninger, 1927
Subfamily Paussinae Latreille, 1806
Tribe Metriini LeConte, 1853
Tribe Mystropomini Horn, 1881
Tribe Ozaenini Hope, 1838
Tribe Paussini Latreille, 1806
†Subtribe Arthropteritina Luna de Carvalho, 1961
Subtribe Carabidomemnina Wasmann, 1928
Subtribe Cerapterina Billberg, 1820
†Subtribe Eopaussina Luna de Carvalho, 1951
Subtribe Heteropaussina Janssens, 1950
Subtribe Homopterina Wasmann, 1920
Subtribe Paussina Latreille, 1806
Subtribe Pentaplatarthrina Jeannel, 1946
Tribe Protopaussini Gestro, 1892
Subfamily Brachininae Bonelli, 1810
Tribe Brachinini Bonelli, 1810
Subtribe Aptinina Gistel, 1848
Subtribe Brachinina Bonelli, 1810
Subtribe Mastacina Erwin, 1970
Subtribe Pheropsophina Jeannel, 1949
Tribe Crepidogastrini Jeannel, 1949
Subfamily Harpalinae Bonelli, 1810
Tribe Abacetini Chaudoir, 1873
Tribe Amorphomerini Sloane, 1923
Tribe Anthiini Bonelli, 1813
Tribe Atranini Horn, 1881
Tribe Bascanini Basilewsky, 1953
Tribe Calophaenini Jeannel, 1948
Tribe Catapieseini Bates, 1882
Tribe Chaetodactylini Tschitschérine, 1903
Tribe Chaetogenyini Emden, 1958
Tribe Chlaeniini Brullé, 1834
Subtribe Callistina Laporte, 1834
Subtribe Chlaeniina Brullé, 1834
Tribe Cnemalobini Germain, 1911
Tribe Cratocerini Lacordaire, 1854
Tribe Ctenodactylini Laporte, 1834
Tribe Cuneipectini Sloane, 1907
Tribe Cyclosomini Laporte, 1834
Subtribe Cyclosomina Laporte, 1834
Subtribe Masoreina Chaudoir, 1871
Tribe Dercylini Sloane, 1923
Tribe Drimostomatini Chaudoir, 1872
Tribe Dryptini Bonelli, 1810
Tribe Enoicini Basilewsky, 1985
Tribe Galeritini Kirby, 1825
Subtribe Galeritina Kirby, 1825
Subtribe Planetina Jedlička, 1941
Tribe Geobaenini Péringuey, 1896
Tribe Ginemini Ball and Shpeley, 2002
Tribe Glyptini Horn, 1881
Tribe Graphipterini Latreille, 1802
Tribe Harpalini Bonelli, 1810
Subtribe Anisodactylina Lacordaire, 1854 nomen protectum
Subtribe Harpalina Bonelli, 1810
Subtribe Pelmatellina Bates, 1882
Subtribe Stenolophina Kirby, 1837
Tribe Helluonini Hope, 1838
Subtribe Helluonina Hope, 1838
Subtribe Omphrina Jedlička, 1941
Tribe Hexagoniini Horn, 1881 (1834)
Tribe Idiomorphini Bates, 1891
Tribe Lachnophorini LeConte, 1853
Subtribe Lachnophorina LeConte, 1853
Subtribe Selinina Jeannel, 1948
Tribe Lebiini Bonelli, 1810
Subtribe Actenonycina Bates, 1871
Subtribe Agrina Kirby, 1837
Subtribe Apenina Ball, 1983
Subtribe Calleidina Chaudoir, 1873
Subtribe Celaenephina Habu, 1982
Subtribe Cymindidina Laporte, 1834
Subtribe Demetriadina Bates, 1886
Subtribe Dromiusina Bonelli, 1810
Subtribe Gallerucidiina Chaudoir, 1872
Subtribe Lebiina Bonelli, 1810
Subtribe Metallicina Basilewsky, 1984
Subtribe Nemotarsina Bates, 1883
Subtribe Pericalina Hope, 1838
Subtribe Pseudotrechina Basilewsky, 1984
Subtribe Sugimotoina Habu, 1975
Subtribe Trichina Basilewsky, 1984
Tribe Licinini Bonelli, 1810
Subtribe Dicaelina Laporte, 1834
Subtribe Dicrochilina Ball, 1992
Subtribe Lestignathina Ball, 1992
Subtribe Licinina Bonelli, 1810
Tribe Melanchitonini Jeannel, 1948
Tribe Microcheilini Jeannel, 1948
Tribe Morionini Brullé, 1835
Tribe Odacanthini Laporte, 1834
Tribe Omphreini Ganglbauer, 1891
Tribe Oodini LaFerté-Sénectère, 1851
Tribe Orthogoniini Schaum, 1857
Tribe Panagaeini Bonelli, 1810
Subtribe Brachygnathina Basilewsky, 1946
Subtribe Panagaeina Bonelli, 1810
Subtribe Tefflina Basilewsky, 1946
Tribe Peleciini Chaudoir, 1880
Subtribe Agonicina Sloane, 1920
Subtribe Peleciina Chaudoir, 1880
Tribe Pentagonicini Bates, 1873
Tribe Perigonini Horn, 1881 nomen protectum
Tribe Physocrotaphini Chaudoir, 1863
Tribe Platynini Bonelli, 1810
Tribe Pseudomorphini Hope, 1838
Tribe Pterostichini Bonelli, 1810
Subtribe Abacomorphina Tschitschérine, 1902
Subtribe Euchroina Chaudoir, 1874
Subtribe Metiina Straneo, 1951
Subtribe Microcephalina Tschitschérine, 1898
Subtribe Pterostichina Bonelli, 1810
Tribe Sphodrini Laporte, 1834
Subtribe Atranopsina Baehr, 1982
Subtribe Calathina Laporte, 1834
Subtribe Dolichina Brullé, 1834
Subtribe Pristosiina Lindroth, 1956
Subtribe Sphodrina Laporte, 1834
Subtribe Synuchina Lindroth, 1956
Tribe Xenaroswellianini Erwin, 2007
Tribe Zabrini Bonelli, 1810
Subtribe Amarina Zimmermann, 1832
Subtribe Zabrina Bonelli, 1810
Tribe Zuphiini Bonelli, 1810
Subtribe Dicrodontina Machado, 1992
Subtribe Leleupidiina Basilewsky, 1951
Subtribe Metazuphiina Mateu, 1992
Subtribe Mischocephalina Mateu, 1992
Subtribe Patriziina Basilewsky, 1953
Subtribe Zuphiina Bonelli, 1810
Family Haliplidae Aubé, 1836
†Family Triaplidae Ponomarenko, 1977
†Family Colymbotethidae Ponomarenko, 1994
†Family Parahygrobiidae Ponomarenko, 1977
†Family Coptoclavidae Ponomarenko, 1961
†Subfamily Necronectinae Ponomarenko, 1977
†Subfamily Charonoscaphinae Ponomarenko, 1977
†Subfamily Coptoclavinae Ponomarenko, 1961
†Subfamily Coptoclaviscinae Soriano, Ponomarenko and Delclos, 2007
†Subfamily Hispanoclavinae Soriano, Ponomarenko and Delclos, 2007
†Family Liadytidae Ponomarenko, 1977
Family Meruidae Spangler and Steiner, 2005
Family Noteridae Thomson, 1860
Subfamily Noterinae Thomson, 1860
Tribe Neohydrocoptini Zalat, Saleh, Angus and Kaschef, 2000
Tribe Noterini Thomson, 1860
Tribe Pronoterini Nilsson, 2005
Tribe Tonerini Miller, 2009
Subfamily Notomicrinae Zimmermann, 1919
Subfamily Phreatodytinae Uéno, 1957
Family Amphizoidae LeConte, 1853
Family Aspidytidae Ribera, Beutel, Balke and Vogler, 2002
Family Hygrobiidae Régimbart, 1879 (1837)
Family Dytiscidae Leach, 1815
Subfamily Agabinae Thomson, 1867
Subfamily Colymbetinae Erichson, 1837
Tribe Anisomeriini Brinck, 1948
Tribe Carabdytini Pederzani, 1995
Tribe Colymbetini Erichson, 1837
Subfamily Copelatinae Branden, 1885
Subfamily Coptotominae Branden, 1885
Subfamily Dytiscinae Leach, 1815
Tribe Aciliini Thomson, 1867
Tribe Aubehydrini Guignot, 1942
Tribe Cybisterini Sharp, 1880
Tribe Dytiscini Leach, 1815
Tribe Eretini Crotch, 1873
Tribe Hydaticini Sharp, 1880
Tribe Hyderodini Miller, 2000
Subfamily Hydrodytinae Miller, 2001
Subfamily Hydroporinae Aubé, 1836
Tribe Bidessini Sharp, 1880
Tribe Carabhydrini Watts, 1978
Tribe Hydroporini Aubé, 1836
Tribe Hydrovatini Sharp, 1880
Tribe Hygrotini Portevin, 1929
Tribe Hyphydrini Gistel, 1848
Tribe Laccornini Wolfe and Roughley, 1990
Tribe Methlini Branden, 1885
†Tribe Schistomerini Palmer, 1957
Tribe Vatellini Sharp, 1880
Subfamily Laccophilinae Gistel, 1848
Tribe Agabetini Branden, 1885
Tribe Laccophilini Gistel, 1848
Subfamily Lancetinae Branden, 1885
Subfamily Matinae Branden, 1885
†Subfamily Palaeogyrininae Schlechtendal, 1894
†Subfamily Liadytiscinae Prokin and Ren, 2010
Suborder POLYPHAGA
Series STAPHYLINIFORMIA
Superfamily Hydrophiloidea Latreille, 1802
Family Hydrophilidae Latreille, 1802
Subfamily Helophorinae Leach, 1815
Subfamily Epimetopinae Zaitzev, 1908
Subfamily Georissinae Laporte, 1840
Subfamily Hydrochinae Thomson, 1859
Subfamily Spercheinae Erichson, 1837
Subfamily Horelophinae Hansen, 1991
Subfamily Horelophopsinae Hansen, 1997
Subfamily Hydrophilinae Latreille, 1802
Tribe Anacaenini Hansen, 1991
Tribe Berosini Mulsant, 1844
Tribe Chaetarthriini Bedel, 1881
Tribe Hydrophilini Latreille, 1802
Subtribe Acidocerina Zaitzev, 1908
Subtribe Globuloseina García, 2001
Subtribe Hydrobiusina Mulsant, 1844
Subtribe Hydrophilina Latreille, 1802
Tribe Laccobiini Houlbert, 1922
Tribe Sperchopsini Hansen, 1991
Subfamily Sphaeridiinae Latreille, 1802
Tribe Andotypini Hansen, 1991
Tribe Borborophorini Hansen, 1991
Tribe Coelostomatini Heyden, 1891 (1890)
Tribe Megasternini Mulsant, 1844
Tribe Omicrini Smetana, 1975
Tribe Protosternini Hansen, 1991
Tribe Rygmodini Orchymont, 1916
Tribe Sphaeridiini Latreille, 1802
Tribe Tormissini Hansen, 1991
Family Sphaeritidae Shuckard, 1839
Family Synteliidae Lewis, 1882
Family Histeridae Gyllenhal, 1808
Subfamily Niponiinae Fowler, 1912
Subfamily Abraeinae MacLeay, 1819
Tribe Abraeini MacLeay, 1819
Tribe Acritini Wenzel, 1944
Tribe Acritomorphini Wenzel, 1944
Tribe Plegaderini Portevin, 1929
Tribe Teretriini Bickhardt, 1914
Subfamily Trypeticinae Bickhardt, 1913
Subfamily Trypanaeinae Marseul, 1857
Subfamily Saprininae Blanchard, 1845
Subfamily Dendrophilinae Reitter, 1909
Tribe Anapleini Olexa, 1982
Tribe Bacaniini Kryzhanovskij, 1976
Tribe Dendrophilini Reitter, 1909
Tribe Paromalini Reitter, 1909
Subfamily Onthophilinae MacLeay, 1819
Subfamily Tribalinae Bickhardt, 1914
Subfamily Histerinae Gyllenhal, 1808
Tribe Exosternini Bickhardt, 1914
Tribe Histerini Gyllenhal, 1808
Tribe Hololeptini Hope, 1840
Tribe Omalodini Kryzhanovskij, 1972
Tribe Platysomatini Bickhardt, 1914
Subfamily Haeteriinae Marseul, 1857
Tribe Haeteriini Marseul, 1857
Tribe Nymphistrini Tishechkin, 2007
Tribe Synoditulini Tishechkin, 2007
Subfamily Chlamydopsinae Bickhardt, 1914
Superfamily Staphylinoidea Latreille, 1802
Family Hydraenidae Mulsant, 1844
Subfamily Orchymontiinae Perkins, 1997
Subfamily Prosthetopinae Perkins, 1994
Tribe Coelometoponini Perkins, 2005
Tribe Nucleotopini Perkins, 1994
Tribe Parasthetopini Perkins, 1994
Tribe Prosthetopini Perkins, 1994
Tribe Protosthetopini Perkins, 1994
Tribe Pterosthetopini Perkins, 1994
Subfamily Hydraeninae Mulsant, 1844
Tribe Hydraenidini Perkins, 1980
Tribe Hydraenini Mulsant, 1844
Tribe Limnebiini Mulsant, 1844
Tribe Madagastrini Perkins, 1997
Tribe Parhydraenini Perkins, 1997
Subfamily Ochthebiinae Thomson, 1859
Tribe Ochthebiini Thomson, 1859
Subtribe Enicocerina Perkins, 1997
Subtribe Meropathina Perkins, 1997
Subtribe Neochthebiina Perkins, 1997
Subtribe Ochthebiina Thomson, 1859
Subtribe Protochthebiina Perkins, 1997
Tribe Ochtheosini Perkins, 1997
Family Ptiliidae Erichson, 1845
Subfamily Ptiliinae Erichson, 1845
Tribe Discheramocephalini Grebennikov, 2009
Tribe Nanosellini Barber, 1924
Tribe Ptenidiini Flach, 1889
Tribe Ptiliini Erichson, 1845
Tribe Ptinellini Reitter, 1906 (1891)
Subfamily Cephaloplectinae Sharp, 1883
Subfamily Acrotrichinae Reitter, 1909 (1856)
Family Agyrtidae Thomson, 1859
Subfamily Agyrtinae Thomson, 1859
Subfamily Necrophilinae Newton, 1997
Subfamily Pterolomatinae Thomson, 1862
Family Leiodidae Fleming, 1821
Subfamily Camiarinae Jeannel, 1911
Tribe Agyrtodini Jeannel, 1936
Tribe Camiarini Jeannel, 1911
Tribe Neopelatopini Jeannel, 1962
Subfamily Catopocerinae Hatch, 1927 (1880)
Tribe Catopocerini Hatch, 1927 (1880)
Tribe Glacicavicolini Westcott, 1968
Subfamily Leiodinae Fleming, 1821
Tribe Agathidiini Westwood, 1838
Tribe Estadiini Portevin, 1914
Tribe Leiodini Fleming, 1821
Tribe Pseudoliodini Portevin, 1926
Tribe Scotocryptini Reitter, 1884
Tribe Sogdini Lopatin, 1961
Subfamily Coloninae Horn, 1880 (1859)
Subfamily Cholevinae Kirby, 1837
Tribe Anemadini Hatch, 1928
Subtribe Anemadina Hatch, 1928
Subtribe Eocatopina Jeannel, 1936
Subtribe Eunemadina Newton, 1998
Subtribe Nemadina Jeannel, 1936
Subtribe Paracatopina Jeannel, 1936
Tribe Cholevini Kirby, 1837
Subtribe Catopina Chaudoir, 1845
Subtribe Cholevina Kirby, 1837
Tribe Eucatopini Jeannel, 1921
Tribe Leptodirini Lacordaire, 1854 (1849)
Subtribe Anthroherponina Jeannel, 1910
Subtribe Bathysciina Horn, 1880
Subtribe Bathysciotina Guéorguiev, 1974
Subtribe Leptodirina Lacordaire, 1854 (1849)
Subtribe Pholeuina Reitter, 1886
Subtribe Platycholeina Horn, 1880
Subtribe Spelaeobatina Guéorguiev, 1974
Tribe Oritocatopini Jeannel, 1936
Tribe Ptomaphagini Jeannel, 1911
Subtribe Baryodirina Perreau, 2000
Subtribe Ptomaphagina Jeannel, 1911
Subtribe Ptomaphaginina Szymczakowski, 1964
Tribe Sciaphyini Perreau, 2000
Subfamily Platypsyllinae Ritsema, 1869
Family Silphidae Latreille, 1806
Subfamily Silphinae Latreille, 1806
Subfamily Nicrophorinae Kirby, 1837
Family Staphylinidae Latreille, 1802
Subfamily Glypholomatinae Jeannel, 1962
Subfamily Microsilphinae Crowson, 1950
Subfamily Omaliinae MacLeay, 1825
Tribe Anthophagini Thomson, 1859
Tribe Aphaenostemmini Peyerimhoff, 1914
Tribe Corneolabiini Steel, 1950
Tribe Coryphiini Jakobson, 1908
Subtribe Boreaphilina Zerche, 1990
Subtribe Coryphiina Jakobson, 1908
Tribe Eusphalerini Hatch, 1957
Tribe Hadrognathini Portevin, 1929
Tribe Omaliini MacLeay, 1825
Subfamily Empelinae Newton and Thayer, 1992
Subfamily Proteininae Erichson, 1839
Tribe Anepiini Steel, 1966
Tribe Austrorhysini Newton and Thayer, 1995
Tribe Nesoneini Steel, 1966
Tribe Proteinini Erichson, 1839
Tribe Silphotelini Newton and Thayer, 1995
Subfamily Micropeplinae Leach, 1815
Subfamily Neophoninae Fauvel, 1905
Subfamily Dasycerinae Reitter, 1887
Subfamily Protopselaphinae Newton and Thayer, 1995
Subfamily Pselaphinae Latreille, 1802
Supertribe Batrisitae Reitter, 1882
Tribe Amauropini Jeannel, 1948
Tribe Batrisini Reitter, 1882
Subtribe Ambicocerina Leleup, 1970
Subtribe Batrisina Reitter, 1882
Subtribe Leupeliina Jeannel, 1954
Subtribe Stilipalpina Jeannel, 1954
Tribe Thaumastocephalini Poggi, Nonveiller, Colla, Pavićević and Rada, 2001
Supertribe Clavigeritae Leach, 1815
Tribe Clavigerini Leach, 1815
Subtribe Apoderigerina Jeannel, 1954
Subtribe Clavigerina Leach, 1815
Subtribe Clavigerodina Schaufuss, 1882
Subtribe Disarthricerina Jeannel, 1949
Subtribe Hoplitoxenina Célis, 1969
Subtribe Lunillina Célis, 1969
Subtribe Mastigerina Jeannel, 1954
Subtribe Miroclavigerina Jeannel, 1949
Subtribe Neocerina Jeannel, 1954
Subtribe Radamina Jeannel, 1954
Subtribe Thysdariina Jeannel, 1954
Tribe Colilodionini Besuchet, 1991
Tribe Tiracerini Besuchet, 1986
Supertribe Euplectitae Streubel, 1839
Tribe Bythinoplectini Schaufuss, 1890
Subtribe Bythinoplectina Schaufuss, 1890
Subtribe Pyxidicerina Raffray, 1904
Tribe Dimerini Raffray, 1908
Tribe Euplectini Streubel, 1839
Tribe Jubini Raffray, 1904
Tribe Mayetiini Winkler, 1925
Tribe Metopiasini Raffray, 1904
Subtribe Metopiasina Raffray, 1904
Subtribe Rhinoscepsina Bowman, 1934
Tribe Trichonychini Reitter, 1882
Subtribe Bibloporina Park, 1951
Subtribe Panaphantina Jeannel, 1950
Subtribe Trichonychina Reitter, 1882
Subtribe Trimiina Bowman, 1934
Tribe Trogastrini Jeannel, 1949
Subtribe Phtegnomina Park, 1951
Subtribe Rhexiina Park, 1951
Subtribe Trogastrina Jeannel, 1949
Supertribe Faronitae Reitter, 1882
Supertribe Goniaceritae Reitter, 1882 (1872)
Tribe Arnylliini Jeannel, 1952
Tribe Barrosellini Leleup, 1973
Tribe Brachyglutini Raffray, 1904
Subtribe Baradina Park, 1951
Subtribe Brachyglutina Raffray, 1904
Subtribe Decarthrina Park, 1951
Subtribe Eupseniina Park, 1951
Tribe Bythinini Raffray, 1890
Subtribe Bythinina Raffray, 1890
Subtribe Machaeritina Jeannel, 1950
Subtribe Xenobythina Jeannel, 1950
Tribe Cyathigerini Schaufuss, 1872
Tribe Goniacerini Reitter, 1882 (1872)
Tribe Imirini Jeannel, 1949
Tribe Iniocyphini Park, 1951
Subtribe Iniocyphina Park, 1951
Subtribe Natypleurina Newton and Thayer, 1992
Tribe Machadoini Jeannel, 1951
Tribe Proterini Jeannel, 1949
Tribe Pygoxyini Reitter, 1909
Tribe Speleobamini Park, 1951
Tribe Tychini Raffray, 1904
Tribe Valdini Park, 1953
Supertribe Pselaphitae Latreille, 1802
Tribe Arhytodini Raffray, 1890
Tribe Attapseniini Bruch, 1933
Tribe Ctenistini Blanchard, 1845
Tribe Hybocephalini Raffray, 1890
Tribe Odontalgini Jeannel, 1949
Tribe Pachygastrodini Leleup, 1969
Tribe Phalepsini Jeannel, 1949
Tribe Pselaphini Latreille, 1802
Tribe Schistodactylini Raffray, 1890
Tribe Tmesiphorini Jeannel, 1949
Tribe Tyrini Reitter, 1882
Subtribe Centrophthalmina Jeannel, 1949
Subtribe Janusculina Cerruti, 1970
Subtribe Somatipionina Jeannel, 1949
Subtribe Tyrina Reitter, 1882
Subfamily Phloeocharinae Erichson, 1839
Subfamily Olisthaerinae Thomson, 1858
Subfamily Tachyporinae MacLeay, 1825
Tribe Deropini Smetana, 1983
Tribe Megarthropsini Cameron, 1919
Tribe Mycetoporini Thomson, 1859
Tribe Tachyporini MacLeay, 1825
Tribe Vatesini Seevers, 1958
Subfamily Trichophyinae Thomson, 1858
Subfamily Habrocerinae Mulsant and Rey, 1876
Subfamily Aleocharinae Fleming, 1821
Tribe Actocharini Bernhauer and Schubert, 1911
Tribe Aenictoteratini Kistner, 1993
Tribe Akatastopsisini Pace, 2000
Tribe Aleocharini Fleming, 1821
Subtribe Aleocharina Fleming, 1821
Subtribe Compactopediina Kistner, 1970
Subtribe Hodoxenina Kistner, 1970
Tribe Athetini Casey, 1910
Subtribe Athetina Casey, 1910
Subtribe Coptotermoeciina Kistner and Pasteels, 1970
Subtribe Microceroxenina Kistner, 1970
Subtribe Nasutiphilina Kistner, 1970
Subtribe Schistogeniina Fenyes, 1918
Subtribe Taxicerina Lohse, 1989
Subtribe Termitotelina Kistner, 1970
Subtribe Thamiaraeina Fenyes, 1921
Tribe Autaliini Thomson, 1859
Tribe Cordobanini Bernhauer, 1910
Tribe Corotocini Fenyes, 1918
Subtribe Abrotelina Seevers, 1957
Subtribe Corotocina Fenyes, 1918
Subtribe Eburniogastrina Jacobson, Kistner and Pasteels, 1986
Subtribe Nasutitellina Jacobson, Kistner and Pasteels, 1986
Subtribe Sphuridaethina Pace, 1988
Subtribe Termitocharina Seevers, 1957
Subtribe Termitocupidina Jacobson, Kistner and Pasteels, 1986
Subtribe Termitogastrina Bernhauer and Scheerpeltz, 1926
Subtribe Termitoiceina Jacobson, Kistner and Pasteels, 1986
Subtribe Termitopithina Jacobson, Kistner and Pasteels, 1986
Subtribe Termitoptochina Fenyes, 1921
Subtribe Timeparthenina Fenyes, 1921
Tribe Crematoxenini Mann, 1921
Tribe Cryptonotopseini Pace, 2003
Tribe Deinopsini Sharp, 1883
Tribe Diestotini Mulsant and Rey, 1871
Tribe Diglottini Jakobson, 1909
Tribe Digrammini Fauvel, 1900
Tribe Dorylogastrini Wasmann, 1916
Tribe Dorylomimini Wasmann, 1916
Tribe Drepanoxenini Kistner and Watson, 1972
Tribe Ecitocharini Seevers, 1965
Tribe Ecitogastrini Fenyes, 1918
Tribe Eusteniamorphini Bernhauer and Scheerpeltz, 1926
Tribe Falagriini Mulsant and Rey, 1873
Tribe Feldini Kistner, 1972
Tribe Gymnusini Heer, 1839
Tribe Himalusini Klimaszewski, Pace and Center, 2010
Tribe Homalotini Heer, 1839
Subtribe Bolitocharina Thomson, 1859
Subtribe Dinardopsina Bernhauer and Scheerpeltz, 1926
Subtribe Gyrophaenina Kraatz, 1856
Subtribe Homalotina Heer, 1839
Subtribe Silusina Fenyes, 1918
Tribe Hoplandriini Casey, 1910
Subtribe Hoplandriina Casey, 1910
Subtribe Platandriina Hanley, 2002
Subtribe Pseudoplandriina Hanley, 2002
Tribe Hygronomini Thomson, 1859
Subtribe Hygronomina Thomson, 1859
Subtribe Saphoglossina Bernhauer and Scheerpeltz, 1926
Tribe Hypocyphtini Laporte, 1835
Tribe Leucocraspedini Fenyes, 1921
Tribe Liparocephalini Fenyes, 1918
Tribe Lomechusini Fleming, 1821
Subtribe Aenictobiina Kistner, 1997
Subtribe Lomechusina Fleming, 1821
Subtribe Myrmedoniina Thomson, 1867
Subtribe Termitozyrina Seevers, 1957
Tribe Masuriini Cameron, 1939
Tribe Mesoporini Cameron, 1959
Tribe Mimanommatini Wasmann, 1912
Subtribe Dorylophilina Fenyes, 1921
Subtribe Mimanommatina Wasmann, 1912
Tribe Mimecitini Wasmann, 1917
Subtribe Labidopullina Jacobson and Kistner, 1991
Subtribe Leptanillophilina Fenyes, 1918
Subtribe Mimecitina Wasmann, 1917
Subtribe Mimonillina Bernhauer and Scheerpeltz, 1926
Tribe Myllaenini Ganglbauer, 1895
Tribe Oxypodini Thomson, 1859
Subtribe Aphytopodina Bernhauer and Scheerpeltz, 1926
Subtribe Blepharhymenina Klimaszewski and Peck, 1986
Subtribe Dinardina Mulsant and Rey, 1873
Subtribe Meoticina Seevers, 1978
Subtribe Oxypodina Thomson, 1859
Subtribe Tachyusina Thomson, 1859
Tribe Oxypodinini Fenyes, 1921
Tribe Paglini Newton and Thayer, 1992
Tribe Paradoxenusini Bruch, 1937
Tribe Pediculotini Ádám, 1987
Tribe Philotermitini Seevers, 1957
Tribe Phyllodinardini Wasmann, 1916
Tribe Phytosini Thomson, 1867
Tribe Placusini Mulsant and Rey, 1871
Tribe Pronomaeini Mulsant and Rey, 1873
Tribe Pseudoperinthini Cameron, 1939
Tribe Pygostenini Fauvel, 1899
Tribe Sahlbergiini Kistner, 1993
Tribe Sceptobiini Seevers, 1978
Tribe Skatitoxenini Kistner and Pasteels, 1969
Tribe Termitodiscini Wasmann, 1904
Subtribe Athexeniina Pace, 2000
Subtribe Termitodiscina Wasmann, 1904
Tribe Termitohospitini Seevers, 1941
Subtribe Hetairotermitina Seevers, 1957
Subtribe Termitohospitina Seevers, 1941
Tribe Termitonannini Fenyes, 1918
Subtribe Perinthina Bernhauer and Scheerpeltz, 1926
Subtribe Termitonannina Fenyes, 1918
Tribe Termitopaediini Seevers, 1957
Tribe Termitusini Fenyes, 1918
Subtribe Termitospectrina Seevers, 1957
Subtribe Termitusina Fenyes, 1918
Tribe Trichopseniini LeConte and Horn, 1883
Tribe Trilobitideini Fauvel, 1899
Subfamily Trigonurinae Reiche, 1866
Subfamily Apateticinae Fauvel, 1895
Subfamily Scaphidiinae Latreille, 1806
Tribe Cypariini Achard, 1924
Tribe Scaphidiini Latreille, 1806
Tribe Scaphiini Achard, 1924
Tribe Scaphisomatini Casey, 1893
Subfamily Piestinae Erichson, 1839
Subfamily Osoriinae Erichson, 1839
Tribe Eleusinini Sharp, 1887
Tribe Leptochirini Sharp, 1887
Tribe Osoriini Erichson, 1839
Subtribe Osoriina Erichson, 1839
Subtribe Parosoriina Bernhauer and Schubert, 1911
Tribe Thoracophorini Reitter, 1909
Subtribe Clavilispinina Newton and Thayer, 1992
Subtribe Glyptomina Newton and Thayer, 1992
Subtribe Lispinina Bernhauer and Schubert, 1910
Subtribe Thoracophorina Reitter, 1909
Subfamily Oxytelinae Fleming, 1821
Tribe Blediini Ádám, 2001
Tribe Coprophilini Heer, 1839
Tribe EuphaniiniReitter, 1909
Tribe Oxytelini Fleming, 1821
Tribe Planeustomini Jacquelin du Val, 1857
Subfamily Oxyporinae Fleming, 1821
Subfamily Megalopsidiinae Leng, 1920
Subfamily Scydmaeninae Leach, 1815
†Supertribe Hapsomelitae Poinar and Brown, 2004
Supertribe Mastigitae Fleming, 1821
Tribe Clidicini Casey, 1897
Tribe Leptomastacini Casey, 1897
Tribe Mastigini Fleming, 1821
Supertribe Scydmaenitae Leach, 1815
Tribe Cephenniini Reitter, 1882
Tribe Chevrolatiini Reitter, 1882
Tribe Cyrtoscydmini Schaufuss, 1889
Tribe Eutheiini Casey, 1897
Tribe Leptoscydmini Casey, 1897
Tribe Plaumanniolini Costa Lima, 1962
Tribe Scydmaenini Leach, 1815
Subfamily Steninae MacLeay, 1825
Subfamily Euaesthetinae Thomson, 1859
Tribe Alzadaesthetini Scheerpeltz, 1974
Tribe Austroesthetini Cameron, 1944
Tribe Euaesthetini Thomson, 1859
Tribe Fenderiini Scheerpeltz, 1974
Tribe Nordenskioldiini Bernhauer and Schubert, 1911
Tribe Stenaesthetini Bernhauer and Schubert, 1911
Subfamily Solieriinae Newton and Thayer, 1992
Subfamily Leptotyphlinae Fauvel, 1874
Tribe Cephalotyphlini Coiffait, 1963
Tribe Entomoculiini Coiffait, 1957
Tribe Leptotyphlini Fauvel, 1874
Tribe Metrotyphlini Coiffait, 1963
Tribe Neotyphlini Coiffait, 1963
Subfamily Pseudopsinae Ganglbauer, 1895
Subfamily Paederinae Fleming, 1821
Tribe Paederini Fleming, 1821
Subtribe Astenina Hatch, 1957
Subtribe Cryptobiina Casey, 1905
Subtribe Cylindroxystina Bierig, 1943
Subtribe Dolicaonina Casey, 1905
Subtribe Echiasterina Casey, 1905
Subtribe Lathrobiina Laporte, 1835
Subtribe Lithocharina Casey, 1905
Subtribe Medonina Casey, 1905
Subtribe Paederina Fleming, 1821
Subtribe Scopaeina Mulsant and Rey, 1878
Subtribe Stilicina Casey, 1905
Subtribe Stilicopsina Casey, 1905
Tribe Pinophilini Nordmann, 1837
Subtribe Pinophilina Nordmann, 1837
Subtribe Procirrina Bernhauer and Schubert, 1912
Subfamily Staphylininae Latreille, 1802
Tribe Arrowinini Solodovnikov and Newton, 2005
Tribe Diochini Casey, 1906
Tribe Maorothiini Assing, 2000
Tribe Othiini Thomson, 1859
Tribe Platyprosopini Lynch Arribálzaga, 1884
Tribe Staphylinini Latreille, 1802
Subtribe Amblyopinina Seevers, 1944
Subtribe Anisolinina Hayashi, 1993
Subtribe Eucibdelina Sharp, 1889
Subtribe Hyptiomina Casey, 1906
Subtribe Philonthina Kirby, 1837
Subtribe Quediina Kraatz, 1857
Subtribe Staphylinina Latreille, 1802
Subtribe Tanygnathinina Reitter, 1909
Subtribe Xanthopygina Sharp, 1884
Tribe Xantholinini Erichson, 1839
†Subfamily Protactinae Heer, 1847
Series SCARABAEIFORMIA
Superfamily Scarabaeoidea Latreille, 1802
Family Pleocomidae LeConte, 1861
Subfamily Pleocominae LeConte, 1861
†Subfamily Cretocominae Nikolajev, 2002
†Subfamily Archescarabaeinae Nikolajev, 2010
Family Geotrupidae Latreille, 1802
Subfamily Taurocerastinae Germain, 1897
Subfamily Bolboceratinae Mulsant, 1842
Tribe Athyreini Lynch Arribálzaga, 1878
Tribe Bolbelasmini Nikolajev, 1996
Tribe Bolboceratini Mulsant, 1842
Tribe Bolbochromini Nikolajev, 1970
Tribe Eubolbitini Nikolajev, 1970
Tribe Eucanthini Nikolajev, 2003
Tribe Gilletinini Nikolajev, 1990
Tribe Odonteini Shokhin, 2007
Tribe Stenaspidiini Nikolajev, 2003
Subfamily Geotrupinae Latreille, 1802
Tribe Ceratotrupini Zunino, 1984
Tribe Enoplotrupini Paulian, 1945
†Tribe Cretogeotrupini Nikolajev, 1996
Tribe Geotrupini Latreille, 1802
Tribe Lethrini Oken, 1843
Family Belohinidae Paulian, 1959
Family Passalidae Leach, 1815
Subfamily Aulacocyclinae Kaup, 1868
Tribe Aulacocyclini Kaup, 1868
Tribe Ceracupedini Boucher, 2006
Subfamily Passalinae Leach, 1815
Tribe Leptaulacini Kaup, 1871
Tribe Macrolinini Kaup, 1871
Tribe Passalini Leach, 1815
Tribe Proculini Kaup, 1868
Tribe Solenocyclini Kaup, 1871
Family Trogidae MacLeay, 1819
†Subfamily Avitortorinae Nikolajev, 2007
Subfamily Troginae MacLeay, 1819
Subfamily Omorginae Nikolajev, 2005
Family Glaresidae Kolbe, 1905
Family Diphyllostomatidae Holloway, 1972
Family Lucanidae Latreille, 1804
†Subfamily Protolucaninae Nikolajev, 2007
Subfamily Aesalinae MacLeay, 1819
Tribe Aesalini MacLeay, 1819
Tribe Ceratognathini Sharp, 1899
Tribe Nicagini LeConte, 1861
†Subfamily Ceruchitinae Nikolajev, 2006
Subfamily Syndesinae MacLeay, 1819
Subfamily Lampriminae MacLeay, 1819
Tribe Lamprimini MacLeay, 1819
Tribe Streptocerini Kikuta, 1986
Subfamily Lucaninae Latreille, 1804
Tribe Chiasognathini Burmeister, 1847
Tribe Lucanini Latreille, 1804
Tribe Platycerini Mulsant, 1842
Tribe Platyceroidini Paulsen and Hawks, 2008
†Subfamily Paralucaninae Nikolajev, 2000
Family Ochodaeidae Mulsant and Rey, 1871
†Subfamily Cretochodaeinae Nikolajev, 1995
Subfamily Ochodaeinae Mulsant and Rey, 1871
Tribe Enodognathini Scholtz, 1988
Tribe Ochodaeini Mulsant and Rey, 1871
Subfamily Chaetocanthinae Scholtz, 1988
Tribe Chaetocanthini Scholtz, 1988
Tribe Pseudochodaeini Scholtz, 1988
Tribe Synochodaeini Scholtz, 1988
Family Hybosoridae Erichson, 1847
†Subfamily Mimaphodiinae Nikolajev, 2007
Subfamily Anaidinae Nikolajev, 1996
Subfamily Ceratocanthinae Martínez, 1968
Tribe Ceratocanthini Martínez, 1968
Tribe Ivieolini Howden and Gill, 2000
Tribe Scarabatermitini Nikolajev, 1999
Subfamily Hybosorinae Erichson, 1847
Subfamily Liparochrinae Ocampo, 2006
Subfamily Pachyplectrinae Ocampo, 2006
Family Glaphyridae MacLeay, 1819
Subfamily Glaphyrinae MacLeay, 1819
Subfamily Amphicominae Blanchard, 1845
†Subfamily Cretoglaphyrinae Nikolajev, 2005
Family Scarabaeidae Latreille, 1802
†Subfamily Lithoscarabaeinae Nikolajev, 1992
Subfamily Chironinae Blanchard, 1845
Subfamily Aegialiinae Laporte, 1840
Subfamily Eremazinae Iablokoff-Khnzorian, 1977
Subfamily Aphodiinae Leach, 1815
Tribe Aphodiini Leach, 1815
Subtribe Aphodiina Leach, 1815
Subtribe Didactyliina Pittino, 1985
Subtribe Proctophanina Stebnicka and Howden, 1995
Tribe Corythoderini Schmidt, 1910
Tribe Eupariini Schmidt, 1910 nomen protectum
Tribe Odontolochini Stebnicka and Howden, 1996
Tribe Odochilini Rakovič, 1987
Tribe Psammodiini Mulsant, 1842
Subtribe Phycocina Landin, 1960
Subtribe Psammodiina Mulsant, 1842
Subtribe Rhyssemina Pittino and Mariani, 1986
Tribe Rhyparini Schmidt, 1910
Tribe Stereomerini Howden and Storey, 1992
Tribe Termitoderini Tangelder and Krikken, 1982
Subfamily Aulonocneminae Janssens, 1946
Subfamily Termitotroginae Wasmann, 1918
Subfamily Scarabaeinae Latreille, 1802
Tribe Ateuchini Perty, 1830
Subtribe Ateuchina Perty, 1830
Subtribe Scatimina Vaz-de-Mello, 2008
Tribe Coprini Leach, 1815
Tribe Deltochilini Lacordaire, 1856
Tribe Eucraniini Burmeister, 1873
Tribe Gymnopleurini Lacordaire, 1856
Tribe Oniticellini Kolbe, 1905
Subtribe Drepanocerina van Lansberge, 1875
Subtribe Eurysternina Vulcano, Martínez and Pereira, 1961
Subtribe Helictopleurina Janssens, 1946
Subtribe Oniticellina Kolbe, 1905
Tribe Onitini Laporte, 1840
Tribe Onthophagini Burmeister, 1846
Tribe Phanaeini Hope, 1838
Tribe Scarabaeini Latreille, 1802
Tribe Sisyphini Mulsant, 1842
†Subfamily Prototroginae Nikolajev, 2000
†Subfamily Cretoscarabaeinae Nikolajev, 1995
Subfamily Dynamopodinae Arrow, 1911
Tribe Dynamopodini Arrow, 1911
Tribe Thinorycterini Semenov and Reichardt, 1925
Subfamily Phaenomeridinae Erichson, 1847
Subfamily Orphninae Erichson, 1847
Tribe Aegidiini Paulian, 1984
Tribe Orphnini Erichson, 1847
Subfamily Allidiostomatinae Arrow, 1940
Subfamily Aclopinae Blanchard, 1850
Tribe Aclopini Blanchard, 1850
†Tribe Holcorobeini Nikolajev, 1992
Tribe Phaenognathini Iablokoff-Khnzorian, 1977
Subfamily Melolonthinae Leach, 1819
Tribe Ablaberini Blanchard, 1850
Tribe Automoliini Britton, 1978
Tribe Chasmatopterini Lacordaire, 1856
Tribe Colymbomorphini Blanchard, 1850
Tribe Comophorinini Britton, 1957
†Tribe Cretomelolonthini Nikolajev, 1998
Tribe Dichelonychini Burmeister, 1855
Tribe Diphucephalini Laporte, 1840
Tribe Diphycerini Medvedev, 1952
Tribe Diplotaxini Kirby, 1837
Tribe Euchirini Hope, 1840
Tribe Heteronychini Lacordaire, 1856
Tribe Hopliini Latreille, 1829
Subtribe Hopliina Latreille, 1829
Subtribe Pachycnemina Laporte, 1840
Tribe Lichniini Burmeister, 1844
Tribe Liparetrini Burmeister, 1855
Tribe Macrodactylini Kirby, 1837
Tribe Maechidiini Burmeister, 1855
Tribe Melolonthini Leach, 1819
Subtribe Enariina Dewailly, 1950
Subtribe Heptophyllina Medvedev, 1951
Subtribe Leucopholina Burmeister, 1855
Subtribe Melolonthina Leach, 1819
Subtribe Pegylina Lacroix, 1989
Subtribe Rhizotrogina Burmeister, 1855
Subtribe Schizonychina Burmeister, 1855
Tribe Oncerini LeConte, 1861
Tribe Pachypodini Erichson, 1840
Tribe Pachytrichini Burmeister, 1855
Tribe Phyllotocidiini Britton, 1957
Tribe Podolasiini Howden, 1997
Tribe Scitalini Britton, 1957
Tribe Sericini Kirby, 1837
Subtribe Phyllotocina Burmeister, 1855
Subtribe Sericina Kirby, 1837
Subtribe Trochalina Brenske, 1898
Tribe Sericoidini Erichson, 1847
Tribe Systellopini Sharp, 1877
Tribe Tanyproctini Erichson, 1847
Subtribe Macrophyllina Burmeister, 1855
Subtribe Tanyproctina Erichson, 1847
Subfamily Rutelinae MacLeay, 1819
Tribe Adoretini Burmeister, 1844
Subtribe Adoretina Burmeister, 1844
Subtribe Adorrhinyptiina Arrow, 1917
Subtribe Pachyrhinadoretina Ohaus, 1912
Subtribe Prodoretina Ohaus, 1912
Subtribe Trigonostomusina Ohaus, 1912
Tribe Alvarengiini Frey, 1975
Tribe Anatistini Lacordaire, 1856
Tribe Anomalini Streubel, 1839 nomen protectum
Subtribe Anisopliina Burmeister, 1844
Subtribe Anomalina Streubel, 1839 nomen protectum
Subtribe Isopliina Péringuey, 1902
Subtribe Leptohopliina Potts, 1974
Subtribe Popilliina Ohaus, 1918
Tribe Anoplognathini MacLeay, 1819
Subtribe Anoplognathina MacLeay, 1819
Subtribe Brachysternina Burmeister, 1844
Subtribe Phalangogoniina Ohaus, 1918
Subtribe Platycoeliina Burmeister, 1844
Subtribe Schizognathina Ohaus, 1918
Tribe Geniatini Burmeister, 1844
Tribe Rutelini MacLeay, 1819
Subtribe Areodina Burmeister, 1844
Subtribe Desmonychina Arrow, 1917
Subtribe Didrepanephorina Ohaus, 1918
Subtribe Heterosternina Bates, 1888 nomen protectum
Subtribe Lasiocalina Ohaus, 1918
Subtribe Oryctomorphina Burmeister, 1847
Subtribe Parastasiina Burmeister, 1844
Subtribe Rutelina MacLeay, 1819
Subfamily Dynastinae MacLeay, 1819
Tribe Agaocephalini Burmeister, 1847
Tribe Cyclocephalini Laporte, 1840
Tribe Dynastini MacLeay, 1819
Tribe Hexodontini Lacordaire, 1856
Tribe Oryctini Mulsant, 1842
Tribe Oryctoderini Endrödi, 1966
Tribe Pentodontini Mulsant, 1842
Subtribe Cheiroplatina Carne, 1957
Subtribe Dipelicina Carne, 1957
Subtribe Pentodontina Mulsant, 1842
Subtribe Pseudoryctina Carne, 1957
Tribe Phileurini Burmeister, 1847
Subtribe Cryptodina Burmeister and Schaum, 1840
Subtribe Phileurina Burmeister, 1847
Subfamily Cetoniinae Leach, 1815
Tribe Cetoniini Leach, 1815
Subtribe Cetoniina Leach, 1815
Subtribe Euphoriina Horn, 1880
Subtribe Leucocelina Kraatz, 1882
Tribe Cremastocheilini Burmeister and Schaum, 1841
Subtribe Aspilina Krikken, 1984
Subtribe Coenochilina Burmeister, 1842
Subtribe Cremastocheilina Burmeister and Schaum, 1841
Subtribe Cymophorina Krikken, 1984
Subtribe Genuchina Krikken, 1984
Subtribe Goliathopsidina Krikken, 1984
Subtribe Heterogeniina Krikken, 1984
Subtribe Lissogeniina Krikken, 1984
Subtribe Macromina Burmeister and Schaum, 1840
Subtribe Nyassinina Krikken, 1984
Subtribe Oplostomina Krikken, 1984
Subtribe Pilinurgina Krikken, 1984
Subtribe Spilophorina Krikken, 1984
Subtribe Telochilina Krikken, 1984
Subtribe Trichoplina Krikken, 1984
Subtribe Trogodina Krikken, 1984
Tribe Diplognathini Burmeister, 1842
Tribe Goliathini Latreille, 1829
Subtribe Coryphocerina Burmeister, 1842
Subtribe Dicronocephalina Krikken, 1984
Subtribe Goliathina Latreille, 1829
Subtribe Ichnestomatina Burmeister, 1842
Tribe Gymnetini Kirby, 1827
Subtribe Blaesiina Schoch, 1895
Subtribe Gymnetina Kirby, 1827
Tribe Phaedimini Schoch, 1894
Tribe Schizorhinini Burmeister, 1842
Subtribe Lomapterina Burmeister, 1842
Subtribe Schizorhinina Burmeister, 1842
Tribe Stenotarsiini Kraatz, 1880
Subtribe Anochiliina Krikken, 1984
Subtribe Coptomiina Schenkling, 1921
Subtribe Chromoptiliina Krikken, 1984
Subtribe Doryscelina Schenkling, 1921
Subtribe Euchroeina Paulian and Descarpentries, 1982
Subtribe Heterophanina Schoch, 1894
Subtribe Heterosomatina Krikken, 1984
Subtribe Pantoliina Krikken, 1984
Subtribe Parachiliina Krikken, 1984
Subtribe Stenotarsiina Kraatz, 1880
Tribe Taenioderini Mikšić, 1976
Subtribe Chalcotheina Mikšić, 1976
Subtribe Taenioderina Mikšić, 1976
Tribe Trichiini Fleming, 1821
Subtribe Cryptodontina Lacordaire, 1856
Subtribe Incina Burmeister, 1842
Subtribe Osmodermatina Schenkling, 1922
Subtribe Platygeniina Krikken, 1984
Subtribe Trichiina Fleming, 1821
Tribe Valgini Mulsant, 1842
Subtribe Microvalgina Kolbe, 1904
Subtribe Valgina Mulsant, 1842
Tribe Xiphoscelidini Burmeister, 1842
†Family Coprinisphaeridae Genise, 2004
†Family Pallichnidae Genise, 2004
Series ELATERIFORMIA
Superfamily Scirtoidea Fleming, 1821
Family Decliniidae Nikitsky, Lawrence, Kirejtshuk and Gratshev, 1994
Family Eucinetidae Lacordaire, 1857
Family Clambidae Fischer von Waldheim, 1821
Subfamily Calyptomerinae Crowson, 1955
Subfamily Acalyptomerinae Crowson, 1979
Subfamily Clambinae Fischer von Waldheim, 1821
Family Scirtidae Fleming, 1821
Subfamily Scirtinae Fleming, 1821
Subfamily Nipponocyphoninae Lawrence and Yoshitomi, 2007
Subfamily Stenocyphoninae Lawrence and Yoshitomi, 2007
†Family Elodophthalmidae Kirejtshuk and Azar, 2008
†Family Mesocinetida Kirejtshuk and Ponomarenko, 2010
Superfamily Dascilloidea Guérin-Méneville, 1843 (1834)
Family Dascillidae Guérin-Méneville, 1843 (1834)
Subfamily Dascillinae Guérin-Méneville, 1843 (1834)
Tribe Cinnabariini Pic, 1914
Tribe Dascillini Guérin-Méneville, 1843 (1834)
Subfamily Karumiinae Escalera, 1913
Tribe Emmitini Escalera, 1914
Tribe Escalerinini Paulus, 1972
Tribe Genecerini Pic, 1914
Tribe Karumiini Escalera, 1913
Family Rhipiceridae Latreille, 1834
Superfamily Buprestoidea Leach, 1815
Family Schizopodidae LeConte, 1859
Subfamily Schizopodinae LeConte, 1859
Tribe Dystaxiini Théry, 1929
†Tribe Electrapatini Iablokoff-Khnzorian, 1962
Tribe Schizopodini LeConte, 1859
Family Buprestidae Leach, 1815
Subfamily Julodinae Lacordaire, 1857
Subfamily Polycestinae Lacordaire, 1857
Tribe Acmaeoderini Kerremans, 1893
Subtribe Acmaeoderina Kerremans, 1893
Subtribe Acmaeoderoidina Cobos, 1955
Subtribe Nothomorphina Cobos, 1955
Tribe Astraeini Cobos, 1980
Tribe Bulini Bellamy, 1995
Tribe Haplostethini LeConte, 1861
Tribe Paratracheini Cobos, 1980
Tribe Perucolini Cobos, 1980
Tribe Polycestini Lacordaire, 1857
Subtribe Polycestina Lacordaire, 1857
Subtribe Xenopseina Volkovitsh, 2008
Tribe Polyctesini Cobos, 1955
Tribe Prospherini Cobos, 1980
Tribe Ptosimini Kerremans, 1903
Tribe Thrincopygini LeConte, 1861
Tribe Tyndaridini Cobos, 1955
Subtribe Mimicoclytrinina Bellamy, 2003
Subtribe Pseudacherusiina Cobos, 1980
Subtribe Tylaucheniina Cobos, 1959
Subtribe Tyndaridina Cobos, 1955
Tribe Xyroscelidini Cobos, 1955
Subfamily Galbellinae Reitter, 1911
Subfamily Chrysochroinae Laporte, 1835
Tribe Chrysochroini Laporte, 1835
Subtribe Chalcophorina Lacordaire, 1857 (1848)
Subtribe Chrysochroina Laporte, 1835
Subtribe Eucallopistina Bellamy, 2003
Tribe Dicercini Gistel, 1848
Subtribe Dicercina Gistel, 1848
Subtribe Haplotrinchina Holyński, 1993
Subtribe Hippomelanina Holyński, 1993
Subtribe Pseudoperotina Tôyama, 1987
Tribe Evidini Tôyama, 1987
Tribe Paraleptodemini Cobos, 1975
Subtribe Euchromatina Holyński, 1993
Subtribe Euplectaleciina Holyński, 1993
Subtribe Hypoprasina Holyński, 1993
Subtribe Paraleptodemina Cobos, 1975
Subtribe Pristipterina Holyński, 1993
Tribe Paratassini Bílý and Volkovitsh, 1996
Tribe Poecilonotini Jakobson, 1913
Subtribe Poecilonotina Jakobson, 1913
Subtribe Nesotrinchina Bílý, Kubáň and Volkovitsh, 2009
Tribe Sphenopterini Lacordaire, 1857
Tribe Vadonaxiini Descarpentries, 1970
Subfamily Buprestinae Leach, 1815
Tribe Actenodini Gistel, 1848
Tribe Anthaxiini Gory and Laporte, 1839
Tribe Bubastini Obenberger, 1920
Tribe Buprestini Leach, 1815
Subtribe Agaeocerina Nelson, 1982
Subtribe Buprestina Leach, 1815
Subtribe Lamprocheilina Holyński, 1993
Subtribe Trachykelina Holyński, 1988
Tribe Chrysobothrini Gory and Laporte, 1836
Tribe Coomaniellini Bílý, 1974
Tribe Curidini Holyński, 1988
Subtribe Anilarina Bílý, 2000
Subtribe Curidina Holyński, 1988
Subtribe Neocuridina Holyński, 1988
Tribe Epistomentini Levey, 1978
Tribe Exagistini Tôyama, 1987
Tribe Julodimorphini Kerremans, 1903
Tribe Kisanthobiini Richter, 1949
Tribe Maoraxiini Holyński, 1984
Tribe Melanophilini Bedel, 1921
Tribe Melobaseini Bílý, 2000
Tribe Mendizabaliini Cobos, 1968
Tribe Nascionini Holyński, 1988
Tribe Phrixiini Cobos, 1975
Tribe Pterobothrini Volkovitsh, 2001
Tribe Stigmoderini Lacordaire, 1857
Tribe Thomassetiini Bellamy, 1987
Subtribe Philanthaxiina Holyński, 1988
Subtribe Thomassetiina Bellamy, 1987
Tribe Trigonogeniini Cobos, 1956
Tribe Xenorhipidini Cobos, 1986
Subtribe Trichinorhipidina Bellamy, 2006
Subtribe Xenorhipidina Cobos, 1986
Subfamily Agrilinae Laporte, 1835
Tribe Agrilini Laporte, 1835
Subtribe Agrilina Laporte, 1835
Subtribe Amorphosternina Cobos, 1974
Subtribe Amyiina Holyński, 1993
Subtribe Rhaeboscelidina Cobos, 1976
Tribe Aphanisticini Jacquelin du Val, 1859
Subtribe Anthaxomorphina Holyński, 1993
Subtribe Aphanisticina Jacquelin du Val, 1859
Subtribe Cylindromorphina Portevin, 1931
Subtribe Cylindromorphoidina Cobos, 1979
Subtribe Germaricina Cobos, 1979
Tribe Coraebini Bedel, 1921
Subtribe Amorphosomatina Majer, 2000
Subtribe Cisseina Majer, 2000
Subtribe Clematina Majer, 2000
Subtribe Coraebina Bedel, 1921
Subtribe Dismorphina Cobos, 1990
Subtribe Ethoniina Majer, 2000
Subtribe Geraliina Cobos, 1988
Subtribe Meliboeina Majer, 2000
Subtribe Synechocerina Majer, 2000
Subtribe Toxoscelina Majer, 2000
Tribe Tracheini Laporte, 1835
Subtribe Brachina LeConte, 1861
Subtribe Leiopleurina Holyński, 1993
Subtribe Pachyschelina Böving and Craighead, 1931
Subtribe Tracheina Laporte, 1835
†Subfamily Parathyreinae Alexeev, 1994
Superfamily Byrrhoidea Latreille, 1804
Family Byrrhidae Latreille, 1804
Subfamily Byrrhinae Latreille, 1804
Tribe Byrrhini Latreille, 1804
Tribe Exomellini Casey, 1914
Tribe Morychini El Moursy, 1961
Tribe Pedilophorini Casey, 1912
Tribe Simplocariini Mulsant and Rey, 1869
Subfamily Syncalyptinae Mulsant and Rey, 1869
Tribe Microchaetini Paulus, 1973
Tribe Syncalyptini Mulsant and Rey, 1869
Subfamily Amphicyrtinae LeConte, 1861
Family Elmidae Curtis, 1830
Subfamily Larainae LeConte, 1861
Tribe Laraini LeConte, 1861
Tribe Potamophilini Mulsant and Rey, 1872
Subfamily Elminae Curtis, 1830
Tribe Ancyronychini Ganglbauer, 1904
Tribe Elmini Curtis, 1830
Subtribe Elmina Curtis, 1830
Subtribe Stenelmina Mulsant and Rey, 1872
Tribe Macronychini Gistel, 1848
Family Dryopidae Billberg, 1820 (1817)
Family Lutrochidae Kasap and Crowson, 1975
Family Limnichidae Erichson, 1846
Subfamily Hyphalinae Britton, 1971
Subfamily Limnichinae Erichson, 1846
Tribe Limnichini Erichson, 1846
Tribe Wooldridgeini Spangler, 1999
Subfamily Cephalobyrrhinae Champion, 1925
Subfamily Thaumastodinae Champion, 1924
Family Heteroceridae MacLeay, 1825
Subfamily Elythomerinae Pacheco, 1964
Subfamily Heterocerinae MacLeay, 1825
Tribe Augylini Pacheco, 1964
Tribe Heterocerini MacLeay, 1825
Tribe Micilini Pacheco, 1964
Tribe Tropicini Pacheco, 1964
Family Psephenidae Lacordaire, 1854
Subfamily Afroeubriinae Lee, Satô, Shepard and Jäch, 2007
Subfamily Eubriinae Lacordaire, 1857
Subfamily Eubrianacinae Jakobson, 1913
Subfamily Psephenoidinae Bollow, 1938
Subfamily Psepheninae Lacordaire, 1854
Family Cneoglossidae Champion, 1897
Family Ptilodactylidae Laporte, 1836
Subfamily Anchytarsinae Champion, 1897
Subfamily Cladotominae Pic, 1914
Subfamily Aploglossinae Champion, 1897
Subfamily Araeopidiinae Lawrence, 1991
Subfamily Ptilodactylinae Laporte, 1836
Family Podabrocephalidae Pic, 1930
Family Chelonariidae Blanchard, 1845
Family Eulichadidae Crowson, 1973
Family Callirhipidae Emden, 1924
Superfamily Elateroidea Leach, 1815
Family Rhinorhipidae Lawrence, 1988
Family Artematopodidae Lacordaire, 1857
Subfamily Electribiinae Crowson, 1975
Subfamily Allopogoniinae Crowson, 1973
Subfamily Artematopodinae Lacordaire, 1857
Tribe Artematopodini Lacordaire, 1857
Tribe Ctesibiini Crowson, 1973
Tribe Macropogonini LeConte, 1861
Family Brachypsectridae LeConte and Horn, 1883
Family Cerophytidae Latreille, 1834
Family Eucnemidae Eschscholtz, 1829
Subfamily Perothopinae Lacordaire, 1857
Subfamily Phyllocerinae Reitter, 1905
Tribe Anelastini Reitter, 1911
Tribe Phyllocerini Reitter, 1905
Subfamily Pseudomeninae Muona, 1993
Tribe Pseudomenini Muona, 1993
Tribe Schizophilini Muona, 1993
Subfamily Palaeoxeninae Muona, 1993
Subfamily Phlegoninae Muona, 1993
Subfamily Anischiinae Fleutiaux, 1936
Subfamily Melasinae Fleming, 1821
Tribe Calyptocerini Muona, 1993
Tribe Ceballosmelasini Muona, 1993
Tribe Dirhagini Reitter, 1911
Tribe Epiphanini Muona, 1993
Tribe Hylocharini Jacquelin du Val, 1859
Tribe Melasini Fleming, 1821
Subtribe Compsocnemina Muona, 1993
Subtribe Melasina Fleming, 1821
Tribe Neocharini Muona, 1993
Tribe Xylobiini Reitter, 1911
Subfamily Eucneminae Eschscholtz, 1829
Tribe Dendrocharini Fleutiaux, 1920
Tribe Dyscharachthini Muona, 1993
Tribe Entomosatopini Muona, 1993
Tribe Eucnemini Eschscholtz, 1829
Tribe Galbitini Muona, 1991
Tribe Mesogenini Muona, 1993
Tribe Muonajini Özdikmen, 2008
Tribe Perrotiini Muona, 1993
Tribe Phaenocerini Muona, 1993
Tribe Proutianini Muona, 1993
Subfamily Macraulacinae Fleutiaux, 1923
Tribe Anelastidini Muona, 1993
Tribe Echthrogasterini Cobos, 1965
Tribe Euryptychini Mamaev, 1976
Tribe Jenibuntorini Muona, 1993
Tribe Macraulacini Fleutiaux, 1923
Tribe Nematodini Leiler, 1976
Tribe Oisocerini Muona, 1993
Tribe Orodotini Muona, 1993
†Tribe Throscogeniini Iablokoff-Khnzorian, 1962
Family Throscidae Laporte, 1840 nomen protectum
†Family Praelateriidae Dolin, 1973
Family Elateridae Leach, 1815
Subfamily Cebrioninae Latreille, 1802
Subfamily Agrypninae Candèze, 1857 nomen protectum
Tribe Agrypnini Candèze, 1857 nomen protectum
Tribe Anaissini Golbach, 1984
†Tribe Cryptocardiini Dolin, 1980
Tribe Euplinthini Costa, 1975
Subtribe Cleidecostina Johnson, 2002
Subtribe Compsoplinthina Costa, 1975
Subtribe Euplinthina Costa, 1975
Tribe Hemirhipini Candèze, 1857
Tribe Oophorini Gistel, 1848
Tribe Platycrepidiini Costa and Casari-Chen, 1993
Tribe Pseudomelanactini Arnett, 1967
Tribe Pyrophorini Candèze, 1863
Subtribe Hapsodrilina Costa, 1975
Subtribe Nyctophyxina Costa, 1975
Subtribe Pyrophorina Candèze, 1863
Tribe Tetralobini Laporte, 1840
Subfamily Thylacosterninae Fleutiaux, 1920
Subfamily Lissominae Laporte, 1835
Subfamily Semiotinae Jakobson, 1913
Subfamily Campyloxeninae Costa, 1975
Subfamily Pityobiinae Hyslop, 1917
Subfamily Oxynopterinae Candèze, 1857
Subfamily Dendrometrinae Gistel, 1848
Tribe Crepidomenini Candèze, 1863
Tribe Dendrometrini Gistel, 1848
Subtribe Dendrometrina Gistel, 1848
Subtribe Denticollina Stein and Weise, 1877 (1848)
Subtribe Hemicrepidiina Champion, 1896
Tribe Dimini Candèze, 1863
Tribe Hypnoidini Schwarz, 1906 (1860)
Tribe Pleonomini Semenov and Pjatakova, 1936
Tribe Prosternini Gistel, 1856 nomen protectum
Tribe Senodoniini Schenkling, 1927
Subfamily Negastriinae Nakane and Kishii, 1956
Tribe Negastriini Nakane and Kishii, 1956
Tribe Quasimusini Schimmel and Tarnawski, 2009
Subtribe Loebliquasimusina Schimmel and Tarnawski, 2009
Subtribe Quasimusina Schimmel and Tarnawski, 2009
Subtribe Striatoquasimusina Schimmel and Tarnawski, 2009
Subtribe Wittmeroquasimusina Schimmel and Tarnawski, 2009
Subfamily Elaterinae Leach, 1815
Tribe Agriotini Laporte, 1840
Subtribe Agriotina Laporte, 1840
Subtribe Cardiorhinina Candèze, 1863
Tribe Ampedini Gistel, 1848
Tribe Dicrepidiini Thomson, 1858
Tribe Elaterini Leach, 1815
Tribe Megapenthini Gurjeva, 1973
Tribe Melanotini Candèze, 1859 (1848)
Tribe Odontonychini Girard, 1973
Tribe Physorhinini Candèze, 1859
Tribe Pomachiliini Candèze, 1859
Tribe Synaptini Gistel, 1856
Subfamily Cardiophorinae Candèze, 1859
Subfamily Hemiopinae Fleutiaux, 1941
Subfamily Physodactylinae Lacordaire, 1857
Subfamily Eudicronychinae Girard, 1971
Subfamily Subprotelaterinae Fleutiaux, 1920
Subfamily Morostomatinae Dolin, 2000
†Subfamily Protagrypninae Dolin, 1973
†Tribe Desmatini Dolin, 1975
†Tribe Hypnomorphini Dolin, 1975
†Tribe Protagrypnini Dolin, 1973
Family Plastoceridae Crowson, 1972
Family Drilidae Blanchard, 1845
Subfamily Drilinae Blanchard, 1845
Subfamily Thilmaninae Kazantsev, 2004
Tribe Euanomini Kazantsev, 2010
Tribe Thilmanini Kazantsev, 2004
Family Omalisidae Lacordaire, 1857
†Family Berendtimiridae Winkler, 1987
Family Lycidae Laporte, 1836
Subfamily Libnetinae Bocák and Bocáková, 1990
Subfamily Dictyopterinae Houlbert, 1922
Tribe Dictyopterini Houlbert, 1922
Tribe Lycoprogenthini Bocák and Bocáková, 2008
Tribe Taphini Bocák and Bocáková, 1990
Subfamily Lyropaeinae Bocák and Bocáková, 1989
Tribe Alyculini Bocák and Bocáková, 2008
Tribe Antennolycini Bocák and Bocáková, 2008
Tribe Lyropaeini Bocák and Bocáková, 1989
Tribe Miniduliticolini Kazantsev, 2003
Tribe Platerodrilini Kazantsev, 2004
Subfamily Ateliinae Kleine, 1929
Tribe Ateliini Kleine, 1929
Tribe Dilophotini Kleine, 1929
Subfamily Lycinae Laporte, 1836
Tribe Calochromini Lacordaire, 1857
Tribe Calopterini Green, 1949
Subtribe Acroleptina Bocáková, 2005
Subtribe Calopterina Green, 1949
Tribe Conderini Bocák and Bocáková, 1990
Tribe Dihammatini Bocák and Bocáková, 2008
Tribe Erotini LeConte, 1881
Tribe Eurrhacini Bocáková, 2005
Tribe Leptolycini Leng and Mutchler, 1922
Tribe Lycini Laporte, 1836
Tribe Lyponiini Bocák and Bocáková, 1990
Tribe Macrolycini Kleine, 1929
Tribe Melanerotini Kazantsev, 2010
Tribe Metriorrhynchini Kleine, 1926
Subtribe Hemiconderinina Bocák and Bocáková, 1990
Subtribe Metriorrhynchina Kleine, 1926
Subtribe Trichalina Kleine, 1929
Tribe Platerodini Kleine, 1929
Tribe Slipinskiini Bocák and Bocáková, 1992
Tribe Thonalmini Kleine, 1933
Subfamily Dexorinae Bocák and Bocáková, 1989
Family Telegeusidae Leng, 1920
Family Phengodidae LeConte, 1861
Subfamily Phengodinae LeConte, 1861
Subfamily Mastinocerinae LeConte, 1881
Subfamily Penicillophorinae Paulus, 1975
Family Rhagophthalmidae Olivier, 1907
Family Lampyridae Rafinesque, 1815
Subfamily Psilocladinae McDermott, 1964
Subfamily Amydetinae Olivier, 1907
Tribe Amydetini Olivier, 1907
Tribe Vestini McDermott, 1964
Subfamily Lampyrinae Rafinesque, 1815
Tribe Cratomorphini Green, 1948
Tribe Lamprocerini Olivier, 1907
Tribe Lamprohizini Kazantsev, 2010
Tribe Lampyrini Rafinesque, 1815
Tribe Lucidotini Lacordaire, 1857
Subtribe Dadophorina Olivier, 1907
Subtribe Lamprigerina McDermott, 1964
Subtribe Lucidotina Lacordaire, 1857
Subtribe Photinina LeConte, 1881
Tribe Pleotomini Summers, 1874
Subfamily Luciolinae Lacordaire, 1857
Tribe Curtosini McDermott, 1964
Tribe Luciolini Lacordaire, 1857
Subfamily Photurinae Lacordaire, 1857
Family Omethidae LeConte, 1861
Subfamily Omethinae LeConte, 1861
Subfamily Matheteinae LeConte, 1881
Subfamily Driloniinae Crowson, 1972
Family Cantharidae Imhoff, 1856 (1815)
Subfamily Cantharinae Imhoff, 1856 (1815)
Tribe Cantharini Imhoff, 1856 (1815)
Tribe Podabrini Gistel, 1856
Subfamily Silinae Mulsant, 1862
Tribe Silini Mulsant, 1862
Tribe Tytthonyxini Arnett, 1962
Subfamily Dysmorphocerinae Brancucci, 1980
Subfamily Malthininae Kiesenwetter, 1852
Tribe Malchinini Brancucci, 1980
Tribe Malthinini Kiesenwetter, 1852
Tribe Malthodini Böving and Craighead, 1931
Subfamily Chauliognathinae LeConte, 1861
Tribe Chauliognathini LeConte, 1861
Tribe Ichthyurini Champion, 1915
Subfamily Cydistinae Paulus, 1972
Subfamily Pterotinae LeConte, 1861
Subfamily Ototretinae McDermott, 1964
Subfamily Ototretadrilinae Crowson, 1972
†Subfamily Lasiosynidae Kirejtshuk, Chang, Ren and Kun, 2010
Series DERODONTIFORMIA
Superfamily Derodontoidea LeConte, 1861
Family Derodontidae LeConte, 1861
Subfamily Peltasticinae LeConte, 1861
Subfamily Derodontinae LeConte, 1861
Subfamily Laricobiinae Mulsant and Rey, 1864
Family Nosodendridae Erichson, 1846
Family Jacobsoniidae Heller, 1926
Series BOSTRICHIFORMIA
Superfamily Bostrichoidea Latreille, 1802
Family Dermestidae Latreille, 1804
Subfamily Dermestinae Latreille, 1804
Tribe Dermestini Latreille, 1804
Tribe Marioutini Jakobson, 1913
Subfamily Thorictinae Agassiz, 1846
Tribe Thaumaphrastini Anderson, 1949
Tribe Thorictini Agassiz, 1846
Subfamily Orphilinae LeConte, 1861
Subfamily Trinodinae Casey, 1900
†Tribe Cretonodini Kirejtshuk and Azar, 2009
Tribe Thylodriini Semenov, 1909
Tribe Trinodini Casey, 1900
Tribe Trinoparvini Háva, 2010
Subfamily Attageninae Laporte, 1840
Tribe Attagenini Laporte, 1840
Tribe Egidyellini Semenov, 1914
Subfamily Megatominae Leach, 1815
Tribe Anthrenini Gistel, 1848
Tribe Megatomini Leach, 1815
Family Endecatomidae LeConte, 1861
Family Bostrichidae Latreille, 1802
Subfamily Dysidinae Lesne, 1921
Subfamily Polycaoninae Lesne, 1896
Subfamily Bostrichinae Latreille, 1802
Tribe Apatini Billberg, 1820
Tribe Bostrichini Latreille, 1802
Tribe Dinapatini Lesne, 1910
Tribe Sinoxylini Marseul, 1857
Tribe Xyloperthini Lesne, 1921
Subfamily Psoinae Blanchard, 1851
Subfamily Dinoderinae Thomson, 1863
Subfamily Lyctinae Billberg, 1820
Tribe Lyctini Billberg, 1820
Tribe Trogoxylini Lesne, 1921
Subfamily Euderiinae Lesne, 1934
Family Ptinidae Latreille, 1802
Subfamily Eucradinae LeConte, 1861
Tribe Eucradini LeConte, 1861
Tribe Hedobiini Mulsant and Rey, 1868
Subfamily Ptininae Latreille, 1802
Tribe Gibbiini Jacquelin du Val, 1860
Tribe Meziini Bellés, 1985
Tribe Ptinini Latreille, 1802
Tribe Sphaericini Portevin, 1931
Subfamily Dryophilinae Gistel, 1848
Tribe Dryophilini Gistel, 1848
Tribe Ptilineurini Böving, 1927
Subfamily Ernobiinae Pic, 1912
Subfamily Anobiinae Fleming, 1821
Subfamily Ptilininae Shuckard, 1839
Subfamily Alvarenganiellinae Viana and Martínez, 1971
Subfamily Xyletininae Gistel, 1848
Tribe Lasiodermini Böving, 1927
Tribe Metholcini Zahradník, 2009
Tribe Xyletinini Gistel, 1848
Subfamily Dorcatominae Thomson, 1859
Subfamily Mesocoelopodinae Mulsant and Rey, 1864
Tribe Tricorynini White, 1971
Tribe Mesocoelopodini Mulsant and Rey, 1864
Series CUCUJIFORMIA
Superfamily Lymexyloidea Fleming, 1821
Family Lymexylidae Fleming, 1821
Subfamily Hylecoetinae Germar, 1818
Subfamily Lymexylinae Fleming, 1821
Subfamily Atractocerinae Laporte, 1840
Subfamily Melittommatinae Wheeler, 1986
Superfamily Cleroidea Latreille, 1802
Family Phloiophilidae Kiesenwetter, 1863
Family Trogossitidae Latreille, 1802
Subfamily Peltinae Latreille, 1806
Tribe Ancyronini Kolibáč, 2006
Tribe Colydiopeltini Kolibáč, 2006
Tribe Decamerini Crowson, 1964
Tribe Lophocaterini Crowson, 1964 nomen protectum
Tribe Peltini Latreille, 1806
Tribe Thymalini Léveillé, 1888
Subfamily Trogossitinae Latreille, 1802
Tribe Calityini Reitter, 1922
Tribe Egoliini Lacordaire, 1854
Tribe Gymnochilini Lacordaire, 1854
Tribe Larinotini Ślipiński, 1992
†Tribe Lithostomatini Kolibáč and Huang, 2008
Tribe Trogossitini Latreille, 1802
Family Chaetosomatidae Crowson, 1952
Family Metaxinidae Kolibáč, 2004
Family Thanerocleridae Chapin, 1924
Subfamily Zenodosinae Kolibáč, 1992
Subfamily Thaneroclerinae Chapin, 1924
Tribe Isoclerini Kolibáč, 1992
Tribe Thaneroclerini Chapin, 1924
Tribe Viticlerini Winkler, 1982
Family Cleridae Latreille, 1802
Subfamily Tillinae Fischer von Waldheim, 1813
Subfamily Hydnocerinae Spinola, 1844
Tribe Callimerini Kolibáč, 1998
Tribe Hydnocerini Spinola, 1844
Tribe Lemidiini Kolibáč, 1998
Subfamily Clerinae Latreille, 1802
Subfamily Korynetinae Laporte, 1836
Family Acanthocnemidae Crowson, 1964
Family Phycosecidae Crowson, 1952
Family Prionoceridae Lacordaire, 1857
Tribe Lobonychini Majer, 1987
Tribe Prionocerini Lacordaire, 1857
Family Mauroniscidae Majer, 1995
Family Melyridae Leach, 1815
Subfamily Rhadalinae LeConte, 1861
Subfamily Melyrinae Leach, 1815
Tribe Arthrobrachini Majer, 1987
Tribe Astylini Pic, 1929
Tribe Cerallini Pic, 1929
Tribe Melyrini Leach, 1815
Subfamily Dasytinae Laporte, 1840
Tribe Chaetomalachiini Majer, 1987
Tribe Danaceini Thomson, 1859
Tribe Dasytini Laporte, 1840
Tribe Gietellini Constantin and Menier, 1987
Tribe Listrini Majer, 1990
Subfamily Malachiinae Fleming, 1821
Tribe Amalthocini Majer, 2002
Tribe Attalomimini Majer, 1995
Tribe Carphurini Champion, 1923
Tribe Lemphini Wittmer, 1976
Tribe Malachiini Fleming, 1821
Tribe Pagurodactylini Constantin, 2001
Superfamily Cucujoidea Latreille, 1802
†Family Parandrexidae Kirejtshuk, 1994
†Family Sinisilvanidae Hong, 2002
Family Boganiidae Sen Gupta and Crowson, 1966
Subfamily Paracucujinae Endrödy-Younga and Crowson, 1986
Subfamily Boganiinae Sen Gupta and Crowson, 1966
Family Byturidae Gistel, 1848
Subfamily Platydascillinae Pic, 1914
Subfamily Byturinae Gistel, 1848
Family Helotidae Chapuis, 1876
Family Protocucujidae Crowson, 1954
Family Sphindidae Jacquelin du Val, 1860
Subfamily Protosphindinae Sen Gupta and Crowson, 1979
Subfamily Odontosphindinae Sen Gupta and Crowson, 1979
Subfamily Sphindiphorinae Sen Gupta and Crowson, 1979
Subfamily Sphindinae Jacquelin du Val, 1860
Family Biphyllidae LeConte, 1861
Family Erotylidae Latreille, 1802
Subfamily Xenoscelinae Ganglbauer, 1899
Subfamily Pharaxonothinae Crowson, 1952
Subfamily Loberinae Bruce, 1951
Subfamily Languriinae Hope, 1840
Tribe Hapalipini Leschen, 2003
Tribe Languriini Hope, 1840
Tribe Thallisellini Sen Gupta, 1968
Subfamily Cryptophilinae Casey, 1900
Tribe Cryptophilini Casey, 1900
Tribe Empocryptini Leschen, 2003
Tribe Toramini Sen Gupta, 1967
Subfamily Erotylinae Latreille, 1802
Tribe Dacnini Gistel, 1848
Tribe Encaustini Crotch, 1876
Tribe Erotylini Latreille, 1802
Tribe Megalodacnini Sen Gupta, 1970
Tribe Tritomini Curtis, 1834
Family Monotomidae Laporte, 1840
Subfamily Rhizophaginae Redtenbacher, 1845
Subfamily Monotominae Laporte, 1840
Tribe Europini Sen Gupta, 1988
Tribe Lenacini Crowson, 1952
Tribe Monotomini Laporte, 1840
†Tribe Rhizophtomini Kirejtshuk and Azar, 2009
Tribe Thionini Crowson, 1952
Family Hobartiidae Sen Gupta and Crowson, 1966
Family Cryptophagidae Kirby, 1826
Subfamily Cryptophaginae Kirby, 1826
Tribe Caenoscelini Casey, 1900
Tribe Cryptophagini Kirby, 1826
Tribe Picrotini Crowson, 1980
Subfamily Atomariinae LeConte, 1861
Tribe Atomariini LeConte, 1861
Tribe Cryptafricini Leschen, 1996
Tribe Hypocoprini Reitter, 1879
Family Agapythidae Sen Gupta and Crowson, 1969
Family Priasilphidae Crowson, 1973
Family Phloeostichidae Reitter, 1911
Family Silvanidae Kirby, 1837
Subfamily Brontinae Blanchard, 1845
Tribe Brontini Blanchard, 1845
Tribe Telephanini LeConte, 1861
Subfamily Silvaninae Kirby, 1837
Family Cucujidae Latreille, 1802
Family Myraboliidae Lawrence and Britton, 1991
Family Cavognathidae Sen Gupta and Crowson, 1966
Family Lamingtoniidae Sen Gupta and Crowson, 1969
Family Passandridae Blanchard, 1845
Family Phalacridae Leach, 1815
Subfamily Phaenocephalinae Matthews, 1899
Subfamily Phalacrinae Leach, 1815
Family Propalticidae Crowson, 1952
Family Laemophloeidae Ganglbauer, 1899
Family Tasmosalpingidae Lawrence and Britton, 1991
Family Cyclaxyridae Gimmel, Leschen and Ślipiński, 2009
Family Kateretida Kirby, 1837
Family Nitidulidae Latreille, 1802
Subfamily Calonecrinae Kirejtshuk, 1982
Subfamily Maynipeplinae Kirejtshuk, 1998
Subfamily Epuraeinae Kirejtshuk, 1986
Tribe Epuraeini Kirejtshuk, 1986
Tribe Taenioncini Kirejtshuk, 1998
Subfamily Carpophilinae Erichson, 1842
Subfamily Amphicrossinae Kirejtshuk, 1986
Subfamily Meligethinae Thomson, 1859
Subfamily Nitidulinae Latreille, 1802
Tribe Cychramini Gistel, 1848
Tribe Cychramptodini Kirejtshuk and Lawrence, 1992
Tribe Cyllodini Everts, 1898
Tribe Lawrencerosini Kirejtshuk, 1991
Tribe Mystropini Murray, 1864
Tribe Nitidulini Latreille, 1802
Subfamily Cillaeinae Kirejtshuk and Audisio, 1986
Subfamily Cryptarchinae Thomson, 1859
Tribe Arhinini Kirejtshuk, 1987
Tribe Cryptarchini Thomson, 1859
Tribe Eucalosphaerini Kirejtshuk, 1987
Tribe Platyarchini Kirejtshuk, 1998
Subfamily Cybocephalinae Jacquelin du Val, 1858
Family Smicripidae Horn, 1880
Family Bothrideridae Erichson, 1845
Subfamily Teredinae Seidlitz, 1888
Tribe Sosylopsini Dajoz, 1980
Tribe Sysolini Ślipiński and Pal, 1985
Tribe Teredini Seidlitz, 1888
Subfamily Xylariophilinae Pal and Lawrence, 1986
Subfamily Anommatinae Ganglbauer, 1899
Subfamily Bothriderinae Erichson, 1845
Family Cerylonidae Billberg, 1820
Subfamily Euxestinae Grouvelle, 1908
Subfamily Loeblioryloninae Ślipiński, 1990
Subfamily Ostomopsinae Sen Gupta and Crowson, 1973
Subfamily Murmidiinae Jacquelin du Val, 1858
Subfamily Ceryloninae Billberg, 1820
Family Alexiidae Imhoff, 1856
Family Discolomatidae Horn, 1878
Subfamily Notiophyginae Jakobson, 1915
Tribe Dystheamonini John, 1954
Tribe Notiophygini Jakobson, 1915
Tribe Pachyplacini John, 1954
Subfamily Discolomatinae Horn, 1878
Subfamily Aphanocephalinae Jakobson, 1904
Subfamily Cephalophaninae John, 1954
Subfamily Pondonatinae John, 1954
Family Endomychidae Leach, 1815
Subfamily Merophysiinae Seidlitz, 1872
Subfamily Pleganophorinae Jacquelin du Val, 1858
Subfamily Anamorphinae Strohecker, 1953
Subfamily Leiestinae Thomson, 1863
Subfamily Mycetaeinae Jacquelin du Val, 1857
Subfamily Eupsilobiinae Casey, 1895
Subfamily Xenomycetinae Strohecker, 1962
Subfamily Danascelinae Tomaszewska, 2000
Subfamily Endomychinae Leach, 1815
Subfamily Epipocinae Gorham, 1873
Subfamily Stenotarsinae Chapuis, 1876
Subfamily Lycoperdininae Bromhead, 1838
Family Coccinellidae Latreille, 1807
Subfamily Microweiseinae Leng, 1920
Tribe Microweiseini Leng, 1920
Tribe Serangiini Pope, 1962
Tribe Sukunahikonini Kamiya, 1960
Subfamily Coccinellinae Latreille, 1807
Tribe Argentipilosini Gordon and de Almeida, 1991
Tribe Aspidimerini Mulsant, 1850
Tribe Azyini Mulsant, 1850
Tribe Brachiacanthini Mulsant, 1850
Tribe Carinodulini Gordon, Pakaluk and Ślipiński, 1989
Tribe Cephaloscymnini Gordon, 1985
Tribe Chilocorini Mulsant, 1846
Tribe Chnoodini Mulsant, 1850
Tribe Coccidulini Mulsant, 1846
Tribe Coccinellini Latreille, 1807
Tribe Cranophorini Mulsant, 1850
Tribe Cryptognathini Mulsant, 1850
Tribe Cynegetini Thomson, 1866
Tribe Diomini Gordon, 1999
Tribe Epilachnini Mulsant, 1846
Tribe Epivertini Pang and Mao, 1979
Tribe Eremochilini Gordon and Vanderberg, 1987
Tribe Hyperaspidini Mulsant, 1846
Tribe Limnichopharini Miyatake, 1994
Tribe Monocorynini Miyatake, 1988
Tribe Noviini Mulsant, 1846
Tribe Ortaliini Mulsant, 1850
Tribe Oryssomini Gordon, 1974
Tribe Platynaspini Mulsant, 1846
Tribe Plotinini Miyatake, 1994
Tribe Poriini Mulsant, 1850
Tribe Scymnillini Casey, 1899
Tribe Scymnini Mulsant, 1846
Tribe Selvadiini Gordon, 1985
Tribe Shirozuellini Sasaji, 1967
Tribe Stethorini Dobzhansky, 1924
Tribe Sticholotidini Weise, 1901
Tribe Telsimiini Casey, 1899
Tribe Tetrabrachini Kapur, 1948
Family Corylophidae LeConte, 1852
Subfamily Periptyctinae Ślipiński, Lawrence and Tomaszewska, 2001
Subfamily Corylophinae LeConte, 1852
Tribe Aenigmaticini Casey, 1900
Tribe Cleidostethini Bowestead, Booth, Ślipiński and Lawrence, 2001
Tribe Corylophini LeConte, 1852
Tribe Foadiini Ślipiński, Tomaszewska and Lawrence, 2009
Tribe Orthoperini Jacquelin du Val, 1857
Tribe Parmulini Poey, 1854
Tribe Peltinodini Paulian, 1950
Tribe Rypobiini Paulian, 1950
Tribe Sericoderini Matthews, 1888
Tribe Teplinini Pakaluk, Ślipiński and Lawrence, 1994
Family Akalyptoischiidae Lord, Hartley, Lawrence, McHugh and Miller, 2010
Family Latridiidae Erichson, 1842
Subfamily Latridiinae Erichson, 1842
Subfamily Corticariinae Curtis, 1829
†Subfamily Tetrameropseinae Kirejtshuk and Azar, 2008
Superfamily Tenebrionoidea Latreille, 1802
Family Mycetophagidae Leach, 1815
Subfamily Esarcinae Reitter, 1882
Subfamily Mycetophaginae Leach, 1815
Tribe Mycetophagini Leach, 1815
Tribe Typhaeini Thomson, 1863
Subfamily Bergininae Leng, 1920
Family Archeocrypticidae Kaszab, 1964
Family Pterogeniidae Crowson, 1953
Family Ciidae Leach, 1819
Subfamily Sphindociinae Lawrence, 1974
Subfamily Ciinae Leach, 1819
Tribe Ciini Leach, 1819
Tribe Orophiini Thomson, 1863
Tribe Xylographellini Kawanabe and Miyatake, 1996
Subtribe Syncosmetina Lopes-Andrade, 2008
Subtribe Xylographellina Kawanabe and Miyatake, 1996
Family Tetratomidae Billberg, 1820
Subfamily Tetratominae Billberg, 1820
Subfamily Piseninae Miyatake, 1960
Subfamily Penthinae Lacordaire, 1859
Subfamily Hallomeninae Gistel, 1848
Subfamily Eustrophinae Gistel, 1848
Tribe Eustrophini Gistel, 1848
Tribe Holostrophini Nikitsky, 1998
Family Melandryidae Leach, 1815
Subfamily Melandryinae Leach, 1815
Tribe Anisoxiellini Nikitsky, 2007
Tribe Dircaeini Kirby, 1837
Tribe Hypulini Gistel, 1848
Tribe Melandryini Leach, 1815
Tribe Orchesiini Mulsant, 1856
Tribe Serropalpini Latreille, 1829
Tribe Xylitini Thomson, 1864
Tribe Zilorini Desbrochers des Loges, 1900
Subfamily Osphyinae Mulsant, 1856 (1839)
Family Mordellidae Latreille, 1802
Subfamily Ctenidiinae Franciscolo, 1951
Subfamily Mordellinae Latreille, 1802
Tribe Conaliini Ermisch, 1956
Tribe Mordellini Latreille, 1802
Tribe Mordellistenini Ermisch, 1941
Tribe Reynoldsiellini Franciscolo, 1957
Tribe Stenaliini Franciscolo, 1955
Family Ripiphoridae Gemminger, 1870 (1855)
Subfamily Ptilophorinae Gerstaecker, 1855
Subfamily Pelecotominae Seidlitz, 1875
Subfamily Hemirhipidiinae Heller, 1921
Subfamily Ripidiinae Gerstaecker, 1855
Tribe Eorhipidiini Iablokoff-Khnzorian, 1986
Tribe Ripidiini Gerstaecker, 1855
Subfamily Ripiphorinae Gemminger, 1870 (1855)
Tribe Macrosiagonini Heyden, 1908
Tribe Ripiphorini Gemminger, 1870 (1855)
Family Zopheridae Solier, 1834
Subfamily Colydiinae Billberg, 1820
Tribe Acropini Sharp, 1894
Tribe Adimerini Sharp, 1894
Tribe Colydiini Billberg, 1820
Tribe Gempylodini Sharp, 1893
Tribe Nematidiini Horn, 1878
Tribe Orthocerini Blanchard, 1845
Tribe Rhagoderini Horn, 1878
Tribe Rhopalocerini Reitter, 1911
Tribe Synchitini Erichson, 1845
Subfamily Zopherinae Solier, 1834
Tribe Latometini Ślipiński and Lawrence, 1999
Tribe Monommatini Blanchard, 1845
Tribe Phellopsini Ślipiński and Lawrence, 1999
Tribe Pycnomerini Erichson, 1845
Tribe Usechini Horn, 1867
Tribe Zopherini Solier, 1834
Family Ulodidae Pascoe, 1869
Family Promecheilidae Lacordaire, 1859
Family Chalcodryidae Watt, 1974
Family Trachelostenidae Lacordaire, 1859
Family Tenebrionidae Latreille, 1802
Subfamily Zolodininae Watt, 1975
Subfamily Lagriinae Latreille, 1825 (1820)
Tribe Adeliini Kirby, 1828
Tribe Belopini Reitter, 1917
Tribe Chaerodini Doyen, Matthews and Lawrence, 1990
Tribe Cossyphini Latreille, 1802
Tribe Goniaderini Lacordaire, 1859
Tribe Laenini Seidlitz, 1895
Tribe Lagriini Latreille, 1825 (1820)
Subtribe Lagriina Latreille, 1825 (1820)
Subtribe Statirina Blanchard, 1845
Tribe Lupropini Ardoin, 1958
Tribe Pycnocerini Lacordaire, 1859 nomen protectum
Subfamily Nilioninae Oken, 1843
Subfamily Phrenapatinae Solier, 1834
Tribe Archaeoglenini Watt, 1975
Tribe Penetini Lacordaire, 1859
Tribe Phrenapatini Solier, 1834
Subfamily Pimeliinae Latreille, 1802
Tribe Adelostomini Solier, 1834
Tribe Adesmiini Lacordaire, 1859 nomen protectum
Tribe Akidini Billberg, 1820
Tribe Anepsiini LeConte, 1862
Tribe Asidini Fleming, 1821
Tribe Boromorphini Skopin, 1978
Tribe Branchini LeConte, 1862
Tribe Caenocrypticini Koch, 1958
Tribe Ceratanisini Gebien, 1937
Tribe Cnemeplatiini Jacquelin du Val, 1861
Subtribe Actizetina Watt, 1992
Subtribe Cnemeplatiina Jacquelin du Val, 1861
Subtribe Rondoniellina Ferrer and Moragues, 2000
Subtribe Thorictosomatina Watt, 1992
Tribe Cnemodinini Gebien, 1910
Tribe Coniontini Waterhouse, 1858
Tribe Cossyphodini Wasmann, 1899
Subtribe Cossyphodina Wasmann, 1899
Subtribe Cossyphoditina Basilewsky, 1950
Subtribe Esemephina Steiner, 1980
Subtribe Paramellonina Andreae, 1961
Tribe Cryptochilini Solier, 1841
Subtribe Calognathina Lacordaire, 1859
Subtribe Cryptochilina Solier, 1841
Subtribe Homebiina Endrödy-Younga, 1989
Subtribe Horatomina Koch, 1955
Subtribe Vansoniina Koch, 1955
Tribe Cryptoglossini LeConte, 1862 nomen protectum
Tribe Edrotini Lacordaire, 1859
Tribe Elenophorini Solier, 1837
Tribe Epitragini Blanchard, 1845 nomen protectum
Tribe Erodiini Billberg, 1820 nomen protectum
Tribe Evaniosomini Lacordaire, 1859
Tribe Falsomycterini Gebien, 1910
Tribe Idisiini Medvedev, 1973
Tribe Klewariini Gebien, 1910
Tribe Kuhitangiini Medvedev, 1962
Tribe Lachnogyini Seidlitz, 1894
Subtribe Lachnodactylina Reitter, 1904
Subtribe Lachnogyina Seidlitz, 1894
Subtribe Netuschiliina Ferrer and Yvinec, 2004
Tribe Leptodini Lacordaire, 1859
Tribe Nycteliini Solier, 1834
Tribe Nyctoporini Lacordaire, 1859
Tribe Phrynocarenini Gebien, 1928
Tribe Physogasterini Lacordaire, 1859
Tribe Pimeliini Latreille, 1802
Tribe Praociini Eschscholtz, 1829
Tribe Sepidiini Eschscholtz, 1829
Subtribe Hypomelina Koch, 1955
Subtribe Molurina Solier, 1834
Subtribe Oxurina Koch, 1955
Subtribe Phanerotomeina Koch, 1958
Subtribe Sepidiina Eschscholtz, 1829
Subtribe Trachynotina Koch, 1955
Tribe Stenosini Schaum, 1859 (1834)
Tribe Tentyriini Eschscholtz, 1831
Tribe Thinobatini Lacordaire, 1859
Tribe Trilobocarini Lacordaire, 1859
Tribe Vacronini Gebien, 1910
Tribe Zophosini Solier, 1834
Subfamily Tenebrioninae Latreille, 1802
Tribe Acropteronini Doyen, 1989
Tribe Alphitobiini Reitter, 1917
Tribe Amarygmini Gistel, 1848
Tribe Amphidorini LeConte, 1862
Tribe Apocryphini Lacordaire, 1859
Tribe Blaptini Leach, 1815
Subtribe Blaptina Leach, 1815
Subtribe Gnaptorina Medvedev, 2001
Subtribe Gnaptorinina Medvedev, 2001
Subtribe Prosodina Skopin, 1960
Subtribe Remipedellina Semenov, 1907
Tribe Bolitophagini Kirby, 1837 nomen protectum
Tribe Centronopini Doyen, 1989
Tribe Cerenopini Horn, 1870
Tribe Dissonomini Medvedev, 1968
Tribe Eulabini Horn, 1870
Tribe Falsocossyphini Ferrer, 2006
Tribe Heleini Fleming, 1821
Subtribe Asphalina Matthews and Lawrence, 2005
Subtribe Cyphaleina Lacordaire, 1859
Subtribe Heleina Fleming, 1821
Tribe Helopini Latreille, 1802
Subtribe Helopina Latreille, 1802
Subtribe Cylindrinotina Español, 1956
Tribe Helopinini Lacordaire, 1859
Subtribe Aptilina Koch, 1958
Subtribe Helopinina Lacordaire, 1859
Subtribe Micrantereina Reitter, 1917
Subtribe Oncosomina Koch, 1958
Tribe Melanimonini Seidlitz, 1894 (1854)
Tribe Opatrini Brullé, 1832
Subtribe Heterocheirina Koch, 1956
Subtribe Heterotarsina Blanchard, 1845
Subtribe Opatrina Brullé, 1832
Subtribe Neopachypterina Bouchard, Löbl and Merkl, 2007
Tribe Palorini Matthews, 2003
Tribe Pedinini Eschscholtz, 1829
Subtribe Dendarina Mulsant and Rey, 1854
Subtribe Eurynotina Mulsant and Rey, 1854
Subtribe Leichenina Mulsant, 1854
Subtribe Loensina Koch, 1956
Subtribe Melambiina Mulsant and Rey, 1854
Subtribe Pedinina Eschscholtz, 1829
Subtribe Platynotina Mulsant and Rey, 1853
Subtribe Pythiopina Koch, 1953
Tribe Platyscelidini Lacordaire, 1859
Tribe Praeugenini De Moor, 1970
Tribe Rhysopaussini Wasmann, 1896
Tribe Scaurini Billberg, 1820
Tribe Scotobiini Solier, 1838
Tribe Tenebrionini Latreille, 1802
Tribe Titaenini Fauvel, 1905
Tribe Toxicini Oken, 1843
Subtribe Eudysantina Bouchard, Lawrence, Davies and Newton, 2005
Subtribe Nycteropina Lacordaire, 1859
Subtribe Toxicina Oken, 1843
Tribe Triboliini Gistel, 1848
Tribe Ulomini Blanchard, 1845
Subfamily Alleculinae Laporte, 1840
Tribe Alleculini Laporte, 1840
Subtribe Alleculina Laporte, 1840
Subtribe Gonoderina Seidlitz, 1896
Subtribe Mycetocharina Gistel, 1848
Subtribe Xystropodina Solier, 1835
Tribe Cteniopodini Solier, 1835
Subfamily Diaperinae Latreille, 1802
Tribe Crypticini Brullé, 1832
Tribe Diaperini Latreille, 1802
Subtribe Adelinina LeConte, 1862
Subtribe Diaperina Latreille, 1802
Tribe Ectychini Doyen, Matthews and Lawrence, 1990
Tribe Gnathidiini Gebien, 1921
Subtribe Anopidiina Jeannel and Paulian, 1945
Subtribe Gnathidiina Gebien, 1921
Tribe Hyociini Medvedev and Lawrence, 1982
Subtribe Brittonina Medvedev and Lawrence, 1986
Subtribe Hyociina Medvedev and Lawrence, 1982
Subtribe Uptonina Medvedev and Lawrence, 1986
Tribe Hypophlaeini Billberg, 1820
Tribe Leiochrinini Lewis, 1894
Tribe Myrmechixenini Jacquelin du Val, 1858
Tribe Phaleriini Blanchard, 1845
Tribe Scaphidemini Reitter, 1922
Tribe Trachyscelini Blanchard, 1845
Subfamily Stenochiinae Kirby, 1837
Tribe Cnodalonini Oken, 1843
Tribe Stenochiini Kirby, 1837
Tribe Talanini Champion, 1887 (1883)
Family Prostomidae Thomson, 1859
Family Synchroidae Lacordaire, 1859
Family Stenotrachelidae Thomson, 1859
Subfamily Stenotrachelinae Thomson, 1859
Subfamily Cephaloinae LeConte, 1862
Subfamily Nematoplinae LeConte, 1862
Subfamily Stoliinae Nikitsky, 1985
Family Oedemeridae Latreille, 1810
Subfamily Polypriinae Lawrence, 2005
Subfamily Calopodinae Costa, 1852 nomen protectum
Subfamily Oedemerinae Latreille, 1810
Tribe Asclerini Gistel, 1848
Tribe Ditylini Mulsant, 1858
Tribe Nacerdini Mulsant, 1858
Tribe Oedemerini Latreille, 1810
Tribe Stenostomatini Mulsant, 1858
Family Meloidae Gyllenhal, 1810
Subfamily Eleticinae Wellman, 1910
Tribe Derideini Wellman, 1910
Tribe Eleticini Wellman, 1910
Subtribe Eleticina Wellman, 1910
Subtribe Eospastina Selander, 1966
Tribe Ertlianini Selander, 1966
Tribe Spasticini Kaszab, 1959
Subtribe Anthicoxenina Selander, 1966
Subtribe Protomeloina Abdullah, 1965
Subtribe Spasticina Kaszab, 1959
Subtribe Xenospastina Selander, 1966
Subfamily Meloinae Gyllenhal, 1810
Tribe Cerocomini Leach, 1815
Tribe Epicautini Parker and Böving, 1924
Tribe Eupomphini LeConte, 1862
Tribe Lyttini Solier, 1851
Tribe Meloini Gyllenhal, 1810
Tribe Mylabrini Rafinesque, 1815
Tribe Pyrotini MacSwain, 1956
Subfamily Tetraonycinae Böving and Craighead, 1931
Subfamily Nemognathinae Laporte, 1840
Tribe Horiini Latreille, 1802
Tribe Nemognathini Laporte, 1840
Subtribe Nemognathina Laporte, 1840
Subtribe Zonitidina Mulsant, 1857
Subtribe Sitarina Mulsant, 1857
Tribe Stenoderini Selander, 1991
Family Mycteridae Oken, 1843
Subfamily Mycterinae Oken, 1843
Subfamily Eurypinae Thomson, 1860
Subfamily Hemipeplinae Lacordaire, 1854
Family Boridae Thomson, 1859
Subfamily Borinae Thomson, 1859
Subfamily Synercticinae Lawrence and Pollock, 1994
Family Trictenotomidae Blanchard, 1845
Family Pythidae Solier, 1834
Family Pyrochroidae Latreille, 1806
Subfamily Tydessinae Nikitsky, 1986
Subfamily Pilipalpinae Abdullah, 1964
Subfamily Pedilinae Lacordaire, 1859
Subfamily Pyrochroinae Latreille, 1806
Subfamily Agnathinae Lacordaire, 1859
Family Salpingidae Leach, 1815
Subfamily Othniinae LeConte, 1861
Subfamily Prostominiinae Grouvelle, 1914
Subfamily Agleninae Horn, 1878
Subfamily Inopeplinae Grouvelle, 1908
Subfamily Dacoderinae LeConte, 1862
Subfamily Aegialitinae LeConte, 1862
Subfamily Salpinginae Leach, 1815
Family Anthicidae Latreille, 1819
Subfamily Eurygeniinae LeConte, 1862
Tribe Eurygeniini LeConte, 1862
Tribe Ictistygnini Borchmann, 1936
Tribe Mitraelabrini Abdullah, 1969
Subfamily Macratriinae LeConte, 1862
Tribe Macratriini LeConte, 1862
†Tribe Camelomorphini Kirejtshuk and Azar, 2008
Subfamily Steropinae Jacquelin du Val, 1863
Subfamily Copobaeninae Abdullah, 1969
Subfamily Lemodinae Lawrence and Britton, 1991
Subfamily Tomoderinae Bonadona, 1961
Subfamily Anthicinae Latreille, 1819
Tribe Anthicini Latreille, 1819
Tribe Endomiini Kaszab, 1956
Tribe Formicomini Bonadona, 1974
Tribe Microhoriini Bonadona, 1974
Subfamily Notoxinae Stephens, 1829
Family Aderidae Csiki, 1909
Tribe Aderini Csiki, 1909
Subtribe Aderina Csiki, 1909
Subtribe Cnopina Báguena Corella, 1948
Subtribe Gompeliina Bouchard, 2011
Subtribe Syzetoninina Báguena Corella, 1948
Tribe Emelinini Báguena Corella, 1948
Tribe Euglenesini Seidlitz, 1875
Subtribe Euglenesina Seidlitz, 1875
Subtribe Pseudolotelina Báguena Corella, 1948
Tribe Phytobaenini Báguena Corella, 1948
Family Scraptiidae Gistel, 1848
Subfamily Scraptiinae Gistel, 1848
Tribe Allopodini Franciscolo, 1964
Tribe Scraptiini Gistel, 1848
Subfamily Anaspidinae Mulsant, 1856
Tribe Anaspidini Mulsant, 1856
Tribe Anaspimordini Franciscolo, 1954
Tribe Menuthianaspidini Franciscolo, 1972
Tribe Pentariini Franciscolo, 1954
Subfamily Lagrioidinae Abdullah and Abdullah, 1968
Subfamily Afreminae Levey, 1985
Subfamily Ischaliinae Blair, 1920
Superfamily Chrysomeloidea Latreille, 1802
Family Oxypeltidae Lacordaire, 1868
Family Vesperidae Mulsant, 1839
Subfamily Philinae Thomson, 1861
Subfamily Vesperinae Mulsant, 1839
Subfamily Anoplodermatinae Guérin-Méneville, 1840
Tribe Anoplodermatini Guérin-Méneville, 1840
Tribe Hypocephalini Blanchard, 1845
Tribe Mysteriini Prosen, 1960
Family Disteniidae Thomson, 1861
Tribe Cyrtonopini Gressitt, 1940
Tribe Disteniini Thomson, 1861
Tribe Dynamostini Lacordaire, 1868
Tribe Heteropalpini Villiers, 1961
Family Cerambycidae Latreille, 1802
Subfamily Parandrinae Blanchard, 1845
Tribe Erichsoniini Thomson, 1861
Tribe Parandrini Blanchard, 1845
Subfamily Prioninae Latreille, 1802
Tribe Acanthophorini Thomson, 1864
Tribe Aegosomatini Thomson, 1861
Tribe Anacolini Thomson, 1857
Tribe Cacoscelini Thomson, 1861
Tribe Callipogonini Thomson, 1861
Tribe Calocomini Galileo and Martins, 1993
Tribe Cantharocnemini Thomson, 1861
Tribe Ergatini Fairmaire, 1864
Tribe Eurypodini Gahan, 1906 (1868)
Tribe Hopliderini Thomson, 1864
Tribe Macrodontiini Thomson, 1861
Tribe Macrotomini Thomson, 1861
Subtribe Archetypina Lameere, 1912
Subtribe Basitoxina Lameere, 1912
Subtribe Macrotomina Thomson, 1861
Subtribe Platygnathina Gilmour, 1954
Subtribe Xixuthrina Lameere, 1912
Tribe Mallaspini Thomson, 1861
Tribe Mallodonini Thomson, 1861
Tribe Meroscelisini Thomson, 1861
Tribe Prionini Latreille, 1802
Tribe Remphanini Lacordaire, 1868
Tribe Solenopterini Lacordaire, 1868
Tribe Tereticini Lameere, 1913
Tribe Vesperoctenini Vives, 2005
Subfamily Lepturinae Latreille, 1802
Tribe Desmocerini Blanchard, 1845
Tribe Encyclopini LeConte, 1873
Tribe Lepturini Latreille, 1802
Tribe Oxymirini Danilevsky, 1997
Tribe Rhagiini Kirby, 1837
Tribe Rhamnusiini Sama, 2009
Tribe Teledapini Pascoe, 1871
Tribe Sachalinobiini Danilevsky, 2010
Tribe Xylosteini Reitter, 1913
Subfamily Spondylidinae Audinet-Serville, 1832
Tribe Anisarthrini Mamaev and Danilevsky, 1973
Tribe Asemini Thomson, 1861
Tribe Atimiini LeConte, 1873
Tribe Saphanini Gistel, 1848
Tribe Spondylidini Audinet-Serville, 1832
Subfamily Necydalinae Latreille, 1825
Subfamily Dorcasominae Lacordaire, 1868
Subfamily Apatophyseinae Lacordaire, 1869
Subfamily Cerambycinae Latreille, 1802
Tribe Acangassuini Galileo and Martins, 2001
Tribe Achrysonini Lacordaire, 1868
Tribe Agallissini LeConte, 1873
Tribe Alanizini Di Iorio, 2003
Tribe Anaglyptini Lacordaire, 1868 nomen protectum
Tribe Aphanasiini Lacordaire, 1868
Tribe Aphneopini Lacordaire, 1868
Tribe Auxesini Lepesme and Breuning, 1952
Tribe Basipterini Fragoso, Monné and Campos Seabra, 1987
Tribe Bimiini Lacordaire, 1868
Tribe Bothriospilini Lane, 1950
Tribe Brachypteromatini Sama, 2008
Tribe Callichromatini Swainson, 1840
Tribe Callidiini Kirby, 1837
Tribe Callidiopini Lacordaire, 1868
Tribe Cerambycini Latreille, 1802
Subtribe Cerambycina Latreille, 1802
Subtribe Sphallotrichina Martins and Monné, 2005
Tribe Certallini Fairmaire, 1864
Tribe Chlidonini Waterhouse, 1879
Tribe Cleomenini Lacordaire, 1868
Tribe Clytini Mulsant, 1839
Tribe Compsocerini Thomson, 1864
Tribe Coptommatini Lacordaire, 1869
Tribe Curiini LeConte, 1873
Tribe Deilini Fairmaire, 1864
Tribe Dejanirini Lacordaire, 1868
Tribe Diorini Lane, 1950
Tribe Distichocerini Pascoe, 1867
Tribe Dodecosini Aurivillius, 1912
Tribe Dryobiini Arnett, 1962 nomen protectum
Tribe Eburiini Blanchard, 1845
Tribe Ectenessini Martins, 1998
Tribe Elaphidiini Thomson, 1864
Tribe Eligmodermini Lacordaire, 1868
Tribe Erlandiini Aurivillius, 1912
Tribe Eroschemini Lacordaire, 1868
Tribe Eumichthini Linsley, 1940
Tribe Gahaniini Quentin and Villiers, 1969
Tribe Glaucytini Lacordaire, 1868
Tribe Graciliini Mulsant, 1839
Tribe Hesperophanini Mulsant, 1839
Subtribe Daramina Sama, 2008
Subtribe Hesperophanina Mulsant, 1839
Tribe Hesthesini Pascoe, 1867
Tribe Heteropsini Lacordaire, 1868 nomen protectum
Tribe Hexoplini Martins, 2006
Tribe Holopleurini Chemsak and Linsley, 1974
Tribe Holopterini Lacordaire, 1868
Tribe Hyboderini Linsley, 1940
Tribe Hylotrupini Zagajkevich, 1991
Tribe Ibidionini Thomson, 1861
Subtribe Compsina Martins and Galileo, 2007
Subtribe Ibidionina Thomson, 1861
Subtribe Tropidina Martins and Galileo, 2007
Tribe Ideratini Martins and Napp, 2009
Tribe Lissonotini Swainson, 1840
Tribe Luscosmodicini Martins, 2003
Tribe Lygrini Sama, 2008
Tribe Macronini Lacordaire, 1868
Tribe Megacoelini Quentin and Villiers, 1969
Tribe Methiini Thomson, 1860
Tribe Molorchini Gistel, 1848
Tribe Mythodini Lacordaire, 1868
Tribe Necydalopsini Lacordaire, 1868
Tribe Neocorini Martins, 2005
Tribe Neostenini Lacordaire, 1868
Tribe Obriini Mulsant, 1839
Tribe Ochyrini Pascoe, 1871
Tribe Oedenoderini Aurivillius, 1912
Tribe Oemini Lacordaire, 1868
Subtribe Methioidina Martins, 1997
Subtribe Oemina Lacordaire, 1868
Tribe Opsimini LeConte, 1873
Tribe Oxycoleini Martins and Galileo, 2003
Tribe Paraholopterini Martins, 1997
Tribe Phalotini Lacordaire, 1868
Tribe Phlyctaenodini Lacordaire, 1868
Tribe Phoracanthini Newman, 1840
Tribe Phyllarthriini Lepesme and Breuning, 1956
Tribe Piesarthriini McKeown, 1947
Tribe Piezocerini Lacordaire, 1868
Subtribe Haruspicina Martins, 1976
Subtribe Piezocerina Lacordaire, 1868
Tribe Platyarthrini Bates, 1870
Tribe Plectogasterini Quentin and Villiers, 1969
Tribe Plectromerini Nearns and Braham, 2008
Tribe Pleiarthrocerini Lane, 1950
Tribe Protaxini Gahan, 1906
Tribe Prothemini Lacordaire, 1868
Tribe Psebiini Lacordaire, 1868
Tribe Pseudocephalini Aurivillius, 1912 (1861)
Tribe Pseudolepturini Thomson, 1861
Tribe Psilomorphini Lacordaire, 1868
Tribe Pteroplatini Thomson, 1861
Tribe Pyrestini Lacordaire, 1868
Tribe Rhagiomorphini Newman, 1841
Tribe Rhinotragini Thomson, 1861
Tribe Rhopalophorini Blanchard, 1845
Tribe Rosaliini Fairmaire, 1864
Tribe Sestyrini Lacordaire, 1868
Tribe Smodicini Lacordaire, 1868
Tribe Spintheriini Lacordaire, 1869
Tribe Stenhomalini Miroshnikov, 1989
Tribe Stenoderini Pascoe, 1867
Tribe Stenopterini Gistel, 1848
Tribe Strongylurini Lacordaire, 1868
Tribe Tessarommatini Lacordaire, 1868
Tribe Thraniini Gahan, 1906
Tribe Thyrsiini Marinoni and Napp, 1984
Tribe Tillomorphini Lacordaire, 1868
Tribe Torneutini Thomson, 1861
Tribe Trachyderini Dupont, 1836
Subtribe Ancylocerina Thomson, 1864
Subtribe Trachyderina Dupont, 1836
Tribe Tragocerini Pascoe, 1867
Tribe Trichomesiini Aurivillius, 1912
Tribe Tropocalymmatini Lacordaire, 1868
Tribe Typhocesini Lacordaire, 1868
Tribe Unxiini Napp, 2007
Tribe Uracanthini Blanchard, 1853
Tribe Vesperellini Sama, 2008
Tribe Xystrocerini Blanchard, 1845
Subfamily Lamiinae Latreille, 1825
Tribe Acanthocinini Blanchard, 1845
Tribe Acanthoderini Thomson, 1860
Tribe Acmocerini Thomson, 1864
Tribe Acridocephalini Dillon and Dillon, 1959
Tribe Acrocinini Swainson, 1840
Tribe Aderpasini Breuning and Teocchi, 1978
Tribe Aerenicini Lacordaire, 1872
Tribe Agapanthiini Mulsant, 1839
Tribe Amphoecini Breuning, 1951
Tribe Ancitini Aurivillius, 1917
Tribe Ancylonotini Lacordaire, 1869
Tribe Anisocerini Thomson, 1860
Tribe Apomecynini Thomson, 1860
Tribe Astathini Thomson, 1864
Tribe Batocerini Thomson, 1864
Tribe Calliini Thomson, 1864
Tribe Ceroplesini Thomson, 1860
Subtribe Ceroplesina Thomson, 1860
Subtribe Crossotina Thomson, 1864
Tribe Cloniocerini Lacordaire, 1872
Tribe Colobotheini Thomson, 1860
Tribe Compsosomatini Thomson, 1857
Tribe Cyrtinini Thomson, 1864
Tribe Desmiphorini Thomson, 1860
Tribe Dorcadionini Swainson, 1840
Tribe Dorcaschematini Thomson, 1860
Tribe Elytracanthinini Bousquet, 2009
Tribe Enicodini Thomson, 1864
Tribe Eupromerini Galileo and Martins, 1995
Tribe Forsteriini Tippmann, 1960
Tribe Gnomini Thomson, 1860
Tribe Gyaritini Breuning, 1950
Tribe Heliolini Breuning, 1951
Tribe Hemilophini Thomson, 1868 nomen protectum
Tribe Homonoeini Thomson, 1864
Tribe Hyborhabdini Aurivillius, 1911
Tribe Lamiini Latreille, 1825
Tribe Laticraniini Lane, 1959
Tribe Mauesiini Lane, 1956
Tribe Megabasini Thomson, 1860
Tribe Mesosini Mulsant, 1839
Tribe Microcymaturini Breuning and Teocchi, 1985
Tribe Moneilemini Thomson, 1864
Tribe Monochamini Gistel, 1848
Tribe Morimonellini Lobanov, Danilevsky and Murzin, 1981
Tribe Morimopsini Lacordaire, 1869
Tribe Nyctimeniini Gressitt, 1951
Tribe Obereini Thomson, 1864
Tribe Oculariini Breuning, 1950
Tribe Onciderini Thomson, 1860
Tribe Oncideropsidini Aurivillius, 1922
Tribe Onocephalini Thomson, 1860
Tribe Onychogleneini Aurivillius, 1923
Tribe Parmenini Mulsant, 1839
Tribe Petrognathini Blanchard, 1845
Tribe Phacellini Lacordaire, 1872
Tribe Phantasini Kolbe, 1897
Tribe Phrynetini Thomson, 1864
Tribe Phymasternini Teocchi, 1989
Tribe Phytoeciini Mulsant, 1839
Tribe Pogonocherini Mulsant, 1839
Tribe Polyrhaphidini Thomson, 1860
Tribe Pretiliini Martins and Galileo, 1990
Tribe Proctocerini Aurivillius, 1922
Tribe Prosopocerini Thomson, 1864
Tribe Pteropliini Thomson, 1860
Tribe Rhodopinini Gressitt, 1951
Tribe Saperdini Mulsant, 1839
Tribe Stenobiini Breuning, 1950
Tribe Sternotomini Thomson, 1860
Tribe Tapeinini Thomson, 1857
Tribe Tetraopini Thomson, 1860
Tribe Tetraulaxini Breuning and Teocchi, 1977
Tribe Tetropini Portevin, 1927
Tribe Theocrini Lacordaire, 1872
Tribe Tmesisternini Blanchard, 1853
Tribe Tragocephalini Thomson, 1857
Tribe Xenicotelini Matsushita, 1933
Tribe Xenofreini Aurivillius, 1923
Tribe Xenoleini Lacordaire, 1872
Tribe Xylorhizini Lacordaire, 1872
Tribe Zygocerini Thomson, 1864
Family Megalopodidae Latreille, 1802
Subfamily Megalopodinae Latreille, 1802
Subfamily Palophaginae Kuschel and May, 1990
Subfamily Zeugophorinae Böving and Craighead, 1931
Family Orsodacnidae Thomson, 1859
Subfamily Orsodacninae Thomson, 1859
Subfamily Aulacoscelidinae Chapuis, 1874
Family Chrysomelidae Latreille, 1802
Subfamily Sagrinae Leach, 1815
Tribe Carpophagini Chapuis, 1874
Tribe Diaphanopsidini Monrós, 1958
Tribe Megamerini Chapuis, 1874
Tribe Sagrini Leach, 1815
Subfamily Bruchinae Latreille, 1802
Tribe Amblycerini Bridwell, 1932
Subtribe Amblycerina Bridwell, 1932
Subtribe Spermophagina Borowiec, 1987
Tribe Bruchini Latreille, 1802
Subtribe Acanthoscelidina Bridwell, 1946
Subtribe Bruchina Latreille, 1802
Subtribe Megacerina Bridwell, 1946
Tribe Eubaptini Bridwell, 1932
Tribe Kytorhinini Bridwell, 1932
Tribe Pachymerini Bridwell, 1929
Subtribe Caryedontina Bridwell, 1929
Subtribe Caryopemina Bridwell, 1929
Subtribe Pachymerina Bridwell, 1929
Tribe Rhaebini Blanchard, 1845
Subfamily Donaciinae Kirby, 1837
Tribe Donaciini Kirby, 1837
Tribe Haemoniini Chen, 1941
Tribe Plateumarini Böving, 1922
Subfamily Criocerinae Latreille, 1804
Tribe Criocerini Latreille, 1804
Tribe Lemini Gyllenhal, 1813
Tribe Pseudocriocerini Heinze, 1962
Subfamily Cassidinae Gyllenhal, 1813
Tribe Alurnini Chapuis, 1875
Tribe Anisoderini Chapuis, 1875
Tribe Aproidini Weise, 1911
Tribe Arescini Chapuis, 1875
Tribe Aspidimorphini Chapuis, 1875
Tribe Basiprionotini Gressitt, 1952 (1929)
Tribe Botryonopini Chapuis, 1875
Tribe Callispini Chapuis, 1875
Tribe Callohispini Uhmann, 1960
Tribe Cassidini Gyllenhal, 1813
Tribe Cephaloleiini Chapuis, 1875
Tribe Chalepini Weise, 1910
Tribe Coelaenomenoderini Weise, 1911
Tribe Cryptonychini Chapuis, 1875
Tribe Delocraniini Spaeth, 1929
Tribe Dorynotini Monrós and Viana, 1949 (1923)
Tribe Eugenysini Hincks, 1952
Tribe Eurispini Chapuis, 1875
Tribe Exothispini Weise, 1911
Tribe Goniocheniini Spaeth, 1942
Tribe Gonophorini Chapuis, 1875
Tribe Hemisphaerotini Monrós and Viana, 1951 (1929)
Tribe Hispini Gyllenhal, 1813
Tribe Hispoleptini Chapuis, 1875
Tribe Hybosispini Weise, 1910
Tribe Imatidiini Hope, 1840
Tribe Ischyrosonychini Chapuis, 1875
Tribe Leptispini Fairmaire, 1868
Tribe Mesomphaliini Hope, 1840
Tribe Nothosacanthini Gressitt, 1952 (1929)
Tribe Oediopalpini Monrós and Viana, 1947 (1910)
Tribe Omocerini Hincks, 1952 (1923)
Tribe Oncocephalini Chapuis, 1875
Tribe Promecothecini Chapuis, 1875
Tribe Prosopodontini Weise, 1910
Tribe Sceloenoplini Uhmann, 1930
Tribe Spilophorini Chapuis, 1875
Tribe Uroplatini Weise, 1910
Subfamily Chrysomelinae Latreille, 1802
Tribe Chrysomelini Latreille, 1802
Tribe Timarchini Motschulsky, 1860
Subfamily Galerucinae Latreille, 1802
Tribe Alticini Newman, 1834
Tribe Decarthrocerini Laboissière, 1937
Tribe Galerucini Latreille, 1802
Tribe Hylaspini Chapuis, 1875
Tribe Luperini Gistel, 1848
Tribe Metacyclini Chapuis, 1875
Tribe Oidini Laboissière, 1921 (1875)
Subfamily Lamprosomatinae Lacordaire, 1848
Tribe Lamprosomatini Lacordaire, 1848
Tribe Neochlamysini Monrós, 1959
Tribe Sphaerocharini Chapuis, 1874
Subfamily Cryptocephalinae Gyllenhal, 1813
Tribe Clytrini Kirby, 1837
Tribe Cryptocephalini Gyllenhal, 1813
Subtribe Achaenopina Chapuis, 1874
Subtribe Cryptocephalina Gyllenhal, 1813
Subtribe Monachulina Leng, 1920
Subtribe Pachybrachina Chapuis, 1874
Subtribe Stylosomina Chapuis, 1874
Tribe Fulcidacini Jakobson, 1924
Subfamily Eumolpinae Hope, 1840
Tribe Bromiini Baly, 1865 (1863)
Tribe Caryonodini Bechyné, 1951
Tribe Cubispini Monrós, 1954
Tribe Eumolpini Hope, 1840
Tribe Euryopini Chapuis, 1874
Tribe Habrophorini Bechyné and Špringlová de Bechyné, 1969
Tribe Hemydacnini Bechyné, 1951
Tribe Megascelidini Chapuis, 1874
Tribe Merodini Chapuis, 1874
Tribe Pygomolpini Bechyné, 1949
Tribe Rosiroiini Bechyné, 1950
Tribe Typophorini Baly, 1865
Subfamily Spilopyrinae Chapuis, 1874
Subfamily Synetinae LeConte and Horn, 1883
†Subfamily Protoscelidinae Medvedev, 1968
Superfamily Curculionoidea Latreille, 1802
Family Nemonychidae Bedel, 1882
Subfamily Nemonychinae Bedel, 1882
Subfamily Cimberidinae Gozis, 1882
Tribe Cimberidini Gozis, 1882
Tribe Doydirhynchini Pierce, 1916
†Tribe Kuschelomacrini Riedel, 2010
Subfamily Rhinorhynchinae Voss, 1922
Tribe Mecomacerini Kuschel, 1994
Subtribe Brarina Legalov, 2009
Subtribe Mecomacerina Kuschel, 1994
Tribe Rhinorhynchini Voss, 1922
†Subfamily Slonikinae Zherikhin, 1977
†Tribe Slonikini Zherikhin, 1977
†Tribe Ulyaniscini Legalov, 2009
†Subfamily Eccoptarthrinae Arnoldi, 1977
†Subfamily Brenthorrhininae Arnoldi, 1977
†Tribe Brenthorrhinini Arnoldi, 1977
†Tribe Brenthorrhinoidini Legalov, 2003
†Subfamily Distenorrhininae Arnoldi, 1977
†Subfamily Eobelinae Arnoldi, 1977
†Tribe Eobelini Arnoldi, 1977
†Subtribe Eobelina Arnoldi, 1977
†Subtribe Procurculionina Arnoldi, 1977
†Tribe Karataucarini Legalov, 2009
†Tribe Nanophydini Arnoldi, 1977
†Tribe Oxycorynoidini Arnoldi, 1977
†Tribe Probelini Legalov, 2009
†Subfamily Paleocartinae Legalov, 2003
†Tribe Nebrenthorrhinini Legalov, 2007
†Tribe Paleocartini Legalov, 2003
†Subfamily Metrioxenoidinae Legalov, 2009
†Subfamily Cretonemonychinae Gratshev and Legalov, 2009
†Subfamily Selengarhynchinae Gratshev and Legalov, 2009
Family Anthribidae Billberg, 1820
Subfamily Anthribinae Billberg, 1820
Tribe Anthribini Billberg, 1820
Tribe Basitropini Lacordaire, 1865
Tribe Corrhecerini Lacordaire, 1865
Tribe Cratoparini LeConte, 1876
†Tribe Cretanthribini Legalov, 2009
Tribe Decataphanini Lacordaire, 1865
Tribe Discotenini Lacordaire, 1865
Tribe Ecelonerini Lacordaire, 1865
Tribe Ischnocerini Lacordaire, 1865
Tribe Gymnognathini Valentine, 1960
Tribe Jordanthribini Morimoto, 1980
Tribe Mauiini Valentine, 1990
Tribe Mecocerini Lacordaire, 1865
Tribe Mycteini Morimoto, 1972
Tribe Ozotomerini Morimoto, 1972
Tribe Piesocorynini Valentine, 1960
Tribe Platyrhinini Bedel, 1882
Tribe Platystomini Pierce, 1916
Tribe Proscoporhinini Lacordaire, 1865
Tribe Ptychoderini Jekel, 1855
Tribe Sintorini Lacordaire, 1865
Tribe Stenocerini Kolbe, 1895
Tribe Tophoderini Lacordaire, 1865
Tribe Trigonorhinini Valentine, 1999
Tribe Tropiderini Lacordaire, 1865
Tribe Xenocerini Lacordaire, 1865
Tribe Xylinadini Lacordaire, 1865
Tribe Zygaenodini Lacordaire, 1865
Subfamily Choraginae Kirby, 1819
Tribe Apolectini Lacordaire, 1865
Tribe Araecerini Lacordaire, 1865
Tribe Cisanthribini Zimmerman, 1994
Tribe Choragini Kirby, 1819
Tribe Valenfriesiini Alonso-Zarazaga and Lyal, 1999
Tribe Xenorchestini Lacordaire, 1865
Subfamily Urodontinae Thomson, 1859
†Family Ulyanidae Zherikhin, 1993
Family Belidae Schönherr, 1826
Subfamily Belinae Schönherr, 1826
Tribe Agnesiotidini Zimmerman, 1994
Tribe Belini Schönherr, 1826
Subtribe Belina Schönherr, 1826
Subtribe Homalocerina Legalov, 2009
Tribe Pachyurini Kuschel, 1959
Subfamily Oxycoryninae Schönherr, 1840
Tribe Aglycyderini Wollaston, 1864
Tribe Alloxycorynini Legalov, 2009
Tribe Distenorrhinoidini Legalov, 2009
Tribe Metrioxenini Voss, 1953
Subtribe Afrocorynina Voss, 1957
Subtribe Metrioxenina Voss, 1953
Subtribe Zherichinixenina Legalov, 2009
Tribe Oxycorynini Schönherr, 1840
Subtribe Allocorynina Sharp, 1890
Subtribe Oxycorynina Schönherr, 1840
Subtribe Oxycraspedina Marvaldi and Oberprieler, 2006
Family Caridae Thompson, 1992
Subfamily Carinae Thompson, 1992
Subfamily Chilecarinae Legalov, 2009
Tribe Carodini Legalov, 2009
Tribe Chilecarini Legalov, 2009
†Subfamily Baissorhynchinae Zherikhin, 1993
Family Attelabidae Billberg, 1820
Subfamily Attelabinae Billberg, 1820
Tribe Attelabini Billberg, 1820
Subtribe Attelabina Billberg, 1820
Subtribe Euscelina Voss, 1925
Subtribe Euscelophilina Voss, 1925
Subtribe Henicolabina Legalov, 2007
Subtribe Himatolabina Legalov, 2003
Subtribe Hybolabina Voss, 1925
Subtribe Isolabina Legalov, 2007
Subtribe Lagenoderina Voss, 1925
Subtribe Lamprolabina Voss, 1925
Subtribe Metocalolabina Legalov, 2003
Subtribe Omolabina Legalov, 2003
Subtribe Paramecolabina Legalov, 2003
Subtribe Phialodina Legalov, 2003
Subtribe Phymatolabina Voss, 1925
Subtribe Phymatopsinina Legalov, 2003
Subtribe Pleurolabina Legalov, 2003
Tribe Euopini Voss, 1925
Tribe Pilolabini Voss, 1925
Subfamily Apoderinae Jekel, 1860
Tribe Apoderini Jekel, 1860
Tribe Clitostylini Voss, 1929
Subtribe Allapoderina Legalov, 2003
Subtribe Clitostylina Voss, 1929
Subtribe Pseudophrysina Legalov, 2003
Tribe Hoplapoderini Voss, 1926
Subtribe Afroapoderina Legalov, 2003
Subtribe Hoplapoderina Voss, 1926
Subtribe Paratomapoderina Legalov, 2003
Tribe Trachelophorini Voss, 1926
Subfamily Rhynchitinae Gistel, 1848
Tribe Auletini Desbrochers des Loges, 1908
Subtribe Auletina Desbrochers des Loges, 1908
Subtribe Auletobiina Legalov, 2001
Subtribe Guineauletina Legalov, 2003
Subtribe Mandelschtamiina Legalov, 2003
Subtribe Pseudauletina Voss, 1933
Subtribe Pseudomesauletina Legalov, 2003
Tribe Auletorhinini Voss, 1935
Tribe Byctiscini Voss, 1923
Subtribe Byctiscina Voss, 1923
Subtribe Listrobyctiscina Legalov, 2003
Subtribe Svetlanaebyctiscina Legalov, 2003
Tribe Cesauletini Legalov, 2003
Tribe Deporaini Voss, 1929
Subtribe Chonostropheina Morimoto, 1962
Subtribe Deporaina Voss, 1929
Tribe Minurini Legalov, 2003
Tribe Rhinocartini Voss, 1931
Tribe Rhynchitini Gistel, 1848
Subtribe Acritorrhynchitina Legalov, 2007
Subtribe Anisomerinina Legalov, 2003
Subtribe Eugnamptina Voss, 1930
Subtribe Lasiorhynchitina Legalov, 2003
Subtribe Perrhynchitina Legalov, 2003
Subtribe Rhynchitallina Legalov, 2003
Subtribe Rhynchitina Gistel, 1848
Subtribe Temnocerina Legalov, 2003
Subfamily Isotheinae Scudder, 1893
Tribe Isotheini Scudder, 1893
Subtribe Depasophilina Legalov, 2003
†Subtribe Isotheina Scudder, 1893
†Tribe Toxorhynchini Scudder, 1893
Subfamily Pterocolinae Lacordaire, 1865
Family Brentidae Billberg, 1820
Subfamily Brentinae Billberg, 1820
Tribe Brentini Billberg, 1820
Subtribe Arrhenodina Lacordaire, 1865
Subtribe Brentina Billberg, 1820
Subtribe Eremoxenina Semenov, 1892
Tribe Cyladini Schönherr, 1823
Tribe Cyphagogini Kolbe, 1892
Subtribe Atopobrentina Damoiseau, 1965
Subtribe Cyphagogina Kolbe, 1892
†Subtribe Dominibrentina Poinar, 2009
Subtribe Hoplopisthiina Senna and Calabresi, 1919
Subtribe Stereodermina Sharp, 1895
Tribe Pholidochlamydini Damoiseau, 1962
Tribe Taphroderini Lacordaire, 1865
Tribe Trachelizini Lacordaire, 1865
Subtribe Acratina Alonso-Zarazaga, Lyal, Bartolozzi and Sforzi, 1999
Subtribe Ithystenina Lacordaire, 1865
Subtribe Microtrachelizina Zimmerman, 1994
Subtribe Pseudoceocephalina Kleine, 1922
Subtribe Rhyticephalina Kleine, 1922
Subtribe Trachelizina Lacordaire, 1865
Subtribe Tychaeina Schoenfeldt, 1908
Tribe Ulocerini Schönherr, 1823
Subfamily Eurhynchinae Lacordaire, 1863
†Tribe Axelrodiellini Legalov, 2009
Tribe Eurhynchini Lacordaire, 1863
Subfamily Apioninae Schönherr, 1823
Supertribe Apionitae Schönherr, 1823
Tribe Apionini Schönherr, 1823
Subtribe Apionina Schönherr, 1823
Subtribe Aplemonina Kissinger, 1968
Subtribe Aspidapiina Alonso-Zarazaga, 1990
Subtribe Catapiina Alonso-Zarazaga, 1990
Subtribe Ceratapiina Alonso-Zarazaga, 1990
Subtribe Exapiina Alonso-Zarazaga, 1990
Subtribe Ixapiina Alonso-Zarazaga, 1990
Subtribe Kalcapiina Alonso-Zarazaga, 1990
Subtribe Malvapiina Alonso-Zarazaga, 1990
Subtribe Metapiina Alonso-Zarazaga, 1990
Subtribe Oxystomatina Alonso-Zarazaga, 1990
Subtribe Piezotrachelina Voss, 1959
Subtribe Prototrichapiina Wanat, 1995
Subtribe Synapiina Alonso-Zarazaga, 1990
Subtribe Trichapiina Alonso-Zarazaga, 1990
Tribe Chilapiini Wanat, 2001
Tribe Noterapiini Kissinger, 2004
Tribe Podapiini Wanat, 2001
Tribe Rhinorhynchidiini Zimmerman, 1994
Supertribe Antliarhinitae Schönherr, 1823
Supertribe Cybebitae Lacordaire, 1863
Supertribe Mecolenitae Wanat, 2001
Supertribe Myrmacicelitae Zimmerman, 1994
Tribe Lispotheriini Wanat, 2001
Tribe Myrmacicelini Zimmerman, 1994
Supertribe Rhadinocybitae Alonso-Zarazaga, 1992
Tribe Notapionini Zimmerman, 1994
Tribe Rhadinocybini Alonso-Zarazaga, 1992
Supertribe Tanaitae Schönherr, 1839
Subfamily Ithycerinae Schönherr, 1823
Subfamily Microcerinae Lacordaire, 1863
Subfamily Nanophyinae Gistel, 1848
Tribe Corimaliini Alonso-Zarazaga, 1989
Tribe Nanophyini Gistel, 1848
Family Dryophthoridae Schönherr, 1825
Subfamily Dryophthorinae Schönherr, 1825
Subfamily Cryptodermatinae Bovie, 1908
Subfamily Orthognathinae Lacordaire, 1865
Tribe Orthognathini Lacordaire, 1865
Tribe Rhinostomini LeConte, 1874
Subfamily Rhynchophorinae Schönherr, 1833
Tribe Diocalandrini Zimmerman, 1993
Tribe Litosomini Lacordaire, 1865
Tribe Ommatolampini Lacordaire, 1865
Tribe Polytini Zimmerman, 1993
Tribe Rhynchophorini Schönherr, 1833
Tribe Sphenophorini Lacordaire, 1865
Subfamily Stromboscerinae Lacordaire, 1865
Family Brachyceridae Billberg, 1820
Subfamily Brachycerinae Billberg, 1820
Tribe Brachycerini Billberg, 1820
Tribe Byrsopini Germar, 1829
Subfamily Cryptolarynginae Schalkwyk, 1966
Subfamily Erirhininae Schönherr, 1825
Tribe Aonychini Zimmerman, 1993
Tribe Arthrostenini Reitter, 1913
†Tribe Cretuliini Legalov, 2009
Tribe Erirhinini Schönherr, 1825
Tribe Himasthlophallini Zherikhin, 1991
Tribe Stenopelmini LeConte, 1876
Tribe Tadiini Zimmerman, 1993
Tribe Tanysphyrini Gistel, 1848
Subfamily Ocladiinae Lacordaire, 1865
Tribe Desmidophorini Morimoto, 1962
Tribe Ocladiini Lacordaire, 1865
Subfamily Raymondionyminae Reitter, 1913
Tribe Myrtonymini Kuschel, 1990
Tribe Raymondionymini Reitter, 1913
Family Curculionidae Latreille, 1802
Subfamily Curculioninae Latreille, 1802
Tribe Acalyptini Thomson, 1859
Subtribe Acalyptina Thomson, 1859
Subtribe Derelomina Lacordaire, 1865
Subtribe Notolomina Franz, 2006
Subtribe Phyllotrogina Franz, 2006
Subtribe Staminodeina Franz, 2006
Tribe Acentrusini Alonso-Zarazaga, 2005
Tribe Ancylocnemidini Voss, 1962
Tribe Anthonomini Thomson, 1859
Tribe Camarotini Schönherr, 1833
Subtribe Camarotina Schönherr, 1833
Subtribe Prionomerina Lacordaire, 1863
Tribe Ceratopodini Lacordaire, 1863
Tribe Cionini Schönherr, 1825
Tribe Cranopoeini Kuschel, 2009
Tribe Cryptoplini Lacordaire, 1863
Tribe Curculionini Latreille, 1802
Subtribe Curculionina Latreille, 1802
Subtribe Pseudobalaninina Heller, 1925
Subtribe Timolina Heller, 1925
Tribe Diabathrariini Lacordaire, 1863
Tribe Ellescini Thomson, 1859
Subtribe Dorytomina Bedel, 1886
Subtribe Ellescina Thomson, 1859
Tribe Erodiscini Lacordaire, 1863
Tribe Eugnomini Lacordaire, 1863
Subtribe Eugnomina Lacordaire, 1863
Subtribe Meriphina Marshall, 1937
Tribe Gonipterini Lacordaire, 1863
Tribe Mecinini Gistel, 1848
Tribe Nerthopini Lacordaire, 1865
Tribe Otidocephalini Lacordaire, 1863
Tribe Piazorhinini Lacordaire, 1863
Tribe Prionobrachiini Hustache, 1938
Tribe Pyropini Lacordaire, 1865
Tribe Rhamphini Rafinesque, 1815
Subtribe Dinorhopalina Voss, 1936
Subtribe Ixalmina Voss, 1936
Subtribe Rhamphina Rafinesque, 1815
Subtribe Tachygonina Lacordaire, 1865
Tribe Smicronychini Seidlitz, 1891 nomen protectum
Tribe Sphaeriopoeini Kuschel, 2003
Tribe Storeini Lacordaire, 1863
Tribe Styphlini Jekel, 1861
Tribe Tychiini Gistel, 1848
Subtribe Demimaeina Voss, 1937
Subtribe Lignyodina Bedel, 1883
Subtribe Ochyromerina Voss, 1935
Subtribe Tychiina Gistel, 1848
Tribe Ulomascini Lacordaire, 1865
Tribe Viticiini Morimoto, 1983
Subfamily Bagoinae Thomson, 1859 nomen protectum
Subfamily Baridinae Schönherr, 1836
Tribe Ambatini Lacordaire, 1863
Tribe Anopsilini Bondar, 1942
Tribe Apostasimerini Schönherr, 1844
Subtribe Apostasimerina Schönherr, 1844
Subtribe Madopterina Lacordaire, 1865
Subtribe Thaliabaridina Bondar, 1943
Subtribe Torcina Bondar, 1943
Subtribe Zygobaridina Pierce, 1907
Tribe Baridini Schönherr, 1836
Subtribe Baridina Schönherr, 1836
Subtribe Coelonertina Casey, 1922
Subtribe Coleomerina Casey, 1922
Subtribe Diorymerina Jekel, 1865
Subtribe Eurhinina Lacordaire, 1865
Tribe Madarini Jekel, 1865
Subtribe Barymerina Lacordaire, 1865
Subtribe Eutoxina Champion, 1908
Subtribe Leptoschoinina Lacordaire, 1865
Subtribe Madarina Jekel, 1865
Subtribe Tonesiina Alonso-Zarazaga and Lyal, 1999
Tribe Neosharpiini Hoffmann, 1956
Tribe Nertinini Voss, 1954
Tribe Optatini Champion, 1907
Tribe Pantotelini Lacordaire, 1865
Subtribe Cyrionychina Casey, 1922
Subtribe Pantotelina Lacordaire, 1865
Tribe Peridinetini Lacordaire, 1865
Subfamily Ceutorhynchinae Gistel, 1848
Tribe Ceutorhynchini Gistel, 1848
Tribe Cnemogonini Colonnelli, 1979
Tribe Egriini Pajni and Kohli, 1982
Tribe Hypohypurini Colonnelli, 2004
Tribe Hypurini Schultze, 1902
Tribe Lioxyonychini Colonnelli, 1984
Tribe Mecysmoderini Wagner, 1938
Tribe Mononychini LeConte, 1876
Tribe Phytobiini Gistel, 1848
Tribe Scleropterini Schultze, 1902
Subfamily Conoderinae Schönherr, 1833
Tribe Arachnopodini Lacordaire, 1865
Tribe Campyloscelini Schönherr, 1845
Subtribe Campyloscelina Schönherr, 1845
Subtribe Corynemerina Hustache, 1929
Subtribe Phaenomerina Faust, 1898
Tribe Conoderini Schönherr, 1833
Tribe Coryssomerini Thomson, 1859
Tribe Coryssopodini Lacordaire, 1865
Tribe Lechriopini Lacordaire, 1865
Tribe Lobotrachelini Lacordaire, 1865
Tribe Mecopini Lacordaire, 1865
Tribe Menemachini Lacordaire, 1865
Tribe Othippiini Morimoto, 1962
Tribe Peloropodini Hustache, 1932
Tribe Piazurini Lacordaire, 1865
Tribe Sphadasmini Lacordaire, 1865
Tribe Trichodocerini Champion, 1906
Tribe Zygopini Lacordaire, 1865
Subfamily Cossoninae Schönherr, 1825
Tribe Acamptini LeConte, 1876
Tribe Acanthinomerini Voss, 1972
Tribe Allomorphini Folwaczny, 1973
Tribe Aphyllurini Voss, 1955
Tribe Araucariini Kuschel, 1966
Tribe Choerorhinini Folwaczny, 1973
Tribe Cossonini Schönherr, 1825
Tribe Cryptommatini Voss, 1972
Tribe Dryotribini LeConte, 1876
Tribe Microxylobiini Voss, 1972
Tribe Nesiobiini Alonso-Zarazaga and Lyal, 1999
Tribe Neumatorini Folwaczny, 1973
Tribe Onychiini Chapuis, 1869
Tribe Onycholipini Wollaston, 1873
Tribe Pentarthrini Lacordaire, 1865
Tribe Proecini Voss, 1956
Tribe Pseudapotrepini Champion, 1909
Tribe Rhyncolini Gistel, 1848
Subtribe Phloeophagina Voss, 1955
Subtribe Pseudomimina Voss, 1939
Subtribe Rhyncolina Gistel, 1848
Tribe Tapiromimini Voss, 1972
Subfamily Cryptorhynchinae Schönherr, 1825
Tribe Aedemonini Faust, 1898
Tribe Camptorhinini Lacordaire, 1865
Tribe Cryptorhynchini Schönherr, 1825
Subtribe Cryptorhynchina Schönherr, 1825
Subtribe Mecistostylina Lacordaire, 1865
Subtribe Tylodina Lacordaire, 1865
Tribe Gasterocercini Zherikhin, 1991
Tribe Psepholacini Lacordaire, 1865
Tribe Sophrorhinini Lacordaire, 1865
Tribe Torneumatini Bedel, 1884
Subfamily Cyclominae Schönherr, 1826
Tribe Amycterini Waterhouse, 1854
Tribe Aterpini Lacordaire, 1863 nomen protectum
Subtribe Aterpina Lacordaire, 1863 nomen protectum
Subtribe Rhadinosomina Lacordaire, 1863
Tribe Cyclomini Schönherr, 1826
Tribe Dichotrachelini Hoffmann, 1957
Tribe Hipporhinini Lacordaire, 1863
Tribe Listroderini LeConte, 1876
Tribe Notiomimetini Wollaston, 1873
Tribe Rhythirrinini Lacordaire, 1863
Subfamily Entiminae Schönherr, 1823
Tribe Agraphini Horn, 1876
Tribe Alophini LeConte, 1874
Tribe Anomophthalmini Morrone, 1998
Tribe Anypotactini Champion, 1911
Tribe Blosyrini Lacordaire, 1863
Tribe Brachyderini Schönherr, 1826
Tribe Celeuthetini Lacordaire, 1863
Subtribe Celeuthetina Lacordaire, 1863
Subtribe Isopterina Morimoto and Kojima, 2001
Tribe Cneorhinini Lacordaire, 1863
Tribe Cratopodini Hustache, 1919
Tribe Cylydrorhinini Lacordaire, 1863
Tribe Cyphicerini Lacordaire, 1863
Subtribe Acanthotrachelina Marshall, 1944
Subtribe Cyphicerina Lacordaire, 1863
Subtribe Mylacorrhinina Reitter, 1913
Subtribe Myllocerina Pierce, 1913
Subtribe Phytoscaphina Lacordaire, 1863
Tribe Ectemnorhinini Lacordaire, 1863
Tribe Elytrurini Marshall, 1956
Tribe Embrithini Marshall, 1942
Tribe Entimini Schönherr, 1823
Tribe Episomini Lacordaire, 1863
Tribe Eudiagogini LeConte, 1874
Tribe Eupholini Günther, 1943
Tribe Eustylini Lacordaire, 1863
Tribe Geonemini Gistel, 1848
Tribe Holcorhinini Desbrochers des Loges, 1898
Tribe Hormorini Horn, 1876
Tribe Laparocerini Lacordaire, 1863
Tribe Leptostethini Lacordaire, 1863
Tribe Lordopini Schönherr, 1823
Tribe Mesostylini Reitter, 1913
Tribe Myorhinini Marseul, 1863
Tribe Nastini Reitter, 1913
Tribe Naupactini Gistel, 1848 nomen protectum
Tribe Nothognathini Marshall, 1916
Tribe Omiini Shuckard, 1839
Tribe Oosomini Lacordaire, 1863
Tribe Ophryastini Lacordaire, 1863
Tribe Ophtalmorrhynchini Hoffmann, 1965
Tribe Otiorhynchini Schönherr, 1826
Tribe Ottistirini Heller, 1925
Tribe Pachyrhynchini Schönherr, 1826
Tribe Peritelini Lacordaire, 1863
Tribe Phyllobiini Schönherr, 1826
Tribe Polycatini Marshall, 1956
Tribe Polydrusini Schönherr, 1823
Tribe Premnotrypini Kuschel, 1956
†Tribe Pristorhynchini Heer, 1847
Tribe Prypnini Lacordaire, 1863
Tribe Psallidiini Lacordaire, 1863
Tribe Rhyncogonini Sharp, 1919
Tribe Sciaphilini Sharp, 1891
Tribe Sitonini Gistel, 1848
Tribe Tanymecini Lacordaire, 1863
Subtribe Piazomiina Reitter, 1913
Subtribe Tainophthalmina Desbrochers des Loges, 1873
Subtribe Tanymecina Lacordaire, 1863
Tribe Tanyrhynchini Schönherr, 1826
Tribe Thecesternini Lacordaire, 1863
Tribe Trachyphloeini Gistel, 1848
Subtribe Trachyphilina Voss, 1948
Subtribe Trachyphloeina Gistel, 1848
Tribe Tropiphorini Marseul, 1863
Tribe Typhlorhinini Kuschel, 1954
Subfamily Hyperinae Marseul, 1863 (1848)
Tribe Cepurini Capiomont, 1867
Tribe Hyperini Marseul, 1863 (1848)
Subfamily Lixinae Schönherr, 1823
Tribe Cleonini Schönherr, 1826 nomen protectum
Tribe Lixini Schönherr, 1823
Tribe Rhinocyllini Lacordaire, 1863
Subfamily Mesoptiliinae Lacordaire, 1863
Tribe Carciliini Pierce, 1916
Tribe Laemosaccini Lacordaire, 1865
Tribe Magdalidini Pascoe, 1870 nomen protectum
Tribe Mesoptiliini Lacordaire, 1863
Subfamily Molytinae Schönherr, 1823
Tribe Anoplini Bedel, 1883
Tribe Amalactini Lacordaire, 1863
Tribe Aminyopini Voss, 1956
Tribe Amorphocerini Voss, 1939
Tribe Anchonini Imhoff, 1856
Tribe Brachyceropseini Aurivillius, 1926
Tribe Cholini Schönherr, 1825
Subtribe Cholina Schönherr, 1825
Subtribe Cholomina Vaurie, 1974
Subtribe Rhinastina Vaurie, 1973
Tribe Cleogonini Gistel, 1848
Tribe Conotrachelini Jekel, 1865
Tribe Cycloterini Lacordaire, 1863
Subtribe Cycloterina Lacordaire, 1863
Subtribe Thrombosternina Voss, 1965
Tribe Dinomorphini Lacordaire, 1863
Tribe Emphyastini Lacordaire, 1863
Tribe Euderini Lacordaire, 1865
Tribe Galloisiini Morimoto, 1962
Tribe Guioperini Lacordaire, 1865
Tribe Hylobiini Kirby, 1837
Subtribe Epistrophina Marshall, 1932
Subtribe Hylobiina Kirby, 1837
Tribe Ithyporini Lacordaire, 1865
Subtribe Colobodina Voss, 1958
Subtribe Ithyporina Lacordaire, 1865
Subtribe Sclerocardiina Lacordaire, 1865
Tribe Itini Reitter, 1913
Tribe Juanorhinini Aurivillius, 1931
Tribe Lepyrini Kirby, 1837
Tribe Lithinini Lacordaire, 1863
Subtribe Lithinina Lacordaire, 1863
Subtribe Rhytidophloeina Voss, 1963
Tribe Lymantini Lacordaire, 1865
Tribe Mecysolobini Reitter, 1913
Tribe Metatygini Pascoe, 1888
Tribe Molytini Schönherr, 1823
Subtribe Leiosomatina Reitter, 1913
Subtribe Molytina Schönherr, 1823
Subtribe Plinthina Lacordaire, 1863
Subtribe Typoderina Voss, 1965
Tribe Nettarhinini Lacordaire, 1865
Tribe Pacholenini Lacordaire, 1863
Tribe Paipalesomini Marshall, 1932
Tribe Petalochilini Lacordaire, 1863
Tribe Phoenicobatini Champion, 1914
Tribe Phrynixini Kuschel, 1964
Tribe Pissodini Gistel, 1848
Subtribe Cotasteromimina Morimoto, 1962
Subtribe Orthorhinina Jekel, 1865
Subtribe Pissodina Gistel, 1848
Tribe Sternechini Lacordaire, 1863
Tribe Styanacini Chûjô and Voss, 1960
Tribe Trachodini Gistel, 1848
Tribe Trigonocolini Lacordaire, 1863
Tribe Trypetidini Lacordaire, 1865
Subfamily Orobitidinae Thomson, 1859
Subfamily Xiphaspidinae Marshall, 1920
Subfamily Scolytinae Latreille, 1804
Tribe Amphiscolytini Mandelshtam and Beaver, 2003
Tribe Bothrosternini Blandford, 1896
Tribe Cactopinini Chamberlin, 1939
Tribe Carphodicticini Wood, 1971
Tribe Coptonotini Chapuis, 1869
Tribe Corthylini LeConte, 1876
Subtribe Corthylina LeConte, 1876
Subtribe Pityophthorina Eichhoff, 1878
Tribe Cryphalini Lindemann, 1877
Tribe Crypturgini LeConte, 1876
†Tribe Cylindrobrotini Kirejtshuk, Azar, Beaver, Mandelshtam and Nel, 2009
Tribe Diamerini Hagedorn, 1909
Tribe Dryocoetini Lindemann, 1877
Tribe Hexacolini Eichhoff, 1878
Tribe Hylastini LeConte, 1876
Tribe Hylesinini Erichson, 1836
Tribe Hylurgini Gistel, 1848
Tribe Hyorrhynchini Hopkins, 1915
Tribe Hypoborini Nüsslin, 1911
Tribe Ipini Bedel, 1888
Tribe Micracidini LeConte, 1876
Tribe Phloeosinini Nüsslin, 1912
Tribe Phloeotribini Chapuis, 1869
Tribe Phrixosomatini Wood, 1978
Tribe Polygraphini Chapuis, 1869
Tribe Premnobiini Browne, 1962
Tribe Scolytini Latreille, 1804
Tribe Scolytoplatypodini Blandford, 1893
Tribe Xyleborini LeConte, 1876
Tribe Xyloctonini Eichhoff, 1878
Tribe Xyloterini LeConte, 1876
Subfamily Platypodinae Shuckard, 1839
Tribe Mecopelmini Thompson, 1992
Tribe Platypodini Shuckard, 1839
Tribe Schedlariini Wood and Bright, 1992
Tribe Tesserocerini Strohmeyer, 1914
Subtribe Diapodina Strohmeyer, 1914
Subtribe Tesserocerina Strohmeyer, 1914
Catalogue of Coleoptera family-group names
†Suborder
PROTOCOLEOPTERA
†Superfamily
Tshekardocoleoidea
Rohdendorf, 1944
Tshekardocoleidae Rohdendorf, 1944: 252 [stem: Tshekardocole-]. Type genus: Tshekardocoleus Rohdendorf, 1944.
†Family
Tshekardocoleidae
Rohdendorf, 1944
Tshekardocoleidae Rohdendorf, 1944: 252 [stem: Tshekardocole-]. Type genus: Tshekardocoleus Rohdendorf, 1944.
Uralocoleidae Zalesskiy, 1947: 857 [stem: Uralocole-]. Type genus: Uralocoleus Zalessky, 1947.
†Family
Labradorocoleidae
Ponomarenko, 1969
Labradorocoleidae Ponomarenko, 1969b: 307 [stem: Labradorocole-]. Type genus: Labradorocoleus Ponomarenko, 1969.
†Family
Oborocoleidae
Kukalová, 1969
Oborocoleidae Kukalová, 1969: 155 [stem: Oborocole-]. Type genus: Oborocoleus Kukalová, 1969.
†Superfamily
Permocupedoidea
Martynov, 1933
Permocupidae Martynov, 1933: 85 [stem: Permocuped-]. Type genus: Permocupes Martynov, 1933.
†Family
Permocupedidae
Martynov, 1933
Permocupidae Martynov, 1933: 85 [stem: Permocuped-]. Type genus: Permocupes Martynov, 1933. Comment: incorrect original stem formation, not in prevailing usage.
Kaltanocoleidae Rohdendorf, 1961: 397 [stem: Kaltanocole-]. Type genus: Kaltanocoleus Rohdendorf, 1961.
†Family
Taldycupedidae
Rohdendorf, 1961
Taldycupidae Rohdendorf, 1961: 412 [stem: Taldycuped-]. Type genus: Taldycupes Rohdendorf, 1961. Comment: usage of Trachycupidae by Fujiyama (1973: 375) was in error for Taldycupidae (Ponomarenko pers. comm. August 2009); incorrect original stem formation, not in prevailing usage.
†Superfamily
Permosynoidea
Tillyard, 1924
Permosynidae Tillyard, 1924: 431 [stem: Permosyn-]. Type genus: Permosyne Tillyard, 1924.
†Family
Ademosynidae
Ponomarenko, 1968
Ademosynidae Ponomarenko, 1968: 128 [stem: Ademosyn-]. Type genus: Ademosyne Handlirsch, 1906.
†Family
Permosynidae
Tillyard, 1924
Permosynidae Tillyard, 1924: 431 [stem: Permosyn-]. Type genus: Permosyne Tillyard, 1924.
Family
Crowsoniellidae
Iablokoff-Khnzorian, 1983
Crowsoniellidae Iablokoff-Khnzorian, 1983: 65 [stem: Crowsoniell-]. Type genus: Crowsoniella Pace, 1975.
Family
Cupedidae
Laporte, 1836
Cupesidae Laporte, 1836: 56 [stem: Cuped-]. Type genus: Cupes Fabricius, 1801.
Subfamily
Priacminae
Crowson, 1962
Priacmini Crowson, 1962: 152 [stem: Priacm-]. Type genus: Priacma J. L. LeConte, 1874.
†Subfamily
Mesocupedinae
Ponomarenko, 1969
Mesocupedini Ponomarenko, 1969a: 105 [stem: Mesocuped-]. Type genus: Mesocupes Martynov, 1926.
Subfamily
Cupedinae
Laporte, 1836
Cupesidae Laporte, 1836: 56 [stem: Cuped-]. Type genus: Cupes Fabricius, 1802. Comment: incorrect original stem formation, not in prevailing usage.
Family
Micromalthidae
Barber, 1913
Micromalthidae Barber, 1913: 185 [stem: Micromalth-]. Type genus: Micromalthus J. L. LeConte, 1878.
Family
Ommatidae
Sharp and Muir, 1912
Ommadidae Sharp and Muir, 1912: 521 [stem: Ommat-]. Type genus: Omma Newman, 1839.
†Subfamily
Brochocoleinae
Hong, 1982
Brochocoleidae Hong, 1982: 100 [stem: Brochocole-]. Type genus: Brochocoleus Hong, 1982.
Subfamily
Tetraphalerinae
Crowson, 1962
Tetraphalerini Crowson, 1962: 152 [stem: Tetraphaler-]. Type genus: Tetraphalerus C. O. Waterhouse, 1901.
Subfamily
Ommatinae
Sharp and Muir, 1912
Ommadidae Sharp and Muir, 1912: 521 [stem: Ommat-]. Type genus: Omma Newman, 1839.
†Tribe
Lithocupedini
Ponomarenko, 1969
Lithocupedini Ponomarenko, 1969a: 82 [stem: Lithocuped-]. Type genus: Lithocupes Ponomarenko, 1966.
†Tribe
Notocupedini
Ponomarenko, 1966
Notocupedini Ponomarenko, 1966: 60 [stem: Notocup-]. Type genus: Notocupes Ponomarenko, 1966.
Tribe
Ommatini
Sharp and Muir, 1912
Ommadidae Sharp and Muir, 1912: 521 [stem: Ommat-]. Type genus: Omma Newman, 1839. Comment: incorrect original stem formation, not in prevailing usage.
Family
Jurodidae
Ponomarenko, 1985
Jurodidae Ponomarenko, 1985: 53 [stem: Jurod-]. Type genus: Jurodes Ponomarenko, 1985.
Sikhotealiniidae Lafer, 1996: 390 [stem: Sikhotealini-]. Type genus: Sikhotealinia Lafer, 1996.
†Family
Triadocupedidae
Ponomarenko, 1966
Triadocupedinae Ponomarenko, 1966: 48 [stem: Triadocuped-]. Type genus: Triadocupes Ponomarenko, 1966.
†Family
Magnocoleidae
Hong, 1998
Magnocoleidae Hong, 1998: 41 [stem: Magnocole-]. Type genus: Magnocoleus Hong, 1998.
†Family
Obrieniidae
Zherikhin and Gratshev, 1994
Obrieniidae Zherikhin and Gratshev, 1994: 51 [stem: Obrieni-]. Type genus: Obrienia Zherikhin and Gratshev, 1994. Comment: precedence (Obrieniidae Zherikhin and Gratshev, 1994 vs Kararhynchidae Zherikhin and Gratshev, 1994) given to taxon originally proposed at the higher rank (Art. 24.1).
†Subfamily
Kararhynchinae
Zherikhin and Gratshev, 1994
Kararhynchinae Zherikhin and Gratshev, 1994: 58 [stem: Kararhynch-]. Type genus: Kararhynchus Zherikhin and Gratshev, 1994.
†Tribe
Kararhynchini
Zherikhin and Gratshev, 1994
Kararhynchinae Zherikhin and Gratshev, 1994: 58 [stem: Kararhynch-]. Type genus: Kararhynchus Zherikhin and Gratshev, 1994.
†Tribe
Kenderlykaini
Legalov, 2009
Kenderlykanini Legalov, 2009c: 285 [stem: Kenderlyka-]. Type genus: Kenderlyka Zherikhin and Gratshev, 1994. Comment: incorrect original stem formation, not in prevailing usage.
†Subfamily
Obrieniinae
Zherikhin and Gratshev, 1994
Obrieniidae Zherikhin and Gratshev, 1994: 51 [stem: Obrieni-]. Type genus: Obrienia Zherikhin and Gratshev, 1994.
†Superfamily
Asiocoleoidea
Rohdendorf, 1961
Asiocoleidae Rohdendorf, 1961: 396 [stem: Asiocole-]. Type genus: Asiocoleus Rohdendorf, 1961.
†Family
Asiocoleidae
Rohdendorf, 1961
Asiocoleidae Rohdendorf, 1961: 396 [stem: Asiocole-]. Type genus: Asiocoleus Rohdendorf, 1961.
†Family
Tricoleidae
Ponomarenko, 1969
Tricoleidae Ponomarenko, 1969a: 138 [stem: Tricole-]. Type genus: Tricoleus Ponomarenko, 1969.
†Superfamily
Rhombocoleoidea
Rohdendorf, 1961
Rhombocoleidae Rohdendorf, 1961: 432 [stem: Rhombocole-]. Type genus: Rhombocoleus Rohdendorf, 1961.
†Family
Rhombocoleidae
Rohdendorf, 1961
Rhombocoleidae Rohdendorf, 1961: 432 [stem: Rhombocole-]. Type genus: Rhombocoleus Rohdendorf, 1961.
†Superfamily
Schizophoroidea
Ponomarenko, 1968
Schizophoridae Ponomarenko, 1968: 130 [stem: Schizophor-]. Type genus: Schizophorus Ponomarenko, 1968. Comment: usage of this name conserved over Schizocoleoidea Rohdendorf, 1961 (Art. 35.5).
†Family
Schizophoridae
Ponomarenko, 1968
Schizophoridae Ponomarenko, 1968: 130 [stem: Schizophor-]. Type genus: Schizophorus Ponomarenko, 1968.
†Family
Catiniidae
Ponomarenko, 1968
Catiniidae Ponomarenko, 1968: 137 [stem: Catini-]. Type genus: Catinius Ponomarenko, 1968.
†Family
Schizocoleidae
Rohdendorf, 1961
Curculiopsidae Martynov, 1937: 39 [stem: Curculiopse-]. Type genus: Curculiopsis Martynov, 1937 [preoccupied genus name, not Curculiopsis Handlirsch, 1907 [fossil Curculionoidea]; syn. of Aenigmocoleus Rohdendorf, 1961 or Rossocoleus Rohdendorf, 1961]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Schizocoleidae Rohdendorf, 1961: 438 [stem: Schizocole-]. Type genus: Schizocoleus Rohdendorf, 1961.
Superfamily
Lepiceroidea
Hinton, 1936 (1882)
Lepiceridae Hinton, 1936: 473 [stem: Lepicer-]. Type genus: Lepicerus Motschulsky, 1855. Comment: use of family-group name conserved over Cyathoceroidea Sharp, 1882 (Art. 40.2) (see Lawrence and Newton 1995: 804).
Family
Lepiceridae
Hinton, 1936 (1882)
Cyathoceridae Sharp, 1882: 141 [stem: Cyathocer-]. Type genus: Cyathocerus Sharp, 1882 [syn. of Lepicerus Motschulsky, 1855]. Comment: use of younger name Lepiceridae Hinton, 1936 conserved over this name (Art. 40.2) (see Lawrence and Newton 1995: 804).
Lepiceridae Hinton, 1936: 473 [stem: Lepicer-]. Type genus: Lepicerus Motschulsky, 1855. Comment: use of family-group name conserved over Cyathoceridae Sharp, 1882 (Art. 40.2) (see Lawrence and Newton 1995: 804).
†Haplochelidae Kirejtshuk and Poinar, 2006: 156 [stem: Haplochel-]. Type genus: Haplochelus Kirejtshuk and Poinar, 2006. Comment: synonymy with Lepiceridae by Ge et al. (2010: 336).
Superfamily
Sphaeriusoidea
Erichson, 1845
Sphaerina Erichson, 1845: 38 [stem: Sphaerius-]. Type genus: Sphaerius Waltl, 1838 [originally placed on the Official Index of Rejected and Invalid Generic Names in Zoology by the Commission (ICZN 1985d) but later placed on the Official List of Generic Names in Zoology (ICZN 2000)]. Comment: correct stem determined to be Sphaerius- and Sphaeriusidae Erichson, 1845 placed on the Official List of Family-Group Names in Zoology (ICZN 2000).
Family
Torridincolidae
Steffan, 1964
Torridincolidae Steffan, 1964: 199 [stem: Torridincol-]. Type genus: Torridincola Steffan, 1964.
Subfamily
Torridincolinae
Steffan, 1964
Torridincolidae Steffan, 1964: 199 [stem: Torridincol-]. Type genus: Torridincola Steffan, 1964.
Ptyopterinae M. Abdullah, 1974: 961 [stem: Ptyopteryg-]. Type genus: Ptyopteryx H. Reichardt and C. Costa, 1967 [preoccupied genus name, not Ptyopteryx Kolenati, 1848 [Trichoptera]; syn. of Iapir Py-Daniel et al., 1993]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Subfamily
Deleveinae
Endrödy-Younga, 1997
Deleveinae Endrödy-Younga, 1997: 317 [stem: Deleve-]. Type genus: Delevea H. Reichardt, 1976.
Family
Hydroscaphidae
LeConte, 1874
Hydroscaphidae J. L. LeConte, 1874a: 45 [stem: Hydroscaph-]. Type genus: Hydroscapha J. L. LeConte, 1874.
Family
Sphaeriusidae
Erichson, 1845
Sphaerina Erichson, 1845: 38 [stem: Sphaerius-]. Type genus: Sphaerius Waltl, 1838 [originally placed on the Official Index of Rejected and Invalid Generic Names in Zoology by the Commission (ICZN 1985d) but later placed on the Official List of Generic Names in Zoology (ICZN 2000)]. Comment: family-group name and its type genus were rejected by the Commission (ICZN 1985d) but rescinded afterwards (ICZN 2000); family-group name placed on the Official List of Family-Group Names in Zoology and correct stem ruled to be Sphaerius- (ICZN 2000).
Microsporidae Crotch, 1873a: 78 [stem: Microspor-]. Type genus: Microsporus Kolenati, 1846 [placed on the Official List of Generic Names in Zoology (ICZN 1985d)]. Comment: name originally placed on the Official List of Family-Group Names in Zoology as “Microsporidae H. Reichardt, 1976” (ICZN 1985d) but author and year subsequently corrected to Crotch, 1873 (ICZN 2000).
†Family
Tritarsidae
Hong, 2002
Tritarsusidae Hong, 2002: 102 [stem: Tritars-]. Type genus: Tritarsus Hong, 2002. Comment: incorrect original stem formation, not in prevailing usage.
Family
Gyrinidae
Latreille, 1810
Gyrinites Latreille, 1810: 141 [stem: Gyrin-]. Type genus: Gyrinus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subfamily
Spanglerogyrinae
Folkerts, 1979
Spanglerogyrinae Folkerts, 1979: 7 [stem: Spanglerogyr-]. Type genus: Spanglerogyrus Folkerts, 1979.
Subfamily
Gyrininae
Latreille, 1810
Gyrinites Latreille, 1810: 141 [stem: Gyrin-]. Type genus: Gyrinus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Enhydrini
Régimbart, 1882
Enhydrini Régimbart, 1882: 392 [stem: Enhydr-]. Type genus: Enhydrus Laporte, 1834 [placed on the Official List of Generic Names in Zoology (ICZN 1964)]. Comment: usage of this name conserved over Dineutini Desmarest, 1851 (Art. 35.5).
Subtribe
Dineutina
Desmarest, 1851
Dineutides Desmarest, 1851: 223 [stem: Dineut-]. Type genus: Dineutes W. S. MacLeay, 1825.
Subtribe
Enhydrina
Régimbart, 1882
Enhydrini Régimbart, 1882: 392 [stem: Enhydr-]. Type genus: Enhydrus Laporte, 1834 [placed on the Official List of Generic Names in Zoology (ICZN 1964)]. Comment: junior homonym of Enhydrini Gray, 1825 (type genus Enhydra Fleming, 1822) in Mammalia; an application was recently submitted to the Commission by Özdikmen and Darilmaz (2010; see Appendix 6) to remove the homonymy (Art. 55.3.1).
Tribe
Gyrinini
Latreille, 1810
Gyrinites Latreille, 1810: 141 [stem: Gyrin-]. Type genus: Gyrinus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subtribe
Gyrinina
Latreille, 1810
Gyrinites Latreille, 1810: 141 [stem: Gyrin-]. Type genus: Gyrinus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subtribe
Heterogyrina
Brinck, 1956
Heterogyrini P. Brinck, 1956: 37, in key [stem: Heterogyr-]. Type genus: Heterogyrus Legros, 1953.
Tribe
Orectochilini
Régimbart, 1882
Orectochilini Régimbart, 1882: 391 [stem: Orectochil-]. Type genus: Orectochilus Dejean, 1833.
Family
Trachypachidae
Thomson, 1857
Trachypachini C. G. Thomson, 1857: 5 [stem: Trachypach-]. Type genus: Trachypachus Motschulsky, 1844.
†Subfamily
Eodromeinae
Ponomarenko, 1977
Eodromeinae Ponomarenko, 1977: 46 [stem: Eodrome-]. Type genus: Eodromeus Ponomarenko, 1977. Comment: the type genus originally included four species: antiquus, stenalis, dissectus and major; Ponomarenko (1977: 66) designated “fasciatus” as the type species of the genus which was a lapsus calami for dissectus (see Carpenter 1992: 291); this genus is considered here as being available in Ponomarenko (1977: 66) and not Ponomarenko (in Carpenter 1992: 291); this action enables us to treat the family-group name Eodromeinae as available from the original description in Ponomarenko (1977: 46).
Leptopodocoleidae Hong, 1982: 118 [stem: Leptopodocole-]. Type genus: Leptopodocoleus Hong, 1982.
Subfamily
Trachypachinae
Thomson, 1857
Trachypachini C. G. Thomson, 1857: 5 [stem: Trachypach-]. Type genus: Trachypachus Motschulsky, 1844.
*Systolosomini Erwin, 1985: 467 [stem: Systolosomat-]. Type genus: Systolosoma Solier, 1849. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Systolosomini Erwin, 1991: 4, in key [stem: Systolosomat-]. Type genus: Systolosoma Solier, 1849. Comment: incorrect original stem formation, not in prevailing usage.
Family
Rhysodidae
Laporte, 1840
Rhysodites Laporte, 1840a: 291 [stem: Rhysod-]. Type genus: Rhysodes Germar, 1822.
Tribe
Leoglymmiini
Bell and Bell, 1978
Leoglymmiina R. T. Bell and J. R. Bell, 1978: 53 [stem: Leoglymmi-]. Type genus: Leoglymmius Bell and Bell, 1978.
Tribe
Dhysorini
Bell and Bell, 1978
Dhysorina R. T. Bell and J. R. Bell, 1978: 53 [stem: Dhysor-]. Type genus: Dhysores Grouvelle, 1903.
Tribe
Medisorini
Bell and Bell, 1987
Medisorina R. T. Bell and J. R. Bell, 1987: 287 [stem: Medisor-]. Type genus: Medisores Bell and Bell, 1987.
Tribe
Rhysodini
Laporte, 1840
Rhysodites Laporte, 1840a: 291 [stem: Rhysod-]. Type genus: Rhysodes Germar, 1822. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 326, as Rhyssodeoidae [incorrect stem formation]), generally accepted as in R. T. Bell (2003: 78, as Rhysodidae).
Tribe
Clinidiini
Bell and Bell, 1978
Clinidiina R. T. Bell and J. R. Bell, 1978: 59 [stem: Clinidi-]. Type genus: Clinidium Kirby, 1830.
Tribe
Omoglymmiini
Bell and Bell, 1978
Omoglymmiina R. T. Bell and J. R. Bell, 1978: 66 [stem: Omoglymmi-]. Type genus: Omoglymmius Ganglbauer, 1891.
Tribe
Sloanoglymmiini
Bell and Bell, 1991
Sloanoglymmiina R. T. Bell and J. R. Bell, 1991: 183 [stem: Sloanoglymmi-]. Type genus: Sloanoglymmius Bell and Bell, 1991.
Family
Carabidae
Latreille, 1802
Carabici Latreille, 1802: 80 [stem: Carab-]. Type genus: Carabus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1950)]. Comment: First Reviser (Carabidae Latreille, 1802 vs Cicindelidae Latreille, 1802 vs Elaphridae Latreille, 1802) not determined, current usage maintained.
†Subfamily
Protorabinae
Ponomarenko, 1977
Protorabinae Ponomarenko, 1977: 71 [stem: Protorab-]. Type genus: Protorabus Ponomarenko, 1977.
†Subfamily
Conjunctiinae
Ponomarenko, 1977
Conjunctiini Ponomarenko, 1977: 85 [stem: Conjuncti-]. Type genus: Conjunctia Ponomarenko, 1977.
Subfamily
Nebriinae
Laporte, 1834
Nebriidae Laporte, 1834b: 90 [stem: Nebri-]. Type genus: Nebria Latreille, 1802.
Tribe
Nebriini
Laporte, 1834
Nebriidae Laporte, 1834b: 90 [stem: Nebri-]. Type genus: Nebria Latreille, 1802.
Tribe
Notiokasiini
Kavanaugh and Nègre, 1983
*Notiokasiini Erwin, 1979: 578 [stem: Notiokasi-]. Type genus: Notiokasis Kavanaugh and Nègre, 1983. Comment: family-group name unavailable (Art. 11.7.1.1): not based on an available genus name at the time.
Notiokasiini Kavanaugh and Nègre, 1983: 551 [stem: Notiokasi-]. Type genus: Notiokasis Kavanaugh and Nègre, 1983.
Tribe
Notiophilini
Motschulsky, 1850
*Notiophiles Motschulsky, 1849: 54 [stem: Notiophil-]. Type genus: Notiophilus Duméril, 1805. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Notiophili Motschulsky, 1850: 16 [stem: Notiophil-]. Type genus: Notiophilus Duméril, 1805.
Tribe
Opisthiini
Dupuis, 1912
Opisthiinae Dupuis, 1912: 1 [stem: Opisthi-]. Type genus: Opisthius Kirby, 1837.
Tribe
Pelophilini
Kavanaugh, 1996
Pelophilini Kavanaugh, 1996: 35 [stem: Pelophil-]. Type genus: Pelophila Dejean, 1821.
Subfamily
Cicindinae
Csiki, 1927
Cicindini Csiki, 1927: 425 [stem: Cicind-]. Type genus: Cicindis Bruch, 1908.
Subfamily
Cicindelinae
Latreille, 1802
Cicindeletae Latreille, 1802: 77 [stem: Cicindel-]. Type genus: Cicindela Linné, 1758.
Tribe
Amblycheilini
Csiki, 1903
Amblychilinae Csiki, 1903: 124 [stem: Amblycheil-]. Type genus: Amblycheila Say, 1830 [as Amblychila, unjustified emendation of type genus name by Agassiz (1846b), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Omites W. Horn, 1907: 466 [stem: Om-]. Type genus: Omus Eschscholtz, 1829.
Tribe
Cicindelini
Latreille, 1802
Cicindeletae Latreille, 1802: 77 [stem: Cicindel-]. Type genus: Cicindela Linnaeus, 1758.
Subtribe
Apteroessina
Rivalier, 1971
Apteroessina Rivalier, 1971: 143 [stem: Apteroess-]. Type genus: Apteroessa Hope, 1838.
Subtribe
Cicindelina
Latreille, 1802
Cicindeletae Latreille, 1802: 77 [stem: Cicindel-]. Type genus: Cicindela Linnaeus, 1758.
Subtribe
Dromicina
Thomson, 1859
Dromicitae J. Thomson, 1859: 89 [stem: Dromic-]. Type genus: Dromica Dejean, 1826.
Caledonicini W. Horn, 1893: 324 [stem: Caledonic-]. Type genus: Caledonica Chaudoir, 1861.
Odontochilini W. Horn, 1899: 41 [stem: Odontocheil-]. Type genus: Odontocheila Laporte, 1834 [as Odontochila, unjustified emendation of type genus name by Agassiz (1846b), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Prepusini W. Horn, 1899: 44 [stem: Prepus-]. Type genus: Prepusa Chaudoir, 1850 [syn. of Eulampra Chaudoir, 1848].
Euryodini W. Horn, 1899: 37 [stem: Euryod-]. Type genus: Euryoda Lacordaire, 1842 [syn. of Prothyma Hope, 1838].
Prothymini W. Horn, 1906: 86 [stem: Prothym-]. Type genus: Prothyma Hope, 1838.
Subtribe
Iresiina
Rivalier, 1971
Euprosopini W. Horn, 1893: 324 [stem: Euprosop-]. Type genus: Euprosopus Dejean, 1825.
Eucalliini W. Horn, 1893: 324 [stem: Eucalli-]. Type genus: Eucallia Guérin-Méneville, 1844 [syn. of Callidema Guérin-Méneville, 1843].
Distypsiderini W. Horn, 1893: 324 [stem: Distipsider-]. Type genus: Distipsidera Westwood, 1837 [as Distypsidera, unjustified emendation of type genus name by Gemminger and Harold (1868a: 32), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Iresina Rivalier, 1971: 138 [stem: Iresi-]. Type genus: Iresia Dejean, 1829. Comment: although this is not the oldest name for the subtribe, we recommend that an application be sent to the Commission in order to conserve usage of Iresiina Rivalier, 1971 over the three older names proposed by W. Horn in 1893 which have not been used as valid recently to our knowledge; incorrect original stem formation, not in prevailing usage.
Subtribe
Theratina
Horn, 1893
Theratidae W. Horn, 1893: 324 [stem: Therat-]. Type genus: Therates Latreille, 1816.
Tribe
Collyridini
Brullé, 1834
Collyriens Brullé, 1834: 96 [stem: Collyrid-]. Type genus: Collyris Fabricius, 1801.
Subtribe
Collyridina
Brullé, 1834
Collyriens Brullé, 1834: 96 [stem: Collyrid-]. Type genus: Collyris Fabricius, 1801. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hope (1838a: 30, as Collyridae [incorrect stem formation]), generally accepted as in Puchkov and Matalin (2003: 116, as Collyridina); incorrect original stem formation, not in prevailing usage.
Tracheloniadae Gistel, 1850: 75 [stem: Tracheloni-]. Type genus: Trachelonia Gistel, 1850 [syn. of Collyris Fabricius, 1801].
*Colliurides Motschulsky, 1855: 34 [stem: Colliur-]. Type genus: Colliuris sensu Latreille, 1802 [not Colliuris DeGeer, 1774; syn. of Collyris Fabricius, 1801]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Subtribe
Tricondylina
Naviaux, 1991
Tricondylina Naviaux, 1991: 219 [stem: Tricondyl-]. Type genus: Tricondyla Latreille, 1822.
Tribe
Ctenostomatini
Laporte, 1834
Ctenostomidae Laporte, 1834b: 38 [stem: Ctenostomat-]. Type genus: Ctenostoma Klug, 1821. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Manticorini
Laporte, 1834
Manticoridae Laporte, 1834b: 33 [stem: Manticor-]. Type genus: Manticora Fabricius, 1781.
Tribe
Megacephalini
Laporte, 1834
Megacephalidae Laporte, 1834b: 33 [stem: Megacephal-]. Type genus: Megacephala Latreille, 1802.
Oxycheilites J. Thomson, 1857a: 17 [stem: Oxycheil-]. Type genus: Oxycheila Dejean, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Chaudoir (1861a: 326, as Oxychilini [incorrect stem formation]), generally accepted as in Fleutiaux (1892a: 13, as Oxychilini [incorrect stem formation]); the stem Oxychil- should not be used for this taxon in order to avoid homonymy with Oxychilini Hesse, 1927 (type genus Oxychilus Fitzinger 1833), currently being used as valid in Mollusca.
Platychilidae W. Horn, 1893: 325 [stem: Platychil-]. Type genus: Platychile W. S. MacLeay, 1825.
Tetrachae Leng and Mutchler, 1916: 683 [stem: Tetrach-]. Type genus: Tetracha Hope, 1838.
Subfamily
Carabinae
Latreille, 1802
Carabici Latreille, 1802: 80 [stem: Carab-]. Type genus: Carabus Linnaeus, 1758.
Tribe
Carabini
Latreille, 1802
Carabici Latreille, 1802: 80 [stem: Carab-]. Type genus: Carabus Linnaeus, 1758.
Calosomii Bonelli, 1810: Tabula Synoptica [stem: Calosomat-]. Type genus: Calosoma Weber, 1801. Comment: incorrect original stem formation, not in prevailing usage.
Proceridae Laporte, 1834b: 87 [stem: Procer-]. Type genus: Procerus Dejean, 1821.
Callisthenisidae Gistel, 1848: [2] [stem: Callisthen-]. Type genus: Callisthenes Fischer von Waldheim, 1820. Comment: incorrect original stem formation, not in prevailing usage.
*Cechenogenici Morawitz, 1889: 40 [stem: Cechen-]. Type genus: Cechenus Fischer von Waldheim, 1822. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun.
*Tribacogenici Morawitz, 1889: 40 [stem: Tribac-]. Type genus: Tribax Fischer von Waldheim, 1817. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun.
*Procrustogenici Morawitz, 1889: 40 [stem: Procrust-]. Type genus: Procrustes Bonelli, 1810. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun.
Procrustogenici Roeschke, 1898: 285 [stem: Procrust-]. Type genus: Procrustes Bonelli, 1810.
Aplothoracina Lapouge, 1927: 45 [stem: Aplothorac-]. Type genus: Aplothorax G. R. Waterhouse, 1842.
*Callistheniens Lapouge, 1927: 47 [stem: Callisthen-]. Type genus: Callisthenes Fischer von Waldheim, 1820. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899; family-group name proposed as new without reference to Callisthenisidae Gistel, 1848.
Cechenogenici Csiki, 1927: 110 [stem: Cechen-]. Type genus: Cechenus Fischer von Waldheim, 1822.
Tribacogenici Csiki, 1927: 121 [stem: Tribac-]. Type genus: Tribax Fischer von Waldheim, 1817.
Megadontici Csiki, 1927: 60 [stem: Megodont-]. Type genus: Megodontus Solier, 1848 [as Megadontus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Ceroglossini
Lapouge, 1927
Ceroglossina Lapouge, 1927: 45 [stem: Cerogloss-]. Type genus: Ceroglossus Solier, 1848.
Tribe
Cychrini
Perty, 1830
Cychrii Perty, 1830: 6 [stem: Cychr-]. Type genus: Cychrus Fabricius, 1794. Comment: name previously attributed to Laporte (1834b: 86).
Tribe
Pamborini
Hope, 1838
Pamboridae Hope, 1838a: 47 [stem: Pambor-]. Type genus: Pamborus Latreille, 1812.
Subfamily
Loricerinae
Bonelli, 1810
Loricerides Bonelli, 1810: Tabula Synoptica [stem: Loricer-]. Type genus: Loricera Latreille, 1802.
Subfamily
Omophroninae
Bonelli, 1810
Omophronii Bonelli, 1810: Tabula Synoptica [stem: Omophron-]. Type genus: Omophron Latreille, 1802.
Scolyti Motschulsky, 1850: 91 [stem: Scolyt-]. Type genus: Scolytus Fabricius, 1790 [preoccupied genus name, not Scolytus Geoffroy, 1762 [Coleoptera: Curculionidae]; syn. of Omophron Latreille, 1802]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; Scolytinae/-ini Latreille, 1804 are currently used as valid in Curculionidae.
Epactiini Fauvel, 1888: 1 [stem: Epacti-]. Type genus: Epactius Schneider, 1791 [syn. of Omophron Latreille, 1802].
Subfamily
Elaphrinae
Latreille, 1802
Elaphrii Latreille, 1802: 81 [stem: Elaphr-]. Type genus: Elaphrus Fabricius, 1775.
Subfamily
Migadopinae
Chaudoir, 1861
Migadopidae Chaudoir, 1861b: 510 [stem: Migadop-]. Type genus: Migadops G. R. Waterhouse, 1842.
Tribe
Amarotypini
Erwin, 1985
Amarotypini Erwin, 1985: 468 [stem: Amarotyp-]. Type genus: Amarotypus H. W. Bates, 1872.
Tribe
Migadopini
Chaudoir, 1861
Migadopidae Chaudoir, 1861b: 510 [stem: Migadop-]. Type genus: Migadops G. R. Waterhouse, 1842.
Subtribe
Aquilicina
Moret, 2005
Aquilicina Moret, 2005: 30 [stem: Aquilic-]. Type genus: Aquilex Moret, 1989.
Subtribe
Migadopina
Chaudoir, 1861
Migadopidae Chaudoir, 1861b: 510 [stem: Migadop-]. Type genus: Migadops G. R. Waterhouse, 1842.
Monolobinae Jeannel, 1938: 10, in key [stem: Monolob-]. Type genus: Monolobus Solier, 1849.
Loxomeriformes Erwin, 1985: 446 [stem: Loxomer-]. Type genus: Loxomerus Chaudoir, 1842.
Subfamily
Hiletinae
Schiødte, 1848
Hiletini Schiødte, 1848: 69 [stem: Hilet-]. Type genus: Hiletus Schiødte, 1848.
Camaragnathini Csiki, 1927: 341 [stem: Camaragnath-]. Type genus: Camaragnathus Bertrand-Bocandé, 1849 [syn. of Hiletus Schiødte, 1847].
Subfamily
Scaritinae
Bonelli, 1810
Scaritides Bonelli, 1810: Tabula Synoptica [stem: Scarit-]. Type genus: Scarites Fabricius, 1775.
Tribe
Carenini
MacLeay, 1887
Carenides W. J. MacLeay, 1887: 117 [stem: Caren-]. Type genus: Carenum Bonelli, 1813.
Tribe
Clivinini
Rafinesque, 1815
Clivinidia Rafinesque, 1815: 109 [stem: Clivin-]. Type genus: Clivina Latreille, 1802.
Subtribe
Ardistomina
Putzeys, 1867
Ardistomides Putzeys, 1867: 200 [stem: Ardistom-]. Type genus: Ardistomis Putzeys, 1846. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Csiki (1927: 547, as Ardistomina), generally accepted as in Balkenohl (2003: 219, as Ardistomina); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Ardistomid-).
Subtribe
Clivinina
Rafinesque, 1815
Clivinidia Rafinesque, 1815: 109 [stem: Clivin-]. Type genus: Clivina Latreille, 1802.
Reicheina Jeannel, 1957: 141, in key [stem: Reichei-]. Type genus: Reicheia Saulcy, 1862. Comment: incorrect original stem formation, not in prevailing usage.
Italodytina Jeannel, 1957: 141, in key [stem: Italodyt-]. Type genus: Italodytes Müller, 1938.
Reicheiina Basilewsky, 1980: 293 [stem: Reichei-]. Type genus: Reicheia Saulcy, 1862. Comment: family-group name proposed as new without reference to Reicheiina Jeannel, 1957.
Subtribe
Forcipatorina
Bänninger, 1938
*Oxystomides Putzeys, 1867: 12 [stem: Oxystom-]. Type genus: Oxystomus Dejean, 1825 [preoccupied genus name, not Oxystomus Fischer von Waldheim, 1803 [Mammalia]; syn. of Forcipator Maindron, 1904]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Putzeys (1867).
Oxystomina Csiki, 1927: 491. Type genus: Oxystomus Dejean, 1825 [preoccupied genus name, not Oxystomus Fischer von Waldheim, 1803 [Mammalia]; syn. of Forcipator Maindron, 1904]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Forcipatorina Bänninger, 1938: 83 [stem: Forcipator-]. Type genus: Forcipator Maindron, 1904.
Tribe
Dalyatini
Mateu, 2002
Dalyatinae Mateu, 2002: 67 [stem: Dalyat-]. Type genus: Dalyat Mateu, 2002.
Tribe
Dyschiriini
Kolbe, 1880
Dyschiriini Kolbe, 1880: 266 [stem: Dyschiri-]. Type genus: Dyschirius Bonelli, 1810.
†Tribe
Palaeoaxinidiini
McKay, 1991
Palaeoaxinidini McKay, 1991: 10 [stem: Palaeoaxinidi-]. Type genus: Palaeoaxinidium McKay, 1991. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Pasimachini
Putzeys, 1867
Pasimachides Putzeys, 1867: 3 [stem: Pasimach-]. Type genus: Pasimachus Bonelli, 1813. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Csiki (1927: 444, as Pasimachina).
Tribe
Promecognathini
LeConte, 1853
Promecognathi J. L. LeConte, 1853b: 394 [stem: Promecognath-]. Type genus: Promecognathus Chaudoir, 1846.
Axinidiini Basilewsky, 1963: 307 [stem: Axinidi-]. Type genus: Axinidium Sturm, 1843.
Tribe
Salcediini
Alluaud, 1930
Salcediini Alluaud, 1930a: 21 [stem: Salcedi-]. Type genus: Salcedia Fairmaire, 1899.
Subtribe
Androzelmina
Bell, 1998
Androzelmina R. T. Bell, 1998: 271 [stem: Androzelm-]. Type genus: Androzelma Dostal, 1993.
Subtribe
Salcediina
Alluaud, 1930 (1929)
Zelmides Andrewes, 1929: 209, 416 [stem: Zelm-]. Type genus: Zelma Andrewes, 1920 [syn. of Salcedia Fairmaire, 1899]. Comment: use of younger name Salcediina Alluaud, 1930 conserved over this name (Art. 40.2).
Salcediini Alluaud, 1930a: 21 [stem: Salcedi-]. Type genus: Salcedia Fairmaire, 1899. Comment: name proposed to replace Zelmini Andrewes, 1929 because of the synonymy of the type genus; use of Salcediina conserved over Zelmina Andrewes, 1929 (Art. 40.2).
Subtribe
Solenogenyina
Bell, 1998
Solenogenyina R. T. Bell, 1998: 271 [stem: Solenogeny-]. Type genus: Solenogenys Westwood, 1859.
Tribe
Scaritini
Bonelli, 1810
Scaritides Bonelli, 1810: Tabula Synoptica [stem: Scarit-]. Type genus: Scarites Fabricius, 1775.
Subtribe
Acanthoscelina
Csiki, 1927
Acanthoscelina Csiki, 1927: 489 [stem: Acanthoscel-]. Type genus: Acanthoscelis Dejean, 1825. Comment: the incorrect stem formation is in prevailing usage (should be Acanthoscelid-) and is maintained (Art. 29.3.1.1); usage of the stem Acanthoscel- avoids homonymy with Acanthoscelidina Bridwell, 1946 (type genus Acanthoscelides Schilsky, 1905) in Chrysomelidae.
Subtribe
Corintascarina
Basilewsky, 1973
Corintascarini Basilewsky, 1973a: 10 [stem: Corintascar-]. Type genus: Corintascaris Basilewsky, 1952. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Corintascarid-).
Subtribe
Dyscherina
Basilewsky, 1973
Dyscherina Basilewsky, 1973a: 101 [stem: Dyscher-]. Type genus: Dyscherus Chaudoir, 1855.
Subtribe
Ochyropina
Basilewsky, 1973
Ochyropini Basilewsky, 1973a: 9 [stem: Ochyrop-]. Type genus: Ochyropus Schiødte, 1847. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Ochyropod-).
Subtribe
Oxylobina
Andrewes, 1929
Oxylobides Andrewes, 1929: 209, 292 [stem: Oxylob-]. Type genus: Oxylobus Chaudoir, 1855.
Subtribe
Scapterina
Putzeys, 1867
Scaptérides Putzeys, 1867: 7 [stem: Scapter-]. Type genus: Scapterus Dejean, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Csiki (1927: 490, as Scapterina), generally accepted as in Balkenohl (2003: 231, as Scapterina).
Passalidiina Csiki, 1927: 489 [stem: Passalidi-]. Type genus: Passalidius Chaudoir, 1863.
Subtribe
Scaritina
Bonelli, 1810
Scaritides Bonelli, 1810: Tabula Synoptica [stem: Scarit-]. Type genus: Scarites Fabricius, 1775.
Subtribe
Storthodontina
Jeannel, 1946
Storthodontini Jeannel, 1946: 237, in key [stem: Storthodont-]. Type genus: Storthodontus Chaudoir, 1855.
Subfamily
Broscinae
Hope, 1838
Broschidae Hope, 1838a: 80 [stem: Brosc-]. Type genus: Broscus Panzer, 1813.
Tribe
Broscini
Hope, 1838
Broschidae Hope, 1838a: 80 [stem: Brosc-]. Type genus: Broscus Panzer, 1813.
Subtribe
Axonyina
Roig-Juñent, 2000
Axonyina Roig-Juñent, 2000: 18 [stem: Axony-]. Type genus: Axonya Andrewes, 1923.
Subtribe
Baripodina
Jeannel, 1941
Barypitae Jeannel, 1941: 287, in key [stem: Baripod-]. Type genus: Baripus Dejean, 1828 [as Barypus, unjustified emendation of type genus name by Agassiz (1846b: 43), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Broscina
Hope, 1838
Broschidae Hope, 1838a: 80 [stem: Brosc-]. Type genus: Broscus Panzer, 1813 [as Broschus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Cephalotida Heer, 1838: 12 [stem: Cephalot-]. Type genus: Cephalotes Bonelli, 1810 [preoccupied genus name, not Cephalotes Latreille, 1802 [Hymenoptera]; syn. of Broscus Panzer, 1813]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Zacotini G. H. Horn, 1881: 169 [stem: Zacot-]. Type genus: Zacotus J. L. LeConte, 1869.
Subtribe
Creobiina
Jeannel, 1941
Creobitae Jeannel, 1941: 287, in key [stem: Creobi-]. Type genus: Creobius Guérin-Méneville, 1838. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Nothobroscina
Roig-Juñent, 2000
Nothobroscina Roig-Juñent, 2000: 32 [stem: Nothobrosc-]. Type genus: Nothobroscus Roig-Juñent and Ball, 1995.
Subfamily
Apotominae
LeConte, 1853
Apotomi J. L. LeConte, 1853b: 370 [stem: Apotom-]. Type genus: Apotomus Illiger, 1807.
Subfamily
Siagoninae
Bonelli, 1813
Siagones Bonelli, 1813: 24 [stem: Siagon-]. Type genus: Siagona Latreille, 1804.
Tribe
Enceladini
Horn, 1881
Enceladini G. H. Horn, 1881: 118 [stem: Encelad-]. Type genus: Enceladus Bonelli, 1813.
Tribe
Lupercini
Lecordier, 1977
Lupercini Lecordier, 1977: 628 [stem: Luperc-]. Type genus: Luperca Laporte, 1840.
Tribe
Siagonini
Bonelli, 1813
Siagones Bonelli, 1813: 24 [stem: Siagon-]. Type genus: Siagona Latreille, 1804.
Subfamily
Melaeninae
Csiki, 1933
Coscinides Chaudoir, 1876b: 115 [stem: Coscini-]. Type genus: Coscinia Dejean, 1831 [preoccupied genus name, not Coscinia Hübner, 1819 [Lepidoptera]; syn. of Cymbionotum Baudi, 1864]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Granigerini Bedel, 1900: 24 [stem: Graniger-]. Type genus: Graniger sensu Chaudoir, 1876 [not Graniger Motschulsky, 1864; syn. of Cymbionotum Baudi, 1864]. Comment: based on a misidentified type genus; an application will need to be submitted to the Commission to suppress this name for the Principles of Priority and Homonymy (Art. 65.2.1) if Granigerini Antoine, 1959 in Harpalinae: Harpalini: Harpalina is to be used as valid in the future.
Melaeninae Csiki, 1933: 1650 [stem: Melaen-]. Type genus: Melaenus Dejean, 1831. Comment: published 26 May 1933; name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Lorenz (1998b: 149) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Cymbionotini Andrewes, 1933: 3 [stem: Cymbionot-]. Type genus: Cymbionotum Baudi di Selve, 1864. Comment: published 30 June 1933.
Subfamily
Gehringiinae
Darlington, 1933
Gehringiini Darlington, 1933: 110 [stem: Gehringi-]. Type genus: Gehringia Darlington, 1933.
Tribe
Gehringiini
Darlington, 1933
Gehringiini Darlington, 1933: 110 [stem: Gehringi-]. Type genus: Gehringia Darlington, 1933.
Subtribe
Gehringiina
Darlington, 1933
Gehringiini Darlington, 1933: 110 [stem: Gehringi-]. Type genus: Gehringia Darlington, 1933.
Subtribe
Helenaeina
Deuve, 2007
Helenaeina Deuve, 2007: 217 [stem: Helenae-]. Type genus: Helenaea Schatzmayr and Koch, 1934.
Subfamily
Trechinae
Bonelli, 1810
Trechii Bonelli, 1810: Tabula Synoptica [stem: Trech-]. Type genus: Trechus Clairville, 1806.
Tribe
Bembidiini
Stephens, 1827
Bembidiidae Stephens, 1827: 5 [stem: Bembidi-]. Type genus: Bembidion Latreille, 1802 [as Bembidium, unjustified emendation of type genus name by Gyllenhal (1810: 12), not in prevailing usage].
Subtribe
Anillina
Jeannel, 1937
Anillini Jeannel, 1937: 244, in key [stem: Anill-]. Type genus: Anillus Jacquelin du Val, 1851. Comment: precedence (Anillina Jeannel, 1937 vs Scotodipnina Jeannel, 1937) given to taxon originally proposed at the higher rank (Art. 24.1).
Scotodipnina Jeannel, 1937: 265, in key [stem: Scotodipn-]. Type genus: Scotodipnus Schaum, 1860.
Typhlocharina Jeanne, 1973: 95, in key [stem: Typhlocharit-]. Type genus: Typhlocharis Dieck, 1869. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Bembidiina
Stephens, 1827
Bembidiidae Stephens, 1827: 5 [stem: Bembidi-]. Type genus: Bembidion Latreille, 1802 [as Bembidium, unjustified emendation of type genus name by Gyllenhal (1810), not in prevailing usage].
Peryphidae Kirby, 1837: 52 [stem: Peryph-]. Type genus: Peryphus Dejean, 1821.
Subtribe
Tachyina
Motschulsky, 1862
Tachyaires Motschulsky, 1862: 24 [stem: Tachy-]. Type genus: Tachys Dejean, 1821 [placed on the Official List of Generic Names in Zoology (ICZN 1990c)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jeannel (1941: 401, 422, as Tachyini), generally accepted as in J. K. Park et al. (2006: 91, as Tachyini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Tache-, see Alonso-Zarazaga 2007).
Micratopini Casey, 1914a: 42 [stem: Micratop-]. Type genus: Micratopus Casey, 1914.
Limnastini Jeannel, 1937: 245, in key [stem: Lymnast-]. Type genus: Lymnastis Motschulsky, 1862 [as Limnastis, unjustified emendation of type genus name by Ganglbauer (1891a: 181), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Xystosomina
Erwin, 1994
Xystosomina Erwin, 1994: 560 [stem: Xystosom-]. Type genus: Xystosomus Schaum, 1863.
Tribe
Horologionini
Jeannel, 1949
Horologionidae Jeannel, 1949b: 91 [stem: Horologion-]. Type genus: Horologion J. M. Valentine, 1932. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Horologi-).
Tribe
Pogonini
Laporte, 1834
Pogonidae Laporte, 1834b: 70 [stem: Pogon-]. Type genus: Pogonus Dejean, 1821.
Pogonopsini Bedel, 1900: 20 [stem: Pogonopse-]. Type genus: Pogonopsis Bedel, 1898. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Trechini
Bonelli, 1810
Trechii Bonelli, 1810: Tabula Synoptica [stem: Trech-]. Type genus: Trechus Clairville, 1806.
Subtribe
Aepina
Fowler, 1887
Aëpyes Fowler, 1887: 123 [stem: Aep-]. Type genus: Aepus Samouelle, 1819.
Temnostegini Enderlein, 1909: 376 [stem: Temnosteg-]. Type genus: Temnostega Enderlein, 1905.
Subtribe
Cnidina
Jeannel, 1958
Cnidina Jeannel, 1958b: 733 [stem: Cnid-]. Type genus: Cnides Motschulsky, 1862. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Casale and Laneyrie (1982: 9, as Cnidini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Subtribe
Perileptina
Sloane, 1903
Perileptides Sloane, 1903: 583 [stem: Perilept-]. Type genus: Perileptus Schaum, 1860. Comment: Perileptides Sloane, 1903 was treated as Latin and available in Madge (1989: 466) but the valid name for this taxon was listed as Perilepti Jeannel, 1922 by Csiki (1928) and Perileptina Jeannel, 1922 by Lorenz (2005); we follow the usage of Madge (1989).
Ochthephilini Jeannel, 1922: 165 [stem: Ochthephil-]. Type genus: Ochthephilus Nietner, 1857 [preoccupied genus name, not Ochthephilus Mulsant and Rey, 1856 [Coleoptera: Staphylinidae]; syn. of Perileptus Schaum, 1860]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Subtribe
Plocamotrechina
Jeannel, 1960
*Plocamotrechina Jeannel, 1958b: 733 [stem: Plocamotrech-]. Type genus: Plocamotrechus Jeannel, 1926 [syn. of Pachydesus Motschulsky, 1864]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Plocamotrechina Jeannel, 1960: 53, in key [stem: Plocamotrech-]. Type genus: Plocamotrechus Jeannel, 1926 [syn. of Pachydesus Motschulsky, 1864].
Subtribe
Trechina
Bonelli, 1810
Trechii Bonelli, 1810: Tabula Synoptica [stem: Trech-]. Type genus: Trechus Clairville, 1806.
*Aphaenopses Fauvel, 1888: 7 [stem: Aphoenop-]. Type genus: Aphoenops Bonvouloir, 1861 [as Aphaenops, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Homaloderini Jeannel, 1926: 397, in key [stem: Omaloder-]. Type genus: Omalodera Blanchard, 1842 [as Homalodera, unjustified emendation of type genus name by Gemminger and Harold (1868a: 389), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Trechodina
Jeannel, 1926
Trechodini Jeannel, 1926: 469 [stem: Trechod-]. Type genus: Trechodes Blackburn, 1901.
Thalassophili Csiki, 1928: 233 [stem: Thalassophil-]. Type genus: Thalassophilus Wollaston, 1854.
Tribe
Zolini
Sharp, 1886
Zolini Sharp, 1886a: 371 [stem: Zol-]. Type genus: Zolus Sharp, 1886.
Subtribe
Chalteniina
Roig-Juñent and Cicchino, 2001
Chalteniina Roig-Juñent and Cicchino, 2001: 658 [stem: Chalteni-]. Type genus: Chaltenia Roig-Juñent and Cicchino, 2001.
Subtribe
Sinozolina
Deuve, 1997
Sinozolini Deuve, 1997: 35 [stem: Sinozol-]. Type genus: Sinozolus Bedel, 1898.
Subtribe
Zolina
Sharp, 1886
Zolini Sharp, 1886a: 371 [stem: Zol-]. Type genus: Zolus Sharp, 1886.
Merizodini Sloane, 1920: 139 [stem: Merizodont-]. Type genus: Merizodus Solier, 1849. Comment: incorrect original stem formation, not in prevailing usage.
*Oopterini Jeannel, 1938: 45 [stem: Oopter-]. Type genus: Oopterus Guérin-Méneville, 1841. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Oopterini Jeannel, 1940: 93 [stem: Oopter-]. Type genus: Oopterus Guérin-Méneville, 1841.
Subfamily
Patrobinae
Kirby, 1837
Patrobidae Kirby, 1837: 50 [stem: Patrob-]. Type genus: Patrobus Dejean, 1821.
Tribe
Lissopogonini
Zamotajlov, 2000
Lissopogonini Zamotajlov, 2000: 266 [stem: Lissopogon-]. Type genus: Lissopogonus Andrewes, 1923.
Zolinopatrobina Deuve and Tian, 2001: 421 [stem: Zolinopatrob-]. Type genus: Zolinopatrobus Deuve and Tian, 2001 [syn. of Lissopogonus Andrewes, 1923].
Tribe
Patrobini
Kirby, 1837
Patrobidae Kirby, 1837: 50 [stem: Patrob-]. Type genus: Patrobus Dejean, 1821.
Subtribe
Deltomerina
Chaudoir, 1871
Deltomeridae Chaudoir, 1871a: 51 [stem: Deltomer-]. Type genus: Deltomerus Motschulsky, 1850.
Subtribe
Deltomerodina
Zamotajlov, 2002
Deltomerodina Zamotajlov, 2002: 83 [stem: Deltomerod-]. Type genus: Deltomerodes Deuve, 1992.
Subtribe
Patrobina
Kirby, 1837
Patrobidae Kirby, 1837: 50 [stem: Patrob-]. Type genus: Patrobus Dejean, 1821.
Subtribe
Platidiolina
Zamotajlov and Lafer, 2001
Platidiolini Zamotajlov and Lafer, 2001: 411 [stem: Platidiol-]. Type genus: Platidiolus Chaudoir, 1878.
Subfamily
Psydrinae
LeConte, 1853
Psydri J. L. LeConte, 1853b: 393 [stem: Psydr-]. Type genus: Psydrus J. L. LeConte, 1846.
Tribe
Amblytelini
Blackburn, 1892
Amblytelides Blackburn, 1892: 85 [stem: Amblytel-]. Type genus: Amblytelus Erichson, 1842. Comment: the junior homonym Amblytelinae Viereck, 1918 (type genus Amblyteles Wesmael, 1845) is available in Hymenoptera: Ichneumonidae, it is presently considered a synonym of Ichneumonini Latreille, 1802; both family-group names are based on the same stem; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Mecyclothoracini
Jeannel, 1940
Mecyclothoracitae Jeannel, 1940: 97 [stem: Mecyclothorac-]. Type genus: Mecyclothorax Sharp, 1903. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Lorenz (1998a: 222) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Tribe
Meonini
Sloane, 1898
Meonides Sloane, 1898: 470 [stem: Meon-]. Type genus: Meonis Laporte, 1867. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Meonid-).
Tribe
Moriomorphini
Sloane, 1890
Moriomorphini Sloane, 1890: 646 [stem: Moriomorph-]. Type genus: Moriomorpha Laporte, 1867.
Melisoderides Sloane, 1898: 470 [stem: Melisoder-]. Type genus: Melisodera Westwood, 1835.
Tribe
Psydrini
LeConte, 1853
Psydri J. L. LeConte, 1853b: 393 [stem: Psydr-]. Type genus: Psydrus J. L. LeConte, 1846.
Nomiidae Gozis, 1875: 3 [stem: Nomi-]. Type genus: Nomius Laporte, 1835. Comment: the younger name Nomiidae Robertson, 1904 (type genus Nomia Latreille, 1804) has been used in Hymenoptera and a Case was recently submitted to the Commission to emend the beetle family-group name to Nomiusidae (Art. 55.3) in order to remove the homonymy (Engel and Bouchard 2009).
Tribe
Tropopterini
Sloane, 1898
Tropopterides Sloane, 1898: 470 [stem: Tropopter-]. Type genus: Tropopterus Solier, 1849.
Subfamily
Nototylinae
Bänninger, 1927
Tylonotini Schaum, 1863: 74 [stem: Tylonot-]. Type genus: Tylonotus Schaum, 1863 [preoccupied genus name, not Tylonotus Haldeman, 1847 [Coleoptera: Cerambycidae], not Tylonotus Fieber, 1858 [Hemiptera]; syn. of Nototylus Gemminger and Harold, 1868]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Nototylini Bänninger, 1927: 177 [stem: Nototyl-]. Type genus: Nototylus Gemminger and Harold, 1868.
Subfamily
Paussinae
Latreille, 1806
Paussili Latreille, 1806: 234 [stem: Pauss-]. Type genus: Paussus Linnaeus, 1775.
Tribe
Metriini
LeConte, 1853
Metrii J. L. LeConte, 1853b: 394 [stem: Metri-]. Type genus: Metrius Eschscholtz, 1829.
Tribe
Mystropomini
Horn, 1881
Mystropomini G. H. Horn, 1881: 116 [stem: Mystropom-]. Type genus: Mystropomus Chaudoir, 1848.
Tribe
Ozaenini
Hope, 1838
Ozaenidae Hope, 1838a: 107 [stem: Ozaen-]. Type genus: Ozaena A. G. Olivier, 1812.
*Tropopsitos Solier, 1849: 179 [stem: Tropopse-]. Type genus: Tropopsis Solier, 1849. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Pseudozaenini Sloane, 1905: 704 [stem: Pseudozaen-]. Type genus: Pseudozaena Laporte, 1834.
Eustrini Jeannel, 1946: 48 [stem: Eustr-]. Type genus: Eustra Schmidt-Göbel, 1846.
Pachytelini Jeannel, 1946: 47, in key [stem: Pachytel-]. Type genus: Pachyteles Perty, 1830.
Physeitae Jeannel, 1946: 47, in key [stem: Physe-]. Type genus: Physea Brullé, 1834.
Tribe
Paussini
Latreille, 1806
Paussili Latreille, 1806: 234 [stem: Pauss-]. Type genus: Paussus Linnaeus, 1775.
†Subtribe
Arthropteritina
Luna de Carvalho, 1961
Arthropteritina Luna de Carvalho, 1961: 3 [stem: Arthropterit-]. Type genus: Arthropterites Wasmann, 1926.
Subtribe
Carabidomemnina
Wasmann, 1928
*Carabidomemninen Kolbe, 1927a: 178 [stem: Carabidomemn-]. Type genus: Carabidomemnus Kolbe, 1924. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
*Eohomopterinen Kolbe, 1927b: 214 [stem: Eohomopter-]. Type genus: Eohomopterus Wasmann, 1920. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899; the latinized form Eohomopterini was used by Nagel (1987: 33, 54, 55, 56) but it is unavailable because it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Carabidomemninae Wasmann, 1928: 271 [stem: Carabidomemn-]. Type genus: Carabidomemnus Kolbe, 1924.
Subtribe
Cerapterina
Billberg, 1820
Cerapterides Billberg, 1820a: 47 [stem: Cerapter-]. Type genus: Cerapterus Swederus, 1788.
Megalopaussinae Wasmann, 1920: 111 [stem: Megalopauss-]. Type genus: Megalopaussus Lea, 1906.
*Arthropterinen Kolbe, 1927b: 211 [stem: Arthropter-]. Type genus: Arthropterus W. S. MacLeay, 1838. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Arthropterini Wasmann, 1928: 274 [stem: Arthropter-]. Type genus: Arthropterus W. S. MacLeay, 1838. Comment: Arthropteridae Fieber, 1861 has been used in Hemiptera but it is not based on a genus and is therefore unavailable; Arthropterini Wasmann, 1928 is a junior homonym of Arthropteridae Jordan, 1923 proposed in Pisces (type genus Arthropterus Agassiz, 1843, preoccupied genus name replaced by Arthrobatis Whitley 1940); the Pisces name is permanently invalid since its type genus is a junior homonym; an application to the Commission is needed to conserve the Coleoptera name if it is to be used as valid.
*Mesarthropterina Nagel, 1987: 60 [stem: Mesarthropter-]. Type genus: Mesarthropterus Wasmann, 1926. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
†Subtribe
Eopaussina
Luna de Carvalho, 1951
*Eopaussina Darlington, 1950: 84 [stem: Eopauss-]. Type genus: Eopaussus Wasmann, 1926. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Eopaussinae Luna de Carvalho, 1951: 22 [stem: Eopauss-]. Type genus: Eopaussus Wasmann, 1926.
Subtribe
Heteropaussina
Janssens, 1950
Pleuropterini Wasmann, 1920: 111 [stem: Pleuropter-]. Type genus: Pleuropterus Westwood, 1841 [preoccupied genus name, not Pleuropterus Burnett, 1829 [Mammalia]; syn. of Janssenius Luna de Carvalho, 1951]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Heteropaussines Janssens, 1950: 5 (footnote) [stem: Heteropauss-]. Type genus: Heteropaussus J. Thomson, 1860.
Janssenini Luna de Carvalho, 1951: 18 [stem: Jansseni-]. Type genus: Janssenius Luna de Carvalho, 1951. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Homopterina
Wasmann, 1920
Homopterinae Wasmann, 1920: 111 [stem: Homopter-]. Type genus: Homopterus Westwood, 1841.
Subtribe
Paussina
Latreille, 1806
Paussili Latreille, 1806: 234 [stem: Pauss-]. Type genus: Paussus Linnaeus, 1775.
Platyrhopalini Jeannel, 1946: 65 [stem: Platyrhopal-]. Type genus: Platyrhopalus Westwood, 1833.
Enneapaussini Jeannel, 1946: 62 [stem: Enneapauss-]. Type genus: Enneapaussus Jeannel, 1946.
Ceratoderina Darlington, 1950: 107 [stem: Ceratoder-]. Type genus: Ceratoderus Westwood, 1841.
Hylotorini Luna de Carvalho, 1951: 50 [stem: Hylotor-]. Type genus: Hylotorus Dalman, 1823.
Leleupaussina Luna de Carvalho, 1989: 430 [stem: Leleupauss-]. Type genus: Leleupaussus Luna de Carvalho, 1962.
Subtribe
Pentaplatarthrina
Jeannel, 1946
*Pentaplatarthrinen Kolbe, 1927b: 214 [stem: Pentaplatarthr-]. Type genus: Pentaplatarthrus Westwood, 1833. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Pentaplatarthrini Jeannel, 1946: 65 [stem: Pentaplatarthr-]. Type genus: Pentaplatarthrus Westwood, 1833.
Hexaplatarthrina Luna de Carvalho, 1961: 3 [stem: Hexaplatarthr-]. Type genus: Hexaplatarthrus Jeannel, 1955.
Tribe
Protopaussini
Gestro, 1892
Protopaussini Gestro, 1892: 707 [stem: Protopauss-]. Type genus: Protopaussus Gestro, 1892.
Subfamily
Brachininae
Bonelli, 1810
Brachinii Bonelli, 1810: Tabula Synoptica [stem: Brachin-]. Type genus: Brachinus Weber, 1801.
Tribe
Brachinini
Bonelli, 1810
Brachinii Bonelli, 1810: Tabula Synoptica [stem: Brachin-]. Type genus: Brachinus Weber, 1801.
Subtribe
Aptinina
Gistel, 1848
Aptinidae Gistel, 1848: [2] [stem: Aptin-]. Type genus: Aptinus Bonelli, 1810. Comment: Aptemidae Gistel (1856a: 356) is probably a misspelling of Aptinidae (see Madge 1989: 460).
Styphlomerini Habu, 1984: 133 [stem: Styphlomer-]. Type genus: Styphlomerus Chaudoir, 1875.
Subtribe
Brachinina
Bonelli, 1810
Brachinii Bonelli, 1810: Tabula Synoptica [stem: Brachin-]. Type genus: Brachinus Weber, 1801.
Subtribe
Mastacina
Erwin, 1970
Mastacina Erwin, 1970: 28, 32 [stem: Mastac-]. Type genus: Mastax Fischer von Waldheim, 1828.
Subtribe
Pheropsophina
Jeannel, 1949
Pheropsophini Jeannel, 1949c: 1083, in key [stem: Pheropsoph-]. Type genus: Pheropsophus Solier, 1833.
Tribe
Crepidogastrini
Jeannel, 1949
Crepidogastritae Jeannel, 1949c: 1080 [stem: Crepidogastr-]. Type genus: Crepidogaster Boheman, 1848.
Subfamily
Harpalinae
Bonelli, 1810
Harpalii Bonelli, 1810: Tabula Synoptica [stem: Harpal-]. Type genus: Harpalus Latreille, 1802. Comment: First Reviser (Harpalinae Bonelli, 1810 vs Dryptinae Bonelli, 1810 vs Lebiinae Bonelli, 1810 vs Licininae Bonelli, 1810 vs Panagaeinae Bonelli, 1810 vs Platyninae Bonelli, 1810 vs Pterostichinae Bonelli, 1810 vs Zabrinae Bonelli, 1810 vs Zuphiinae Bonelli, 1810) not determined, current usage maintained; Graphipterinae Latreille, 1802 has precedence over this taxon, however, Harpalinae Bonelli, 1810 is in prevailing usage and is maintained (Art. 35.5).
Tribe
Abacetini
Chaudoir, 1873
Abacétides Chaudoir, 1873a: 5 [stem: Abacet-]. Type genus: Abacetus Dejean, 1828. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1873: 224, as Abacetinae), generally accepted as in Bousquet (2003: 346, as Abacetini).
Celioschesini Jeannel, 1948b: 442 [stem: Celioschese-]. Type genus: Celioschesis Tschitschérine, 1898 [syn. of Aristopus LaFerté-Sénectère, 1853]. Comment: incorrect original stem formation, not in prevailing usage.
Loxandrina Erwin and Sims, 1984: 383, in key [stem: Loxandr-]. Type genus: Loxandrus J. L. LeConte, 1852.
Loxandrini Bousquet and Larochelle, 1993: 31 [stem: Loxandr-]. Type genus: Loxandrus J. L. LeConte, 1852. Comment: family-group name proposed as new without reference to Loxandrina Erwin and Sims, 1984.
Tribe
Amorphomerini
Sloane, 1923
Trimerinae Alluaud, 1922: 500 [stem: Trimer-]. Type genus: Trimerus Chaudoir, 1878 [preoccupied genus name, not Trimerus Green, 1832 [Trilobita]; syn. of Amorphomerus Sloane, 1923]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Amorphomerini Sloane, 1923a: 249 [stem: Amorphomer-]. Type genus: Amorphomerus Sloane, 1923.
Tribe
Anthiini
Bonelli, 1813
Anthies Bonelli, 1813: 18 [stem: Anthi-]. Type genus: Anthia Weber, 1801.
Polyhirmi Rousseau, 1905: 3, in key [stem: Polyhirm-]. Type genus: Polyhirma Chaudoir, 1850.
Cypholobini G. Strohmeyer, 1928: 287 [stem: Cypholob-]. Type genus: Cypholoba Chaudoir, 1850.
Tribe
Atranini
Horn, 1881
Atrani G. H. Horn, 1881: 145 [stem: Atran-]. Type genus: Atranus J. L. LeConte, 1848.
Tribe
Bascanini
Basilewsky, 1953
Bascanini Basilewsky, 1953a: 165 [stem: Bascan-]. Type genus: Bascanus Péringuey, 1896.
Tribe
Calophaenini
Jeannel, 1948
*Calophaenidae Jeannel, 1942: 1017 [stem: Calophaen-]. Type genus: Calophaena Klug, 1821. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Calophaenidae Jeannel, 1948b: 378 [stem: Calophaen-]. Type genus: Calophaena Klug, 1821.
Tribe
Catapieseini
Bates, 1882
Catapiesinae H. W. Bates, 1882: 90 [stem: Catapiese-]. Type genus: Catapiesis Solier, 1835. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Chaetodactylini
Tschitschérine, 1903
Chaetodactylini Tschitschérine, 1903: 157 [stem: Chaetodactyl-]. Type genus: Chaetodactyla Tschitschérine, 1897.
Tribe
Chaetogenyini
Emden, 1958
Chaetogenyina Emden, 1958: 24 [stem: Chaetogeny-]. Type genus: Chaetogenys Emden, 1958.
Tribe
Chlaeniini
Brullé, 1834
Chlaenides Brullé, 1834: 123 [stem: Chlaeni-]. Type genus: Chlaenius Bonelli, 1810.
Subtribe
Callistina
Laporte, 1834
Callistidae Laporte, 1834b: 80 [stem: Callist-]. Type genus: Callistus Bonelli, 1810. Comment: published before 9 August 1834.
Eusynetadae Gistel, 1856a: 356 [stem: Eusynet-]. Type genus: Eusyneta Gistel, 1856 [syn. of Callistus Bonelli, 1810]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Chlaeniina
Brullé, 1834
Chlaenides Brullé, 1834: 123 [stem: Chlaeni-]. Type genus: Chlaenius Bonelli, 1810. Comment: published before 2 August 1834; original vernacular name available (Art. 11.7.2): first used in latinized form by Erichson (1837: 96, as Chlaeniini), generally accepted as in Lorenz (2005: 328, as Chlaeniini); incorrect original stem formation, not in prevailing usage.
Lissaucheniidae Gistel, 1848: [2] [stem: Lissaucheni-]. Type genus: Lissauchenius W. S. MacLeay, 1825.
Rhopalomelini Alluaud, 1930b: 105 [stem: Rhopalomel-]. Type genus: Rhopalomelus Boheman, 1848.
Chlaeniodini Jeannel, 1949c: 777 [stem: Chlaeniod-]. Type genus: Chlaeniodus Jeannel, 1949.
Eccoptomenini Jeannel, 1949c: 821 [stem: Eccoptomen-]. Type genus: Eccoptomenus Chaudoir, 1850.
Chlaenionini Jeannel, 1949c: 776, in key [stem: Chlaenion-]. Type genus: Chlaenionus Kuntzen, 1913.
Procletini Basilewsky, 1950b: 49 [stem: Proclet-]. Type genus: Procletus Péringuey, 1896.
Pleroticini Basilewsky, 1950b: 50 [stem: Plerotic-]. Type genus: Pleroticus Péringuey, 1896.
Callistoidini Basilewsky, 1950b: 51 [stem: Callistoid-]. Type genus: Callistoides Motschulsky, 1865.
Harpaglossini Basilewsky, 1950b: 52 [stem: Harpagloss-]. Type genus: Harpaglossus Motschulsky, 1858.
Leptodinodini Basilewsky and Grundmann, 1955: 205 [stem: Leptodinod-]. Type genus: Leptodinodes Jeannel, 1949.
Chlaenioctenini Basilewsky and Grundmann, 1955: 204 [stem: Chlaeniocten-]. Type genus: Chlaenioctenus H. W. Bates, 1892.
Brachylobini Basilewsky and Grundmann, 1955: 204 [stem: Brachylob-]. Type genus: Brachylobus Chaudoir, 1876.
Tribe
Cnemalobini
Germain, 1911
Cnémacanthides Lacordaire, 1854a: 237 [stem: Cnemacanth-]. Type genus: Cnemacanthus sensu Brullé, 1834 [not Cnemacanthus Gray, 1832; syn. of Cnemalobus Guérin-Méneville, 1838]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Schaum (1860: 353, as Cnemacanthidae), generally accepted as in Broun (1880: 7, as Cnemacanthidae); based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).
Cnemalobini Germain, 1911: 53 [stem: Cnemalob-]. Type genus: Cnemalobus Guérin-Méneville, 1838.
Cnemalobini Bousquet and Larochelle, 1993: 27 [stem: Cnemalob-]. Type genus: Cnemalobus Guérin-Méneville, 1838. Comment: proposed as a new without reference to Cnemalobini Germain, 1911.
Tribe
Cratocerini
Lacordaire, 1854
Cratocérides Lacordaire, 1854a: 257 [stem: Cratocer-]. Type genus: Cratocerus Dejean, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Murray (1858: 343, as Cratoceridae), generally accepted as in Csiki (1929: 493, as Cratoceri).
Tribe
Ctenodactylini
Laporte, 1834
Ctenodactylidae Laporte, 1834b: 45 [stem: Ctenodactyl-]. Type genus: Ctenodactyla Dejean, 1825.
*Leptotrachélides Chaudoir, 1848: 52 [stem: Leptotrachel-]. Type genus: Leptotrachelus Latreille, 1829. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized, also originally proposed in synonymy with Rhagocrepides (see Madge 1989: 464).
Rhagocrepides Chaudoir, 1848: 52 [stem: Rhagocrepid-]. Type genus: Rhagocrepis Eschscholtz, 1829 [syn. of Leptotrachelus Latreille, 1829]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by and generally accepted as in Chaudoir (1861b: 528, as Rhagocrepidae); incorrect original stem formation, not in prevailing usage.
Tribe
Cuneipectini
Sloane, 1907
Cuneipectini Sloane, 1907a: 358 [stem: Cuneipect-]. Type genus: Cuneipectus Sloane, 1907. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Cuneipector-).
Tribe
Cyclosomini
Laporte, 1834
Cyclosomidae Laporte, 1834b: 69 [stem: Cyclosom-]. Type genus: Cyclosomus Latreille, 1829.
Subtribe
Cyclosomina
Laporte, 1834
Cyclosomidae Laporte, 1834b: 69 [stem: Cyclosom-]. Type genus: Cyclosomus Latreille, 1829.
Tétragonodérides Chaudoir, 1871b: 111 [stem: Tetragonoder-]. Type genus: Tetragonoderus Dejean, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by and generally accepted as in Chaudoir (1876a: 2, as Tetragonoderidae).
Sarothrocrepidae Chaudoir, 1876a: 83 [stem: Sarothrocrepid-]. Type genus: Sarothrocrepis Chaudoir, 1850. Comment: incorrect original stem formation, not in prevailing usage; Chaudoir also used the spelling Sarothrocrépides in his original publication on page 80.
Subtribe
Masoreina
Chaudoir, 1871
*Somoplatides Chaudoir, 1846: 511 [stem: Somoplat-]. Type genus: Somoplatus Dejean, 1829. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chaudoir (1846) and generally accepted as valid; this name was first used in latinized form by Carus and Engelmann (1861: 1806 [index], as Somoplatidae) referring to Chaudoir’s paper, but the name was not used as valid; this name was treated as “vernacular, not scientific” by Ball (1979: 77); Basilewsky (1984: 527, as Somoplatini) used this taxon as valid but did not refer to Chaudoir’s original vernacular name, Somoplatini Basilewsky, 1984 is also unavailable since it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); Lorenz (2005: 451) listed this name as as “Somoplatides Chaudoir 1846 [suppr.]”.
Mazoréides Chaudoir, 1871b: 111 [stem: Masore-]. Type genus: Masoreus Dejean, 1821 [implicit use of Mazoreus as the type genus, which is an incorrect subsequent spelling of the type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Chaudoir (1876: 2, as Masoreidae), generally accepted as in Lorenz (2005: 451, as Masoreina); incorrect stem formation, not in prevailing usage.
Corsyrini Ganglbauer, 1891b: 53 [stem: Corsyr-]. Type genus: Corsyra Dejean, 1825.
Aephnidiina Jakobson, 1907: 390 [stem: Aephnidi-]. Type genus: Aephnidius W. S. MacLeay, 1825.
Anaulacini Csiki, 1932b: 1287 [stem: Anaulac-]. Type genus: Anaulacus W. S. MacLeay, 1825. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Blackwelder (1944: 52) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Discopterini Jedlička, 1941: 6, in key [stem: Discopter-]. Type genus: Discoptera Semenov, 1889.
Tribe
Dercylini
Sloane, 1923
Dercylini Sloane, 1923b: 250a [stem: Dercyl-]. Type genus: Dercylus Laporte, 1832.
Dercylidae Jeannel, 1948b: 626 [stem: Dercyl-]. Type genus: Dercylus Laporte, 1832. Comment: family-group name proposed as new without reference to Dercylini Sloane, 1923.
Tribe
Drimostomatini
Chaudoir, 1872
Drimostomidas Chaudoir, 1872a: 283 [stem: Drimostomat-]. Type genus: Drimostoma Dejean, 1830 [syn. of Caelostomus W. S. MacLeay, 1825]. Comment: the original spelling Drimostomidas is the accusative (Latin) form of Drimostomidae and was recognized as such by Madge (1989: 463); original incorrect original stem formation, not in prevailing usage.
Caelostomina Burgeon, 1935: 194 [stem: Caelostom-]. Type genus: Caelostomus W. S. MacLeay, 1825.
Cyrtolaina Whitehead and Ball, 1975: 595 [stem: Cyrtola-]. Type genus: Cyrtolaus H. W. Bates, 1882.
Tribe
Dryptini
Bonelli, 1810
Dryptinae Bonelli, 1810: Tabula Synoptica [stem: Drypt-]. Type genus: Drypta Latreille, 1797.
Tribe
Enoicini
Basilewsky, 1985
Enoicini Basilewsky, 1985: 16 [stem: Enoic-]. Type genus: Enoicus Péringuey, 1896.
Tribe
Galeritini
Kirby, 1825
Galeritidae Kirby, 1825: 564 [stem: Galerit-]. Type genus: Galerita Fabricius, 1801 [Galerita Gouan, 1770 [Aves] has been suppressed both for the Principle of Priority and the Principle of Homonymy (ICZN 1968b); Galerita Fabricius, 1801 placed on the Official List of Generic Names in Zoology (ICZN 1968b)].
Subtribe
Galeritina
Kirby, 1825
Galeritidae Kirby, 1825: 564 [stem: Galerit-]. Type genus: Galerita Fabricius, 1801 [Galerita Gouan, 1770 [Aves] has been suppressed both for the Principle of Priority and the Principle of Homonymy (ICZN 1968b); Galerita Fabricius, 1801 placed on the Official List of Generic Names in Zoology (ICZN 1968b)]. Comment: family-group name previously attributed to J. L. LeConte (1853b) or Lacordaire (1854a); name placed on the Official List of Family-Group Names in Zoology (ICZN 1968b, as Galeritini Lacordaire, 1854).
Galeritinini Jeannel, 1949c: 1057 [stem: Galeritin-]. Type genus: Galeritina Jeannel, 1949 [syn. of Galerita Fabricius, 1801]. Comment: replacement name for “Galeritini Lacordaire, 1854” because of the homonymy of the type genus; name placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1968b).
Galeritulini Jedlička, 1963: 279, in key [stem: Galeritul-]. Type genus: Galeritula Strand, 1936 [syn. of Galerita Fabricius, 1801]. Comment: replacement name for “Galeritini Lacordaire, 1854” because of the homonymy of the type genus; name placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1968b).
Subtribe
Planetina
Jedlička, 1941
Planetini Jedlička, 1941: 7, in key [stem: Planet-]. Type genus: Planetes W. S. MacLeay, 1825.
Tribe
Geobaenini
Péringuey, 1896
Geobaenides Péringuey, 1896: 469 [stem: Geobaen-]. Type genus: Geobaenus Dejean, 1829.
Tribe
Ginemini
Ball and Shpeley, 2002
Ginemini Ball and Shpeley, 2002: 77 [stem: Ginem-]. Type genus: Ginema Ball and Shpeley, 2002.
Tribe
Glyptini
Horn, 1881
Glypti G. H. Horn, 1881: 179 [stem: Glypt-]. Type genus: Glyptus Brullé, 1835 [Glyptus Hoffmannsegg, 1818 placed on the Official Index of Rejected and Invalid Generic Names in Zoology and Glyptus Brullé, 1835 placed on the Official List of Generic Names in Zoology (ICZN 1985b)].
Tribe
Graphipterini
Latreille, 1802
Graphipterides Latreille, 1802: 83 [stem: Graphipter-]. Type genus: Graphipterus Latreille, 1802. Comment: usage of Harpalinae Bonelli, 1810 is maintained over Graphipterinae Latreille, 1802 (Art. 35.5).
Tribe
Harpalini
Bonelli, 1810
Harpalii Bonelli, 1810: Tabula Synoptica [stem: Harpal-]. Type genus: Harpalus Latreille, 1802.
Subtribe
Anisodactylina
Lacordaire, 1854
nomen protectum
Eurytrichini J. L. LeConte, 1847: 376 [stem: Eurytrich-]. Type genus: Eurytrichus J. L. LeConte, 1847 [syn. of Anisotarsus Chaudoir, 1837]. Comment: nomen oblitum (Appendix 1).
Anisodactylides Lacordaire, 1854a: 268 [stem: Anisodactyl-]. Type genus: Anisodactylus Dejean, 1829. Comment: nomen protectum (see Appendix 1); original vernacular name available (Art. 11.7.2): first used in latinized form by Murray (1858: 346, as Anisodactylidae), generally accepted as in Lorenz (2005: 348, as Anisodactylini).
Anisotarsi Csiki, 1932a: 1039 [stem: Anisotars-]. Type genus: Anisotarsus Chaudoir, 1837. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Blackwelder (1944: 46) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
*Geopini Csiki, 1932a: 1026 [stem: Geopin-]. Type genus: Geopinus J. L. LeConte, 1847. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Gnathaphani Schauberger, 1934: 104, 106 [stem: Gnathaphan-]. Type genus: Gnathaphanus W. S. MacLeay, 1825. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Harpalina
Bonelli, 1810
Harpalii Bonelli, 1810: Tabula Synoptica [stem: Harpal-]. Type genus: Harpalus Latreille, 1802. Comment: First Reviser found (Harpalini Bonelli, 1810 vs Ditomini Bonelli, 1810) is Noonan (1976: 28).
Ditomici Bonelli, 1810: Tabula Synoptica [stem: Ditom-]. Type genus: Ditomus Bonelli, 1810.
Acinopidae Laporte, 1834b: 67 [stem: Acinopod-]. Type genus: Acinopus Dejean, 1821. Comment: incorrect original stem formation, not in prevailing usage.
Ophonidae Laporte, 1834b: 68 [stem: Ophon-]. Type genus: Ophonus Dejean, 1821 [placed on the Official List of Generic Names in Zoology (ICZN 1990c)].
Cratognathidae Laporte, 1834b: 70 [stem: Cratognath-]. Type genus: Cratognathus Dejean, 1829.
Stenomorphidae Laporte, 1834b: 71 [stem: Stenomorph-]. Type genus: Stenomorphus Dejean, 1831.
Daptini J. L. LeConte, 1847: 371 [stem: Dapt-]. Type genus: Daptus Fischer von Waldheim, 1823.
Amblystomini Fauvel, 1889: 17 [stem: Amblystom-]. Type genus: Amblystomus Erichson, 1837.
Trichopselaphini Tschitschérine, 1900: 351 [stem: Trichopselaph-]. Type genus: Trichopselaphus Chaudoir, 1843.
Selenophorini Casey, 1914b: 134 [stem: Selenophor-]. Type genus: Selenophorus Dejean, 1829.
Pachycarina Stichel, 1923: 81 [stem: Pachycar-]. Type genus: Pachycarus Solier, 1835.
*Dioctini Csiki, 1932a: 1023a [stem: Dioct-]. Type genus: Dioctes Ménétriés, 1849 [syn. of Machozetus Chaudoir, 1850]. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently.
Diorychi Csiki, 1932a: 1193 [stem: Diorych-]. Type genus: Dioryche W. S. MacLeay, 1825. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Blackwelder (1944: 48) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
*Machozeti Csiki, 1932a: 1023 [stem: Machozet-]. Type genus: Machozetus Chaudoir, 1850. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Bradybaeni Csiki, 1932a: 1187 [stem: Bradybaen-]. Type genus: Bradybaenus Dejean, 1829. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Noonan (1976: 28) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1); Bradybaenidae Pilsbry, 1939 (type genus Bradybaena Beck, 1837) is currently used as valid in Mollusca, both family-group names have a correct stem; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
*Euryderi Csiki, 1932a: 1081 [stem: Euryder-]. Type genus: Euryderus J. L. LeConte, 1846. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Heteracanthi Csiki, 1932a: 1085 [stem: Heteracanth-]. Type genus: Heteracantha Brullé, 1834. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Barysomi Csiki, 1932a: 1192 [stem: Barysom-]. Type genus: Barysomus Dejean, 1829. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Machozeti Schauberger, 1934: 99 [stem: Machozet-]. Type genus: Machozetus Chaudoir, 1850.
Trichotichnini Jeannel, 1942: 624 [stem: Trichotichn-]. Type genus: Trichotichnus Morawitz, 1863.
Bleusei Antoine, 1959: 386 [stem: Bleuse-]. Type genus: Bleusea Bedel, 1896.
Eriotomi Antoine, 1959: 354 [stem: Eriotom-]. Type genus: Eriotomus Piochard de la Brûlerie, 1873 [syn. of Oedesis Motschulsky, 1850].
Granigeri Antoine, 1959: 357 [stem: Graniger-]. Type genus: Graniger Motschulsky, 1864. Comment: an application will need to be submitted to the Commission to suppress Granigerini Bedel, 1900 (based on the misidentified type genus Graniger sensu Chaudoir, 1876) for the Principles of Priority and Homonymy (Art. 65.2.1) if this name is to be used as valid.
Cratacanthi Lindroth, 1968: 742 [stem: Cratacanth-]. Type genus: Cratacanthus Dejean, 1829.
*Carterophonini Jeanne, 1971: 11 [stem: Carterophon-]. Type genus: Carterophonus Ganglbauer, 1891 [syn. of Graniger Motschulsky, 1864]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Pelmatellina
Bates, 1882
Pelmatellinae H. W. Bates, 1882: 67 [stem: Pelmatell-]. Type genus: Pelmatellus H. W. Bates, 1882.
Subtribe
Stenolophina
Kirby, 1837
Stenolophidae Kirby, 1837: 46 [stem: Stelonoph-]. Type genus: Stenolophus Dejean, 1821.
Polpochilinae H. W. Bates, 1891a: 10 [stem: Polpochil-]. Type genus: Polpochila Solier, 1849.
Acupalpini Tschitschérine, 1900: 351 [stem: Acupalp-]. Type genus: Acupalpus Latreille, 1829.
Cratocarini Casey, 1914b: 48 [stem: Cratocar-]. Type genus: Cratocara J. L. LeConte, 1863 [syn. of Polpochila Solier, 1849].
Pachytracheli Csiki, 1932a: 1082 [stem: Pachytrachel-]. Type genus: Pachytrachelus Chaudoir, 1852. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Jeannel (1948b: 720, as Pachytrachelini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
*Agonoderi Csiki, 1932a: 1188 [stem: Agonoder-]. Type genus: Agonoderus Dejean, 1829. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Bradycelli Csiki, 1932a: 1222 [stem: Bradycell-]. Type genus: Bradycellus Erichson, 1837. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Anoplogenii Schauberger, 1937: 272 [stem: Anoplogeni-]. Type genus: Anoplogenius Chaudoir, 1852 [syn. of Loxoncus Schmidt-Göbel, 1846].
*Dichirotrichi F. Burmeister, 1939: 186 [stem: Dicheirotrich-]. Type genus: Dicheirotrichus Jacquelin du Val, 1855 [as Dichirotrichus, unjustified emendation of type genus name by Gemminger and Harold (1868a: 262), not in prevailing usage]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage (see Madge 1989: 463).
Bradycellini Jeannel, 1942: 700 [stem: Bradycell-]. Type genus: Bradycellus Erichson, 1837.
Hippolaetina Basilewsky, 1951b: 258 [stem: Hippoloet-]. Type genus: Hippoloetis Laporte, 1835 [as Hippolaetis, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Anthracini Schuler, 1970: 114 [stem: Anthrac-]. Type genus: Anthracus Motschulsky, 1850.
Tribe
Helluonini
Hope, 1838
Helluonidae Hope, 1838a: 110 [stem: Helluon-]. Type genus: Helluo Bonelli, 1813.
Subtribe
Helluonina
Hope, 1838
Helluonidae Hope, 1838a: 110 [stem: Helluon-]. Type genus: Helluo Bonelli, 1813.
Subtribe
Omphrina
Jedlička, 1941
Omphrini Jedlička, 1941: 6, in key [stem: Omphr-]. Type genus: Omphra Dejean, 1825.
Helluomorphina H. Reichardt, 1974: 226 [stem: Helluomorph-]. Type genus: Helluomorpha Laporte, 1834.
Tribe
Hexagoniini
Horn, 1881 (1834)
Trigonodactyliens Brullé, 1834: 127 [stem: Trigonodactyl-]. Type genus: Trigonodactyla Dejean, 1831 [syn. of Hexagonia Kirby, 1825]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Desmarest (1851: 56, as Trigonodactylidae); use of the younger name Hexagoniini G. H. Horn, 1881 conserved over this name (Art. 40.2).
Hexagoniae G. H. Horn, 1881: 146 [stem: Hexagoni-]. Type genus: Hexagonia Kirby, 1825. Comment: use of family-group name conserved over Trigonodactylini Brullé, 1834 (Art. 40.2).
Tribe
Idiomorphini
Bates, 1891
Idiomorphinae H. W. Bates, 1891b: cccxxxiii [stem: Idiomorph-]. Type genus: Idiomorphus Chaudoir, 1846.
Perochnoristhinae Basilewsky, 1973b: 224 [stem: Perochnoristh-]. Type genus: Perochnoristhus Basilewsky, 1973.
Tribe
Lachnophorini
LeConte, 1853
Lachnophori J. L. LeConte, 1853b: 375 [stem: Lachnophor-]. Type genus: Lachnophorus Dejean, 1831.
Subtribe
Lachnophorina
LeConte, 1853
Lachnophori J. L. LeConte, 1853b: 375 [stem: Lachnophor-]. Type genus: Lachnophorus Dejean, 1831.
Anchonodérides Lacordaire, 1854a: 373 [stem: Anchonoder-]. Type genus: Anchonoderus Reiche, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in H. W. Bates (1871: 30, as Anchonoderinae).
Eucaeri J. L. LeConte, 1861: 22 [stem: Eucaer-]. Type genus: Eucaerus J. L. LeConte, 1853.
Egini G. H. Horn, 1881: 152 [stem: Eg-]. Type genus: Ega Laporte, 1834.
Subtribe
Selinina
Jeannel, 1948
Selinini Jeannel, 1948b: 743, in key [stem: Selin-]. Type genus: Selina Motschulsky, 1858. Comment: the junior homonym Selinina Koch, 1956 (type genus Selinus Mulsant and Rey, 1853) is available in Tenebrionidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Lebiini
Bonelli, 1810
Lebiotae Bonelli, 1810: Tabula Synoptica [stem: Lebi-]. Type genus: Lebia Latreille, 1802. Comment: First Reviser (Lebiini Bonelli, 1810 vs Dromiusini Bonelli, 1810) not determined, current usage maintained.
Subtribe
Actenonycina
Bates, 1871
Actenonycinae H. W. Bates, 1871: 30 [stem: Actenonyc-]. Type genus: Actenonyx A. White, 1846. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Actenonych-).
Subtribe
Agrina
Kirby, 1837
Agridae Kirby, 1837: 13 [stem: Agr-]. Type genus: Agra Fabricius, 1801.
Subtribe
Apenina
Ball, 1983
Apenina Ball, 1983: 516 [stem: Apen-]. Type genus: Apenes J. L. LeConte, 1851.
*Trymosternini Zaballos and Jeanne, 1994: 116 [stem: Trymostern-]. Type genus: Trymosternus Chaudoir, 1873. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Platytarini Zaballos and Jeanne, 1994: 116 [stem: Platytar-]. Type genus: Platytarus Fairmaire, 1850. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Calleidina
Chaudoir, 1873
Callidides Chaudoir, 1873b: 97 [stem: Calleid-]. Type genus: Calleida Dejean, 1824 [as Callida, unjustified emendation of type genus name by Agassiz (1846b: 58), not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1873: 311, as Calleidinae), generally accepted as in Csiki (1932b: 1436, as Callidi [incorrect stem formation]); incorrect stem formation, not in prevailing usage.
Plochionidae Gozis, 1875: 3 [stem: Plochion-]. Type genus: Plochionus Dejean, 1821.
Anomotarina Habu, 1967: 117 [stem: Anomotar-]. Type genus: Anomotarus Chaudoir, 1875.
Subtribe
Celaenephina
Habu, 1982
Celaenephina Habu, 1982: 111 [stem: Celaeneph-]. Type genus: Celaenephes Schmidt-Göbel, 1846.
Subtribe
Cymindidina
Laporte, 1834
Cymindidae Laporte, 1834b: 46 [stem: Cymindid-]. Type genus: Cymindis Latreille, 1806. Comment: incorrect original stem formation, not in prevailing usage.
Taridae Gistel, 1848: [2] [stem: Tar-]. Type genus: Tarus Clairville, 1806 [syn. of Cymindis Latreille, 1806].
Pseudomasoreini Jeannel, 1942: 1039, in key [stem: Pseudomasore-]. Type genus: Pseudomasoreus Desbrochers des Loges, 1904.
Subtribe
Demetriadina
Bates, 1886
Demetriinae H. W. Bates, 1886: 207 [stem: Demetriad-]. Type genus: Demetrias Bonelli, 1810. Comment: incorrect original stem formation, not in prevailing usage.
Peliocypini Basilewsky, 1984: 553 [stem: Peliocypad-]. Type genus: Peliocypas Schmidt-Göbel, 1846. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Dromiusina
Bonelli, 1810
Dromiei Bonelli, 1810: Tabula Synoptica [stem: Dromius-]. Type genus: Dromius Bonelli, 1810 [placed on the Official List of Generic Names in Zoology (ICZN 2006b)]. Comment: Dromiidae Bonelli, 1810 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology, correct stem ruled to be Dromius- and Dromiusidae Bonelli, 1810 placed on the Official List of Family-Group Names in Zoology (ICZN 2006b).
Lichnasthenitae J. Thomson, 1858: 35 [stem: Lichnasthen-]. Type genus: Lichnasthenus J. Thomson, 1858. Comment: J. Thomson (1858: 35) used the spelling Lichnastenitae but corrected the name to Lichnasthenitae in the same publication in the index (page 458) and in the errata (page [472, unn.]), the corrected spelling is considered a justified emendation (see Madge 1989: 464).
Lionychidae Jeannel, 1948b: 378, in key [stem: Lionych-]. Type genus: Lionychus Wissmann, 1846.
Singilini Jeannel, 1949c: 915 [stem: Singil-]. Type genus: Singilis Rambur, 1837.
*Syntomini Jeanne, 1972: 101 [stem: Syntom-]. Type genus: Syntomus Hope, 1838. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Metadromiina Basilewsky, 1984: 545 [stem: Metadromi-]. Type genus: Metadromius Bedel, 1907.
Metaxymorphina Basilewsky, 1984: 551 [stem: Metaxymorph-]. Type genus: Metaxymorphus Chaudoir, 1850.
Singiliomimina Basilewsky, 1984: 552 [stem: Singiliomim-]. Type genus: Singiliomimus Péringuey, 1896.
Subtribe
Gallerucidiina
Chaudoir, 1872
Gallerucidiae Chaudoir, 1872b: 416 [stem: Gallerucidi-]. Type genus: Gallerucidia Chaudoir, 1872.
Lebidiina Jakobson, 1907: 394 [stem: Lebidi-]. Type genus: Lebidia Morawitz, 1862.
Subtribe
Lebiina
Bonelli, 1810
Lebiotae Bonelli, 1810: Tabula Synoptica [stem: Lebi-]. Type genus: Lebia Latreille, 1802.
Encratidae Gistel, 1856a: 355 [stem: Encrat-]. Type genus: Encrates Gistel, 1848 [syn. of Lebia Latreille, 1802].
Lampriadae Chaudoir, 1871b: 115 [stem: Lampri-]. Type genus: Lamprias Bonelli, 1810. Comment: incorrect original stem formation, not in prevailing usage.
Physodérides Chaudoir, 1877: 213 [stem: Physoder-]. Type genus: Physodera Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1883: 207, as Physoderinae), generally accepted as in Lorenz (2005: 489, as Physoderina).
Subtribe
Metallicina
Basilewsky, 1984
Metallicini Basilewsky, 1984: 542 [stem: Metallic-]. Type genus: Metallica Chaudoir, 1872.
Subtribe
Nemotarsina
Bates, 1883
Nemotarsinae H. W. Bates, 1883: 173 [stem: Nemotars-]. Type genus: Nemotarsus J. L. LeConte, 1853.
Subtribe
Pericalina
Hope, 1838
Pericallidae Hope, 1838a: 105 [stem: Perical-]. Type genus: Pericalus W. S. MacLeay, 1825. Comment: incorrect original stem formation, not in prevailing usage.
*Mormolycites Blanchard, 1845a: 366 [stem: Mormolyc-]. Type genus: Mormolyce Hagenbach, 1825. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845a).
Costodérides Chaudoir, 1848: 116 [stem: Coptoder-]. Type genus: Coptodera Dejean, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1871: 30, as Coptoderinae), generally accepted as in Handlirsch (1925: 547, as Coptoderini); incorrect stem formation, not in prevailing usage.
Mormolycidae Desmarest, 1851: 62 [stem: Mormolyc-]. Type genus: Mormolyce Hagenbach, 1825.
Thyréoptérides Chaudoir, 1869: 113 [stem: Thyreopter-]. Type genus: Thyreopterus Dejean, 1831. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Bertkau (1878: 417, as Thyreopterini), generally accepted as in Jeannel (1950b: 165, as Thyreopteridae).
Eucheilinae H. W. Bates, 1883: 168 [stem: Eucheil-]. Type genus: Eucheila Dejean, 1829 [Eucheila is an incorrect subsequent spelling of the original spelling Eucheyla, in prevailing usage and so deemed to be the correct original spelling (Art. 33.3.1)].
Miscelini Sloane, 1907b: 473 [stem: Miscel-]. Type genus: Miscelus Klug, 1834.
Periglossiinae Liebke, 1929: 247 [stem: Periglossi-]. Type genus: Periglossium Liebke, 1929 [syn. of Inna Putzeys, 1861].
Catascopi Csiki, 1932b: 1352 [stem: Catascop-]. Type genus: Catascopus Kirby, 1823. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Blackwelder (1944: 57) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Miscelini Jeannel, 1949c: 1006 [stem: Miscel-]. Type genus: Miscelus Klug, 1834. Comment: family-group name proposed as new without reference to Miscelini Sloane, 1907.
Lobodontini Jeannel, 1949c: 1007, in key [stem: Lobodont-]. Type genus: Lobodontus Chaudoir, 1842.
Thysanotini Jeannel, 1949c: 947, in key [stem: Thysanot-]. Type genus: Thysanotus Chaudoir, 1848.
Somotrichini Mateu, 1963: 125, in key [stem: Somotrich-]. Type genus: Somotrichus Seidlitz, 1887.
Subtribe
Pseudotrechina
Basilewsky, 1984
Pseudotrechini Basilewsky, 1984: 555 [stem: Pseudotrech-]. Type genus: Pseudotrechus Rosenhauer, 1856.
Subtribe
Sugimotoina
Habu, 1975
Sugimotoina Habu, 1975: 77 [stem: Sugimoto-]. Type genus: Sugimotoa Habu, 1975.
Subtribe
Trichina
Basilewsky, 1984
Trichini Basilewsky, 1984: 549 [stem: Trich-]. Type genus: Trichis Klug, 1832.
Tribe
Licinini
Bonelli, 1810
Licinii Bonelli, 1810: Tabula Synoptica [stem: Licin-]. Type genus: Licinus Latreille, 1802.
Subtribe
Dicaelina
Laporte, 1834
Dicoelidae Laporte, 1834b: 83 [stem: Dicael-]. Type genus: Dicaelus Bonelli, 1813 [as Dicoelus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Rembidae Gistel, 1848: [2] [stem: Remb-]. Type genus: Rembus W. S. MacLeay, 1825 [preoccupied genus name, not Rembus Germar, 1824 [Coleoptera: Curculionidae]; syn. of Diplocheila Brullé, 1834]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
*Submerini Lafer, 1989: 205 [stem: Submer-]. Type genus: Submera Habu, 1956. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Dicrochilina
Ball, 1992
Dicrochilina Ball, 1992: 347 [stem: Dicrochil-]. Type genus: Dicrochile Guérin-Méneville, 1846.
Subtribe
Lestignathina
Ball, 1992
Lestignathina Ball, 1992: 347 [stem: Lestignath-]. Type genus: Lestignathus Erichson, 1842.
Subtribe
Licinina
Bonelli, 1810
Licinii Bonelli, 1810: Tabula Synoptica [stem: Licin-]. Type genus: Licinus Latreille, 1802.
Badistidae Gistel, 1856a: 357 [stem: Badister-]. Type genus: Badister Clairville, 1806 [as Badistes, unjustified emendation of type genus name by Agassiz (1846b: 42), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Badistritae Jeannel, 1942: 999 [stem: Badister-]. Type genus: Badister Clairville, 1806 [as Badistes, unjustified emendation of type genus name by Agassiz (1846b: 42), not in prevailing usage]. Comment: family-group name proposed as new without reference to Badistidae Gistel, 1856; incorrect original stem formation, not in prevailing usage.
Tribe
Melanchitonini
Jeannel, 1948
Melanodini Alluaud, 1916: 228 [stem: Melanod-]. Type genus: Melanodes Chaudoir, 1876 [preoccupied type genus, not Melanodes Guenée, 1857 [Lepidoptera]; syn. of Melanchiton Basilewsky, 1946]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Melanchitonitae Jeannel, 1948b: 627 [stem: Melanchiton-]. Type genus: Melanchiton Basilewsky, 1946. Comment: replacement name for Melanodini Alluaud, 1916.
Tribe
Microcheilini
Jeannel, 1948
Microchilitae Jeannel, 1948b: 616 [stem: Microcheil-]. Type genus: Microcheila Brullé, 1834 [as Microchila, unjustified emendation of type genus name by Agassiz (1846b: 233), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Morionini
Brullé, 1835
Morioniens Brullé, 1835: 36 [stem: Morion-]. Type genus: Morion Latreille, 1810. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hope (1838a: 109, as Morionidae [based on “Morio Latreille”, an incorrect subsequent spellling of the type genus name, see Madge (1989: 465)], generally accepted as in Lorenz (2005: 247, as Morionini).
Tribe
Odacanthini
Laporte, 1834
Odacanthidae Laporte, 1834b: 40 [stem: Odacanth-]. Type genus: Odacantha Paykull, 1798.
Casnoniae J. L. LeConte, 1861: 21 [stem: Cosnani-]. Type genus: Cosnania Dejean, 1821 [as Casnonia, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Colliurini Bedel, 1910: 72 [stem: Colliur-]. Type genus: Colliuris DeGeer, 1774.
Lasiocerini Jeannel, 1948b: 747, in key [stem: Lasiocer-]. Type genus: Lasiocera Dejean, 1831.
Tribe
Omphreini
Ganglbauer, 1891
Omphreini Ganglbauer, 1891b: 26 [stem: Omphre-]. Type genus: Omphreus Dejean, 1828.
Tribe
Oodini
LaFerté-Sénectère, 1851
Oodites LaFerté-Sénectère, 1851: 266 [stem: Ood-]. Type genus: Oodes Bonelli, 1810. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Bedel (1879: 52, as Oodini), generally accepted as in Lorenz (2005: 324, as Oodini).
Thryptocerini Jeannel, 1949c: 841 [stem: Thryptocer-]. Type genus: Thryptocerus Chaudoir, 1878.
Sphaerodini Jeannel, 1949c: 829, in key [stem: Sphoerod-]. Type genus: Sphoerodes Chaudoir, 1883 [as Sphaerodes, incorrect subsequent spelling of type genus name, not in prevailaing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Simoini Basilewsky, 1953a: 153 [stem: Simo-]. Type genus: Simous Chaudoir, 1882.
Tribe
Orthogoniini
Schaum, 1857
Orthogoniden Schaum, 1857b: 308 [stem: Orthogoni-]. Type genus: Orthogonius W. S. MacLeay, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Schaum (1860: 774, as Orthogonini [incorrect stem formation]), generally accepted as in Lorenz (2005: 391, as Orthogoniinae); incorrect original stem formation, not in prevailing usage.
Tribe
Panagaeini
Bonelli, 1810
Panagaeides Bonelli, 1810: Tabula Synoptica [stem: Panagae-]. Type genus: Panagaeus Latreille, 1802.
Subtribe
Brachygnathina
Basilewsky, 1946
Brachygnathini Basilewsky, 1946: 7 [stem: Brachygnath-]. Type genus: Brachygnathus Perty, 1830.
Subtribe
Panagaeina
Bonelli, 1810
Panagaeides Bonelli, 1810: Tabula Synoptica [stem: Panagae-]. Type genus: Panagaeus Latreille, 1802.
Subtribe
Tefflina
Basilewsky, 1946
Tefflini Basilewsky, 1946: 7 [stem: Teffl-]. Type genus: Tefflus Leach, 1819.
Tribe
Peleciini
Chaudoir, 1880
Pélécides Chaudoir, 1880: 317 [stem: Peleci-]. Type genus: Pelecium Kirby, 1819.
Subtribe
Agonicina
Sloane, 1920
Agonicini Sloane, 1920: 129 [stem: Agonic-]. Type genus: Agonica Sloane, 1920.
Subtribe
Peleciina
Chaudoir, 1880
Pélécides Chaudoir, 1880: 317 [stem: Peleci-]. Type genus: Pelecium Kirby, 1819. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by G. H. Horn (1881: 170, as Peleciini), generally accepted as in Csiki (1932b: 1285, as Peleciini); incorrect original stem formation, not in prevailing usage.
Disphaericini Sloane, 1923a: 248 [stem: Dispheric-]. Type genus: Disphericus G. R. Waterhouse, 1842 [as Disphaericus, unjustified emendation of type genus name by Agassiz (1846b: 127), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Pentagonicini
Bates, 1873
Pentagonicinae H. W. Bates, 1873: 320 [stem: Pentagonic-]. Type genus: Pentagonica Schmidt-Göbel, 1846.
Scopodinae H. W. Bates, 1874b: 275 [stem: Scopod-]. Type genus: Scopodes Erichson, 1842.
Tribe
Perigonini
Horn, 1881
nomen protectum
Trechichinae H. W. Bates, 1873: 281 [stem: Trechic-]. Type genus: Trechicus J. L. LeConte, 1853 [as Trechichus, incorrect subsequent spelling of type genus name, not in prevailing usage; subgenus of Perigona Laporte, 1835]. Comment: nomen oblitum (see Appendix 1); incorrect original stem formation, not in prevailing usage.
Perigonae G. H. Horn, 1881: 143 [stem: Perigon-]. Type genus: Perigona Laporte, 1835. Comment: nomen protectum (see Appendix 1).
Tribe
Physocrotaphini
Chaudoir, 1863
Physocrotaphides Chaudoir, 1863: 303 [stem: Physocrotaph-]. Type genus: Physocrotaphus Parry, 1849. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1892: 388, as Physocrotaphinae), generally accepted as in B. P. Moore (1998: 369, as Physocrotaphini).
Helluodini Csiki, 1932b: 1571 [stem: Helluod-]. Type genus: Helluodes Westwood, 1847. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Darlington (1968: 222) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Tribe
Platynini
Bonelli, 1810
Platynii Bonelli, 1810: Tabula Synoptica [stem: Platyn-]. Type genus: Platynus Bonelli, 1810. Comment: First Reviser found (Platynina Bonelli, 1810 vs Anchomenina Bonelli, 1810) is Jakobson (1906: 317).
Anchomenii Bonelli, 1810: Tabula Synoptica [stem: Anchomen-]. Type genus: Anchomenus Bonelli, 1810.
Sericodiadae Kirby, 1837: 14 [stem: Sericod-]. Type genus: Sericoda Kirby, 1837.
Agonidae Kirby, 1837: 23 [stem: Agonum-]. Type genus: Agonum Bonelli, 1810 [placed on the Official List of Generic Names in Zoology (ICZN 1996d)]. Comment: senior homonym of Agonidae Swainson, 1839 (type genus Agonus Bloch and Schneider, 1801) in Pisces; Agonidae Kirby, 1837 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1996d), stem emended to Agonum- and Agonumidae Kirby, 1837 placed on the Official List of Family-Group Names in Zoology (ICZN 1996d).
Colpodidas Chaudoir, 1872a: 285 [stem: Colpod-]. Type genus: Colpodes W. S. MacLeay, 1825. Comment: the original spelling Colpodidas is the accusative (Latin) form of and Colpodidae was recognized as such by Madge (1989: 463); Colpodidae Poche, 1913 (type genus Colpoda Müller, 1773) has been used as valid in Protista as recently as in 2006; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Agelaeina Jakobson, 1907: 334 [stem: Agelae-]. Type genus: Agelaea Gené, 1839.
Prosphodrini J. M. Valentine, 1987: 74 [stem: Prosphodr-]. Type genus: Prosphodrus Britton, 1959.
Meleagrosini Morvan, 2004: 2 [stem: Meleagr-]. Type genus: Meleagros Kirschenhofer, 1999. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Pseudomorphini
Hope, 1838
Heteromorphidae Hope, 1838a: 108 [stem: Pseudomorph-]. Type genus: Pseudomorpha Kirby, 1823 [as Heteromorpha, incorrect original spelling of type genus name, not in prevailing usage]. Comment: this family-group name is based on an incorrect original spelling of the name of its type genus (Heteromorpha) and must be corrected (ICZN 1999a: Article 35.4.1); Kirby (1823a) originally used two different names for the type genus, Pseudomorpha (in the description on p. 98) and Heteromorpha (in the explanation of the figures on p. 109). Both names are considered different original spellings of the same name. Kirby reissued the paper later the same year in another journal, without the plate, and so only Pseudomorpha was used; as such, Kirby (1823b) is the “First Reviser” and Pseudomorpha is the correct original spelling (ICZN 1999a: Article 24.2.4), therefore Heteromorphidae Hope, 1838 is corrected to Pseudomorphidae Hope, 1838.
Tribe
Pterostichini
Bonelli, 1810
Pterostichii Bonelli, 1810: Tabula Synoptica [stem: Pterostich-]. Type genus: Pterostichus Bonelli, 1810.
Subtribe
Abacomorphina
Tschitschérine, 1902
Abacomorphini Tschitschérine, 1902: 507 [stem: Abacomorph-]. Type genus: Abacomorphus Chaudoir, 1878. Comment: First Reviser (Abacomorphina Tschitschérine, 1902 vs Sphodrosomina Tschitschérine, 1902) not determined, current usage maintained.
Sphodrosomini Tschitschérine, 1902: 507 [stem: Sphodrosom-]. Type genus: Sphodrosomus Perroud and Montrouzier, 1864.
Subtribe
Euchroina
Chaudoir, 1874
Euchroides Chaudoir, 1874: 16 [stem: Euchro-]. Type genus: Euchroa Brullé, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Tschitschérine (1899: 84, as Euchroini), generally accepted as in Frania and Ball (2007: 12, as Euchroina).
Subtribe
Metiina
Straneo, 1951
Antarctiides Lacordaire, 1854a: 336 [stem: Antarcti-]. Type genus: Antarctia Dejean, 1828 [preoccupied genus name, not Antarctia Hübner, 1820 [Lepidoptera]; syn. of Metius Curtis, 1838]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Reed (1874: 58, as Antarctiinae), generally accepted as in Jeannel (1942: 734, as Antarctiitae); permanently invalid (Art. 39): based on preoccupied type genus.
Metiini Straneo, 1951: 56 [stem: Meti-]. Type genus: Metius Curtis, 1838.
Kuschelinini Jeannel, 1967: 437 [stem: Kuschelin-]. Type genus: Kuschelinus Straneo, 1963.
Subtribe
Microcephalina
Tschitschérine, 1898
Microcephalides Tschitschérine, 1898: 46 [stem: Microcephal-]. Type genus: Microcephalus Dejean, 1828. Comment: the older name Microcephali Latreille, 1825: 345 is unavailable (Art. 11.7.1.1) because it is not based on an available generic name [descriptive name for a group of Staphylinidae including Tachyporus and other genera with relatively small heads].
Tichoniini Semenov, 1904a: 201 [stem: Tichoni-]. Type genus: Tichonia Semenov, 1904 [preoccupied genus name, not Tichonia Hübner, 1826 [Lepidoptera]; syn. of Microcephalus Dejean, 1828]. Comment: replacement name for Microcephalini Tschitschérine, 1898 because of the homonymy of the type genus; permanently invalid (Art. 39): based on preoccupied type genus.
Tichonillina Emden, 1958: 24 [stem: Tichonill-]. Type genus: Tichonilla Strand, 1942 [syn. of Microcephalus Dejean, 1828].
Subtribe
Pterostichina
Bonelli, 1810
Pterostichii Bonelli, 1810: Tabula Synoptica [stem: Pterostich-]. Type genus: Pterostichus Bonelli, 1810. Comment: First Reviser (Pterostichina Bonelli, 1810 vs Poecilina Bonelli, 1810 vs Molopina Bonelli, 1810) not determined, current usage maintained.
Poecilii Bonelli, 1810: Tabula Synoptica [stem: Poecil-]. Type genus: Poecilus Bonelli, 1810.
Molopides Bonelli, 1810: Tabula Synoptica [stem: Molop-]. Type genus: Molops Bonelli, 1810.
Féroniens Dejean, 1825: 3 [stem: Feroni-]. Type genus: Feronia Latreille, 1816 [syn. of Poecilus Bonelli, 1810]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Thon (1827: 15, as Feroniae), generally accepted as in Schaum (1857a: 136, as Feronidae).
Trogonotomidae Laporte, 1834b: 75 [stem: Trigonotom-]. Type genus: Trigonotoma Dejean, 1828. Comment: the original spelling was Trogonotomidae, but this was apparently a typographical error since both the vernacular spelling (Trigonotomites) and the genus name (Trigonotoma) were spelled with the stem “Trigonotom-” on the same page.
Catadromiens Brullé, 1834: 277, 328 [stem: Catadrom-]. Type genus: Catadromus W. S. MacLeay, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Desmarest (1851: 143, as Catadromidae).
Thaliadae Hope, 1838a: 71 [stem: Thali-]. Type genus: Thalia Hope, 1838 [preoccupied genus name, not Thalia Blumenbach, 1827 [Tunicata]; syn. of Poecilus Bonelli, 1810]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Stomidae Chaudoir, 1846: 514 [stem: Stomid-]. Type genus: Stomis Clairville, 1806. Comment: incorrect original stem formation, not in prevailing usage.
Rhathyminae H. W. Bates, 1891b: cccxxxiv [stem: Rathym-]. Type genus: Rathymus Dejean, 1831 [as Rhathymus, unjustified emendation of type genus name by Agassiz (1846b: 321), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Trigonognathides Tschitschérine, 1898: 66 [stem: Trigonognath-]. Type genus: Trigonognatha Motschulsky, 1858.
Platysmatini Tschitschérine, 1899: 84 [stem: Platysmat-]. Type genus: Platysma Bonelli, 1810.
Cyrtoderini Tschitschérine, 1902: 506 [stem: Cyrtoder-]. Type genus: Cyrtoderus Hope, 1842.
Darodiliini Tschitschérine, 1902: 506 [stem: Darodili-]. Type genus: Darodilia Laporte, 1867.
Deliniini Tschitschérine, 1902: 506 [stem: Delini-]. Type genus: Delinius Westwood, 1864.
Cyphosomatini Tschitschérine, 1902: 507 [stem: Cyphosomat-]. Type genus: Cyphosoma Hope, 1842 [preoccupied genus name, not Cyphosoma Mannerheim, 1837 [Coleoptera: Buprestidae]; syn. of Cratogaster Blanchard, 1853]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Myadina Jakobson, 1907: 334 [stem: Myad-]. Type genus: Myas Sturm, 1826.
Cratogastri Csiki, 1929: 524 [stem: Cratogastr-]. Type genus: Cratogaster Blanchard, 1843.
Abaxini Schuler, 1970: 115 [stem: Abac-]. Type genus: Abax Bonelli, 1810. Comment: incorrect original stem formation, not in prevailing usage.
Aristochroodini Sciaky, 1996: 437 [stem: Aristochrood-]. Type genus: Aristochroodes Marcilhac, 1993.
Tribe
Sphodrini
Laporte, 1834
Sphodridae Laporte, 1834b: 78 [stem: Sphodr-]. Type genus: Sphodrus Clairville, 1806. Comment: published before 9 August 1834; First Reviser found (Sphodrini Laporte, 1834 vs Calathini Laporte, 1834) is Habu (1978: 296).
Subtribe
Atranopsina
Baehr, 1982
Atranopsina Baehr, 1982: 265 [stem: Atranops-]. Type genus: Atranopsis Baehr, 1982. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Atranopse-).
*Platyderina Baehr, 1982: 265 [stem: Platyder-]. Type genus: Platyderes Stephens, 1827. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Calathina
Laporte, 1834
Calathidae Laporte, 1834b: 71 [stem: Calath-]. Type genus: Calathus Bonelli, 1810.
Subtribe
Dolichina
Brullé, 1834
Dolichiens Brullé, 1834: 295 [stem: Dolich-]. Type genus: Dolichus Bonelli, 1810. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hope (1838a: 72, as Dolichidae), generally accepted as in Lorenz (2005: 399, as Dolichina); published “31 Dec” 1834, therefore Sphodrini Laporte, 1834 and Calathini Laporte, 1834 have priority over this name for the tribe name.
Subtribe
Pristosiina
Lindroth, 1956
Pristosiae Lindroth, 1956: 489, in key [stem: Pristosi-]. Type genus: Pristosia Motschulsky, 1865.
*Eucalathi Lindroth, 1956: 489, in key [stem: Eucalath-]. Type genus: Eucalathus H. W. Bates, 1883 [syn. of Pristosia Motschulsky, 1865]. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently.
Subtribe
Sphodrina
Laporte, 1834
Sphodridae Laporte, 1834b: 78 [stem: Sphodr-]. Type genus: Sphodrus Clairville, 1806.
Pristonychidae Gistel, 1848: [2] [stem: Pristonych-]. Type genus: Pristonychus Dejean, 1828.
*Laemostenina Baehr, 1982: 265 [stem: Laemosten-]. Type genus: Laemostenus Bonelli, 1810. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Synuchina
Lindroth, 1956
Synuchi Lindroth, 1956: 489, in key [stem: Synuch-]. Type genus: Synuchus Gyllenhal, 1810.
Tribe
Xenaroswellianini
Erwin, 2007
Xenaroswellanini Erwin, 2007: 563 [stem: Xenaroswellian-]. Type genus: Xenaroswelliana Erwin, 2007. Comment: the original spelling Xenaroswellanini is considered a lapsus calami and the stem is corrected to Xenaroswellian- here.
Tribe
Zabrini
Bonelli, 1810
Zabrides Bonelli, 1810: Tabula Synoptica [stem: Zabr-]. Type genus: Zabrus Clairville, 1806.
Subtribe
Amarina
Zimmermann, 1832
Amaroiden C. Zimmermann, 1832: 6 [stem: Amar-]. Type genus: Amara Bonelli, 1810. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Laporte (1834b: 78, as Amaridae), generally accepted as in Hieke (2003: 547, as Amarina).
Isopleuridae Kirby, 1837: 49 [stem: Isopleur-]. Type genus: Isopleurus Kirby, 1837 [syn. of Celia C. Zimmermann, 1832].
Agronomaeidae Gistel, 1848: [2] [stem: Agronom-]. Type genus: Agronoma Gistel, 1848 [Carabus familiaris Duftschmid, 1812, one of the species originally included, is here chosen as type species of Agronoma Gistel, 1848; syn. nov. of Amara Bonelli, 1810]. Comment: syn. nov.; incorrect original stem formation, not in prevailing usage.
Pangeteidae Gistel, 1856a: 358 [stem: Panget-]. Type genus: Pangetes Gistel, 1856 [syn. of Amara Bonelli, 1810]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Zabrina
Bonelli, 1810
Zabrides Bonelli, 1810: Tabula Synoptica [stem: Zabr-]. Type genus: Zabrus Clairville, 1806.
Tribe
Zuphiini
Bonelli, 1810
Zuphietae Bonelli, 1810: Tabula Synoptica [stem: Zuphi-]. Type genus: Zuphium Latreille, 1806.
Subtribe
Dicrodontina
Machado, 1992
Dicrodontini Machado, 1992: 569 [stem: Dicrodont-]. Type genus: Dicrodontus Chaudoir, 1872.
Subtribe
Leleupidiina
Basilewsky, 1951
Leleupidiini Basilewsky, 1951a: 178 [stem: Leleupidi-]. Type genus: Leleupidia Basilewsky, 1951.
Subtribe
Metazuphiina
Mateu, 1992
Metazuphiina Mateu, 1992: 198, in key [stem: Metazuphi-]. Type genus: Metazuphium Mateu, 1992.
Subtribe
Mischocephalina
Mateu, 1992
Mischocephalina Mateu, 1992: 198, in key [stem: Mischocephal-]. Type genus: Mischocephalus Chaudoir, 1863. Comment: this name was incorrectly spelled Mischocephaliina in the key on page 198 but correctly spelled Mischocephalina on p. 203 of the same publication.
Subtribe
Patriziina
Basilewsky, 1953
Patriziini Basilewsky, 1953b: 266 [stem: Patrizi-]. Type genus: Patrizia Alluaud, 1931.
Subtribe
Zuphiina
Bonelli, 1810
Zuphietae Bonelli, 1810: Tabula Synoptica [stem: Zuphi-]. Type genus: Zuphium Latreille, 1806.
*Polystichides Chaudoir, 1863: 308 [stem: Polistich-]. Type genus: Polistichus Bonelli, 1810 [as Polystichus, unjustified emendation of type genus name by Agassiz (1846b: 301), not in prevailing usage]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chaudoir (1863); incorrect original spelling of family-group name, not in prevailing usage.
Polystichinae H. W. Bates, 1886: 199 [stem: Polistich-]. Type genus: Polistichus Bonelli, 1810 [as Polystichus, unjustified emendation of type genus name by Agassiz (1846b: 301), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Family
Haliplidae
Aubé, 1836
Haliplides Aubé, 1836: 15 [stem: Halipl-]. Type genus: Haliplus Latreille, 1802. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Erichson (1837: 183, as Haliplini), generally accepted as in Lawrence and Newton (1995: 807, as Haliplidae).
Peltodytinae Böving and Craighead, 1931: 17 [stem: Peltodyt-]. Type genus: Peltodytes Régimbart, 1879.
Brychiini Ádám, 1996: 15 [stem: Brychi-]. Type genus: Brychius C. G. Thomson, 1859.
†Family
Triaplidae
Ponomarenko, 1977
Triaplidae Ponomarenko, 1977: 17 [stem: Triapl-]. Type genus: Triaplus Ponomarenko, 1977.
†Family
Colymbotethidae
Ponomarenko, 1994
Colymbotethidae Ponomarenko, 1994: 188 [stem: Colymboteth-]. Type genus: Colymbotethis Ponomarenko, 1994.
†Family
Parahygrobiidae
Ponomarenko, 1977
Parahygrobiidae Ponomarenko, 1977: 19 [stem: Parahygrobi-]. Type genus: Parahygrobia Ponomarenko, 1977.
†Family
Coptoclavidae
Ponomarenko, 1961
Coptoclavidae Ponomarenko, 1961: 68 [stem: Coptoclav-]. Type genus: Coptoclava Ping, 1928.
†Subfamily
Necronectinae
Ponomarenko, 1977
Necronectinae Ponomarenko, 1977: 22 [stem: Necronect-]. Type genus: Necronectes Ponomarenko, 1977 [syn. of Timarchopsis Brauer, Redtenbacher and Ganglbauer, 1889].
†Subfamily
Charonoscaphinae
Ponomarenko, 1977
Charonoscaphinae Ponomarenko, 1977: 32 [stem: Charonoscaph-]. Type genus: Charonoscapha Ponomarenko, 1977.
†Subfamily
Coptoclavinae
Ponomarenko, 1961
Coptoclavidae Ponomarenko, 1961: 68 [stem: Coptoclav-]. Type genus: Coptoclava Ping, 1928.
†Subfamily
Coptoclaviscinae
Soriano, Ponomarenko and Delclos, 2007
Coptoclaviscinae Soriano et al., 2007: 532 [stem: Coptoclavisc-]. Type genus: Coptoclavisca Ponomarenko, 1987.
†Subfamily
Hispanoclavinae
Soriano, Ponomarenko and Delclos, 2007
Hispanoclavinae Soriano et al., 2007: 527 [stem: Hispanoclav-]. Type genus: Hispanoclava Soriano, Ponomarenko and Delclos, 2007.
†Family
Liadytidae
Ponomarenko, 1977
Liadytidae Ponomarenko, 1977: 37 [stem: Liadyt-]. Type genus: Liadytes Ponomarenko, 1977.
Family
Meruidae
Spangler and Steiner, 2005
Meruidae Spangler and Steiner, 2005: 351 [stem: Meru-]. Type genus: Meru Spangler and Steiner, 2005.
Family
Noteridae
Thomson, 1860
Noterides C. G. Thomson, 1860: 34 [stem: Noter-]. Type genus: Noterus Clairville, 1806.
Subfamily
Noterinae
Thomson, 1860
Noterides C. G. Thomson, 1860: 34 [stem: Noter-]. Type genus: Noterus Clairville, 1806.
Tribe
Neohydrocoptini
Zalat, Saleh, Angus and Kaschef, 2000
Neohydrocoptini Zalat et al., 2000: 11 [stem: Neohydrocopt-]. Type genus: Neohydrocoptus Satô, 1972.
Tribe
Noterini
Thomson, 1860
Noterides C. G. Thomson, 1860: 34 [stem: Noter-]. Type genus: Noterus Clairville, 1806.
Hydrocanthini Sharp, 1880: cxlviii [stem: Hydrocanth-]. Type genus: Hydrocanthus Say, 1823.
Suphisini Sharp, 1880: cxlviii [stem: Suphid-]. Type genus: Suphis Aubé, 1836. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Pronoterini
Nilsson, 2005
Pronoterini Nilsson, 2005: 90 [stem: Pronoter-]. Type genus: Pronoterus Sharp, 1882.
Tribe
Tonerini
Miller, 2009
Tonerini K. B. Miller, 2009: 201 [stem: Toner-]. Type genus: Tonerus K. B. Miller, 2009.
Subfamily
Notomicrinae
Zimmermann, 1919
Notomicrini A. Zimmermann, 1919: 110 [stem: Notomicr-]. Type genus: Notomicrus Sharp, 1882.
Subfamily
Phreatodytinae
Uéno, 1957
Phreatodytidae Uéno, 1957: 251 [stem: Phreatodyt-]. Type genus: Phreatodytes Uéno, 1957.
Family
Amphizoidae
LeConte, 1853
Amphizoidae J. L. LeConte, 1853a: 227 [stem: Amphizo-]. Type genus: Amphizoa J. L. LeConte, 1853.
Family
Aspidytidae
Ribera, Beutel, Balke and Vogler, 2002
Aspidytidae Ribera et al., 2002: 2354 [stem: Aspidyt-]. Type genus: Aspidytes Ribera, Beutel, Balke and Vogler, 2002.
Family
Hygrobiidae
Régimbart, 1879 (1837)
Pelobiinae Erichson, 1837: 182 [stem: Paelobi-]. Type genus: Paelobius Schönherr, 1808 [as Pelobius, unjustified emendation of type genus name by Erichson (1832: 45); syn. of Hygrobia Latreille, 1804]. Comment: use of younger family-group name Hygrobiidae Régimbart, 1879 conserved (Art. 40.2) (see Lawrence and Newton 1995: 808); incorrect original stem formation, not in prevailing usage.
Hygrobiinae Régimbart, 1879: 449 [stem: Hygrobi-]. Type genus: Hygrobia Latreille, 1804 [the original spelling of the type genus, Hygriobia, has been corrected to Hygrobia and Hygrobia Latreille, 1804 placed on the Official List of Generic Names in Zoology (ICZN 1954)]. Comment: use of this name conserved over the older name Paelobiidae Erichson, 1837 (Art. 40.2) (see Lawrence and Newton 1995: 808).
Family
Dytiscidae
Leach, 1815
Dyticides Leach, 1815: 84 [stem: Dytisc-]. Type genus: Dytiscus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1961e)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1961e, as Dytiscidae Leach, 1817).
Subfamily
Agabinae
Thomson, 1867
Agabides C. G. Thomson, 1867: 84 [stem: Agab-]. Type genus: Agabus Leach, 1817.
Agabinini Crotch, 1873d: 385 [stem: Agabin-]. Type genus: Agabinus Crotch, 1873 [syn. of Platambus C. G. Thomson, 1859].
Ilybii Acloque, 1896: 81 [stem: Ilybi-]. Type genus: Ilybius Erichson, 1832.
Hydronebriini Guignot, 1948: 168, in key [stem: Hydronebri-]. Type genus: Hydronebrius Jakovlev, 1897. Comment: printed in the issue of November 1948; this family-group name was also used in the same year by C. Brinck (1948 [December]: 112, as Hydronebriini).
Hydrotrupinae Roughley, 2000: 52 [stem: Hydrotrup-]. Type genus: Hydrotrupes Sharp, 1882.
Subfamily
Colymbetinae
Erichson, 1837
Colymbetini Erichson, 1837: 149 [stem: Colymbet-]. Type genus: Colymbetes Clairville, 1806.
Tribe
Anisomeriini
Brinck, 1948
Anisomeriini C. Brinck, 1948: 112 [stem: Anisomeri-]. Type genus: Anisomeria C. Brinck, 1943.
Tribe
Carabdytini
Pederzani, 1995
Carabdytini Pederzani, 1995: 45, in key [stem: Carabdyt-]. Type genus: Carabdytes Balke, Hendrich and Wewalka, 1992.
Tribe
Colymbetini
Erichson, 1837
Colymbetini Erichson, 1837: 149 [stem: Colymbet-]. Type genus: Colymbetes Clairville, 1806.
Cymatopterini Portevin, 1929: 198, in key [stem: Cymatopter-]. Type genus: Cymatopterus Dejean, 1833 [syn. of Colymbetes Clairville, 1806].
Subfamily
Copelatinae
Branden, 1885
Copelatini Branden, 1885: 82 [stem: Copelat-]. Type genus: Copelatus Erichson, 1832. Comment: First Reviser (Copelatini Branden, 1885 vs Aglymbini Branden, 1885 vs Lacconectini Branden, 1885) not determined, current usage maintained.
Aglymbini Branden, 1885: 87 [stem: Aglymb-]. Type genus: Aglymbus Sharp, 1882.
Lacconectini Branden, 1885: 87 [stem: Lacconect-]. Type genus: Lacconectus Motschulsky, 1855.
Subfamily
Coptotominae
Branden, 1885
Coptotomini Branden, 1885: 88 [stem: Coptotom-]. Type genus: Coptotomus Say, 1830.
Coptotomini C. Brinck, 1948: 112, 116 [stem: Coptotom-]. Type genus: Coptotomus Say, 1830. Comment: family-group name proposed as new without reference to Coptotomini Branden, 1885.
Subfamily
Dytiscinae
Leach, 1815
Dyticides Leach, 1815: 84 [stem: Dytisc-]. Type genus: Dytiscus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1961e)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1961e, as Dytiscidae Leach, 1817).
Tribe
Aciliini
Thomson, 1867
Aciliides C. G. Thomson, 1867: 84 [stem: Acili-]. Type genus: Acilius Leach, 1817 [placed on the Official List of Generic Names in Zoology (ICZN 1961e)].
Thermonectini Sharp, 1880: cl [stem: Thermonect-]. Type genus: Thermonectus Dejean, 1833.
Graphoderini Houlbert, 1934: 126 [stem: Graphoder-]. Type genus: Graphoderus Dejean, 1833 [placed on the Official List of Generic Names in Zoology (ICZN 1961d)].
Tribe
Aubehydrini
Guignot, 1942
Aubehydrinae Guignot, 1942: 11 [stem: Aubehydr-]. Type genus: Aubehydrus Guignot, 1942 [syn. of Notaticus A. Zimmermann, 1928].
Tribe
Cybisterini
Sharp, 1880
Cybistrini Sharp, 1880: cl [stem: Cybister-]. Type genus: Cybister Curtis, 1827. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Dytiscini
Leach, 1815
Dyticides Leach, 1815: 84 [stem: Dytisc-]. Type genus: Dytiscus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1961e)]. Comment: incorrect original stem formation, not in prevailing usage; name placed on the Official List of Family-Group Names in Zoology (ICZN 1961e, as Dytiscidae Leach, 1817).
Tribe
Eretini
Crotch, 1873
Eretini Crotch, 1873d: 385 [stem: Eret-]. Type genus: Eretes Laporte, 1833.
Eunectini Portevin, 1929: 208 [stem: Eunect-]. Type genus: Eunectes Erichson, 1832 [preoccupied genus name, not Eunectes Wagler, 1830 [Reptilia]; syn. of Eretes Laporte, 1833]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Hydaticini
Sharp, 1880
Hydaticini Sharp, 1880: cl [stem: Hydatic-]. Type genus: Hydaticus Leach, 1817.
Tribe
Hyderodini
Miller, 2000
Hyderodini K. B. Miller, 2000: 175 [stem: Hyderod-]. Type genus: Hyderodes Hope, 1838.
Subfamily
Hydrodytinae
Miller, 2001
Hydrodytinae K. B. Miller, 2001: 76 [stem: Hydrodyt-]. Type genus: Hydrodytes K. B. Miller, 2001.
Subfamily
Hydroporinae
Aubé, 1836
Hydroporides Aubé, 1836: 15 [stem: Hydropor-]. Type genus: Hydroporus Clairville, 1806.
Tribe
Bidessini
Sharp, 1880
Bidessini Sharp, 1880: cxlviii [stem: Bidess-]. Type genus: Bidessus Sharp, 1882.
Tribe
Carabhydrini
Watts, 1978
Carabhydrini Watts, 1978: 26 [stem: Carabhydr-]. Type genus: Carabhydrus Watts, 1978.
Tribe
Hydroporini
Aubé, 1836
Hydroporides Aubé, 1836: 15 [stem: Hydropor-]. Type genus: Hydroporus Clairville, 1806. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Erichson (1837: 166, as Hydroporini), generally accepted as in Nilsson (2003: 54, as Hydroporinae).
Hydrocoptini Branden, 1885: 13 [stem: Hydrocopt-]. Type genus: Hydrocoptus Motschulsky, 1853 [syn. of Hydroporus Clairville, 1806].
Sternopriscini Branden, 1885: 38 [stem: Sternoprisc-]. Type genus: Sternopriscus Sharp, 1882.
Siettitiini Smrz, 1982: 289 [stem: Siettiti-]. Type genus: Siettitia Abeille de Perrin, 1904.
Deronectini Galewski, 1994: 98, in key [stem: Deronect-]. Type genus: Deronectes Sharp, 1882.
Tribe
Hydrovatini
Sharp, 1880
Hydrovatini Sharp, 1880: cxlviii [stem: Hydrovat-]. Type genus: Hydrovatus Motschulsky, 1853.
Tribe
Hygrotini
Portevin, 1929
Hygrotini Portevin, 1929: 180 [stem: Hygrot-]. Type genus: Hygrotus Stephens, 1828.
Tribe
Hyphydrini
Gistel, 1848
Hyphydriidae Gistel, 1848: [2] [stem: Hyphydr-]. Type genus: Hyphydrus Illiger, 1802. Comment: incorrect original stem formation, not in prevailing usage.
Actobaenidae Gistel, 1856a: 355 [stem: Actobaen-]. Type genus: Actobaena Gistel, 1856 [syn. of Hyphydrus Illiger, 1802].
Pachydrini Biström et al., 1997: 66 [stem: Pachydr-]. Type genus: Pachydrus Sharp, 1882.
Tribe
Laccornini
Wolfe and Roughley, 1990
Laccornini Wolfe and Roughley, 1990: 302 [stem: Laccorn-]. Type genus: Laccornis Gozis, 1914. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Laccornith-).
Tribe
Methlini
Branden, 1885
Methlidae Branden, 1885: 65 [stem: Methl-]. Type genus: Methles Sharp, 1882. Comment: precedence (Methlini Branden, 1885 vs Celinini Branden, 1885) given to taxon originally proposed at the higher rank (Art. 24.1).
Celinini Branden, 1885: 65 [stem: Celin-]. Type genus: Celina Aubé, 1837.
†Tribe
Schistomerini
Palmer, 1957
Schistomerini Palmer, 1957: 259 [stem: Schistomer-]. Type genus: Schistomerus Palmer, 1957.
Tribe
Vatellini
Sharp, 1880
Vatellini Sharp, 1880: cxlviii [stem: Vatell-]. Type genus: Vatellus Aubé, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1992)].
Subfamily
Laccophilinae
Gistel, 1848
Laccophilidae Gistel, 1848: [1] [stem: Laccophil-]. Type genus: Laccophilus Leach, 1815.
Tribe
Agabetini
Branden, 1885
Agabetini Branden, 1885: 87 [stem: Agabet-]. Type genus: Agabetes Crotch, 1873.
Tribe
Laccophilini
Gistel, 1848
Laccophilidae Gistel, 1848: [1] [stem: Laccophil-]. Type genus: Laccophilus Leach, 1817.
Subfamily
Lancetinae
Branden, 1885
Lancetini Branden, 1885: 88 [stem: Lancet-]. Type genus: Lancetes Sharp, 1882.
Subfamily
Matinae
Branden, 1885
Matini Branden, 1885: 88 [stem: Mat-]. Type genus: Matus Aubé, 1836.
†Subfamily
Palaeogyrininae
Schlechtendal, 1894
Palaeogyrinidae Schlechtendal, 1894: 200 [stem: Palaeogyrin-]. Type genus: Palaeogyrinus Schlechtendal, 1894.
†Subfamily
Liadytiscinae
Prokin and Ren, 2010
Liadytiscinae Prokin and Ren, 2010: 48 [stem: Liadytisc-]. Type genus: Liadytiscus Prokin and Ren, 2010.
Superfamily
Hydrophiloidea
Latreille, 1802
Hydrophilii Latreille, 1802: 136 [stem: Hydrophil-]. Type genus: Hydrophilus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: First Reviser (Hydrophiloidea Latreille, 1802 vs Sphaeridioidea Latreille, 1802) not determined, current usage maintained.
Family
Hydrophilidae
Latreille, 1802
Hydrophilii Latreille, 1802: 136 [stem: Hydrophil-]. Type genus: Hydrophilus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: First Reviser (Hydrophilidae Latreille, 1802 vs Sphaeridiidae Latreille, 1802) not determined, current usage maintained.
Subfamily
Helophorinae
Leach, 1815
Helopherida Leach, 1815: 95 [stem: Helophor-]. Type genus: Helophorus Fabricius, 1775 [the original spelling Elophorus Fabricius, 1775 was considered an incorrect original spelling and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1993b); Helophorus Fabricius, 1775 was placed on the Official List of Generic Names in Zoology (ICZN 1993b)]. Comment: published in April 1815; incorrect original stem formation, not in prevailing usage; this family-group name was also used in the same year by Rafinesque (1815 [between April 1815 and 21 July 1815]: 112, as Elophoria).
Subfamily
Epimetopinae
Zaitzev, 1908
Epimetopina Zaitzev, 1908: 353 [stem: Epimetop-]. Type genus: Epimetopus Lacordaire, 1854.
Subfamily
Georissinae
Laporte, 1840
Géorissites Laporte, 1840b: 44 [stem: Georiss-]. Type genus: Georissus Latreille, 1809 [placed on the Official List of Generic Names in Zoology (ICZN 1998)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heer (1841a: 471, as Georissida), generally accepted as in Hansen (2004: 42, as Georissidae).
Subfamily
Hydrochinae
Thomson, 1859
Hydrochidae C. G. Thomson, 1859: 15 [stem: Hydroch-]. Type genus: Hydrochus Leach, 1817.
Subfamily
Spercheinae
Erichson, 1837
Spercheini Erichson, 1837: 193 [stem: Sperche-]. Type genus: Spercheus Kugelann, 1798.
Subfamily
Horelophinae
Hansen, 1991
Horelophinae Hansen, 1991: 104 [stem: Horeloph-]. Type genus: Horelophus Orchymont, 1913.
Subfamily
Horelophopsinae
Hansen, 1997
Horelophopsinae Hansen, 1997: 108 [stem: Horelophops-]. Type genus: Horelophopsis Hansen, 1997. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Horelophopse-).
Subfamily
Hydrophilinae
Latreille, 1802
Hydrophilii Latreille, 1802: 136 [stem: Hydrophil-]. Type genus: Hydrophilus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Anacaenini
Hansen, 1991
Anacaenini Hansen, 1991: 129 [stem: Anacaen-]. Type genus: Anacaena C. G. Thomson, 1859.
Tribe
Berosini
Mulsant, 1844
Bérosaires Mulsant, 1844: 97 [stem: Beros-]. Type genus: Berosus Leach, 1817 [placed on the Official List of Generic Names in Zoology (ICZN 1990a)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. G. Thomson (1859: 17, as Berosina), generally accepted as in Hansen (2004: 46, as Berosini).
Derallini Bertrand, 1967: 86, in key [stem: Derall-]. Type genus: Derallus Sharp, 1882.
Tribe
Chaetarthriini
Bedel, 1881
*Cyllidiaires Mulsant, 1844: 143 [stem: Cyllidi-]. Type genus: Cyllidium Erichson, 1837 [syn. of Chaetarthria Stephens, 1835]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Chaetarthriini Bedel, 1881: 314 [stem: Chaetarthri-]. Type genus: Chaetarthria Stephens, 1835.
Amphiopitae Kuwert, 1890: 120 [stem: Amphiop-]. Type genus: Amphiops Erichson, 1843.
Tribe
Hydrophilini
Latreille, 1802
Hydrophilii Latreille, 1802: 136 [stem: Hydrophil-]. Type genus: Hydrophilus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subtribe
Acidocerina
Zaitzev, 1908
*Philhydrates Mulsant, 1844: 131 [stem: Philydr-]. Type genus: Philydrus Solier, 1834 [as Philhydrus, incorrect subsequent spelling of type genus name, not in prevailing usage; preoccupied genus name, not Philydrus Duftschmid, 1805 [Coleoptera: Elmidae]; syn. of Enochrus C. G. Thomson, 1859]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant (1844); incorrect original stem formation, not in prevailing usage.
Philhydrida J. E. LeConte, 1849: 27 [stem: Philydr-]. Type genus: Philydrus Solier, 1834 [as Philhydrus, incorrect subsequent spelling of type genus name, not in prevailing usage; preoccupied genus name, not Philydrus Duftschmid, 1805 [Coleoptera: Elmidae]; syn. of Enochrus C. G. Thomson, 1859]. Comment: incorrect original stem formation, not in prevailing usage; permanently invalid (Art. 39): based on preoccupied type genus.
Helopeltini G. H. Horn, 1873: 118 [stem: Helopelt-]. Type genus: Helopeltis G. H. Horn, 1873 [preoccupied genus name, not Helopeltis Signoret, 1858 [Hemiptera]; syn. of Helobata Bergroth, 1888]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Acidocerini Zaitzev, 1908: 353 [stem: Acidocer-]. Type genus: Acidocerus Klug, 1855.
Helocharae Orchymont, 1919: 147 [stem: Helochar-]. Type genus: Helochares Mulsant, 1844 [placed on the Official List of Generic Names in Zoology (ICZN 1964)]. Comment: senior homonym of Helocharini Metcalf, 1965 in Homoptera (type genus Helochara Fitch, 1851), the homopteran name does not appear to be available (Art. 13.1).
Subtribe
Globuloseina
García, 2001
Globulina García, 2001: 153 [stem: Globulose-]. Type genus: Globulosis García, 2001. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Hydrobiusina
Mulsant, 1844
Hydrobiaires Mulsant, 1844: 116 [stem: Hydrobius-]. Type genus: Hydrobius Leach, 1815 [placed on the Official List of Generic Names in Zoology (ICZN 1990a)]. Comment: spelling emended to Hydrobius- to avoid homonymy with Hydrobiidae Troschel, 1857 in Mollusca (type genus Hydrobia Hartmann, 1821) (ICZN 2003a); Hydrobiusina Mulsant, 1844 placed on the Official List of Family-Group Names in Zoology (ICZN 2003a); Hydrobiina Mulsant, 1844 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 2003a); the family-group name Hydrobia was used earlier by Rafinesque (1815: 110) but since the list of genera did not include Hydrobius, it was probably not based on that genus name.
Subtribe
Hydrophilina
Latreille, 1802
Hydrophilii Latreille, 1802: 136 [stem: Hydrophil-]. Type genus: Hydrophilus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Hydatophilidae Gistel, 1856a: 353 [stem: Hydatophil-]. Type genus: Hydatophilus Gistel, 1856 [syn. of Hydrophilus Geoffroy, 1762]. Comment: published by 18 February 1856; this family-group name was also used in the same year by Gistel (1856b [“31 December”]: 95, as Hydatophilida).
Tribe
Laccobiini
Houlbert, 1922
Laccobiini Houlbert, 1922a: 154, in key [stem: Laccobi-]. Type genus: Laccobius Erichson, 1837. Comment: family-group name previously attributed to Bertrand (1954: 439; see Hansen 2004).
Oocyclini Hansen, 1991: 136 [stem: Oocycl-]. Type genus: Oocyclus Sharp, 1882.
Tribe
Sperchopsini
Hansen, 1991
Sperchopsini Hansen, 1991: 108 [stem: Sperchops-]. Type genus: Sperchopsis J. L. LeConte, 1861. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Sperchopse-).
Subfamily
Sphaeridiinae
Latreille, 1802
Sphaeridiota Latreille, 1802: 135 [stem: Sphaeridi-]. Type genus: Sphaeridium Fabricius, 1775.
Tribe
Andotypini
Hansen, 1991
Andotypini Hansen, 1991: 186 [stem: Andotyp-]. Type genus: Andotypus Spangler, 1979.
Tribe
Borborophorini
Hansen, 1991
Borborophorini Hansen, 1991: 190 [stem: Borborophor-]. Type genus: Borborophorus Hansen, 1990.
Tribe
Coelostomatini
Heyden, 1891 (1890)
*Cyclonotides Motschulsky, 1857: 74 [stem: Cyclonot-]. Type genus: Cyclonotum Erichson, 1837 [syn. of Coelostoma Brullé, 1835]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1857).
Cyclonotides G. H. Horn, 1890: 281 [stem: Cyclonot-]. Type genus: Cyclonotum Erichson, 1837 [syn. of Coelostoma Brullé, 1835]. Comment: use of younger family-group name Coelostomatini L. Heyden, 1891 conserved (Art. 40.2).
Coelostomitae L. Heyden, 1891: 71 [stem: Coelostomat-]. Type genus: Coelostoma Brullé, 1835. Comment: incorrect original stem formation, not in prevailing usage; use of family-group name conserved over Cyclonotini G. H. Horn, 1890 (Art. 40.2) (see Hansen 1991); Hansen (1991: 282) mentions that “Cyclonotaires Rey, 1886” is the oldest name for this tribe but he used Art. 40b of the then Code to conserve usage of Coelostomatini L. Heyden, 1891; we could not find any instance where the requirements of Art. 11.7.2 were met and therefore treat the vernacular name Cyclonotaires as unavailable.
Cyllomina Zaitzev, 1908: 400 [stem: Cylomat-]. Type genus: Cyloma Sharp, 1872 [as Cylloma, unjustified emendation of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Megasternini
Mulsant, 1844
Mégasternaires Mulsant, 1844: 186 [stem: Megastern-]. Type genus: Megasternum Mulsant, 1844 [placed on the Official List of Generic Names in Zoology (ICZN 1981b)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by G. H. Horn (1890: 307, as Megasterni), generally accepted as in Hansen (2004: 61, as Megasternini).
Cercyones G. H. Horn, 1890: 287 [stem: Cercyon-]. Type genus: Cercyon Leach, 1817.
Tribe
Omicrini
Smetana, 1975
Omicrini Smetana, 1975: 155 [stem: Omicr-]. Type genus: Omicrus Sharp, 1879.
Tribe
Protosternini
Hansen, 1991
Protosternini Hansen, 1991: 212 [stem: Protostern-]. Type genus: Protosternum Sharp, 1890.
Tribe
Rygmodini
Orchymont, 1916
Rygmodini Orchymont, 1916: 238 [stem: Rygmod-]. Type genus: Rygmodus A. White, 1846.
Rygmodini Orchymont, 1919: 105 [stem: Rygmod-]. Type genus: Rygmodus A. White, 1846. Comment: family-group name proposed as new without reference to Rygmodini Orchymont, 1916.
Tribe
Sphaeridiini
Latreille, 1802
Sphaeridiota Latreille, 1802: 135 [stem: Sphaeridi-]. Type genus: Sphaeridium Fabricius, 1775. Comment: Latreille also used the incorrect original spelling Sphéridiote on page 138 of the same work.
Tribe
Tormissini
Hansen, 1991
Tormissini Hansen, 1991: 181 [stem: Tormiss-]. Type genus: Tormissus Broun, 1893.
Family
Sphaeritidae
Shuckard, 1839
Sphaeritidae Shuckard, 1839a: 149 [stem: Sphaerit-]. Type genus: Sphaerites Duftschmid, 1805.
Family
Synteliidae
Lewis, 1882
Synteliidae Lewis, 1882: 137 [stem: Synteli-]. Type genus: Syntelia Westwood, 1864.
Family
Histeridae
Gyllenhal, 1808
Histeroides Gyllenhal, 1808: 74 [stem: Hister-]. Type genus: Hister Linnaeus, 1758.
Subfamily
Niponiinae
Fowler, 1912
Niponiidae Fowler, 1912: 93 [stem: Niponi-]. Type genus: Niponius Lewis, 1885.
Subfamily
Abraeinae
MacLeay, 1819
Abreidae W. S. MacLeay, 1819: 25 [stem: Abrae-]. Type genus: Abraeus Leach, 1817.
Tribe
Abraeini
MacLeay, 1819
Abreidae W. S. MacLeay, 1819: 25 [stem: Abrae-]. Type genus: Abraeus Leach, 1817.
Tribe
Acritini
Wenzel, 1944
Acritini Wenzel, 1944: 55, in key [stem: Acrit-]. Type genus: Acritus J. L. LeConte, 1853.
Tribe
Acritomorphini
Wenzel, 1944
Acritomorphini Wenzel, 1944: 55, in key [stem: Acritomorph-]. Type genus: Acritomorphus Wenzel, 1944.
Tribe
Plegaderini
Portevin, 1929
Plegaderini Portevin, 1929: 601, in key [stem: Plegader-]. Type genus: Plegaderus Erichson, 1834.
Tribe
Teretriini
Bickhardt, 1914
Teretriinae Bickhardt, 1914: 306 [stem: Teretri-]. Type genus: Teretrius Erichson, 1834.
Subfamily
Trypeticinae
Bickhardt, 1913
Trypeticinae Bickhardt, 1913: 166 [stem: Trypetic-]. Type genus: Trypeticus Marseul, 1864.
Subfamily
Trypanaeinae
Marseul, 1857
Trypanéens Marseul, 1857b: 148 [stem: Trypanae-]. Type genus: Trypanaeus Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jakobson (1910: 638, as Trypanaeina), generally accepted as in Newton and Thayer (1992: 25, as Trypanaeinae); incorrect stem formation, not in prevailing usage.
Subfamily
Saprininae
Blanchard, 1845
Saprinites Blanchard, 1845a: 276 [stem: Saprin-]. Type genus: Saprinus Erichson, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Fairmaire and Laboulbène (1855: 273, as Saprinii), generally accepted as in Mazur (2004: 90, as Saprininae).
Myrmetini Portevin, 1929: 593 [stem: Myrmet-]. Type genus: Myrmetes Marseul, 1862.
Subfamily
Dendrophilinae
Reitter, 1909
Dendrophilini Reitter, 1909: 288 [stem: Dendrophil-]. Type genus: Dendrophilus Leach, 1817. Comment: First Reviser (Dendrophilinae Reitter, 1909 vs Paromalinae Reitter, 1909) not determined, current usage maintained.
Tribe
Anapleini
Olexa, 1982
Anapleini Olexa, 1982: 38 [stem: Anaple-]. Type genus: Anapleus G. H. Horn, 1873.
Tribe
Bacaniini
Kryzhanovskij, 1976
*Bacaniini Vienna, 1974: 273 [stem: Bacani-]. Type genus: Bacanius J. L. LeConte, 1853. Comment: unavailable name (Art. 13.1): proposed after 1930 without description or bibliographic reference to such a description.
Bacaniini Kryzhanovskij, 1976: 266 [stem: Bacani-]. Type genus: Bacanius J. L. LeConte, 1853.
Tribe
Dendrophilini
Reitter, 1909
Dendrophilini Reitter, 1909: 288 [stem: Dendrophil-]. Type genus: Dendrophilus Leach, 1817.
Tribe
Paromalini
Reitter, 1909
Paromalini Reitter, 1909: 287 [stem: Paromal-]. Type genus: Paromalus Erichson, 1834.
Subfamily
Onthophilinae
MacLeay, 1819
Onthophilidae W. S. MacLeay, 1819: 25 [stem: Onthophil-]. Type genus: Onthophilus Leach, 1817.
Scolytini Jakobson, 1911a: 652 [stem: Scolyt-]. Type genus: Scolytus Müller, 1764 [preoccupied genus name, not Scolytus Geoffroy, 1762 [Coleoptera: Curculionidae]; syn. of Onthophilus Leach, 1817]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Subfamily
Tribalinae
Bickhardt, 1914
Tribalini Bickhardt, 1914: 307 [stem: Tribal-]. Type genus: Tribalus Erichson, 1834.
Subfamily
Histerinae
Gyllenhal, 1808
Histeroides Gyllenhal, 1808: 74 [stem: Hister-]. Type genus: Hister Linnaeus, 1758.
Tribe
Exosternini
Bickhardt, 1914
Exosternini Bickhardt, 1914: 308 [stem: Exostern-]. Type genus: Exosternus Lewis, 1902.
Tribe
Histerini
Gyllenhal, 1808
Histeroides Gyllenhal, 1808: 74 [stem: Hister-]. Type genus: Hister Linnaeus, 1758. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Histr-).
Tribe
Hololeptini
Hope, 1840
Hololeptidae Hope, 1840a: 106 [stem: Hololept-]. Type genus: Hololepta Paykull, 1811. Comment: W. S. MacLeay (1819: 25) published the name Omoleptidae, presumably as a lapsus calami for Hololeptidae, but he did not cite the type genus, nor did he use the family-group name subsequently; in the absence of validating evidence we treat Omoleptidae as a nomen dubium.
Tribe
Omalodini
Kryzhanovskij, 1972
*Omalodini A. N. Reichardt, 1941: 37 [stem: Omalod-]. Type genus: Omalodes Dejean, 1833. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Omalodini Kryzhanovskij, 1972: 19 [stem: Omalod-]. Type genus: Omalodes Dejean, 1833.
Tribe
Platysomatini
Bickhardt, 1914
Platysomini Bickhardt, 1914: 307 [stem: Platysomat-]. Type genus: Platysoma Leach, 1817. Comment: incorrect original stem formation, not in prevailing usage; correction of original spelling by Mazur (1973); the corrected spelling should be used in order to avoid homonymy with Platysomidae Young, 1866 (type genus Platysomus Agassiz, 1833) in Pisces.
Althanini Cooman, 1939: 138 [stem: Althan-]. Type genus: Althanus Lewis, 1903.
Subfamily
Haeteriinae
Marseul, 1857
Hétériens Marseul, 1857b: 148 [stem: Haeteri-]. Type genus: Haeterius Dejean, 1833 [as Hetaerius, incorrect subsequent spelling of type genus name, not in prevailing usage].
Tribe
Haeteriini
Marseul, 1857
Hétériens Marseul, 1857b: 148 [stem: Haeteri-]. Type genus: Haeterius Dejean, 1833 [as Hetaerius, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Schmidt (1885: 283, as Hetaeriini [incorrect stem formation]), generally accepted as in Mazur (2004: 75, as Haeteriinae); incorrect stem formation, not in prevailing usage, correction of original spelling by Bousquet and Laplante (1999: 104).
Tribe
Nymphistrini
Tishechkin, 2007
Nymphisterini Tishechkin, 2007: 51 [stem: Nymphistr-]. Type genus: Nymphister Reichensperger, 1933. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Synoditulini
Tishechkin, 2007
Synoditulini Tishechkin, 2007: 50 [stem: Synoditul-]. Type genus: Synoditulus Reichensperger, 1924.
Subfamily
Chlamydopsinae
Bickhardt, 1914
Chlamydopsini Bickhardt, 1914: 308 [stem: Chlamydops-]. Type genus: Chlamydopsis Westwood, 1869. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Chlamydopse-).
Superfamily
Staphylinoidea
Latreille, 1802
Staphyliniae Latreille, 1802: 124 [stem: Staphylin-]. Type genus: Staphylinus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1959a)]. Comment: First Reviser (Staphylinoidea Latreille, 1802 vs Pselaphoidea Latreille, 1802) not determined, current usage maintained.
Family
Hydraenidae
Mulsant, 1844
Hydraenaires Mulsant, 1844: 50 [stem: Hydraen-]. Type genus: Hydraena Kugelann, 1794.
Subfamily
Orchymontiinae
Perkins, 1997
Orchymontinae Perkins, 1997: 189 [stem: Orchymonti-]. Type genus: Orchymontia Broun, 1919. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Prosthetopinae
Perkins, 1994
Prosthetopinae Perkins, 1994: 7 [stem: Prosthetop-]. Type genus: Prosthetops C. O. Waterhouse, 1879. Comment: precedence (Nucleotopinae Perkins, 1994 vs Parasthetopinae Perkins, 1994 vs Prosthetopinae Perkins, 1994 vs Protosthetopinae Perkins, 1994 vs Pterosthetopinae Perkins, 1994) given to taxon originally proposed at the higher rank (Art. 24.1).
Tribe
Coelometoponini
Perkins, 2005
Coelometoponini Perkins, 2005: 9 [stem: Coelometopon-]. Type genus: Coelometopon Janssens, 1972. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Coelometop-).
Tribe
Nucleotopini
Perkins, 1994
Nucleotopini Perkins, 1994: 8, in key [stem: Nucleotop-]. Type genus: Nucleotops Perkins and Balfour-Browne, 1994.
Tribe
Parasthetopini
Perkins, 1994
Parasthetopini Perkins, 1994: 8, in key [stem: Parasthetop-]. Type genus: Parasthetops Perkins and Balfour-Browne, 1994.
Tribe
Prosthetopini
Perkins, 1994
Prosthetopinae Perkins, 1994: 7 [stem: Prosthetop-]. Type genus: Prosthetops C. O. Waterhouse, 1879.
Tribe
Protosthetopini
Perkins, 1994
Protosthetopini Perkins, 1994: 8, in key [stem: Protosthetop-]. Type genus: Protosthetops Perkins, 1994.
Tribe
Pterosthetopini
Perkins, 1994
Pterosthetopini Perkins, 1994: 9, in key [stem: Pterosthetop-]. Type genus: Pterosthetops Perkins, 1994.
Subfamily
Hydraeninae
Mulsant, 1844
Hydraenaires Mulsant, 1844: 50 [stem: Hydraen-]. Type genus: Hydraena Kugelann, 1794. Comment: First Reviser (Hydraeninae Mulsant, 1844 vs Limnebiinae Mulsant, 1844) not determined, current usage maintained.
Tribe
Hydraenidini
Perkins, 1980
Hydraenidini Perkins, 1980: 414 [stem: Hydraenid-]. Type genus: Hydraenida Germain, 1901.
Tribe
Hydraenini
Mulsant, 1844
Hydraenaires Mulsant, 1844: 50 [stem: Hydraen-]. Type genus: Hydraena Kugelann, 1794. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1848: [1], as Hydraenaeidae [incorrect stem formation]), generally accepted as in Jäch (2004: 102, as Hydraenini).
Tribe
Limnebiini
Mulsant, 1844
Limnébiaires Mulsant, 1844: 88 [stem: Limnebi-]. Type genus: Limnebius Leach, 1815. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. G. Thomson (1859: 14, as Limnebiidae), generally accepted as in Jäch (2004: 110, as Limnebiini).
Tribe
Madagastrini
Perkins, 1997
Madagastrini Perkins, 1997: 176 [stem: Madagastr-]. Type genus: Madagaster Perkins, 1997.
Tribe
Parhydraenini
Perkins, 1997
Parhydraenini Perkins, 1997: 170 [stem: Parhydraen-]. Type genus: Parhydraena Orchymont, 1937.
Subfamily
Ochthebiinae
Thomson, 1859
Ochtebiidae C. G. Thomson, 1859: 15 [stem: Ochthebi-]. Type genus: Ochthebius Leach, 1815 [placed on the Official List of Generic Names in Zoology (ICZN 1991a)].
Tribe
Ochthebiini
Thomson, 1859
Ochtebiidae C. G. Thomson, 1859: 15 [stem: Ochthebi-]. Type genus: Ochthebius Leach, 1815 [placed on the Official List of Generic Names in Zoology (ICZN 1991a)].
Subtribe
Enicocerina
Perkins, 1997
Enicocerina Perkins, 1997: 157 [stem: Enicocer-]. Type genus: Enicocerus Stephens, 1829.
Subtribe
Meropathina
Perkins, 1997
Meropathina Perkins, 1997: 143 [stem: Meropath-]. Type genus: Meropathus Enderlein, 1901.
Subtribe
Neochthebiina
Perkins, 1997
Neochthebina Perkins, 1997: 152 [stem: Neochthebi-]. Type genus: Neochthebius Orchymont, 1932. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Ochthebiina
Thomson, 1859
Ochtebiidae C. G. Thomson, 1859: 15 [stem: Ochthebi-]. Type genus: Ochthebius Leach, 1815 [as Ochtebius, incorrect subsequent spelling of type genus name, not in prevailing usage; Ochthebius placed on the Official List of Generic Names in Zoology (ICZN 1991a)]. Comment: incorrect original stem formation, not in prevailing usage.
Ochthebiinae Perkins, 1980: 414 [stem: Ochthebi-]. Type genus: Ochthebius Leach, 1815 [placed on the Official List of Generic Names in Zoology (ICZN 1991a)]. Comment: family-group name proposed as new without reference to Ochtebiidae C. G. Thomson, 1859.
Subtribe
Protochthebiina
Perkins, 1997
Protochthebiina Perkins, 1997: 152 [stem: Protochthebi-]. Type genus: Protochthebius Perkins, 1997.
Tribe
Ochtheosini
Perkins, 1997
Ochtheosini Perkins, 1997: 123 [stem: Ochtheos-]. Type genus: Ochtheosus Perkins, 1997.
Family
Ptiliidae
Erichson, 1845
Ptilina Erichson, 1845: 15 [stem: Ptili-]. Type genus: Ptilium Gyllenhal, 1827 [placed on the Official List of Generic Names in Zoology (ICZN 1984b)].
Subfamily
Ptiliinae
Erichson, 1845
Ptilina Erichson, 1845: 15 [stem: Ptili-]. Type genus: Ptilium Gyllenhal, 1827 [placed on the Official List of Generic Names in Zoology (ICZN 1984b)].
Tribe
Discheramocephalini
Grebennikov, 2009
Discheramocephalini Grebennikov, 2009: 115 [stem: Discheramocephal-]. Type genus: Discheramocephalus C. Johnson, 2007.
Tribe
Nanosellini
Barber, 1924
Nanosellinae Barber, 1924: 170 [stem: Nanosell-]. Type genus: Nanosella Motschulsky, 1869.
Tribe
Ptenidiini
Flach, 1889
Ptenidiini Flach, 1889: 489 [stem: Ptenidi-]. Type genus: Ptenidium Erichson, 1845 [placed on the Official List of Generic Names in Zoology and given precedence over the generic name Anisarthria Stephens, 1830 whenever the two names are considered synonyms (ICZN 1984b)].
Tribe
Ptiliini
Erichson, 1845
*Ptilina Heer, 1843: 60 [stem: Ptili-]. Type genus: Ptilium Gyllenhal, 1827 [as a synonym of Trichopteryx Kirby, 1826; Ptilium Gyllenhal, 1827 placed on the Official List of Generic Names in Zoology (ICZN 1984b)]. Comment: family-group name unavailable (Art. 11.7.1.1): not based on a genus used as valid at the time; incorrect original stem formation, not in prevailing usage.
Ptilina Erichson, 1845: 15 [stem: Ptili-]. Type genus: Ptilium Gyllenhal, 1827 [placed on the Official List of Generic Names in Zoology (ICZN 1984b)]. Comment: published 28 May 1845; incorrect original stem formation, not in prevailing usage; this family-group name was also used in the same year by Motschulsky (1845 [after 6 August]: 504, as Ptilien); Motschulsky’s name, originally proposed in a vernacular form, was treated as available by Lawrence and Newton (1995: 818).
Ptilopteriidae Gistel, 1848: [4] [stem: Ptilopteri-]. Type genus: Ptilopterium Gistel, 1848 [syn. of Ptilium Gyllenhal, 1827].
Actidiini Portevin, 1929: 568, in key [stem: Actidi-]. Type genus: Actidium A. Matthews, 1868.
Tribe
Ptinellini
Reitter, 1906 (1891)
Neuglenini Reitter, 1891: 146 [stem: Neuglen-]. Type genus: Neuglenes C. G. Thomson, 1859 [syn. of Ptinella Motschulsky, 1844].
Ptinellini Reitter, 1906: 259 [stem: Ptinell-]. Type genus: Ptinella Motschulsky, 1844 [placed on the Official List of Generic Names in Zoology (ICZN 1985b)]. Comment: name proposed to replace Neuglenini Reitter, 1891 because of the synonymy of the type genus; use of younger family-group name Ptinellini Reitter, 1906 conserved (Art. 40.2).
*Pterycini Dybas, 1966: 16, 44 [stem: Pteryg-]. Type genus: Pteryx A. Matthews, 1858. Comment: incorrect original stem formation, not in prevailing usage; the earlier usage of “pterycine group” by Dybas (1955: 562) is unavailable because it is not a noun (Art. 11.7.1.1); also this name has been used subsequently by Hall (2003: 85) but the name Pterycini has not been made available yet (see Hall 2005: 257).
Subfamily
Cephaloplectinae
Sharp, 1883
Cephaloplectinae Sharp, 1883: 295 [stem: Cephaloplect-]. Type genus: Cephaloplectus Sharp, 1883.
Limulodinae Ganglbauer, 1898: 297 [stem: Limulod-]. Type genus: Limulodes A. Matthews, 1866.
Subfamily
Acrotrichinae
Reitter, 1909 (1856)
Trichopterygia Erichson, 1845: 13 [stem: Trichopteryg-]. Type genus: Trichopteryx Kirby, 1826 [preoccupied genus name, not Trichopteryx Hübner, 1825 [Lepidoptera]; syn. of Acrotrichis Motschulsky, 1848]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Cleopteriidae Gistel, 1856a: 360 [stem: Cleopteri-]. Type genus: Cleopterium Gistel, 1856 [syn. of Acrotrichis Motschulsky, 1848]. Comment: use of younger family-group name Acrotrichinae Reitter, 1909 conserved (Art. 40.2) (see Newton and Thayer 1992: 44).
Acrotrichini Reitter, 1909: 272 [stem: Acrotrich-]. Type genus: Acrotrichis Motschulsky, 1848. Comment: use of family-group name conserved over Cleopteriinae Gistel, 1856 (Art. 40.2) (see Newton and Thayer 1992: 44).
Nephanini Portevin, 1929: 573, in key [stem: Nephan-]. Type genus: Nephanes C. G. Thomson, 1859 [placed on the Official List of Generic Names in Zoology (ICZN 1985b)].
Family
Agyrtidae
Thomson, 1859
Agyrtidae C. G. Thomson, 1859: 57 [stem: Agyrt-]. Type genus: Agyrtes Frölich, 1799.
Subfamily
Agyrtinae
Thomson, 1859
*Agyrtes Motschulsky, 1849: 57 [stem: Agyrt-]. Type genus: Agyrtes Frölich, 1799. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Agyrtidae C. G. Thomson, 1859: 57 [stem: Agyrt-]. Type genus: Agyrtes Frölich, 1799.
Lyrosomini G. H. Horn, 1880b: 247 [stem: Lyrosomat-]. Type genus: Lyrosoma Mannerheim, 1853. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 33).
Lendomini Bradley, 1930: 159, in key [stem: Lendom-]. Type genus: Lendomus Casey, 1924 [syn. of Agyrtes Frölich, 1799].
Subfamily
Necrophilinae
Newton, 1997
*Necrophilini Jeannel, 1936: 10 [stem: Necrophil-]. Type genus: Necrophilus Latreille, 1829. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Necrophilinae Newton, 1997: 127 [stem: Necrophil-]. Type genus: Necrophilus Latreille, 1829. Comment: the type genus of the older names Necrophilidae Gistel, 1848: [4] and Necrophilidae Gistel, 1856a: 362 cannot be determined (either Necrophilus Latreille, 1829 [Agyrtidae] or Necrophila Kirby and Spence, 1828 [Silphidae]) and therefore those names are considered unavailable (also see comments in Newton and Thayer (1992: 23) and Newton (1997: 147)).
Subfamily
Pterolomatinae
Thomson, 1862
Pterolomina C. G. Thomson, 1862: 20 [stem: Pterolomat-]. Type genus: Pteroloma Gyllenhal, 1827. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 33).
Family
Leiodidae
Fleming, 1821
Leiodesidae Fleming, 1821: 51 [stem: Leiod-]. Type genus: Leiodes Latreille, 1797. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Camiarinae
Jeannel, 1911
Camiarinae Jeannel, 1911: 192, in key [stem: Camiar-]. Type genus: Camiarus Sharp, 1878.
Tribe
Agyrtodini
Jeannel, 1936
Agyrtodini Jeannel, 1936: 99 [stem: Agyrtod-]. Type genus: Agyrtodes Portevin, 1907.
*†Guvanocoleina Perkovsky, 2002: 16 [stem: Gurvanocole-]. Type genus: Gurvanocoleus Ponomarenko, 1986. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Tribe
Camiarini
Jeannel, 1911
Camiarinae Jeannel, 1911: 192, in key [stem: Camiar-]. Type genus: Camiarus Sharp, 1878.
*†Mesecanina Perkovsky, 2002: 15 [stem: Mesecan-]. Type genus: Mesecanus Newton, 1982. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Neopelatopini
Jeannel, 1962
Neopelatopini Jeannel, 1962b: 487 [stem: Neopelatop-]. Type genus: Neopelatops Jeannel, 1936.
Subfamily
Catopocerinae
Hatch, 1927 (1880)
Catopocerini Hatch, 1927: 4, in key [stem: Catopocer-]. Type genus: Catopocerus Motschulsky, 1869. Comment: use of family-group name conserved over Pinodytinae G. H. Horn, 1880 (Art. 40.2) (see Newton and Thayer 1992: 34).
Tribe
Catopocerini
Hatch, 1927 (1880)
Pinodytini G. H. Horn, 1880b: 248 [stem: Pinodyt-]. Type genus: Pinodytes G. H. Horn, 1880 [syn. of Catopocerus Motschulsky, 1869]. Comment: use of younger family-group name Catopocerini Hatch, 1927 conserved (Art. 40.2) (see Newton and Thayer 1992: 34).
Catopocerini Hatch, 1927: 4, in key [stem: Catopocer-]. Type genus: Catopocerus Motschulsky, 1869. Comment: new name for Pinodytini G. H. Horn, 1880 because of synonymy of type genus; use of family-group name conserved over Pinodytini G. H. Horn, 1880 (Art. 40.2) (see Newton and Thayer 1992: 34).
Tribe
Glacicavicolini
Westcott, 1968
Glacicavicolinae Westcott, 1968: 1 [stem: Glacicavicol-]. Type genus: Glacicavicola Westcott, 1968.
Subfamily
Leiodinae
Fleming, 1821
Leiodesidae Fleming, 1821: 51 [stem: Leiod-]. Type genus: Leiodes Latreille, 1797.
Tribe
Agathidiini
Westwood, 1838
*Anisotomidae Stephens, 1828: 99 [stem: Anisotom-]. Type genus: Anisotoma Panzer, 1797. Comment: family-group name unavailable (Art. 11.7.1.1): not based on a genus used as valid at the time; see Newton and Thayer (1992: 20).
Agathidiidae Westwood, 1838: 10 [stem: Agathidi-]. Type genus: Agathidium Illiger, 1798.
Anisotomidae Reitter, 1884: 6 [stem: Anisotom-]. Type genus: Anisotoma Panzer, 1797. Comment: an application will need to be submitted to the Commission to suppress Anisotomidae Erichson, 1845 (based on the misidentified type genus Anisotoma sensu Schmidt, 1841) for the Principles of Priority and Homonymy (Art. 65.2.1) if this name is to be used as valid.
Tribe
Estadiini
Portevin, 1914
Estadiini Portevin, 1914: 199 [stem: Estadi-]. Type genus: Estadia Fairmaire, 1903 [syn. of Dietta Sharp, 1876].
Tribe
Leiodini
Fleming, 1821
Leiodesidae Fleming, 1821: 51 [stem: Leiod-]. Type genus: Leiodes Latreille, 1797. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 34).
Anisotomidae Erichson, 1845: 41 [stem: Anisotom-]. Type genus: Anisotoma sensu Schmidt, 1841 [not Anisotoma Panzer, 1797; syn. of Leiodes Latreille, 1797]. Comment: based on a misidentified type genus; an application will need to be submitted to the Commission to suppress this name for the Principles of Priority and Homonymy (Art. 65.2.1) if Anisotomidae Reitter, 1884 in Leiodinae: Agathidiini is to be used as valid in the future.
*Cyrtusina Perkovsky, 1997b: 168 [stem: Cyrtus-]. Type genus: Cyrtusa Erichson, 1842. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Hypoliodina Perkovsky, 2002: 15 [stem: Hypoliod-]. Type genus: Hypoliodes Portevin, 1908. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Pseudoliodini
Portevin, 1926
Pseudoliodini Portevin, 1926: 75 [stem: Pseudoliod-]. Type genus: Pseudoliodes Portevin, 1926 [syn. of Pseudcolenis Reitter, 1884].
Dermatohomoeini Hlisnikovský, 1963: 311 [stem: Dermatohomoe-]. Type genus: Dermatohomoeus Hlisnikovský, 1963.
Tribe
Scotocryptini
Reitter, 1884
Scotocryptini Reitter, 1884: 91 [stem: Scotocrypt-]. Type genus: Scotocryptus Girard, 1874.
Tribe
Sogdini
Lopatin, 1961
Sogdiidae Lopatin, 1961: 121 [stem: Sogd-]. Type genus: Sogda Lopatin, 1961. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 36).
Triarthrini Jeannel, 1962b: 486, in key [stem: Triarthr-]. Type genus: Triarthron Märkel, 1840. Comment: junior homonym of Triarthridae Ulrich, 1930 (type genus Triarthrus Green, 1832) in Trilobita; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Hydnobiini Jeannel, 1962b: 492 [stem: Hydnobi-]. Type genus: Hydnobius Schmidt, 1841.
*Stereina Perkovsky, 2002: 15 [stem: Stere-]. Type genus: Stereus Wollaston, 1857 [preoccupied genus name, not Stereus Mannerheim, 1846 [Coleoptera: Curculionidae]; syn. of Deltocnemis Sahlberg, 1886]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subfamily
Coloninae
Horn, 1880 (1859)
Myloechina C. G. Thomson, 1859: 60 [stem: Myloech-]. Type genus: Myloechus Latreille, 1807 [syn. of Colon Herbst, 1797]. Comment: use of younger family-group name Coloninae G. H. Horn, 1880 conserved (Art. 40.2); see Newton and Thayer (1992: 36).
Colones G. H. Horn, 1880b: 266 [stem: Colon-]. Type genus: Colon Herbst, 1797 [the original spelling Kolon was placed on the Official Index of Rejected and Suppressed Generic Names in Zoology, Colon was considered the correct original spelling of the genus and placed on the Official List of Generic Names in Zoology (ICZN 1995b)]. Comment: use of family-group name conserved over Myloechinae C. G. Thomson, 1859 (Art. 40.2); see Newton and Thayer (1992: 36).
Subfamily
Cholevinae
Kirby, 1837
Cholevidae Kirby, 1837: 108 [stem: Cholev-]. Type genus: Choleva Latreille, 1797.
Tribe
Anemadini
Hatch, 1928
Anemadina Hatch, 1928: 159 [stem: Anemad-]. Type genus: Anemadus Reitter, 1884.
Subtribe
Anemadina
Hatch, 1928
Anemadina Hatch, 1928: 159 [stem: Anemad-]. Type genus: Anemadus Reitter, 1884.
Anemadinae Jeannel, 1936: 179 [stem: Anemad-]. Type genus: Anemadus Reitter, 1884. Comment: family-group name proposed as new without reference to Anemadina Hatch, 1928.
Subtribe
Eocatopina
Jeannel, 1936
Eocatopina Jeannel, 1936: 124, in key [stem: Eocatop-]. Type genus: Eocatops Peyerimhoff, 1924.
Subtribe
Eunemadina
Newton, 1998
Eunemadina Newton, 1998: 103 [stem: Eunemad-]. Type genus: Eunemadus Portevin, 1914.
Subtribe
Nemadina
Jeannel, 1936
Nemadinae Jeannel, 1936: 96 [stem: Nemad-]. Type genus: Nemadus C. G. Thomson, 1867.
Subtribe
Paracatopina
Jeannel, 1936
Paracatopini Jeannel, 1936: 181 [stem: Paracatop-]. Type genus: Paracatops Portevin, 1907.
Tribe
Cholevini
Kirby, 1837
Cholevidae Kirby, 1837: 108 [stem: Cholev-]. Type genus: Choleva Latreille, 1797.
Subtribe
Catopina
Chaudoir, 1845
Catopides Chaudoir, 1845: 195 [stem: Catop-]. Type genus: Catops Paykull, 1798. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 68, as Catopidae), generally accepted as in Perreau (2004: 136, as Catopina).
Subtribe
Cholevina
Kirby, 1837
Cholevidae Kirby, 1837: 108 [stem: Cholev-]. Type genus: Choleva Latreille, 1797.
Tribe
Eucatopini
Jeannel, 1921
Eucatopini Jeannel, 1921: 233 [stem: Eucatop-]. Type genus: Eucatops Portevin, 1903.
Tribe
Leptodirini
Lacordaire, 1854 (1849)
Leptodérides Lacordaire, 1854b: 195 [stem: Leptodir-]. Type genus: Leptodirus Schmidt, 1832 [as Leptoderus, unjustified emendation of type genus name by F. Schmidt (1852), not in prevailing usage]. Comment: use of family-group name conserved over Stagobiini Schiødte, 1849 (Art. 40.2); see Newton and Thayer (1992: 35).
Subtribe
Anthroherponina
Jeannel, 1910
Antroherpona Jeannel, 1910: 25 [stem: Antroherpon-]. Type genus: Anthroherpon Reitter, 1889 [as Antroherpon, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Hadesiini Absolon, 1913: 108 [stem: Hadesi-]. Type genus: Hadesia J. Müller, 1911.
Antroherponina Guéorguiev, 1974: 841, in key [stem: Antroherpon-]. Type genus: Anthroherpon Reitter, 1889 [as Antroherpon, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: family-group name proposed as new without reference to Antroherpona Jeannel, 1910.
Subtribe
Bathysciina
Horn, 1880
Bathysciae G. H. Horn, 1880b: 251 [stem: Bathysci-]. Type genus: Bathyscia Schiødte, 1847.
Oriotini Reitter, 1889b: 296 [stem: Oryot-]. Type genus: Oryotus Miller, 1856 [as Oriotus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Bathysciina Guéorguiev, 1974: 841, in key [stem: Bathysci-]. Type genus: Bathyscia Schiødte, 1847. Comment: family-group name proposed as new without reference to Bathysciae G. H. Horn, 1880.
Neotropospeonellina Perkovsky, 1997a: 59 [stem: Neotropospeonell-]. Type genus: Neotropospeonella Pace, 1985 [syn. of Oryotus Miller, 1856].
*Aphaobiina Perkovsky, 2002: 16 [stem: Aphaobi-]. Type genus: Aphaobius Abeille de Perrin, 1878. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Bathysciotina
Guéorguiev, 1974
Bathysciotina Guéorguiev, 1974: 841, in key [stem: Bathysciot-]. Type genus: Bathysciotes Jeannel, 1910.
Subtribe
Leptodirina
Lacordaire, 1854 (1849)
Stagobiinae Schiødte, 1849: 16 [stem: Stagobi-]. Type genus: Stagobius Schiødte, 1847 [syn. of Leptodirus Schmidt, 1832]. Comment: use of younger family-group name Leptodirina Lacordaire, 1854 conserved (Art. 40.2); see Newton and Thayer (1992: 35).
Leptodérides Lacordaire, 1854b: 195 [stem: Leptodir-]. Type genus: Leptodirus Schmidt, 1832 [as Leptoderus, unjustified emendation of type genus name by Schmidt (1852), not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kraatz (1859a: 35, as Leptoderidae), generally accepted as in Perreau (2004: 147, as Leptodirini); use of family-group name conserved over Stagobiina Schiødte, 1849 (Art. 40.2); see Newton and Thayer (1992: 35); incorrect original stem formation, not in prevailing usage.
Leptodirina Guéorguiev, 1974: 841, in key [stem: Leptodir-]. Type genus: Leptodirus Schmidt, 1832. Comment: family-group name proposed as new without reference to Leptodérides Lacordaire, 1854.
*Laneyriellina Perkovsky, 2002: 16 [stem: Laneyriell-]. Type genus: Laneyriella Guéorguiev, 1976. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Pholeuina
Reitter, 1886
Pholeuones Reitter, 1886: 314 [stem: Pholeu-]. Type genus: Pholeuon Hampe, 1856. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 35); current spelling maintained (Art. 29.5): corrected stem formation in prevailing usage (should be Pholeuont-).
Ghidiniina Guéorguiev, 1974: 841, in key [stem: Ghidini-]. Type genus: Ghidinia Pavan, 1938 [syn. of Boldoria Jeannel, 1924].
*Coiffaitiolina Perkovsky, 2002: 16 [stem: Coiffaitiol-]. Type genus: Coiffaitiola Jeannel, 1955. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Platycholeina
Horn, 1880
Platycholei G. H. Horn, 1880b: 251 [stem: Platychole-]. Type genus: Platycholeus G. H. Horn, 1880.
Subtribe
Spelaeobatina
Guéorguiev, 1974
Spelaeobatina Guéorguiev, 1974: 841, in key [stem: Spelaeobat-]. Type genus: Spelaeobates Müller, 1901.
Tribe
Oritocatopini
Jeannel, 1936
Oritocatopini Jeannel, 1936: 116 [stem: Oritocatop-]. Type genus: Oritocatops Jeannel, 1921.
Tribe
Ptomaphagini
Jeannel, 1911
Ptomaphagini Jeannel, 1911: 193 [stem: Ptomaphag-]. Type genus: Ptomaphagus Hellwig, 1795.
Subtribe
Baryodirina
Perreau, 2000
Baryodirina Perreau, 2000: 24 [stem: Baryodir-]. Type genus: Baryodirus Perreau, 2000.
Subtribe
Ptomaphagina
Jeannel, 1911
Ptomaphagini Jeannel, 1911: 193 [stem: Ptomaphag-]. Type genus: Ptomaphagus Hellwig, 1795.
Subtribe
Ptomaphaginina
Szymczakowski, 1964
Ptomaphaginini Szymczakowski, 1964: 66 [stem: Ptomaphagin-]. Type genus: Ptomaphaginus Portevin, 1914.
Tribe
Sciaphyini
Perreau, 2000
Sciaphyini Perreau, 2000: 23 [stem: Sciaphy-]. Type genus: Sciaphyes Jeannel, 1910.
Subfamily
Platypsyllinae
Ritsema, 1869
Platypsyllidae Ritsema, 1869: 38 [stem: Platypsyll-]. Type genus: Platypsyllus Ritsema, 1869.
Leptinidae J. L. LeConte, 1872: 802 [stem: Leptin-]. Type genus: Leptinus Müller, 1817. Comment: this family-group name has been attributed to J. L. LeConte (1866 [error for 1867]) in the literature, however J. L. LeConte (1867: 368) did not give a name to the new taxon and therefore it is not available; see Newton and Thayer (1992: 37).
Family
Silphidae
Latreille, 1806
Silphales Latreille, 1806: 1 [stem: Silph-]. Type genus: Silpha Linnaeus, 1758.
Subfamily
Silphinae
Latreille, 1806
Silphales Latreille, 1806: 1 [stem: Silph-]. Type genus: Silpha Linnaeus, 1758.
Necrodisidae Gistel, 1848: [4] [stem: Necrod-]. Type genus: Necrodes Leach, 1815. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Nicrophorinae
Kirby, 1837
Necrophoridae Kirby, 1837: 95 [stem: Nicrophor-]. Type genus: Nicrophorus Fabricius, 1775 [as Necrophorus, unjustified emendation of type genus name by Thunberg (1789), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Family
Staphylinidae
Latreille, 1802
Staphyliniae Latreille, 1802: 124 [stem: Staphylin-]. Type genus: Staphylinus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1959a)]. Comment: placed on the Official List of Family-Group Names in Zoology (as Staphylinidae Latreille, [1803-1804]) and “Staphylinii Latreille, [1803-1804]” placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1959a); First Reviser (Staphylinidae Latreille, 1802 vs Pselaphidae Latreille, 1802) not determined, current usage maintained.
Subfamily
Glypholomatinae
Jeannel, 1962
Glypholomini Jeannel, 1962b: 482 [stem: Glypholomat-]. Type genus: Glypholoma Jeannel, 1962. Comment: originally proposed as a tribe of Silphidae; incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 58).
Subfamily
Microsilphinae
Crowson, 1950
*Micragyrtini Blackwelder, 1944: 84 [stem: Micragyrt-]. Type genus: Micragyrtes Champion, 1918 [syn. of Microsilpha Broun, 1886]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); originally proposed as a tribe of Leiodidae.
Microsilphinae Crowson, 1950: 279, in key [stem: Microsilph-]. Type genus: Microsilpha Broun, 1886. Comment: originally proposed as a subfamily of Silphidae.
Micragyrtini Jeannel, 1962b: 484 [stem: Micragyrt-]. Type genus: Micragyrtes Champion, 1918 [syn. of Microsilpha Broun, 1886]. Comment: originally proposed as a tribe of Silphidae.
Subfamily
Omaliinae
MacLeay, 1825
Omalidae W. S. MacLeay, 1825: 49 [stem: Omali-]. Type genus: Omalium Gravenhorst, 1802.
Tribe
Anthophagini
Thomson, 1859
Anthophagides C. G. Thomson, 1859: 48 [stem: Anthophag-]. Type genus: Anthophagus Gravenhorst, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 2004d)].
Brathinidae J. L. LeConte, 1861: 48 [stem: Brathin-]. Type genus: Brathinus J. L. LeConte, 1852. Comment: originally proposed as a subfamily of Silphidae.
*Lestévates Mulsant and Rey, 1880: 8 [stem: Lestev-]. Type genus: Lesteva Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 2004d)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant and Rey (1880).
Lestevina Jakobson, 1908: 450 [stem: Lestev-]. Type genus: Lesteva Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 2004d)].
Tribe
Aphaenostemmini
Peyerimhoff, 1914
Aphaenostemmini Peyerimhoff, 1914: 248, in key [stem: Aphaenostemm-]. Type genus: Aphaenostemmus Peyerimhoff, 1914.
Tribe
Corneolabiini
Steel, 1950
Corneolabiini Steel, 1950: 54 [stem: Corneolabi-]. Type genus: Corneolabium Steel, 1950.
Tribe
Coryphiini
Jakobson, 1908
Coryphiina Jakobson, 1908: 452 [stem: Coryphi-]. Type genus: Coryphium Stephens, 1834 [placed on the Official List of Generic Names in Zoology (ICZN 1990b)].
Subtribe
Boreaphilina
Zerche, 1990
*Boréaphilaires Mulsant and Rey, 1880: 391 [stem: Boreaphil-]. Type genus: Boreaphilus Sahlberg, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant and Rey (1880).
Boreaphilina Zerche, 1990: 22, in key [stem: Boreaphil-]. Type genus: Boreaphilus Sahlberg, 1832. Comment: family-group name proposed as new without reference to Boréaphilaires Mulsant and Rey, 1880.
Subtribe
Coryphiina
Jakobson, 1908
Coryphiina Jakobson, 1908: 452 [stem: Coryphi-]. Type genus: Coryphium Stephens, 1834 [placed on the Official List of Generic Names in Zoology (ICZN 1990b)].
Tribe
Eusphalerini
Hatch, 1957
*Anthobiates Mulsant and Rey, 1880: 290 [stem: Anthobi-]. Type genus: Anthobium sensu Erichson, 1840 [not Anthobium Leach, 1819; syn. of Eusphalerum Kraatz, 1857]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant and Rey (1880).
Anthobiini Portevin, 1929: 450 [stem: Anthobi-]. Type genus: Anthobium sensu Erichson, 1840 [not Anthobium Leach, 1819; syn. of Eusphalerum Kraatz, 1857]. Comment: based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).
Eusphalerini Hatch, 1957: 82 [stem: Eusphaler-]. Type genus: Eusphalerum Kraatz, 1857. Comment: new name for Anthobiini; although this is not the oldest name for the tribe, we recommend that an application be submitted to the Commission to suppress the older name Anthobiini Poertevin, 1929 because it is based on a misidentified type genus (Art. 65.2.1).
Tribe
Hadrognathini
Portevin, 1929
*Eugnathates Mulsant and Rey, 1880: 386 [stem: Eugnath-]. Type genus: Eugnathus Mulsant and Rey, 1851 [preoccupied genus name, not Eugnathus Schönherr, 1833 [Coleoptera: Curculionidae] or Eugnathus Agassiz, 1836 [Pisces]; syn. of Hadrognathus Schaum, 1852]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if found to be available then permanently invalid (Art. 39): based on preoccupied type genus.
Hadrognathini Portevin, 1929: 431 [stem: Hadrognath-]. Type genus: Hadrognathus Schaum, 1852.
Tribe
Omaliini
MacLeay, 1825
Omalidae W. S. MacLeay, 1825: 49 [stem: Omali-]. Type genus: Omalium Gravenhorst, 1802. Comment: incorrect original stem formation, not in prevailing usage.
Micralymmates Mulsant and Rey, 1880: 3 [stem: Micralymmat-]. Type genus: Micralymma Westwood, 1838. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Portevin (1929: 443, as Micralymmini), generally accepted as in Newton and Thayer (1992: 58, as Micralymmini); incorrect original stem formation, not in prevailing usage.
Tetradeli Fauvel, 1904: 90 [stem: Tetradel-]. Type genus: Tetradelus Fauvel, 1904.
Arpediopsini Cameron, 1917a: 123 [stem: Arpediopse-]. Type genus: Arpediopsis Cameron, 1917 [preoccupied genus name, not Arpediopsis Ganglbauer, 1895 [Coleoptera: Staphylinidae: Omaliinae: Anthophagini]; syn. of Crymus Fauvel, 1904]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Arpediomimi Cameron, 1917b: 277 [stem: Arpediomim-]. Type genus: Arpediomimus Cameron, 1917 [syn. of Crymus Fauvel, 1904]. Comment: replacement name for Arpediopsini Cameron, 1917; incorrect original stem formation, not in prevailing usage.
Phloeonomini Ádám, 2001: 231 [stem: Phloeonom-]. Type genus: Phloeonomus Heer, 1839. Comment: syn. nov.
Subfamily
Empelinae
Newton and Thayer, 1992
*Empelidae M. Abdullah, 1969b: 683 [stem: Empel-]. Type genus: Empelus J. L. LeConte, 1861. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Empelinae Newton and Thayer, 1992: 25 [stem: Empel-]. Type genus: Empelus J. L. LeConte, 1861.
Subfamily
Proteininae
Erichson, 1839
Proteinini Erichson, 1839a: 641 [stem: Protein-]. Type genus: Proteinus Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 1969a)].
Tribe
Anepiini
Steel, 1966
Anepiini Steel, 1966: 306 [stem: Anepi-]. Type genus: Anepius Blackburn, 1902.
Tribe
Austrorhysini
Newton and Thayer, 1995
Austrorhysini Newton and Thayer, 1995: 298 [stem: Austrorhys-]. Type genus: Austrorhysus Steel, 1966.
Tribe
Nesoneini
Steel, 1966
Nesoneini Steel, 1966: 292 [stem: Nesone-]. Type genus: Nesoneus Bernhauer, 1939.
Tribe
Proteinini
Erichson, 1839
Proteinini Erichson, 1839a: 641 [stem: Protein-]. Type genus: Proteinus Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 1969a)]. Comment: this family-group name was also proposed in the same year by Erichson (1839b: 31, as Proteinini) and Heer (1839b: 4, as Proteinina); for comments about the priority of these works see Newton and Thayer (1992: 24).
*Phléobiens Mulsant and Rey, 1876: 209 [stem: Phloeobi-]. Type genus: Phloeobium sensu Erichson, 1840 [not Phloeobium Dejean, 1833; syn. of Metopsia Wollaston, 1854]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant and Rey, 1876; incorrect original stem formation, not in prevailing usage.
Phloeobiinae Fowler, 1888: 431 [stem: Phloeobi-]. Type genus: Phloeobium sensu Erichson, 1840 [not Phloeobium Dejean, 1833; syn. of Metopsia Wollaston, 1854]. Comment: based on a misidentified type genus.
Megarthrini Joy, 1932: 93, in key [stem: Megarthr-]. Type genus: Megarthrus Curtis, 1829.
Metopsiinae Tottenham, 1954: 8, in key [stem: Metopsi-]. Type genus: Metopsia Wollaston, 1854.
Pteronini Arnett, 1961: 238, in key [stem: Pteroni-]. Type genus: Pteronius Blackwelder, 1952 [syn. of Proteinus Latreille, 1797]. Comment: incorrect original stem formation, not in prevailing usage.
Pteroniinae I. Moore, 1964: 85, in key [stem: Pteroni-]. Type genus: Pteronius Blackwelder, 1952 [syn. of Proteinus Latreille, 1797]. Comment: proposed as new without reference to Pteronini Arnett, 1961.
Tribe
Silphotelini
Newton and Thayer, 1995
Silphotelini Newton and Thayer, 1995: 297 [stem: Silphotel-]. Type genus: Silphotelus Broun, 1895.
Subfamily
Micropeplinae
Leach, 1815
Micropeplida Leach, 1815: 90 [stem: Micropepl-]. Type genus: Micropeplus Latreille, 1809.
Micropeplida Heer, 1839a: 169 [stem: Micropepl-]. Type genus: Micropeplus Latreille, 1809. Comment: family-group name proposed as new without reference to Micropeplida Leach, 1815; this family-group name was also used in the same year by Heer (1839b: 4, as Micropeplida); for comments about the priority of these works see Newton and Thayer (1992: 24).
Subfamily
Neophoninae
Fauvel, 1905
Neophoni Fauvel, 1905a: 98 [stem: Neophon-]. Type genus: Neophonus Fauvel, 1905.
Subfamily
Dasycerinae
Reitter, 1887
Dasycerini Reitter, 1887: 8, in key [stem: Dasycer-]. Type genus: Dasycerus Brongniart, 1800. Comment: originally proposed as a new tribe of Latridiidae; the earlier usage of the Dasycerinae by Swainson (1840: 293) was not based on a type genus and is therefore unavailable.
Subfamily
Protopselaphinae
Newton and Thayer, 1995
Protopselaphinae Newton and Thayer, 1995: 227 [stem: Protopselaph-]. Type genus: Protopselaphus Newton and Thayer, 1995.
Subfamily
Pselaphinae
Latreille, 1802
Pselaphii Latreille, 1802: 239 [stem: Pselaph-]. Type genus: Pselaphus Herbst, 1791.
Supertribe
Batrisitae
Reitter, 1882
Batrisini Reitter, 1882b: 183 [stem: Batris-]. Type genus: Batrisus Aubé, 1833.
Tribe
Amauropini
Jeannel, 1948
Amauropsini Jeannel, 1948a: 1 [stem: Amaurop-]. Type genus: Amaurops Fairmaire, 1851. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 37).
Tribe
Batrisini
Reitter, 1882
Batrisini Reitter, 1882b: 183 [stem: Batris-]. Type genus: Batrisus Aubé, 1833.
Subtribe
Ambicocerina
Leleup, 1970
Ambicocerina Leleup, 1970: 309 [stem: Ambicocer-]. Type genus: Ambicocerus Leleup, 1970.
Subtribe
Batrisina
Reitter, 1882
Batrisini Reitter, 1882b: 183 [stem: Batris-]. Type genus: Batrisus Aubé, 1833.
Oropygiina Jeannel, 1949a: 113, in key [stem: Orropygi-]. Type genus: Orropygia Raffray, 1910 [as Oropygia, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 37).
Trabisina Jeannel, 1949a: 114, in key [stem: Trabis-]. Type genus: Trabisus Raffray, 1890 [syn. of Atheropterus Raffray, 1882].
Subtribe
Leupeliina
Jeannel, 1954
Leupeliina Jeannel, 1954c: 106 [stem: Leupeli-]. Type genus: Leupelia Jeannel, 1954.
Subtribe
Stilipalpina
Jeannel, 1954
Stilipalpina Jeannel, 1954c: 118 [stem: Stilipalp-]. Type genus: Stilipalpus Jeannel, 1951.
Tribe
Thaumastocephalini
Poggi, Nonveiller, Colla, Pavićević and Rada, 2001
Thaumastocephalini Poggi et al., 2001: 5, in key [stem: Thaumastocephal-]. Type genus: Thaumastocephalus Poggi et al., 2001.
Supertribe
Clavigeritae
Leach, 1815
Clavigerides Leach, 1815: 117 [stem: Claviger-]. Type genus: Claviger Preyssler, 1790.
Tribe
Clavigerini
Leach, 1815
Clavigerides Leach, 1815: 117 [stem: Claviger-]. Type genus: Claviger Preyssler, 1790.
Subtribe
Apoderigerina
Jeannel, 1954
Apoderigerini Jeannel, 1954a: 310 [stem: Apoderiger-]. Type genus: Apoderiger Wasmann, 1897.
Subtribe
Clavigerina
Leach, 1815
Clavigerides Leach, 1815: 117 [stem: Claviger-]. Type genus: Claviger Preyssler, 1790.
Adranites Desmarest, 1857: 144 [stem: Adran-]. Type genus: Adranes J. L. LeConte, 1849.
Adraniini O. Park, 1951: 58, in key [stem: Adran-]. Type genus: Adranes J. L. LeConte, 1849. Comment: family-group name proposed as new without reference to Adranites Desmarest, 1857; incorrect original stem formation, not in prevailing usage.
Subtribe
Clavigerodina
Schaufuss, 1882
Clavigerodini L. W. Schaufuss, 1882a: 205 [stem: Clavigerod-]. Type genus: Clavigerodes Raffray, 1877.
Commatocerini L. W. Schaufuss, 1882b: 349 [stem: Commatocer-]. Type genus: Commatocerus Raffray, 1882 [syn. of Fustiger J. L. LeConte, 1866].
Clavigeropsini L. W. Schaufuss, 1890: in table [stem: Clavigeropse-]. Type genus: Clavigeropsis Raffray, 1882. Comment: incorrect original stem formation, not in prevailing usage.
Fustigerini Jeannel, 1949a: 31 [stem: Fustiger-]. Type genus: Fustiger J. L. LeConte, 1866.
Neoceratopsini Célis, 1970: 260 [stem: Neoceratopse-]. Type genus: Neoceratopsis Jeannel, 1956. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Disarthricerina
Jeannel, 1949
Diarthricerini Jeannel, 1949a: 29, in key [stem: Disarthricer-]. Type genus: Disarthricerus Raffray, 1895 [as Diarthricerus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Hoplitoxenina
Célis, 1969
Hoplitoxenini Célis, 1969: 418 [stem: Hoplitoxen-]. Type genus: Hoplitoxenus Jeannel, 1960.
Dimerometopini Célis, 1970: 244 [stem: Dimerometop-]. Type genus: Dimerometopus Célis, 1970.
Subtribe
Lunillina
Célis, 1969
Lunillini Célis, 1969: 416 [stem: Lunill-]. Type genus: Lunilla Jeannel, 1957.
Subtribe
Mastigerina
Jeannel, 1954
Mastigerini Jeannel, 1954a: 291, in key [stem: Mastiger-]. Type genus: Mastiger Motschulsky, 1851.
Subtribe
Miroclavigerina
Jeannel, 1949
Miroclavigerini Jeannel, 1949a: 29, in key [stem: Miroclaviger-]. Type genus: Miroclaviger Wasmann, 1893.
Subtribe
Neocerina
Jeannel, 1954
Neocerini Jeannel, 1954a: 316 [stem: Neocer-]. Type genus: Neocerus Wasmann, 1893. Comment: First Reviser (Neocerina Jeannel, 1954 vs Theocerina Jeannel, 1954) not determined, current usage maintained.
Theocerini Jeannel, 1954a: 314 [stem: Theocer-]. Type genus: Theocerus Raffray, 1897.
Subtribe
Radamina
Jeannel, 1954
Radamini Jeannel, 1954a: 319 [stem: Radam-]. Type genus: Radama Raffray, 1883.
Subtribe
Thysdariina
Jeannel, 1954
Thysdariini Jeannel, 1954a: 332 [stem: Thysdari-]. Type genus: Thysdarius Fairmaire, 1904.
Tribe
Colilodionini
Besuchet, 1991
Colilodionini Besuchet, 1991: 514 [stem: Colilodion-]. Type genus: Colilodion Besuchet, 1991.
Tribe
Tiracerini
Besuchet, 1986
Articerides Desmarest, 1857: 145 [stem: Articer-]. Type genus: Articerus sensu Hope, 1845 [not Articerus Dalman, 1826; syn. of Tiracerus Besuchet, 1986]. Comment: based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).
Tiracerini Besuchet, 1986: 263 [stem: Tiracer-]. Type genus: Tiracerus Besuchet, 1986.
Supertribe
Euplectitae
Streubel, 1839
Euplectidae Streubel, 1839: 135 [stem: Euplect-]. Type genus: Euplectus Leach, 1817.
Tribe
Bythinoplectini
Schaufuss, 1890
Bythinoplectini L. W. Schaufuss, 1890: in table [stem: Bythinoplect-]. Type genus: Bythinoplectus Reitter, 1882.
Subtribe
Bythinoplectina
Schaufuss, 1890
Bythinoplectini L. W. Schaufuss, 1890: in table [stem: Bythinoplect-]. Type genus: Bythinoplectus Reitter, 1882.
Zethini L. W. Schaufuss, 1890: in table [stem: Zeth-]. Type genus: Zethus L. W. Schaufuss, 1872 [preoccupied genus name, not Zethus Fabricius, 1805 [Hymenoptera], not Zethus Pander, 1830 [Trilobita]; syn. of Zethopsus Reitter, 1880]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Zethopsina Jeannel, 1952a: 51, in key [stem: Zethops-]. Type genus: Zethopsus Reitter, 1880.
Subtribe
Pyxidicerina
Raffray, 1904
Pyxidicerini Raffray, 1904: 489, in key [stem: Pyxidicer-]. Type genus: Pyxidicerus Motschulsky, 1863.
Tribe
Dimerini
Raffray, 1908
Dimerini Raffray, 1908: 412 [stem: Dimer-]. Type genus: Dimerus Fiori, 1899 [syn. of Octomicrus L. W. Schaufuss, 1877]. Comment: published before 17 March 1908; this family-group name was also used in the same year by Bernhauer (1908 [23 March]: 327, as Dimerini); use of family-group name Dimerini Raffray, 1908 (based on synonym) conserved (Art. 40.2); see Newton and Thayer (1992: 39).
Octomicrini Jeannel, 1952a: 43 [stem: Octomicr-]. Type genus: Octomicrus L. W. Schaufuss, 1877.
Tribe
Euplectini
Streubel, 1839
Euplectidae Streubel, 1839: 135 [stem: Euplect-]. Type genus: Euplectus Leach, 1817.
Mitracephalini O. Park, 1951: 64, in key [stem: Mitracephal-]. Type genus: Mitracephala Raffray, 1890 [preoccupied genus name, not Mitracephala J. Thomson, 1859 [Coleoptera: Scarabaeidae]; syn. of Mitrametopus Raffray, 1911]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Mitrametopina O. Park, 1952: 87 [stem: Mitrametop-]. Type genus: Mitrametopus Raffray, 1911. Comment: replacement name for Mitracephalini O. Park, 1951 because of the homonymy of the type genus.
Tribe
Jubini
Raffray, 1904
Jubini Raffray, 1904: 489, in key [stem: Jub-]. Type genus: Jubus L. W. Schaufuss, 1872.
Auxenocerini Jeannel, 1962a: 319 [stem: Auxenocer-]. Type genus: Auxenocerus Jeannel, 1962.
Tribe
Mayetiini
Winkler, 1925
Mayetiini A. Winkler, 1925: 348 [stem: Mayeti-]. Type genus: Mayetia Mulsant and Rey, 1875.
Tribe
Metopiasini
Raffray, 1904
Metopiini Raffray, 1904: 490, in key [stem: Metopias-]. Type genus: Metopias Gory, 1832 [placed on the Official List of Generic Names in Zoology (ICZN 1994d)]. Comment: Metopias- established as the correct stem and Metopiasini Raffray, 1904 placed on the Official List of Family-Group Names in Zoology (ICZN 1994d).
Subtribe
Metopiasina
Raffray, 1904
Metopiini Raffray, 1904: 490, in key [stem: Metopias-]. Type genus: Metopias Gory, 1832 [placed on the Official List of Generic Names in Zoology (ICZN 1994d)]. Comment: correct original stem Metopi- modified to Metopias- and Metopiasini Raffray, 1904 placed on the Official List of Family-Group Names in Zoology (ICZN 1994d); Metopiini Raffray, 1904 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1994d).
Subtribe
Rhinoscepsina
Bowman, 1934
Rhinoscepsii Bowman, 1934: 8 [stem: Rhinosceps-]. Type genus: Rhinoscepsis J. L. LeConte, 1878. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Rhinoscepse-).
Tribe
Trichonychini
Reitter, 1882
Trichonyides Reitter, 1882b: 194 [stem: Trichonych-]. Type genus: Trichonyx Chaudoir, 1845.
Subtribe
Bibloporina
Park, 1951
Bibloporini O. Park, 1951: 64, in key [stem: Biblopor-]. Type genus: Bibloporus C. G. Thomson, 1859.
Bibloporellina Jeannel, 1952a: 92 [stem: Bibloporell-]. Type genus: Bibloporellus Jeannel, 1949.
Subtribe
Panaphantina
Jeannel, 1950
Panaphantina Jeannel, 1950a: 76 [stem: Panaphant-]. Type genus: Panaphantus Kiesenwetter, 1858.
Trisignina O. Park and Schuster, 1955: 1 [stem: Trisign-]. Type genus: Trisignis O. Park and Schuster, 1955.
Acetaliini Jeannel, 1958a: 81 [stem: Acetali-]. Type genus: Acetalius Sharp, 1883.
Bibloplectina Jeannel, 1959: 96, in key [stem: Bibloplect-]. Type genus: Bibloplectus Reitter, 1881.
Subtribe
Trichonychina
Reitter, 1882
Trichonyides Reitter, 1882b: 194 [stem: Trichonych-]. Type genus: Trichonyx Chaudoir, 1845. Comment: incorrect original stem formation, not in prevailing usage.
Raffrayina Jeannel, 1949a: 76 [stem: Raffrayi-]. Type genus: Raffrayia Reitter, 1882. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Jeannel (1964: 65, as Raffrayiini).
Ranavalini Jeannel, 1954a: 183 [stem: Ranaval-]. Type genus: Ranavala Raffray, 1898.
Chrestomerina Jeannel, 1962a: 344 [stem: Chrestomer-]. Type genus: Chrestomera Jeannel, 1962.
Pteracmini Jeannel, 1962a: 347 [stem: Pteracmet-]. Type genus: Pteracmes Raffray, 1890. Comment: incorrect original stem formation, not in prevailing usage.
Trimiodytina Jeannel, 1964: 39 [stem: Trimiodyt-]. Type genus: Trimiodytes Raffray, 1897.
Subtribe
Trimiina
Bowman, 1934
Trimii Bowman, 1934: 8 [stem: Trimi-]. Type genus: Trimium Aubé, 1833.
Trimiina Jeannel, 1950a: 139 [stem: Trimi-]. Type genus: Trimium Aubé, 1833. Comment: family-group name proposed as new without reference to Trimii Bowman, 1934.
Tribe
Trogastrini
Jeannel, 1949
Trogastrini Jeannel, 1949a: 41, in key [stem: Trogastr-]. Type genus: Trogaster Sharp, 1874.
Subtribe
Phtegnomina
Park, 1951
Phtegnomini O. Park, 1951: 64, in key [stem: Phtegnom-]. Type genus: Phtegnomus Raffray, 1890.
Subtribe
Rhexiina
Park, 1951
Rhexini O. Park, 1951: 63, in key [stem: Rhexi-]. Type genus: Rhexius J. L. LeConte, 1849. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 39).
Subtribe
Trogastrina
Jeannel, 1949
Trogastrini Jeannel, 1949a: 41, in key [stem: Trogastr-]. Type genus: Trogaster Sharp, 1874.
Supertribe
Faronitae
Reitter, 1882
Faronides Reitter, 1882b: 194 [stem: Faron-]. Type genus: Faronus Aubé, 1844.
Supertribe
Goniaceritae
Reitter, 1882 (1872)
Goniacerides Reitter, 1882b: 188 [stem: Goniacer-]. Type genus: Goniacerus Motschulsky, 1855. Comment: use of family-group name conserved over Goniastitae L. W. Schaufuss, 1872 (Art. 40.2) (see Newton and Thayer 1992: 39).
Tribe
Arnylliini
Jeannel, 1952
Arnylliini Jeannel, 1952b: 100 [stem: Arnylli-]. Type genus: Arnyllium Reitter, 1884.
Tribe
Barrosellini
Leleup, 1973
Barrosellini Leleup, 1973: 81 [stem: Barrosell-]. Type genus: Barrosellus Jeannel, 1951.
Tribe
Brachyglutini
Raffray, 1904
Brachyglutini Raffray, 1904: 490, in key [stem: Brachyglut-]. Type genus: Brachygluta C. G. Thomson, 1859.
Subtribe
Baradina
Park, 1951
Baradiini O. Park, 1951: 62, in key [stem: Barad-]. Type genus: Barada Raffray, 1891 [syn. of Euphalepsus Reitter, 1883]. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 40).
Subtribe
Brachyglutina
Raffray, 1904
Bryaxes J. L. LeConte, 1861: 57 [stem: Bryaxe-]. Type genus: Bryaxis Leach, 1817 [preoccupied genus name, not Bryaxis Kugelann, 1794 [Coleoptera: Staphylinidae: Pselaphinae: Bythinini]; placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1969b); syn. of Rybaxis Saulcy, 1876]. Comment: permanently invalid (Art. 39): based on preoccupied and suppressed type genus; incorrect original stem formation, not in prevailing usage.
Brachyglutini Raffray, 1904: 490, in key [stem: Brachyglut-]. Type genus: Brachygluta C. G. Thomson, 1859. Comment: replacement name for Bryaxes J. L. LeConte, 1861.
Reichenbachiina Jakobson, 1910: 577 [stem: Reichenbachi-]. Type genus: Reichenbachia Leach, 1826.
Globina Jeannel, 1959: 462, in key [stem: Glob-]. Type genus: Globa Raffray, 1887.
Halorabyxina Leleup, 1969a: 138 [stem: Halorabyx-]. Type genus: Halorabyxis Jeannel, 1954 [syn. of Physoplectus Reitter, 1882].
Pselaptina O. Park, 1976: 48 [stem: Pselapt-]. Type genus: Pselaptus J. L. LeConte, 1880.
Subtribe
Decarthrina
Park, 1951
Decarthronini O. Park, 1951: 61, in key [stem: Decarthr-]. Type genus: Decarthron Brendel, 1865. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 40).
Subtribe
Eupseniina
Park, 1951
Eupseniini O. Park, 1951: 61, in key [stem: Eupseni-]. Type genus: Eupsenius J. L. LeConte, 1849.
Tribe
Bythinini
Raffray, 1890
Bythinini Raffray, 1890: 83, in key [stem: Bythin-]. Type genus: Bythinus Leach, 1817 [placed on the Official List of Generic Names in Zoology (ICZN 1969b)].
Subtribe
Bythinina
Raffray, 1890
Bythinini Raffray, 1890: 83, in key [stem: Bythin-]. Type genus: Bythinus Leach, 1817 [placed on the Official List of Generic Names in Zoology (ICZN 1969b)].
Bryaxina Jakobson, 1910: 579 [stem: Bryaxe-]. Type genus: Bryaxis Kugelann, 1794 [placed on the Official List of Generic Names in Zoology (ICZN 1969b)]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Machaeritina
Jeannel, 1950
Machaeritina Jeannel, 1950a: 168 [stem: Machaerit-]. Type genus: Machaerites Miller, 1855.
Subtribe
Xenobythina
Jeannel, 1950
Xenobythina Jeannel, 1950a: 201 [stem: Xenobyth-]. Type genus: Xenobythus Peyerimhoff, 1901.
Tribe
Cyathigerini
Schaufuss, 1872
Cyathigerini L. W. Schaufuss, 1872: 245 [stem: Cyathiger-]. Type genus: Cyathiger King, 1865 [syn. of Plagiophorus Motschulsky, 1851].
Tribe
Goniacerini
Reitter, 1882 (1872)
Goniastini L. W. Schaufuss, 1872: 245 [stem: Goniast-]. Type genus: Goniastes Westwood, 1870. Comment: use of younger family-group name Goniacerini Reitter, 1882 conserved (Art. 40.2) (see Newton and Thayer 1992: 40).
Goniacerides Reitter, 1882b: 188 [stem: Goniacer-]. Type genus: Goniacerus Motschulsky, 1855. Comment: use of family-group name conserved over Goniastini L. W. Schaufuss, 1872 (Art. 40.2) (see Newton and Thayer 1992: 40).
Simini L. W. Schaufuss, 1890: in table [stem: Sim-]. Type genus: Simus Raffray, 1882 [preoccupied genus name, not Simus Bonaparte, 1838 [Reptilia] or Simus Hodgson, 1841 [Aves]; syn. of Ipsimus Reitter, 1885]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Listriophorini L. W. Schaufuss, 1890: in table [stem: Listriophor-]. Type genus: Listriophorus L. W. Schaufuss, 1872.
Tribe
Imirini
Jeannel, 1949
Mirini Raffray, 1917: 110 [stem: Mir-]. Type genus: Mirus Saulcy, 1877 [preoccupied genus name, not Mirus Albers, 1850 [Mollusca]; syn. of Imirus Reitter, 1885]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; junior homonym of Miridae Hahn, 1833 (type genus Miris Fabricius, 1794) in Hemiptera.
Imirini Jeannel, 1949a: 41, in key [stem: Imir-]. Type genus: Imirus Reitter, 1885.
Tribe
Iniocyphini
Park, 1951
Iniocyphini O. Park, 1951: 60, in key [stem: Iniocyph-]. Type genus: Iniocyphus Raffray, 1912.
Subtribe
Iniocyphina
Park, 1951
Iniocyphini O. Park, 1951: 60, in key [stem: Iniocyph-]. Type genus: Iniocyphus Raffray, 1912. Comment: First Reviser found (Iniocyphina Park, 1951 vs Dalmodiina Park, 1951) is Chandler (2001: 377).
Dalmodiini O. Park, 1951: 61, in key [stem: Dalmod-]. Type genus: Dalmodes Reitter, 1882. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 41).
Subtribe
Natypleurina
Newton and Thayer, 1992
Tanypleurini Jeannel, 1949a: 79 [stem: Tanypleur-]. Type genus: Tanypleurus Raffray, 1890 [preoccupied genus name, not Tanypleurus Steenstrup and Luetken, 1861 [Crustacea]; syn. of Natypleurus Newton and Thayer, 1992]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Natypleurina Newton and Thayer, 1992: 41 [stem: Natypleur-]. Type genus: Natypleurus Newton and Thayer, 1992. Comment: replacement name for Tanypleurini Jeannel, 1949 because of the homonymy of the type genus.
Tribe
Machadoini
Jeannel, 1951
Machadoini Jeannel, 1951: 105 [stem: Machado-]. Type genus: Machadous Jeannel, 1951.
Tribe
Proterini
Jeannel, 1949
Proterini Jeannel, 1949a: 41, in key [stem: Proter-]. Type genus: Proterus Raffray, 1897.
Tribe
Pygoxyini
Reitter, 1909
Pygoxyini Reitter, 1909: 202 [stem: Pygoxy-]. Type genus: Pygoxyon Reitter, 1880.
Tribe
Speleobamini
Park, 1951
Speleobamini O. Park, 1951: 51 [stem: Speleobam-]. Type genus: Speleobama O. Park, 1951.
Tribe
Tychini
Raffray, 1904
Tychini Raffray, 1904: 490, in key [stem: Tych-]. Type genus: Tychus Leach, 1817.
Tribe
Valdini
Park, 1953
Valdiini O. Park, 1953: 261 [stem: Vald-]. Type genus: Valda Casey, 1894. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 41).
Supertribe
Pselaphitae
Latreille, 1802
Pselaphii Latreille, 1802: 239 [stem: Pselaph-]. Type genus: Pselaphus Herbst, 1791.
Tribe
Arhytodini
Raffray, 1890
Arhytodini Raffray, 1890: 84, in key [stem: Arhytod-]. Type genus: Arhytodes Reitter, 1882 [syn. of Rhytus Westwood, 1870].
Holozodini Raffray, 1900: 518 [stem: Holozod-]. Type genus: Holozodus Fairmaire, 1898.
Tribe
Attapseniini
Bruch, 1933
Attapseniini Bruch, 1933a: 26 [stem: Attapseni-]. Type genus: Attapsenius Bruch, 1933.
Tribe
Ctenistini
Blanchard, 1845
Cténistites Blanchard, 1845a: 306 [stem: Ctenist-]. Type genus: Ctenistes Reichenbach, 1816. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Reitter (1882b: 183, as Ctenistini), generally accepted as in Chandler (2001: 484, as Ctenistini).
Chenniides Reitter, 1882b: 183 [stem: Chenni-]. Type genus: Chennium Latreille, 1807.
Tribe
Hybocephalini
Raffray, 1890
Hybocephalini Raffray, 1890: 83, in key [stem: Hybocephal-]. Type genus: Hybocephalus Motschulsky, 1851.
Mestogastrina Jakobson, 1910: 585 [stem: Mestogastr-]. Type genus: Mestogaster Schmidt-Göbel, 1838.
Tribe
Odontalgini
Jeannel, 1949
Odontalgini Jeannel, 1949a: 177 [stem: Odontalg-]. Type genus: Odontalgus Raffray, 1877.
Tribe
Pachygastrodini
Leleup, 1969
Pachygastrodini Leleup, 1969b: 282 [stem: Pachygastrod-]. Type genus: Pachygastrodes Leleup, 1969.
Tribe
Phalepsini
Jeannel, 1949
Phalepsini Jeannel, 1949a: 208 [stem: Phaleps-]. Type genus: Phalepsus Westwood, 1870.
Tribe
Pselaphini
Latreille, 1802
Pselaphii Latreille, 1802: 239 [stem: Pselaph-]. Type genus: Pselaphus Herbst, 1791.
Tribe
Schistodactylini
Raffray, 1890
Schistodactylini Raffray, 1890: 84, in key [stem: Schistodactyl-]. Type genus: Schistodactylus Raffray, 1883.
Tribe
Tmesiphorini
Jeannel, 1949
Tmesiphorini Jeannel, 1949a: 202 [stem: Tmesiphor-]. Type genus: Tmesiphorus J. L. LeConte, 1849.
Tribe
Tyrini
Reitter, 1882
Tyrides Reitter, 1882b: 184 [stem: Tyr-]. Type genus: Tyrus Aubé, 1833.
Subtribe
Centrophthalmina
Jeannel, 1949
Centrophthalmina Jeannel, 1949a: 209, in key [stem: Centrophthalm-]. Type genus: Centrophthalmus Schmidt-Göbel, 1838.
Petanopini Jeannel, 1954b: 102, in key [stem: Petanop-]. Type genus: Petanops Jeannel, 1954 [syn. of Daveyia Lea, 1912].
Subtribe
Janusculina
Cerruti, 1970
Janusculina Cerruti, 1970: 123 [stem: Januscul-]. Type genus: Janusculus Cerruti, 1970.
Subtribe
Somatipionina
Jeannel, 1949
Somatipionini Jeannel, 1949a: 208 [stem: Somatipion-]. Type genus: Somatipion L. W. Schaufuss, 1877.
Hamotini O. Park, 1951: 67, in key [stem: Hamot-]. Type genus: Hamotus Aubé, 1844.
Subtribe
Tyrina
Reitter, 1882
Tyrides Reitter, 1882b: 184 [stem: Tyr-]. Type genus: Tyrus Aubé, 1833.
Chalcoplectini Oke, 1925: 13 [stem: Chalcoplect-]. Type genus: Chalcoplectus Oke, 1925.
Ceophyllini O. Park, 1951: 67, in key [stem: Ceophyll-]. Type genus: Ceophyllus J. L. LeConte, 1849.
Subfamily
Phloeocharinae
Erichson, 1839
Phloeocharini Erichson, 1839a: 612 [stem: Phloeochar-]. Type genus: Phloeocharis Mannerheim, 1830. Comment: this family-group name was also used in the same year by Erichson (1839b: 31, as Phloeocharini) and Heer (1839b: 5, as Phloeocharina); for comments about the priority of these works see Newton and Thayer (1992: 24); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Phloeocharit-).
Scotodytidae Saulcy, 1870: 90 [stem: Scotodyt-]. Type genus: Scotodytes Saulcy, 1865 [syn. of Phloeocharis Mannerheim, 1830].
Subfamily
Olisthaerinae
Thomson, 1858
Olisthaerini C. G. Thomson, 1858: 38 [stem: Olisthaer-]. Type genus: Olisthaerus Dejean, 1833.
Subfamily
Tachyporinae
MacLeay, 1825
Tachyporidae W. S. MacLeay, 1825: 49 [stem: Tachypor-]. Type genus: Tachyporus Gravenhorst, 1802.
Tribe
Deropini
Smetana, 1983
Deropsini Smetana, 1983: 272 [stem: Derop-]. Type genus: Derops Sharp, 1889. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 66).
Tribe
Megarthropsini
Cameron, 1919
Megarthropsini Cameron, 1919: 231 [stem: Megarthrops-]. Type genus: Megarthropsis Cameron, 1919. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Megarthropse-).
Tribe
Mycetoporini
Thomson, 1859
Mycetoporides C. G. Thomson, 1859: 46 [stem: Mycetopor-]. Type genus: Mycetoporus Mannerheim, 1830 [placed on the Official List of Generic Names in Zoology (ICZN 1993c)].
Bolitobii G. H. Horn, 1877: 83 [stem: Bolitobi-]. Type genus: Bolitobius Leach, 1819.
Tribe
Tachyporini
MacLeay, 1825
Tachinidae Fleming, 1821: 49 [stem: Tachinus-]. Type genus: Tachinus Gravenhorst, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1993f)]. Comment: senior homonym of Tachinariae Robineau-Desvoidy, 1830 [Diptera: Tachinidae], stem emended to Tachinus- and Tachinusidae Fleming, 1821 placed on the Official List of Family-Group Names in Zoology (ICZN 1993f); Tachinidae Fleming, 1821 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1993f).
Tachyporidae W. S. MacLeay, 1825: 49 [stem: Tachypor-]. Type genus: Tachyporus Gravenhorst, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1993f)]. Comment: family-group name given precedence over Tachinusidae Fleming, 1821 and placed on the Official List of Family-Group Names in Zoology (ICZN 1993f).
Symmixini Bernhauer, 1915: 56 [stem: Symmix-]. Type genus: Symmixus Bernhauer, 1915.
Euconosomini Cameron, 1918: 216 [stem: Euconosomat-]. Type genus: Euconosoma Cameron, 1918. Comment: incorrect original stem formation, not in prevailing usage.
Conosomini Jeannel and Jarrige, 1949: 335 [stem: Conosomat-]. Type genus: Conosoma Kraatz, 1857 [preoccupied genus name, not Conosoma Lenz, 1794 [Diptera]; syn. of Tachinus Gravenhorst, 1802]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Sepedophilini Ádám, 2001: 199 [stem: Sepedophil-]. Type genus: Sepedophilus Gistel, 1856. Comment: syn. nov.
Tribe
Vatesini
Seevers, 1958
Xenocephalini Wasmann, 1887: 411 [stem: Xenocephal-]. Type genus: Xenocephalus Wasmann, 1887 [preoccupied genus name, not Xenocephalus Kaup, 1858 [Pisces]; replaced by Wasmannotherium Bernhauer, 1921; syn. of Vatesus Sharp, 1876]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Vatesini Seevers, 1958: 183 [stem: Vates-]. Type genus: Vatesus Sharp, 1876. Comment: replacement name for Xenocephalini Wasmann, 1887 because of the homonymy of the type genus.
Subfamily
Trichophyinae
Thomson, 1858
Trichophyini C. G. Thomson, 1858: 30 [stem: Trichophy-]. Type genus: Trichophya Mannerheim, 1830.
Subfamily
Habrocerinae
Mulsant and Rey, 1876
Habrocériens Mulsant and Rey, 1876: 210 [stem: Habrocer-]. Type genus: Habrocerus Erichson, 1839. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by G. H. Horn (1877: 83, as Habroceri), generally accepted as in Newton and Thayer (1992: 56, as Habrocerinae).
Subfamily
Aleocharinae
Fleming, 1821
Aleocharidae Fleming, 1821: 49 [stem: Aleochar-]. Type genus: Aleochara Gravenhorst, 1802. Comment: First Reviser (Aleocharinae Fleming, 1821 vs Lomechusinae Fleming, 1821) not determined, current usage maintained.
Tribe
Actocharini
Bernhauer and Schubert, 1911
Actochari Bernhauer and Schubert, 1911: 91 [stem: Actochar-]. Type genus: Actocharis Fauvel, 1869. Comment: originally proposed as a subtribe of Oxytelini Fleming, 1821; current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Actocharit-).
Tribe
Aenictoteratini
Kistner, 1993
Aenictoteratini Kistner, 1993: 242 [stem: Aenictoterat-]. Type genus: Aenictoteras W. M. Wheeler, 1932.
Tribe
Akatastopsisini
Pace, 2000
Akatastopsisini Pace, 2000a: 112 [stem: Akatastopsis-]. Type genus: Akatastopsis Pace, 2000.
Tribe
Aleocharini
Fleming, 1821
Aleocharidae Fleming, 1821: 49 [stem: Aleochar-]. Type genus: Aleochara Gravenhorst, 1802.
Subtribe
Aleocharina
Fleming, 1821
Aleocharidae Fleming, 1821: 49 [stem: Aleochar-]. Type genus: Aleochara Gravenhorst, 1802.
Piochardiae Fenyes, 1918: 20 [stem: Piochardi-]. Type genus: Piochardia Heyden, 1870.
Subtribe
Compactopediina
Kistner, 1970
Compactopedina Kistner, 1970b: 18 [stem: Compactopedi-]. Type genus: Compactopedia Kistner, 1970. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 47).
Subtribe
Hodoxenina
Kistner, 1970
Hodoxenina Kistner, 1970a: 12 [stem: Hodoxen-]. Type genus: Hodoxenus Kistner, 1970.
Tribe
Athetini
Casey, 1910
Athetae Casey, 1910: 2 [stem: Athet-]. Type genus: Atheta C. G. Thomson, 1858 [placed on the Official List of Generic Names in Zoology (ICZN 1961b)]. Comment: notice of a new application for the conservation of this name over the older name Callicerini Jakobson, 1908 by Gusarov was recently published in the Bulletin of Zoological Nomenclature (2010: 270; also see Appendix 6).
Subtribe
Athetina
Casey, 1910
Callicerina Jakobson, 1908: 448 [stem: Callicer-]. Type genus: Callicerus Gravenhorst, 1802. Comment: junior homonym of Callicerina Rondani, 1856 (type genus Callicera Panzer, 1806) in Diptera: Syrphidae; notice of a new application for the suppression of this name by Gusarov was recently published in the Bulletin of Zoological Nomenclature (2010: 270; also see Appendix 6).
Athetae Casey, 1910: 2 [stem: Athet-]. Type genus: Atheta C. G. Thomson, 1858 [placed on the Official List of Generic Names in Zoology (ICZN 1961b)]. Comment: notice of a new application for the conservation of this name over the older name Callicerina Jakobson, 1908 by Gusarov was recently published in the Bulletin of Zoological Nomenclature (2010: 270; also see Appendix 6).
Strigotae Casey, 1910: 176 [stem: Strigot-]. Type genus: Strigota Casey, 1910.
Plagiarthrini Cameron, 1926: 184 [stem: Plagiarthrin-]. Type genus: Plagiarthrina Keys, 1920 [syn. of Atheta C. G. Thomson, 1858]. Comment: incorrect original stem formation, not in prevailing usage.
Ischnopodini Hatch, 1957: 141 [stem: Ischnopod-]. Type genus: Ischnopoda sensu Westwood, 1838 [not Ischnopoda Stephens, 1835 (see ICZN 1961b); syn. of Acrotona C. G. Thomson, 1859]. Comment: based on misidentified type genus.
*Xenotae Seevers, 1978: 113 [stem: Xenot-]. Type genus: Xenota Mulsant and Rey, 1873 [subgenus of Atheta C. G. Thomson, 1858]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Acrotonae Seevers, 1978: 97 [stem: Acroton-]. Type genus: Acrotona C. G. Thomson, 1859 [placed on the Official List of Generic Names in Zoology (ICZN 1961b)].
Geostibae Seevers, 1978: 126 [stem: Geostib-]. Type genus: Geostiba C. G. Thomson, 1858 [placed on the Official List of Generic Names in Zoology (ICZN 2005a)].
Dimetrotae Seevers, 1978: 102 [stem: Dimetrot-]. Type genus: Dimetrota Mulsant and Rey, 1873 [subgenus of Atheta C. G. Thomson, 1858].
*Hydrosmectina Muona, 1979: 23 [stem: Hydrosmect-]. Type genus: Hydrosmecta C. G. Thomson, 1858. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Trichomicrina Muona, 1979: 23 [stem: Trichomicr-]. Type genus: Trichomicra Brundin, 1945. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Dadobiina Muona, 1979: 23 [stem: Dadobi-]. Type genus: Dadobia C. G. Thomson, 1858. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Amischina Muona, 1979: 25 [stem: Amisch-]. Type genus: Amischa C. G. Thomson, 1858. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Coptotermoeciina
Kistner and Pasteels, 1970
*Coptotermoeciina Seevers, 1957: 248 [stem: Coptotermoeci-]. Type genus: Coptotermoecia Oke, 1933. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Coptotermoeciina Kistner and Pasteels, 1970: 86 [stem: Coptotermoeci-]. Type genus: Coptotermoecia Oke, 1933.
Subtribe
Microceroxenina
Kistner, 1970
Microceroxenina Kistner, 1970c: 10 [stem: Microceroxen-]. Type genus: Microceroxenus Kistner, 1970.
Subtribe
Nasutiphilina
Kistner, 1970
Nasutiphilina Kistner, 1970d: 500 [stem: Nasutiphil-]. Type genus: Nasutiphilus Kistner, 1970.
Subtribe
Schistogeniina
Fenyes, 1918
Schistogeniae Fenyes, 1918: 18 [stem: Schistogeni-]. Type genus: Schistogenia Kraatz, 1857.
Subtribe
Taxicerina
Lohse, 1989
Taxicerina Lohse, 1989: 210 [stem: Taxicer-]. Type genus: Taxicera Mulsant and Rey, 1873.
Subtribe
Termitotelina
Kistner, 1970
*Termitotelina Seevers, 1957: 250 [stem: Termitotel-]. Type genus: Termitotelus Wasmann, 1908. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Termitotelina Kistner, 1970a: 4 [stem: Termitotel-]. Type genus: Termitotelus Wasmann, 1908.
Subtribe
Thamiaraeina
Fenyes, 1921
Thamiaraeini Fenyes, 1921: 34 [stem: Thamiarae-]. Type genus: Thamiaraea C. G. Thomson, 1858.
Tribe
Autaliini
Thomson, 1859
Autaliides C. G. Thomson, 1859: 30 [stem: Autali-]. Type genus: Autalia Leach, 1819.
Ophioglossae Fenyes, 1918: 18 [stem: Ophiogloss-]. Type genus: Ophioglossa Fauvel, 1866.
Rhopalogastra Fenyes, 1918: 17 [stem: Rhopalogastr-]. Type genus: Rhopalogastrum Bernhauer, 1912.
Tribe
Cordobanini
Bernhauer, 1910
Cordobanini Bernhauer, 1910: 386 [stem: Cordoban-]. Type genus: Cordobanus Bernhauer, 1910.
Tribe
Corotocini
Fenyes, 1918
Corotocini Fenyes, 1918: 61 [stem: Corotoc-]. Type genus: Corotoca Schiødte, 1847.
Subtribe
Abrotelina
Seevers, 1957
Abrotelina Seevers, 1957: 121 [stem: Abrotel-]. Type genus: Abroteles Casey, 1889.
Subtribe
Corotocina
Fenyes, 1918
Corotocini Fenyes, 1918: 61 [stem: Corotoc-]. Type genus: Corotoca Schiødte, 1847.
Termitomimini Fenyes, 1921: 33 [stem: Termitomim-]. Type genus: Termitomimus Trägårdh, 1907.
Subtribe
Eburniogastrina
Jacobson, Kistner and Pasteels, 1986
Eburniogastrina Jacobson et al., 1986: 27 [stem: Eburniogastr-]. Type genus: Eburniogaster Seevers, 1938.
Subtribe
Nasutitellina
Jacobson, Kistner and Pasteels, 1986
Nasutitellina Jacobson et al., 1986: 95 [stem: Nasutitell-]. Type genus: Nasutitella Pasteels, 1967.
Subtribe
Sphuridaethina
Pace, 1988
Sphuridaethina Pace, 1988: 980 [stem: Sphuridaeth-]. Type genus: Sphuridaethes Pace, 1988.
Subtribe
Termitocharina
Seevers, 1957
Termitocharina Seevers, 1957: 64, in key [stem: Termitochar-]. Type genus: Termitochara Wasmann, 1893.
Subtribe
Termitocupidina
Jacobson, Kistner and Pasteels, 1986
Termitocupidina Jacobson et al., 1986: 35 [stem: Termitocupid-]. Type genus: Termitocupidus Jacobson et al., 1986.
Subtribe
Termitogastrina
Bernhauer and Scheerpeltz, 1926
Termitogastri Bernhauer and Scheerpeltz, 1926: 734 [stem: Termitogastr-]. Type genus: Termitogaster Casey, 1889.
Termitellici Jacobson et al., 1986: 47 [stem: Termitell-]. Type genus: Termitella Wasmann, 1911. Comment: proposed as an infratribe, a rank not used here.
Termitogastrici Jacobson et al., 1986: 58 [stem: Termitogastr-]. Type genus: Termitogaster Casey, 1889. Comment: family-group name proposed as a new taxon, without reference to Termitogastri Bernhauer and Scheerpeltz, 1926; proposed as an infratribe, a rank not used here.
Subtribe
Termitoiceina
Jacobson, Kistner and Pasteels, 1986
Termitoiceina Jacobson et al., 1986: 84 [stem: Termitoice-]. Type genus: Termitoiceus Silvestri, 1901.
Subtribe
Termitopithina
Jacobson, Kistner and Pasteels, 1986
Termitopithina Jacobson et al., 1986: 80 [stem: Termitopith-]. Type genus: Termitopithus Seevers, 1957.
Subtribe
Termitoptochina
Fenyes, 1921
Termitoptochini Fenyes, 1921: 33 [stem: Termitoptoch-]. Type genus: Termitoptochus Silvestri, 1911.
Termitoptochina Jacobson et al., 1986: 96 [stem: Termitoptoch-]. Type genus: Termitoptochus Silvestri, 1911. Comment: family-group name proposed as new without reference to Termitoptochini Fenyes, 1921.
Subtribe
Timeparthenina
Fenyes, 1921
Timeparthenini Fenyes, 1921: 34 [stem: Timeparthen-]. Type genus: Timeparthenus Silvestri, 1901.
Tribe
Crematoxenini
Mann, 1921
Crematoxenini Mann, 1921: 547 [stem: Crematoxen-]. Type genus: Crematoxenus Mann, 1921.
Pulicomorphini Mann, 1924: 87 [stem: Pulicomorph-]. Type genus: Pulicomorpha Mann, 1924.
Philacamatini Bruch, 1933b: 206 [stem: Philacamat-]. Type genus: Philacamatus Bruch, 1933.
Tribe
Cryptonotopseini
Pace, 2003
Cryptonotopsisini Pace, 2003: 38 [stem: Cryptonotopse-]. Type genus: Cryptonotopsis Pace, 2003. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Deinopsini
Sharp, 1883
Deinopsini Sharp, 1883: 294 [stem: Deinops-]. Type genus: Deinopsis A. Matthews, 1838. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Deinopse-).
Adinopsini Cameron, 1919: 242 [stem: Adinopse-]. Type genus: Adinopsis Cameron, 1919. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Diestotini
Mulsant and Rey, 1871
Diestotates Mulsant and Rey, 1871c: 96 [stem: Diestot-]. Type genus: Diestota Mulsant and Rey, 1870. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Lohse (1989: 186, as Diestotini), generally accepted as in Newton and Thayer (1992: 49, as Diestotini).
Elachistarthronini Notman, 1920: 714 [stem: Elachistarthr-]. Type genus: Elachistarthron Notman, 1920 [syn. of Diestota Mulsant and Rey, 1870]. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 49).
Tribe
Diglottini
Jakobson, 1909
Diglossaires Mulsant and Rey, 1873a: 73 [stem: Digloss-]. Type genus: Diglossa Haliday, 1837 [preoccupied genus name, not Diglossa Wagler, 1832 [Aves]; syn. of Diglotta Champion, 1899]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Ganglbauer (1895: 313, as Diglossini); permanently invalid (Art. 39): based on preoccupied type genus.
Diglottina Jakobson, 1909: 529 [stem: Diglott-]. Type genus: Diglotta Champion, 1899. Comment: published 4 March 1909; this family-group name was also used in the same year by Eichelbaum (1909 [before 26 December]: 204, as Diglottini).
Tribe
Digrammini
Fauvel, 1900
Digrammini Fauvel, 1900: 123 [stem: Digramm-]. Type genus: Digrammus Fauvel, 1900.
Tribe
Dorylogastrini
Wasmann, 1916
Dorylogastrini Wasmann, 1916a: 103 [stem: Dorylogastr-]. Type genus: Dorylogaster Wasmann, 1904.
Tribe
Dorylomimini
Wasmann, 1916
Dorylomimini Wasmann, 1916a: 99 [stem: Dorylomim-]. Type genus: Dorylomimus Wasmann, 1902.
Tribe
Drepanoxenini
Kistner and Watson, 1972
Drepanoxenini Kistner and Watson, 1972: 2 [stem: Drepanoxen-]. Type genus: Drepanoxenus Kistner and Watson, 1972.
Tribe
Ecitocharini
Seevers, 1965
Ecitocharini Seevers, 1965: 287 [stem: Ecitochar-]. Type genus: Ecitochara Wasmann, 1887.
Tribe
Ecitogastrini
Fenyes, 1918
Ecitogastrini Fenyes, 1918: 74 [stem: Ecitogastr-]. Type genus: Ecitogaster Wasmann, 1899.
Tribe
Eusteniamorphini
Bernhauer and Scheerpeltz, 1926
Eusteniamorphini Bernhauer and Scheerpeltz, 1926: 517 [stem: Eusteniamorph-]. Type genus: Eusteniamorpha Cameron, 1920.
Tribe
Falagriini
Mulsant and Rey, 1873
Falagriates Mulsant and Rey, 1873b: 8 [stem: Falagri-]. Type genus: Falagria Leach, 1819. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1874 [Gatt.]: 71, as Falagriina), generally accepted as in Newton and Thayer (1992: 50, as Falagriini).
Tribe
Feldini
Kistner, 1972
*Feldina Seevers, 1957: 236 [stem: Feld-]. Type genus: Felda Blackwelder, 1952. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Feldini Kistner, 1972: 2 [stem: Feld-]. Type genus: Felda Blackwelder, 1952.
Tribe
Gymnusini
Heer, 1839
Gymnusida Heer, 1839a: 302 [stem: Gymnus-]. Type genus: Gymnusa Gravenhorst, 1806. Comment: this family-group name was also used in the same year by Heer (1839b: 49, as Gymnusida); for comments about the priority of these works see Newton and Thayer (1992: 24).
Tribe
Himalusini
Klimaszewski, Pace and Center, 2010
Himalusini Klimaszewski et al., 2010: 3 [stem: Himalus-]. Type genus: Himalusa Pace, 2006.
Tribe
Homalotini
Heer, 1839
Homalotida Heer, 1839a: 305 [stem: Homalot-]. Type genus: Homalota Mannerheim, 1830.
Subtribe
Bolitocharina
Thomson, 1859
Bolitocharides C. G. Thomson, 1859: 31 [stem: Bolitochar-]. Type genus: Bolitochara Mannerheim, 1830 [placed on the Official List of Generic Names in Zoology (ICZN 1961a)]. Comment: First Revisers found (Bolitocharini C. G. Thomson, 1859 vs Euryusini C. G. Thomson, 1859) are Newton and Thayer (1992: 50); Bolitocharini C. G. Thomson, 1859 placed on the Official List of Family-Group Names in Zoology (ICZN 1961a); the original spelling Bolitocharides C. G. Thomson, 1859 and several subsequent spellings of this name placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1961a).
Euryusides C. G. Thomson, 1859: 40 [stem: Euryus-]. Type genus: Euryusa Erichson, 1837.
Sipaliae Casey, 1910: 167 [stem: Sipali-]. Type genus: Sipalia Mulsant and Rey, 1853 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 2005d)]. Comment: permanently invalid (Art. 39): based on suppressed type genus.
Heterotae Fenyes, 1918: 18 [stem: Heterot-]. Type genus: Heterota Mulsant and Rey, 1873.
Leptusae Fenyes, 1918: 18 [stem: Leptus-]. Type genus: Leptusa Kraatz, 1856 [placed on the Official List of Generic Names in Zoology (ICZN 2005d)].
Nanoglossae Fenyes, 1918: 20 [stem: Nanogloss-]. Type genus: Nanoglossa Fauvel, 1868 [subgenus of Leptusa Kraatz, 1856].
Ditropaliini Hatch, 1957: 134, in key, 147 [stem: Ditropali-]. Type genus: Ditropalia Casey, 1906 [syn. of Bolitochara Mannerheim, 1830]. Comment: this taxon was named correctly Ditropaliini in the text on p. 147 but incorrect as Bolitocharini in the key on p. 134.
Subtribe
Dinardopsina
Bernhauer and Scheerpeltz, 1926
Dinardopses Bernhauer and Scheerpeltz, 1926: 525, 804 [stem: Dinardops-]. Type genus: Dinardopsis Bruch, 1917. Comment: the erroneous spelling Dinardopsis was used in the catalogue on p. 525 but this was corrected to Dinardopses on page 804 of the same work; current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Dinardopse-).
Subtribe
Gyrophaenina
Kraatz, 1856
Gyrophaenini Kraatz, 1856: 351 [stem: Gyrophaen-]. Type genus: Gyrophaena Mannerheim, 1830.
Subtribe
Homalotina
Heer, 1839
Homalotida Heer, 1839a: 305 [stem: Homalot-]. Type genus: Homalota Mannerheim, 1830. Comment: this family-group name was also used in the same year by Heer (1839b: 50, as Homalotida); for comments about the priority of these works see Newton and Thayer (1992: 24).
Thecturotae Fenyes, 1918: 18 [stem: Thecturot-]. Type genus: Thecturota Casey, 1893.
*Cypheae Seevers, 1978: 272 [stem: Cyphe-]. Type genus: Cyphea Fauvel, 1863. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Silusina
Fenyes, 1918
Silusae Fenyes, 1918: 17 [stem: Silus-]. Type genus: Silusa Erichson, 1837.
Tribe
Hoplandriini
Casey, 1910
Hoplandriae Casey, 1910: 170 [stem: Hoplandri-]. Type genus: Hoplandria Kraatz, 1857.
Subtribe
Hoplandriina
Casey, 1910
Hoplandriae Casey, 1910: 170 [stem: Hoplandri-]. Type genus: Hoplandria Kraatz, 1857.
Subtribe
Platandriina
Hanley, 2002
Platandriina Hanley, 2002: 317 [stem: Platandri-]. Type genus: Platandria Casey, 1893.
Subtribe
Pseudoplandriina
Hanley, 2002
Pseudoplandriina Hanley, 2002: 317 [stem: Pseudoplandri-]. Type genus: Pseudoplandria Fenyes, 1921.
Tribe
Hygronomini
Thomson, 1859
Hygronomides C. G. Thomson, 1859: 31 [stem: Hygronom-]. Type genus: Hygronoma Erichson, 1837.
Subtribe
Hygronomina
Thomson, 1859
Hygronomides C. G. Thomson, 1859: 31 [stem: Hygronom-]. Type genus: Hygronoma Erichson, 1837.
Subtribe
Saphoglossina
Bernhauer and Scheerpeltz, 1926
Saphoglossae Bernhauer and Scheerpeltz, 1926: 521 [stem: Saphogloss-]. Type genus: Saphoglossa Sharp, 1883.
Tribe
Hypocyphtini
Laporte, 1835
Hypocyphtidae Laporte, 1835a: 135 [stem: Hypocypht-]. Type genus: Hypocyphtus Gyllenhal, 1827 [syn. of Cypha Leach, 1819].
Oligotides C. G. Thomson, 1859: 30 [stem: Oligot-]. Type genus: Oligota Mannerheim, 1830.
Nematoscelini Fenyes, 1921: 33 [stem: Nematoscelid-]. Type genus: Nematoscelis Wollaston, 1867. Comment: incorrect original stem formation, not in prevailing usage.
*Cyphinae Lohse, 1974: 7 [stem: Cyph-]. Type genus: Cypha Leach, 1819. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently; also see Cyphini Lacordaire, 1863 (type genus Cyphus Germar, 1824) in Coleoptera: Curculionidae.
Tribe
Leucocraspedini
Fenyes, 1921
Leucocraspedini Fenyes, 1921: 34 [stem: Leucocrasped-]. Type genus: Leucocraspedum Kraatz, 1859.
Tribe
Liparocephalini
Fenyes, 1918
Liparocephali Fenyes, 1918: 18 [stem: Liparocephal-]. Type genus: Liparocephalus Mäklin, 1853.
Tribe
Lomechusini
Fleming, 1821
Lomechusidae Fleming, 1821: 49 [stem: Lomechus-]. Type genus: Lomechusa Gravenhorst, 1806.
Subtribe
Aenictobiina
Kistner, 1997
Aenictobiina Kistner, 1997: 174 [stem: Aenictobi-]. Type genus: Aenictobia Seevers, 1953.
Subtribe
Lomechusina
Fleming, 1821
Lomechusidae Fleming, 1821: 49 [stem: Lomechus-]. Type genus: Lomechusa Gravenhorst, 1806.
Xenodusae Seevers, 1978: 155 [stem: Xenodus-]. Type genus: Xenodusa Wasmann, 1894.
Subtribe
Myrmedoniina
Thomson, 1867
Myrmedoniides C. G. Thomson, 1867: 209 [stem: Myrmedoni-]. Type genus: Myrmedonia Erichson, 1837 [syn. of Zyras Stephens, 1835].
*Myrméciates Mulsant and Rey, 1873b: 98 [stem: Myrmoeci-]. Type genus: Myrmoecia Mulsant and Rey, 1873. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Zyrini Bradley, 1930: 83 [stem: Zyr-]. Type genus: Zyras Stephens, 1835 [placed on the Official List of Generic Names in Zoology (ICZN 1961a)].
*Ecitoporae Seevers, 1978: 13 [stem: Ecitopor-]. Type genus: Ecitopora Wasmann, 1887. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Tetradoniae Seevers, 1978: 13 [stem: Tetradoni-]. Type genus: Tetradonia Wasmann, 1894. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Dinocorynae Seevers, 1978: 13 [stem: Dinocoryn-]. Type genus: Dinocoryna Casey, 1893. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Termitozyrina
Seevers, 1957
Termitozyrina Seevers, 1957: 62, in key [stem: Termitozyr-]. Type genus: Termitozyras Seevers, 1957.
Tribe
Masuriini
Cameron, 1939
Masuriini Cameron, 1939: 24 [stem: Masuri-]. Type genus: Masuria Cameron, 1928.
Tribe
Mesoporini
Cameron, 1959
Mesoporinae Cameron, 1959: 119 [stem: Mesopor-]. Type genus: Mesoporus Cameron, 1959.
Tribe
Mimanommatini
Wasmann, 1912
Mimanommatinae Wasmann, 1912: 478 [stem: Mimanommat-]. Type genus: Mimanomma Wasmann, 1912.
Subtribe
Dorylophilina
Fenyes, 1921
Dorylophilini Fenyes, 1921: 34 [stem: Dorylophil-]. Type genus: Dorylophila Wasmann, 1904.
Deremini Seevers, 1965: 294 [stem: Derem-]. Type genus: Derema Fauvel, 1899.
Subtribe
Mimanommatina
Wasmann, 1912
Mimanommatinae Wasmann, 1912: 478 [stem: Mimanommat-]. Type genus: Mimanomma Wasmann, 1912.
Tribe
Mimecitini
Wasmann, 1917
Mimecitonini Wasmann, 1917: 325 [stem: Mimecit-]. Type genus: Mimeciton Wasmann, 1893.
Subtribe
Labidopullina
Jacobson and Kistner, 1991
Labidopullina Jacobson and Kistner, 1991: 7 [stem: Labidopull-]. Type genus: Labidopullus Borgmeier, 1958.
Subtribe
Leptanillophilina
Fenyes, 1918
Leptanillophilini Fenyes, 1918: 59 [stem: Leptanillophil-]. Type genus: Leptanillophilus Holmgren, 1908.
Subtribe
Mimecitina
Wasmann, 1917
Mimecitonini Wasmann, 1917: 325 [stem: Mimecit-]. Type genus: Mimeciton Wasmann, 1893. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 52).
Subtribe
Mimonillina
Bernhauer and Scheerpeltz, 1926
Mimonillae Bernhauer and Scheerpeltz, 1926: 518 [stem: Mimonill-]. Type genus: Mimonilla Wasmann, 1913.
Tribe
Myllaenini
Ganglbauer, 1895
Myllaenini Ganglbauer, 1895: 317 [stem: Myllaen-]. Type genus: Myllaena Erichson, 1837.
Dimonomerini Cameron, 1933: 103 [stem: Dimonomer-]. Type genus: Dimonomera Cameron, 1933.
Tribe
Oxypodini
Thomson, 1859
Oxypodides C. G. Thomson, 1859: 36 [stem: Oxypod-]. Type genus: Oxypoda Mannerheim, 1830 [placed on the Official List of Generic Names in Zoology (ICZN 1957)]. Comment: First Revisers found (Oxypodini C. G. Thomson, 1859 vs Tachyusini C. G. Thomson, 1859 vs Ocaleini C. G. Thomson, 1859 vs Phloeoporini C. G. Thomson, 1859) are Newton and Thayer (1992: 53); name placed on the Official List of Family-Group Names in Zoology (ICZN 1957).
Subtribe
Aphytopodina
Bernhauer and Scheerpeltz, 1926
Aphytopi Bernhauer and Scheerpeltz, 1926: 740 [stem: Aphytopod-]. Type genus: Aphytopus Sharp, 1886. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 53).
Subtribe
Blepharhymenina
Klimaszewski and Peck, 1986
*Blepharhymeni Seevers, 1978: 82 [stem: Blepharhymen-]. Type genus: Blepharhymenus Solier, 1849. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Blepharrhymeni Klimaszewski and Peck, 1986: 58 [stem: Blepharhymen-]. Type genus: Blepharhymenus Solier, 1849 [as Blepharrhymenus, unjustified emendation of genus name by Gemminger and Harold (1868b: 505), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 53).
Subtribe
Dinardina
Mulsant and Rey, 1873
Dinardaires Mulsant and Rey, 1873a: 6 [stem: Dinard-]. Type genus: Dinarda Leach, 1819. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Wasmann (1904a: 218, as Dinardini), generally accepted as in Newton and Thayer (1992: 53, as Dinardina).
*Homéusates Mulsant and Rey, 1874: 286 [stem: Homoeus-]. Type genus: Homoeusa Kraatz, 1856. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Decusini Fenyes, 1918: 19 [stem: Decus-]. Type genus: Decusa Casey, 1900.
Subtribe
Meoticina
Seevers, 1978
Meoticae Seevers, 1978: 78 [stem: Meotic-]. Type genus: Meotica Mulsant and Rey, 1873.
Subtribe
Oxypodina
Thomson, 1859
Oxypodides C. G. Thomson, 1859: 36 [stem: Oxypod-]. Type genus: Oxypoda Mannerheim, 1830 [placed on the Official List of Generic Names in Zoology (ICZN 1957)]. Comment: First Reviser (Oxypodina C. G. Thomson, 1859 vs Ocaleina C. G. Thomson, 1859 vs Phloeoporina C. G. Thomson, 1859) not determined, current usage maintained; placed on the Official List of Family-Group Names in Zoology (ICZN 1957).
Ocaleides C. G. Thomson, 1859: 38 [stem: Ocale-]. Type genus: Ocalea Erichson, 1837.
Phloeoporides C. G. Thomson, 1859: 33 [stem: Phloeopor-]. Type genus: Phloeopora Erichson, 1837.
Ocyusates Mulsant and Rey, 1874: 286 [stem: Ocyus-]. Type genus: Ocyusa Kraatz, 1856. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Fenyes (1918: 20, as Ocyusae), generally accepted as in Newton and Thayer (1992: 53, as Ocyusina).
Calodérates Mulsant and Rey, 1874: 286 [stem: Caloder-]. Type genus: Calodera Mannerheim, 1830. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Fenyes (1918: 20, as Caloderae), generally accepted as in Ádám (2001:142, as Caloderini).
Microglottae Fenyes, 1918: 20 [stem: Microglott-]. Type genus: Microglotta Kraatz, 1862 [syn. of Haploglossa Kraatz, 1856].
Phloeoporini Cameron, 1939: 562 [stem: Phloeopor-]. Type genus: Phloeopora Erichson, 1837. Comment: proposed as new without reference to Phloeoporides C. G. Thomson, 1859.
Subtribe
Tachyusina
Thomson, 1859
Tachyusides C. G. Thomson, 1859: 34 [stem: Tachyus-]. Type genus: Tachyusa Erichson, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1961b)].
Tribe
Oxypodinini
Fenyes, 1921
Oxypodinini Fenyes, 1918: 18 [stem: Oxypodin-]. Type genus: Oxypodinus Bernhauer, 1901.
Heterotaxini Fenyes, 1921: 33 [stem: Heterotax-]. Type genus: Heterotaxus Bernhauer, 1915.
Tribe
Paglini
Newton and Thayer, 1992
Pachyglossini Fenyes, 1918: 60 [stem: Pachygloss-]. Type genus: Pachyglossa Fauvel, 1868 [preoccupied genus name, not Pachyglossa Hodgson, 1843 [Aves]; syn. of Pagla Blackwelder, 1952]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Paglini Newton and Thayer, 1992: 54 [stem: Pagl-]. Type genus: Pagla Blackwelder, 1952. Comment: replacement name for Pachyglossini Fenyes, 1918 because of the homonymy of the type genus.
Tribe
Paradoxenusini
Bruch, 1937
Paradoxenusini Bruch, 1937: 354 [stem: Paradoxenus-]. Type genus: Paradoxenusa Bruch, 1937.
Tribe
Pediculotini
Ádám, 1987
Pediculotini Ádám, 1987: 156 [stem: Pediculot-]. Type genus: Pediculota Ádám, 1987.
Tribe
Philotermitini
Seevers, 1957
Philotermitini Seevers, 1957: 63, in key [stem: Philotermit-]. Type genus: Philotermes Kraatz, 1857.
Tribe
Phyllodinardini
Wasmann, 1916
Phyllodinardini Wasmann, 1916a: 105 [stem: Phyllodinard-]. Type genus: Phyllodinarda Wasmann, 1916.
Tribe
Phytosini
Thomson, 1867
Phytosides C. G. Thomson, 1867: 206 [stem: Phytos-]. Type genus: Phytosus Curtis, 1838.
Tribe
Placusini
Mulsant and Rey, 1871
Placusates Mulsant and Rey, 1871c: 102 [stem: Placus-]. Type genus: Placusa Erichson, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Fenyes (1918: 17, as Placusae), generally accepted as in Newton and Thayer (1992: 54, as Placusini).
*Euvirae Seevers, 1978: 272 [stem: Euvir-]. Type genus: Euvira Sharp, 1883. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Pronomaeini
Mulsant and Rey, 1873
Pronoméates Mulsant and Rey, 1873b: 8 [stem: Pronomae-]. Type genus: Pronomaea Erichson, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Ganglbauer (1895: 315, as Pronomaeini), generally accepted as in Newton and Thayer (1992: 54, as Pronomaeini); incorrect original stem formation, not in prevailing usage.
Tribe
Pseudoperinthini
Cameron, 1939
Pseudoperinthinae Cameron, 1939: 1 [stem: Pseudoperinth-]. Type genus: Pseudoperinthus Wasmann, 1916.
Tribe
Pygostenini
Fauvel, 1899
Pygostenini Fauvel, 1899: 5 [stem: Pygosten-]. Type genus: Pygostenus Kraatz, 1858.
Sympolemonini Fenyes, 1918: 51 [stem: Sympolemont-]. Type genus: Sympolemon Wasmann, 1900. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Sahlbergiini
Kistner, 1993
Sahlbergiini Kistner, 1993: 315 [stem: Sahlbergi-]. Type genus: Sahlbergius Bernhauer, 1927.
Tribe
Sceptobiini
Seevers, 1978
Sceptobiini Seevers, 1978: 148 [stem: Sceptobi-]. Type genus: Sceptobius Sharp, 1883.
Tribe
Skatitoxenini
Kistner and Pasteels, 1969
Skatitoxenini Kistner and Pasteels, 1969: 1190 [stem: Skatitoxen-]. Type genus: Skatitoxenus Kistner and Pasteels, 1969.
Tribe
Termitodiscini
Wasmann, 1904
Termitodiscini Wasmann, 1904b: 656 [stem: Termitodisc-]. Type genus: Termitodiscus Wasmann, 1899.
Subtribe
Athexeniina
Pace, 2000
Athexenina Pace, 2000b: 336, in key [stem: Athexeni-]. Type genus: Athexenia Pace, 1999. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Termitodiscina
Wasmann, 1904
Termitodiscini Wasmann, 1904b: 656 [stem: Termitodisc-]. Type genus: Termitodiscus Wasmann, 1899.
Tribe
Termitohospitini
Seevers, 1941
Termitohospini Seevers, 1941: 331 [stem: Termitohospit-]. Type genus: Termitohospes Seevers, 1941.
Subtribe
Hetairotermitina
Seevers, 1957
Hetairotermitina Seevers, 1957: 191 [stem: Hetairotermit-]. Type genus: Hetairotermes Cameron, 1920.
Subtribe
Termitohospitina
Seevers, 1941
Termitohospini Seevers, 1941: 331 [stem: Termitohospit-]. Type genus: Termitohospes Seevers, 1941. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Seevers (1957: 191).
Tribe
Termitonannini
Fenyes, 1918
Termitonannini Fenyes, 1918: 75 [stem: Termitonann-]. Type genus: Termitonannus Wasmann, 1902.
Subtribe
Perinthina
Bernhauer and Scheerpeltz, 1926
Perinthi Bernhauer and Scheerpeltz, 1926: 521 [stem: Perinth-]. Type genus: Perinthus Casey, 1889.
*Poduroideae Scheerpeltz, 1934: 1537 [stem: Poduroid-]. Type genus: Poduroides Mann, 1926. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Termitonannina
Fenyes, 1918
Termitonannini Fenyes, 1918: 75 [stem: Termitonann-]. Type genus: Termitonannus Wasmann, 1902.
Tribe
Termitopaediini
Seevers, 1957
Termitopaediini Seevers, 1957: 214 [stem: Termitopaedi-]. Type genus: Termitopaedia Wasmann, 1911.
Termitondina Seevers, 1957: 63, in key [stem: Termitond-]. Type genus: Termitonda Seevers, 1957. Comment: as Termitendina, name spelled correctly on page 238.
Tribe
Termitusini
Fenyes, 1918
Termitusae Fenyes, 1918: 18 [stem: Termitus-]. Type genus: Termitusa Wasmann, 1905.
Subtribe
Termitospectrina
Seevers, 1957
Termitospectrina Seevers, 1957: 191 [stem: Termitospectr-]. Type genus: Termitospectrum Mann, 1926.
Subtribe
Termitusina
Fenyes, 1918
Termitusae Fenyes, 1918: 18 [stem: Termitus-]. Type genus: Termitusa Wasmann, 1905.
Tribe
Trichopseniini
LeConte and Horn, 1883
Trichopsenii J. L. LeConte and G. H. Horn, 1883: 100 [stem: Trichopseni-]. Type genus: Trichopsenius G. H. Horn, 1877.
Termitopsenini Wasmann, 1916b: 196 [stem: Termitopseni-]. Type genus: Termitopsenius Wasmann, 1902. Comment: incorrect original stem formation, not in prevailing usage.
Schizelythrinae Kemner, 1925: 122 [stem: Schizelythr-]. Type genus: Schizelythron Kemner, 1925.
Tribe
Trilobitideini
Fauvel, 1899
Trilobitideidae Fauvel, 1899: 3 [stem: Trilobitide-]. Type genus: Trilobitideus Raffray, 1898.
Subfamily
Trigonurinae
Reiche, 1866
Trigonurides Reiche, 1866: 642 [stem: Trigonur-]. Type genus: Trigonurus Mulsant, 1847.
Subfamily
Apateticinae
Fauvel, 1895
Apateticae Fauvel, 1895: 190 [stem: Apatetic-]. Type genus: Apatetica Westwood, 1848.
Subfamily
Scaphidiinae
Latreille, 1806
Scaphidilia Latreille, 1806: 3 [stem: Scaphidi-]. Type genus: Scaphidium A. G. Olivier, 1790.
Tribe
Cypariini
Achard, 1924
Cypariini Achard, 1924: 28 [stem: Cypari-]. Type genus: Cyparium Erichson, 1845.
Tribe
Scaphidiini
Latreille, 1806
Scaphidilia Latreille, 1806: 3 [stem: Scaphidi-]. Type genus: Scaphidium A. G. Olivier, 1790. Comment: incorrect original stem formation, not in prevailing usage.
Cerambyciscaphini Pic, 1915: 30 [stem: Cerambyciscaph-]. Type genus: Cerambyciscapha Pic, 1915.
Diateliitae Achard, 1924: 28 [stem: Diateli-]. Type genus: Diatelium Pascoe, 1863.
Tribe
Scaphiini
Achard, 1924
Scaphiitae Achard, 1924: 27 [stem: Scaphi-]. Type genus: Scaphium Kirby, 1837.
Tribe
Scaphisomatini
Casey, 1893
Scaphisomini Casey, 1893: 511 [stem: Scaphisomat-]. Type genus: Scaphisoma Leach, 1815. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 64).
Heteroscaphini Achard, 1914: 395 [stem: Heteroscaph-]. Type genus: Heteroscapha Achard, 1914 [syn. of Bironium Csiki, 1909].
Cyparellini Achard, 1924: 28 [stem: Cyparell-]. Type genus: Cyparella Achard, 1924 [syn. of Baeocera Erichson, 1845].
Baeoceritae Achard, 1924: 30 [stem: Baeocer-]. Type genus: Baeocera Erichson, 1845 [placed on the Official List of Generic Names in Zoology (ICZN 1982)].
Sciatrophitae Achard, 1924: 30 [stem: Sciatroph-]. Type genus: Sciatrophes Blackburn, 1903 [syn. of Baeocera Erichson, 1845].
Baeoceridiitae Achard, 1924: 30 [stem: Baeoceridi-]. Type genus: Baeoceridium Reitter, 1889.
Scaphicomitae Achard, 1924: 31 [stem: Scaphicom-]. Type genus: Scaphicoma Motschulsky, 1863.
Toxidiini Achard, 1924: 31 [stem: Toxidi-]. Type genus: Toxidium J. L. LeConte, 1860.
Subfamily
Piestinae
Erichson, 1839
Piestini Erichson, 1839b: 31 [stem: Piest-]. Type genus: Piestus Gravenhorst, 1806.
Prognathites Blanchard, 1845a: 290 [stem: Prognath-]. Type genus: Prognathus Berthold, 1827 [as Prognata, incorrect subsequent spelling of type genus name not in prevailing usage; syn. of Siagonium Kirby and Spence, 1815]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Blanchard (1853: 54, as Prognathitae).
*Siagoniini Crowson, 1980: 289 [stem: Siagoni-]. Type genus: Siagonium Kirby and Spence, 1815. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subfamily
Osoriinae
Erichson, 1839
Osorini Erichson, 1839b: 30 [stem: Osori-]. Type genus: Osorius Latreille, 1829.
Tribe
Eleusinini
Sharp, 1887
Eleusinina Sharp, 1887: 728 [stem: Eleusin-]. Type genus: Eleusis Laporte, 1835.
Tribe
Leptochirini
Sharp, 1887
Leptochirina Sharp, 1887: 733 [stem: Leptochir-]. Type genus: Leptochirus Germar, 1824.
Tribe
Osoriini
Erichson, 1839
Osorini Erichson, 1839b: 30 [stem: Osori-]. Type genus: Osorius Latreille, 1829.
Subtribe
Osoriina
Erichson, 1839
Osorini Erichson, 1839b: 30 [stem: Osori-]. Type genus: Osorius Latreille, 1829. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by J. L. LeConte (1861: 68).
Subtribe
Parosoriina
Bernhauer and Schubert, 1911
Parosorii Bernhauer and Schubert, 1911: 146 [stem: Parosori-]. Type genus: Parosorius Bernhauer, 1904.
Tribe
Thoracophorini
Reitter, 1909
Thoracophorinae Reitter, 1909: 199 [stem: Thoracophor-]. Type genus: Thoracophorus Motschulsky, 1837 [unjustified emendation of original type genus name Thoraxophorus by Erichson (1840a: 908); unjustified emendation in prevailing usage, treated as justified emendation (Art. 33.2.3.1)].
Subtribe
Clavilispinina
Newton and Thayer, 1992
Paralispini Blackwelder, 1942: 79 [stem: Paralispin-]. Type genus: Paralispinus Bernhauer, 1921 [preoccupied genus name, not Paralispinus Eichelbaum, 1913 [Coleoptera: Staphylinidae]; syn. of Clavilispinus Bernhauer, 1926]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Clavilispinina Newton and Thayer, 1992: 59 [stem: Clavilispin-]. Type genus: Clavilispinus Bernhauer, 1926. Comment: replacement name for Paralispini Blackwelder, 1942 because of the homonymy of the type genus.
Subtribe
Glyptomina
Newton and Thayer, 1992
Caloceri Blackwelder, 1942: 78 [stem: Calocer-]. Type genus: Calocerus Fauvel, 1891 [preoccupied genus name, not Calocerus J. L. LeConte, 1853 [Coleoptera: Elateridae]; syn. of Glyptoma Erichson, 1839]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Glyptomina Newton and Thayer, 1992: 59 [stem: Glyptom-]. Type genus: Glyptoma Erichson, 1839. Comment: replacement name for Caloceri Blackwelder, 1942 because of the homonymy of the type genus; current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Glyptomat-).
Subtribe
Lispinina
Bernhauer and Schubert, 1910
Lispini Bernhauer and Schubert, 1910: 19 [stem: Lispin-]. Type genus: Lispinus Erichson, 1839.
Subtribe
Thoracophorina
Reitter, 1909
Thoracophorinae Reitter, 1909: 199 [stem: Thoracophor-]. Type genus: Thoracophorus Motschulsky, 1837 [unjustified emendation of original type genus name Thoraxophorus by Erichson (1840a: 908); unjustified emendation in prevailing usage, treated as justified emendation (Art. 33.2.3.1)]. Comment: based on corrected spelling of type genus.
Subfamily
Oxytelinae
Fleming, 1821
Oxytelidae Fleming, 1821: 49 [stem: Oxytel-]. Type genus: Oxytelus Gravenhorst, 1802.
Tribe
Blediini
Ádám, 2001
Blediini Ádám, 2001: 216 [stem: Bledi-]. Type genus: Bledius Leach, 1819.
Tribe
Coprophilini
Heer, 1839
Coprophilina Heer, 1839a: 198 [stem: Coprophil-]. Type genus: Coprophilus Latreille, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1993a)]. Comment: this family-group name was also used in the same year by Erichson (1839b: 30, as Coprophilini) and Heer (1839b: 13, as Coprophilida); for comments about the priority of these works see Newton and Thayer (1992: 24).
*Homalotriquitos Solier, 1849: 321 [stem: Homalotrich-]. Type genus: Homalotrichus Solier, 1849. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Toxoderi Bernhauer and Schubert, 1911: 91 [stem: Toxoder-]. Type genus: Toxoderus Fauvel, 1900 [syn. of Homalotrichus Solier, 1849]. Comment: junior homonym of Toxoderini Saussure, 1869 (type genus Toxodera Audinet-Serville, 1837) in Mantodea: Mantidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Euphaniini
Reitter, 1909
*Pholidiens Mulsant and Rey, 1876: 209 [stem: Pholid-]. Type genus: Pholidus Mulsant and Rey, 1856 [preoccupied genus name, not Pholidus Rafinesque, 1815 [Pisces], or Pholidus Gray, 1840 [Aves]; syn. of Euphanias Fairmaire and Laboulbène, 1856]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant and Rey (1876).
Pholidini Acloque, 1896: 145 [stem: Pholid-]. Type genus: Pholidus Mulsant and Rey, 1856 [preoccupied genus name, not Pholidus Rafinesque, 1815 [Pisces], or Pholidus Gray, 1840 [Aves]; syn. of Euphanias Fairmaire and Laboulbène, 1856]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Euphaniae Reitter, 1909: 16 [stem: Euphani-]. Type genus: Euphanias Fairmaire and Laboulbène, 1856. Comment: precedence (Euphaniini Reitter, 1909 vs Deleasterini Reitter, 1909) given to taxon originally proposed at the higher rank (Art. 24.1).
Deleasterini Reitter, 1909: 164 [stem: Deleaster-]. Type genus: Deleaster Erichson, 1839.
Syntomiinae Böving and Craighead, 1931: 28 [stem: Syntomi-]. Type genus: Syntomium Curtis, 1828.
Tribe
Oxytelini
Fleming, 1821
Oxytelidae Fleming, 1821: 49 [stem: Oxytel-]. Type genus: Oxytelus Gravenhorst, 1802.
Thinobiides J. Sahlberg, 1876: 242 [stem: Thinobi-]. Type genus: Thinobius Kiesenwetter, 1844.
Trogophléaires Mulsant and Rey, 1878b: 688 [stem: Trogophloe-]. Type genus: Trogophloeus Mannerheim, 1830 [syn. of Carpelimus Leach, 1819]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1889 [Gatt.]: 90, as Trogophloeina), generally accepted as in Johansen (1914: 533, as Trogophloeina); incorrect original stem formation, not in prevailing usage.
Apocellaria Lynch Arribálzaga, 1884: 344 [stem: Apocell-]. Type genus: Apocellus Erichson, 1839.
Ecitoclimacini Borgmeier, 1934: 452 [stem: Ecitoclimac-]. Type genus: Ecitoclimax Borgmeier, 1934. Comment: originally proposed as a tribe of Aleocharinae.
Torrentomi Bierig, 1934: 213 [stem: Torrentom-]. Type genus: Torrentomus Bierig, 1934 [syn. of Thinobius Kiesenwetter, 1844].
Trigonobregmini Scheerpeltz, 1944: 170, in key [stem: Trigonobregmat-]. Type genus: Trigonobregma Scheerpeltz, 1944. Comment: incorrect original stem formation, not in prevailing usage; placement following Herman (2001).
Carpelimini Hatch, 1957: 85, in key [stem: Carpelim-]. Type genus: Carpelimus Leach, 1819.
*Thinodromini Gildenkov, 2000: 56 [stem: Thinodrom-]. Type genus: Thinodromus Kraatz, 1858. Comment: name unavailable (Art. 16.1): name not indicated as intentionally new.
Aploderini Ádám, 2001: 218 [stem: Aploder-]. Type genus: Aploderus Stephens, 1834.
Tribe
Planeustomini
Jacquelin du Val, 1857
Planeustomites Jacquelin du Val, 1857a: 58 [stem: Planeustom-]. Type genus: Planeustomus Jacquelin du Val, 1857. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Ádám (2001: 208, as Planeustomini).
Acrognathini Reitter, 1909: 164 [stem: Acrognath-]. Type genus: Acrognathus Erichson, 1839 [preoccupied genus name, not Acrognathus Agassiz, 1836 [Pisces]; syn. of Manda Blackwelder, 1952]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Mandini Gildenkov, 2003: 35 [stem: Mand-]. Type genus: Manda Blackwelder, 1952. Comment: replacement name for Acrognathini Reitter, 1909 because of the homonymy of the type genus.
Subfamily
Oxyporinae
Fleming, 1821
Oxyporidae Fleming, 1821: 49 [stem: Oxypor-]. Type genus: Oxyporus Fabricius, 1775.
Subfamily
Megalopsidiinae
Leng, 1920
Megalopini Erichson, 1839b: 30 [stem: Megalop-]. Type genus: Megalops Erichson, 1839 [preoccupied genus name, not Megalops Lacepède, 1803 [Pisces], not Megalops Rafinesque, 1815 [Pisces]; syn. of Megalopinus Eichelbaum, 1915]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Megalopsidiini Leng, 1920: 98 [stem: Megalopsidi-]. Type genus: Megalopsidia Leng, 1918 [syn. of Megalopinus Eichelbaum, 1915].
Aulacotrachelinae L. Benick, 1920: 1 [stem: Aulacotrachel-]. Type genus: Aulacotrachelus Benick, 1920 [syn. of Megalopinus Eichelbaum, 1915]. Comment: replacement name for Megalopinae Erichson, 1839 because of the homonymy of the type genus.
Stylopodinae Blackwelder, 1943: 202 [stem: Stylopod-]. Type genus: Stylopodus Benick, 1917 [syn. of Megalopinus Eichelbaum, 1915]. Comment: name proposed to replace Megalopinae Erichson, 1839 and Megalopsidiinae Leng, 1920 because of the synonymy of the type genus.
*Megalopininae Puthz, 1967: 192 [stem: Megalopin-]. Type genus: Megalopinus Eichelbaum, 1915. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Megalopininae Naomi, 1986: 344 [stem: Megalopin-]. Type genus: Megalopinus Eichelbaum, 1915. Comment: name proposed to replace Megalopinae Erichson, 1839 and Megalopsidiinae Leng, 1920 because of the synonymy of the type genus.
Subfamily
Scydmaeninae
Leach, 1815
Scydmaenides Leach, 1815: 92 [stem: Scydmaen-]. Type genus: Scydmaenus Latreille, 1802. Comment: placement follows Grebennikov and Newton (2009).
†Supertribe
Hapsomelitae
Poinar and Brown, 2004
Hapsomelinae Poinar and Brown, 2004: 790 [stem: Hapsomel-]. Type genus: Hapsomela Poinar and Brown, 2004.
Supertribe
Mastigitae
Fleming, 1821
Mastigoidae Fleming, 1821: 49 [stem: Mastig-]. Type genus: Mastigus Latreille, 1802. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Clidicini
Casey, 1897
Clidicini Casey, 1897: 541 [stem: Clidic-]. Type genus: Clidicus Laporte, 1833.
Tribe
Leptomastacini
Casey, 1897
Leptomastacini Casey, 1897: 541 [stem: Leptomastac-]. Type genus: Leptomastax Pirazzoli, 1855.
Tribe
Mastigini
Fleming, 1821
Mastigoidae Fleming, 1821: 49 [stem: Mastig-]. Type genus: Mastigus Latreille, 1802. Comment: incorrect original stem formation, not in prevailing usage.
Supertribe
Scydmaenitae
Leach, 1815
Scydmaenides Leach, 1815: 92 [stem: Scydmaen-]. Type genus: Scydmaenus Latreille, 1802.
Tribe
Cephenniini
Reitter, 1882
Cephenniini Reitter, 1882c: 142 [stem: Cephenni-]. Type genus: Cephennium Müller and Kunze, 1822.
Anisosphaeridae Tömösváry, 1883: 128 [stem: Anisosphaer-]. Type genus: Anisosphaera Tömösváry, 1883 [syn. of Cephennium Müller and Kunze, 1822]. Comment: family-group name originally based on larva only.
Tribe
Chevrolatiini
Reitter, 1882
Chevrolatini Reitter, 1882c: 142 [stem: Chevrolati-]. Type genus: Chevrolatia Jacquelin du Val, 1859. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Cyrtoscydmini
Schaufuss, 1889
Cyrtoscydmini L. W. Schaufuss, 1889: 2 [stem: Cyrtoscydm-]. Type genus: Cyrtoscydmus Motschulsky, 1869 [syn. of Stenichnus C. G. Thomson, 1859].
Glandulariidae L. W. Schaufuss, 1889: 3 [stem: Glandulari-]. Type genus: Glandularia L. W. Schaufuss, 1889 [syn. of Euconnus (Napochus) C. G. Thomson, 1859].
Euconnini Casey, 1897: 354 [stem: Euconn-]. Type genus: Euconnus C. G. Thomson, 1859.
Opresini Casey, 1897: 354 [stem: Opres-]. Type genus: Opresus Casey, 1897 [syn. of Microscydmus Saulcy and Croissandeau, 1893].
Lophioderini Casey, 1897: 356 [stem: Lophioder-]. Type genus: Lophioderus Casey, 1897.
Stenichnini Ganglbauer, 1898: 25 [stem: Stenichn-]. Type genus: Stenichnus C. G. Thomson, 1859.
Neuraphini Csiki, 1909b: 18 [stem: Neuraph-]. Type genus: Neuraphes C. G. Thomson, 1859 [this name is an incorrect subsequent spelling of Nevraphes, in prevailing usage and so deemed to be the correct original spelling (Art. 33.3.1)].
Syndicini Csiki, 1919: 17 [stem: Syndic-]. Type genus: Syndicus Motschulsky, 1851.
Sciacharini Csiki, 1919: 69 [stem: Sciacharit-]. Type genus: Sciacharis Broun, 1893. Comment: incorrect original stem formation, not in prevailing usage.
Siamitini H. Franz, 1989: 44 [stem: Siamit-]. Type genus: Siamites Franz, 1989.
Tribe
Eutheiini
Casey, 1897
Eutheiini Casey, 1897: 507 [stem: Euthei-]. Type genus: Eutheia Stephens, 1830.
Ascydmini Casey, 1897: 355 [stem: Ascydm-]. Type genus: Ascydmus Casey, 1897 [syn. of Euthiconus Reitter, 1882].
Tribe
Leptoscydmini
Casey, 1897
Leptoscydmini Casey, 1897: 355 [stem: Leptoscydm-]. Type genus: Leptoscydmus Casey, 1897.
Tribe
Plaumanniolini
Costa Lima, 1962
Plaumanniolinae Costa Lima, 1962: 415 [stem: Plaumanniol-]. Type genus: Plaumanniola Costa Lima, 1962. Comment: originally proposed as a subfamily of Ptinidae.
Tribe
Scydmaenini
Leach, 1815
Scydmaenides Leach, 1815: 92 [stem: Scydmaen-]. Type genus: Scydmaenus Latreille, 1802.
Eumicrini Reitter, 1882c: 192 [stem: Eumicr-]. Type genus: Eumicrus Laporte, 1833 [syn. of Scydmaenus Latreille, 1802].
Subfamily
Steninae
MacLeay, 1825
Stenidae W. S. MacLeay, 1825: 49 [stem: Sten-]. Type genus: Stenus Latreille, 1797. Comment: the younger name Steninae Fraser and Purves, 1960 (type genus Steno Gray, 1846) in Mammalia: Delphinidae is unavailable according to Newton and Thayer (1992: 66).
Subfamily
Euaesthetinae
Thomson, 1859
Euaesthetina C. G. Thomson, 1859: 42 [stem: Euaesthet-]. Type genus: Euaesthetus Gravenhorst, 1806.
Tribe
Alzadaesthetini
Scheerpeltz, 1974
Alzadaesthetini Scheerpeltz, 1974: 102, in key [stem: Alzadaesthet-]. Type genus: Alzadaesthetus Kistner, 1961.
Tribe
Austroesthetini
Cameron, 1944
Austroaesthetini Cameron, 1944: 69 [stem: Austroesthet-]. Type genus: Austroesthetus Oke, 1933 [as Austroaesthetus, unjustified emendation of genus name not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Euaesthetini
Thomson, 1859
Euaesthetina C. G. Thomson, 1859: 42 [stem: Euaesthet-]. Type genus: Euaesthetus Gravenhorst, 1806.
Tamotini Coiffait, 1984: 353 [stem: Tamot-]. Type genus: Tamotus L. W. Schaufuss, 1872.
Tribe
Fenderiini
Scheerpeltz, 1974
Fenderiini Scheerpeltz, 1974: 103, in key [stem: Fenderi-]. Type genus: Fenderia Hatch, 1957.
Tribe
Nordenskioldiini
Bernhauer and Schubert, 1911
Nordenskioeldiini Bernhauer and Schubert, 1911: 186 [stem: Nordenskioldi-]. Type genus: Nordenskioldia Sahlberg, 1880 [as Nordenskioeldia, incorrect subsequent spelling of genus name not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage (see Newton and Thayer 1992: 56).
Tribe
Stenaesthetini
Bernhauer and Schubert, 1911
Stenaesthetini Bernhauer and Schubert, 1911: 186 [stem: Stenaesthet-]. Type genus: Stenaesthetus Sharp, 1874.
Subfamily
Solieriinae
Newton and Thayer, 1992
Fisognatitos Solier, 1849: 303 [stem: Physognath-]. Type genus: Physognathus Solier, 1849 [preoccupied genus name, not Physognathus Agassiz, 1846 [Reptilia]; syn. of Solierius Bernhauer, 1921]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Kraatz (1859b: 3, as Physognathites [treated as Latin]); permanently invalid (Art. 39): based on preoccupied type genus.
Solieriinae Newton and Thayer, 1992: 27 [stem: Solieri-]. Type genus: Solierius Bernhauer, 1921. Comment: replacement name for Physognathinae Solier, 1849 because of the homonymy of the type genus.
Subfamily
Leptotyphlinae
Fauvel, 1874
Leptotyphli Fauvel, 1874: 35 [stem: Leptotyphl-]. Type genus: Leptotyphlus Fauvel, 1874.
Tribe
Cephalotyphlini
Coiffait, 1963
Cephalotyphlini Coiffait, 1963: 380, in key [stem: Cephalotyphl-]. Type genus: Cephalotyphlus Coiffait, 1955.
Tribe
Entomoculiini
Coiffait, 1957
Entomoculini Coiffait, 1957: 61 [stem: Entomoculi-]. Type genus: Entomoculia Croissandeau, 1891. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 56).
Tribe
Leptotyphlini
Fauvel, 1874
Leptotyphli Fauvel, 1874: 35 [stem: Leptotyphl-]. Type genus: Leptotyphlus Fauvel, 1874.
Leptotyphlini Coiffait, 1957: 61 [stem: Leptotyphl-]. Type genus: Leptotyphlus Fauvel, 1874. Comment: family-group name proposed as new without reference to Leptotyphli Fauvel, 1874.
Tribe
Metrotyphlini
Coiffait, 1963
Metrotyphlini Coiffait, 1963: 381 [stem: Metrotyphl-]. Type genus: Metrotyphlus Coiffait, 1959.
Tribe
Neotyphlini
Coiffait, 1963
Neotyphlini Coiffait, 1963: 381, in key [stem: Neotyphl-]. Type genus: Neotyphlus Coiffait, 1959.
Subfamily
Pseudopsinae
Ganglbauer, 1895
Pseudopsini Ganglbauer, 1895: 690 [stem: Pseudops-]. Type genus: Pseudopsis Newman, 1834. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Pseudopse-).
Subfamily
Paederinae
Fleming, 1821
Poederidae Fleming, 1821: 49 [stem: Paeder-]. Type genus: Paederus Fabricius, 1775.
Tribe
Paederini
Fleming, 1821
Poederidae Fleming, 1821: 49 [stem: Paeder-]. Type genus: Paederus Fabricius, 1775.
Subtribe
Astenina
Hatch, 1957
Suniina Sharp, 1886b: 591 [stem: Suni-]. Type genus: Sunius sensu Erichson, 1839 [not Sunius Stephens, 1829; syn. of Astenus Dejean, 1833]. Comment: based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).
Astenina Hatch, 1957: 151, in key [stem: Asten-]. Type genus: Astenus Dejean, 1833. Comment: although this is not the oldest name for the subtribe, we recommend that an application be submitted to the Commission to suppress Suniina Sharp, 1886 because it is based on a misidentified type genus (Art. 65.2.1).
Subtribe
Cryptobiina
Casey, 1905
Cryptobia Casey, 1905: 21 [stem: Cryptobi-]. Type genus: Cryptobium Mannerheim, 1830 [syn. of Ochthephilum Stephens, 1829]. Comment: the younger name Cryptobiinae Hollande, 1952 (type genus Cryptobia Leidy, 1846) in Protozoa: Bodonidae is unavailable (see Newton and Thayer 1992: 61).
Cryptobiina Bordoni, 1975: 420 [stem: Cryptobi-]. Type genus: Cryptobium Mannerheim, 1830 [syn. of Ochthephilum Stephens, 1829]. Comment: family-group name proposed as new without reference to Cryptobia Casey, 1905.
Subtribe
Cylindroxystina
Bierig, 1943
Cylindroxystini Bierig, 1943: 158 [stem: Cylindroxyst-]. Type genus: Cylindroxystus Bierig, 1943.
Subtribe
Dolicaonina
Casey, 1905
Gnatimenitos Solier, 1849: 326 [stem: Gnathymen-]. Type genus: Gnathymenus Solier, 1849. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Ádám (2001: 115, as Gnathymenini); incorrect original stem formation, not in prevailing usage; this name was treated as unavailable by Newton and Thayer (1992: 61) which led to their use of Dolicaonina Casey, 1905 as the valid name for this subtribe; all subsequent authors have followed Newton and Thayer (1992: 61) except for the subsequent use of “Gnathymenini (Solier, 1849)” as valid by Ádám (2001: 115) which made Solier’s name available according to our criteria of availability for names originally proposed in vernacular form; here we continue to use Dolicaonina Casey, 1905 as the valid name for this subtribe and recommend that an application be submitted to the Commission to suppress the name Gnathymenina Solier, 1849.
Dolicaones Casey, 1905: 56 [stem: Dolicaon-]. Type genus: Dolicaon Laporte, 1835. Comment: see comments under the name Gnatimenitos Solier, 1849 above.
Leptobii Bordoni, 1980: 170 [stem: Leptobi-]. Type genus: Leptobium Casey, 1905. Comment: unnecessary replacement name for “Dolicaina Bordoni, 1975”.
Subtribe
Echiasterina
Casey, 1905
Echiasteres Casey, 1905: 245 [stem: Echiaster-]. Type genus: Echiaster Erichson, 1839.
Subtribe
Lathrobiina
Laporte, 1835
Lathrobidae Laporte, 1835a: 117 [stem: Lathrobi-]. Type genus: Lathrobium Gravenhorst, 1802. Comment: incorrect original stem formation, not in prevailing usage.
Sphaeronia Casey, 1905: 54 [stem: Sphaeron-]. Type genus: Sphaeronum Sharp, 1876 [as Sphaeronium, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Lithocharina
Casey, 1905
Lithochares Casey, 1905: 146 [stem: Lithochar-]. Type genus: Lithocharis Dejean, 1833. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Lithocharit-).
Lithocharina Bordoni, 1974: 324 [stem: Lithochar-]. Type genus: Lithocharis Dejean, 1833. Comment: family-group name proposed as new without reference to Lithochares Casey, 1905.
Subtribe
Medonina
Casey, 1905
Medones Casey, 1905: 20 [stem: Medon-]. Type genus: Medon Stephens, 1833. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Medont-).
Medina Bordoni, 1975: 420 [stem: Medon-]. Type genus: Medon Stephens, 1833. Comment: family-group name proposed as new without reference to Medones Casey, 1905; incorrect original stem formation, not in prevailing usage (should be Medont-).
Acanthoglossi Coiffait, 1982: 10 [stem: Acanthogloss-]. Type genus: Acanthoglossa Kraatz, 1859.
Subtribe
Paederina
Fleming, 1821
Poederidae Fleming, 1821: 49 [stem: Paeder-]. Type genus: Paederus Fabricius, 1775 [as Poederus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage, see comments in Newton and Thayer (1992: 60).
Geopaederidae Gistel, 1848: [13] [stem: Geopaeder-]. Type genus: Geopaederus Gistel, 1848 [syn. of Paederus Fabricius, 1775].
Subtribe
Scopaeina
Mulsant and Rey, 1878
*Poliodontidos Solier, 1849: 303 [stem: Polyodont-]. Type genus: Polyodontus Solier, 1849 [preoccupied genus name, not Polyodontus Eysenhardt, 1818 [Vermes]; syn. of Scopaeus Erichson, 1839]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if found to be available then permanently invalid (Art. 39): based on preoccupied type genus; Polyodontidae Bonaparte, 1838 (type genus Polyodon Lacepède, 1797) is available in Pisces; incorrect original stem formation, not in prevailing usage.
Scopéates Mulsant and Rey, 1878a: 178 [stem: Scopae-]. Type genus: Scopaeus Erichson, 1839. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1889 [Gatt.]: 92, as Scopaeina), generally accepted as in Newton and Thayer (1992: 62, as Scopaeina); incorrect original stem formation, not in prevailing usage.
Subtribe
Stilicina
Casey, 1905
Stilici Casey, 1905: 218 [stem: Stilic-]. Type genus: Stilicus Berthold, 1827 [syn. of Rugilus Leach, 1819].
Rugilina Hatch, 1957: 151, in key [stem: Rugil-]. Type genus: Rugilus Leach, 1819.
Subtribe
Stilicopsina
Casey, 1905
Stilicopses Casey, 1905: 230 [stem: Stilicops-]. Type genus: Stilicopsis Sachse, 1852. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Stilicopse-).
Xenasteres Bierig, 1939: 179 [stem: Xenaster-]. Type genus: Xenaster Bierig, 1939 [preoccupied genus name, not Xenaster Simonwitsch, 1871 [Echinodermata]; the nomenclatural status of Xenaster will be addressed in the near future by L. Herman (pers. comm. 2010) therefore we refrain from proposing a new replacement name here]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
*Stamnoderes Blackwelder, 1944: 126 [stem: Stamnoder-]. Type genus: Stamnoderus Sharp, 1886. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Pinophilini
Nordmann, 1837
Pinophiliniformes Nordmann, 1837: 6 [stem: Pinophil-]. Type genus: Pinophilus Gravenhorst, 1802.
Subtribe
Pinophilina
Nordmann, 1837
Pinophiliniformes Nordmann, 1837: 6 [stem: Pinophil-]. Type genus: Pinophilus Gravenhorst, 1802.
Subtribe
Procirrina
Bernhauer and Schubert, 1912
Procirri Bernhauer and Schubert, 1912: 197 [stem: Procirr-]. Type genus: Procirrus Latreille, 1829.
Subfamily
Staphylininae
Latreille, 1802
Staphyliniae Latreille, 1802: 124 [stem: Staphylin-]. Type genus: Staphylinus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1959a)].
Tribe
Arrowinini
Solodovnikov and Newton, 2005
Arrowinini Solodovnikov and Newton, 2005: 420 [stem: Arrowin-]. Type genus: Arrowinus Bernhauer, 1935.
Tribe
Diochini
Casey, 1906
Diochi Casey, 1906: 429 [stem: Dioch-]. Type genus: Diochus Erichson, 1839.
Diochinae I. Moore, 1964: 86, in key [stem: Dioch-]. Type genus: Diochus Erichson, 1839. Comment: family-group name proposed as new without reference to Diochi Casey, 1906.
Tribe
Maorothiini
Assing, 2000
Maorothiini Assing, 2000: 16 [stem: Maorothi-]. Type genus: Maorothius Assing, 2000.
Tribe
Othiini
Thomson, 1859
Othiides C. G. Thomson, 1859: 26 [stem: Othi-]. Type genus: Othius Stephens, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1983c)].
Atrecini Hatch, 1957: 172, in key [stem: Atrec-]. Type genus: Atrecus Jacquelin du Val, 1856.
Tribe
Platyprosopini
Lynch Arribálzaga, 1884
Platyprosoparia Lynch Arribálzaga, 1884: 165 [stem: Platyprosop-]. Type genus: Platyprosopus Mannerheim, 1830.
Platyprosopinae I. Moore, 1964: 86, in key [stem: Platyprosop-]. Type genus: Platyprosopus Mannerheim, 1830. Comment: family-group name proposed as new without reference to Platyprosopinae Lynch Arribálzaga, 1884.
Tribe
Staphylinini
Latreille, 1802
Staphyliniae Latreille, 1802: 124 [stem: Staphylin-]. Type genus: Staphylinus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1959a)]. Comment: placed on the Official List of Family-Group Names in Zoology (as Staphylinidae Latreille, [1803-1804]) (ICZN 1959a).
Subtribe
Amblyopinina
Seevers, 1944
Amblyopininae Seevers, 1944: 157 [stem: Amblyopin-]. Type genus: Amblyopinus Solsky, 1875.
Heterothopsi Coiffait, 1978: 300 [stem: Heterothop-]. Type genus: Heterothops Stephens, 1829. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 65); placement based on Chatzimanolis et al. (2010).
Subtribe
Anisolinina
Hayashi, 1993
Anisolinina Hayashi, 1993: 288 [stem: Anisolin-]. Type genus: Anisolinus Sharp, 1889. Comment: originally proposed as a subtribe of Philonthini Kirby, 1837.
Subtribe
Eucibdelina
Sharp, 1889
Eucibdelini Sharp, 1889: 112 [stem: Eucibdel-]. Type genus: Eucibdelus Kraatz, 1859.
Subtribe
Hyptiomina
Casey, 1906
Hyptiomae Casey, 1906: 361 [stem: Hyptiom-]. Type genus: Hyptioma Casey, 1906 [syn. of Holisus Erichson, 1839]. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Hyptiomat-).
*Holisi Blackwelder, 1944: 143 [stem: Holis-]. Type genus: Holisus Erichson, 1839. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Holisina Newton, 1988: 259 [stem: Holis-]. Type genus: Holisus Erichson, 1839.
Subtribe
Philonthina
Kirby, 1837
Philonthidae Kirby, 1837: 91 [stem: Philonth-]. Type genus: Philonthus Curtis, 1829.
*Rémates Mulsant and Rey, 1876: 596 [stem: Rem-]. Type genus: Remus sensu C. G. Thomson, 1859 [not Remus Holme, 1837; syn. of Erichsonius Fauvel, 1874]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; also based on a misidentified type genus.
Craspedomeri Bernhauer, 1911: 88 [stem: Craspedomer-]. Type genus: Craspedomerus Bernhauer, 1911.
Subtribe
Quediina
Kraatz, 1857
Quediiformes Kraatz, 1857: 473 [stem: Quedi-]. Type genus: Quedius Stephens, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1996c)]. Comment: use of family-group name given precedence over Platycnemini Nordmann 1837 and "Quediini Kraatz, [1857]" placed on the Official List of Family-Group Names in Zoology (ICZN 1996c).
*Acylophorini Scheerpeltz, 1968: 97 [stem: Acylophor-]. Type genus: Acylophorus Nordmann, 1837. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Acylophorini Outerelo and Gamarra, 1985: 48, in key [stem: Acylophor-]. Type genus: Acylophorus Nordmann, 1837.
Subtribe
Staphylinina
Latreille, 1802
Staphyliniae Latreille, 1802: 124 [stem: Staphylin-]. Type genus: Staphylinus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1959a)]. Comment: placed on the Official List of Family-Group Names in Zoology (as Staphylinidae Latreille, [1803-1804]) and “Staphylinii Latreille, [1803-1804]” placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1959a).
Creophilidae Kirby, 1837: 95 [stem: Creophil-]. Type genus: Creophilus Leach, 1819 [Creophilus (attributed to Samouelle, 1819) placed on the Official List of Generic Names in Zoology (ICZN 1959a)].
Thinopininae Böving and Craighead, 1931: 30 [stem: Thinopin-]. Type genus: Thinopinus J. L. LeConte, 1852.
Ocypina Hatch, 1957: 173, in key [stem: Ocypod-]. Type genus: Ocypus Leach, 1819. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Newton and Thayer (1992: 65).
Subtribe
Tanygnathinina
Reitter, 1909
Tanygnathinini Reitter, 1909: 105 [stem: Tanygnathin-]. Type genus: Tanygnathinus Reitter, 1909 [syn. of Atanygnathus Jakobson, 1909].
Tanygnathini Casey, 1915b: 424 [stem: Tanygnath-]. Type genus: Tanygnathus Erichson, 1839 [preoccupied genus name, not Tanygnathus Wagler, 1832 [Aves]; syn. of Atanygnathus Jakobson, 1909]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Atanygnathini Lohse, 1964: 220 [stem: Atanygnath-]. Type genus: Atanygnathus Jakobson, 1909.
Subtribe
Xanthopygina
Sharp, 1884
Platycnemidiformes Nordmann, 1837: 6 [stem: Platycnem-]. Type genus: Platycnemus Nordmann, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1996c); syn. of Haematodes Laporte, 1835]. Comment: younger name Xanthopygina Sharp, 1884 given precedence over this name (ICZN 1996c); placed on the Official List of Family-Group Names in Zoology (ICZN 1996c, as Platycnemini Nordmann, 1837).
Xanthopygina Sharp, 1884: 342 [stem: Xanthopyg-]. Type genus: Xanthopygus Kraatz, 1857. Comment: family-group name given precedence over Platycnemini Nordmann, 1837 and placed on the Official List of Family-Group Names in Zoology (ICZN 1996c).
Triacri Bernhauer, 1931: 84 [stem: Triacr-]. Type genus: Triacrus Nordmann, 1837.
Tribe
Xantholinini
Erichson, 1839
Agraeformes Nordmann, 1837: 7 [stem: Agrod-]. Type genus: Agrodes Nordmann, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1996c); syn. of Plochionocerus Dejean, 1833]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1996c, as Agrodini Nordmann, 1837); younger name Xantholinini Erichson, 1839 given precedence over this name (ICZN 1996c).
Gyrohypnidae Kirby, 1837: 88 [stem: Gyrohypn-]. Type genus: Gyrohypnus Leach, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 1983c, as Gyrohypnus Samouelle, 1819)]. Comment: placed on the Official List of Family-Group Names in Zoology (ICZN 1996c, as Gyrohypnini Kirby, 1837); younger name Xantholinini Erichson, 1839 given precedence over this name (ICZN 1996c).
Xantholinini Erichson, 1839b: 28 [stem: Xantholin-]. Type genus: Xantholinus Dejean, 1821 [placed on the Official List of Generic Names in Zoology (ICZN 1983c)]. Comment: family-group name given precedence over Agrodini Nordmann, 1837 and Gyrohypnini Kirby, 1837 and placed on the Official List of Family-Group Names in Zoology (ICZN 1996c).
Araeocnemes Casey, 1906: 359 [stem: Araeocnem-]. Type genus: Araeocnemus Nordmann, 1837 [as Araeocnemis, incorrect subsequent spelling of type genus name, not in prevailing usage; syn. of Plochionocerus Dejean, 1833].
Metoponci Casey, 1906: 360 [stem: Metoponc-]. Type genus: Metoponcus Kraatz, 1857 [syn. of Zeteotomus Jacquelin du Val, 1856].
†Subfamily
Protactinae
Heer, 1847
Protactiden Heer, 1847: 28 [stem: Protact-]. Type genus: Protactus Heer, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Bronn (1848: 1045, as Protactidae), generally accepted as in Herman (2001: 3839, as Protactinae).
Staphylinoidea
incertae sedis
Hameedini M. Abdullah and Quadri, 1968: 310 [stem: Hameedi-]. Type genus: Hameedia M. Abdullah and Quadri, 1968. Comment: incorrect original stem formation, not in prevailing usage; originally proposed as a tribe of Staphylinidae: Oxyporinae; correct placement uncertain (see Newton and Thayer 1992: 67).
Superfamily
Scarabaeoidea
Latreille, 1802
Scarabaeïdes Latreille, 1802: 144 [stem: Scarabae-]. Type genus: Scarabaeus Linnaeus, 1758. Comment: First Reviser (Scarabaeoidea Latreille, 1802 vs Geotrupoidea Latreille, 1802) not determined, current usage maintained.
Family
Pleocomidae
LeConte, 1861
Pleocomini J. L. LeConte, 1861: 128 [stem: Pleocom-]. Type genus: Pleocoma J. L. LeConte, 1856.
Subfamily
Pleocominae
LeConte, 1861
Pleocomini J. L. LeConte, 1861: 128 [stem: Pleocom-]. Type genus: Pleocoma J. L. LeConte, 1856.
†Subfamily
Cretocominae
Nikolajev, 2002
Cretocomini Nikolajev, 2002: 53 [stem: Cretocom-]. Type genus: Cretocoma Nikolajev, 2002.
†Subfamily
Archescarabaeinae
Nikolajev, 2010
Archescarabaeinae Nikolajev, 2010: 69 [stem: Archescarabae-]. Type genus: Archescarabaeus Nikolajev, 2010.
Family
Geotrupidae
Latreille, 1802
Geotrupini Latreille, 1802: 142 [stem: Geotrup-]. Type genus: Geotrupes Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 1955b)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1955c, as Geotrupini Latreille, 1806).
Subfamily
Taurocerastinae
Germain, 1897
Taurocerastidae Germain, 1897: 288 [stem: Taurocerast-]. Type genus: Taurocerastes Philippi, 1866.
Subfamily
Bolboceratinae
Mulsant, 1842
Bolbocéraires Mulsant, 1842: 347 [stem: Bolbocerat-]. Type genus: Bolboceras Kirby, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 2006a)].
Tribe
Athyreini
Lynch Arribálzaga, 1878
*Athyréites Blanchard, 1845a: 221 [stem: Athyre-]. Type genus: Athyreus W. S. MacLeay, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 1985a)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845a) (see A. B. T. Smith 2006: 148).
Athyreitae Lynch Arribálzaga, 1878: 145 [stem: Athyre-]. Type genus: Athyreus W. S. MacLeay, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 1985a)]. Comment: name previously attributed to Howden and Martínez (1963).
Athyreini Howden and Martínez, 1963: 346 [stem: Athyre-]. Type genus: Athyreus W. S. MacLeay, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 1985a)]. Comment: family-group name proposed as new without reference to Athyréides Blanchard, 1845 or Athyreitae Lynch Arribálzaga, 1878.
Tribe
Bolbelasmini
Nikolajev, 1996
Bolbelasmini Nikolajev, 1996: 96 [stem: Bolbelasm-]. Type genus: Bolbelasmus Boucomont, 1911.
Tribe
Bolboceratini
Mulsant, 1842
Bolbocéraires Mulsant, 1842: 347 [stem: Bolbocerat-]. Type genus: Bolboceras Kirby, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 2006a)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Boucomont (1911: 333, as Bolbocerinae [incorrect stem formation]), generally accepted as in A. B. T. Smith (2006: 148, as Bolboceratini); concept of Bolboceras Kirby, 1819 fixed by the Commission (ICZN 2006a); incorrect original stem formation, not in prevailing usage.
Australobolbini Nikolajev, 1996: 96 [stem: Australobolb-]. Type genus: Australobolbus Howden and Cooper, 1977.
Tribe
Bolbochromini
Nikolajev, 1970
Bolbochromini Nikolajev, 1970: 34 [stem: Bolbochrom-]. Type genus: Bolbochromus Boucomont, 1909.
Tribe
Eubolbitini
Nikolajev, 1970
Eubolbitini Nikolajev, 1970: 33 [stem: Eubolbit-]. Type genus: Eubolbitus Reitter, 1892.
Tribe
Eucanthini
Nikolajev, 2003
Eucanthini Nikolajev, 2003b: 209 [stem: Eucanth-]. Type genus: Eucanthus Westwood, 1848.
Tribe
Gilletinini
Nikolajev, 1990
Gilletinini Nikolajev, 1990: 99 [stem: Gilletin-]. Type genus: Gilletinus Boucomont, 1932.
Tribe
Odonteini
Shokhin, 2007
Odonteini Shokhin, 2007: 111 [stem: Odonte-]. Type genus: Odonteus Samouelle, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 2006a)]. Comment: replacement name for Bolboceratini of authors because the concept of Bolboceras Kirby, 1819 was fixed differently by the Commission (ICZN 2006a).
Tribe
Stenaspidiini
Nikolajev, 2003
Stenaspidiini Nikolajev, 2003a: 190, in key [stem: Stenaspidi-]. Type genus: Stenaspidius Westwood, 1848.
Subfamily
Geotrupinae
Latreille, 1802
Geotrupini Latreille, 1802: 142 [stem: Geotrup-]. Type genus: Geotrupes Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 1955b)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1955c, as Geotrupini Latreille, 1806).
Tribe
Ceratotrupini
Zunino, 1984
Ceratotrupini Zunino, 1984: 89 [stem: Ceratotrup-]. Type genus: Ceratotrupes Jekel, 1866.
Tribe
Enoplotrupini
Paulian, 1945
Enoplotrupini Paulian, 1945: 40, in key [stem: Enoplotrup-]. Type genus: Enoplotrupes Lucas, 1869.
Chromogeotrupini Zunino, 1984: 30 [stem: Chromogeotrup-]. Type genus: Chromogeotrupes Bovo and Zunino, 1983.
†Tribe
Cretogeotrupini
Nikolajev, 1996
Cretogeotrupini Nikolajev, 1996: 97 [stem: Cretogeotrup-]. Type genus: Cretogeotrupes Nikolajev, 1992.
Tribe
Geotrupini
Latreille, 1802
Geotrupini Latreille, 1802: 142 [stem: Geotrup-]. Type genus: Geotrupes Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 1955b)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1955c, as Geotrupini Latreille, 1806).
Tribe
Lethrini
Oken, 1843
Lethren Oken, 1843: 484 [stem: Lethr-]. Type genus: Lethrus Scopoli, 1777. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Tulk (1847: 614, as Lethri); name previously attributed to Mulsant and Rey (1871a).
Family
Belohinidae
Paulian, 1959
Belohininae Paulian, 1959: 40 [stem: Belohin-]. Type genus: Belohina Paulian, 1959.
Family
Passalidae
Leach, 1815
Passalida Leach, 1815: 100 [stem: Passal-]. Type genus: Passalus Fabricius, 1792.
Subfamily
Aulacocyclinae
Kaup, 1868
Aulacocyclinae Kaup, 1868a: 4 [stem: Aulacocycl-]. Type genus: Aulacocyclus Kaup, 1868.
Tribe
Aulacocyclini
Kaup, 1868
Aulacocyclinae Kaup, 1868a: 4 [stem: Aulacocycl-]. Type genus: Aulacocyclus Kaup, 1868.
Tribe
Ceracupedini
Boucher, 2006
Ceracupini Boucher, 2006: 319 [stem: Ceracuped-]. Type genus: Ceracupes Kaup, 1871. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Passalinae
Leach, 1815
Passalida Leach, 1815: 100 [stem: Passal-]. Type genus: Passalus Fabricius, 1792.
Tribe
Leptaulacini
Kaup, 1871
Leptaulaceae Kaup, 1871: 28 [stem: Leptaulac-]. Type genus: Leptaulax Kaup, 1868.
Tribe
Macrolinini
Kaup, 1871
Macrolineae Kaup, 1871: 42 [stem: Macrolin-]. Type genus: Macrolinus Kaup, 1868. Comment: First Reviser found (Macrolinini Kaup, 1871 vs Aceraiini Kaup, 1871) is Gravely (1918: 76).
Aceraiae Kaup, 1871: 47 [stem: Acerai-]. Type genus: Aceraius Kaup, 1868.
Eriocnemiae Kaup, 1871: 35 [stem: Eriocnemid-]. Type genus: Eriocnemis Kaup, 1868 [preoccupied genus name, not Eriocnemis Reichenbach, 1849 [Aves]; syn. of Pelopides Kuwert, 1896]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Gonatinae Kuwert, 1891: 169 [stem: Gonat-]. Type genus: Gonatas Kaup, 1871.
Mastachilinae Kuwert, 1891: 167 [stem: Mastachil-]. Type genus: Mastachilus Kaup, 1868.
Pharochilinae Kuwert, 1891: 166 [stem: Pharochil-]. Type genus: Pharochilus Kaup, 1868.
Tarquininae Kuwert, 1891: 164 [stem: Tarquini-]. Type genus: Tarquinius Kuwert, 1891. Comment: incorrect original stem formation, not in prevailing usage.
Vellejinae Kuwert, 1891: 167 [stem: Vellej-]. Type genus: Vellejus Kaup, 1871 [preoccupied genus name, not Vellejus Mannerheim, 1830 [Coleoptera: Staphylinidae], not Vellejus Stål, 1865 [Hemiptera]; syn. of Labienus Kaup, 1871]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Aureliinae Kuwert, 1896: 230 [stem: Aureli-]. Type genus: Aurelius Kuwert, 1891.
Lachinae Kuwert, 1896: 230 [stem: Lach-]. Type genus: Laches Kaup, 1871 [preoccupied genus name, not Laches Gistel, 1848 [Hymenoptera], not Laches Thorell, 1869 [Arachnida]; syn. of Aceraius Kaup, 1868]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Pelopinae Kuwert, 1896: 229 [stem: Pelopid-]. Type genus: Pelopides Kuwert, 1896. Comment: incorrect original stem formation, not in prevailing usage.
Pleurariinae Kuwert, 1896: 224 [stem: Pleurari-]. Type genus: Pleurarius Kaup, 1868.
Gnaphalocneminae Gravely, 1914: 194 [stem: Gnaphalocnemid-]. Type genus: Gnaphalocnemis Heller, 1900 [syn. of Pelopides Kuwert, 1896]. Comment: incorrect original stem formation, not in prevailing usage.
Austropassalinae Mjöberg, 1917: 11 [stem: Austropassal-]. Type genus: Austropassalus Mjöberg, 1917.
Tribe
Passalini
Leach, 1815
Passalida Leach, 1815: 100 [stem: Passal-]. Type genus: Passalus Fabricius, 1792.
Neleinae Kaup, 1869: 28 [stem: Nele-]. Type genus: Neleus Kaup, 1869 [preoccupied genus name, not Neleus Desbonne and Schramm, 1867 [Crustacea]; syn. of Passalus Fabricius, 1792]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Pertinaceae Kaup, 1871: 89 [stem: Pertinac-]. Type genus: Pertinax Kaup, 1869.
Phoroneae Kaup, 1871: 97 [stem: Phorone-]. Type genus: Phoroneus Kaup, 1869 [preoccupied genus name, not Phoroneus Stål, 1865 [Hemiptera]; syn. of Passalus Fabricius, 1792]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Mitrorhinae Kuwert, 1891: 190 [stem: Mitrorhin-]. Type genus: Mitrorhinus Kaup, 1871.
Paxillinae Kuwert, 1891: 182 [stem: Paxill-]. Type genus: Paxillus W. S. MacLeay, 1819.
Petrejinae Kuwert, 1891: 176 [stem: Petrej-]. Type genus: Petrejus Kaup, 1869.
Nelidinae Kuwert, 1896: 222 [stem: Neleid-]. Type genus: Neleides Kaup, 1869. Comment: incorrect original stem formation, not in prevailing usage.
Ptichopinae Kuwert, 1896: 224 [stem: Ptichopod-]. Type genus: Ptichopus Kaup, 1869. Comment: incorrect original stem formation, not in prevailing usage.
Rhodacanthopinae Kuwert, 1896: 222 [stem: Rhodocanthopod-]. Type genus: Rhodocanthopus Kaup, 1871. Comment: incorrect original stem formation, not in prevailing usage.
Vatiniinae Kuwert, 1896: 226 [stem: Vatini-]. Type genus: Vatinius Kaup, 1869 [preoccupied genus name, not Vatinius Stål, 1865 [Hemiptera]; syn. of Passalus Fabricius, 1792]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Proculini
Kaup, 1868
Proculinae Kaup, 1868b: 8 [stem: Procul-]. Type genus: Proculus Kaup, 1868.
Pseudacantheae Kaup, 1871: 73 [stem: Pseudacanth-]. Type genus: Pseudacanthus Kaup, 1869.
Oileinae Kuwert, 1891: 192 [stem: Oile-]. Type genus: Oileus Kaup, 1869.
Sertorinae Kuwert, 1891: 175 [stem: Sertori-]. Type genus: Sertorius Kaup, 1871 [preoccupied genus name, not Sertorius Stål, 1866 [Hemiptera]; syn. of Arrox Zang, 1905]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Unduliferinae Kuwert, 1891: 176 [stem: Undulifer-]. Type genus: Undulifer Kaup, 1869.
Veturinae Kuwert, 1891: 173 [stem: Veturi-]. Type genus: Veturius Kaup, 1871. Comment: incorrect original stem formation, not in prevailing usage.
Popiliinae Kuwert, 1896: 221 [stem: Popili-]. Type genus: Popilius Kaup, 1871.
Proculejinae Kuwert, 1896: 221 [stem: Proculej-]. Type genus: Proculejus Kaup, 1868.
Spuriinae Kuwert, 1896: 221 [stem: Spuri-]. Type genus: Spurius Kaup, 1871.
Vindicinae Kuwert, 1896: 227 [stem: Vindic-]. Type genus: Vindex Kaup, 1871.
Tribe
Solenocyclini
Kaup, 1871
Solenocycleae Kaup, 1871: 24 [stem: Solenocycl-]. Type genus: Solenocyclus Kaup, 1868. Comment: First Reviser found (Solenocyclini Kaup, 1871 vs Stephanocephalini Kaup, 1871) is Boucher (2006).
Stephanocephaleae Kaup, 1871: 78 [stem: Stephanocephal-]. Type genus: Stephanocephalus Kaup, 1868.
Ciceroninae Kuwert, 1891: 183 [stem: Ciceroni-]. Type genus: Ciceronius Kaup, 1871. Comment: incorrect original stem formation, not in prevailing usage.
Erionominae Kuwert, 1891: 176 [stem: Erionom-]. Type genus: Erionomus Kaup, 1868.
Flamininae Kuwert, 1891: 185 [stem: Flamini-]. Type genus: Flaminius Kuwert, 1891. Comment: incorrect original stem formation, not in prevailing usage.
Semicyclinae Kuwert, 1891: 177 [stem: Semicycl-]. Type genus: Semicyclus Kaup, 1871.
Family
Trogidae
MacLeay, 1819
Trogidae W. S. MacLeay, 1819: 59 [stem: Trog-]. Type genus: Trox Fabricius, 1775.
†Subfamily
Avitortorinae
Nikolajev, 2007
Avitortorinae Nikolajev, 2007a: 110 [stem: Avitortor-]. Type genus: Avitortor Ponomarenko, 1977.
Subfamily
Troginae
MacLeay, 1819
Trogidae W. S. MacLeay, 1819: 59 [stem: Trog-]. Type genus: Trox Fabricius, 1775.
Phoberidae Gistel, 1848: [5] [stem: Phober-]. Type genus: Phoberus W. S. MacLeay, 1819.
Subfamily
Omorginae
Nikolajev, 2005
Omorgini Nikolajev, 2005a: 322 [stem: Omorg-]. Type genus: Omorgus Erichson, 1847.
Family
Glaresidae
Kolbe, 1905
Glaresini Kolbe, 1905: 543 [stem: Glares-]. Type genus: Glaresis Erichson, 1848.
Family
Diphyllostomatidae
Holloway, 1972
Diphyllostomatidae Holloway, 1972: 38 [stem: Diphyllostomat-]. Type genus: Diphyllostoma Fall, 1901.
Family
Lucanidae
Latreille, 1804
Lucanides Latreille, 1804c: 149 [stem: Lucan-]. Type genus: Lucanus Scopoli, 1763.
†Subfamily
Protolucaninae
Nikolajev, 2007
Protolucaninae Nikolajev, 2007a: 18 [stem: Protolucan-]. Type genus: Protolucanus Nikolajev, 2007.
Subfamily
Aesalinae
MacLeay, 1819
Aesalidae W. S. MacLeay, 1819: 102 [stem: Aesal-]. Type genus: Aesalus Fabricius, 1801.
Tribe
Aesalini
MacLeay, 1819
Aesalidae W. S. MacLeay, 1819: 102 [stem: Aesal-]. Type genus: Aesalus Fabricius, 1801.
Tribe
Ceratognathini
Sharp, 1899
Ceratognathini Sharp, 1899a: 194 [stem: Ceratognath-]. Type genus: Ceratognathus Westwood, 1838.
Tribe
Nicagini
LeConte, 1861
Nicagini J. L. LeConte, 1861: 130 [stem: Nicag-]. Type genus: Nicagus J. L. LeConte, 1861.
†Subfamily
Ceruchitinae
Nikolajev, 2006
Ceruchitinae Nikolajev, 2006: 133 [stem: Ceruchit-]. Type genus: Ceruchites Statz, 1952.
Subfamily
Syndesinae
MacLeay, 1819
Syndesidae W. S. MacLeay, 1819: 103 [stem: Syndes-]. Type genus: Syndesus W. S. MacLeay, 1819.
Sinodendriens Mulsant, 1842: 600 [stem: Sinodendr-]. Type genus: Sinodendron Hellwig, 1791. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Imhoff (1856: xii, as Sinodendridae), generally accepted as in Ratcliffe (2002: 8, as Sinodendrini).
Ceruchites Jacquelin du Val, 1859: 4 [stem: Ceruch-]. Type genus: Ceruchus W. S. MacLeay, 1819. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 121, as Ceruchini), generally accepted as in Ratcliffe (2002: 8, as Ceruchini).
Subfamily
Lampriminae
MacLeay, 1819
Lamprimidae W. S. MacLeay, 1819: 97 [stem: Lamprim-]. Type genus: Lamprima Latreille, 1804.
Tribe
Lamprimini
MacLeay, 1819
Lamprimidae W. S. MacLeay, 1819: 97 [stem: Lamprim-]. Type genus: Lamprima Latreille, 1804.
Tribe
Streptocerini
Kikuta, 1986
Streptocerini Kikuta, 1986: 131 [stem: Streptocer-]. Type genus: Streptocerus Fairmaire, 1850.
Subfamily
Lucaninae
Latreille, 1804
Lucanides Latreille, 1804c: 149 [stem: Lucan-]. Type genus: Lucanus Scopoli, 1763.
Tribe
Chiasognathini
Burmeister, 1847
Chiasognathidae H. C. C. Burmeister, 1847: 334 [stem: Chiasognath-]. Type genus: Chiasognathus Stephens, 1831.
Tribe
Lucanini
Latreille, 1804
Lucanides Latreille, 1804c: 149 [stem: Lucan-]. Type genus: Lucanus Scopoli, 1763. Comment: published 7 March 1804; this family-group name was also used in the same year by Latreille (1804b [between 19 August and 17 September]: 234, as Lucanides).
Figulidae H. C. C. Burmeister, 1847: 428 [stem: Figul-]. Type genus: Figulus W. S. MacLeay, 1819.
Corypticidae Gistel, 1848: [5] [stem: Coryptic-]. Type genus: Corypticus Sturm, 1843 [the genus was spelled Coryptius (Sturm 1843: 136) and Corypticus (Sturm 1843: 347) in the paper making the genus name available but the latter spelling is in prevailing usage and is the correct spelling].
Dorcidae Parry, 1864: 86 [stem: Dorc-]. Type genus: Dorcus W. S. MacLeay, 1819.
Cladognathidae Parry, 1870: 75 [stem: Cladognath-]. Type genus: Cladognathus H. C. C. Burmeister, 1847 [syn. of Prosopocoilus Hope, 1845].
Odontolabidae Parry, 1870: 106 [stem: Odontolabid-]. Type genus: Odontolabis Hope, 1842. Comment: incorrect original stem formation, not in prevailing usage.
Nigidiini Jakobson, 1911b: 142 [stem: Nigidi-]. Type genus: Nigidius W. S. MacLeay, 1819. Comment: Nigidiini was attributed to Benesh (1960) and treated as unavailable by A. B. T. Smith (2006: 154).
Leptinopterini Jakobson, 1911b: 142 [stem: Leptinopter-]. Type genus: Leptinopterus Hope, 1838 [unjustified emendation of Leptynopterus by Hope (1845: 5), in prevailing usage and so deemed to be a justified emendation (Article 33.2.3.1)].
Rhaetulinae Miwa, 1931: 323 [stem: Rhaetul-]. Type genus: Rhaetulus Westwood, 1871.
Penichrolucaninae Arrow, 1950: 233 [stem: Penichrolucan-]. Type genus: Penichrolucanus Deyrolle, 1863.
*Chalcodinae Didier and Séguy, 1953: 91 [stem: Chalcod-]. Type genus: Chalcodes H. C. C. Burmeister, 1847. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Dendeziini Benesh, 1955b: 72 [stem: Dendezi-]. Type genus: Dendezia Basilewsky, 1952.
Lissotini Benesh, 1955b: 73 [stem: Lissotet-]. Type genus: Lissotes Westwood, 1855. Comment: incorrect original stem formation, not in prevailing usage.
Sclerostomini Benesh, 1955a: 97 [stem: Sclerostom-]. Type genus: Sclerostomus H. C. C. Burmeister, 1847.
Scortizini Benesh, 1955a: 103 [stem: Scortiz-]. Type genus: Scortizus Westwood, 1834.
*Prosopocoilini Benesh, 1960: 50 [stem: Prosopocoil-]. Type genus: Prosopocoilus Hope, 1845. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Pseudodorcini Benesh, 1960: 97 [stem: Pseudodorc-]. Type genus: Pseudodorcus Parry, 1870. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Rhyssonotini Benesh, 1960: 148 [stem: Ryssonot-]. Type genus: Ryssonotus W. S. MacLeay, 1819 [as Rhyssonotus, unjustified emendation of type genus name by Agassiz (1846b: 329), not in prevailing usage]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
*Chalcodinae J. P. Lacroix, 1979: 258 [stem: Chalcod-]. Type genus: Chalcodes H. C. C. Burmeister, 1847. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Pholidotini Kikuta, 1986: 131 [stem: Pholidot-]. Type genus: Pholidotus W. S. MacLeay, 1819 [preoccupied genus name, not Pholidotus Brisson, 1762 [Mammalia]; syn. of Casignetus W. S. MacLeay, 1819]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
*Aegini Maes, 1992b: 97 [stem: Aeg-]. Type genus: Aegus W. S. MacLeay, 1819. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); Aegidae A. White, 1850 (type genus Aega Leach, 1815) is currently used as valid in Isopoda and therefore anyone wishing to base a new family-group name on the genus Aegus W. S. MacLeay should change the stem in order to avoid homonymy with the isopod name.
*Allotopini Maes, 1992a: 56 [stem: Allotop-]. Type genus: Allotopus Albers, 1894. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Casignetini Maes, 1992b: 61 [stem: Casignet-]. Type genus: Casignetus W. S. MacLeay, 1819. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); Casignetini Falkovitsh, 1972 (type genus Casigneta Wallengren, 1881) proposed in Lepidoptera is permanently invalid since it is based on a preoccupied type genus name.
*Colophonini Maes, 1992b: 107 [stem: Colophon-]. Type genus: Colophon Gray, 1832. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Cyclommatini Maes, 1992a: 56 [stem: Cyclommat-]. Type genus: Cyclommatus Parry, 1863. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Homoderini Maes, 1992b: 85 [stem: Homoder-]. Type genus: Homoderus Parry, 1863. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Casignetini Reid, 1999: 175 [stem: Casignet-]. Type genus: Casignetus W. S. MacLeay, 1819. Comment: replacement name for Pholidotini Kikuta, 1986 because of the homonymy of the type genus.
Brasilucanini Nikolajev, 1999a: 171 [stem: Brasilucan-]. Type genus: Brasilucanus Vulcano and Pereira, 1961.
Tribe
Platycerini
Mulsant, 1842
Platycéraires Mulsant, 1842: 593 [stem: Platycer-]. Type genus: Platycerus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1856a: 365, as Platyceridae), generally accepted as in Paulsen and Hawks (2008: 1, as Platycerini).
Systenocerini Portevin, 1931: 2, in key [stem: Systenocer-]. Type genus: Systenocerus Weise, 1883 [syn. of Platycerus Geoffroy, 1762].
Tribe
Platyceroidini
Paulsen and Hawks, 2008
Platyceroidini Paulsen and Hawks, 2008: 2 [stem: Platyceroid-]. Type genus: Platyceroides Benesh, 1946.
†Subfamily
Paralucaninae
Nikolajev, 2000
Paralucaninae Nikolajev, 2000b: S328 [stem: Paralucan-]. Type genus: Paralucanus Nikolajev, 2000.
Family
Ochodaeidae
Mulsant and Rey, 1871
Ochodéens Mulsant and Rey, 1871b: 493 [stem: Ochodae-]. Type genus: Ochodaeus Dejean, 1821.
†Subfamily
Cretochodaeinae
Nikolajev, 1995
Cretochodaeini Nikolajev, 1995a: 78 [stem: Cretochodae-]. Type genus: Cretochodaeus Nikolajev, 1995.
Subfamily
Ochodaeinae
Mulsant and Rey, 1871
Ochodéens Mulsant and Rey, 1871b: 493 [stem: Ochodae-]. Type genus: Ochodaeus Dejean, 1821.
Tribe
Enodognathini
Scholtz, 1988
Endognathini Scholtz, 1988: 228 [stem: Enodognath-]. Type genus: Enodognathus Benderitter, 1920 [as Endognathus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Scholtz (1991: 30).
Tribe
Ochodaeini
Mulsant and Rey, 1871
Ochodéens Mulsant and Rey, 1871b: 493 [stem: Ochodae-]. Type genus: Ochodaeus Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Arrow (1904: 747, as Ochodaeinae), generally accepted as in A. B. T. Smith (2006: 155, as Ochodaeidae); incorrect original stem formation, not in prevailing usage.
Subfamily
Chaetocanthinae
Scholtz, 1988
Chaetocanthinae Scholtz, 1988: 231 [stem: Chaetocanth-]. Type genus: Chaetocanthus Péringuey, 1901. Comment: precedence (Chaetocanthinae Scholtz, 1988 vs Pseudochodaeinae Scholtz, 1988 vs Synochodaeinae Scholtz, 1988) given to taxon originally proposed at the higher rank (Art. 24.1).
Tribe
Chaetocanthini
Scholtz, 1988
Chaetocanthinae Scholtz, 1988: 231 [stem: Chaetocanth-]. Type genus: Chaetocanthus Péringuey, 1901.
Tribe
Pseudochodaeini
Scholtz, 1988
Pseudochodaeini Scholtz, 1988: 235 [stem: Pseudochodae-]. Type genus: Pseudochodaeus Carlson and Ritcher, 1974.
Tribe
Synochodaeini
Scholtz, 1988
Synochodaeini Scholtz, 1988: 237 [stem: Synochodae-]. Type genus: Synochodaeus Kolbe, 1907.
Family
Hybosoridae
Erichson, 1847
Hybosoridae Erichson, 1847a: 104 [stem: Hybosor-]. Type genus: Hybosorus W. S. MacLeay, 1819.
†Subfamily
Mimaphodiinae
Nikolajev, 2007
Mimaphodiinae Nikolajev, 2007b: 47 [stem: Mimaphodi-]. Type genus: Mimaphodius Nikolajev, 2007.
Subfamily
Anaidinae
Nikolajev, 1996
Anaidini Nikolajev, 1996: 94 [stem: Anaid-]. Type genus: Anaides Westwood, 1842.
Cryptogeniini Howden, 2001: 199 [stem: Cryptogeni-]. Type genus: Cryptogenius Westwood, 1842.
Subfamily
Ceratocanthinae
Martínez, 1968
Ceratocanthini Martínez, 1968: 14 [stem: Ceratocanth-]. Type genus: Ceratocanthus A. White, 1842.
Tribe
Ceratocanthini
Martínez, 1968
Acanthocérides Lacordaire, 1856: 155 [stem: Acanthocer-]. Type genus: Acanthocerus W. S. MacLeay, 1819 [preoccupied genus name, not Acanthocerus Palisot de Beauvois, 1818 [Hemiptera]; syn. of Ceratocanthus A. White, 1842]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte (1861: 129, as Acanthocerini); permanently invalid (Art. 39): based on preoccupied type genus.
Ceratocanthini Martínez, 1968: 14 [stem: Ceratocanth-]. Type genus: Ceratocanthus A. White, 1842. Comment: replacement name for Acanthocerini Lacordaire, 1856 because of the homonymy of the type genus.
Tribe
Ivieolini
Howden and Gill, 2000
Ivieolini Howden and Gill, 2000: 315 [stem: Ivieol-]. Type genus: Ivieolus Howden and Gill, 1988.
Tribe
Scarabatermitini
Nikolajev, 1999
Scarabatermitini Nikolajev, 1999b: 175 [stem: Scarabatermit-]. Type genus: Scarabatermes Howden, 1973.
Subfamily
Hybosorinae
Erichson, 1847
Hybosoridae Erichson, 1847a: 104 [stem: Hybosor-]. Type genus: Hybosorus W. S. MacLeay, 1819.
Subfamily
Liparochrinae
Ocampo, 2006
Liparochrinae Ocampo, 2006: 29 [stem: Liparochr-]. Type genus: Liparochrus Erichson, 1848.
Subfamily
Pachyplectrinae
Ocampo, 2006
Pachyplectrinae Ocampo, 2006: 30 [stem: Pachyplectr-]. Type genus: Pachyplectrus J. L. LeConte, 1874.
Family
Glaphyridae
MacLeay, 1819
Glaphyridae W. S. MacLeay, 1819: 76 [stem: Glaphyr-]. Type genus: Glaphyrus Latreille, 1802.
Subfamily
Glaphyrinae
MacLeay, 1819
Glaphyridae W. S. MacLeay, 1819: 76 [stem: Glaphyr-]. Type genus: Glaphyrus Latreille, 1802.
Subfamily
Amphicominae
Blanchard, 1845
Amphicomites Blanchard, 1845a: 211 [stem: Amphicom-]. Type genus: Amphicoma Latreille, 1807. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Blanchard (1850: 52, as Amphicomitae); name treated as unavailable by A. B. T. Smith (2006: 157).
†Subfamily
Cretoglaphyrinae
Nikolajev, 2005
Cretoglaphyrini Nikolajev, 2005b: 70 [stem: Cretoglaphyr-]. Type genus: Cretoglaphyrus Nikolajev, 2005.
Family
Scarabaeidae
Latreille, 1802
Scarabaeïdes Latreille, 1802: 144 [stem: Scarabae-]. Type genus: Scarabaeus Linnaeus, 1758.
†Subfamily
Lithoscarabaeinae
Nikolajev, 1992
Lithoscarabaeinae Nikolajev, 1992: 76 [stem: Lithoscarabae-]. Type genus: Lithoscarabaeus Nikolajev, 1992.
Subfamily
Chironinae
Blanchard, 1845
Chironites Blanchard, 1845a: 225 [stem: Chiron-]. Type genus: Chiron W. S. MacLeay, 1819. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Harold (1868: 278, as Chironidae), generally accepted as in A. B. T. Smith (2006: 157, as Chironinae).
Subfamily
Aegialiinae
Laporte, 1840
Aegialites Laporte, 1840b: 99 [stem: Aegiali-]. Type genus: Aegialia Latreille, 1807. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Harold (1868: 278, as Aegialidae [incorrect stem formation]), generally accepted as in A. B. T. Smith (2006: 157, as Aegialiinae); incorrect original stem formation, not in prevailing usage.
Silluviinae Landin, 1950: 3 [stem: Silluvi-]. Type genus: Silluvia Landin, 1950.
Saprini Nikolajev, 2008: 149 [stem: Sapr-]. Type genus: Saprus Blackburn, 1904 [preoccupied genus name, not Saprus Gistel, 1856 [Coleoptera: Sphaeriusidae]; syn. of Sapriniana Strand, 1917]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Subfamily
Eremazinae
Iablokoff-Khnzorian, 1977
Eremazini Iablokoff-Khnzorian, 1977 [3 October]: 168 [stem: Eremaz-]. Type genus: Eremazus Mulsant, 1851.
Eremazina Stebnicka, 1977 [“31 December”]: 412 [stem: Eremaz-]. Type genus: Eremazus Mulsant, 1851. Comment: proposed as new, without reference to Eremazini Iablokoff-Khnzorian, 1977.
Subfamily
Aphodiinae
Leach, 1815
Aphodida Leach, 1815: 97 [stem: Aphodi-]. Type genus: Aphodius Illiger, 1798.
Tribe
Aphodiini
Leach, 1815
Aphodida Leach, 1815: 97 [stem: Aphodi-]. Type genus: Aphodius Illiger, 1798.
Subtribe
Aphodiina
Leach, 1815
Aphodida Leach, 1815: 97 [stem: Aphodi-]. Type genus: Aphodius Illiger, 1798.
*Ammoeciates Mulsant, 1842: 301 [stem: Ammoeci-]. Type genus: Ammoecius Mulsant, 1842. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Plagiogonates Mulsant and Rey, 1871a: 609 [stem: Plagiogon-]. Type genus: Plagiogonus Mulsant, 1842. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Heptaulacates Mulsant and Rey, 1871a: 585 [stem: Heptaulac-]. Type genus: Heptaulacus Mulsant, 1842. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Subtribe
Didactyliina
Pittino, 1985
Didactyliina Pittino, 1985: 270, in key [stem: Didactyli-]. Type genus: Didactylia d’Orbigny, 1896.
Subtribe
Proctophanina
Stebnicka and Howden, 1995
Proctophanini Stebnicka and Howden, 1995: 742 [stem: Proctophan-]. Type genus: Proctophanes Harold, 1861.
Tribe
Corythoderini
Schmidt, 1910
Corythoderina A. Schmidt, 1910a: 137 [stem: Corythoder-]. Type genus: Corythoderus Klug, 1845. Comment: the earliest known publication date of this work is 31 December 1910 (see Evenhuis, 1994: 55); although Corythoderina A. Schmidt (1910b: 93) was published on 30 September 1910, we use A. Schmidt, 1910a as the correct original work since “Corythoderina Schmidt 1910: 137” was used as the correct citation for this name by A. Schmidt himself (1910b: 93).
Tribe
Eupariini
Schmidt, 1910
nomen protectum
Ataenidae Harold, 1868: 278 [stem: Ataeni-]. Type genus: Ataenius Harold, 1867 [placed on the Official List of Generic Names in Zoology (ICZN 2010a)]. Comment: nomen oblitum (see A. B. T. Smith 2006: 158).
*Hexalates Mulsant and Rey, 1871a: 605 [stem: Hexal-]. Type genus: Hexalus Mulsant and Rey, 1871 [syn. of Ataenius Harold, 1867]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Eupariina A. Schmidt, 1910a: 102 [stem: Eupari-]. Type genus: Euparia Lepeletier and Audinet-Serville, 1828. Comment: nomen protectum (see A. B. T. Smith 2006: 158); senior homonym of Eupariini B. D. Valentine, 1960 (type genus Euparius Schönherr, 1823) in Anthribidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1); the earliest known publication date of this work is 31 December 1910 (see Evenhuis, 1994: 55); although Eupariina A. Schmidt (1910b: 71) was published on 30 September 1910, we use A. Schmidt, 1910a as the correct original work since “Eupariina Schmidt 1910: 102” was used as the correct citation for this name by A. Schmidt himself (1910b: 71).
Lomanoxiini Stebnicka, 1999: 280 [stem: Lomanoxi-]. Type genus: Lomanoxia Martínez, 1951.
Tribe
Odontolochini
Stebnicka and Howden, 1996
Odontolochini Stebnicka and Howden, 1996: 99 [stem: Odontoloch-]. Type genus: Odontolochus Schmidt, 1916.
Tribe
Odochilini
Rakovič, 1987
Odochilini Rakovič, 1987: 29 [stem: Odochil-]. Type genus: Odochilus Harold, 1877.
Tribe
Psammodiini
Mulsant, 1842
Psammodiaires Mulsant, 1842: 317 [stem: Psammodi-]. Type genus: Psammodius Fallén, 1807.
Subtribe
Phycocina
Landin, 1960
Phycochi Landin, 1960: 59 [stem: Phycoc-]. Type genus: Phycocus Broun, 1886 [as Phycochus, incorrect subsequent spelling of type genus name, not in prevailing usage (see Stebnicka 2001)]. Comment: incorrect original stem formation, not in prevailing usage.
Phycochini Rakovič and Král, 1997: 59 [stem: Phycoc-]. Type genus: Phycocus Broun, 1886 [as Phycochus, incorrect subsequent spelling of type genus name, not in prevailing usage (see Stebnicka 2001)]. Comment: incorrect original stem formation, not in prevailing usage; family-group name proposed as new without reference to Phycochi Landin, 1960.
Subtribe
Psammodiina
Mulsant, 1842
Psammodiaires Mulsant, 1842: 317 [stem: Psammodi-]. Type genus: Psammodius Fallén, 1807. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Harold (1868: 278, as Psammodidae [incorrect stem formation]), generally accepted as in A. B. T. Smith (2006: 159, as Psammodiini).
Pleurophorates Mulsant, 1842: 304 [stem: Pleurophor-]. Type genus: Pleurophorus Mulsant, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Ádám (1994: 15, as Pleurophorini).
Psammobiina Reitter, 1909: 303 [stem: Psammobi-]. Type genus: Psammobius Heer, 1841 [syn. of Psammodius Fallén, 1807].
Subtribe
Rhyssemina
Pittino and Mariani, 1986
Rhyssemina Pittino and Mariani, 1986: 17, in key [stem: Rhyssem-]. Type genus: Rhyssemus Mulsant, 1842.
Tribe
Rhyparini
Schmidt, 1910
Rhyparina A. Schmidt, 1910a: 130 [stem: Rhypar-]. Type genus: Rhyparus Westwood, 1845 [Rhyparus is an unjustified emendation of Ryparus Westwood, 1845 by Agassiz (1846b: 328), in prevailing usage, and so deemed to be a justified emendation (Article 33.2.3.1); the emended spelling avoids homonymy with Ryparus Spinola, 1844 [Coleoptera: Cleridae] (see A. B. T. Smith 2006: 159)]. Comment: the earliest known publication date of this work is 31 December 1910 (see Evenhuis, 1994: 55); although Rhyparina A. Schmidt (1910b: 91) was published on 30 September 1910, we use A. Schmidt, 1910a as the correct original work since “Rhyparina Schmidt 1910: 130” was used as the correct citation for this name by A. Schmidt himself (1910b: 91).
Tribe
Stereomerini
Howden and Storey, 1992
Stereomerini Howden and Storey, 1992: 1811 [stem: Stereomer-]. Type genus: Stereomera Arrow, 1905.
Tribe
Termitoderini
Tangelder and Krikken, 1982
Termitoderini Tangelder and Krikken, 1982: 10, in key [stem: Termitoder-]. Type genus: Termitoderus Mateu, 1966.
Subfamily
Aulonocneminae
Janssens, 1946
Aulonocneminae Janssens, 1946: 7, in key [stem: Aulonocnem-]. Type genus: Aulonocnemis Klug, 1837. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Aulonocnemid-).
Subfamily
Termitotroginae
Wasmann, 1918
Termitotrogini Wasmann, 1918: 4 [stem: Termitotrog-]. Type genus: Termitotrox Reichensperger, 1915.
Subfamily
Scarabaeinae
Latreille, 1802
Scarabaeïdes Latreille, 1802: 144 [stem: Scarabae-]. Type genus: Scarabaeus Linnaeus, 1758.
Tribe
Ateuchini
Perty, 1830
Ateuchidae Perty, 1830: 38 [stem: Ateuch-]. Type genus: Ateuchus Weber, 1801.
Subtribe
Ateuchina
Perty, 1830
Ateuchidae Perty, 1830: 38 [stem: Ateuch-]. Type genus: Ateuchus Weber, 1801. Comment: name previously attributed to Laporte (1840b: 63); use of subtribes follow Vaz-de-Mello (2008).
Choerididae Harold, 1867: 9 [stem: Choeridi-]. Type genus: Choeridium Lepeletier and Audinet-Serville, 1828.
Pinotinae Kolbe, 1905: 548 [stem: Pinot-]. Type genus: Pinotus Erichson, 1847.
Dichotomiini Pereira, 1954: 55 [stem: Dichotomi-]. Type genus: Dichotomius Hope, 1838.
Subtribe
Scatimina
Vaz-de-Mello, 2008
Scatimina Vaz-de-Mello, 2008: 10 [stem: Scatim-]. Type genus: Scatimus Erichson, 1847.
Ateuchini
incertae sedis
Demarziellini Balthasar, 1961: 178 [stem: Demarziell-]. Type genus: Demarziella Balthasar, 1961. Comment: family-group taxon not originally described but available (Art. 13.5) (see Matthews and Stebnicka 1986); current placement follows Vaz-de-Mello (2008).
Tribe
Coprini
Leach, 1815
*Coprides Baudet-Lafarge, 1809: 44 [stem: Copr-]. Type genus: Copris Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Baudet-Lafarge (1809).
Coprides Leach, 1815: 96 [stem: Copr-]. Type genus: Copris Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Coprid-).
Coptodactylini Paulian, 1933: 67 [stem: Coptodactyl-]. Type genus: Coptodactyla H. C. C. Burmeister, 1846.
Tribe
Deltochilini
Lacordaire, 1856
Deltochilides Lacordaire, 1856: 78 [stem: Deltochil-]. Type genus: Deltochilum Eschscholtz, 1822. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 125, as Deltochila), generally accepted as in Germain (1903: 354, as Deltochilidae); First Reviser (Deltochilini Lacordaire, 1856 vs Mentophilini Lacordaire, 1856 vs Scatonomini Lacordaire, 1856) not determined, current usage maintained.
Minthophilides Lacordaire, 1856: 80 [stem: Mentophil-]. Type genus: Mentophilus Laporte, 1840 [as Minthophilus, both Mentophilus (p. 74) and Minthophilus (p. 63) were used in the original publication by Laporte (1840b), although Mentophilus has been used as the correct spelling in recent literature, we could not determine the First Reviser]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Gerstaecker (1861: 464, as Minthophilidae); name treated as unavailable by A. B. T. Smith (2006: 160); incorrect original stem formation, not in prevailing usage.
Scatonomides Lacordaire, 1856: 87 [stem: Scatonom-]. Type genus: Scatonomus Erichson, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte (1861: 125, as Scatonomi); transferred from Ateuchini by Vaz-de-Mello (2008).
Coprobiadae H. C. C. Burmeister, 1873: 407 [stem: Coprobi-]. Type genus: Coprobius Latreille, 1829.
Canthonides van Lansberge, 1875a: 184 [stem: Canthon-]. Type genus: Canthon Hoffmannsegg, 1817. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1905: 530, as Canthoninae), generally accepted as in A. B. T. Smith (2006: 160, as Canthonini).
Epilissides van Lansberge, 1875a: 188 [stem: Epiliss-]. Type genus: Epilissus Dejean, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Shipp (1894: 255, as Epilissini).
Epirinides van Lansberge, 1875a: 189 [stem: Epirin-]. Type genus: Epirinus Dejean, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Bertkau (1875: 335, as Epirini); name treated as unavailable by A. B. T. Smith (2006: 160).
Panelini Arrow, 1931: 404 [stem: Panel-]. Type genus: Panelus Lewis, 1895.
Tribe
Eucraniini
Burmeister, 1873
Eucraniadae H. C. C. Burmeister, 1873: 405 [stem: Eucrani-]. Type genus: Eucranium Brullé, 1837.
Ennearabdini Pereira and Martínez, 1956: 238 [stem: Ennearabd-]. Type genus: Ennearabdus van Lansberge, 1874.
Tribe
Gymnopleurini
Lacordaire, 1856
Gymnopleurides Lacordaire, 1856: 72 [stem: Gymnopleur-]. Type genus: Gymnopleurus Illiger, 1803. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 124, as Gymnopleuri), generally accepted as in A. B. T. Smith (2006: 160, as Gymnopleurini).
Tribe
Oniticellini
Kolbe, 1905
Oniticellini Kolbe, 1905: 547 [stem: Oniticell-]. Type genus: Oniticellus Dejean, 1821. Comment: usage of this name over Drepanocerini conserved (Art. 35.5) (see A. B. T. Smith 2006).
Subtribe
Drepanocerina
van Lansberge, 1875
Drèpanocérides van Lansberge, 1875b: 14 [stem: Drepanocer-]. Type genus: Drepanocerus Kirby, 1828. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1905: 531, as Drepanocerini), generally accepted as in A. B. T. Smith (2006: 161, as Drepanocerini); although Drepanocerini has priority over Oniticellini, the latter is in prevailing usage at the tribal level and must not be displaced by the older name (Art. 35.5) (see A. B. T. Smith 2006).
Subtribe
Eurysternina
Vulcano, Martínez and Pereira, 1961
Eurysternini Vulcano et al., 1961: 268 [stem: Eurystern-]. Type genus: Eurysternus Dalman, 1824. Comment: current placement based on Génier (2009).
Subtribe
Helictopleurina
Janssens, 1946
Helictopleurides Janssens, 1946: 11, in key [stem: Helictopleur-]. Type genus: Helictopleurus d’Orbigny, 1915.
Subtribe
Oniticellina
Kolbe, 1905
Oniticellini Kolbe, 1905: 547 [stem: Oniticell-]. Type genus: Oniticellus Dejean, 1821.
Tribe
Onitini
Laporte, 1840
Onitides Laporte, 1840b: 88 [stem: Onit-]. Type genus: Onitis Fabricius, 1798. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Shipp (1894: 255, as Onitidae), generally accepted as in A. B. T. Smith (2006: 161, as Onitini); although Fabricius originally treated the type genus name as masculine, it appears in Greek dictionaries as a feminine noun (with stem Onitid-); we recommend that an application be submitted to the Commission to establish the correct gender of Onitis and the correct stem based on that genus.
Tribe
Onthophagini
Burmeister, 1846
Onthophagidae H. C. C. Burmeister, 1846: [1] [stem: Onthophag-]. Type genus: Onthophagus Latreille, 1802.
Alloscelides Janssens, 1946: 10, in key [stem: Alloscel-]. Type genus: Alloscelus Boucomont, 1923.
Tribe
Phanaeini
Hope, 1838
Phanaeidae Hope, 1838a: 321 [stem: Phanae-]. Type genus: Phanaeus W. S. MacLeay, 1819.
Gromphina Zunino, 1985a: 22 [stem: Gromphad-]. Type genus: Gromphas Brullé, 1837. Comment: published 20 November 1985; this family-group name was also used in the same year by Zunino (1985b [20 December]: 107, as Gromphina); incorrect original stem formation, not in prevailing usage.
Tribe
Scarabaeini
Latreille, 1802
Scarabaeïdes Latreille, 1802: 144 [stem: Scarabae-]. Type genus: Scarabaeus Linnaeus, 1758.
Pachysomides Ferreira, 1953: 5, in key [stem: Pachysomat-]. Type genus: Pachysoma W. S. MacLeay, 1821. Comment: incorrect original stem formation, not in prevailing usage.
Actinophorini Ádám, 2003: 130 [stem: Actinophor-]. Type genus: Actinophorus Creutzer, 1799 [syn. of Scarabaeus Linnaeus, 1758].
Tribe
Sisyphini
Mulsant, 1842
Sisyphaires Mulsant, 1842: 41 [stem: Sisyph-]. Type genus: Sisyphus Latreille, 1807 [the original spelling of the type genus is Sisyphe, however, the incorrect subsequent spelling Sisyphus is in prevailing usage and should be considered the correct original spelling (Art. 33.3.1) (see A. B. T. Smith 2006)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Shipp (1894: 255, as Sisyphinae), generally accepted as in A. B. T. Smith (2006: 162, as Sisyphini).
†Subfamily
Prototroginae
Nikolajev, 2000
Prototroginae Nikolajev, 2000a: 63 [stem: Prototrog-]. Type genus: Prototrox Nikolajev, 2000.
†Subfamily
Cretoscarabaeinae
Nikolajev, 1995
Cretoscarabaeinae Nikolajev, 1995b: 147 [stem: Cretoscarabae-]. Type genus: Cretoscarabaeus Nikolajev, 1995.
Subfamily
Dynamopodinae
Arrow, 1911
Dynamopinae Arrow, 1911: 611 [stem: Dynamopod-]. Type genus: Dynamopus Semenov, 1895.
Tribe
Dynamopodini
Arrow, 1911
Dynamopinae Arrow, 1911: 611 [stem: Dynamopod-]. Type genus: Dynamopus Semenov, 1895. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Lawrence and Newton (1995).
Tribe
Thinorycterini
Semenov and A. N. Reichardt, 1925
Thinorycterina Semenov and A. N. Reichardt, 1925: 86 [stem: Thinorycter-]. Type genus: Thinorycter Semenov and A. N. Reichardt, 1925.
Subfamily
Phaenomeridinae
Erichson, 1847
Phaenomerini Erichson, 1847b: 655 [stem: Phaenomerid-]. Type genus: Phaenomeris Hope, 1833 [placed on the Official List of Generic Names in Zoology (ICZN 1962)]. Comment: stem emended from Phaenomer- to Phaenomerid- thus removing it from homonymy with another family-group name in Curculionidae and placed on the Official List of Family-Group Names in Zoology (ICZN 1962, as Phaenomerididae Ohaus, 1913); in this ruling (ICZN 1962) ‘‘Ohaus, 1913’’ was erroneously given as the original author of this family-group name (see A. B. T. Smith 2006).
Subfamily
Orphninae
Erichson, 1847
Orphnidae Erichson, 1847a: 111 [stem: Orphn-]. Type genus: Orphnus W. S. MacLeay, 1819.
Tribe
Aegidiini
Paulian, 1984
Aegidiinae Paulian, 1984: 68 [stem: Aegidi-]. Type genus: Aegidium Westwood, 1845.
Tribe
Orphnini
Erichson, 1847
Orphnidae Erichson, 1847a: 111 [stem: Orphn-]. Type genus: Orphnus W. S. MacLeay, 1819.
Hybalidae Marseul, 1857a: 83 [stem: Hybal-]. Type genus: Hybalus Dejean, 1833. Comment: name previously attributed to Jacquelin du Val (1859: 31, as Hybalites) and treated as unavailable by A. B. T. Smith (2006: 163).
Subfamily
Allidiostomatinae
Arrow, 1940
Idiostominae Arrow, 1904: 747 [stem: Idiostomat-]. Type genus: Idiostoma Arrow, 1904 [preoccupied genus name, not Idiostoma Walsingham, 1882 [Lepidoptera]; syn. of Allidiostoma Arrow, 1940]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Allidiostomidae Arrow, 1940: 16 [stem: Allidiostomat-]. Type genus: Allidiostoma Arrow, 1940. Comment: replacement name for Idiostominae Arrow, 1904 because of the homonymy of the type genus; incorrect original stem formation, not in prevailing usage; correction of stem by Lawrence and Newton (1995).
Subfamily
Aclopinae
Blanchard, 1850
Aclopitae Blanchard, 1850: 96 [stem: Aclop-]. Type genus: Aclopus Erichson, 1835.
Tribe
Aclopini
Blanchard, 1850
Aclopitae Blanchard, 1850: 96 [stem: Aclop-]. Type genus: Aclopus Erichson, 1835.
†Tribe
Holcorobeini
Nikolajev, 1992
Holcorobeini Nikolajev, 1992: 81 [stem: Holcorobe-]. Type genus: Holcorobeus Nikritin, 1977.
Tribe
Phaenognathini
Iablokoff-Khnzorian, 1977
Phaenognathini Iablokoff-Khnzorian, 1977: 137, 172 [stem: Phaenognath-]. Type genus: Phaenognatha Hope, 1842.
Subfamily
Melolonthinae
Leach, 1819
Melolonthidae Leach, 1819: 189 [stem: Melolonth-]. Type genus: Melolontha Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Ablaberini
Blanchard, 1850
Ablaberitae Blanchard, 1850: 100 [stem: Ablaber-]. Type genus: Ablabera Dejean, 1833.
Camentini Machatschke, 1959: 738 [stem: Cament-]. Type genus: Camenta Erichson, 1847.
Tribe
Automoliini
Britton, 1978
Caulobiina H. C. C. Burmeister, 1855: 204 [stem: Caulobi-]. Type genus: Caulobius Le Guillou, 1844 [preoccupied genus name, not Caulobius Duponchel, 1838 [Lepidoptera]; syn. of Deuterocaulobius Dalla Torre, 1912]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Automolini Britton, 1957: 72 [stem: Automol-]. Type genus: Automolus H. C. C. Burmeister, 1855 [preoccupied genus name, not Automolus Reichenbach, 1853 [Aves]; syn. of Automolius Britton, 1978]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Automoliini Britton, 1978: 7 [stem: Automoli-]. Type genus: Automolius Britton, 1978. Comment: replacement name for Automolini Britton, 1957 because of the homonymy of the type genus.
Tribe
Chasmatopterini
Lacordaire, 1856
Chasmatoptérides Lacordaire, 1856: 220 [stem: Chasmatopter-]. Type genus: Chasmatopterus Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1887: 130, as Chasmatopteridae), generally accepted as in A. B. T. Smith (2006: 164, as Chasmatopterini).
Tribe
Colymbomorphini
Blanchard, 1850
Colymbomorphitae Blanchard, 1850: 97 [stem: Colymbomorph-]. Type genus: Colymbomorpha Blanchard, 1850.
Stethaspididae H. C. C. Burmeister, 1855: 218 [stem: Stethaspid-]. Type genus: Stethaspis Hope, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1983b)]. Comment: A. B. T. Smith (2006) used “Stethaspini” as the valid name for this tribe, however, the fact that the genus Colymbomorpha is considered to be in this tribe, e.g., Houston and Weir (1992), brings the name Colymbomorphini into synonymy with Stethaspini and Xylonichini; Colymbomorphini has nomenclatural priority and should be considered the valid name for this tribe.
Xylonychini Britton, 1957: 9 [stem: Xylonich-]. Type genus: Xylonichus Boisduval, 1835 [as Xylonychus, incorrect subsequent spelling of type genus name, not in prevailing usage (see A. B. T. Smith 2006: 166)]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Comophorinini
Britton, 1957
Comophorini Britton, 1957: 10 [stem: Comophorin-]. Type genus: Comophorina Strand, 1928. Comment: incorrect original stem formation, not in prevailing usage.
†Tribe
Cretomelolonthini
Nikolajev, 1998
Cretomelolonthini Nikolajev, 1998: 80 [stem: Cretomelolonth-]. Type genus: Cretomelolontha Nikolajev, 1998.
Tribe
Dichelonychini
Burmeister, 1855
Dichelonychidae H. C. C. Burmeister, 1855: 70 [stem: Dichelonych-]. Type genus: Dichelonyx Harris, 1827.
Tribe
Diphucephalini
Laporte, 1840
Diphucéphalites Laporte, 1840b: 145 [stem: Diphucephal-]. Type genus: Diphucephala Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 125, as Diphycocephaloidae [incorrect stem formation]), generally accepted as in A. B. T. Smith (2006: 165, as Diphucephalini).
Tribe
Diphycerini
Medvedev, 1952
Diphycerini S. I. Medvedev, 1952: 186 [stem: Diphycer-]. Type genus: Diphycerus Deyrolle and Fairmaire, 1878.
Tribe
Diplotaxini
Kirby, 1837
Diplotaxidae Kirby, 1837: 129 [stem: Diplotax-]. Type genus: Diplotaxis Kirby, 1837. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Diplotaxe-).
Apogoniitae Blanchard, 1851a: 228 [stem: Apogoni-]. Type genus: Apogonia Kirby, 1819.
Liogenitae Blanchard, 1851a: 166 [stem: Liogeny-]. Type genus: Liogenys Guérin-Méneville, 1831. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Euchirini
Hope, 1840
Eucheiridae Hope, 1840b: 300 [stem: Euchir-]. Type genus: Euchirus Kirby, 1828 [as Eucheirus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Heteronychini
Lacordaire, 1856
Hétéronycides Lacordaire, 1856: 225 [stem: Heteronych-]. Type genus: Heteronyx Guérin-Méneville, 1831. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Bertkau (1890: 277, as Heteronychini), generally accepted as in A. B. T. Smith (2006: 167, as Heteronychini); incorrect original stem formation, not in prevailing usage.
Tribe
Hopliini
Latreille, 1829
Hoplides Latreille, 1829a: 563 [stem: Hopli-]. Type genus: Hoplia Illiger, 1803.
Subtribe
Hopliina
Latreille, 1829
Hoplides Latreille, 1829a: 563 [stem: Hopli-]. Type genus: Hoplia Illiger, 1803. Comment: incorrect original stem formation, not in prevailing usage.
Dicheliden Oken, 1843: 483 [stem: Dichel-]. Type genus: Dichelus Lepeletier and Audinet-Serville, 1828. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Tulk (1847: 614, as Dichelidae).
Gymnolomidae H. C. C. Burmeister, 1844: 138 [stem: Gymnolomat-]. Type genus: Gymnoloma Dejean, 1833. Comment: incorrect original stem formation, not in prevailing usage.
Heterochelidae H. C. C. Burmeister, 1844: 86 [stem: Heterochel-]. Type genus: Heterochelus H. C. C. Burmeister, 1844.
Lepisiidae H. C. C. Burmeister, 1844: 166 [stem: Lepisi-]. Type genus: Lepisia Lepeletier and Audinet-Serville, 1828.
Scelophysides Péringuey, 1902: 624 [stem: Scelophys-]. Type genus: Scelophysa H. C. C. Burmeister, 1844.
*Madahopliini M. Lacroix, 1997: 21 [stem: Madahopli-]. Type genus: Madahoplia Lacroix, 1998. Comment: unavailable family-group name, not based on available genus name at the time (see A. B. T. Smith 2006).
Subtribe
Pachycnemina
Laporte, 1840
Pachycnémides Laporte, 1840b: 155 [stem: Pachycnem-]. Type genus: Pachycnema Lepeletier and Audinet-Serville, 1828. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Burmeister (1844: 53, as Pachycnemidae), generally accepted as in A. B. T. Smith (2006: 170, as Pachycnemina).
Lepitrichiden Oken, 1843: 483 [stem: Lepitrich-]. Type genus: Lepitrix Lepeletier and Audinet-Serville, 1828. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Tulk (1847: 614, as Lepitrichidae).
Anisonychidae H. C. C. Burmeister, 1844: 35 [stem: Anisonych-]. Type genus: Anisonyx Latreille, 1807. Comment: Lucas (1920: 3) listed “Anisochelidae H. C. C. Burmeister” in error for “Anisonychidae H. C. C. Burmeister”; this name is a senior homonym of Anisonychini Legalov, 2003 (type genus Anisonychus Voss, 1927) in Attelabidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Lichniini
Burmeister, 1844
Lichniadae H. C. C. Burmeister, 1844: 8 [stem: Lichni-]. Type genus: Lichnia Erichson, 1835.
Tribe
Liparetrini
Burmeister, 1855
Liparetridae H. C. C. Burmeister, 1855: 187 [stem: Liparetr-]. Type genus: Liparetrus Guérin-Méneville, 1831.
Haplonychidae H. C. C. Burmeister, 1855: 224 [stem: Haplonych-]. Type genus: Haplonycha Dejean, 1836 [unjustified emendation of Aplonycha by Agassiz (1846b: 29), in prevailing usage and so deemed to be a justified emendation (Article 33.2.3.1) (see A. B. T. Smith 2006: 166); syn. of Colpochila Erichson, 1843]. Comment: this name is a senior homonym of Haplonychini Lacordaire, 1865 (type genus Haplonyx Schönherr, 1836) proposed in Curculionidae and now a synonym of Cryptoplini Lacordaire, 1863; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Allarini Britton, 1955: 125 [stem: Allar-]. Type genus: Allara Britton, 1955.
Colpochilini Britton, 1957: 10 [stem: Colpochil-]. Type genus: Colpochila Erichson, 1843.
Tribe
Macrodactylini
Kirby, 1837
Macrodactylidae Kirby, 1837: 133 [stem: Macrodactyl-]. Type genus: Macrodactylus Dejean, 1821.
Ceraspididae H. C. C. Burmeister, 1855: 91 [stem: Ceraspid-]. Type genus: Ceraspis Lepeletier and Audinet-Serville, 1828.
Dicraniadae H. C. C. Burmeister, 1855: 65 [stem: Dicrani-]. Type genus: Dicrania Lepeletier and Audinet-Serville, 1828.
Isonychidae H. C. C. Burmeister, 1855: 22 [stem: Isonych-]. Type genus: Isonychus Mannerheim, 1829.
Microcraniadae H. C. C. Burmeister, 1855: 75 [stem: Microcrani-]. Type genus: Microcrania H. C. C. Burmeister, 1855.
Plectridae H. C. C. Burmeister, 1855: 80 [stem: Plectr-]. Type genus: Plectris Lepeletier and Audinet-Serville, 1828.
*Philochlénides Lacordaire, 1856: 256 [stem: Philochloeni-]. Type genus: Philochloenia Dejean, 1833 [as Philochlaenia, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Clavipalpides Lacordaire, 1856: 267 [stem: Clavipalp-]. Type genus: Clavipalpus Laporte, 1832. Comment: published before 29 March 1856; original vernacular name available (Art. 11.7.2): first used in latinized form by Imhoff (1856 [before 25 December]: xi, as Clavipalpidae), generally accepted as in G. H. Horn (1880e: 147, as Clavipalpides [treated as Latin]).
Tribe
Maechidiini
Burmeister, 1855
Maechidiina H. C. C. Burmeister, 1855: 208 [stem: Maechidi-]. Type genus: Maechidius W. S. MacLeay, 1819.
Tribe
Melolonthini
Leach, 1819
Melolonthidae Leach, 1819: 189 [stem: Melolonth-]. Type genus: Melolontha Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subtribe
Enariina
Dewailly, 1950
Enarina Dewailly, 1950: 323 [stem: Enari-]. Type genus: Enaria Erichson, 1847. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Heptophyllina
Medvedev, 1951
Heptophyllini S. I. Medvedev, 1951: 197 [stem: Heptophyll-]. Type genus: Heptophylla Motschulsky, 1858.
Subtribe
Leucopholina
Burmeister, 1855
Leucopholidae H. C. C. Burmeister, 1855: 285 [stem: Leucophol-]. Type genus: Leucopholis Dejean, 1833. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Leucopholid-).
Subtribe
Melolonthina
Leach, 1819
*Mélolonthides Baudet-Lafarge, 1809: 12 [stem: Melolonth-]. Type genus: Melolontha Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Baudet-Lafarge (1809).
Melolonthidae Leach, 1819: 189 [stem: Melolonth-]. Type genus: Melolontha Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: published in June 1819; this family-group name was also used in the same year by W. S. MacLeay (1819 [November]: 79, as Melolonthidae).
Polyphyllidae H. C. C. Burmeister, 1855: 397 [stem: Polyphyll-]. Type genus: Polyphylla Harris, 1841.
Psilonychides Péringuey, 1904: 184 [stem: Psilonych-]. Type genus: Psilonychus H. C. C. Burmeister, 1855.
Subtribe
Pegylina
Lacroix, 1989
Pegylini M. Lacroix, 1989: 115 [stem: Pegyl-]. Type genus: Pegylis Erichson, 1847. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Pegylid-).
Subtribe
Rhizotrogina
Burmeister, 1855
Rhizotrogidae H. C. C. Burmeister, 1855: 308 [stem: Rhizotrog-]. Type genus: Rhizotrogus Latreille, 1825.
Tostegopterae J. L. LeConte, 1861: 139 [stem: Tostegopter-]. Type genus: Tostegoptera Blanchard, 1851.
Subtribe
Schizonychina
Burmeister, 1855
Schizonychidae H. C. C. Burmeister, 1855: 265 [stem: Schizonych-]. Type genus: Schizonycha Dejean, 1833.
Tribe
Oncerini
LeConte, 1861
Oncerini J. L. LeConte, 1861: 133 [stem: Oncer-]. Type genus: Oncerus J. L. LeConte, 1856.
Tribe
Pachypodini
Erichson, 1840
Pachypoden Erichson, 1840b: 29 [stem: Pachypod-]. Type genus: Pachypus Dejean, 1821 [nomen protectum; this genus name is a junior homonym of Pachypus Billberg, 1820 nomen oblitum; we provide references to support the conservation of Pachypus Dejean, 1821 as the valid name for this genus (Art. 23.9.1) (see Appendix 1)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Erichson (1847a: 100, as Pachypoda), generally accepted as in A. B. T. Smith (2006: 164, as Pachypodini).
Tribe
Pachytrichini
Burmeister, 1855
Pachytrichiadae H. C. C. Burmeister, 1855: 241 [stem: Pachytrich-]. Type genus: Pachytricha Hope, 1841.
Tribe
Phyllotocidiini
Britton, 1957
Phyllotocidiini Britton, 1957: 58 [stem: Phyllotocidi-]. Type genus: Phyllotocidium Blackburn, 1898.
Tribe
Podolasiini
Howden, 1997
Lasiopodes J. L. LeConte, 1856: 282 [stem: Lasiopod-]. Type genus: Lasiopus J. L. LeConte, 1856 [preoccupied genus name, not Lasiopus Schönherr, 1823 [Coleoptera: Curculionidae], not Lasiopus Geoffroy, 1835 [Mammalia]; syn. of Podolasia Harold, 1869]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Podolasiini Howden, 1997: 224 [stem: Podolasi-]. Type genus: Podolasia Harold, 1869.
Tribe
Scitalini
Britton, 1957
Scitalini Britton, 1957: 10 [stem: Scital-]. Type genus: Scitala Erichson, 1842.
Tribe
Sericini
Kirby, 1837
Sericidae Kirby, 1837: 128 [stem: Seric-]. Type genus: Serica W. S. MacLeay, 1819.
Subtribe
Phyllotocina
Burmeister, 1855
Phyllotocidae H. C. C. Burmeister, 1855: 182 [stem: Phyllotoc-]. Type genus: Phyllotocus Fisher von Waldheim, 1823.
Subtribe
Sericina
Kirby, 1837
Sericidae Kirby, 1837: 128 [stem: Seric-]. Type genus: Serica W. S. MacLeay, 1819. Comment: published 23 October 1837; this family-group name was also proposed in the same year by Hope (1837 [“31 December”]: 73, 107, as Sericidae); Hope (1840a: 112) refers to the publication by Kirby (1837) which is further evidence that the publication by Kirby was published first (see A. B. T. Smith 2006: 165).
Omalopliites Blanchard, 1845a: 212 [stem: Omalopli-]. Type genus: Omaloplia Schönherr, 1817. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1848: [5], as Omalopliaeidae [incorrect stem formation]), generally accepted as in Blanchard (1853: 129, as Omalopliitae); Omalopliites Blanchard, 1845 treated as unavailable and the first available name based on this type genus listed as Homalopliina H. C. C. Burmeister, 1855 by A. B. T. Smith (2006: 165).
Astaenidae H. C. C. Burmeister, 1855: 123 [stem: Astaen-]. Type genus: Astaena Erichson, 1847.
Subtribe
Trochalina
Brenske, 1898
Trochalinae Brenske, 1898: 354 [stem: Trochal-]. Type genus: Trochalus Laporte, 1832.
Tribe
Sericoidini
Erichson, 1847
Sericoideae Erichson, 1847a: 102 [stem: Sericoid-]. Type genus: Sericoides Guérin-Méneville, 1840.
Tribe
Systellopini
Sharp, 1877
Systellopides Sharp, 1877: 311 [stem: Systellop-]. Type genus: Systellopus Sharp, 1877. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Systellopod-).
Tribe
Tanyproctini
Erichson, 1847
Tanyproctini Erichson, 1847b: 653 [stem: Tanyproct-]. Type genus: Tanyproctus Ménétriés, 1832.
Subtribe
Macrophyllina
Burmeister, 1855
Macrophyllidae H. C. C. Burmeister, 1855: 447 [stem: Macrophyll-]. Type genus: Macrophylla Hope, 1837 [syn. of Aegostheta Dejean, 1833]. Comment: senior homonym of Macrophyllina Gray, 1866 (type genus Macrophyllum Gray, 1838) in Mammalia; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Aegosthetini Lacroix, 2007: 203 [stem: Aegosthet-]. Type genus: Aegostheta Dejean, 1833.
Subtribe
Tanyproctina
Erichson, 1847
Tanyproctini Erichson, 1847b: 653 [stem: Tanyproct-]. Type genus: Tanyproctus Ménétriés, 1832.
Elaphoceritae Blanchard, 1851a: 164 [stem: Elaphocer-]. Type genus: Elaphocera Géné, 1836. Comment: Elaphocerini no longer has priority over Pachydemini through a reversal of precedence (A. B. T. Smith 2006: 168).
Pachydemidae H. C. C. Burmeister, 1855: 437 [stem: Pachydem-]. Type genus: Pachydema Laporte, 1832. Comment: name conserved over Elaphocerini Blanchard, 1851 as a nomen protectum and treated as valid by A. B. T. Smith (2006: 168).
Achloidae H. C. C. Burmeister, 1855: 465 [stem: Achlo-]. Type genus: Achloa Erichson, 1840.
Cephalotrichiadae H. C. C. Burmeister, 1855: 433 [stem: Cephalotrichi-]. Type genus: Cephalotrichia Hope, 1837.
*Leptopodidae H. C. C. Burmeister, 1855: 428 [stem: Leptopod-]. Type genus: Leptopus Waltl, 1838 [preoccupied genus name, not Leptopus Latreille, 1809 [Hemiptera], not Leptopus Rafinesque, 1814 [Pisces], not Leptopus Lamarck, 1818 [Crustacea], and not Leptopus Fallén, 1823 [Diptera]; syn. of Elaphocera Gené, 1836]. Comment: family-group name unavailable (Art. 11.7.1.1): not based on a genus used as valid at the time.
Sparrmannini Péringuey, 1904: 170 [stem: Sparrmanni-]. Type genus: Sparrmannia Laporte, 1840 [nomen protectum; this genus name is a junior synonym of Leocaeta Dejean, 1833 nomen oblitum and Cephalotrichia Hope, 1837 nomen oblitum; we provide references to support the conservation of Sparrmannia as the valid name for this genus (Art. 23.9.1) (see Appendix 1); Sparrmannia is an incorrect subsequent spelling of Sparmannia Laporte, 1840, in prevailing usage and so deemed to be the correct original spelling (see Evans 1989)]. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Rutelinae
MacLeay, 1819
Rutelidae W. S. MacLeay, 1819: 69 [stem: Rutel-]. Type genus: Rutela Latreille, 1802. Comment: First Reviser (Rutelinae MacLeay, 1819 vs Anoplognathinae MacLeay, 1819) not determined, current usage maintained.
Tribe
Adoretini
Burmeister, 1844
Adoretidae H. C. C. Burmeister, 1844: 466 [stem: Adoret-]. Type genus: Adoretus Dejean, 1833.
Subtribe
Adoretina
Burmeister, 1844
Adoretidae H. C. C. Burmeister, 1844: 466 [stem: Adoret-]. Type genus: Adoretus Dejean, 1833.
Adorodociina Ohaus, 1912: 151 [stem: Adorodoci-]. Type genus: Adorodocia Brenske, 1893.
Adoroleptina Ohaus, 1912: 151 [stem: Adorolept-]. Type genus: Adoroleptus Brenske, 1893.
Pseudadoretina Ohaus, 1912: 151 [stem: Pseudadoret-]. Type genus: Pseudadoretus Semenov, 1889.
Scaphorhinadoretina Ohaus, 1912: 151 [stem: Scaphorhinadoret-]. Type genus: Scaphorhinadoretus Ohaus, 1912. Comment: erroneously cited as a new genus originally but correctly listed as a subtribe on p. 426 of the same work.
Subtribe
Adorrhinyptiina
Arrow, 1917
Adorrhinyptiini Arrow, 1917: 273 [stem: Adorrhinypti-]. Type genus: Adorrhinyptia Arrow, 1917.
Subtribe
Pachyrhinadoretina
Ohaus, 1912
Pachyrhinadoretina Ohaus, 1912: 151 [stem: Pachyrhinadoret-]. Type genus: Pachyrhinadoretus Ohaus, 1912.
Subtribe
Prodoretina
Ohaus, 1912
Prodoretina Ohaus, 1912: 151 [stem: Prodoret-]. Type genus: Prodoretus Brenske, 1893.
Subtribe
Trigonostomusina
Ohaus, 1912
Trigonostomina Ohaus, 1912: 151 [stem: Trigonostomus-]. Type genus: Trigonostomus Brenske, 1893 [placed on the Official List of Generic Names in Zoology (ICZN 2009b)]. Comment: Trigonostomus- determined to be the correct stem of this name to avoid homonymy with Trigonostomidae Graff, 1905 (type genus Trigonostomum Schmidt, 1852) in Platyhelminthes and Trigonostomusina Ohaus, 1912 placed on the Official List of Family-group Names in Zoology (ICZN 2009b).
Tribe
Alvarengiini
Frey, 1975
*Pachylides Lacordaire, 1856: 394 [stem: Pachyl-]. Type genus: Pachylus H. C. C. Burmeister, 1847 [preoccupied genus name, not Pachylus C. L. Koch, 1839 [Arachnida]; syn. of Ottokelleria d’Andretta and Martínez, 1957]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if discovered to be available then permanently invalid (Art. 39): based on preoccupied type genus; Pachylinae Sørensen, 1884 (type genus Pachylus C. L. Koch, 1839) is used as valid in Arachnida.
Alvarengiini Frey, 1975: 84 [stem: Alvarengi-]. Type genus: Alvarengius Frey, 1975.
Tribe
Anatistini
Lacordaire, 1856
Anatistides Lacordaire, 1856: 321 [stem: Anatist-]. Type genus: Anatista Brême, 1844. Comment: published before 29 March 1856; original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Imhoff (1856 [before 25 December]: xi, as Anatistidae).
Spodochlamydini Ohaus, 1918: 166 [stem: Spodochlamyd-]. Type genus: Spodochlamys H. C. C. Burmeister, 1855.
Tribe
Anomalini
Streubel, 1839
nomen protectum
Anomalidae Streubel, 1839: 136 [stem: Anomal-]. Type genus: Anomala Samouelle, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 1989d); Anomala von Block, 1799 was suppressed for the purposes of the Principle of Priority and the Principle of Homonymy (ICZN 1989d)]. Comment: nomen protectum (see A. B. T. Smith 2006: 173).
Subtribe
Anisopliina
Burmeister, 1844
Anisopliadae H. C. C. Burmeister, 1844: 208 [stem: Anisopli-]. Type genus: Anisoplia Schönherr, 1817. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Anomalina
Streubel, 1839
nomen protectum
Euchloridae Hope, 1839: 67 [stem: Euchlor-]. Type genus: Euchlora W. S. MacLeay, 1819. Comment: nomen oblitum (see A. B. T. Smith 2006: 173).
Anomalidae Streubel, 1839: 136 [stem: Anomal-]. Type genus: Anomala Samouelle, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 1989d); Anomala von Block, 1799 was suppressed for the purposes of the Principle of Priority and the Principle of Homonymy (ICZN 1989d)]. Comment: nomen protectum (see A. B. T. Smith 2006: 173).
Phyllurgaeidae Gistel, 1848: [5] [stem: Phyllurg-]. Type genus: Phyllurga Gistel, 1848 [syn. of Euchlora W. S. MacLeay, 1819].
Dilophochilina Ohaus, 1918: 166 [stem: Dilophochil-]. Type genus: Dilophochila H. W. Bates, 1888.
Subtribe
Isopliina
Péringuey, 1902
Isopliini Péringuey, 1902: 564 [stem: Isopli-]. Type genus: Isoplia H. C. C. Burmeister, 1855.
Subtribe
Leptohopliina
Potts, 1974
Lepothopliini Potts, 1974: 152 [stem: Leptohopli-]. Type genus: Leptohoplia Saylor, 1935. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Popilliina
Ohaus, 1918
Popilliina Ohaus, 1918: 133 [stem: Popilli-]. Type genus: Popillia Dejean, 1821.
Tribe
Anoplognathini
MacLeay, 1819
Anoplognathidae W. S. MacLeay, 1819: 81 [stem: Anoplognath-]. Type genus: Anoplognathus Leach, 1815.
Subtribe
Anoplognathina
MacLeay, 1819
Anoplognathidae W. S. MacLeay, 1819: 81 [stem: Anoplognath-]. Type genus: Anoplognathus Leach, 1815.
Subtribe
Brachysternina
Burmeister, 1844
Brachysternidae H. C. C. Burmeister, 1844: 455 [stem: Brachystern-]. Type genus: Brachysternus Guérin-Méneville, 1831.
Subtribe
Phalangogoniina
Ohaus, 1918
Phalangogoniina Ohaus, 1918: 176 [stem: Phalangogoni-]. Type genus: Phalangogonia H. C. C. Burmeister, 1844.
Subtribe
Platycoeliina
Burmeister, 1844
Platycoeliidae H. C. C. Burmeister, 1844: 451 [stem: Platycoeli-]. Type genus: Platycoelia Dejean, 1833.
Subtribe
Schizognathina
Ohaus, 1918
Schizognathina Ohaus, 1918: 174 [stem: Schizognath-]. Type genus: Schizognathus Fischer von Waldheim, 1823.
Tribe
Geniatini
Burmeister, 1844
Geniatidae H. C. C. Burmeister, 1844: 478 [stem: Geniat-]. Type genus: Geniates Kirby, 1819. Comment: First Reviser (Geniatini H. C. C. Burmeister, 1844 vs Leucothyreini H. C. C. Burmeister, 1844) not determined, current usage maintained.
Leucothyreidae H. C. C. Burmeister, 1844: 485 [stem: Leucothyre-]. Type genus: Leucothyreus W. S. MacLeay, 1819.
Tribe
Rutelini
MacLeay, 1819
Rutelidae W. S. MacLeay, 1819: 69 [stem: Rutel-]. Type genus: Rutela Latreille, 1802.
Subtribe
Areodina
Burmeister, 1844
Areodidae H. C. C. Burmeister, 1844: 423 [stem: Areod-]. Type genus: Areoda W. S. MacLeay, 1819.
Subtribe
Desmonychina
Arrow, 1917
Desmonychinae Arrow, 1917: 359 [stem: Desmonych-]. Type genus: Desmonyx Arrow, 1907. Comment: spelled Desmonycinae pp. xiii, 359, 382, but spelled correctly Desmonychinae p. 3 [in key].
Subtribe
Didrepanephorina
Ohaus, 1918
Didrepanophorina Ohaus, 1918: 14 [stem: Didrepanephor-]. Type genus: Didrepanephorus Wood-Mason, 1878 [as Didrepanophorus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Heterosternina
Bates, 1888
nomen protectum
Macropni G. H. Horn, 1867b: 398 [stem: Macropn-]. Type genus: Macropnus G. H. Horn, 1867. Comment: nomen oblitum (see A. B. T. Smith 2006: 171).
Heterosterninae H. W. Bates, 1888: 286 [stem: Heterostern-]. Type genus: Heterosternus Dupont, 1832. Comment: nomen protectum (see A. B. T. Smith 2006: 171).
Subtribe
Lasiocalina
Ohaus, 1918
Lasiocalina Ohaus, 1918: 30 [stem: Lasiocal-]. Type genus: Lasiocala Blanchard, 1851.
Subtribe
Oryctomorphina
Burmeister, 1847
Oryctomorphidae H. C. C. Burmeister, 1847: 28 [stem: Oryctomorph-]. Type genus: Oryctomorphus Guérin-Méneville, 1831.
Subtribe
Parastasiina
Burmeister, 1844
Parastasiidae H. C. C. Burmeister, 1844: 368 [stem: Parastasi-]. Type genus: Parastasia Westwood, 1841.
Subtribe
Rutelina
MacLeay, 1819
Rutelidae W. S. MacLeay, 1819: 69 [stem: Rutel-]. Type genus: Rutela Latreille, 1802.
Chasmodiidae H. C. C. Burmeister, 1844: 333 [stem: Chasmodi-]. Type genus: Chasmodia W. S. MacLeay, 1819.
Chrysophoridae H. C. C. Burmeister, 1844: 412 [stem: Chrysophor-]. Type genus: Chrysophora Dejean, 1821.
Macraspididae H. C. C. Burmeister, 1844: 343 [stem: Macraspid-]. Type genus: Macraspis W. S. MacLeay, 1819.
Pelidnotidae H. C. C. Burmeister, 1844: 388 [stem: Pelidnot-]. Type genus: Pelidnota W. S. MacLeay, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 2003b)].
Antichirides Lacordaire, 1856: 341 [stem: Anticheir-]. Type genus: Anticheira Eschscholtz, 1818 [as Antichira, unjustified emendation of type genus name by Agassiz (1846b: 27), not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in H. W. Bates (1888: 262, as Antichirina); incorrect original stem formation, not in prevailing usage.
Plusiotina H. W. Bates, 1888: 276 [stem: Plusiotid-]. Type genus: Plusiotis H. C. C. Burmeister, 1844. Comment: incorrect original stem formation, not in prevailing usage.
Fruhstorferiina Ohaus, 1918: 43 [stem: Fruhstorferi-]. Type genus: Fruhstorferia Kolbe, 1894.
Subfamily
Dynastinae
MacLeay, 1819
Dynastidae W. S. MacLeay, 1819: 64 [stem: Dynast-]. Type genus: Dynastes W. S. MacLeay, 1819.
Tribe
Agaocephalini
Burmeister, 1847
Agaocephalidae H. C. C. Burmeister, 1847: 280 [stem: Agaocephal-]. Type genus: Agaocephala Lepeletier and Audinet-Serville, 1828 [as Agacephala, incorrect subsequent spelling for Agacephala, in prevailing usage and so deemed to be the correct original spelling (Art. 33.3.1) (see A. B. T. Smith 2006)].
Tribe
Cyclocephalini
Laporte, 1840
Cyclocephalites Laporte, 1840b: 124 [stem: Cyclocephal-]. Type genus: Cyclocephala Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Imhoff (1856: xi, as Cyclocephalidae), generally accepted as in A. B. T. Smith (2006: 175, as Cyclocephalini).
Chalepidae H. C. C. Burmeister, 1847: 71 [stem: Chalep-]. Type genus: Chalepus W. S. MacLeay, 1819 [preoccupied genus name, not Chalepus Thunberg, 1805 [Coleoptera: Chrysomelidae]; syn. of Dyscinetus Harold, 1869]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; this family-group name is a senior homonym of Chalepini Weise, 1910 (type genus Chalepus Thunberg, 1805) currently used as valid in Chrysomelidae.
Peltonotini Arrow, 1917: 27 [stem: Peltonot-]. Type genus: Peltonotus H. C. C. Burmeister, 1847.
Acrobolbiina Ohaus, 1918: 13 [stem: Acrobolbi-]. Type genus: Acrobolbia Ohaus, 1912.
Tribe
Dynastini
MacLeay, 1819
Dynastidae W. S. MacLeay, 1819: 64 [stem: Dynast-]. Type genus: Dynastes W. S. MacLeay, 1819.
Xylotrupidae Hope, 1838b: 319 [stem: Xylotrup-]. Type genus: Xylotrupes Hope, 1837.
Megasominae Swainson, 1840: 210 [stem: Megasomat-]. Type genus: Megasoma Kirby, 1825. Comment: family-group name attributed to Imhoff (1856) in A. B. T. Smith (2006: 175); incorrect original stem formation, not in prevailing usage.
Tribe
Hexodontini
Lacordaire, 1856
Hexodontides Lacordaire, 1856: 391 [stem: Hexodont-]. Type genus: Hexodon A. G. Olivier, 1789. Comment: published before 29 March 1856; original vernacular name available (Art. 11.7.2): first used in latinized form by Imhoff (1856 [before 25 December]: xi, as Hexodontidae), generally accepted as in Arrow (1937: 4, as Hexodontini).
Tribe
Oryctini
Mulsant, 1842
Oryctésaires Mulsant, 1842: 372 [stem: Oryct-]. Type genus: Oryctes Illiger, 1798. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by MacDonald (1845: 185, as Oryctesiae [incorrect stem formation]), generally accepted as in A. B. T. Smith (2006: 175, as Oryctini); incorrect original stem formation, not in prevailing usage.
Megaceridae H. C. C. Burmeister, 1847: 212 [stem: Megacerat-]. Type genus: Megaceras Hope, 1837. Comment: incorrect original stem formation, not in prevailing usage.
Strategidae H. C. C. Burmeister, 1847: 87 [stem: Strateg-]. Type genus: Strategus Kirby, 1828.
Tribe
Oryctoderini
Endrödi, 1966
Oryctoderini Endrödi, 1966: 25 [stem: Oryctoder-]. Type genus: Oryctoderus Boisduval, 1835.
Tribe
Pentodontini
Mulsant, 1842
Pentodonaires Mulsant, 1842: 381 [stem: Pentodont-]. Type genus: Pentodon Hope, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 2003b)].
Subtribe
Cheiroplatina
Carne, 1957
Cheiroplatina Carne, 1957: 61 [stem: Cheiroplat-]. Type genus: Cheiroplatys Hope, 1837. Comment: Casey (1915a) cites “Cheiroplatids” several times within the Pentodontini without formally erecting a subtribe; current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Cheiroplate-).
Subtribe
Dipelicina
Carne, 1957
Dipelicina Carne, 1957: 117 [stem: Dipelic-]. Type genus: Dipelicus Hope, 1843.
Subtribe
Pentodontina
Mulsant, 1842
Pentodonaires Mulsant, 1842: 381 [stem: Pentodont-]. Type genus: Pentodon Hope, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 2003b)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1888: 314, as Pentodontinae), generally accepted as in A. B. T. Smith (2006: 176, as Pentodontina); incorrect original stem formation, not in prevailing usage.
Calicnémiens Mulsant, 1842: 386 [stem: Calicnemid-]. Type genus: Calicnemis Laporte, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by MacDonald (1845: 185, as Callicnemiae [incorrect stem formation]) and generally accepted as in Acloque (1896: 267, as Calicnemisii) and Houlbert (1922b: 225, as Callicnemini [incorrect stem formation]); name attributed to Blanchard (1845a) and treated as unavailable by A. B. T. Smith (2006: 168); incorrect original stem formation, not in prevailing usage.
Bothynidae H. C. C. Burmeister, 1847: 90 [stem: Bothyn-]. Type genus: Bothynus Hope, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 2008a)].
Pimelopodea H. C. C. Burmeister, 1847: 172 [stem: Pimelopod-]. Type genus: Pimelopus Erichson, 1842.
Podalgidae H. C. C. Burmeister, 1847: 90 [stem: Podalg-]. Type genus: Podalgus H. C. C. Burmeister, 1847 [placed on the Official List of Generic Names in Zoology (ICZN 2004a)].
Metanastina Carne, 1957: 32 [stem: Metanast-]. Type genus: Metanastes Arrow, 1911.
Subtribe
Pseudoryctina
Carne, 1957
Pseudoryctina Carne, 1957: 121 [stem: Pseudoryct-]. Type genus: Pseudoryctes Sharp, 1873.
Tribe
Phileurini
Burmeister, 1847
Phileuridae H. C. C. Burmeister, 1847: 138 [stem: Phileur-]. Type genus: Phileurus Latreille, 1807. Comment: usage of this name conserved over Cryptodini H. C. C. Burmeister and Schaum, 1840 (Art. 35.5).
Subtribe
Cryptodina
Burmeister and Schaum, 1840
Cryptodinae H. C. C. Burmeister and Schaum, 1840: 360 [stem: Cryptod-]. Type genus: Cryptodus MacLeay, 1819. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Cryptodont-); conservation of original stem avoids homonymy problems with Cryptodontidae Dall, 1895 (type genus Cryptodon Turton, 1822) in Mollusca: Bivalvia and Cryptodontina Lacordaire, 1856 (type genus Cryptodontes H. C. C. Burmeister, 1847) in Scarabaeidae: Cetoniinae: Trichiini.
Subtribe
Phileurina
Burmeister, 1847
Phileuridae H. C. C. Burmeister, 1847: 138 [stem: Phileur-]. Type genus: Phileurus Latreille, 1807.
Subfamily
Cetoniinae
Leach, 1815
Cetonida Leach, 1815: 99 [stem: Cetoni-]. Type genus: Cetonia Fabricius, 1775. Comment: A. B. T. Smith (2006: 177) noted that although Cetoniinae has priority over Melolonthinae, the latter is in prevailing usage at the family level and must not be displaced by the older name under Article 35.5 (only relevant in cases where authors consider Melolonthidae to be a family containing the subfamily Cetoniinae).
Tribe
Cetoniini
Leach, 1815
Cetonida Leach, 1815: 99 [stem: Cetoni-]. Type genus: Cetonia Fabricius, 1775.
Subtribe
Cetoniina
Leach, 1815
Cetonida Leach, 1815: 99 [stem: Cetoni-]. Type genus: Cetonia Fabricius, 1775.
Elaphini Schoch, 1894: 175 [stem: Elaphin-]. Type genus: Elaphinis H. C. C. Burmeister, 1842.
Glycyphanae Schoch, 1894: 175 [stem: Glycyphan-]. Type genus: Glycyphana H. C. C. Burmeister, 1842.
Pachnodii Péringuey, 1907: 371 [stem: Pachnod-]. Type genus: Pachnoda H. C. C. Burmeister, 1842.
Tephraeides Schenkling, 1921: 313 [stem: Tephrae-]. Type genus: Tephraea H. C. C. Burmeister, 1842.
Subtribe
Euphoriina
Horn, 1880
Euphoriae G. H. Horn, 1880c: 397 [stem: Euphori-]. Type genus: Euphoria H. C. C. Burmeister, 1842. Comment: name attributed to Schoch (1894) by A. B. T. Smith (2006: 182).
Subtribe
Leucocelina
Kraatz, 1882
Leucoceliden Kraatz, 1882b: 65 [stem: Leucocel-]. Type genus: Leucocelis H. C. C. Burmeister, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Schoch (1894: 213, as Leucocelidae), generally accepted as in A. B. T. Smith (2006: 182, as Leucocelina); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Leucocelid-).
Tribe
Cremastocheilini
Burmeister and Schaum, 1841
Cremastochilidae H. C. C. Burmeister and Schaum, 1841: 243 [stem: Cremastocheil-]. Type genus: Cremastocheilus Knoch, 1801. Comment: usage of this name conserved over Macromini H. C. C. Burmeister and Schaum, 1840 (Art. 35.5).
Subtribe
Aspilina
Krikken, 1984
Aspilina Krikken, 1984: 25, in key [stem: Aspil-]. Type genus: Aspilus Westwood in Schaum, 1848.
Subtribe
Coenochilina
Burmeister, 1842
Coenochilidae H. C. C. Burmeister, 1842: 148 [stem: Coenochil-]. Type genus: Coenochilus Schaum, 1841.
Subtribe
Cremastocheilina
Burmeister and Schaum, 1841
Cremastochilidae H. C. C. Burmeister and Schaum, 1841: 243 [stem: Cremastocheil-]. Type genus: Cremastocheilus Knoch, 1801. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Cymophorina
Krikken, 1984
Cymophorina Krikken, 1984: 23, in key [stem: Cymophor-]. Type genus: Cymophorus Kirby, 1827.
Subtribe
Genuchina
Krikken, 1984
Genuchina Krikken, 1984: 21, in key [stem: Genuch-]. Type genus: Genuchus Kirby, 1825.
Subtribe
Goliathopsidina
Krikken, 1984
Goliathopsidina Krikken, 1984: 25, in key [stem: Goliathopsid-]. Type genus: Goliathopsis Janson, 1881. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Goliathopse-).
Subtribe
Heterogeniina
Krikken, 1984
Heterogeniina Krikken, 1984: 25, in key [stem: Heterogeni-]. Type genus: Heterogenius Moser, 1911.
Subtribe
Lissogeniina
Krikken, 1984
Lissogeniina Krikken, 1984: 25, in key [stem: Lissogeni-]. Type genus: Lissogenius Schaum, 1845.
Subtribe
Macromina
Burmeister and Schaum, 1840
Macrominae H. C. C. Burmeister and Schaum, 1840: 360 [stem: Macrom-]. Type genus: Macroma Gory and Percheron, 1833 [syn. of Campsiura Hope, 1831]. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Macromat-).
Subtribe
Nyassinina
Krikken, 1984
Nyassinina Krikken, 1984: 25, in key [stem: Nyassin-]. Type genus: Nyassinus Westwood, 1879.
Subtribe
Oplostomina
Krikken, 1984
Oplostomina Krikken, 1984: 23, in key [stem: Oplostom-]. Type genus: Oplostomus W. S. MacLeay, 1838.
Subtribe
Pilinurgina
Krikken, 1984
Pilinurgina Krikken, 1984: 25, in key [stem: Pilinurg-]. Type genus: Pilinurgus H. C. C. Burmeister, 1842.
Subtribe
Spilophorina
Krikken, 1984
Spilophorina Krikken, 1984: 25, in key [stem: Spilophor-]. Type genus: Spilophorus Westwood in Schaum, 1848. Comment: this family-group name is a junior homonym of Spilophorini Chapuis, 1875 (type genus Spilophora Boheman, 1850) currently used as valid in Chrysomelidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Subtribe
Telochilina
Krikken, 1984
Telochilina Krikken, 1984: 21, in key [stem: Telochil-]. Type genus: Telochilus Krikken, 1975.
Subtribe
Trichoplina
Krikken, 1984
Trichoplina Krikken, 1984: 23, in key [stem: Trichopl-]. Type genus: Trichoplus H. C. C. Burmeister, 1842.
Subtribe
Trogodina
Krikken, 1984
Trogodina Krikken, 1984: 27, in key [stem: Trogod-]. Type genus: Trogodes Westwood, 1874.
Tribe
Diplognathini
Burmeister, 1842
Diplognathidae H. C. C. Burmeister, 1842: 617 [stem: Diplognath-]. Type genus: Diplognatha Gory and Percheron, 1833.
Porphyronotii Péringuey, 1907: 371 [stem: Porphyronot-]. Type genus: Porphyronota H. C. C. Burmeister, 1842.
Tribe
Goliathini
Latreille, 1829
Goliathides Latreille, 1829a: 571 [stem: Goliath-]. Type genus: Goliathus Lamarck, 1801. Comment: name attributed to Griffith and Pidgeon (1832: 492) in A. B. T. Smith (2006: 181).
Subtribe
Coryphocerina
Burmeister, 1842
Coryphoceridae H. C. C. Burmeister, 1842: 215 [stem: Coryphocer-]. Type genus: Coryphocera H. C. C. Burmeister, 1842.
Heterorrhinidae Kraatz, 1880a: 21 [stem: Heterorhin-]. Type genus: Heterorhina Westwood, 1842 [as Heterorrhina, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Ceratorrhinidae Kraatz, 1880a: 18 [stem: Ceratorhin-]. Type genus: Ceratorhina Westwood, 1843 [as Ceratorrhina, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Bothrorrhinae Schoch, 1894: 173 [stem: Bothrorrhin-]. Type genus: Bothrorrhina H. C. C. Burmeister, 1842.
Gnathoceridae Schoch, 1894: 170 [stem: Gnathocer-]. Type genus: Gnathocera Kirby, 1825.
Ischnosceli Schoch, 1894: 170 [stem: Ischnoscelid-]. Type genus: Ischnoscelis H. C. C. Burmeister, 1842. Comment: incorrect original stem formation, not in prevailing usage.
Rhomborrhinae Schoch, 1894: 171 [stem: Rhomborhin-]. Type genus: Rhomborhina Hope, 1837 [as Rhomborrhina, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tmesorrhinae Schoch, 1894: 170 [stem: Tmesorrhin-]. Type genus: Tmesorrhina Westwood, 1842.
Coelorrhinae Schoch, 1895: iii [stem: Coelorrhin-]. Type genus: Coelorrhina Hope, 1841 [Coelorrhina is an incorrect subsequent spelling for Caelorrhina in prevailing usage, and so deemed to be the correct original spelling (Article 33.3.1) (see A. B. T. Smith 2006: 181)].
Mecynorrhinina Schenkling, 1921: 15 [stem: Mecynorhin-]. Type genus: Mecynorhina Hope, 1837 [as Mecynorrhina, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Stephanorrhinina Schenkling, 1921: 35 [stem: Stephanorrhin-]. Type genus: Stephanorrhina H. C. C. Burmeister, 1842.
Subtribe
Dicronocephalina
Krikken, 1984
Dicronocephalina Krikken, 1984: 37, in key [stem: Dicronocephal-]. Type genus: Dicronocephalus Hope, 1831 [Dicronocephalus is an incorrect subsequent spelling for Dicranocephalus, since the incorrect subsequent spelling is in prevailing usage, it is now considered the correct original spelling of the name under Article 33.3.1; the original spelling placed the name in homonymy with Dicranocephalus Hahn, 1826 (Hemiptera) but usage of the subsequent spelling as the correct spelling avoids this homonymy problem (see A. B. T. Smith 2006)].
Subtribe
Goliathina
Latreille, 1829
Goliathides Latreille, 1829a: 571 [stem: Goliath-]. Type genus: Goliathus Lamarck, 1801. Comment: name attributed to Griffith and Pidgeon (1832) in A. B. T. Smith (2006: 181).
Hypselogeniae Schoch, 1894: 169 [stem: Hypselogeni-]. Type genus: Hypselogenia H. C. C. Burmeister, 1840.
Subtribe
Ichnestomatina
Burmeister, 1842
Ischnostomidae H. C. C. Burmeister, 1842: 600 [stem: Ichnestomat-]. Type genus: Ichnestoma Gory and Percheron, 1833 [as Ichnostoma, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Gymnetini
Kirby, 1827
Gymnetidae Kirby, 1827: 150 [stem: Gymnet-]. Type genus: Gymnetis W. S. MacLeay, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 1967a)].
Subtribe
Blaesiina
Schoch, 1895
Blaesiae Schoch, 1895: iii [stem: Blaesi-]. Type genus: Blaesia H. C. C. Burmeister, 1842.
Subtribe
Gymnetina
Kirby, 1827
Gymnetidae Kirby, 1827: 150 [stem: Gymnet-]. Type genus: Gymnetis W. S. MacLeay, 1819. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Gymnetid-).
Clinteriidae Kraatz, 1882a: 49 [stem: Clinteri-]. Type genus: Clinteria H. C. C. Burmeister, 1842.
Stethodesmae Schoch, 1894: 172 [stem: Stethodesmat-]. Type genus: Stethodesma Bainbridge, 1841. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Phaedimini
Schoch, 1894
Phaedimi Schoch, 1894: 169 [stem: Phaedim-]. Type genus: Phaedimus Westwood, 1841.
Tribe
Schizorhinini
Burmeister, 1842
Schizorrhinidae H. C. C. Burmeister, 1842: 530 [stem: Schizorhin-]. Type genus: Schizorhina Kirby, 1825. Comment: First Reviser (Lomapterini H. C. C. Burmeister, 1842 vs Macronotini H. C. C. Burmeister, 1842 vs Schizorrhinini H. C. C. Burmeister, 1842) not determined, current usage maintained.
Subtribe
Lomapterina
Burmeister, 1842
Lomapteridae H. C. C. Burmeister, 1842: 310 [stem: Lomapter-]. Type genus: Lomaptera Gory and Percheron, 1833. Comment: First Reviser (Lomapterina H. C. C. Burmeister, 1842 vs Macronotina H. C. C. Burmeister, 1842) not determined, current usage maintained.
Macronotidae H. C. C. Burmeister, 1842: 318 [stem: Macronot-]. Type genus: Macronota Hoffmannsegg, 1817.
Subtribe
Schizorhinina
Burmeister, 1842
Schizorrhinidae H. C. C. Burmeister, 1842: 530 [stem: Schizorhin-]. Type genus: Schizorhina Kirby, 1825 [as Schizorrhina, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Diaphoniadae Kraatz, 1880b: 195 [stem: Diaphoni-]. Type genus: Diaphonia Newman, 1840.
Eupoecilidae Kraatz, 1880b: 188 [stem: Eupoecil-]. Type genus: Eupoecila H. C. C. Burmeister, 1842.
Hemipharidae Kraatz, 1880b: 182 [stem: Hemiphar-]. Type genus: Hemipharis H. C. C. Burmeister, 1842.
Tribe
Stenotarsiini
Kraatz, 1880
Stenotarsiden Kraatz, 1880b: 182 [stem: Stenotarsi-]. Type genus: Stenotarsia H. C. C. Burmeister, 1842.
Subtribe
Anochiliina
Krikken, 1984
*Anochiliens Pouillaude, 1917: 54 [stem: Anochili-]. Type genus: Anochilia H. C. C. Burmeister, 1842. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Anochiliina Krikken, 1984: 31, in key [stem: Anochili-]. Type genus: Anochilia H. C. C. Burmeister, 1842.
Subtribe
Coptomiina
Schenkling, 1921
Coptomiini Schenkling, 1921: 147 [stem: Coptomi-]. Type genus: Coptomia H. C. C. Burmeister, 1842.
Subtribe
Chromoptiliina
Krikken, 1984
Chromoptiliina Krikken, 1984: 31, in key [stem: Chromoptili-]. Type genus: Chromoptilia Westwood, 1842.
Subtribe
Doryscelina
Schenkling, 1921
*Dorysceliens Pouillaude, 1917: 53, in key [stem: Doryscel-]. Type genus: Doryscelis Dejean, 1836. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Doryscelina Schenkling, 1921: 148 [stem: Doryscel-]. Type genus: Doryscelis Dejean, 1836. Comment: name attributed to Krikken (1984) by A. B. T. Smith (2006: 180); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Doryscelid-).
Subtribe
Euchroeina
Paulian and Descarpentries, 1982
*Euchroeens Pouillaude, 1917: 64 [stem: Euchroe-]. Type genus: Euchroea H. C. C. Burmeister, 1842. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899; the author also used the incorrect spelling Euchaeeus in the key on page 54.
Euchroeina Paulian and Descarpentries, 1982: 5 [stem: Euchroe-]. Type genus: Euchroea H. C. C. Burmeister, 1842. Comment: the name Euchroeidae Dahlbom, 1854 (type genus Euchroeus Latreille, 1809) is available in Hymenoptera; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Subtribe
Heterophanina
Schoch, 1894
Heterophanae Schoch, 1894: 173 [stem: Heterophan-]. Type genus: Heterophana H. C. C. Burmeister, 1842.
Subtribe
Heterosomatina
Krikken, 1984
Heterosomatina Krikken, 1984: 29, in key [stem: Heterosomat-]. Type genus: Heterosoma Schaum, 1845.
Subtribe
Pantoliina
Krikken, 1984
*Pantoliens Pouillaude, 1917: 54 [stem: Pantoli-]. Type genus: Pantolia H. C. C. Burmeister, 1842. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Pantoliina Krikken, 1984: 33, in key [stem: Pantoli-]. Type genus: Pantolia H. C. C. Burmeister, 1842.
Subtribe
Parachiliina
Krikken, 1984
Parachiliina Krikken, 1984: 33, in key [stem: Parachili-]. Type genus: Parachilia H. C. C. Burmeister, 1842.
Subtribe
Stenotarsiina
Kraatz, 1880
Stenotarsiden Kraatz, 1880b: 182 [stem: Stenotarsi-]. Type genus: Stenotarsia H. C. C. Burmeister, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jakobson (1915: 961, as Stenotarsina [incorrect stem formation]), generally accepted as in A. B. T. Smith (2006: 179, as Stenotarsiini); incorrect original stem formation, not in prevailing usage.
Tribe
Taenioderini
Mikšić, 1976
Taenioderina Mikšić, 1976: 29 [stem: Taenioder-]. Type genus: Taeniodera H. C. C. Burmeister, 1842. Comment: First Reviser found (Taenioderini Mikšić, 1976 vs Chalcotheini Mikšić, 1976) is Krikken (1984: 63).
Subtribe
Chalcotheina
Mikšić, 1976
Chalcotheina Mikšić, 1976: 357 [stem: Chalcothe-]. Type genus: Chalcothea H. C. C. Burmeister, 1842.
Subtribe
Taenioderina
Mikšić, 1976
Taenioderina Mikšić, 1976: 29 [stem: Taenioder-]. Type genus: Taeniodera H. C. C. Burmeister, 1842.
Tribe
Trichiini
Fleming, 1821
Trichiadae Fleming, 1821: 50 [stem: Trichi-]. Type genus: Trichius Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 2004c)]. Comment: Trichiinae Lozek, 1956 (type genus Trichia Hartmann, 1840) proposed in Mollusca: Gastropoda is a junior homonym of this family-group name; Trichiinae Lozek, 1956 was recently placed on the Official index of Rejected and Invalid Family-Group Names in Zoology (ICZN 2004c).
Subtribe
Cryptodontina
Lacordaire, 1856
Cryptodontides Lacordaire, 1856: 462 [stem: Cryptodont-]. Type genus: Cryptodontes H. C. C. Burmeister, 1847. Comment: published before 29 March 1856; original vernacular name available (Art. 11.7.2): first used in latinized form by Imhoff (1856 [before 25 December]: xii, as Cryptodontidae), generally accepted as in A. B. T. Smith (2006: 177, as Cryptodontina); the name Cryptodontidae Dall, 1895 (type genus Cryptodon Turton, 1822) is available in Mollusca: Bivalvia while the name Cryptodontidae Owen, 1860 used in Therapsida is unavailable because it is not based on an available genus-group name; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Subtribe
Incina
Burmeister, 1842
Incadae H. C. C. Burmeister, 1842: 704 [stem: Inc-]. Type genus: Inca Lepeletier and Audinet-Serville, 1828.
Subtribe
Osmodermatina
Schenkling, 1922
Osmodermini Schenkling, 1922: 3 [stem: Osmodermat-]. Type genus: Osmoderma Lepeletier and Audinet-Serville, 1828 [placed on the Official List of Generic Names in Zoology (ICZN 2007)]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Platygeniina
Krikken, 1984
Platygeniini Krikken, 1984: 18, in key [stem: Platygeni-]. Type genus: Platygenia W. S. MacLeay, 1819.
Subtribe
Trichiina
Fleming, 1821
*Trichides Baudet-Lafarge, 1809: 34 [stem: Trichi-]. Type genus: Trichius Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 2004c)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Baudet-Lafarge (1809); incorrect original stem formation, not in prevailing usage.
Trichiadae Fleming, 1821: 50 [stem: Trichi-]. Type genus: Trichius Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 2004c)].
Paniscidae Gistel, 1848: [5] [stem: Panisc-]. Type genus: Paniscus Gistel, 1848 [preoccupied genus name, not Paniscus Schrank, 1802 [Hymenoptera]; Gistel (1848: [5]) originally included Scarabaeus fasciatus Linnaeus, 1758 and Trichius zonatus Germar, 1831 in his genus Paniscus, we hereby select Scarabaeus fasciatus Linnaeus, 1758 as the type species of Paniscus Gistel, 1848; syn. nov. of Trichius Fabricius, 1775]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Elpidides Péringuey, 1907: 314 [stem: Elpid-]. Type genus: Elpidus Péringuey, 1907.
Myodermini Péringuey, 1907: 313 [stem: Myoderm-]. Type genus: Myodermum H. C. C. Burmeister and Schaum, 1840.
Tribe
Valgini
Mulsant, 1842
Valguaires Mulsant, 1842: 519 [stem: Valg-]. Type genus: Valgus Scriba, 1790.
Subtribe
Microvalgina
Kolbe, 1904
Microvalginae Kolbe, 1904: 10 [stem: Microvalg-]. Type genus: Microvalgus Kraatz, 1883.
Subtribe
Valgina
Mulsant, 1842
Valguaires Mulsant, 1842: 519 [stem: Valg-]. Type genus: Valgus Scriba, 1790. Comment: published before 6 August 1842; original vernacular name available (Art. 11.7.2): used in latinized form by several authors, generally accepted as in A. B. T. Smith (2006: 177, as Valgini); this family-group name was also used in the same year by Burmeister (1842 [before 28 December]: 718, as Valgidae); incorrect original stem formation, not in prevailing usage.
Acanthovalginae Kolbe, 1904: 11 [stem: Acanthovalg-]. Type genus: Acanthovalgus Kraatz, 1895.
Cosmovalginae Kolbe, 1904: 11 [stem: Cosmovalg-]. Type genus: Cosmovalgus Kolbe, 1897.
Dasyvalginae Kolbe, 1904: 11 [stem: Dasyvalg-]. Type genus: Dasyvalgus Kolbe, 1904.
Ischnovalginae Kolbe, 1904: 9 [stem: Ischnovalg-]. Type genus: Ischnovalgus Kolbe, 1897.
Sphinctovalginae Kolbe, 1904: 9 [stem: Sphinctovalg-]. Type genus: Sphinctovalgus Kolbe, 1904.
Tribe
Xiphoscelidini
Burmeister, 1842
Xiphoscelideae H. C. C. Burmeister, 1842: 613 [stem: Xiphoscelid-]. Type genus: Xiphoscelis H. C. C. Burmeister, 1842. Comment: incorrect original stem formation, not in prevailing usage.
†Family
Coprinisphaeridae
Genise, 2004
Coprinisphaeridae Genise, 2004: 426 [stem: Coprinisphaer-]. Type genus: Coprinisphaera Sauer, 1955 [placed on the Official List of Generic Names in Zoology and given precedence over Fontanai Roselli, 1939 (ICZN 2008b)]. Comment: ichnotaxon based on dung beetle burrows.
†Family
Pallichnidae
Genise, 2004
Pallichnidae Genise, 2004: 432 [stem: Pallichn-]. Type genus: Pallichnus Retallack, 1984. Comment: ichnotaxon based on dung beetle burrows.
Superfamily
Scirtoidea
Fleming, 1821
Scirtesidae Fleming, 1821: 50 [stem: Scirt-]. Type genus: Scirtes Illiger, 1807. Comment: as pointed out by Lawrence and Newton (1995: 838) the name Scirtoidea Fleming, 1821 has priority over Clamboidea Fischer von Waldheim, 1821.
Family
Decliniidae
Nikitsky, Lawrence, Kirejtshuk and Gratshev, 1994
Decliniidae Nikitsky et al., 1994: 7 [stem: Declini-]. Type genus: Declinia Nikitsky et al., 1994.
Family
Eucinetidae
Lacordaire, 1857
Eucinétides Lacordaire, 1857: 281 [stem: Eucinet-]. Type genus: Eucinetus Germar, 1818. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 181, as Eucinetini), generally accepted as in Hansen (1996: 135, as Eucinetidae).
Cryptomeridae Broun, 1893: 1358 [stem: Cryptomer-]. Type genus: Cryptomera Broun, 1893 [syn. of Eucinetus Germar, 1818].
Family
Clambidae
Fischer von Waldheim, 1821
Clambini Fischer von Waldheim, 1821: 52 [stem: Clamb-]. Type genus: Clambus Fischer von Waldheim, 1821.
Subfamily
Calyptomerinae
Crowson, 1955
Calyptomeridae Crowson, 1955: 11 [stem: Calyptomer-]. Type genus: Calyptomerus Redtenbacher, 1849.
Subfamily
Acalyptomerinae
Crowson, 1979
Acalyptomerinae Crowson, 1979: 612, in key [stem: Acalyptomer-]. Type genus: Acalyptomerus Crowson, 1979.
Subfamily
Clambinae
Fischer von Waldheim, 1821
Clambini Fischer von Waldheim, 1821: 52 [stem: Clamb-]. Type genus: Clambus Fischer von Waldheim, 1821.
Family
Scirtidae
Fleming, 1821
Scirtesidae Fleming, 1821: 50 [stem: Scirt-]. Type genus: Scirtes Illiger, 1807.
†Sinodryopitidae Hong, 2002: 107 [stem: Sinodryopit-]. Type genus: Sinodryopites Hong, 2002. Comment: originally described in Byrrhoidea; synonymy with Scirtidae by Kirejtshuk and Azar (2008: 29).
Subfamily
Scirtinae
Fleming, 1821
Scirtesidae Fleming, 1821: 50 [stem: Scirt-]. Type genus: Scirtes Illiger, 1807. Comment: incorrect original stem formation, not in prevailing usage.
Cyphonidae Stephens, 1829a: 11 [stem: Cyphon-]. Type genus: Cyphon Paykull, 1799.
Elodiidae Shuckard, 1839b: 41 [stem: Elod-]. Type genus: Elodes Latreille, 1797. Comment: incorrect original stem formation, not in prevailing usage.
Atopidini Pic, 1914: 16 [stem: Atopid-]. Type genus: Atopida A. White, 1846.
Subfamily
Nipponocyphoninae
Lawrence and Yoshitomi, 2007
Nipponocyphoninae Lawrence and Yoshitomi, 2007: 522 [stem: Nipponocyphon-]. Type genus: Nipponocyphon Lawrence and Yoshitomi, 2007.
Subfamily
Stenocyphoninae
Lawrence and Yoshitomi, 2007
Stenocyphoninae Lawrence and Yoshitomi, 2007: 522 [stem: Stenocyphon-]. Type genus: Stenocyphon Lawrence, 2001.
†Family
Elodophthalmidae
Kirejtshuk and Azar, 2008
Elodophthalmidae Kirejtshuk and Azar, 2008: 24 [stem: Elodophthalm-]. Type genus: Elodophthalmus Kirejtshuk and Azar, 2008.
†Famil
y Mesocinetidae
Kirejtshuk and Ponomarenko, 2010
Mesocinetidae Kirejtshuk and Ponomarenko, 2010: 304 [stem: Mesocinet-]. Type genus: Mesocinetus Ponomarenko, 1986.
Superfamily
Dascilloidea
Guérin-Méneville, 1843 (1834)
Dascillidae Guérin-Méneville, 1843: 193 [stem: Dascill-]. Type genus: Dascillus Latreille, 1797. Comment: usage of younger name conserved over Atopoidea Laporte, 1834 (Art. 40.2); although Rhipiceroidea is also an older name for this superfamily, its use for a taxon including the families Dascilidae (including Karumiinae) and Rhipiceridae is likely to cause some confusion and we therefore continue to use Dascilloidea as valid; the concept of Rhipiceroidea has varied among authors, Crowson (1953) used Rhipiceroidea for the families Rhipiceridae and Callirhipidae, which are not now considered to form a monophyletic group, and Dascilloidea for Dascillidae plus those families now included in Scirtoidea; the family group name Sandalidae (which is a junior synonym of Rhipiceridae) was used by Craighead (1921) and Böving and Craighead (1931) for Rhipiceridae in the strict sense, while Emden (1924, 1931, 1933) used the same name for a family which also included those genera now placed in Callirhipidae; Crowson (1971, 1973b) proposed a reconstituted Dascilloidea for Dascillidae plus Rhipiceridae (sensu stricto) and excluded Callirhipidae from Rhipiceroidea and placed it in another superfamily Artematopoidea (along with Artematopidae (now Artematopodidae) and Brachypsectridae); finally, the evidence for placing Dascillidae and Rhipiceridae in the same superfamily is not convincing (especially when the larvae are taken into account) so it is quite possible that Dascilloidea in the sense of Crowson (1971) may cease to exist when more evidence is presented.
Family
Dascillidae
Guérin-Méneville, 1843 (1834)
Dascillidae Guérin-Méneville, 1843: 193 [stem: Dascill-]. Type genus: Dascillus Latreille, 1797.
Subfamily
Dascillinae
Guérin-Méneville, 1843 (1834)
Dascillidae Guérin-Méneville, 1843: 193 [stem: Dascill-]. Type genus: Dascillus Latreille, 1797.
Tribe
Cinnabariini
Pic, 1914
Cinnabariini Pic, 1914: 15 [stem: Cinnabari-]. Type genus: Cinnabarium Fairmaire, 1895 [syn. of Coptocera Murray, 1868].
Tribe
Dascillini
Guérin-Méneville, 1843 (1834)
Atopites Laporte, 1834a: 227 [stem: Atop-]. Type genus: Atopa Paykull, 1799 [syn. of Dascillus Latreille, 1797]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Laporte (1836: 21, as Atopidae).
Dascillidae Guérin-Méneville, 1843: 193 [stem: Dascill-]. Type genus: Dascillus Latreille, 1797. Comment: name proposed to replace Atopidae Laporte, 1834 because of the synonymy of the type genus; usage of younger name conserved over Atopini Laporte, 1834 (Art. 40.2) (see Lawrence and Newton 1995).
Subfamily
Karumiinae
Escalera, 1913
Karuminae Escalera, 1913: 320 [stem: Karumi-]. Type genus: Karumia Escalera, 1913.
Tribe
Emmitini
Escalera, 1914
Emminae Escalera, 1913: 318 [stem: Emm-]. Type genus: Emma Escalera, 1913 [preoccupied genus name, not Emma Gray, 1843 [Bryozoa]; syn. of Emmita Escalera, 1914]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Emmitinae Escalera, 1914a: 349 [stem: Emmit-]. Type genus: Emmita Escalera, 1914. Comment: replacement name for Emminae Escalera, 1913 because of the homonymy of the type genus.
Tribe
Escalerinini
Paulus, 1972
Escalerini Paulus, 1972a: 49, in key [stem: Escalerin-]. Type genus: Escalerina Bolívar, 1926. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Genecerini
Pic, 1914
Genecerini Pic, 1914: 16 [stem: Genecer-]. Type genus: Genecerus Walker, 1871.
Tribe
Karumiini
Escalera, 1913
Karuminae Escalera, 1913: 320 [stem: Karumi-]. Type genus: Karumia Escalera, 1913. Comment: incorrect original stem formation, not in prevailing usage.
Zarudniolidae Semenov and Martynov, 1925: 74 [stem: Zarudniol-]. Type genus: Zarudniola Semenov and Martynov, 1925 [syn. of Karumia Escalera, 1913].
Family
Rhipiceridae
Latreille, 1834
Rhipicerides Latreille, 1834: 167 [stem: Rhipicer-]. Type genus: Rhipicera Latreille, 1816. Comment: published in issue 1 of volume 3 of the Annales de la Société Entomologique de France; this family-group name was also used in the same year by Laporte (1834a: 226, as Rhipicérites) in issue 2 of volume 3 of the same journal.
Sandalidae Jakobson, 1913: 729 [stem: Sandal-]. Type genus: Sandalus Knoch, 1801.
Superfamily
Buprestoidea
Leach, 1815
Buprestides Leach, 1815: 85 [stem: Buprest-]. Type genus: Buprestis Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1994e)].
Family
Schizopodidae
LeConte, 1859
Schizopodidae J. L. LeConte, 1859b: 122 [stem: Schizopod-]. Type genus: Schizopus J. L. LeConte, 1858 [placed on the Official List of Generic Names in Zoology (ICZN 1993d)].
Subfamily
Schizopodinae
LeConte, 1859
Schizopodidae J. L. LeConte, 1859b: 122 [stem: Schizopod-]. Type genus: Schizopus J. L. LeConte, 1858 [placed on the Official List of Generic Names in Zoology (ICZN 1993d)].
Tribe
Dystaxiini
Théry, 1929
Dystaxini Théry, 1929: 60 [stem: Dystaxi-]. Type genus: Dystaxia J. L. LeConte, 1866. Comment: incorrect original stem formation, not in prevailing usage.
Glyptoscelimorphini Cobos, 1963: 354 [stem: Glyptoscelimorph-]. Type genus: Glyptoscelimorpha G. H. Horn, 1893.
†Tribe
Electrapatini
Iablokoff-Khnzorian, 1962
Electrapatidae Iablokoff-Khnzorian, 1962: 87 [stem: Electrapat-]. Type genus: Electrapate Iablokoff-Khnzorian, 1962.
Tribe
Schizopodini
LeConte, 1859
Schizopodidae J. L. LeConte, 1859b: 122 [stem: Schizopod-]. Type genus: Schizopus J. L. LeConte, 1858 [Schizopus J. L. LeConte, 1858 has precedence over Schizopus Claparéde et Lachman, 1858 and was placed on the Official List of Generic Names in Zoology (ICZN 1993d)].
Family
Buprestidae
Leach, 1815
Buprestides Leach, 1815: 85 [stem: Buprest-]. Type genus: Buprestis Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1994e)].
Subfamily
Julodinae
Lacordaire, 1857
Julodides Lacordaire, 1857: 10 [stem: Julod-]. Type genus: Julodis Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 154, as Julodini), generally accepted as in Bellamy (2008a: 23, as Julodinae).
Sternocerini Csiki, 1904: 132 [stem: Sternocer-]. Type genus: Sternocera Eschscholtz, 1829.
Amblysternini Cobos, 1955: 22, in key [stem: Amblystern-]. Type genus: Amblysterna Saunders, 1871.
Subfamily
Polycestinae
Lacordaire, 1857
Polycestides Lacordaire, 1857: 61 [stem: Polycest-]. Type genus: Polycesta Dejean, 1833.
Tribe
Acmaeoderini
Kerremans, 1893
Acmaeoderini Kerremans, 1893: 112 [stem: Acmaeoder-]. Type genus: Acmaeodera Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 2005b)].
Subtribe
Acmaeoderina
Kerremans, 1893
Acmaeoderini Kerremans, 1893: 112 [stem: Acmaeoder-]. Type genus: Acmaeodera Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 2005b)].
Subtribe
Acmaeoderoidina
Cobos, 1955
Acmaeoderoidini Cobos, 1955: 15 [stem: Acmaeoderoid-]. Type genus: Acmaeoderoides Van Dyke, 1942.
Subtribe
Nothomorphina
Cobos, 1955
Notomorphini Cobos, 1955: 17 [stem: Nothomorph-]. Type genus: Nothomorpha Saunders, 1871 [as Notomorpha, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: Cobos (1955: 23) also used the spelling Nothomorphini in his original paper, this is considered a misspelling since he listed the type genus name as Notomorpha; incorrect original stem formation, not in prevailing usage.
Acmaeoderini
incertae sedis
*Odetteina Volkovitsh, 2001: 52, 91 [stem: Odette-]. Type genus: Odettea Baudon, 1966. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Astraeini
Cobos, 1980
Astraeusini Cobos, 1980: 28 [stem: Astrae-]. Type genus: Astraeus Laporte and Gory, 1838 [placed on the Official List of Generic Names in Zoology (ICZN 1966)]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Bulini
Bellamy, 1995
Bulisina Bellamy, 1995: 173 [stem: Bul-]. Type genus: Bulis Laporte and Gory, 1838. Comment: incorrect original stem formation, not in prevailing usage; stem correction by Bellamy (1996: 222).
Tribe
Haplostethini
LeConte, 1861
Haplostethini J. L. LeConte, 1861: 155 [stem: Haplosteth-]. Type genus: Haplostethus J. L. LeConte, 1860.
Mastogenini J. L. LeConte and G. H. Horn, 1883: 199 [stem: Mastogeni-]. Type genus: Mastogenius Solier, 1849. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Paratracheini
Cobos, 1980
Paratrachysae Cobos, 1980: 47 [stem: Paratrache-]. Type genus: Paratrachys Saunders, 1873. Comment: here we adopt the stem Paratrache- since the correct stem for the genus Trachys, with the same ending, was recently determined to be Trache- (ICZN 2009a).
Tribe
Perucolini
Cobos, 1980
Perucolini Cobos, 1980: 81 [stem: Perucol-]. Type genus: Perucola Théry, 1925.
Tribe
Polycestini
Lacordaire, 1857
Polycestides Lacordaire, 1857: 61 [stem: Polycest-]. Type genus: Polycesta Dejean, 1833.
Subtribe
Polycestina
Lacordaire, 1857
Polycestides Lacordaire, 1857: 61 [stem: Polycest-]. Type genus: Polycesta Dejean, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Stein (1868: 62, as Polycestini), generally accepted as in Bellamy (2008a: 23, as Polycestini).
Subtribe
Xenopseina
Volkovitsh, 2008
Xenopsina Volkovitsh, 2008: 628 [stem: Xenopse-]. Type genus: Xenopsis Saunders, 1867. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Polyctesini
Cobos, 1955
Polyctesini Cobos, 1955: 6 [stem: Polyctes-]. Type genus: Polyctesis Marseul, 1865. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Polyctese-).
Tribe
Prospherini
Cobos, 1980
Prospheresini Cobos, 1980: 84 [stem: Prospher-]. Type genus: Prospheres Saunders, 1868. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Ptosimini
Kerremans, 1903
Ptosimites Kerremans, 1903: 37 [stem: Ptosim-]. Type genus: Ptosima Dejean, 1833.
Tribe
Thrincopygini
LeConte, 1861
Thrincopygini J. L. LeConte, 1861: 154 [stem: Thrincopyg-]. Type genus: Thrincopyge J. L. LeConte, 1858.
Tribe
Tyndaridini
Cobos, 1955
Tyndarini Cobos, 1955: 11 [stem: Tyndarid-]. Type genus: Tyndaris J. Thomson, 1857.
Subtribe
Mimicoclytrinina
Bellamy, 2003
Acherusini Cobos, 1955: 24, in key [stem: Acherusi-]. Type genus: Acherusia Laporte and Gory, 1838 [preoccupied genus name, not Acherusia Costa, 1834 [Crustacea]; syn. of Mimicoclytrina Bellamy, 2003]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Mimicoclytrinina Bellamy, 2003: 25 [stem: Mimicoclytrin-]. Type genus: Mimicoclytrina Bellamy, 2003. Comment: replacement name for Acherusiina Cobos, 1955 because of the homonymy of the type genus.
Subtribe
Pseudacherusiina
Cobos, 1980
Pseudoacherusini Cobos, 1980: 78 [stem: Pseudacherusi-]. Type genus: Pseudacherusia Kerremans, 1905. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Tylaucheniina
Cobos, 1959
Tylacheniae Cobos, 1959: 4 [stem: Tylaucheni-]. Type genus: Tylauchenia H. C. C. Burmeister, 1872 [syn. of Ocypetes Saunders, 1871]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Tyndaridina
Cobos, 1955
Tyndarini Cobos, 1955: 11 [stem: Tyndarid-]. Type genus: Tyndaris J. Thomson, 1857. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Xyroscelidini
Cobos, 1955
Xiroscelini Cobos, 1955: 19 [stem: Xyroscelid-]. Type genus: Xyroscelis Saunders, 1868 [as Xiroscelis, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Galbellinae
Reitter, 1911
Galbellinae Reitter, 1911: 178 [stem: Galbell-]. Type genus: Galbella Westwood, 1848.
Subfamily
Chrysochroinae
Laporte, 1835
Chrysochroidae Laporte, 1835b: 158 [stem: Chrysochro-]. Type genus: Chrysochroa Dejean, 1833.
Tribe
Chrysochroini
Laporte, 1835
Chrysochroidae Laporte, 1835b: 158 [stem: Chrysochro-]. Type genus: Chrysochroa Dejean, 1833.
Subtribe
Chalcophorina
Lacordaire, 1857 (1848)
Anaglyptisidae Gistel, 1848: [5] [stem: Anaglypt-]. Type genus: Anaglyptes Gistel, 1848 [syn. of Chalcophora Dejean, 1833]. Comment: senior homonym of Anaglyptini Lacordaire, 1868 (type genus Anaglyptus Mulsant, 1839) in Cerambycidae; incorrect original stem formation, not in prevailing usage; the younger name Chalcophorina Lacordaire, 1857 is conserved over this name (Art. 40.2).
Chalcophorides Lacordaire, 1857: 14 [stem: Chalcophor-]. Type genus: Chalcophora Dejean, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 151, as Chalcophorae), generally accepted as in Bellamy (2008a: 23, as Chalcophorina); name conserved over the older name Anaglyptina Gistel, 1848 (Art. 40.2).
*Chrysodemides H. Deyrolle, 1865: 11 [stem: Chrysodem-]. Type genus: Chrysodema Laporte and Gory, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 2004b)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to H. Deyrolle (1865).
Chrysodémides Kerremans, 1892: 49 [stem: Chrysodem-]. Type genus: Chrysodema Laporte and Gory, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 2004b)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Skinner (1892: 124, as Chrysodemidae).
Chalcophorellini Tôyama, 1986: 189 [stem: Chalcophorell-]. Type genus: Chalcophorella Kerremans, 1903.
Iridotaenini Tôyama, 1987: 5 [stem: Iridotaeni-]. Type genus: Iridotaenia Deyrolle, 1864 [placed on the Official List of Generic Names in Zoology (ICZN 2004b)]. Comment: incorrect original stem formation, not in prevailing usage.
Lampropeplina Holyński, 1993: 22 [stem: Lampropepl-]. Type genus: Lampropepla Fairmaire, 1904 [syn. of Madecassia Kerremans, 1903].
Subtribe
Chrysochroina
Laporte, 1835
Chrysochroidae Laporte, 1835b: 158 [stem: Chrysochro-]. Type genus: Chrysochroa Dejean, 1833.
Catoxanthina Jakobson, 1913: 772 [stem: Catoxanth-]. Type genus: Catoxantha Solier, 1833.
Subtribe
Eucallopistina
Bellamy, 2003
Callopistina Kurosawa, 1990: 63 [stem: Callopist-]. Type genus: Callopistus Deyrolle, 1864 [preoccupied genus name, not Callopistus Say, 1831 [Coleoptera: Curculionidae]; syn. of Eucallopistus Bellamy, 2003]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Eucallopistina Bellamy, 2003: 31 [stem: Eucallopist-]. Type genus: Eucallopistus Bellamy, 2003. Comment: replacement name for Callopistina Kurosawa, 1990 because of the homonymy of the type genus.
Tribe
Dicercini
Gistel, 1848
Dicercaeidae Gistel, 1848: [5] [stem: Dicerc-]. Type genus: Dicerca Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1994e)]. Comment: First Reviser found (Dicercini Gistel, 1848 vs Polybothrisini Gistel, 1848) is Bellamy (2008b: 819).
Subtribe
Dicercina
Gistel, 1848
Dicercaeidae Gistel, 1848: [5] [stem: Dicerc-]. Type genus: Dicerca Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1994e)]. Comment: incorrect original stem formation, not in prevailing usage.
Polybothrisidae Gistel, 1848: [5] [stem: Polybothrid-]. Type genus: Polybothris Spinola, 1837. Comment: incorrect original stem formation, not in prevailing usage.
Psiloptérides Lacordaire, 1857: 26 [stem: Psilopter-]. Type genus: Psiloptera Dejean, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1858: 72, as Psilopteritae), generally accepted as in Bellamy (2002: 61, as Psilopterini).
Capnodina Jakobson, 1913: 779 [stem: Capnod-]. Type genus: Capnodis Eschscholtz, 1829.
Subtribe
Haplotrinchina
Holyński, 1993
Haplotrinchina Holyński, 1993: 27 [stem: Haplotrinch-]. Type genus: Haplotrinchus Kerremans, 1903.
Subtribe
Hippomelanina
Holyński, 1993
Hippomelanina Holyński, 1993: 24 [stem: Hippomelan-]. Type genus: Hippomelas Gory and Laporte, 1837.
Subtribe
Pseudoperotina
Tôyama, 1987
Pseudoperotina Tôyama, 1987: 4 [stem: Pseudoperot-]. Type genus: Pseudoperotis Obenberger, 1936.
Tribe
Evidini
Tôyama, 1987
Evidini Tôyama, 1987: 6 [stem: Evid-]. Type genus: Evides Dejean, 1833.
Tribe
Paraleptodemini
Cobos, 1975
Paraleptodemini Cobos, 1975: 88 [stem: Paraleptodem-]. Type genus: Paraleptodema Obenberger, 1936 [syn. of Cinyra Laporte and Gory, 1837].
Subtribe
Euchromatina
Holyński, 1993
Euchromatina Holyński, 1993: 23 [stem: Euchromat-]. Type genus: Euchroma Dejean, 1833.
Subtribe
Euplectaleciina
Holyński, 1993
Euplectaleciina Holyński, 1993: 24 [stem: Euplectaleci-]. Type genus: Euplectalecia Obenberger, 1924.
Subtribe
Hypoprasina
Holyński, 1993
Hypoprasina Holyński, 1993: 23 [stem: Hypopras-]. Type genus: Hypoprasis Fairmaire and Germain, 1864. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Hypoprase-).
Subtribe
Paraleptodemina
Cobos, 1975
Paraleptodemini Cobos, 1975: 88 [stem: Paraleptodem-]. Type genus: Paraleptodema Obenberger, 1936 [syn. of Cinyra Laporte and Gory, 1837].
Cinyrini Cobos, 1979a: 226 [stem: Cinyr-]. Type genus: Cinyra Gory and Laporte, 1837.
Subtribe
Pristipterina
Holyński, 1993
Pristipterina Holyński, 1993: 23 [stem: Pristipter-]. Type genus: Pristiptera Dejean, 1833 [syn. of Pelecopselaphus Solier, 1833].
Tribe
Paratassini
Bílý and Volkovitsh, 1996
Paratassini Bílý and Volkovitsh, 1996: 329 [stem: Paratass-]. Type genus: Paratassa Marseul, 1882.
Tribe
Poecilonotini
Jakobson, 1913
Poecilonotina Jakobson, 1913: 773 [stem: Poecilonot-]. Type genus: Poecilonota Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1996a)].
Subtribe
Poecilonotina
Jakobson, 1913
Poecilonotina Jakobson, 1913: 773 [stem: Poecilonot-]. Type genus: Poecilonota Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1996a)].
Subtribe
Nesotrinchina
Bílý, Kubáň and Volkovitsh, 2009
Nesotrinchina Bílý et al., 2009: 750 [stem: Nesotrinch-]. Type genus: Nesotrinchus Obenberger, 1924.
Tribe
Sphenopterini
Lacordaire, 1857
Sphénoptérides Lacordaire, 1857: 68 [stem: Sphenopter-]. Type genus: Sphenoptera Dejean, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Stein (1868: 62, as Sphenopterini), generally accepted as in Bellamy (2008a: 23, as Sphenopterini).
Tribe
Vadonaxiini
Descarpentries, 1970
Vadonaxiini Descarpentries, 1970: 188 [stem: Vadonaxi-]. Type genus: Vadonaxia Descarpentries, 1970.
Subfamily
Buprestinae
Leach, 1815
Buprestides Leach, 1815: 85 [stem: Buprest-]. Type genus: Buprestis Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1994e)].
Tribe
Actenodini
Gistel, 1848
Actenodeidae Gistel, 1848: [5] [stem: Actenod-]. Type genus: Actenodes Dejean, 1833 [placed on the Official List of Generic Names in Zoology (ICZN 2002b)]. Comment: name previously attributed to Kerremans (1890) in the literature; incorrect original stem formation, not in prevailing usage.
Belionotina Jakobson, 1913: 793 [stem: Belionot-]. Type genus: Belionota Eschscholtz, 1829.
Tribe
Anthaxiini
Gory and Laporte, 1839
Anthaxidae Gory and Laporte, 1839: unnumbered page [stem: Anthaxi-]. Type genus: Anthaxia Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 2002c)]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Bubastini
Obenberger, 1920
Bubastini Obenberger, 1920: 89 [stem: Bubast-]. Type genus: Bubastes Laporte and Gory, 1838.
Tribe
Buprestini
Leach, 1815
Buprestides Leach, 1815: 85 [stem: Buprest-]. Type genus: Buprestis Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1994e)].
Subtribe
Agaeocerina
Nelson, 1982
Agaeocerini Nelson, 1982: 440 [stem: Agaeocer-]. Type genus: Agaeocera Saunders, 1871.
Subtribe
Buprestina
Leach, 1815
Buprestides Leach, 1815: 85 [stem: Buprest-]. Type genus: Buprestis Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1994e)]. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Buprestid-).
Ancylochirina Jakobson, 1913: 787 [stem: Ancylochir-]. Type genus: Ancylochira Eschscholtz, 1829 [subgenus of Buprestis Linnaeus, 1758].
Subtribe
Lamprocheilina
Holyński, 1993
Lamprocheilina Holyński, 1993: 13 [stem: Lamprocheil-]. Type genus: Lamprocheila Saunders, 1871.
Subtribe
Trachykelina
Holyński, 1988
Trachykelina Holyński, 1988: 51, in key [stem: Trachykel-]. Type genus: Trachykele Marseul, 1865.
Tribe
Chrysobothrini
Gory and Laporte, 1836
Chrysobothridae Gory and Laporte, 1836: [1] [stem: Chrysobothr-]. Type genus: Chrysobothris Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1994e)]. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Chrysobothrid-).
Tribe
Coomaniellini
Bílý, 1974
Coomaniellini Bílý, 1974: 41 [stem: Coomaniell-]. Type genus: Coomaniella Bourgoin, 1924.
Tribe
Curidini
Holyński, 1988
Curidina Holyński, 1988: 52, in key [stem: Curid-]. Type genus: Curis Gory and Laporte, 1837 [syn. of Selagis Dejean, 1836 (also see ICZN 2008c)]. Comment: First Reviser (Curidini Holyński, 1988 vs Neocuridini Holyński, 1988) not determined, current usage maintained.
Subtribe
Anilarina
Bílý, 2000
Anilarini Bílý, 2000: 113 [stem: Anilar-]. Type genus: Anilara Saunders, 1868.
Subtribe
Curidina
Holyński, 1988
Curidina Holyński, 1988: 52, in key [stem: Curid-]. Type genus: Curis Gory and Laporte, 1837 [syn. of Selagis Dejean, 1836 (also see ICZN 2008c)].
Subtribe
Neocuridina
Holyński, 1988
Neocuridina Holyński, 1988: 52, in key [stem: Neocurid-]. Type genus: Neocuris Saunders, 1868.
Tribe
Epistomentini
Levey, 1978
Epistomentini Levey, 1978: 155 [stem: Epistoment-]. Type genus: Epistomentis Solier, 1849.
Tribe
Exagistini
Tôyama, 1987
Exagistini Tôyama, 1987: 2 [stem: Exagist-]. Type genus: Exagistus Deyrolle, 1864.
Tribe
Julodimorphini
Kerremans, 1903
Julodimorphites Kerremans, 1903: 16 [stem: Julodimorph-]. Type genus: Julodimorpha Harold, 1869.
Tribe
Kisanthobiini
Richter, 1949
Kisanthobiini Richter, 1949: 215 [stem: Kisanthobi-]. Type genus: Kisanthobia Marseul, 1865.
Tribe
Maoraxiini
Holyński, 1984
Maoraxiini Holyński, 1984: 106 [stem: Maoraxi-]. Type genus: Maoraxia Obenberger, 1937.
Tribe
Melanophilini
Bedel, 1921
Melanophilini Bedel, 1921: 171 [stem: Melanophil-]. Type genus: Melanophila Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1996b)].
Tribe
Melobaseini
Bílý, 2000
Melobasini Bílý, 2000: 113 [stem: Melobase-]. Type genus: Melobasis Gory and Laporte, 1837. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Mendizabaliini
Cobos, 1968
Mendizabaliini Cobos, 1968: 19 [stem: Mendizabali-]. Type genus: Mendizabalia Cobos, 1957.
Tribe
Nascionini
Holyński, 1988
Nascionina Holyński, 1988: 51, in key [stem: Nascion-]. Type genus: Nascio Laporte and Gory, 1838 [placed on the Official List of Generic Names in Zoology (ICZN 2002b)].
Tribe
Phrixiini
Cobos, 1975
Phrixiini Cobos, 1975: 102 [stem: Phrixi-]. Type genus: Phrixia Deyrolle, 1864.
Tribe
Pterobothrini
Volkovitsh, 2001
Pterobothrini Volkovitsh, 2001: 86, 103 [stem: Pterobothr-]. Type genus: Pterobothris Fairmaire and Germain, 1858. Comment: current spelling maintained (Art. 29.3.1.1): incorrect original stem formation in prevailing usage (should be Pterobothrid-).
Tribe
Stigmoderini
Lacordaire, 1857
Stigmodérides Lacordaire, 1857: 52 [stem: Stigmoder-]. Type genus: Stigmodera Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Wallace (1860: 183, as Stigmoderidae), generally accepted as in Bellamy (2008a: 23, as Stigmoderini).
Tribe
Thomassetiini
Bellamy, 1987
Thomassetiini Bellamy, 1987: 223 [stem: Thomasseti-]. Type genus: Thomassetia Théry, 1928.
Subtribe
Philanthaxiina
Holyński, 1988
Philanthaxiina Holyński, 1988: 51, in key [stem: Philanthaxi-]. Type genus: Philanthaxia Deyrolle, 1864.
Subtribe
Thomassetiina
Bellamy, 1987
Thomassetiini Bellamy, 1987: 223 [stem: Thomasseti-]. Type genus: Thomassetia Théry, 1928.
Tribe
Trigonogeniini
Cobos, 1956
Trigonogeniini Cobos, 1956: 72 [stem: Trigonogeni-]. Type genus: Trigonogenium Harold, 1869.
Tribe
Xenorhipidini
Cobos, 1986
Xenorhipini Cobos, 1986: 136 [stem: Xenorhipid-]. Type genus: Xenorhipis J. L. LeConte, 1866. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Trichinorhipidina
Bellamy, 2006
Trichinorhipidina Bellamy, 2006: 139 [stem: Trichinorhipid-]. Type genus: Trichinorhipis Barr, 1948.
Subtribe
Xenorhipidina
Cobos, 1986
Xenorhipini Cobos, 1986: 136 [stem: Xenorhipid-]. Type genus: Xenorhipis J. L. LeConte, 1866. Comment: incorrect original stem formation, not in prevailing usage.
Buprestinae
incertae sedis
†Glaphyropteridae Pongrácz, 1935: 541 [stem: Glaphyropter-]. Type genus: Glaphyroptera Heer, 1852. Comment: the older name Glaphyropteridae Brauer, 1852 is a collective name for most Neuroptera in the present sense and it is not based on a genus name (Ponomarenko 2009 pers. comm.).
Subfamily
Agrilinae
Laporte, 1835
Agrilidae Laporte, 1835b: 165 [stem: Agril-]. Type genus: Agrilus Curtis, 1825. Comment: First Reviser (Agrilinae Laporte, 1835 vs Tracheinae Laporte, 1835) not determined, current usage maintained.
Tribe
Agrilini
Laporte, 1835
Agrilidae Laporte, 1835b: 165 [stem: Agril-]. Type genus: Agrilus Curtis, 1825.
Subtribe
Agrilina
Laporte, 1835
Agrilidae Laporte, 1835b: 165 [stem: Agril-]. Type genus: Agrilus Curtis, 1825.
Subtribe
Amorphosternina
Cobos, 1974
Amorphosternae Cobos, 1974: 69 [stem: Amorphostern-]. Type genus: Amorphosternus Deyrolle, 1864.
Subtribe
Amyiina
Holyński, 1993
Amyiina Holyński, 1993: 14 [stem: Amyi-]. Type genus: Amyia Saunders, 1871.
Subtribe
Rhaeboscelidina
Cobos, 1976
*Rhaeboscelini Böving and Craighead, 1931: 77 [stem: Rhaeboscelid-]. Type genus: Rhaeboscelis Chevrolat, 1838. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Rhaeboscelidi Cobos, 1976: 20 [stem: Rhaeboscelid-]. Type genus: Rhaeboscelis Chevrolat, 1838.
Tribe
Aphanisticini
Jacquelin du Val, 1859
Aphanisticites Jacquelin du Val, 1859: 104 [stem: Aphanistic-]. Type genus: Aphanisticus Latreille, 1810.
Subtribe
Anthaxomorphina
Holyński, 1993
Anthaxomorphina Holyński, 1993: 32 [stem: Anthaxomorph-]. Type genus: Anthaxomorphus Deyrolle, 1864.
Subtribe
Aphanisticina
Jacquelin du Val, 1859
Aphanisticites Jacquelin du Val, 1859: 104 [stem: Aphanistic-]. Type genus: Aphanisticus Latreille, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Acloque (1896: 280, as Aphanisticii), generally accepted as in Hansen (1996: 137, as Aphanisticini).
Subtribe
Cylindromorphina
Portevin, 1931
Cylindromorphini Portevin, 1931: 335 [stem: Cylindromorph-]. Type genus: Cylindromorphus Kiesenwetter, 1857.
Subtribe
Cylindromorphoidina
Cobos, 1979
Cylindromorphoidini Cobos, 1979b: 420, in key [stem: Cylindromorphoid-]. Type genus: Cylindromorphoides Kerremans, 1903.
Subtribe
Germaricina
Cobos, 1979
Germaricini Cobos, 1979b: 420, in key [stem: Germaric-]. Type genus: Germarica Blackburn, 1887.
Tribe
Coraebini
Bedel, 1921
Coraebini Bedel, 1921: 170 [stem: Coraeb-]. Type genus: Coraebus Gory and Laporte, 1839.
Subtribe
Amorphosomatina
Majer, 2000
Amorphosomina Majer, 2000: 210 [stem: Amorphosomat-]. Type genus: Amorphosoma Laporte, 1835. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Cisseina
Majer, 2000
Cisseina Majer, 2000: 203 [stem: Cisse-]. Type genus: Cisseis Gory and Laporte, 1839 [syn. of Diphucrania Dejean, 1833 (see ICZN 2008c)].
Subtribe
Clematina
Majer, 2000
Clemina Majer, 2000: 215 [stem: Clemat-]. Type genus: Clema Semenov, 1900. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Coraebina
Bedel, 1921
Coraebini Bedel, 1921: 170 [stem: Coraeb-]. Type genus: Coraebus Gory and Laporte, 1839.
Subtribe
Dismorphina
Cobos, 1990
Dismorphina Cobos, 1990: 542 [stem: Dismorph-]. Type genus: Dismorpha Gistel, 1848.
Subtribe
Ethoniina
Majer, 2000
Ethoniina Majer, 2000: 201 [stem: Ethoni-]. Type genus: Ethonion Kubáň, 2000.
Subtribe
Geraliina
Cobos, 1988
Geraliina Cobos, 1988: 10 [stem: Gerali-]. Type genus: Geralius Harold, 1869.
Subtribe
Meliboeina
Majer, 2000
Meliboeina Majer, 2000: 213 [stem: Meliboe-]. Type genus: Meliboeus Deyrolle, 1864.
Subtribe
Synechocerina
Majer, 2000
Synechocerina Majer, 2000: 214 [stem: Synechocer-]. Type genus: Synechocera Deyrolle, 1864.
Subtribe
Toxoscelina
Majer, 2000
Toxoscelina Majer, 2000: 207 [stem: Toxoscel-]. Type genus: Toxoscelus Deyrolle, 1864.
Tribe
Tracheini
Laporte, 1835
Trachisidae Laporte, 1835b: 166 [stem: Trache-]. Type genus: Trachys Fabricius, 1801 [placed on the Official List of Generic Names in Zoology (ICZN 2009a)]. Comment: correct stem ruled to be Trache- and Tracheidae Laporte, 1835 placed on the Official List of Family-Group Names in Zoology (ICZN 2009a).
Subtribe
Brachina
LeConte, 1861
Braches J. L. LeConte, 1861: 156 [stem: Brach-]. Type genus: Brachys Dejean, 1833. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Brache-).
Brachyini Cobos, 1979b: 417, in key [stem: Brach-]. Type genus: Brachys Dejean, 1833. Comment: family-group name proposed as a new taxon, without reference to Braches J. L. LeConte, 1861; incorrect original stem formation, not in prevailing usage.
Subtribe
Leiopleurina
Holyński, 1993
Leiopleurina Holyński, 1993: 32 [stem: Leiopleur-]. Type genus: Leiopleura Deyrolle, 1864.
Subtribe
Pachyschelina
Böving and Craighead, 1931
Pachyschelinae Böving and Craighead, 1931: 49, in key [stem: Pachyschel-]. Type genus: Pachyschelus Solier, 1833.
Subtribe
Tracheina
Laporte, 1835
Trachisidae Laporte, 1835b: 166 [stem: Trache-]. Type genus: Trachys Fabricius, 1801 [placed on the Official List of Generic Names in Zoology (ICZN 2009a)]. Comment: published before 6 May 1835; correct stem ruled to be Trache- and Tracheidae Laporte, 1835 placed on the Official List of Family-Group Names in Zoology (ICZN 2009a); Trachisidae Laporte, 1835 deemed to be an incorrect original spelling and placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology along with subsequent family-group names based on Trachys Fabricius (ICZN 2009a); the correct spelling Tracheidae was first used by Gistel (1848: [5]); this family-group name was also used in the same year by Solier (1835c [before 4 May]: c, as Trachysides).
Phytoteradae Gistel, 1856a: 366 [stem: Phytoter-]. Type genus: Phytotera Gistel, 1856 [syn. of Trachys Fabricius, 1801]. Comment: incorrect original stem formation, not in prevailing usage.
†Subfamily
Parathyreinae
Alexeev, 1994
Parathyreinae Alexeev, 1994: 10 [stem: Parathyre-]. Type genus: Parathyrea Alexeev, 1994.
Superfamily
Byrrhoidea
Latreille, 1804
Byrrhii Latreille, 1804c: 146 [stem: Byrrh-]. Type genus: Byrrhus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Family
Byrrhidae
Latreille, 1804
Byrrhii Latreille, 1804c: 146 [stem: Byrrh-]. Type genus: Byrrhus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subfamily
Byrrhinae
Latreille, 1804
Byrrhii Latreille, 1804c: 146 [stem: Byrrh-]. Type genus: Byrrhus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Byrrhini
Latreille, 1804
Byrrhii Latreille, 1804c: 146 [stem: Byrrh-]. Type genus: Byrrhus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: published 7 March 1804; this family-group name was also used in the same year by Latreille (1804a [between 19 August and 17 September]: 190, as Byrrhii).
*Tylicini Johnson, 1991: 160 [stem: Tylic-]. Type genus: Tylicus Casey, 1912 [syn. of Arctobyrrhus Munster, 1902]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Exomellini
Casey, 1914
Exomellini Casey, 1914a: 378 [stem: Exomell-]. Type genus: Exomella Casey, 1914.
Tribe
Morychini
El Moursy, 1961
Morychini El Moursy, 1961: 11 [stem: Morych-]. Type genus: Morychus Erichson, 1847.
Tribe
Pedilophorini
Casey, 1912
Pedilophorini Casey, 1912: 4 [stem: Pedilophor-]. Type genus: Pedilophorus Steffahny, 1843.
Tribe
Simplocariini
Mulsant and Rey, 1869
Simplocariates Mulsant and Rey, 1869: 151 [stem: Simplocari-]. Type genus: Simplocaria Stephens, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1985c)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Casey (1912: 14, as Simplocariini), generally accepted as in P. J. Johnson (2002b: 115, as Simplocariini).
Lioonini Leng, 1920: 193 [stem: Lio-]. Type genus: Lioon Casey, 1912. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Syncalyptinae
Mulsant and Rey, 1869
Syncalyptaires Mulsant and Rey, 1869: 31 [stem: Syncalypt-]. Type genus: Syncalypta Dillwyn, 1829 [syn. of Chaetophora Kirby and Spence, 1817].
Tribe
Microchaetini
Paulus, 1973
Microchaetini Paulus, 1973: 353, in key [stem: Microchaet-]. Type genus: Microchaetes Hope, 1834.
Tribe
Syncalyptini
Mulsant and Rey, 1869
Syncalyptaires Mulsant and Rey, 1869: 31 [stem: Syncalypt-]. Type genus: Syncalypta Dillwyn, 1829 [syn. of Chaetophora Kirby and Spence, 1817]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Portevin (1931: 290, as Syncalyptini), generally accepted as in Hansen (1996: 138, as Syncalyptinae).
Subfamily
Amphicyrtinae
LeConte, 1861
Amphicyrtini J. L. LeConte, 1861: 111 [stem: Amphicyrt-]. Type genus: Amphicyrta Erichson, 1843.
Family
Elmidae
Curtis, 1830
Elmidae Curtis, 1830: pl. 294 [stem: Elm-]. Type genus: Elmis Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1995d)]. Comment: name placed on the Official List of Family-Group Names in Zoology and correct stem ruled to be Elm- (ICZN 1995d).
Subfamily
Larainae
LeConte, 1861
Larini J. L. LeConte, 1861: 116 [stem: Lara-]. Type genus: Lara J. L. LeConte, 1852 [placed on the Official List of Generic Names in Zoology (ICZN 1988g)].
Tribe
Laraini
LeConte, 1861
Larini J. L. LeConte, 1861: 116 [stem: Lara-]. Type genus: Lara J. L. LeConte, 1852 [placed on the Official List of Generic Names in Zoology (ICZN 1988g)]. Comment: Laraini J. L. LeConte, 1861 placed on the Official List of Family-Group Names in Zoology and correct stem ruled to be Lara- to avoid homonymy with Laridae Rafinesque-Schmaltz, 1815 (type genus Larus Linnaeus, 1758) in Aves (ICZN 1988g); Larini J. L. LeConte, 1861 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1988g).
Tribe
Potamophilini
Mulsant and Rey, 1872
*Potamophiles Motschulsky, 1849: 54 [stem: Potamophil-]. Type genus: Potamophilus Germar, 1811. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Potamophilaires Mulsant and Rey, 1872b: 11 [stem: Potamophil-]. Type genus: Potamophilus Germar, 1811. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Ganglbauer (1904: 99, as Potamophilini), generally accepted as in Jäch et al. (2006: 432, as Potamophilini).
Subfamily
Elminae
Curtis, 1830
Elmidae Curtis, 1830: pl. 294 [stem: Elm-]. Type genus: Elmis Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1995d)]. Comment: placed on the Official List of Family-Group Names in Zoology and correct stem ruled to be Elm- (ICZN 1995d).
Tribe
Ancyronychini
Ganglbauer, 1904
Ancyronychini Ganglbauer, 1904: 108 [stem: Ancyronych-]. Type genus: Ancyronyx Erichson, 1847.
Tribe
Elmini
Curtis, 1830
Elmidae Curtis, 1830: pl. 294 [stem: Elm-]. Type genus: Elmis Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1995d)]. Comment: Elmidae Curtis, 1830 placed on the Official List of Family-Group Names in Zoology and correct stem ruled to be Elm- (ICZN 1995d).
Subtribe
Elmina
Curtis, 1830
*Limniidae Stephens, 1828: 104 [stem: Limni-]. Type genus: Limnius Illiger, 1802. Comment: unavailable family-group name, not based on genus used as valid at the time (see Jäch 1994).
*Limniidae Stephens, 1829a: 5 [stem: Limni-]. Type genus: Limnius Illiger, 1802. Comment: unavailable family-group name, not based on genus used as valid at the time.
Elmidae Curtis, 1830: pl. 294 [stem: Elm-]. Type genus: Elmis Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1995d)]. Comment: Elmidae Curtis, 1830 placed on the Official List of Family-Group Names in Zoology and correct stem ruled to be Elm- (ICZN 1995d).
Limniidae Hope, 1838a: 153 [stem: Limni-]. Type genus: Limnius Illiger, 1802. Comment: family-group name previously attributed to C. G. Thomson (1859) in the literature.
Subtribe
Stenelmina
Mulsant and Rey, 1872
Stenelmisates Mulsant and Rey, 1872a: 49 [stem: Stenelm-]. Type genus: Stenelmis Dufour, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Bollow (1941: 3, as Stenelmini), generally accepted as in Jäch et al. (2006: 437, as Stenelmina); incorrect original stem formation, not in prevailing usage.
Tribe
Macronychini
Gistel, 1848
Macronychidae Gistel, 1848: [1] [stem: Macronych-]. Type genus: Macronychus P. W. J. Müller, 1806. Comment: family-group name previously attributed to Mulsant and Rey (1872) in the literature.
Family
Dryopidae
Billberg, 1820 (1817)
Parnidea Leach, 1817: 88 [stem: Parn-]. Type genus: Parnus Fabricius, 1792 [syn. of Dryops A. G. Olivier, 1791]. Comment: younger name Dryopidae Billberg, 1820 conserved (Art. 40.2) (see Lawrence and Newton 1995).
Dryopides Billberg, 1820a: 38 [stem: Dryop-]. Type genus: Dryops A. G. Olivier, 1791. Comment: younger name conserved over Parnidae Leach, 1817 (Art. 40.2) (see Lawrence and Newton 1995).
Pelonominae Böving and Craighead, 1931: 45, in key [stem: Pelonom-]. Type genus: Pelonomus Erichson, 1847.
Chiloeidae Dajoz, 1973: 179 [stem: Chiloe-]. Type genus: Chiloea Dajoz, 1973 [syn. of Sosteamorphus Hinton, 1936].
Family
Lutrochidae
Kasap and Crowson, 1975
*Lutrochidae Hinton, 1971: 297 [stem: Lutroch-]. Type genus: Lutrochus Erichson, 1847. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Lutrochidae Kasap and Crowson, 1975: 442 [stem: Lutroch-]. Type genus: Lutrochus Erichson, 1847.
Family
Limnichidae
Erichson, 1846
Limnichini Erichson, 1846: 465 [stem: Limnich-]. Type genus: Limnichus Dejean, 1821.
Subfamily
Hyphalinae
Britton, 1971
Hyphalinae Britton, 1971: 88 [stem: Hyphal-]. Type genus: Hyphalus Britton, 1971.
Subfamily
Limnichinae
Erichson, 1846
Limnichini Erichson, 1846: 465 [stem: Limnich-]. Type genus: Limnichus Dejean, 1821.
Tribe
Limnichini
Erichson, 1846
Limnichini Erichson, 1846: 465 [stem: Limnich-]. Type genus: Limnichus Dejean, 1821.
Botriophorates Mulsant and Rey, 1869: 160 [stem: Bothriophor-]. Type genus: Bothriophorus Mulsant and Rey, 1852 [as Botriophorus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: family-group name also spelled Botriaphorates in the original publication (p. 173); original vernacular name available (Art. 11.7.2): first used in latinized form by Ganglbauer (1904: 51, as Bothriophorini), generally accepted as in Shepard (2002: 125, as Bothriophorini); incorrect original stem formation, not in prevailing usage.
Tribe
Wooldridgeini
Spangler, 1999
Wooldridgeini Spangler, 1999: 181 [stem: Wooldridge-]. Type genus: Wooldridgeus Spangler, 1998.
Subfamily
Cephalobyrrhinae
Champion, 1925
Cephalobyrrhinae Champion, 1925: 174 [stem: Cephalobyrrh-]. Type genus: Cephalobyrrhus Pic, 1923.
Subfamily
Thaumastodinae
Champion, 1924
Thaumastodinae Champion, 1924: 25 [stem: Thaumastod-]. Type genus: Thaumastodus Champion, 1924 [syn. of Pseudeucinetus Heller, 1921].
Family
Heteroceridae
MacLeay, 1825
Heteroceridae W. S. MacLeay, 1825: 34 [stem: Heterocer-]. Type genus: Heterocerus Fabricius, 1792.
Subfamily
Elythomerinae
Pacheco, 1964
Elythomerini Pacheco, 1964: 120 [stem: Elythomer-]. Type genus: Elythomerus C. O. Waterhouse, 1874.
Subfamily
Heterocerinae
MacLeay, 1825
Heteroceridae W. S. MacLeay, 1825: 34 [stem: Heterocer-]. Type genus: Heterocerus Fabricius, 1792.
Tribe
Augylini
Pacheco, 1964
Augyliini Pacheco, 1964: 19 [stem: Augyl-]. Type genus: Augyles Schiødte, 1866. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Heterocerini
MacLeay, 1825
Heteroceridae W. S. MacLeay, 1825: 34 [stem: Heterocer-]. Type genus: Heterocerus Fabricius, 1792.
Tribe
Micilini
Pacheco, 1964
Micilini Pacheco, 1964: 32 [stem: Micil-]. Type genus: Micilus Mulsant and Rey, 1872.
Tribe
Tropicini
Pacheco, 1964
Tropicini Pacheco, 1964: 101 [stem: Tropic-]. Type genus: Tropicus Pacheco, 1964.
Family
Psephenidae
Lacordaire, 1854
Pséphénides Lacordaire, 1854b: 497 [stem: Psephen-]. Type genus: Psephenus Haldeman, 1853.
Subfamily
Afroeubriinae
Lee, Satô, Shepard and Jäch, 2007
Afroeubriinae Lee, Satô, Shepard and Jäch, 2007: 532 [stem: Afroeubri-]. Type genus: Afroeubria Villiers, 1961.
Subfamily
Eubriinae
Lacordaire, 1857
Eubriades Lacordaire, 1857: 283 [stem: Eubri-]. Type genus: Eubria Germar, 1818. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 180, as Eubriini), generally accepted as in Hansen (1996: 139, as Eubriinae).
Subfamily
Eubrianacinae
Jakobson, 1913
Placonychini G. H. Horn, 1880a: 110 [stem: Placonych-]. Type genus: Placonycha G. H. Horn, 1880 [syn. of Eubrianax Kiesenwetter, 1874]. Comment: usage of the younger name Eubrianacinae Jakobson, 1913 conserved over this name (Art. 40.2) (see Lawrence and Newton 1995: 846).
Eubrianacini Jakobson, 1913: 723 [stem: Eubrianac-]. Type genus: Eubrianax Kiesenwetter, 1874. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Eubrianact-); use of this name conserved over Placonychinae G. H. Horn, 1880 (Art. 40.2) (see Lawrence and Newton 1995: 846).
Subfamily
Psephenoidinae
Bollow, 1938
Psephenoidini Bollow, 1938: 156 [stem: Psephenoid-]. Type genus: Psephenoides Gahan, 1914.
Subfamily
Psepheninae
Lacordaire, 1854
Eurypalpini J. L. LeConte, 1852a: 41 [stem: Eurypalp-]. Type genus: Eurypalpus J. L. LeConte, 1852 [preoccupied genus name, not Eurypalpus Macquart, 1835 [Diptera]; syn. of Psephenus Haldeman, 1853]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Pséphénides Lacordaire, 1854b: 497 [stem: Psephen-]. Type genus: Psephenus Haldeman, 1853. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Imhoff (1856: xv, as Psephenidae), generally accepted as in Hansen (1996: 139, as Psephenidae).
Family
Cneoglossidae
Champion, 1897
Cneoglossini Champion, 1897: 594 [stem: Cneogloss-]. Type genus: Cneoglossa Guérin-Méneville, 1843.
Family
Ptilodactylidae
Laporte, 1836
Ptilodactylidae Laporte, 1836: 21 [stem: Ptilodactyl-]. Type genus: Ptilodactyla Illiger, 1807.
Subfamily
Anchytarsinae
Champion, 1897
Coloboderides Erichson, 1847a: 174 [stem: Coloboder-]. Type genus: Colobodera Klug, 1838 [syn. of Daemon Laporte, 1836].
Anchytarsini Champion, 1897: 593 [stem: Anchytars-]. Type genus: Anchytarsus Guérin-Méneville, 1843. Comment: Coloboderinae Erichson, 1847 is the oldest name for this tribe, however since Coloboderinae has not been used as valid after 1899 to our knowledge, we believe that usage of Anchytarsinae Champion, 1897 should be conserved and an application be submitted to the Commission to suppress the older name.
*Epilichadinae Lawrence and Stribling, 1992: 19 [stem: Epilichad-]. Type genus: Epilichas A. White, 1859. Comment: although Ivie (2002: 136) treated this name as available we do not believe that requirements of Art. 13.1 were met and we therefore treat this name here as unavailable.
Subfamily
Cladotominae
Pic, 1914
Cladotomini Pic, 1914: 45 [stem: Cladotom-]. Type genus: Cladotoma Westwood, 1836.
Subfamily
Aploglossinae
Champion, 1897
Aploglossinae Champion, 1897: 623 [stem: Aplogloss-]. Type genus: Aploglossa Guérin-Méneville, 1849.
Subfamily
Araeopidiinae
Lawrence, 1991
Araeopidiinae Lawrence, 1991: 250, in key [stem: Araeopidi-]. Type genus: Araeopidius Cockerell, 1906.
Subfamily
Ptilodactylinae
Laporte, 1836
Ptilodactylidae Laporte, 1836: 21 [stem: Ptilodactyl-]. Type genus: Ptilodactyla Illiger, 1807.
Family
Podabrocephalidae
Pic, 1930
Podabrocephalidae Pic, 1930: 314 [stem: Podabrocephal-]. Type genus: Podabrocephalus Pic, 1913.
Family
Chelonariidae
Blanchard, 1845
Chelonariites Blanchard, 1845b: 70 [stem: Chelonari-]. Type genus: Chelonarium Fabricius, 1801. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Imhoff (1856: 140, as Chelonarii), generally accepted as in Satô (2006: 454, as Chelonariidae).
Family
Eulichadidae
Crowson, 1973
*Lichadiden Kolbe, 1908: 249 [stem: Lichad-]. Type genus: Lichas Westwood, 1853 [preoccupied genus name, not Lichas Dalman, 1827 [Trilobita]; syn. of Eulichas Jakobson, 1913]. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Lichadidae Forbes, 1926: 102 [stem: Lichad-]. Type genus: Lichas Westwood, 1853 [preoccupied genus name, not Lichas Dalman, 1827 [Trilobita]; syn. of Eulichas Jakobson, 1913]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Eulichadidae Crowson, 1973b: 237 [stem: Eulichad-]. Type genus: Eulichas Jakobson, 1913.
Family
Callirhipidae
Emden, 1924
Zenoini J. L. LeConte, 1866a: 50 [stem: Zeno-]. Type genus: Zenoa Say, 1835. Comment: this name is older than Callirhipidae Emden, 1924 however we recommend that an application be submitted to conserve the younger name.
Callirhipini Emden, 1924: 87 [stem: Callirhip-]. Type genus: Callirhipis Latreille, 1829. Comment: the oldest name for this family is Zenoidae LeConte, 1866, however, as pointed out by Lawrence and Newton (1995: 849) an application to the Commission is needed in order to preserve the broadly accepted younger name Callirhipidae Emden, 1924; current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Callirhipid-).
Superfamily
Elateroidea
Leach, 1815
Elaterides Leach, 1815: 85 [stem: Elater-]. Type genus: Elater Linnaeus, 1758. Comment: the oldest available name for this superfamily is Cebrionoidea Latreille, 1802, however Elateroidea Leach, 1815 is maintained here pending resolution of an application to the ICZN being prepared by P. J. Johnson (see Lawrence and Newton 1995: 849; P. J. Johnson pers. comm. 2009).
Family
Rhinorhipidae
Lawrence, 1988
Rhinorhipidae Lawrence, 1988: 3 [stem: Rhinorhip-]. Type genus: Rhinorhipus Lawrence, 1988.
Family
Artematopodidae
Lacordaire, 1857
Artématopides Lacordaire, 1857: 260 [stem: Artematopod-]. Type genus: Artematopus Perty, 1832.
Subfamily
Electribiinae
Crowson, 1975
Electropogonini Crowson, 1975: 77 [stem: Electribi-]. Type genus: Electribius Crowson, 1973. Comment: incorrect original stem formation, not in prevailing usage; Electropogonini (based on Electropogon) in Crowson’s work has been treated as a lapsus calami for Electribiidae based on Electribius (see Lawrence and Newton 1995: 850).
Subfamily
Allopogoniinae
Crowson, 1973
Allopogonini Crowson, 1973b: 231, in key [stem: Allopogoni-]. Type genus: Allopogonia Cockerell, 1906. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Artematopodinae
Lacordaire, 1857
Artématopides Lacordaire, 1857: 260 [stem: Artematopod-]. Type genus: Artematopus Perty, 1832.
Tribe
Artematopodini
Lacordaire, 1857
Artématopides Lacordaire, 1857: 260 [stem: Artematopod-]. Type genus: Artematopus Perty, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1897: 586, as Artematopinae [incorrect stem formation]), generally accepted as in Lawrence and Newton (1995: 849, as Artematopodidae); incorrect original stem formation, not in prevailing usage.
Tribe
Ctesibiini
Crowson, 1973
Ctesibiinae Crowson, 1973b: 228, in key [stem: Ctesibi-]. Type genus: Ctesibius Champion, 1897.
Tribe
Macropogonini
LeConte, 1861
Macropogonini J. L. LeConte, 1861: 178 [stem: Macropogon-]. Type genus: Macropogon Motschulsky, 1845.
Eurypogonidae Böving and Craighead, 1931: 45, in key [stem: Eurypogon-]. Type genus: Eurypogon Motschulsky, 1859.
Family
Brachypsectridae
LeConte and Horn, 1883
Brachypsectrini J. L. LeConte and G. H. Horn, 1883: 170 [stem: Brachypsectr-]. Type genus: Brachypsectra J. L. LeConte, 1874.
Family
Cerophytidae
Latreille, 1834
Cerophytides Latreille, 1834: 133 [stem: Cerophyt-]. Type genus: Cerophytum Latreille, 1809.
Family
Eucnemidae
Eschscholtz, 1829
Eucnemides Eschscholtz, 1829a: 10 [stem: Eucnem-]. Type genus: Eucnemis Ahrens, 1812. Comment: the name Melasidae Fleming, 1821 has priority over this name, however Muona and Alaruikka (2007: 32) mentioned that an application has been sent to the Commission to conserve usage of Eucnemidae over the older name Melasidae for reasons of stability, we follow current usage until the case is resolved (also see Appendix 6).
Subfamily
Perothopinae
Lacordaire, 1857
Pérothopides Lacordaire, 1857: 128 [stem: Perothop-]. Type genus: Perothops Eschscholtz, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 162, as Perothopini), generally accepted as in Lawrence and Newton (1995: 851, as Perothopinae).
Subfamily
Phyllocerinae
Reitter, 1905
Phylloceridae Reitter, 1905: 4 [stem: Phyllocer-]. Type genus: Phyllocerus Lepeletier and Audinet-Serville, 1825.
Tribe
Anelastini
Reitter, 1911
Anelastini Reitter, 1911: 202 [stem: Anelast-]. Type genus: Anelastes Kirby, 1819.
Tribe
Phyllocerini
Reitter, 1905
Phylloceridae Reitter, 1905: 4 [stem: Phyllocer-]. Type genus: Phyllocerus Lepeletier and Audinet-Serville, 1825.
Subfamily
Pseudomeninae
Muona, 1993
Pseudomeninae Muona, 1993: 41 [stem: Pseudomen-]. Type genus: Pseudomenes Fleutiaux, 1902. Comment: precedence (Pseudomeninae Muona, 1993 vs Schizophilinae Muona, 1993) given to taxon originally proposed at the higher rank (Art. 24.1).
Tribe
Pseudomenini
Muona, 1993
* Pseudomeninae Muona, 1991a: 167 [stem: Pseudomen-]. Type genus: Pseudomenes Fleutiaux, 1902. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Pseudomeninae Muona, 1993: 41 [stem: Pseudomen-]. Type genus: Pseudomenes Fleutiaux, 1902.
Tribe
Schizophilini
Muona, 1993
*Schizophilini Muona, 1991a: 167 [stem: Schizophil-]. Type genus: Schizophilus Bonvouloir, 1871. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Schizophilini Muona, 1993: 42 [stem: Schizophil-]. Type genus: Schizophilus Bonvouloir, 1871.
Subfamily
Palaeoxeninae
Muona, 1993
Palaeoxeninae Muona, 1993: 42 [stem: Palaeoxen-]. Type genus: Palaeoxenus G. H. Horn, 1891.
Subfamily
Phlegoninae
Muona, 1993
Phlegoninae Muona, 1993: 42 [stem: Phlegon-]. Type genus: Phlegon Laporte, 1840. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Phlegont-).
Subfamily
Anischiinae
Fleutiaux, 1936
Anischinae Fleutiaux, 1936: 292 [stem: Anischi-]. Type genus: Anischia Fleutiaux, 1896. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Melasinae
Fleming, 1821
Melasidae Fleming, 1821: 49 [stem: Melas-]. Type genus: Melasis A. G. Olivier, 1790.
Tribe
Calyptocerini
Muona, 1993
*Calyptocerini Muona, 1991a: 167 [stem: Calyptocer-]. Type genus: Calyptocerus Guérin-Méneville, 1843. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Calyptocerini Muona, 1993: 43 [stem: Calyptocer-]. Type genus: Calyptocerus Guérin-Méneville, 1843.
Tribe
Ceballosmelasini
Muona, 1993
*Ceballosmelasini Muona, 1991a: 167 [stem: Ceballosmelas-]. Type genus: Ceballosmelasis Cobos, 1964. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Ceballosmelasini Muona, 1993: 42 [stem: Ceballosmelas-]. Type genus: Ceballosmelasis Cobos, 1964.
Tribe
Dirhagini
Reitter, 1911
Dirrhagini Reitter, 1911: 202 [stem: Dirhag-]. Type genus: Dirhagus Latreille, 1834 [as Dirrhagus, incorrect subsequent spelling of type genus name, not in prevailing usage; syn. of Microrhagus Dejean, 1833]. Comment: incorrect original stem formation, not in prevailing usage.
Microrhaginae Fleutiaux, 1919: 112 [stem: Microrhag-]. Type genus: Microrhagus Dejean, 1833.
Arhipini Cobos, 1965: 396 [stem: Arrhipid-]. Type genus: Arrhipis Bonvouloir, 1871 [as Arhipis, incorrect subsequent spelling type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Epiphanini
Muona, 1993
*Epiphanini Muona, 1991a: 167 [stem: Epiphan-]. Type genus: Epiphanis Eschscholtz, 1829. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Epiphanini Muona, 1993: 45 [stem: Epiphan-]. Type genus: Epiphanis Eschscholtz, 1829.
Tribe
Hylocharini
Jacquelin du Val, 1859
Hylocharites Jacquelin du Val, 1859: 119 [stem: Hylochar-]. Type genus: Hylochares Latreille, 1834. Comment: original vernacular name available (Art. 11.7.2): generally accepted as in Muona (2007: 84, as Hylocharini); family-group names with the incorrectly formed stem Hylochar- have been used in Aves (type genus Hylocharis Boie, 1831), the correct stem for the bird family-group name is Hylocharit- (see Bock 1994: 143).
Hylocharini Cobos, 1965: 369 [stem: Hylochar-]. Type genus: Hylochares Latreille, 1834. Comment: proposed as new without reference to Hylocharini Jacquelin du Val, 1859.
Tribe
Melasini
Fleming, 1821
Melasidae Fleming, 1821: 49 [stem: Melas-]. Type genus: Melasis A. G. Olivier, 1790.
Subtribe
Compsocnemina
Muona, 1993
Compsocnemina Muona, 1993: 43 [stem: Compsocnem-]. Type genus: Compsocnemis Bonvouloir, 1871. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Compsocnemid-).
Subtribe
Melasina
Fleming, 1821
Melasidae Fleming, 1821: 49 [stem: Melas-]. Type genus: Melasis A. G. Olivier, 1790.
Tribe
Neocharini
Muona, 1993
*Neocharini Muona, 1991a: 167 [stem: Neochar-]. Type genus: Neocharis Sharp, 1887. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Neocharini Muona, 1993: 44 [stem: Neochar-]. Type genus: Neocharis Sharp, 1887. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Neocharit-).
Tribe
Xylobiini
Reitter, 1911
Xylobiini Reitter, 1911: 203 [stem: Xylobi-]. Type genus: Xylobius Latreille, 1834 [syn. of Xylophilus Mannerheim, 1823].
Subfamily
Eucneminae
Eschscholtz, 1829
Eucnemides Eschscholtz, 1829a: 10 [stem: Eucnem-]. Type genus: Eucnemis Ahrens, 1812.
Tribe
Dendrocharini
Fleutiaux, 1920
Dendrocharini Fleutiaux, 1920: 100, in key [stem: Dendrochar-]. Type genus: Dendrocharis Guérin-Méneville, 1843. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Dendrocharit-).
Tribe
Dyscharachthini
Muona, 1993
*Dyscharachthini Muona, 1991a: 167 [stem: Dyscharachth-]. Type genus: Dyscharachthis Blackburn, 1900. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Dyscharachthini Muona, 1993: 49 [stem: Dyscharachth-]. Type genus: Dyscharachthis Blackburn, 1900. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Dyscharachthent-).
Tribe
Entomosatopini
Muona, 1993
*Entomosatopini Muona, 1991a: 167 [stem: Entomosatop-]. Type genus: Entomosatopus Bonvouloir, 1871. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Entomosatopini Muona, 1993: 48 [stem: Entomosatop-]. Type genus: Entomosatopus Bonvouloir, 1871.
Tribe
Eucnemini
Eschscholtz, 1829
Eucnemides Eschscholtz, 1829a: 10 [stem: Eucnem-]. Type genus: Eucnemis Ahrens, 1812. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Eucnemid-).
Gastraulaci Fleutiaux, 1902: 648 [stem: Gastraulac-]. Type genus: Gastraulacus Guérin-Méneville, 1843.
Tribe
Galbitini
Muona, 1991
*Galbites Blanchard, 1845b: 71 [stem: Galb-]. Type genus: Galba Latreille, 1829 [preoccupied genus name, not Galba Schrank, 1803 [Mollusca]; syn. of Galbites Fleutiaux, 1918]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if found to be availabe in the future then permanently invalid (Art. 39): based on preoccupied type genus.
Pterotarsini Cobos, 1965: 294 [stem: Pterotars-]. Type genus: Pterotarsus sensu Guérin-Méneville, 1838 [syn. of Galbites Fleutiaux, 1918]. Comment: proposed as new without reference to Pterotarsini Fleutiaux, 1902 in Elateridae; based on misidentified type genus; name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).
*Galbitini Muona, 1991a: 167 [stem: Galbit-]. Type genus: Galbites Fleutiaux, 1918. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Galbitini Muona, 1991b: 19 [stem: Galbit-]. Type genus: Galbites Fleutiaux, 1918.
Tribe
Mesogenini
Muona, 1993
*Mesogenini Muona, 1991a: 167 [stem: Mesogen-]. Type genus: Mesogenus Bonvouloir, 1871. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Mesogenini Muona, 1993: 49 [stem: Mesogen-]. Type genus: Mesogenus Bonvouloir, 1871.
Tribe
Muonajini
Özdikmen, 2008
*Yangini Muona, 1991a: 167 [stem: Yang-]. Type genus: Yanga Muona, 1993 [preoccupied genus name, not Yanga Distant, 1904 [Hemiptera]; syn. of Muonaja Özdikmen, 2008]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Yangini Muona, 1993: 48 [stem: Yang-]. Type genus: Yanga Muona, 1993 [preoccupied genus name, not Yanga Distant, 1904 [Hemiptera]; syn. of Muonaja Özdikmen, 2008]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Muonajini Özdikmen, 2008: 675 [stem: Muonaj-]. Type genus: Muonaja Özdikmen, 2008. Comment: replacement name for Yangini Muona, 1993 because of the homonymy of the type genus.
Tribe
Perrotiini
Muona, 1993
*Perrotini Muona, 1991a: 167 [stem: Perroti-]. Type genus: Perrotius Fleutiaux, 1938. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Perrotini Muona, 1993: 47 [stem: Perroti-]. Type genus: Perrotius Fleutiaux, 1938. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Phaenocerini
Muona, 1993
* Phaenocerini Muona, 1991a: 167 [stem: Phaenocer-]. Type genus: Phaenocerus Bonvouloir, 1871. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Phaenocerini Muona, 1993: 47 [stem: Phaenocer-]. Type genus: Phaenocerus Bonvouloir, 1871.
Tribe
Proutianini
Muona, 1993
*Proutianini Muona, 1991a: 167 [stem: Proutian-]. Type genus: Proutianus Muona, 1993. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Proutianini Muona, 1993: 47 [stem: Proutian-]. Type genus: Proutianus Muona, 1993.
Subfamily
Macraulacinae
Fleutiaux, 1923
Macraulacinae Fleutiaux, 1923: 304 [stem: Micraulac-]. Type genus: Macraulacus Bonvouloir, 1872.
Tribe
Anelastidini
Muona, 1993
Anelastidini Muona, 1993: 52 [stem: Anelastid-]. Type genus: Anelastidius Jacquelin du Val, 1863. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Anelastidi-).
Tribe
Echthrogasterini
Cobos, 1965
Echthrogasterini Cobos, 1965: 369 [stem: Echthrogaster-]. Type genus: Echthrogaster Blackburn, 1900.
Tribe
Euryptychini
Mamaev, 1976
Euryptychini Mamaev, 1976: 154 [stem: Euryptych-]. Type genus: Euryptychus J. L. LeConte, 1852.
Tribe
Jenibuntorini
Muona, 1993
*Jenibuntorini Muona, 1991a: 167 [stem: Jenibuntor-]. Type genus: Jenibuntor Muona, 1993. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Jenibuntorini Muona, 1993: 51 [stem: Jenibuntor-]. Type genus: Jenibuntor Muona, 1993.
Tribe
Macraulacini
Fleutiaux, 1923
Macraulacinae Fleutiaux, 1923: 304 [stem: Macraulac-]. Type genus: Macraulacus Bonvouloir, 1872.
Fornaxini Cobos, 1965: 294 [stem: Fornac-]. Type genus: Fornax Laporte, 1835. Comment: incorrect original stem formation, not in prevailing usage.
Dromaeolini Leiler, 1976: 48 [stem: Dromaeol-]. Type genus: Dromaeolus Kiesenwetter, 1858.
Tribe
Nematodini
Leiler, 1976
Nematodini Leiler, 1976: 48 [stem: Nematod-]. Type genus: Nematodes Berthold, 1827.
Tribe
Oisocerini
Muona, 1993
Oisocerini Muona, 1993: 51 [stem: Oisocer-]. Type genus: Oisocerus Bonvouloir, 1868.
Tribe
Orodotini
Muona, 1993
Cryptostomidae Laporte, 1835b: 181 [stem: Cryptostomat-]. Type genus: Cryptostoma Dejean, 1821 [preoccupied genus name, not Cryptostoma Blainville, 1818 [Mollusca]; syn. of Ceratogonys Perty, 1830]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
*Orodotini Muona, 1991a: 167 [stem: Orodot-]. Type genus: Orodotes Bonvouloir, 1871. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Orodotini Muona, 1993: 52 [stem: Orodot-]. Type genus: Orodotes Bonvouloir, 1871.
†Tribe
Throscogeniini
Iablokoff-Khnzorian, 1962
Throscogeniinae Iablokoff-Khnzorian, 1962: 81 [stem: Throscogeni-]. Type genus: Throscogenius Iablokoff-Khnzorian, 1962.
Family
Throscidae
Laporte, 1840
nomen protectum
Stereolia Rafinesque, 1815: 112 [stem: Stereol-]. Type genus: Stereolus Rafinesque, 1815 [unjustified emendation of Throscus Latreille, 1797 not in prevailing usage; syn. of Trixagus Kugelann, 1794]. Comment: nomen oblitum (see Appendix 1).
Throscites Laporte, 1840a: 228 [stem: Throsc-]. Type genus: Throscus Latreille, 1797 [syn. of Trixagus Kugelann, 1794]. Comment: nomen protectum (see Appendix 1); original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 369, as Throscoidae), generally accepted as in P. J. Johnson (2002b: 158, as Throscidae).
Trixagidae Gistel, 1848: [4] [stem: Trixag-]. Type genus: Trixagus Kugelann, 1794.
Potergini Cobos, 1961: 5 [stem: Poterg-]. Type genus: Potergus Bonvouloir, 1871.
†Family
Praelateriidae
Dolin, 1973
Praelateriidae Dolin, 1973: 78 [stem: Praelateri-]. Type genus: Praelaterium Dolin, 1973.
Family
Elateridae
Leach, 1815
Elaterides Leach, 1815: 85 [stem: Elater-]. Type genus: Elater Linnaeus, 1758. Comment: the oldest available name for this family is Cebrionidae Latreille, 1802, however Elateridae Leach, 1815 is maintained here pending resolution of an application to the ICZN being prepared by P. J. Johnson (see Lawrence and Newton 1995: 852; P. J. Johnson pers. comm. 2009).
Subfamily
Cebrioninae
Latreille, 1802
Cebrionates Latreille, 1802: 97 [stem: Cebrion-]. Type genus: Cebrio A. G. Olivier, 1790.
Plastocerini J. L. LeConte, 1861: 172 [stem: Plastocer-]. Type genus: Plastocerus sensu J. L. LeConte, 1853 [not Plastocerus Schaum, 1852; syn. of Octinodes Candèze, 1863]. Comment: based on a misidentified type genus; an application should be submitted to the Commission to suppress this name for the Principles of Priority and Homonymy (Art. 65.2.1) since Plastoceridae Crowson, 1972 is currently used as valid in Elateroidea.
Aplastinae Stibick, 1979: 175 [stem: Aplast-]. Type genus: Aplastus J. L. LeConte, 1859.
Cebriognathinae Paulus, 1981: 264, in key [stem: Cebriognath-]. Type genus: Cebriognathus Chobaut, 1899.
Subfamily
Agrypninae
Candèze, 1857
nomen protectum
Agrypnides Candèze, 1857: 17 [stem: Agrypn-]. Type genus: Agrypnus Eschscholtz, 1829. Comment: nomen protectum (see Appendix 1); Agrypninae Candèze, 1857 given precedence for subfamily name over Oophorinae Gistel, 1848 (Art. 35.5).
Tribe
Agrypnini
Candèze, 1857
nomen protectum
Adeloceraeidae Gistel, 1848: [5] [stem: Adelocer-]. Type genus: Adelocera Latreille, 1829. Comment: nomen oblitum (see Appendix 1); incorrect original stem formation, not in prevailing usage.
Pangauradae Gistel, 1856a: 366 [stem: Pangaur-]. Type genus: Pangaura Gistel, 1856 [syn. of Lacon Laporte, 1838]. Comment: nomen oblitum (see Appendix 1); incorrect original stem formation, not in prevailing usage.
Agrypnides Candèze, 1857: 17 [stem: Agrypn-]. Type genus: Agrypnus Eschscholtz, 1829. Comment: published before 29 June 1857; nomen protectum (see Appendix 1); original vernacular name available (Art. 11.7.2): first used in latinized form by Kiesenwetter (1858: 230, as Agrypnini), generally accepted as in Lawrence and Newton (1995: 854, as Agrypninae); this family-group name was also proposed in the same year by Lacordaire (1857 [before 25 May]: 138, as Agrypnides) (see Bibliographic notes in Introduction).
*Octocryptites Candèze, 1892: 486 [stem: Octocrypt-]. Type genus: Octocryptus Candèze, 1892. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Candèze (1892).
Adelocerini Buysson, 1893: 15 [stem: Adelocer-]. Type genus: Adelocera Latreille, 1829. Comment: family-group name proposed as new without reference to Adeloceridae Gistel, 1848.
Octocryptini Schwarz, 1906: 31 [stem: Octocrypt-]. Type genus: Octocryptus Candèze, 1892.
Cavicoxumidae Pic, 1928: 21 [stem: Cavicox-]. Type genus: Cavicoxum Pic, 1928 [syn. of Agraeus Candèze, 1857]. Comment: incorrect original stem formation, not in prevailing usage.
Laconini Dajoz, 1964: 60 [stem: Lacon-]. Type genus: Lacon Laporte, 1838.
Tribe
Anaissini
Golbach, 1984
Alampina C. Costa, 1975: 53 [stem: Alamp-]. Type genus: Alampes Champion, 1895 [preoccupied genus name, not Alampes Horváth, 1884 [Hemiptera]; syn. of Peralampes P. J. Johnson, 2002]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Anaissini Golbach, 1984: 81 [stem: Anaiss-]. Type genus: Anaissus Candèze, 1857.
†Tribe
Cryptocardiini
Dolin, 1980
Cryptocardiini Dolin, 1980: 74 [stem: Cryptocardi-]. Type genus: Cryptocardius Dolin, 1980.
Tribe
Euplinthini
Costa, 1975
Euplinthina C. Costa, 1975: 66 [stem: Euplinth-]. Type genus: Euplinthus C. Costa, 1975. Comment: we act as First Revisers (Compsoplinthini C. Costa, 1975 vs Euplinthini C. Costa, 1975).
Subtribe
Cleidecostina
Johnson, 2002
Heligmini C. Costa, 1975: 53 [stem: Heligm-]. Type genus: Heligmus Candèze, 1865 [preoccupied genus name, not Heligmus Dujardin, 1845 [Nematoda]; syn. of Cleidecosta P. J. Johnson, 2002]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Cleidecostini P. J. Johnson, 2002a: 16 [stem: Cleidecost-]. Type genus: Cleidecosta P. J. Johnson, 2002. Comment: replacement name for Heligmini C. Costa, 1975 because of the homonymy of the type genus.
Subtribe
Compsoplinthina
Costa, 1975
Compsoplinthina C. Costa, 1975: 71 [stem: Compsoplinth-]. Type genus: Compsoplinthus C. Costa, 1975.
Subtribe
Euplinthina
Costa, 1975
Euplinthina C. Costa, 1975: 66 [stem: Euplinth-]. Type genus: Euplinthus C. Costa, 1975.
Tribe
Hemirhipini
Candèze, 1857
Hémirhipides Candèze, 1857: 199 [stem: Hemirhip-]. Type genus: Hemirhipus Berthold, 1827. Comment: published before 29 June; original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1858: 75, as Hemirhipitae), generally accepted as in P. J. Johnson (2002b: 169, as Hemirhipini); this family-group name was also proposed in the same year by Lacordaire (1857 [before 25 May]: 148, as Hémirhipides) (see Bibliographic notes in Introduction); First Reviser (Hemirhipini Candèze, 1857 vs Chalcolepidiini Candèze, 1857) not determined, current usage maintained.
Chalcolépidiides Candèze, 1857: 257 [stem: Chalcolepidi-]. Type genus: Chalcolepidius Eschscholtz, 1829. Comment: published before 29 June 1857; original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 164, as Chalcolepidiini), generally accepted as in Hyslop (1917: 252, as Chalcolepidina [incorrect stem formation]); this family-group name was also proposed in the same year by Lacordaire (1857 [before 25 May]: 153, as Chalcolépidiides) (see Bibliographic notes in Introduction).
Alaites Candèze, 1874: 112 [stem: Ala-]. Type genus: Alaus Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1894: 269, as Alaini), generally accepted as in Heyne and Taschenberg (1905: 152, as Alaini).
Ludioctenina Jakobson, 1913: 755 [stem: Ludiocten-]. Type genus: Ludioctenus Fairmaire, 1893.
Alauinae Laurent, 1974: 16 [stem: Ala-]. Type genus: Alaus Eschscholtz, 1829. Comment: family-group name proposed as new without reference to Alaites Candèze, 1874; incorrect original stem formation, not in prevailing usage.
Tribe
Oophorini
Gistel, 1848
Oophoridae Gistel, 1848: [5] [stem: Oophor-]. Type genus: Oophorus Eschscholtz, 1833 [syn. of Aeolus Eschscholtz, 1829]. Comment: family-group name attributed to Gistel (1856a: 367) in recent literature.
Monocrépidiites Candèze, 1859: 176 [stem: Monocrepidi-]. Type genus: Monocrepidius Eschscholtz, 1829 [syn. of Conoderus Eschscholtz, 1829]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 167, as Monocrepidii), generally accepted as in Burakowski et al. (1985: 147, as Monocrepidiinae).
Drasteriini Houlbert, 1912: 184 [stem: Drasteri-]. Type genus: Drasterius Eschscholtz, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1987b)]. Comment: the junior homonym Drasteriini Wiltshire, 1976 (type genus Drasteria Hübner, 1818) is available in Lepidoptera: Noctuidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
*Aeolina Jakobson, 1913: 747 [stem: Aeol-]. Type genus: Aeolus Eschscholtz, 1829. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently.
Conoderinae Fleutiaux, 1919: 58 [stem: Conoder-]. Type genus: Conoderus Eschscholtz, 1829. Comment: the older name Conoderinae Schönherr, 1833 (type genus Conoderes Schönherr, 1833) is curently used as valid in Coleoptera: Curculionidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Pachyderinae Fleutiaux, 1919: 57 [stem: Pachyder-]. Type genus: Pachyderes Guérin-Méneville, 1830.
Tribe
Platycrepidiini
Costa and Casari-Chen, 1993
Eudactylites Candèze, 1859: 153 [stem: Eudactyl-]. Type genus: Eudactylus Sallé, 1855 [preoccupied genus name, not Eudactylus Fitzinger, 1843 [Reptilia]; syn. of Platycrepidius Candèze, 1859]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1895: 337, as Eudactylini), generally accepted as in Hyslop (1917: 259, as Eudactylini); permanently invalid (Art. 39): based on preoccupied type genus.
Platycrepidiini C. Costa and Casari-Chen, 1993: 62 [stem: Platycrepidi-]. Type genus: Platycrepidius Candèze, 1859.
Tribe
Pseudomelanactini
Arnett, 1967
Pseudomelanactini Arnett, 1967: 111 [stem: Pseudomelanact-]. Type genus: Pseudomelanactes Mathieu, 1961 [syn. of Anthracalaus Fairmaire, 1889].
Tribe
Pyrophorini
Candèze, 1863
Pyrophorites Candèze, 1863: 3 [stem: Pyrophor-]. Type genus: Pyrophorus Billberg, 1820.
Subtribe
Hapsodrilina
Costa, 1975
Hapsodrilina C. Costa, 1975: 88 [stem: Hapsodril-]. Type genus: Hapsodrilus C. Costa, 1975.
Subtribe
Nyctophyxina
Costa, 1975
Nyctophyxina C. Costa, 1975: 85 [stem: Nyctophyx-]. Type genus: Nyctophysis C. Costa, 1975. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Nyctophyse-).
Subtribe
Pyrophorina
Candèze, 1863
Pyrophorites Candèze, 1863: 3 [stem: Pyrophor-]. Type genus: Pyrophorus Billberg, 1820. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1896: 463, as Pyrophorini), generally accepted as in P. J. Johnson (2002b: 169, as Pyrophorini).
Tribe
Tetralobini
Laporte, 1840
Tetralobites Laporte, 1840a: 230 [stem: Tetralob-]. Type genus: Tetralobus Lepeletier and Audinet-Serville, 1828. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Blanchard (1853: 84, as Tetralobitae), generally accepted as in Lawrence and Newton (1995: 853, as Tetralobinae).
Phyllophoridae Hope, 1842: 73 [stem: Phyllophor-]. Type genus: Phyllophorus Hope, 1842 [preoccupied genus name, not Phyllophorus Grube, 1840 [Echinodermata]; syn. of Tetralobus Lepeletier and Audinet-Serville, 1828]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Piezophyllini Laurent, 1967: 85 [stem: Piezophyll-]. Type genus: Piezophyllus Hope, 1842.
Subfamily
Thylacosterninae
Fleutiaux, 1920
Soleniscinae Lameere, 1900: 377 [stem: Solenisc-]. Type genus: Soleniscus Bonvouloir, 1875 [preoccupied genus name, not Soleniscus Meek and Worthen, 1860 [Mollusca]; syn. of Cussolenis Fleutiaux, 1918]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Pterotarsini Fleutiaux, 1902: 648 [stem: Pterotars-]. Type genus: Pterotarsus Guérin-Méneville, 1829. Comment: although this name has priority over Thylacosterninae Fleutiaux, 1920, an application will be submitted to the Commission by J. Muona and H. Silfverberg to conserve the well-established younger name.
Thylacosterninae Fleutiaux, 1920: 94 [stem: Thylacostern-]. Type genus: Thylacosternus Bonvouloir, 1875. Comment: an application will be submitted to the Commission by J. Muona and H. Silfverberg (pers. comm. 2010) to conserve prevailing usage of Thylacosterninae Fleutiaux, 1920 over the older name Pterotarsinae Fleutiaux, 1902.
Balginae Fleutiaux, 1926: 30 [stem: Balg-]. Type genus: Balgus Fleutiaux, 1920.
Cussolenitae Cobos, 1961: 3 [stem: Cussolen-]. Type genus: Cussolenis Fleutiaux, 1918.
Subfamily
Lissominae
Laporte, 1835
Lissomidae Laporte, 1835b: 178 [stem: Lissom-]. Type genus: Lissomus Dalman, 1824.
Drapetini J. L. LeConte, 1863: 44 [stem: Drapet-]. Type genus: Drapetes Dejean, 1821. Comment: Drapetini Collin, 1961 (type genus Drapetis Meigen, 1822) has been used as valid to this day in Diptera although Sabrosky (1999: 118) pointed out that the correct stem for the Diptera name is Drapetid-.
Protelateridae Schwarz, 1902: 365 [stem: Protelater-]. Type genus: Protelater Sharp, 1877.
Oestodini Hyslop, 1917: 251 [stem: Oestod-]. Type genus: Oestodes J. L. LeConte, 1853.
Athoomorphinae Laurent, 1966: 818 [stem: Athoomorph-]. Type genus: Athoomorphus Schwarz, 1898.
Drapetini Dolin, 1975b: 1627, in key [stem: Drapet-]. Type genus: Drapetes Dejean, 1821. Comment: proposed as new without reference to Drapetini J. L. LeConte, 1863.
Sphaenelaterini Stibick, 1979: 179 [stem: Sphaenelater-]. Type genus: Sphaenelater Schwarz, 1902.
Subfamily
Semiotinae
Jakobson, 1913
Semiotina Jakobson, 1913: 736 [stem: Semiot-]. Type genus: Semiotus Eschscholtz, 1829.
Semiotinae Golbach, 1970: 320 [stem: Semiot-]. Type genus: Semiotus Eschscholtz, 1829. Comment: family-group name proposed as new without reference to Semiotina Jakobson, 1913.
Subfamily
Campyloxeninae
Costa, 1975
Campyloxeninae C. Costa, 1975: 114 [stem: Campyloxen-]. Type genus: Campyloxenus Fairmaire, 1860.
Subfamily
Pityobiinae
Hyslop, 1917
Pityobini Hyslop, 1917: 249 [stem: Pityobi-]. Type genus: Pityobius J. L. LeConte, 1853. Comment: incorrect original stem formation, not in prevailing usage.
Rostricephalinae Fleutiaux, 1947: 238, in key [stem: Rostricephal-]. Type genus: Rostricephalus Fleutiaux, 1918.
Adzusini Kishii, 1989: 2 [stem: Adzus-]. Type genus: Adzusa Kishii, 1957.
Subfamily
Oxynopterinae
Candèze, 1857
Oxynoptérides Candèze, 1857: 355 [stem: Oxynopter-]. Type genus: Oxynopterus Hope, 1842. Comment: published before 29 June 1857; original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1905: 155, as Oxynopterini), generally accepted as in Johnson (2002b: 169, as Oxynopterini); this family-group name was also proposed in the same year by Lacordaire (1857 [before 25 May]: 158, as Oxynoptérides) (see Bibliographic notes in Introduction); First Revisor (Melanactinae Candèze, 1857 vs Oxynopterinae Candèze, 1857) not determined, current usage maintained.
Mélanactides Candèze, 1857: 182 [stem: Melanact-]. Type genus: Melanactes J. L. LeConte, 1853. Comment: published before 29 June 1857; original vernacular name available (Art. 11.7.2): first used in latinized form by Holmgren (1899: 199, as Melanactidae), generally accepted as in Stibick (1979: 164, as Melanactinae); this family-group name was also proposed in the same year by Lacordaire (1857 [before 25 May]: 144, as Mélanactides) (see Bibliographic notes in Introduction).
Asaphites Candèze, 1863: 207 [stem: Asaph-]. Type genus: Asaphes Kirby, 1837 [preoccupied genus name, not Asaphes Walker, 1834 [Hymenoptera]; syn. of Hemicrepidius Germar, 1839]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Heyne and Taschenberg (1905: 163, as Asaphini); permanently invalid (Art. 39): based on preoccupied type genus; the older name Asaphidae H. C. C. Burmeister, 1843 (type genus Asaphus Brongniart 1822) is currently used as valid in Trilobita; the younger name Asaphinae Ashmead, 1904 (type genus Asaphes Walker, 1834) is currently used as valid in Hymenoptera: Pteromalidae.
Campsosterninae Fleutiaux, 1927a: 104 [stem: Campsostern-]. Type genus: Campsosternus Latreille, 1834. Comment: published 25 April 1927; this family-group name was also used in the same year by Fleutiaux (1927b [“31 December”]: 108, as Campsosterninae).
Pectocerini Gurjeva, 1974: 107, in key [stem: Pectocer-]. Type genus: Pectocera Hope, 1842.
Subfamily
Dendrometrinae
Gistel, 1848
Dendrometridae Gistel, 1848: [5] [stem: Dendrometr-]. Type genus: Dendrometrus Gistel, 1848 [syn. of Limonius Eschscholtz, 1829]. Comment: First Reviser found (Denticollinae Stein and Weise, 1877 (1848) vs Dendrometrinae Gistel, 1848) is Sánchez-Ruiz (1996: 74).
Tribe
Crepidomenini
Candèze, 1863
Crépidoménites Candèze, 1863: 190 [stem: Crepidomen-]. Type genus: Crepidomenus Erichson, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1905: 163, as Crepidomenini), generally accepted as in Calder (1978: 295, as Crepidomeninae).
Tribe
Dendrometrini
Gistel, 1848
Dendrometridae Gistel, 1848: [5] [stem: Dendrometr-]. Type genus: Dendrometrus Gistel, 1848 [syn. of Limonius Eschscholtz, 1829]. Comment: First Reviser found (Denticollini Stein and Weise, 1877 (1848) vs Dendrometrini Gistel, 1848) is Sánchez-Ruiz (1996: 75).
Subtribe
Dendrometrina
Gistel, 1848
Dendrometridae Gistel, 1848: [5] [stem: Dendrometr-]. Type genus: Dendrometrus Gistel, 1848 [syn. of Limonius Eschscholtz, 1829].
Athoites Candèze, 1859: 4 [stem: Atho-]. Type genus: Athous Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 170, as Athoi), generally accepted as in Burakowski et al. (1985: 189, as Athoinae).
Limoniina Jakobson, 1913: 755 [stem: Limoni-]. Type genus: Limonius Eschscholtz, 1829.
Subtribe
Denticollina
Stein and Weise, 1877 (1848)
Campylidae Gistel, 1848: [5] [stem: Campyl-]. Type genus: Campylus Fischer von Waldheim, 1824 [syn. of Denticollis Piller and Mitterpacher, 1783]. Comment: usage of the younger name Denticollina Stein and Weise, 1877 conserved over this name (Art. 40.2).
Denticollini Stein and Weise, 1877: 96 [stem: Denticoll-]. Type genus: Denticollis Piller and Mitterpacher, 1783. Comment: younger name conserved over Campylina Gistel, 1848 (Art. 40.2).
Lepturoidini Schwarz, 1906: 3, in key [stem: Lepturoid-]. Type genus: Lepturoides Herbst, 1784 [syn. of Denticollis Piller and Mitterpacher, 1783].
Subtribe
Hemicrepidiina
Champion, 1896
Hemicrepidiini Champion, 1896: 477 [stem: Hemicrepidi-]. Type genus: Hemicrepidius Germar, 1839.
Tribe
Dimini
Candèze, 1863
Dimites Candèze, 1863: 237 [stem: Dim-]. Type genus: Dima Charpentier, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1896: 476, as Dimini), generally accepted as in Gurjeva (1974: 107, as Diminae); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Dimat-).
Beliophorina Jakobson, 1913: 737 [stem: Beliophor-]. Type genus: Beliophorus Eschscholtz, 1829.
Peniini Dolin, 1990: 17 [stem: Peni-]. Type genus: Penia Laporte, 1836.
Tribe
Hypnoidini
Schwarz, 1906 (1860)
Cryptohypnites Candèze, 1860: 50 [stem: Cryptohypn-]. Type genus: Cryptohypnus Eschscholtz, 1830 [syn. of Hypnoidus Dillwyn, 1829]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 166, as Cryptohypni), generally accepted as in Burakowski et al. (1985: 214, as Cryptohypninae); Hypnoidini Schwarz, 1906 conserved over this name (Art. 40.2).
Hypnoidini Schwarz, 1906: 150 [stem: Hypnoid-]. Type genus: Hypnoidus Dillwyn, 1829. Comment: name proposed to replace Cryptohypnites Candèze, 1860 because of the synonymy of the type genus; younger name conserved over Cryptohypnini Candèze, 1860 (Art. 40.2).
Hypolithinae Fleutiaux, 1928: 252 [stem: Hypolith-]. Type genus: Hypolithus Eschscholtz, 1829. Comment: name proposed to replace Cryptohypnites Candèze, 1860 because of the synonymy of the type genus.
*Prisahypini Stibick, 1976: 197 [stem: Prisahypn-]. Type genus: Prisahypnus Stibick, 1979. Comment: unavailable family-group name, not based on available genus name; incorrect original stem formation, not in prevailing usage.
Prisahypini Stibick, 1979: 166 [stem: Prisahypn-]. Type genus: Prisahypnus Stibick, 1979. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Pleonomini
Semenov and Pjatakova, 1936
Pleonomini Semenov and Pjatakova, 1936: 103 [stem: Pleonom-]. Type genus: Pleonomus Ménétriés, 1849. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Stibick (1979: 176) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Tribe
Prosternini
Gistel, 1856
nomen protectum
Diacanthidae Gistel, 1848: [5] [stem: Diacanth-]. Type genus: Diacanthus Latreille, 1834 [syn. of Selatosomus Stephens, 1830]. Comment: nomen oblitum (see Appendix 1).
Prosternidae Gistel, 1856a: 367 [stem: Prostern-]. Type genus: Prosternon Latreille, 1834. Comment: nomen protectum (see Appendix 1).
Corymbitini J. L. LeConte, 1861: 169 [stem: Corymbit-]. Type genus: Corymbites Latreille, 1834 [syn. of Ctenicera Latreille, 1829].
Ctenicerina Jakobson, 1913: 736 [stem: Ctenicer-]. Type genus: Ctenicera Latreille, 1829. Comment: family-group name previously attributed to Fleutiaux (1936) in the literature; “Ctenicerini Fleutiaux, 1936” used as valid instead of Prosternini Gistel, 1856 by Cate (2007: 33, 173).
Ctenicerinae Neboiss, 1956: 47 [stem: Ctenicer-]. Type genus: Ctenicera Latreille, 1829. Comment: name proposed to replace Corymbitinae J. L. LeConte, 1861 because of the synonymy of the type genus; family-group name proposed as new without reference to Ctenicerina Jakobson, 1913 or Ctenicerinae Fleutiaux, 1936.
Tribe
Senodoniini
Schenkling, 1927
Allotriites Candèze, 1863: 225 [stem: Allotri-]. Type genus: Allotrius Laporte, 1840 [preoccupied genus name, not Allotrius Temminck, 1835 [Aves]; syn. of Senodonia Laporte, 1838]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1896: 489, as Allotriini), generally accepted as in Heyne and Taschenberg (1905: 163, as Allotriini); permanently invalid (Art. 39): based on preoccupied type genus.
Senodoniinae Schenkling, 1927: 417 [stem: Senodoni-]. Type genus: Senodonia Laporte, 1838.
Senodoniini Dolin, 2000: 19 [stem: Senodoni-]. Type genus: Senodonia Laporte, 1838. Comment: family-group name proposed as new without reference to Senodoniinae Schenkling, 1927.
Subfamily
Negastriinae
Nakane and Kishii, 1956
Negastriinae Nakane and Kishii, 1956: 203 [stem: Negastri-]. Type genus: Negastrius C. G. Thomson, 1859.
Tribe
Negastriini
Nakane and Kishii, 1956
Negastriinae Nakane and Kishii, 1956: 203 [stem: Negastri-]. Type genus: Negastrius C. G. Thomson, 1859.
Tribe
Quasimusini
Schimmel and Tarnawski, 2009
Quasimusini Schimmel and Tarnawski, 2009: 17 [stem: Quasimus-]. Type genus: Quasimus Gozis, 1886.
Subtribe
Loebliquasimusina
Schimmel and Tarnawski, 2009
Loebliquasina Schimmel and Tarnawski, 2009: 18 [stem: Loebliquas-]. Type genus: Loebliquasis Dolin, 1997. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Quasimusina
Schimmel and Tarnawski, 2009
Quasimusina Schimmel and Tarnawski, 2009: 19 [stem: Quasimus-]. Type genus: Quasimus Gozis, 1886.
Subtribe
Striatoquasimusina
Schimmel and Tarnawski, 2009
Striatoquasina Schimmel and Tarnawski, 2009: 20 [stem: Striatoquasimus-]. Type genus: Striatoquasimus Schimmel and Tarnawski, 2009. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Wittmeroquasimusina
Schimmel and Tarnawski, 2009
Wittmeroquasina Schimmel and Tarnawski, 2009: 20 [stem: Wittmeroquasimus-]. Type genus: Wittmeroquasimus Dolin, 1997. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Elaterinae
Leach, 1815
Elaterides Leach, 1815: 85 [stem: Elater-]. Type genus: Elater Linnaeus, 1758.
Tribe
Agriotini
Laporte, 1840
Agriotites Laporte, 1840a: 233 [stem: Agriot-]. Type genus: Agriotes Eschscholtz, 1829.
Subtribe
Agriotina
Laporte, 1840
Agriotites Laporte, 1840a: 233 [stem: Agriot-]. Type genus: Agriotes Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1896: 511, as Agriotini), generally accepted as in Hansen (1996: 143, as Agriotini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Agriotet-).
Subtribe
Cardiorhinina
Candèze, 1863
Cardiorhinites Candèze, 1863: 247 [stem: Cardiorhin-]. Type genus: Cardiorhinus Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1896: 495, as Cardiorhini), generally accepted as in Heyne and Taschenberg (1905: 164, as Cardiorhinini).
Tribe
Ampedini
Gistel, 1848
Ampedidae Gistel, 1848: [5] [stem: Amped-]. Type genus: Ampedus Dejean, 1833. Comment: family-group name previously attributed to Gistel (1856a: 367), e.g., P. J. Johnson (2002b: 168).
Tribe
Dicrepidiini
Thomson, 1858
Dicrepidiitae J. Thomson, 1858: 75 [stem: Dicrepidi-]. Type genus: Dicrepidius Eschscholtz, 1829.
Tribe
Elaterini
Leach, 1815
Elaterides Leach, 1815: 85 [stem: Elater-]. Type genus: Elater Linnaeus, 1758.
Arneidae Gistel, 1848: [6] [stem: Arne-]. Type genus: Arneus Gistel, 1848 [syn. of Sericus Eschscholtz, 1829].
Steatoderidae Gistel, 1848: [5] [stem: Steatoder-]. Type genus: Steatoderus Dejean, 1833 [syn. of Elater Linnaeus, 1758].
Amphilabridae Gistel, 1856a: 367 [stem: Amphilabr-]. Type genus: Amphilabris Gistel, 1834 [syn. of Sericus Eschscholtz, 1829].
Ludiides Lacordaire, 1857: 197 [stem: Ludi-]. Type genus: Ludius Berthold, 1827 [syn. of Elater Linnaeus, 1758]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte (1861: 168, as Ludii); the junior homonym Ludiini Aurivillius, 1904 (type genus Ludia Wallengren, 1865) in Lepidoptera: Saturniidae was replaced by Micragonini Cockerell, 1914 (see Oberprieler 1997: 146).
Hypodésites Candèze, 1863: 242 [stem: Hypodese-]. Type genus: Hypodesis Latreille, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1896: 490, as Hypodesini), generally accepted as in Heyne and Taschenberg (1905: 164, as Hypodesini); incorrect original stem formation, not in prevailing usage.
Sericosomina Hyslop, 1917: 258 [stem: Sericosom-]. Type genus: Sericosomus Dejean, 1833 [syn. of Sericus Eschscholtz, 1829].
*Dolerosomini Dolin, 1975b: 1632 [stem: Dolerosom-]. Type genus: Dolerosomus Motschulsky, 1859. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Megapenthini
Gurjeva, 1973
Megapenthini Gurjeva, 1973: 448 [stem: Megapenth-]. Type genus: Megapenthes Kiesenwetter, 1858.
Tribe
Melanotini
Candèze, 1859 (1848)
Cratonychidae Gistel, 1848: [5] [stem: Cratonych-]. Type genus: Cratonychus Dejean, 1833 [syn. of Melanotus Eschscholtz, 1829]. Comment: use of Melanotini Candèze, 1859 converved over this name (Art. 40.2).
Mélanotites Candèze, 1859: 4 [stem: Melanot-]. Type genus: Melanotus Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 169, as Melanoti), generally accepted as in Hansen (1996: 143, as Melanotini); name conserved over Cratonychini Gistel, 1848 (Art. 40.2) (see Sánchez-Ruiz 1996: 167).
Tribe
Odontonychini
Girard, 1973
Odontonychini Girard, 1973: 276 [stem: Odontonych-]. Type genus: Odontonychus Candèze, 1897.
Tribe
Physorhinini
Candèze, 1859
Physorhinites Candèze, 1859: 384 [stem: Physorhin-]. Type genus: Physorhinus Germar, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 167, as Physorhini), generally accepted as in Burakowski et al. (1985: 114, as Physorhininae).
Tribe
Pomachiliini
Candèze, 1859
Pomachiliites Candèze, 1859: 4 [stem: Pomachili-]. Type genus: Pomachilius Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1895: 402, as Pomachiliini), generally accepted as in Burakowski et al. (1985: 131, as Pomachiliinae).
Tribe
Synaptini
Gistel, 1856
Dairaeidae Gistel, 1848: [5] [stem: Dair-]. Type genus: Daira Gistel, 1848 [preoccupied genus name, not Daira Milne-Edwards, 1830 [Crustacea]; syn. of Synaptus Eschscholtz, 1829]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Synaptidae Gistel, 1856a: 366 [stem: Synapt-]. Type genus: Synaptus Eschscholtz, 1829.
Adrastites Candèze, 1863: 448 [stem: Adrast-]. Type genus: Adrastus Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1896: 533, as Adrastini), generally accepted as in Burakowski et al. (1985: 133, as Adrastinae).
Subfamily
Cardiophorinae
Candèze, 1859
Cardiophorites Candèze, 1859: 4 [stem: Cardiophor-]. Type genus: Cardiophorus Eschscholtz, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 166, as Cardiophori), generally accepted as in Burakowski et al. (1985: 227, as Cardiophorinae).
Aphrici J. L. LeConte, 1861: 173 [stem: Aphric-]. Type genus: Aphricus J. L. LeConte, 1854.
Aptopina Jakobson, 1913: 760 [stem: Aptopod-]. Type genus: Aptopus Eschscholtz, 1829. Comment: incorrect original stem formation, not in prevailing usage.
Esthesopinae Fleutiaux, 1919: 76 [stem: Esthesopod-]. Type genus: Esthesopus Eschscholtz, 1829. Comment: incorrect original stem formation, not in prevailing usage.
Nyctorini Semenov and Pjatakova, 1936: 102 [stem: Nyctor-]. Type genus: Nyctor Semenov and Pjatakova, 1936 [syn. of Cardiophorus Eschscholtz, 1829].
Subfamily
Hemiopinae
Fleutiaux, 1941
Hemiopinae Fleutiaux, 1941: 31 [stem: Hemiop-]. Type genus: Hemiops Laporte, 1838.
Subfamily
Physodactylinae
Lacordaire, 1857
Physodactylides Lacordaire, 1857: 236 [stem: Physodactyl-]. Type genus: Physodactylus Fischer von Waldheim, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Fleutiaux (1892b: 404, as Physodactylini), generally accepted as in Lawrence and Newton (1995: 855, as Physodactylinae).
Toxognathinae Fleutiaux, 1941: 34 [stem: Toxognath-]. Type genus: Toxognathus Fairmaire, 1878.
Subfamily
Eudicronychinae
Girard, 1971
Dicronychidae Schwarz, 1897: 11 [stem: Dicronych-]. Type genus: Dicronychus Laporte, 1840 [preoccupied genus name, not Dicronychus Brullé, 1832 [Coleoptera: Elateridae: Cardiophorinae]; syn. of Eudicronychus Méquignon, 1931]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Eudicronychinae Girard, 1971: 645 [stem: Eudicronych-]. Type genus: Eudicronychus Méquignon, 1931. Comment: description by indication (distinguishing characters given in Schwarz (1907: 2, as Dicronychidae)).
Subfamily
Subprotelaterinae
Fleutiaux, 1920
Subprotelaterinae Fleutiaux, 1920: 99 [stem: Subprotelater-]. Type genus: Subprotelater Fleutiaux, 1916.
Subfamily
Morostomatinae
Dolin, 2000
Morostominae Dolin, 2000: 18 [stem: Morostomat-]. Type genus: Morostoma Candèze, 1879. Comment: incorrect original stem formation, not in prevailing usage.
†Subfamily
Protagrypninae
Dolin, 1973
Protagrypnini Dolin, 1973: 74 [stem: Protagrypn-]. Type genus: Protagrypnus Dolin, 1973.
†Tribe
Desmatini
Dolin, 1975
Desmatini Dolin, 1975a: 60 [stem: Desmat-]. Type genus: Desmatus Dolin, 1975.
†Tribe
Hypnomorphini
Dolin, 1975
Hypnomorphini Dolin, 1975a: 54 [stem: Hypnomorph-]. Type genus: Hypnomorphus Dolin, 1975.
†Tribe
Protagrypnini
Dolin, 1973
Protagrypnini Dolin, 1973: 74 [stem: Protagrypn-]. Type genus: Protagrypnus Dolin, 1973.
Family
Plastoceridae
Crowson, 1972
Plastoceridae Crowson, 1972: 37, in key [stem: Plastocer-]. Type genus: Plastocerus Schaum, 1852. Comment: an application needs to be submitted to the Commission to suppress Plastoceri J. L. LeConte, 1861 (based on the misidentified type genus Plastocerus sensu J. L. LeConte, 1853) for the Principles of Priority and Homonymy (Art. 65.2.1) to conserve this name as valid.
Family
Drilidae
Blanchard, 1845
Drilites Blanchard, 1845b: 53 [stem: Dril-]. Type genus: Drilus A. G. Olivier, 1790.
Subfamily
Drilinae
Blanchard, 1845
Drilites Blanchard, 1845b: 53 [stem: Dril-]. Type genus: Drilus A. G. Olivier, 1790. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1848: [11], as Drilidae), generally accepted as in Hansen (1996: 144, as Drilidae).
Subfamily
Thilmaninae
Kazantsev, 2004
Thilmaninae Kazantsev, 2004b: 240 [stem: Thilman-]. Type genus: Thilmanus Baudi di Selve, 1871. Comment: transferred from Lycidae by Bocák and Brlik (2008).
Tribe
Euanomini
Kazantsev, 2010
Euanomini Kazantsev, 2010a: 55 [stem: Euanom-]. Type genus: Euanoma Reitter, 1889.
Tribe
Thilmanini
Kazantsev, 2004
Thilmaninae Kazantsev, 2004b: 240 [stem: Thilman-]. Type genus: Thilmanus Baudi di Selve, 1871.
Family
Omalisidae
Lacordaire, 1857
*Homalises Motschulsky, 1849: 55 [stem: Omalis-]. Type genus: Omalisus Geoffroy, 1762 [as Homalisus, unjustified emendation of type genus name by Illiger (1801: 139), not in prevailing usage; placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Homalisides Lacordaire, 1857: 303 [stem: Omalis-]. Type genus: Omalisus Geoffroy, 1762 [as Homalisus, unjustified emendation of type genus name by Illiger (1801: 139), not in prevailing usage; placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kiesenwetter (1860: 442, as Homalisidae [incorrect stem formation]), generally accepted as in Lawrence and Newton (1995: 856, as Omalisidae); incorrect original stem formation, not in prevailing usage.
†Family
Berendtimiridae
Winkler, 1987
Berendtimiridae J. R. Winkler, 1987: 52 [stem: Berendtimir-]. Type genus: Berendtimirus J. R. Winkler, 1987.
Family
Lycidae
Laporte, 1836
Lycusidae Laporte, 1836: 25 [stem: Lyc-]. Type genus: Lycus Fabricius, 1787.
Subfamily
Libnetinae
Bocák and Bocáková, 1990
Libnetinina Bocák and Bocáková, 1990: 652 [stem: Libnet-]. Type genus: Libnetis C. O. Waterhouse, 1878. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Dictyopterinae
Houlbert, 1922
Dictyopterini Houlbert, 1922a: 338 [stem: Dictyopter-]. Type genus: Dictyoptera Latreille, 1829.
Tribe
Dictyopterini
Houlbert, 1922
Dictyopterini Houlbert, 1922a: 338 [stem: Dictyopter-]. Type genus: Dictyoptera Latreille, 1829. Comment: family-group name previously attributed to Kleine (1929: 226); although Houlbert (1922a) uses the tribe name Lycini for a group of genera that includes Dictyoptera Latreille in the main text (page 240), this was certainly a mistake since the tribe name Dictyopterini is correctly used in the “Index alphabétique” (page 319) and in the “Table systématique” (page 338).
Tribe
Lycoprogenthini
Bocák and Bocáková, 2008
Lycoprogenthini Bocák and Bocáková, 2008: 709 [stem: Lycoprogenth-]. Type genus: Lycoprogenthes Pic, 1915.
Tribe
Taphini
Bocák and Bocáková, 1990
Taphinina Bocák and Bocáková, 1990: 650 [stem: Taph-]. Type genus: Taphes C. O. Waterhouse, 1878. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Lyropaeinae
Bocák and Bocáková, 1989
Lyropaeini Bocák and Bocáková, 1989: 718 [stem: Lyropae-]. Type genus: Lyropaeus C. O. Waterhouse, 1878.
Tribe
Alyculini
Bocák and Bocáková, 2008
Alyculini Bocák and Bocáková, 2008: 710 [stem: Alycul-]. Type genus: Alyculus Kazantsev, 1999.
Tribe
Antennolycini
Bocák and Bocáková, 2008
Antennolycini Bocák and Bocáková, 2008: 710 [stem: Antennolyc-]. Type genus: Antennolycus Bocák and Bocáková, 1999.
Tribe
Lyropaeini
Bocák and Bocáková, 1989
Lyropaeini Bocák and Bocáková, 1989: 718 [stem: Lyropae-]. Type genus: Lyropaeus C. O. Waterhouse, 1878.
Paralycinae L. N. Medvedev and Kazantsev, 1992: 59 [stem: Paralyc-]. Type genus: Paralycus L. N. Medvedev and Kazantsev, 1992 [syn. of Lyropaeus C. O. Waterhouse, 1878].
Tribe
Miniduliticolini
Kazantsev, 2003
Miniduliticolini Kazantsev, 2003: 20 [stem: Miniduliticol-]. Type genus: Miniduliticola Kazantsev, 2003.
Tribe
Platerodrilini
Kazantsev, 2004
*Duliticolinae Mjöberg, 1925: 140 [stem: Duliticol-]. Type genus: Duliticola Mjöberg, 1925 [syn. of Platerodrilus Pic, 1921]. Comment: unavailable family-group name (Art. 8.3): nomenclatural act disclaimed.
*Duliticolinae Kazantsev, 2003: 19 [stem: Duliticol-]. Type genus: Duliticola Mjöberg, 1925 [syn. of Platerodrilus Pic, 1921]. Comment: family-group name proposed as a new taxon but unavailable (Art. 11.7.1.1) because it was based on a type genus which was considered a synonym of Platerodrilus Pic, 1921 by the author.
Platerodrilini Kazantsev, 2004b: 241 [stem: Platerodril-]. Type genus: Platerodrilus Pic, 1921.
Subfamily
Ateliinae
Kleine, 1929
Atelinae Kleine, 1929: 222 [stem: Ateli-]. Type genus: Atelius C. O. Waterhouse, 1878. Comment: First Revisers found (Ateliinae Kleine, 1928 vs Dilophotinae Kleine, 1928) are Bocák and Bocáková (2008: 712).
Tribe
Ateliini
Kleine, 1929
Atelinae Kleine, 1929: 222 [stem: Ateli-]. Type genus: Atelius C. O. Waterhouse, 1878. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Dilophotini
Kleine, 1929
Dilophotinae Kleine, 1929: 222 [stem: Dilophot-]. Type genus: Dilophotes C. O. Waterhouse, 1879. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Dilophotet-).
Subfamily
Lycinae
Laporte, 1836
Lycusidae Laporte, 1836: 25 [stem: Lyc-]. Type genus: Lycus Fabricius, 1787.
Tribe
Calochromini
Lacordaire, 1857
Calochromides Lacordaire, 1857: 301 [stem: Calochrom-]. Type genus: Calochromus Guérin-Méneville, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gorham (1880: 1, as Calochrominae), generally accepted as in Hansen (1996: 144, as Calochrominae).
Lygistopteri J. L. LeConte, 1881: 27 [stem: Lygistopter-]. Type genus: Lygistopterus Dejean, 1833.
Tribe
Calopterini
Green, 1949
Calopterini Green, 1949: 56, in key [stem: Calopter-]. Type genus: Calopteron Laporte, 1838.
Subtribe
Acroleptina
Bocáková, 2005
Acroleptina Bocáková, 2005: 445 [stem: Acrolept-]. Type genus: Acroleptus Bourgeois, 1886.
Subtribe
Calopterina
Green, 1949
*Calopterini Kleine, 1933: 20 [stem: Calopter-]. Type genus: Calopteron Laporte, 1838. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Calopterini Green, 1949: 56, in key [stem: Calopter-]. Type genus: Calopteron Laporte, 1838.
Tribe
Conderini
Bocák and Bocáková, 1990
Conderini Bocák and Bocáková, 1990: 643 [stem: Conder-]. Type genus: Conderis C. O. Waterhouse, 1879. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Conderid-).
Tribe
Dihammatini
Bocák and Bocáková, 2008
Dihammatini Bocák and Bocáková, 2008: 716 [stem: Dihammat-]. Type genus: Dihammatus C. O. Waterhouse, 1879.
Tribe
Erotini
LeConte, 1881
Erotes J. L. LeConte, 1881: 23 [stem: Erot-]. Type genus: Eros Newman, 1838.
Flagraxina Kazantsev, 2004a: 35, in key [stem: Flagrax-]. Type genus: Flagrax Kazantsev, 1992.
Aferotini Kazantsev, 2004a: 4 [stem: Aferot-]. Type genus: Aferos Kazantsev, 1992.
Tribe
Eurrhacini
Bocáková, 2005
Eurrhacina Bocáková, 2005: 444 [stem: Eurrhac-]. Type genus: Eurrhacus C. O. Waterhouse, 1879.
Tribe
Leptolycini
Leng and Mutchler, 1922
Leptolycini Leng and Mutchler, 1922: 430 [stem: Leptolyc-]. Type genus: Leptolycus Leng and Mutchler, 1922.
Tribe
Lycini
Laporte, 1836
Lycusidae Laporte, 1836: 25 [stem: Lyc-]. Type genus: Lycus Fabricius, 1787. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Lyponiini
Bocák and Bocáková, 1990
Lyponiinina Bocák and Bocáková, 1990: 652 [stem: Lyponi-]. Type genus: Lyponia C. O. Waterhouse, 1878. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Macrolycini
Kleine, 1929
Macrolycinae Kleine, 1929: 222 [stem: Macrolyc-]. Type genus: Macrolycus C. O. Waterhouse, 1878.
Tribe
Melanerotini
Kazantsev, 2010
Melanerotini Kazantsev, 2010b: 195 [stem: Melanerot-]. Type genus: Melaneros Fairmaire, 1877.
Tribe
Metriorrhynchini
Kleine, 1926
Metriorrhynchinae Kleine, 1926: 97 [stem: Metriorrhynch-]. Type genus: Metriorrhynchus Guérin-Méneville, 1838.
Subtribe
Hemiconderinina
Bocák and Bocáková, 1990
Hemiconderinina Bocák and Bocáková, 1990: 645 [stem: Hemiconderin-]. Type genus: Hemiconderis Kleine, 1926. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Hemiconderid-).
Subtribe
Metriorrhynchina
Kleine, 1926
Metriorrhynchinae Kleine, 1926: 97 [stem: Metriorrhynch-]. Type genus: Metriorrhynchus Guérin-Méneville, 1838 [Metriorrhynchus is an incorrect subsequent spelling by Gemminger (1869: 1629) of the original name Metriorhynchus in prevailing usage and so deemed to be the correct original spelling (Art. 33.3.1)]. Comment: First Reviser (Metriorrhynchina Kleine, 1926 vs Dilolycina Kleine, 1926) not determined, current usage maintained.
Dilolycinae Kleine, 1926: 186 [stem: Dilolyc-]. Type genus: Dilolycus Kleine, 1926. Comment: one of the original spellings of this family-group name was Haplothoracinae on page 95 (based on the new genus Haplothorax Kleine), the spelling of the family-group and genus-group names were changed to Dilolycinae and Dilolycus on page 186 and in the “Corrigenda” on page 188 of the same work because Haplothorax Agassiz, 1846 (unjustified emendation of Aplothorax G. R. Waterhouse, 1841) was already available in Carabidae; Dilolycinae is therefore treated as the correct original spelling of this family-group name (Art. 19.2).
Cladophorinae Kleine, 1929: 222 [stem: Cladophor-]. Type genus: Cladophorus Guérin-Méneville, 1830.
Subtribe
Trichalina
Kleine, 1929
Trichalinae Kleine, 1929: 222 [stem: Trichal-]. Type genus: Trichalus C. O. Waterhouse, 1877.
Tribe
Platerodini
Kleine, 1929
Platerodinae Kleine, 1929: 222 [stem: Platerod-]. Type genus: Plateros Bourgeois, 1879. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Platerot-) (see Kazantsev 2010c: 278).
Tribe
Slipinskiini
Bocák and Bocáková, 1992
Slipinskiinina Bocák and Bocáková, 1992: 256 [stem: Slipinski-]. Type genus: Slipinskia Bocák and Bocáková, 1992. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Thonalmini
Kleine, 1933
Thonalmini Kleine, 1933: 18 [stem: Thonalm-]. Type genus: Thonalmus Bourgeois, 1883. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Blackwelder (1945: 343, as Thonalmini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1) (see Bocák and Bocáková 2008: 717).
Subfamily
Dexorinae
Bocák and Bocáková, 1989
*Dexorini Kleine, 1933: 113 [stem: Dexor-]. Type genus: Dexoris C. O. Waterhouse, 1878. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Dexorinae Bocák and Bocáková, 1989: 718 [stem: Dexor-]. Type genus: Dexoris C. O. Waterhouse, 1878.
Family
Telegeusidae
Leng, 1920
Telegeusidae Leng, 1920: 152 [stem: Telegeus-]. Type genus: Telegeusis G. H. Horn, 1895. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Telegeuse-).
Family
Phengodidae
LeConte, 1861
Phengodini J. L. LeConte, 1861: 185 [stem: Phengod-]. Type genus: Phengodes Illiger, 1807.
Subfamily
Phengodinae
LeConte, 1861
Phengodini J. L. LeConte, 1861: 185 [stem: Phengod-]. Type genus: Phengodes Illiger, 1807.
Pseudophengodidae Pic, 1930: 319 [stem: Pseudophengod-]. Type genus: Pseudophengodes Pic, 1930.
Subfamily
Mastinocerinae
LeConte, 1881
Mastinocerini J. L. LeConte, 1881: 40 [stem: Mastinocer-]. Type genus: Mastinocerus Solier, 1849.
Subfamily
Penicillophorinae
Paulus, 1975
Penicillophorini Paulus, 1975: 80 [stem: Penicillophor-]. Type genus: Penicillophorus Paulus, 1975.
Family
Rhagophthalmidae
Olivier, 1907
Rhagophthalmidae E. Olivier, 1907: 63 [stem: Rhagophthalm-]. Type genus: Rhagophthalmus Motschulsky, 1853.
Family
Lampyridae
Rafinesque, 1815
Lampyria Rafinesque, 1815: 110 [stem: Lampyr-]. Type genus: Lampyris Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)].
Subfamily
Psilocladinae
McDermott, 1964
Psilocladina McDermott, 1964: 12, in key [stem: Psiloclad-]. Type genus: Psilocladus Blanchard, 1846.
Cyphonocerinae Crowson, 1972: 55, in key [stem: Cyphonocer-]. Type genus: Cyphonocerus Kiesenwetter, 1879.
Subfamily
Amydetinae
Olivier, 1907
Amydetini E. Olivier, 1907: 48 [stem: Amydet-]. Type genus: Amydetes Hoffmansegg, 1807.
Tribe
Amydetini
Olivier, 1907
Amydetini E. Olivier, 1907: 48 [stem: Amydet-]. Type genus: Amydetes Hoffmansegg, 1807.
Megalophthalmini E. Olivier, 1907: 46 [stem: Megalophthalm-]. Type genus: Megalophthalmus Gray, 1832 [preoccupied genus name, not Megalophthalmus Leach, 1830 [Crustacea]; syn. of Magnoculus McDermott, 1964]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Vestini
McDermott, 1964
Vestina McDermott, 1964: 12, in key [stem: Vest-]. Type genus: Vesta Laporte, 1833.
Subfamily
Lampyrinae
Rafinesque, 1815
Lampyria Rafinesque, 1815: 110 [stem: Lampyr-]. Type genus: Lampyris Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)].
Tribe
Cratomorphini
Green, 1948
Cratomorphi Green, 1948: 68, in key [stem: Cratomorph-]. Type genus: Cratomorphus Motschulsky, 1853.
Tribe
Lamprocerini
Olivier, 1907
Lamprocerini E. Olivier, 1907: 7 [stem: Lamprocer-]. Type genus: Lamprocera Laporte, 1833.
Tribe
Lamprohizini
Kazantsev, 2010
Lamprohizini Kazantsev, 2010d: 189 [stem: Lamprohiz-]. Type genus: Lamprohiza Motschulsky, 1853.
Tribe
Lampyrini
Rafinesque, 1815
Lampyria Rafinesque, 1815: 110 [stem: Lampyr-]. Type genus: Lampyris Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)]. Comment: family-group name previously attributed to Latreille (1816: 236); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Lampyrid-).
Tribe
Lucidotini
Lacordaire, 1857
Lucidotides Lacordaire, 1857: 310 [stem: Lucidot-]. Type genus: Lucidota Laporte, 1833.
Subtribe
Dadophorina
Olivier, 1907
Dadophorini E. Olivier, 1907: 26 [stem: Dadophor-]. Type genus: Dadophora E. Olivier, 1907.
Subtribe
Lamprigerina
McDermott, 1964
Lamprigerina McDermott, 1964: 12, in key [stem: Lampriger-]. Type genus: Lamprigera Motschulsky, 1853.
Subtribe
Lucidotina
Lacordaire, 1857
Lucidotides Lacordaire, 1857: 310 [stem: Lucidot-]. Type genus: Lucidota Laporte, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 184, as Lucidotae), generally accepted as in Gorham (1881: 29, as Lucidotides [treated as Latin]).
Pristolycini J. R. Winkler, 1953: 409 [stem: Pristolyc-]. Type genus: Pristolycus Gorham, 1883.
Pristolycini Kazantsev, 2010d: 205 [stem: Pristolyc-]. Type genus: Pristolycus Gorham, 1883. Comment: name proposed as new without reference to Pristolycini J. R. Winkler, 1953.
Subtribe
Photinina
LeConte, 1881
*Phosphaenaires Mulsant, 1862: 116 [stem: Phosphaen-]. Type genus: Phosphaenus Laporte, 1833. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant (1862).
Photini J. L. LeConte, 1881: 30 [stem: Photin-]. Type genus: Photinus Laporte, 1833. Comment: incorrect original stem formation, not in prevailing usage; this name is a senior homonym of Photininae Giglio-Tos, 1915 (type genus Photina Burmeister, 1838) in Mantodea; this case was submitted to the Commission to remove the homonymy by emending the stem of the mantid name (see Svenson and Branham 2007: 243).
Phosphaenides E. Olivier, 1884: 4 [stem: Phosphaen-]. Type genus: Phosphaenus Laporte, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Ganglbauer (1886: 280, as Phosphaenini).
Tribe
Pleotomini
Summers, 1874
Pleotomini Summers, 1874: 91 [stem: Pleotom-]. Type genus: Pleotomus J. L. LeConte, 1861.
Calyptocephalina Jakobson, 1911a: 667 [stem: Calyptocephal-]. Type genus: Calyptocephalus Gray, 1832.
Pleotomi Green, 1948: 68, in key [stem: Pleotom-]. Type genus: Pleotomus J. L. LeConte, 1861. Comment: family-group name proposed as new without reference to Pleotomini Summers, 1874.
Subfamily
Luciolinae
Lacordaire, 1857
Luciolides Lacordaire, 1857: 333 [stem: Luciol-]. Type genus: Luciola Laporte, 1833.
Tribe
Curtosini
McDermott, 1964
Curtosini McDermott, 1964: 47 [stem: Curtos-]. Type genus: Curtos Motschulsky, 1845. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Curt-).
Tribe
Luciolini
Lacordaire, 1857
Luciolides Lacordaire, 1857: 333 [stem: Luciol-]. Type genus: Luciola Laporte, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 184, as Luciolini), generally accepted as in Lawrence and Newton (1995: 859, as Luciolinae).
Subfamily
Photurinae
Lacordaire, 1857
Photurides Lacordaire, 1857: 338 [stem: Photur-]. Type genus: Photuris J. L. LeConte, 1851. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1906: 177, as Photurini), generally accepted as in Lawrence and Newton (1995: 859, as Photurinae); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Photurid-).
Lampyridae
incertae sedis
Cheguevariini Kazantsev, 2007: 370 [stem: Cheguevari-]. Type genus: Cheguevaria Kazantsev, 2007.
Family
Omethidae
LeConte, 1861
Omethes J. L. LeConte, 1861: 187 [stem: Ometh-]. Type genus: Omethes J. L. LeConte, 1861.
Subfamily
Omethinae
LeConte, 1861
Omethes J. L. LeConte, 1861: 187 [stem: Ometh-]. Type genus: Omethes J. L. LeConte, 1861.
Subfamily
Matheteinae
LeConte, 1881
Mathetei J. L. LeConte, 1881: 29 [stem: Mathete-]. Type genus: Matheteus J. L. LeConte, 1874.
Subfamily
Driloniinae
Crowson, 1972
Driloniinae Crowson, 1972: 58, in key [stem: Driloni-]. Type genus: Drilonius Kiesenwetter, 1874.
Family
Cantharidae
Imhoff, 1856 (1815)
Cantharidae Imhoff, 1856: [2] 69 [stem: Canthar-]. Type genus: Cantharis Linnaeus, 1758. Comment: this family-group name was used by many authors prior to Imhoff’s usage, these were based on a the misidentified type genus Cantharis (syn. of Lytta Fabricius, 1775) and are therefore not used as valid for this group (see Lawrence and Newton 1995:860); this case should be referred to the Commission to suppress any use of Cantharidae prior to Imhoff (1856); usage of the younger name Cantharidae Imhoff, 1856 over Telephoridae Leach, 1815 is conserved (Art. 40.2) (see Lawrence and Newton 1995: 860).
Subfamily
Cantharinae
Imhoff, 1856 (1815)
Cantharidae Imhoff, 1856: [2] 69 [stem: Canthar-]. Type genus: Cantharis Linnaeus, 1758. Comment: usage of the younger name over Telephorinae Leach, 1815 is conserved (Art. 40.2) (see Lawrence and Newton 1995: 860).
Tribe
Cantharini
Imhoff, 1856 (1815)
Telephorides Leach, 1815: 85 [stem: Telephor-]. Type genus: Telephorus Schaeffer, 1766 [syn. of Cantharis Linnaeus, 1758]. Comment: usage of Cantharidae/-inae/-ini over this name is conserved (Art. 40.2) (see Lawrence and Newton 1995: 860).
Cantharidae Imhoff, 1856: [2] 69 [stem: Canthar-]. Type genus: Cantharis Linnaeus, 1758. Comment: usage of the younger name over Telephorini Leach, 1815 is conserved (Art. 40.2) (see Lawrence and Newton 1995: 860); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Cantharid-).
Tribe
Podabrini
Gistel, 1856
Podabridae Gistel, 1856a: 385 [stem: Podabr-]. Type genus: Podabrus Dejean, 1833. Comment: family-group name attributed to J. L. LeConte (1881: 45) in recent literature although the same author had used the family-group name Podabri twenty years earlier (J. L. LeConte, 1861: 188).
Subfamily
Silinae
Mulsant, 1862
Siliaires Mulsant, 1862: 342 [stem: Sil-]. Type genus: Silis Charpentier, 1825.
Tribe
Silini
Mulsant, 1862
Siliaires Mulsant, 1862: 342 [stem: Sil-]. Type genus: Silis Charpentier, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gorham (1881: 91, as Silini), generally accepted as in Hansen (1996: 145, as Silinae).
Tribe
Tytthonyxini
Arnett, 1962
Tytthonyini Arnett, 1962a: 537 [stem: Tytthonyx-]. Type genus: Tytthonyx J. L. LeConte, 1851. Comment: spelling changed to Tytthonyxini by Wittmer (1970: 42); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Tytthonych-).
Subfamily
Dysmorphocerinae
Brancucci, 1980
Dysmorphocerinae Brancucci, 1980: 292 [stem: Dysmorphocer-]. Type genus: Dysmorphocerus Solier, 1849.
Subfamily
Malthininae
Kiesenwetter, 1852
Malthinen Kiesenwetter, 1852: 239 [stem: Malthin-]. Type genus: Malthinus Latreille, 1806.
Tribe
Malchinini
Brancucci, 1980
Malchinini Brancucci, 1980: 313 [stem: Malchin-]. Type genus: Malchinus Kiesenwetter, 1863.
Tribe
Malthinini
Kiesenwetter, 1852
Malthinen Kiesenwetter, 1852: 239 [stem: Malthin-]. Type genus: Malthinus Latreille, 1806. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 187, as Malthini), generally accepted as in Lawrence and Newton (1995: 860, as Malthininae); this family-group name was also used in the same year by Motschulsky (1852: 1, as Malthinides) but this vernacular name is unavailable (Art. 11.7.2) because it is not generally attributed to Motschulsky (1852).
Tribe
Malthodini
Böving and Craighead, 1931
Malthodinae Böving and Craighead, 1931: 48, in key [stem: Malthod-]. Type genus: Malthodes Kiesenwetter, 1852.
Malthodini Brancucci, 1980: 307 [stem: Malthod-]. Type genus: Malthodes Kiesenwetter, 1852. Comment: family-group name proposed as new without reference to Malthodinae Böving and Craighead, 1931.
Subfamily
Chauliognathinae
LeConte, 1861
Chauliognathini J. L. LeConte, 1861: 186 [stem: Chauliognath-]. Type genus: Chauliognathus Hentz, 1829.
Tribe
Chauliognathini
LeConte, 1861
Chauliognathini J. L. LeConte, 1861: 186 [stem: Chauliognath-]. Type genus: Chauliognathus Hentz, 1829.
Tribe
Ichthyurini
Champion, 1915
Ichthyurini Champion, 1915: 128 [stem: Ichthyur-]. Type genus: Ichthyurus Westwood, 1848.
Subfamily
Cydistinae
Paulus, 1972
Cydistinae Paulus, 1972a: 48, in key [stem: Cydist-]. Type genus: Cydistus Bourgeois, 1885.
Subfamily
Pterotinae
LeConte, 1861
Pterotini J. L. LeConte, 1861: 185 [stem: Pterot-]. Type genus: Pterotus J. L. LeConte, 1859.
Subfamily
Ototretinae
McDermott, 1964
Ototretinae McDermott, 1964: 11, in key [stem: Ototret-]. Type genus: Ototreta E. Olivier, 1900 [syn. of Drilaster Kiesenwetter, 1879].
Subfamily
Ototretadrilinae
Crowson, 1972
Ototretadrilinae Crowson, 1972: 55, in key [stem: Ototretadril-]. Type genus: Ototretadrilus Pic, 1921.
†Subfamily
Lasiosynidae
Kirejtshuk, Chang, Ren and Kun, 2010
Lasiosynidae Kirejtshuk et al., 2010: 68 [stem: Lasiosyn-]. Type genus: Lasiosyne Tan et al., 2007.
Superfamily
Derodontoidea
LeConte, 1861
Derodontidae J. L. LeConte, 1861: 100 [stem: Derodont-]. Type genus: Derodontus J. L. LeConte, 1861. Comment: name conserved over Nosodendroidea Erichson, 1846 (Art. 35.5).
Family
Derodontidae
LeConte, 1861
Derodontidae J. L. LeConte, 1861: 100 [stem: Derodont-]. Type genus: Derodontus J. L. LeConte, 1861. Comment: First Reviser found (Derodontidae J. L. LeConte, 1861 vs Peltasticidae J. L. LeConte, 1861) is Schenkling (1915: 3).
Subfamily
Peltasticinae
LeConte, 1861
Peltasticidae J. L. LeConte, 1861: 88 [stem: Peltastic-]. Type genus: Peltastica Mannerheim, 1852.
Subfamily
Derodontinae
LeConte, 1861
Derodontidae J. L. LeConte, 1861: 100 [stem: Derodont-]. Type genus: Derodontus J. L. LeConte, 1861.
Subfamily
Laricobiinae
Mulsant and Rey, 1864
Laricobiens Mulsant and Rey, 1864a: 374 [stem: Laricobi-]. Type genus: Laricobius Rosenhauer, 1846. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Ganglbauer (1899: 766, as Laricobiidae), generally accepted as in Hansen (1996: 146, as Laricobiinae).
Family
Nosodendridae
Erichson, 1846
Nosodendrini Erichson, 1846: 465 [stem: Nosodendr-]. Type genus: Nosodendron Latreille, 1804.
Family
Jacobsoniidae
Heller, 1926
Jacobsoniidae Heller, 1926: 127 [stem: Jacobsoni-]. Type genus: Jacobsonium Heller, 1926 [syn. of Sarothrias Grouvelle, 1918].
Sarothriidae Crowson, 1955: 75 [stem: Sarothri-]. Type genus: Sarothrias Grouvelle, 1918. Comment: replacement name for Jacobsoniidae Heller, 1926 because of synonymy of the type genus.
Derolathriinae Sen Gupta, 1979: 692 [stem: Derolathr-]. Type genus: Derolathrus Sharp, 1900. Comment: incorrect original stem formation, not in prevailing usage.
Superfamily
Bostrichoidea
Latreille, 1802
Bostrichini Latreille, 1802: 202 [stem: Bostrich-]. Type genus: Bostrichus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: First Reviser (Bostrichoidea Latreille, 1802 vs Ptinoidea Latreille, 1802) not determined, current usage maintained.
Family
Dermestidae
Latreille, 1804
Dermestini Latreille, 1804c: 142 [stem: Dermest-]. Type genus: Dermestes Linnaeus, 1758.
Subfamily
Dermestinae
Latreille, 1804
Dermestini Latreille, 1804c: 142 [stem: Dermest-]. Type genus: Dermestes Linnaeus, 1758.
Tribe
Dermestini
Latreille, 1804
Dermestini Latreille, 1804c: 142 [stem: Dermest-]. Type genus: Dermestes Linnaeus, 1758. Comment: published 7 March 1804; this family-group name was also used in the same year by Latreille (1804a [between 19 August and 17 September]: 233, as Dermestini).
Tribe
Marioutini
Jakobson, 1913
Marioutini Jakobson, 1913: 826 [stem: Mariout-]. Type genus: Mariouta Pic, 1899.
Rhopalosilphinae Arrow, 1929: 98 [stem: Rhopalosilph-]. Type genus: Rhopalosilpha Arrow, 1929. Comment: originally proposed as a subfamily of Silphidae.
Subfamily
Thorictinae
Agassiz, 1846
Thorictides Agassiz, 1846a: 162 [stem: Thorict-]. Type genus: Thorictus Germar, 1834.
Tribe
Thaumaphrastini
Anderson, 1949
Thaumaphrastinae W. H. Anderson, 1949: 127 [stem: Thaumaphrast-]. Type genus: Thaumaphrastus Blaisdell, 1927 [syn. of Thorictodes Reitter, 1875].
Tribe
Thorictini
Agassiz, 1846
Thorictides Agassiz, 1846a: 162 [stem: Thorict-]. Type genus: Thorictus Germar, 1834.
Subfamily
Orphilinae
LeConte, 1861
Orphili J. L. LeConte, 1861: 109 [stem: Orphil-]. Type genus: Orphilus Erichson, 1846.
Subfamily
Trinodinae
Casey, 1900
Trinodini Casey, 1900: 139 [stem: Trinod-]. Type genus: Trinodes Dejean, 1821.
†Tribe
Cretonodini
Kirejtshuk and Azar, 2009
Cretonodini Kirejtshuk and Azar, 2009: 121 [stem: Cretonod-]. Type genus: Cretonodes Kirejtshuk and Azar, 2009.
Tribe
Thylodriini
Semenov, 1909
Thelydriini Semenov, 1909: xxv [stem: Thylodri-]. Type genus: Thylodrias Motschulsky, 1839 [as Thelydrias, unjustified emendation of type genus name by Agassiz (1846b: 370), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Trichelodini Peacock, 1978: 344 [stem: Trichelod-]. Type genus: Trichelodes Carter, 1935.
Tribe
Trinodini
Casey, 1900
Trinodini Casey, 1900: 139 [stem: Trinod-]. Type genus: Trinodes Dejean, 1821.
Tribe
Trinoparvini
Háva, 2010
Trinoparvini Háva, 2010: 56 [stem: Trinoparv-]. Type genus: Trinoparvus Háva, 2004.
Subfamily
Attageninae
Laporte, 1840
Attagénites Laporte, 1840b: 35 [stem: Attagen-]. Type genus: Attagenus Latreille, 1802.
Tribe
Attagenini
Laporte, 1840
Attagénites Laporte, 1840b: 35 [stem: Attagen-]. Type genus: Attagenus Latreille, 1802. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 108, as Attageni), generally accepted as in Hansen (1996: 147, as Attageninae).
Tribe
Egidyellini
Semenov, 1914
Epidyellini Semenov, 1914: 15 [stem: Egidyell-]. Type genus: Egidyella Reitter, 1899 [as Epidyella, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Megatominae
Leach, 1815
Megatomida Leach, 1815: 94 [stem: Megatom-]. Type genus: Megatoma Herbst, 1791.
Tribe
Anthrenini
Gistel, 1848
Anthrenidae Gistel, 1848: [5] [stem: Anthren-]. Type genus: Anthrenus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
*Trogodermates Mulsant and Rey, 1867a: 120 [stem: Trogodermat-]. Type genus: Trogoderma Dejean, 1821. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Tribe
Megatomini
Leach, 1815
Megatomida Leach, 1815: 94 [stem: Megatom-]. Type genus: Megatoma Herbst, 1791.
Ctesiini Rees, 1943: 12 [stem: Ctesi-]. Type genus: Ctesias Stephens, 1830.
Family
Endecatomidae
LeConte, 1861
Endecatomini J. L. LeConte, 1861: 207 [stem: Endecatom-]. Type genus: Endecatomus Mellié, 1847.
Family
Bostrichidae
Latreille, 1802
Bostrichini Latreille, 1802: 202 [stem: Bostrich-]. Type genus: Bostrichus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subfamily
Dysidinae
Lesne, 1921
Dysididae Lesne, 1921b: 286 [stem: Dysid-]. Type genus: Dysides Perty, 1832.
Apoleoninae Gardner, 1933: 3 [stem: Apoleont-]. Type genus: Apoleon Gorham, 1885. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Polycaoninae
Lesne, 1896
Polycaoninae Lesne, 1896: 96 [stem: Polycaon-]. Type genus: Polycaon Laporte, 1836.
Subfamily
Bostrichinae
Latreille, 1802
Bostrichini Latreille, 1802: 202 [stem: Bostrich-]. Type genus: Bostrichus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Apatini
Billberg, 1820
Apatides Billberg, 1820a: 47 [stem: Apat-]. Type genus: Apate Fabricius, 1775 [nomen protectum (see Borowski and Węgrzynowicz 2009)]. Comment: this family-group name was also used in the same year by Billberg (1820b: 394, as Apatides).
Ligniperdidae Jacobi, 1906: 139 [stem: Ligniperd-]. Type genus: Ligniperda Pallas, 1772 [nomen oblitum (see Borowski and Węgrzynowicz 2009); syn. of Apate Fabricius, 1775].
Chileniidae Lesne, 1921b: 287 [stem: Chileni-]. Type genus: Chilenius Lesne, 1921.
Bostrychopsini Lesne, 1921b: 288 [stem: Bostrychopse-]. Type genus: Bostrychopsis Lesne, 1899 [syn. of Apate Fabricius, 1775]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Bostrichini
Latreille, 1802
Bostrichini Latreille, 1802: 202 [stem: Bostrich-]. Type genus: Bostrichus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Apatidini Bradley, 1930: 207, in key [stem: Apatid-]. Type genus: Apatides Casey, 1898.
Lichenophanini Portevin, 1931: 470, in key [stem: Lichenophan-]. Type genus: Lichenophanes Lesne, 1899.
Tribe
Dinapatini
Lesne, 1910
Dinapatinae Lesne, 1910: 471 [stem: Dinapat-]. Type genus: Dinapate G. H. Horn, 1886.
Tribe
Sinoxylini
Marseul, 1857
Sinoxylidae Marseul, 1857a: 107 [stem: Sinoxyl-]. Type genus: Sinoxylon Duftschmid, 1825.
Tribe
Xyloperthini
Lesne, 1921
Xyloperthini Lesne, 1921b: 288 [stem: Xyloperth-]. Type genus: Xylopertha Guérin-Méneville, 1845.
Subfamily
Psoinae
Blanchard, 1851
Psoitas Blanchard, 1851b: 434 [stem: Pso-]. Type genus: Psoa Herbst, 1797. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 208, as Psoini), generally accepted as in Ivie (2002: 241, as Psoinae).
Subfamily
Dinoderinae
Thomson, 1863
Dinoderina C. G. Thomson, 1863: 201 [stem: Dinoder-]. Type genus: Dinoderus Stephens, 1830.
Subfamily
Lyctinae
Billberg, 1820
Lyctides Billberg, 1820a: 48 [stem: Lyct-]. Type genus: Lyctus Fabricius, 1792.
Tribe
Lyctini
Billberg, 1820
Lyctides Billberg, 1820a: 48 [stem: Lyct-]. Type genus: Lyctus Fabricius, 1792.
Tribe
Trogoxylini
Lesne, 1921
Trogoxylini Lesne, 1921a: 231 [stem: Trogoxyl-]. Type genus: Trogoxylon J. L. LeConte, 1862.
Tristariini Lesne, 1921b: 287 [stem: Tristari-]. Type genus: Tristaria Reitter, 1878.
Subfamily
Euderiinae
Lesne, 1934
Euderiitae Lesne, 1934: 392 [stem: Euderi-]. Type genus: Euderia Broun, 1880.
Family
Ptinidae
Latreille, 1802
Ptiniores Latreille, 1802: 112 [stem: Ptin-]. Type genus: Ptinus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology (ICZN 1995a)]. Comment: Ptinidae Latreille, 1802 placed on the Official List of Family-Group Names in Zoology (ICZN 1995a); several subfamilies in Ptinidae contain tribes introduced in the literature by R. E. White (1982), those tribal names were not described originally and have not been made available subsequently; although some of White’s tribes have been used as available and valid names recently, e.g., Zahradník (2007), they are not available; in subfamilies containing tribes first proposed by R. E. White (1982), we have decided not to use tribal names at all in order to avoid adding to the confusion, the necessary phylogenetic work needed to establish a meaningful tribal classification in those subfamilies is currently under way (K. Philips pers. comm. 2009).
Subfamily
Eucradinae
LeConte, 1861
Eucradini J. L. LeConte, 1861: 202 [stem: Eucrad-]. Type genus: Eucrada J. L. LeConte, 1861.
Tribe
Eucradini
LeConte, 1861
Eucradini J. L. LeConte, 1861: 202 [stem: Eucrad-]. Type genus: Eucrada J. L. LeConte, 1861.
Tribe
Hedobiini
Mulsant and Rey, 1868
Hédobiaires Mulsant and Rey, 1868b: 23 [stem: Hedobi-]. Type genus: Hedobia Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1889 [Gatt.]: 116, as Hedobiini), generally accepted as in Philips (2002: 254, as Hedobiini).
Subfamily
Ptininae
Latreille, 1802
Ptiniores Latreille, 1802: 112 [stem: Ptin-]. Type genus: Ptinus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology (ICZN 1995a)]. Comment: Ptinidae Latreille, 1802 placed on the Official List of Family-Group Names in Zoology (ICZN 1995a).
Tribe
Gibbiini
Jacquelin du Val, 1860
Gibbiites Jacquelin du Val, 1860: 211 [stem: Gibbi-]. Type genus: Gibbium Scopoli, 1777. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kiesenwetter (1877: 44, as Gibbiini), generally accepted as in Bellés (1985: 13, as Gibbiinae).
Tribe
Meziini
Bellés, 1985
Meziini Bellés, 1985: 37, in key [stem: Mezi-]. Type genus: Mezium Curtis, 1828.
Tribe
Ptinini
Latreille, 1802
Ptiniores Latreille, 1802: 112 [stem: Ptin-]. Type genus: Ptinus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology (ICZN 1995a)]. Comment: Ptinidae Latreille, 1802 placed on the Official List of Family-Group Names in Zoology (ICZN 1995a).
Tribe
Sphaericini
Portevin, 1931
Sphaericini Portevin, 1931: 494 [stem: Sphaeric-]. Type genus: Sphaericus Wollaston, 1854.
Ptininae
incertae sedis
Gnostidae Gemminger and Harold, 1868: 700 [stem: Gnost-]. Type genus: Gnostus Westwood, 1855.
Ectrephidae Wasmann, 1894: 121 [stem: Ectreph-]. Type genus: Ectrephes Pascoe, 1866.
Subfamily
Dryophilinae
Gistel, 1848
Dryophilidae Gistel, 1848: [6] [stem: Dryophil-]. Type genus: Dryophilus Chevrolat, 1832.
Tribe
Dryophilini
Gistel, 1848
Dryophilidae Gistel, 1848: [6] [stem: Dryophil-]. Type genus: Dryophilus Chevrolat, 1832. Comment: family-group name previously attributed to J. L. LeConte (1861: 205) in the literature.
Dryobiadae Gistel, 1856a: 368 [stem: Dryobi-]. Type genus: Dryobia Gistel, 1856 [syn. of Dryophilus Chevrolat, 1832]. Comment: senior homonym of Dryobiini Arnett, 1962 (type genus Dryobius J. L. LeConte, 1850) used as valid in Cerambycidae; nomen oblitum (see Bousquet et al. 2009: 45); incorrect original stem formation, not in prevailing usage.
Tribe
Ptilineurini
Böving, 1927
Ptilineurini Böving, 1927a: 56 [stem: Ptilineur-]. Type genus: Ptilineurus Reitter, 1901.
Subfamily
Ernobiinae
Pic, 1912
Cosmoceroideos Solier, 1849: 476 [stem: Cosmocer-]. Type genus: Cosmocerus Solier, 1849 [preoccupied genus name, not Cosmocerus Guérin-Méneville, 1844 [Coleoptera: Cerambycidae]; syn. of Cerocosmus Gemminger, 1873]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; Lawrence and Newton (1995: 864) used the latinized form Cosmocerinae and treated Solier’s name as available therefore we treat this name as available but permanently invalid.
Ernobiinae Pic, 1912: 12 [stem: Ernobi-]. Type genus: Ernobius C. G. Thomson, 1859. Comment: First Reviser found (Ernobiinae Pic, 1912 vs Cerocosminae Pic, 1912) is White (1974: 419).
Cerocosminae Pic, 1912: 45 [stem: Cerocosm-]. Type genus: Cerocosmus Gemminger, 1873.
Xestobiini Böving, 1927a: 56 [stem: Xestobi-]. Type genus: Xestobium Motschulsky, 1845.
*Ozognathini R. E. White, 1982: 2 [stem: Ozognath-]. Type genus: Ozognathus J. L. LeConte, 1861. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subfamily
Anobiinae
Fleming, 1821
Anobiumedae Fleming, 1821: 50 [stem: Anobi-]. Type genus: Anobium Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1976)]. Comment: incorrect original stem formation, not in prevailing usage.
*Euceratocerini R. E. White, 1982: 7 [stem: Euceratocer-]. Type genus: Euceratocerus J. L. LeConte, 1874. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Colposternini R. E. White, 1982: 9 [stem: Colpostern-]. Type genus: Colposternus Fall, 1905. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Gastrallini R. E. White, 1982: 9 [stem: Gastrall-]. Type genus: Gastrallus Jacquelin du Val, 1860. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Hadrobregmini R. E. White, 1982: 15 [stem: Hadrobregm-]. Type genus: Hadrobregmus C. G. Thomson, 1859. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Nicobiini R. E. White, 1982: 10 [stem: Nicobi-]. Type genus: Nicobium J. L. LeConte, 1861. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Stegobiini R. E. White, 1982: 11 [stem: Stegobi-]. Type genus: Stegobium Motschulsky, 1860. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subfamily
Ptilininae
Shuckard, 1839
Ptilinidae Shuckard, 1839b: 45 [stem: Ptilin-]. Type genus: Ptilinus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Sclerasteidae Gistel, 1856a: 368 [stem: Sclerast-]. Type genus: Sclerastes Gistel, 1856 [this genus originally included “pectinicornis” and “costatus Gy”, we here chose Ptilinus costatus Gyllenhal, 1827 as the type of Sclerastes; syn. nov. of Ptilinus Geoffroy, 1762]. Comment: syn. nov.; incorrect original stem formation, not in prevailing usage.
Subfamily
Alvarenganiellinae
Viana and Martínez, 1971
Alvarenganiellinae Viana and Martínez, 1971: 121 [stem: Alvarenganiell-]. Type genus: Alvarenganiella Viana and Martínez, 1971.
Subfamily
Xyletininae
Gistel, 1848
Xyletinidae Gistel, 1848: [6] [stem: Xyletin-]. Type genus: Xyletinus Latreille, 1810 [placed on the Official List of Generic Names in Zoology (ICZN 1971)].
Tribe
Lasiodermini
Böving, 1927
Lasiodermini Böving, 1927a: 56 [stem: Lasioderm-]. Type genus: Lasioderma Stephens, 1835. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Lasiodermat-).
Tribe
Metholcini
Zahradník, 2009
Metholcini Zahradník, 2009: 180 [stem: Metholc-]. Type genus: Metholcus Jacquelin du Val, 1860.
Tribe
Xyletinini
Gistel, 1848
Xyletinidae Gistel, 1848: [6] [stem: Xyletin-]. Type genus: Xyletinus Latreille, 1810 [placed on the Official List of Generic Names in Zoology (ICZN 1971)].
Vrilettini Böving, 1927a: 56 [stem: Vrillett-]. Type genus: Vrilletta J. L. LeConte, 1874. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Dorcatominae
Thomson, 1859
Dorcatomina C. G. Thomson, 1859: 90 [stem: Dorcatom-]. Type genus: Dorcatoma Herbst, 1791 [the original spelling Dorkatoma was placed on the Official Index of Rejected and Suppressed Generic Names in Zoology, Dorcatoma was chosen as correct original spelling and placed on the Official List of Generic Names in Zoology (ICZN 1995b)].
Caenocarini Böving, 1927a: 57 [stem: Caenocar-]. Type genus: Caenocara C. G. Thomson, 1859. Comment: also misspelled as Coenecarini (page 56) in the original publication.
*Calymmaderini R. E. White, 1982: 22 [stem: Calymmader-]. Type genus: Calymmaderus Solier, 1849. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Cryptoramorphini R. E. White, 1982: 25 [stem: Cryptoramorph-]. Type genus: Cryptoramorphus R. E. White, 1966. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Petaliini R. E. White, 1982: 23 [stem: Petali-]. Type genus: Petalium J. L. LeConte, 1861. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); the tribe Petaliini Tillyard, 1917 in Odonata (type genus Petalia Hagen, 1854, a junior homonym of Petalia Gray, 1838 [Mammalia]) is available but permanently invalid because it is based on a preoccupied type genus.
*Prothecini R. E. White, 1982: 25 [stem: Prothec-]. Type genus: Protheca J. L. LeConte, 1865. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subfamily
Mesocoelopodinae
Mulsant and Rey, 1864
Mésocoelopaires Mulsant and Rey, 1864b: 311 [stem: Mesocoelopod-]. Type genus: Mesocoelopus Jacquelin du Val, 1860.
Tribe
Tricorynini
White, 1971
Tricoryninae R. E. White, 1971: 1301 [stem: Tricoryn-]. Type genus: Tricorynus G. R. Waterhouse, 1849.
Tribe
Mesocoelopodini
Mulsant and Rey, 1864
Mésocoelopaires Mulsant and Rey, 1864b: 311 [stem: Mesocoelopod-]. Type genus: Mesocoelopus Jacquelin du Val, 1860. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kiesenwetter (1877: 153, as Mesocoelopini [incorrect stem formation]), generally accepted as in Philips (2002: 258, as Mesocoelopodinae); incorrect original stem formation, not in prevailing usage.
Mesothini Portevin, 1931: 489 [stem: Mesothet-]. Type genus: Mesothes Mulsant and Rey, 1864. Comment: incorrect original stem formation, not in prevailing usage.
Ptinidae
incertae sedis
Fabiinae Martínez and Viana, 1964: 7 [stem: Fabi-]. Type genus: Fabia Martínez and Viana, 1964.
Superfamily
Lymexyloidea
Fleming, 1821
Lymoxylonidae Fleming, 1821: 49 [stem: Lymexyl-]. Type genus: Lymexylon Fabricius, 1775. Comment: Lymexyloidea given precedence for superfamily name over Hylecoetoidea Germar, 1818 (Art. 35.5).
Family
Lymexylidae
Fleming, 1821
Lymoxylonidae Fleming, 1821: 49 [stem: Lymexyl-]. Type genus: Lymexylon Fabricius, 1775. Comment: Lymexylidae given precedence for family name over Hylecoetidae Germar, 1818 (Art. 35.5).
Subfamily
Hylecoetinae
Germar, 1818
Hylecoeti Germar, 1818: 344 [stem: Hylecoet-]. Type genus: Hylecoetus Latreille, 1806. Comment: name previously attributed to Gistel (1856a).
Subfamily
Lymexylinae
Fleming, 1821
Lymoxylonidae Fleming, 1821: 49 [stem: Lymexyl-]. Type genus: Lymexylon Fabricius, 1775 [as Lymoxylon, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: published in July 1821; incorrect original stem formation, not in prevailing usage; this family-group name was also used in the same year by Fischer von Waldheim (1821 [“31 December”]: 37, as Lymexyla).
Subfamily
Atractocerinae
Laporte, 1840
Atractocérites Laporte, 1840a: 290 [stem: Atractocer-]. Type genus: Atractocerus Palisot de Beauvois, 1802. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Brues and Melander (1932: 23, as Atractoceridae), generally accepted as in Cuccodoro (2007: 363, as Atractocerinae).
Subfamily
Melittommatinae
Wheeler, 1986
Melittomminae Q. D. Wheeler, 1986: 160 [stem: Melittommat-]. Type genus: Melittomma Murray, 1867. Comment: incorrect original stem formation, not in prevailing usage.
Superfamily
Cleroidea
Latreille, 1802
Clerii Latreille, 1802: 110 [stem: Cler-]. Type genus: Clerus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)]. Comment: First Reviser (Cleroidea Latreille, 1802 vs Trogossitoidea Latreille, 1802) not determined, current usage maintained.
Family
Phloiophilidae
Kiesenwetter, 1863
Phloeophilini Kiesenwetter, 1863: 626, in key [stem: Phloiophil-]. Type genus: Phloiophilus Stephens, 1830 [as Phloeophilus, unjustified emendation of type genus name by Agassiz (1846b: 286), not in prevailing usage; Phloeophilus Agassiz, 1846 is a junior homonym of Phloeophilus Schönherr, 1833 [Coleoptera: Anthribidae]]. Comment: incorrect original stem formation, not in prevailing usage; using the original spelling would place Phloeophilini Lacordaire, 1865 [Coleoptera: Anthribidae] into homonymy with this name.
Family
Trogossitidae
Latreille, 1802
Trogossitarii Latreille, 1802: 159 [stem: Trogossit-]. Type genus: Trogossita A. G. Olivier, 1790 [syn. of Tenebroides Piller and Mitterpacher, 1783].
Subfamily
Peltinae
Latreille, 1806
Peltides Latreille, 1806: 8 [stem: Pelt-]. Type genus: Peltis Kugelann, 1792 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Ancyronini
Kolibáč, 2006
Ancyronini Kolibáč, 2006: 127 [stem: Ancyron-]. Type genus: Ancyrona Reitter, 1876.
Tribe
Colydiopeltini
Kolibáč, 2006
Colydiopeltini Kolibáč, 2006: 126 [stem: Colydiopelt-]. Type genus: Colydiopeltis Ślipiński, 1992.
Tribe
Decamerini
Crowson, 1964
Decamerinae Crowson, 1964: 287 [stem: Decamer-]. Type genus: Decamerus Solier, 1849.
Tribe
Lophocaterini
Crowson, 1964
nomen protectum
Lycoptini Casey, 1890: 311 [stem: Lycopt-]. Type genus: Lycoptis Casey, 1890. Comment: nomen oblitum (see Appendix 1).
Lophocaterinae Crowson, 1964: 297 [stem: Lophocater-]. Type genus: Lophocateres Olliff, 1883. Comment: nomen protectum (see Appendix 1).
Tribe
Peltini
Latreille, 1806
Peltides Latreille, 1806: 8 [stem: Pelt-]. Type genus: Peltis Kugelann, 1792 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: name previously attributed to Kirby (1837).
Ostomini Harold, 1876: 169 [stem: Ostomat-]. Type genus: Ostoma Laicharting, 1781. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Thymalini
Léveillé, 1888
*Thymalites Blanchard, 1845a: 277 [stem: Thymal-]. Type genus: Thymalus Latreille, 1802. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845a).
Thymalini Léveillé, 1888: 444 [stem: Thymal-]. Type genus: Thymalus Latreille, 1802.
Rentoniinae Crowson, 1966: 120 [stem: Rentoni-]. Type genus: Rentonium Crowson, 1966.
Protopeltini Crowson, 1966: 120 [stem: Protopelt-]. Type genus: Protopeltis Crowson, 1964.
†Meligethiellinae Kirejtshuk and Ponomarenko, 1990: 79 [stem: Meligethiell-]. Type genus: Meligethiella L. N. Medvedev, 1969.
Subfamily
Trogossitinae
Latreille, 1802
Trogossitarii Latreille, 1802: 159 [stem: Trogossit-]. Type genus: Trogossita A. G. Olivier, 1790 [syn. of Tenebroides Piller and Mitterpacher, 1783].
Tribe
Calityini
Reitter, 1922
Calityni Reitter, 1922a: 66 [stem: Cality-]. Type genus: Calitys C. G. Thomson, 1859. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Egoliini
Lacordaire, 1854
Égoliides Lacordaire, 1854b: 334 [stem: Egoli-]. Type genus: Egolia Erichson, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Ganglbauer (1899: 419, as Egoliides [treated as Latin]), generally accepted as in Kolibáč (2006: 119, as Egoliini).
Tribe
Gymnochilini
Lacordaire, 1854
Gymnochilides Lacordaire, 1854b: 344 [stem: Gymnochil-]. Type genus: Gymnochila Klug, 1844. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1858: 43, as Gymnochilitae), generally accepted as in Burakowski et al. (1986: 118, as Gymnochilinae).
Leperini Reitter, 1876a: 55 [stem: Leperin-]. Type genus: Leperina Erichson, 1844. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Larinotini
Ślipiński, 1992
Larinotinae Ślipiński, 1992: 443 [stem: Larinot-]. Type genus: Larinotus Carter and Zeck, 1937.
†Tribe
Lithostomatini
Kolibáč and Huang, 2008
Lithostomini Kolibáč and Huang, 2008: 142 [stem: Lithostomat-]. Type genus: Lithostoma Martynov, 1926. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Trogossitini
Latreille, 1802
Trogossitarii Latreille, 1802: 159 [stem: Trogossit-]. Type genus: Trogossita A. G. Olivier, 1790 [syn. of Tenebroides Piller and Mitterpacher, 1783].
Nemosomida Leach, 1815: 110 [stem: Nemozomat-]. Type genus: Nemozoma Latreille, 1804 [as Nemosoma, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Temnochilini Léveillé, 1888: 432 [stem: Temnoscheil-]. Type genus: Temnoscheila Westwood, 1830 [as Temnochila, unjustified emendation of type genus name by Erichson (1844: 449), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tenebrioidini Ganglbauer, 1899: 420 [stem: Tenebroid-]. Type genus: Tenebroides Piller and Mitterpacher, 1783 [as Tenebrioides, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Family
Chaetosomatidae
Crowson, 1952
Chaetosomatidae Crowson, 1952: 66 [stem: Chaetosomat-]. Type genus: Chaetosoma Westwood, 1851 [this genus is a junior homonym of Chaetosoma Chevrolat, 1843 in Cerambycidae; Chaetosoma Chevrolat, 1843 is also a senior objective synonym of the well-established name Apodasya Pascoe, 1863]. Comment: this name is a junior homonym of Chaetosomatidae Claus, 1872 (type genus Chaetosoma Claparède, 1863, which is a junior homonym of Chaetosoma Westwood, 1851) in Nematoda; the nematode family-group name is permanently invalid (Art. 39) because it is based on a junior homonym, the valid name for this nematode family is Draconematidae Filipjev, 1918; furthermore, the type genus of Chaetosomatidae Crowson is a junior homonym of Chaetosoma Chevrolat, 1843 in Cerambycidae; an application was recently submitted by Bousquet and Bouchard (2010) to suppress Chaetosoma Chevrolat, 1843 for the Principles of Priority and Homonymy and Chaetosomatidae Claus, 1872 for the Principle of Priority, therefore conserving the names Chaetosomatidae Crowson and Chaetosoma Westwood (see Appendix 6).
Family
Metaxinidae
Kolibáč, 2004
Metaxinidae Kolibáč, 2004: 247 [stem: Metaxin-]. Type genus: Metaxina Broun, 1909.
Family
Thanerocleridae
Chapin, 1924
Thaneroclerinae Chapin, 1924: 251 [stem: Thanerocler-]. Type genus: Thaneroclerus Westwood, 1838.
Subfamily
Zenodosinae
Kolibáč, 1992
Zenodosini Kolibáč, 1992: 307 [stem: Zenodos-]. Type genus: Zenodosus Wolcott, 1910.
Subfamily
Thaneroclerinae
Chapin, 1924
Thaneroclerinae Chapin, 1924: 251 [stem: Thanerocler-]. Type genus: Thaneroclerus Westwood, 1838.
Tribe
Isoclerini
Kolibáč, 1992
Isoclerina Kolibáč, 1992: 315 [stem: Isocler-]. Type genus: Isoclerus Lewis, 1892.
Tribe
Thaneroclerini
Chapin, 1924
Thaneroclerinae Chapin, 1924: 251 [stem: Thanerocler-]. Type genus: Thaneroclerus Westwood, 1838.
Tribe
Viticlerini
Winkler, 1982
Viticlerini J. R. Winkler, 1982: 529, in key [stem: Viticler-]. Type genus: Viticlerus Miyatake, 1977.
Family
Cleridae
Latreille, 1802
Clerii Latreille, 1802: 110 [stem: Cler-]. Type genus: Clerus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)]. Comment: see Opitz (2010) for an alternative subfamilial classification within this family.
Subfamily
Tillinae
Fischer von Waldheim, 1813
Tillii Fischer von Waldheim, 1813: 232 [stem: Till-]. Type genus: Tillus A. G. Olivier, 1790. Comment: family-group name previously attributed to Leach (1815).
Cylidrina Reitter, 1894: 38 [stem: Cylidr-]. Type genus: Cylidrus Latreille, 1829.
*Monophyllini Böving and Craighead, 1931: 78 [stem: Monophyll-]. Type genus: Monophylla Spinola, 1841. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subfamily
Hydnocerinae
Spinola, 1844
Hydnocéroïdes Spinola, 1844: Tableau générique (3) [stem: Hydnocer-]. Type genus: Hydnocera Newman, 1838 [syn. of Phyllobaenus Dejean, 1833].
Tribe
Callimerini
Kolibáč, 1998
Callimerini Kolibáč, 1998: 165 [stem: Callimer-]. Type genus: Callimerus Gorham, 1876.
Tribe
Hydnocerini
Spinola, 1844
Hydnocéroïdes Spinola, 1844: Tableau générique (3) [stem: Hydnocer-]. Type genus: Hydnocera Newman, 1838 [syn. of Phyllobaenus Dejean, 1833]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Desmarest (1857: 263, as Hydnoceridae), generally accepted as in Kolibáč (1998: 127, as Hydnocerinae).
Phyllobénides Lacordaire, 1857: 466 [stem: Phyllobaen-]. Type genus: Phyllobaenus Dejean, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by and generally accepted as in Gorham (1876: 59, as Phyllobenides [treated as Latin]); incorrect original stem formation, not in prevailing usage.
Tribe
Lemidiini
Kolibáč, 1998
Lemidiini Kolibáč, 1998: 183 [stem: Lemidi-]. Type genus: Lemidia Spinola, 1841.
Subfamily
Clerinae
Latreille, 1802
Clerii Latreille, 1802: 110 [stem: Cler-]. Type genus: Clerus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)].
Notoxii Sturm, 1826: 38 [stem: Notox-]. Type genus: Notoxus sensu Fabricius, 1775 [not Notoxus Geoffroy, 1762; syn. of Opilo Latreille, 1802]. Comment: based on a misidentified type genus; an application should be submitted to the Commission to suppress this name for the Principles of Priority and Homonymy (Art. 65.2.1) since Notoxinae Stephens, 1829 is currently used as valid in Anthicidae.
Prioceridae Laporte, 1836: 33 [stem: Priocer-]. Type genus: Priocera Kirby, 1819.
*Trichodites Blanchard, 1845b: 84 [stem: Trichod-]. Type genus: Trichodes Herbst, 1792 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845b).
Opilonidae Gistel, 1848: [6] [stem: Opilon-]. Type genus: Opilo Latreille, 1802.
Thanasimiidae Gistel, 1848: [6] [stem: Thanasim-]. Type genus: Thanasimus Latreille, 1806. Comment: incorrect original stem formation, not in prevailing usage.
Dendroplaneteidae Gistel, 1856a: 395 [stem: Dendroplanet-]. Type genus: Dendroplanetes Gistel, 1856 [syn. of Opilo Latreille, 1802]. Comment: also spelled Dondroplaneteidae in the same publication on page 368; incorrect original stem formation, not in prevailing usage.
Trichodini Portevin, 1931: 457, in key [stem: Trichod-]. Type genus: Trichodes Herbst, 1792 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)]. Comment: the older name originally proposed as Trichodina Maitland, 1851 (type genus Trichoda Müller, 1773) is available in Protozoa: Ciliophora; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Dieropsinae J. R. Winkler, 1964: 317 [stem: Dieropse-]. Type genus: Dieropsis Gahan, 1908. Comment: incorrect original stem formation, not in prevailing usage.
Cleropiestinae J. R. Winkler, 1980: 437 [stem: Cleropiest-]. Type genus: Cleropiestus Fairmaire, 1889.
Anthicoclerinae Opitz, 2010: 58 [stem: Anthicocler-]. Type genus: Anthicoclerus Schenkling, 1906.
Subfamily
Korynetinae
Laporte, 1836
Corynetidae Laporte, 1836: 34 [stem: Korynet-]. Type genus: Korynetes Herbst, 1792 [as Corynetes, unjustified emendation of type genus name by Paykull (1798: 274), not in prevailing usage; Korynetes Herbst, 1792 placed on the Official List of Generic Names in Zoology and Corynetes Paykull, 1798 placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1961c)]. Comment: incorrect original stem formation, not in prevailing usage.
Ichnoïdes Spinola, 1841: 71 [stem: Ichne-]. Type genus: Ichnea Laporte, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 193, as Ichneoidae), treated as available by Opitz and Herman (2009: 183, as Ichneinae); nomen oblitum, Epiphloeinae Kuwert, 1893 conserved as valid subfamily over this name by Opitz and Herman (2009).
Platynoptéroïdes Spinola, 1844: Tableau générique (4) [stem: Platynopter-]. Type genus: Platynoptera Chevrolat, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Desmarest (1857: 269, as Platynopteridae).
Necrobiaeidae Gistel, 1848: [6] [stem: Necrobi-]. Type genus: Necrobia A. G. Olivier, 1795 [placed on the Official List of Generic Names in Zoology (ICZN 1961c)]. Comment: incorrect original stem formation, not in prevailing usage.
Enopliidae Gistel, 1848: [6] [stem: Enopli-]. Type genus: Enoplium Latreille, 1802.
Tarsosténites Jacquelin du Val, 1860: 198 [stem: Tarsosten-]. Type genus: Tarsostenus Spinola, 1844. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Böving and Craighead (1931: 57, as Tarsosteninae), generally accepted as in Opitz (2002: 279, as Tarsostenninae [incorrect stem formation]).
Epiphlöinen Kuwert, 1893: 492 [stem: Epiphloe-]. Type genus: Epiphloeus Spinola, 1841. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Schenkling (1910: 113, as Epiphloeini), generally accepted as in Opitz and Herman (2009: 183, as Epiphloeinae); nomen protectum (see Opitz and Herman (2009: 184)); although Kuwert apparently originally published his name as a synonym of Phyllobaenides Lacordaire, 1857, the name Epiphloeini was treated as valid by several authors, e.g., Chapin (1924: 165), before 1961 which made the name available (Art. 11.6.1) (see Opitz and Herman 2009: 184).
Dermestoidini Jakobson, 1911a: 719 [stem: Dermestoid-]. Type genus: Dermestoides Schaeffer, 1771.
Orthopleurinae Böving and Craighead, 1931: 56, in key [stem: Orthopleur-]. Type genus: Orthopleura Spinola, 1844.
Neorthopleurinae Opitz, 2009: 138 [stem: Neorthopleur-]. Type genus: Neorthopleura Barr, 1976.
Peloniinae Opitz, 2010: 97 [stem: Peloni-]. Type genus: Pelonium Spinola, 1844.
Family
Acanthocnemidae
Crowson, 1964
Acanthocneminae Crowson, 1964: 317, in key [stem: Acanthocnem-]. Type genus: Acanthocnemus Perris, 1866.
Family
Phycosecidae
Crowson, 1952
Phycosecidae Crowson, 1952: 117 [stem: Phycosec-]. Type genus: Phycosecis Pascoe, 1875. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Phycosecid-).
Family
Prionoceridae
Lacordaire, 1857
Prionocérides Lacordaire, 1857: 411 [stem: Prionocer-]. Type genus: Prionocerus Perty, 1831. Comment: Prionocérites was used earlier by Laporte (1840a: 275) but this name was treated as a lapsus calami for Priocérites Laporte (1834b: 33) by Lawrence and Newton (1995: 871).
Tribe
Lobonychini
Majer, 1987
Lobonychini Majer, 1987: 790, in key [stem: Lobonych-]. Type genus: Lobonyx Jacquelin du Val, 1859.
Tribe
Prionocerini
Lacordaire, 1857
Prionocérides Lacordaire, 1857: 411 [stem: Prionocer-]. Type genus: Prionocerus Perty, 1831. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Ritsema (1884: 494, as Prionocerini), generally accepted as in M. C. Thomas (2008: 445, as Prionoceridae); the earlier usage of Prionocerites by Laporte (1840a: 285) was in error for Priocerites in Cleridae: Clerinae (see Lawrence and Newton 1995: 871); Prionocerini Savchenko, 1966 (type genus Prionocera Loew, 1844) is available in Diptera; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Family
Mauroniscidae
Majer, 1995
Mauroniscidae Majer, 1995a: 340 [stem: Mauronisc-]. Type genus: Mauroniscus Bourgeois, 1911. Comment: published 30 April 1995; this family-group name was also used in the same year by Majer (1995b [1 December]: 57, as Mauroniscidae).
Family
Melyridae
Leach, 1815
Melyrides Leach, 1815: 87 [stem: Melyr-]. Type genus: Melyris Fabricius, 1775.
Subfamily
Rhadalinae
LeConte, 1861
Rhadalini J. L. LeConte, 1861: 194 [stem: Rhadal-]. Type genus: Rhadalus J. L. LeConte, 1852.
Haplocnémates Mulsant and Rey, 1868a: 181 [stem: Aplocnem-]. Type genus: Aplocnemus Stephens, 1830 [as Haplocnemus, unjustified emendation of type genus name by Agassiz (1846b: 29), not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Crowson (1964: 316, as Haplocneminae [incorrect stem formation]), generally accepted as in Majer (1987: 800, as Aplocnemini); incorrect original stem formation, not in prevailing usage.
Microjulistini Majer, 1987: 797, in key [stem: Microjulist-]. Type genus: Microjulistus Reitter, 1889.
Pelecophorini Majer, 1987: 797, in key [stem: Pelecophor-]. Type genus: Pelecophora Lepeletier and Audinet-Serville, 1825.
Subfamily
Melyrinae
Leach, 1815
Melyrides Leach, 1815: 87 [stem: Melyr-]. Type genus: Melyris Fabricius, 1775.
Tribe
Arthrobrachini
Majer, 1987
Arthrobrachini Majer, 1987: 793, in key [stem: Arthrobrach-]. Type genus: Arthrobrachus Solier, 1849.
Tribe
Astylini
Pic, 1929
Astylini Pic, 1929: 1 [stem: Astyl-]. Type genus: Astylus Laporte, 1836.
Tribe
Cerallini
Pic, 1929
Cerallini Pic, 1929: 13 [stem: Cerall-]. Type genus: Cerallus Jacquelin du Val, 1859.
Tribe
Melyrini
Leach, 1815
Melyrides Leach, 1815: 87 [stem: Melyr-]. Type genus: Melyris Fabricius, 1775. Comment: published April 1815; this family-group name was also used in the same year by Rafinesque (1815 [between April and 21 July]: 110, as Melyria).
Zygiidae Jakobson, 1911a: 687 [stem: Zygi-]. Type genus: Zygia Fabricius, 1775 [syn. of Melyris Fabricius, 1775].
Subfamily
Dasytinae
Laporte, 1840
Dasydites Laporte, 1840a: 280 [stem: Dasyt-]. Type genus: Dasytes Paykull, 1799.
Tribe
Chaetomalachiini
Majer, 1987
Chaetomalachiini Majer, 1987: 799, in key [stem: Chaetomalachi-]. Type genus: Chaetomalachius Kraatz, 1882.
Tribe
Danaceini
Thomson, 1859
Danacaeina C. G. Thomson, 1859: 108 [stem: Danace-]. Type genus: Danacea Laporte, 1838 [as Danacaea, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Amauronoinini Majer, 1987: 799, in key [stem: Amauronioid-]. Type genus: Amauronioides Champion, 1923 [syn. of Pseudamauronia Pic, 1915]. Comment: incorrect original stem formation, not in prevailing usage.
Danacaeomimini Majer, 1987: 799, in key [stem: Danacaeomim-]. Type genus: Danacaeomimus Champion, 1922.
Tribe
Dasytini
Laporte, 1840
Dasydites Laporte, 1840a: 280 [stem: Dasyt-]. Type genus: Dasytes Paykull, 1799. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 117, as Dasytetoidae [incorrect stem formation]), generally accepted as in Pic (1937: 1, as Dasytinae); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Dasytet-).
Colpothisidae Gistel, 1848: [11] [stem: Colpoth-]. Type genus: Colpothis Gistel, 1848 [syn. of Dasytes Paykull, 1799]. Comment: incorrect original stem formation, not in prevailing usage.
*Hénicopaires Mulsant and Rey, 1868a: 264 [stem: Enicopod-]. Type genus: Enicopus Stephens, 1830 [as Henicopus, unjustified emendation of type genus name by Agassiz (1846b: 138), not in prevailing usage]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant and Rey (1868a); incorrect original stem formation, not in prevailing usage.
Henicopini Escalera, 1927: 7 [stem: Enicopod-]. Type genus: Enicopus Stephens, 1830 [as Henicopus, unjustified emendation of type genus name by Agassiz (1846b: 138), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Gietellini
Constantin and Menier, 1987
Gietellinae Constantin and Menier, 1987: 62 [stem: Gietell-]. Type genus: Gietella Constantin and Menier, 1987.
Tribe
Listrini
Majer, 1990
Listrini Majer, 1990: 371 [stem: Listr-]. Type genus: Listrus Motschulsky, 1859.
Subfamily
Malachiinae
Fleming, 1821
Malachiusidae Fleming, 1821: 50 [stem: Malachi-]. Type genus: Malachius Fabricius, 1775.
Tribe
Amalthocini
Majer, 2002
Amalthocinae Majer, 2002: 186 [stem: Amalthoc-]. Type genus: Amalthocus Fairmaire, 1886.
Tribe
Attalomimini
Majer, 1995
Attalomimidae Majer, 1995a: 378 [stem: Attalomim-]. Type genus: Attalomimus Wittmer, 1976.
Tribe
Carphurini
Champion, 1923
Carphurinae Champion, 1923: 2 [stem: Carphur-]. Type genus: Carphurus Erichson, 1840.
Tribe
Lemphini
Wittmer, 1976
Lemphini Wittmer, 1976: 427 [stem: Lemph-]. Type genus: Lemphus Erichson, 1840.
Tribe
Malachiini
Fleming, 1821
Malachiusidae Fleming, 1821: 50 [stem: Malachi-]. Type genus: Malachius Fabricius, 1775. Comment: incorrect original stem formation, not in prevailing usage.
Tamulidae Gistel, 1848: [11] [stem: Tamul-]. Type genus: Tamulus Gistel, 1848 [syn. of Malachius Fabricius, 1775].
Polycystophoridae Gistel, 1856a: 385 [stem: Polycystophor-]. Type genus: Polycystophorus Gistel, 1856 [this genus was included by Gistel in his new family Polycystophoridae along with one genus now classified in Cantharidae (Malthinus Latreille) and four genera of Malachiinae (Anthocomus Erichson, Ebaeus Erichson, Charopus Erichson and Troglops Erichson); several available species names were originally included in Polycystophorus (Gistel, 1856a: 385), we hereby select Cantharis aeneus Linnaeus, 1758 as the type species; syn. nov. of Malachius Fabricius, 1775]. Comment: syn. nov.
Apalochraires Mulsant and Rey, 1867b: 19 [stem: Apalochr-]. Type genus: Apalochrus Erichson, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Houlbert (1922a: 250, as Apalochrini).
*Anthocomates Mulsant and Rey, 1867b: 127 [stem: Anthocom-]. Type genus: Anthocomus Erichson, 1840. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Troglopates Mulsant and Rey, 1867b: 281 [stem: Troglop-]. Type genus: Troglops Erichson, 1840. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Caulautaires Abeille de Perrin, 1890: 244 [stem: Colot-]. Type genus: Colotes Erichson, 1840 [as Caulautes, unjustified emendation of type genus name by Abeille de Perrin (1890), not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Bertkau (1891: 304, as Colotini), generally accepted as in Hansen (1996: 152, as Colotini); incorrect original stem formation, not in prevailing usage; Colotini Larsen, 1983 (type genus Colotis Hübner, 1819) does not appear to be available in Lepidoptera: Pieridae; if the Lepidoptera name is found to be available then this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Attalaires Abeille de Perrin, 1891: 364 [stem: Attal-]. Type genus: Attalus Erichson, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hatch (1962: 87, as Attalina), generally accepted as in Mayor (2007: 419, as Attalini).
Laiina Jakobson, 1911a: 688 [stem: Lai-]. Type genus: Laius Guérin-Méneville, 1838.
Illopina Jakobson, 1911a: 688 [stem: Illop-]. Type genus: Illops Erichson, 1840.
Paratini Portevin, 1931: 426, in key [stem: Paratin-]. Type genus: Paratinus Abeille de Perrin, 1891 [syn. of Apalochrus Erichson, 1840]. Comment: the correct spelling Paratinini was used in the same work on page 442.
Ebaeini Portevin, 1931: 426, in key [stem: Ebae-]. Type genus: Ebaeus Erichson, 1840.
Collopina Hatch, 1962: 88, in key [stem: Collop-]. Type genus: Collops Erichson, 1840.
Tribe
Pagurodactylini
Constantin, 2001
Pagurodactylinae Constantin, 2001: 6 [stem: Pagurodactyl-]. Type genus: Pagurodactylus Gorham, 1900.
Melyridae
incertae sedis
Elosomatidae Jakobson, 1915: 993 [stem: Elosomat-]. Type genus: Elosoma Motschulsky, 1845. Comment: transferred from Salpingidae by Zerche (2004), correct placement within the family uncertain.
Superfamily
Cucujoidea
Latreille, 1802
Cucujipes Latreille, 1802: 210 [stem: Cucuj-]. Type genus: Cucujus Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: First Reviser (Cucujoidea Latreille, 1802 vs Erotyloidea Latreille, 1802 vs Nitiduloidea Latreille, 1802) not determined, current usage maintained.
†Family
Parandrexidae
Kirejtshuk, 1994
Parandrexidae Kirejtshuk, 1994: 57 [stem: Parandrex-]. Type genus: Parandrexis Martynov, 1926. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Parandrexe-).
†Family
Sinisilvanidae
Hong, 2002
Sinisilvanidae Hong, 2002: 132 [stem: Sinisilvan-]. Type genus: Sinisilvana Hong, 2002.
Family
Boganiidae
Sen Gupta and Crowson, 1966
Boganiidae Sen Gupta and Crowson, 1966: 63 [stem: Bogani-]. Type genus: Boganium Sen Gupta and Crowson, 1966.
Subfamily
Paracucujinae
Endrödy-Younga and Crowson, 1986
Paracucujinae Endrödy-Younga and Crowson, 1986: 255 [stem: Paracucuj-]. Type genus: Paracucujus Sen Gupta and Crowson, 1966.
Athertoniini Crowson, 1990: 91 [stem: Athertoni-]. Type genus: Athertonium Crowson, 1990.
Subfamily
Boganiinae
Sen Gupta and Crowson, 1966
Boganiidae Sen Gupta and Crowson, 1966: 63 [stem: Bogani-]. Type genus: Boganium Sen Gupta and Crowson, 1966.
Family
Byturidae
Gistel, 1848
Byturidae Gistel, 1848: [3] [stem: Bytur-]. Type genus: Byturus Latreille, 1797.
Subfamily
Platydascillinae
Pic, 1914
Platydascillidae Pic, 1914: 20 [stem: Platydascill-]. Type genus: Platydascillus Everts, 1909.
Subfamily
Byturinae
Gistel, 1848
Byturidae Gistel, 1848: [3] [stem: Bytur-]. Type genus: Byturus Latreille, 1797. Comment: family-group name previously attributed to Jacquelin du Val (1858: 211).
Family
Helotidae
Chapuis, 1876
Hélotides Chapuis, 1876: 15 [stem: Helot-]. Type genus: Helota W. S. MacLeay, 1825. Comment: published before 1 May 1876; original vernacular name available (Art. 11.7.2): used in latinized form by several authors subsequently, generally accepted as in Lawrence and Newton (1995: 875, as “Helotidae Reitter, 1876/Chapuis, 1876”); this family-group name was also used in the same year by Reitter (1876a [before 15 August]: 5, as Helotidae); this is a junior homonym of Helotidae Adams et al., 1854 (type genus Helotes Cuvier, 1829) in Pisces, this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Family
Protocucujidae
Crowson, 1954
Protocucujidae Crowson, 1954: 60 [stem: Protocucuj-]. Type genus: Protocucujus Crowson, 1954.
Family
Sphindidae
Jacquelin du Val, 1860
Sphindides Jacquelin du Val, 1860: 224 [stem: Sphind-]. Type genus: Sphindus Dejean, 1821 [placed on the Official List of Generic Names in Zoology (ICZN 1997)]. Comment: this name (as Sphindidae Jacquelin du Val, [1861]) was placed on the Official List of Family-Group Names in Zoology and ruled under the plenary power that this and other family-group names based on Sphindus Dejean, 1821 are to be given precedence over Aspidiphoridae Kiesenwetter, 1877 (1859) and other family-group names based on Aspidiphorus Dejean, 1821 whenever their type genera are placed in the same family-group taxon (ICZN 1997).
Subfamily
Protosphindinae
Sen Gupta and Crowson, 1979
Protosphindinae Sen Gupta and Crowson, 1979: 179, in key [stem: Protosphind-]. Type genus: Protosphindus Sen Gupta and Crowson, 1979.
Subfamily
Odontosphindinae
Sen Gupta and Crowson, 1979
Odontosphindini Sen Gupta and Crowson, 1979: 180, in key [stem: Odontosphind-]. Type genus: Odontosphindus Sen Gupta and Crowson, 1979.
Subfamily
Sphindiphorinae
Sen Gupta and Crowson, 1979
Sphindiphorini Sen Gupta and Crowson, 1979: 180, in key [stem: Sphindiphor-]. Type genus: Sphindiphorus Sen Gupta and Crowson, 1979.
Subfamily
Sphindinae
Jacquelin du Val, 1860
Coniporina C. G. Thomson, 1859: 90 [stem: Conipor-]. Type genus: Coniporus C. G. Thomson, 1859 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1997); syn. of Aspidiphorus Dejean, 1821]. Comment: permanently invalid (Art. 39): based on suppressed type genus.
Sphindides Jacquelin du Val, 1860: 224 [stem: Sphind-]. Type genus: Sphindus Dejean, 1821 [placed on the Official List of Generic Names in Zoology (ICZN 1997)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kiesenwetter (1877: 7, as Sphindini), generally accepted as in Lawrence and Newton (1995: 872, as Sphindidae).
Aspidiphoridae Kiesenwetter, 1877: 198 [stem: Aspidiphor-]. Type genus: Aspidiphorus Dejean, 1821 [Arpidiphorus, the original spelling of the type genus, as well as Aspidophorus Agassiz, 1846 were placed on the Official Index of Rejected and Invalid Generic Names in Zoology and Aspidiphorus was placed on the Official List of Generic Names in Zoology (ICZN 1997)]. Comment: this name (as Aspidiphoridae Kiesenwetter, 1877 (1859)) was placed on the Official List of Family-Group Names in Zoology and ruled under the plenary power that this and other family-group names based on Aspidiphorus are not to be given priority over Sphindidae and other family-group names based on Sphindus Dejean, 1821 whenever their type genera are placed in the same family-group taxon (ICZN 1997).
Eurysphindinae Sen Gupta and Crowson, 1979: 180, in key [stem: Eurysphind-]. Type genus: Eurysphindus J. L. LeConte, 1878.
Family
Biphyllidae
LeConte, 1861
Diphyllidae J. L. LeConte, 1861: 105 [stem: Biphyll-]. Type genus: Biphyllus Dejean, 1821 [as Diphyllus, unjustified emendation of type genus name by Agassiz (1846b: 47), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage (see Cline and Shockley 2010).
Family
Erotylidae
Latreille, 1802
Erotilenae Latreille, 1802: 233 [stem: Erotyl-]. Type genus: Erotylus Fabricius, 1775.
Subfamily
Xenoscelinae
Ganglbauer, 1899
Xenoscelini Ganglbauer, 1899: 649 [stem: Xenoscel-]. Type genus: Xenoscelis Wollaston, 1864. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Xenoscelid-).
Eicolyctini Vogt, 1967: 103 [stem: Eicolyct-]. Type genus: Eicolyctus J. R. Sahlberg, 1919 [syn. of Zavaljus Reitter, 1880].
Loberonothini Sen Gupta and Crowson, 1969a: 127 [stem: Loberonoth-]. Type genus: Loberonotha Sen Gupta and Crowson, 1969.
Subfamily
Pharaxonothinae
Crowson, 1952
Setariini Casey, 1900: 77 [stem: Setari-]. Type genus: Setaria Mulsant and Rey, 1863 [preoccupied genus name, not Setaria Viborg, 1795 [Vermes], not Setaria Oken, 1815 [Vermes], not Setaria Blyth, 1844 [Aves]; syn. of Setariola Jakobson, 1915]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Pharaxonothinae Crowson, 1952: 127 [stem: Pharaxonoth-]. Type genus: Pharaxonotha Reitter, 1875. Comment: First Reviser found (Pharaxonothinae Crowson, 1952 vs Setariolinae Crowson, 1952) is Leschen (2003: 35).
Setariolinae Crowson, 1952: 127 [stem: Setariol-]. Type genus: Setariola Jakobson, 1915.
Xenoscelinini Sen Gupta and Crowson, 1971: 5, in key [stem: Xenoscelin-]. Type genus: Xenoscelinus Grouvelle, 1910 [syn. of Cathartocryptus Sharp, 1886].
Subfamily
Loberinae
Bruce, 1951
Loberinae Bruce, 1951: 4 [stem: Lober-]. Type genus: Loberus J. L. LeConte, 1861. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Sen Gupta (1968a: 1, as Loberini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Subfamily
Languriinae
Hope, 1840
Languiridae Hope, 1840a: 190 [stem: Languri-]. Type genus: Languria Latreille, 1802 [as Languiria, incorrect subsequent spelling of type genus name, not in prevailing usage].
Tribe
Hapalipini
Leschen, 2003
Hapalipini Leschen, 2003: 38 [stem: Hapalip-]. Type genus: Hapalips Reitter, 1877.
Tribe
Languriini
Hope, 1840
Languiridae Hope, 1840a: 190 [stem: Languri-]. Type genus: Languria Latreille, 1802 [as Languiria, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; family-group name previously attributed to Crotch (1873c) by Pakaluk et al. (1994) and subsequent authors; usage of «Wiedeman, 1823» as the correct author and year for this family-group name in the literature is incorrect (see Pakaluk et al. 1994).
Cladoxeninae Arrow, 1925: 166 [stem: Cladoxen-]. Type genus: Cladoxena Motschulsky, 1866.
Tribe
Thallisellini
Sen Gupta, 1968
Thallisellini Sen Gupta, 1968b: 470 [stem: Thallisell-]. Type genus: Thallisella Crotch, 1876.
Subfamily
Cryptophilinae
Casey, 1900
Cryptophilini Casey, 1900: 77 [stem: Cryptophil-]. Type genus: Cryptophilus Reitter, 1874.
Tribe
Cryptophilini
Casey, 1900
Cryptophilini Casey, 1900: 77 [stem: Cryptophil-]. Type genus: Cryptophilus Reitter, 1874.
Tribe
Empocryptini
Leschen, 2003
Empocryptini Leschen, 2003: 41 [stem: Empocrypt-]. Type genus: Empocryptus Sharp, 1900.
Tribe
Toramini
Sen Gupta, 1967
Toraminae Sen Gupta, 1967: 168 [stem: Toram-]. Type genus: Toramus Grouvelle, 1916.
Subfamily
Erotylinae
Latreille, 1802
Erotilenae Latreille, 1802: 233 [stem: Erotyl-]. Type genus: Erotylus Fabricius, 1775.
Tribe
Dacnini
Gistel, 1848
*Engidites Latreille, 1829a: 506 [stem: Eng-]. Type genus: Engis Paykull, 1800 [syn. of Dacne Latreille, 1797]. Comment: family-group name unavailable (Art. 11.7.1.1): not based on a genus used as valid at the time.
Dacneidae Gistel, 1848: [3] [stem: Dacn-]. Type genus: Dacne Latreille, 1797. Comment: incorrect original stem formation, not in prevailing usage.
Cryptodacnini Sen Gupta, 1970: 101, in key [stem: Cryptodacn-]. Type genus: Cryptodacne Sharp, 1878.
Tribe
Encaustini
Crotch, 1876
Engidae W. S. MacLeay, 1825: 40 [stem: Engid-]. Type genus: Engis sensu W. S. MacLeay, 1825 [not Engis Paykull, 1800; syn. of Encaustes Lacordaire, 1842]. Comment: based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).
Encaustini Crotch, 1876: 476 [stem: Encaust-]. Type genus: Encaustes Lacordaire, 1842. Comment: published after February 1876; this family-group name was also used in the same year by Chapuis (1876 [before 1 May]: 46, as Encaustites).
Tribe
Erotylini
Latreille, 1802
Erotilenae Latreille, 1802: 233 [stem: Erotyl-]. Type genus: Erotylus Fabricius, 1775. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Megalodacnini
Sen Gupta, 1970
Megalodacnini Sen Gupta, 1970: 100, in key [stem: Megalodacn-]. Type genus: Megalodacne Crotch, 1873.
Tribe
Tritomini
Curtis, 1834
Tritomidae Curtis, 1834: plate 498 [stem: Tritom-]. Type genus: Tritoma Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Triplacinae Erichson, 1847a: 179 [stem: Triplac-]. Type genus: Triplax Herbst, 1793.
Renaniinae Chûjô, 1941: 10 [stem: Renani-]. Type genus: Renania Lewis, 1887.
Cyrtotriplacina Chûjô, 1969: 201 [stem: Cyrtotriplac-]. Type genus: Cyrtotriplax Crotch, 1873 [syn. of Tritoma Fabricius, 1775].
Family
Monotomidae
Laporte, 1840
Monotomites Laporte, 1840b: 377 [stem: Monotom-]. Type genus: Monotoma Herbst, 1793.
Subfamily
Rhizophaginae
Redtenbacher, 1845
Rhizophagi L. Redtenbacher, 1845: 125 [stem: Rhizophag-]. Type genus: Rhizophagus Herbst, 1793 [the original spelling Ryzophagus was placed on the Official Index of Rejected and Suppressed Generic Names in Zoology, Rhizophagus was considered the correct original spelling of the genus and placed on the Official List of Generic Names in Zoology (ICZN 1995b)].
Subfamily
Monotominae
Laporte, 1840
Monotomites Laporte, 1840b: 377 [stem: Monotom-]. Type genus: Monotoma Herbst, 1793.
Tribe
Europini
Sen Gupta, 1988
Europini Sen Gupta, 1988: 14, in key [stem: Europ-]. Type genus: Europs Wollaston, 1854.
Tribe
Lenacini
Crowson, 1952
Lenacinae Crowson, 1952: 121 [stem: Lenac-]. Type genus: Lenax Sharp, 1877.
Tribe
Monotomini
Laporte, 1840
Monotomites Laporte, 1840b: 377 [stem: Monotom-]. Type genus: Monotoma Herbst, 1793 [nomen protectum; this genus name is a junior homonym of Monotoma Panzer, 1792 nomen oblitum; we provide references to support the conservation of Monotoma Herbst, 1793 as the valid name for this genus (Art. 23.9.1) (see Appendix 1)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 238, as Monotomoidae), generally accepted as in Pakaluk et al. (1994: 230, as Monotomidae).
†Tribe
Rhizophtomini
Kirejtshuk and Azar, 2009
Rhizophtominae Kirejtshuk and Azar, 2009: 123 [stem: Rhizophtom-]. Type genus: Rhizophtoma Kirejtshuk and Azar, 2009.
Tribe
Thionini
Crowson, 1952
Thioninae Crowson, 1952: 121 [stem: Thion-]. Type genus: Thione Sharp, 1899.
Family
Hobartiidae
Sen Gupta and Crowson, 1966
Hobartiini Sen Gupta and Crowson, 1966: 65, in key [stem: Hobarti-]. Type genus: Hobartius Sen Gupta and Crowson, 1966.
Family
Cryptophagidae
Kirby, 1826
Cryptophagidae Kirby, 1826: 504 [stem: Cryptophag-]. Type genus: Cryptophagus Herbst, 1792.
Subfamily
Cryptophaginae
Kirby, 1826
Cryptophagidae Kirby, 1826: 504 [stem: Cryptophag-]. Type genus: Cryptophagus Herbst, 1792.
Tribe
Caenoscelini
Casey, 1900
Caenoscelini Casey, 1900: 103 [stem: Caenoscel-]. Type genus: Caenoscelis C. G. Thomson, 1863. Comment: First Reviser found (Caenoscelini Casey, 1900 vs Sternodeini Casey, 1900) is Leschen (1996: 607); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Caenoscelid-).
Sternodeini Casey, 1900: 103 [stem: Sternode-]. Type genus: Sternodea Reitter, 1875.
Tribe
Cryptophagini
Kirby, 1826
Ips Latreille, 1802: 129 [stem: Ip-]. Type genus: Ips sensu Latreille, 1802 [not Ips DeGeer, 1775; syn. of Cryptophagus Herbst, 1792]. Comment: although the spelling of the family-group names is identical to the spelling of the type genus, it is clear, based on other evidence from the same work, that Latreille was using it as a family-group name (see Lawrence and Newton 1995: 913); based on a misidentified type genus; name treated here as invalid until an application is submitted to the Commission to suppress it for the Principles of Priority and Homonymy (Art. 65.2.1); also see Ipini Bedel, 1888 in Curculionidae: Scolytinae.
Cryptophagidae Kirby, 1826: 504 [stem: Cryptophag-]. Type genus: Cryptophagus Herbst, 1792 [the original spelling Kryptophagus was placed on the Official Index of Rejected and Suppressed Generic Names in Zoology, Cryptophagus was considered the correct original spelling of the genus and placed on the Official List of Generic Names in Zoology (ICZN 1995b)].
Telmatophilides Jacquelin du Val, 1858: 209 [stem: Telmatophil-]. Type genus: Telmatophilus Heer, 1841. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 98, as Telmatophilini), generally accepted as in Telnov (2004: 73, as Telmatophilini).
Antherophagi J. L. LeConte, 1861: 98 [stem: Antherophag-]. Type genus: Antherophagus Dejean, 1821.
Paramecosomina Reitter, 1875: 4 [stem: Paramecosomat-]. Type genus: Paramecosoma Curtis, 1833. Comment: incorrect original stem formation, not in prevailing usage.
Catopochrotidae Reitter, 1889a: 289 [stem: Catopochrot-]. Type genus: Catopochrotus Reitter, 1889.
Emphyli Casey, 1900: 86 [stem: Emphyl-]. Type genus: Emphylus Erichson, 1846 [syn. of Spavius Motschulsky, 1844].
Spaniophaeni Casey, 1900: 86 [stem: Spaniophaen-]. Type genus: Spaniophaenus Reitter, 1875.
Tribe
Picrotini
Crowson, 1980
Picrotini Crowson, 1980: 283 [stem: Picrot-]. Type genus: Picrotus Sharp, 1886.
*Cryptosomatulini Crowson, 1980: 284 [stem: Cryptosomatul-]. Type genus: Cryptosomatula Bruce, 1940. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Cryptosomatulini Leschen, 1996: 605 [stem: Cryptosomatul-]. Type genus: Cryptosomatula Bruce, 1940.
Subfamily
Atomariinae
LeConte, 1861
Atomariini J. L. LeConte, 1861: 99 [stem: Atomari-]. Type genus: Atomaria Stephens, 1829.
Tribe
Atomariini
LeConte, 1861
Atomariini J. L. LeConte, 1861: 99 [stem: Atomari-]. Type genus: Atomaria Stephens, 1829.
Ephistemini Casey, 1900: 104 [stem: Ephistem-]. Type genus: Ephistemus Stephens, 1829.
*Salltiini Crowson, 1980: 284 [stem: Sallti-]. Type genus: Salltius Broun, 1893. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Atomaroidini Lyubarsky, 1998: 71 [stem: Atomaroid-]. Type genus: Atomaroides Lyubarsky, 1989. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Cryptafricini
Leschen, 1996
Cryptafricini Leschen, 1996: 623 [stem: Cryptafric-]. Type genus: Cryptafricus Leschen, 1996.
*Microphagini Lyubarsky, 1998: 39, in key [stem: Microphag-]. Type genus: Microphagus Leschen, 1996. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Scytomariini Lyubarsky, 1998: 71 [stem: Scytomari-]. Type genus: Scytomaria Lyubarsky, 1998 [syn. of Anitamaria Leschen, 1996]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Hypocoprini
Reitter, 1879
Hypocoprini Reitter, 1879: 74 [stem: Hypocopr-]. Type genus: Hypocoprus Motschulsky, 1839.
*Alfieriellinae Crowson, 1980: 282 [stem: Alfieriell-]. Type genus: Alfieriella Wittmer, 1935. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Hypophagina Lyubarsky, 1998: 71 [stem: Hypophag-]. Type genus: Hypophagus Lyubarsky, 1989. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Family
Agapythidae
Sen Gupta and Crowson, 1969
Agapythinae Sen Gupta and Crowson, 1969b: 579, in key [stem: Agapyth-]. Type genus: Agapytho Broun, 1921.
Family
Priasilphidae
Crowson, 1973
Priasilphinae Crowson, 1973a: 56 [stem: Priasilph-]. Type genus: Priasilpha Broun, 1893.
Family
Phloeostichidae
Reitter, 1911
Phloeostichini Reitter, 1911: 44 [stem: Phloeostich-]. Type genus: Phloeostichus W. Redtenbacher, 1842.
Hymaeinae Sen Gupta and Crowson, 1966: 65, in key [stem: Hymae-]. Type genus: Hymaea Pascoe, 1869.
Family
Silvanidae
Kirby, 1837
Sylvanidae Kirby, 1837: 110 [stem: Silvan-]. Type genus: Silvanus Latreille, 1804.
Subfamily
Brontinae
Blanchard, 1845
Brontites Blanchard, 1845b: 134 [stem: Bront-]. Type genus: Brontes Fabricius, 1801.
Tribe
Brontini
Blanchard, 1845
Brontites Blanchard, 1845b: 134 [stem: Bront-]. Type genus: Brontes Fabricius, 1801. Comment: published before 11 June 1845; original vernacular name available (Art. 11.7.2): first used in latinized form by Erichson (1845 [before 31 June]: 304, as Brontini), treated as available by Lawrence and Newton (1995: 876).
Dendrophagidae Gistel, 1848: [8] [stem: Dendrophag-]. Type genus: Dendrophagus Schönherr, 1809.
Uleiotaeidae Gistel, 1848: [10] [stem: Uleiot-]. Type genus: Uleiota Latreille, 1797. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Telephanini
LeConte, 1861
Telephanidae J. L. LeConte, 1861: 96 [stem: Telephan-]. Type genus: Telephanus Erichson, 1845.
Pseudophanini J. L. LeConte, 1861: 96 [stem: Pseudophan-]. Type genus: Pseudophanus J. L. LeConte, 1859 [syn. of Cryptamorpha Wollaston, 1854].
Psammoecini Reitter, 1879: 74 [stem: Psammoec-]. Type genus: Psammoecus Latreille, 1829.
Cryptamorphini Casey, 1884: 102 [stem: Cryptamorph-]. Type genus: Cryptamorpha Wollaston, 1854.
Subfamily
Silvaninae
Kirby, 1837
Sylvanidae Kirby, 1837: 110 [stem: Silvan-]. Type genus: Silvanus Latreille, 1804. Comment: incorrect original stem formation, not in prevailing usage.
Family
Cucujidae
Latreille, 1802
Cucujipes Latreille, 1802: 210 [stem: Cucuj-]. Type genus: Cucujus Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
*Biophloces Motschulsky, 1849: 60 [stem: Biophloe-]. Type genus: Biophloeus Dejean, 1835 [nomen oblitum; this genus name is a senior synonym of the well-established name Pediacus Shuckard, 1839 nomen protectum; however since Biophloeus Dejean has not been used as valid after 1899 to our knowledge, we provide references to support the conservation of Pediacus as the valid name for this genus (Art. 23.9.1) (see Appendix 1)]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Earophilidae Gistel, 1856a: 375 [stem: Earophil-]. Type genus: Earophilus Gistel, 1856 [syn. of Cucujus Fabricius, 1775].
Family
Myraboliidae
Lawrence and Britton, 1991
Myraboliinae Lawrence and Britton, 1991: 650 [stem: Myraboli-]. Type genus: Myrabolia Reitter, 1876.
Family
Cavognathidae
Sen Gupta and Crowson, 1966
Cavognathinae Sen Gupta and Crowson, 1966: 65, in key [stem: Cavognath-]. Type genus: Cavognatha Crowson, 1964 [syn. of Taphropiestes Reitter, 1875].
Family
Lamingtoniidae
Sen Gupta and Crowson, 1969
Lamingtoniidae Sen Gupta and Crowson, 1969a: 125 [stem: Lamingtoni-]. Type genus: Lamingtonium Sen Gupta and Crowson, 1969.
Family
Passandridae
Blanchard, 1845
Passandrites Blanchard, 1845b: 134 [stem: Passandr-]. Type genus: Passandra Dalman, 1817. Comment: published before 11 June 1845; original vernacular name available (Art. 11.7.2): first used in latinized form by Erichson (1845 [before June 31]: 304, as Passandrini), treated as available by Lawrence and Newton (1995: 876).
Ancistriinae Sharp, 1899b: 541 [stem: Ancistri-]. Type genus: Ancistria Erichson, 1845.
Catogenini Grouvelle, 1916: 6 [stem: Catogen-]. Type genus: Catogenus Westwood, 1830.
Passandrellini Grouvelle, 1916: 6 [stem: Passandrell-]. Type genus: Passandrella Grouvelle, 1916.
Scalidiini Grouvelle, 1916: 6 [stem: Scalidi-]. Type genus: Scalidia Erichson, 1845.
Laemotmetini Kessel, 1921: 35 [stem: Laemotmet-]. Type genus: Laemotmetus Gerstaecker, 1871 [syn. of Aulonosoma Motschulsky, 1858].
Family
Phalacridae
Leach, 1815
Phalacrurida Leach, 1815: 116 [stem: Phalacr-]. Type genus: Phalacrus Paykull, 1800.
Subfamily
Phaenocephalinae
Matthews, 1899
Phaenocephalidae A. Matthews, 1899: 205 [stem: Phaenocephal-]. Type genus: Phaenocephalus Wollaston, 1873.
Subfamily
Phalacrinae
Leach, 1815
Phalacrurida Leach, 1815: 116 [stem: Phalacr-]. Type genus: Phalacrus Paykull, 1800. Comment: incorrect original stem formation, not in prevailing usage.
Idiobiidae Gistel, 1856a: 383 [stem: Idiobi-]. Type genus: Idiobius Gistel, 1856 [syn. of Olibrus Erichson, 1845].
Eustilbini Guillebeau, 1892: 149 [stem: Eustilb-]. Type genus: Eustilbus Sharp, 1888.
Olibrini Guillebeau, 1892: 147 [stem: Olibr-]. Type genus: Olibrus Erichson, 1845.
Tolyphini Guillebeau, 1892: 147 [stem: Tolyph-]. Type genus: Tolyphus Erichson, 1845.
Biophytini Guillebeau, 1894: 276, in key [stem: Biophyt-]. Type genus: Biophytus Guillebeau, 1894.
Megapalpini Guillebeau, 1894: 278, in key [stem: Megapalp-]. Type genus: Megapalpus Guillebeau, 1893 [preoccupied genus name, not Megapalpus Macquart, 1834 [Diptera]; syn. of Megistopalpus Guillebeau, 1895]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Ochrolitini Guillebeau, 1894: 278, in key [stem: Ochrolit-]. Type genus: Ochrolitus Sharp, 1889.
Stilbini Jakobson, 1915: 948 [stem: Stilb-]. Type genus: Stilbus Seidlitz, 1872.
Family
Propalticidae
Crowson, 1952
Propalticidae Crowson, 1952: 117 [stem: Propaltic-]. Type genus: Propalticus Sharp, 1879.
Family
Laemophloeidae
Ganglbauer, 1899
Laemophloeini Ganglbauer, 1899: 606 [stem: Laemophloe-]. Type genus: Laemophloeus Dejean, 1835.
Nartheciinae Grouvelle, 1908: 453 [stem: Nartheci-]. Type genus: Narthecius J. L. LeConte, 1861.
Family
Tasmosalpingidae
Lawrence and Britton, 1991
Tasmosalpinginae Lawrence and Britton, 1991: 651 [stem: Tasmosalping-]. Type genus: Tasmosalpingus Lea, 1918.
Family
Cyclaxyridae
Gimmel, Leschen and Ślipiński, 2009
*Cyclaxyrinae Crowson, 1984: 259 [stem: Cyclaxyr-]. Type genus: Cyclaxyra Broun, 1893 [placed on the Official List of Generic Names in Zoology (ICZN 1988a)]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Cyclaxyridae Gimmel et al., 2009: 513 [stem: Cyclaxyr-]. Type genus: Cyclaxyra Broun, 1893 [placed on the Official List of Generic Names in Zoology (ICZN 1988a)]. Comment: although this family-group name was used several times before 2009, all previous uses of the name were unavailable (see Gimmel et al. 2009: 512, 513).
Family
Kateretidae
Kirby, 1837
Catheretidae Kirby, 1837: 107 [stem: Kateret-]. Type genus: Kateretes Herbst, 1793 [as Catheretes, incorrect subsequent spelling of type genus name, not in prevailing usage; Kateretes Herbst, 1793 placed on Official List of Generic Names in Zoology (ICZN 1999b)]. Comment: incorrect original stem formation, not in prevailing usage; Kateretidae chosen as correct spelling of the family-group name, given precedence over Brachypteridae Erichson, 1845 and placed on Official List of Family-Group Names in Zoology (ICZN 1999b, as Kateretidae Erichson in Agassiz, 1846); discovery of the older available name proposed by Kirby (1837) removes priority problems with Brachypterinae.
Brachypterinae Erichson, 1845: 125 [stem: Brachypter-]. Type genus: Brachypterus Kugelann, 1794 [placed on the Official List of Generic Names in Zoology (ICZN 1999b)]. Comment: placed on Official List of Family-Group Names in Zoology (ICZN 1999b).
Cercidae Desmarest, 1851: 291 [stem: Cerc-]. Type genus: Cercus Latreille, 1797 [syn. of Kateretes Herbst, 1793].
Family
Nitidulidae
Latreille, 1802
Nitidulariae Latreille, 1802: 131 [stem: Nitidul-]. Type genus: Nitidula Fabricius, 1775. Comment: the classification of Nitidulidae follows Kirejtshuk (2008).
Subfamily
Calonecrinae
Kirejtshuk, 1982
Calonecrinae Kirejtshuk, 1982: 117 [stem: Calonecr-]. Type genus: Calonecrus J. Thomson, 1857.
Subfamily
Maynipeplinae
Kirejtshuk, 1998
Maynipeplinae Kirejtshuk, 1998b: 540 [stem: Maynipepl-]. Type genus: Maynipeplus Kirejtshuk, 1998.
Subfamily
Epuraeinae
Kirejtshuk, 1986
Epuraeini Kirejtshuk, 1986: 27 [stem: Epurae-]. Type genus: Epuraea Erichson, 1843.
Tribe
Epuraeini
Kirejtshuk, 1986
Epuraeini Kirejtshuk, 1986: 27 [stem: Epurae-]. Type genus: Epuraea Erichson, 1843.
Tribe
Taenioncini
Kirejtshuk, 1998
Taenioncini Kirejtshuk, 1998a: 322 [stem: Taenionc-]. Type genus: Taenioncus Kirejtshuk, 1984.
Subfamily
Carpophilinae
Erichson, 1842
Carpophilinae Erichson, 1842: 148 [stem: Carpophil-]. Type genus: Carpophilus Stephens, 1829.
Subfamily
Amphicrossinae
Kirejtshuk, 1986
Amphicrossini Kirejtshuk, 1986: 28 [stem: Amphicross-]. Type genus: Amphicrossus Erichson, 1843.
Subfamily
Meligethinae
Thomson, 1859
Meligethina C. G. Thomson, 1859: 67 [stem: Meligeth-]. Type genus: Meligethes Stephens, 1830.
Subfamily
Nitidulinae
Latreille, 1802
Nitidulariae Latreille, 1802: 131 [stem: Nitidul-]. Type genus: Nitidula Fabricius, 1775.
Tribe
Cychramini
Gistel, 1848
Cychramidae Gistel, 1848: [4] [stem: Cychram-]. Type genus: Cychramus Kugelann, 1794.
Tribe
Cychramptodini
Kirejtshuk and Lawrence, 1992
Cychramptodini Kirejtshuk and Lawrence, 1992: 29, in key [stem: Cychramptod-]. Type genus: Cychramptodes Reitter, 1878.
Tribe
Cyllodini
Everts, 1898
Strongylinae Erichson, 1843: 227 [stem: Strongyl-]. Type genus: Strongylus Herbst, 1792 [preoccupied genus name, not Strongylus O. F. Müller, 1780 [Nematoda]; syn. of Cyllodes Erichson, 1843]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; Strongylidae Baird, 1853 (type genus Strongylus O. F. Müller, 1780) is available in Nematoda.
Cyllodini Everts, 1898: 469, in key [stem: Cyllod-]. Type genus: Cyllodes Erichson, 1843.
Amborotubini Leschen and Carlton, 2004: 450 [stem: Amborotub-]. Type genus: Amborotubus Leschen and Carlton, 2004.
Tribe
Lawrencerosini
Kirejtshuk, 1991
Lawrencerosini Kirejtshuk, 1991: 863 [stem: Lawrenceros-]. Type genus: Lawrencerosus Kirejtshuk, 1991.
Tribe
Mystropini
Murray, 1864
Mystropidae Murray, 1864: 411 [stem: Mystrop-]. Type genus: Mystrops Erichson, 1843.
Tribe
Nitidulini
Latreille, 1802
Nitidulariae Latreille, 1802: 131 [stem: Nitidul-]. Type genus: Nitidula Fabricius, 1775.
Thalycrina C. G. Thomson, 1859: 68 [stem: Thalycr-]. Type genus: Thalycra Erichson, 1843.
Pocadiini Seidlitz, 1872 [Gatt.]: 32 [stem: Pocadi-]. Type genus: Pocadius Erichson, 1843.
Prometopinae Böving and Craighead, 1931: 37, in key [stem: Prometopi-]. Type genus: Prometopia Erichson, 1843. Comment: the spelling Prometopinae in the key on page 37 is considered a lapsus calami since the correct spelling Prometopiinae was used in the same publication in the conspectus on page 74 and in the caption for plate 35.
Orvoenini Dajoz, 1980b: 191 [stem: Orvoeni-]. Type genus: Orvoenia Dajoz, 1980 [syn. of Megauchenia W. S. MacLeay, 1825]. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Cillaeinae
Kirejtshuk and Audisio, 1986
Cillaeinae Kirejtshuk and Audisio, 1986: 219 [stem: Cillae-]. Type genus: Cillaeus Laporte, 1835.
Subfamily
Cryptarchinae
Thomson, 1859
Cryptarchina C. G. Thomson, 1859: 69 [stem: Cryptarch-]. Type genus: Cryptarcha Shuckard, 1839.
Tribe
Arhinini
Kirejtshuk, 1987
Arhinini Kirejtshuk, 1987: 63 [stem: Arhin-]. Type genus: Arhina Murray, 1876.
Tribe
Cryptarchini
Thomson, 1859
Cryptarchina C. G. Thomson, 1859: 69 [stem: Cryptarch-]. Type genus: Cryptarcha Shuckard, 1839.
Pityophagini Reitter, 1891: 163 [stem: Pityophag-]. Type genus: Pityophagus Shuckard, 1839.
Glischrochilini Iablokoff-Khnzorian, 1966: 314 [stem: Glischrochil-]. Type genus: Glischrochilus Reitter, 1873.
Tribe
Eucalosphaerini
Kirejtshuk, 1987
Eucalosphaerini Kirejtshuk, 1987: 80 [stem: Eucalosphaer-]. Type genus: Eucalosphaera Jelinek, 1978.
Tribe
Platyarchini
Kirejtshuk, 1998
Platyarchini Kirejtshuk, 1998a: 41, in key [stem: Platyarch-]. Type genus: Platyarcha Kirejtshuk, 1987.
Subfamily
Cybocephalinae
Jacquelin du Val, 1858
Cybocéphalites Jacquelin du Val, 1858: 151 [stem: Cybocephal-]. Type genus: Cybocephalus Erichson, 1844. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. G. Thomson (1862: 123, as Cybocephalidae), generally accepted as in Pakaluk et al. (1994: 230, as Cybocephalinae).
Family
Smicripidae
Horn, 1880
Smicripini G. H. Horn, 1880d: 268 [stem: Smicrip-]. Type genus: Smicrips J. L. LeConte, 1878.
Tisiphoninae Sharp, 1900a: 578 [stem: Tisiphon-]. Type genus: Tisiphone Reitter, 1876 [preoccupied genus name, not Tisiphone Hübner, 1816 [Lepidoptera]; syn. of Smicrips J. L. LeConte, 1878]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Family
Bothrideridae
Erichson, 1845
Bothriderini Erichson, 1845: 287 [stem: Bothrider-]. Type genus: Bothrideres Dejean, 1835.
Subfamily
Teredinae
Seidlitz, 1888
Teredini Seidlitz, 1888 [Gatt.]: 57 [stem: Tered-]. Type genus: Teredus Dejean, 1835.
Tribe
Sosylopsini
Dajoz, 1980
Sosylopsini Dajoz, 1980a: 158 [stem: Sosylops-]. Type genus: Sosylopsis Grouvelle, 1910. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Sosylopse-).
Tribe
Sysolini
Ślipiński and Pal, 1985
Sysolini Ślipiński and Pal, 1985: 40 [stem: Sysol-]. Type genus: Sysolus Grouvelle, 1908.
Tribe
Teredini
Seidlitz, 1888
Teredini Seidlitz, 1888 [Gatt.]: 57 [stem: Tered-]. Type genus: Teredus Dejean, 1835.
Subfamily
Xylariophilinae
Pal and Lawrence, 1986
Xylariophilinae Pal and Lawrence, 1986: 208, in key [stem: Xylariophil-]. Type genus: Xylariophilus Pal and Lawrence, 1986.
Subfamily
Anommatinae
Ganglbauer, 1899
Anommatini Ganglbauer, 1899: 894 [stem: Anommat-]. Type genus: Anommatus Wesmael, 1835.
Subfamily
Bothriderinae
Erichson, 1845
Bothriderini Erichson, 1845: 287 [stem: Bothrider-]. Type genus: Bothrideres Dejean, 1835.
Deretaphrini G. H. Horn, 1878: 578 [stem: Deretaphr-]. Type genus: Deretaphrus Newman, 1842.
Dastarcini Reitter, 1922a: 39 [stem: Dastarc-]. Type genus: Dastarcus Walker, 1858.
Family
Cerylonidae
Billberg, 1820
Cerylonides Billberg, 1820a: 47 [stem: Cerylon-]. Type genus: Cerylon Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1995c)]. Comment: Cerylonidae Billberg, 1820 placed on the Official List of Family-Group Names in Zoology (ICZN 1995c).
Subfamily
Euxestinae
Grouvelle, 1908
Euxestinae Grouvelle, 1908: 397, in key [stem: Euxest-]. Type genus: Euxestus Wollaston, 1858.
Pachyochthesinae Reitter, 1911: 105 [stem: Pachyochth-]. Type genus: Pachyochthes Reitter, 1897 [syn. of Hypodacne J. L. LeConte, 1875]. Comment: incorrect original stem formation, not in prevailing usage.
Metacerylini Heinze, 1944: 21 [stem: Metacerylon-]. Type genus: Metacerylon Grouvelle, 1906. Comment: incorrect original stem formation, not in prevailing usage.
Cycloxenini Jeannel and Paulian, 1945: 57, in key [stem: Cycloxen-]. Type genus: Cycloxenus Arrow, 1925.
Tachyoryctidiini Jeannel and Paulian, 1945: 57, in key [stem: Tachyoryctidi-]. Type genus: Tachyoryctidium Jeannel and Paulian, 1945.
Subfamily
Loeblioryloninae
Ślipiński, 1990
Loebliorylininae Ślipiński, 1990: 81 [stem: Loebliorylon-]. Type genus: Loebliorylon Ślipiński, 1990. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Ostomopsinae
Sen Gupta and Crowson, 1973
Ostomopsini Sen Gupta and Crowson, 1973: 400 [stem: Ostomops-]. Type genus: Ostomopsis Scott, 1922. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Ostomopse-).
Subfamily
Murmidiinae
Jacquelin du Val, 1858
Murmidiides Jacquelin du Val, 1858: 227 [stem: Murmidi-]. Type genus: Murmidius Leach, 1822. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 78, as Murmidiidae), generally accepted as in Pakaluk et al. (1994: 232, as Murmidiinae).
Subfamily
Ceryloninae
Billberg, 1820
Cerylonides Billberg, 1820a: 47 [stem: Cerylon-]. Type genus: Cerylon Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1995c)]. Comment: Cerylonidae Billberg, 1820a placed on the Official List of Family-Group Names in Zoology (ICZN 1995c); this family-group name was also used in the same year by Billberg (1820b: 394, as Cerylonides).
Pleosomides Fauvel, 1891: 162 [stem: Ploeosomat-]. Type genus: Ploeosoma Wollaston, 1854. Comment: incorrect original stem formation, not in prevailing usage.
Lapethinae Sharp, 1894: 445 [stem: Lapeth-]. Type genus: Lapethus Casey, 1890 [syn. of Mychocerus Erichson, 1845].
Mychocerinae Sharp, 1895b: 494 [stem: Mychocer-]. Type genus: Mychocerus sensu J. L. LeConte, 1869 [syn. of Mychocerinus Ślipiński, 1990]. Comment: based on a misidentified type genus.
Aculagnathidae Oke, 1932: 22 [stem: Aculagnath-]. Type genus: Aculagnathus Oke, 1932.
Dolosidae Dajoz, 1963: 96 [stem: Dolos-]. Type genus: Dolosus Dajoz, 1963 [subgenus of Thyroderus Sharp, 1885].
Family
Alexiidae
Imhoff, 1856
Alexiadae Imhoff, 1856: [2] 151 [stem: Alexi-]. Type genus: Alexia Stephens, 1833 [syn. of Sphaerosoma Stephens, 1832].
Sphaerosominae Ganglbauer, 1899: 913 [stem: Sphaerosomat-]. Type genus: Sphaerosoma Stephens, 1832. Comment: incorrect original stem formation, not in prevailing usage.
Family
Discolomatidae
Horn, 1878
Discolomidae G. H. Horn, 1878: 557 [stem: Discolomat-]. Type genus: Discoloma Erichson, 1845. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Notiophyginae
Jakobson, 1915
Notiophygidae Jakobson, 1915: 920 [stem: Notiophyg-]. Type genus: Notiophygus Gory, 1834.
Tribe
Dystheamonini
John, 1954
Dystheamonini John, 1954: 42 [stem: Dystheamon-]. Type genus: Dystheamon Grouvelle, 1927.
Tribe
Notiophygini
Jakobson, 1915
Notiophygidae Jakobson, 1915: 920 [stem: Notiophyg-]. Type genus: Notiophygus Gory, 1834.
Tribe
Pachyplacini
John, 1954
Pachyplacini John, 1954: 38 [stem: Pachyplac-]. Type genus: Pachyplacus John, 1935.
Subfamily
Discolomatinae
Horn, 1878
Discolomidae G. H. Horn, 1878: 557 [stem: Discolomat-]. Type genus: Discoloma Erichson, 1845. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Aphanocephalinae
Jakobson, 1904
Aphanocephalidae Jakobson, 1904: 273 [stem: Aphanocephal-]. Type genus: Aphanocephalus Wollaston, 1873. Comment: name previously attributed to Grouvelle (1912: 198); usage of Aphaenocephalidae by Ganglbauer (1903: 316) in Staphylinoidea is a lapsus calami for Phaenocephalidae (now in Phalacridae) according to Lawrence and Newton (1995).
Subfamily
Cephalophaninae
John, 1954
Cephalophaninae John, 1954: 64 [stem: Cephalophan-]. Type genus: Cephalophanus John, 1940.
Subfamily
Pondonatinae
John, 1954
Pondonatinae John, 1954: 19 [stem: Pondonat-]. Type genus: Pondonatus John, 1954.
Family
Endomychidae
Leach, 1815
Endomychides Leach, 1815: 116 [stem: Endomych-]. Type genus: Endomychus Panzer, 1795.
Subfamily
Merophysiinae
Seidlitz, 1872
Merophysiini Seidlitz, 1872 [Gatt.]: 39 [stem: Merophysi-]. Type genus: Merophysia Lucas, 1852.
Holoparamecini Seidlitz, 1888 [Gatt.]: 57 [stem: Holoparamec-]. Type genus: Holoparamecus Curtis, 1833.
Subfamily
Pleganophorinae
Jacquelin du Val, 1858
Pléganophorides Jacquelin du Val, 1858: 186 [stem: Pleganophor-]. Type genus: Pleganophorus Hampe, 1855. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gerstaecker (1861: 419, as Pleganophoridae), generally accepted as in Pakaluk et al. (1994: 233, as Pleganophorinae).
Trochoidéites Chapuis, 1876: 146 [stem: Trochoide-]. Type genus: Trochoideus Westwood, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Ganglbauer (1899: 926, as Trochoideinae); the earlier use of Trochoïdeidae by Gorham (1874: 185) is not considered to be a proposal for a new family-group taxon but was rather used as a hypothetical example addressed to the editors of the journal.
Subfamily
Anamorphinae
Strohecker, 1953
Anamorphini Strohecker, 1953: 15, in key [stem: Anamorph-]. Type genus: Anamorphus J. L. LeConte, 1878.
Mychotheninae Sasaji, 1978: 8 [stem: Mychothen-]. Type genus: Mychothenus Strohecker, 1953.
Acritosomatinae Pakaluk and Ślipiński, 1995: 328 [stem: Acritosomat-]. Type genus: Acritosoma Pakaluk and Ślipiński, 1995.
Subfamily
Leiestinae
Thomson, 1863
Leiestina C. G. Thomson, 1863: 306 [stem: Leiest-]. Type genus: Leiestes Chevrolat, 1836.
Rhanes J. L. LeConte and G. H. Horn, 1883: 120 [stem: Rhanid-]. Type genus: Rhanis J. L. LeConte, 1854 [preoccupied genus name, not Rhanis C. L. Koch, 1846 [Arachnida]; syn. of Rhanidea Strohecker, 1953]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Phymaphorina Jakobson, 1915: 961 [stem: Phymaphor-]. Type genus: Phymaphora Newman, 1838.
Subfamily
Mycetaeinae
Jacquelin du Val, 1857
Mycetéides Jacquelin du Val, 1857b: 102 [stem: Mycetae-]. Type genus: Mycetaea Stephens, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gerstaecker (1861: 419, as Mycetaeidae), generally accepted as in Pakaluk et al. (1994: 233, as Mycetaeinae); incorrect original stem formation, not in prevailing usage.
Subfamily
Eupsilobiinae
Casey, 1895
Eupsilobiini Casey, 1895: 452 [stem: Eupsilobi-]. Type genus: Eupsilobius Casey, 1895 [syn. of Eidoreus Sharp, 1885].
Cerasommatidiidae Brèthes, 1925: 199 [stem: Cerasommatidi-]. Type genus: Cerasommatidia Brèthes, 1925.
Eidoreinae Sasaji, 1986: 235 [stem: Eidore-]. Type genus: Eidoreus Sharp, 1885.
Subfamily
Xenomycetinae
Strohecker, 1962
Xenomycetinae Strohecker, 1962: 801, in key [stem: Xenomycet-]. Type genus: Xenomycetes G. H. Horn, 1880.
Subfamily
Danascelinae
Tomaszewska, 2000
Danascelinae Tomaszewska, 2000: 494, in key [stem: Danascel-]. Type genus: Danascelis Tomaszewska, 1999. Comment: current spelling maintained (Art. 29.3.1.1): incorrect original stem formation in prevailing usage (should be Danascelid-).
Subfamily
Endomychinae
Leach, 1815
Endomychides Leach, 1815: 116 [stem: Endomych-]. Type genus: Endomychus Panzer, 1795.
*Agaricophiles Motschulsky, 1849: 58 [stem: Agaricophil-]. Type genus: Agaricophilus Motschulsky, 1837. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Cyclotomidae Imhoff, 1856: [2] 151 [stem: Cyclotom-]. Type genus: Cyclotoma Mulsant, 1851.
*Agaricophilinae Lawrence, 1991: 484 [stem: Agaricophil-]. Type genus: Agaricophilus Motschulsky, 1837. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); proposed as new without reference to Agaricophiles Motschulsky, 1849.
Subfamily
Epipocinae
Gorham, 1873
Epipocidae Gorham, 1873: 20 [stem: Epipoc-]. Type genus: Epipocus Germar, 1843.
Subfamily
Stenotarsinae
Chapuis, 1876
Sténotarsites Chapuis, 1876: 125 [stem: Stenotars-]. Type genus: Stenotarsus Perty, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Csiki (1901: 37, as Stenotarsini), generally accepted as in Shockley et al. (2009: 73, as Stenotarsinae).
Subfamily
Lycoperdininae
Bromhead, 1838
Lycoperdinadae Bromhead, 1838: 419 [stem: Lycoperdin-]. Type genus: Lycoperdina Latreille, 1807. Comment: name previously attributed to L. Redtenbacher (1844); incorrect original stem formation, not in prevailing usage.
Eumorphidae Gistel, 1848: [10] [stem: Eumorph-]. Type genus: Eumorphus Weber, 1801.
Dapsini Gerstaecker, 1858: 170 [stem: Daps-]. Type genus: Dapsa Latreille, 1829.
Corynomalidae Gorham, 1873: 14 [stem: Corynomal-]. Type genus: Corynomalus Gerstaecker, 1857.
Amphicini Csiki, 1910: 25 [stem: Amphic-]. Type genus: Amphix Laporte, 1840.
Beccariini Arrow, 1925: 278 [stem: Beccari-]. Type genus: Beccaria Gorham, 1885 [preoccupied genus name, not Beccaria Bourguignat, 1883 [Mollusca]; syn. of Beccariola Arrow, 1943]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Amphisternini Strohecker, 1964: 319 [stem: Amphistern-]. Type genus: Amphisternus Germar, 1843.
Family
Coccinellidae
Latreille, 1807
Coccinellidae Latreille, 1807: 70 [stem: Coccinell-]. Type genus: Coccinella Linnaeus, 1758.
Subfamily
Microweiseinae
Leng, 1920
Microweiseini Leng, 1920: 213 [stem: Microweise-]. Type genus: Microweisea Cockerell, 1903.
Tribe
Microweiseini
Leng, 1920
Microweiseini Leng, 1920: 213 [stem: Microweise-]. Type genus: Microweisea Cockerell, 1903.
Tribe
Serangiini
Pope, 1962
*Serangiini Blackwelder, 1945: 450 [stem: Serangi-]. Type genus: Serangium Blackburn, 1889. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Serangiini Pope, 1962: 627 [stem: Serangi-]. Type genus: Serangium Blackburn, 1889.
Tribe
Sukunahikonini
Kamiya, 1960
Sukunahikonini Kamiya, 1960: 24 [stem: Sukunahikon-]. Type genus: Sukunahikona Kamiya, 1960 [syn. of Scymnomorphus Weise, 1897].
Scotoscymninae Duverger, 2003: 63 [stem: Scotoscymn-]. Type genus: Scotoscymnus Weise, 1901. Comment: name proposed to replace Sukunahikonini Kamiya, 1960 because of the synonymy of the type genus.
Subfamily
Coccinellinae
Latreille, 1807
Coccinellidae Latreille, 1807: 70 [stem: Coccinell-]. Type genus: Coccinella Linnaeus, 1758.
Tribe
Argentipilosini
Gordon and de Almeida, 1991
Argentipilosini Gordon and de Almeida, 1991: 150 [stem: Argentipilos-]. Type genus: Argentipilosa Gordon and de Almeida, 1991.
Tribe
Aspidimerini
Mulsant, 1850
Aspidiméraires Mulsant, 1850: 943 [stem: Aspidimer-]. Type genus: Aspidimerus Mulsant, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jakobson (1915: 969, as Aspidimerina), generally accepted as in Pakaluk et al. (1994: 234, as Aspidimerini).
Tribe
Azyini
Mulsant, 1850
Azyaires Mulsant, 1850: 927 [stem: Azy-]. Type genus: Azya Mulsant, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Crotch (1874: 279, as Azyae), generally accepted as in Pakaluk et al. (1994: 233, as Azyini).
*Bucolites Chapuis, 1876: 237 [stem: Bucol-]. Type genus: Bucolus Mulsant, 1850. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Tribe
Brachiacanthini
Mulsant, 1850
Brachyacanthaires Mulsant, 1850: 520 [stem: Brachiacanth-]. Type genus: Brachiacantha Chevrolat, 1836 [as Brachyacantha, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Duverger (1990: 143, as Brachiacanthadini [incorrect stem formation]), generally accepted as in Pakaluk et al. (1994: 249, as Brachiacanthini); incorrect original stem formation, not in prevailing usage.
Tribe
Carinodulini
Gordon, Pakaluk and Ślipiński, 1989
Carinodulini Gordon et al., 1989: 360 [stem: Carinodul-]. Type genus: Carinodula Gordon et al., 1989.
Tribe
Cephaloscymnini
Gordon, 1985
Cephaloscymnini Gordon, 1985: 66 [stem: Cephaloscymn-]. Type genus: Cephaloscymnus Crotch, 1872.
Tribe
Chilocorini
Mulsant, 1846
Chilocoriens Mulsant, 1846: 166 [stem: Chilocor-]. Type genus: Chilocorus Leach, 1815. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1885: 239, as Chilocorini), generally accepted as in Pakaluk et al. (1994: 234, as Chilocorini).
*Exochomaires Mulsant, 1850: 465 [stem: Exochom-]. Type genus: Exochomus Redtenbacher, 1843. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Tribe
Chnoodini
Mulsant, 1850
Chnoodiens Mulsant, 1850: 907 [stem: Chnood-]. Type genus: Chnoodes Chevrolat, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Sicard (1909: 103, as Chnoodini); name incorrectly treated as unavailable by Pakaluk et al. (1994: 248).
*Siolaires Mulsant, 1850: 931 [stem: Siol-]. Type genus: Siola Mulsant, 1850. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Exoplectrides Crotch, 1874: xiii [stem: Exoplectr-]. Type genus: Exoplectra Chevrolat, 1836.
Sumniini Hoang, 1982: 138 [stem: Sumni-]. Type genus: Sumnius Weise, 1892.
Tribe
Coccidulini
Mulsant, 1846
Cocciduliens Mulsant, 1846: 266 [stem: Coccidul-]. Type genus: Coccidula Kugelann, 1798. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1848: [10], as Coccidulaeidae [incorrect stem formation]), generally accepted as in Pakaluk et al. (1994: 233, as Coccidulini).
*Rhizobiates Mulsant, 1846: 261 [stem: Rhyzobi-]. Type genus: Rhyzobius Stephens, 1829 [as Rhizobius, unjustified emendation of type genus name by Agassiz (1846b: 327), not in prevailing usage]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant (1846); incorrect original stem formation, not in prevailing usage.
Rhizobiina C. G. Thomson, 1866: 328 [stem: Rhyzobi-]. Type genus: Rhyzobius Stephens, 1829 [as Rhizobius, unjustified emendation of type genus name by Agassiz (1846b: 327), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; the name Rhizobiinae Buckton, 1883 (type genus Rhizobius Burmeister, 1835) is available in Hemiptera.
Tribe
Coccinellini
Latreille, 1807
Coccinellidae Latreille, 1807: 70 [stem: Coccinell-]. Type genus: Coccinella Linnaeus, 1758.
*Adoniates Mulsant, 1846: 35 [stem: Adoni-]. Type genus: Adonia Mulsant, 1846 [syn. of Hippodamia Chevrolat, 1836]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Hippodamiaires Mulsant, 1846: 30 [stem: Hippodami-]. Type genus: Hippodamia Chevrolat, 1836. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Micraspiaires Mulsant, 1846: 162 [stem: Micraspid-]. Type genus: Micraspis Chevrolat, 1836. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
*Mysiates Mulsant, 1846: 125 [stem: Myzi-]. Type genus: Myzia Mulsant, 1846 [as Mysia, alternative original spelling of type genus name; we follow Kovář (2007: 620) in using Myzia as the correct spelling for this genus]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Halyziaires Mulsant, 1846: 123 [stem: Halyzi-]. Type genus: Halyzia Mulsant, 1846. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Pakaluk et al. (1994: 234, as Halyziini).
Alesiaires Mulsant, 1850: 343 [stem: Alesi-]. Type genus: Alesia Mulsant, 1850 [syn. of Micraspis Chevrolat, 1836]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Mader (1934: 297, as Alesiini), generally accepted as in Mader (1954: 93, as Alesiina).
*Cariaires Mulsant, 1850: 228 [stem: Cari-]. Type genus: Caria Mulsant, 1850 [preoccupied genus name, not Caria Hübner, 1823[ Lepidoptera]; syn. of Megalocaria Crotch, 1874]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Casey (1899: 72, as Cariini), but not generally accepted as valid; if name found to be available then permanently invalid (Art. 39): based on preoccupied type genus.
Coelophoraires Mulsant, 1850: 374 [stem: Coelophor-]. Type genus: Coelophora Mulsant, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Mader (1954: 93, as Coelophorina).
Cydoniaires Mulsant, 1850: 429 [stem: Cydoni-]. Type genus: Cydonia Mulsant, 1850 [syn. of Cheilomenes Chevrolat, 1836]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Mader (1954: 93, as Cydoniina).
Discotomaires Mulsant, 1850: 214 [stem: Discotom-]. Type genus: Discotoma Mulsant, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Korschevsky (1932: 577, as Discotomini), generally accepted as in Pakaluk et al. (1994: 234, as Discotomini).
Tytthaspides Crotch, 1874: xiii [stem: Tytthaspid-]. Type genus: Tytthaspis Crotch, 1874.
Synonychini Weise, 1885: 7 [stem: Synonych-]. Type genus: Synonycha Chevrolat, 1836.
Psylloborini Casey, 1899: 73 [stem: Psyllobor-]. Type genus: Psyllobora Chevrolat, 1836.
Anisostictina Jakobson, 1915: 969 [stem: Anisostict-]. Type genus: Anisosticta Chevrolat, 1836.
Anisolemniina Mader, 1954: 93, in key [stem: Anisolemni-]. Type genus: Anisolemnia Crotch, 1874.
Bulaeini Savoyskaya, 1969: 102 [stem: Bulae-]. Type genus: Bulaea Mulsant, 1850.
Singhikaliini Miyatake, 1972: 96 [stem: Singhikali-]. Type genus: Singhikalia Kapur, 1963.
Tribe
Cranophorini
Mulsant, 1850
Cranophoraires Mulsant, 1850: 939 [stem: Cranophor-]. Type genus: Cranophorus Mulsant, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Casey (1899: 74, as Cranophorini), generally accepted as in Pakaluk et al. (1994: 233, as Cranophorini).
Tribe
Cryptognathini
Mulsant, 1850
Cryptognathaires Mulsant, 1850: 496 [stem: Cryptognath-]. Type genus: Cryptognatha Mulsant, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gordon (1971: 181, as Cryptognathini), generally accepted as in Pakaluk et al. (1994: 249, as Cryptognathini).
*Pentiliaires Mulsant, 1850: 501 [stem: Pentili-]. Type genus: Pentilia Mulsant, 1850. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant (1846).
Pentiliini Casey, 1899: 74 [stem: Pentili-]. Type genus: Pentilia Mulsant, 1850.
Oeneini Casey, 1899: 74 [stem: Oene-]. Type genus: Oeneis Mulsant, 1850 [preoccupied genus name, not Oeneis Hübner, 1819 [Lepidoptera]; syn. of Delphastus Casey, 1899]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; the taxon Oeneini Wheeler, 1903 (type genus Oeneis Hübner, 1819) is available in Lepidoptera.
Tribe
Cynegetini
Thomson, 1866
Cynegetides C. G. Thomson, 1866: 374 [stem: Cyneget-]. Type genus: Cynegetis Chevrolat, 1836. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Cynegetid-).
Madaini Gordon, 1975: 206 [stem: Mad-]. Type genus: Mada Mulsant, 1850. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Diomini
Gordon, 1999
Diomini Gordon, 1999: 4 [stem: Diom-]. Type genus: Diomus Mulsant, 1850.
Tribe
Epilachnini
Mulsant, 1846
Epilachniens Mulsant, 1846: 190 [stem: Epilachn-]. Type genus: Epilachna Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Ganglbauer (1898: 947, as Epilachninae), generally accepted as in Pakaluk et al. (1994: 234, as Epilachnini).
*Chnootribaires Mulsant, 1850: 697 [stem: Chnootrib-]. Type genus: Chnootriba Chevrolat, 1836. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Subcoccinellini Jakobson, 1915: 968 [stem: Subcoccinell-]. Type genus: Subcoccinella Huber, 1841.
Tribe
Epivertini
Pang and Mao, 1979
Epivertini Pang and Mao, 1979: 158 [stem: Epivert-]. Type genus: Epiverta Dieke, 1947.
Tribe
Eremochilini
Gordon and Vanderberg, 1987
Eremochilini Gordon and Vanderberg, 1987: 6 [stem: Eremochil-]. Type genus: Eremochilus Weise, 1912.
Tribe
Hyperaspidini
Mulsant, 1846
Hypéraspiens Mulsant, 1846: 177 [stem: Hyperaspid-]. Type genus: Hyperaspis Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1885: 5, as Hyperaspini), generally accepted as in Pakaluk et al. (1994: 233, as Hyperaspini); incorrect original stem formation, not in prevailing usage.
*Thalassaires Mulsant, 1850: 505 [stem: Thalass-]. Type genus: Thalassa Mulsant, 1850. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Tiphysaires Mulsant, 1850: 516 [stem: Tiphys-]. Type genus: Tiphysa Mulsant, 1850. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Tribe
Limnichopharini
Miyatake, 1994
Limnichopharini Miyatake, 1994: 269 [stem: Limnichophar-]. Type genus: Limnichopharus Miyatake, 1994.
Tribe
Monocorynini
Miyatake, 1988
Monocorynini Miyatake, 1988: 28 [stem: Monocoryn-]. Type genus: Monocoryna Gorham, 1885.
Tribe
Noviini
Mulsant, 1846
Noviaires Mulsant, 1846: 213 [stem: Novi-]. Type genus: Novius Mulsant, 1846 [the original spelling used by Mulsant was Nomius (p. 213), however, the spelling was changed to Novius by Mulsant in the “Addenda et errata” of the same work and is therefore considered a justified emendation (Art. 19.2)]. Comment: the original spelling of this family-group name was Nomiaires (based on the new genus Nomius Mulsant, 1846), the spelling of the family-group and genus-group names were changed to Noviaires and Novius in the “Addenda et errata” of the same work, Noviaires is therefore treated as the correct original spelling of this family-group name (Art. 19.2); original vernacular name available (Art. 11.7.2): first used in latinized form by Ganglbauer (1899: 977, as Noviini), generally accepted as in Pakaluk et al. (1994: 233, as Noviini).
*Rodoliaires Mulsant, 1850: 901 [stem: Rodoli-]. Type genus: Rodolia Mulsant, 1850. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Tribe
Ortaliini
Mulsant, 1850
Ortaliens Mulsant, 1850: 892 [stem: Ortali-]. Type genus: Ortalia Mulsant, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Crotch (1874: 274, as Ortaliae), generally accepted as in Pakaluk et al. (1994: 234, as Ortaliini).
Tribe
Oryssomini
Gordon, 1974
Oryssomini Gordon, 1974: 146 [stem: Oryssom-]. Type genus: Oryssomus Mulsant, 1850.
Tribe
Platynaspini
Mulsant, 1846
Platynaspiaires Mulsant, 1846: 215 [stem: Platynasp-]. Type genus: Platynaspis Redtenbacher, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Casey (1899: 109, as Platynaspini), generally accepted as in Pakaluk et al. (1994: 234, as Platynaspini); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Platynaspid-).
Tribe
Plotinini
Miyatake, 1994
Plotinini Miyatake, 1994: 279 [stem: Plotin-]. Type genus: Plotina Lewis, 1896.
Tribe
Poriini
Mulsant, 1850
Poriens Mulsant, 1850: 884 [stem: Pori-]. Type genus: Poria Mulsant, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Gorham (1894: 206, as Poriides [treated as Latin]).
Tribe
Scymnillini
Casey, 1899
Scymnillini Casey, 1899: 74 [stem: Scymnill-]. Type genus: Scymnillus G. H. Horn, 1895.
Zilini Gordon, 1985: 74 [stem: Zil-]. Type genus: Zilus Mulsant, 1850. Comment: name proposed to replace Scymnillini Casey, 1899 because of the synonymy of the type genus at that time.
Tribe
Scymnini
Mulsant, 1846
Scymniaires Mulsant, 1846: 189 [stem: Scymn-]. Type genus: Scymnus Kugelann, 1794. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1885: 5, as Scymnini), generally accepted as in Pakaluk et al. (1994: 233, as Scymnini); incorrect original stem formation, not in prevailing usage.
*Cryptolaemaires Mulsant, 1853: 267 [stem: Cryptolaem-]. Type genus: Cryptolaemus Mulsant, 1853. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Tribe
Selvadiini
Gordon, 1985
Selvadiini Gordon, 1985: 347 [stem: Selvadi-]. Type genus: Selvadius Casey, 1899.
Tribe
Shirozuellini
Sasaji, 1967
Shirozuellini Sasaji, 1967: 23 [stem: Shirozuell-]. Type genus: Shirozuella Sasaji, 1967.
Ghaniini Ahmad, 1973: 449 [stem: Ghani-]. Type genus: Ghanius Ahmad, 1973.
Tribe
Stethorini
Dobzhansky, 1924
Stethorini Dobzhansky, 1924: 20 [stem: Stethor-]. Type genus: Stethorus Weise, 1885.
Tribe
Sticholotidini
Weise, 1901
Pharini Casey, 1899: 74 [stem: Phar-]. Type genus: Pharus Mulsant, 1850 [preoccupied genus name, not Pharus Gray, 1840 [Mollusca]; syn. of Pharoscymnus Bedel, 1906]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; this family-group name was also proposed in the same year by Ganglbauer (1899: 954, as Pharini), priority could not be established between those two works; the name Pharinae Adams and Adams, 1856 in Mollusca (type genus Pharus Gray, 1840) is available.
Sticholotini Weise, 1901: 430 [stem: Sticholotid-]. Type genus: Sticholotis Crotch, 1874. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Gordon (1977: 186).
Clanini Weise, 1901: 430 [stem: Clan-]. Type genus: Clanis Mulsant, 1850 [preoccupied genus name, not Clanis Hübner, 1819 [Lepidoptera]; syn. of Jauravia Motschulsky, 1858]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Coelopterini Weise, 1906: column 369 [stem: Coelopter-]. Type genus: Coelopterus Mulsant and Rey, 1852. Comment: name previously attributed to Della Beffa (1912: 171, as Coleopterini [incorrect stem formation]) in the literature.
Tribe
Telsimiini
Casey, 1899
Telsimiini Casey, 1899: 74 [stem: Telsimi-]. Type genus: Telsimia Casey, 1899.
Tribe
Tetrabrachini
Kapur, 1948
Lithophilidae Imhoff, 1856: [2] 151 [stem: Lithophil-]. Type genus: Lithophilus Fröhlich, 1799 [preoccupied genus name, not Lithophilus Schneider, 1791 [Coleoptera: Carabidae]; syn. of Tetrabrachys Kapur, 1948]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tetrabrachinae Kapur, 1948: 320 [stem: Tetrabrach-]. Type genus: Tetrabrachys Kapur, 1948. Comment: replacement name for Lithophilinae Imhoff, 1856 because of the homonymy of the type genus; current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Tetrabrache-).
Family
Corylophidae
LeConte, 1852
Corylophi J. L. LeConte, 1852b: 141 [stem: Coryloph-]. Type genus: Corylophus Stephens, 1833.
Subfamily
Periptyctinae
Ślipiński, Lawrence and Tomaszewska, 2001
Periptyctinae Ślipiński et al., 2001: 312 [stem: Periptyct-]. Type genus: Periptyctus Blackburn, 1825.
Subfamily
Corylophinae
LeConte, 1852
Corylophi J. L. LeConte, 1852b: 141 [stem: Coryloph-]. Type genus: Corylophus Stephens, 1833.
Tribe
Aenigmaticini
Casey, 1900
Aenigmaticini Casey, 1900: 61 [stem: Aenigmatic-]. Type genus: Aenigmaticum A. Matthews, 1888.
Tribe
Cleidostethini
Bowestead, Booth, Ślipiński and Lawrence, 2001
Cleidostethini Bowestead et al., 2001: 323 [stem: Cleidosteth-]. Type genus: Cleidostethus Arrow, 1929.
Tribe
Corylophini
LeConte, 1852
Corylophi J. L. LeConte, 1852b: 141 [stem: Coryloph-]. Type genus: Corylophus Stephens, 1833.
Tribe
Foadiini
Ślipiński, Tomaszewska and Lawrence, 2009
Foadiini Ślipiński et al., 2009: 422 [stem: Foadi-]. Type genus: Foadia Pakaluk, 1985.
Tribe
Orthoperini
Jacquelin du Val, 1857
Orthopérites Jacquelin du Val, 1857b: 100 [stem: Orthoper-]. Type genus: Orthoperus Stephens, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. G. Thomson (1859: 63, as Orthoperidae), generally accepted as in Ślipiński et al. (2009: 428, as Orthoperini).
Tribe
Parmulini
Poey, 1854
Clypeastres L. Redtenbacher, 1845: 122 [stem: Clypeaster-]. Type genus: Clypeaster Dejean, 1821 [preoccupied genus name, not Clypeaster Lamarck, 1801 [Echinodermata]; syn. of Clypastraea Haldeman, 1842]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage; Clypeasteridae L. Agassiz, 1835 (type genus Clypeaster Lamarck, 1801) is available in Echinodermata.
Parmulini Poey, 1854: 323 [stem: Parmul-]. Type genus: Parmulus Gundlach, 1854 [syn. of Clypastraea Haldeman, 1842].
Saciina A. Matthews, 1888: 103 [stem: Saci-]. Type genus: Sacium J. L. LeConte, 1852 [syn. of Clypastraea Haldeman, 1842].
Arthrolipinae Böving and Craighead, 1931: 36, in key [stem: Arthrolip-]. Type genus: Arthrolips Wollaston, 1854.
Tribe
Peltinodini
Paulian, 1950
Peltinoditae Paulian, 1950: 19 [stem: Peltinod-]. Type genus: Peltinodes Paulian, 1950 [syn. of Holopsis Broun, 1883]. Comment: precedence (Peltinodini Paulian, 1950 vs Corylophodini Paulian, 1950) given to taxon originally proposed at the higher rank (Art. 24.1).
Corylophodini Paulian, 1950: 21, in key [stem: Corylophod-]. Type genus: Corylophodes A. Matthews, 1885 [syn. of Holopsis Broun, 1883].
Tribe
Rypobiini
Paulian, 1950
Rhypobiini Paulian, 1950: 48 [stem: Rypobi-]. Type genus: Rypobius J. L. LeConte, 1852 [as Rhypobius, unjustified emendation of type genus name by Gemminger and Harold (1876: 3818), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Gloeosomatini Bowestead, 1999: 128 [stem: Gloeosomat-]. Type genus: Gloeosoma Wollaston, 1854.
Tribe
Sericoderini
Matthews, 1888
Sericoderina A. Matthews, 1888: 103 [stem: Sericoder-]. Type genus: Sericoderus Stephens, 1829.
Tribe
Teplinini
Pakaluk, Ślipiński and Lawrence, 1994
Peltinini Paulian, 1950: 21, in key [stem: Peltin-]. Type genus: Peltinus Mulsant, 1861 [preoccupied genus name, not Peltinus Rafinesque, 1815 [Coleoptera: Trogossitidae]; syn. of Teplinus Pakaluk et al., 1994]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Teplinini Pakaluk et al., 1994: 251 [stem: Teplin-]. Type genus: Teplinus Pakaluk et al., 1994. Comment: replacement name for Peltinini Paulian, 1950 because of the homonymy of the type genus.
Family
Akalyptoischiidae
Lord, Hartley, Lawrence, McHugh and Miller, 2010
Akalyptoischiidae Lord et al., 2010: 761 [stem: Akalyptoischi-]. Type genus: Akalyptoischion Andrews, 1976.
Family
Latridiidae
Erichson, 1842
Lathridien Erichson, 1842: 122 [stem: Latridi-]. Type genus: Latridius Herbst, 1793 [as Lathridius, unjustified emendation of type genus name by Illiger (1802), not in prevailing usage]. Comment: the younger name Latridiidae has been used widely for this family in recent literature although Pakaluk et al. (1994: 252) and a small number of subsequent authors, e.g., Míka (2000), Lassau et al. (2005), have used the older name Corticariidae; an application was recently submitted to the Commission in order to conserve usage of the well-established name Latridiidae (Bousquet et al. 2010; see Appendix 6).
Subfamily
Latridiinae
Erichson, 1842
Lathridien Erichson, 1842: 122 [stem: Latridi-]. Type genus: Latridius Herbst, 1793 [as Lathridius, unjustified emendation of type genus name by Illiger (1802), not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by L. Redtenbacher (1845: 123, as Lathridii), generally accepted as in Lawrence and Newton (1995: 886, as Latridiidae); incorrect original stem formation, not in prevailing usage.
Subfamily
Corticariinae
Curtis, 1829
Corticaridae Curtis, 1829: pl. 283 [stem: Corticari-]. Type genus: Corticaria Marsham, 1802. Comment: incorrect original stem formation, not in prevailing usage; although this is the oldest name for the family, an application was recently submitted by Bousquet et al. (2010) to conserve usage of the well-established name Latridiidae (see Appendix 6).
*Melanophthalmidae Arnett, 1962b: 835 [stem: Melanophthalm-]. Type genus: Melanophthalma Motschulsky, 1866. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently; name listed by Arnett as a synonym of Latridiidae and attributed to “auct.”, we could find an earlier usage of this name.
†Subfamily
Tetrameropseinae
Kirejtshuk and Azar, 2008
Tetrameropsinae Kirejtshuk and Azar, 2008: 36 [stem: Tetrameropse-]. Type genus: Tetrameropsis Kirejtshuk and Azar, 2008. Comment: incorrect original stem formation, not in prevailing usage.
Superfamily
Tenebrionoidea
Latreille, 1802
Tenebrionites Latreille, 1802: 165 [stem: Tenebrion-]. Type genus: Tenebrio Linnaeus, 1758. Comment: First Revisers found (Tenebrionoidea Latreille, 1802 vs Mordelloidea Latreille, 1802) are Alonso-Zarazaga and Mansilla-Castrillo (1988: 19).
Family
Mycetophagidae
Leach, 1815
Mycetophagida Leach, 1815: 110 [stem: Mycetophag-]. Type genus: Mycetophagus Fabricius, 1792.
Subfamily
Esarcinae
Reitter, 1882
Esarcini Reitter, 1882a: 115 [stem: Esarc-]. Type genus: Esarcus Reiche, 1864.
Subfamily
Mycetophaginae
Leach, 1815
Mycetophagida Leach, 1815: 110 [stem: Mycetophag-]. Type genus: Mycetophagus Fabricius, 1792.
Tribe
Mycetophagini
Leach, 1815
Mycetophagida Leach, 1815: 110 [stem: Mycetophag-]. Type genus: Mycetophagus Fabricius, 1792.
Tritomidae Crotch, 1873a: 78 [stem: Tritom-]. Type genus: Tritoma Geoffroy, 1762 [senior homonym of Tritoma Fabricius, 1775 [Coleoptera: Erotylidae] but placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1994a); syn. of Mycetophagus Fabricius, 1792]. Comment: permanently invalid (Art. 39): based on suppressed type genus; Tritomidae was also used the same year by Crotch (1873b: 42).
Triphyllini Harold, 1880: 757 [stem: Triphyll-]. Type genus: Triphyllus Dejean, 1821.
Tribe
Typhaeini
Thomson, 1863
Typhaeina C. G. Thomson, 1863: 241 [stem: Typhae-]. Type genus: Typhaea Stephens, 1829.
Typhaeini Nikitsky, 1993: 165 [stem: Typhae-]. Type genus: Typhaea Stephens, 1829. Comment: family-group name proposed as new without reference to Typhaeina C. G. Thomson, 1863.
Subfamily
Bergininae
Leng, 1920
Bergini Leng, 1920: 246 [stem: Bergin-]. Type genus: Berginus Erichson, 1846. Comment: incorrect original stem formation, not in prevailing usage.
Family
Archeocrypticidae
Kaszab, 1964
Archeocrypticini Kaszab, 1964: 361 [stem: Archeocryptic-]. Type genus: Archeocrypticus Kaszab, 1964.
Family
Pterogeniidae
Crowson, 1953
Pterogeniidae Crowson, 1953: 38 [stem: Pterogeni-]. Type genus: Pterogenius Candèze, 1861.
Family
Ciidae
Leach, 1819
Cisidae Leach, 1819: 206 [stem: Ci-]. Type genus: Cis Latreille, 1797. Comment: incorrect original stem formation, not in prevailing usage; Gistel (1848: [6], 1856a: 368) used the name Microtrocteidae for a family that included only the genus Cis Latreille, we could not find any genus name on which Microtrocteidae could be based on therefore this family-group name is unavailable.
Subfamily
Sphindociinae
Lawrence, 1974
Sphindociinae Lawrence, 1974b: 9 [stem: Sphindoci-]. Type genus: Sphindocis Fall, 1917. Comment: published 28 June 1974; this family-group name was also used in the same year by Lawrence (1974a [22 January]: 24, as Sphindociinae) but the older name is unavailable because it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subfamily
Ciinae
Leach, 1819
Cisidae Leach, 1819: 206 [stem: Ci-]. Type genus: Cis Latreille, 1797.
Tribe
Ciini
Leach, 1819
Cisidae Leach, 1819: 206 [stem: Ci-]. Type genus: Cis Latreille, 1797. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Orophiini
Thomson, 1863
Orophiina C. G. Thomson, 1863: 195 [stem: Orophi-]. Type genus: Orophius L. Redtenbacher, 1848 [syn. of Octotemnus Mellié, 1847].
Octotemnidae Reitter, 1878: 21 [stem: Octotemn-]. Type genus: Octotemnus Mellié, 1847.
Rhopalodontini Everts, 1898: 517, in key [stem: Ropalodont-]. Type genus: Ropalodontus Mellié, 1847 [as Rhopalodontus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: the fossil Reptilia name Rhopalodontidae Seeley, 1894 (type genus Rhopalodon Fischer von Waldheim, 1841) is available, therefore we recomment using the spelling Ropalodont- for the ciid name in order to avoid homonymy problems; incorrect original stem formation, not in prevailing usage.
Tribe
Xylographellini
Kawanabe and Miyatake, 1996
Xylographellini Kawanabe and Miyatake, 1996: 125 [stem: Xylographell-]. Type genus: Xylographella Miyatake, 1985.
Subtribe
Syncosmetina
Lopes-Andrade, 2008
Syncosmetina Lopes-Andrade, 2008: 40 [stem: Syncosmet-]. Type genus: Syncosmetus Sharp, 1891.
Subtribe
Xylographellina
Kawanabe and Miyatake, 1996
Xylographellini Kawanabe and Miyatake, 1996: 125 [stem: Xylographell-]. Type genus: Xylographella Miyatake, 1985.
Family
Tetratomidae
Billberg, 1820
Tetratomaedes Billberg, 1820a: 34 [stem: Tetratom-]. Type genus: Tetratoma Fabricius, 1790.
Subfamily
Tetratominae
Billberg, 1820
Tetratomaedes Billberg, 1820a: 34 [stem: Tetratom-]. Type genus: Tetratoma Fabricius, 1790.
Subfamily
Piseninae
Miyatake, 1960
Pisenini Miyatake, 1960: 124 [stem: Pisen-]. Type genus: Pisenus Casey, 1900.
Subfamily
Penthinae
Lacordaire, 1859
Penthides Lacordaire, 1859: 456 [stem: Penth-]. Type genus: Penthe Newman, 1838. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by G. H. Horn (1888: 43, as Penthini), generally accepted as in Young and Pollock (2002: 415, as Penthinae).
Subfamily
Hallomeninae
Gistel, 1848
Hallomenidae Gistel, 1848: [11] [stem: Hallomen-]. Type genus: Hallomenus Panzer, 1793.
*Dryalates Mulsant, 1856b: 44 [stem: Dryal-]. Type genus: Dryala Mulsant, 1856 [syn. of Hallomenus Panzer, 1793]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Mycétomiens Mulsant, 1856b: 103 [stem: Mycetomat-]. Type genus: Mycetoma Dejean, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Desbrochers des Loges (1900: 16, as Mycetomini); incorrect original stem formation, not in prevailing usage.
Subfamily
Eustrophinae
Gistel, 1848
Eustrophidae Gistel, 1848: [10] [stem: Eustroph-]. Type genus: Eustrophus Illiger, 1802.
Tribe
Eustrophini
Gistel, 1848
Eustrophidae Gistel, 1848: [10] [stem: Eustroph-]. Type genus: Eustrophus Illiger, 1802.
Tribe
Holostrophini
Nikitsky, 1998
Holostrophini Nikitsky, 1998: 39 [stem: Holostroph-]. Type genus: Holostrophus G. H. Horn, 1888.
Family
Melandryidae
Leach, 1815
Melyandrida Leach, 1815: 104 [stem: Melandry-]. Type genus: Melandrya Fabricius, 1801.
Subfamily
Melandryinae
Leach, 1815
Melyandrida Leach, 1815: 104 [stem: Melandry-]. Type genus: Melandrya Fabricius, 1801.
Tribe
Anisoxiellini
Nikitsky, 2007
Anisoxiellini Nikitsky, 2007: 58 [stem: Anisoxiell-]. Type genus: Anisoxiella Nikitsky, 1989.
Tribe
Dircaeini
Kirby, 1837
Dircaeidae Kirby, 1837: 240 [stem: Dircae-]. Type genus: Dircaea Fabricius, 1798.
Tribe
Hypulini
Gistel, 1848
Hypulidae Gistel, 1848: [11] [stem: Hypul-]. Type genus: Hypulus Paykull, 1798.
Maroliini Portevin, 1934: 57, in key [stem: Maroli-]. Type genus: Marolia Mulsant, 1856.
Tribe
Melandryini
Leach, 1815
Melyandrida Leach, 1815: 104 [stem: Melandry-]. Type genus: Melandrya Fabricius, 1801. Comment: incorrect original stem formation, not in prevailing usage.
*Phryganophiles Motschulsky, 1849: 58 [stem: Phryganophil-]. Type genus: Phryganophilus Sahlberg, 1833. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Hylepnigalionidae Gistel, 1856a: 384 [stem: Hylepnigalion-]. Type genus: Hylepnigalio Gistel, 1856 [this genus originally included the species caraboides Linnaeus and caniculatus (without author name); the type species of Hylepnigalio Gistel, 1856 is here considered to be Chrysomela caraboides Linnaeus, 1760 by monotypy; syn. nov. of Melandrya Fabricius, 1801]. Comment: syn. nov.
Tribe
Orchesiini
Mulsant, 1856
Orchésiens Mulsant, 1856b: 27 [stem: Orchesi-]. Type genus: Orchesia Latreille, 1807. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. G. Thomson (1859: 119, as Orchesiina), generally accepted as in Nikitsky and Pollock (2008: 68, as Orchesiini).
Tribe
Serropalpini
Latreille, 1829
Serropalpides Latreille, 1829b: 43 [stem: Serropalp-]. Type genus: Serropalpus Hellenius, 1786.
Tribe
Xylitini
Thomson, 1864
Xylitina C. G. Thomson, 1864: 316 [stem: Xylit-]. Type genus: Xylita Paykull, 1798.
Tribe
Zilorini
Desbrochers des Loges, 1900
Zilorini Desbrochers des Loges, 1900: 2, in key [stem: Zilor-]. Type genus: Zilora Mulsant, 1856. Comment: spelled Ziloriini in description on page 2 (in key), but correctly spelled Zilorini on p. 17 of the same work.
Zilorini Nikitsky, 2007: 60 [stem: Zilor-]. Type genus: Zilora Mulsant, 1856. Comment: family-group name proposed as new without reference to Zilorini Desbrochers des Loges, 1900.
Subfamily
Osphyinae
Mulsant, 1856 (1839)
Nothidae Shuckard, 1839b: 51 [stem: Noth-]. Type genus: Nothus A. G. Olivier, 1811 [syn. of Osphya Illiger, 1807]. Comment: Osphyinae Mulsant, 1856 conserved over this name (Art. 40.2) (see Lawrence and Newton 1995: 888).
*Osphies Motschulsky, 1849: 59 [stem: Osphy-]. Type genus: Osphya Illiger, 1807. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849); incorrect original stem formation, not in prevailing usage.
Osphyens Mulsant, 1856b: 108 [stem: Osphy-]. Type genus: Osphya Illiger, 1807. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1875 [Gatt.]: 103, as Osphyini), generally accepted as in Pollock (2002: 421, as Osphyinae); name conserved over the older Nothinae Shuckard, 1839 (Art. 40.2) (see Lawrence and Newton 1995: 888).
Conopalpiens Mulsant, 1856b: 105 [stem: Conopalp-]. Type genus: Conopalpus Gyllenhal, 1810. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. G. Thomson (1864: 313, as Conopalpina), generally accepted as in Hansen (1996: 171, as Conopalpini).
Family
Mordellidae
Latreille, 1802
Mordellonae Latreille, 1802: 183 [stem: Mordell-]. Type genus: Mordella Linnaeus, 1758.
†Praemordellinae Ščegoleva-Barovskaya, 1929: 27 [stem: Praemordell-]. Type genus: Praemordella Ščegoleva-Barovskaya, 1929.
†Liaoximordellidae Wang, 1993: 87 [stem: Liaoximordell-]. Type genus: Liaoximordella Wang, 1993.
Subfamily
Ctenidiinae
Franciscolo, 1951
Ctenidiinae Franciscolo, 1951: 56 [stem: Ctenidi-]. Type genus: Ctenidia Laporte, 1840.
Subfamily
Mordellinae
Latreille, 1802
Mordellonae Latreille, 1802: 183 [stem: Mordell-]. Type genus: Mordella Linnaeus, 1758.
Tribe
Conaliini
Ermisch, 1956
Conaliini Ermisch, 1956: 273 [stem: Conali-]. Type genus: Conalia Mulsant and Rey, 1858.
Tribe
Mordellini
Latreille, 1802
Mordellonae Latreille, 1802: 183 [stem: Mordell-]. Type genus: Mordella Linnaeus, 1758.
Tribe
Mordellistenini
Ermisch, 1941
Mordellistenini Ermisch, 1941: 714 [stem: Mordellisten-]. Type genus: Mordellistena A. Costa, 1854.
Tribe
Reynoldsiellini
Franciscolo, 1957
Reynoldsiellini Franciscolo, 1957: 237 [stem: Reynoldsiell-]. Type genus: Reynoldsiella Ray, 1930.
Tribe
Stenaliini
Franciscolo, 1955
Stenaliini Franciscolo, 1955: 177 [stem: Stenali-]. Type genus: Stenalia Mulsant, 1856.
Family
Ripiphoridae
Gemminger, 1870 (1855)
Rhipidophoridae Gemminger, 1870: 2117 [stem: Ripiphor-]. Type genus: Ripiphorus Bosc, 1791 [as Rhipidophorus, unjustified emendation of Ripiphorus Bosc, 1791 by Perty (1831: xix), not in prevailing usage]. Comment: name conserved over the older Myoditidae Gerstaecker, 1855 (Art. 40.2) (see Lawrence and Newton 1995: 889, as “Myoditini Costa, 1853”); First Reviser (Ripiphoridae Gemminger, 1870 (1855) vs Ptilophoridae Gerstaecker, 1855 vs Ripidiidae Gerstaecker, 1855) not determined, prevailing usage maintained.
Subfamily
Ptilophorinae
Gerstaecker, 1855
*Ptilophores Motschulsky, 1849: 59 [stem: Ptilophor-]. Type genus: Ptilophorus Dejean, 1834. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Ptilophorini Gerstaecker, 1855: 2 [stem: Ptilophor-]. Type genus: Ptilophorus Dejean, 1834.
Évaniocérides Lacordaire, 1859: 618 [stem: Evaniocer-]. Type genus: Evaniocera Guérin-Méneville, 1835 [syn. of Ptilophorus Dejean, 1834]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 276, as Evaniocerini), generally accepted as in J. L. LeConte and G. H. Horn (1883: xxxvii, as Evaniocerini).
Subfamily
Pelecotominae
Seidlitz, 1875
Pelecotomini Seidlitz, 1875 [Gatt.]: 104 [stem: Pelecotom-]. Type genus: Pelecotoma Fischer von Waldheim, 1808.
Micholaeminae Viana, 1971: 69 [stem: Micholaem-]. Type genus: Micholaemus Viana, 1971.
Subfamily
Hemirhipidiinae
Heller, 1921
Hemirhipidiini Heller, 1921: 168 [stem: Hemirhipidi-]. Type genus: Hemirhipidius Heller, 1921 [syn. of Nephrites Shuckard, 1838].
Nephritinae Selander, 1957: 101 [stem: Nephrit-]. Type genus: Nephrites Shuckard, 1838. Comment: name proposed to replace Hemirhipidiini Heller because of the synonymy of the type genus, Hemirhipidiinae is in prevailing usage (see Lawrence and Newton, 1995: 889).
Subfamily
Ripidiinae
Gerstaecker, 1855
Rhipidiini Gerstaecker, 1855: 14 [stem: Ripidi-]. Type genus: Ripidius Thunberg, 1806 [as Rhipidius, unjustified emendation of type genus name by Agassiz (1846b: 327), not in prevailing usage].
Tribe
Eorhipidiini
Iablokoff-Khnzorian, 1986
Eorhipidiini Iablokoff-Khnzorian, 1986: 92 [stem: Eorhipidi-]. Type genus: Eorhipidius Iablokoff-Khnzorian, 1986.
Tribe
Ripidiini
Gerstaecker, 1855
Rhipidiini Gerstaecker, 1855: 14 [stem: Ripidi-]. Type genus: Ripidius Thunberg, 1806 [as Rhipidius, unjustified emendation of type genus name by Agassiz (1846b: 327), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; spelling of family-group and type genus names based on Krell (1996).
Subfamily
Ripiphorinae
Gemminger, 1870 (1855)
Rhipidophoridae Gemminger, 1870: 2117 [stem: Ripiphor-]. Type genus: Ripiphorus Bosc, 1791 [as Rhipidophorus, unjustified emendation of Ripiphorus Bosc, 1791 by Perty (1831: xix), not in prevailing usage]. Comment: name conserved over the older Myoditinae Gerstaecker, 1855 (Art. 40.2) (see Lawrence and Newton 1995: 889, as “Myoditini Costa, 1853”).
Tribe
Macrosiagonini
Heyden, 1908
Rhipiphorites Laporte, 1840b: 261 [stem: Ripiphor-]. Type genus: Ripiphorus sensu Fabricius, 1792 [as Rhipiphorus, incorrect subsequent spelling of the type genus name, not in prevailing usage; not Ripiphorus Bosc, 1791; syn. of Metoecus Dejean, 1834]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Gerstaecker (1855: 17, as Rhipiphorini [incorrect stem formation]); based on a misidentified type genus; name treated here as invalid until an application is submitted to the Commission to suppress it for the Principles of Priority and Homonymy (Art. 65.2.1); also see Ripiphorini Gemminger, 1870.
Macrosiagonini L. Heyden, 1908: 45 [stem: Macrosiagon-]. Type genus: Macrosiagon Hentz, 1830. Comment: although this is not the oldest name for the tribe, we recommend that an application be sent to the Commission to suppress Ripiphorini Laporte, 1840 because it is based on a misidentified type genus (Art. 65.2.1).
Tribe
Ripiphorini
Gemminger, 1870 (1855)
*Mioditini A. Costa, 1853: 2 [stem: Myodit-]. Type genus: Myodites Latreille, 1829 [syn. of Ripiphorus Bosc, 1791]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally treated as valid attributed to A. Costa, 1853; incorrect original stem formation, not in prevailing usage.
Myoditini Gerstaecker, 1855: 15 [stem: Myodit-]. Type genus: Myodites Latreille, 1829 [syn. of Ripiphorus Bosc, 1791]. Comment : usage of younger name Ripiphorini Gemminger, 1870 conserved over this name (Art. 40.2) (see Lawrence and Newton 1995: 889, as “Myoditini Costa, 1853”).
Rhipidophoridae Gemminger, 1870: 2117 [stem: Ripiphor-]. Type genus: Ripiphorus Bosc, 1791 [as Rhipidophorus, unjustified emendation of type genus name by Perty (1831: xix), not in prevailing usage]. Comment: name conserved over the older Myoditini Gerstaecker, 1855 (Art. 40.2) (see Lawrence and Newton 1995: 889, as “Myoditini Costa, 1853”); incorrect original stem formation, not in prevailing usage; as mentioned by Lawrence and Newton (1995: 889) a number of authors used Ripiphoridae before Gemminger, however, their family-group names were based on Ripiphorus sensu Fabricius, 1792, which is a synonym of Metoecus Dejean, 1834; spelling of family-group and type genus names based on Krell (1996); we recommend that an application be submitted to the Commission to suppress Ripiphorites Laporte, 1840 for the Principles of Priority and Homonymy (Art. 65.2.1).
Family
Zopheridae
Solier, 1834
Zophérites Solier, 1834: 505 [stem: Zopher-]. Type genus: Zopherus Gray, 1832. Comment: usage of this name conserved over Colydiidae Billberg, 1820 (Art. 35.5).
Subfamily
Colydiinae
Billberg, 1820
Colydiides Billberg, 1820b: 394 [stem: Colydi-]. Type genus: Colydium Fabricius, 1792 [placed on the Official List of Generic Names in Zoology (ICZN 1995c)]. Comment: name placed on Official List of Family-Group Names in Zoology (as Colydiidae Erichson, 1842) and given precedence over Orthocerinae Blanchard, 1845 and Cerylonidae Billberg, 1820 whenever their type genera are placed in the same family-group taxon (ICZN 1995c); usage of the younger name Zopheridae Solier, 1834 conserved over this name (Art. 35.5).
Tribe
Acropini
Sharp, 1894
Acropinae Sharp, 1894: 444 [stem: Acrop-]. Type genus: Acropis H. C. C. Burmeister, 1840. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Acropid-).
Tribe
Adimerini
Sharp, 1894
Adimeridae Sharp, 1894: 441 [stem: Adimer-]. Type genus: Adimerus Sharp, 1894.
Monoedidae Schaeffer, 1911: 114 [stem: Monoed-]. Type genus: Monoedus G. H. Horn, 1882.
Tribe
Colydiini
Billberg, 1820
Colydiides Billberg, 1820b: 394 [stem: Colydi-]. Type genus: Colydium Fabricius, 1792 [placed on the Official List of Generic Names in Zoology (ICZN 1995c)].
Tribe
Gempylodini
Sharp, 1893
Gempylodini Sharp, 1893: 256 [stem: Gempylod-]. Type genus: Gempylodes Pascoe, 1863.
Tribe
Nematidiini
Horn, 1878
Nematidii G. H. Horn, 1878: 573 [stem: Nematidi-]. Type genus: Nematidium Erichson, 1845.
Tribe
Orthocerini
Blanchard, 1845
Sarrotriides Billberg, 1820a: 9 [stem: Sarrotri-]. Type genus: Sarrotrium Illiger, 1798 [syn. of Orthocerus Latreille, 1797]. Comment: the same name was also published in the same year by Billberg (1820b: 390); Sarrotriidae Billberg, 1820 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1995c).
Orthocérites Blanchard, 1845b: 29 [stem: Orthocer-]. Type genus: Orthocerus Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 1995c)]. Comment: Orthocerini Blanchard, 1845 placed on Official List of Family-group names in Zoology (ICZN 1995c).
Euglochidae Gistel, 1856a: 382 [stem: Euglochin-]. Type genus: Euglochis Gistel, 1856 [syn. of Orthocerus Latreille, 1797]. Comment: published 18 February 1856; this family-group name was also used in the same year by Gistel (1856b [“31 December”]: 180, as Euglochida); incorrect original stem formation, not in prevailing usage.
Tribe
Rhagoderini
Horn, 1878
Rhagoderini G. H. Horn, 1878: 557 [stem: Rhagoder-]. Type genus: Rhagodera Mannerheim, 1843.
Tribe
Rhopalocerini
Reitter, 1911
Apistini Ganglbauer, 1899: 873 [stem: Apist-]. Type genus: Apistus Motschulsky, 1840 [Apistus is an unjustified emendation of type genus name, originally spelled Apeistus, by Agassiz (1846b: 28), in prevailing usage; the emended name is a junior homonym of Apistus Cuvier, 1829 [Pisces]; Apeistus Motschulsky, 1840 and Apistus Agassiz, 1846 were placed on Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1986b); syn. of Rhopalocerus Redtenbacher, 1842]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; placed on Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1986b).
Rhopalocerini Reitter, 1911: 108 [stem: Rhopalocer-]. Type genus: Rhopalocerus Redtenbacher, 1842 [placed on the Official List of Generic Names in Zoology (ICZN 1986b)]. Comment: placed on the Official List of Family-Group Names in Zoology (ICZN 1986b).
Tribe
Synchitini
Erichson, 1845
Synchitini Erichson, 1845: 254 [stem: Synchit-]. Type genus: Synchita Hellwig, 1792. Comment: published before 31 June 1845; this family-group name was also used in the same year by L. Redtenbacher (1845 [before September]: 123, as Synchitae).
Ditomidae Imhoff, 1856: [2] 159 [stem: Bitom-]. Type genus: Bitoma Herbst, 1793 [as Ditoma, unjustified emendation of type genus name by Illiger (1807), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; stem of the family-group name corrected according to Art. 35.4.1 and also to remove from potential homonymy with the older family-group name originally proposed as Ditomici Bonelli, 1810 (type genus Ditomus Bonelli, 1810) in the family Carabidae.
Coxelini Seidlitz, 1872 [Gatt.]: 38 [stem: Coxel-]. Type genus: Coxelus Dejean, 1821.
Langelandiina Fowler, 1889: 192 [stem: Langelandi-]. Type genus: Langelandia Aubé, 1843.
Megataphrini Casey, 1890: 309 [stem: Megataphr-]. Type genus: Megataphrus Casey, 1890.
Tarphiinae Sharp, 1894: 444 [stem: Tarphi-]. Type genus: Tarphius Erichson, 1845.
Corticini Ganglbauer, 1899: 870 [stem: Cortic-]. Type genus: Corticus Latreille, 1829.
Diodesmini Reitter, 1911: 110 [stem: Diodesmat-]. Type genus: Diodesma Latreille, 1829. Comment: incorrect original stem formation, not in prevailing usage.
Trachypholini Grouvelle, 1911: 121 [stem: Trachypholid-]. Type genus: Trachypholis Erichson, 1845. Comment: incorrect original stem formation, not in prevailing usage.
Endophloeini Reitter, 1922a: 17 [stem: Endophloe-]. Type genus: Endophloeus Dejean, 1834.
Priolomini Dajoz, 1980a: 127 [stem: Priolom-]. Type genus: Priolomus Erichson, 1845.
Subfamily
Zopherinae
Solier, 1834
Zophérites Solier, 1834: 505 [stem: Zopher-]. Type genus: Zopherus Gray, 1832.
Tribe
Latometini
Ślipiński and Lawrence, 1999
Latometini Ślipiński and Lawrence, 1999: 9, in key [stem: Latomet-]. Type genus: Latometus Erichson, 1842.
Tribe
Monommatini
Blanchard, 1845
Monommites Blanchard, 1845b: 16 [stem: Monommat-]. Type genus: Monomma Klug, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 246, as Monommidae [incorrect stem formation]), generally accepted as in Lawrence and Newton (1995: 890, as Monommatidae); incorrect original stem formation, not in prevailing usage.
Tribe
Phellopsini
Ślipiński and Lawrence, 1999
Phellopsini Ślipiński and Lawrence, 1999: 10, in key [stem: Phellops-]. Type genus: Phellopsis J. L. LeConte, 1862. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Phellopse-).
Tribe
Pycnomerini
Erichson, 1845
Pycnomerini Erichson, 1845: 290 [stem: Pycnomer-]. Type genus: Pycnomerus Erichson, 1842.
Tribe
Usechini
Horn, 1867
Usechini G. H. Horn, 1867a: 294 [stem: Usech-]. Type genus: Usechus Motschulsky, 1845.
Tribe
Zopherini
Solier, 1834
Zophérites Solier, 1834: 505 [stem: Zopher-]. Type genus: Zopherus Gray, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 392, as Zopheroidae), generally accepted as in Lawrence and Newton (1995: 891, as Zopheridae).
Nosodermini Casey, 1907: 280 [stem: Nosodermat-]. Type genus: Nosoderma Solier, 1841 [preoccupied genus name, not Nosoderma Guérin-Méneville, 1838 [Coleoptera: Zopheridae: Zopherinae: Zopherini]; syn. of Verodes Casey, 1907; see Foley and Ivie (2007) for comments on type genus]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; all subsequent uses of Nosodermini were based on the junior homonym Nosoderma Solier, 1841 (see Foley and Ivie 2007); incorrect original stem formation, not in prevailing usage.
Zopherosini Casey, 1907: 522 [stem: Zopherose-]. Type genus: Zopherosis A. White, 1859. Comment: incorrect original stem formation, not in prevailing usage.
Family
Ulodidae
Pascoe, 1869
Ulodinae Pascoe, 1869a: 31 [stem: Ulod-]. Type genus: Ulodes Erichson, 1842.
Merycidae Crowson, 1953: 37 [stem: Meryc-]. Type genus: Meryx Latreille, 1802.
Family
Promecheilidae
Lacordaire, 1859
Proméchilides Lacordaire, 1859: 698 [stem: Promecheil-]. Type genus: Promecheilus Solier, 1851 [as Promechilus, unjustified emendation of type genus name by Lacordaire (1859: 700), not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Handlirsch (1925: 594, as Promecheilinae), generally accepted as in Lawrence et al. (2010: 563, as Promecheilidae); incorrect original stem formation, not in prevailing usage.
Perimylopidae St. George, 1939: 212 [stem: Perimylop-]. Type genus: Perimylops Müller, 1884.
*Parahelopinae Watt, 1975: 424 [stem: Parahelop-]. Type genus: Parahelops C. O. Waterhouse, 1876. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such description (Art. 13.1); this name was used several times subsequently, e.g., Elgueta and Arriagada (1989: 37), Lawrence and Britton (1991: 665), Lawrence (1994: 339), but this taxon name remains unavailable (see Lawrence and Newton 1995: 892).
Family
Chalcodryidae
Watt, 1974
Chalcodryidae Watt, 1974: 24 [stem: Chalcodry-]. Type genus: Chalcodrya Redtenbacher, 1868.
Family
Trachelostenidae
Lacordaire, 1859
Trachélosténides Lacordaire, 1859: 567 [stem: Trachelosten-]. Type genus: Trachelostenus Solier, 1851. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1898: 319, as Trachelostenini), generally accepted as in Lawrence and Newton (1995: 892 as Trachelostenidae).
Family
Tenebrionidae
Latreille, 1802
Tenebrionites Latreille, 1802: 165 [stem: Tenebrion-]. Type genus: Tenebrio Linnaeus, 1758. Comment: First Reviser (Tenebrionidae Latreille, 1802 vs Pimeliidae Latreille, 1802 vs Diaperidae Latreille, 1802) not determined, current usage maintained.
Subfamily
Zolodininae
Watt, 1975
Zolodininae Watt, 1975: 401 [stem: Zolodin-]. Type genus: Zolodinus Blanchard, 1853.
Subfamily
Lagriinae
Latreille, 1825 (1820)
Lagriariae Latreille, 1825: 381 [stem: Lagri-]. Type genus: Lagria Fabricius, 1775. Comment: use of younger family-group name conserved over Lachninae Billberg, 1820 (Art. 40.2) (see Bouchard et al. 2005); the older name Cossyphinae Latreille, 1802 is not given precedence over this name because the current placement of Cossyphini in this subfamily is uncertain (see Bouchard et al. 2005).
Tribe
Adeliini
Kirby, 1828
Adeliadae Kirby, 1828: 525 [stem: Adeli-]. Type genus: Adelium Kirby, 1819. Comment: family-group name attributed to Hope (1840a: 188) in recent literature; this is a senior homonym of Adeliini Viereck, 1918 (type genus Adelius Haliday, 1833) which is considered a valid tribe in the Hymenoptera: Braconidae: Cheloninae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
*Apatèlates Mulsant and Rey, 1859: 87 [stem: Apatel-]. Type genus: Apatelus Mulsant and Rey, 1859. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 208, as Apatelini), but not generally accepted as valid; if evidence is found in the future that would lead to the treatment of Mulsant and Rey’s name as available, this would threaten the younger name originally proposed as Apatelinae Grote, 1883 (type genus Apatele Hübner, 1822) in Lepidoptera which has been used as valid in recent literature.
Tribe
Belopini
Reitter, 1917
Calcariens Mulsant, 1854: 268 [stem: Calcar-]. Type genus: Calcar Dejean, 1821 [preoccupied genus name, not Calcar de Montfort, 1810 [Mollusca]; syn. of Centorus Mulsant, 1854]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Seidlitz (1895: 647, as Calcarina); permanently invalid (Art. 39): based on preoccupied type genus.
Belopinae Reitter, 1917: 59 [stem: Belop-]. Type genus: Belopus Gebien, 1911. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Belopod-).
Tribe
Chaerodini
Doyen, Matthews and Lawrence, 1990
Chaerodini Doyen et al., 1990: 239 [stem: Chaerod-]. Type genus: Chaerodes A. White, 1846.
Tribe
Cossyphini
Latreille, 1802
Cossyphores Latreille, 1802: 164 [stem: Cossyph-]. Type genus: Cossyphus A. G. Olivier, 1791. Comment: not given precedence over Lagriinae Latreille, 1825 because its current placement is uncertain (see Bouchard et al. 2005).
Tribe
Goniaderini
Lacordaire, 1859
Goniadérides Lacordaire, 1859: 390 [stem: Goniader-]. Type genus: Goniadera Perty, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1895: 614, as Goniaderina), generally accepted as in Gebien (1911: 467, as Goniaderinae).
*Phobéliides Lacordaire, 1859: 393 [stem: Phobeli-]. Type genus: Phobelius Blanchard, 1845. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 212, as Phobeliini), but not generally accepted as valid.
Eschatoporini Blaisdell, 1906: 78 [stem: Eschatopori-]. Type genus: Eschatoporis Blaisdell, 1906. Comment: incorrect original stem formation, not in prevailing usage.
Phobeliina Ardoin, 1961: 33 [stem: Phobeli-]. Type genus: Phobelius Blanchard, 1845.
*Anaedini Skopin, 1964: 7 [stem: Anaed-]. Type genus: Anaedus Blanchard, 1845. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Laenini
Seidlitz, 1895
Laenina Seidlitz, 1895: 669 [stem: Laen-]. Type genus: Laena Dejean, 1821.
Tribe
Lagriini
Latreille, 1825 (1820)
Lagriariae Latreille, 1825: 381 [stem: Lagri-]. Type genus: Lagria Fabricius, 1775. Comment: use of younger family-group name conserved over Lachnini Billberg, 1820 (Art. 40.2) (see Bouchard et al. 2005).
Subtribe
Lagriina
Latreille, 1825 (1820)
*Lagrien Oken, 1817: 1180 [stem: Lagri-]. Type genus: Lagria Fabricius, 1775. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Oken (1817).
Lachnaedes Billberg, 1820a: 34 [stem: Lachn-]. Type genus: Lachna Billberg, 1820 [syn. of Lagria Fabricius, 1775]. Comment: the family-group names Lachnidae/-inae/-ini Herrich-Schaeffer 1854 (based on Lachnus H. C. C. Burmeister 1835) are currently used as valid in Hemiptera, Lachninae (incorrectly attributed to Passerini (1863)) was placed on the Official List of Family-Group Names in Zoology (ICZN 1956); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Lagriariae Latreille, 1825: 381 [stem: Lagri-]. Type genus: Lagria Fabricius, 1775. Comment: use of younger family-group name conserved over Lachnina Billberg, 1820 (Art. 40.2) (see Bouchard et al. 2005).
Loubacantini Bonadona, 1959: 1034 [stem: Loubacant-]. Type genus: Loubacantus Bonadona, 1959 [syn. of Entypodera Gerstaecker, 1871]. Comment: originally proposed in Anthicidae, transfer to Lagriinae by Bonadona (1984).
Subtribe
Statirina
Blanchard, 1845
Statyrites Blanchard, 1845b: 39 [stem: Statir-]. Type genus: Statira Lepeletier and Audinet-Serville, 1828 [as Statyra, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 246, as Statyrini), generally accepted as in Bouchard et al. (2005: 501, as Statyrina); incorrect original stem formation, not in prevailing usage.
*Hystérarthrides Lacordaire, 1869: 231 [stem: Hysterarthr-]. Type genus: Hysterarthron J. Thomson, 1864. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 240, as Hysterarthrini), but not generally accepted as valid; type genus transferred from Cerambycidae by Ritsema (1892: 54).
Tribe
Lupropini
Ardoin, 1958
Lupropsini Ardoin, 1958: 59 [stem: Luprop-]. Type genus: Luprops Hope, 1833. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Pycnocerini
Lacordaire, 1859
nomen protectum
Chiroscelidae Hope, 1840a: 127 [stem: Chiroscelid-]. Type genus: Chiroscelis Lamarck, 1804. Comment: published before 30 September 1840; Bouchard et al. (2005: 524) treated this name and Chiroscelites Laporte (1840b [before 26 December]: 216) as nomina oblita; incorrect original stem formation, not in prevailing usage.
Pycnocérides Lacordaire, 1859: 399 [stem: Pycnocer-]. Type genus: Pycnocerus Westwood, 1841. Comment: nomen protectum (see Bouchard et al. 2005: 524); original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1887: 50, as Pycnoceridae), generally accepted as in Bouchard et al. (2005: 501, as Pycnocerini).
Prioscelina Skopin, 1964: 30, in key [stem: Prioscelid-]. Type genus: Prioscelis Hope, 1840. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Nilioninae
Oken, 1843
Nilioniden Oken, 1843: 484 [stem: Nilion-]. Type genus: Nilio Latreille, 1802 [incorrect subsequent spelling of Nilion by Latreille (1804b: 333), incorrect subsequent spelling in prevailing usage, treated as correct original spelling (Art. 33.3.1)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Tulk (1847: 614, as Niliondae [incorrect stem formation]); family-group name previously attributed to Lacordaire (1859: 518).
Subfamily
Phrenapatinae
Solier, 1834
Phrépatides Solier, 1834: 488 [stem: Phrenapat-]. Type genus: Phrenapates Gray, 1832.
Tribe
Archaeoglenini
Watt, 1975
Archaeoglenini Watt, 1975: 412 [stem: Archaeoglen-]. Type genus: Archaeoglenes Broun, 1893.
Tribe
Penetini
Lacordaire, 1859
Pénétides Lacordaire, 1859: 318 [stem: Penet-]. Type genus: Peneta Lacordaire, 1859. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1894: 545, as Penetina), generally accepted as in Bouchard et al. (2005: 501, as Penetini).
Phthorini Boddy, 1965: 144 [stem: Phtor-]. Type genus: Phtora Mulsant, 1854 [as Phthora, incorrect subsequent spelling of type genus name, not in prevailing usage; Phtora is a junior homonym of Phtora Germar, 1836 [Coleoptera: Tenebrionidae: Diaperinae]; syn. of Clamoris Gozis, 1886]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Phrenapatini
Solier, 1834
Phrépatides Solier, 1834: 488 [stem: Phrenapat-]. Type genus: Phrenapates Gray, 1832 [as Phrepates, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 287, as Phrenapatoidae), generally accepted as in Bouchard et al. (2005: 501, as Phrenapatinae/-ini); incorrect original stem formation, not in prevailing usage.
Subfamily
Pimeliinae
Latreille, 1802
Pimeliariae Latreille, 1802: 166 [stem: Pimeli-]. Type genus: Pimelia Fabricius, 1775.
Tribe
Adelostomini
Solier, 1834
Adélostomites Solier, 1834: 502 [stem: Adelostom-]. Type genus: Adelostoma Duponchel, 1827. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 8, as Adelostomoidae), generally accepted as in Bouchard et al. (2005: 501, as Adelostomini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Adelostomat-).
*Eurychorites Solier, 1837b: 153 [stem: Eurychor-]. Type genus: Eurychora Thunberg, 1791. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Solier (1837).
Eurychoridae Hope, 1840a: 121 [stem: Eurychor-]. Type genus: Eurychora Thunberg, 1791.
Tribe
Adesmiini
Lacordaire, 1859
nomen protectum
*Macropodites Solier, 1834: 501 [stem: Macropod-]. Type genus: Macropoda Solier, 1835. Comment: unavailable family-group name, not based on an available genus name.
*Macropodites Solier, 1835b: 509 [stem: Macropod-]. Type genus: Macropoda Solier, 1835. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally accepted as valid.
*Adesmiites Blanchard, 1845b: 4 [stem: Adesmi-]. Type genus: Adesmia Fischer von Waldheim, 1822. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845b).
Macropodoidae Agassiz, 1846b: 221 [stem: Macropod-]. Type genus: Macropoda Solier, 1835. Comment: nomen oblitum (Appendix 1); this name is a junior homonym of Macropodidae Gray, 1821 in Mammalia (type genus Macropus Shaw and Nodder, 1790) which is on the Official List of Family-Group Names in Zoology.
Megagenianos Solier, 1851: 124 [stem: Megageni-]. Type genus: Megagenius Solier, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1887: 50, as Megageniidae), generally accepted as in Bouchard et al. (2007: 386, as Megageniini); nomen oblitum, Adesmiini Lacordaire, 1859 conserved over this name by Bouchard et al. (2007: 386).
Adesmiides Lacordaire, 1859: 22 [stem: Adesmi-]. Type genus: Adesmia Fischer von Waldheim, 1822. Comment: nomen protectum (Appendix 1); original vernacular name available (Art. 11.7.2): first used in latinized form by Baudi di Selve (1875: 22, as Adesmidae [incorrect stem formation]), generally accepted as in Bouchard et al. (2005: 502); also Adesmiini Lacordaire, 1859 conserved over Megageniini Solier, 1851 by Bouchard et al. (2007: 386).
Épiphysides Lacordaire, 1859: 29 [stem: Epiphys-]. Type genus: Epiphysa Dejean, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 213, as Epiphysini), generally accepted as in Gebien (1910a: 82, as Epiphysinae).
Tribe
Akidini
Billberg, 1820
Acides Billberg, 1820a: 32 [stem: Akid-]. Type genus: Akis Herbst, 1799 [as Acis, unjustified emendation of type genus by Billberg (1820a), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Anepsiini
LeConte, 1862
Anepsiini J. L. LeConte, 1862: 215 [stem: Anepsi-]. Type genus: Anepsius J. L. LeConte, 1851.
Batuliini G. H. Horn, 1870: 270 [stem: Batuli-]. Type genus: Batulius J. L. LeConte, 1851.
Anchommini G. H. Horn, 1878: 558 [stem: Anchommat-]. Type genus: Anchomma J. L. LeConte, 1858 [genus originally proposed in Colydiidae]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Asidini
Fleming, 1821
Asidadae Fleming, 1821: 51 [stem: Asid-]. Type genus: Asida Latreille, 1802.
*Machlides Lacordaire, 1859: 155 [stem: Machl-]. Type genus: Machla Herbst, 1799 [earlier usage of the type genus name by Lichtenstein (1796) was suppressed for nomenclatural purposes (ICZN 1995e); this genus is currently known under the name Pseudomachla Wilke, 1921]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form and attributed to the original author but not used as valid then (Bouchard et al. 2005: 506).
Astroti G. H. Horn, 1870: 289 [stem: Astrot-]. Type genus: Astrotus J. L. LeConte, 1858.
Craniotini J. L. LeConte and G. H. Horn, 1883: 361 [stem: Craniot-]. Type genus: Craniotus J. L. LeConte, 1851.
Machlini Chatanay, 1914: 1 [stem: Machl-]. Type genus: Machla Herbst, 1799 [earlier usage of the type genus name by Lichtenstein (1796) was suppressed for nomenclatural purposes (ICZN 1995e); this genus is currently known under the name Pseudomachla Wilke, 1921].
Parecatini Chatanay, 1914: 2 [stem: Parecat-]. Type genus: Parecatus Fairmaire, 1900.
Tribe
Boromorphini
Skopin, 1978
Boromorphini Skopin, 1978: 228, in key [stem: Boromorph-]. Type genus: Boromorphus Wollaston, 1854.
Tribe
Branchini
LeConte, 1862
Branchini J. L. LeConte, 1862: 222 [stem: Branch-]. Type genus: Branchus J. L. LeConte, 1862.
Tribe
Caenocrypticini
Koch, 1958
Caenocrypticini Koch, 1958: 121 [stem: Caenocryptic-]. Type genus: Caenocrypticus Gebien, 1920.
Tribe
Ceratanisini
Gebien, 1937
Apolitina Seidlitz, 1895: 666 [stem: Apolit-]. Type genus: Apolites Jacquelin du Val, 1861 [preoccupied genus name, not Apolites Sundevall, 1835 [Aves]; syn. of Idastrandiella Strand, 1929]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Anisocerini Reitter, 1906: 477 [stem: Anisocer-]. Type genus: Anisocerus Faldermann, 1837 [preoccupied genus name, not Anisocerus Audinet-Serville, 1835 [Coleoptera: Cerambycidae]; syn. of Ceratanisus Gemminger, 1870]. Comment: junior homonym of Anisocerini J. Thomson, 1860 [Coleoptera: Cerambycidae]; permanently invalid (Art. 39): based on preoccupied type genus.
Ceratanisini Gebien, 1937: 791 [stem: Ceratanis-]. Type genus: Ceratanisus Gemminger, 1870.
Tribe
Cnemeplatiini
Jacquelin du Val, 1861
Cnéméplatiites Jacquelin du Val, 1861: 286 [stem: Cnemeplati-]. Type genus: Cnemeplatia A. Costa, 1847.
Subtribe
Actizetina
Watt, 1992
Actizetina Watt, 1992: 297 [stem: Actizet-]. Type genus: Actizeta Pascoe, 1875.
Subtribe
Cnemeplatiina
Jacquelin du Val, 1861
*Autocérides Lacordaire, 1859: 279 [stem: Autocer-]. Type genus: Autocera Wollaston, 1857 [syn. of Cnemeplatia A. Costa, 1847]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 208, as Autocerini), but not generally accepted as valid.
Cnéméplatiites Jacquelin du Val, 1861: 286 [stem: Cnemeplati-]. Type genus: Cnemeplatia A. Costa, 1847. Comment: Bouchard et al. (2005: 504) considered Jacquelin du Val’s name as unavailable based on the requirements of Art. 11.7.2, however, the recent usage of «Cnemeplatiini Jacquelin du Val, 1861” as a valid taxon by Löbl et al. (2008: 140) made this name available.
Cnemeplatiini Csiki, 1953: 117 [stem: Cnemeplati-]. Type genus: Cnemeplatia A. Costa, 1847. Comment: proposed as new without reference to Cnéméplatiites Jacquelin du Val, 1861.
Subtribe
Rondoniellina
Ferrer and Moragues, 2000
Rondoniellina Ferrer and Moragues, 2000: 100 [stem: Rondoniell-]. Type genus: Rondoniella Kaszab, 1970.
Subtribe
Thorictosomatina
Watt, 1992
Thorictosomatina Watt, 1992: 296 [stem: Thorictosomat-]. Type genus: Thorictosoma Lea, 1919.
Tribe
Cnemodinini
Gebien, 1910
Cnemodini G. H. Horn, 1870: 266 [stem: Cnemodont-]. Type genus: Cnemodus G. H. Horn, 1870 [preoccupied genus name, not Cnemodus Herrich-Schaeffer, 1850 [Hemiptera]; syn. of Cnemodinus Cockerell, 1906]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Cnemodininae Gebien, 1910a: 4 [stem: Cnemodin-]. Type genus: Cnemodinus Cockerell, 1906.
Tribe
Coniontini
Waterhouse, 1858
Coniontidae G. R. Waterhouse, 1858: 59 [stem: Coniont-]. Type genus: Coniontis Eschscholtz, 1829. Comment: family-group name previously attributed to Lacordaire (1859)/Schaum (1859) (see Bouchard et al. 2005: 502).
Coelini Casey, 1907: 500 [stem: Coel-]. Type genus: Coelus Eschscholtz, 1829.
Eusatti Doyen, 1984: 11 [stem: Eusatt-]. Type genus: Eusattus J. L. LeConte, 1851.
Tribe
Cossyphodini
Wasmann, 1899
Cossyphodidae Wasmann, 1899: 161 [stem: Cossyphod-]. Type genus: Cossyphodes Westwood, 1851. Comment: downgraded from a subfamily of Tenebrionidae to a tribe of Pimeliinae by Matthews et al. (2010).
Subtribe
Cossyphodina
Wasmann, 1899
Cossyphodidae Wasmann, 1899: 161 [stem: Cossyphod-]. Type genus: Cossyphodes Westwood, 1851.
Subtribe
Cossyphoditina
Basilewsky, 1950
Cossyphoditinae Basilewsky, 1950a: 187 [stem: Cossyphodit-]. Type genus: Cossyphodites Brauns, 1901.
Subtribe
Esemephina
Steiner, 1980
Esemephini Steiner, 1980: 391 [stem: Esemeph-]. Type genus: Esemephe Steiner, 1980.
Subtribe
Paramellonina
Andreae, 1961
Paramelloninae Andreae, 1961: 200 [stem: Paramellon-]. Type genus: Paramellon C. O. Waterhouse, 1882. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Paramellont-).
Tribe
Cryptochilini
Solier, 1841
Cryptochilites Solier, 1841: 248 [stem: Cryptochil-]. Type genus: Cryptochile Latreille, 1829.
Subtribe
Calognathina
Lacordaire, 1859
Calognathides Lacordaire, 1859: 85 [stem: Calognath-]. Type genus: Calognathus Guérin-Méneville, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1887: 50, as Calognathidae), generally accepted as in Gebien (1910a: 117, as Calognathinae).
Subtribe
Cryptochilina
Solier, 1841
Cryptochilites Solier, 1841: 248 [stem: Cryptochil-]. Type genus: Cryptochile Latreille, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1887: 50, as Cryptochilidae), generally accepted as in Bouchard et al. (2005: 502, as Cryptochilini).
Subtribe
Homebiina
Endrödy-Younga, 1989
Homebiina Endrödy-Younga, 1989: 124 [stem: Homebi-]. Type genus: Homebius Endrödy-Younga, 1989.
Subtribe
Horatomina
Koch, 1955
Horatomina Koch, 1955: 14 [stem: Horatom-]. Type genus: Horatoma Solier, 1840.
Subtribe
Vansoniina
Koch, 1955
Vansonini Koch, 1955: 12 [stem: Vansoni-]. Type genus: Vansonium Koch, 1950. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Cryptoglossini
LeConte, 1862
nomen protectum
Centrioptérides Lacordaire, 1859: 134 [stem: Centriopter-]. Type genus: Centrioptera Mannerheim, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 220, as Centriopterae), generally accepted as in Bouchard et al. (2005: 502, as Centriopterini); nomen oblitum (see Aalbu 2006: 57).
Cryptoglossini J. L. LeConte, 1862: 220 [stem: Cryptogloss-]. Type genus: Cryptoglossa Solier, 1837. Comment: nomen protectum (see Aalbu 2006: 57).
Tribe
Edrotini
Lacordaire, 1859
Édrotides Lacordaire, 1859: 31 [stem: Edrot-]. Type genus: Edrotes J. L. LeConte, 1851. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Casey (1907: 279, as Edrotini), generally accepted as in Bouchard et al. (2005: 502, as Edrotini).
Triorophi J. L. LeConte and G. H. Horn, 1883: 362 [stem: Trioroph-]. Type genus: Triorophus J. L. LeConte, 1851.
Auchmobii J. L. LeConte and G. H. Horn, 1883: 362 [stem: Auchmobi-]. Type genus: Auchmobius J. L. LeConte, 1851.
Trimytini Casey, 1907: 278 [stem: Trimytid-]. Type genus: Trimytis Leconte, 1851. Comment: incorrect original stem formation, not in prevailing usage.
Eurymetoponini Casey, 1907: 278 [stem: Eurymetop-]. Type genus: Eurymetopon Eschscholtz, 1831. Comment: incorrect original stem formation, not in prevailing usage.
Trientomini Casey, 1907: 278 [stem: Trientom-]. Type genus: Trientoma Solier, 1835.
Tribe
Elenophorini
Solier, 1837
Elénophorites Solier, 1837a: 638 [stem: Elenophor-]. Type genus: Elenophorus Dejean, 1821 [syn. of Leptoderis Billberg, 1820]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Schaum (1859: 66, as Elenophoridae), generally accepted as in Bouchard et al. (2005: 502, as Elenophorini).
Tribe
Epitragini
Blanchard, 1845
nomen protectum
Lygophilia Rafinesque, 1815: 113 [stem: Lygophil-]. Type genus: Lygophilus Rafinesque, 1815 [syn. of Epitragus Latreille, 1802]. Comment: nomen oblitum (see Bouchard et al. 2007: 386).
*Épitragites Solier, 1834: 490 [stem: Epitrag-]. Type genus: Epitragus Latreille, 1802. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Solier (1834).
Épitragites Blanchard, 1845b: 16 [stem: Epitrag-]. Type genus: Epitragus Latreille, 1802. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Erichson (1847a: 117, as Epitragii), generally accepted as in Bouchard et al. (2005: 502, as Epitragini); nomen protectum (see Bouchard et al. 2007: 386).
Tribe
Erodiini
Billberg, 1820
nomen protectum
Cephaceria Rafinesque, 1815: 113 [stem: Cephacer-]. Type genus: Cephacerus Rafinesque, 1815 [syn. of Erodius Fabricius, 1775]. Comment: nomen oblitum (see Bouchard et al. 2007: 386).
Erodiides Billberg, 1820a: 32 [stem: Erodi-]. Type genus: Erodius Fabricius, 1775. Comment: nomen protectum (see Bouchard et al. 2007: 386); this family-group name was also used in the same year by Billberg (1820b: 392, as Erodiides).
*Estenogenianos Solier, 1851: 138 [stem: Stenogeni-]. Type genus: Stenogenius Solier, 1834 [unavailable genus name, proposed in synonymy and not subsequently made available]. Comment: family-group name unavailable (Art. 11.7.1.1): not based on an available genus name at the time; incorrect original stem formation, not in prevailing usage.
*Arthrodeiden Koch, 1943: 483 [stem: Arthrode-]. Type genus: Arthrodeis Solier, 1834. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Tribe
Evaniosomini
Lacordaire, 1859
Évaniosomides Lacordaire, 1859: 73 [stem: Evaniosom-]. Type genus: Evaniosomus Guérin-Méneville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Gebien (1910a: 22, as Evaniosominae).
Tribe
Falsomycterini
Gebien, 1910
Falsomycterinae Gebien, 1910b: 177 [stem: Falsomycter-]. Type genus: Falsomycterus Pic, 1907.
Tribe
Idisiini
Medvedev, 1973
Idisiini G. S. Medvedev, 1973: 644 [stem: Idisi-]. Type genus: Idisia Pascoe, 1866.
Tribe
Klewariini
Gebien, 1910
Klewariinae Gebien, 1910a: 36 [stem: Klewari-]. Type genus: Klewaria Reitter, 1910.
Tribe
Kuhitangiini
Medvedev, 1962
Kuhitangiinae G. S. Medvedev, 1962: 1184 [stem: Kuhitangi-]. Type genus: Kuhitangia G. S. Medvedev, 1962.
Tribe
Lachnogyini
Seidlitz, 1894
Lachnogyini Seidlitz, 1894: 490 [stem: Lachnogy-]. Type genus: Lachnogya Ménétriés, 1849. Comment: subtribal classification according to G. S. Medvedev (2006).
Subtribe
Lachnodactylina
Reitter, 1904
Lachnodactylina Reitter, 1904: 182 [stem: Lachnodactyl-]. Type genus: Lachnodactylus Seidlitz, 1898.
Subtribe
Lachnogyina
Seidlitz, 1894
Lachnogyini Seidlitz, 1894: 490 [stem: Lachnogy-]. Type genus: Lachnogya Ménétriés, 1849.
Subtribe
Netuschiliina
Ferrer and Yvinec, 2004
Netuschiliina Ferrer and Yvinec, 2004: 48 [stem: Netuschili-]. Type genus: Netuschilia Reitter, 1904. Comment: also incorrectly spelled as Netuschilina in the original publication on page 48 (First Revisers are Bouchard et al. 2005: 504).
Tribe
Leptodini
Lacordaire, 1859
Leptodides Lacordaire, 1859: 108 [stem: Leptod-]. Type genus: Leptodes Dejean, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Baudi di Selve (1875: 74, as Leptodidae), generally accepted as in Gebien (1910a: 91, as Leptodinae).
Tribe
Nycteliini
Solier, 1834
Nyctélites Solier, 1834: 502 [stem: Nycteli-]. Type genus: Nyctelia Latreille, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hope (1840a: 127, as Nyctelidae [incorrect stem formation]), generally accepted as in Bouchard et al. (2005: 502, as Nycteliini); incorrect original stem formation, not in prevailing usage.
Tribe
Nyctoporini
Lacordaire, 1859
Nyctoporides Lacordaire, 1859: 130 [stem: Nyctopor-]. Type genus: Nyctoporis Eschscholtz, 1831. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 219, as Nyctoporini), generally accepted as in Bouchard et al. (2005: 502, as Nyctoporini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Nyctoporid-).
Tribe
Phrynocarenini
Gebien, 1928
Phrynocareninae Gebien, 1928: 105 [stem: Phrynocaren-]. Type genus: Phrynocarenum Gebien, 1928.
Tribe
Physogasterini
Lacordaire, 1859
Physogastérides Lacordaire, 1859: 206 [stem: Physogaster-]. Type genus: Physogaster Lacordaire, 1830 [this genus name has been credited to Guérin-Méneville (1834: 2) in the literature, however the description of Physogaster mendocinus by Lacordaire (1830a: 276) makes the genus name available from that author and year]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. C. C. Burmeister (1875: 488, as Physogasteridae), generally accepted as in Bouchard et al. (2005: 502, as Physogasterini).
Tribe
Pimeliini
Latreille, 1802
Pimeliariae Latreille, 1802: 166 [stem: Pimeli-]. Type genus: Pimelia Fabricius, 1775.
Pimidinia Rafinesque, 1815: 113 [stem: Pimidi-]. Type genus: Pimidia Rafinesque, 1815 [syn. of Pimelia Fabricius, 1775]. Comment: incorrect original stem formation, not in prevailing usage.
*Platyopes Motschulsky, 1849: 58 [stem: Platyop-]. Type genus: Platyope Fischer von Waldheim, 1820. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Platyopidae Semenov, 1893a: 260 [stem: Platyop-]. Type genus: Platyope Fischer von Waldheim, 1820. Comment: family-group name proposed as new without reference to Platyopes Motschulsky, 1849; Platyopidae Huene, 1931 has been used in amphibians (type genus Platyops Twelvetrees, 1880) but this name is permanently invalid since it is based on a preoccupied genus name.
Leucolaephusini Pierre, 1961: 558 [stem: Leucolaeph-]. Type genus: Leucolaephus Lucas, 1859. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Praociini
Eschscholtz, 1829
Praocidae Eschscholtz, 1829b: 5 [stem: Praoci-]. Type genus: Praocis Eschscholtz, 1829. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Sepidiini
Eschscholtz, 1829
Sepidiae Eschscholtz, 1829b: 4 [stem: Sepidi-]. Type genus: Sepidium Fabricius, 1775.
Subtribe
Hypomelina
Koch, 1955
Hypomelina Koch, 1955: 36 [stem: Hypomel-]. Type genus: Hypomelus Solier, 1843.
Subtribe
Molurina
Solier, 1834
Molurites Solier, 1834: 505 [stem: Molur-]. Type genus: Moluris Latreille, 1802. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Schaum (1859: 68, as Moluridae), generally accepted as in Bouchard et al. (2005: 502, as Molurina); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Molurid-).
*Psammodoiden Koch, 1953a: 138 [stem: Psammod-]. Type genus: Psammodes Kirby, 1819. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899; Psammodidae has been used in the Scarabaeidae literature but this is based on an incorrect stem formation (type genus Psammodius Fallén, 1807, stem Psammodi-).
*Phrynocoloiden Koch, 1953a: 138 [stem: Phrynocol-]. Type genus: Phrynocolus Lacordaire, 1859. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Subtribe
Oxurina
Koch, 1955
Oxurina Koch, 1955: 34 [stem: Oxur-]. Type genus: Oxura Kirby, 1819.
Subtribe
Phanerotomeina
Koch, 1958
*Phanerotomoide Koch, 1953a: 138 [stem: Phanerotom-]. Type genus: Phanerotoma Solier, 1843 [preoccupied genus name, not Phanerotoma Wesmael, 1838 [Hymenoptera]; syn. of Phanerotomea Koch, 1957]. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun.
Phanerotomina Koch, 1955: 37 [stem: Phanerotom-]. Type genus: Phanerotoma Solier, 1843 [preoccupied genus name, not Phanerotoma Wesmael, 1838 [Hymenoptera]; syn. of Phanerotomea Koch, 1957]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; Phanerotomini Baker, 1925 (type genus Phanerotoma Wesmael, 1838) is available in Hymenoptera: Braconidae.
Phanerotomeina Koch, 1958: 58 [stem: Phanerotome-]. Type genus: Phanerotomea Koch, 1958. Comment: replacement name for Phanerotomina Koch, 1955 because of the homonymy of the type genus.
Subtribe
Sepidiina
Eschscholtz, 1829
Sepidiae Eschscholtz, 1829b: 4 [stem: Sepidi-]. Type genus: Sepidium Fabricius, 1775.
Subtribe
Trachynotina
Koch, 1955
*Trachynotides Brullé, 1832: 189 [stem: Trachynot-]. Type genus: Trachynotus Latreille, 1828. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Brullé (1832).
Trachynotina Koch, 1955: 34, in key [stem: Trachynot-]. Type genus: Trachynotus Latreille, 1828. Comment: Trachynotidae Schrammen, 1924 in Porifera is permanently invalid because it was based on the junior homonym Trachynotus Schrammen, 1924; the fish name Trachynotinae Gill, 1861 (type genus Trachynotus Agassiz, 1846, an unjustified emendation of Trachinotus Lacepède, 1801 and also a junior homonym of Trachynotus Latreille, 1828) was later corrected to Trachinotinae, which is used as valid today; Trachynotoidae Förster, 1869 has also been used in ichneumonid literature but this name is permanently invalid since it was based on the junior homonym Trachynotus Gravenhorst, 1829.
Tribe
Stenosini
Schaum, 1859 (1834)
Tagénites Solier, 1834: 503 [stem: Tageni-]. Type genus: Tagenia Latreille, 1802 [syn. of Stenosis Herbst, 1799]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hope (1840a: 127, as Tagenidae), generally accepted as in G. R. Waterhouse (1845: 30, as Tageniidae [incorrect stem formation]; use of younger name Stenosini conserved (Art. 40.2) (see Bouchard et al. 2005: 523); incorrect original stem formation, not in prevailing usage.
Stenosidae Schaum, 1859: 66 [stem: Stenos-]. Type genus: Stenosis Herbst, 1799. Comment: published before end of February 1859; this family-group name was also used in the same year by Lacordaire (1859 [before 27 June]: 101, as Sténosides); use of family-group name conserved over Tagenini Solier, 1834 (Art. 40.2) (see Bouchard et al. 2005: 523); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Stenose-).
Platamodina Reitter, 1900: 82 [stem: Platamod-]. Type genus: Platamodes Ménétriés, 1849.
Typhlusechini Casey, 1907: 281 [stem: Typhlusech-]. Type genus: Typhlusechus Linell, 1897.
Araeoschizini Casey, 1907: 484 [stem: Araeoschiz-]. Type genus: Araeoschizus J. L. LeConte, 1851.
Dichillina Reitter, 1916: 137, in key [stem: Dichill-]. Type genus: Dichillus Jacquelin du Val, 1860.
Harvengina Ferrer, 2004: 370 [stem: Harvengi-]. Type genus: Harvengia Ferrer, 2004. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Tentyriini
Eschscholtz, 1831
Tentyridae Eschscholtz, 1831: 4 [stem: Tentyri-]. Type genus: Tentyria Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 2010c)]. Comment: incorrect original stem formation, not in prevailing usage.
Gnathosiides Lacordaire, 1859: 33 [stem: Gnathosi-]. Type genus: Gnathosia Fischer von Waldheim, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 213, as Gnathosiini), generally accepted as in Skopin (1979: 170, as Gnathosiina).
*Hypéropides Lacordaire, 1859: 60 [stem: Hyperop-]. Type genus: Hyperops Eschscholtz, 1831. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Capnisini Casey, 1907: 279 [stem: Capnis-]. Type genus: Capnisa Dejean, 1836 [syn. of Gnathosia Fisher von Waldheim, 1821].
Pachycerina Skopin, 1979: 170, in key [stem: Pachycer-]. Type genus: Pachycera Eschscholtz, 1831 [preoccupied genus name, not Pachycera Billberg, 1820 [Hemiptera]; syn. of Oedenocera Reiche, 1862]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Himatismina Skopin, 1979: 170, in key [stem: Himatism-]. Type genus: Himatismus Erichson, 1843.
Tribe
Thinobatini
Lacordaire, 1859
Thinobatides Lacordaire, 1859: 63 [stem: Thinobat-]. Type genus: Thinobatis Eschscholtz, 1831. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 214, as Thinobatini), generally accepted as in Bouchard et al. (2005: 502, as Thinobatini); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Thinobatid-).
Tribe
Trilobocarini
Lacordaire, 1859
Tribolocarides Lacordaire, 1859: 69 [stem: Trilobocar-]. Type genus: Trilobocara Solier, 1851 [as Tribolocara, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Kolbe (1887: 51, as Tribolocarini); incorrect stem formation, not in prevailing usage.
Salaxini Casey, 1907: 282 [stem: Salac-]. Type genus: Salax Guérin-Méneville, 1834. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Vacronini
Gebien, 1910
Vacroninae Gebien, 1910a: 118 [stem: Vacron-]. Type genus: Vacronus Casey, 1907 [syn. of Alaephus G. H. Horn, 1870].
*Eupsophulites Kwieton, 1982: 96 [stem: Eupsophul-]. Type genus: Eupsophulus Cockerell, 1906. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Zophosini
Solier, 1834
Zophosites Solier, 1834: 597 [stem: Zophos-]. Type genus: Zophosis Latreille, 1802. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Stein (1868: 78, as Zophosini), generally accepted as in Bouchard et al. (2005: 501, as Zophosini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Zophose-).
*Onychosites A. Deyrolle, 1867: 79 [stem: Onychose-]. Type genus: Onychosis Deyrolle, 1867. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
*Cardiosites A. Deyrolle, 1867: 79 [stem: Cardiose-]. Type genus: Cardiosis Deyrolle, 1867. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Dactylocalcarini Gebien, 1938a: 46 [stem: Dactylocalcar-]. Type genus: Dactylocalcar Gebien, 1938.
Subfamily
Tenebrioninae
Latreille, 1802
Tenebrionites Latreille, 1802: 165 [stem: Tenebrion-]. Type genus: Tenebrio Linnaeus, 1758. Comment: First Reviser (Tenebrioninae Latreille, 1802 vs Helopinae Latreille, 1802) not determined, current usage maintained
Tribe
Acropteronini
Doyen, 1989
Acropteronini Doyen, 1989: 288 [stem: Acropteron-]. Type genus: Acropteron Perty, 1832. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Acropter-).
Tribe
Alphitobiini
Reitter, 1917
Alphitobiini Reitter, 1917: 58 [stem: Alphitobi-]. Type genus: Alphitobius Stephens, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1975)].
Tribe
Amarygmini
Gistel, 1848
Amarygmiidae Gistel, 1848: [10] [stem: Amarygm-]. Type genus: Amarygmus Dalman, 1823. Comment: incorrect original stem formation, not in prevailing usage.
Mégacanthides Lacordaire, 1859: 467 [stem: Megacanth-]. Type genus: Megacantha Westwood, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Quedenfeldt (1885: 20, as Megacanthidae).
Méracanthides Lacordaire, 1859: 464 [stem: Meracanth-]. Type genus: Meracantha Kirby, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 240, as Meracanthini), generally accepted as in Gebien (1911: 567, as Meracanthinae).
Megacanthina Ardoin, 1962: 960 [stem: Megacanth-]. Type genus: Megacantha Westwood, 1843. Comment: family-group name proposed as new without reference to Mégacanthides Lacordaire, 1859.
Tribe
Amphidorini
LeConte, 1862
*Nycterinoides Solier, 1851: 210 [stem: Nycterin-]. Type genus: Nycterinus Eschscholtz, 1829. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Solier (1851).
*Embaphionides Lacordaire, 1859: 151 [stem: Embaphi-]. Type genus: Embaphion Say, 1824. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., D. B. Thomas (2005: 549, as Embaphionini), but not generally attributed to Lacordaire (1859); Embaphionini, as used by D. B. Thomas (2005) is unavailable because it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Amphidorae J. L. LeConte, 1862: 239 [stem: Amphidor-]. Type genus: Amphidora Eschscholtz, 1829.
Eleodiini Blaisdell, 1909: 27 [stem: Eleod-]. Type genus: Eleodes Eschscholtz, 1829. Comment: incorrect original stem formation, not in prevailing usage.
Eleodopsinae Blaisdell, 1939: 51 [stem: Eleodopse-]. Type genus: Eleodopsis Blaisdell, 1939 [syn. of Eleodes Eschscholtz, 1829]. Comment: incorrect original stem formation, not in prevailing usage.
Lariversiina La Rivers, 1948: 98 [stem: Lariversi-]. Type genus: Lariversius Blaisdell, 1947.
Trogloderina La Rivers, 1948: 98 [stem: Trogloder-]. Type genus: Trogloderus J. L. LeConte, 1879.
*Nycterini Doyen et al., 1990: 244 [stem: Nycterin-]. Type genus: Nycterinus Eschscholtz, 1829. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Tribe
Apocryphini
Lacordaire, 1859
Apocryphides Lacordaire, 1859: 432 [stem: Apocryph-]. Type genus: Apocrypha Eschscholtz, 1831. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 217, as Apocryphini), generally accepted as in Gebien (1911: 503, as Apocryphinae).
Diplocyrthini Escalera, 1914b: 355 [stem: Diplocyrt-]. Type genus: Diplocyrtus Quendenfeldt, 1887 [as Diplocyrthus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Blaptini
Leach, 1815
Blapsida Leach, 1815: 101 [stem: Blapt-]. Type genus: Blaps Fabricius, 1775.
Subtribe
Blaptina
Leach, 1815
Blapsida Leach, 1815: 101 [stem: Blapt-]. Type genus: Blaps Fabricius, 1775. Comment: although the original stem formation (Blaps-) was correct, the stem Blapt- has been used since it was first used by C. G. Thomson (1859: 114, as Blaptidae) and is also used here; an application to the Commission is needed to conserve the current spelling.
Subtribe
Gnaptorina
Medvedev, 2001
Gnaptorina G. S. Medvedev, 2001: 29 [stem: Gnaptor-]. Type genus: Gnaptor Brullé, 1832.
Subtribe
Gnaptorinina
Medvedev, 2001
Gnaptorinina G. S. Medvedev, 2001: 31 [stem: Gnaptorin-]. Type genus: Gnaptorina Reitter, 1887.
Subtribe
Prosodina
Skopin, 1960
Prosodina Skopin, 1960: 48 [stem: Prosod-]. Type genus: Prosodes Eschscholtz, 1829.
Subtribe
Remipedellina
Semenov, 1907
Remipedellini Semenov, 1907a: 259 [stem: Remipedell-]. Type genus: Remipedella Semenov, 1907. Comment: this family-group name was also used in the same year by Semenov (1907b: 176, as Remipedellini); priority for the two publications could not be established although Remipedellini Semenov, 1907b could not be considered available since the type genus Remipedella was made available only in Semenov (1907a).
Tribe
Bolitophagini
Kirby, 1837
nomen protectum
Eledonaedes Billberg, 1820b: 392 [stem: Eledon-]. Type genus: Eledona Latreille, 1797. Comment: nomen oblitum (see Appendix 1); the name Eledoninae (type genus Eledone Leach, 1817) is used as valid in cephalopods (earliest usage in cephalopods found is Eledonidae Rochebrune, 1884).
Bolitophagidae Kirby, 1837: 236 [stem: Bolitophag-]. Type genus: Bolitophagus Illiger, 1798. Comment: nomen protectum (see Appendix 1).
Rhipidandri J. L. LeConte, 1862: 236 [stem: Rhipidandr-]. Type genus: Rhipidandrus J. L. LeConte, 1862.
Eutomides Lacordaire, 1865: 369 [stem: Eutom-]. Type genus: Eutomus Lacordaire, 1865 [syn. of Rhipidandrus J. L. LeConte, 1862]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Ferrari (1867: 3, as Eutomides [treated as Latin]); name treated as unavailable by Bouchard et al. (2005: 508) but recently considered available by Alonso-Zarazaga and Lyal (2009: 7); taxon originally described in Curculionidae: Scolytinae.
Tribe
Centronopini
Doyen, 1989
Centronopini Doyen, 1989: 284 [stem: Centronop-]. Type genus: Centronopus Solier, 1848. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Centronopod-).
Tribe
Cerenopini
Horn, 1870
Cerenopi G. H. Horn, 1870: 325 [stem: Cerenop-]. Type genus: Cerenopus J. L. LeConte, 1851. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Cerenopod-).
Tribe
Dissonomini
Medvedev, 1968
*Hétérophilates Mulsant and Rey, 1859: 6 [stem: Heterophyl-]. Type genus: Heterophylus Mulsant and Rey, 1859 [preoccupied genus name, not Heterophylus Klug, 1833 [Coleoptera: Tenebrionidae: Diaperinae]; syn. of Dissonomus Jacquelin du Val, 1861]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if found to be available then permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
*Dissonomites Jacquelin du Val, 1861: 280 [stem: Dissonom-]. Type genus: Dissonomus Jacquelin du Val, 1861. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Jacquelin du Val (1861).
Dissonomini G. S. Medvedev, 1968: 211 [stem: Dissonom-]. Type genus: Dissonomus Jacquelin du Val, 1861.
Tribe
Eulabini
Horn, 1870
Eulabes G. H. Horn, 1870: 323 [stem: Eulab-]. Type genus: Eulabis Eschscholtz, 1829.
Tribe
Falsocossyphini
Ferrer, 2006
Falsocossyphini Ferrer, 2006: 77 [stem: Falsocossyph-]. Type genus: Falsocossyphus Pic, 1916.
Tribe
Heleini
Fleming, 1821
Heleadae Fleming, 1821: 51 [stem: Hele-]. Type genus: Helea Latreille, 1816.
Subtribe
Asphalina
Matthews and Lawrence, 2005
Asphalina E. G. Matthews and Lawrence, 2005: 544 [stem: Asphal-]. Type genus: Asphalus Pascoe, 1868.
Subtribe
Cyphaleina
Lacordaire, 1859
Cyphaléides Lacordaire, 1859: 407 [stem: Cyphale-]. Type genus: Cyphaleus Westwood, 1841. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1866b: 470, as Cyphaleinae), generally accepted as in Bouchard et al. (2005: 502, as Cyphaleina); precedence (Cyphaleini Lacordaire, 1859 vs Nyctozoilini Lacordaire, 1859) given to taxon originally proposed at the higher rank (Art. 24.1).
Nyctozoïlides Lacordaire, 1859: 349 [stem: Nyctozoil-]. Type genus: Nyctozoilus Guérin-Méneville, 1830. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Bates (1872b: 98, as Nyctozoilides [treated as Latin]), generally accepted as in Carter (1911: 138, as Nyctozoilides [treated as Latin]).
Subtribe
Heleina
Fleming, 1821
Heleadae Fleming, 1821: 51 [stem: Hele-]. Type genus: Helea Latreille, 1804. Comment: incorrect original stem formation, not in prevailing usage.
Briseinae Carter, 1924: 33 [stem: Bris-]. Type genus: Brises Pascoe, 1869. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Helopini
Latreille, 1802
Helopii Latreille, 1802: 176 [stem: Helop-]. Type genus: Helops Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 2009d)].
Subtribe
Helopina
Latreille, 1802
Helopii Latreille, 1802: 176 [stem: Helop-]. Type genus: Helops Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 2009d)].
Hypulia Rafinesque, 1815: 114 [stem: Hypul-]. Type genus: Hypulus Rafinesque, 1815 [syn. of Helops Fabricius, 1775].
*Enoplopites Solier, 1848: 155 [stem: Enoplopod-]. Type genus: Enoplopus Solier, 1848. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Solier (1848); incorrect original stem formation, not in prevailing usage.
*Hédyphanes Motschulsky, 1849: 57 [stem: Hedyphan-]. Type genus: Hedyphanes Fischer von Waldheim, 1820. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Enoplopini Reitter, 1917: 62 [stem: Enoplopod-]. Type genus: Enoplopus Solier, 1848. Comment: incorrect original stem formation, not in prevailing usage.
Nephodini Reitter, 1917: 63 [stem: Nephod-]. Type genus: Nephodes Blanchard, 1845.
Hedyphanina Reitter, 1922b: 6 [stem: Hedyphan-]. Type genus: Hedyphanes Fischer von Waldheim, 1820.
*Stenotrichini Blaisdell, 1939: 50 [stem: Stenotrich-]. Type genus: Stenotrichus J. L. LeConte, 1862 [syn. of Helops Fabricius, 1775]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Cylindrinotina
Español, 1956
*Xanthomini Antoine, 1949: 162 [stem: Xanthom-]. Type genus: Xanthomus Mulsant, 1854. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Ectromopsini Antoine, 1949: 162 [stem: Ectromopse-]. Type genus: Ectromopsis Antoine, 1949. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Cylindronotini Español, 1956: 84 [stem: Cylindrinot-]. Type genus: Cylindrinotus Faldermann, 1837 [as Cylindronotus, unjustified emendation of type genus name by Agassiz (1846b: 111), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Bouchard et al. (2005: 509).
Tribe
Helopinini
Lacordaire, 1859
Hélopinides Lacordaire, 1859: 457 [stem: Helopin-]. Type genus: Helopinus Solier, 1848.
Subtribe
Aptilina
Koch, 1958
Aptilina Koch, 1958: 139 [stem: Aptil-]. Type genus: Aptila Fåhraeus, 1870.
Subtribe
Helopinina
Lacordaire, 1859
Hélopinides Lacordaire, 1859: 457 [stem: Helopin-]. Type genus: Helopinus Solier, 1848. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Bertkau (1875: 356, as Helopinini), generally accepted as in Gebien (1911: 563, as Helopininae).
Drosochrini Koch, 1958: 133 [stem: Drosochr-]. Type genus: Drosochrus Erichson, 1843. Comment: unnecessary replacement name for Helopinini Lacordaire, 1859.
Subtribe
Micrantereina
Reitter, 1917
Micrantereini Reitter, 1917: 60 [stem: Micrantere-]. Type genus: Micrantereus Solier, 1848.
Subtribe
Oncosomina
Koch, 1958
Oncosomina Koch, 1958: 134 [stem: Oncosom-]. Type genus: Oncosoma Westwood, 1843 [unjustified emendation of Ogcosoma by Agassiz (1846b: 257), in prevailing usage, treated as justified emendation (Art. 33.2.3.1)]. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Oncosomat-).
Tribe
Melanimonini
Seidlitz, 1894 (1854)
Microzoumates Mulsant, 1854: 176 [stem: Microzo-]. Type genus: Microzoum Dejean, 1834 [syn. of Melanimon Steven 1829]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1889 [Gatt.]: 128, as Microzoina), generally accepted as in Bouchard et al. (2005: 523, as Microzoini); use of Melanimonini Seidlitz, 1894 conserved (Art. 40.2) (see Bouchard et al. 2005: 523); incorrect original stem formation, not in prevailing usage.
Melanimonina Seidlitz, 1894: 449 [stem: Melanimon-]. Type genus: Melanimon Steven, 1829. Comment: use of younger family-group name conserved (Art. 40.2) (see Bouchard et al. 2005).
Tribe
Opatrini
Brullé, 1832
Opatrites Brullé, 1832: 213 [stem: Opatr-]. Type genus: Opatrum Fabricius, 1775.
Subtribe
Heterocheirina
Koch, 1956
Heterocheirini Koch, 1956: 43 [stem: Heterocheir-]. Type genus: Heterocheira Lacordaire, 1859.
Subtribe
Heterotarsina
Blanchard, 1845
Hétérotarsites Blanchard, 1845b: 14 [stem: Heterotars-]. Type genus: Heterotarsus Latreille, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 231, as Heterotarsini), generally accepted as in Bouchard et al. (2005: 502, as Heterotarsina).
Subtribe
Opatrina
Brullé, 1832
Opatrites Brullé, 1832: 213 [stem: Opatr-]. Type genus: Opatrum Fabricius, 1775. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Shuckard (1839b: 49, as Opatridae), generally accepted as in Bouchard et al. (2005: 502, as Opatrini).
*Gonocéphalites Solier, 1834: 498 [stem: Gonocephal-]. Type genus: Gonocephalum Solier, 1834. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Solier (1834).
*Blapstinoides Solier, 1851: 231 [stem: Blapstin-]. Type genus: Blapstinus Dejean, 1821. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Solier (1851).
Blapstinites Mulsant and Rey, 1853: 258 [stem: Blapstin-]. Type genus: Blapstinus Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte (1862: 227, as Blapstini).
Stizopides Lacordaire, 1859: 258 [stem: Stizopod-]. Type genus: Stizopus Erichson, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 207, as Stizopini), generally accepted as in Gebien (1938b: 72 [393], as Stizopini); incorrect original stem formation, not in prevailing usage.
Sclérides Lacordaire, 1859: 263 [stem: Scler-]. Type genus: Scleron Hope, 1840 [syn. of Sclerum Dejean, 1834]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1890 [Gatt.]: 129, as Sclerina), generally accepted as in Seidlitz (1894: 415, as Sclerina).
*Penthicaires Mulsant and Rey, 1859: 5 [stem: Penthic-]. Type genus: Penthicus Faldermann, 1836. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 208, as Penthicini, under the valid tribe name Opatrini), but not generally accepted as valid.
*Caediaires Mulsant and Rey, 1859: 124 [stem: Caedi-]. Type genus: Caedius Blanchard, 1845. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 208, as Caediini, under the valid tribe name Opatrini), but not generally accepted as valid.
*Blacodaires Mulsant and Rey, 1859: 93 [stem: Blacod-]. Type genus: Blacodes Blanchard, 1845 [syn. of Blenosia Laporte, 1840]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 208, as Blacodini, under the valid tribe name Opatrini), but not generally accepted as valid.
*Clitobiates Mulsant and Rey, 1859: 141 [stem: Clitobi-]. Type genus: Clitobius Mulsant and Rey, 1859. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 208, as Clitobiini, under the valid tribe name Opatrini), but not generally accepted as valid.
Gonocéphalates Mulsant and Revelière, 1861: 154 [stem: Gonocephal-]. Type genus: Gonocephalum Solier, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Gerstaecker (1861: 483, as Gonocephalidae).
*Dilamites Jacquelin du Val, 1861: 279 [stem: Dilam-]. Type genus: Dilamus Jacquelin du Val, 1861. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Ammobiini Desbrochers des Loges, 1902b: 1 [stem: Ammobi-]. Type genus: Ammobius Guérin-Méneville, 1844.
Emmallina Koch, 1956: 51 [stem: Emmal-]. Type genus: Emmalus Erichson, 1843 [as Emmallus, unjustified emendation of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Stenolamina Koch, 1956: 49 [stem: Stenolam-]. Type genus: Stenolamus Gebien, 1920.
Subtribe
Neopachypterina
Bouchard, Löbl and Merkl, 2007
Pachyptérates Mulsant and Rey, 1859: 83 [stem: Pachypter-]. Type genus: Pachypterus Lucas, 1846 [preoccupied genus name, not Pachypterus Swainson, 1839 [Pisces]; syn. of Neopachypterus Bouchard et al., 2007]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Desbrochers des Loges (1901: 146, as Pachypterini [actually spelled Pachnephorini but this name was corrected to Pachypterini later by Desbroches des Loges (1902a: 85)]; permanently invalid (Art. 39): based on preoccupied type genus.
Pachypterini G. S. Medvedev, 1968: 247 [stem: Pachypter-]. Type genus: Pachypterus Lucas, 1846 [preoccupied genus name, not Pachypterus Swainson, 1839 [Pisces]; syn. of Neopachypterus Bouchard et al., 2007]. Comment: family-group name proposed as new without reference to Pachyptérates Mulsant and Rey, 1859; permanently invalid (Art. 39): based on preoccupied type genus.
Neopachypterina Bouchard et al., 2007: 386 [stem: Neopachypter-]. Type genus: Neopachypterus Bouchard et al., 2007. Comment: replacement name for Pachyptérates Mulsant and Rey, 1859 and Pachypterini G. S. Medvedev, 1968 because of the homonymy of the type genus.
Tribe
Palorini
Matthews, 2003
*Palorinae E. G. Matthews, 2003a: 50 [stem: Palor-]. Type genus: Palorus Mulsant, 1854. Comment: unavailable name (Art. 16): not explicitly indicated as new.
Palorinae E. G. Matthews, 2003b: 7 [stem: Palor-]. Type genus: Palorus Mulsant, 1854. Comment: validation of Palorinae Matthews (2003a).
Tribe
Pedinini
Eschscholtz, 1829
Pediniden Eschscholtz, 1829b: 4 [stem: Pedin-]. Type genus: Pedinus Latreille, 1797.
Subtribe
Dendarina
Mulsant and Rey, 1854
Pandarites Mulsant and Rey, 1854: 153 [stem: Pandar-]. Type genus: Dendarus Dejean, 1821 [as Pandarus, unjustified emendation of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gerstaecker (1854: 226, as Pandaridae [incorrect stem formation]), and generally accepted as in Seidlitz (1889[Gatt.]: 126, as Dendarina).
*Omocratates Mulsant and Rey, 1854: 266, in key [stem: Omocrat-]. Type genus: Omocrates Mulsant, 1854 [preoccupied genus name, not Omocrates H. C. C. Burmeister, 1844 [Coleoptera: Scarabaeidae]; syn. of Phylan Dejean, 1821]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if found to be available then permanently invalid (Art. 39): based on preoccupied type genus.
*Micrositates Mulsant and Rey, 1854: 274 [stem: Microsit-]. Type genus: Micrositus Mulsant and Rey, 1854. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Isocérates Mulsant and Rey, 1854: 188 [stem: Isocer-]. Type genus: Isocerus Dejean, 1821 [preoccupied genus name, not Isocerus Illiger, 1802 [Coleoptera: Cerambycidae]; syn. of Neoisocerus Bouchard et al., 2005]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if found to be available then permanently invalid (Art. 39): based on preoccupied type genus.
*Héliopathaires Mulsant and Rey, 1854: 265 [stem: Heliopat-]. Type genus: Heliopates Dejean, 1834 [as Heliopathes, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Phylacides Lacordaire, 1859: 270 [stem: Phylac-]. Type genus: Phylax Brullé, 1832 [syn. of Dendarus Dejean, 1821]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Seidlitz (1893: 220, as Phylacina).
Bioplanina A. N. Reichardt, 1936: 24 [stem: Bioplanet-]. Type genus: Bioplanes Mulsant, 1854. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Eurynotina
Mulsant and Rey, 1854
Eurynotaires Mulsant and Rey, 1854: 156 [stem: Eurynot-]. Type genus: Eurynotus Kirby, 1819. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Koch (1956: 25, as Eurynotina), generally accepted as in Bouchard et al. (2005: 502, as Eurynotina).
Psectropini Kaszab, 1941: 33 [stem: Psectropod-]. Type genus: Psectropus sensu Kaszab, 1941 [not Psectropus Solier, 1848; syn. of Schyzochelus Koch, 1954]. Comment: based on misidentified type genus; incorrect original stem formation, not in prevailing usage.
Oncotini Koch, 1953c: 267, in key [stem: Oncot-]. Type genus: Oncotus Blanchard, 1845.
Schyzoschelina Koch, 1956: 25 [stem: Schyzoschel-]. Type genus: Schyzoschelus Koch, 1954.
Subtribe
Leichenina
Mulsant, 1854
Leichenaires Mulsant, 1854: 179 [stem: Leichen-]. Type genus: Leichenum Blanchard, 1845. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Casey (1890: 391, as Leichenini), generally accepted as in Bouchard et al. (2005: 502, as Leichenina).
Subtribe
Loensina
Koch, 1956
*Loensini Koch, 1955: 1 [stem: Loens-]. Type genus: Loensus Lucas, 1920. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Loensini Koch, 1956: 402 [stem: Loens-]. Type genus: Loensus Lucas, 1920.
Subtribe
Melambiina
Mulsant and Rey, 1854
Melambiates Mulsant and Rey, 1854: 267 [stem: Melambi-]. Type genus: Melambius Mulsant and Rey, 1854. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Español (1945: 226, as Melambina [incorrect stem formation]), generally accepted as in Bouchard et al. (2005: 502, as Melambiina).
Litoborinae Antoine, 1941: 19-21 [stem: Litobor-]. Type genus: Litoborus Mulsant and Rey, 1854.
Zadenina Koch, 1956: 279, in key [stem: Zaden-]. Type genus: Zadenos Laporte, 1840.
Subtribe
Pedinina
Eschscholtz, 1829
Pediniden Eschscholtz, 1829b: 4 [stem: Pedin-]. Type genus: Pedinus Latreille, 1797. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by G. R. Waterhouse (1845: 32, as Pedinidae), generally accepted as in Bouchard et al. (2005: 502, as Pedinini).
Subtribe
Platynotina
Mulsant and Rey, 1853
*Hétéroscélites Solier, 1836: 502 [stem: Heteroscelid-]. Type genus: Heteroscelis Latreille, 1829 [preoccupied genus name, not Heteroscelis Latreille, 1829 [Hemiptera]; syn. of Anomalipus Guérin-Méneville, 1831]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if found to be available then permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Platynotaires Mulsant and Rey, 1853: 263 [stem: Platynot-]. Type genus: Platynotus Fabricius, 1801. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 226, as Platynotini), generally accepted as in Bouchard et al. (2005: 502, as Platynotina); First Reviser (Platynotina Mulsant and Rey, 1853 vs Trigonopodina Mulsant and Rey, 1853) not determined, current usage maintained.
Trigonopaires Mulsant and Rey, 1853: 104 [stem: Trigonopod-]. Type genus: Trigonopus Mulsant and Rey, 1853. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aalbu (2006: 70, as Trigonopina); incorrect original stem formation, not in prevailing usage.
Gonopides Lacordaire, 1859: 255 [stem: Gonopod-]. Type genus: Gonopus Latreille, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 207, as Gonopini), generally accepted as in Gebien (1938a: 90, as Gonopini); incorrect original stem formation, not in prevailing usage.
Platynotini Koch, 1953c: 268, in key [stem: Platynot-]. Type genus: Platynotus Fabricius, 1801. Comment: family-group name proposed as new without reference to Platynotaires Mulsant and Rey, 1853.
Anomalipina Koch, 1954: 427 [stem: Anomalipod-]. Type genus: Anomalipus Guérin-Méneville, 1831 [unjustified emendation of Anomalipes Guérin-Méneville, 1831 by Lacordaire, 1859; unjustified emendation in prevailing usage, treated as justified emendation (Art. 33.2.3.1)]. Comment: incorrect original stem formation, not in prevailing usage.
*anchophthalmoid Koch, 1956: 71, in key [stem: Anchophthalm-]. Type genus: Anchophthalmus Gerstaecker, 1854. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun.
*selinoid Koch, 1956: 70, in key [stem: Selin-]. Type genus: Selinus Mulsant and Rey, 1853. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun; Selinini Jeannel, 1948 (type genus Selina Motschulsky, 1858) is currently used as a valid in Carabidae.
*melanocratoid Iwan, 1996: 379, 380-381 [stem: Melanocrat-]. Type genus: Melanocratus Fairmaire, 1895. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun.
Subtribe
Pythiopina
Koch, 1953
Pythiopini Koch, 1953b: 245 [stem: Pythiop-]. Type genus: Pythiopus Koch, 1953. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Pythiopod-).
Tribe
Platyscelidini
Lacordaire, 1859
Platyscélides Lacordaire, 1859: 229 [stem: Platyscelid-]. Type genus: Platyscelis Latreille, 1818 [placed on the Official List of Generic Names in Zoology (ICZN 1993e)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1889 [Gatt.]: 126, as Platyscelina [incorrect stem formation]), generally accepted as in Bouchard et al. (2005: 502, as Platyscelidini); incorrect original stem formation, not in prevailing usage; correction of stem by Egorov (1990: 401) in order to avoid homonymy with Platyscelidae Bate, 1862 in Crustacea.
Tribe
Praeugenini
De Moor, 1970
*Praogenini Ferreira, 1965: 312 [stem: Praeugen-]. Type genus: Praeugena Laporte, 1840 [as Praogena, unjustified emendation of type genus name, not in prevailing usage]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Praeugenina De Moor, 1970: 4 [stem: Praeugen-]. Type genus: Praeugena Laporte, 1840.
Tribe
Rhysopaussini
Wasmann, 1896
Rhysopaussidae Wasmann, 1896: 613 [stem: Rhysopauss-]. Type genus: Rhysopaussus Wasmann, 1896.
Tribe
Scaurini
Billberg, 1820
Scaurides Billberg, 1820a: 31 [stem: Scaur-]. Type genus: Scaurus Fabricius, 1775. Comment: this family-group name was also used in the same year by Billberg (1820b: 392, as Scaurides).
Tribe
Scotobiini
Solier, 1838
Scotobites Solier, 1838: 39 [stem: Scotobi-]. Type genus: Scotobius Germar, 1824. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 336, as Scotobioidae), generally accepted as in Bouchard et al. (2005: 502, as Scotobiini); incorrect original stem formation, not in prevailing usage.
Tribe
Tenebrionini
Latreille, 1802
Tenebrionites Latreille, 1802: 165 [stem: Tenebrion-]. Type genus: Tenebrio Linnaeus, 1758.
Biuini Skopin, 1978: 224, in key [stem: Bi-]. Type genus: Bius Dejean, 1834. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Titaenini
Fauvel, 1905
Titaenini Fauvel, 1905b: 209 [stem: Titaen-]. Type genus: Titaena Erichson, 1842.
Tribe
Toxicini
Oken, 1843
Toxiciden Oken, 1843: 484 [stem: Toxic-]. Type genus: Toxicum Latreille, 1802.
Subtribe
Eudysantina
Bouchard, Lawrence, Davies and Newton, 2005
Dysantinae Gebien, 1922: 289 [stem: Dysant-]. Type genus: Dysantes Pascoe, 1871 [preoccupied genus name, not Dysantes Foerster, 1868 [Hymenoptera]; syn. of Eudysantes Bouchard et al., 2005]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Eudysantina Bouchard et al., 2005: 508 [stem: Eudysant-]. Type genus: Eudysantes Bouchard et al., 2005. Comment: replacement name for Dysantina Gebien, 1922 because of the homonymy of the type genus.
Subtribe
Nycteropina
Lacordaire, 1859
Nyctéropides Lacordaire, 1859: 388 [stem: Nycterop-]. Type genus: Nycteropus Klug, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 211, as Nycteropini), generally accepted as in Bouchard et al. (2005: 502, as Nycteropina).
Subtribe
Toxicina
Oken, 1843
Toxiciden Oken, 1843: 484 [stem: Toxic-]. Type genus: Toxicum Latreille, 1802. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Tulk (1847: 614, as Toxicidae); family-group name previously attributed to Lacordaire (1859).
Anthraciini Houlbert, 1922b: 185, in key [stem: Anthraci-]. Type genus: Anthracias Dejean, 1834 [syn. of Cryphaeus Klug, 1833].
Tribe
Triboliini
Gistel, 1848
Triboliidae Gistel, 1848: [4] [stem: Triboli-]. Type genus: Tribolium W. S. MacLeay, 1825 [placed on the Official List of Generic Names in Zoology (ICZN 1988e)].
Tribe
Ulomini
Blanchard, 1845
Ulomites Blanchard, 1845b: 16 [stem: Ulom-]. Type genus: Uloma Dejean, 1821 [placed on the Official List of Generic Names in Zoology (ICZN 1975)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Blanchard (1853: 164, as Ulomitae), generally accepted as in Bouchard et al. (2005: 502, as Ulomini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Ulomat-).
*Oligocaroides Solier, 1851: 225 [stem: Oligocar-]. Type genus: Oligocara Solier, 1848. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Alégoriides Lacordaire, 1859: 325 [stem: Alegori-]. Type genus: Alegoria Laporte, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Seidlitz (1894: 545, as Alegoriina).
Neopsectropinae Kaszab, 1941: 30 [stem: Neopsectropod-]. Type genus: Neopsectropus Kaszab, 1941. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Alleculinae
Laporte, 1840
Alléculites Laporte, 1840b: 242 [stem: Allecul-]. Type genus: Allecula Fabricius, 1801. Comment: given precedence over the older names Xystropodinae Solier, 1835 and Cteniopodinae Solier, 1835 (Art. 35.5) (see Bouchard et al. 2005).
Tribe
Alleculini
Laporte, 1840
Alléculites Laporte, 1840b: 242 [stem: Allecul-]. Type genus: Allecula Fabricius, 1801. Comment: given precedence over the older name Xystropodini Solier, 1835 (Art. 35.5) (see Bouchard et al. 2005).
Subtribe
Alleculina
Laporte, 1840
Alléculites Laporte, 1840b: 242 [stem: Allecul-]. Type genus: Allecula Fabricius, 1801. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Packard (1874: iii, as Alleculidae), generally accepted as in Bouchard et al. (2005: 502, as Alleculina).
*Cylindrothorides Lacordaire, 1859: 494 [stem: Cylindrothor-]. Type genus: Cylindrothorus Solier, 1844. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Upinellae J. L. LeConte, 1866b: 137 [stem: Upinell-]. Type genus: Upinella Mulsant, 1856.
Subtribe
Gonoderina
Seidlitz, 1896
Gonoderina Seidlitz, 1896: 83 [stem: Gonoder-]. Type genus: Gonodera Mulsant, 1856.
Pseudocistelini Portevin, 1934: 39, in key [stem: Pseudocistel-]. Type genus: Pseudocistela Crotch, 1873.
Subtribe
Mycetocharina
Gistel, 1848
Mycetocharisidae Gistel, 1848: [10] [stem: Mycetochar-]. Type genus: Mycetochara Berthold, 1827 [as Mycetochares, incorrect subsequent spelling not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Xystropodina
Solier, 1835
Xystropides Solier, 1835a: 229 [stem: Xystropod-]. Type genus: Xystropus Solier, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 390, as Xystropodidae), generally accepted as in Bouchard et al. (2005: 502, as Xystropodina); incorrect original stem formation, not in prevailing usage.
Lystronychides Lacordaire, 1859: 512 [stem: Lystronych-]. Type genus: Lystronychus Latreille, 1829 [incorrect subsequent spelling of Lystronichus, incorrect spelling in current usage, treated as correct original spelling (Art. 33.3.1)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte and G. H. Horn (1883: 390, as Lystronychi).
Tribe
Cteniopodini
Solier, 1835
Cisteleniae Latreille, 1802: 188 [stem: Cistel-]. Type genus: Cistela Fabricius, 1775 [preoccupied genus name, not Cistela Geoffroy, 1762 [Coleoptera: Byrrhidae]; syn. of Cteniopus Solier, 1835; Geoffroy’s generic name was suppressed for the purposes of the Principle of Priority but not for the Principle of Homonymy (ICZN 1994a)]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Cténiopites Solier, 1835a: 245 [stem: Cteniopod-]. Type genus: Cteniopus Solier, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 107, as Cteniopodidae), generally accepted as in Bouchard et al. (2005: 502, as Cteniopodini); incorrect original stem formation, not in prevailing usage.
Omophliens Mulsant, 1856a: 65 [stem: Omophl-]. Type genus: Omophlus Dejean, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1896: 28, as Omophlini), generally accepted as in Ogloblin and Znojko (1950: 9, as Omophlinae).
Phibalidae Gistel, 1856a: 384 [stem: Phibal-]. Type genus: Phibalus Gistel, 1856 [the genus Phibalus Gistel, 1856 originally included only one species therefore the type species is Chrysomela pubescens Linnaeus, 1758 by monotypy; this genus has not been used subsequently to our knowledge; syn. nov. of Omophlus Dejean, 1834]. Comment: syn. nov.
Petriidae Semenov, 1893b: 359 [stem: Petri-]. Type genus: Petria Semenov, 1894.
Podontini Portevin, 1934: 45, in key [stem: Podont-]. Type genus: Podonta Solier, 1835.
Subfamily
Diaperinae
Latreille, 1802
Diaperialae Latreille, 1802: 161 [stem: Diaper-]. Type genus: Diaperis Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Crypticini
Brullé, 1832
Crypticites Brullé, 1832: 190 [stem: Cryptic-]. Type genus: Crypticus Latreille, 1816. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. G. Thomson (1859: 114, as Crypticina), generally accepted as in Bouchard et al. (2005: 503, as Crypticini).
Oochrotini Desbrochers des Loges, 1901: 143, in key [stem: Oochrot-]. Type genus: Oochrotus Lucas, 1852.
Tribe
Diaperini
Latreille, 1802
Diaperialae Latreille, 1802: 161 [stem: Diaper-]. Type genus: Diaperis Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subtribe
Adelinina
LeConte, 1862
*Sitophagiens Mulsant, 1854: 263 [stem: Sitophag-]. Type genus: Sitophagus Mulsant, 1854. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Alphitophagida Gistel, 1856b: 185 [stem: Alphitophag-]. Type genus: Alphitophagus Stephens, 1832. Comment: although this is the oldest name for the subtribe, we recommend that an application be submitted to the Commission to conserve usage of the name Adelinina J. L. LeConte, 1862.
*Gnathocérites Jacquelin du Val, 1861: 304 [stem: Gnatocer-]. Type genus: Gnatocerus Thunberg, 1814 [as Gnathocerus, incorrect subsequent spelling of type genus name, not in prevailing usage; Gnatocerus Thunberg, 1814 placed on the Official List of Generic Names in Zoology (ICZN 1975)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Jacquelin du Val (1861); incorrect original stem formation, not in prevailing usage; Gnathoceridae Schoch, 1894 is available in Scarabaeidae.
Adelinini J. L. LeConte, 1862: 237 [stem: Adelin-]. Type genus: Adelina Dejean, 1835. Comment: this well-established family-group name is threatened by the discovery of the older name Alphitophagida Gistel, 1856; although using Reversal of Precedence (Art. 23.9.1) to conserve usage of Adelinina J. L. LeConte, 1862 would be preferable, this can not be done because we cannot find 25 references in which this name has been used in the last 50 years; current usage is maintained here and an application should be submitted to the Commission in order to conserve Adelinina J. L. LeConte, 1862.
Schedarosini Reitter, 1876b: 42 [stem: Schedaros-]. Type genus: Schedarosus Reitter, 1876 [syn. of Adelina Dejean, 1835]. Comment: originally placed in the family Cucujidae.
Doliemini Reitter, 1917: 58 [stem: Doliem-]. Type genus: Doliema Pascoe, 1860 [syn. of Adelina Dejean, 1835].
Gnathocerini Skopin, 1978: 228, in key [stem: Gnatocer-]. Type genus: Gnatocerus Thunberg, 1814 [as Gnathocerus, incorrect subsequent spelling of type genus name, not in prevailing usage; Gnatocerus Thunberg, 1814 placed on the Official List of Generic Names in Zoology (ICZN 1975)]. Comment: incorrect original stem formation, not in prevailing usage; Gnathoceridae Schoch, 1894 is available in Scarabaeidae.
Subtribe
Diaperina
Latreille, 1802
Diaperialae Latreille, 1802: 161 [stem: Diaper-]. Type genus: Diaperis Geoffroy, 1762 [placed on the on Official List of Generic Names in Zoology (ICZN 1994a)].
Pentaphyllaires Mulsant, 1854: 196 [stem: Pentaphyll-]. Type genus: Pentaphyllus Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 236, as Pentaphylli), generally accepted as in Silfverberg (1992: 61, as Pentaphyllini); the name Pentaphyllinae Schindewolf, 1942 (type genus Pentaphyllum de Koninck, 1872) is used as valid in Anthozoa; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Platydeminae Reitter, 1917: 61 [stem: Platydem-]. Type genus: Platydema Laporte and Brullé, 1831.
*Hoplocephalini Kwieton, 1982: 98 [stem: Hoplocephal-]. Type genus: Hoplocephala Laporte and Brullé, 1831 [based on unjustified emendation of type genus name; originally proposed as Oplocephala; emended name in prevailing usage, treated as a justified emendation (Art. 33.2.3.1); syn. of Neomida Latreille, 1829]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Ectychini
Doyen, Matthews and Lawrence, 1990
Ectychini Doyen et al., 1990: 247 [stem: Ectych-]. Type genus: Ectyche Pascoe, 1869.
Tribe
Gnathidiini
Gebien, 1921
Gnathidiinae Gebien, 1921: 41 [stem: Gnathidi-]. Type genus: Gnathidium Gebien, 1921.
Subtribe
Anopidiina
Jeannel and Paulian, 1945
Anopidiini Jeannel and Paulian, 1945: 62 [stem: Anopidi-]. Type genus: Anopidium Jeannel and Paulian, 1945.
Subtribe
Gnathidiina
Gebien, 1921
Gnathidiinae Gebien, 1921: 41 [stem: Gnathidi-]. Type genus: Gnathidium Gebien, 1921.
Tribe
Hyociini
Medvedev and Lawrence, 1982
Hyocini G. S. Medvedev and Lawrence, 1982: 548 [stem: Hyoci-]. Type genus: Hyocis Pascoe, 1866.
Subtribe
Brittonina
Medvedev and Lawrence, 1986
Brittonina G. S. Medvedev and Lawrence, 1986: 574 [stem: Britton-]. Type genus: Brittona G. S. Medvedev and Lawrence, 1986.
Subtribe
Hyociina
Medvedev and Lawrence, 1982
Hyocini G. S. Medvedev and Lawrence, 1982: 548 [stem: Hyoci-]. Type genus: Hyocis Pascoe, 1866. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Uptonina
Medvedev and Lawrence, 1986
Uptonina G. S. Medvedev and Lawrence, 1986: 581 [stem: Upton-]. Type genus: Uptona G. S. Medvedev and Lawrence, 1986.
Tribe
Hypophlaeini
Billberg, 1820
Hypophlaeides Billberg, 1820a: 33 [stem: Hypophlae-]. Type genus: Hypophlaeus Fabricius, 1790 [syn. of Corticeus Piller and Mitterpacher, 1783].
Corticeini Boddy, 1965: 144 [stem: Cortice-]. Type genus: Corticeus Piller and Mitterpacher, 1783.
Tribe
Leiochrinini
Lewis, 1894
Leiochrininae Lewis, 1894: 390 [stem: Leiochrin-]. Type genus: Leiochrinus Westwood, 1883.
Tribe
Myrmechixenini
Jacquelin du Val, 1858
Myrméchixénites Jacquelin du Val, 1858: 223 [stem: Myrmechixen-]. Type genus: Myrmechixenus Chevrolat, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Marseul (1863: 111, as Myrmekixeni [incorrect stem formation]), generally accepted as in Bouchard et al. (2005: 503, as Myrmechixenini); originally placed in the family Mycetaeidae (syn. of Endomychidae).
Tribe
Phaleriini
Blanchard, 1845
Phalériides Blanchard, 1845b: 29 [stem: Phaleri-]. Type genus: Phaleria Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1975)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1848: [10], as Phaleriadae [incorrect stem formation]), generally accepted as in Bouchard et al. (2005: 503, as Phaleriini).
Sepedonastidae Gistel, 1856a: 382 [stem: Sepedonast-]. Type genus: Sepedonastes Gistel, 1856 [syn. of Phaleria Latreille, 1802].
Cataphronetini Reitter, 1917: 57 [stem: Cataphronet-]. Type genus: Cataphronetis Lucas, 1846 [syn. of Phtora Germar, 1836].
Tribe
Scaphidemini
Reitter, 1922
Scaphidemini Reitter, 1922b: 2 [stem: Scaphidem-]. Type genus: Scaphidema Redtenbacher, 1849.
Tribe
Trachyscelini
Blanchard, 1845
Trachyscélides Blanchard, 1845b: 28 [stem: Trachyscel-]. Type genus: Trachyscelis Latreille, 1809. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by G. R. Waterhouse (1858: 60, as Trachyscelidae), generally accepted as in Bouchard et al. (2005: 503, as Trachyscelini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Trachyscelid-); conservation of the original stem avoids potential homonymy with family-group names based on the type genus Trachyscelida G. H. Horn, 1893 in Chrysomelidae.
Subfamily
Stenochiinae
Kirby, 1837
Stenochiadae Kirby, 1837: 238 [stem: Stenochi-]. Type genus: Stenochia Kirby, 1819.
Tribe
Cnodalonini
Oken, 1843
Cnodaliden Oken, 1843: 484 [stem: Cnodalon-]. Type genus: Cnodalon Latreille, 1797. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Tulk (1847: 614, as Cnodalidae); family-group name previously attributed to Gistel (1856a: 382); the correct stem based on Cnodalon is Cnodal-, therefore the original stem was correctly formed; however, since this family-group name has not been attributed to Oken (1843) in the past and the accepted stem has been Cnodalon- to this date, we have kept the currently accepted stem; an application to the Commission is necessary to conserve Cnodalon- as the correct stem for this taxon.
Coelometopidae Schaum, 1859: 71 [stem: Coelometop-]. Type genus: Coelometopus Solier, 1848. Comment: published before February 31, 1859; this family-group name was also used in the same year by Lacordaire (1859 [before 27 June]: 358, as Coelométopides).
Upidae C. G. Thomson, 1859: 116 [stem: Upid-]. Type genus: Upis Fabricius, 1792. Comment: incorrect original stem formation, not in prevailing usage.
Eutélides Lacordaire, 1859: 354 [stem: Eutel-]. Type genus: Eutelus Solier, 1843 [preoccupied genus name, not Eutelus Walker, 1834 [Hymenoptera]; syn. of Nodotelus Koch, 1950]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1866b: 469, as Eutelinae), generally accepted as in Gebien (1911: 431, as Eutelinae); permanently invalid (Art. 39): based on preoccupied type genus; Eutelini Ashmead, 1904 (type genus Eutelus Walker, 1834) is available in Hymenoptera.
Misolampides Lacordaire, 1859: 440 [stem: Misolamp-]. Type genus: Misolampus Latreille, 1807. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by and generally accepted as in F. Bates (1873: 373, as Misolampides [treated as Latin]).
Catapiestides Lacordaire, 1859: 381 [stem: Catapiest-]. Type genus: Catapiestus Perty, 1831. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Seidlitz (1895: 614, as Catapiestina).
*Camarides Desmarest, 1860: 165 [stem: Camari-]. Type genus: Camaria Lepeletier and Audinet-Serville, 1828. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Desmarest (1860).
Polypleuri J. L. LeConte, 1862: 229 [stem: Polypleur-]. Type genus: Polypleurus Eschscholtz, 1831.
Misolampidiini Reitter, 1917: 60, in key [stem: Misolampidi-]. Type genus: Misolampidius Solsky, 1876.
Menephilini Reitter, 1920: 15 [stem: Menephil-]. Type genus: Menephilus Mulsant, 1854.
Stenophanini Reitter, 1922b: 2, 3 [stem: Stenophan-]. Type genus: Stenophanes Solsky, 1876.
Hegemonini Reitter, 1922b: 3 [stem: Hegemon-]. Type genus: Hegemona Laporte, 1840.
Camariinae Anonymous, 1924: 164 [stem: Camari-]. Type genus: Camaria Lepeletier and Audinet-Serville, 1828. Comment: this was a translation of the vernacular name “Camariinen” used by Gebien (1919); Gebien’s family-group name is not available since it was proposed in a vernacular form after 1899 (Art. 11.7.2); for comments about authorship of Camariinae see Evenhuis et al. (2008: 4).
Nodotelini Koch, 1950: 67 [stem: Nodotel-]. Type genus: Nodotelus Koch, 1950. Comment: replacement name for Eutelini Lacordaire, 1859 because of the homonymy of the type genus.
*Thesileini Kaszab, 1982: 29 [stem: Thesile-]. Type genus: Thesilea Haag-Rutenberg, 1878. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Stenochiini
Kirby, 1837
Stenochiadae Kirby, 1837: 238 [stem: Stenochi-]. Type genus: Stenochia Kirby, 1819. Comment: incorrect original stem formation, not in prevailing usage.
Strongyliides Lacordaire, 1859: 478 [stem: Strongyli-]. Type genus: Strongylium Kirby, 1819 [name conserved over Strongylium Ditmar, 1809 [Myxomycota] (see Bouchard et al. 2005)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 241, as Strongyliini), generally accepted as in Aalbu et al. (2002: 483, as Strongyliini).
Tribe
Talanini
Champion, 1887 (1883)
Dignamptini J. L. LeConte and G. H. Horn, 1883: 385 [stem: Dignampt-]. Type genus: Dignamptus J. L. LeConte, 1878 [syn. of Talanus Jacquelin du Val, 1857].
Talanides Champion, 1887: 321 [stem: Talan-]. Type genus: Talanus Jacquelin du Val, 1857. Comment: family-group name conserved over Dignamptini J. L. LeConte and G. H. Horn, 1883 (Art. 40.2) (see Bouchard et al. 2005).
Tenebrionidae
incertae sedis
Betschiini Dajoz, 1980a: 135 [stem: Betschi-]. Type genus: Betschia Dajoz, 1980. Comment: originally placed in the family Colydiidae, the type genus was transferred to Tenebrionidae by Ivie and Ślipiński (1990: 18); placement uncertain although probably close to Gnathidiini (Ivie pers. comm. 2008).
Family
Prostomidae
Thomson, 1859
*Mégagnathes Motschulsky, 1849: 60 [stem: Megagnath-]. Type genus: Megagnathus Dejean, 1821 [preoccupied genus name, not Megagnathus Billberg, 1820 [Coleoptera: Cerambycidae]; syn. of Prostomis Latreille, 1819]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; if found to be available then permanently invalid (Art. 39): based on preoccupied type genus.
Prostomidae C. G. Thomson, 1859: 84 [stem: Prostom-]. Type genus: Prostomis Latreille, 1819. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Prostomid-).
Family
Synchroidae
Lacordaire, 1859
Synchroïdes Lacordaire, 1859: 544 [stem: Synchro-]. Type genus: Synchroa Newman, 1838. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 249, as Synchroae), generally accepted as in Lawrence and Newton (1995: 894, as Synchroidae).
Mallodryini G. H. Horn, 1888: 43 [stem: Mallodry-]. Type genus: Mallodrya G. H. Horn, 1888.
Family
Stenotrachelidae
Thomson, 1859
Stenotrachelidae C. G. Thomson, 1859: 124 [stem: Stenotrachel-]. Type genus: Stenotrachelus Berthold, 1827.
Subfamily
Stenotrachelinae
Thomson, 1859
Stenotrachelidae C. G. Thomson, 1859: 124 [stem: Stenotrachel-]. Type genus: Stenotrachelus Berthold, 1827.
Subfamily
Cephaloinae
LeConte, 1862
*Cephalaonides Motschulsky, 1860: 140 [stem: Cephalo-]. Type genus: Cephaloon Newman, 1838 [as Cephalaon, incorrect subsequent spelling of type genus name corrected by Motschulsky to Cephaloon on pp. 140 in same work]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1860).
Cephaloidae J. L. LeConte, 1862: 259 [stem: Cephalo-]. Type genus: Cephaloon Newman, 1838.
Subfamily
Nematoplinae
LeConte, 1862
Nematopli J. L. LeConte, 1862: 264 [stem: Nematopl-]. Type genus: Nematoplus J. L. LeConte, 1855.
Subfamily
Stoliinae
Nikitsky, 1985
Stoliinae Nikitsky, 1985: 32, in key [stem: Stoli-]. Type genus: Stolius Lewis, 1895.
Family
Oedemeridae
Latreille, 1810
Oedemerites Latreille, 1810: 216 [stem: Oedemer-]. Type genus: Oedemera A. G. Olivier, 1789.
Subfamily
Polypriinae
Lawrence, 2005
*Polypriidae Triplehorn and N. F. Johnson, 2005: 441 [stem: Polypri-]. Type genus: Polypria Chevrolat, 1874. Comment: unavailable family-group name, name published after 1999 and not explicitly indicated as new (Art. 16.1).
Polypriinae Lawrence, 2005: 671, in key [stem: Polypri-]. Type genus: Polypria Chevrolat, 1874.
Subfamily
Calopodinae
Costa, 1852
nomen protectum
Sparedriidae Gistel, 1848: [11] [stem: Sparedr-]. Type genus: Sparedrus Dejean, 1821. Comment: nomen oblitum (see Appendix 1); incorrect original stem formation, not in prevailing usage.
Calopini A. Costa, 1852: 4 [stem: Calopod-]. Type genus: Calopus Fabricius, 1775. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Semenov (1894: 449, as Calopodidae), generally accepted as in Švihla (2008: 353, as Calopodinae); nomen protectum (see Appendix 1); incorrect original stem formation, not in prevailing usage.
Catachirotidae Gistel, 1856a: 384 [stem: Catachirotet-]. Type genus: Catachirotes Gistel, 1856 [syn. of Calopus Fabricius, 1775]. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Oedemerinae
Latreille, 1810
Oedemerites Latreille, 1810: 216 [stem: Oedemer-]. Type genus: Oedemera A. G. Olivier, 1789.
Tribe
Asclerini
Gistel, 1848
Ascleraeidae Gistel, 1848: [11] [stem: Ascler-]. Type genus: Asclera Stephens, 1839. Comment: incorrect original stem formation, not in prevailing usage.
Ganglbaueriidae Semenov, 1894: 450 [stem: Ganglebaueri-]. Type genus: Ganglbaueria Semenov, 1891.
*Hypasclerini Macnamara, 1971: 164 [stem: Hypascler-]. Type genus: Hypasclera Kirsch, 1866. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Hypasclerini Švihla, 1986: 161 [stem: Hypascler-]. Type genus: Hypasclera Kirsch, 1866. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently.
*Danacerinae Švihla, 1986: 161 [stem: Danerc-]. Type genus: Danerces Westwood, 1875. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently; incorrect original stem formation, not in prevailing usage.
*Oxacini Švihla, 1986: 161 [stem: Oxacid-]. Type genus: Oxacis J. L. LeConte, 1866. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently; incorrect original stem formation, not in prevailing usage.
Tribe
Ditylini
Mulsant, 1858
*Dityles Motschulsky, 1849: 59 [stem: Dityl-]. Type genus: Ditylus Fischer von Waldheim, 1817. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Ditylates Mulsant, 1858: 100 [stem: Dityl-]. Type genus: Ditylus Fischer von Waldheim, 1817. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Semenov (1894: 450, as Ditylidae), generally accepted as in Kriska (2002: 517, as Ditylini).
Tribe
Nacerdini
Mulsant, 1858
Nacerdates Mulsant, 1858: 104 [stem: Nacerd-]. Type genus: Nacerdes Dejean, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Semenov (1894: 450, as Nacerdidae), generally accepted as in Hansen (1996: 175, as Nacerdini).
Tribe
Oedemerini
Latreille, 1810
Oedemerites Latreille, 1810: 216 [stem: Oedemer-]. Type genus: Oedemera A. G. Olivier, 1789.
Oncomeradae Gistel, 1856a: 384 [stem: Oncomer-]. Type genus: Oncomera Stephens, 1829 [subgenus of Oedemera A. G. Olivier, 1789; placed on the Official List of Generic Names in Zoology (ICZN 1988f)]. Comment: incorrect original stem formation, not in prevailing usage.
*Oncomerinini Švihla, 1986: 161 [stem: Oncomerin-]. Type genus: Oncomerina Seidlitz, 1899 [syn. of Oncomera Stephens, 1829]. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently.
Tribe
Stenostomatini
Mulsant, 1858
Sténostomates Mulsant, 1858: 228 [stem: Stenostomat-]. Type genus: Stenostoma Latreille, 1810. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Semenov (1894: 450, as Stenostomatidae), generally accepted as in Švihla (2008: 369, as Stenostomini [incorrect stem formation]); incorrect original stem formation, not in prevailing usage.
Family
Meloidae
Gyllenhal, 1810
Melooides Gyllenhal, 1810: 481 [stem: Melo-]. Type genus: Meloe Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1999c)]. Comment: given precedence over the older name Horiidae Latreille, 1802 and placed on the Official List of Family-Group Names in Zoology (ICZN 1999c).
Subfamily
Eleticinae
Wellman, 1910
Eleticides Wellman, 1910: 221 [stem: Eletic-]. Type genus: Eletica Dejean, 1834. Comment: First Reviser (Eleticinae Wellman, 1910 vs Derideinae Wellman, 1910) not determined, current usage maintained.
Tribe
Derideini
Wellman, 1910
Derideides Wellman, 1910: 222 [stem: Deride-]. Type genus: Deridea Westwood, 1875.
Tribe
Eleticini
Wellman, 1910
Eleticides Wellman, 1910: 221 [stem: Eletic-]. Type genus: Eletica Dejean, 1834.
Subtribe
Eleticina
Wellman, 1910
Eleticides Wellman, 1910: 221 [stem: Eletic-]. Type genus: Eletica Dejean, 1834.
Subtribe
Eospastina
Selander, 1966
Eospastina Selander, 1966: 474 [stem: Eospast-]. Type genus: Eospasta Selander, 1966.
Tribe
Ertlianini
Selander, 1966
Ertliini Kaszab, 1959: 71 [stem: Ertli-]. Type genus: Ertlia Borchmann, 1942 [preoccupied genus name, not Ertlia Aurivillius, 1907 [Coleoptera: Cerambycidae]; syn. of Morphozonitis Pic, 1922]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Ertlianina Selander, 1966: 478 [stem: Ertlian-]. Type genus: Ertliana Selander, 1964 [syn. of Morphozonitis Pic, 1922]. Comment: replacement name for Ertliini Kaszab, 1959 because of the homonymy of the type genus; although Selander (1991: 78) used Morphozonitina Kaszab, 1969 as the valid name for this taxon (because of the synonymy of Ertliana Selander with Morphozonitis Pic) the Principle of Priority applies here and therefore the oldest available family-group name Ertlianini should be used as valid instead.
Morphozonitini Kaszab, 1969: 242 [stem: Morphozonitid-]. Type genus: Morphozonitis Pic, 1922. Comment: replacement name for Ertliini Kaszab, 1959 because of the homonymy of the type genus and Ertlianina Selander, 1966 because of the synonymy of the type genus; incorrect original stem formation, not in prevailing usage.
Tribe
Spasticini
Kaszab, 1959
Spasticina Kaszab, 1959: 72 [stem: Spastic-]. Type genus: Spastica Lacordaire, 1859.
Subtribe
Anthicoxenina
Selander, 1966
Anthicoxenina Selander, 1966: 470 [stem: Anthicoxen-]. Type genus: Anthicoxenus Fairmaire and Germain, 1860.
Subtribe
Protomeloina
Abdullah, 1965
Protomeloini M. Abdullah, 1965b: 43 [stem: Protomelo-]. Type genus: Protomeloe M. Abdullah, 1965.
Subtribe
Spasticina
Kaszab, 1959
Spasticina Kaszab, 1959: 72 [stem: Spastic-]. Type genus: Spastica Lacordaire, 1859.
Subtribe
Xenospastina
Selander, 1966
Xenospastina Selander, 1966: 460, in key [stem: Xenospast-]. Type genus: Xenospasta Selander, 1966.
Subfamily
Meloinae
Gyllenhal, 1810
Melooides Gyllenhal, 1810: 481 [stem: Melo-]. Type genus: Meloe Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1999c)].
Tribe
Cerocomini
Leach, 1815
Cerocomatida Leach, 1815: 105 [stem: Cerocom-]. Type genus: Cerocoma Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Epicautini
Parker and Böving, 1924
Macrobases J. L. LeConte, 1862: 272 [stem: Macrobase-]. Type genus: Macrobasis J. L. LeConte, 1862 [subgenus of Epicauta Dejean, 1834]. Comment: incorrect original stem formation, not in prevailing usage.
Apterospastides Wellman, 1910: 221 [stem: Apterospast-]. Type genus: Apterospasta J. L. LeConte, 1862 [syn. of Macrobasis J. L. LeConte, 1862].
Epicautini Parker and Böving, 1924: 25 [stem: Epicaut-]. Type genus: Epicauta Dejean, 1834. Comment: family-group name previously attributed to Denier (1935); this name is threatened by the two older names Macrobaseini J. L. LeConte, 1862 and Apterospastini Wellman, 1910; to maintain nomenclatural stability, we believe that an application should be submitted to the Commission to conserve usage of the well-established name Epicautini.
*Henoini Böving and Craighead, 1931: 79 [stem: Heno-]. Type genus: Henous Haldeman, 1852 [syn. of Epicauta Dejean, 1834]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Eupomphini
LeConte, 1862
Eupomphae J. L. LeConte, 1862: 274 [stem: Eupomph-]. Type genus: Eupompha J. L. LeConte, 1858. Comment: First Reviser found (Eupomphini J. L. LeConte, 1862 vs Phodagini J. L. LeConte, 1862) is Selander (1991: 78, as Eupomphina).
Phodagae J. L. LeConte, 1862: 274 [stem: Phodag-]. Type genus: Phodaga J. L. LeConte, 1858.
Calospastides Wellman, 1910: 221 [stem: Calospast-]. Type genus: Calospasta J. L. LeConte, 1862 [syn. of Eupompha J. L. LeConte, 1858].
Gynaecomelodides Wellman, 1910: 221 [stem: Gynaecomelo-]. Type genus: Gynaecomeloe Wellman, 1910 [syn. of Cordylospasta G. H. Horn, 1875]. Comment: incorrect original stem formation, not in prevailing usage.
Cordylospastides Wellman, 1910: 221 [stem: Cordylospast-]. Type genus: Cordylospasta G. H. Horn, 1875.
Cysteodemides Wellman, 1910: 221 [stem: Cysteodem-]. Type genus: Cysteodemus J. L. LeConte, 1851.
Tegroderini L. S. Dillon, 1952: 373 [stem: Tegroder-]. Type genus: Tegrodera J. L. LeConte, 1851.
Megetrina Kaszab, 1959: 80 [stem: Megetr-]. Type genus: Megetra J. L. LeConte, 1859.
Tribe
Lyttini
Solier, 1851
Cantharidiae Latreille, 1802: 185 [stem: Canthar-]. Type genus: Cantharis sensu Latreille, 1802 [not Cantharis Linnaeus, 1758; syn. of Lytta Fabricius, 1775]. Comment: based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principles of Priority and Homonymy (Art. 65.2.1); also see Cantharidae Imhoff, 1856.
*Lyttes Motschulsky, 1849: 59 [stem: Lytt-]. Type genus: Lytta Fabricius, 1775. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Lyttoides Solier, 1851: 278 [stem: Lytt-]. Type genus: Lytta Fabricius, 1775. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1856a: 385, as Lyttadae [incorrect stem formation]), generally accepted as in Selander (1991: 78, as Lyttina); although this is not the oldest name for the tribe, we recommend that an application be submitted to the Commission to suppress Cantharini Latreille, 1802 because it is based on a misidentified type genus (Art. 65.2.1).
*Alosimates Mulsant, 1857: 358 [stem: Alosim-]. Type genus: Alosimus Mulsant, 1857. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Selander (1991: 65, as Alosimina), but not generally accepted as valid; Selander’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Lydini Wellman, 1908: 424 [stem: Lyd-]. Type genus: Lydus Dejean, 1821. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently; Wellman correctly pointed out that the family-group for a taxon based on Lydus should be Lydini, however he did not want to use this family-group name since the name Lydidae Newman, 1834 (type genus Lyda Fabricius, 1805) is available for a group of Hymenoptera: Symphyta.
Sybarides Wellman, 1910: 221 [stem: Sybare-]. Type genus: Sybaris Stephens, 1832. Comment: incorrect original stem formation, not in prevailing usage.
Lydina Kaszab, 1959: 82 [stem: Lyd-]. Type genus: Lydus Dejean, 1821. Comment: Kaszab proposed the subtribe name Lydina without reference to the comment by Wellman (1908) and his family-group name has been used as valid in Meloidae since (e.g., Turco et al. 2006); the name Lydidae Newman, 1834 (type genus Lyda Fabricius, 1805) is available for a group of Hymenoptera: Symphyta; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Prolyttini Selander, 1960: 25 [stem: Prolytt-]. Type genus: Prolytta Kaszab, 1959.
Tribe
Meloini
Gyllenhal, 1810
Melooides Gyllenhal, 1810: 481 [stem: Melo-]. Type genus: Meloe Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1999c)].
Proscarabaeidae Gistel, 1848: [11] [stem: Proscarabae-]. Type genus: Proscarabaeus Schrank, 1781 [syn. of Meloe Linnaeus, 1758].
Tribe
Mylabrini
Rafinesque, 1815
Mylabria Rafinesque, 1815: 114 [stem: Mylabr-]. Type genus: Mylabris Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1995a)]. Comment: family-group name previously attributed to Billberg (1820a); Brullé (1832: 190) uses the family-group name “Corynites” for a group containing the genera Sarrotrium Illiger, 1798, Orthocerus Latreille, 1797, Chiroscelis Lamarck, 1804, Toxicum Latreille, 1802 and Boros Herbst, 1797, although Brullé could have based his family-group name on Coryna Billberg, 1813 (which is a junior homonym and a synonym of Hycleus Latreille, 1817 in Mylabrini) we cannot be sure and therefore we have not considered this a separate family-group name.
Zonabrini Wellman, 1908: 424 [stem: Zonabr-]. Type genus: Zonabris Harold, 1879 [syn. of Mylabris Fabricius, 1775].
Calydina Kaszab, 1960: 126 [stem: Calyd-]. Type genus: Calydus Reitter, 1896.
Tribe
Pyrotini
MacSwain, 1956
Pyrotini MacSwain, 1956: 67 [stem: Pyrot-]. Type genus: Pyrota Dejean, 1834.
Subfamily
Tetraonycinae
Böving and Craighead, 1931
Tetraonycidae Böving and Craighead, 1931: 59, in key [stem: Tetraonyc-]. Type genus: Tetraonyx Latreille, 1809. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Tetraonych-).
Meloetyphlini Borgmeier, 1937: 248 [stem: Meloetyphl-]. Type genus: Meloetyphlus C. O. Waterhouse, 1872.
Subfamily
Nemognathinae
Laporte, 1840
Nemognathites Laporte, 1840b: 280 [stem: Nemognath-]. Type genus: Nemognatha Illiger, 1807 [as Nemognathus, unjustified emendation of type genus name by Latreille (1829b), not in prevailing usage; Nemognatha Illiger, 1807 placed on the Official List of Generic Names in Zoology (ICZN 1999c)]. Comment: Nemognathinae Laporte, 1840 placed on the Official List of Family-Group Names in Zoology and given precedence over Horiinae Latreille, 1802 (ICZN 1999c).
Tribe
Horiini
Latreille, 1802
Horiales Latreille, 1802: 182 [stem: Hori-]. Type genus: Horia Fabricius, 1787 [placed on the Official List of Generic Names in Zoology (ICZN 1999c)]. Comment: Horiidae Latreille, 1802 placed on the Official List of Family-Group Names in Zoology (ICZN 1999c); Meloidae Gyllenhal, 1810 and Nemognathinae Laporte, 1840 given precedence over this name (ICZN 1999c).
Cissitini Kaszab, 1966: 187 [stem: Cissit-]. Type genus: Cissites Latreille, 1804. Comment: we accept the character about parameres given by Kaszab (1966: 187) to characterize this tribe as sufficient to fulfill the requirements of Art. 13.1; Selander (1991: 66) treated this name as unavailable.
Tribe
Nemognathini
Laporte, 1840
Nemognathites Laporte, 1840b: 280 [stem: Nemognath-]. Type genus: Nemognatha Illiger, 1807 [as Nemognathus, unjustified emendation of type genus name by Latreille (1829b), not in prevailing usage; Nemognatha Illiger, 1807 placed on the Official List of Generic Names in Zoology (ICZN 1999c)].
Subtribe
Nemognathina
Laporte, 1840
Nemognathites Laporte, 1840b: 280 [stem: Nemognath-]. Type genus: Nemognatha Illiger, 1807 [as Nemognathus, unjustified emendation of type genus name by Latreille (1829b), not in prevailing usage; Nemognatha Illiger, 1807 placed on the Official List of Generic Names in Zoology (ICZN 1999c)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1862: 270, as Nemognathini), generally accepted as in Hansen (1996: 176, as Nemognathinae).
Hornii Dugès, 1889: 213 [stem: Horni-]. Type genus: Hornia Riley, 1877.
Tricraniides Wellman, 1910: 222 [stem: Tricrani-]. Type genus: Tricrania J. L. LeConte, 1860.
Subtribe
Zonitidina
Mulsant, 1857
*Zonites Motschulsky, 1849: 59 [stem: Zonitid-]. Type genus: Zonitis Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1999c)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849); incorrect original stem formation, not in prevailing usage.
Zonitaires Mulsant, 1857: 322 [stem: Zonitid-]. Type genus: Zonitis Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1999c)]. Comment: incorrect original stem formation, not in prevailing usage; Zonitidinae Mulsant, 1857 placed on the Official List of Family-Group Names in Zoology (ICZN 1999c); this family-group name is a senior homonym of Zonitidae Mörch, 1864 in Mollusca (based on Zonites Montfort, 1810) which is used as valid in recent literature (see Bologna and Pinto 1997); stem of family-group name changed to Zonitid- by the Commission (ICZN 1999c) to remove from homonymy with Mörch’s name.
*Leptopalpina Kaszab, 1959: 102, Fig. 97 [stem: Leptopalp-]. Type genus: Leptopalpus Guérin-Méneville, 1834. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Leptopalpina Kaszab, 1963: 138 [stem: Leptopalp-]. Type genus: Leptopalpus Guérin-Méneville, 1834.
*Palaestrina Kaszab, 1969: 244 [stem: Palaestr-]. Type genus: Palaestra Laporte, 1840. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subtribe
Sitarina
Mulsant, 1857
*Apales Motschulsky, 1849: 59 [stem: Apal-]. Type genus: Apalus Fabricius, 1775. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Sitarates Mulsant, 1857: 393 [stem: Sitar-]. Type genus: Sitaris Latreille, 1802. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Dugès (1886: 580, as Sitarini), generally accepted as in Selander (1991: 79, as Sitarina).
Apalides Parker and Böving, 1924: 33 [stem: Apal-]. Type genus: Apalus Fabricius, 1775.
Stenoriides Parker and Böving, 1924: 32 [stem: Stenori-]. Type genus: Stenoria Mulsant, 1857.
Tribe
Stenoderini
Selander, 1991
*Stenoderini Selander, 1964: 1057 [stem: Stenoder-]. Type genus: Stenodera Eschscholtz, 1818. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Stenoderini Selander, 1991: 77 [stem: Stenoder-]. Type genus: Stenodera Eschscholtz, 1818. Comment: junior homonym of Stenoderini Pascoe, 1867 (type genus Stenoderus Dejean, 1821) currently used as valid in Cerambycidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Family
Mycteridae
Oken, 1843
Mycteriden Oken, 1843: 484 [stem: Mycter-]. Type genus: Mycterus Clairville, 1798.
Subfamily
Mycterinae
Oken, 1843
Rhinomaceridae Fleming, 1821: 50 [stem: Rhinomacr-]. Type genus: Rhinomacer Fabricius, 1781 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 2005c); syn. of Mycterus Clairville, 1798]. Comment: permanently invalid (Art. 39): based on suppressed type genus; incorrect original stem formation, not in prevailing usage.
Mycteriden Oken, 1843: 484 [stem: Mycter-]. Type genus: Mycterus Clairville, 1798. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Tulk (1847: 614, as Mycteridae); name previously attributed to Blanchard (1845b: 97).
Artaxidae Gistel, 1848: [8] [stem: Artax-]. Type genus: Artaxus Gistel, 1848 [syn. of Mycterus Clairville, 1798].
Subfamily
Eurypinae
Thomson, 1860
Eurypitae J. Thomson, 1860b: 63 [stem: Euryp-]. Type genus: Eurypus Kirby, 1819. Comment: the correct stem based on the type genus Eurypus is Eurypod-, however, we prefer to conserve the incorrect original stem formation for this name (Art. 29.5) to avoid homoymy with Eurypodini Gahan, 1906 (type genus Eurypoda W. Saunders, 1853) currently used as valid in Cerambycidae.
Lacconotini J. L. LeConte, 1862: 254 [stem: Lacconot-]. Type genus: Lacconotus J. L. LeConte, 1862.
Batobiina Seidlitz, 1917a: 88 [stem: Batobi-]. Type genus: Batobius Fairmaire and Germar, 1863.
Thisiina Seidlitz, 1917a: 91 [stem: Thisi-]. Type genus: Thisias Champion, 1889.
*Stilpnonotinae Blackwelder, 1945: 503 [stem: Stilpnonot-]. Type genus: Stilpnonotus Gray, 1832. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Subfamily
Hemipeplinae
Lacordaire, 1854
Hémipéplides Lacordaire, 1854b: 404 [stem: Hemipepl-]. Type genus: Hemipeplus Latreille, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1861: 96, as Hemipeplidae), generally accepted as in Pollock (2002: 532, as Hemipeplinae).
Family
Boridae
Thomson, 1859
Boridae C. G. Thomson, 1859: 117 [stem: Bor-]. Type genus: Boros Herbst, 1797.
Subfamily
Borinae
Thomson, 1859
Boridae C. G. Thomson, 1859: 117 [stem: Bor-]. Type genus: Boros Herbst, 1797.
Subfamily
Synercticinae
Lawrence and Pollock, 1994
Synercticinae Lawrence and Pollock, 1994: 36, in key [stem: Synerctic-]. Type genus: Synercticus Newman, 1842.
Family
Trictenotomidae
Blanchard, 1845
Tricténotomides Blanchard, 1845b: 137 [stem: Trictenotom-]. Type genus: Trictenotoma Gray, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by F. Smith (1851: 18, as Trictenotomidae), generally accepted as in Lawrence and Newton (1995: 897, as Trictenotomidae).
Family
Pythidae
Solier, 1834
Pythites Solier, 1834: 496 [stem: Pyth-]. Type genus: Pytho Latreille, 1797. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. G. Thomson (1859: 123, as Pythonidae [incorrect stem formation]), generally accepted as in Hansen (1996: 176, as Pythidae).
Enoptisidae Gistel, 1848: [11] [stem: Enopt-]. Type genus: Enoptes Gistel, 1848 [syn. of Pytho Latreille, 1797]. Comment: incorrect original stem formation, not in prevailing usage.
Ischyomiides Champion, 1886: 258 [stem: Ischyomi-]. Type genus: Ischyomius Chevrolat, 1878.
Osphyoplesiini Reitter, 1917: 59, in key [stem: Osphyoplesi-]. Type genus: Osphyoplesius A. Winkler, 1915.
Anaplopinae M. Abdullah, 1966: 147 [stem: Anaplopod-]. Type genus: Anaplopus Blackburn, 1890. Comment: incorrect original stem formation, not in prevailing usage.
Family
Pyrochroidae
Latreille, 1806
Pyrochroides Latreille, 1806: 199 [stem: Pyrochro-]. Type genus: Pyrochroa Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subfamily
Tydessinae
Nikitsky, 1986
Tydessini Nikitsky, 1986: 1188 [stem: Tydess-]. Type genus: Tydessa Peacock, 1982.
Subfamily
Pilipalpinae
Abdullah, 1964
Pilipalpini M. Abdullah, 1964: 4 [stem: Pilipalp-]. Type genus: Pilipalpus Fairmaire, 1876.
Techmessinae Paulus, 1972b: 84, in key [stem: Techmess-]. Type genus: Techmessa F. Bates, 1874.
Subfamily
Pedilinae
Lacordaire, 1859
Pédilides Lacordaire, 1859: 574 [stem: Pedil-]. Type genus: Pedilus Fischer von Waldheim, 1820. Comment: published before 27 June 1859; original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1859a [December]: 46, as Pedilidae), generally accepted as in Lawrence and Newton (1995: 898, as Pedilinae).
Neopedilini M. Abdullah, 1969a: 354 [stem: Neopedil-]. Type genus: Neopedilus M. Abdullah, 1969 [syn. of Pedilus Fischer von Waldheim, 1820].
Subfamily
Pyrochroinae
Latreille, 1806
Pyrochroides Latreille, 1806: 199 [stem: Pyrochro-]. Type genus: Pyrochroa Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Anthomanisidae Gistel, 1848: [11] [stem: Anthoman-]. Type genus: Anthomanes Gistel, 1848 [syn. of Pyrochroa Geoffroy, 1762]. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Agnathinae
Lacordaire, 1859
Agnathides Lacordaire, 1859: 531 [stem: Agnath-]. Type genus: Agnathus Germar, 1818. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1875 [Gatt.]: 100, as Agnathini), generally accepted as in Young (2002: 542, as Agnathinae).
Cononotini J. L. LeConte, 1862: 256 [stem: Cononot-]. Type genus: Cononotus J. L. LeConte, 1851.
Family
Salpingidae
Leach, 1815
Salpingides Leach, 1815: 106 [stem: Salping-]. Type genus: Salpingus Illiger, 1802.
Subfamily
Othniinae
LeConte, 1861
Othniidae J. L. LeConte, 1861: 102 [stem: Othni-]. Type genus: Othnius J. L. LeConte, 1861 [syn. of Elacatis Pascoe, 1860].
Elacatidae Cockerell, 1906: 242 [stem: Elacat-]. Type genus: Elacatis Pascoe, 1860. Comment: junior homonym of Elacatinae Gill, 1861 (type genus Elacate Cuvier, 1831) proposed in Pisces; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Subfamily
Prostominiinae
Grouvelle, 1914
Prostominini Grouvelle, 1914: 152 [stem: Prostomini-]. Type genus: Prostominia Reitter, 1889. Comment: incorrect original stem formation, not in prevailing usage.
*Trogocryptinae Crowson, 1953: 51 [stem: Trogocrypt-]. Type genus: Trogocryptus Sharp, 1900. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); this name has been used subsequently, e.g., Lawrence (1977: 43, 1980: 307), Lawrence and Newton (1995: 900), but it has not been made available.
Subfamily
Agleninae
Horn, 1878
Agleni G. H. Horn, 1878: 573 [stem: Aglen-]. Type genus: Aglenus Erichson, 1845.
Subfamily
Inopeplinae
Grouvelle, 1908
Inopeplini Grouvelle, 1908: 459, in key [stem: Inopepl-]. Type genus: Inopeplus F. Smith, 1851.
Subfamily
Dacoderinae
LeConte, 1862
Dacoderini J. L. LeConte, 1862: 216 [stem: Dacoder-]. Type genus: Dacoderus J. L. LeConte, 1858.
Tretothoracidae Lea, 1910: 210 [stem: Tretothorac-]. Type genus: Tretothorax Lea, 1910.
Subfamily
Aegialitinae
LeConte, 1862
Aegialitidae J. L. LeConte, 1862: 241 [stem: Aegialit-]. Type genus: Aegialites Mannerheim, 1853.
Eurystethidae Seidlitz, 1916: 127 [stem: Eurysteth-]. Type genus: Eurystethes Seidlitz, 1916 [syn. of Aegialites Mannerheim, 1853].
Subfamily
Salpinginae
Leach, 1815
Salpingides Leach, 1815: 106 [stem: Salping-]. Type genus: Salpingus Illiger, 1802.
*Rhinosimites Solier, 1834: 496 [stem: Rhinosim-]. Type genus: Rhinosimus Latreille, 1802. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Solier (1834).
Rhinosimidae Hope, 1840a: 134 [stem: Rhinosim-]. Type genus: Rhinosimus Latreille, 1802.
Lissodemina Seidlitz, 1917b: 422 [stem: Lissodem-]. Type genus: Lissodema Curtis, 1833.
Istrisiini Nikitsky, 1992: 483, in key [stem: Istrisi-]. Type genus: Istrisia Lewis, 1895.
Family
Anthicidae
Latreille, 1819
Anthicites Latreille, 1819: 363 [stem: Anthic-]. Type genus: Anthicus Paykull, 1798.
Subfamily
Eurygeniinae
LeConte, 1862
Eurygenii J. L. LeConte, 1862: 264 [stem: Eurygeni-]. Type genus: Eurygenius LaFerté-Sénectère, 1849.
Tribe
Eurygeniini
LeConte, 1862
Eurygenii J. L. LeConte, 1862: 264 [stem: Eurygeni-]. Type genus: Eurygenius LaFerté-Sénectère, 1849.
Tribe
Ictistygnini
Borchmann, 1936
Ictistygninae Borchmann, 1936: 534 [stem: Ictistygn-]. Type genus: Ictistygna Pascoe, 1866.
Tribe
Mitraelabrini
Abdullah, 1969
Mitraelabrini M. Abdullah, 1969a: 350 [stem: Mitraelabr-]. Type genus: Mitraelabrus Solier, 1851.
Subfamily
Macratriinae
LeConte, 1862
Macratriini J. L. LeConte, 1862: 265 [stem: Macratri-]. Type genus: Macratria Newman, 1838.
Tribe
Macratriini
LeConte, 1862
Macratriini J. L. LeConte, 1862: 265 [stem: Macratri-]. Type genus: Macratria Newman, 1838.
†Tribe
Camelomorphini
Kirejtshuk and Azar, 2008
Camelomorphini Kirejtshuk and Azar, 2008: 40 [stem: Camelomorph-]. Type genus: Camelomorpha Kirejtshuk and Azar, 2008.
Subfamily
Steropinae
Jacquelin du Val, 1863
Stéropites Jacquelin du Val, 1863: 365 [stem: Sterop-]. Type genus: Steropes Steven, 1806. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jakobson (1915: 1022, as Steropina), generally accepted as in Lawrence and Newton (1995: 901, as Steropinae); the older name Steropinae Dana, 1854 is available in copepods (type genus Sterope Goodsir, 1845) although it was recently treated as a nomen oblitum by Huys (2009: 28); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Subfamily
Copobaeninae
Abdullah, 1969
Copobaeninae M. Abdullah, 1969a: 334 [stem: Copobaen-]. Type genus: Copobaenus Fairmaire and Germain, 1863.
Subfamily
Lemodinae
Lawrence and Britton, 1991
*Lemodinae Lawrence, 1977: 43 [stem: Lemod-]. Type genus: Lemodes Boheman, 1858. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Lemodinae Lawrence and Britton, 1991: 603, in key [stem: Lemod-]. Type genus: Lemodes Boheman, 1858.
Subfamily
Tomoderinae
Bonadona, 1961
Tomoderini Bonadona, 1961: 11 [stem: Tomoder-]. Type genus: Tomoderus LaFerté-Sénectère, 1849.
Subfamily
Anthicinae
Latreille, 1819
Anthicites Latreille, 1819: 363 [stem: Anthic-]. Type genus: Anthicus Paykull, 1798.
Tribe
Anthicini
Latreille, 1819
Anthicites Latreille, 1819: 363 [stem: Anthic-]. Type genus: Anthicus Paykull, 1798.
Tribe
Endomiini
Kaszab, 1956
Endomiini Kaszab, 1956: 45, in key [stem: Endomi-]. Type genus: Endomia Laporte, 1840.
Tribe
Formicomini
Bonadona, 1974
Formicomini Bonadona, 1974: 110, in key [stem: Formicom-]. Type genus: Formicomus LaFerté-Sénectère, 1849.
Tribe
Microhoriini
Bonadona, 1974
Microhorini Bonadona, 1974: 110, in key [stem: Microhori-]. Type genus: Microhoria Chevrolat, 1877. Comment: incorrect original stem formation, not in prevailing usage.
Liparoderini Bonadona, 1990: 20 [stem: Liparoder-]. Type genus: Liparoderus LaFerté-Sénectère, 1849.
Subfamily
Notoxinae
Stephens, 1829
Notoxidae Stephens, 1829b: 254 [stem: Notox-]. Type genus: Notoxus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: an application needs to be submitted to the Commission to suppress Notoxii Sturm, 1826 (based on the misidentified type genus Notoxus sensu Fabricius, 1775) for the Principles of Priority and Homonymy (Art. 65.2.1) to conserve this name as valid.
Family
Aderidae
Csiki, 1909
Aderidae Csiki, 1909a: 6 [stem: Ader-]. Type genus: Aderus Stephens, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1989e)]. Comment: name given precedence over Euglenesidae Seidlitz, 1875 (ICZN 1989e).
†Circaeidae Iablokoff-Khnzorian, 1961: 209 [stem: Circae-]. Type genus: Circaeus Iablokoff-Khnzorian, 1961.
Tribe
Aderini
Csiki, 1909
Aderidae Csiki, 1909a: 6 [stem: Ader-]. Type genus: Aderus Stephens, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1989e)]. Comment: name given precedence over Euglenesidae Seidlitz, 1875 and placed on the Official List of Family-Group Names in Zoology (ICZN 1989e, as Aderidae A. Winkler (1927)).
Subtribe
Aderina
Csiki, 1909
Xylophilidae Shuckard, 1839b: 47 [stem: Xylophil-]. Type genus: Xylophilus Latreille, 1825 [preoccupied genus name, not Xylophilus Mannerheim, 1823 [Coleoptera: Eucnemidae]; syn. of Aderus Stephen, 1829]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Hylophilidae Pic, 1900: 754 [stem: Hylophil-]. Type genus: Hylophilus Berthold, 1827 [preoccupied genus name, not Hylophilus Temminck, 1822 [Aves]; syn. of Aderus Stephens, 1829]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Aderidae Csiki, 1909a: 6 [stem: Ader-]. Type genus: Aderus Stephens, 1829 [placed on the Official List of Generic Names in Zoology (ICZN 1989e)]. Comment: placed on the Official List of Family-Group Names in Zoology (ICZN 1989e, as Aderidae A. Winkler (1927)).
Subtribe
Cnopina
Báguena Corella, 1948
Cnopina Báguena Corella, 1948: 57 [stem: Cnop-]. Type genus: Cnopus Champion, 1893.
Subtribe
Gompeliina
Bouchard, 2011
Olotelina Báguena Corella, 1948: 58 [stem: Olotel-]. Type genus: Olotelus Mulsant and Rey, 1866 [preoccupied genus name, not Olotelus Solier, 1851 [Coleoptera: Elateridae]; syn. of Gompelia Alonso-Zarazaga, 2010]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Gompeliina Bouchard, 2011 nom. nov. [stem: Gompeli-]. Type genus: Gompelia Alonso-Zarazaga, 2010. Comment: replacement name for Olotelina Báguena Corella, 1948 because of the homonymy of the type genus.
Subtribe
Syzetoninina
Báguena Corella, 1948
Syzetoninini Báguena Corella, 1948: 41 [stem: Syzetonin-]. Type genus: Syzetoninus Blackburn, 1891.
Tribe
Emelinini
Báguena Corella, 1948
Emelinini Báguena Corella, 1948: 36 [stem: Emelin-]. Type genus: Emelinus Casey, 1895.
Tribe
Euglenesini
Seidlitz, 1875
Euglenini Seidlitz, 1875 [Gatt.]: 106 [stem: Euglenes-]. Type genus: Euglenes Westwood, 1830 [placed on the Official List of Generic Names in Zoology (ICZN 1989e)].
Subtribe
Euglenesina
Seidlitz, 1875
Euglenini Seidlitz, 1875 [Gatt.]: 106 [stem: Euglenes-]. Type genus: Euglenes Westwood, 1830 [placed on the Official List of Generic Names in Zoology (ICZN 1989e)]. Comment: the younger name Aderidae Csiki, 1909 (listed as Aderidae A. Winkler, 1927) was given precedence for the family name, Euglenes- was ruled to be the correct stem of this family-group name and Euglenesidae Seidlitz, 1875 was placed on the Official List of Family-Group Names in Zoology (ICZN 1989e); Euglenidae Seidlitz, 1875 was placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1989e).
Subtribe
Pseudolotelina
Báguena Corella, 1948
Pseudolotelina Báguena Corella, 1948: 40 [stem: Pseudolotel-]. Type genus: Pseudolotelus Pic, 1901.
Tribe
Phytobaenini
Báguena Corella, 1948
Phytobaenini Báguena Corella, 1948: 30 [stem: Phytobaen-]. Type genus: Phytobaenus Sahlberg, 1834.
Family
Scraptiidae
Gistel, 1848
Scraptiaeidae Gistel, 1848: [11] [stem: Scrapti-]. Type genus: Scraptia Latreille, 1807.
Subfamily
Scraptiinae
Gistel, 1848
Scraptiaeidae Gistel, 1848: [11] [stem: Scrapti-]. Type genus: Scraptia Latreille, 1807. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Allopodini
Franciscolo, 1964
Allopodini Franciscolo, 1964: 176 [stem: Allopod-]. Type genus: Allopoda J. L. LeConte, 1866.
Tribe
Scraptiini
Gistel, 1848
Scraptiaeidae Gistel, 1848: [11] [stem: Scrapti-]. Type genus: Scraptia Latreille, 1807. Comment: family-group name previously attributed to Mulsant (1856)/Gistel (1856); incorrect original stem formation, not in prevailing usage.
Subfamily
Anaspidinae
Mulsant, 1856
Anaspiens Mulsant, 1856c: 85 [stem: Anaspid-]. Type genus: Anaspis Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)].
Tribe
Anaspidini
Mulsant, 1856
Anaspiens Mulsant, 1856c: 85 [stem: Anaspid-]. Type genus: Anaspis Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. G. Thomson (1859: 121, as Anaspina [incorrect stem formation]), generally accepted as in Lawrence and Newton (1995: 903, as Anaspidinae); incorrect original stem formation, not in prevailing usage.
Tribe
Anaspimordini
Franciscolo, 1954
Anaspimordini Franciscolo, 1954: 65, in key [stem: Anaspimord-]. Type genus: Anaspimorda Ermisch, 1950.
Tribe
Menuthianaspidini
Franciscolo, 1972
*Cyrtoscraptiini Franciscolo, 1964: 176 [stem: Cyrtoscrapti-]. Type genus: Cyrtoscraptia Franciscolo, 1964 [unavailable genus name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); syn. of Menuthianaspis Franciscolo, 1972]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), based on unavailable type genus (Art. 12.2.4).
Menuthianaspidini Franciscolo, 1972: 123 [stem: Menuthianaspid-]. Type genus: Menuthianaspis Franciscolo, 1972.
Tribe
Pentariini
Franciscolo, 1954
Pentariini Franciscolo, 1954: 64, in key [stem: Pentari-]. Type genus: Pentaria Mulsant, 1856.
Subfamily
Lagrioidinae
Abdullah and Abdullah, 1968
Lagrioidini M. Abdullah and A. Abdullah, 1968: 73 [stem: Lagrioid-]. Type genus: Lagrioida Fairmaire and Germain, 1860.
Subfamily
Afreminae
Levey, 1985
Afreminae Levey, 1985: 420 [stem: Afrem-]. Type genus: Afremus Levey, 1985.
Subfamily
Ischaliinae
Blair, 1920
Ischaliinae Blair, 1920: 134 [stem: Ischali-]. Type genus: Ischalia Pascoe, 1860.
Superfamily
Chrysomeloidea
Latreille, 1802
Chrysomelinae Latreille, 1802: 220 [stem: Chrysomel-]. Type genus: Chrysomela Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1984c)]. Comment: Švácha and Danilevsky (1987: 17) recognized Vesperinae, Oxypeltinae and Disteniinae at the family level, and later Švácha et al. (1997: 361) placed Anoplodermatinae and Philinae as subfamilies of Vesperidae; although the classification is based primarily on larval characters, it is also supported by adult features, as pointed out by Švácha et al. (1997), as well as by some earlier workers, e.g., Gahan (1906), Linsley (1961), Crowson (1981), Saito (1990); this classification will also be used in the upcoming volume 3 of the “Handbook of Zoology: Coleoptera, beetles” series; First Reviser (Chrysomeloidea Latreille, 1802 vs Megalopodoidea Latreille, 1802 vs Cerambycoidea Latreille, 1802) not determined, current usage maintained.
Family
Oxypeltidae
Lacordaire, 1868
Oxypeltides Lacordaire, 1868: 461 [stem: Oxypelt-]. Type genus: Oxypeltus Blanchard, 1851. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 254, as Oxypeltini).
*Cheloderidos Germain, 1900: 86 [stem: Cheloder-]. Type genus: Cheloderus Gray, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Family
Vesperidae
Mulsant, 1839
Vespéraires Mulsant, 1839: 214 [stem: Vesper-]. Type genus: Vesperus Dejean, 1821.
Subfamily
Philinae
Thomson, 1861
Philitae J. Thomson, 1861: 297 [stem: Phil-]. Type genus: Philus Saunders, 1853.
Subfamily
Vesperinae
Mulsant, 1839
Vespéraires Mulsant, 1839: 214 [stem: Vesper-]. Type genus: Vesperus Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 240, as Vesperini), generally accepted as in Villiers (1978: 67, as Vesperini).
Subfamily
Anoplodermatinae
Guérin-Méneville, 1840
Anoplodermiens Guérin-Méneville, 1840: 276 [stem: Anoplodermat-]. Type genus: Anoploderma Guérin-Méneville, 1840.
Tribe
Anoplodermatini
Guérin-Méneville, 1840
Anoplodermiens Guérin-Méneville, 1840: 276 [stem: Anoplodermat-]. Type genus: Anoploderma Guérin-Méneville, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1861: 277, as Anoplodermitae [incorrect stem formation]), generally accepted as in Monné (1994c: 9, as Anoplodermatinae); incorrect original stem formation, not in prevailing usage.
Cherrocriinae Prosen, 1960: 90 [stem: Cherrocri-]. Type genus: Cherrocrius Berg, 1898.
Tribe
Hypocephalini
Blanchard, 1845
Hypocéphaliens Blanchard, 1845b: 135 [stem: Hypocephal-]. Type genus: Hypocephalus Desmarest, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Imhoff (1856 [2]: 170, as Hypocephalidae), generally accepted as in Lameere (1913: 94, as Hypocephali).
Tribe
Mysteriini
Prosen, 1960
Mysterinae Prosen, 1960: 90 [stem: Mysteri-]. Type genus: Mysteria J. Thomson, 1860. Comment: incorrect original stem formation, not in prevailing usage.
Family
Disteniidae
Thomson, 1861
Distenitae J. Thomson, 1861: 181 [stem: Disteni-]. Type genus: Distenia Lepeletier and Audinet-Serville, 1828.
Tribe
Cyrtonopini
Gressitt, 1940
Cyrtonopini Gressitt, 1940: 27 [stem: Cyrtonop-]. Type genus: Cyrtonops A. White, 1853.
Tribe
Disteniini
Thomson, 1861
*Cométites Blanchard, 1845b: 163 [stem: Comet-]. Type genus: Cometes Lepeletier and Audinet-Serville, 1828. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; Monné and Santos-Silva (2008: 261) considered this name a nomen oblitum, this action was unnecessary since Cométites Blanchard is not an available name.
Distenitae J. Thomson, 1861: 181 [stem: Disteni-]. Type genus: Distenia Lepeletier and Audinet-Serville, 1828. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Dynamostini
Lacordaire, 1868
Dynamostides Lacordaire, 1868: 196 [stem: Dynamost-]. Type genus: Dynamostes Pascoe, 1857. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 238, as Dynamostini), generally accepted as in Santos-Silva and Martins (2004: 145, as Dynamostini).
Tribe
Heteropalpini
Villiers, 1961
Heteropalpini Villiers, 1961: 385, in key [stem: Heteropalp-]. Type genus: Heteropalpus Buquet, 1843.
Family
Cerambycidae
Latreille, 1802
Cerambicini Latreille, 1802: 211 [stem: Cerambyc-]. Type genus: Cerambyx Linnaeus, 1758. Comment: First Reviser (Cerambycidae Latreille, 1802 vs Prionidae Latreille, 1802 vs Lepturidae Latreille, 1802) not determined, current usage maintained.
Subfamily
Parandrinae
Blanchard, 1845
Parandrides Blanchard, 1845b: 134 [stem: Parandr-]. Type genus: Parandra Latreille, 1802.
Tribe
Erichsoniini
Thomson, 1861
Erichsonitae J. Thomson, 1861: 274 [stem: Erichsoni-]. Type genus: Erichsonia Westwood, 1849. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Parandrini
Blanchard, 1845
Parandrides Blanchard, 1845b: 134 [stem: Parandr-]. Type genus: Parandra Latreille, 1802. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1848: [8], as Parandraeidae [incorrect stem formation]), generally accepted as in Lameere (1913: 4, as Parandrae).
Subfamily
Prioninae
Latreille, 1802
Prionii Latreille, 1802: 212 [stem: Prion-]. Type genus: Prionus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Acanthophorini
Thomson, 1864
Acanthophoritae J. Thomson, 1864: 289 [stem: Acanthophor-]. Type genus: Acanthophorus Audinet-Serville, 1832.
Tribe
Aegosomatini
Thomson, 1861
Aegosomitae J. Thomson, 1861: 308 [stem: Aegosomat-]. Type genus: Aegosoma Audinet-Serville, 1832. Comment: incorrect original stem formation, not in prevailing usage.
*Catypnides Lacordaire, 1868: 62 [stem: Catypn-]. Type genus: Catypnes Pascoe, 1864. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Jamwoninae Kolbe, 1897: 294 [stem: Jamwon-]. Type genus: Jamwonus Harold, 1879.
Megopides Lameere, 1912: 181 [stem: Megopid-]. Type genus: Megopis Audinet-Serville, 1832. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Anacolini
Thomson, 1857
Anacolites J. Thomson, 1857b: 10 [stem: Anacol-]. Type genus: Anacolus Berthold, 1827. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1861: 302, as Anacolitae), generally accepted as in Galileo (1987: 482, as Anacolini).
*Poecilosomides Lacordaire, 1868: 185 [stem: Poekilosomat-]. Type genus: Poekilosoma Audinet-Serville, 1832 [as Poecilosoma, unjustified emendation of type genus name by Agassiz (1846b: 301), not in prevailing usage; unjustified emendation also junior homonym of Poecilosoma Hübner, 1819]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1868); incorrect original stem formation, not in prevailing usage.
Poecilosomi H. W. Bates, 1869: 49 [stem: Poekilosomat-]. Type genus: Poekilosoma Audinet-Serville, 1832 [as Poecilosoma, unjustified emendation of type genus name by Agassiz (1846b: 301), not in prevailing usage; unjustified emendation also junior homonym of Poecilosoma Hübner, 1819]. Comment: incorrect original stem formation, not in prevailing usage.
Erythraeninae H. W. Bates, 1875: 52 [stem: Erythraen-]. Type genus: Erythraenus H. W. Bates, 1875.
*Sobarines Lameere, 1901: 320 [stem: Sobar-]. Type genus: Sobarus Harold, 1879 [preoccupied genus name, not Sobarus Loew, 1855 [Diptera]; a replacement name is needed]. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
*Délochiliens Lameere, 1912: 57 [stem: Delocheil-]. Type genus: Delocheilus J. Thomson, 1860 [as Delochilus, unjustified emendation of type genus name by Gemminger (1872: 2777), not in prevailing usage]. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Sobari Lameere, 1913: 85 [stem: Sobar-]. Type genus: Sobarus Harold, 1879 [preoccupied genus name, not Sobarus Loew, 1855 [Diptera]; a replacement name is needed]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Delochili Lameere, 1913: 85 [stem: Delocheil-]. Type genus: Delocheilus J. Thomson, 1860 [as Delochilus, unjustified emendation of type genus name by Gemminger and Harold (1872), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Cacoscelini
Thomson, 1861
*Notophysites Blanchard, 1845b: 138 [stem: Notophyse-]. Type genus: Notophysis Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845b); incorrect original stem formation, not in prevailing usage.
Cacoscelitae J. Thomson, 1861: 325 [stem: Cacoscel-]. Type genus: Cacosceles Newman, 1838.
Colpodérides Lacordaire, 1868: 133 [stem: Colpoder-]. Type genus: Colpoderus Audinet-Serville, 1832 [syn. of Notophysis Audinet-Serville, 1832]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Pascoe (1869b: 673, as Colpoderinae); this name was incorrectly credited to Pascoe, 1869 by Bousquet et al. (2009: 115).
Nothophysini Lameere, 1903a: 4, in key [stem: Notophyse-]. Type genus: Notophysis Audinet-Serville, 1832 [as Nothophysis, unjustified emendation of type genus name by Scudder (1882: 226), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Callipogonini
Thomson, 1861
Callipogonitae J. Thomson, 1861: 323 [stem: Callipogon-]. Type genus: Callipogon Audinet-Serville, 1832.
Anacanthitae J. Thomson, 1864: 285 [stem: Anacanth-]. Type genus: Anacanthus Audinet-Serville, 1832 [preoccupied genus name, not Anacanthus Gray, 1831 [Pisces]; syn. of Chorenta Gistel, 1848]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Enoploceritae J. Thomson, 1864: 290 [stem: Enoplocer-]. Type genus: Enoplocerus Audinet-Serville, 1832.
Orthomegitae J. Thomson, 1864: 294 [stem: Orthomegal-]. Type genus: Orthomegas Audinet-Serville, 1832. Comment: incorrect original stem formation, not in prevailing usage.
Ctenoscelitae J. Thomson, 1864: 295 [stem: Ctenoscelid-]. Type genus: Ctenoscelis Audinet-Serville, 1832. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Calocomini
Galileo and Martins, 1993
Calocomini Galileo and Martins, 1993: 81 [stem: Calocom-]. Type genus: Calocomus Audinet-Serville, 1832.
Tribe
Cantharocnemini
Thomson, 1861
Cantharocnemitae J. Thomson, 1861: 274 [stem: Cantharocnem-]. Type genus: Cantharocnemis Audinet-Serville, 1832. Comment: current spelling maintained (Art. 29.3.1.1): incorrect original stem formation in prevailing usage (should be Cantharocnemid-).
*Scéléocanthides Lacordaire, 1868: 34 [stem: Sceleocanth-]. Type genus: Sceleocantha Newman, 1840. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 236, as Sceleocanthini), but not generally accepted as valid.
Tribe
Ergatini
Fairmaire, 1864
Ergatites Fairmaire, 1864: 117 [stem: Ergat-]. Type genus: Ergates Audinet-Serville, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1873: 286, as Ergatini), generally accepted as in Linsley (1962: 24, as Ergatini).
Tribe
Eurypodini
Gahan, 1906 (1868)
Zaracides Lacordaire, 1868: 131 [stem: Zarac-]. Type genus: Zarax Pascoe, 1867 [syn. of Eurypoda W. Saunders, 1853]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Pascoe (1869b: 672, as Zaracinae); usage of the younger name Eurypodini Gahan, 1906 conserved over this name (Art. 40.2).
Eurypodini Gahan, 1906: 27 [stem: Eurypod-]. Type genus: Eurypoda W. Saunders, 1853. Comment: name proposed to replace Zaracini Lacordaire, 1868 because of the synonymy of the type genus; use of family-group name conserved over Zaracini Lacordaire, 1868 (Art. 40.2); also see comments under Eurypinae Thomson, 1860 in Mycteridae.
Tribe
Hopliderini
Thomson, 1864
Hoplideritae J. Thomson, 1864: 290 [stem: Hoplider-]. Type genus: Hoplideres Audinet-Serville, 1832.
Tribe
Macrodontiini
Thomson, 1861
Macrodontitae J. Thomson, 1861: 324 [stem: Macrodonti-]. Type genus: Macrodontia Lacordaire, 1830. Comment: incorrect original stem formation, not in prevailing usage.
Acanthinoderitae J. Thomson, 1864: 294 [stem: Acanthinoder-]. Type genus: Acanthinodera Hope, 1833.
*Ancistrotides Lacordaire, 1868: 81 [stem: Ancistrot-]. Type genus: Ancistrotus Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1868).
*Acalodegmites J. Thomson, 1877: 261 [stem: Acalodegm-]. Type genus: Acalodegma Thomson, 1877. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Ancistrotini Lameere, 1919: 90 [stem: Ancistrot-]. Type genus: Ancistrotus Audinet-Serville, 1832.
Tribe
Macrotomini
Thomson, 1861
Macrotomitae J. Thomson, 1861: 312 [stem: Macrotom-]. Type genus: Macrotoma Audinet-Serville, 1832.
Subtribe
Archetypina
Lameere, 1912
Archetypi Lameere, 1912: 180 [stem: Archetyp-]. Type genus: Archetypus J. Thomson, 1861.
Subtribe
Basitoxina
Lameere, 1912
*Mécosarthrines Lameere, 1903b: 307 [stem: Mecosarthr-]. Type genus: Mecosarthron Buquet, 1840. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Basitoxi Lameere, 1912: 180 [stem: Basitox-]. Type genus: Basitoxus Audinet-Serville, 1832.
Mecosarthrini Melzer, 1919: 35 [stem: Mecosarthr-]. Type genus: Mecosarthron Buquet, 1840.
Subtribe
Macrotomina
Thomson, 1861
Macrotomitae J. Thomson, 1861: 312 [stem: Macrotom-]. Type genus: Macrotoma Audinet-Serville, 1832 [Macrotoma Audinet-Serville, 1832 [July] is a junior homonym of Macrotoma Laporte, 1832 [April], which is a synonym of Longina Wiedemann, 1830 in Diptera; Heffern et al. (2006) applied the Reversal of Precedence (Art. 23.9) to qualify Macrotoma Audinet-Serville as nomen protectum].
*Aulacopides Lacordaire, 1868: 101 [stem: Aulacopod-]. Type genus: Aulacopus Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1868); incorrect original stem formation, not in prevailing usage.
Aulacopinae Kolbe, 1897: 295 [stem: Aulacopod-]. Type genus: Aulacopus Audinet-Serville, 1832. Comment: incorrect original stem formation, not in prevailing usage.
*Cnémoplitiens Lameere, 1904: 1 [stem: Cnemoplit-]. Type genus: Cnemoplites Newman, 1842. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Cnemoplitinae Schröder, 1905: vii [stem: Cnemoplit-]. Type genus: Cnemoplites Newman, 1842. Comment: see Bousquet et al. (2009: 17) for comments about the authorship of this name.
Prinobiini Vives, 2000: 84 [stem: Prinobi-]. Type genus: Prinobius Mulsant, 1842. Comment: replacement name for Macrotomini Thomson, 1861 because of the homonymy of the type genus.
Subtribe
Platygnathina
Gilmour, 1954
Platygnathina Gilmour, 1954: 33 [stem: Platygnath-]. Type genus: Platygnathus Audinet-Serville, 1832. Comment: as pointed out by Bousquet et al. (2009: 18) Gilmour (1954: 33) did not provide a description of his new taxon but his new name is available because it was used as valid before 2000, e.g., Ferreira and Veiga Ferreira (1959a: 34, as "Platygnathina Gilmour, 1954") and was not rejected by an author who, after 1960 and before 2000, expressly applied Article 13 of the then current editions of the Code (Article 13.2.1).
Platygnathini Quentin and Villiers, 1975: 25 [stem: Platygnath-]. Type genus: Platygnathus Audinet-Serville, 1832. Comment: family-group name proposed as new without reference to Platygnathina Gilmour, 1954.
Subtribe
Xixuthrina
Lameere, 1912
Xixuthri Lameere, 1912: 181 [stem: Xixuthr-]. Type genus: Xixuthrus J. Thomson, 1864.
Tribe
Mallaspini
Thomson, 1861
Mallaspitae J. Thomson, 1861: 302 [stem: Mallasp-]. Type genus: Mallaspis Audinet-Serville, 1832. Comment: current spelling maintained (Art. 29.3.1.1): incorrect original stem formation in prevailing usage (should be Mallaspid-).
*Pyrodides Lacordaire, 1868: 174 [stem: Pyrod-]. Type genus: Pyrodes Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1868).
Pyrodini Harold, 1879: 165 [stem: Pyrod-]. Type genus: Pyrodes Audinet-Serville, 1832.
Tribe
Mallodonini
Thomson, 1861
Mallodonitae J. Thomson, 1861: 318 [stem: Mallodon-]. Type genus: Mallodon Lacordaire, 1830. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Mallodont-); we treat this group as a valid tribe as in Drumont and Komiya (2010: 91).
*Sténodontines Lameere, 1902: 66 [stem: Stenodont-]. Type genus: Stenodontes Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Stenodontini Lameere, 1903a: 54 [stem: Stenodont-]. Type genus: Stenodontes Audinet-Serville, 1832. Comment: homonym of Stenodontina Schmiedeknecht, 1903 (type genus Stenodontus Berthoumieu, 1897) in Hymenoptera; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Meroscelisini
Thomson, 1861
Meroscelisitae J. Thomson, 1861: 299 [stem: Meroscelis-]. Type genus: Meroscelisus Audinet-Serville, 1832.
*Tragosomites Fairmaire, 1864: 119 [stem: Tragosomat-]. Type genus: Tragosoma Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Fairmaire (1864); incorrect original stem formation, not in prevailing usage.
Tragosomitae J. Thomson, 1864: 286 [stem: Tragosomat-]. Type genus: Tragosoma Audinet-Serville, 1832. Comment: incorrect original stem formation, not in prevailing usage.
Clostérides Lacordaire, 1868: 149 [stem: Closter-]. Type genus: Closterus Audinet-Serville, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 676, as Closterinae), generally accepted as in Lameere (1913: 81, as Closteri).
*Monodesmides Lacordaire, 1868: 157 [stem: Monodesm-]. Type genus: Monodesmus Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1868).
Monodesminae Gahan, 1890: 299 [stem: Monodesm-]. Type genus: Monodesmus Audinet-Serville, 1832.
Luluina Gilmour, 1956: 222 [stem: Lulu-]. Type genus: Lulua Burgeon, 1931.
Tribe
Prionini
Latreille, 1802
Prionii Latreille, 1802: 212 [stem: Prion-]. Type genus: Prionus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
*Prioceria Rafinesque, 1815: 116 [stem: Priocerat-]. Type genus: Prioceras Rafinesque, 1815 [as pointed out by Bousquet et al. (2009: 19) Rafinesque (1815: 116) listed the genus “Prioceras R. sp. do.” following the genus “Prionus Fabr.” in his subfamily Prioceria, the abbreviations after the genus name Prioceras mean that this is a new genus by the author “R[afinesque].” and that the new genus includes some species that were included in Prionus previously “sp. do.”; Rafinesque did not list which species he included in his new genus Prioceras and we are not aware of any subsequent validation of this name; Prioceras Rafinesque is also listed as a nomen nudum in Neave (1940: 889) and Sherborn (1929: 5148)]. Comment: family-group name unavailable (Art. 11.7.1.1): not based on an available genus name at the time; incorrect original stem formation, not in prevailing usage.
*Cyrtognathites Blanchard, 1845b: 138 [stem: Cyrtognath-]. Type genus: Cyrtognathus Faldermann, 1835 [subgenus of Dorysthenes Vigors, 1826]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845b).
*Psalidognathites Blanchard, 1845b: 138 [stem: Psalidognath-]. Type genus: Psalidognathus Gray, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845b).
Cyrthognathitae J. Thomson, 1861: 328 [stem: Cyrtognath-]. Type genus: Cyrtognathus Faldermann, 1835 [as Cyrthognathus, incorrect subsequent spelling of type genus name, not in prevailing usage; subgenus of Dorysthenes Vigors, 1826]. Comment: incorrect original stem formation, not in prevailing usage.
Psalidognathitae J. Thomson, 1861: 331 [stem: Psalidognath-]. Type genus: Psalidognathus Gray, 1832.
Prionommitae J. Thomson, 1861: 327 [stem: Prionommat-]. Type genus: Prionomma A. White, 1853. Comment: incorrect original stem formation, not in prevailing usage.
Orthosomitae J. Thomson, 1864: 284 [stem: Orthosomat-]. Type genus: Orthosoma Audinet-Serville, 1832. Comment: incorrect original stem formation, not in prevailing usage.
Pithoclitae J. Thomson, 1864: 291 [stem: Pithocl-]. Type genus: Pithocles J. Thomson, 1864 [syn. of Derobrachus Audinet-Serville, 1832].
Derobrachitae J. Thomson, 1864: 291 [stem: Derobrach-]. Type genus: Derobrachus Audinet-Serville, 1832.
Titanitae J. Thomson, 1864: 292 [stem: Titan-]. Type genus: Titanus Audinet-Serville, 1832.
Aulacoceritae J. Thomson, 1864: 292 [stem: Aulacocer-]. Type genus: Aulacocerus A. White, 1853.
*Psalidocoptides Lacordaire, 1868: 38 [stem: Psalidocopt-]. Type genus: Psalidocoptus A. White, 1856. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Polyarthrides Lacordaire, 1868: 44 [stem: Polyarthr-]. Type genus: Polyarthron Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1868).
*Micropsalides Lacordaire, 1868: 42 [stem: Micropsalid-]. Type genus: Micropsalis Burmeister, 1865 [preoccupied genus name, not Micropsalis Meyer, 1859 [Crustacea]; syn. of Apterocaulus Fairmaire, 1864]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1868); incorrect original stem formation, not in prevailing usage.
Polyarthrini Gounelle, 1911: 326 [stem: Polyarthr-]. Type genus: Polyarthron Audinet-Serville, 1832. Comment: Polyarthridae Daday, 1893 (type genus Polyarthra Ehrenberg, 1834) is available in Rotifera; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Micropsalini Gounelle, 1911: 326 [stem: Micropsalid-]. Type genus: Micropsalis Burmeister, 1865 [preoccupied genus name, not Micropsalis Meyer, 1859 [Crustacea]; syn. of Apterocaulus Fairmaire, 1864]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Tribe
Remphanini
Lacordaire, 1868
Remphanides Lacordaire, 1868: 103 [stem: Remphan-]. Type genus: Remphan G. R. Waterhouse, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 667, as Remphaninae), generally accepted as in Drumont and Komiya (2010: 95, as Remphanini); this name was incorrectly credited to Pascoe, 1869 by Bousquet et al. (2009: 18); we treat this group as a valid tribe as in Drumont and Komiya (2010: 95).
Rhaphipodi Lameere, 1912: 181 [stem: Rhaphipod-]. Type genus: Rhaphipodus Audinet-Serville, 1832.
Tribe
Solenopterini
Lacordaire, 1868
Solénoptérides Lacordaire, 1868: 180 [stem: Solenopter-]. Type genus: Solenoptera Audinet-Serville, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1873: 286, as Solenopterini), generally accepted as in Monné (1995c: 37, as Solenopterini).
*Dérancistrines Lameere, 1909: 1 [stem: Derancistr-]. Type genus: Derancistrus Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Derancistrini Lameere, 1912: 181 [stem: Derancistr-]. Type genus: Derancistrus Audinet-Serville, 1832.
Tribe
Tereticini
Lameere, 1913
*Téréticiens Lameere, 1912: 72 [stem: Teretic-]. Type genus: Tereticus C. O. Waterhouse, 1879. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Teretici Lameere, 1913: 87 [stem: Teretic-]. Type genus: Tereticus C. O. Waterhouse, 1879.
Tribe
Vesperoctenini
Vives, 2005
Vesperoctenini Vives, 2005: 438 [stem: Vesperocten-]. Type genus: Vesperoctenus H. W. Bates, 1891.
Subfamily
Lepturinae
Latreille, 1802
Lepturetae Latreille, 1802: 218 [stem: Leptur-]. Type genus: Leptura Linnaeus, 1758.
Tribe
Desmocerini
Blanchard, 1845
Desmocérites Blanchard, 1845b: 163 [stem: Desmocer-]. Type genus: Desmocerus Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1861: 159, as Desmoceritae), generally accepted as in Monné (1995b: 1, as Desmocerini).
Tribe
Encyclopini
LeConte, 1873
Encyclopini J. L. LeConte, 1873: 326 [stem: Enclyclop-]. Type genus: Encyclops Newman, 1838.
Tribe
Lepturini
Latreille, 1802
Lepturetae Latreille, 1802: 218 [stem: Leptur-]. Type genus: Leptura Linnaeus, 1758.
*Grammoptérates Mulsant, 1863b: 569 [stem: Grammopter-]. Type genus: Grammoptera Audinet-Serville, 1835. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Strangalini Zagajkevich, 1991: 96 [stem: Strangali-]. Type genus: Strangalia Audinet-Serville, 1835. Comment: family-group name unavailable (Art. 11.6): originally published as synonym and not made available subsequently; incorrect original stem formation, not in prevailing usage.
Tribe
Oxymirini
Danilevsky, 1997
Oxymirini Danilevsky, 1997: 8 [stem: Oxymir-]. Type genus: Oxymirus Mulsant, 1862.
Tribe
Rhagiini
Kirby, 1837
Rhagiadae Kirby, 1837: 178 [stem: Rhagi-]. Type genus: Rhagium Fabricius, 1775.
*Toxotaires Mulsant, 1839: 230 [stem: Toxot-]. Type genus: Toxotus Dejean, 1821 [syn. of Stenocorus Geoffroy, 1762]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant (1839).
*Pachytes Motschulsky, 1849: 60 [stem: Pachyt-]. Type genus: Pachyta Dejean, 1821. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).
Stenocoritae J. Thomson, 1861: 156 [stem: Stenocor-]. Type genus: Stenocorus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: an application will need to be submitted to the Commission to suppress Stenocoridae Hope, 1834 (based on the misidentified type genus Stenocorus sensu Hope, 1834) for the Principles of Priority and Homonymy (Art. 65.2.1) if this name is to be used as valid.
Toxoti J. L. LeConte and G. H. Horn, 1883: 313 [stem: Toxot-]. Type genus: Toxotus Dejean, 1821 [syn. of Stenocorus Geoffroy, 1762]. Comment: junior homonym of Toxotidae Günther, 1860 (type genus Toxotes Cuvier and Cloquet, 1816) currently used as valid in Pisces; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Pachytini Portevin, 1934: 119, in key [stem: Pachyt-]. Type genus: Pachyta Dejean, 1821.
*Enoploderini Danilevsky, 1997: 9 [stem: Enoploder-]. Type genus: Enoploderes Faldermann, 1837. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Rhamnusiini
Sama, 2009
*Rhamnusiini Danilevsky, 1997: 9 [stem: Rhamnusi-]. Type genus: Rhamnusium Latreille, 1829. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Rhamnusiini Sama, 2009b: 383 [stem: Rhamnusi-]. Type genus: Rhamnusium Latreille, 1829.
Tribe
Teledapini
Pascoe, 1871
Teledapinae Pascoe, 1871: 268 [stem: Teledap-]. Type genus: Teledapus Pascoe, 1871.
Tribe
Sachalinobiini
Danilevsky, 2010
Sachalinobiini Danilevsky, 2010: 43 [stem: Sachalinobi-]. Type genus: Sachalinobia Jakobson, 1899.
Tribe
Xylosteini
Reitter, 1913
Xylosteina Reitter, 1913a: 5 [stem: Xyloste-]. Type genus: Xylosteus Frivaldszky, 1838.
Subfamily
Spondylidinae
Audinet-Serville, 1832
Spondylii Audinet-Serville, 1832: 123 [stem: Spondylid-]. Type genus: Spondylis Fabricius, 1775.
Tribe
Anisarthrini
Mamaev and Danilevsky, 1973
*Anisarthrites Fairmaire, 1864: 124 [stem: Anisarthr-]. Type genus: Anisarthron Dejean, 1835. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Fairmaire (1864).
Anisarthronini Mamaev and Danilevsky, 1973: 1260, in key [stem: Anisarthr-]. Type genus: Anisarthron Dejean, 1835. Comment: family-group name proposed as new without reference to Anisarthrites Fairmaire, 1864; incorrect original stem formation, not in prevailing usage.
Tribe
Asemini
Thomson, 1861
Asemitae J. Thomson, 1861: 259 [stem: Asem-]. Type genus: Asemum Eschscholtz, 1830.
*Criomorphates Mulsant, 1863a: 421 [stem: Criomorph-]. Type genus: Criomorphus Mulsant, 1839 [preoccupied genus name, not Criomorphus Curtis, 1831 [Hemiptera]; placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1988b); syn. of Tetropium Kirby, 1837]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant (1863).
*Criocéphalites Fairmaire, 1864: 125 [stem: Criocephal-]. Type genus: Criocephalus Mulsant, 1839 [syn. of Arhopalus Audinet-Serville, 1834]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Fairmaire (1864).
Tetropiina Seidlitz, 1891 [Gatt.]: 179 [stem: Tetropi-]. Type genus: Tetropium Kirby, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1988b)].
Criocephalinae Sharp, 1905: 147 [stem: Criocephal-]. Type genus: Criocephalus Mulsant, 1839 [syn. of Arhopalus Audinet-Serville, 1834].
Criomorphini Portevin, 1927: 20, in key [stem: Criomorph-]. Type genus: Criomorphus Mulsant, 1839 [preoccupied genus name, not Criomorphus Curtis, 1831 [Hemiptera]; placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1988b); syn. of Tetropium Kirby, 1837]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Nothorhinini Zagajkevich, 1991: 110 [stem: Nothorhin-]. Type genus: Nothorhina Redtenbacher, 1845. Comment: placed in synonymy with Asemini by Vives and Alonso-Zarazaga (2000: 569).
Tribe
Atimiini
LeConte, 1873
Atimiini J. L. LeConte, 1873: 322 [stem: Atimi-]. Type genus: Atimia Haldeman, 1847.
Tribe
Saphanini
Gistel, 1848
Saphanidae Gistel, 1848: [8] [stem: Saphan-]. Type genus: Saphanus Audinet-Serville, 1834.
Michthysomini J. L. LeConte, 1873: 332 [stem: Michthisomat-]. Type genus: Michthisoma J. L. LeConte, 1850 [as Michthysoma, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Spondylidini
Audinet-Serville, 1832
Spondylii Audinet-Serville, 1832: 123 [stem: Spondylid-]. Type genus: Spondylis Fabricius, 1775. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Necydalinae
Latreille, 1825
Necydalides Latreille, 1825: 401 [stem: Necydal-]. Type genus: Necydalis Linnaeus, 1758.
Subfamily
Dorcasominae
Lacordaire, 1868
Dorcasomides Lacordaire, 1868: 456 [stem: Dorcasom-]. Type genus: Dorcasomus Audinet-Serville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 299, as Dorcasominae), generally accepted as in Aurivillius (1912: 251, as Dorcasomini).
Subfamily
Apatophyseinae
Lacordaire, 1869
Apatophysides Lacordaire, 1869: 234 [stem: Apatophyse-]. Type genus: Apatophysis Chevrolat, 1860. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 240, as Apatophysini), generally accepted as in Švácha and Danilevsky (1988: 125, as Apatophyseinae); incorrect original stem formation, not in prevailing usage; valid status given here as in Danilevsky (2010: 48).
Subfamily
Cerambycinae
Latreille, 1802
Cerambicini Latreille, 1802: 211 [stem: Cerambyc-]. Type genus: Cerambyx Linnaeus, 1758.
Tribe
Acangassuini
Galileo and Martins, 2001
Acangassuini Galileo and Martins, 2001: 95 [stem: Acangassu-]. Type genus: Acangassu Galileo and Martins, 2001. Comment: this name was incorrectly treated as unavailable by Bousquet et al. (2009).
Tribe
Achrysonini
Lacordaire, 1868
Achrysonides Lacordaire, 1868: 231 [stem: Achryson-]. Type genus: Achryson Audinet-Serville, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Murray (1870: 169, as Achrysonidae), generally accepted as in Aurivillius (1912: 39, as Achrysonini); current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Achrys-).
Tribe
Agallissini
LeConte, 1873
Agallissini J. L. LeConte, 1873: 321 [stem: Agalliss-]. Type genus: Agallissus Dalman, 1823.
Tribe
Alanizini
Di Iorio, 2003
Alanizini Di Iorio, 2003: 1 [stem: Alaniz-]. Type genus: Alanizus Di Iorio, 2003.
Tribe
Anaglyptini
Lacordaire, 1868
nomen protectum
Anaglyptides Lacordaire, 1868: 404 [stem: Anaglypt-]. Type genus: Anaglyptus Mulsant, 1839. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1873: 319, as Anaglypti), generally accepted as in Linsley (1964: 173, as Anaglyptini); junior homonym of Anaglyptidae Gistel, 1848 (type genus Anaglyptes Gistel, 1848; syn. of Chalcophora Dejean, 1833) in Coleoptera: Buprestidae; this name was treated as a nomen protectum by Bousquet et al. (2009: 41).
Tribe
Aphanasiini
Lacordaire, 1868
Aphanasiides Lacordaire, 1868: 367 [stem: Aphanasi-]. Type genus: Aphanasium Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 139, as Aphanasiini).
Tribe
Aphneopini
Lacordaire, 1868
Aphnéopides Lacordaire, 1868: 421 [stem: Aphneop-]. Type genus: Aphneope Pascoe, 1863. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 155, as Aphneopini).
Tribe
Auxesini
Lepesme and Breuning, 1952
*Auxésides Lacordaire, 1872: 463 [stem: Auxes-]. Type genus: Auxesis J. Thomson, 1858. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1872).
Auxesina Lepesme and Breuning, 1952: 140 [stem: Auxes-]. Type genus: Auxesis J. Thomson, 1858. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Auxese-).
Psathyrini Quentin, 1954: 103, in key [stem: Psathyr-]. Type genus: Psathyrus J. Thomson, 1857.
Tribe
Basipterini
Fragoso, Monné and Campos Seabra, 1987
Basipterini Fragoso et al., 1987: 201 [stem: Basipter-]. Type genus: Basiptera J. Thomson, 1864.
Tribe
Bimiini
Lacordaire, 1868
Bimiides Lacordaire, 1868: 464 [stem: Bimi-]. Type genus: Bimia A. White, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 254, as Bimiini).
Sibyllini Cerda, 1973: 115 [stem: Sibyll-]. Type genus: Sibylla J. Thomson, 1858 [preoccupied genus name, not Sibylla Stål, 1856 [Orthoptera]; the valid name for this genus has recently been given as Sybilla J. Thomson, 1864 by Monné and Bezark (2009: 30) which is an incorrect subsequent spelling of Sibylla J. Thomson, 1858; syn. of Zehra Özdikmen, 2008]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Bothriospilini
Lane, 1950
Bothriospilinae Lane, 1950: 370 [stem: Bothriospil-]. Type genus: Bothriospila Aurivillius, 1923.
Tribe
Brachypteromatini
Sama, 2008
Brachypteromini Sama, 2008: 229 [stem: Brachypteromat-]. Type genus: Brachypteroma Heyden, 1863. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Callichromatini
Swainson, 1840
Callichrominae Swainson, 1840: 293 [stem: Callichromat-]. Type genus: Callichroma Latreille, 1816. Comment: incorrect original stem formation, not in prevailing usage; this family-group name was incorrectly credited to Swainson and Shuckard (1840) by Bousquet et al. (2009: 42).
Terambidae Gistel, 1848: [8] [stem: Teramb-]. Type genus: Terambus Gistel, 1848 [syn. of Aromia Audinet-Serville, 1834].
Tribe
Callidiini
Kirby, 1837
Callidiadae Kirby, 1837: 170 [stem: Callidi-]. Type genus: Callidium Fabricius, 1775 [for comments on problems with the authorship and type species of this type genus see Bousquet et al. (2009: 42)].
*Phymatodates Mulsant, 1863a: 397 [stem: Phymatod-]. Type genus: Phymatodes Mulsant, 1839 [placed on the Official List of Generic Names in Zoology (ICZN 1989a)]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Tribe
Callidiopini
Lacordaire, 1868
Callidiopsides Lacordaire, 1868: 340 [stem: Callidiop-]. Type genus: Callidiopis A. White, 1855 [as Callidiopsis, unjustified emendation of type-genus by Lacordaire (1868: 356), not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 535, as Callidiopsinae [incorrect stem formation]), generally accepted as in Aurivillius (1912: 115, as Callidiopini); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Callidiopid-).
Tribe
Cerambycini
Latreille, 1802
Cerambicini Latreille, 1802: 211 [stem: Cerambyc-]. Type genus: Cerambyx Linnaeus, 1758.
Subtribe
Cerambycina
Latreille, 1802
Cerambicini Latreille, 1802: 211 [stem: Cerambyc-]. Type genus: Cerambyx Linnaeus, 1758. Comment: Gistel (1856a: 375) used the name Ceratambycidae but this was based on the incorrect subsequent spelling of the type genus name (incorrectly given as Caratambyx in the same work on page 375 but later corrected to Ceratambyx by Gistel (1856b: 13)) and was therefore not proposed as a new family-group name.
Subtribe
Sphallotrichina
Martins and Monné, 2005
Sphallotrichina Martins and Monné, 2005: 2 [stem: Sphallotrich-]. Type genus: Sphallotrichus Fragoso, 1982. Comment: this name was incorrectly treated as unavailable by Bousquet et al. (2009).
Tribe
Certallini
Fairmaire, 1864
Cartallites Fairmaire, 1864: 149 [stem: Certall-]. Type genus: Certallum Dejean, 1821 [as Cartallum, unjustified emendation of type genus name by Audinet-Serville (1834: 94), not in prevailing usage]. Comment: published early September 1864; original vernacular name available (Art. 11.7.2): first used in a latinized form by Sama (1990: 287, as Certallini), generally accepted as in Vives (2000: 155, as Certallini); incorrect original stem formation, not in prevailing usage.
Pytheitae J. Thomson, 1864: 153 [stem: Pythe-]. Type genus: Pytheus Newman, 1840. Comment: published before October 1864.
*Erionispites Chapuis, 1875: 301 [stem: Erionisp-]. Type genus: Erionispa Chapuis, 1875 [syn. of Pytheus Newman, 1840]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Erionispidae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); name written with an accent as Érionispites on page 301 of the original publication, but without accent in the key and for the description; type genus transferred from Chrysomelidae by Lameere (1885) and treated as a synonym of Pytheus Newman, 1840.
Tribe
Chlidonini
Waterhouse, 1879
Chlidoninae C. O. Waterhouse, 1879b: 320 [stem: Chlidon-]. Type genus: Chlidones C. O. Waterhouse, 1879.
Tribe
Cleomenini
Lacordaire, 1868
Cléoménides Lacordaire, 1868: 405 [stem: Cleomen-]. Type genus: Cleomenes J. Thomson, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 554, as Cleomeninae), generally accepted as in Adlbauer et al. (2010: 163, as Cleomenini); this name was incorrectly credited to Pascoe (1869) by Bousquet et al. (2009: 55); we treat this group as a valid tribe as in Adlbauer et al. (2010: 163).
Tribe
Clytini
Mulsant, 1839
Clytaires Mulsant, 1839: 70 [stem: Clyt-]. Type genus: Clytus Laicharting, 1784. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1848: [8], as Clytiidae [incorrect stem formation]), generally accepted as in Aurivillius (1912: 358, as Clytini).
Neoclytitae J. Thomson, 1861: 219 [stem: Neoclyt-]. Type genus: Neoclytus J. Thomson, 1861.
Cyllenitae J. Thomson, 1864: 184 [stem: Cyllen-]. Type genus: Cyllene Newman, 1840 [preoccupied genus name, not Cyllene Gray, 1834 [Mollusca]; syn. of Megacyllene Casey, 1912]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Compsocerini
Thomson, 1864
Compsoceritae J. Thomson, 1864: 260 [stem: Compsocer-]. Type genus: Compsocerus Audinet-Serville, 1834 [for comments on problems with the authorship and type species of this type genus see Bousquet et al. (2009: 44)].
Tribe
Coptommatini
Lacordaire, 1869
Coptommides Lacordaire, 1869: 221 [stem: Coptommat-]. Type genus: Coptomma Newman, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 488, as Coptommatini); First Reviser (Coptommatini Lacordaire, 1869 vs Navomorphini Lacordaire, 1869) not determined, current usage maintained.
Navomorphides Lacordaire, 1869: 223 [stem: Navomorph-]. Type genus: Navomorpha A. White, 1855 [syn. of Coptomma Newman, 1840]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 488, as Navomorphini).
Tribe
Curiini
LeConte, 1873
Curii J. L. LeConte, 1873: 304 [stem: Curi-]. Type genus: Curius Newman, 1840.
Tribe
Deilini
Fairmaire, 1864
*Déilates Mulsant, 1863b: 190 [stem: Deil-]. Type genus: Deilus Audinet-Serville, 1834. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant (1863).
Déilates Fairmaire, 1864: 154 [stem: Deil-]. Type genus: Deilus Audinet-Serville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 294, as Deilini).
Tribe
Dejanirini
Lacordaire, 1868
Déjanirides Lacordaire, 1868: 460 [stem: Dejanir-]. Type genus: Dejanira J. Thomson, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 561, as Dejanirinae), generally accepted as in Aurivillius (1912: 253, as Dejanirini).
Tribe
Diorini
Lane, 1950
Diorinae Lane, 1950: 373 [stem: Dior-]. Type genus: Diorus A. White, 1853.
Tribe
Distichocerini
Pascoe, 1867
*Distichocérites Blanchard, 1845b: 144 [stem: Distichocer-]. Type genus: Distichocera Kirby, 1819. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845b).
Distichocerinae Pascoe, 1867a: 125 [stem: Distichocer-]. Type genus: Distichocera Kirby, 1819.
Tribe
Dodecosini
Aurivillius, 1912
Dodecosini Aurivillius, 1912: 132 [stem: Dodecos-]. Type genus: Dodecosis H. W. Bates, 1867. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Dodecose-).
Olexandrellaeini Zajciw, 1960: 605 [stem: Olexandrell-]. Type genus: Olexandrella Zajciw, 1959. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Dryobiini
Arnett, 1962
nomen protectum
Dryobiini Arnett, 1962c: 861 [stem: Dryobi-]. Type genus: Dryobius J. L. LeConte, 1850. Comment: nomen protectum (see Bousquet et al. 2009: 45); this is a junior homonym of Dryobiadae Gistel, 1856a: 368 (type genus Dryobia Gistel, 1856) which is a junior synonym of Dryophilidae Gistel, 1848.
Tribe
Eburiini
Blanchard, 1845
Éburiites Blanchard, 1845b: 145 [stem: Eburi-]. Type genus: Eburia Lacordaire, 1830. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1861: 237, as Eburitae), generally accepted as in Monné (1993a: 20, as Eburiini).
Tribe
Ectenessini
Martins, 1998
Ectenessini Martins, 1998: 82 [stem: Ecteness-]. Type genus: Ectenessa H. W. Bates, 1885.
Tribe
Elaphidiini
Thomson, 1864
Elaphidionitae J. Thomson, 1864: 235 [stem: Elaphidi-]. Type genus: Elaphidion Audinet-Serville, 1834. Comment: incorrect original stem formation, not in prevailing usage; Ivie (1985: 303) pointed out that the correct stem based on Elaphidion is Elaphidi-; both Elaphidiini and Elaphidionini have been used in recent literature, we prefer to use the correct spelling of the stem here.
Sphérionides Lacordaire, 1868: 312 [stem: Sphaeri-]. Type genus: Sphaerion Audinet-Serville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 239, as Sphaerionini), generally accepted as in Aurivillius (1912: 96, as Sphaerionini); incorrect original stem formation, not in prevailing usage; junior homonym of Sphaeriidae Deshayes, 1855 (type genus Sphaerium Scopoli, 1877) in Mollusca and Sphaerina Erichson, 1845 [incorrect original spelling] (type genus Sphaerius Waltl, 1838) in Myxophaga, the stem of the beetle family-group name was recently emended to Sphaerius- (ICZN 2000); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Stenosphenini J. L. LeConte, 1873: 316 [stem: Stenosphen-]. Type genus: Stenosphenus Haldeman, 1847.
Tribe
Eligmodermini
Lacordaire, 1868
Éligmodermides Lacordaire, 1868: 337 [stem: Eligmoderm-]. Type genus: Eligmoderma J. Thomson, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 114, as Eligmodermini); current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Eligmodermat-).
Tribe
Erlandiini
Aurivillius, 1912
Erlandiini Aurivillius, 1912: 12 [stem: Erlandi-]. Type genus: Erlandia Aurivillius, 1904.
Tribe
Eroschemini
Lacordaire, 1868
Éroschémides Lacordaire, 1868: 515 [stem: Eroschem-]. Type genus: Eroschema Pascoe, 1859. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1872a: 185, as Eroscheminae), generally accepted as in Aurivillius (1912: 287, as Eroschemini); current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Eroschemat-).
Tribe
Eumichthini
Linsley, 1940
Eumichthini Linsley, 1940: 368 [stem: Eumichth-]. Type genus: Eumichthus J. L. LeConte, 1873.
Tribe
Gahaniini
Quentin and Villiers, 1969
Gahaniini Quentin and Villiers, 1969: 615, in key [stem: Gahani-]. Type genus: Gahania Distant, 1907.
Tribe
Glaucytini
Lacordaire, 1868
Glaucytides Lacordaire, 1868: 405 [stem: Glaucyt-]. Type genus: Glaucytes J. Thomson, 1857. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 650, as Glaucytinae), generally accepted as in Aurivillius (1912: 435, as Glaucytini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Glaucytet-).
Tribe
Graciliini
Mulsant, 1839
Graciliaires Mulsant, 1839: 99 [stem: Gracili-]. Type genus: Gracilia Audinet-Serville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1873: 300, as Graciliinae), generally accepted as in Monné (1993b: 9, as Graciliini).
Tribe
Hesperophanini
Mulsant, 1839
Hespérophanaires Mulsant, 1839: 61 [stem: Hesperophan-]. Type genus: Hesperophanes Dejean, 1835.
Subtribe
Daramina
Sama, 2008
Daramina Sama, 2008: 224 [stem: Daram-]. Type genus: Daramus Fairmaire, 1892.
Subtribe
Hesperophanina
Mulsant, 1839
Hespérophanaires Mulsant, 1839: 61 [stem: Hesperophan-]. Type genus: Hesperophanes Dejean, 1835 [see Vives and Alonso-Zarazaga (2000: 657) and Bousquet et al. (2009: 47) for a discussion of problems with the type species of this genus]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869: 523, as Hesperophaninae), generally accepted as in Monné (1993a: 1, as Hesperophanini).
Cerasphoritae J. Thomson, 1861: 234 [stem: Cerasphor-]. Type genus: Cerasphorus Audinet-Serville, 1834.
Tribe
Hesthesini
Pascoe, 1867
Hesthesinae Pascoe, 1867a: 127 [stem: Hesthes-]. Type genus: Hesthesis Newman, 1840. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Hesthese-).
Tribe
Heteropsini
Lacordaire, 1868
nomen protectum
Dichophyiaeidae Gistel, 1848: [8] [stem: Dichophyi-]. Type genus: Dichophyia Gistel, 1848. Comment: nomen oblitum (see Bousquet et al. 2009: 48). Comment: incorrect original stem formation, not in prevailing usage.
Hétéropsides Lacordaire, 1868: 405 [stem: Heterops-]. Type genus: Heterops Blanchard, 1842. Comment: nomen protectum (see Bousquet et al. 2009: 48); original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1872a: 179, as Heteropsinae), generally accepted as in Aurivillius (1912: 438, as Heteropsini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Heterop-); this name was incorrectly attributed to Lacordaire (1869) by Bousquet et al. (2009: 48).
Tribe
Hexoplini
Martins, 2006
Hexoplonini Martins, 2006: 22 [stem: Hexopl-]. Type genus: Hexoplon J. Thomson, 1864. Comment: incorrect original stem formation, not in prevailing usage; this name was incorrectly treated as unavailable by Bousquet et al. (2009).
Tribe
Holopleurini
Chemsak and Linsley, 1974
Holopleurini Chemsak and Linsley, 1974: 183 [stem: Holopleur-]. Type genus: Holopleura J. L. LeConte, 1873.
Tribe
Holopterini
Lacordaire, 1868
Holoptérides Lacordaire, 1868: 393 [stem: Holopter-]. Type genus: Holopterus Blanchard, 1851. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Lucas (1920: xxiii, as Holopterini), generally accepted as in Aurivillius (1912: 148, as Holopterini); this tribe was transferred to the subfamily Lepturinae by Vitali (2002: 32) but placed back into the Cerambycinae by Monné (2005a: 301).
Tribe
Hyboderini
Linsley, 1940
Hyboderini Linsley, 1940: 371 [stem: Hyboder-]. Type genus: Hybodera J. L. LeConte, 1873.
Tribe
Hylotrupini
Zagajkevich, 1991
Hylotrupini Zagajkevich, 1991: 67 [stem: Hylotrup-]. Type genus: Hylotrupes Audinet-Serville, 1834.
Tribe
Ibidionini
Thomson, 1861
Ibidionitae J. Thomson, 1861: 199 [stem: Ibidion-]. Type genus: Ibidion Gory, 1833.
Subtribe
Compsina
Martins and Galileo, 2007
Compsina Martins and Galileo, 2007: 6, in key [stem: Comps-]. Type genus: Compsa Perty, 1832. Comment: this name was incorrectly treated as unavailable by Bousquet et al. (2009); junior homonym of Compsini Pierce, 1913 (type genus Compsus Schönherr, 1823) in Curculionidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Subtribe
Ibidionina
Thomson, 1861
Ibidionitae J. Thomson, 1861: 199 [stem: Ibidion-]. Type genus: Ibidion Gory, 1833. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Ibidi-).
*Sydacini Martins, 2003a: 204 [stem: Sydac-]. Type genus: Sydax Lacordaire, 1868. Comment: name unavailable (Art. 16.1): name not indicated as intentionally new; this taxon was originally described by Martins (1997a: 8-9) but not named.
Subtribe
Tropidina
Martins and Galileo, 2007
Tropidina Martins and Galileo, 2007: 7 [stem: Tropid-]. Type genus: Tropidion J. Thomson, 1867. Comment: this name was incorrectly treated as unavailable by Bousquet et al. (2009); incorrect original spelling maintained (should be Tropidi-) in order to avoid homonymy with Tropidiini Hull, 1949 (type genus Tropidia Meigen, 1822) available in Diptera: Syrphidae.
Tribe
Ideratini
Martins and Napp, 2009
Ideratini Martins and Napp, 2009: 216 [stem: Iderat-]. Type genus: Ideratus J. Thomson, 1864.
Tribe
Lissonotini
Swainson, 1840
Lissonotinae Swainson, 1840: 289 [stem: Lissonot-]. Type genus: Lissonotus Dalman, 1817. Comment: this family-group name was incorrectly credited to Swainson and Shuckard (1840) by Bousquet et al. (2009: 49); the junior homonym Lissonotini Förster, 1869 (type genus Lissonota Gravenhorst, 1829) is available in Hymenoptera: Ichneumonidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Luscosmodicini
Martins, 2003
Luscosmodicini Martins, 2003b: 30 [stem: Luscosmodic-]. Type genus: Luscosmodicum Martins, 1970. Comment: this name was incorrectly treated as unavailable by Bousquet et al. (2009).
Tribe
Lygrini
Sama, 2008
Lygrini Sama, 2008: 222 [stem: Lygr-]. Type genus: Lygrus Fåhraeus, 1872.
Tribe
Macronini
Lacordaire, 1868
Enchapteritae J. Thomson, 1861: 151 [stem: Enchopter-]. Type genus: Enchoptera Saunders, 1850 [as Enchaptera, incorrect subsequent spelling type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; this name has precedence over Macronini Lacordaire, 1868 but has not been used as a valid name after 1899 to our knowledge, although we were unable to find 25 references to conserve usage of Macronini (Art. 23.9.2), we believe the name Macronini should be conserved for this group and an application should be submitted to the Commission (Art. 23.9.3).
Macronides Lacordaire, 1868: 414 [stem: Macron-]. Type genus: Macrones Newman, 1841. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1871: 268, as Macroninae), generally accepted as in Aurivillius (1912: 153, as Macronini); although this is not the oldest name for the tribe, we recommend that an application be submitted to the Commission to conserve usage of the well-established name Macronini Lacordaire, 1868.
Tribe
Megacoelini
Quentin and Villiers, 1969
Megacoelini Quentin and Villiers, 1969: 615, in key [stem: Megacoel-]. Type genus: Megacoelus Lacordaire, 1868.
Tribe
Methiini
Thomson, 1860
Methiitae J. Thomson, 1860a: 127 [stem: Methi-]. Type genus: Methia Newman, 1842.
Tribe
Molorchini
Gistel, 1848
Molorchidae Gistel, 1848: [9] [stem: Molorch-]. Type genus: Molorchus Fabricius, 1792 [see Bousquet (2008: 620) for a discussion of the type species]. Comment: name previously attributed to Mulsant (1862) but was also used by Marseul (1857a: 166, as Molorchidae) prior to Mulsant’s name.
Tribe
Mythodini
Lacordaire, 1868
Mythodides Lacordaire, 1868: 418 [stem: Mythod-]. Type genus: Mythodes J. Thomson, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1871: 268, as Mythodinae), generally accepted as in Aurivillius (1912: 154, as Mythodini).
Tribe
Necydalopsini
Lacordaire, 1868
Nécydalopsides Lacordaire, 1868: 493 [stem: Necydalops-]. Type genus: Necydalopsis Blanchard, 1851. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 275, as Necydalopsini); current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Necydalopse-).
Tribe
Neocorini
Martins, 2005
Neocorini Martins, 2005: 240 [stem: Neocor-]. Type genus: Neocorus J. Thomson, 1864. Comment: this name was incorrectly treated as unavailable by Bousquet et al. (2009).
Tribe
Neostenini
Lacordaire, 1868
Néosténides Lacordaire, 1868: 363 [stem: Neosten-]. Type genus: Neostenus Pascoe, 1857. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1871: 268, as Neosteninae), generally accepted as in Aurivillius (1912: 138, as Neostenini).
Tribe
Obriini
Mulsant, 1839
Obriaires Mulsant, 1839: 95 [stem: Obri-]. Type genus: Obrium Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1848: [9], as Obriidae), generally accepted as in Monné (1993a: 11, as Obriini).
Tribe
Ochyrini
Pascoe, 1871
Ochyrinae Pascoe, 1871: 273 [stem: Ochyr-]. Type genus: Ochyra Pascoe, 1871.
Tribe
Oedenoderini
Aurivillius, 1912
Oedenoderini Aurivillius, 1912: 358 [stem: Oedenoder-]. Type genus: Oedenoderus Chevrolat, 1858.
Tribe
Oemini
Lacordaire, 1868
Oemides Lacordaire, 1868: 216 [stem: Oem-]. Type genus: Oeme Newman, 1840.
Subtribe
Methioidina
Martins, 1997
Methioidina Martins, 1997a: 119 [stem: Methioid-]. Type genus: Methioides Chemsak and Linsley, 1967.
Subtribe
Oemina
Lacordaire, 1868
*Malacoptérites Blanchard, 1845b: 147 [stem: Malacopter-]. Type genus: Malacopterus Audinet-Serville, 1833. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Oemides Lacordaire, 1868: 216 [stem: Oem-]. Type genus: Oeme Newman, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 498, as Oeminae), generally accepted as in Aurivillius (1912: 26, as Oemini).
Tribe
Opsimini
LeConte, 1873
Opsimi J. L. LeConte, 1873: 294 [stem: Opsim-]. Type genus: Opsimus Mannerheim, 1843.
Tribe
Oxycoleini
Martins and Galileo, 2003
Oxycoleini Martins and Galileo, 2003: 52 [stem: Oxycole-]. Type genus: Oxycoleus Lacordaire, 1868. Comment: this name was incorrectly treated as unavailable by Bousquet et al. (2009).
Tribe
Paraholopterini
Martins, 1997
Paraholopterini Martins, 1997b: 201 [stem: Paraholopter-]. Type genus: Paraholopterus Cerda and Cekalovic, 1987.
Tribe
Phalotini
Lacordaire, 1868
Phalotides Lacordaire, 1868: 495 [stem: Phalot-]. Type genus: Phalota Pascoe, 1863. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 276, as Phalotini).
Tribe
Phlyctaenodini
Lacordaire, 1868
Phlycténodides Lacordaire, 1868: 370 [stem: Phlyctaenod-]. Type genus: Phlyctaenodes Newman, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Aurivillius (1912: 140, as Phlyctaenodini), generally accepted as in Monné (1993b: 20, as Phlyctaenodini); incorrect original stem formation, not in prevailing usage.
Tribe
Phoracanthini
Newman, 1840
Stenocoridae Hope, 1834: 106 [stem: Stenocor-]. Type genus: Stenocorus sensu Hope, 1835 [not Stenocorus Geoffroy, 1762; syn. of Phoracantha Newman, 1840]. Comment: based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principles of Priority and Homonymy (Art. 65.2.1); also see Stenocoritae J. Thomson, 1861 in Lepturinae: Rhagiini.
Phoracanthidae Newman, 1840: 2 [stem: Phoracanth-]. Type genus: Phoracantha Newman, 1840.
Tribe
Phyllarthriini
Lepesme and Breuning, 1956
Phyllarthriini Lepesme and Breuning, 1956: 287 [stem: Phyllarthri-]. Type genus: Phyllarthrius Hope, 1843.
Tribe
Piesarthriini
McKeown, 1947
Piesarthrini McKeown, 1947: 55 [stem: Piesarthri-]. Type genus: Piesarthrius Hope, 1834. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Gressitt (1959: 84, as Piesarthini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1); incorrect original stem formation, not in prevailing usage.
Tribe
Piezocerini
Lacordaire, 1868
Piézocérides Lacordaire, 1868: 324 [stem: Piezocer-]. Type genus: Piezocera Audinet-Serville, 1834.
Subtribe
Haruspicina
Martins, 1976
Haruspicina Martins, 1976: 199 [stem: Haruspic-]. Type genus: Haruspex J. Thomson, 1864.
Subtribe
Piezocerina
Lacordaire, 1868
Piézocérides Lacordaire, 1868: 324 [stem: Piezocer-]. Type genus: Piezocera Audinet-Serville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 102, as Piezocerini).
Zelliboriinae Lane, 1951: 5 [stem: Zellibori-]. Type genus: Zelliboria Lane, 1951.
Tribe
Platyarthrini
Bates, 1870
*Coelarthrides Lacordaire, 1868: 405 [stem: Caelomarth-]. Type genus: Caelomarthron J. Thomson, 1860 [as Coelarthron, unjustified emendation of type genus name by Lacordaire (1869: 142), not in prevailing usage; syn. of Platyarthron Guérin-Méneville, 1844]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally accepted as valid; subsequent usage of Coelarthrinae by Lucas (1920: 17) and Caelarthrinae by Ferreira and Veiga Ferreira (1959b: 331) did not validate this name because Lacordaire’s taxon was listed as a synonym of Platyarthrini H. W. Bates; Coelarthridae was used as valid by Ienistea (1986: 30) but it was not attributed to Lacordaire (1868); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Platyarthrinae H. W. Bates, 1870: 419 [stem: Platyarthr-]. Type genus: Platyarthron Guérin-Méneville, 1844.
Tribe
Plectogasterini
Quentin and Villiers, 1969
Plectogasterini Quentin and Villiers, 1969: 615, in key [stem: Plectogaster-]. Type genus: Plectogaster C. O. Waterhouse, 1881. Comment: Bousquet et al. (2009: 52) erroneously listed Plectogastrini as the original spelling.
Tribe
Plectromerini
Nearns and Braham, 2008
Plectromerini Nearns and Braham, 2008: 19 [stem: Plectromer-]. Type genus: Plectromerus Haldeman, 1847 [see Bousquet et al. (2009: 53) about problems with the type species of this genus].
Tribe
Pleiarthrocerini
Lane, 1950
Pleiarthrocerinae Lane, 1950: 371 [stem: Pleiarthrocer-]. Type genus: Pleiarthrocerus Bruch, 1915.
Tribe
Protaxini
Gahan, 1906
Protaxini Gahan, 1906: 92 [stem: Protax-]. Type genus: Protaxis Gahan, 1906. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Protaxe-).
Tribe
Prothemini
Lacordaire, 1868
Prothémides Lacordaire, 1868: 524 [stem: Prothem-]. Type genus: Prothema Pascoe, 1856. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 578, as Protheminae), generally accepted as in Aurivillius (1912: 291, as Prothemini).
Tribe
Psebiini
Lacordaire, 1868
*Leptidéites Fairmaire, 1864: 148 [stem: Leptide-]. Type genus: Leptidea Mulsant, 1839 [preoccupied genus name, not Leptidea Billberg, 1820 [Lepidoptera]; syn. of Nathrius Bréthes, 1916]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Fairmaire (1864).
Psébiides Lacordaire, 1868: 479 [stem: Psebi-]. Type genus: Psebium Pascoe, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 299, as Psebiinae), generally accepted as in Aurivillius (1912: 261, as Psebiini).
Leptideina Reitter, 1913a: 24 [stem: Leptide-]. Type genus: Leptidea Mulsant, 1839 [preoccupied genus name, not Leptidea Billberg, 1820 [Lepidoptera]; syn. of Nathrius Bréthes, 1916]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; Leptideini Verity, 1947 (type genus Leptidea Billberg, 1820) is available in Lepidoptera.
Cambaiinae Lane, 1951: 12 [stem: Cambai-]. Type genus: Cambaia Lane, 1951 [syn. of Paraleptidea Gounelle, 1913].
Nathriini Arnett, 1962c: 860 [stem: Nathri-]. Type genus: Nathrius Brèthes, 1916.
Tribe
Pseudocephalini
Aurivillius, 1912 (1861)
Ametrocephalitae J. Thomson, 1861: 256 [stem: Ametrocephal-]. Type genus: Ametrocephala Blanchard, 1851 [syn. of Pseudocephalus Newman, 1842]. Comment: use of younger name Pseudocephalini Aurivillius, 1912 conserved over this name (Art. 40.2).
Pseudocephalini Aurivillius, 1912: 154 [stem: Pseudocephal-]. Type genus: Pseudocephalus Newman, 1842. Comment: name proposed to replace Ametrocephalini J. Thomson, 1861 because of the synonymy of the type genus; usage of this name conserved over Ametrocephalini Thomson, 1861 (Art. 40.2).
Tribe
Pseudolepturini
Thomson, 1861
Pseudolepturitae J. Thomson, 1861: 146 [stem: Pseudoleptur-]. Type genus: Pseudoleptura J. Thomson, 1861. Comment: we treat this group as a valid tribe as in Adlbauer et al. (2010: 200).
Erythrinae Pascoe, 1866a: 227 [stem: Erythr-]. Type genus: Erythrus A. White, 1853.
Tribe
Psilomorphini
Lacordaire, 1868
Psilomorphides Lacordaire, 1868: 392 [stem: Psilomorph-]. Type genus: Psilomorpha Saunders, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 148, as Psilomorphini).
Tribe
Pteroplatini
Thomson, 1861
Pteroplatitae J. Thomson, 1861: 254 [stem: Pteroplat-]. Type genus: Pteroplatus Buquet, 1840.
Tribe
Pyrestini
Lacordaire, 1868
Pyresthides Lacordaire, 1868: 518 [stem: Pyrest-]. Type genus: Pyrestes Pascoe, 1857 [as Pyresthes, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Aurivillius (1912: 288, as Pyrestini), generally accepted as in Bousquet et al. (2009: 54, as Pyrestini); incorrect original stem formation, not in prevailing usage.
Tribe
Rhagiomorphini
Newman, 1841
Rhagiomorphidae Newman, 1841: 34 [stem: Rhagiomorph-]. Type genus: Rhagiomorpha Newman, 1840.
Tribe
Rhinotragini
Thomson, 1861
Rhinotragitae J. Thomson, 1861: 177 [stem: Rhinotrag-]. Type genus: Rhinotragus Germar, 1824.
Tribe
Rhopalophorini
Blanchard, 1845
Rhopalophorites Blanchard, 1845b: 152 [stem: Rhopalophor-]. Type genus: Rhopalophora Audinet-Serville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Blanchard (1853: 268, as Rhopalophoritae), generally accepted as in Monné (1994a: 1, as Rhopalophorini).
Tribe
Rosaliini
Fairmaire, 1864
Rosaliites Fairmaire, 1864: 137 [stem: Rosali-]. Type genus: Rosalia Audinet-Serville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1873: 310, as Rosaliini), generally accepted as in Linsley (1964: 4, as Rosaliini).
Tribe
Sestyrini
Lacordaire, 1868
Sestyrides Lacordaire, 1868: 405 [stem: Sestyr-]. Type genus: Sestyra Pascoe, 1867. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 643, as Sestyrinae), generally accepted as in Aurivillius (1912: 424, as Sestyrini).
Tribe
Smodicini
Lacordaire, 1868
Smodicides Lacordaire, 1868: 405 [stem: Smodic-]. Type genus: Smodicum Haldeman, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1873: 294, as Smodici), generally accepted as in Aurivillius (1912: 12, as Smodicini); this name was incorrectly attributed to Lacordaire (1869) by Bousquet et al. (2009: 55).
Tribe
Spintheriini
Lacordaire, 1869
Spinthériides Lacordaire, 1869: 219 [stem: Spintheri-]. Type genus: Spintheria J. Thomson, 1861. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 487, as Spintheriini).
Tribe
Stenhomalini
Miroshnikov, 1989
Stenhomalini Miroshnikov, 1989: 742 [stem: Stenhomal-]. Type genus: Stenhomalus A. White, 1855.
Tribe
Stenoderini
Pascoe, 1867
*Sténodérites Blanchard, 1845b: 163 [stem: Stenoder-]. Type genus: Stenoderus Dejean, 1821. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845b).
Syllitae J. Thomson, 1864: 138 [stem: Syllit-]. Type genus: Syllitus Pascoe, 1859. Comment: this name has precedence over Stenoderini Pascoe, 1867 but has not been used as a valid name after 1899 to our knowledge, unfortunately, we were unable to provide 25 references to conserve usage of Stenoderini (Art. 23.9.2) although we believe the name Stenoderini should be conserved for this group until an application to the ICZN is submitted (Art. 23.9.3).
Stenoderinae Pascoe, 1867b: 311 [stem: Stenoder-]. Type genus: Stenoderus Dejean, 1821. Comment: senior homonym of Stenoderini Selander, 1991 (type genus Stenodera Eschscholtz, 1818) currently used as valid in Meloidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1); also, this is not the oldest available name for this tribe, see comments under Syllitae Thomson, 1864.
*Ptérosténides Lacordaire, 1868: 410 [stem: Pterosten-]. Type genus: Pterostenus Laporte, 1840 [syn. of Stenoderus Dejean, 1821]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally accepted as valid; subsequent usage of Pterosteninae by Aurivillius (1912: 150) and Lucas (1920: 53) did not validate this name because Lacordaire’s taxon was listed as a synonym of Stenoderini.
Calliprasonini McKeown, 1947: 71 [stem: Callipras-]. Type genus: Calliprason A. White, 1843. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Gressitt (1959: 148, as Calliprasonini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1); incorrect original stem formation, not in prevailing usage.
Tribe
Stenopterini
Gistel, 1848
Stenopteridae Gistel, 1848: [9] [stem: Stenopter-]. Type genus: Stenopterus Illiger, 1804.
Tribe
Strongylurini
Lacordaire, 1868
Strongylurides Lacordaire, 1868: 379 [stem: Strongylur-]. Type genus: Strongylurus Hope, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 548, as Strongylurinae), generally accepted as in Aurivillius (1912: 144, as Strongylurini).
Tribe
Tessarommatini
Lacordaire, 1868
Tessarommides Lacordaire, 1868: 378 [stem: Tessarommat-]. Type genus: Tessaromma Newman, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 143, as Tessarommatini); name also spelled Tessérommides in original publication on page 204 (key); incorrect original stem formation, not in prevailing usage.
Tribe
Thraniini
Gahan, 1906
Thraniini Gahan, 1906: 236 [stem: Thrani-]. Type genus: Thranius Pascoe, 1859.
Tribe
Thyrsiini
Marinoni and Napp, 1984
Thyrsiini Marinoni and Napp, 1984: 44 [stem: Thyrsi-]. Type genus: Thyrsia Dalman, 1819.
Tribe
Tillomorphini
Lacordaire, 1868
Tillomorphides Lacordaire, 1868: 405 [stem: Tillomorph-]. Type genus: Tillomorpha Blanchard, 1851. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 554, as Tillomorphinae), generally accepted as in Adlbauer et al. (2010: 206, as Tillomorphini); this name was incorrectly credited to Pascoe, 1869 by Bousquet et al. (2009: 56).
Epipedocerini Gahan, 1906: 305 [stem: Epipedocer-]. Type genus: Epipedocera Chevrolat, 1863.
Tribe
Torneutini
Thomson, 1861
Torneutitae J. Thomson, 1861: 272 [stem: Torneut-]. Type genus: Torneutes Reich, 1838.
Thaumasidae J. Thomson, 1864: 313 [stem: Thaumas-]. Type genus: Thaumasus Reiche, 1853.
Tribe
Trachyderini
Dupont, 1836
Trachydérides Dupont, 1836: 1 [stem: Trachyder-]. Type genus: Trachyderes Dalman, 1817.
Subtribe
Ancylocerina
Thomson, 1864
Ancyloceritae J. Thomson, 1864: 210 [stem: Ancylocer-]. Type genus: Ancylocera Audinet-Serville, 1834.
Subtribe
Trachyderina
Dupont, 1836
Trachydérides Dupont, 1836: 1 [stem: Trachyder-]. Type genus: Trachyderes Dalman, 1817. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1861: 209, as Trachyderitae), generally accepted as in Monné (1994b: 16, as Trachyderini).
Purpuricenitae J. Thomson, 1861: 203 [stem: Purpuricen-]. Type genus: Purpuricenus Dejean, 1821.
Tylositae J. Thomson, 1861: 205 [stem: Tylose-]. Type genus: Tylosis J. L. LeConte, 1850. Comment: incorrect original stem formation, not in prevailing usage.
Sphaenothecitae J. Thomson, 1861: 212 [stem: Sphaenothec-]. Type genus: Sphaenothecus Dupont, 1838.
Megaderitae J. Thomson, 1861: 213 [stem: Megader-]. Type genus: Megaderus Dejean, 1821.
Eriphitae J. Thomson, 1864: 200 [stem: Eriph-]. Type genus: Eriphus Audinet-Serville, 1834.
Pteracanthitae J. Thomson, 1864: 255 [stem: Pteracanth-]. Type genus: Pteracantha Newman, 1838.
Metopocoïlitae J. Thomson, 1864: 255 [stem: Metopocoil-]. Type genus: Metopocoilus Audinet-Serville, 1832.
Sternacanthitae J. Thomson, 1864: 259 [stem: Sternacanth-]. Type genus: Sternacanthus Audinet-Serville, 1832.
Tropidosomitae J. Thomson, 1864: 256 [stem: Tropidosomat-]. Type genus: Tropidosoma Perty, 1832. Comment: incorrect original stem formation, not in prevailing usage.
Poecilopéplides Lacordaire, 1868: 404 [stem: Poecilopepl-]. Type genus: Poecilopeplus Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 449, as Poecilopeplini).
*Dorcacérides Lacordaire, 1868: 404 [stem: Dorcacer-]. Type genus: Dorcacerus Dejean, 1821. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1868).
Sténaspides Lacordaire, 1868: 404 [stem: Stenaspid-]. Type genus: Stenaspis Audinet-Serville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 653, as Stenaspidinae), generally accepted as in Aurivillius (1912: 457, as Stenaspini); incorrect original stem formation, not in prevailing usage.
Paristémiides Lacordaire, 1868: 404 [stem: Paristemi-]. Type genus: Paristemia Westwood, 1841. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte (1873: 309, as Paristemiini).
Dorcacerinae H. W. Bates, 1870: 430 [stem: Dorcacer-]. Type genus: Dorcacerus Dejean, 1821. Comment: Aurivillius (1912: 476) used the spelling Dorcadocerini but this was based on Docadocerus, which is an incorrect subsequent spelling of the type genus name.
Tribe
Tragocerini
Pascoe, 1867
Tragocerinae Pascoe, 1867a: 125 [stem: Tragocer-]. Type genus: Tragocerus Latreille, 1829.
Tribe
Trichomesiini
Aurivillius, 1912
Trichomesiini Aurivillius, 1912: 276 [stem: Trichomesi-]. Type genus: Trichomesia Pascoe, 1859.
Tribe
Tropocalymmatini
Lacordaire, 1868
Tropocalymmides Lacordaire, 1868: 408 [stem: Tropocalymmat-]. Type genus: Tropocalymma J. Thomson, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 150, as Tropocalymmatini); incorrect original stem formation, not in prevailing usage.
Tribe
Typhocesini
Lacordaire, 1868
Typhocésides Lacordaire, 1868: 539 [stem: Typhoces-]. Type genus: Typhocesis Pascoe, 1863. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1912: 296, as Typhocesini).
Tribe
Unxiini
Napp, 2007
Unxiini Napp, 2007: 312 [stem: Unxi-]. Type genus: Unxia J. Thomson, 1861.
Tribe
Uracanthini
Blanchard, 1853
*Uracantitas Blanchard, 1851b: 475 [stem: Uracanth-]. Type genus: Uracanthus Hope, 1833 [Uracanthus is an incorrect subsequent spelling of Uracantha Hope, 1833 (p. 64), first used by Hope (1834: 108), in prevailing usage and so deemed to be the correct original spelling (Art. 33.3.1)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1851b); incorrect original stem formation, not in prevailing usage.
Uracanthitae Blanchard, 1853: 264 [stem: Uracanth-]. Type genus: Uracanthus Hope, 1833 [Uracanthus is an incorrect subsequent spelling of Uracantha Hope, 1833 (p. 64), first used by Hope (1834: 108), in prevailing usage and so deemed to be the correct original spelling (Art. 33.3.1)].
Rhinophthalmitae J. Thomson, 1861: 152 [stem: Rhinophthalm-]. Type genus: Rhinophthalmus J. Thomson, 1861.
Tribe
Vesperellini
Sama, 2008
Vesperellini Sama, 2008: 227 [stem: Vesperell-]. Type genus: Vesperella Dayrem, 1933.
Tribe
Xystrocerini
Blanchard, 1845
Xystrocérites Blanchard, 1845b: 147 [stem: Xystrocer-]. Type genus: Xystrocera Audinet-Serville, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1861: 249, as Xystroceritae), generally accepted as in Martins and Carvalho (1984: 214, as Xystrocerini).
Subfamily
Lamiinae
Latreille, 1825
Lamiariae Latreille, 1825: 401 [stem: Lami-]. Type genus: Lamia Fabricius, 1775.
Tribe
Acanthocinini
Blanchard, 1845
*Aedilaires Mulsant, 1839: 142 [stem: Aedil-]. Type genus: Aedilis Audinet-Serville, 1835. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Acanthocinites Blanchard, 1845b: 154 [stem: Acanthocin-]. Type genus: Acanthocinus Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Blanchard (1853: 289, as Acanthocinitae), generally accepted as in Monné (1995a: 1, as Acanthocinini).
Trypanidiitae J. Thomson, 1860a: 7 [stem: Trypanidi-]. Type genus: Trypanidius Blanchard, 1846.
Dectitae J. Thomson, 1860a: 127 [stem: Dect-]. Type genus: Dectes J. L. LeConte, 1852.
*Astynomaires Mulsant, 1863b: 286 [stem: Astynom-]. Type genus: Astynomus Dejean, 1835. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Mulsant (1863b).
Lagocheirinae H. W. Bates, 1863: 100 [stem: Lagocheir-]. Type genus: Lagocheirus Dejean, 1835.
Liopi J. L. LeConte, 1873: 338 [stem: Leiopod-]. Type genus: Leiopus Audinet-Serville, 1835 [as Liopus, unjustified emendation of type genus name by Agassiz (1846b: 204), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Graphisurini Leng, 1920: 283 [stem: Graphisur-]. Type genus: Graphisurus Kirby, 1837.
Astynomini Portevin, 1927: 39 [stem: Astynom-]. Type genus: Astynomus Dejean, 1835.
Tribe
Acanthoderini
Thomson, 1860
Acanthoderitae J. Thomson, 1860a: 5 [stem: Acanthoder-]. Type genus: Acanthoderes Audinet-Serville, 1835. Comment: First Reviser (Acanthoderini J. Thomson, 1860 vs Dryoctenini J. Thomson, 1860 vs Oreoderini J. Thomson, 1860) not determined, current usage maintained.
Dryoctenitae J. Thomson, 1860a: 29 [stem: Dryocten-]. Type genus: Dryoctenes Audinet-Serville, 1835.
Oreoderitae J. Thomson, 1860a: 29 [stem: Oreoder-]. Type genus: Oreodera Audinet-Serville, 1835.
Hoplosiae J. L. LeConte and G. H. Horn, 1883: 326 [stem: Oplosi-]. Type genus: Oplosia Mulsant, 1863 [as Hoplosia, unjustified emendation of type genus name by Fairmaire (1864), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Acmocerini
Thomson, 1864
Acmoceritae J. Thomson, 1864: 57 [stem: Acmocer-]. Type genus: Acmocera Dejean, 1835.
Tribe
Acridocephalini
Dillon and Dillon, 1959
Acridocephalidi E. S. Dillon and L. S. Dillon, 1959a: 49 [stem: Acridocephal-]. Type genus: Acridocephala Chevrolat, 1855.
Tribe
Acrocinini
Swainson, 1840
Acrocininae Swainson, 1840: 287 [stem: Acrocin-]. Type genus: Acrocinus Illiger, 1806. Comment: this family-group name was incorrectly credited to Swainson and Shuckard (1840) by Bousquet et al. (2009: 24).
Tribe
Aderpasini
Breuning and Teocchi, 1978
Aderpasini Breuning and Teocchi, 1978: 142 [stem: Aderpas-]. Type genus: Aderpas J. Thomson, 1864.
Tribe
Aerenicini
Lacordaire, 1872
Aerénicides Lacordaire, 1872: 897 [stem: Aerenic-]. Type genus: Aerenica Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Harold (1872: 248, as Aerenicinae), generally accepted as in Aurivillius (1923: 596, as Aerenicini).
Tribe
Agapanthiini
Mulsant, 1839
Agapanthaires Mulsant, 1839: 172 [stem: Agapanthi-]. Type genus: Agapanthia Audinet-Serville, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1884: 255, as Agapanthinae [incorrect stem formation]), generally accepted as in Aurivillius (1923: 458, as Agapanthiini); incorrect original stem formation, not in prevailing usage.
Hippopsitae J. Thomson, 1860a: 123 [stem: Hippopse-]. Type genus: Hippopsis Lepeletier and Audinet-Serville, 1828. Comment: incorrect original stem formation, not in prevailing usage.
Nemotragitae J. Thomson, 1864: 93 [stem: Nemotrag-]. Type genus: Nemotragus Westwood, 1843.
Anauxesitae J. Thomson, 1864: 94 [stem: Anauxese-]. Type genus: Anauxesis J. Thomson, 1857. Comment: incorrect original stem formation, not in prevailing usage.
Aprosopitae J. Thomson, 1864: 95 [stem: Aprosop-]. Type genus: Aprosopus Guérin-Méneville, 1844.
Aegoprepinae Pascoe, 1871: 277 [stem: Aegoprep-]. Type genus: Aegoprepes Pascoe, 1871.
Pachypézides Lacordaire, 1872: 691 [stem: Pachypez-]. Type genus: Pachypeza Audinet-Serville, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in L. S. Dillon and E. S. Dillon (1945: 12, as Pachypezini).
Spalacopsides Lacordaire, 1872: 701 [stem: Spalacopse-]. Type genus: Spalacopsis Newman, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gahan (1890: 325, as Spalacopsinae), generally accepted as in Aurivillius (1923: 360, as Spalacopsini); incorrect original stem formation, not in prevailing usage.
Didymonychini Aurivillius, 1922b: 31 [stem: Didymonych-]. Type genus: Didymonycha Aurivillius, 1922 [syn. of Amillarus J. Thomson, 1857].
*Amillarinen Aurivillius, 1926a: 22 [stem: Amillar-]. Type genus: Amillarus J. Thomson, 1857. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Hippopsiconini L. S. Dillon and E. S. Dillon, 1945: 11 [stem: Hippopsicon-]. Type genus: Hippopsicon J. Thomson, 1858.
Tribe
Amphoecini
Breuning, 1951
Amphoecini Breuning, 1951a: 5 [stem: Amphoec-]. Type genus: Amphoecus Montrouzier, 1861.
Tribe
Ancitini
Aurivillius, 1917
Ancitini Aurivillius, 1917: 28 [stem: Ancit-]. Type genus: Ancita J. Thomson, 1864.
Tribe
Ancylonotini
Lacordaire, 1869
Ancylonotides Lacordaire, 1869: 391 [stem: Ancylonot-]. Type genus: Ancylonotus Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1871: 268, as Ancylonotinae), generally accepted as in Aurivillius (1922a: 152, as Ancylonotini).
Tribe
Anisocerini
Thomson, 1860
Anisoceritae J. Thomson, 1860a: 31 [stem: Anisocer-]. Type genus: Anisocerus Lacordaire, 1830.
Onychoceritae J. Thomson, 1864: 19 [stem: Onychocer-]. Type genus: Onychocerus Lacordaire, 1830.
Platysternides Lacordaire, 1872: 729 [stem: Platystern-]. Type genus: Platysternus Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1923: 371, as Platysternini); junior homonym of the turtle family Platysternidae Gray, 1869 (type genus Platysternon Gray, 1831) which is currently used as valid; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Apomecynini
Thomson, 1860
Apomecynitae J. Thomson, 1860a: 66 [stem: Apomecyn-]. Type genus: Apomecyna Dejean, 1821. Comment: name incorrectly spelled Apomecinitae on page 3 but correct spelling used on pages 42, 66 and 68 of the original publication.
Adétides Lacordaire, 1872: 592 [stem: Adet-]. Type genus: Adetus J. L. LeConte, 1852. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 288, as Adetini).
*Agennopsides Lacordaire, 1872: 592 [stem: Agennopse-]. Type genus: Agennopsis J. Thomson, 1857 [syn. of Adetus J. L. LeConte, 1852]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Ptéricoptides Lacordaire, 1872: 601 [stem: Ptericopt-]. Type genus: Ptericoptus Lacordaire, 1830. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gestro (1876: 140, as Ptericoptini), generally accepted as in Aurivillius (1922a: 294, as Ptericoptini).
Ectatosiides Lacordaire, 1872: 708 [stem: Ectatosi-]. Type genus: Ectatosia Pascoe, 1857. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1923: 363, as Ectatosiini).
Ischiolonchides Lacordaire, 1872: 709 [stem: Ischiolonch-]. Type genus: Ischioloncha J. Thomson, 1860. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1923: 364, as Ischiolonchini).
Tribe
Astathini
Thomson, 1864
*Tétraophthalmites Blanchard, 1845b: 160 [stem: Tetraophthalm-]. Type genus: Tetraophthalmus Dejean, 1835. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Sama (2010: 56, as Tetraophthalmini) but not generally accepted as valid.
Astathitae J. Thomson, 1864: 117 [stem: Astath-]. Type genus: Astathes Newman, 1842 [syn. of Tetraophthalmus Dejean, 1835]. Comment: published before October 1864; this family-group name was also used in the same year by Pascoe (1864 [before 3 October]: 8, as Asthateinae [incorrect stem formation]); Pascoe (1864: 81, 85) refers to the publication by J. Thomson (1864) which is further evidence that the publication by Thomson was published first; this tribe name was incorrectly credited to Pascoe (1864) by Bousquet et al. (2009: 26).
Tribe
Batocerini
Thomson, 1864
Batoceritae J. Thomson, 1864: 74 [stem: Batocer-]. Type genus: Batocera Dejean, 1835.
Tribe
Calliini
Thomson, 1864
Callitae J. Thomson, 1864: 123 [stem: Calli-]. Type genus: Callia Audinet-Serville, 1835. Comment: incorrect original stem formation, not in prevailing usage.
Gryllicides Lacordaire, 1872: 902 [stem: Gryllic-]. Type genus: Gryllica J. Thomson, 1860. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by H. W. Bates (1874a: 234, as Gryllicinae), generally accepted as in Aurivillius (1923: 604, as Gryllicini).
Tribe
Ceroplesini
Thomson, 1860
Ceroplesitae J. Thomson, 1860a: 95 [stem: Ceroples-]. Type genus: Ceroplesis Audinet-Serville, 1835 [placed on the Official List of Generic Names in Zoology (ICZN 1986c)].
Subtribe
Ceroplesina
Thomson, 1860
Ceroplesitae J. Thomson, 1860a: 95 [stem: Ceroples-]. Type genus: Ceroplesis Audinet-Serville, 1835 [placed on the Official List of Generic Names in Zoology (ICZN 1986c)].
Subtribe
Crossotina
Thomson, 1864
Crossotitae J. Thomson, 1864: 64 [stem: Crossot-]. Type genus: Crossotus Audinet-Serville, 1835. Comment: downgraded to subtribe by Sama (2008: 235).
Écyroschémides Lacordaire, 1872: 503 [stem: Ecyroschemat-]. Type genus: Ecyroschema J. Thomson, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 241, as Ecyroschemini); incorrect original stem formation, not in prevailing usage.
Hécyridides Lacordaire, 1872: 517 [stem: Hecyrid-]. Type genus: Hecyrida J. Thomson, 1860 [syn. of Hecyra J. Thomson, 1857]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 318, as Hecyridinae), generally accepted as in Aurivillius (1922a: 243, as Hecyrini).
Corynofreinae Aurivillius, 1911a: 37 [stem: Corynofre-]. Type genus: Corynofrea Aurivillius, 1911.
Tribe
Cloniocerini
Lacordaire, 1872
Cloniocérides Lacordaire, 1872: 590 [stem: Cloniocer-]. Type genus: Cloniocerus Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 287, as Cloniocerini).
Tribe
Colobotheini
Thomson, 1860
Colobotheitae J. Thomson, 1860a: 18 [stem: Colobothe-]. Type genus: Colobothea Lepeletier and Audinet-Serville, 1825 [Colobothea is an incorrect subsequent spelling of the original spelling Colobotea, in prevailing usage and so deemed to be the correct original spelling (Art. 33.3.1)].
Tribe
Compsosomatini
Thomson, 1857
Compsosomites J. Thomson, 1857b: 70 [stem: Compsosomat-]. Type genus: Compsosoma Lacordaire, 1830. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1860a: 34, as Compsosomitae), generally accepted as in Aurivillius (1923: 336, as Compsosomatini); incorrect original stem formation, not in prevailing usage.
Aereneites J. Thomson, 1868: 92 [stem: Aerene-]. Type genus: Aerenea J. Thomson, 1857 [Aerenea is an incorrect subsequent spelling of the original spelling Aerenaea, introduced by J. Thomson (1860a: 34), in prevailing usage and so deemed to be the correct original spelling (Art. 33.3.1)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gounelle (1908: 7, as Aereneinae), generally accepted as in Aurivillius (1923: 338, as Aereneini).
Tribe
Cyrtinini
Thomson, 1864
Cyrtinitae J. Thomson, 1864: 41 [stem: Cyrtin-]. Type genus: Cyrtinus J. L. LeConte, 1852.
Acanthomerosternoplonini Tippmann, 1956: 10 [stem: Acanthomerosternopl-]. Type genus: Acanthomerosternoplon Tippmann, 1955 [syn. of Omosarotes Pascoe, 1860]. Comment: incorrect original stem formation, not in prevailing usage.
Scopadini Villiers, 1980: 587 [stem: Scopad-]. Type genus: Scopadus Pascoe, 1857.
Tribe
Desmiphorini
Thomson, 1860
Desmiphoritae J. Thomson, 1860a: 74 [stem: Desmiphor-]. Type genus: Desmiphora Audinet-Serville, 1835.
*Anaesthétites Fairmaire, 1864: 166 [stem: Anaesthet-]. Type genus: Anaesthetis Dejean, 1835. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Métonides Lacordaire, 1869: 387 [stem: Metont-]. Type genus: Meton Pascoe, 1859. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 150, as Metonini); incorrect original stem formation, not in prevailing usage.
Hebesecinae Pascoe, 1871: 277 [stem: Hebesecid-]. Type genus: Hebesecis Pascoe, 1865. Comment: incorrect original stem formation, not in prevailing usage.
Amymomides Lacordaire, 1872: 468 [stem: Amymom-]. Type genus: Amymoma Pascoe, 1866 [preoccupied genus name, not Amymoma Latreille, 1797 [Crustacea]; syn. of Neoamymoma Marinoni, 1977]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 218, as Amymomini); permanently invalid (Art. 39): based on preoccupied type genus.
Crinotarsides Lacordaire, 1872: 475 [stem: Crinotars-]. Type genus: Crinotarsus Blanchard, 1853. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 229, as Crinotarsini).
Épicastides Lacordaire, 1872: 490 [stem: Epicast-]. Type genus: Epicasta J. Thomson, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 237, as Epicastini).
Apodasyides Lacordaire, 1872: 623 [stem: Apodasy-]. Type genus: Apodasya Pascoe, 1863 [the genus name Chaetosoma Chevrolat, 1843: 367 is a senior objective synonym of Apodasya Pascoe and should be used as valid based on the Principle of Priority; Chaetosoma Chevrolat, 1843 is also a senior homonym of the well-established Chaetosoma Westwood, 1851, which is the type genus of Chaetosomatidae Crowson, 1952 used as valid in Cucujoidea; because the discovery of the available name Chaetosoma Chevrolat, 1843 causes problems for well-established names in Cucujoidea and Cerambycidae, an application was recently submitted by Bousquet and Bouchard (2010) to suppress it for the Principles of Priority and Homonymy (see Appendix 6)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 321, as Apodasinae [incorrect stem formation]), generally accepted as in Aurivillius (1922a: 305, as Apodasyini).
Nédinides Lacordaire, 1872: 635 [stem: Nedin-]. Type genus: Nedine J. Thomson, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 317, as Nedinini).
Estolides Lacordaire, 1872: 636 [stem: Estol-]. Type genus: Estola Fairmaire and Germain, 1859. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1873: 340, as Estolae), generally accepted as in Aurivillius (1922a: 317, as Estolini).
Psenocerini J. L. LeConte, 1873: 333 [stem: Psenocer-]. Type genus: Psenocerus J. L. LeConte, 1852.
Eupogonii J. L. LeConte, 1873: 342 [stem: Eupogoni-]. Type genus: Eupogonius J. L. LeConte, 1852.
Essisini Aurivillius, 1917: 44 [stem: Essis-]. Type genus: Essisus Pascoe, 1866.
Velorini Aurivillius, 1917: 32 [stem: Velor-]. Type genus: Velora J. Thomson, 1864.
Tribe
Dorcadionini
Swainson, 1840
Dorcadioninae Swainson, 1840: 290 [stem: Dorcadi-]. Type genus: Dorcadion Dalman, 1817. Comment: this family-group name was incorrectly credited to Swainson and Shuckard (1840) by Bousquet et al. (2009: 29); current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Dorcadi-; see Vives and Alonso-Zarazaga 2000: 659); tribe placed in synonymy with Lamiini by Sama (2008: 233).
Dorcadodiidae Gistel, 1856a: 376 [stem: Dorcadodi-]. Type genus: Dorcadodium Gistel, 1856 [this name is a senior synonym of Carinatodorcadion Breuning, 1943 (see Vives and Alonso-Zarazaga 2000: 659 for type species designation of Dorcadodium Gistel); we could not find enough references to treat Dorcadodium Gistel, 1856 as a nomen oblitum, however we believe that an application should be submitted to the Commission to preserve usage of Carinatodorcadion Breuning, 1943].
Tribe
Dorcaschematini
Thomson, 1860
Dorcaschemitae J. Thomson, 1860a: 107 [stem: Dorcaschemat-]. Type genus: Dorcaschema Haldeman, 1847. Comment: incorrect original stem formation, not in prevailing usage; name spelled Dorchaschemitae on page 104 but spelled Dorcaschemitae on pages 4 and 107 of the original publication.
Protonarthronitae J. Thomson, 1864: 57 [stem: Protonarthr-]. Type genus: Protonarthron J. Thomson, 1858. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Elytracanthinini
Bousquet, 2009
Elytracanthinae Lane, 1955: 281 [stem: Elytracanth-]. Type genus: Elytracantha Lane, 1955 [preoccupied type genus, not Elytracantha Kleine, 1915 [Coleoptera: Brentidae]; syn. of Elytracanthina Monné, 2005]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Elytracanthinini Bousquet, 2009: 30 [stem: Elytracanthin-]. Type genus: Elytracanthina Monné, 2005. Comment: replacement name for Elytracanthinae Lane, 1955 because of the homonymy of the type genus.
Tribe
Enicodini
Thomson, 1864
Enicoditae J. Thomson, 1864: 36 [stem: Enicod-]. Type genus: Enicodes Gray, 1832. Comment: First Reviser (Enicodini J. Thomson, 1864 vs Nemaschematini J. Thomson, 1864 vs Leptonotini J. Thomson, 1864) not determined, current usage maintained.
Nemaschemitae J. Thomson, 1864: 36 [stem: Nemaschemat-]. Type genus: Nemaschema J. Thomson, 1861. Comment: incorrect original stem formation, not in prevailing usage.
Leptonotitae J. Thomson, 1864: 36 [stem: Leptonot-]. Type genus: Leptonota J. Thomson, 1861.
*Énotides Lacordaire, 1872: 487 [stem: Enotet-]. Type genus: Enotes J. Thomson, 1864. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Tribe
Eupromerini
Galileo and Martins, 1995
Eupromerini Galileo and Martins, 1995: 132 [stem: Eupromer-]. Type genus: Eupromera Westwood, 1845.
Tribe
Forsteriini
Tippmann, 1960
Hebestolitae J. Thomson, 1864: 107 [stem: Hebestol-]. Type genus: Hebestola Blanchard, 1851 [preoccupied genus name, not Hebestola Haldeman, 1847 [Coleoptera: Cerambycidae: Lamiinae: Monochamini]; syn. of Neohebestola Marinoni, 1977]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Forsteriini Tippmann, 1960: 210 [stem: Forsteri-]. Type genus: Forsteria Tippmann, 1960 [syn. of Falsamblesthis Breuning, 1959].
Falsamblesthiini Gilmour, 1961: 131 [stem: Falsamblesth-]. Type genus: Falsamblesthis Breuning, 1959. Comment: name proposed to replace Forsteriini Tippmann, 1960 because of the synonymy of the type genus; however, because the name was proposed after 1960, it cannot be maintained (Art. 40.2); incorrect original stem formation, not in prevailing usage.
Tribe
Gnomini
Thomson, 1860
Gnomitae J. Thomson, 1860a: 105 [stem: Gnom-]. Type genus: Gnoma Fabricius, 1801.
Tribe
Gyaritini
Breuning, 1950
Gyaritini Breuning, 1950c: 27 [stem: Gyarit-]. Type genus: Gyaritus Pascoe, 1858.
Tribe
Heliolini
Breuning, 1951
Heliolini Breuning, 1951a: 8 [stem: Heliol-]. Type genus: Heliolus Fauvel, 1907.
Tribe
Hemilophini
Thomson, 1868
nomen protectum
Amphionychitae J. Thomson, 1860a: 63 [stem: Amphionych-]. Type genus: Amphionycha Dejean, 1835 [syn. of Adesmus Lepeletier and Audinet-Serville, 1825]. Comment: nomen oblitum (see Bousquet et al. 2009: 31).
Hemilophitae J. Thomson, 1868: 189 [stem: Hemiloph-]. Type genus: Hemilophus Audinet-Serville, 1835. Comment: nomen protectum (see Bousquet et al. 2009: 31).
Itesini Lepesme, 1943: 137 [stem: It-]. Type genus: Ites C. O. Waterhouse, 1880. Comment: incorrect original stem formation, not in prevailing usage; the tribe Itini Reitter, 1913 (type genusIta Tournier, 1878), which is based on the same stem, is currently used as valid in Coleoptera: Curculionidae.
Tribe
Homonoeini
Thomson, 1864
Homonaeitae J. Thomson, 1864: 35 [stem: Homonoe-]. Type genus: Homonoea Newman, 1842 [as Homonaea, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Bumétopides Lacordaire, 1872: 477 [stem: Bumetopi-]. Type genus: Bumetopia Pascoe, 1858. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gestro (1876: 140, as Bumetopini [incorrect stem formation]), generally accepted as in Aurivillius (1922a: 231, as Bumetopini [incorrect stem formation]); incorrect original stem formation, not in prevailing usage.
Tribe
Hyborhabdini
Aurivillius, 1911
Hyborhabdinae Aurivillius, 1911b: 22 [stem: Hyborhabd-]. Type genus: Hyborhabdus Aurivillius, 1911.
Tribe
Lamiini
Latreille, 1825
Lamiariae Latreille, 1825: 401 [stem: Lami-]. Type genus: Lamia Fabricius, 1775.
Pachystolaeidae Gistel, 1848: [9] [stem: Pachystol-]. Type genus: Pachystola Dejean, 1835 [syn. of Lamia Fabricius, 1775]. Comment: incorrect original stem formation, not in prevailing usage.
Phrissomitae J. Thomson, 1860a: 22 [stem: Phrissomat-]. Type genus: Phrissoma Dejean, 1835. Comment: incorrect original stem formation, not in prevailing usage; J. Thomson originally spelled the family-group name Phryssomitae on page 25, however the spelling Phrissomitae was used on pages 2 and 22 in the same publication; tribe placed in synonymy with Lamiini by Sama (2008: 233).
Morimitae J. Thomson, 1864: 77 [stem: Morim-]. Type genus: Morimus Brullé, 1832.
Potemnemini Aurivillius, 1922a: 117 [stem: Potemnem-]. Type genus: Potemnemus J. Thomson, 1864.
Tribe
Laticraniini
Lane, 1959
Laticraniinae Lane, 1959: 312 [stem: Laticrani-]. Type genus: Laticranium Lane, 1959.
Tribe
Mauesiini
Lane, 1956
Mauesinae Lane, 1956: 19 [stem: Mauesi-]. Type genus: Mauesia Lane, 1956. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Megabasini
Thomson, 1860
Megabasitae J. Thomson, 1860a: 30 [stem: Megabas-]. Type genus: Megabasis Audinet-Serville, 1835. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Megabase-).
Tribe
Mesosini
Mulsant, 1839
Mésosaires Mulsant, 1839: 165 [stem: Mesos-]. Type genus: Mesosa Latreille, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gistel (1848: [9], as Mesosaeidae [incorrect stem formation]), generally accepted as in López-Pérez (2006: 59, as Mesosini).
Tribe
Microcymaturini
Breuning and Teocchi, 1985
Microcymaturini Breuning and Teocchi, 1985: 155 [stem: Microcymatur-]. Type genus: Microcymatura Breuning, 1950.
Tribe
Moneilemini
Thomson, 1864
Moneilemitae J. Thomson, 1864: 43 [stem: Moneilem-]. Type genus: Moneilema Say, 1824. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Moneilemat-).
Tribe
Monochamini
Gistel, 1848
Monohammidae Gistel, 1848: [9] [stem: Monocham-]. Type genus: Monochamus Dejean, 1821 [as Monohammus, unjustified emendation of type genus name by Dejean (1835: 340), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Taeniotitae J. Thomson, 1864: 76 [stem: Taeniot-]. Type genus: Taeniotes Audinet-Serville, 1835.
Agnitae J. Thomson, 1864: 83 [stem: Agni-]. Type genus: Agnia Newman, 1842. Comment: incorrect original stem formation, not in prevailing usage.
Geranitae J. Thomson, 1864: 93 [stem: Gerani-]. Type genus: Gerania Audinet-Serville, 1835. Comment: incorrect original stem formation, not in prevailing usage.
Ptychodes J. L. LeConte, 1873: 335 [stem: Ptychod-]. Type genus: Ptychodes Audinet-Serville, 1835.
Goes J. L. LeConte, 1873: 335 [stem: Goet-]. Type genus: Goes J. L. LeConte, 1852. Comment: incorrect original stem formation, not in prevailing usage.
Docohammidi E. S. Dillon and L. S. Dillon, 1959b: 7 [stem: Docohamm-]. Type genus: Docohammus Aurivillius, 1908.
Tribe
Morimonellini
Lobanov, Danilevsky and Murzin, 1981
Morimonellini Lobanov et al., 1981: 790 [stem: Morimonell-]. Type genus: Morimonella Podaný, 1979. Comment: description by indication (distinguishing characters given in Podany (1979: 43, as Oligorchini)).
Tribe
Morimopsini
Lacordaire, 1869
Morimopsides Lacordaire, 1869: 289 [stem: Morimops-]. Type genus: Morimopsis J. Thomson, 1857. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 64, as Morimopsini); current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Morimopse-).
Tribe
Nyctimeniini
Gressitt, 1951
Nyctimenitae J. Thomson, 1864: 94 [stem: Nyctimen-]. Type genus: Nyctimene J. Thomson, 1857 [preoccupied genus name, not Nyctimene Borkenhausen, 1797 [Mammalia]; syn. of Nyctimenius Gressitt, 1951]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Nyctimeniini Gressitt, 1951: 629 [stem: Nyctimeni-]. Type genus: Nyctimenius Gressitt, 1951. Comment: replacement name for Nyctimenini J. Thomson, 1864 because of the homonymy of the type genus.
Tribe
Obereini
Thomson, 1864
Obereitae J. Thomson, 1864: 117 [stem: Obere-]. Type genus: Oberea Dejean, 1835. Comment: published before October 1864; this family-group name was also used in the same year by Pascoe (1864 [before 3 October]: 8, as Obereinae); Pascoe (1864: 81, 85) refers to the publication by J. Thomson (1864) which is further evidence that the publication by Thomson was published first; this tribe name was incorrectly credited to Pascoe (1864) by Bousquet et al. (2009: 33).
Tribe
Oculariini
Breuning, 1950
Oculariini Breuning, 1950a: 263 [stem: Oculari-]. Type genus: Ocularia Jordan, 1894.
Tribe
Onciderini
Thomson, 1860
Oncideritae J. Thomson, 1860a: 38 [stem: Oncider-]. Type genus: Oncideres Lacordaire, 1830 [Oncideres is an incorrect subsequent spelling of Oncyderes Lacordaire, 1830, introduced by Audinet-Serville (1835: 67), in prevailing usage and attributed to Lacordaire (1830b), e.g., Monné (2005b: 571), and so deemed to be the correct original spelling (Art. 33.3.1)]. Comment: First Reviser (Onciderini J. Thomson, 1860 vs Hypsiomatini J. Thomson, 1860) not determined, current usage maintained.
Hypsiomitae J. Thomson, 1860a: 109 [stem: Hypsiomat-]. Type genus: Hypsioma Audinet-Serville, 1835. Comment: incorrect original stem formation, not in prevailing usage.
Hypselominae Pascoe, 1864: 7 [stem: Hypselom-]. Type genus: Hypselomus Perty, 1832.
Tribe
Oncideropsidini
Aurivillius, 1922
Oncideropsidini Aurivillius, 1922c: 165 [stem: Oncideropsid-]. Type genus: Oncideropsis Aurivillius, 1922. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Oncideropse-).
Tribe
Onocephalini
Thomson, 1860
Onocephalitae J. Thomson, 1860a: 120 [stem: Onocephal-]. Type genus: Onocephala J. Thomson, 1857.
Tribe
Onychogleneini
Aurivillius, 1923
Onychogleneini Aurivillius, 1923: 513 [stem: Onychoglene-]. Type genus: Onychoglenea Aurivillius, 1922.
Tribe
Parmenini
Mulsant, 1839
Parménaires Mulsant, 1839: 118 [stem: Parmen-]. Type genus: Parmena Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1864: 38, as Parmenitae), generally accepted as in Villiers (1978: 449, as Parmenini).
Hexathricitae J. Thomson, 1864: 38 [stem: Hexatrich-]. Type genus: Hexatricha A. White, 1846 [as Hexathrica, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: the spelling Hexarthricitae (page 38) was corrected to Hexathricitae on page 339 and in the errata (page 483) of the same work; the corrected spelling represents an incorrect original stem formation, not in prevailing usage.
*Dorcadidides Lacordaire, 1869: 257 [stem: Dorcadid-]. Type genus: Dorcadida A. White, 1846. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 241, as Dorcadidini), but not generally accepted as valid.
Tribe
Petrognathini
Blanchard, 1845
Pétrognathites Blanchard, 1845b: 160 [stem: Petrognath-]. Type genus: Petrognatha Leach, 1819. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 316, as Petrognathinae), generally accepted as in Aurivillius (1922a: 205, as Petrognathini).
*Omacanthides Lacordaire, 1872: 447 [stem: Omacanth-]. Type genus: Omacantha Audinet-Serville, 1835 [syn. of Petrognatha Leach, 1819]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 243, as Omacanthini), but not generally accepted as valid; Omacanthidae was used as valid by Ienistea (1986: 30) but it was not attributed to Lacordaire (1872); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Phacellini
Lacordaire, 1872
Phacellides Lacordaire, 1872: 664 [stem: Phacell-]. Type genus: Phacellus Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gounelle (1908: 7, as Phacellinae), generally accepted as in Aurivillius (1923: 339, as Phacellini).
Tribe
Phantasini
Kolbe, 1897
*Phantasides Lacordaire, 1869: 285 [stem: Phantas-]. Type genus: Phantasis J. Thomson, 1860. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1869).
Phantasinae Kolbe, 1897: 306 [stem: Phantas-]. Type genus: Phantasis J. Thomson, 1860. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Phantase-).
Phantasini Hunt and Breuning, 1957: 51 [stem: Phantas-]. Type genus: Phantasis J. Thomson, 1860. Comment: family group name proposed as new without reference to Phantasinae Kolbe, 1897; current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Phantase-).
Tribe
Phrynetini
Thomson, 1864
Phrynetitae J. Thomson, 1864: 71 [stem: Phrynet-]. Type genus: Phryneta Dejean, 1835.
Tribe
Phymasternini
Teocchi, 1989
Phymasternini Teocchi, 1989: 4 [stem: Phymastern-]. Type genus: Phymasterna Laporte, 1840 [for comments on problems with the authorship and type species of this type genus see Bousquet et al. (2009: 35)].
Tribe
Phytoeciini
Mulsant, 1839
Phytoeciaires Mulsant, 1839: 191 [stem: Phytoeci-]. Type genus: Phytoecia Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1864: 8, as Phytoeciinae), generally accepted as in Villiers (1978: 521, as Phytoeciini).
Tribe
Pogonocherini
Mulsant, 1839
Pogonochéraires Mulsant, 1839: 151 [stem: Pogonocher-]. Type genus: Pogonocherus Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1873: 340, as Pogonocherini), generally accepted as in Villiers (1978: 465, as Pogonocherini).
*Exocentrites Fairmaire, 1864: 157 [stem: Exocentr-]. Type genus: Exocentrus Dejean, 1835. Comment: published early September 1864; original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Fairmaire (1864).
Exocentrinae Pascoe, 1864: 7 [stem: Exocentr-]. Type genus: Exocentrus Dejean, 1835. Comment: published 3 October 1864.
Zaploi J. L. LeConte and G. H. Horn, 1883: 327 [stem: Zaplo-]. Type genus: Zaplous J. L. LeConte, 1878.
Tribe
Polyrhaphidini
Thomson, 1860
Polyrhaphitae J. Thomson, 1860a: 30 [stem: Polyrhaphid-]. Type genus: Polyrhaphis Audinet-Serville, 1835. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Pretiliini
Martins and Galileo, 1990
Pretiliini Martins and Galileo, 1990: 705, in key [stem: Pretili-]. Type genus: Pretilia H. W. Bates, 1866.
Tribe
Proctocerini
Aurivillius, 1922
*Cliniides Lacordaire, 1872: 424 [stem: Clini-]. Type genus: Clinia J. Thomson, 1857 [syn. of Proctocera Chevrolat, 1855]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Breuning (1950d: 411, as Cliniini), but not generally accepted as valid; Cliniidae was used as valid by Ienistea (1986: 30) but it was not attributed to Lacordaire (1872); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Proctocerini Aurivillius, 1922a: 182 [stem: Proctocer-]. Type genus: Proctocera Chevrolat, 1855.
Tribe
Prosopocerini
Thomson, 1864
Prosopoceritae J. Thomson, 1864: 72 [stem: Prosopocer-]. Type genus: Prosopocera Blanchard, 1845.
Tribe
Pteropliini
Thomson, 1860
Pteropliitae J. Thomson, 1860a: 73 [stem: Pteropli-]. Type genus: Pteroplius Lacordaire, 1830 [as Pteroplia, incorrect subsequent spelling of type genus name, not in prevailing usage; Pteroplius is an incorrect subsequent spelling of Pterhoplius Lacordaire, 1830, introduced by Audinet-Serville (1835: 65), in prevailing usage and attributed to Lacordaire (1830b), e.g., Monné (2005b: 295), and so deemed to be the correct original spelling (Art. 33.3.1)].
Abrynitae J. Thomson, 1864: 44 [stem: Abryn-]. Type genus: Abryna Newman, 1842.
Protorhopalitae J. Thomson, 1864: 69 [stem: Protorhopal-]. Type genus: Protorhopala J. Thomson, 1860.
Niphoninae Pascoe, 1864: 56 [stem: Niphon-]. Type genus: Niphona Mulsant, 1839.
Ataxiides Lacordaire, 1872: 597 [stem: Ataxi-]. Type genus: Ataxia Haldeman, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1873: 344, as Ataxiini), generally accepted as in Aurivillius (1922a: 291, as Ataxiini).
Emphytoeciides Lacordaire, 1872: 713 [stem: Emphytoeci-]. Type genus: Emphytoecia Fairmaire and Germain, 1860. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1923: 365, as Emphytoeciini).
Baroeides Lacordaire, 1872: 439 [stem: Barae-]. Type genus: Baraeus J. Thomson, 1858 [as Baroeus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: also spelled Baréides on page 414 of the same work; original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 206, as Baraeini).
Atossides Lacordaire, 1872: 496 [stem: Atoss-]. Type genus: Atossa J. Thomson, 1864 [syn. of Grammoechus J. Thomson, 1864]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 149, as Atossini).
Metagnomini Aurivillius, 1925: 13 [stem: Metagnom-]. Type genus: Metagnoma Aurivillius, 1925.
Tribe
Rhodopinini
Gressitt, 1951
Rhodopides Lacordaire, 1872: 450 [stem: Rhodopid-]. Type genus: Rhodopis J. Thomson, 1857 [preoccupied genus name, not Rhodopis Reichenbach, 1854 [Aves]; syn. of Rhodopina Gressit, 1951]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 210, as Rhodopini); incorrect original stem formation, not in prevailing usage; permanently invalid (Art. 39): based on preoccupied type genus.
Rhodopinini Gressitt, 1951: 439 [stem: Rhodopin-]. Type genus: Rhodopina Gressitt, 1951. Comment: replacement name for Rhodopini Lacordaire, 1872 because of the homonymy of the type genus.
Tribe
Saperdini
Mulsant, 1839
Saperdaires Mulsant, 1839: 181 [stem: Saperd-]. Type genus: Saperda Fabricius, 1775 [placed on the Official List of Generic Names in Zoology (ICZN 1980)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Blanchard (1853: 299, as Saperditae), generally accepted as in Aurivillius (1923: 468, as Saperdini).
Gleneïtae J. Thomson, 1864: 123 [stem: Glene-]. Type genus: Glenea Newman, 1842 [for comments on problems with the authorship and type species of this type genus see Bousquet et al. (2009: 37)].
Tribe
Stenobiini
Breuning, 1950
Stenobiini Breuning, 1950b: 305 [stem: Stenobi-]. Type genus: Stenobia Lacordaire, 1872.
Tribe
Sternotomini
Thomson, 1860
*Stellognathites Blanchard, 1845b: 158 [stem: Stellognath-]. Type genus: Stellognatha Dejean, 1835. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Sternotomitae J. Thomson, 1860a: 87 [stem: Sternotom-]. Type genus: Sternotomis Percheron, 1836 [Sternodonta Dejean, 1835, a senior synonym of Sternotomis Percheron, 1836, is a nomen oblitum and Sternotomis is a nomen protectum following Sama (2009a: 24)]. Comment: current spelling maintained (Art. 29.3.1.1): incorrect original stem formation in prevailing usage (should be Sternotomid-).
Tribe
Tapeinini
Thomson, 1857
Tapeinites J. Thomson, 1857b: 41 [stem: Tapein-]. Type genus: Tapeina Lepeletier and Audinet-Serville, 1828. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1860a: 73, as Tapeinitae), generally accepted as in Aurivillius (1922a: 236, as Tapeinini).
Tribe
Tetraopini
Thomson, 1860
Tetraopesitae J. Thomson, 1860a: 66 [stem: Tetraop-]. Type genus: Tetraopes Schönherr, 1817. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Tetraulaxini
Breuning and Teocchi, 1977
Tetraulaxini Breuning and Teocchi, 1977: 881 [stem: Tetraulax-]. Type genus: Tetraulax Jordan, 1903. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Tetraulac-).
Tribe
Tetropini
Portevin, 1927
*Polyopsiates Mulsant, 1863b: 340 [stem: Polyopsi-]. Type genus: Polyopsia Mulsant, 1839 [syn. of Tetrops Stephens, 1829]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Tétropides Planet, 1924: 326 [stem: Tetrop-]. Type genus: Tetrops Stephens, 1829 [for comments on problems with the authorship and type species of this type genus see Bousquet et al. (2009: 38)]. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899; the argument of Sama (2008: 240) that Planet’s name is available, even if published after 1900 in a vernacular form, because it was used as valid in a latinized form and credited to Planet (1924) by Vives (2000: 508) is incorrect (see Bousquet et al. 2009: 38).
Tetropini Portevin, 1927: 39, in key [stem: Tetrop-]. Type genus: Tetrops Stephens, 1829 [for comments on problems with the authorship and type species of this type genus see Bousquet et al. (2009: 38)].
Tribe
Theocrini
Lacordaire, 1872
Théocrides Lacordaire, 1872: 494 [stem: Theocr-]. Type genus: Theocris J. Thomson, 1858. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gahan (1890: 322, as Theocrinae), generally accepted as in Aurivillius (1922a: 238, as Theocridini [incorrect stem formation]).
Tribe
Tmesisternini
Blanchard, 1853
Tmesisternitae Blanchard, 1853: 274 [stem: Tmesistern-]. Type genus: Tmesisternus Latreille, 1829.
Spingnothitae J. Thomson, 1864: 31 [stem: Sphingnot-]. Type genus: Sphingnotus Perroud, 1855 [as Spingnothus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Ichthyosomitae J. Thomson, 1864: 33 [stem: Ichthyosom-]. Type genus: Ichthyosomus Boisduval, 1835.
*Arsysiides Lacordaire, 1872: 479 [stem: Arsysi-]. Type genus: Arsysia Pascoe, 1867 [syn. of Trigonoptera Perroud, 1855]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Gestro (1876: 140, as Arsysiini), but not generally accepted as valid; Arsysiidae was used as valid by Ienistea (1986: 30) but it was not attributed to Lacordaire (1872); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Trigonopterini Aurivillius, 1922a: 229 [stem: Trigonopter-]. Type genus: Trigonoptera Perroud, 1855. Comment: replacement name for Arsysiides Lacordaire, 1872 because of the synonymy of the type genus.
Tribe
Tragocephalini
Thomson, 1857
Tragocephalites J. Thomson, 1857b: 26 [stem: Tragocephal-]. Type genus: Tragocephala Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. Thomson (1860a: 88, as Tragocephalitae), generally accepted as in Aurivillius (1922a: 171, as Tragocephalini).
Tribe
Xenicotelini
Matsushita, 1933
Xenicotelini Matsushita, 1933: 346 [stem: Xenicotel-]. Type genus: Xenicotela H. W. Bates, 1884.
Tribe
Xenofreini
Aurivillius, 1923
Xenofreini Aurivillius, 1923: 375 [stem: Xenofre-]. Type genus: Xenofrea H. W. Bates, 1885. Comment: this name has been attributed to H. W. Bates (1885: 373) in the literature but we did not find such a name in the Biologia Centrali-Americana or other publications by the same author.
Tribe
Xenoleini
Lacordaire, 1872
Xénoléides Lacordaire, 1872: 460 [stem: Xenole-]. Type genus: Xenolea J. Thomson, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Aurivillius (1922a: 216, as Xenoleini).
Tribe
Xylorhizini
Lacordaire, 1872
Xylorhizides Lacordaire, 1872: 443 [stem: Xylorhiz-]. Type genus: Xylorhiza Dejean, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 316, as Xylorrhizinae [incorrect stem formation]), generally accepted as in Aurivillius (1922a: 208, as Xylorhizini).
Tribe
Zygocerini
Thomson, 1864
Zygoceritae J. Thomson, 1864: 87 [stem: Zygocer-]. Type genus: Zygocera Erichson, 1842.
Disterninae Pascoe, 1871: 268 [stem: Distern-]. Type genus: Disterna J. Thomson, 1864.
Family
Megalopodidae
Latreille, 1802
Megalopides Latreille, 1802: 227 [stem: Megalopod-]. Type genus: Megalopus Fabricius, 1801.
Subfamily
Megalopodinae
Latreille, 1802
Megalopides Latreille, 1802: 227 [stem: Megalopod-]. Type genus: Megalopus Fabricius, 1801. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Palophaginae
Kuschel and May, 1990
Palophaginae Kuschel and May, 1990: 699 [stem: Palophag-]. Type genus: Palophagus Kuschel, 1990.
Subfamily
Zeugophorinae
Böving and Craighead, 1931
Zeugophorinae Böving and Craighead, 1931: 63, in key [stem: Zeugophor-]. Type genus: Zeugophora Kunze, 1818 [placed on the Official List of Generic Names in Zoology (ICZN 1986a)].
Family
Orsodacnidae
Thomson, 1859
Orsodachnidae C. G. Thomson, 1859: 154 [stem: Orsodacn-]. Type genus: Orsodacne Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 2002a)].
Subfamily
Orsodacninae
Thomson, 1859
Orsodachnidae C. G. Thomson, 1859: 154 [stem: Orsodacn-]. Type genus: Orsodacne Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 2002a)]. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Aulacoscelidinae
Chapuis, 1874
Aulacoscélites Chapuis, 1874: 54 [stem: Aulacoscelid-]. Type genus: Aulacoscelis Duponchel et Chevrolat, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 245, as Aulacoscelinae), generally accepted as in Monrós (1959a: 18, as Aulacoscelidinae); incorrect original stem formation, not in prevailing usage; a recent application by Santiago-Blay (2008) to conserve usage of the stem Aulacoscel- was not approved by the Commission (ICZN 2010b).
Family
Chrysomelidae
Latreille, 1802
Chrysomelinae Latreille, 1802: 220 [stem: Chrysomel-]. Type genus: Chrysomela Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1984c)]. Comment: First Reviser (Chrysomelidae Latreille, 1802 vs Bruchidae Latreille, 1802 vs Galerucidae Latreille, 1802) not determined, current usage maintained.
Subfamily
Sagrinae
Leach, 1815
Sagrida Leach, 1815: 113 [stem: Sagr-]. Type genus: Sagra Fabricius, 1792.
Tribe
Carpophagini
Chapuis, 1874
Carpophagites Chapuis, 1874: 36 [stem: Carpophag-]. Type genus: Carpophagus W. S. MacLeay, 1827. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby (1903: 4, as Carpophaginae), generally accepted as in Clavareau (1913a: 6, as Carpophagini); the older name Carpophaginae Selby, 1835 (type genus Carpophaga Selby, 1835) is available in Aves (see Bock 1994: 139); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Diaphanopsidini
Monrós, 1958
Diaphanopsidini Monrós, 1958a: 7 [stem: Diaphanopsid-]. Type genus: Diaphanops Schönherr, 1845. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Seeno and Wilcox (1982: 17, 18) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Diaphanop-).
Tribe
Megamerini
Chapuis, 1874
Mégamérites Chapuis, 1874: 30 [stem: Megamer-]. Type genus: Megamerus W. S. MacLeay, 1827. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby (1903: 2, as Megamerinae), generally accepted as in Clavareau (1913a: 4, as Megamerini); First Reviser (Megamerini Chapuis, 1874 vs Ametallini Chapuis, 1874 vs Mecynoderini Chapuis, 1874) not determined, current usage maintained.
Amétallites Chapuis, 1874: 46 [stem: Ametall-]. Type genus: Ametalla Hope, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby (1903: 1, as Ametallinae), generally accepted as in Clavareau (1913a: 12, as Ametallini).
Mécynodérites Chapuis, 1874: 44 [stem: Mecynoder-]. Type genus: Mecynodera Hope, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby (1903: 1, as Mecynoderinae), generally accepted as in Clavareau (1913a: 11, as Mecynoderini).
Tribe
Sagrini
Leach, 1815
Sagrida Leach, 1815: 113 [stem: Sagr-]. Type genus: Sagra Fabricius, 1792.
Subfamily
Bruchinae
Latreille, 1802
Bruchelae Latreille, 1802: 192 [stem: Bruch-]. Type genus: Bruchus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology and older name Bruchus Geoffroy, 1762 placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1995a)]. Comment: Bruchidae Latreille, 1802 placed on the Official List of Family-Group Names in Zoology (ICZN 1995a).
Tribe
Amblycerini
Bridwell, 1932
Amblycerinae Bridwell, 1932: 103, in key [stem: Amblycer-]. Type genus: Amblycerus Thunberg, 1815.
Subtribe
Amblycerina
Bridwell, 1932
Spermophagidae Crotch, 1873b: 93 [stem: Spermophag-]. Type genus: Spermophagus sensu G. H. Horn, 1873 [not Spermophagus Schönherr, 1833; syn. of Amblycerus Thunberg, 1815]. Comment: based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principles of Priority and Homonymy (Art. 65.2.1); also see Spermophagini Borowiec, 1987 below.
Amblycerinae Bridwell, 1932: 103, in key [stem: Amblycer-]. Type genus: Amblycerus Thunberg, 1815. Comment: although this is the oldest name for the tribe, we recommend that an application be sent to the Commission in order to conserve usage of Amblycerina Bridwell, 1932 because the older name Spermophagina Crotch, 1873 is based on a misidentified type genus (Art. 65.2.1); MAAZ and CHCL will submit an application to the Commission in order to conserve the current concept of Amblycerus Thunberg, 1815 following the discovery of an overlooked type species designation that would alter the concept of the genus (also see Alonso-Zarazaga and Lyal (1999: 8, 23)).
Subtribe
Spermophagina
Borowiec, 1987
Spermophagini Borowiec, 1987: 27 [stem: Spermophag-]. Type genus: Spermophagus Schönherr, 1833. Comment: family-group name proposed as new without reference to Spermophagidae Crotch, 1873; an application needs to be submitted to the Commission to suppress Spermophagidae Crotch, 1873 (based on the misidentified type genus Spermophagus sensu Horn, 1873) for the Principles of Priority and Homonymy (Art. 65.2.1) to conserve this name as valid.
Tribe
Bruchini
Latreille, 1802
Bruchelae Latreille, 1802: 192 [stem: Bruch-]. Type genus: Bruchus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology and older name Bruchus Geoffroy, 1762 placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1995a)]. Comment: Bruchidae Latreille, 1802 placed on the Official List of Family-Group Names in Zoology (ICZN 1995a).
Subtribe
Acanthoscelidina
Bridwell, 1946
Acanthoscelidini Bridwell, 1946: 54, in key [stem: Acanthoscelid-]. Type genus: Acanthoscelides Schilsky, 1905. Comment: First Reviser (Acanthoscelidina Bridwell, 1946 vs Bruchidiina Bridwell, 1946) not determined, current usage maintained; description of family-group name not unequivocal (Art. 13.1.1) but name treated as available (Art. 13.2.1); Bridwell (1946: 53) mentioned that he wanted to establish the two new tribes Bruchidiini and Acanthoscelidini but he followed by saying that it was “premature to attempt a diagnosis” of these tribes; both tribal names were included in his key (p. 54) but their identification was not unequivocal; Bradley (1947: 37) noted that the new tribes Bruchidiini and Acanthoscelidini were not diagnosed adequately by Bridwell (1946), however he did not cite Article 13 of the then current edition of the Code of Zoological nomenclature to treat those names as unavailable; since Bruchidiini and Acanthoscelidini have been used as valid and attributed to Bridwell (1946) by subsequent authors, e.g., Bottimer (1968: 1028 and 1015 respectively), we have also treated those names as available here.
Bruchidiini Bridwell, 1946: 54, in key [stem: Bruchidi-]. Type genus: Bruchidius Schilsky, 1905. Comment: description of family-group name not unequivocal (Art. 13.1.1) but name treated as available (Art. 13.2.1); see comment under Acanthoscelidini Bridwell, 1946.
Subtribe
Bruchina
Latreille, 1802
Bruchelae Latreille, 1802: 192 [stem: Bruch-]. Type genus: Bruchus Linnaeus, 1767 [placed on the Official List of Generic Names in Zoology and older name Bruchus Geoffroy, 1762 placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1995a)]. Comment: Bruchidae Latreille, 1802 placed on the Official List of Family-Group Names in Zoology (ICZN 1995a).
Lariidae Bedel, 1901: 341 [stem: Lari-]. Type genus: Laria Scopoli, 1763 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1995a); syn. of Bruchus Linnaeus, 1767]. Comment: permanently invalid (Art. 39): based on suppressed type genus.
Subtribe
Megacerina
Bridwell, 1946
Megacerini Bridwell, 1946: 54, in key [stem: Megacer-]. Type genus: Megacerus Fahraeus, 1839. Comment: Megacerini Viret, 1961 (type genus Megaceros Owen, 1844) is used as valid in Mammalia: Cervidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Eubaptini
Bridwell, 1932
Eubaptinae Bridwell, 1932: 103, in key [stem: Eubapt-]. Type genus: Eubaptus Lacordaire, 1845.
Tribe
Kytorhinini
Bridwell, 1932
Kytorhininae Bridwell, 1932: 103, in key [stem: Kytorhin-]. Type genus: Kytorhinus Fischer von Waldheim, 1809.
Tribe
Pachymerini
Bridwell, 1929
Pachymerinae Bridwell, 1929: 142 [stem: Pachymer-]. Type genus: Pachymerus Thunberg, 1805. Comment: precedence (Pachymerini Bridwell, 1929 vs Caryedini Brimwell, 1929 vs Caryopemini Brimwell, 1929) given to taxon originally proposed at the higher rank (Art. 24.1).
Subtribe
Caryedontina
Bridwell, 1929
Caryedini Bridwell, 1929: 143 [stem: Caryedont-]. Type genus: Caryedon Schönherr, 1823. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Decelle (1966).
Subtribe
Caryopemina
Bridwell, 1929
Caryopemini Bridwell, 1929: 143 [stem: Caryopem-]. Type genus: Caryopemon Jekel, 1855. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Caryopemon-).
Subtribe
Pachymerina
Bridwell, 1929
Pachymerinae Bridwell, 1929: 142 [stem: Pachymer-]. Type genus: Pachymerus Thunberg, 1805.
Tribe
Rhaebini
Blanchard, 1845
Rhaebites Blanchard, 1845b: 180 [stem: Rhaeb-]. Type genus: Rhaebus Fischer von Waldheim, 1824. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kraatz (1879: 278, as Rhaebini), generally accepted as in Anton (2010: 353, as Rhaebini); Blanchard (1845b: 193) also used the spelling Rhoebites.
Subfamily
Donaciinae
Kirby, 1837
Donaciadae Kirby, 1837: 222 [stem: Donaci-]. Type genus: Donacia Fabricius, 1775.
Tribe
Donaciini
Kirby, 1837
Donaciadae Kirby, 1837: 222 [stem: Donaci-]. Type genus: Donacia Fabricius, 1775.
Donacociadae Gistel, 1856a: 377 [stem: Donacoci-]. Type genus: Donacocia Gistel, 1856 [although this genus has been considered a senior syn. of Plateumaris C. G. Thomson, 1859 by some authors, Löbl and Silfverberg (2010: 64) recently chose “brevicornis Ahrens, 1810” as the type species for this genus; syn. of Donacia Fabricius, 1775]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Haemoniini
Chen, 1941
Haemoniini Chen, 1941: 8 [stem: Haemoni-]. Type genus: Haemonia Dejean, 1821 [syn. of Macroplea Smouelle, 1819].
Tribe
Plateumarini
Böving, 1922
Plateumarini Böving, 1922: 50 [stem: Plateumar-]. Type genus: Plateumaris C. G. Thomson, 1859. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Plateumarid-).
Plateumarini Askevold, 1990: 633 [stem: Plateumar-]. Type genus: Plateumaris C. G. Thomson, 1859. Comment: family-group name proposed as new without reference to Plateumarini Böving, 1922; current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Plateumarid-).
Subfamily
Criocerinae
Latreille, 1804
Criocerides Latreille, 1804c: 159 [stem: Criocer-]. Type genus: Crioceris Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Criocerini
Latreille, 1804
Criocerides Latreille, 1804c: 159 [stem: Criocer-]. Type genus: Crioceris Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: published 7 March 1804; this family-group name was also used in the same year by Latreille (1804d [between 19 August and 17 September]: 324, as Criocerides); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Criocerid-).
Tribe
Lemini
Gyllenhal, 1813
Lemoideae Gyllenhal, 1813: 632 [stem: Lem-]. Type genus: Lema Fabricius, 1798 [placed on the Official List of Generic Names in Zoology (ICZN 1970a)].
Tribe
Pseudocriocerini
Heinze, 1962
Pseudocriocerini Heinze, 1962: 198, in key [stem: Pseudocriocer-]. Type genus: Pseudocrioceris Pic, 1916. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Pseudocriocerid-).
Subfamily
Cassidinae
Gyllenhal, 1813
Cassideae Gyllenhal, 1813: 434 [stem: Cassid-]. Type genus: Cassida Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1974b)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1974b, as Cassidinae Stephens, 1831); First Reviser found (Cassidinae Gyllenhal, 1813 vs Hispinae Gyllenhal, 1813) is Shuckard (1839b: 68).
Tribe
Alurnini
Chapuis, 1875
Alurnites Chapuis, 1875: 292 [stem: Alurn-]. Type genus: Alurnus Fabricius, 1775. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1910: 69, as Alurnini), generally accepted as in Staines (2002: 742, as Alurnini).
*Sphaeropalpites Chapuis, 1875: 359 [stem: Sphaeropalp-]. Type genus: Sphaeropalpus Guérin-Méneville, 1844 [syn. of Platyauchenia Sturm, 1843]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Platyaucheniitae Spaeth, 1929b: 113 [stem: Platyaucheni-]. Type genus: Platyauchenia Sturm, 1843.
Tribe
Anisoderini
Chapuis, 1875
Anisodérites Chapuis, 1875: 294 [stem: Anisoder-]. Type genus: Anisodera Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1911: 40, as Anisoderini), generally accepted as in Würmli (1975: 10, as Anisoderini).
*Lasiochilini Gressitt, 1950: 62 [stem: Lasiochil-]. Type genus: Lasiochila Weise, 1916. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); the author apparently used this family-group name in error for Anisoderini (see Gressitt 1953: 126); Lasiochilinae Carayon, 1972 (type genus Lasiochilus Reuter, 1871) has been used as valid in Hemiptera.
Tribe
Aproidini
Weise, 1911
Aprioidini Weise, 1911: 41 [stem: Aproid-]. Type genus: Aproida Pascoe, 1863.
Tribe
Arescini
Chapuis, 1875
Arescites Chapuis, 1875: 298 [stem: Aresc-]. Type genus: Arescus Perty, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1910: 69, as Arescini), generally accepted as in Staines (2002: 741, as Arescini).
Tribe
Aspidimorphini
Chapuis, 1875
Aspidimorphites Chapuis, 1875: 406 [stem: Aspidimorph-]. Type genus: Aspidimorpha Hope, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Spaeth (1914: (129), as Aspidormophitae [incorrect stem formation]), generally accepted as in Hincks (1952: 336, as Aspidomorphini).
Tribe
Basiprionotini
Gressitt, 1952 (1914)
*Prioptérites Chapuis, 1875: 367 [stem: Priopter-]. Type genus: Prioptera Hope, 1840 [syn. of Basiprionota Chevrolat, 1836]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875).
Priopteritae Spaeth, 1914: (129) [stem: Priopter-]. Type genus: Prioptera Hope, 1840 [syn. of Basiprionota Chevrolat, 1836]. Comment: usage of Basiprionotini Gressitt, 1952 conserved over this name (Art. 40.2).
Basiprionotini Gressitt, 1952: 444 [stem: Basiprionot-]. Type genus: Basiprionota Chevrolat, 1836. Comment: published 8 December 1952; name proposed to replace Priopterini because of synonymy of the type genus; this family-group name was also proposed in the same year by Hincks (1952 [31 December]: 329, in key, as Basiprionotini); usage of this name over the older name Priopterini Spaeth, 1929 conserved (Art. 40.2).
Epistictinini Hincks, 1952: 329, in key [stem: Epistictin-]. Type genus: Epistictina Hincks, 1950.
Tribe
Botryonopini
Chapuis, 1875
Botryonopites Chapuis, 1875: 291 [stem: Botryonop-]. Type genus: Botryonopa Guérin-Méneville, 1840 [Botryonopa is a incorrect subsequent spelling of the original name Bothryonopa, in prevailing usage and so deemed to be the correct original spelling (Art. 33.3.1); for correct author and year of type genus see Staines (2010: 172)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1911: 39, as Botryonopini), generally accepted as in Würmli (1975: 8, as Botryonopini).
Tribe
Callispini
Chapuis, 1875
Callispites Chapuis, 1875: 269 [stem: Callisp-]. Type genus: Callispa Baly, 1859. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1911: 41, as Callispini), generally accepted as in Würmli (1975: 12, as Callispini).
*Hispodontites Chapuis, 1875: 284 [stem: Hispodont-]. Type genus: Hispodonta Baly, 1859. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Tribe
Callohispini
Uhmann, 1960
Callohispini Uhmann, 1960: 60 [stem: Callohisp-]. Type genus: Callohispa Uhmann, 1960.
Tribe
Cassidini
Gyllenhal, 1813
Cassideae Gyllenhal, 1813: 434 [stem: Cassid-]. Type genus: Cassida Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1974b)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1974b, as Cassidinae Stephens, 1831).
Evaspidotidae Gistel, 1856a: 381 [stem: Evaspist-]. Type genus: Evaspistes Gistel, 1856 [syn. of Cassida Linnaeus, 1758]. Comment: incorrect original stem formation, not in prevailing usage.
*Chiridites Chapuis, 1875: 405 [stem: Chirid-]. Type genus: Chirida Chapuis, 1875. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Hybosites Chapuis, 1875: 380 [stem: Hybos-]. Type genus: Hybosa Boheman, 1855 [Hybosa has been credited to Duponchel and Chevrolat, 1842, e.g., Seeno and Wilcox (1982: 178), but the name is not available from that reference since there is no description nor available species included; this genus was first made available by Boheman (1855: 1)]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Basiptites Chapuis, 1875: 379 [stem: Basipt-]. Type genus: Basipta Chevrolat, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Hincks (1952: 337, as Basiptini).
Coptocyclitae Spaeth and Reitter, 1926: 7, in key [stem: Coptocycl-]. Type genus: Coptocycla Chevrolat, 1836. Comment: this family-group name was also used in the same year by Spaeth (1926: 1, as Coptocyclitae) but we could not establish priority.
Charidotitae Spaeth, 1942: 40 [stem: Charidotid-]. Type genus: Charidotis Boheman, 1854. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Hincks (1952: 343, as Charidotini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Tribe
Cephaloleiini
Chapuis, 1875
Céphaloléites Chapuis, 1875: 277 [stem: Cephalolei-]. Type genus: Cephaloleia Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1910: 69, as Cephaloleiini), generally accepted as in Borowiec (1995: 553, as Cephaloleiini); incorrect original stem formation, not in prevailing usage.
Tribe
Chalepini
Weise, 1910
Chalepini Weise, 1910: 69 [stem: Chalep-]. Type genus: Chalepus Thunberg, 1805. Comment: Chalepini Weise, 1910 is a junior homonym of Chalepidae H. C. C. Burmeister, 1847 (type genus Chalepus W. S. MacLeay, 1819) in Scarabaeidae; the older scarab family-group name is permanently invalid because it is based on a preoccupied type genus; an application to the Commission is needed to conserve usage of Chalepini Weise, 1910.
Tribe
Coelaenomenoderini
Weise, 1911
Coelaenomenoderini Weise, 1911: 51 [stem: Coelaenomenoder-]. Type genus: Coelaenomenodera Blanchard, 1845.
Pharangispini Uhmann, 1940: 122 [stem: Pharangisp-]. Type genus: Pharangispa Maulik, 1929. Comment: proposed after 1930 without description but available because accompanied by a bibliographic reference (Maulik 1929: 233) to a description (Art. 13.1).
Tribe
Cryptonychini
Chapuis, 1875
Cryptonychites Chapuis, 1875: 286 [stem: Cryptonych-]. Type genus: Cryptonychus Gyllenhal, 1817. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1911: 45, as Cryptonychini), generally accepted as in Würmli (1975: 21, as Cryptonychini).
Tribe
Delocraniini
Spaeth, 1929
Delocraniitae Spaeth, 1929b: 113 [stem: Delocrani-]. Type genus: Delocrania Guérin-Méneville, 1844.
Tribe
Dorynotini
Monrós and Viana, 1949 (1923)
*Batonotites Chapuis, 1875: 377 [stem: Batonot-]. Type genus: Batonota Hope, 1839 [syn. of Dorynota Chevrolat, 1836]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875).
Batonotitae Spaeth, 1923: 66 [stem: Batonot-]. Type genus: Batonota Hope, 1839 [syn. of Dorynota Chevrolat, 1836]. Comment: usage of Dorynotini Monrós and Viana, 1949 conserved over this name (Art. 40.2).
Dorynotini Monrós and Viana, 1949: 392 [stem: Dorynot-]. Type genus: Dorynota Chevrolat, 1836. Comment: name proposed to replace “Batonotites” because of synonymy of the type genus; usage of younger name conserved over Batonotini Spaeth, 1923 (Art. 40.2).
Tribe
Eugenysini
Hincks, 1952
*Calaspideitae Spaeth, 1942: 18 [stem: Calaspide-]. Type genus: Calaspidea Hope, 1840 [syn. of Eugenysa Chevrolat, 1836]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Eugenysini Hincks, 1952: 329, in key [stem: Eugenys-]. Type genus: Eugenysa Chevrolat, 1836.
Tribe
Eurispini
Chapuis, 1875
Eurispites Chapuis, 1875: 264 [stem: Eurisp-]. Type genus: Eurispa Baly, 1859. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1911: 45, as Eurispini), generally accepted as in Uhmann (1960: 61, as Eurispini).
Tribe
Exothispini
Weise, 1911
Exothispini Weise, 1911: 51 [stem: Exothisp-]. Type genus: Exothispa Kolbe, 1897.
Tribe
Goniocheniini
Spaeth, 1942
Goniocheniitae Spaeth, 1942: 17 [stem: Goniocheni-]. Type genus: Goniochenia Weise, 1896. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Viana (1964: 217, as Goniocheniici) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Tribe
Gonophorini
Chapuis, 1875
Gonophorites Chapuis, 1875: 303 [stem: Gonophor-]. Type genus: Gonophora Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1911: 54, as Gonophorini), generally accepted as in Würmli (1975: 47, as Gonophorini).
*Wallacéites Chapuis, 1875: 281 [stem: Wallace-]. Type genus: Wallacea Baly, 1859 [this genus is a senior homonym of Wallacea Doleschall, 1859 (see Woodley 2002: 410)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Wallaceiad[a]e [incorrect stem formation] was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Hemisphaerotini
Monrós and Viana, 1951 (1929)
Porphyraspitae Spaeth, 1929a: 29 [stem: Porphyraspid-]. Type genus: Porphyraspis Hope, 1840 [syn. of Hemisphaerota Chevrolat, 1836]. Comment: this family-group name was also used in the same year by Spaeth (1929b: 113); usage of younger name Hemisphaerotini Monrós and Viana, 1951 conserved over this name (Art. 40.2); incorrect original stem formation, not in prevailing usage.
Hemisphaerotini Monrós and Viana, 1951: 370 [stem: Hemisphaerot-]. Type genus: Hemisphaerota Chevrolat, 1836. Comment: usage of this name conserved over Porphyraspidini Spaeth, 1929 (Art. 40.2).
Tribe
Hispini
Gyllenhal, 1813
Hispoideae Gyllenhal, 1813: 448 [stem: Hisp-]. Type genus: Hispa Linnaeus, 1767.
*Monochirites Chapuis, 1875: 330 [stem: Monochir-]. Type genus: Monochirus Chapuis, 1875 [preoccupied genus name, not Monochirus Rafinesque, 1814 [Pisces]; syn. of Hispellinus Weise, 1898]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875).
*Trichispites Chapuis, 1875: 331 [stem: Trichisp-]. Type genus: Trichispa Chapuis, 1875. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Monochirini Weise, 1905: 317 [stem: Monochir-]. Type genus: Monochirus Chapuis, 1875 [preoccupied genus name, not Monochirus Rafinesque, 1814 [Pisces]; syn. of Hispellinus Weise, 1897]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Hispoleptini
Chapuis, 1875
Hispoleptites Chapuis, 1875: 283 [stem: Hispolept-]. Type genus: Hispopleptis Baly, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Uhmann (1940: 121, as Hispoleptini), generally accepted as in Staines (2002: 777, as Hispoleptini).
Tribe
Hybosispini
Weise, 1910
Hybosispini Weise, 1910: 69 [stem: Hybosisp-]. Type genus: Hybosispa Weise, 1910.
Tribe
Imatidiini
Hope, 1840
Imatidiidae Hope, 1840a: 152 [stem: Imatidi-]. Type genus: Imatidium Fabricius, 1801. Comment: name previously attributed to Chapuis (1875).
Tribe
Ischyrosonychini
Chapuis, 1875
Ischyrosonychites Chapuis, 1875: 382 [stem: Ischyrosonych-]. Type genus: Ischyrosonyx Sturm, 1843 [syn. of Eurypedus Gistel, 1834]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Spaeth (1942: 31, as Ischyronycitae [incorrect stem formation]), generally accepted as in Borowiec (1995: 556, as Ischyrosonychini).
*Physonotitae Spaeth, 1942: 32 [stem: Physonot-]. Type genus: Physonota Boheman, 1854. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Asterizini Hincks, 1952: 330, in key [stem: Asteriz-]. Type genus: Asteriza Chevrolat, 1836. Comment: Hincks (1952: 336) and Seeno and Wilcox (1982: 171) mention “Asterizitae Spaeth”, we could not find any earlier usage of Asterizitae by Spaeth.
Tribe
Leptispini
Fairmaire, 1868
Leptispites Fairmaire, 1868: 258 [stem: Leptisp-]. Type genus: Leptispa Baly, 1859. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1911: 44, as Lepthispini [incorrect stem formation]), generally accepted as in Würmli (1975: 18, as Leptispini).
Tribe
Mesomphaliini
Hope, 1840
Mesomphalidae Hope, 1840a: 160 [stem: Mesomphali-]. Type genus: Mesomphalia Hope, 1839. Comment: incorrect original stem formation, not in prevailing usage.
*Chélymorphites Chapuis, 1875: 402 [stem: Chelymorph-]. Type genus: Chelymorpha Chevrolat, 1836. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Elytrogonites Chapuis, 1875: 403 [stem: Elytrogon-]. Type genus: Elytrogona Chevrolat, 1836. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Omoplatites Chapuis, 1875: 397 [stem: Omoplat-]. Type genus: Omoplata Hope, 1840 [syn. of Echoma Chevrolat, 1836]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Stolaini Hincks, 1952: 329, in key [stem: Stolad-]. Type genus: Stolas Billberg, 1820. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Nothosacanthini
Gressitt, 1952 (1929)
*Hoplionotites Chapuis, 1875: 357 [stem: Hoplionot-]. Type genus: Hoplionota Hope, 1840 [syn. of Notosacantha Chevrolat, 1836]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875).
Hoplionotitae Spaeth, 1929b: 113 [stem: Hoplionot-]. Type genus: Hoplionota Hope, 1840 [syn. of Notosacantha Chevrolat, 1836]. Comment: usage of younger name Nothosacanthini Gressitt, 1952 conserved over this name (Art. 40.2).
Notosacanthina Gressitt, 1952: 444 [stem: Notosacanth-]. Type genus: Notosacantha Chevrolat, 1836. Comment: published 8 December 1952; name proposed to replace Hoplionotini Spaeth, 1929 because of the synonymy of the type genus; this family-group name was also proposed in the same year by Hincks (1952 [31 December]: 328, in key, as Notosacanthini); usage of this name conserved over Hoplionotini Spaeth, 1929 (Art. 40.2).
Tribe
Oediopalpini
Monrós and Viana, 1947 (1910)
Amplipalpini Weise, 1910: 69 [stem: Amplipalp-]. Type genus: Amplipalpa Harold, 1875 [syn. of Oediopalpa Baly, 1859]. Comment: younger name Oediopalpini Monrós and Viana, 1947 conserved over this name (Art. 40.2).
Oediopalpini Monrós and Viana, 1947: 150, in key [stem: Oediopalp-]. Type genus: Oediopalpa Baly, 1859. Comment: name proposed to replace Amplipalpini Weise, 1910 because of synonymy of the type genus; usage of this name conserved over the older name Amplipalpini Weise, 1910 (Art. 40.2).
Tribe
Omocerini
Hincks, 1952 (1923)
*Tauromites Chapuis, 1875: 372 [stem: Taurom-]. Type genus: Tauroma Hope, 1839 [syn. of Omocerus Chevrolat, 1835]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875).
Tauromitae Spaeth, 1923: 66 [stem: Taurom-]. Type genus: Tauroma Hope, 1839 [syn. of Omocerus Chevrolat, 1835]. Comment: younger name Omocerini Hincks, 1952 conserved over this name (Art. 40.2).
Omocerini Hincks, 1952: 329, in key [stem: Omocer-]. Type genus: Omocerus Chevrolat, 1835. Comment: name proposed to replace Tauromini Spaeth, 1923 because of synonymy of the type genus; usage of younger name conserved over Tauromini Spaeth, 1923 (Art. 40.2).
Tribe
Oncocephalini
Chapuis, 1875
Oncocéphalites Chapuis, 1875: 308 [stem: Oncocephal-]. Type genus: Onchocephala Guérin-Méneville, 1844 [this genus name was first made available by Guérin-Méneville (1844: 280-281) although most authors credit Chevrolat (1847: 110, spelled Oncocephalus) as the correct author and date of publication of this genus]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1911: 50, as Oncocephalini), generally accepted as in Świętojańska et al. (2006: 49, as Oncocephalini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Onchocephal-).
Choeridionini Weise, 1911: 49 [stem: Chaeridion-]. Type genus: Chaeridiona Baly, 1869 [as Choeridiona, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Promecothecini
Chapuis, 1875
Promecothécites Chapuis, 1875: 300 [stem: Promecothec-]. Type genus: Promecotheca Chevrolat, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1911: 53, as Promecothecini), generally accepted as in Gressitt (1950: 81, as Promecothecini).
Tribe
Prosopodontini
Weise, 1910
Prosopodontini Weise, 1910: 69 [stem: Prosopodont-]. Type genus: Prosopodonta Baly, 1885.
Tribe
Sceloenoplini
Uhmann, 1930
*Céphalodontites Chapuis, 1875: 313 [stem: Cephalodont-]. Type genus: Cephalodonta Chevrolat, 1842 [this name was recently attributed to “Chevrolat, 1843” (e.g., Staines (2002: 748)) but it was made available for the first time by Chevrolat (1842: 272); syn. of Sceloenopla Chevrolat, 1836]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875).
Cephalodontinae Gestro, 1906: 548 [stem: Cephalodont-]. Type genus: Cephalodonta Chevrolat, 1842 [this name was recently attributed to “Chevrolat, 1843” (e.g., Staines (2002: 748)) but it was made available for the first time by Chevrolat (1842: 272); syn. of Sceloenopla Chevrolat, 1836]. Comment: usage of the younger name Sceloenoplini Uhmann, 1930 conserved over this name (Art. 40.2).
Sceloenoplini Uhmann, 1930: 238 [stem: Sceloenopl-]. Type genus: Sceloenopla Chevrolat, 1836. Comment: although this is not the oldest name for the tribe, we recommend that an application be sent to the Commission in order to conserve usage of Sceloenoplini Uhmann, 1930.
Tribe
Spilophorini
Chapuis, 1875
Spilophorites Chapuis, 1875: 364 [stem: Spilophor-]. Type genus: Spilophora Boheman, 1850. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Spaeth (1929a: 29, as Spilophoritae), generally accepted as in Borowiec (1995: 553, as Spilophorini); this name is a senior homonym of Spilophorina Krikken, 1984 (type genus Spilophorus Schaum, 1848) currently used as valid in Scarabaeidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Uroplatini
Weise, 1910
*Octotomites Chapuis, 1875: 310 [stem: Octotom-]. Type genus: Octotoma Dejean, 1836. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Octotomidae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Uroplatini Weise, 1910: 69 [stem: Uroplat-]. Type genus: Uroplata Chevrolat, 1836 [“Uroplata Chevrolat, 1835” placed on the Official List of Generic Names in Zoology (ICZN 1985f)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1985f, as Uroplatini Leng, 1920).
Subfamily
Chrysomelinae
Latreille, 1802
Chrysomelinae Latreille, 1802: 220 [stem: Chrysomel-]. Type genus: Chrysomela Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1984c)]. Comment: see Kippenberg (2010b) for an alternative tribal classification within this subfamily.
Tribe
Chrysomelini
Latreille, 1802
Chrysomelinae Latreille, 1802: 220 [stem: Chrysomel-]. Type genus: Chrysomela Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1984c)].
Eleiaeidae Gistel, 1848: [9] [stem: Elei-]. Type genus: Eleia Gistel, 1848 [syn. of Chrysomela Linnaeus, 1758]. Comment: incorrect original stem formation, not in prevailing usage.
Prasocurisidae Gistel, 1848: [10] [stem: Prasocur-]. Type genus: Prasocuris Latreille, 1802. Comment: incorrect original stem formation, not in prevailing usage.
Chloëmeladae Gistel, 1856a: 379 [stem: Chloemel-]. Type genus: Chloemela Gistel, 1856 [syn. of Chrysomela Linnaeus, 1758]. Comment: incorrect original stem formation, not in prevailing usage.
Doryphores Motschulsky, 1860: 181 [stem: Doryphor-]. Type genus: Doryphora Illiger, 1807. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gerstaecker (1862: 405, as Doryphorae), generally accepted as in Riley et al. (2002: 639, as Doryphorina).
Gonioctènes Motschulsky, 1860: 179 [stem: Goniocten-]. Type genus: Gonioctena Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gerstaecker (1862: 405, as Gonioctenae), generally accepted as in Warchalowski (1994: 100, as Gonioctenina).
Linaeines Motschulsky, 1860: 196 [stem: Lin-]. Type genus: Lina Latreille, 1829 [syn. of Chrysomela Linnaeus, 1758]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Gerstaecker (1862: 406, as Linae).
Ovosomes Motschulsky, 1860: 211 [stem: Ovosomat-]. Type genus: Ovosoma Motschulsky, 1860. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Gerstaecker (1862: 406, as Ovosomae); incorrect original stem formation, not in prevailing usage.
Paropsines Motschulsky, 1860: 192 [stem: Paropse-]. Type genus: Paropsis A. G. Olivier, 1807. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gerstaecker (1862: 405, as Paropsinae), generally accepted as in Riley et al. (2002: 639, as Paropsina); incorrect original stem formation, not in prevailing usage.
Phrathorines Motschulsky, 1860: 216 [stem: Phrator-]. Type genus: Phratora Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gerstaecker (1862: 406, as Phratorinae), generally accepted as in Warchalowski (1994: 139, as Phratorina); incorrect original stem formation, not in prevailing usage.
*Australicites Chapuis, 1874: 428 [stem: Australic-]. Type genus: Australica Chevrolat, 1836 [this name is sometimes treated as a nomen nudum (Reid 2006: 55), however it is available since four available species names were included by Chevrolat (1836)]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Clidonotites Chapuis, 1874: 414 [stem: Clidonot-]. Type genus: Clidonotus Chapuis, 1874. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1874); Clidonotidae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1874); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Colaspidémites Chapuis, 1874: 364 [stem: Colaspidem-]. Type genus: Colaspidema Laporte, 1833. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1874); Colaspidemidae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1874); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Cyrtonites Chapuis, 1874: 364 [stem: Cyrton-]. Type genus: Cyrtonus Latreille, 1829. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Elytrosphoerites Chapuis, 1874: 406 [stem: Elytrosphoer-]. Type genus: Elytrosphoera Blanchard, 1845. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; name incorrectly spelled Elytrosphaerites in the index of the same work (page 449).
Entomoscélites Chapuis, 1874: 418 [stem: Entomoscelid-]. Type genus: Entomoscelis Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte and G. H. Horn (1883: 344, as Entomoscelides [treated as Latin]), generally accepted as in Warchalowski (1994: 154, as Entomoscelina); incorrect original stem formation, not in prevailing usage.
*Lycariites Chapuis, 1874: 420 [stem: Lycari-]. Type genus: Lycaria Stål, 1857. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Phyllocharites Chapuis, 1874: 422 [stem: Phyllocharit-]. Type genus: Phyllocharis Dalman, 1824. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Weise (1915: 436, as Phyllocharini); incorrect original stem formation, not in prevailing usage.
*Pyxites Chapuis, 1874: 438 [stem: Pyxid-]. Type genus: Pyxis Chevrolat, 1843. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.
Phyllodectae J. L. LeConte and G. H. Horn, 1883: 344 [stem: Phyllodect-]. Type genus: Phyllodecta Kirby, 1837 [subgenus of Phratora Chevrolat, 1836].
Dicranosternini Weise, 1915: 436 [stem: Dicranostern-]. Type genus: Dicranosterna Motschulsky, 1860.
Phaedonini Weise, 1915: 435 [stem: Phaedon-]. Type genus: Phaedon Latreille, 1829.
Zygogrammini Weise, 1915: 435 [stem: Zygogrammat-]. Type genus: Zygogramma Chevrolat, 1836. Comment: incorrect original stem formation, not in prevailing usage.
Chrysolinina Chen, 1936: 64, in key [stem: Chrysolin-]. Type genus: Chrysolina Motschulsky, 1860 [placed on the Official List of Generic Names in Zoology (ICZN 1984c)].
Barymelini Bechyné, 1948: 295 [stem: Barymel-]. Type genus: Barymela Weise, 1910.
Doryphorini Bechyné, 1950a: 115, in key [stem: Doryphor-]. Type genus: Doryphora Illiger, 1807. Comment: family-group name proposed as new without reference to Doryphores Motschulsky, 1860.
Monarditini Bechyné, 1953: 85 [stem: Monardit-]. Type genus: Monardita Bechyné, 1948.
Oreinini Bechyné, 1958: 218, nota [stem: Orein-]. Type genus: Oreina Chevrolat, 1836. Comment: the older name Oreinini Bleeker, 1863 (type genus Oreinus M’Clelland, 1839) is available in Pisces; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
*Phytodectini Bechyné, 1980: 58 [stem: Phytodect-]. Type genus: Phytodecta Kirby, 1837 [syn. of Gonioctena Chevrolat, 1836]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Hispostomini Daccordi, 1981: 181, in key [stem: Hispostomat-]. Type genus: Hispostoma Weise, 1907. Comment: incorrect original stem formation, not in prevailing usage.
Sphaeratrixini Daccordi, 1981: 181, in key [stem: Sphaeratrix-]. Type genus: Sphaeratrix Gistel, 1848. Comment: the correct stem based on Sphaeratrix is unclear since Gistel did not specify the etymology of his genus therefore we accept the original stem as correct.
*Gastrophysina Steinhausen, 2001: 47 [stem: Gastrophys-]. Type genus: Gastrophysa Chevrolat, 1836. Comment: unavailable family-group name, proposed after 1999 without explicit intention (Art. 16.1).
Gastrophysina Kippenberg, 2010a: 68 [stem: Gastrophys-]. Type genus: Gastrophysa Chevrolat, 1836.
Tribe
Timarchini
Motschulsky, 1860
Timarchaeines Motschulsky, 1860: 187 [stem: Timarch-]. Type genus: Timarcha Samouelle, 1819. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Gerstaecker (1862: 405, as Timarchae), generally accepted as in Burakowski et al. (1990: 115, as Timarchini); incorrect original stem formation, not in prevailing usage.
Subfamily
Galerucinae
Latreille, 1802
Galerucae Latreille, 1802: 228 [stem: Galeruc-]. Type genus: Galeruca Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Alticini
Newman, 1834
Halticites Newman, 1834: 421 [stem: Altic-]. Type genus: Altica Geoffroy, 1762 [as Haltica, unjustified emendation of type genus name by Illiger (1801), not in prevailing usage; Altica placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: incorrect original stem formation, not in prevailing usage.
Hesperidae Swainson, 1840: 310 [stem: Hesper-]. Type genus: Hespera Weise, 1889.
Longitarses Maehler, 1850: 7 [stem: Longitars-]. Type genus: Longitarsus Latreille, 1829.
Plectroscelides C. G. Thomson, 1866: 213 [stem: Plectroscelid-]. Type genus: Plectroscelis Chevrolat, 1836. Comment: incorrect original stem formation, not in prevailing usage.
*Acrocryptites Chapuis, 1875: 36 [stem: Acrocrypt-]. Type genus: Acrocrypta Baly, 1862. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Acrocryptidae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Amphimélites Chapuis, 1875: 34 [stem: Amphimel-]. Type genus: Amphimela Chapuis, 1875. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently used in latinized form and treated as valid.
Aphthonites Chapuis, 1875: 69 [stem: Aphthon-]. Type genus: Aphthona Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte and G. H. Horn (1883: 352, as Aphthonae), generally accepted as in Bechyné (1968: 1708, as Aphthonini).
Arsipodites Chapuis, 1875: 37 [stem: Arsipod-]. Type genus: Arsipoda Erichson, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte and G. H. Horn (1883: 353, as Arsipodes [treated as Latin]), generally accepted as in Bechyné and Špringlová de Bechyné (1973: 26, as Arsipodini).
Aspicélites Chapuis, 1875: 75 [stem: Aspicel-]. Type genus: Aspicela Dejean, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in G. H. Horn (1889: 166, as Aspicelae).
Blépharidites Chapuis, 1875: 26 [stem: Blepharid-]. Type genus: Blepharida Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte and G. H. Horn (1883: 350, as Blepharidae [incorrect stem formation]), generally accepted as in Bechyné (1968: 1724, as Blepharidini).
Crépidodérites Chapuis, 1875: 51 [stem: Crepidoder-]. Type genus: Crepidodera Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte and G. H. Horn (1883: 350, as Crepidoderae).
*Diamphidiites Chapuis, 1875: 24 [stem: Diamphidi-]. Type genus: Diamphidia Gerstaecker, 1855. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Diamphidiini and Diamphidiidae were subsequently used as valid by Bechyné (1980: 58) and Ienistea (1986: 31) but those names were not attributed to Chapuis (1875); Bechyné’s and Ienistea’s names are also unavailable, they were proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Diboliites Chapuis, 1875: 137 [stem: Diboli-]. Type genus: Dibolia Latreille, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte and G. H. Horn (1883: 350, as Diboliae), generally accepted as in Bechyné (1997: 206, as Diboliini).
*Élithiites Chapuis, 1875: 21 [stem: Elithi-]. Type genus: Elithia Chapuis, 1875. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); this name was originally spelled Elithiites in the key but Élithiides in description (both p. 21); Elithiidae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Lacticites Chapuis, 1875: 123 [stem: Lactic-]. Type genus: Lactica Erichson, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte and G. H. Horn (1883: 350, as Lacticae).
Mniophilites Chapuis, 1875: 129 [stem: Mniophil-]. Type genus: Mniophila Stephens, 1831. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte and G. H. Horn (1883: 350, as Mniophilae).
Monoplatites Chapuis, 1875: 91 [stem: Monoplat-]. Type genus: Monoplatus Chevrolat, 1846 [this genus is usually credited to Clark, 1860, e.g., Seeno and Wilcox (1982: 141) and Linzmeier and Konstantinov (2009), but the name was first made available with a short description by Chevrolat (1846: 333); syn. of Sphaeronychus Dejean, 1836]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte and G. H. Horn (1883: 350, as Monoplati), generally accepted as in Bechyné and Špringlová de Bechyné (1975: 132, as Monoplatini); treated as a valid tribe and given precedence over Sphaeronychini (Art. 40.1) by Linzmeier and Konstantinov (2009: 657).
*Nonarthrites Chapuis, 1875: 141 [stem: Nonarthr-]. Type genus: Nonarthra Baly, 1862. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Nonarthridae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Oedionychites Chapuis, 1875: 81 [stem: Oedionych-]. Type genus: Oedionychis Latreille, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte and G. H. Horn (1883: 349, as Oedionyches [treated as Latin]), generally accepted as in Bechyné (1968: 1703, as Oedionychini).
Oxygonites Chapuis, 1875: 43 [stem: Oxygon-]. Type genus: Oxygona Chevrolat, 1847 [this genus has been attributed to “Chevrolat, 1837” (see Seeno and Wilcox 1982: 132) but the name is not available from Dejean’s second and third editions of his catalogue; it was first made available by Chevrolat (1847: 368)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Jacoby (1884a: 34, as Oxygoninae).
*Procalites Chapuis, 1875: 175 [stem: Procal-]. Type genus: Procalus Clark, 1865. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Procalidae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Psylliodites Chapuis, 1875: 140 [stem: Psylliod-]. Type genus: Psylliodes Latreille, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in J. L. LeConte and G. H. Horn (1883: 349, as Psylliodes [treated as Latin]).
Chaetocnemae J. L. LeConte and G. H. Horn, 1883: 354 [stem: Chaetocnem-]. Type genus: Chaetocnema Stephens, 1831.
Disonychae J. L. LeConte and G. H. Horn, 1883: 351 [stem: Disonych-]. Type genus: Disonycha Chevrolat, 1836.
Euplectrosceles G. H. Horn, 1889: 167, in key [stem: Euplectroscelid-]. Type genus: Euplectroscelis Crotch, 1873. Comment: incorrect original stem formation, not in prevailing usage.
Pseudolampses G. H. Horn, 1889: 166, in key [stem: Pseudolampse-]. Type genus: Pseudolampsis Horn, 1889. Comment: incorrect original stem formation, not in prevailing usage.
Systenae G. H. Horn, 1889: 167, in key [stem: Systen-]. Type genus: Systena Chevrolat, 1836.
Luperalticini Leng, 1920: 301 [stem: Luperaltic-]. Type genus: Luperaltica Crotch, 1873.
Octogonotini Weise, 1921: 151 [stem: Octogonot-]. Type genus: Octogonotes Drapiez, 1820.
Serraticollini B. E. White, 1942: 17 [stem: Serraticoll-]. Type genus: Serraticollis B. E. White, 1942.
Sphaeronychini Bechyné and Špringlová de Bechyné, 1960: 7 [stem: Sphaeronych-]. Type genus: Sphaeronychus Dejean, 1836 [Sphaeronychus is an incorrect subsequent spelling of Sphraeronychus Dejean, 1836, in prevailing usage, treated as correct original spelling (Art. 33.3.1)]. Comment: this taxon was treated as a valid tribe (which included Monoplatini) by Furth (2007: 90).
Disonychina Bechyné and Špringlová de Bechyné, 1966: 142, in key [stem: Disonych-]. Type genus: Disonycha Chevrolat, 1836. Comment: family-group name proposed as new without reference to Disonychae J. L. LeConte and G. H. Horn, 1883.
Cacoscelini Bechyné, 1968: 1715 [stem: Cacoscelid-]. Type genus: Cacoscelis Chevrolat, 1836. Comment: Cacoscelini J. Thomson, 1861 (type genus Cacosceles Newman, 1838) is available in Cerambycidae and used as valid; the correct stem Cacoscelid- should be used for the alticine name in the future in order to avoid homonymy with the cerambycid name.
*Hermaeophagina Bechyné, 1968: 1698 [stem: Hermaeophag-]. Type genus: Hermaeophaga Foudras, 1859. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Longitarsini Bechyné, 1968: 1708 [stem: Longitars-]. Type genus: Longitarsus Latreille, 1829. Comment: family-group name proposed as new without reference to Longitarses Maehler, 1850.
Lypneina Bechyné, 1968: 1717 [stem: Lypne-]. Type genus: Lypnea Baly, 1876.
Podagricini Bechyné, 1968: 1718 [stem: Podagric-]. Type genus: Podagrica Chevrolat, 1836.
*Sphaerodermini Bechyné, 1968: 1702 [stem: Sphaerodermat-]. Type genus: Sphaeroderma Stephens, 1831. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Wittmeralticina Bechyné, 1968: 1717 [stem: Wittmeraltic-]. Type genus: Wittmeraltica Bechyné, 1956.
Psilaphina Bechyné and Špringlová de Bechyné, 1973: 27, in key [stem: Psilaph-]. Type genus: Psilapha Clark, 1865.
Diphaulacini Bechyné and Špringlová de Bechyné, 1975: 116 [stem: Diphaulac-]. Type genus: Diphaulaca Chevrolat, 1836.
Manobiini Bechyné and Špringlová de Bechyné, 1975: 132, in key [stem: Manobi-]. Type genus: Manobia Jacoby, 1885.
Marcapatiini Bechyné and Špringlová de Bechyné, 1975: 131 [stem: Marcapati-]. Type genus: Marcapatia Bechyné, 1958.
Monomacrina Bechyné and Špringlová de Bechyné, 1975: 27 [stem: Monomacr-]. Type genus: Monomacra Chevrolat, 1836.
Systenini Bechyné and Špringlová de Bechyné, 1975: 132, in key [stem: Systen-]. Type genus: Systena Chevrolat, 1836. Comment: family-group name proposed as new without reference to Systenae G. H. Horn, 1889.
*Phygasiini Bechyné, 1980: 58 [stem: Phygasi-]. Type genus: Phygasia Dejean, 1836. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Ocnoscelina Seeno and Wilcox, 1982: 127 [stem: Ocnoscelid-]. Type genus: Ocnoscelis Erichson, 1847. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Hermaeophagina Bechyné, 1997: 116 [stem: Hermaeophag-]. Type genus: Hermaeophaga Foudras, 1859.
Ocnoscelina Bechyné, 1997: 124 [stem: Ocnoscelid-]. Type genus: Ocnoscelis Erichson, 1847. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Decarthrocerini
Laboissière, 1937
Decarthrocerina Laboissière, 1937: 29 [stem: Decarthrocer-]. Type genus: Decarthrocera Laboissière, 1937.
Tribe
Galerucini
Latreille, 1802
Galerucae Latreille, 1802: 228 [stem: Galeruc-]. Type genus: Galeruca Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Coelomérites Chapuis, 1875: 196 [stem: Coelomer-]. Type genus: Coelomera Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Leng (1920: 296, as Coelomerini), generally accepted as in Riley et al. (2002: 639, as Coelomerites [treated as Latin]).
Atysites Chapuis, 1875: 192 [stem: Atys-]. Type genus: Atysa Baly, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Leng (1920: 296, as Atysini), generally accepted as in Riley et al. (2002: 639, as Atysites [treated as Latin]).
Apophyliites Chapuis, 1875: 182 [stem: Apophyli-]. Type genus: Apophylia Duponchel and Chevrolat, 1841 [this genus name has been attributed to “J. Thomson, 1858”, e.g., Bezdĕk (2003: 71), however it was made available for the first time with a description by Duponchel and Chevrolat (1841: 31)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1923: 124, as Apophydiini [incorrect stem formation]), generally accepted as in Seeno and Wilcox (1982: 96, as Apophyliites [treated as Latin]).
Rupiliites Chapuis, 1875: 213 [stem: Rupili-]. Type genus: Rupilia Clark, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Jacoby (1899: 83, as Rupilinae [incorrect stem formation]).
Schematizites Chapuis, 1875: 195 [stem: Schematiz-]. Type genus: Schematiza Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Wilcox (1971: 101, as Schematizites [treated as Latin]), generally accepted as in Riley et al. (2002: 639, as Schematizites [treated as Latin]).
Mombasicites Laboissière, 1922: 230 [stem: Mombasic-]. Type genus: Mombasica Fairmaire, 1888.
Chorini Weise, 1923: 124 [stem: Chorin-]. Type genus: Chorina Baly, 1866. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Weise (1924: 179).
Leptosonychini Weise, 1924: 180 [stem: Leptosonych-]. Type genus: Leptosonyx Weise, 1885.
Theonina Laboissière, 1934: 7, in key [stem: Theon-]. Type genus: Theone Gistel, 1857.
Tribe
Hylaspini
Chapuis, 1875
Hylaspites Chapuis, 1875: 237 [stem: Hylasp-]. Type genus: Hylaspes Baly, 1865. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Laboissière (1934: 8, as Hylaspina), generally accepted as in Silfverberg (1990: 120, as Hylaspini); First Reviser found (Hylaspini Chapuis, 1875 vs Agelasticini Chapuis, 1875) is Silfverberg (1990: 120).
Sermylites Chapuis, 1875: 224 [stem: Sermyl-]. Type genus: Sermyla Chapuis, 1875 [preoccupied genus name, not Sermyla Walker 1854 [Lepidoptera], not Sermyla Adams, 1854 [Mollusca]; syn. of Sermylassa Reitter, 1912]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby (1884a: 64, as Sermylinae), generally accepted as in Seeno and Wilcox (2002: 104, as Sermylites [treated as Latin]); permanently invalid (Art. 39): based on preoccupied type genus.
Antiphites Chapuis, 1875: 232 [stem: Antiph-]. Type genus: Antipha Baly, 1865 [preoccupied genus name, not Antipha Walker, 1855 [Lepidoptera]; syn. of Dercetina Gressitt and Kimoto, 1963]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Seeno and Wilcox (1982: 103, as Antiphites [treated as Latin]); permanently invalid (Art. 39): based on preoccupied type genus.
Agelasticites Chapuis, 1875: 167 [stem: Agelastic-]. Type genus: Agelastica Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Weise (1903a: 209, as Agelasticites [treated as Latin]), generally accepted as in Burakowski et al. (1991: 23, as Agelasticini).
Agelasini Leng, 1920: 298 [stem: Agelas-]. Type genus: Agelasa sensu Horn, 1893 [not Agelasa Motschulsky, 1860; syn. of Sermylassa Reitter, 1912]. Comment: based on a misidentified type genus.
Bonesiites Laboissière, 1926: 91 [stem: Bonesi-]. Type genus: Bonesia Baly, 1865.
Capulini Ogloblin, 1936: 341 [stem: Capul-]. Type genus: Capula Jakobson, 1925.
Gallerucidini Gressitt and Kimoto, 1963: 390, in key [stem: Gallerucid-]. Type genus: Gallerucida Motschulsky, 1861.
Sermylassini Mroczkowski, 1991: 22 [stem: Sermylass-]. Type genus: Sermylassa Reitter, 1912. Comment: replacement name for Sermylites Chapuis, 1875 and Agelasini Leng, 1920 because of the homonymy of the type genus.
Tribe
Luperini
Gistel, 1848
Luperiidae Gistel, 1848: [9] [stem: Luper-]. Type genus: Luperus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1984a)]. Comment: name previously attributed to Chapuis (1875) in the literature; also see Beenen (2010: 464) for an alternative classification within this tribe; incorrect original stem formation, not in prevailing usage.
Aulacophorites Chapuis, 1875: 158 [stem: Aulacophor-]. Type genus: Aulacophora Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Laboissière (1934: 8, as Aulacophorina), generally accepted as in Seeno and Wilcox (1982: 97, as Aulacophorites [treated as Latin]).
Diabroticites Chapuis, 1875: 165 [stem: Diabrotic-]. Type genus: Diabrotica Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Leng (1920: 297, as Diabroticini), generally accepted as in Riley et al. (2002: 639, as Diabroticina).
Cérotomites Chapuis, 1875: 229 [stem: Cerotom-]. Type genus: Cerotoma Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Leng (1920: 298, as Cerotomini), generally accepted as in Riley et al. (2002: 639, as Cerotomites [treated as Latin]).
Platyxanthites Chapuis, 1875: 243 [stem: Platyxanth-]. Type genus: Platyxantha Baly, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Jacoby (1884b: 226, as Platyxanthinae).
*Théopéites Chapuis, 1875: 241 [stem: Theope-]. Type genus: Theopea Baly, 1864. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Theopeidae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); Theopeina Clench, 1955 (type genus Theope Doubleday, 1847) is available in Lepidoptera.
Scelidites Chapuis, 1875: 184 [stem: Scelid-]. Type genus: Scelida Chapuis, 1875. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby (1884a: 62, as Scelidinae), generally accepted as in Riley et al. (2002: 639, as Scelidites [treated as Latin]).
Phyllobroticites Chapuis, 1875: 163 [stem: Phyllobrotic-]. Type genus: Phyllobrotica Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Leng (1920: 297, as Phyllobroticini), generally accepted as in Riley et al. (2002: 639, as Phyllobroticites [treated as Latin]).
*Agetocérites Chapuis, 1875: 177 [stem: Agetocer-]. Type genus: Agetocera Hope, 1831 [Agetocera is an unjustified emendation of the original genus name Aegelocerus Hope, 1831 by Hope (1840a: 170); because Agetocera is in prevailing usage and has been attributed to the original author and date, e.g. Bezdĕk (2010a: 73, 2010c: 464) it is treated here as a justified emendation (Art. 33.2.3.1)]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Agetoceridae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Mimastrites Chapuis, 1875: 178 [stem: Mimastr-]. Type genus: Mimastra Baly, 1865. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby (1884a: 67, as Mimastrinae), generally accepted as in Weise (1903b: 334, as Mimastrites [treated as Latin]).
Ornithognathites Chapuis, 1875: 176 [stem: Ornithognath-]. Type genus: Ornithognathus J. Thomson, 1858. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Harold (1877: 110, as Ornithognathinae), generally accepted as in Seeno and Wilcox (1982: 97, as Ornithognathites [treated as Latin]).
Monoleptites Chapuis, 1875: 234 [stem: Monolept-]. Type genus: Monolepta Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby (1884a: 61, as Monoleptinae), generally accepted as in Riley et al. (2002: 639, as Monoleptites [treated as Latin]).
Cerophysites Chapuis, 1875: 181 [stem: Cerophys-]. Type genus: Cerophysa Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Seeno and Wilcox (1982: 97, as Cerophysides [treated as Latin]).
Phyllecthrites G. H. Horn, 1893: 60, in key [stem: Phyllecthr-]. Type genus: Phyllecthris Dejean, 1836.
Rhaphidopalpini Weise, 1902: 140 [stem: Raphidopalp-]. Type genus: Raphidopalpa Chevrolat, 1836 [as Rhaphidopalpa, unjustified emendation of type genus name by Rosenhauer (1856: 325), not in prevailing usage; syn. of Aulacophora Chevrolat, 1836]. Comment: incorrect original stem formation, not in prevailing usage.
Androlyperini Leng, 1920: 298 [stem: Androlyper-]. Type genus: Androlyperus Crotch, 1873.
Idacanthites Laboissière, 1921: 63 [stem: Idacanth-]. Type genus: Idacantha Fairmaire, 1869.
Hyperacanthites Laboissière, 1924: 142 [stem: Hyperacanth-]. Type genus: Hyperacantha Chapuis, 1879.
Megalognathites Laboissière, 1926: 185 [stem: Megalognath-]. Type genus: Megalognatha Baly, 1878.
*Exosomites Wilcox, 1965: 93 [stem: Exosomat-]. Type genus: Exosoma Jacoby, 1903. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
*Trachyscelidites Wilcox, 1972: 430 [stem: Trachyscelid-]. Type genus: Trachyscelida G. H. Horn, 1893. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Adoxiites Wilcox, 1973: 433 [stem: Adoxi-]. Type genus: Adoxia Broun, 1880. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Eumeleptites Wilcox, 1973: 447 [stem: Eumelept-]. Type genus: Eumelepta Jacoby, 1892. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Xenodites Wilcox, 1973: 604 [stem: Xenod-]. Type genus: Xenoda Baly, 1877. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
*Doryscites Wilcox, 1973: 606 [stem: Dorysc-]. Type genus: Doryscus Jacoby, 1887. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Metacyclini
Chapuis, 1875
Métacyclites Chapuis, 1875: 212 [stem: Metacycl-]. Type genus: Metacycla Baly, 1861. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Leng (1920: 298, as Metacyclini), generally accepted as in Riley et al. (2002: 639, as Metacyclini).
Stictocemites Laboissière, 1922: 220 [stem: Stictocem-]. Type genus: Stictocema Jacoby, 1906.
*Exorini Wilcox, 1965: 179 [stem: Exor-]. Type genus: Exora Chevrolat, 1836. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Oidini
Laboissière, 1921 (1875)
Adoriites Chapuis, 1875: 155 [stem: Adori-]. Type genus: Adorium Fabricius, 1801 [syn. of Oides Weber, 1801]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lea (1925: 16, as Adoriites [treated as Latin]); usage of younger name Oidini Laboissière, 1921 conserved (Art. 40.2).
Oïdites Laboissière, 1921: 38 [stem: Oid-]. Type genus: Oides Weber, 1801. Comment: usage of this name conserved over Adoriini Chapuis, 1875 (Art. 40.2).
Subfamily
Lamprosomatinae
Lacordaire, 1848
Lamprosomideae Lacordaire, 1848: 559 [stem: Lamprosomat-]. Type genus: Lamprosoma Kirby, 1819.
Tribe
Lamprosomatini
Lacordaire, 1848
Lamprosomideae Lacordaire, 1848: 559 [stem: Lamprosomat-]. Type genus: Lamprosoma Kirby, 1819. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Neochlamysini
Monrós, 1959
Neochlamysini Monrós, 1959b: 29 [stem: Neochlamys-]. Type genus: Neochlamys Jacoby, 1882. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Neochlamyd-).
Tribe
Sphaerocharini
Chapuis, 1874
Sphoerocharides Chapuis, 1874: 206 [stem: Sphaerochar-]. Type genus: Sphaerocharis Lacordaire, 1848 [as Sphoerocharis, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 248, as Sphaerocharini), generally accepted as in Monrós (1956: 32, as Sphaerocharini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Sphaerocharit-).
Subfamily
Cryptocephalinae
Gyllenhal, 1813
Cryptocephaloideae Gyllenhal, 1813: 582 [stem: Cryptocephal-]. Type genus: Cryptocephalus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Tribe
Clytrini
Kirby, 1837
Clythridae Kirby, 1837: 207 [stem: Clytr-]. Type genus: Clytra Laicharting, 1781 [as Clythra, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Megalostomideae Lacordaire, 1848: 486 [stem: Megalostomid-]. Type genus: Megalostomis Chevrolat, 1836. Comment: incorrect original stem formation, not in prevailing usage.
Babideae Lacordaire, 1848: 394 [stem: Babi-]. Type genus: Babia Chevrolat, 1836. Comment: incorrect original stem formation, not in prevailing usage.
Ischiopachites Chapuis, 1874: 151 [stem: Ischiopache-]. Type genus: Ischiopachys Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby and Clavareau (1906: 75, as Ischiopachitae), generally accepted as in Riley et al. (2002: 639, as Ischiopachina); incorrect original stem formation, not in prevailing usage.
Melolonthini Bedel, 1891: 106 [stem: Melolonth-]. Type genus: Melolontha Geoffroy, 1762 [preoccupied genus name, not Melolontha Fabricius, 1775 [Coleoptera: Scarabaeidae]; placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1994a); syn. of Clytra Laicharting, 1781]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Eoclytrini Monrós, 1958b: 35 [stem: Eoclytr-]. Type genus: Eoclytra Monrós, 1958.
Arateini Moldenke, 1981: 85, in key [stem: Arate-]. Type genus: Aratea Lacordaire, 1848.
Tribe
Cryptocephalini
Gyllenhal, 1813
Cryptocephaloideae Gyllenhal, 1813: 582 [stem: Cryptocephal-]. Type genus: Cryptocephalus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subtribe
Achaenopina
Chapuis, 1874
Achaenopites Chapuis, 1874: 171 [stem: Achaenop-]. Type genus: Achaenops Suffrian, 1857. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Clavareau (1913b: 109, as Achaenopini).
Subtribe
Cryptocephalina
Gyllenhal, 1813
Cryptocephaloideae Gyllenhal, 1813: 582 [stem: Cryptocephal-]. Type genus: Cryptocephalus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].
Subtribe
Monachulina
Leng, 1920
*Monachiden Suffrian, 1863: 79 [stem: Monach-]. Type genus: Monachus Chevrolat, 1836 [preoccupied genus name, not Monachus Fleming, 1822 [Mammalia], not Monachus Kaup, 1829 [Aves]; syn. of Lexiphanes Gistel, 1848]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Suffrian (1863).
Monachites Chapuis, 1874: 172 [stem: Monach-]. Type genus: Monachus Chevrolat, 1836 [preoccupied genus name, not Monachus Fleming, 1822 [Mammalia], not Monachus Kaup, 1829 [Aves]; syn. of Lexiphanes Gistel, 1848]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte and G. H. Horn (1883: 342, as Monachi), generally accepted as in Liu (1935: 286, as Monachini); permanently invalid (Art. 39): based on preoccupied type genus; Monachinae Trouessart, 1897 (type genus Monachus Fleming, 1822) is available in Mammalia.
Monachulini Leng, 1920: 290 [stem: Monachul-]. Type genus: Monachulus Leng, 1918 [syn. of Lexiphanes Gistel, 1848].
Subtribe
Pachybrachina
Chapuis, 1874
*Pachybrachiden Suffrian, 1863: 79 [stem: Pachybrach-]. Type genus: Pachybrachis Chevrolat, 1836. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Suffrian (1863).
Pachybrachites Chapuis, 1874: 163 [stem: Pachybrach-]. Type genus: Pachybrachis Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte and G. H. Horn (1883: 342, as Pachybrachi), generally accepted as in Burakowski et al. (1990: 63, as Pachybrachini).
Subtribe
Stylosomina
Chapuis, 1874
Stylosomites Chapuis, 1874: 162 [stem: Stylosom-]. Type genus: Stylosomus Suffrian, 1848. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Sharp (1876: 99, as Stylosomites [treated as Latin]), generally accepted as in Liu (1935: 292, as Stylosomini).
Tribe
Fulcidacini
Jakobson, 1924
*Chlamytes Blanchard, 1845b: 186 [stem: Chlamyd-]. Type genus: Chlamys Knoch, 1801 [preoccupied genus name, not Chlamys Bolten, 1798 [Mollusca]; syn. of Chlamisus Rafinesque, 1815]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Blanchard (1845b); incorrect original stem formation, not in prevailing usage.
Chlamydeae Lacordaire, 1848: 636 [stem: Chlamyd-]. Type genus: Chlamys Knoch, 1801 [preoccupied genus name, not Chlamys Bolten, 1798 [Mollusca]; syn. of Chlamisus Rafinesque, 1815]. Comment: incorrect original stem formation, not in prevailing usage; permanently invalid (Art. 39): based on preoccupied type genus.
Fulcidacina Jakobson, 1924: 239 [stem: Fulcidac-]. Type genus: Fulcidax Clavareau, 1913 [for comments on the authorship of this genus see Bezdĕk (2010b: 76); syn. of Poropleura Lacordaire, 1848]. Comment: we use Fulcidacini Jakobson, 1924 as valid for this taxon as in Bezdĕk (2010b: 76).
Chlamisinae Gressitt, 1946: 84 [stem: Chlamis-]. Type genus: Chlamisus Rafinesque, 1815. Comment: replacement name for Chlamydinae Lacordaire, 1848 because of the homonymy of the type genus.
Subfamily
Eumolpinae
Hope, 1840
Eumolpidae Hope, 1840a: 162 [stem: Eumolp-]. Type genus: Eumolpus Kugelann, 1798 [an application to suppress Eumolpus Kugelann, 1798 and conserve Eumolpus Weber, 1801 was recently submitted to the Commission by Moseyko et al. (2010)].
Tribe
Bromiini
Baly, 1865 (1863)
Adoxinae Baly, 1863: 146 [stem: Adox-]. Type genus: Adoxus Kirby, 1837 [syn. of Bromius Chevrolat, 1836]. Comment: use of younger name Bromiini Baly, 1865 conserved over this name (Art. 40.2).
Heteraspinae Baly, 1863: 147 [stem: Heteraspid-]. Type genus: Heteraspis Chevrolat, 1836 [this genus (incorrectly attributed to “Chevrolat, 1837”) has been treated as a synonym of Scelodonta Westwood, 1837 in the literature; Heteraspis was in fact made available by Chevrolat (1836: 413) and is a senior synonym of Scelodonta Westwood, 1837 (also see Löbl 2010: 83)]. Comment: although this is the oldest name for the tribe, we recommend that an application be sent to the Commission in order to conserve usage of Bromiini Baly, 1865; incorrect original stem formation, not in prevailing usage.
Bromiinae Baly, 1865: 438 [stem: Bromi-]. Type genus: Bromius Chevrolat, 1836 [an application to conserve Bromius Chevrolat, 1836, threatened by the older name Eumolpus Kugelann, 1798, was recently submitted to the Commission by Moseyko et al. (2010)]. Comment: use of family-group name conserved over Adoxiini Baly, 1863 (Art. 40.2).
Myochroinae Baly, 1865: 433 [stem: Myochro-]. Type genus: Myochrous Erichson, 1847.
Leprotites Chapuis, 1874: 268 [stem: Leprotet-]. Type genus: Leprotes Baly, 1863 [syn. of Fidia Motschulsky, 1860]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Lefèvre (1885: 71, as Leprotitae), generally accepted as in Riley et al. (2002: 639, as Leprotites [treated as Latin]); incorrect original stem formation, not in prevailing usage.
Pseudocolaspites Chapuis, 1874: 287 [stem: Pseudocolaspid-]. Type genus: Pseudocolaspis Laporte, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 84, as Pseudocolaspitae); incorrect original stem formation, not in prevailing usage.
Scelodontites Chapuis, 1874: 266 [stem: Scelodont-]. Type genus: Scelodonta Westwood, 1838 [syn. of Heteraspis Chevrolat, 1836]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Lefèvre (1885: 67, as Scelodontitae), generally accepted as in Riley et al. (2002: 639, as Scelodontites [treated as Latin]).
Tomyrites Chapuis, 1874: 264 [stem: Tomyri-]. Type genus: Tomyris Chapuis, 1874 [preoccupied genus name, not Tomyris Eichald, 1831 [Reptilia]; syn. of Eboo Reid, 1993]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 64, as Tomyritae); permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
*Goniopleurites Chapuis, 1875: 247 [stem: Goniopleur-]. Type genus: Goniopleura Westwood, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875); Goniopleuridae was used as valid by Ienistea (1986: 31) but it was not attributed to Chapuis (1875); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); usage of the name Goniopleuridae dates back at least to the 1850’s in Trilobita, however it is based on the junior homonym Goniopleura Hawle and Corda 1847 and the trilobite family-group name is therefore permanently invalid (Art. 39).
Eubrachini Jacoby, 1908: 432 [stem: Eubrach-]. Type genus: Eubrachis sensu Baly, 1878 [not Eubrachis Dejean, 1836; syn. of Macrocoma Chapuis, 1874]. Comment: based on misidentified type genus.
Nerissini Kuntzen, 1912: 42 [stem: Neriss-]. Type genus: Nerissus Chapuis, 1874.
Cynoini Clavareau, 1914: 109 [stem: Cynoi-]. Type genus: Cyno T. A. Marshall, 1865. Comment: incorrect original stem formation, not in prevailing usage.
Odontionopini Clavareau, 1914: 63 [stem: Odontionop-]. Type genus: Odontionopa Erichson, 1842 [preoccupied genus name, not Odontionopa Chevrolat, 1836 [Coleoptera: Chrysomelidae: Eumolpinae: Euryopini]; syn. of Eboo Reid, 1993]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Trichochryseini Clavareau, 1914: 82 [stem: Trichochryse-]. Type genus: Trichochrysea Baly, 1861.
Lypesthini Chûjô, 1956: 87 [stem: Lypesthet-]. Type genus: Lypesthes Baly, 1863 [conservation of usage of this name over Fidia Motschulsky, 1860 rejected by Commission (ICZN 2009c); syn. of Fidia Motschulsky, 1860]. Comment: name proposed to replace Leprotini Chapuis, 1874 because of the synonymy of the type genus; incorrect original stem formation, not in prevailing usage.
Ebooina Reid, 1993: 61 [stem: Eboo-]. Type genus: Eboo Reid, 1993. Comment: replacement name for Tomyrites Chapuis, 1874 because of the homonymy of the type genus.
Tribe
Caryonodini
Bechyné, 1951
Caryonodini Bechyné, 1951: 265 [stem: Caryonod-]. Type genus: Caryonoda Bechyné, 1951.
Tribe
Cubispini
Monrós, 1954
Cubispini Monrós, 1954: 26 [stem: Cubisp-]. Type genus: Cubispa Barber, 1946.
Tribe
Eumolpini
Hope, 1840
Eumolpidae Hope, 1840a: 162 [stem: Eumolp-]. Type genus: Eumolpus Kugelann, 1798 [an application to suppress Eumolpus Kugelann, 1798 and conserve Eumolpus Weber, 1801 was recently submitted to the Commission by Moseyko et al. (2010)]. Comment: First Reviser (Eumolpini Hope, 1840 vs Colaspidini Hope, 1840) not determined, current usage maintained.
Colaspidae Hope, 1840a: 163 [stem: Colaspid-]. Type genus: Colaspis Fabricius, 1801. Comment: incorrect original stem formation, not in prevailing usage.
Corynodina T. A. Marshall, 1865: 29 [stem: Corynod-]. Type genus: Corynodes Hope, 1840 [syn. of Platycorynus Chevrolat, 1836].
Chalcophanites Chapuis, 1874: 256 [stem: Chalcophan-]. Type genus: Chalcophana Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 50, as Chalcophanitae).
Edusites Chapuis, 1874: 306 [stem: Edus-]. Type genus: Edusa Chapuis, 1874 [preoccupied genus name, not Edusa Gistel, 1848 [Tunicata], not Edusa Albers, 1860 [Gastropoda] and not Edusa Martens 1860 [Mollusca]; syn. of Edusella Chapuis, 1874]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 111, as Edusitae); permanently invalid (Art. 39): based on preoccupied type genus.
Endocéphalites Chapuis, 1874: 343 [stem: Endocephal-]. Type genus: Endocephalus Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 154, as Endocephalitae).
Iphiméites Chapuis, 1874: 230 [stem: Iphime-]. Type genus: Iphimeis Baly, 1864. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Lefèvre (1885: 12, as Iphimeitae), generally accepted as in Riley et al. (2002: 639, as Iphimeites [treated as Latin]).
Chrysodinitae Lefèvre, 1885: 5 [stem: Chrysodin-]. Type genus: Chrysodina Baly, 1864 [syn. of Spintherophyta Dejean, 1836].
Edusellini Clavareau, 1914: 121 [stem: Edusell-]. Type genus: Edusella Chapuis, 1874.
Colaspoidini Chen, 1940: 488 [stem: Colaspoid-]. Type genus: Colaspoides Laporte, 1833.
Tribe
Euryopini
Chapuis, 1874
Euryopites Chapuis, 1874: 302 [stem: Euryop-]. Type genus: Euryope Dalman, 1824. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 108, as Euryopitae).
Odontionopites Lefèvre, 1876: 301 [stem: Odontionop-]. Type genus: Odontionopa Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 65, as Odontionopitae).
Prasoideini Clavareau, 1914: 65 [stem: Prasoide-]. Type genus: Prasoidea Weise, 1907 [syn. of Odontionopa Chevrolat, 1836].
*Colasposomini Bechyné, 1957: 7, 8 [stem: Colasposomat-]. Type genus: Colasposoma Laporte, 1833. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Colasposomini Špringlová de Bechyné, 1960: 3 [stem: Colasposomat-]. Type genus: Colasposoma Laporte, 1833. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Habrophorini
Bechyné and Špringlová de Bechyné, 1969
Habrophorini Bechyné and Špringlová de Bechyné, 1969: 75 [stem: Habrophor-]. Type genus: Habrophora Erichson, 1847.
Tribe
Hemydacnini
Bechyné, 1951
Hemydacnini Bechyné, 1951: 91 [stem: Hemydacn-]. Type genus: Hemydacne Jacoby, 1897.
Tribe
Megascelidini
Chapuis, 1874
Mégascélides Chapuis, 1874: 82 [stem: Megascelid-]. Type genus: Megascelis Sturm, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jacoby and Clavareau (1905: 1, as Megascelidae [incorrect stem formation]), generally accepted as in Riley et al. (2002: 639, as Megascelidini); incorrect original stem formation, not in prevailing usage.
Tribe
Merodini
Chapuis, 1874
Merodites Chapuis, 1874: 327 [stem: Merod-]. Type genus: Meroda Baly, 1860. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 128, as Meroditae).
Tribe
Pygomolpini
Bechyné, 1949
Pygomolpini Bechyné, 1949: 532 [stem: Pygomolp-]. Type genus: Pygomolpus Bechyné, 1949.
Tribe
Rosiroiini
Bechyné, 1950
Rosiroiini Bechyné, 1950b: 148 [stem: Rosiroi-]. Type genus: Rosiroia Bechyné, 1950.
Tribe
Typophorini
Baly, 1865
Typophorinae Baly, 1865: 433 [stem: Typophor-]. Type genus: Typophorus Chevrolat, 1836.
Callisinites Chapuis, 1874: 263 [stem: Callisin-]. Type genus: Callisina Baly, 1860. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 63, as Callisinitae).
Métachromites Chapuis, 1874: 295 [stem: Metachromat-]. Type genus: Metachroma Chevrolat, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Lefèvre (1885: 92, as Metachromitae), generally accepted as in Riley et al. (2002: 639, as Metachromites [treated as Latin]); incorrect original stem formation, not in prevailing usage.
Nodostomites Chapuis, 1874: 261 [stem: Nodostomat-]. Type genus: Nodostoma Motschulsky, 1860 [syn. of Basilepta Baly, 1860]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 56, as Nodostomitae); incorrect original stem formation, not in prevailing usage.
Pagriites Lefèvre, 1884: lxvii [stem: Pagri-]. Type genus: Pagria Lefèvre, 1884. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Lefèvre (1885: 62, as Pagriitae).
Cheirideitae Lefèvre, 1885: 67 [stem: Cheiride-]. Type genus: Cheiridea Baly, 1878.
Nodinini Chen, 1940: 487 [stem: Nodin-]. Type genus: Nodina Motschulsky, 1858.
Basileptini Chûjô, 1956: 9 [stem: Basilept-]. Type genus: Basilepta Baly, 1860. Comment: name proposed to replace Nodostomini Chapuis, 1874 because of the synonymy of the type genus.
Nodini Selman, 1965: 145 [stem: Nod-]. Type genus: Noda Chevrolat, 1836 [preoccupied genus name, not Noda Schellenberg 1803 [Diptera]; syn. of Brachypnoea Gistel, 1848]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Eumolpinae
incertae sedis
*Eupalini Verma et al., 2005: 164 [stem: Eupal-]. Type genus: Eupales Lefèvre, 1885 [although Floricola Gistel, 1848 is an older available name for this genus, almost all authors have used Eupales Lefèvre, 1885 in the last 125 years; we use Eupales Lefèvre, 1885 as valid here pending a vote on its conservation following a recent submission to the Commission by Jolivet and Verma (Bulletin of Zoological Nomenclature 2009: 204; also see Appendix 6)]. Comment: unavailable family-group name, proposed after 1999 without explicit intention (Art. 16.1); included here as “incertae sedis” as was done in the most recent volume of the Catalogue of Palaearctic Coleoptera (see Löbl 2010: 83); notice of a new application for the conservation of Eupales Lefèvre, 1885 and Eupalini Verma et al. submitted by Jolivet and Verma was recently published in the Bulletin of Zoological Nomenclature (2009: 204; also see Appendix 6).
Subfamily
Spilopyrinae
Chapuis, 1874
Spilopyrites Chapuis, 1874: 259 [stem: Spilopyr-]. Type genus: Spilopyra Baly, 1860. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Lefèvre (1885: 56, as Spilopyritae), generally accepted as in Reid (2000: 852, as Spilopyrinae); First Reviser found (Spilopyrinae Chapuis, 1874 vs Stenomelinae Chapuis, 1874) is Reid (2000: 852).
Stenomèlites Chapuis, 1874: 421 [stem: Stenomel-]. Type genus: Stenomela Erichson, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Monrós (1958c: 146, as Stenomelini).
Hornibiini Crowson, 1946: 79, in key [stem: Hornibi-]. Type genus: Hornibius Fairmaire, 1888 [syn. of Hornius Fairmaire, 1885].
Subfamily
Synetinae
LeConte and Horn, 1883
Synetae J. L. LeConte and G. H. Horn, 1883: 338 [stem: Synet-]. Type genus: Syneta Dejean, 1835.
Synetidae Edwards, 1953: 29 [stem: Synet-]. Type genus: Syneta Dejean, 1835. Comment: family-group name proposed as new without reference to Synetae J. L. LeConte and G. H. Horn, 1883.
†Subfamily
Protoscelidinae
Medvedev, 1968
Protoscelinae L. N. Medvedev, 1968: 155 [stem: Protoscelid-]. Type genus: Protoscelis L. N. Medvedev, 1968. Comment: incorrect original stem formation, not in prevailing usage.
Superfamily
Curculionoidea
Latreille, 1802
Curculionites Latreille, 1802: 195 [stem: Curculion-]. Type genus: Curculio Linnaeus, 1758.
Family
Nemonychidae
Bedel, 1882
Nemonychidae Bedel, 1882: 3, in key [stem: Nemonych-]. Type genus: Nemonyx Redtenbacher, 1845 [placed on the Official List of Generic Names in Zoology (ICZN 2005c)]. Comment: given precedence over Cimberididae Gozis, 1882 and placed on the Official List of Family-Group Names in Zoology (ICZN 2005c).
Subfamily
Nemonychinae
Bedel, 1882
Nemonychidae Bedel, 1882: 3, in key [stem: Nemonych-]. Type genus: Nemonyx Redtenbacher, 1845 [placed on the Official List of Generic Names in Zoology (ICZN 2005c)]. Comment: placed on the Official List of Family-Group Names in Zoology (ICZN 2005c).
Subfamily
Cimberidinae
Gozis, 1882
Cimberidae Gozis, 1882: 58 [stem: Cimberid-]. Type genus: Cimberis Gozis, 1881 [placed on the Official List of Generic Names in Zoology (ICZN 2005c)]. Comment: Cimberidae Gozis, 1882 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology and Cimberididae Gozis, 1882 placed on the Official List of Family-Group Names in Zoology (ICZN 2005c).
Tribe
Cimberidini
Gozis, 1882
Rhinomacerides Schönherr, 1823: column 1136 [stem: Rhinomacr-]. Type genus: Rhinomacer sensu A. G. Olivier, 1807 [not Rhinomacer Fabricius, 1781; syn. of Cimberis Gozis, 1881]. Comment: placed on the Official Index of Rejected and Invalid Family-Group Names (ICZN 2005c).
Cimberidae Gozis, 1882: 58 [stem: Cimberid-]. Type genus: Cimberis Gozis, 1881 [placed on the Official List of Generic Names in Zoology (ICZN 2005c)]. Comment: Cimberidae Gozis, 1882 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology and Cimberididae Gozis, 1882 placed on the Official List of Family-Group Names in Zoology (ICZN 2005c); incorrect original stem formation, not in prevailing usage.
*Neocimberini O’Brien and Wibmer, 1982: 3 [stem: Neocimberid-]. Type genus: Neocimberis O’Brien and Wibmer, 1982 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 2005c)]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.
Tribe
Doydirhynchini
Pierce, 1916
Doydirhynchoidea Pierce, 1916: 463, in key [stem: Doydirhynch-]. Type genus: Doydirhynchus Dejean, 1821.
†Tribe
Kuschelomacrini
Riedel, 2010
Kuschelomacerini Riedel, 2010: 31 [stem: Kuschelomacr-]. Type genus: Kuschelomacer Riedel, 2010. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Rhinorhynchinae
Voss, 1922
Rhinorhynchini Voss, 1922: 17 [stem: Rhinorhynch-]. Type genus: Rhinorhynchus Sharp, 1882.
Tribe
Mecomacerini
Kuschel, 1994
Mecomacerini Kuschel, 1994: 576, in key [stem: Mecomacer-]. Type genus: Mecomacer Kuschel, 1954. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Mecomacr-).
Subtribe
Brarina
Legalov, 2009
Brarina Legalov, 2009b: 208 [stem: Brar-]. Type genus: Brarus Kuschel, 1997.
Subtribe
Mecomacerina
Kuschel, 1994
*Mecomacerini May, 1993: 15, in key [stem: Mecomacer-]. Type genus: Mecomacer Kuschel, 1954. Comment: unavailable family-group name, description shared with Rhynchitoplesiini, not unequivocal (Art. 13.1.1) (see Alonso-Zarazaga and Lyal 1999: 27).
*Rhynchitoplesiini May, 1993: 15, in key [stem: Rhynchitoplesi-]. Type genus: Rhynchitoplesius Voss, 1952. Comment: unavailable family-group name, description shared with Mecomacerini, not unequivocal (Art. 13.1.1) (see Alonso-Zarazaga and Lyal 1999: 27).
Mecomacerini Kuschel, 1994: 576, in key [stem: Mecomacer-]. Type genus: Mecomacer Kuschel, 1954. Comment: current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Mecomacr-).
Tribe
Rhinorhynchini
Voss, 1922
Rhinorhynchini Voss, 1922: 17 [stem: Rhinorhynch-]. Type genus: Rhinorhynchus Sharp, 1882.
Rhynchitomacerini May, 1993: 15, in key [stem: Rhynchitomacr-]. Type genus: Rhynchitomacer Voss, 1937. Comment: incorrect original stem formation, not in prevailing usage.
†Subfamily
Slonikinae
Zherikhin, 1977
Slonikinae Zherikhin, 1977: 179 [stem: Slonik-]. Type genus: Slonik Zherikhin, 1977.
†Tribe
Slonikini
Zherikhin, 1977
Slonikinae Zherikhin, 1977: 179 [stem: Slonik-]. Type genus: Slonik Zherikhin, 1977.
†Tribe
Ulyaniscini
Legalov, 2009
Ulyaniscini Legalov, 2009a: 128 [stem: Ulyanisc-]. Type genus: Ulyanisca Gratschev, 1998.
†Subfamily
Eccoptarthrinae
Arnoldi, 1977
Eccoptarthrini Arnoldi, 1977: 169 [stem: Eccoptarthr-]. Type genus: Eccoptarthrus Arnoldi, 1977.
Mesophyletinae Poinar, 2006: 879 [stem: Mesophylet-]. Type genus: Mesophyletis Poinar, 2006. Comment: Poinar (2008: 262) unnecessarily revalidated this name on the basis of Art. 16.2, the criteria of availability were met in the original description in 2006 where the name of the type genus was cited.
†Subfamily
Brenthorrhininae
Arnoldi, 1977
Brenthorrhininae Arnoldi, 1977: 172 [stem: Brenthorrhin-]. Type genus: Brenthorrhinus Arnoldi, 1977. Comment: placement according to Legalov (2009c: 289).
†Tribe
Brenthorrhinini
Arnoldi, 1977
Brenthorrhininae Arnoldi, 1977: 172 [stem: Brenthorrhin-]. Type genus: Brenthorrhinus Arnoldi, 1977.
†Tribe
Brenthorrhinoidini
Legalov, 2003
Brenthorrhinoidini Legalov, 2003: 88 [stem: Brenthorrhinoid-]. Type genus: Brenthorrhinoides Gratshev and Zherikhin, 1996.
†Subfamily
Distenorrhininae
Arnoldi, 1977
Distenorrhinini Arnoldi, 1977: 170 [stem: Distenorrhin-]. Type genus: Distenorrhinus Arnoldi, 1977. Comment: placement according to Legalov (2009c: 290).
†Subfamily
Eobelinae
Arnoldi, 1977
Eobelidae Arnoldi, 1977: 144 [stem: Eobel-]. Type genus: Eobelus Arnoldi, 1977. Comment: precedence (Oxycorynoidinae Arnoldi, 1977 vs Nanophydinae Arnoldi, 1977 vs Eobelinae Arnoldi, 1977) given to taxon originally proposed at the higher rank (Art. 24.1).
†Tribe
Eobelini
Arnoldi, 1977
Eobelidae Arnoldi, 1977: 144 [stem: Eobel-]. Type genus: Eobelus Arnoldi, 1977. Comment: precedence (Eobelini Arnoldi, 1977 vs Procurculionini Arnoldi, 1977) given to taxon originally proposed at the higher rank (Art. 24.1).
†Subtribe
Eobelina
Arnoldi, 1977
Eobelidae Arnoldi, 1977: 144 [stem: Eobel-]. Type genus: Eobelus Arnoldi, 1977.
†Subtribe
Procurculionina
Arnoldi, 1977
Procurculionini Arnoldi, 1977: 157 [stem: Procurculion-]. Type genus: Procurculio Arnoldi, 1977. Comment: First Reviser (Eccoptothoracina Arnoldi, 1977 vs Procurculionina Arnoldi, 1977) not determined, current usage maintained.
Eccoptothoracini Arnoldi, 1977: 158 [stem: Eccoptothorac-]. Type genus: Eccoptothorax Arnoldi, 1977.
†Tribe
Karataucarini
Legalov, 2009
Karataucarini Legalov, 2009c: 288 [stem: Karataucar-]. Type genus: Karataucar Legalov, 2009.
†Tribe
Nanophydini
Arnoldi, 1977
Nanophydinae Arnoldi, 1977: 173 [stem: Nanophyd-]. Type genus: Nanophydes Arnoldi, 1977.
†Tribe
Oxycorynoidini
Arnoldi, 1977
Oxycorynoidinae Arnoldi, 1977: 159 [stem: Oxycorynoid-]. Type genus: Oxycorynoides Arnoldi, 1977.
†Tribe
Probelini
Legalov, 2009
Probelini Legalov, 2009c: 287 [stem: Probel-]. Type genus: Probelus Arnoldi, 1977.
†Subfamily
Paleocartinae
Legalov, 2003
Paleocartini Legalov, 2003: 78 [stem: Paleocart-]. Type genus: Paleocartus Legalov, 2003.
†Tribe
Nebrenthorrhinini
Legalov, 2007
Nebrenthorrhinina Legalov, 2007: 34 [stem: Nebrenthorrhin-]. Type genus: Nebrenthorrhinus Legalov, 2003. Comment: placement according to Legalov (2009c: 288).
†Tribe
Paleocartini
Legalov, 2003
Paleocartini Legalov, 2003: 78 [stem: Paleocart-]. Type genus: Paleocartus Legalov, 2003. Comment: placement according to Legalov (2009c: 288).
†Subfamily
Metrioxenoidinae
Legalov, 2009
Metrioxenoidinae Legalov, 2009c: 288 [stem: Metrioxenoid-]. Type genus: Metrioxenoides Gratshev et al., 1998.
†Subfamily
Cretonemonychinae
Gratshev and Legalov, 2009
Cretonemonychinae Gratshev and Legalov, 2009: 412 [stem: Cretonemonych-]. Type genus: Cretonemonyx Gratshev and Legalov, 2009.
†Subfamily
Selengarhynchinae
Gratshev and Legalov, 2009
Selengarhynchinae Gratshev and Legalov, 2009: 414 [stem: Selengarhynch-]. Type genus: Selengarhynchus Gratshev and Legalov, 2009.
Family
Anthribidae
Billberg, 1820
Anthribides Billberg, 1820a: 39 [stem: Anthrib-]. Type genus: Anthribus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: placed on the Official List of Family-Group Names in Zoology and given priority over Choragidae Kirby, 1819 (ICZN 1994b).
Subfamily
Anthribinae
Billberg, 1820
Anthribides Billberg, 1820a: 39 [stem: Anthrib-]. Type genus: Anthribus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: placed on the Official List of Family-Group Names in Zoology and given priority over Choragidae Kirby, 1819 (ICZN 1994b).
Tribe
Anthribini
Billberg, 1820
Anthribides Billberg, 1820a: 39 [stem: Anthrib-]. Type genus: Anthribus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)]. Comment: placed on the Official List of Family-Group Names in Zoology and given priority over Choragidae Kirby, 1819 (ICZN 1994b).
Brachytarsina C. G. Thomson, 1859: 128 [stem: Brachytars-]. Type genus: Brachytarsus Schönherr, 1823 [syn. of Anthribus Geoffroy, 1762].
Brachytarsini Pierce, 1930: 22, in key [stem: Brachytars-]. Type genus: Brachytarsus Schönherr, 1823 [syn. of Anthribus Geoffroy, 1762]. Comment: family-group name proposed as new without reference to Brachytarsina C. G. Thomson, 1859.
Tribe
Basitropini
Lacordaire, 1865
Basitropides Lacordaire, 1865: 566 [stem: Basitrop-]. Type genus: Basitropis Jekel, 1855. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Stein (1868: 115, as Basitropini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 28, as Basitropidini [incorrect stem formation]); Alonso-Zarazaga and Lyal (1999: 28) used the spelling Basitropidini but the same authors subsequently (2002: 4) reverted to Basitropini; precedence (Basitropini Lacordaire, 1865 vs Eugonini Lacordaire, 1865) given to taxon originally proposed at the higher rank (Art. 24.1).
Eugonides Lacordaire, 1865: 569 [stem: Eugon-]. Type genus: Eugonus Schönherr, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Sharp (1873: 32, as Eugonides [treated as Latin]); synonymy with Basitropini by Alonso-Zarazaga and Lyal (2002: 4).
Phaenithonini Pierce, 1930: 4, in key [stem: Phaenithon-]. Type genus: Phaenithon Schönherr, 1826.
Tribe
Corrhecerini
Lacordaire, 1865
Corrhécérides Lacordaire, 1865: 547 [stem: Corrhecer-]. Type genus: Corrhecerus Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Morimoto (1981: 78, as Corrhecerini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 29, as Corrhecerini).
Nessiarini Morimoto, 1972: 38, in key [stem: Nessiar-]. Type genus: Nessiara Pascoe, 1860.
Tribe
Cratoparini
LeConte, 1876
Cratopares J. L. LeConte, 1876: 403 [stem: Cratopar-]. Type genus: Cratoparis Dejean, 1834 [syn. of Euparius Schönherr, 1823]. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Cratoparent-).
Eupariini B. D. Valentine, 1960: 49, in key [stem: Eupari-]. Type genus: Euparius Schönherr, 1823. Comment: junior homonym of Eupariini A. Schmidt, 1910 (type genus Euparia Lepeletier and Serville, 1828) in Scarabaeidae.
†Tribe
Cretanthribini
Legalov, 2009
Cretanthribini Legalov, 2009c: 291 [stem: Cretanthrib-]. Type genus: Cretanthribus Legalov, 2009.
Tribe
Decataphanini
Lacordaire, 1865
Décataphanides Lacordaire, 1865: 556 [stem: Decataphan-]. Type genus: Decataphanes Labram and Imhoff, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 289, as Decatophaninae [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 30, as Decataphanini).
Tribe
Discotenini
Lacordaire, 1865
Discoténides Lacordaire, 1865: 500 [stem: Discoten-]. Type genus: Discotenes Labram and Imhoff, 1841. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pierce (1930: 4, as Discotenini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 30, as Discotenini).
Tribe
Ecelonerini
Lacordaire, 1865
Ecélonérides Lacordaire, 1865: 562 [stem: Eceloner-]. Type genus: Ecelonerus Schönherr, 1839. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Morimoto (1972: 38, as Ecelonerini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 30, as Ecelonerini).
Tribe
Ischnocerini
Lacordaire, 1865
Ischnocérides Lacordaire, 1865: 504 [stem: Ischnocer-]. Type genus: Ischnocerus Schönherr, 1839. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1876: 393, as Ischnoceri), generally accepted as in Alonso-Zarazaga and Lyal (2002: 5, as Ischnocerini); the Hymenoptera name Ischnocerini Clement, 1938 (type genus Ischnoceros Gravenhorst, 1829) is apparently unavailable.
Meconemini Pierce, 1930: 4, in key [stem: Meconem-]. Type genus: Meconemus Labram and Imhoff, 1838.
Tribe
Gymnognathini
Valentine, 1960
Gymnognathini B. D. Valentine, 1960: 48, in key [stem: Gymnognath-]. Type genus: Gymnognathus Schönherr, 1826.
Tribe
Jordanthribini
Morimoto, 1980
Jordanthribini Morimoto, 1980: 16 [stem: Jordanthrib-]. Type genus: Jordanthribus Zimmerman, 1938.
Tribe
Mauiini
Valentine, 1990
Mauiini B. D. Valentine, 1990: 235, in key [stem: Maui-]. Type genus: Mauia Blackburn, 1885.
Tribe
Mecocerini
Lacordaire, 1865
Mécocérides Lacordaire, 1865: 493 [stem: Mecocer-]. Type genus: Mecocerus Schönherr, 1833 [placed on the Official List of Generic Names in Zoology (ICZN 1972); syn. of Acanthothorax Gaede, 1832]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 288, as Mecocerinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 31, as Mecocerini).
Phloeophilides Lacordaire, 1865: 517 [stem: Phloeophil-]. Type genus: Phloeophilus Schönherr, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Olliff (1891: 75, as Phloeophilides [treated as Latin]); the older name Phloeophilini Kiesenwetter, 1863 [Cleroidea] was based on an unjustified emendation, the correct stem for the cleroid family-group name is Phloiophil-.
Cappadocini Alonso-Zarazaga and Lyal, 1999: 28 [stem: Cappadoc-]. Type genus: Cappadox Alonso-Zarazaga and Lyal, 1999 [syn. of Phloeophilus Schönherr, 1833].
Tribe
Mycteini
Morimoto, 1972
Mycteini Morimoto, 1972: 38, in key [stem: Mycte-]. Type genus: Mycteis Pascoe, 1860.
Tribe
Ozotomerini
Morimoto, 1972
Ozotomerini Morimoto, 1972: 37, in key [stem: Ozotomer-]. Type genus: Ozotomerus Perroud, 1853.
Tribe
Piesocorynini
Valentine, 1960
Piesocorynini B. D. Valentine, 1960: 49, in key [stem: Piesocoryn-]. Type genus: Piesocorynus Dejean, 1834.
Tribe
Platyrhinini
Bedel, 1882
Platyrrhinidae Bedel, 1882: 3, in key [stem: Platyrhin-]. Type genus: Platyrhinus Clairville, 1798. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Platystomini
Pierce, 1916
Anthotribidae Gistel, 1856a: 375 [stem: Anthotrib-]. Type genus: Anthotribus Gistel, 1856 [preoccupied genus name, not Anthotribus Hoffmann, 1803 [Coleoptera: Anthribidae: Anthribinae: Anthribini]; syn. of Platystomos Schneider, 1791]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Platystomoidea Pierce, 1916: 463, in key [stem: Platystom-]. Type genus: Platystomos Schneider, 1791. Comment: the type genus for this family-group name was in fact proposed as a replacement name for Anthribus sensu Fabricius, 1790, however it was never used in that sense subsequently (Alonso-Zarazaga and Lyal 1999: 33); an application will be submitted by MAAZ and CHCL to conserve the current concept of Platystomos Schneider, 1791 and its associated family-group name.
Tribe
Proscoporhinini
Lacordaire, 1865
Proscoporhinides Lacordaire, 1865: 544 [stem: Proscoporhin-]. Type genus: Proscoporhinus Montrouzier, 1861. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Ienistea (1986: 32, as Prosoporhinidae [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 33, as Proscoporhinini).
Tribe
Ptychoderini
Jekel, 1855
Ptychoderidae Jekel, 1855: 70 [stem: Ptychoder-]. Type genus: Ptychoderes Schönherr, 1823.
*Phloeotragides Lacordaire, 1865: 486 [stem: Phloeotrag-]. Type genus: Phloeotragus Schönherr, 1823. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1865) and used as valid.
Phloeotraginae Kolbe, 1897: 288 [stem: Phloeotrag-]. Type genus: Phloeotragus Schönherr, 1823.
Tribe
Sintorini
Lacordaire, 1865
Sintorides Lacordaire, 1865: 510 [stem: Sintor-]. Type genus: Sintor Schönherr, 1839. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Morimoto (1972: 38, as Sintorini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 34, as Sintorini).
Tribe
Stenocerini
Kolbe, 1895
Stenocerinarum Kolbe, 1895: 381 [stem: Stenocer-]. Type genus: Stenocerus Schönherr, 1826. Comment: Stenocerinarum is thegenitive form of Stenocerinae and was recognized as such by Alonso-Zarazaga and Lyal (1999: 34).
Allandrini Pierce, 1930: 18 [stem: Allandr-]. Type genus: Allandrus J. L. LeConte, 1876.
Tribe
Tophoderini
Lacordaire, 1865
Tophodérides Lacordaire, 1865: 499 [stem: Tophoder-]. Type genus: Tophoderes Dejean, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 288, as Tophoderinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 34, as Tophoderini).
Tribe
Trigonorhinini
Valentine, 1999
Trigonorhinini B. D. Valentine, 1999: 287, in key [stem: Trigonorhin-]. Type genus: Trigonorhinus Wollaston, 1861.
Tribe
Tropiderini
Lacordaire, 1865
Tropidérides Lacordaire, 1865: 484 [stem: Tropider-]. Type genus: Tropideres Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Stein (1868: 115, as Tropiderini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 35, as Tropiderini); precedence (Tropiderini Lacordaire, 1865 vs Acorynini Lacordaire, 1865) given to taxon originally proposed at the higher rank (Art. 24.1).
Acorynides Lacordaire, 1865: 512 [stem: Acoryn-]. Type genus: Acorynus Schönherr, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 289, as Acoryninae), generally accepted as in Morimoto (1980: 22, as Acorynini).
Eurymycterini Pierce, 1930: 15 [stem: Eurymycter-]. Type genus: Eurymycter J. L. LeConte, 1876.
Tribe
Xenocerini
Lacordaire, 1865
Xénocerides Lacordaire, 1865: 558 [stem: Xenocer-]. Type genus: Xenocerus Schönherr, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 235, as Xenocerini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 35, as Xenocerini).
Tribe
Xylinadini
Lacordaire, 1865
Xylinadides Lacordaire, 1865: 560 [stem: Xylinad-]. Type genus: Xylinades Latreille, 1828 [syn. of Xylinada Berthold, 1827]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 289, as Xylinadinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 36, as Xylinadini).
Tribe
Zygaenodini
Lacordaire, 1865
Zygénodides Lacordaire, 1865: 542 [stem: Zygaenod-]. Type genus: Zygaenodes Pascoe, 1859 [syn. of Exechesops Schönherr, 1847]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 289, as Zygaenodinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 36, as Zygaenodini); incorrect original stem formation, not in prevailing usage.
Hormisci J. L. LeConte, 1876: 396 [stem: Ormisc-]. Type genus: Ormiscus G. R. Waterhouse, 1845 [as Hormiscus, unjustified emendation of type genus name by Agassiz (1846b: 263), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Choraginae
Kirby, 1819
Choragidae Kirby, 1819: 447 [stem: Chorag-]. Type genus: Choragus Kirby, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 1994b)]. Comment: placed on the Official List of Family-Group Names in Zoology and not given priority over Anthribidae Billberg, 1820 (ICZN 1994b).
Tribe
Apolectini
Lacordaire, 1865
Apolectides Lacordaire, 1865: 554 [stem: Apolect-]. Type genus: Apolecta Pascoe, 1859. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1897: 289, as Apolectinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 28, as Apolectini); Apolectidae Jordan, 1923 (type genus Apolectus Cuvier, 1832) is available in Pisces although it is based on a preoccupied genus name and therefore permanently invalid; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Tribe
Araecerini
Lacordaire, 1865
Araeocérides Lacordaire, 1865: 588 [stem: Araecer-]. Type genus: Araecerus Schönherr, 1823 [as Araeocerus, unjustified emendation of type genus name by Schönherr (1839), not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Stein (1868: 115, as Araeocerini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 37, as Araecerini); incorrect original stem formation, not in prevailing usage.
Tribe
Cisanthribini
Zimmerman, 1994
Cisanthribini Zimmerman, 1994a: 232 [stem: Cisanthrib-]. Type genus: Cisanthribus Zimmerman, 1938.
Tribe
Choragini
Kirby, 1819
Choragidae Kirby, 1819: 447 [stem: Chorag-]. Type genus: Choragus Kirby, 1819 [placed on the Official List of Generic Names in Zoology (ICZN 1994b)]. Comment: not given priority over Anthribidae Billberg, 1820, placed on the Official List of Family-Group Names in Zoology (ICZN 1994b).
Tribe
Valenfriesiini
Alonso-Zarazaga and Lyal, 1999
Notioxénides Lacordaire, 1865: 593 [stem: Notioxen-]. Type genus: Notioxenus Wollaston, 1861 [preoccupied genus name, not Notioxenus Motschulsky, 1858 [Coleoptera: Carabidae]; syn. of Valenfriesia Alonso-Zarazaga and Lyal, 1999]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pierce (1930: 31, as Notioxenini), generally accepted as in Morimoto (1978a: 18, as Notioxenini); permanently invalid (Art. 39): based on preoccupied type genus.
Valenfriesiini Alonso-Zarazaga and Lyal, 1999: 38 [stem: Valenfriesi-]. Type genus: Valenfriesia Alonso-Zarazaga and Lyal, 1999. Comment: replacement name for Notioxenini Lacordaire, 1865 because of the homonymy of the type genus.
Tribe
Xenorchestini
Lacordaire, 1865
Xénorchestides Lacordaire, 1865: 595 [stem: Xenorchest-]. Type genus: Xenorchestes Wollaston, 1854. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1876: 408, as Xenorchestini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 38, as Xenorchestini).
Homoeoderides Wollaston, 1870: 23 [stem: Homoeoder-]. Type genus: Homoeodera Wollaston, 1870.
Subfamily
Urodontinae
Thomson, 1859
Urodontides C. G. Thomson, 1859: 128 [stem: Urodont-]. Type genus: Urodon Schönherr, 1823 [syn. of Bruchela Dejean, 1821].
Bruchelidae Pierce, 1916: 463 [stem: Bruchel-]. Type genus: Bruchela Dejean, 1821.
†Family
Ulyanidae
Zherikhin, 1993
Ulyanidae Zherikhin, 1993: 26 [stem: Ulyan-]. Type genus: Ulyana Zherikhin, 1993.
Family
Belidae
Schönherr, 1826
Belides Schönherr, 1826: 73 [stem: Bel-]. Type genus: Belus Schönherr, 1823 [syn. of Rhinotia Kirby, 1819].
Subfamily
Belinae
Schönherr, 1826
Belides Schönherr, 1826: 73 [stem: Bel-]. Type genus: Belus Schönherr, 1823 [syn. of Rhinotia Kirby, 1819].
Tribe
Agnesiotidini
Zimmerman, 1994
Agnesiotidini Zimmerman, 1994a: 258 [stem: Agnesiotid-]. Type genus: Agnesiotis Pascoe, 1870.
Tribe
Belini
Schönherr, 1826
Belides Schönherr, 1826: 73 [stem: Bel-]. Type genus: Belus Schönherr, 1823 [syn. of Rhinotia Kirby, 1819].
Subtribe
Belina
Schönherr, 1826
Belides Schönherr, 1826: 73 [stem: Bel-]. Type genus: Belus Schönherr, 1823 [syn. of Rhinotia Kirby, 1819].
Subtribe
Homalocerina
Legalov, 2009
Homalocerina Legalov, 2009d: 308 [stem: Homalocer-]. Type genus: Homalocerus Schönherr, 1839.
Tribe
Pachyurini
Kuschel, 1959
Pachyurini Kuschel, 1959a: 253, in key [stem: Pachyur-]. Type genus: Pachyura Hope, 1834.
Subfamily
Oxycoryninae
Schönherr, 1840
Oxycorynides Schönherr, 1840: 581 [stem: Oxycoryn-]. Type genus: Oxycorynus Chevrolat, 1832.
Tribe
Aglycyderini
Wollaston, 1864
Aglycyderidae Wollaston, 1864: 384 [stem: Aglycyder-]. Type genus: Aglycyderes Westwood, 1864.
Proterhinides Fauvel, 1891: 154 [stem: Proterhin-]. Type genus: Proterhinus Sharp, 1878. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Sharp (1899a: 298, as Proterhinidae).
Platycephalitae Paulian, 1944: 118 [stem: Platycephal-]. Type genus: Platycephala Montrouzier, 1861 [preoccupied genus name, not Platycephala Fallén, 1820 [Diptera]; syn. of Aralius Kuschel, 1990]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; also Platycephalidae Gill, 1872 (type genus Platycephalus Bloch, 1795) is available in Pisces.
Tribe
Alloxycorynini
Legalov, 2009
Alloxycorynini Legalov, 2009d: 313 [stem: Alloxycoryn-]. Type genus: Alloxycorynus Voss, 1957.
Tribe
Distenorrhinoidini
Legalov, 2009
Distenorrhinoidini Legalov, 2009d: 313 [stem: Distenorrhinoid-]. Type genus: Distenorrhinoides Gratshev and Zherikhin, 2000.
Tribe
Metrioxenini
Voss, 1953
Metrioxenini Voss, 1953: 124, in key [stem: Metrioxen-]. Type genus: Metrioxena Pascoe, 1870.
Subtribe
Afrocorynina
Voss, 1957
Afrocorynini Voss, 1957: 102, in key [stem: Afrocoryn-]. Type genus: Afrocorynus G. A. K. Marshall, 1955.
Hispodini Voss, 1957: 102, in key [stem: Hispod-]. Type genus: Hispodes G. A. K. Marshall, 1955.
Subtribe
Metrioxenina
Voss, 1953
Metrioxenini Voss, 1953: 124, in key [stem: Metrioxen-]. Type genus: Metrioxena Pascoe, 1870.
Subtribe
Zherichinixenina
Legalov, 2009
Zherichinixenina Legalov, 2009d: 310 [stem: Zherichinixen-]. Type genus: Zherichinixena Legalov, 2009.
Tribe
Oxycorynini
Schönherr, 1840
Oxycorynides Schönherr, 1840: 581 [stem: Oxycoryn-]. Type genus: Oxycorynus Chevrolat, 1832.
Subtribe
Allocorynina
Sharp, 1890
Allocoryninae Sharp, 1890: 45 [stem: Allocoryn-]. Type genus: Allocorynus Sharp, 1890 [syn. of Rhopalotria Chevrolat, 1878].
Subtribe
Oxycorynina
Schönherr, 1840
Oxycorynides Schönherr, 1840: 581 [stem: Oxycoryn-]. Type genus: Oxycorynus Chevrolat, 1832.
Subtribe
Oxycraspedina
Marvaldi and Oberprieler, 2006
Oxycraspedina Marvaldi and Oberprieler, 2006: 460 [stem: Oxycrasped-]. Type genus: Oxycraspedus Kuschel, 1955.
Family
Caridae
Thompson, 1992
Carinae Thompson, 1992: 882 [stem: Car-]. Type genus: Car Blackburn, 1897.
Subfamily
Carinae
Thompson, 1992
Carinae Thompson, 1992: 882 [stem: Car-]. Type genus: Car Blackburn, 1897.
Caridae Zimmerman, 1994a: 499 [stem: Car-]. Type genus: Car Blackburn, 1897. Comment: family-group name proposed as new without reference to Carinae Thompson, 1992.
Carinae Kuschel, 1995: 18 [stem: Car-]. Type genus: Car Blackburn, 1897. Comment: family-group name proposed as new without reference to Carinae Thompson, 1992 or Caridae Zimmerman, 1994.
Subfamily
Chilecarinae
Legalov, 2009
Chilecarini Legalov, 2009a: 125 [stem: Chilecar-]. Type genus: Chilecar Kuschel, 1992.
Tribe
Carodini
Legalov, 2009
Carodesina Legalov, 2009a: 126 [stem: Carod-]. Type genus: Carodes Zimmerman, 1994. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Chilecarini
Legalov, 2009
Chilecarini Legalov, 2009a: 125 [stem: Chilecar-]. Type genus: Chilecar Kuschel, 1992.
†Subfamily
Baissorhynchinae
Zherikhin, 1993
Baissorhynchini Zherikhin, 1993: 30 [stem: Baissorhynch-]. Type genus: Baissorhynchus Zherikhin, 1977.
†Abrocarina Legalov, 2009c: 291 [stem: Abrocar-]. Type genus: Abrocar Liu and Ren, 2006.
Family
Attelabidae
Billberg, 1820
Attelabides Billberg, 1820a: 39 [stem: Attelab-]. Type genus: Attelabus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1983a)].
Subfamily
Attelabinae
Billberg, 1820
Attelabides Billberg, 1820a: 39 [stem: Attelab-]. Type genus: Attelabus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1983a)].
Tribe
Attelabini
Billberg, 1820
Attelabides Billberg, 1820a: 39 [stem: Attelab-]. Type genus: Attelabus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1983a)].
Subtribe
Attelabina
Billberg, 1820
Attelabides Billberg, 1820a: 39 [stem: Attelab-]. Type genus: Attelabus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1983a)]. Comment: this family-group name was also used in the same year by Billberg (1820b: 393, as Attelabides).
Subtribe
Euscelina
Voss, 1925
Euscelina Voss, 1925: 32 [stem: Euscel-]. Type genus: Euscelus Schönherr, 1833.
Alleuscelina Legalov, 2003: 411 [stem: Alleuscel-]. Type genus: Alleuscelus Voss, 1937. Comment: proposed as a subtribe of Euscelini.
Clinolabina Legalov, 2003: 410 [stem: Clinolab-]. Type genus: Clinolabus Jekel, 1860. Comment: proposed as a subtribe of Euscelini.
Subtribe
Euscelophilina
Voss, 1925
Euscelophilina Voss, 1925: 29 [stem: Euscelophil-]. Type genus: Euscelophilus Voss, 1925.
Subtribe
Henicolabina
Legalov, 2007
Henicolabina Legalov, 2007: 282 [stem: Henicolab-]. Type genus: Henicolabus Voss, 1925.
Subtribe
Himatolabina
Legalov, 2003
Himatolabina Legalov, 2003: 424 [stem: Himatolab-]. Type genus: Himatolabus Jekel, 1860. Comment: proposed as a subtribe of Hybolabini.
Subtribe
Hybolabina
Voss, 1925
Hybolabina Voss, 1925: 191 [stem: Hybolab-]. Type genus: Hybolabus Jekel, 1860.
Subtribe
Isolabina
Legalov, 2007
Isolabina Legalov, 2007: 282 [stem: Isolab-]. Type genus: Isolabus Voss, 1925.
Subtribe
Lagenoderina
Voss, 1925
Lagenoderina Voss, 1925: 206 [stem: Lagenoder-]. Type genus: Lagenoderus A. White, 1841.
Subtribe
Lamprolabina
Voss, 1925
Lamprolabina Voss, 1925: 213 [stem: Lamprolab-]. Type genus: Lamprolabus Jekel, 1860.
Subtribe
Metocalolabina
Legalov, 2003
Metocalolabina Legalov, 2003: 433 [stem: Metocalolab-]. Type genus: Metocalolabus Legalov, 2003.
Subtribe
Omolabina
Legalov, 2003
Omolabina Legalov, 2003: 426 [stem: Omolab-]. Type genus: Omolabus Jekel, 1860. Comment: proposed as a subtribe of Hybolabini.
Subtribe
Paramecolabina
Legalov, 2003
Paramecolabina Legalov, 2003: 439 [stem: Paramecolab-]. Type genus: Paramecolabus Jekel, 1860.
Subtribe
Phialodina
Legalov, 2003
Phialodina Legalov, 2003: 437 [stem: Phialod-]. Type genus: Phialodes Roelofs, 1874.
Subtribe
Phymatolabina
Voss, 1925
Phymatolabina Voss, 1925: 199 [stem: Phymatolab-]. Type genus: Phymatolabus Jekel, 1860.
Subtribe
Phymatopsinina
Legalov, 2003
Phymatopsinina Legalov, 2003: 456 [stem: Phymatopsin-]. Type genus: Phymatopsinus Voss, 1925. Comment: proposed as a subtribe of Lagenoderini.
Subtribe
Pleurolabina
Legalov, 2003
Pleurolabina Legalov, 2003: 460 [stem: Pleurolab-]. Type genus: Pleurolabus Jekel, 1860. Comment: proposed as a subtribe of Lagenoderini.
Tribe
Euopini
Voss, 1925
Euopsini Voss, 1925: 291 [stem: Euop-]. Type genus: Euops Schönherr, 1839. Comment: incorrect original stem formation, not in prevailing usage.
Archeuopsina Legalov, 2003: 359 [stem: Archeuop-]. Type genus: Archeuops Legalov, 2003. Comment: proposed as a subtribe of Euopini; incorrect original stem formation, not in prevailing usage.
Suniopsina Legalov, 2003: 364 [stem: Suniop-]. Type genus: Suniops Voss, 1928. Comment: proposed as a subtribe of Euopini; incorrect original stem formation, not in prevailing usage.
Synaptopsina Legalov, 2003: 368 [stem: Synaptop-]. Type genus: Synaptops Jekel, 1860. Comment: proposed as a subtribe of Euopini; incorrect original stem formation, not in prevailing usage.
Ljudmilinina Legalov, 2007: 219 [stem: Ljudmilini-]. Type genus: Ljudmilinius Legalov, 2003. Comment: proposed as a subtribe of Euopini; incorrect original stem formation, not in prevailing usage.
Parasynaptopsisina Legalov, 2007: 227 [stem: Parasynaptopse-]. Type genus: Parasynaptopsis Legalov, 2003. Comment: proposed as a subtribe of Euopini; incorrect original stem formation, not in prevailing usage.
Riedelinina Legalov, 2007: 218 [stem: Riedelini-]. Type genus: Riedelinius Legalov, 2003. Comment: proposed as a subtribe of Euopini; incorrect original stem formation, not in prevailing usage.
Sawadaeuopsina Legalov, 2007: 241 [stem: Sawadaeuop-]. Type genus: Sawadaeuops Legalov, 2003. Comment: proposed as a subtribe of Euopini; incorrect original stem formation, not in prevailing usage.
Tribe
Pilolabini
Voss, 1925
Pilolabini Voss, 1925: 19 [stem: Pilolab-]. Type genus: Pilolabus Jekel, 1860.
Subfamily
Apoderinae
Jekel, 1860
Apoderidae Jekel, 1860: 180 [stem: Apoder-]. Type genus: Apoderus A. G. Olivier, 1807.
Tribe
Apoderini
Jekel, 1860
Apoderidae Jekel, 1860: 180 [stem: Apoder-]. Type genus: Apoderus A. G. Olivier, 1807.
Anisonychina Legalov, 2003: 555 [stem: Anisonych-]. Type genus: Anisonychus Voss, 1927. Comment: junior homonym of Anisonychidae H. C. C. Burmeister, 1844 (type genus Anisonyx Latreille, 1807) available in Scarabaeidae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Centrocorynina Legalov, 2003: 557 [stem: Centrocoryn-]. Type genus: Centrocorynus Jekel, 1860.
Cycnotrachelina Legalov, 2003: 568 [stem: Cycnotrachel-]. Type genus: Cycnotrachelus Jekel, 1860.
Opanassenkoviina Legalov, 2003: 554 [stem: Opanassenkovi-]. Type genus: Opanassenkovius Legalov, 2003.
Pseudocycnotrachelina Legalov, 2003: 525 [stem: Pseudocycnotrachel-]. Type genus: Pseudocycnotrachelus Legalov, 2003.
Tribe
Clitostylini
Voss, 1929
Clitostylini Voss, 1929a: 192 [stem: Clitostyl-]. Type genus: Clitostylus Voss, 1929 [syn. of Trachelismus Motschulsky, 1870].
Subtribe
Allapoderina
Legalov, 2003
Allapoderina Legalov, 2003: 467 [stem: Allapoder-]. Type genus: Allapoderus Voss, 1927.
Subtribe
Clitostylina
Voss, 1929
Clitostylini Voss, 1929a: 192 [stem: Clitostyl-]. Type genus: Clitostylus Voss, 1929 [syn. of Trachelismus Motschulsky, 1870].
Subtribe
Pseudophrysina
Legalov, 2003
Pseudophrysina Legalov, 2003: 476 [stem: Pseudophrys-]. Type genus: Pseudophrysus Legalov, 2003.
Tribe
Hoplapoderini
Voss, 1926
Hoplapoderini Voss, 1926: 14 [stem: Hoplapoder-]. Type genus: Hoplapoderus Jekel, 1860.
Subtribe
Afroapoderina
Legalov, 2003
Afroapoderina Legalov, 2003: 482 [stem: Afroapoder-]. Type genus: Afroapoderus Legalov, 2003.
Subtribe
Hoplapoderina
Voss, 1926
Hoplapoderini Voss, 1926: 14 [stem: Hoplapoder-]. Type genus: Hoplapoderus Jekel, 1860.
Subtribe
Paratomapoderina
Legalov, 2003
Paratomapoderina Legalov, 2003: 488 [stem: Paratomapoder-]. Type genus: Paratomapoderus Voss, 1926.
Tribe
Trachelophorini
Voss, 1926
Trachelophorini Voss, 1926: 14, in key [stem: Trachelophor-]. Type genus: Trachelophorus Jekel, 1860.
Subfamily
Rhynchitinae
Gistel, 1848
Rhynchitisidae Gistel, 1848: [8] [stem: Rhynchit-]. Type genus: Rhynchites Schneider, 1791.
Tribe
Auletini
Desbrochers des Loges, 1908
Auletinidae Desbrochers des Loges, 1908: 10, in key [stem: Aulet-]. Type genus: Auletes Schönherr, 1826. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Auletina
Desbrochers des Loges, 1908
Auletinidae Desbrochers des Loges, 1908: 10, in key [stem: Aulet-]. Type genus: Auletes Schönherr, 1826. Comment: incorrect original stem formation, not in prevailing usage.
Auletini Pierce, 1913: 365, in key [stem: Aulet-]. Type genus: Auletes Schönherr, 1826. Comment: family-group name proposed as new without reference to Auletinidae Desbrochers des Loges, 1908.
Subtribe
Auletobiina
Legalov, 2001
Auletobiina Legalov, 2001: 37 [stem: Auletobi-]. Type genus: Auletobius Desbrochers des Loges, 1869.
Subtribe
Guineauletina
Legalov, 2003
Guineauletina Legalov, 2003: 108 [stem: Guineaulet-]. Type genus: Guineauletes Legalov, 2003.
Subtribe
Mandelschtamiina
Legalov, 2003
Mandelschtamiina Legalov, 2003: 105 [stem: Mandelschtami-]. Type genus: Mandelschtamius Legalov, 2003.
Subtribe
Pseudauletina
Voss, 1933
Pseudauletina Voss, 1933b: 110, in key [stem: Pseudaulet-]. Type genus: Pseudauletes Voss, 1922.
Subtribe
Pseudomesauletina
Legalov, 2003
Pseudomesauletina Legalov, 2003: 113 [stem: Pseudomesaulet-]. Type genus: Pseudomesauletes Legalov, 2001.
Tribe
Auletorhinini
Voss, 1935
Auletorhinini Voss, 1935b: 509 [stem: Auletorhin-]. Type genus: Auletorhinus Voss, 1935.
Tribe
Byctiscini
Voss, 1923
Byctiscini Voss, 1923: 510 [stem: Byctisc-]. Type genus: Byctiscus C. G. Thomson, 1859.
Subtribe
Byctiscina
Voss, 1923
Byctiscini Voss, 1923: 510 [stem: Byctisc-]. Type genus: Byctiscus C. G. Thomson, 1859.
Subtribe
Listrobyctiscina
Legalov, 2003
Listrobyctiscina Legalov, 2003: 337 [stem: Listrobyctisc-]. Type genus: Listrobyctiscus Voss, 1923.
Subtribe
Svetlanaebyctiscina
Legalov, 2003
Svetlanaebyctiscina Legalov, 2003: 323 [stem: Svetlanaebyctisc-]. Type genus: Svetlanaebyctiscus Legalov, 2001.
Tribe
Cesauletini
Legalov, 2003
Cesauletini Legalov, 2003: 134 [stem: Cesaulet-]. Type genus: Cesauletes Hamilton, 1983. Comment: proposed as a tribe of the supertribe Rhynchititae.
Tribe
Deporaini
Voss, 1929
Deporaini Voss, 1929b: 28 [stem: Depora-]. Type genus: Deporaus Samouelle, 1819.
Subtribe
Chonostropheina
Morimoto, 1962
Chonostropheina Morimoto, 1962a: 30, in key [stem: Chonostrophe-]. Type genus: Chonostropheus Prell, 1924.
Subtribe
Deporaina
Voss, 1929
Deporaini Voss, 1929b: 28 [stem: Depora-]. Type genus: Deporaus Samouelle, 1819.
Tribe
Minurini
Legalov, 2003
Minurini Legalov, 2003: 133 [stem: Minur-]. Type genus: Minurus G. R. Waterhouse, 1842. Comment: proposed as a tribe of the supertribe Rhynchititae.
Tribe
Rhinocartini
Voss, 1931
Rhinocartini Voss, 1931: 162, in key [stem: Rhinocart-]. Type genus: Rhinocartus Voss, 1922.
*Rhynchitallini Voss, 1960: 415 [stem: Rhynchitall-]. Type genus: Rhynchitallus Voss, 1960. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Proteugnamptini Legalov, 2003: 80 [stem: Proteugnamptini]. Type genus: Proteugnamptus Voss, 1939. Comment: proposed as a tribe in supertribe Rhinocartitae.
Auletanini Legalov, 2003: 84 [stem: Auletan-]. Type genus: Auletanus Voss, 1922. Comment: proposed as a tribe in the supertribe Rhinocartitae.
†Sanyrevilleina Legalov, 2003: 85 [stem: Sanyreville-]. Type genus: Sanyrevilleus Gratshev and Zherikhin, 2000. Comment: proposed as a subtribe of Auletanini; raised to tribal level by Legalov (2007).
Proteugnamptini Legalov, 2003: 80 [stem: Proteugnampt-]. Type genus: Proteugnamptus Voss, 1939. Comment: proposed as a tribe in the supertribe Rhinocartitae.
Vossicartini Legalov, 2003: 79 [stem: Vossicart-]. Type genus: Vossicartus Legalov, 2003.
Eosalacina Legalov, 2007: 30 [stem: Eosalac-]. Type genus: Eosalacus Legalov, 2007. Comment: proposed as a subtribe of Proteugnamptini.
Parauletanina Legalov, 2007: 31 [stem: Parauletan-]. Type genus: Parauletanus Legalov, 2007. Comment: proposed as a subtribe of Sanyrevilleini.
Tribe
Rhynchitini
Gistel, 1848
Rhynchitisidae Gistel, 1848: [8] [stem: Rhynchit-]. Type genus: Rhynchites Schneider, 1791. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Acritorrhynchitina
Legalov, 2007
Acritorrhynchitina Legalov, 2007: 70 [stem: Acritorrhynchit-]. Type genus: Acritorrhynchites Voss, 1941.
Subtribe
Anisomerinina
Legalov, 2003
Anisomerinina Legalov, 2003: 223 [stem: Anisomerin-]. Type genus: Anisomerinus Voss, 1933.
Subtribe
Eugnamptina
Voss, 1930
Eugnamptina Voss, 1930: 67 [stem: Eugnampt-]. Type genus: Eugnamptus Schönherr, 1839.
Subtribe
Lasiorhynchitina
Legalov, 2003
Lasiorhynchitina Legalov, 2003: 202 [stem: Lasiorhynchit-]. Type genus: Lasiorhynchites Jekel, 1860.
Subtribe
Perrhynchitina
Legalov, 2003
Perrhynchitina Legalov, 2003: 218 [stem: Perrhynchit-]. Type genus: Perrhynchites Voss, 1953.
Subtribe
Rhynchitallina
Legalov, 2003
Rhynchitallina Legalov, 2003: 226 [stem: Rhynchitall-]. Type genus: Rhynchitallus Voss, 1960.
Subtribe
Rhynchitina
Gistel, 1848
Rhynchitisidae Gistel, 1848: [8] [stem: Rhynchit-]. Type genus: Rhynchites Schneider, 1791. Comment: incorrect original stem formation, not in prevailing usage.
Rhynchitini Pierce, 1913: 365, in key [stem: Rhynchit-]. Type genus: Rhynchites Schneider, 1791. Comment: family-group name proposed as new without reference to Rhynchitisidae Gistel, 1848.
Subtribe
Temnocerina
Legalov, 2003
Temnocerina Legalov, 2003: 207 [stem: Temnocer-]. Type genus: Temnocerus Thunberg, 1815.
Subfamily
Isotheinae
Scudder, 1893
Isotheinae Scudder, 1893: 16 [stem: Isothe-]. Type genus: Isothea Scudder, 1893. Comment: precedence (Isotheinae Scudder, 1893 vs Toxorhynchinae Scudder, 1893) given to taxon originally proposed at the higher rank (Art. 24.1).
Tribe
Isotheini
Scudder, 1893
Isotheinae Scudder, 1893: 16 [stem: Isothe-]. Type genus: Isothea Scudder, 1893.
Subtribe
Depasophilina
Legalov, 2003
Depasophilina Legalov, 2003: 166 [stem: Depasophil-]. Type genus: Depasophilus Voss, 1922.
†Subtribe
Isotheina
Scudder, 1893
Isotheinae Scudder, 1893: 16 [stem: Isothe-]. Type genus: Isothea Scudder, 1893.
†Tribe
Toxorhynchini
Scudder, 1893
Toxorhynchini Scudder, 1893: 23 [stem: Toxorhynch-]. Type genus: Toxorhynchus Scudder, 1893.
Subfamily
Pterocolinae
Lacordaire, 1865
Ptérocolides Lacordaire, 1865: 190 [stem: Pterocol-]. Type genus: Pterocolus Schönherr, 1833 [preoccupied genus name, not Pterocolus Say, 1831 [Coleoptera: Attelabidae: Pterocolinae]; syn. of Pterocolus Say, 1831]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Pterocolinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 44, as Pterocolinae); permanently invalid (Art. 39): based on preoccupied type genus; although strictly speaking the type genus is a junior homonym, and thus cannot be used as valid, to preserve stability, we are considering this as a valid name here; an application to the Commission is necessary to officially conserve usage of this family-group name.
Family
Brentidae
Billberg, 1820
Brenthides Billberg, 1820a: 40 [stem: Brent-]. Type genus: Brentus Fabricius, 1787 [as Brenthus, unjustified emendation of type genus name by Illiger (1801), not in prevailing usage].
Subfamily
Brentinae
Billberg, 1820
Brenthides Billberg, 1820a: 40 [stem: Brent-]. Type genus: Brentus Fabricius, 1787 [as Brenthus, unjustified emendation of type genus name by Illiger (1801), not in prevailing usage].
Tribe
Brentini
Billberg, 1820
Brenthides Billberg, 1820a: 40 [stem: Brent-]. Type genus: Brentus Fabricius, 1787 [as Brenthus, unjustified emendation of type genus name by Illiger (1801), not in prevailing usage].
Subtribe
Arrhenodina
Lacordaire, 1865
Arrhénodides Lacordaire, 1865: 425 [stem: Arrhenod-]. Type genus: Arrenodes sensu Lacordaire, 1865 [as Arrhenodes, incorrect subsequent spelling of type genus name, not in prevailing usage; not Arrenodes Schönherr 1823; syn. of Estenorhinus Lacordaire, 1865]. Comment: based on misidentified type genus; MAAZ and CHCL will submit an application to designate as type genus Arrenodes Schönherr, 1823 and correct the name of the tribe to Arrenodini; First Reviser (Arrhenodina Lacordaire, 1865 vs Belopherina Lacordaire, 1865 vs Belorhynchina Lacordaire, 1865 vs Eutrachelina Lacordaire, 1865) not determined, current usage maintained.
Bélophérides Lacordaire, 1865: 433 [stem: Belopher-]. Type genus: Belopherus Schönherr, 1833 [syn. of Belorhynchus Berthold, 1827]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1872: 317, as Belopherinae), generally accepted as in Sharp (1895a: 53, as Belopherina).
Bélorhynchides Lacordaire, 1865: 437 [stem: Belorhynch-]. Type genus: Belorhynchus Berthold, 1827. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form generally accepted as in Sharp (1895a: 63, as Belorhynchina); placed in synonymy with Arrhenodini by Alonso-Zarazaga and Lyal (2002: 10).
Eutrachélides Lacordaire, 1865: 438 [stem: Eutrachel-]. Type genus: Eutrachelus Berthold, 1827. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Senna (1892: 179, as Eutrachelinae), generally accepted as in Sharp (1900b: 385, as Eutrachelides [treated as Latin]).
Orychodi Senna, 1895: 213 [stem: Orychod-]. Type genus: Orychodes Pascoe, 1862.
Eupsalini Muizon, 1960: 170 [stem: Eupsalid-]. Type genus: Eupsalis sensu Kleine, 1927 [not Eupsalis Lacordaire, 1865; syn. of Orfilaia Haedo Rosii, 1955]. Comment: based on a misidentified type genus; incorrect original stem formation, not in prevailing usage.
Subtribe
Brentina
Billberg, 1820
Brenthides Billberg, 1820a: 40 [stem: Brent-]. Type genus: Brentus Fabricius, 1787 [as Brenthus, unjustified emendation of type genus name by Illiger (1801), not in prevailing usage].
Subtribe
Eremoxenina
Semenov, 1892
Amorphocephalides Power, 1879: 478 [stem: Amorphocephal-]. Type genus: Amorphocephalus Schönherr, 1840 [preoccupied genus name, not Amorphocephalus Bowdich, 1825 [Pisces]; syn. of Amorphocephala Damoiseau, 1966]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Eremoxenidae Semenov, 1892: 440 [stem: Eremoxen-]. Type genus: Eremoxenus Semenov, 1892.
Paussobrenthini Gestro, 1919: 270 [stem: Paussobrenth-]. Type genus: Paussobrenthus Gestro, 1919.
Tribe
Cyladini
Schönherr, 1823
Cylades Schönherr, 1823: column 1137 [stem: Cylad-]. Type genus: Cylas Latreille, 1802.
Tribe
Cyphagogini
Kolbe, 1892
Cyphagoginae Kolbe, 1892: 162 [stem: Cyphagog-]. Type genus: Cyphagogus Parry, 1849.
Subtribe
Atopobrentina
Damoiseau, 1965
Atopobrentini Damoiseau, 1965: 1 [stem: Atopobrent-]. Type genus: Atopobrentus Damoiseau, 1965.
Subtribe
Cyphagogina
Kolbe, 1892
Cyphagoginae Kolbe, 1892: 162 [stem: Cyphagog-]. Type genus: Cyphagogus Parry, 1849.
*Calodrominen Kolbe, 1916: 50 [stem: Calodrom-]. Type genus: Calodromus Guérin-Méneville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899.
Calodromini Kleine, 1922a: 148 [stem: Calodrom-]. Type genus: Calodromus Guérin-Méneville, 1832.
†Subtribe
Dominibrentina
Poinar, 2009
Dominibrentini Poinar, 2009: 51 [stem: Dominibrent-]. Type genus: Dominibrentus Poinar, 2009.
Subtribe
Hoplopisthiina
Senna and Calabresi, 1919
Hoplopisthi Senna and Calabresi, 1919: 65 [stem: Hoplopisthi-]. Type genus: Hoplopisthius Senna, 1892. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Stereodermina
Sharp, 1895
Stereodermina Sharp, 1895a: 7 [stem: Stereoderm-]. Type genus: Stereodermus Lacordaire, 1865.
Tribe
Pholidochlamydini
Damoiseau, 1962
Pholidochlamydinae Damoiseau, 1962: 26 [stem: Pholidochlamyd-]. Type genus: Pholidochlamys Lacordaire, 1865. Comment: the original spelling Pholidochlomydinae on page 26 is considered a lapsus calami since the the type genus is spelled correctly throughout and the family-group name is spelled correctly on page 18.
Tribe
Taphroderini
Lacordaire, 1865
Taphrodérides Lacordaire, 1865: 406 [stem: Taphroder-]. Type genus: Taphroderes Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1872: 317, as Taphroderinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 52, as Taphroderinae).
Ischnomérides Lacordaire, 1865: 414 [stem: Ischnomer-]. Type genus: Ischnomerus Schönherr, 1840 [preoccupied genus name, not Ischnomerus Labram and Imhoff, 1838 [Coleoptera: Curculionidae: Trachelizini]; syn. of Aulacoderes Chevrolat, 1839]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Schoenfeldt (1908: 2, as Ischnomeridae); permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Trachelizini
Lacordaire, 1865
Trachélizides Lacordaire, 1865: 417 [stem: Tracheliz-]. Type genus: Trachelizus Dejean, 1834. Comment: First Reviser (Trachelizini Lacordaire, 1865 vs Ithystenini Lacordaire, 1865) not determined, current usage maintained.
Subtribe
Acratina
Alonso-Zarazaga, Lyal, Bartolozzi and Sforzi, 1999
Némocéphalides Lacordaire, 1865: 459 [stem: Nemocephal-]. Type genus: Nemocephalus sensu Lacordaire, 1865 [not Nemocephalus Guérin-Méneville, 1827; syn. of Neacratus Alonso-Zarazaga et al., 1999]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Kolbe (1883: 79, as Nemocephali); based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).
Acratini Alonso-Zarazaga et al., 1999: 53 [stem: Acrat-]. Type genus: Acratus Lacordaire, 1865.
Subtribe
Ithystenina
Lacordaire, 1865
Ithysténides Lacordaire, 1865: 464 [stem: Ithysten-]. Type genus: Ithystenus Pascoe, 1862. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1872: 317, as Ithysteninae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 54, as Ithystenini).
Ischnomeri Kolbe, 1883: 74 [stem: Ischnomer-]. Type genus: Ischnomerus Labram and Imhoff, 1838.
Leptorrhynchidae Schoenfeldt, 1908: 69 [stem: Leptorhynch-]. Type genus: Leptorhynchus Guérin-Méneville, 1838 [as Leptorrhynchus, incorrect subsequent spelling of type genus name, not in prevailing usage; preoccupied genus name, not Leptorhynchus Clift, 1828 [Reptilia]; syn. of Ithystenus Pascoe, 1862]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Ozodocerini Jakobson, 1911b: 142 [stem: Ozodecer-]. Type genus: Ozodecerus Chevrolat, 1839. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Microtrachelizina
Zimmerman, 1994
Microtrachelizina Zimmerman, 1994b: 182 [stem: Microtracheliz-]. Type genus: Microtrachelizus Senna, 1893.
Subtribe
Pseudoceocephalina
Kleine, 1922
Céocéphalides Lacordaire, 1865: 444 [stem: Ceocephal-]. Type genus: Ceocephalus sensu Lacordaire, 1865 [not Ceocephalus Guérin-Méneville, 1833; Ceocephalus sensu Lacordaire, 1865 is a mixture of Orphanobrentus Damoiseau, 1962 and Pseudoceocephalus Kleine, 1920 (see Alonso-Zarazaga and Lyal 1999: 55)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Sharp (1900b: 385, as Ceocephalides [treated as Latin]); based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).
Uropterini Jakobson, 1911b: 142 [stem: Uropter-]. Type genus: Uroptera Berthold, 1827.
Pseudoceocephalidae Kleine, 1922b: 225 [stem: Pseudoceocephal-]. Type genus: Pseudoceocephalus Kleine, 1920. Comment: this family-group name is preceeded by two older names: Ceocephalina Lacordaire, 1865, which is based on a misidentified type genus, and the newly discovered name Uropterina Jakobson, 1911; we recommend that an application be submitted to the Commission to conserve usage of the well-established name Pseudoceocephalina Kleine, 1922.
Subtribe
Rhyticephalina
Kleine, 1922
Rhyticephalini Kleine, 1922c: 163 [stem: Rhyticephal-]. Type genus: Rhyticephalus Chevrolat, 1839.
Subtribe
Trachelizina
Lacordaire, 1865
Trachélizides Lacordaire, 1865: 417 [stem: Tracheliz-]. Type genus: Trachelizus Dejean, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1872: 317, as Trachelizinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 53, as Trachelizini); First Reviser (Trachelizina Lacordaire, 1865 vs Hephebocerina Lacordaire, 1865) not determined, current usage maintained.
Héphébocérides Lacordaire, 1865: 415 [stem: Hephebocer-]. Type genus: Hephebocerus Schönherr, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Pascoe (1872: 317, as Ephebocerinae [incorrect stem formation]).
Anchistenini Schedl, 1961: 198 [stem: Anchiste-]. Type genus: Anchisteus Kolbe, 1883. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Tychaeina
Schoenfeldt, 1908
Tychaeidae Schoenfeldt, 1908: 48 [stem: Tychae-]. Type genus: Tychaeus Fischer von Waldheim, 1823 [syn. of Nemorhinus Schönherr, 1823].
Tribe
Ulocerini
Schönherr, 1823
Ulocerides Schönherr, 1823: column 1137 [stem: Ulocer-]. Type genus: Ulocerus Schönherr, 1823.
Subfamily
Eurhynchinae
Lacordaire, 1863
Eurhynchides Lacordaire, 1863: 527 [stem: Eurhynch-]. Type genus: Eurhynchus Kirby, 1828 [placed on the Official List of Generic Names in Zoology (ICZN 1985e)].
†Tribe
Axelrodiellini
Legalov, 2009
Axeirodiellini Legalov, 2009c: 292 [stem: Axelrodiell-]. Type genus: Axelrodiellus Zherikhin and Gratshev, 2004. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Eurhynchini
Lacordaire, 1863
Eurhynchides Lacordaire, 1863: 527 [stem: Eurhynch-]. Type genus: Eurhynchus Kirby, 1828 [placed on the Official List of Generic Names in Zoology (ICZN 1985e)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Eurhynchinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 56, as Eurhynchidae); Eurhynchinae Lacordaire, 1863 placed on the Official List of Family-Group Names in Zoology (ICZN 1985e).
Eurhinini Kissinger, 1968: 10 [stem: Eurhin-]. Type genus: Eurhinus Kirby, 1819 [preoccupied genus name, not Eurhinus Illiger, 1807 [Coleoptera: Curculionidae: Baridini]; syn. of Eurhynchus Kirby, 1828]. Comment: replacement name for Eurhynchides Lacordaire, 1863 because of the homonymy of the type genus; junior homonym of Eurhinina Lacordaire, 1865 [Curculionidae: Baridinae]; permanently invalid (Art. 39): based on preoccupied type genus.
Subfamily
Apioninae
Schönherr, 1823
Apionides Schönherr, 1823: column 1136 [stem: Apion-]. Type genus: Apion Herbst, 1797. Comment: First Reviser (Apioninae Schönherr, 1823 vs Antliarhininae Schönherr, 1823) not determined, current usage maintained.
Supertribe
Apionitae
Schönherr, 1823
Apionides Schönherr, 1823: column 1136 [stem: Apion-]. Type genus: Apion Herbst, 1797.
Tribe
Apionini
Schönherr, 1823
Apionides Schönherr, 1823: column 1136 [stem: Apion-]. Type genus: Apion Herbst, 1797.
Subtribe
Apionina
Schönherr, 1823
Apionides Schönherr, 1823: column 1136 [stem: Apion-]. Type genus: Apion Herbst, 1797. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Api-).
Oxeostomatidae Gistel, 1856a: 374 [stem: Oxeostom-]. Type genus: Oxeostomum Gistel, 1856 [syn. of Apion Herbst, 1797]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Aplemonina
Kissinger, 1968
Aplemonini Kissinger, 1968: 16 [stem: Aplemon-]. Type genus: Aplemonus Schönherr, 1847.
Subtribe
Aspidapiina
Alonso-Zarazaga, 1990
Aspidapiini Alonso-Zarazaga, 1990: 52 [stem: Aspidapi-]. Type genus: Aspidapion Schilsky, 1901.
Subtribe
Catapiina
Alonso-Zarazaga, 1990
Catapiina Alonso-Zarazaga, 1990: 114 [stem: Catapi-]. Type genus: Catapion Schilsky, 1906.
Subtribe
Ceratapiina
Alonso-Zarazaga, 1990
Ceratapiini Alonso-Zarazaga, 1990: 42 [stem: Ceratapi-]. Type genus: Ceratapion Schilsky, 1901.
Subtribe
Exapiina
Alonso-Zarazaga, 1990
Exapiini Alonso-Zarazaga, 1990: 75 [stem: Exapi-]. Type genus: Exapion Bedel, 1887.
Subtribe
Ixapiina
Alonso-Zarazaga, 1990
Ixapiini Alonso-Zarazaga, 1990: 71 [stem: Ixapi-]. Type genus: Ixapion Roudier and Tempère, 1973.
Subtribe
Kalcapiina
Alonso-Zarazaga, 1990
Kalcapiini Alonso-Zarazaga, 1990: 55 [stem: Kalcapi-]. Type genus: Kalcapion Schilsky, 1906.
Subtribe
Malvapiina
Alonso-Zarazaga, 1990
Malvapiini Alonso-Zarazaga, 1990: 65 [stem: Malvapi-]. Type genus: Malvapion Hoffmann, 1958.
Subtribe
Metapiina
Alonso-Zarazaga, 1990
Metapiini Alonso-Zarazaga, 1990: 62 [stem: Metapi-]. Type genus: Metapion Schilsky, 1906.
Subtribe
Oxystomatina
Alonso-Zarazaga, 1990
Oxystomatini Alonso-Zarazaga, 1990: 111 [stem: Oxystomat-]. Type genus: Oxystoma Duméril, 1805.
Subtribe
Piezotrachelina
Voss, 1959
Piezotrachelini Voss, 1959: 51 [stem: Piezotrachel-]. Type genus: Piezotrachelus Schönherr, 1839.
Subtribe
Prototrichapiina
Wanat, 1995
Prototrichapiini Wanat, 1995: 19 [stem: Prototrichapi-]. Type genus: Prototrichapion Voss, 1959.
Subtribe
Synapiina
Alonso-Zarazaga, 1990
Synapiina Alonso-Zarazaga, 1990: 118 [stem: Synapi-]. Type genus: Synapion Schilsky, 1902.
Subtribe
Trichapiina
Alonso-Zarazaga, 1990
Trichapiina Alonso-Zarazaga, 1990: 116 [stem: Trichapi-]. Type genus: Trichapion Wagner, 1912.
Tribe
Chilapiini
Wanat, 2001
Chilapiitae Wanat, 2001: 366 [stem: Chilapi-]. Type genus: Chilapion Kissinger, 1968.
Tribe
Noterapiini
Kissinger, 2004
Noterapionini Kissinger, 2004: 243 [stem: Noterapi-]. Type genus: Noterapion Kissinger, 2002. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Podapiini
Wanat, 2001
Podapiitae Wanat, 2001: 366 [stem: Podapi-]. Type genus: Podapion Riley, 1883.
Tribe
Rhinorhynchidiini
Zimmerman, 1994
Rhinorhynchidiinae Zimmerman, 1994b: 337 [stem: Rhinorhynchidi-]. Type genus: Rhinorhynchidius Voss, 1922.
Supertribe
Antliarhinitae
Schönherr, 1823
Antliarhinides Schönherr, 1823: column 1137 [stem: Antliarhin-]. Type genus: Antliarhis Billberg, 1820 [as Antliarhinus, incorrect subsequent spelling of type genus name, not in prevailing usage].
Supertribe
Cybebitae
Lacordaire, 1863
Cybébides Lacordaire, 1863: 539 [stem: Cybeb-]. Type genus: Cybebus Schönherr, 1839. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Cybebinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 58, as Cybebini).
Supertribe
Mecolenitae
Wanat, 2001
Mecoleninae Wanat, 2001: 363 [stem: Mecolen-]. Type genus: Mecolenus Schönherr, 1847.
Supertribe
Myrmacicelitae
Zimmerman, 1994
Myrmacicelinae Zimmerman, 1994b: 349 [stem: Myrmacicel-]. Type genus: Myrmacicelus Chevrolat, 1833.
Tribe
Lispotheriini
Wanat, 2001
Lispotheriini Wanat, 2001: 365 [stem: Lispotheri-]. Type genus: Lispotherium Faust, 1891.
Tribe
Myrmacicelini
Zimmerman, 1994
Myrmacicelinae Zimmerman, 1994b: 349 [stem: Myrmacicel-]. Type genus: Myrmacicelus Chevrolat, 1833.
Supertribe
Rhadinocybitae
Alonso-Zarazaga, 1992
Rhadinocybini Alonso-Zarazaga, 1992: 193 [stem: Rhadinocyb-]. Type genus: Rhadinocyba Faust, 1889.
Tribe
Notapionini
Zimmerman, 1994
Notapionini Zimmerman, 1994b: 317 [stem: Notapion-]. Type genus: Notapion Zimmerman, 1994. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Notapi-).
Tribe
Rhadinocybini
Alonso-Zarazaga, 1992
Rhadinocybini Alonso-Zarazaga, 1992: 193 [stem: Rhadinocyb-]. Type genus: Rhadinocyba Faust, 1889.
Supertribe
Tanaitae
Schönherr, 1839
Tanaonides Schönherr, 1839: 447 [stem: Tana-]. Type genus: Tanaos Schönherr, 1826. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Ithycerinae
Schönherr, 1823
Ithycerides Schönherr, 1823: column 1136 [stem: Ithycer-]. Type genus: Ithycerus Schönherr, 1823.
Pachyrhinchidae Kirby, 1837: 203 [stem: Pachyrhynch-]. Type genus: Pachyrhynchus Kirby, 1837 [preoccupied genus name, not Pachyrhynchus Germar, 1824 [Coleoptera: Curculionidae: Otiorhynchini]; syn. of Ithycerus Schönherr, 1823]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
†Gobicarini Legalov, 2009c: 291 [stem: Gobicar-]. Type genus: Gobicar Gratshev and Zherikhin, 1999.
Subfamily
Microcerinae
Lacordaire, 1863
Microcérides Lacordaire, 1863: 20 [stem: Microcer-]. Type genus: Microcerus Schönherr, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Microcerinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 63, as Microcerinae).
*Épisides Lacordaire, 1863: 22 [stem: Epis-]. Type genus: Episus Billberg, 1820. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 224, as Episini), but not generally accepted as valid.
Subfamily
Nanophyinae
Gistel, 1848
Nanophyeidae Gistel, 1848: [7] [stem: Nanophy-]. Type genus: Nanophyes Schönherr, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1989b)].
Tribe
Corimaliini
Alonso-Zarazaga, 1989
Corimaliini Alonso-Zarazaga, 1989: 225 [stem: Corimali-]. Type genus: Corimalia Gozis, 1885.
Tribe
Nanophyini
Gistel, 1848
Nanophyeidae Gistel, 1848: [7] [stem: Nanophy-]. Type genus: Nanophyes Schönherr, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1989b)]. Comment: incorrect original stem formation, not in prevailing usage.
Family
Dryophthoridae
Schönherr, 1825
Dryophthorides Schönherr, 1825: column 588 [stem: Dryophthor-]. Type genus: Dryophthorus Germar, 1824 [placed on the Official List of Generic Names in Zoology (ICZN 1987c)].
Subfamily
Dryophthorinae
Schönherr, 1825
Dryophthorides Schönherr, 1825: column 588 [stem: Dryophthor-]. Type genus: Dryophthorus Germar, 1824 [placed on the Official List of Generic Names in Zoology (ICZN 1987c)].
Subfamily
Cryptodermatinae
Bovie, 1908
Oxyrhynchides Schönherr, 1823: column 1137 [stem: Oxyrhynch-]. Type genus: Oxyrhynchus Schönherr, 1823 [preoccupied genus name, not Oxyrhynchus Leach, 1818 [Pisces]; syn. of Cryptoderma Ritsema, 1885]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Cryptoderminae Bovie, 1908: 1 [stem: Cryptodermat-]. Type genus: Cryptoderma Ritsema, 1885. Comment: replacement name for Oxyrhynchinae Schönherr, 1823 because of the homonymy of the type genus; incorrect original stem formation, not in prevailing usage.
Subfamily
Orthognathinae
Lacordaire, 1865
Orthognathides Lacordaire, 1865: 311 [stem: Orthognath-]. Type genus: Orthognathus Schönherr, 1837.
Tribe
Orthognathini
Lacordaire, 1865
Orthognathides Lacordaire, 1865: 311 [stem: Orthognath-]. Type genus: Orthognathus Schönherr, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 233, as Orthognathini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 64, as Orthognathini).
Sipalides Lacordaire, 1865: 310 [stem: Sipal-]. Type genus: Sipalus Schönherr, 1825 [preoccupied genus name, not Sipalus Fischer, 1813 [Mammalia]; syn. of Sipalinus G. A. K. Marshall, 1943]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Sipalinae), generally accepted as in Heyne and Taschenberg (1907: 233, as Sipalini); permanently invalid (Art. 39): based on preoccupied type genus.
*Sipalininae G. A. K. Marshall, 1953: 117 [stem: Sipalin-]. Type genus: Sipalinus G. A. K. Marshall, 1943. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Sipalininae Zimmerman, 1993: 47, in key [stem: Sipalin-]. Type genus: Sipalinus G. A. K. Marshall, 1943.
Tribe
Rhinostomini
LeConte, 1874
Rhinidae J. L. LeConte, 1874b: 466 [stem: Rhin-]. Type genus: Rhina Latreille, 1806 [preoccupied genus name, not Rhina Schneider, 1801 [Pisces]; syn. of Rhinostomus Rafinesque, 1815]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Rhinostomini Kuschel, 1995: 24, in key [stem: Rhinostom-]. Type genus: Rhinostomus Rafinesque, 1815 [placed on the Official List of Generic Names in Zoology (ICZN 1955a)].
Subfamily
Rhynchophorinae
Schönherr, 1833
Rhynchophorides Schönherr, 1833: 26 [stem: Rhynchophor-]. Type genus: Rhynchophorus Herbst, 1795.
Tribe
Diocalandrini
Zimmerman, 1993
Diocalandrini Zimmerman, 1993: 99 [stem: Diocalandr-]. Type genus: Diocalandra Faust, 1894.
Tribe
Litosomini
Lacordaire, 1865
Calandrina C. G. Thomson, 1859: 145 [stem: Calandr-]. Type genus: Calandra Gistel, 1848 [placed on the Official Index of Invalid and Rejected Generic Names in Zoology (ICZN 1959b); preoccupied genus name, not Calandra Clairville, 1798 [Coleoptera: Curculionidae: Rhynchophorinae: Sphenophorini]; syn. of Sitophilus Schönherr, 1837]. Comment: permanently invalid (Art. 39): based on suppressed type genus.
Litosomides Lacordaire, 1865: 303 [stem: Litosom-]. Type genus: Litosomus Lacordaire, 1865 [syn. of Toxorhinus Lacordaire, 1865]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1895: 224, as Litosominorum), generally accepted as in Alonso-Zarazaga and Lyal (1999: 65, as Litosomini).
Sitophili Csiki, 1936: 68 [stem: Sitophil-]. Type genus: Sitophilus Schönherr, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1959b)]. Comment: name proposed to replace Calandrina C. G. Thomson, 1865 because of the synonymy of the type genus (Calandra auctorum is a synonym of Sitophilus Schönherr, 1837).
Tribe
Ommatolampini
Lacordaire, 1865
Ommatolampides Lacordaire, 1865: 276 [stem: Ommatolamp-]. Type genus: Ommatolampes Schönherr, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 233, as Ommatolampini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 66, as Ommatolampini).
Tribe
Polytini
Zimmerman, 1993
Polytini Zimmerman, 1993: 94 [stem: Polyt-]. Type genus: Polytus Faust, 1894.
Tribe
Rhynchophorini
Schönherr, 1833
Rhynchophorides Schönherr, 1833: 26 [stem: Rhynchophor-]. Type genus: Rhynchophorus Herbst, 1795.
Tribe
Sphenophorini
Lacordaire, 1865
Calandraedes Billberg, 1820a: 40 [stem: Calandr-]. Type genus: Calandra Clairville, 1798 [placed on the Official Index of Invalid and Rejected Generic Names in Zoology (ICZN 1959b)]. Comment: permanently invalid (Art. 39): based on suppressed type genus; several family-group names based on Calandra Clairville, 1798 were placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1959b).
Sphénophorides Lacordaire, 1865: 286 [stem: Sphenophor-]. Type genus: Sphenophorus Schönherr, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1959b)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1876: 330, as Sphenophorini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 66, as Sphenophorini); First Reviser (Sphenophorini Lacordaire, 1865 vs Sphenocorynini Lacordaire, 1865) not determined, current usage maintained.
Sphénocorynides Lacordaire, 1865: 279 [stem: Sphenocoryn-]. Type genus: Sphenocorynes Schönherr, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Aurivillius (1886: 97, as Sphenocoryninae), generally accepted as in Csiki (1936: 20, as Sphenocoryni).
Oxyopisthinae Kolbe, 1899: 5 [stem: Oxyopisth-]. Type genus: Oxyopisthen sensu Lacordaire, 1865 [not Oxyopisthen J. Thomson, 1858; syn. of Korotyaevius Alonso-Zarazaga and Lyal, 1999]. Comment: based on misidentified type genus.
Subfamily
Stromboscerinae
Lacordaire, 1865
Stromboscérides Lacordaire, 1865: 306 [stem: Stromboscer-]. Type genus: Stromboscerus Schönherr, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Strombocerinae [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 68, as Stromboscerinae).
Family
Brachyceridae
Billberg, 1820
Brachycerides Billberg, 1820a: 39 [stem: Brachycer-]. Type genus: Brachycerus A. G. Olivier, 1789.
Subfamily
Brachycerinae
Billberg, 1820
Brachycerides Billberg, 1820a: 39 [stem: Brachycer-]. Type genus: Brachycerus A. G. Olivier, 1789. Comment: the tribal classification used here follows Oberprieler (2010).
Tribe
Brachycerini
Billberg, 1820
Brachycerides Billberg, 1820a: 39 [stem: Brachycer-]. Type genus: Brachycerus A. G. Olivier, 1789.
Protomantiinae Aurivillius, 1886: 21, in key [stem: Protomanti-]. Type genus: Protomantis Schönherr, 1840.
Tribe
Byrsopini
Germar, 1829
Cryptopsides Schönherr, 1826: 65 [stem: Cryptop-]. Type genus: Cryptops Schönherr, 1823 [preoccupied genus name, not Cryptops Leach, 1814 [Chilopoda]; syn. of Byrsops Germar, 1829]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Byrsoptides Germar, 1829: 358 [stem: Byrsop-]. Type genus: Byrsops Germar, 1829. Comment: incorrect original stem formation, not in prevailing usage.
Byrsopsides Schönherr, 1833: 14 [stem: Byrsop-]. Type genus: Byrsops Schönherr, 1833 [preoccupied genus name, not Byrsops Germar, 1829 [Coleoptera: Curculionidae: Byrsopini]; syn. of Byrsops Germar, 1829]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Brotheinae G. A. K. Marshall, 1907: 89 [stem: Brothe-]. Type genus: Brotheus Stephens, 1829. Comment: junior homonym of Brotheini Simon, 1879 in Scorpiones (type genus Brotheas C. L. Koch, 1837); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Brotheusini Alonso-Zarazaga and Lyal, 2006: 24 [stem: Brotheus-]. Type genus: Brotheus Stephens, 1829. Comment: unnecessary replacement name for Brotheinae Marshall, 1907 because of the homonymy of the family-group name; Brotheinae/-ini/-ina Simon, 1879 (type genus Brotheas C. L. Koch, 1837) has been used as valid in Scorpiones in recent literature, e.g., Prendini and W. C. Wheeler (2005); the stem used by Alonso-Zarazaga and Lyal (2006) was chosen in accordance with Art. 29.6.
Subfamily
Cryptolarynginae
Schalkwyk, 1966
Cryptopharynginae G. A. K. Marshall, 1957: 18 [stem: Cryptopharyng-]. Type genus: Cryptopharynx G. A. K. Marshall, 1957 [preoccupied genus name, not Cryptopharynx Kahl, 1928 [Protozoa]; syn. of Cryptolarynx Schalkwyk, 1966]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Cryptolarynginae Schalkwyk, 1966: 745 [stem: Cryptolaryng-]. Type genus: Cryptolarynx Schalkwyk, 1966. Comment: replacement name for Cryptopharynginae G. A. K. Marshall, 1957 because of the homonymy of the type genus.
Periegini Legalov, 2003: 68 [stem: Perieg-]. Type genus: Perieges Schönherr, 1842.
Subfamily
Erirhininae
Schönherr, 1825
Erirhinides Schönherr, 1825: column 582 [stem: Erirhin-]. Type genus: Erirhinus Schönherr, 1825 [syn. of Notaris Germar, 1817].
Tribe
Aonychini
Zimmerman, 1993
Aonychini Zimmerman, 1993: 136, in key [stem: Aonych-]. Type genus: Aonychus Schönherr, 1844. Comment: junior homonym of Aonychini Davis, 1978 in Mammalia (type genus Aonyx Lesson, 1827); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Aonychusini Zimmerman, 1999: 72 [stem: Aonychus-]. Type genus: Aonychus Schönherr, 1844. Comment: unnecessary replacement name for Aonychini Zimmerman, 1993 because of the homonymy of the family-group name; the stem used by Zimmerman (1999) was chosen in accordance with Art. 29.6.
Tribe
Arthrostenini
Reitter, 1913
Arthrostenini Reitter, 1913b: 57, in key [stem: Arthrosten-]. Type genus: Arthrostenus Schönherr, 1826.
†Tribe
Cretuliini
Legalov, 2009
Cretuliini Legalov, 2009c: 293 [stem: Cretuli-]. Type genus: Cretulio Zherikhin, 1993.
Tribe
Erirhinini
Schönherr, 1825
Erirhinides Schönherr, 1825: column 582 [stem: Erirhin-]. Type genus: Erirhinus Schönherr, 1825 [syn. of Notaris Germar, 1817].
Rhynchaenides Latreille, 1828: 599 [stem: Rhynchaen-]. Type genus: Rhynchaenus sensu Latreille, 1828 [not Rhynchaenus Clairville, 1798; syn. of Notaris Germar, 1817]. Comment: based on a misidentified type genus; an application will need to be submitted to the Commission to suppress this name for the Principles of Priority and Homonymy (Art. 65.2.1) if Rhyncahenidae Blanchard, 1853 in Curculioninae: Rhamphini: Rhamphina is to be used as valid in the future.
Notarini Zumpt, 1929: 216 [stem: Notar-]. Type genus: Notaris sensu Tournier, 1874 [not Notaris Germar, 1817; syn. of Tournotaris Alonso-Zarazaga and Lyal, 1999]. Comment: based on a misidentified type genus.
Notodermina Voss, 1952: 196, in note [stem: Notoderm-]. Type genus: Notodermus Desbrochers des Loges, 1875 [syn. of Procas Stephens, 1831].
Tribe
Himasthlophallini
Zherikhin, 1991
Himasthlophallini Zherikhin, 1991: 31 [stem: Himasthlophall-]. Type genus: Himasthlophallus Egorov and Zherikhin, 1991.
Tribe
Stenopelmini
LeConte, 1876
Stenopelmi J. L. LeConte, 1876: 179 [stem: Stenopelm-]. Type genus: Stenopelmus Schönherr, 1835.
Lissorhoptrinae Böving and Craighead, 1931: 67, in key [stem: Lissorhoptr-]. Type genus: Lissorhoptrus J. L. LeConte, 1876.
Tribe
Tadiini
Zimmerman, 1993
Tadiini Zimmerman, 1993: 136, in key [stem: Tadi-]. Type genus: Tadius Pascoe, 1885.
Tribe
Tanysphyrini
Gistel, 1848
Tanysphyridae Gistel, 1848: [8] [stem: Tanysphyr-]. Type genus: Tanysphyrus Germar, 1817.
Brachypi J. L. LeConte, 1876: 180 [stem: Brachypod-]. Type genus: Brachypus Schönherr, 1826 [preoccupied genus name, not Brachypus Meyer, 1814 [Aves]; syn. of Brachygyius G. A. K. Marshall, 1939]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Subfamily
Ocladiinae
Lacordaire, 1865
Ocladiides Lacordaire, 1865: 79 [stem: Ocladi-]. Type genus: Ocladius Schönherr, 1825.
Tribe
Desmidophorini
Morimoto, 1962
Desmidophorinae Morimoto, 1962a: 32, in key [stem: Desmidophor-]. Type genus: Desmidophorus Dejean, 1835.
Tribe
Ocladiini
Lacordaire, 1865
Ocladiides Lacordaire, 1865: 79 [stem: Ocladi-]. Type genus: Ocladius Schönherr, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hustache (1925: 9, as Ocladiini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 63, as Ocladiini).
Subfamily
Raymondionyminae
Reitter, 1913
Raymondionymini Reitter, 1913b: 58, in key [stem: Raymondionym-]. Type genus: Raymondionymus Wollaston, 1873.
Tribe
Myrtonymini
Kuschel, 1990
Myrtonymina Kuschel, 1990: 80 [stem: Myrtonym-]. Type genus: Myrtonymus Kuschel, 1990.
Tribe
Raymondionymini
Reitter, 1913
Raymondionymini Reitter, 1913b: 58, in key [stem: Raymondionym-]. Type genus: Raymondionymus Wollaston, 1873.
Family
Curculionidae
Latreille, 1802
Curculionites Latreille, 1802: 195 [stem: Curculion-]. Type genus: Curculio Linnaeus, 1758.
Subfamily
Curculioninae
Latreille, 1802
Curculionites Latreille, 1802: 195 [stem: Curculion-]. Type genus: Curculio Linnaeus, 1758.
Tribe
Acalyptini
Thomson, 1859
Acalyptina C. G. Thomson, 1859: 143 [stem: Acalypt-]. Type genus: Acalyptus Schönherr, 1833.
Subtribe
Acalyptina
Thomson, 1859
Acalyptina C. G. Thomson, 1859: 143 [stem: Acalypt-]. Type genus: Acalyptus Schönherr, 1833.
Subtribe
Derelomina
Lacordaire, 1865
Dérélomides Lacordaire, 1865: 9 [stem: Derelom-]. Type genus: Derelomus Schönherr, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Marseul (1866: 109, as Derelomidae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 77, as Derelomini).
Hoplorrhinina Champion, 1903: 277 [stem: Hoplorhin-]. Type genus: Hoplorhinus Chevrolat, 1878 [as Hoplorrhinus, incorrect subsequent spelling of type genus name, not in prevailing usage; syn. of Celetes Schönherr, 1836]. Comment: incorrect original stem formation, not in prevailing usage.
Celetini Voss, 1954: 344 [stem: Celet-]. Type genus: Celetes Schönherr, 1836. Comment: description by indication (distinguishing characters given in Champion (1903: 277, as Hoplorrhinina)); name proposed to replace Hoplorhinini Champion, 1903 because of synonymy of the type genus; Celetini is not in prevailing usage (Art. 40.2.1).
Subtribe
Notolomina
Franz, 2006
Notolomina N. M. Franz, 2006: 276 [stem: Notolom-]. Type genus: Notolomus J. L. LeConte, 1876.
Subtribe
Phyllotrogina
Franz, 2006
Phyllotrogina N. M. Franz, 2006: 276 [stem: Phyllotrog-]. Type genus: Phyllotrox Schönherr, 1843.
Subtribe
Staminodeina
Franz, 2006
Staminodeina N. M. Franz, 2006: 277 [stem: Staminode-]. Type genus: Staminodeus Franz, 2001.
Tribe
Acentrusini
Alonso-Zarazaga, 2005
Acentrina Seidlitz, 1890: 608 [stem: Acentr-]. Type genus: Acentrus Schönherr, 1845 [preoccupied genus name, not Acentrus Duponchel, 1839 [Coleoptera: Curculionidae: Curculioninae: Acentrusini]; syn. of Acentrus Duponchel, 1839]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Acentrusini Alonso-Zarazaga, 2005: 92 [stem: Acentrus-]. Type genus: Acentrus Duponchel, 1839. Comment: replacement name for Acentrini Seidlitz, 1890 because of the homonymy of the family-group name; the stem used by Alonso-Zarazaga (2005) was chosen in accordance with Art. 29.6.
Tribe
Ancylocnemidini
Voss, 1962
Ancylocnemini Voss, 1962: 339 [stem: Ancylocnemid-]. Type genus: Ancylocnemis G. A. K. Marshall, 1920. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Anthonomini
Thomson, 1859
Anthonomina C. G. Thomson, 1859: 144 [stem: Anthonom-]. Type genus: Anthonomus Germar, 1817.
*Brachonydes Tournier, 1874: 68, in key [stem: Brachonych-]. Type genus: Brachonyx Schönherr, 1825. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Tournier (1874); Alonso-Zarazaga and Lyal (1999: 74) attributed this name to the correct author but did not use it as a valid taxon; incorrect original stem formation, not in prevailing usage.
Loncophorini Pierce, 1916: 467 [stem: Loncophor-]. Type genus: Loncophorus Chevrolat, 1832.
Bradybatini Pierce, 1919: 26, in key [stem: Bradybat-]. Type genus: Bradybatus Germar, 1824.
Brachonychina Voss, 1944: 38, in key [stem: Brachonych-]. Type genus: Brachonyx Schönherr, 1825.
Tachypterellina Voss, 1944: 38, in key [stem: Tachypterell-]. Type genus: Tachypterellus Fall and Cockerell, 1907 [syn. of Anthonomus Germar, 1817].
Tribe
Camarotini
Schönherr, 1833
Camarotides Schönherr, 1833: 4 [stem: Camarot-]. Type genus: Camarotus Germar, 1817.
Subtribe
Camarotina
Schönherr, 1833
Camarotides Schönherr, 1833: 4 [stem: Camarot-]. Type genus: Camarotus Germar, 1817.
Subtribe
Prionomerina
Lacordaire, 1863
Prionomérides Lacordaire, 1863: 598 [stem: Prionomer-]. Type genus: Prionomerus Schönherr, 1835 [syn. of Odontopus Say, 1831]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Prionomerinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 75, as Prionomerina).
Tribe
Ceratopodini
Lacordaire, 1863
Cératopides Lacordaire, 1863: 589 [stem: Ceratopod-]. Type genus: Ceratopus Schönherr, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Ceratopinae [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 76, as Ceratopodini); incorrect original stem formation, not in prevailing usage.
Tribe
Cionini
Schönherr, 1825
Cionides Schönherr, 1825: column 587 [stem: Cion-]. Type genus: Cionus Clairville, 1798.
Tribe
Cranopoeini
Kuschel, 2009
Cranopoeini Kuschel, 2009: 44 [stem: Cranopoe-]. Type genus: Cranopoeus G. A. K. Marshall, 1931.
Tribe
Cryptoplini
Lacordaire, 1863
Cryptoplides Lacordaire, 1863: 486 [stem: Cryptopl-]. Type genus: Cryptoplus Erichson, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1876: 175, as Cryptopli), generally accepted as in Alonso-Zarazaga and Lyal (1999: 76, as Cryptoplini).
Haplonycides Lacordaire, 1865: 16 [stem: Haplonych-]. Type genus: Haplonyx Schönherr, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Haplonychinae), generally accepted as in Schenkling and G. A. K. Marshall (1936: 1, as Haplonychinae); incorrect original stem formation, not in prevailing usage; this name is a junior homonym of Haplonychini Burmeister, 1855 (type genus Haplonycha Dejean, 1836) proposed in Scarabaeidae and now a synonym of Liparetrini H. C. C. Burmeister, 1855; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1) if either of those names is to be used as valid in the future.
Tribe
Curculionini
Latreille, 1802
Curculionites Latreille, 1802: 195 [stem: Curculion-]. Type genus: Curculio Linnaeus, 1758.
Subtribe
Curculionina
Latreille, 1802
Curculionites Latreille, 1802: 195 [stem: Curculion-]. Type genus: Curculio Linnaeus, 1758.
Balaninidae Gistel, 1848: [7] [stem: Balanin-]. Type genus: Balaninus Germar, 1817 [syn. of Curculio Linnaeus, 1758].
Subtribe
Pseudobalaninina
Heller, 1925
Pseudobalaninina Heller, 1925b: 94, in key [stem: Pseudobalanin-]. Type genus: Pseudobalaninus Faust, 1889.
Subtribe
Timolina
Heller, 1925
Timolina Heller, 1925b: 94, in key [stem: Timol-]. Type genus: Timola Pascoe, 1886.
Tribe
Diabathrariini
Lacordaire, 1863
Diabathrariides Lacordaire, 1863: 407 [stem: Diabathrari-]. Type genus: Diabathrarius Schönherr, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Diabathrariinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 141, as Diabathrariini); transfer from Cyclominae by Oberprieler (2010).
Tribe
Ellescini
Thomson, 1859
Elleschina C. G. Thomson, 1859: 143 [stem: Ellesc-]. Type genus: Ellescus Dejean, 1821 [as Elleschus, unjustified emendation of type genus name by Schönherr (1837), not in prevailing usage].
Subtribe
Dorytomina
Bedel, 1886
Dorytomini Bedel, 1886: 280 [stem: Dorytom-]. Type genus: Dorytomus Germar, 1817.
Subtribe
Ellescina
Thomson, 1859
Elleschina C. G. Thomson, 1859: 143 [stem: Ellesc-]. Type genus: Ellescus Dejean, 1821 [as Elleschus, unjustified emendation of type genus name by Schönherr (1837), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Erodiscini
Lacordaire, 1863
Erodiscides Lacordaire, 1863: 566 [stem: Erodisc-]. Type genus: Erodiscus sensu Lacordaire, 1863 [Lacordaire excluded the type species of Erodiscus Schönherr, 1825 to place it in Toxeutes Schönherr, 1843; the five species that Lacordaire included in Erodiscus are now placed in Sicoderus Vanin, 1986, Prosicoderus Vanin, 1986 and Pimelerodius Vanin, 1986]. Comment: based on a misidentified type genus; MAAZ and CHCL will submit an application for the conservation of this name by designating Erodiscus Schönherr, 1825 as its type genus.
Tribe
Eugnomini
Lacordaire, 1863
Eugnomides Lacordaire, 1863: 499 [stem: Eugnom-]. Type genus: Eugnomus Schönherr, 1847.
Subtribe
Eugnomina
Lacordaire, 1863
Eugnomides Lacordaire, 1863: 499 [stem: Eugnom-]. Type genus: Eugnomus Schönherr, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1876: 174, as Eugnomi), generally accepted as in Alonso-Zarazaga and Lyal (1999: 79, as Eugnomini); First Reviser found (Eugnomina Lacordaire, 1863 vs Scolopterina Lacordaire, 1863) is Voss (1937a: 37).
Scoloptérides Lacordaire, 1863: 565 [stem: Scolopter-]. Type genus: Scolopterus A. White, 1846. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Scolopterinae), generally accepted as in Broun (1880: 472 as Scolopteridae).
Pactolina Voss, 1936: 112, in key [stem: Pactol-]. Type genus: Pactola Pascoe, 1876.
Stephanorhynchina Voss, 1936: 112 [stem: Stephanorhynch-]. Type genus: Stephanorhynchus A. White, 1846.
Subtribe
Meriphina
Marshall, 1937
Meriphinae G. A. K. Marshall, 1937a: 329 [stem: Meriph-]. Type genus: Meriphus Erichson, 1842.
Tribe
Gonipterini
Lacordaire, 1863
Goniptérides Lacordaire, 1863: 391 [stem: Gonipter-]. Type genus: Gonipterus Schönherr, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Gonipterinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 141, as Gonipterini); transfer from Cyclominae by Oberprieler (2010).
Tribe
Mecinini
Gistel, 1848
Mecinidae Gistel, 1848: [7] [stem: Mecin-]. Type genus: Mecinus Germar, 1821.
Gymnetrina C. G. Thomson, 1859: 143 [stem: Gymnetr-]. Type genus: Gymnetron Schönherr, 1825. Comment: junior homonym of Gymnetrinae Swainson, 1839 in Pisces (type genus Gymnetrus Bloch, 1795).
Miarides Tournier, 1874: 66 [stem: Miar-]. Type genus: Miarus Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Pierce (1919: 30, as Miarinae).
Miarini Zherikhin, 1991: 123 [stem: Miar-]. Type genus: Miarus Schönherr, 1826. Comment: family-group name proposed as new without reference to Miarides Tournier, 1874.
Tribe
Nerthopini
Lacordaire, 1865
Nerthopides Lacordaire, 1865: 19 [stem: Nerthop-]. Type genus: Nerthops Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Nerthopinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 80, as Nerthopini); precedence (Nerthopini Lacordaire, 1865 vs Acallopistini Lacordaire, 1865) given to taxon originally proposed at the higher rank (Art. 24.1).
*Microstylides Lacordaire, 1865: 20 [stem: Microstyl-]. Type genus: Microstylus Schönherr, 1847. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1865); subsequent usage of Microstylinae by Hoffmann (1968: 27) and Microstylidae by Ienistea (1986: 33) did not validate this name because these authors did not attribute their names to Lacordaire, 1865; Microstylinae Hoffmann, 1968: 27 and Microstylidae Ienistea, 1986: 33 are also unavailable because they were proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Acallopistides Lacordaire, 1865: 22 [stem: Acallopist-]. Type genus: Acallopistus Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hustache (1932: 371, as Acallopistinae), generally accepted as in Hustache (1934: 3, as Acallopistini).
Archolabinae Voss, 1929a: 200 [stem: Archolab-]. Type genus: Archolabus Voss, 1929.
Tribe
Otidocephalini
Lacordaire, 1863
Otidocéphalides Lacordaire, 1863: 568 [stem: Otidocephal-]. Type genus: Otidocephalus Chevrolat, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Otidocephalinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 80, as Otidocephalini).
Tribe
Piazorhinini
Lacordaire, 1863
Piazorhinides Lacordaire, 1863: 601 [stem: Piazorhin-]. Type genus: Piazorhinus Schönherr, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kirsch (1875: 162, as Piazorhinidae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 81, as Piazorhinini).
Tribe
Prionobrachiini
Hustache, 1938
Prionobrachiina Hustache, 1938: 90, in key [stem: Prionobrachi-]. Type genus: Prionobrachium Faust, 1894.
Tribe
Pyropini
Lacordaire, 1865
Pyropides Lacordaire, 1865: 187 [stem: Pyrop-]. Type genus: Pyropus Schönherr, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Pyropinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 81, as Pyropini).
Tribe
Rhamphini
Rafinesque, 1815
Ramphoria Rafinesque, 1815: 115 [stem: Rhamph-]. Type genus: Rhamphus Clairville, 1798 [as Ramphorus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Dinorhopalina
Voss, 1936
Dinorrhopalina Voss, 1936: 112, in key [stem: Dinorhopal-]. Type genus: Dinorhopala Pascoe, 1860 [as Dinorrhopala, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Ixalmina
Voss, 1936
Ixalmina Voss, 1936: 112, in key [stem: Ixalm-]. Type genus: Ixalma Pascoe, 1871.
Subtribe
Rhamphina
Rafinesque, 1815
Ramphoria Rafinesque, 1815: 115 [stem: Rhamph-]. Type genus: Rhamphus Clairville, 1798 [as Ramphorus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Rhamphides Schönherr, 1823: column 1136 [stem: Rhamph-]. Type genus: Rhamphus Clairville, 1798. Comment: family-group name proposed as new without reference to Ramphoria Rafinesque, 1815.
Orchestides Latreille, 1828: 601 [stem: Orchest-]. Type genus: Orchestes Illiger, 1798.
Rhynchenidae Blanchard, 1853: 240 [stem: Rhynchaen-]. Type genus: Rhynchaenus Clairville, 1798. Comment: incorrect original stem formation, not in prevailing usage; an application will need to be submitted to the Commission to suppress Rhynchaenides Latreille, 1828 (based on the misidentified type genus Rhynchaenus sensu Latreille, 1828) for the Principles of Priority and Homonymy (Art. 65.2.1) if this name is to be used as valid.
Subtribe
Tachygonina
Lacordaire, 1865
Tachygonides Lacordaire, 1865: 167 [stem: Tachygon-]. Type genus: Tachygonus Schönherr, 1833 [syn. of Tachygonus Guérin-Méneville, 1833]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Tachygoninae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 82, as Tachygonina); the type genus for Lacordaire’s Tachygonides is Tachygonus Schönherr, 1833; an application will be submitted by MAAZ and CHCL to the Commission to designate Tachygonus Guérin-Méneville, [1833] as the type genus of this family-group name.
Tribe
Smicronychini
Seidlitz, 1891
nomen protectum
Desmorhines J. L. LeConte, 1876: 167 [stem: Desmorin-]. Type genus: Desmoris J. L. LeConte, 1876 [syn. of Smicronyx Schönherr, 1843]. Comment: nomen oblitum (see Appendix 1); incorrect original stem formation, not in prevailing usage.
Smicronycina Seidlitz, 1891 [Gatt.]: 162 [stem: Smicronych-]. Type genus: Smicronyx Schönherr, 1843. Comment: nomen protectum (see Appendix 1); incorrect original stem formation, not in prevailing usage.
Tribe
Sphaeriopoeini
Kuschel, 2003
Sphaeriopoeini Kuschel, 2003: 63 [stem: Sphaeriopoe-]. Type genus: Sphaeriopoeus Kuschel, 2003.
Tribe
Storeini
Lacordaire, 1863
Storéides Lacordaire, 1863: 494 [stem: Store-]. Type genus: Storeus Schönherr, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kirsch (1877: 170, as Storeidae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 82, as Storeini).
Hybomorphides Lacordaire, 1865: 141 [stem: Hybomorph-]. Type genus: Hybomorphus Saunders and Jekel, 1855. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Hustache (1936: 261, as Hybomorphina).
Tribe
Styphlini
Jekel, 1861
Styphlidae Jekel, 1861: 274 [stem: Styphl-]. Type genus: Styphlus Schönherr, 1826.
*Orthochaetini A. Winkler, 1932: 1544 [stem: Orthochaet-]. Type genus: Orthochaetes Germar, 1824. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Orthocaetina Morimoto, 1962a: 56, in key [stem: Orthochaet-]. Type genus: Orthochaetes Germar, 1824. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Tychiini
Gistel, 1848
Tychiidae Gistel, 1848: [7] [stem: Tychi-]. Type genus: Tychius Germar, 1817.
Subtribe
Demimaeina
Voss, 1937
Demimaeini Voss, 1937b: 144 [stem: Demimae-]. Type genus: Demimaea Pascoe, 1870.
Subtribe
Lignyodina
Bedel, 1884
Lignyodini Bedel, 1884a: 66, in key [stem: Lignyod-]. Type genus: Lignyodes Dejean, 1835.
Subtribe
Ochyromerina
Voss, 1935
Ochyromerina Voss, 1935a: 228 [stem: Ochyromer-]. Type genus: Ochyromera Pascoe, 1874.
Endaeini Voss, 1958: 101, in key [stem: Endae-]. Type genus: Endaeus Schönherr, 1826.
Subtribe
Tychiina
Gistel, 1848
Tychiidae Gistel, 1848: [7] [stem: Tychi-]. Type genus: Tychius Germar, 1817.
Miccotrogidae Gistel, 1856a: 371 [stem: Miccotrog-]. Type genus: Miccotrogus Schönherr, 1825 [syn. of Tychius Germar, 1817].
Sibynidae Marseul, 1863: 239 [stem: Sibini-]. Type genus: Sibinia Germar, 1817 [as Sibynes, unjustified emendation of type genus name by Schönherr (1825), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Ulomascini
Lacordaire, 1865
Ulomascides Lacordaire, 1865: 184 [stem: Ulomasc-]. Type genus: Ulomascus Fairmaire, 1848. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Ulomascinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 86, as Ulomascini).
Tribe
Viticiini
Morimoto, 1983
Viticiinae Morimoto, 1983: 55 [stem: Vitici-]. Type genus: Viticis Lea, 1930. Comment: transfer from Cyclominae by Oberprieler (2010).
Subfamily
Bagoinae
Thomson, 1859
nomen protectum
Lypriidae Gistel, 1848: [7] [stem: Lypr-]. Type genus: Lyprus Schönherr, 1826 [syn. of Bagous Germar, 1817]. Comment: as pointed out by Alonso-Zarazaga and Lyal (2002: 17) this is the oldest available name for the subfamily; Lyprinae Gistel, 1848 was recently considered a nomen oblitum by Colonnelli (2003: 7) however the necessary supporting references were not provided; we hereby consider Lyprinae Gistel 1848 as a nomen oblitum (see supporting references in Appendix 1); incorrect original stem formation, not in prevailing usage.
Bagoina C. G. Thomson, 1859: 135 [stem: Bago-]. Type genus: Bagous Germar, 1817. Comment: nomen protectum (see Appendix 1).
Hydronomides Lacordaire, 1863: 483 [stem: Hydronom-]. Type genus: Hydronomus Schönherr, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1876: 182, as Hydronomi), generally accepted as in Zimmerman (1993: 150, as Hydronomini).
Pseudobagoini Sharp, 1917: 26 [stem: Pseudobago-]. Type genus: Pseudobagous Sharp, 1917.
Subfamily
Baridinae
Schönherr, 1836
Baridides Schönherr, 1836: 636 [stem: Barid-]. Type genus: Baris Germar, 1817 [as Baridius, unjustified emendation of type genus name by Schönherr (1825), not in prevailing usage].
Tribe
Ambatini
Lacordaire, 1863
Ambatides Lacordaire, 1863: 512 [stem: Ambat-]. Type genus: Ambates Schönherr, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Ambatinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 93, as Ambatini).
*Pteracanthides Chevrolat, 1878: 341 [stem: Pteracanth-]. Type genus: Pteracanthus Dejean, 1835. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; a family-group name with the same stem, Pteracanthina J. Thomson, 1864 (type genus Pteracantha Newman, 1838), is available in Coleoptera: Cerambycidae.
Tribe
Anopsilini
Bondar, 1942
Anopsilini Bondar, 1942: 253 [stem: Anopsil-]. Type genus: Anopsilus Kirsch, 1870.
Tribe
Apostasimerini
Schönherr, 1844
Apostasimerides Schönherr, 1844: 1 [stem: Apostasimer-]. Type genus: Apostasimerus Schönherr, 1844. Comment: see Prena (2009: 33) for details on authorship and taxa included in this tribe.
Subtribe
Apostasimerina
Schönherr, 1844
*Apostasimerides Schönherr, 1836: 557 [stem: Apostasimer-]. Type genus: Apostasimerus Schönherr, 1844. Comment: family-group name unavailable (Art. 11.7.1.1): not based on an available genus name at the time.
Apostasimerides Schönherr, 1844: 1 [stem: Apostasimer-]. Type genus: Apostasimerus Schönherr, 1844.
Subtribe
Madopterina
Lacordaire, 1865
Madoptérides Lacordaire, 1865: 243 [stem: Madopter-]. Type genus: Madopterus Schönherr, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kirsch (1870: 218, as Madopteridae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 96, as Madopterini).
Subtribe
Thaliabaridina
Bondar, 1943
Thaliabarini Bondar, 1943b: 131 [stem: Thaliabarid-]. Type genus: Thaliabaris Bondar, 1943. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Torcina
Bondar, 1943
Torcocina Bondar, 1943a: 53 [stem: Torc-]. Type genus: Torcus Casey, 1922. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Zygobaridina
Pierce, 1907
Centrinides Jekel, 1865: 548 [stem: Centrin-]. Type genus: Centrinus Schönherr, 1825. Comment: junior homonym of Centrininae Swainson, 1839 in Pisces (type genus Centrina Cuvier 1817); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Zygobari Pierce, 1907: 381 [stem: Zygobarid-]. Type genus: Zygobaris J. L. LeConte, 1876. Comment: incorrect original stem formation, not in prevailing usage; Alonso-Zarazaga and Lyal (1999: 97) and subsequent workers have used this name as valid instead of the junior homonym Centrinina Jekel, 1865, we continue to use Zygobaridina as valid here.
Limnobarini Casey, 1922: 269 [stem: Limnobarid-]. Type genus: Limnobaris Bedel, 1885. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Baridini
Schönherr, 1836
Baridides Schönherr, 1836: 636 [stem: Barid-]. Type genus: Baris Germar, 1817 [as Baridius, unjustified emendation of type genus name by Schönherr (1825), not in prevailing usage].
Subtribe
Baridina
Schönherr, 1836
Baridides Schönherr, 1836: 636 [stem: Barid-]. Type genus: Baris Germar, 1817 [as Baridius, unjustified emendation of type genus name by Schönherr (1825), not in prevailing usage].
Subtribe
Coelonertina
Casey, 1922
Coelonertini Casey, 1922: 94 [stem: Coelonert-]. Type genus: Coelonertus Solari and Solari, 1906.
Subtribe
Coleomerina
Casey, 1922
Coleomerini Casey, 1922: 84 [stem: Coleomer-]. Type genus: Coleomerus Schönherr, 1836.
Subtribe
Diorymerina
Jekel, 1865
Diorymerides Jekel, 1865: 548 [stem: Diorymer-]. Type genus: Diorymerus Schönherr, 1825.
Subtribe
Eurhinina
Lacordaire, 1865
Eurhinides Lacordaire, 1865: 220 [stem: Eurhin-]. Type genus: Eurhinus Illiger, 1807 [placed on the Official List of Generic Names in Zoology (ICZN 1985e)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pierce (1915: 15, as Eurhininae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 92, as Eurhinina); Eurhinini Lacordaire, 1865 placed on the Official List of Family-Group Names in Zoology (ICZN 1985e).
Tribe
Madarini
Jekel, 1865
Madarides Jekel, 1865: 550 [stem: Madar-]. Type genus: Madarus Schönherr, 1825. Comment: Madarini, published on 25 January 1865, has priority over Barymerini and Leptoschoinini both published by Lacordaire “before 12 December 1865”.
Subtribe
Barymerina
Lacordaire, 1865
Barymérides Lacordaire, 1865: 259 [stem: Barymer-]. Type genus: Barymerus Lacordaire, 1865. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1896: 127, as Barymerinorum), generally accepted as in Alonso-Zarazaga and Lyal (1999: 94, as Barymerina).
Sonnetiini Casey, 1922: 321 [stem: Sonneti-]. Type genus: Sonnetius Casey, 1922.
Subtribe
Eutoxina
Champion, 1908
Eutoxides Champion, 1908: 360 [stem: Eutox-]. Type genus: Eutoxus Schönherr, 1844.
Subtribe
Leptoschoinina
Lacordaire, 1865
Leptoschoïnides Lacordaire, 1865: 236 [stem: Leptoschoin-]. Type genus: Leptoschoinus Dejean, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1896: 126, as Leptoschoenorum [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 94, as Leptoschoinina).
Subtribe
Madarina
Jekel, 1865
Madarides Jekel, 1865: 550 [stem: Madar-]. Type genus: Madarus Schönherr, 1825.
Subtribe
Tonesiina
Alonso-Zarazaga and Lyal, 1999
Lytériides Lacordaire, 1865: 249 [stem: Lyteri-]. Type genus: Lyterius sensu Lacordaire, 1865 [not Lyterius Schönherr, 1844; syn. of Tonesia Casey, 1922]. Comment: based on a misidentified type genus, name treated here as invalid until an application is submitted to the Commission to suppress it for the Principle of Priority (Art. 65.2.1).
Tonesiina Alonso-Zarazaga and Lyal, 1999: 95 [stem: Tonesi-]. Type genus: Tonesia Casey, 1922. Comment: although this is not the oldest name for the subtribe, we recommend that an application be submitted to the Commission to suppress Lyteriini Lacordaire, 1865 because it is based on a misidentified type genus (Art. 65.2.1).
Tribe
Neosharpiini
Hoffmann, 1956
Neosharpini Hoffmann, 1956: 246 [stem: Neosharpi-]. Type genus: Neosharpia Hoffmann, 1956. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Nertinini
Voss, 1954
*Nertides Lacordaire, 1865: 238 [stem: Nert-]. Type genus: Nertus Schönherr, 1844 [preoccupied genus name, not Nertus Boie, 1828 [Aves]; syn. of Nertinus G. A. K. Marshall, 1943]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized and attributed to Lacordaire (1865); if found to be available then permanently invalid (Art. 39): based on preoccupied type genus.
Nertini Bondar, 1949: 223 [stem: Nert-]. Type genus: Nertus Schönherr, 1844 [preoccupied genus name, not Nertus Boie 1828 [Aves]; syn. of Nertinus G. A. K. Marshall, 1943]. Comment: family-group name proposed as new without reference to Nertides Lacordaire, 1865; permanently invalid (Art. 39): based on preoccupied type genus.
Nertinina Voss, 1954: 323 [stem: Nertin-]. Type genus: Nertinus G. A. K. Marshall, 1943.
Tribe
Optatini
Champion, 1907
Optatides Champion, 1907: 185 [stem: Optat-]. Type genus: Optatus Pascoe, 1889.
Tribe
Pantotelini
Lacordaire, 1865
Pantotélides Lacordaire, 1865: 212 [stem: Pantotel-]. Type genus: Pantoteles Schönherr, 1845.
Subtribe
Cyrionychina
Casey, 1922
Cyrionichini Casey, 1922: 10 [stem: Cyrionych-]. Type genus: Cyrionyx Faust, 1896. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Pantotelina
Lacordaire, 1865
Pantotélides Lacordaire, 1865: 212 [stem: Pantotel-]. Type genus: Pantoteles Schönherr, 1845. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Pantotelinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 102, as Pantotelini).
Tribe
Peridinetini
Lacordaire, 1865
Péridinétides Lacordaire, 1865: 209 [stem: Peridinet-]. Type genus: Peridinetus Schönherr, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1886: cliii, as Peridinetinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 102, as Peridinetini).
Subfamily
Ceutorhynchinae
Gistel, 1848
Centorhynchidae Gistel, 1848: [7] [stem: Ceutorhynch-]. Type genus: Ceutorhynchus Germar, 1824 [as Centorhynchus, incorrect subsequent spelling of type genus name, not in prevailing usage; placed on the Official List of Generic Names in Zoology (ICZN 1989c)]. Comment: incorrect original stem formation, not in prevailing usage; First Revisers found (Ceutorhynchinae Gistel, 1848 vs Phytobiinae Gistel, 1848) are Alonso-Zarazaga and Lyal (2002).
Tribe
Ceutorhynchini
Gistel, 1848
Centorhynchidae Gistel, 1848: [7] [stem: Ceutorhynch-]. Type genus: Ceutorhynchus Germar, 1824 [as Centorhynchus, incorrect subsequent spelling of type genus name, not in prevailing usage; placed on the Official List of Generic Names in Zoology (ICZN 1989c)]. Comment: incorrect original stem formation, not in prevailing usage.
Isorhynchides Lacordaire, 1865: 172 [stem: Isorhynch-]. Type genus: Isorhynchus Schönherr, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Isorhynchinae), generally accepted as in Hustache (1934: 72, as Isorhynchini).
Coeliodes J. L. LeConte, 1876: 268 [stem: Coeliod-]. Type genus: Coeliodes Schönherr, 1837 [placed on the Official List of Generic Names in Zoology (ICZN 1989c)].
Poophagidae Schultze, 1902: 226 [stem: Poophag-]. Type genus: Poophagus Schönherr, 1837.
Amalina Wagner, 1936: 166 [stem: Amal-]. Type genus: Amalus Schönherr, 1825.
Phrydiuchina Wagner, 1938: 163 [stem: Phrydiuch-]. Type genus: Phrydiuchus Gozis, 1885.
Oxyonyxina Hoffmann, 1957b: 220 [stem: Oxyonych-]. Type genus: Oxyonyx Faust, 1885. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Cnemogonini
Colonnelli, 1979
Cnemogonini Colonnelli, 1979: 2 [stem: Cnemogon-]. Type genus: Cnemogonus J. L. LeConte, 1876.
Tribe
Egriini
Pajni and Kohli, 1982
Egriini Pajni and Kohli, 1982: 339, in key [stem: Egri-]. Type genus: Egrius Pascoe, 1865.
Egriini Colonnelli, 1984: 209 [stem: Egri-]. Type genus: Egrius Pascoe, 1865. Comment: family-group name proposed as new without reference to Egriini Pajni and Kohli, 1982.
Tribe
Hypohypurini
Colonnelli, 2004
Hypohypurini Colonnelli, 2004: 10 [stem: Hypohypur-]. Type genus: Hypohypurus Hustache, 1920.
Tribe
Hypurini
Schultze, 1902
Hypuridae Schultze, 1902: 209 [stem: Hypur-]. Type genus: Hypurus Rey, 1882.
Tribe
Lioxyonychini
Colonnelli, 1984
Lioxyonyxini Colonnelli, 1984: 209 [stem: Lioxyonych-]. Type genus: Lioxyonyx Hustache, 1933. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Mecysmoderini
Wagner, 1938
Mecysmoderina Wagner, 1938: 170 [stem: Mecysmoder-]. Type genus: Mecysmoderes Schönherr, 1837.
Tribe
Mononychini
LeConte, 1876
Mononychi J. L. LeConte, 1876: 267 [stem: Mononych-]. Type genus: Mononychus Germar, 1824 [placed on the Official List of Generic Names in Zoology (ICZN 1989c)].
Tribe
Phytobiini
Gistel, 1848
Phytobiidae Gistel, 1848: [7] [stem: Phytobi-]. Type genus: Phytobius Schönherr, 1833 [placed on the Official List of Generic Names in Zoology (ICZN 2001)].
Rhinoncides C. G. Thomson, 1865: 231 [stem: Rhinonc-]. Type genus: Rhinoncus Schönherr, 1825 [placed on the Official List of Generic Names in Zoology (ICZN 1989c)].
Tribe
Scleropterini
Schultze, 1902
Scleropteridae Schultze, 1902: 209 [stem: Scleropter-]. Type genus: Scleropterus Schönherr, 1825. Comment: the family-group name Scléropterides was used earlier by Solier (1834) in the Heteromera, however, Solier’s name was not based on a genus name and is therefore unavailable; Agassiz (1846a: 146) incorrectly mentions Scleropterus as the type genus of Solier’s name but this is impossible because the family-group and genus-group names belonged to different orders; Agassiz (1846b: 335) also latinized Solier’s name as Scleropteroidae.
Subfamily
Conoderinae
Schönherr, 1833
Conoderides Schönherr, 1833: 26 [stem: Conoder-]. Type genus: Conoderes Schönherr, 1833.
Tribe
Arachnopodini
Lacordaire, 1865
Arachnopides Lacordaire, 1865: 159 [stem: Arachnopod-]. Type genus: Arachnopus Guérin-Méneville, 1838 [syn. of Arachnobas Boisduval, 1835]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 232, as Arachnopini [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 109, as Arachnopodini); incorrect original stem formation, not in prevailing usage.
Tribe
Campyloscelini
Schönherr, 1845
Campyloscelides Schönherr, 1845: 197 [stem: Campyloscel-]. Type genus: Campyloscelus Schönherr, 1845.
Subtribe
Campyloscelina
Schönherr, 1845
Campyloscelides Schönherr, 1845: 197 [stem: Campyloscel-]. Type genus: Campyloscelus Schönherr, 1845.
Subtribe
Corynemerina
Hustache, 1929
Corynemerinae Hustache, 1929: 531 [stem: Corynemer-]. Type genus: Corynemerus Fåhraeus, 1871.
Subtribe
Phaenomerina
Faust, 1898
Phaenomerina Faust, 1898: 76, in key [stem: Phaenomer-]. Type genus: Phaenomerus Schönherr, 1836 [placed on the Official List of Generic Names in Zoology (ICZN 1962)]. Comment: the stem of the senior homonym based on the Scarabaeidae genus Phaenomeris was emended to Phaeromerid- to remove the homonymy (ICZN 1962); available for family-group taxon below subfamily level and placed on the Official List of Family-Group Names in Zoology (ICZN 1962).
Tribe
Conoderini
Schönherr, 1833
Conoderides Schönherr, 1833: 26 [stem: Conoder-]. Type genus: Conoderes Schönherr, 1833. Comment: the name Conoderinae Fleutiaux, 1919 (type genus Conoderus Eschscholtz, 1829) is available in Elateridae; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Conophorides Schönherr, 1837: 719 [stem: Conophor-]. Type genus: Conophorus Schönherr, 1837 [preoccupied genus name, not Conophorus Meigen, 1803 [Diptera]; syn. of Conoderes Schönherr, 1833]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Coryssomerini
Thomson, 1859
Coryssomerina C. G. Thomson, 1859: 137 [stem: Coryssomer-]. Type genus: Coryssomerus Schönherr, 1825.
Synophthalmides Lacordaire, 1863: 514 [stem: Synophthalm-]. Type genus: Synophthalmus Lacordaire, 1863. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hustache (1932: 373, as Synophthalmini), generally accepted as in Hustache (1934: 49, as Synophthalmini).
Metialmini Hustache, 1932: 374, in key [stem: Metialm-]. Type genus: Metialma Pascoe, 1871.
Tribe
Coryssopodini
Lacordaire, 1865
Coryssopides Lacordaire, 1865: 163 [stem: Coryssopod-]. Type genus: Coryssopus Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 232, as Coryssopini [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 110, as Coryssopodini); incorrect original stem formation, not in prevailing usage.
*Sympiézopides Lacordaire, 1865: 166 [stem: Sympiezopod-]. Type genus: Sympiezopus Schönherr, 1837. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g. Faust (1886b: 367, as Sympiezopinorum [incorrect stem formation]), but not generally attributed to Lacordaire (1865); incorrect original stem formation, not in prevailing usage.
Tribe
Lechriopini
Lacordaire, 1865
Léchriopides Lacordaire, 1865: 149 [stem: Lechriop-]. Type genus: Lechriops Schönherr, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 232, as Lechriopini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 111, as Lechriopini).
Copturidae Desbrochers des Loges, 1892: 68 [stem: Coptur-]. Type genus: Copturus Schönherr, 1825.
Tribe
Lobotrachelini
Lacordaire, 1865
Lobotrachélides Lacordaire, 1865: 172 [stem: Lobotrachel-]. Type genus: Lobotrachelus Schönherr, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Aurivillius (1910: 433, as Lobotrachelinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 111, as Lobotrachelini).
Tribe
Mecopini
Lacordaire, 1865
Mécopides Lacordaire, 1865: 156 [stem: Mecop-]. Type genus: Mecopus Schönherr, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 232, as Mecopini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 111, as Mecopini); current spelling maintained (Art. 29.5): incorrect original stem formation in prevailing usage (should be Mecopod-); if the stem is corrected, it would threaten stability of a widely used name in Orthoptera (see Alonso-Zarazaga and Lyal 1999: 111).
Tribe
Menemachini
Lacordaire, 1865
Ménémachides Lacordaire, 1865: 27 [stem: Menemach-]. Type genus: Menemachus Schönherr, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Mnemachinae [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 112, as Menemachini).
Tribe
Othippiini
Morimoto, 1962
Othippiini Morimoto, 1962a: 47, in key [stem: Othippi-]. Type genus: Othippia Pascoe, 1874.
Tribe
Peloropodini
Hustache, 1932
Peleropini Hustache, 1932: 372, in key [stem: Peloropod-]. Type genus: Peloropus Schönherr, 1836. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Piazurini
Lacordaire, 1865
Piazurides Lacordaire, 1865: 144 [stem: Piazur-]. Type genus: Piazurus Schönherr, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1899: 100, as Piazurinorum), generally accepted as in Alonso-Zarazaga and Lyal (1999: 113, as Piazurini).
Tribe
Sphadasmini
Lacordaire, 1865
Sphadasmides Lacordaire, 1865: 161 [stem: Sphadasm-]. Type genus: Sphadasmus Schönherr, 1844. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1899: 102, as Sphadasminorum), generally accepted as in Alonso-Zarazaga and Lyal (1999: 114, as Sphadasmini).
Tribe
Trichodocerini
Champion, 1906
Trichodocerides Champion, 1906: 713 [stem: Trichodocer-]. Type genus: Trichodocerus Chevrolat, 1879.
Tribe
Zygopini
Lacordaire, 1865
Zygopides Lacordaire, 1865: 142 [stem: Zygop-]. Type genus: Zygops Schönherr, 1825 [placed on the Official List of Generic Names in Zoology (ICZN 1987e)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Zygopinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 114, as Zygopini).
Eccoptinae Pierce, 1919: 30, in key [stem: Eccopt-]. Type genus: Eccoptus Dejean, 1821 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1987e); syn. of Zygops Schönherr, 1825]. Comment: permanently invalid (Art. 39): based on suppressed type genus.
Cylindrocopturini Böving, 1927b: 156, in key [stem: Cylindrocoptur-]. Type genus: Cylindrocopturus Heller, 1895.
Subfamily
Cossoninae
Schönherr, 1825
Cossonides Schönherr, 1825: column 587 [stem: Cosson-]. Type genus: Cossonus Clairville, 1798.
Tribe
Acamptini
LeConte, 1876
Acampti J. L. LeConte, 1876: 238 [stem: Acampt-]. Type genus: Acamptus J. L. LeConte, 1876.
Tribe
Acanthinomerini
Voss, 1972
Acanthinomerini Voss, 1972: 382 [stem: Acanthinomer-]. Type genus: Acanthinomerus Boheman, 1859.
Tribe
Allomorphini
Folwaczny, 1973
Allomorphini Folwaczny, 1973: 68, in key [stem: Allomorph-]. Type genus: Allomorphus Folwaczny, 1968.
Tribe
Aphyllurini
Voss, 1955
Aphyllurina Voss, 1955a: 276 [stem: Aphyllur-]. Type genus: Aphyllura Reitter, 1884.
Tribe
Araucariini
Kuschel, 1966
Araucariini Kuschel, 1966: 4 [stem: Araucari-]. Type genus: Araucarius Kuschel, 1966.
Tribe
Choerorhinini
Folwaczny, 1973
Choerorhini Folwaczny, 1973: 69, in key [stem: Choerorhin-]. Type genus: Choerorhinus Fairmaire, 1858. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Cossonini
Schönherr, 1825
Cossonides Schönherr, 1825: column 587 [stem: Cosson-]. Type genus: Cossonus Clairville, 1798.
Tribe
Cryptommatini
Voss, 1972
Cryptommatini Voss, 1972: 373 [stem: Cryptommat-]. Type genus: Cryptommata Wollaston, 1877.
Tribe
Dryotribini
LeConte, 1876
Dryotribi J. L. LeConte, 1876: 335 [stem: Dryotrib-]. Type genus: Dryotribus G. H. Horn, 1873.
Cotasterinen Faust, 1886a: 28 [stem: Cotaster-]. Type genus: Cotaster Motschulsky, 1851. Comment: also as Cotasteriden on p. 31; original vernacular name available (Art. 11.7.2): first used in latinized form by Sainte-Claire Deville (1923: 63, as Cotastrini [incorrect stem formation]), generally accepted as in Hansen (1996: 209, as Cotasterini).
Tribe
Microxylobiini
Voss, 1972
Microxylobiini Voss, 1972: 431 [stem: Microxylobi-]. Type genus: Microxylobius Chevrolat, 1836.
Tribe
Nesiobiini
Alonso-Zarazaga and Lyal, 1999
Nesiotinae Voss, 1972: 338 [stem: Nesiot-]. Type genus: Nesiotes Wollaston, 1861 [preoccupied genus name, not Nesiotes Martens, 1860 [Mollusca]; syn. of Nesiobius G. A. K. Marshall, 1943]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Nesiobiini Alonso-Zarazaga and Lyal, 1999: 119 [stem: Nesiobi-]. Type genus: Nesiobius G. A. K. Marshall, 1943. Comment: replacement name for Nesiotinae Voss, 1872 because of the homonymy of the type genus.
Tribe
Neumatorini
Folwaczny, 1973
Neumatorini Folwaczny, 1973: 69, in key [stem: Neumator-]. Type genus: Neumatora Normand, 1920.
Tribe
Onychiini
Chapuis, 1869
Onychiidae Chapuis, 1869: 48 [stem: Onychi-]. Type genus: Onychius Chapuis, 1869. Comment: see Alonso-Zarazaga and Lyal (2009: 9) for comments on placement.
Tribe
Onycholipini
Wollaston, 1873
Onycholipides Wollaston, 1873: 454 [stem: Onycholip-]. Type genus: Onycholips Wollaston, 1861.
Stenoscelides Wollaston, 1877: 82, in key [stem: Stenoscelid-]. Type genus: Stenoscelis Wollaston, 1861.
Stereocorynina Voss, 1955b: 232 [stem: Stereocoryn-]. Type genus: Stereocorynes Wollaston, 1873.
Stereocorynini Roudier, 1958: 212 [stem: Stereocoryn-]. Type genus: Stereocorynes Wollaston, 1873. Comment: family-group name proposed as new without reference to Stereocorynina Voss, 1955.
Tribe
Pentarthrini
Lacordaire, 1865
Pentarthrides Lacordaire, 1865: 323 [stem: Pentarthr-]. Type genus: Pentarthrum Wollaston, 1854. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 234, as Pentarthrini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 120, as Pentarthrini).
Tribe
Proecini
Voss, 1956
Proecina Voss, 1956a: 139 [stem: Proec-]. Type genus: Proeces Schönherr, 1837.
Phylloplatypodini Kato, 1998: 77 [stem: Phylloplatypod-]. Type genus: Phylloplatypus Kato, 1998.
Tribe
Pseudapotrepini
Champion, 1909
Pseudapotrepides Champion, 1909: 21 [stem: Pseudapotrep-]. Type genus: Pseudapotrepus Champion, 1909.
Tribe
Rhyncolini
Gistel, 1848
Rhynchalidae Gistel, 1848: [6] [stem: Rhyncol-]. Type genus: Rhyncolus Germar, 1817 [as Rhyncholus, incorrect subsequent spelling of type genus name, not in prevailing usage; Rhyncolus Germar, 1817 placed on the Official List of Generic Names in Zoology (ICZN 1991b)].
Subtribe
Phloeophagina
Voss, 1955
Phloeophagina Voss, 1955b: 228 [stem: Phloeophag-]. Type genus: Phloeophagus Schönherr, 1837.
Subtribe
Pseudomimina
Voss, 1939
Pseudomimini Voss, 1939: 65 [stem: Pseudomim-]. Type genus: Pseudomimus Hartmann, 1904.
Subtribe
Rhyncolina
Gistel, 1848
Rhynchalidae Gistel, 1848: [6] [stem: Rhyncol-]. Type genus: Rhyncolus Germar, 1817 [as Rhyncholus, incorrect subsequent spelling of type genus name, not in prevailing usage; Rhyncolus Germar, 1817 placed on the Official List of Generic Names in Zoology (ICZN 1991b)]. Comment: incorrect original stem formation, not in prevailing usage.
Eremotini Voss, 1939: 65 [stem: Eremot-]. Type genus: Eremotes Wollaston, 1861.
Cotasterosomini Konishi, 1962: 2 [stem: Cotasterosomat-]. Type genus: Cotasterosoma Konishi, 1962. Comment: incorrect original stem formation, not in prevailing usage.
Himatinini Konishi, 1962: 7 [stem: Himatin-]. Type genus: Himatinum Cockerell, 1906 [syn. of Himatium Wollaston, 1873].
Tribe
Tapiromimini
Voss, 1972
Tapiromimini Voss, 1972: 380 [stem: Tapiromim-]. Type genus: Tapiromimus Wollaston, 1877.
Subfamily
Cryptorhynchinae
Schönherr, 1825
Cryptorhynchides Schönherr, 1825: column 585 [stem: Cryptorhynch-]. Type genus: Cryptorhynchus Illiger, 1807 [placed on the Official List of Generic Names in Zoology (ICZN 1967b)]. Comment: Cryptorhynchinae Schönherr, 1825 placed on the Official List of Family-Group Names in Zoology (ICZN 1967b).
Tribe
Aedemonini
Faust, 1898
Acdemonini Faust, 1898: 34 [stem: Aedemon-]. Type genus: Aedemonus Schönherr, 1837 [as Acdemonus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Mecistocerini Morimoto, 1978b: 122 [stem: Mechistocer-]. Type genus: Mechistocerus Fauvel, 1863 [as Mecistocerus, unjustified emendation of type genus name by Pascoe (1870b), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Morimoto (1987).
Derbyiellina Zimmerman, 1994a: 645, in key [stem: Derbyiell-]. Type genus: Derbyiella Lea, 1907.
Tribe
Camptorhinini
Lacordaire, 1865
Camptorhinides Lacordaire, 1865: 86 [stem: Camptorhin-]. Type genus: Camptorhinus Schönherr, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1885: 254, as Camptorhininae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 136, as Camptorhinini).
Tribe
Cryptorhynchini
Schönherr, 1825
Cryptorhynchides Schönherr, 1825: column 585 [stem: Cryptorhynch-]. Type genus: Cryptorhynchus Illiger, 1807 [placed on the Official List of Generic Names in Zoology (ICZN 1967b)]. Comment: Cryptorhynchinae Schönherr, 1825 placed on the Official List of Family-Group Names in Zoology (ICZN 1967b).
Subtribe
Cryptorhynchina
Schönherr, 1825
Cryptorhynchides Schönherr, 1825: column 585 [stem: Cryptorhynch-]. Type genus: Cryptorhynchus Illiger, 1807 [placed on the Official List of Generic Names in Zoology (ICZN 1967b)]. Comment: Cryptorhynchinae Schönherr, 1825 placed on the Official List of Family-Group Names in Zoology (ICZN 1967b).
Cryptorhynchidiini Bradley, 1930: 277, in key [stem: Cryptorhynchidi-]. Type genus: Cryptorhynchidius Pierce, 1919 [syn. of Cryptorhynchus Illiger, 1807].
Subtribe
Mecistostylina
Lacordaire, 1865
Mécistostylides Lacordaire, 1865: 131 [stem: Mecistostyl-]. Type genus: Mecistostylus Lacordaire, 1865. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by G. A. K. Marshall (1916: 10, as Mecistostylini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 129, as Mecistostylina).
Cylindrocosyninae Pascoe, 1875: 55 [stem: Cylindrocoryn-]. Type genus: Cylindrocorynus Schönherr, 1837. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Tylodina
Lacordaire, 1865
Tylodides Lacordaire, 1865: 90 [stem: Tylod-]. Type genus: Tylodes Sahlberg, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1885: 255, as Tylodinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 129, as Tylodina).
Tribe
Gasterocercini
Zherikhin, 1991
Gasterocercini Zherikhin, 1991: 92 [stem: Gasterocerc-]. Type genus: Gasterocercus Laporte and Brullé, 1828.
Tribe
Psepholacini
Lacordaire, 1865
Psépholacides Lacordaire, 1865: 72 [stem: Psepholac-]. Type genus: Psepholax Lacordaire, 1865. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hustache (1936: 56, as Psepholacina), generally accepted as in Alonso-Zarazaga and Lyal (1999: 136, as Psepholacini); First Reviser (Psepholacini Lacordaire, 1865 vs Strongylopterini Lacordaire, 1865 vs Sympiezoscelini Lacordaire, 1865) not determined, current usage maintained.
Strongyloptérides Lacordaire, 1865: 73 [stem: Strongylopter-]. Type genus: Strongylopterus Schönherr, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Hustache (1936: 60, as Strongylopteridina [incorrect stem formation]).
Sympiézoscélides Lacordaire, 1865: 138 [stem: Sympiezoscel-]. Type genus: Sympiezoscelus G. R. Waterhouse, 1853. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Pascoe (1874: 96, as Sympiezoscelides [treated as Latin]).
Tribe
Sophrorhinini
Lacordaire, 1865
Sophrorhinides Lacordaire, 1865: 81 [stem: Sophrorhin-]. Type genus: Sophrorhinus Rouzet, 1855. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by C. O. Waterhouse (1879a: 310, as Sophororhinae [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 137, as Sophororhinini).
Tribe
Torneumatini
Bedel, 1884
Torneumatini Bedel, 1884b: 136, in note [stem: Torneumat-]. Type genus: Torneuma Wollaston, 1860.
Subfamily
Cyclominae
Schönherr, 1826
Cyclomides Schönherr, 1826: 185 [stem: Cyclom-]. Type genus: Cyclomus Schönherr, 1826 [syn. of Epicthonius Schönherr, 1826]. Comment: the classification of this subfamily follows Oberprieler (2010).
Tribe
Amycterini
Waterhouse, 1854
Amycteridae G. R. Waterhouse, 1854: 75, in note [stem: Amycter-]. Type genus: Amycterus Schönherr, 1823.
Euomides Lacordaire, 1863: 315 [stem: Euom-]. Type genus: Euomus Schönherr, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by W. J. MacLeay (1866: 319, as Euomidae), generally accepted as in Schenkling and G. A. K. Marshall (1931a: 27, as Euomini).
Psaliduridae Pierce, 1914: 350 [stem: Phalidur-]. Type genus: Phalidura Fischer von Waldheim, 1823 [as Psalidura, incorrect subsequent spelling of type genus name, not in prevailing usage; syn. of Amycterus Schönherr, 1823]. Comment: incorrect original stem formation, not in prevailing usage.
*Acantholophini Schenkling and G. A. K. Marshall, 1931a: Amyct. 6 [stem: Acantholoph-]. Type genus: Acantholophus Boisduval, 1835. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Aterpini
Lacordaire, 1863
Aterpides Lacordaire, 1863: 410 [stem: Aterp-]. Type genus: Aterpus Schönherr, 1826 [syn. of Aades Schönherr, 1823]. Comment: precedence (Aterpini Lacordaire, 1863 vs Rhadinosomini Lacordaire, 1863) given to taxon originally proposed at the higher rank (Art. 24.1).
Subtribe
Aterpina
Lacordaire, 1863
nomen protectum
Heliomeneidae Gistel, 1848: [8] [stem: Heliomen-]. Type genus: Heliomene Gistel, 1848 [syn. of Chrysolopus Germar, 1817]. Comment: as pointed out by Alonso-Zarazaga and Lyal (2002: 22) this is the oldest available name for the subtribe name; Heliomeneini Gistel, 1848 was recently considered a nomen oblitum by Colonnelli (2003: 7) however the necessary supporting references were not provided; we hereby consider Heliomeneini Gistel, 1848 as a nomen oblitum (see supporting references in Appendix 1); incorrect original stem formation, not in prevailing usage.
Aterpides Lacordaire, 1863: 410 [stem: Aterp-]. Type genus: Aterpus Schönherr, 1826 [syn. of Aades Schönherr, 1823]. Comment: nomen protectum (see Appendix 1); original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Aterpinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 140, as Aterpini).
*Pélororhinides Lacordaire, 1863: 415 [stem: Pelororhin-]. Type genus: Pelororhinus Schönherr, 1834. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 228, as Pelororhinini), Morrone (1997: 101, as Pelorhirhini [incorrect stem formation]), but not generally accepted as valid; Pelorhinidae [incorrect stem formation] was used as valid by Ienistea (1986: 33) but it was not attributed to Lacordaire (1863); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Lophotides Jekel, 1865: 546 [stem: Lophot-]. Type genus: Lophotus Schönherr, 1834 [preoccupied genus name, not Lophotus Giorna, 1809 [Pisces]; syn. of Aegorhinus Erichson, 1834]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Rhinarides Jekel, 1865: 546 [stem: Rhinari-]. Type genus: Rhinaria Kirby, 1819. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Rhadinosomina
Lacordaire, 1863
Rhadinosomides Lacordaire, 1863: 63 [stem: Rhadinosom-]. Type genus: Rhadinosomus Schönherr, 1840. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1875: 55, as Rhadinosominae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 141, as Rhadinosomina).
Rhadinosomini Pierce, 1913: 405, in key [stem: Rhadinosom-]. Type genus: Rhadinosomus Schönherr, 1840. Comment: family-group name proposed as new without reference to Rhadinosomides Lacordaire, 1863.
Tribe
Cyclomini
Schönherr, 1826
Cyclomides Schönherr, 1826: 185 [stem: Cyclom-]. Type genus: Cyclomus Schönherr, 1826 [syn. of Epicthonius Schönherr, 1826].
Somatodides Lacordaire, 1863: 319 [stem: Somatod-]. Type genus: Somatodes Schönherr, 1840 [placed on the Official List of Generic Names in Zoology (ICZN 1994c)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Somatodinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 144, as Somatodini); Somatodinae Lacordaire, 1863 placed on the Official List of Family-Group Names in Zoology (ICZN 1994c).
Tribe
Dichotrachelini
Hoffmann, 1957
Dichotrachelini Hoffmann, 1957a: 60, in note [stem: Dichotrachel-]. Type genus: Dichotrachelus Stierlin, 1853.
Tribe
Hipporhinini
Lacordaire, 1863
Hipporhinides Lacordaire, 1863: 323 [stem: Hipporhin-]. Type genus: Hipporhinus Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Pascoe (1870b: 436, as Hipporhininae).
Gronopini Bedel, 1884a: 68, in key [stem: Gronop-]. Type genus: Gronops Schönherr, 1823.
Gronopina Louw and Oberprieler, 1998: 24 [stem: Gronop-]. Type genus: Gronops Schönherr, 1823. Comment: family-group name proposed as new without reference to Gronopini Bedel, 1884.
Tribe
Listroderini
LeConte, 1876
Listroderi J. L. LeConte, 1876: 124 [stem: Listroder-]. Type genus: Listroderes Schönherr, 1826.
Palaechtini C. Brinck, 1948: 43 [stem: Palaechth-]. Type genus: Palaechthus C. O. Waterhouse, 1884 [as Palaechtus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Notiomimetini
Wollaston, 1873
Notiomimetides Wollaston, 1873: 440 [stem: Notiomimet-]. Type genus: Notiomimetes Wollaston, 1873.
Apheli Csiki, 1936: 109 [stem: Aphel-]. Type genus: Aphela Pascoe, 1865. Comment: description by indication (distinguishing characters given in Wollaston (1873: 440)).
Tribe
Rhythirrinini
Lacordaire, 1863
Rhytirhinides Lacordaire, 1863: 296 [stem: Rhythirrin-]. Type genus: Rhythirrinus Schönherr, 1823 [as Rhytirhinus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1875 [Gatt.]: 112, as Rhytirhinini [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 141, as Rhythirrinini); incorrect original stem formation, not in prevailing usage; First Reviser found (Rhythirrinini Lacordaire, 1863 vs Rhyparosomini Lacordaire, 1863 vs Eupagini Lacordaire, 1863) is Kuschel (1971: 251).
Rhyparosomides Lacordaire, 1863: 327 [stem: Rhyparosom-]. Type genus: Rhyparosomus Schönherr, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Stein (1868: 100, as Rhyparosomini), generally accepted as in Heyne and Taschenberg (1907: 228, as Rhyparosomini).
Eupagides Lacordaire, 1863: 328 [stem: Eupag-]. Type genus: Eupages Schönherr, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Péringuey (1888: 168, as Eupagidae), generally accepted as in Schenkling and G. A. K. Marshall (1929: 47, as Eupagini).
Rhytidorhinides Wollaston, 1865: 307 [stem: Rhytidorhin-]. Type genus: Rhytidorhinus Wollaston, 1864 [preoccupied genus name, not Rhytidorhinus Agassiz, 1846 [Coleoptera: Curculionidae: Cyclominae]; syn. of Rhythirrinus Schönherr, 1823]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Subfamily
Entiminae
Schönherr, 1823
Entimides Schönherr, 1823: column 1138 [stem: Entim-]. Type genus: Entimus Germar, 1817. Comment: First Revisers found (Entiminae Schönherr, 1823 vs Lordopinae Schönherr, 1823 vs Polydrusinae Schönherr, 1823) are Alonso-Zarazaga and Lyal (1999: 144).
Tribe
Agraphini
Horn, 1876
Agraphi G. H. Horn, 1876: 58 [stem: Agraph-]. Type genus: Agraphus Say, 1831.
Tribe
Alophini
LeConte, 1874
Alophini J. L. LeConte, 1874b: 461 [stem: Aloph-]. Type genus: Alophus Schönherr, 1826 [syn. of Graptus Schönherr, 1823].
Tribe
Anomophthalmini
Morrone, 1998
Anomophthalmina Morrone, 1998: 86 [stem: Anomophthalm-]. Type genus: Anomophthalmus Fairmaire, 1884.
Tribe
Anypotactini
Champion, 1911
Anypotactina Champion, 1911: 215 [stem: Anypotact-]. Type genus: Anypotactus Schönherr, 1840.
Tribe
Blosyrini
Lacordaire, 1863
Blosyrides Lacordaire, 1863: 27 [stem: Blosyr-]. Type genus: Blosyrus Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Péringuey (1888: 144, as Blosyridae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 146, as Blosyrini).
Tribe
Brachyderini
Schönherr, 1826
Brachyderides Schönherr, 1826: 94 [stem: Brachyder-]. Type genus: Brachyderes Schönherr, 1823 [placed on the Official List of Generic Names in Zoology (ICZN 1987a)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1987a, as Brachyderinae Schönherr, 1837).
Thylacitidae Kirby, 1837: 202 [stem: Thylacit-]. Type genus: Thylacites Germar, 1817 [placed on the Official Index of Invalid and Rejected Generic Names in Zoology (ICZN 1987a)]. Comment: placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1987a).
Strophosomidae Gistel, 1848: [8] [stem: Strophosom-]. Type genus: Strophosomum Gistel, 1856 [syn. of Strophosoma Billberg, 1820].
Tribe
Celeuthetini
Lacordaire, 1863
Céleuthétides Lacordaire, 1863: 145 [stem: Celeuthet-]. Type genus: Celeuthetes Schönherr, 1842.
Subtribe
Celeuthetina
Lacordaire, 1863
Céleuthétides Lacordaire, 1863: 145 [stem: Celeuthet-]. Type genus: Celeuthetes Schönherr, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1891: 166, as Celeuthetinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 147, as Celeuthetini).
Coptorrhynchina Voss, 1940: 279 [stem: Coptorhynch-]. Type genus: Coptorhynchus sensu Faust, 1897 [not Coptorhynchus Guérin-Méneville, 1841; syn. of Resites Alonso-Zarazaga and Lyal, 1999]. Comment: incorrect original stem formation, not in prevailing usage; based on a misidentified type genus.
Subtribe
Isopterina
Morimoto and Kojima, 2001
Isopterina Morimoto and Kojima, 2001: 274 [stem: Isopter-]. Type genus: Isopterus Faust, 1895.
Tribe
Cneorhinini
Lacordaire, 1863
Cnéorhinides Lacordaire, 1863: 31 [stem: Cneorhin-]. Type genus: Cneorhinus Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Seidlitz (1875 [Gatt.]: 111, as Cneorhinini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 148, as Cneorhinini).
Philopedini Bedel, 1883: 32, in key [stem: Philoped-]. Type genus: Philopedon Schönherr, 1826.
Dermatodini Emden, 1936: 76, in key [stem: Dermatod-]. Type genus: Dermatodes Schönherr, 1840.
Stigmatrachelini Richard, 1983: 8 [stem: Stigmatrachel-]. Type genus: Stigmatrachelus Schönherr, 1840.
Tribe
Cratopodini
Hustache, 1919
Cratopini Hustache, 1919: 473 [stem: Cratopod-]. Type genus: Cratopus Schönherr, 1823. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Cylydrorhinini
Lacordaire, 1863
Cylindrorhinides Lacordaire, 1863: 339 [stem: Cylydrorhin-]. Type genus: Cylydrorhinus Guérin-Méneville, 1838 [as Cylindrorhinus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Cylindrorhininae [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 150, as Cylydrorhinini); incorrect original stem formation, not in prevailing usage.
Tribe
Cyphicerini
Lacordaire, 1863
Cyphicérides Lacordaire, 1863: 220 [stem: Cyphicer-]. Type genus: Cyphicerus Schönherr, 1823. Comment: First Reviser (Cyphicerini Lacordaire, 1863 vs Phytoscaphini Lacordaire, 1863) not determined, current usage maintained.
Subtribe
Acanthotrachelina
Marshall, 1944
Acanthotrachelini G. A. K. Marshall, 1944: 76, in key [stem: Acanthotrachel-]. Type genus: Acanthotrachelus Schönherr, 1842.
Subtribe
Cyphicerina
Lacordaire, 1863
Cyphicérides Lacordaire, 1863: 220 [stem: Cyphicer-]. Type genus: Cyphicerus Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1888: 284, as Cyphicerinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 151, as Cyphicerini).
Corigetini Faust, 1885: 167, in key [stem: Coriget-]. Type genus: Corigetus Desbrochers des Loges, 1872.
Subtribe
Mylacorrhinina
Reitter, 1913
Mylacorrhynchina Reitter, 1913b: 12, in key [stem: Mylacorrhin-]. Type genus: Mylacorrhina Reitter, 1913 [syn. of Altonomus Desbrochers des Loges, 1907]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Myllocerina
Pierce, 1913
Myllocerini Pierce, 1913: 421, in key [stem: Myllocer-]. Type genus: Myllocerus Schönherr, 1826.
Ptochini Reitter, 1913b: 8, in key [stem: Ptoch-]. Type genus: Ptochus Schönherr, 1826 [placed on the Official List of Generic Names in Zoology (ICZN 1990d)].
Subtribe
Phytoscaphina
Lacordaire, 1863
Phytoscaphides Lacordaire, 1863: 229 [stem: Phytoscaph-]. Type genus: Phytoscaphus Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1869: 382, as Phytoscaphi), generally accepted as in Alonso-Zarazaga and Lyal (1999: 154, as Phytoscaphina).
*Oxyophthalmina Voss, 1933a: 30 [stem: Oxyophthalm-]. Type genus: Oxyophthalmus Hochhuth, 1847. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Tribe
Ectemnorhinini
Lacordaire, 1863
Ectemnorhinides Lacordaire, 1863: 562 [stem: Ectemnorhin-]. Type genus: Ectemnorhinus G. R. Waterhouse, 1853. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Ectemnorhinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 155, as Ectemnorhinini).
Canonopsini Dreux and Voisin, 1989: 112 [stem: Canonopse-]. Type genus: Canonopsis C. O. Waterhouse, 1875. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Elytrurini
Marshall, 1956
Elytrurini G. A. K. Marshall, 1956: 5 [stem: Elytrur-]. Type genus: Elytrurus Boisduval, 1835.
Tribe
Embrithini
Marshall, 1942
Embrithini G. A. K. Marshall, 1942: 3 [stem: Embrith-]. Type genus: Embrithes Schönherr, 1842.
Tribe
Entimini
Schönherr, 1823
Entimides Schönherr, 1823: column 1138 [stem: Entim-]. Type genus: Entimus Germar, 1817.
Tribe
Episomini
Lacordaire, 1863
Episomides Lacordaire, 1863: 175 [stem: Episom-]. Type genus: Episomus Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1894: 185, as Episomini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 156, as Episomini).
Episomini Pierce, 1913: 421, in key [stem: Episom-]. Type genus: Episomus Schönherr, 1823. Comment: family-group name proposed as new without reference to Episomides Lacordaire, 1863.
Tribe
Eudiagogini
LeConte, 1874
Promécopides Lacordaire, 1863: 384 [stem: Promecop-]. Type genus: Promecops Schönherr, 1823 [preoccupied genus name, not Promecops Sahlberg, 1823 [Coleoptera: Curculionidae: Entiminae: Eudiagogini]; syn. of Promecops Sahlberg, 1823]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Pascoe (1870b: 436, as Promecopinae); permanently invalid (Art. 39): based on preoccupied type genus.
Eudiagogini J. L. LeConte, 1874b: 454 [stem: Eudiagog-]. Type genus: Eudiagogus Schönherr, 1840. Comment: First Reviser (Eudiagogini J. L. LeConte, 1874 vs Bathyrini J. L. LeConte, 1874) not determined, current usage maintained.
Bathyrini J. L. LeConte, 1874b: 461 [stem: Bathyrin-]. Type genus: Bathyris J. L. LeConte, 1874 [syn. of Colecerus Schönherr, 1840]. Comment: incorrect original stem formation, not in prevailing usage.
Coleocerini Jekel, 1875: 144 [stem: Colecer-]. Type genus: Colecerus Schönherr, 1840 [as Coleocerus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Eupholini
Günther, 1943
*Eupholini Schenkling and G. A. K. Marshall, 1931b: Lept. 62 [stem: Euphol-]. Type genus: Eupholus Guérin-Méneville, 1838 [preoccupied genus name, not Eupholus Boisduval, 1835 [Coleoptera: Curculionidae: Entiminae]; syn. of Eupholus Boisduval, 1835]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).
Eupholini Günther, 1943: 19 [stem: Euphol-]. Type genus: Eupholus Guérin-Méneville, 1838 [preoccupied genus name, not Eupholus Boisduval, 1835 [Coleoptera: Curculionidae: Entiminae]; syn. of Eupholus Boisduval, 1835]. Comment: although this name should be treated as permanently invalid because it is based on a preoccupied type genus (Art. 39), an application will be submitted to the Commission by MAAZ and CHCL to conserve Eupholini Günther, 1943 and designate Eupholus Boisduval, 1835 as its type genus.
Eupholini Alonso-Zarazaga and Lyal, 1999: 157 [stem: Euphol-]. Type genus: Eupholus Boisduval, 1835. Comment: proposed as a new taxon; junior homonym of Eupholini Günther, 1943.
Tribe
Eustylini
Lacordaire, 1863
Eustylides Lacordaire, 1863: 205 [stem: Eustyl-]. Type genus: Eustylus Schönherr, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1886b: 338, as Eustylinorum), generally accepted as in Alonso-Zarazaga and Lyal (1999: 157, as Eustylini).
Exophthalmini G. H. Horn, 1876: 100 [stem: Exophthalm-]. Type genus: Exophthalmus Schönherr, 1823.
Eustylini Pierce, 1913: 421, in key [stem: Eustyl-]. Type genus: Eustylus Schönherr, 1842. Comment: family-group name proposed as new without reference to Eustylides Lacordaire, 1863.
Compsi Pierce, 1913: 406, in key [stem: Comps-]. Type genus: Compsus Schönherr, 1823. Comment: the younger name Compsina Martins and Galileo, 2007 (type genus Compsa Perty, 1832) in Cerambycidae is available; this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Exophthalmodina Voss, 1954: 199, in key [stem: Exophthalmod-]. Type genus: Exophthalmodes Pierce, 1916 [syn. of Exophthalmus Schönherr, 1823].
Tribe
Geonemini
Gistel, 1848
Geonemidae Gistel, 1848: [8] [stem: Geonem-]. Type genus: Geonemus Schönherr, 1833 [placed on the Official List of Generic Names in Zoology (ICZN 1988d)].
Barynotides Lacordaire, 1863: 37 [stem: Barynot-]. Type genus: Barynotus Germar, 1817. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by G. H. Horn (1876: 22, as Barynoti), generally accepted as in Dalla Torre et al. (1936: 32, as Barynotini).
Trigonoscutae J. L. LeConte, 1874b: 456 [stem: Trigonoscut-]. Type genus: Trigonoscuta Motschulsky, 1853.
Epicaeri G. H. Horn, 1876: 18 [stem: Epicaer-]. Type genus: Epicaerus Schönherr, 1834.
Calyptilli G. H. Horn, 1876: 26 [stem: Calyptill-]. Type genus: Calyptillus G. H. Horn, 1876.
Omilei G. H. Horn, 1876: 101 [stem: Omile-]. Type genus: Omileus G. H. Horn, 1876.
Trigonoscutini Pierce, 1913: 405, in key [stem: Trigonoscut-]. Type genus: Trigonoscuta Motschulsky, 1853. Comment: family-group name proposed as new without reference to Trigonoscutae J. L. LeConte, 1874.
Menoetiini Pierce, 1913: 373, in key [stem: Menoeti-]. Type genus: Menoetius Dejean, 1821 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1987f)]. Comment: permanently invalid (Art. 39): based on suppressed type genus.
Calyptillini Pierce, 1913: 405, in key [stem: Calyptill-]. Type genus: Calyptillus G. H. Horn, 1876. Comment: family-group name proposed as new without reference to Calyptilli G. H. Horn, 1876.
Tribe
Holcorhinini
Desbrochers des Loges, 1898
Holcorhinidae Desbrochers des Loges, 1898: 5 [stem: Holcorhin-]. Type genus: Holcorhinus Schönherr, 1826.
Cyclopterini Reitter, 1913b: 9, in key [stem: Cyclopter-]. Type genus: Cyclopterus Marseul, 1871 [preoccupied genus name, not Cyclopterus Linnaeus, 1758 [Pisces]; syn. of Nucterocephalus Desbrochers des Loges, 1897]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; Cyclopteridae Bonaparte, 1831 (type genus Cyclopterus Linnaeus, 1758) is available in Pisces.
Tribe
Hormorini
Horn, 1876
Hormori G. H. Horn, 1876: 23 [stem: Hormor-]. Type genus: Hormorus G. H. Horn, 1876.
Tribe
Laparocerini
Lacordaire, 1863
Laparocérides Lacordaire, 1863: 196 [stem: Laparocer-]. Type genus: Laparocerus Schönherr, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 226, as Laparocerini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 160, as Laparocerini).
Laparocerini Pierce, 1913: 421, in key [stem: Laparocer-]. Type genus: Laparocerus Schönherr, 1834. Comment: family-group name proposed as new without reference to Laparocérides Lacordaire, 1863.
Tribe
Leptostethini
Lacordaire, 1863
Leptostéthides Lacordaire, 1863: 258 [stem: Leptosteth-]. Type genus: Leptostethus G. R. Waterhouse, 1853. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 226, as Leptostethini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 161, as Leptostethini).
Tribe
Lordopini
Schönherr, 1823
Lordopides Schönherr, 1823: column 1142 [stem: Lordop-]. Type genus: Lordops Schönherr, 1823.
Hypsonotidae Jekel, 1853: I.II.3 [stem: Hypsonot-]. Type genus: Hypsonotus Germar, 1824.
Alocorhini Jekel, 1856: 9bis, in key [stem: Alocorhin-]. Type genus: Alocorhinus Sahlberg, 1823.
Elytroxysi Jekel, 1856: 9bis, in key [stem: Elytroxe-]. Type genus: Elytroxys Jekel, 1856. Comment: incorrect original stem formation, not in prevailing usage.
Eurylobi Jekel, 1856: 9bis, in key [stem: Eurylob-]. Type genus: Eurylobus Schönherr, 1826.
Merodonti Jekel, 1856: 9bis, in key [stem: Merodont-]. Type genus: Merodontus Jekel, 1856.
Tomorhini Jekel, 1856: 9bis, in key [stem: Tomorhin-]. Type genus: Tomorhinus Jekel, 1856.
Tribe
Mesostylini
Reitter, 1913
Mesostylini Reitter, 1913b: 8, in key [stem: Mesostyl-]. Type genus: Mesostylus Faust, 1894.
Tribe
Myorhinini
Marseul, 1863
Myorhinidae Marseul, 1863: 223 [stem: Myorhin-]. Type genus: Myorhinus Schönherr, 1826 [syn. of Apsis Germar, 1820]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kiesenwetter (1864: 265, as Myorhinidae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 162, as Myorhinini).
Tribe
Nastini
Reitter, 1913
Nastini Reitter, 1913b: 9, in key [stem: Nast-]. Type genus: Nastus Schönherr, 1842.
Tribe
Naupactini
Gistel, 1848
nomen protectum
Iphiides Schönherr, 1823: column 1139 [stem: Iphi-]. Type genus: Iphius Schönherr, 1823 [syn. of Alceis Billberg, 1820]. Comment: nomen oblitum (see Appendix 1).
Naupactidae Gistel, 1848: [8] [stem: Naupact-]. Type genus: Naupactus Dejean, 1821. Comment: nomen protectum (see Appendix 1).
Cyphides Lacordaire, 1863: 107 [stem: Cyph-]. Type genus: Cyphus Germar, 1824 [syn. of Cyrtomon Schönherr, 1823]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1874b: 458, as Cyphi), generally accepted as in J. L. LeConte and G. H. Horn (1883: 453, as Cyphi).
Macrostyles J. L. LeConte, 1874b: 457 [stem: Macrostyl-]. Type genus: Macrostylus Boheman, 1840.
Symmathetes J. L. LeConte, 1874b: 458 [stem: Symmathet-]. Type genus: Symmathetes Schönherr, 1847 [syn. of Pantomorus Schönherr, 1840].
Artipi G. H. Horn, 1876: 91 [stem: Artipod-]. Type genus: Artipus Schönherr, 1823 [preoccupied genus name, not Artipus Sahlberg, 1823 [Coleoptera: Curculionidae: Entiminae: Naupactini]; syn. of Artipus Sahlberg, 1823]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Platyomina Champion, 1911: 282 [stem: Platyom-]. Type genus: Platyomus Schönherr, 1823 [preoccupied genus name, not Platyomus Sahlberg, 1823 [Coleoptera: Curculionidae: Entiminae: Naupactini]; syn. of Platyomus Sahlberg, 1823]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Alceidini Pierce, 1913: 404, in key [stem: Alceent-]. Type genus: Alceis Billberg, 1820. Comment: incorrect original stem formation, not in prevailing usage.
Glaphyrometopi Pierce, 1913: 406, in key [stem: Glaphyrometop-]. Type genus: Glaphyrometopus Pierce, 1913.
Pseudocyphi Pierce, 1913: 406, in key [stem: Pseudocyph-]. Type genus: Pseudocyphus Schaeffer, 1905 [syn. of Platyomus Sahlberg, 1823].
Neocyphini Hustache, 1919: 476 [stem: Neocyph-]. Type genus: Neocyphus Bedel, 1883 [syn. of Cyrtomon Schönherr, 1823].
Platyomina Voss, 1954: 199, in key [stem: Platyom-]. Type genus: Platyomus Sahlberg, 1823. Comment: family-group name proposed as new without reference to Platyomina Champion, 1911.
Canephorotomina Voss, 1954: 206, in key [stem: Canephorotom-]. Type genus: Canephorotomus Voss, 1954 [syn. of Amitrus Schönherr, 1840].
Pantomorina Voss, 1954: 207, in key [stem: Pantomor-]. Type genus: Pantomorus Schönherr, 1840.
Plectrophorina Voss, 1954: 207, in key [stem: Plectrophor-]. Type genus: Plectrophorus Schönherr, 1826 [preoccupied genus name, not Plectrophorus Férussac, 1819 [Mollusca]; syn. of Plectrophoroides Wibmer and O’Brien, 1986]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Nothognathini
Marshall, 1916
Nothognathides G. A. K. Marshall, 1916: 204, in key [stem: Nothognath-]. Type genus: Nothognathus G. A. K. Marshall, 1916.
Tribe
Omiini
Shuckard, 1839
Omiadae Shuckard, 1839b: 60 [stem: Omi-]. Type genus: Omias Germar, 1817. Comment: senior homonym of Omiini Beck, 1996 in Lepidoptera (type genus Omia Hübner, 1821); this case is to be referred to the Commission to remove the homonymy (Art. 55.3.1).
Mylacini Reitter, 1913b: 9, in key [stem: Mylac-]. Type genus: Mylacus Boheman, 1843 [syn. of Omias Germar, 1817].
Tribe
Oosomini
Lacordaire, 1863
*Oosomides Schönherr, 1823: column 1145 [stem: Oosom-]. Type genus: Oosomus Schönherr, 1826. Comment: family-group name unavailable (Art. 11.7.1.1): not based on an available genus name at the time.
Oosomides Lacordaire, 1863: 164 [stem: Oosom-]. Type genus: Oosomus Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kolbe (1891: 26, as Oosominae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 166, as Oosomini).
Oosomini Pierce, 1913: 421, in key [stem: Oosom-]. Type genus: Oosomus Schönherr, 1826. Comment: family-group name proposed as new without reference to Oosomides Lacordaire, 1863.
Tribe
Ophryastini
Lacordaire, 1863
Ophryastides Lacordaire, 1863: 256 [stem: Ophryast-]. Type genus: Ophryastes Germar, 1829. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1874b: 454, as Ophryastini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 167, as Ophryastini).
Tribe
Ophtalmorrhynchini
Hoffmann, 1965
Ophtalmorynchini Hoffmann, 1965: 1411 [stem: Ophtalmorrhynch-]. Type genus: Ophtalmorrhynchus Hoffmann, 1965. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Otiorhynchini
Schönherr, 1826
Loborhynchides Schönherr, 1823: column 1144 [stem: Loborhynch-]. Type genus: Loborhynchus Schönherr, 1823 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1972); see Lyal and Alonso-Zarazaga (2010) and Appendix 6; syn. of Otiorhynchus Germar, 1822]. Comment: Loborhynchinae Schönherr, 1823 placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1972).
Otiorhynchides Schönherr, 1826: 203 [stem: Otiorhynch-]. Type genus: Otiorhynchus Germar, 1822 [placed on the Official List of Generic Names in Zoology (ICZN 1972, as Otiorhynchus Germar, 1824); see Lyal and Alonso-Zarazaga (2010) and Appendix 6]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1972, as Otiorhynchinae Schönherr, 1826).
Brachyrrhinidae Bedel, 1883: 30 [stem: Brachyrhin-]. Type genus: Brachyrhinus Latreille, 1802 [as Brachyrrhinus, incorrect original spelling of type genus name, not in prevailing usage; Brachyrhinus Latreille, 1802 placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1972); syn. of Cryphiphorus Stierlin, 1883]. Comment: name placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1972).
Tribe
Ottistirini
Heller, 1925
Ottistirini Heller, 1925a: 56, in key [stem: Ottistir-]. Type genus: Ottistira Pascoe, 1872.
Tribe
Pachyrhynchini
Schönherr, 1826
Somatodides Schönherr, 1823: column 1139 [stem: Somatod-]. Type genus: Somatodes Schönherr, 1823 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1994c); syn. of Pachyrhynchus Germar, 1824]. Comment: name placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology as Somatodini Schönherr, 1823 (ICZN 1994c).
Pachyrhynchides Schönherr, 1826: 88 [stem: Pachyrhynch-]. Type genus: Pachyrhynchus Germar, 1824 [placed on the Official List of Generic Names in Zoology (ICZN 1970b)]. Comment: name placed on the Official List of Family-Group Names in Zoology as Pachyrhynchini Schönherr, 1826 (ICZN 1970b).
Tribe
Peritelini
Lacordaire, 1863
Péritélides Lacordaire, 1863: 178 [stem: Peritel-]. Type genus: Peritelus Germar, 1824. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1874b: 455, as Periteli), generally accepted as in Alonso-Zarazaga and Lyal (1999: 171, as Peritelini).
Simoini Pierce, 1913: 421, in key [stem: Simon-]. Type genus: Simo Dejean, 1821. Comment: incorrect original stem formation, not in prevailing usage.
Paraptochi Pierce, 1913: 423, in key [stem: Paraptoch-]. Type genus: Paraptochus Seidlitz, 1868.
Homorythmini Hoffmann, 1950: 154 [stem: Homorhythm-]. Type genus: Homorhythmus Bedel, 1883 [as Homorythmus, incorrect subsequent spelling of type genus name, not in prevailing usage; syn. of Simo Dejean, 1821]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Phyllobiini
Schönherr, 1826
Phyllobides Schönherr, 1826: 178 [stem: Phyllobi-]. Type genus: Phyllobius Germar, 1824 [placed on the Official List of Generic Names in Zoology (ICZN 1981c)]. Comment: Phyllobiini Schönherr, 1826 placed on the Official List of Family-Group Names in Zoology (ICZN 1981c); incorrect original stem formation, not in prevailing usage.
Evotini J. L. LeConte, 1874b: 454 [stem: Evot-]. Type genus: Evotus J. L. LeConte, 1874.
Aphrasti J. L. LeConte, 1874b: 458 [stem: Aphrast-]. Type genus: Aphrastus Say, 1831.
Metacinopinae Reitter, 1913b: 6, in key [stem: Metacinop-]. Type genus: Metacinops Kraatz, 1862.
Tribe
Polycatini
Marshall, 1956
Polycatini G. A. K. Marshall, 1956: 4 [stem: Polycat-]. Type genus: Polycatus Heller, 1913.
Tribe
Polydrusini
Schönherr, 1823
Polydrosides Schönherr, 1823: column 1144 [stem: Polydrus-]. Type genus: Polydrusus Germar, 1817 [as Polydrosus, unjustified emendation of type genus name by Schönherr (1823), not in prevailing usage; syn. of Polydrusus Germar, 1817; Polydrusus Germar, 1817 placed on the Official List of Generic Names in Zoology (ICZN 1981c)]. Comment: placed on the Official List of Family-Group Names in Zoology (ICZN 1981c, as Polydrosini Schönherr, 1823), however, since this family-group name was based on an unjustified emendation of the type genus name, it was corrected (see Art. 32.5.3.2) to Polydrusini by Alonso-Zarazaga and Lyal (1999: 174).
Phyllomanisidae Gistel, 1848: [8] [stem: Phylloman-]. Type genus: Phyllomanes Gistel, 1848 [preoccupied genus name, not Phyllomanes Cabanis, 1847 [Aves]; syn. of Polydrusus Germar, 1817]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Liophloeidae Gistel, 1848: [8] [stem: Liophloe-]. Type genus: Liophloeus Germar, 1817.
Oligocyidae Gistel, 1856a: 373 [stem: Oligoce-]. Type genus: Oligocys Gistel, 1856 [syn. of Liophloeus Germar, 1817]. Comment: incorrect original stem formation, not in prevailing usage.
Scythropides Lacordaire, 1863: 380 [stem: Scythrop-]. Type genus: Scythropus Schönherr, 1826 [syn. of Pachyrhinus Schönherr, 1823]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Marseul (1863: 224, as Scytropidae [incorrect stem formation]), generally accepted as in Hustache (1919: 509, as Scythropini).
Auchmeresthinae Reitter, 1913b: 6, in key [stem: Auchmeresthet-]. Type genus: Auchmeresthes Kraatz, 1862. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Premnotrypini
Kuschel, 1956
Premnotrypini Kuschel, 1956: 187 [stem: Premnotryp-]. Type genus: Premnotrypes Pierce, 1914.
†Tribe
Pristorhynchini
Heer, 1847
Pristorhynchiden Heer, 1847: 190 [stem: Pristorhynch-]. Type genus: Pristorhynchus Heer, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Scudder (1893: 29, as Pristorhynchini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 176, as Pristorhynchini).
Tribe
Prypnini
Lacordaire, 1863
Prypnides Lacordaire, 1863: 135 [stem: Prypn-]. Type genus: Prypnus Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 225, as Prypnini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 176, as Prypnini).
Tribe
Psallidiini
Lacordaire, 1863
Psalidiides Lacordaire, 1863: 138 [stem: Psallidi-]. Type genus: Psallidium Herbst, 1795 [as Psalidium, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Everts (1903: 547, as Psalidiini [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 176, as Psallidiini); incorrect original stem formation, not in prevailing usage.
Tribe
Rhyncogonini
Sharp, 1919
Rhyncogonides Sharp, 1919a: 77 [stem: Rhyncogon-]. Type genus: Rhyncogonus Sharp, 1885.
Tribe
Sciaphilini
Sharp, 1891
Sciaphilina Sharp, 1891: 167 [stem: Sciaphil-]. Type genus: Sciaphilus Schönherr, 1823.
Sciaphilini Pierce, 1913: 405, in key [stem: Sciaphil-]. Type genus: Sciaphilus Schönherr, 1823. Comment: family-group name proposed as new without reference to Sciaphilina Sharp, 1891.
Tribe
Sitonini
Gistel, 1848
Sitonisidae Gistel, 1848: [8] [stem: Siton-]. Type genus: Sitones Schönherr, 1840 [syn. of Sitona Germar, 1817]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Tanymecini
Lacordaire, 1863
Tanymécides Lacordaire, 1863: 82 [stem: Tanymec-]. Type genus: Tanymecus Germar, 1817.
Subtribe
Piazomiina
Reitter, 1913
Piazomiini Reitter, 1913b: 28, in key [stem: Piazomi-]. Type genus: Piazomias Schönherr, 1840.
Subtribe
Tainophthalmina
Desbrochers des Loges, 1873
Tainophthalmidae Desbrochers des Loges, 1873: 426 [stem: Tainophthalm-]. Type genus: Tainophthalmus Desbrochers des Loges, 1873.
Amomphi J. L. LeConte, 1874b: 455 [stem: Amomph-]. Type genus: Amomphus Schönherr, 1848 [syn. of Aspidiotes Schönherr, 1847].
Subtribe
Tanymecina
Lacordaire, 1863
Tanymécides Lacordaire, 1863: 82 [stem: Tanymec-]. Type genus: Tanymecus Germar, 1817. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1874b: 454, as Tanymecini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 178, as Tanymecini).
Hadromerides Jekel, 1865: 545 [stem: Hadromer-]. Type genus: Hadromerus Schönherr, 1834 [preoccupied genus name, not Hadromerus Schönherr, 1823 [Coleoptera: Curculionidae: Entiminae]; syn. of Hadromeropsis Pierce, 1913]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Siderodactylides Jekel, 1865: 551, in note [stem: Siderodactyl-]. Type genus: Siderodactylus Schönherr, 1834 [syn. of Hadromerus Schönherr, 1823].
Pachnaei J. L. LeConte, 1874b: 457 [stem: Pachnae-]. Type genus: Pachnaeus Schönherr, 1826.
Minyomeri G. H. Horn, 1876: 17 [stem: Minyomer-]. Type genus: Minyomerus G. H. Horn, 1876.
Pandeleteini Pierce, 1913: 399, in key [stem: Pandeletei-]. Type genus: Pandeleteius Schönherr, 1834. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Tanyrhynchini
Schönherr, 1826
Tanyrhynchides Schönherr, 1826: 212 [stem: Tanyrhynch-]. Type genus: Tanyrhynchus Schönherr, 1826.
Éremnides Lacordaire, 1863: 220 [stem: Eremn-]. Type genus: Eremnus Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1869: 380, as Eremnini), generally accepted as in Schenkling and G. A. K. Marshall (1931b: Erem. 13, as Eremnini).
Tribe
Thecesternini
Lacordaire, 1863
Thécesternides Lacordaire, 1863: 306 [stem: Thecestern-]. Type genus: Thecesternus Say, 1831. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Sharp (1891: 86, as Thecesterninae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 182, as Thecesternini).
Tribe
Trachyphloeini
Gistel, 1848
Trachyphloeidae Gistel, 1848: [7] [stem: Trachyphloe-]. Type genus: Trachyphloeus Germar, 1817.
Subtribe
Trachyphilina
Voss, 1948
Trachyphilina Voss, 1948: 73, in key [stem: Trachyphil-]. Type genus: Trachyphilus Faust, 1887.
Subtribe
Trachyphloeina
Gistel, 1848
Trachyphloeidae Gistel, 1848: [7] [stem: Trachyphloe-]. Type genus: Trachyphloeus Germar, 1817.
Phyllastolidae Gistel, 1856a: 372 [stem: Phyllastol-]. Type genus: Phyllastolus Gistel, 1856 [syn. of Trachyphloeus Germar, 1817].
Cathormiocerini Reitter, 1913b: 8, in key [stem: Cathormiocer-]. Type genus: Cathormiocerus Schönherr, 1842.
Trachyphloeini Pierce, 1913: 421, in key [stem: Trachyphloe-]. Type genus: Trachyphloeus Germar, 1817. Comment: family-group name proposed as new without reference to Trachyphloeidae Gistel, 1848.
Pseudocneorrhinini Kôno, 1930: 163, in key [stem: Pseudocneorhin-]. Type genus: Pseudocneorhinus Roelofs, 1873 [as Pseudocneorrhinus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Callirhopalini Voss, 1956b: 23 [stem: Callirhopal-]. Type genus: Callirhopalus Hochhuth, 1851.
Tribe
Tropiphorini
Marseul, 1863
Tropiphoridae Marseul, 1863: 220 [stem: Tropiphor-]. Type genus: Tropiphorus Schönherr, 1842 [placed on the Official List of Generic Names in Zoology (ICZN 1988c)]. Comment: Tropiphorini, published on 15 June 1863, is given priority over Strangaliodini, Byrsopagini, Pantopoeini and Synaptonychini which were published by Lacordaire “before 10 August 1863”.
Leptopsides Lacordaire, 1863: 232 [stem: Leptop-]. Type genus: Leptops Schönherr, 1834 [preoccupied genus name, not Leptops Rafinesque, 1820 [Pisces]; syn. of Leptopius Oke, 1951]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870a: 181, as Leptopinae), generally accepted as in Heyne and Taschenberg (1907: 226, as Leptopsini [incorrect stem formation]); permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Strangaliodides Lacordaire, 1863: 234 [stem: Strangaliod-]. Type genus: Strangaliodes Schönherr, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by G. H. Horn (1876: 37, as Strangaliodes [treated as Latin]), generally accepted as in Voss (1954: 237, as Strangaliodini).
Byrsopagides Lacordaire, 1863: 337 [stem: Byrsopag-]. Type genus: Byrsopages Schönherr, 1842. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 228, as Byrsopagini), generally accepted as in Schenkling and G. A. K. Marshall (1929: 57, as Byrsopagini).
Pantopéides Lacordaire, 1863: 346 [stem: Pantopoe-]. Type genus: Pantopoeus Schönherr, 1842 [syn. of Perperus Schönherr, 1842]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Faust (1892a: 179, as Pantopeinarum [incorrect stem formation]); incorrect original stem formation, not in prevailing usage.
Synaptonycides Lacordaire, 1863: 372 [stem: Synaptonych-]. Type genus: Synaptonyx G. R. Waterhouse, 1853. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Wollaston (1877: 158, as Synaptonychides [treated as Latin]); incorrect original stem formation, not in prevailing usage.
Rhigopsini J. L. LeConte, 1874b: 454 [stem: Rhigopse-]. Type genus: Rhigopsis J. L. LeConte, 1874. Comment: incorrect original stem formation, not in prevailing usage, the comment stating the contrary in Alonso-Zarazaga and Lyal (2002: 28) is incorrect.
Dyslobini J. L. LeConte, 1874b: 454 [stem: Dyslob-]. Type genus: Dyslobus J. L. LeConte, 1869.
Phyxeles G. H. Horn, 1876: 56 [stem: Phyxelid-]. Type genus: Phyxelis Schönherr, 1842. Comment: incorrect original stem formation, not in prevailing usage.
Dirotognathini G. H. Horn, 1876: 79 [stem: Dirotognath-]. Type genus: Dirotognathus G. H. Horn, 1876.
Synirmini Bedel, 1883: 32, in key [stem: Synirm-]. Type genus: Synirmus Bedel, 1883 [syn. of Tropiphorus Schönherr, 1842].
Stenocorynini McKeown, 1939: 408 [stem: Stenocoryn-]. Type genus: Stenocorynus Schönherr, 1842. Comment: replacement name for Leptopsides Lacordaire, 1863 because of the homonymy of the type genus.
Leptopiinae Oke, 1951: 24 [stem: Leptopi-]. Type genus: Leptopius Oke, 1951. Comment: replacement name for Leptopsides Lacordaire, 1863 because of the homonymy of the type genus.
Leptosinae G. A. K. Marshall, 1952: 264 [stem: Leptos-]. Type genus: Leptosus G. A. K. Marshall, 1952 [syn. of Leptopius Oke, 1951]. Comment: replacement name for Leptopsides Lacordaire, 1863 because of the homonymy of the type genus.
Tribe
Typhlorhinini
Kuschel, 1954
Typhlorhinini Kuschel, 1954: 287 [stem: Typhlorhin-]. Type genus: Typhlorhinus Kuschel, 1954 [syn. of Hapactorrhynchus Richard, 1953].
Subfamily
Hyperinae
Marseul, 1863 (1848)
Hyperidae Marseul, 1863: 224 [stem: Hyper-]. Type genus: Hypera Germar, 1817. Comment: use of family-group name conserved over Phytonominae Gistel, 1848 (Art. 40.2).
Tribe
Cepurini
Capiomont, 1867
*Haplopides Lacordaire, 1863: 394 [stem: Haplopod-]. Type genus: Haplopus Schönherr, 1840 [preoccupied genus name, not Haplopus Burmeister, 1838 [Orthoptera]; syn. of Haplopodus G. A. K. Marshall, 1946]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Heyne and Taschenberg (1907: 228, as Haplopini) but not generally accepted as valid; Haplopidae was used as valid by Ienistea (1986: 33) but it was not attributed to Lacordaire (1863); Ienistea’s name is also unavailable, it was proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); if found to be available then permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Cépurides Capiomont, 1867: 438 [stem: Cepur-]. Type genus: Cepurus Schönherr, 1834. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Csiki (1934: 3, as Cepurini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 189, as Cepurini).
Tribe
Hyperini
Marseul, 1863 (1848)
Phytonomidae Gistel, 1848: [8] [stem: Phytonom-]. Type genus: Phytonomus Schönherr, 1823 [syn. of Hypera Germar, 1817]. Comment: use of the younger names Hyperinae/-ini Marseul, 1863 are conserved over this name (Art. 40.2).
Hyperidae Marseul, 1863: 224 [stem: Hyper-]. Type genus: Hypera Germar, 1817. Comment: published 15 June 1863; use of family-group name conserved over Phytonomini Gistel, 1848 (Art. 40.2); this family-group name was also used in the same year by Lacordaire (1863 [before 10 August]: 395, as Hypérides).
Coniatina Legalov, 2007: 401 [stem: Coniat-]. Type genus: Coniatus Germar, 1817.
Macrotarrhusina Legalov, 2007: 401 [stem: Macrotarrh-]. Type genus: Macrotarrhus Bedel, 1906. Comment: incorrect original stem formation, not in prevailing usage.
Subfamily
Lixinae
Schönherr, 1823
Lixides Schönherr, 1823: column 1146 [stem: Lix-]. Type genus: Lixus Fabricius, 1801.
Tribe
Cleonini
Schönherr, 1826
nomen protectum
Geomorides Schönherr, 1823: column 1141 [stem: Geomor-]. Type genus: Geomorus Schönherr, 1823 [syn. of Cleonis Dejean, 1821]. Comment: as pointed out by Alonso-Zarazaga and Lyal (1999: 190) this is the oldest available name for the tribe name, however, it has not been used as valid after 1899 and therefore we treat it here as a nomen oblitum (see Appendix 1).
Cleonides Schönherr, 1826: 145 [stem: Cleon-]. Type genus: Cleonus Schönherr, 1826 [syn. of Cleonis Dejean, 1821]. Comment: nomen protectum (see Appendix 1).
Cleonidae Kirby, 1837: 198 [stem: Cleon-]. Type genus: Cleonis Dejean, 1821.
Xerobiadae Gistel, 1856a: 373 [stem: Xerobi-]. Type genus: Xerobia Gistel, 1856 [syn. of Cleonis Dejean, 1821]. Comment: incorrect original stem formation, not in prevailing usage.
*Bothynodérides Chevrolat, 1872: 16 [stem: Bothynoder-]. Type genus: Bothynoderes sensu Chevrolat, 1872 [not Bothynoderes Schönherr, 1823; syn. of Asproparthenis Gozis, 1886]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
*Conorhynchides Chevrolat, 1872: 17 [stem: Conorhynch-]. Type genus: Conorhynchus Motschulsky, 1860. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form, e.g., Handlirsch (1925: 687 [as Cneorhynchides; lapsus calami for Conorhynchides [treated as Latin]), but not generally accepted as valid; Conorhynchinae Gill, 1861 (type genus Conorhynchus Gill, 1861, unjustified emendation of Conorynchus Nozeman, 1758 and junior homonym of Conorhynchus Motschulsky, 1860) is available in Pisces; the Pisces family-group name is permanently invalid since it it based on a preoccupied type genus (Art. 39).
*Cossinodérides Chevrolat, 1872: 18 [stem: Cossinoder-]. Type genus: Cossinoderus Chevrolat, 1872 [syn. of Porocleonus Motschulsky, 1860]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.
Epirhynchini Petri, 1914: 3, in key [stem: Epirrhynch-]. Type genus: Epirrhynchus Schönherr, 1823 [as Epirhynchus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Tribe
Lixini
Schönherr, 1823
Lixides Schönherr, 1823: column 1146 [stem: Lix-]. Type genus: Lixus Fabricius, 1801.
*Lariniden Suffrian, 1848: 62 [stem: Larin-]. Type genus: Larinus Germar, 1824 [preoccupied genus name, not Larinus Dejean, 1821 [Coleoptera: Curculionidae: Lixinae: Lixini]; syn. of Larinus Dejean, 1821]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Suffrian (1848); if found to be available then permanently invalid (Art. 39): based on preoccupied type genus.
Larinidae Gistel, 1848: [7] [stem: Larin-]. Type genus: Larinus Germar, 1824 [preoccupied genus name, not Larinus Dejean, 1821 [Coleoptera: Curculionidae: Lixinae: Lixini]; syn. of Larinus Dejean, 1821]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Phyllonomeidae Gistel, 1856a: 372 [stem: Phyllonome-]. Type genus: Phyllonomeus Gistel, 1856.
Larinini Petri, 1914: 4, in key [stem: Larin-]. Type genus: Larinus Germar, 1824 [preoccupied genus name, not Larinus Dejean, 1821 [Coleoptera: Curculionidae: Lixinae: Lixini]; syn. of Larinus Dejean, 1821]. Comment: family-group name proposed as new without reference to Larinidae Gistel, 1848; permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Rhinocyllini
Lacordaire, 1863
Rhinocyllides Lacordaire, 1863: 433 [stem: Rhinocyll-]. Type genus: Rhinocyllus Germar, 1817. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Faust (1904: 182, as Rhinocyllinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 193, as Rhinocyllini).
Subfamily
Mesoptiliinae
Lacordaire, 1863
Mésoptilides Lacordaire, 1863: 563 [stem: Mesoptili-]. Type genus: Mesoptilius Labram and Imhoff, 1845.
Tribe
Carciliini
Pierce, 1916
Carciliinae Pierce, 1916: 465 [stem: Carcili-]. Type genus: Carcilia Roelofs, 1874.
Tribe
Laemosaccini
Lacordaire, 1865
Lémosacides Lacordaire, 1865: 12 [stem: Laemosacc-]. Type genus: Laemosaccus Schönherr, 1823. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Laemosaccinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 194, as Laemosaccini); incorrect original stem formation, not in prevailing usage.
Tribe
Magdalidini
Pascoe, 1870
nomen protectum
Thamnophilides Schönherr, 1823: column 1136 [stem: Thamnophil-]. Type genus: Thamnophilus Schönherr, 1823 [preoccupied genus name, not Thamnophilus Vieillot, 1816 [Aves]; syn. of Magdalis Germar, 1817]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Scardamyctisidae Gistel, 1848: [7] [stem: Scardamyct-]. Type genus: Scardamyctes Gistel, 1848 [syn. of Magdalis Germar, 1817]. Comment: Scardamyctini Gistel, 1848 was recently considered a nomen oblitum by Colonnelli (2003: 7) however the necessary supporting references were not provided; we hereby consider Scardamyctini Gistel, 1848 as a nomen oblitum (see supporting references in Appendix 1); incorrect original stem formation, not in prevailing usage.
Magdalinidae Gistel, 1856a: 371 [stem: Magdalin-]. Type genus: Magdalinus Germar, 1843 [placed on the Official Index of Rejected and Invalid Generic Names in Zoology by the Commission (ICZN 1955a); syn. of Magdalis Germar, 1817]. Comment: permanently invalid (Art. 39): based on suppressed type genus.
Magdalinae Pascoe, 1870b: 436 [stem: Magdalid-]. Type genus: Magdalis Germar, 1817 [placed on the Official List of Generic Names in Zoology (ICZN 1955a)]. Comment: nomen protectum (see Appendix 1); incorrect original stem formation, not in prevailing usage.
Rhininae Pierce, 1916: 465 [stem: Rhin-]. Type genus: Rhina Latreille, 1802 [preoccupied genus name, not Rhina Schneider, 1801 [Pisces]; syn. of Panus Schönherr, 1823]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Mesoptiliini
Lacordaire, 1863
Mésoptilides Lacordaire, 1863: 563 [stem: Mesoptili-]. Type genus: Mesoptilius Labram and Imhoff, 1845. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Mesoptilinae [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 193, as Mesoptiliini); incorrect original stem formation, not in prevailing usage.
Cnemidophorini Hustache, 1937: 199, in key [stem: Cnemidophor-]. Type genus: Cnemidophorus Schönherr, 1835 [preoccupied genus name, not Cnemidophorus Wagler, 1830 [Reptilia]; syn. of Cnemidontus Schenkling, 1935]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Cnemidontini Kuschel, 1955: 271 [stem: Cnemidont-]. Type genus: Cnemidontus Schenkling, 1935. Comment: replacement name for Cnemidophorini Hustache, 1937 because of the homonymy of the type genus.
Subfamily
Molytinae
Schönherr, 1823
Molytides Schönherr, 1823: column 1142 [stem: Molyt-]. Type genus: Molytes Schönherr, 1823 [syn. of Liparus A. G. Olivier, 1807].
Tribe
Anoplini
Bedel, 1884
Anoplini Bedel, 1884a: 67, in key [stem: Anopl-]. Type genus: Anoplus Germar, 1820.
Tribe
Amalactini
Lacordaire, 1863
Amalactides Lacordaire, 1863: 506 [stem: Amalact-]. Type genus: Amalactus Schönherr, 1835. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Amalactinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 196, as Amalactini).
Tribe
Aminyopini
Voss, 1956
Aminyopini Voss, 1956b: 30, in key [stem: Aminyop-]. Type genus: Aminyops Voss, 1956.
Niphadini Voss, 1963: 5 [stem: Niphad-]. Type genus: Niphades Pascoe, 1871.
Niphadonothina Voss, 1965: 344, in key [stem: Niphadonoth-]. Type genus: Niphadonothus Voss, 1965.
Tribe
Amorphocerini
Voss, 1939
Amorphocerini Voss, 1939: 65 [stem: Amorphocer-]. Type genus: Amorphocerus Schönherr, 1826.
Tribe
Anchonini
Imhoff, 1856
Anchonidae Imhoff, 1856: [2] 213 [stem: Anchon-]. Type genus: Anchonus Schönherr, 1825.
Tribe
Brachyceropseini
Aurivillius, 1926
Brachyceropseinae Aurivillius, 1926b: 2 [stem: Brachyceropse-]. Type genus: Brachyceropsis Aurivillius, 1926. Comment: Alonso-Zarazaga and Lyal (2002: 18) incorrectly changed their previous opinion in Alonso-Zarazaga and Lyal (1999: 103) regarding the correct stem based on this genus name, the correct stem is Brachyceropse-.
Tribe
Cholini
Schönherr, 1825
Cholides Schönherr, 1825: column 584 [stem: Chol-]. Type genus: Cholus Germar, 1824 [placed on the Official List of Generic Names in Zoology (ICZN 1987d)].
Subtribe
Cholina
Schönherr, 1825
Cholides Schönherr, 1825: column 584 [stem: Chol-]. Type genus: Cholus Germar, 1824 [placed on the Official List of Generic Names in Zoology (ICZN 1987d)].
Amerininae Pierce, 1919: 30, in key [stem: Amerin-]. Type genus: Ameris Dejean, 1821.
Subtribe
Cholomina
Vaurie, 1974
Cholomini Vaurie, 1974: 4 [stem: Cholom-]. Type genus: Cholomus Roelofs, 1880.
Subtribe
Rhinastina
Vaurie, 1973
Rhinastini Vaurie, 1973: 4, in key [stem: Rhinast-]. Type genus: Rhinastus Schönherr, 1825.
Tribe
Cleogonini
Gistel, 1848
Cleogonidae Gistel, 1848: [7] [stem: Cleogon-]. Type genus: Cleogonus Schönherr, 1825.
Tribe
Conotrachelini
Jekel, 1865
Conotrachelides Jekel, 1865: 550 [stem: Conotrachel-]. Type genus: Conotrachelus Dejean, 1835.
Echinaspini Blatchley, 1922: 121 [stem: Echinaspid-]. Type genus: Echinaspis Blatchley, 1922 [preoccupied genus name, not Echinaspis Haeckel, 1881 [Protista]; syn. of Microhyus J. L. LeConte, 1876]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.
Tribe
Cycloterini
Lacordaire, 1863
Cyclotérides Lacordaire, 1863: 365 [stem: Cycloter-]. Type genus: Cycloteres Schönherr, 1843.
Subtribe
Cycloterina
Lacordaire, 1863
Cyclotérides Lacordaire, 1863: 365 [stem: Cycloter-]. Type genus: Cycloteres Schönherr, 1843. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 228, as Cycloterini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 200, as Cycloterini).
Subtribe
Thrombosternina
Voss, 1965
Thrombosternina Voss, 1965: 344, in key [stem: Thrombostern-]. Type genus: Thrombosternus G. A. K. Marshall, 1955.
Tribe
Dinomorphini
Lacordaire, 1863
Dinomorphides Lacordaire, 1863: 291 [stem: Dinomorph-]. Type genus: Dinomorphus Perty, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Dinomorphinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 201, as Dinomorphini).
Tribe
Emphyastini
Lacordaire, 1863
Emphiastides Lacordaire, 1863: 510 [stem: Emphyast-]. Type genus: Emphyastes Mannerheim, 1852 [as Emphiastes, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by J. L. LeConte (1876: 137, as Emphyastini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 143, as Emphyastina); incorrect original stem formation, not in prevailing usage.
Phycocoetes J. L. LeConte, 1876: 189 [stem: Phycocoet-]. Type genus: Phycocoetes J. L. LeConte, 1876 [preoccupied genus name, not Phycocoetes Agassiz, 1846 [Pisces]; syn. of Thalasselephas Egorov and Korotyaev, 1976]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Thalasselephantini Alonso-Zarazaga and Lyal, 1999: 208 [stem: Thalasselephant-]. Type genus: Thalasselephas Egorov and Korotyaev, 1976. Comment: replacement name for Phycocoetini J. L. LeConte, 1876 because of the homonymy of the type genus.
Tribe
Euderini
Lacordaire, 1865
Eudérides Lacordaire, 1865: 18 [stem: Euder-]. Type genus: Euderes Schönherr, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Euderinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 201, as Euderini); the junior homonym Euderini Erdös, 1956 (type genus Euderus Haliday, 1844) is available in Hymenoptera: Eulophidae; the name Entiinae Hedqvist, 1974 is now used as valid in Hymenoptera instead of Euderini Erdös, 1956 (see Hansson and Straka 2009: 272).
Tribe
Galloisiini
Morimoto, 1962
Galloisiinae Morimoto, 1962b: 375 [stem: Galloisi-]. Type genus: Galloisia Hustache, 1920.
Tribe
Guioperini
Lacordaire, 1865
Guiopérides Lacordaire, 1865: 78 [stem: Guioper-]. Type genus: Guioperus Perty, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 232, as Guioperini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 201, as Guioperini).
Tribe
Hylobiini
Kirby, 1837
Hylobidae Kirby, 1837: 196 [stem: Hylobi-]. Type genus: Hylobius Germar, 1817.
Subtribe
Epistrophina
Marshall, 1932
Epistrophina G. A. K. Marshall, 1932: 346, in key [stem: Epistroph-]. Type genus: Epistrophus Kirsch, 1868 [preoccupied genus name, not Epistrophus Gistel, 1834 [Coleoptera: Elateridae]; Epistrophus Kirsch, 1868 is currently considered a valid genus name]. Comment: as pointed out by Alonso-Zarazaga and Lyal (1999: 202) this family-group name should be treated as permanently invalid (Art. 39) because it is based on a preoccupied type genus, however, an application will be submitted by MAAZ and CHCL to the Commission for the conservation of the genus Epistrophus Kirsch, 1868 over its unused senior homonym; meanwhile current usage is maintained.
Subtribe
Hylobiina
Kirby, 1837
Hylobidae Kirby, 1837: 196 [stem: Hylobi-]. Type genus: Hylobius Germar, 1817. Comment: incorrect original stem formation, not in prevailing usage.
Heilipinae Faust, 1892b: 202, in key [stem: Heilipod-]. Type genus: Heilipus Germar, 1824. Comment: incorrect original stem formation, not in prevailing usage.
Syphorbina G. A. K. Marshall, 1932: 346, in key [stem: Syphorb-]. Type genus: Syphorbus Pascoe, 1881.
Tribe
Ithyporini
Lacordaire, 1865
Ithyporides Lacordaire, 1865: 50 [stem: Ithypor-]. Type genus: Ithyporus Schönherr, 1836. Comment: precedence (Ithyporini Lacordaire, 1865 vs Sclerocardiini Lacordaire, 1865) given to taxon originally proposed at the higher rank (Art. 24.1).
Subtribe
Colobodina
Voss, 1958
Colobodini Voss, 1958: 50, in key [stem: Colobod-]. Type genus: Colobodes Schönherr, 1837.
Subtribe
Ithyporina
Lacordaire, 1865
Ithyporides Lacordaire, 1865: 50 [stem: Ithypor-]. Type genus: Ithyporus Schönherr, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Kirsch (1870: 192, as Ithyporidae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 203, as Ithyporini).
Subtribe
Sclerocardiina
Lacordaire, 1865
Sclérocardiides Lacordaire, 1865: 317 [stem: Sclerocardi-]. Type genus: Sclerocardius Schönherr, 1847. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 233, as Sclerocardini [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 204, as Sclerocardiina).
Tribe
Itini
Reitter, 1913
Itini Reitter, 1913b: 58, in key [stem: It-]. Type genus: Ita Tournier, 1878.
Tribe
Juanorhinini
Aurivillius, 1931
Juanorhinini Aurivillius, 1931: 465 [stem: Juanorhin-]. Type genus: Juanorhinus Aurivillius, 1926.
Tribe
Lepyrini
Kirby, 1837
Lepyridae Kirby, 1837: 197 [stem: Lepyr-]. Type genus: Lepyrus Germar, 1817.
Tribe
Lithinini
Lacordaire, 1863
Lithinides Lacordaire, 1863: 349 [stem: Lithin-]. Type genus: Lithinus Klug, 1833.
Subtribe
Lithinina
Lacordaire, 1863
Lithinides Lacordaire, 1863: 349 [stem: Lithin-]. Type genus: Lithinus Klug, 1833. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Lithininae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 204, as Lithinini).
Subtribe
Rhytidophloeina
Voss, 1963
Rhytidophloeini Voss, 1963: 3 [stem: Rhytidophloe-]. Type genus: Rhytidophloeus Schönherr, 1842.
Tribe
Lymantini
Lacordaire, 1865
Lymantides Lacordaire, 1865: 328 [stem: Lymant-]. Type genus: Lymantes Schönherr, 1837. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 234, as Lymantini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 205, as Lymantini).
Ithaurinae Kuschel, 1959b: 509 [stem: Ithaur-]. Type genus: Ithaura Pascoe, 1871.
Caecossonina Osella, 1980: 369 [stem: Caecosson-]. Type genus: Caecossonus Gilbert, 1955.
Tribe
Mecysolobini
Reitter, 1913
Alcidides Jekel, 1865: 547 [stem: Alcid-]. Type genus: Alcides Schönherr, 1825 [preoccupied genus name, not Alcides Hübner, 1822 [Lepidoptera]; syn. of Sternuchopsis Heller, 1918]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Mecyslobini Reitter, 1913b: 33, in key [stem: Mecysolob-]. Type genus: Mecysolobus Reitter, 1905 [as Mecyslobus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Alcidodinae G. A. K. Marshall, 1939: 582 [stem: Alcidod-]. Type genus: Alcidodes G. A. K. Marshall, 1939 [syn. of Sternuchopsis Heller, 1918]. Comment: replacement name for Alcidinae Jekel, 1865 because of the homonymy of the type genus.
Tribe
Metatygini
Pascoe, 1888
Metatyginae Pascoe, 1888: 409 [stem: Metatyg-]. Type genus: Metatyges Pascoe, 1865 [syn. of Omophorus Schönherr, 1835].
Omophorinae G. A. K. Marshall, 1917: 195 [stem: Omophor-]. Type genus: Omophorus Schönherr, 1835.
Metatyginae Pierce, 1919: 30, in key [stem: Metatyg-]. Type genus: Metatyges Pascoe, 1865 [syn. of Omophorus Schönherr, 1835]. Comment: family-group name proposed as new without reference to Metatyginae Pascoe, 1888.
Sternechosomini Voss, 1958: 43, in key [stem: Sternechosom-]. Type genus: Sternechosomus Voss, 1958.
Tribe
Molytini
Schönherr, 1823
Molytides Schönherr, 1823: column 1142 [stem: Molyt-]. Type genus: Molytes Schönherr, 1823 [syn. of Liparus A. G. Olivier, 1807].
Subtribe
Leiosomatina
Reitter, 1913
Liosomina Reitter, 1913b: 52, in key [stem: Leiosomat-]. Type genus: Leiosoma Stephens, 1829 [as Liosoma, unjustified emendation of type genus name by Agassiz (1846b: 204), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.
Subtribe
Molytina
Schönherr, 1823
Molytides Schönherr, 1823: column 1142 [stem: Molyt-]. Type genus: Molytes Schönherr, 1823 [syn. of Liparus A. G. Olivier, 1807].
Liparides Latreille, 1828: 597 [stem: Lipar-]. Type genus: Liparus A. G. Olivier, 1807.
Subtribe
Plinthina
Lacordaire, 1863
Plinthides Lacordaire, 1863: 359 [stem: Plinth-]. Type genus: Plinthus sensu Westwood, 1838 [not Plinthus Germar, 1817; syn. of Mitoplinthus Reitter, 1897 but see Alonso-Zarazaga and Lyal (2010) and Appendix 6]. Comment: based on a misidentified type genus; an application was recently submitted to the Commission by Alonso-Zarazaga and Lyal (2010) to keep the name Plinthina in its usual sense, with Plinthus Germar, 1817 as its type genus (see Appendix 6).
Minyopidae Marseul, 1863: 221 [stem: Minyop-]. Type genus: Minyops Schönherr, 1823.
Subtribe
Typoderina
Voss, 1965
Typoderina Voss, 1965: 343, in key [stem: Typoder-]. Type genus: Typoderus G. A. K. Marshall, 1953.
Tribe
Nettarhinini
Lacordaire, 1865
Nettarhinides Lacordaire, 1865: 76 [stem: Nettarhin-]. Type genus: Nettarhinus Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hustache (1936: 62, as Nettarhinina), generally accepted as in Alonso-Zarazaga and Lyal (1999: 206, as Nettarhinini).
Tribe
Pacholenini
Lacordaire, 1863
Pacholénides Lacordaire, 1863: 443 [stem: Pacholen-]. Type genus: Pacholenus Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 229, as Pacholenini), generally accepted as in Alonso-Zarazaga and Lyal (1999: 206, as Pacholenini).
Tribe
Paipalesomini
Marshall, 1932
Paipalesomini G. A. K. Marshall, 1932: 345, in key [stem: Paipalesom-]. Type genus: Paipalesomus Schönherr, 1847 [syn. of Peribleptus Schönherr, 1843].
Tribe
Petalochilini
Lacordaire, 1863
Pétalochilides Lacordaire, 1863: 517 [stem: Petalochil-]. Type genus: Petalochilus Schönherr, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Petalochilinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 206, as Petalochilini).
Épipédides Lacordaire, 1865: 186 [stem: Epiped-]. Type genus: Epipedus Schönherr, 1842 [preoccupied genus name, not Epipedus Spinola, 1837 [Hemiptera]; syn. of Epipedophyes G. A. K. Marshall, 1946]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Pascoe (1870b: 437, as Epipedinae); permanently invalid (Art. 39): based on preoccupied type genus.
Hormopini J. L. LeConte, 1876: 320 [stem: Hormop-]. Type genus: Hormops J. L. LeConte, 1876.
Schoenherriellinae Viana, 1952: 231 [stem: Schoenherriell-]. Type genus: Schoenherriella Viana, 1952 [syn. of Epipedophyes G. A. K. Marshall, 1946]. Comment: replacement name for Epipédides Lacordaire, 1865 because of the homonymy of the type genus.
Epipedophyinae Kuschel, 1955: 270 [stem: Epipedophy-]. Type genus: Epipedophyes G. A. K. Marshall, 1946. Comment: replacement name for Epipédides Lacordaire, 1865 because of the homonymy of the type genus.
Tribe
Phoenicobatini
Champion, 1914
Phoenicobatina Champion, 1914: 416 [stem: Phoenicobat-]. Type genus: Phoenicobates Champion, 1914.
Tribe
Phrynixini
Kuschel, 1964
Phrynixinae Kuschel, 1964: 472 [stem: Phrynix-]. Type genus: Phrynixus Pascoe, 1875.
Tribe
Pissodini
Gistel, 1848
Pissodisidae Gistel, 1848: [7] [stem: Pissod-]. Type genus: Pissodes Germar, 1817.
Subtribe
Cotasteromimina
Morimoto, 1962
Cotasteromimini Morimoto, 1962a: 61, in key [stem: Cotasteromim-]. Type genus: Cotasteromimus Chûjô and Voss, 1960.
Subtribe
Orthorhinina
Jekel, 1865
Orthorhinides Jekel, 1865: 548 [stem: Orthorhin-]. Type genus: Orthorhinus Schönherr, 1825.
Subtribe
Pissodina
Gistel, 1848
Pissodisidae Gistel, 1848: [7] [stem: Pissod-]. Type genus: Pissodes Germar, 1817. Comment: incorrect original stem formation, not in prevailing usage.
Angianides Sharp, 1919b: 151 [stem: Angian-]. Type genus: Angianus Sharp, 1919 [syn. of Vanapa Pouillaude, 1915].
Tribe
Sternechini
Lacordaire, 1863
Sternéchides Lacordaire, 1863: 447 [stem: Sternech-]. Type genus: Sternechus Schönherr, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Champion (1902: 113, as Sternechina), generally accepted as in Alonso-Zarazaga and Lyal (1999: 208, as Sternechini).
Tribe
Styanacini
Chûjô and Voss, 1960
Styanacinae Chûjô and Voss, 1960: 2 [stem: Styanac-]. Type genus: Styanax Pascoe, 1871.
Tribe
Trachodini
Gistel, 1848
Trachodisidae Gistel, 1848: [7] [stem: Trachod-]. Type genus: Trachodes Germar, 1824. Comment: incorrect original stem formation, not in prevailing usage.
Blastophiladae Gistel, 1856a: 370 [stem: Blastophil-]. Type genus: Blastophila Gistel, 1856 [syn. of Trachodes Germar, 1824]. Comment: incorrect original stem formation, not in prevailing usage.
Acicnémides Lacordaire, 1865: 31 [stem: Acicnemid-]. Type genus: Acicnemis Fairmaire, 1849. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Klima (1935: Acic. 1, as Acicnemidinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 196, as Acicnemidini); incorrect original stem formation, not in prevailing usage.
Tribe
Trigonocolini
Lacordaire, 1863
Trigonocolides Lacordaire, 1863: 592 [stem: Trigonocol-]. Type genus: Trigonocolus Lacordaire, 1863. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 436, as Trigonocolinae), generally accepted as in Alonso-Zarazaga and Lyal (1999: 208, as Trigonocolini).
Megarhininae Faust, 1888: 284 [stem: Megarhin-]. Type genus: Megarhinus Schönherr, 1835 [preoccupied genus name, not Megarhinus Rafinesque, 1820 [Pisces]; syn. of Trigonocolus Lacordaire, 1863]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Tribe
Trypetidini
Lacordaire, 1865
Trypétides Lacordaire, 1865: 177 [stem: Trypetid-]. Type genus: Trypetes Schönherr, 1836 [placed on the Official List of Generic Names in Zoology (ICZN 1974a)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1870b: 437, as Trypetinae [incorrect stem formation]), generally accepted as in Alonso-Zarazaga and Lyal (1999: 209, as Trypetidini); incorrect original stem formation, not in prevailing usage; stem based on Trypetes Schönherr, 1836 ruled to be Trypetid- and Trypetidinae placed on the Official List of Family-Group Names in Zoology (ICZN 1974a, as Trypetidinae Pierce, 1919).
Trypetesidae Heller, 1916: 348, in note [stem: Trypetes-]. Type genus: Trypetes Schönherr, 1836 [placed on the Official List of Generic Names in Zoology (ICZN 1974a)]. Comment: emendation for Trypetidae Lacordaire, 1865 to avoid homonymy with Trypetidae Loew, 1861 (type genus Trypeta Meigen, 1803) in Diptera; corrected spelling not adopted since it creates homonymy problem with Trypetesidae Stebbing, 1910 (type genus Trypetesa Norman, 1903) in Crustacea.
Subfamily
Orobitidinae
Thomson, 1859
Orobitina C. G. Thomson, 1859: 138 [stem: Orobitid-]. Type genus: Orobitis Germar, 1817. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Alonso-Zarazaga and Lyal (1999: 210).
Subfamily
Xiphaspidinae
Marshall, 1920
Xiphaspidinae G. A. K. Marshall, 1920: 393 [stem: Xiphaspid-]. Type genus: Xiphaspis G. A. K. Marshall, 1920.
Subfamily
Scolytinae
Latreille, 1804
Scolitarii Latreille, 1804c: 156 [stem: Scolyt-]. Type genus: Scolytus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1963b)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1963b, as Scolytidae Westwood, 1838).
Tribe
Amphiscolytini
Mandelshtam and Beaver, 2003
Amphiscolytini Mandelshtam and Beaver, 2003: 2 [stem: Amphiscolyt-]. Type genus: Amphiscolytus Mandelshtam and Beaver, 2003.
Tribe
Bothrosternini
Blandford, 1896
Bothrosternini Blandford, 1896: 120 [stem: Bothrostern-]. Type genus: Bothrosternus Eichhoff, 1868.
Tribe
Cactopinini
Chamberlin, 1939
Cactopinae Chamberlin, 1939: 243 [stem: Cactopin-]. Type genus: Cactopinus Schwartz, 1899. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Wood (1978: 113).
Tribe
Carphodicticini
Wood, 1971
Carphodicticini Wood, 1971: 19 [stem: Carphodictic-]. Type genus: Carphodicticus Wood, 1971.
Tribe
Coptonotini
Chapuis, 1869
Coptonotidae Chapuis, 1869: 11 [stem: Coptonot-]. Type genus: Coptonotus Chapuis, 1869.
Tribe
Corthylini
LeConte, 1876
Corthyli J. L. LeConte, 1876: 347 [stem: Corthyl-]. Type genus: Corthylus Erichson, 1836.
Subtribe
Corthylina
LeConte, 1876
Corthyli J. L. LeConte, 1876: 347 [stem: Corthyl-]. Type genus: Corthylus Erichson, 1836.
Amphicranidae Eichhoff, 1878: 460 [stem: Amphicran-]. Type genus: Amphicranus Erichson, 1836.
Xyleboripina Reitter, 1913c: 102 [stem: Xyleborip-]. Type genus: Xyleborips Reitter, 1913. Comment: incorrectly spelled Xyloboripina on page 31 of the same work.
Gnathotrichina Balachowsky, 1949: 241 [stem: Gnathotrich-]. Type genus: Gnathotrichus Eichhoff, 1869.
Subtribe
Pityophthorina
Eichhoff, 1878
Pityophthoridae Eichhoff, 1878: 173 [stem: Pityophthor-]. Type genus: Pityophthorus Eichhoff, 1864. Comment: First Revisers found (Pityophthorina Eichhoff, 1878 vs Araptina Eichhoff, 1878) are Wood and Bright (1992: 949).
Araptidae Eichhoff, 1878: 305 [stem: Arapt-]. Type genus: Araptus Eichhoff, 1872.
Tribe
Cryphalini
Lindemann, 1877
Cryphaloideae Lindemann, 1877: 165 [stem: Cryphal-]. Type genus: Cryphalus Erichson, 1836.
Trypophloeinae Nüsslin, 1911: 375 [stem: Trypophloe-]. Type genus: Trypophloeus Fairmaire, 1868.
Ernoporinae Nüsslin, 1911: 375 [stem: Ernopor-]. Type genus: Ernoporus C. G. Thomson, 1859.
*Eidopherinae Murayama, 1954: 200 [stem: Eidophel-]. Type genus: Eidophelus Eichhoff, 1875. Comment: unavailable (Art. 13.1): proposed after 1930 without description or bibliographic reference to such a description; incorrect original stem formation, not in prevailing usage; correction of stem by Wood (1978: 114).
Tribe
Crypturgini
LeConte, 1876
Crypturgi J. L. LeConte, 1876: 387 [stem: Crypturg-]. Type genus: Crypturgus Erichson, 1836.
†Tribe
Cylindrobrotini
Kirejtshuk, Azar, Beaver, Mandelshtam and Nel, 2009
Cylindrobrotini Kirejtshuk et al., 2009: 103 [stem: Cylindrobrot-]. Type genus: Cylindrobrotus Kirejtshuk et al., 2009.
Tribe
Diamerini
Hagedorn, 1909
Diamerinae Hagedorn, 1909: 163 [stem: Diamer-]. Type genus: Diamerus Erichson, 1836.
Strombophorini Schedl, 1959: 16 [stem: Strombophor-]. Type genus: Strombophorus Hagedorn, 1909. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Ferreira (1966: 664, as Strombophorini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).
Sphaerotrypini Murayama, 1963: 66 [stem: Sphaerotryp-]. Type genus: Sphaerotrypes Blandford, 1894.
Tribe
Dryocoetini
Lindemann, 1877
Dryocoetoideae Lindemann, 1877: 165 [stem: Dryocoet-]. Type genus: Dryocoetes Eichhoff, 1864 [placed on the Official List of Generic Names in Zoology (ICZN 1979b)].
Thamnurginae Nüsslin, 1911: 377 [stem: Thamnurg-]. Type genus: Thamnurgus Eichhoff, 1864.
Taphrorychini Reitter, 1913c: 29 [stem: Taphrorych-]. Type genus: Taphrorychus Eichhoff, 1878.
Tribe
Hexacolini
Eichhoff, 1878
Ctenophoridae Chapuis, 1869: 49 [stem: Ctenophor-]. Type genus: Ctenophorus Chapuis, 1869 [preoccupied genus name, not Ctenophorus Fitzinger, 1843 [Reptilia]; syn. of Scolytodes Ferrari, 1867]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Hexacolidae Eichhoff, 1878: 306 [stem: Hexacol-]. Type genus: Hexacolus Eichhoff, 1868. Comment: First Revisers found (Problechilini Eichhoff, 1878 vs Hexacolini Eichhoff, 1878) are Alonso-Zarazaga and Lyal (2009: 58).
Problechilidae Eichhoff, 1878: 167 [stem: Problechil-]. Type genus: Problechilus Eichhoff, 1878.
Erineophilides Hopkins, 1902: 37 [stem: Erineophil-]. Type genus: Erineophilus Hopkins, 1902.
Tribe
Hylastini
LeConte, 1876
Hylastes J. L. LeConte, 1876: 387 [stem: Hylast-]. Type genus: Hylastes Erichson, 1836.
Hylurgopina Balachowsky, 1949: 122 [stem: Hylurgop-]. Type genus: Hylurgops J. L. LeConte, 1876.
Tribe
Hylesinini
Erichson, 1836
Hylesinen Erichson, 1836: 46 [stem: Hylesin-]. Type genus: Hylesinus Fabricius, 1801. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Shuckard (1839b: 63, as Hylesinidae), generally accepted as in Alonso-Zarazaga and Lyal (2009: 63, as Hylesinini); incorrect original stem formation, not in prevailing usage.
Phloeotrupides Lacordaire, 1865: 370 [stem: Phloeotrup-]. Type genus: Phloeotrupes Erichson, 1836. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Chapuis (1869: 11, as Phloeotrupidae).
Phloeobori Blandford, 1893: 426 [stem: Phloeobor-]. Type genus: Phloeoborus Erichson, 1836.
Dactylipalpi Blandford, 1893: 426 [stem: Dactylipalp-]. Type genus: Dactylipalpus Chapuis, 1869.
Alniphagini Murayama, 1963: 29 [stem: Alniphag-]. Type genus: Alniphagus Swaine, 1918.
Tribe
Hylurgini
Gistel, 1848
Hylurgidae Gistel, 1848: [6] [stem: Hylurg-]. Type genus: Hylurgus Latreille, 1806. Comment: status resurrected by Alonso-Zarazaga and Lyal (2009: 69).
Dendroctonides Nüsslin, 1912: 278 [stem: Dendrocton-]. Type genus: Dendroctonus Erichson, 1836 [placed on the Official List of Generic Names in Zoology (ICZN 1963a)].
Xylechinides Nüsslin, 1912: 281 [stem: Xylechin-]. Type genus: Xylechinus Chapuis, 1869.
Tomicini Wood, 1978: 118, in key [stem: Tomic-]. Type genus: Tomicus Latreille, 1802 [placed on the Official List of Generic Names in Zoology (ICZN 1963a)]. Comment: the older name Tomicidae Shuckard, 1839 was based on misidentified type genus (see Alonso-Zarazaga and Lyal 2009); an application will be submitted by MAAZ and CHCL to the Commission to suppress Tomicidae Shuckard, 1839 for the Principles of Priority and Homonymy (Art. 65.2.1).
Tribe
Hyorrhynchini
Hopkins, 1915
Hyorrhynchinae Hopkins, 1915: 225, in key [stem: Hyorrhynch-]. Type genus: Hyorrhynchus Blandford, 1894.
*Sueinae Murayama, 1959: 26 [stem: Sue-]. Type genus: Sueus Murayama, 1951. Comment: unavailable (Art. 13.1): proposed after 1930 without description or bibliographic reference to such a description.
Hyorrhynchini Murayama, 1963: 62 [stem: Hyorrhynch-]. Type genus: Hyorrhynchus Blandford, 1894. Comment: family-group name proposed as new without reference to Hyorrhynchinae Hopkins, 1915.
Sueinae Murayama, 1963: 4 [stem: Sue-]. Type genus: Sueus Murayama, 1951.
Tribe
Hypoborini
Nüsslin, 1911
Hypoborinae Nüsslin, 1911: 376 [stem: Hypobor-]. Type genus: Hypoborus Erichson, 1836.
Chaetophloeini Schedl, 1966: 361 [stem: Chaetophloe-]. Type genus: Chaetophloeus J. L. LeConte, 1876.
Tribe
Ipini
Bedel, 1888
Tomicidae Shuckard, 1839b: 64 [stem: Tomic-]. Type genus: Tomicus sensu Shuckard, 1839 [not Tomicus Latreille, 1802; syn. of Ips DeGeer, 1775]. Comment: based on a misidentified type genus (see Alonso-Zarazaga and Lyal 2009); see also Tomicini Wood, 1978; an application will be submitted by MAAZ and CHCL to the Commission to suppress Tomicidae Shuckard, 1839 for the Principles of Priority and Homonymy (Art. 65.2.1).
Ipini Bedel, 1888: 386 [stem: Ip-]. Type genus: Ips DeGeer, 1775. Comment: an application needs to be submitted to the Commission to suppress Ipini Latreille, 1802 (based on the misidentified type genus Ips sensu Latreille, 1802) for the Principles of Priority and Homonymy (Art. 65.2.1) to conserve this name as valid.
Pityogenina Balachowsky, 1949: 244 [stem: Pityogen-]. Type genus: Pityogenes Bedel, 1888.
Tribe
Micracidini
LeConte, 1876
Micracides J. L. LeConte, 1876: 367 [stem: Micracid-]. Type genus: Micracis J. L. LeConte, 1868. Comment: incorrect original stem formation, not in prevailing usage; stem corrected by Alonso-Zarazaga and Lyal (2009: 81).
Hylocuridae Eichhoff, 1878: 298 [stem: Hylocur-]. Type genus: Hylocurus Eichhoff, 1872.
Tribe
Phloeosinini
Nüsslin, 1912
Phloeosinides Nüsslin, 1912: 289 [stem: Phloeosin-]. Type genus: Phloeosinus Chapuis, 1869 [placed on the Official List of Generic Names in Zoology (ICZN 1981a)].
*Dendrosinini Nunberg, 1967: 314 [stem: Dendrosin-]. Type genus: Dendrosinus Chapuis, 1869. Comment: unavailable (Art. 13.1): proposed after 1930 without description or bibliographic reference to such a description.
Tribe
Phloeotribini
Chapuis, 1869
Phloeotribidae Chapuis, 1869: 42 [stem: Phloeotrib-]. Type genus: Phloeotribus Latreille, 1797 [placed on the Official List of Generic Names in Zoology (ICZN 1979a)].
Phthorophloeides Nüsslin, 1912: 289 [stem: Phthorophloe-]. Type genus: Phthorophloeus Rey, 1885.
Tribe
Phrixosomatini
Wood, 1978
Phrixosomini Wood, 1978: 118, in key [stem: Phrixosomat-]. Type genus: Phrixosoma Blandford, 1897. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Alonso-Zarazaga and Lyal (2009: 91).
Tribe
Polygraphini
Chapuis, 1869
Polygraphidae Chapuis, 1869: 48 [stem: Polygraph-]. Type genus: Polygraphus Erichson, 1836.
Carphoborinae Nüsslin, 1911: 376 [stem: Carphobor-]. Type genus: Carphoborus Eichhoff, 1864.
Tribe
Premnobiini
Browne, 1962
Premnobiini Browne, 1962: 80 [stem: Premnobi-]. Type genus: Premnobius Eichhoff, 1878. Comment: family-group name available (Art. 13.1.2): description by indication (distinguishing characters given in Browne (1961)).
Tribe
Scolytini
Latreille, 1804
Scolitarii Latreille, 1804c: 156 [stem: Scolyt-]. Type genus: Scolytus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1963b)]. Comment: name placed on the Official List of Family-Group Names in Zoology (ICZN 1963b, as Scolytidae Westwood, 1838); incorrect original stem formation, not in prevailing usage.
Eccoptogastridae Gistel, 1848: [6] [stem: Eccoptogastr-]. Type genus: Ekkoptogaster Herbst, 1793 [as Eccoptogaster, incorrect subsequent spelling of type genus name by Gyllenhal (1813), not in prevailing usage; Ekkoptogaster Herbst, 1793 and Eccoptogaster Erichson, 1836 placed on the Official Index of Rejected and Invalid Genus-Group Names in Zoology (ICZN 1963b); syn. of Scolytus Geoffroy, 1762]. Comment: permanently invalid (Art. 39): based on suppressed type genus; Eccoptogasterinae Trédl, 1907 and Eccoptogasterini Reitter, 1906 placed on the Index of Rejected and Invalid Family-Group names in Zoology (ICZN 1963b).
Camptocérides Lacordaire, 1865: 366 [stem: Camptocer-]. Type genus: Camptocerus Dejean, 1821. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Chapuis (1869: 49, as Camptoceridae).
Minulini Reitter, 1913c: 12 [stem: Minul-]. Type genus: Minulus Eggers, 1912.
Tribe
Scolytoplatypodini
Blandford, 1893
Scolytoplatypini Blandford, 1893: 428 [stem: Scolytoplatypod-]. Type genus: Scolytoplatypus C. Schaufuss, 1891. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Blanford (1895: 84).
Taeniocerini Blandford, 1893: 428 [stem: Taeniocer-]. Type genus: Taeniocerus Blandford, 1893 [preoccupied genus name, not Taeniocerus Kaup, 1871 [Coleoptera: Passalidae]; syn. of Scolytoplatypus C. Schaufuss, 1891]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Spongocerinae Hagedorn, 1909: 163 [stem: Spongocer-]. Type genus: Spongocerus Blandford, 1893.
Tribe
Xyleborini
LeConte, 1876
Xylebori J. L. LeConte, 1876: 358 [stem: Xylebor-]. Type genus: Xyleborus Eichhoff, 1864 [placed on the Official List of Generic Names in Zoology (ICZN 1968a)].
Webbinae Hopkins, 1915: 224, in key [stem: Webbi-]. Type genus: Webbia Hopkins, 1915. Comment: incorrect original stem formation, not in prevailing usage.
*Eccoptopterina Browne, 1961: 49 [stem: Eccoptopter-]. Type genus: Eccoptopterus Motschulsky, 1863. Comment: unavailable (Art. 13.1): proposed after 1930 without description or bibliographic reference to such a description.
Tribe
Xyloctonini
Eichhoff, 1878
Xyloctonidae Eichhoff, 1878: 171 [stem: Xylocton-]. Type genus: Xyloctonus Eichhoff, 1872.
Tribe
Xyloterini
LeConte, 1876
Xyloteri J. L. LeConte, 1876: 356 [stem: Xyloter-]. Type genus: Xyloterus Erichson, 1836.
*Trypodendrinae Trédl, 1907: 70 [stem: Trypodendr-]. Type genus: Trypodendron Stephens, 1830. Comment: family-group name unavailable (Art. 11.7.1.1): not based on a genus used as valid at the time.
Trypodendrina Nunberg, 1954: 16 [stem: Trypodendr-]. Type genus: Trypodendron Stephens, 1830.
Subfamily
Platypodinae
Shuckard, 1839
Platypodidae Shuckard, 1839b: 64 [stem: Platypod-]. Type genus: Platypus Herbst, 1793.
Tribe
Mecopelmini
Thompson, 1992
*Mecopelmini Wood, 1966: 45 [stem: Mecopelm-]. Type genus: Mecopelmus Blackman, 1944. Comment: unavailable (Art. 13.1): proposed after 1930 without description or bibliographic reference to such a description.
Mecopelminae Thompson, 1992: 873, in key [stem: Mecopelm-]. Type genus: Mecopelmus Blackman, 1944.
Tribe
Platypodini
Shuckard, 1839
Platypodidae Shuckard, 1839b: 64 [stem: Platypod-]. Type genus: Platypus Herbst, 1793.
Crossotarsariae H. Strohmeyer, 1914: 19 [stem: Crossotars-]. Type genus: Crossotarsus Chapuis, 1865.
Tribe
Schedlariini
Wood and Bright, 1992
Chapuisiides Blandford, 1895: 89 [stem: Chapuisi-]. Type genus: Chapuisia Dugès, 1886 [preoccupied genus name, not Chapuisia Duvivier, 1885 [Coleoptera: Chrysomelidae]; syn. of Schedlarius Wood, 1957]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Schedlarini Wood and Bright, 1992: 1087 [stem: Schedlari-]. Type genus: Schedlarius Wood, 1957. Comment: replacement name for Chapuisiides Blandford, 1895 because of the homonymy of the type genus; incorrect original stem formation, not in prevailing usage; correction of stem and transfer to Platypodinae by Alonso-Zarazaga and Lyal (2009: 17).
Tribe
Tesserocerini
Strohmeyer, 1914
Tesserocerinae H. Strohmeyer, 1914: 19 [stem: Tesserocer-]. Type genus: Tesserocerus Saunders, 1836. Comment: precedence (Tesserocerini H. Strohmeyer, 1914 vs Diapodini Strohmeyer, 1914) given to taxon originally proposed at the higher rank (Art. 24.1).
Subtribe
Diapodina
Strohmeyer, 1914
Diapodariae H. Strohmeyer, 1914: 19 [stem: Diapod-]. Type genus: Diapus Chapuis, 1865.
Genyocerinae Hopkins, 1915: 225, in key [stem: Genyocer-]. Type genus: Genyocerus Motschulsky, 1858.
Subtribe
Tesserocerina
Strohmeyer, 1914
Tesserocerinae H. Strohmeyer, 1914: 19 [stem: Tesserocer-]. Type genus: Tesserocerus Saunders, 1836. Comment: precedence (Tesserocerina H. Strohmeyer, 1914 vs Cenocephalina Strohmeyer, 1914) given to taxon originally proposed at the higher rank (Art. 24.1).
Symmerariae H. Strohmeyer, 1914: 19 [stem: Symmer-]. Type genus: Symmerus Chapuis, 1865 [preoccupied genus name, not Symmerus Walker, 1848 [Diptera]; syn. of Chaetastus Nunberg, 1953]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.
Cenocephalariae H. Strohmeyer, 1914: 19 [stem: Cenocephal-]. Type genus: Cenocephalus Chapuis, 1865.
Platytarsilidae Schedl, 1939: 387 [stem: Platytarsul-]. Type genus: Platytarsulus Schedl, 1935. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Wood and Bright (1992: 1084).
Periomatini Schedl, 1939: 397, in key [stem: Periommat-]. Type genus: Periommatus Chapuis, 1865 [as Periomatus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage; correction of stem by Schedl (1972: 249).
Platypicerinae Nunberg, 1953: 44 [stem: Platypicer-]. Type genus: Platypicerus Nunberg, 1953.
Coleoptera
incertae sedis
Homoeoplastidae Gistel, 1856a: 360. Type genus: Homoeoplastus Gistel, 1856 [Gistel includes three species in his new genus, none of which are described in his work nor attributed to authors as having been described previously; we therefore consider Homoeoplastus unavailable]. Comment: family-group name unavailable (Art. 11.7.1.1): not based on an available genus name.
Plocasteidae Gistel, 1856a: 365. Type genus: Plocastes Gistel, 1856 [the only species originally included in Plocastes is scaraboides; Gistel does not mention the author of the species name, nor does he describe the taxon, we therefore consider Plocastes as unavailable]. Comment: family-group name unavailable (Art. 11.7.1.1): not based on an available genus name.
Serratopalpidae Gistel, 1856a: 384. Type genus: Serratopalpus Gistel, 1856 [Serratopalpus helwigii was the only species originally included in Serratopalpus; this new species was not described in Gistel’s original paper therefore the genus name is considered unavailable]. Comment: family-group name unavailable (Art. 11.7.1.1): not based on an available genus name.