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We synthesize data on all known extant and fossil Coleoptera family-group names for the first time. A catalogue of 4887 family-group names (124 fossil, 4763 extant) based on 4707 distinct genera in Coleoptera is given. A total of 4492 names are available, 183 of which are permanently invalid because they are based on a preoccupied or a suppressed type genus. Names are listed in a classification framework. We recognize as valid 24 superfamilies, 211 families, 541 subfamilies, 1663 tribes and 740 subtribes. For each name, the original spelling, author, year of publication, page number, correct stem and type genus are included. The original spelling and availability of each name were checked from primary literature. A list of necessary changes due to Priority and Homonymy problems, and actions taken, is given. Current usage of names was conserved, whenever possible, to promote stability of the classification.
New synonymies (family-group names followed by genus-group names): Agronomina Gistel, 1848 syn. nov. of Amarina Zimmermann, 1832 (Carabidae), Hylepnigalioini Gistel, 1856 syn. nov. of Melandryini Leach, 1815 (Melandryidae), Polycystophoridae Gistel, 1856 syn. nov. of Malachiinae Fleming, 1821 (Melyridae), Sclerasteinae Gistel, 1856 syn. nov. of Ptilininae Shuckard, 1839 (Ptinidae), Phloeonomini Ádám, 2001 syn. nov. of Omaliini MacLeay, 1825 (Staphylinidae), Sepedophilini Ádám, 2001 syn. nov. of Tachyporini MacLeay, 1825 (Staphylinidae), Phibalini Gistel, 1856 syn. nov. of Cteniopodini Solier, 1835 (Tenebrionidae); Agronoma Gistel 1848 (type species Carabus familiaris Duftschmid, 1812, designated herein) syn. nov. of Amara Bonelli, 1810 (Carabidae), Hylepnigalio Gistel, 1856 (type species Chrysomela caraboides Linnaeus, 1760, by monotypy) syn. nov. of Melandrya Fabricius, 1801 (Melandryidae), Polycystophorus Gistel, 1856 (type species Cantharis aeneus Linnaeus, 1758, designated herein) syn. nov. of Malachius Fabricius, 1775 (Melyridae), Sclerastes Gistel, 1856 (type species Ptilinus costatus Gyllenhal, 1827, designated herein) syn. nov. of Ptilinus Geoffroy, 1762 (Ptinidae), Paniscus Gistel, 1848 (type species Scarabaeus fasciatus Linnaeus, 1758, designated herein) syn. nov. of Trichius Fabricius, 1775 (Scarabaeidae), Phibalus Gistel, 1856 (type species Chrysomela pubescens Linnaeus, 1758, by monotypy) syn. nov. of Omophlus Dejean, 1834 (Tenebrionidae). The following new replacement name is proposed: Gompeliina Bouchard, 2011 nom. nov. for Olotelina Báguena Corella, 1948 (Aderidae).
Reversal of Precedence (Article 23.9) is used to conserve usage of the following names (family-group names followed by genus-group names): Perigonini Horn, 1881 nom. protectum over Trechicini Bates, 1873 nom. oblitum (Carabidae), Anisodactylina Lacordaire, 1854 nom. protectum over Eurytrichina LeConte, 1848 nom. oblitum (Carabidae), Smicronychini Seidlitz, 1891 nom. protectum over Desmorini LeConte, 1876 nom. oblitum (Curculionidae), Bagoinae Thomson, 1859 nom. protectum over Lyprinae Gistel 1848 nom. oblitum (Curculionidae), Aterpina Lacordaire, 1863 nom. protectum over Heliomenina Gistel, 1848 nom. oblitum (Curculionidae), Naupactini Gistel, 1848 nom. protectum over Iphiini Schönherr, 1823 nom. oblitum (Curculionidae), Cleonini Schönherr, 1826 nom. protectum over Geomorini Schönherr, 1823 nom. oblitum (Curculionidae), Magdalidini Pascoe, 1870 nom. protectum over Scardamyctini Gistel, 1848 nom. oblitum (Curculionidae), Agrypninae/-ini Candèze, 1857 nom. protecta over Adelocerinae/-ini Gistel, 1848 nom. oblita and Pangaurinae/-ini Gistel, 1856 nom. oblita (Elateridae), Prosternini Gistel, 1856 nom. protectum over Diacanthini Gistel, 1848 nom. oblitum (Elateridae), Calopodinae Costa, 1852 nom. protectum over Sparedrinae Gistel, 1848 nom. oblitum (Oedemeridae), Adesmiini Lacordaire, 1859 nom. protectum over Macropodini Agassiz, 1846 nom. oblitum (Tenebrionidae), Bolitophagini Kirby, 1837 nom. protectum over Eledonini Billberg, 1820 nom. oblitum (Tenebrionidae), Throscidae Laporte, 1840 nom. protectum over Stereolidae Rafinesque, 1815 nom. oblitum (Throscidae) and Lophocaterini Crowson, 1964 over Lycoptini Casey, 1890 nom. oblitum (Trogossitidae); Monotoma Herbst, 1799 nom. protectum over Monotoma Panzer, 1792 nom. oblitum (Monotomidae); Pediacus Shuckard, 1839 nom. protectum over Biophloeus Dejean, 1835 nom. oblitum (Cucujidae), Pachypus Dejean, 1821 nom. protectum over Pachypus Billberg, 1820 nom. oblitum (Scarabaeidae), Sparrmannia Laporte, 1840 nom. protectum over Leocaeta Dejean, 1833 nom. oblitum and Cephalotrichia Hope, 1837 nom. oblitum (Scarabaeidae).
Beetles, nomenclature, classification, world fauna, family-group names, type genera, stem
Zoological nomenclature affects the work of all zoologists, yet only a minuscule fraction of one percent of [the craziest] zoologists deal directly with problems associated with scientific names of animals (
The accurate use of scientific names in zoology is necessary to minimize nomenclatural instability and allow the maximal retrieval of scientific information from the ever increasing body of literature. Akin to other scientific names such as species- and genus-group names, family-group names are extremely important in the exchange of information about the world we live in. Rules about how to treat family-group names, as determined by the International Commission on Zoological Nomenclature [henceforth the Commission], have been an integral part of zoological nomenclature since the early 1900s (for a historical review of family-group names refer to
The first family-group names based on the stem of their type genus appeared in zoological literature in the early 19th Century (see
The last twenty five years has seen an increase in the number of studies on family-group names of entire groups in zoology (e.g.,
Given the importance of family-group names in the scientific literature, it is rather surprising that still many authors do not cite the author or year of publication of those names correctly, or at all. According to Recommendation 51A of the Code (
Coleoptera are currently the most species-rich group of organisms on this planet with approximately 360 000 described species (
The nomenclature of family-group names in Coleoptera did not receive much attention until the treatment of Geadephaga names by
The vast body of scientific literature dealing with beetles has been a deterrent to producing a complete review of all Coleoptera family-group names in the past. This publication was only made possible by the collaboration of several coleopterists.
The specific objectives of this study are to: 1) establish the first comprehensive list of all Coleoptera family-group names with information on type genus, author(s), year of publication and complete bibliographical references; 2) assess the availability and validity of each name using rules laid out in the most recent Code of Zoological Nomenclature (
This publication should be seen as the starting point of what could eventually serve as the basis for a submission of Part of a List of Available names in Zoology (Art. 79). With this goal in mind, we ask all coleopterists to either send us, or to publish, corrections or differences of opinions in the months and years to come. It is our intention to update our list of family-group names as new data are published and to provide a second edition of this work in approximately five years.
Methods Criteria of availabilityDecisions about the availability and validity of each Coleoptera family-group name in our catalogue were made following the process outline in Figure 1. First we established whether the name was available or unavailable based on the criteria of availability summarized below. If a name first proposed on a particular type genus was determined to be unavailable then searches were conducted to establish if a family-group name based on the same type genus was made available subsequently. If so then this available name was entered in our catalogue. From the pool of available names, we removed those that are permanently invalid (Art. 39). All the remaining available names were then separated into those that are valid based on the classification used here (at any rank from subtribe to superfamily) and those that are invalid (i.e., synonyms).
Figure 1. Overview of the process used to determine the availability and validity of family-group names in Coleoptera (modified from Bouchet and Rocroi 2005). The number of names for each category is given in parentheses.
In order to be available, a family-group name proposed before 1931 must to be a scientific name (i.e., in latinized form) in the nominative plural based on the stem of an available genus name then used as valid in the new suprageneric taxon (Art. 11.7).
In addition to the criteria of availability mentioned above, new family-group names proposed between 1931 and 1999 had to be described in words, or be associated with a bibliographic reference to such a description, in order to be considered available (Arts 13, 15). In works published after 1930 which contained more than one use of a new family-group name, we selected a page where a description was clearly associated with the new taxon for the catalogue. It should be noted that replacement names proposed in that time period are available without description (Art. 13.1.3). Additionally, “a family-group name first published after 1930 and before 1961 which does not satisfy the provisions of Article 13.1 is available from its original publication only if it was used as valid before 2000, and also was not rejected by an author who, after 1960 and before 2000, expressly applied Article 13 of the then current editions of the Code” (Art. 13.2.1). Names proposed between 1931 and 1960 can also be considered available if the description of the new family-group name and a single new genus-group name is combined (Art. 13.5). Finally, family-group names proposed since 2000 have to be explicitly indicated as new and the name of the type genus has to be clearly cited in order to be available (Art. 16). Based on our interpretation of Article 16.2, any new family-group name proposed after 1999 in a paper in which the name of the type genus is clearly cited in connection with it (although maybe not explicitly with a formula such as “Type genus = Aus Doe, 2010”) is available.
One of the most difficult tasks while working on this review was to decide on the most consistent and objective way to apply Article 11.7.2, which deals with the availability of names that were originally proposed in a vernacular form. Vernacular names are generally not treated as scientific names in zoological nomenclature (see Recommendation 11A). However, the International Commission on Zoological Nomenclature has made a single, but rather restricted, exception that applies to family-group names. According to the Code, a vernacular name “…is available with its original author and date only if it has been latinized by later authors and has been generally accepted as valid by authors interested in the group concerned and as dating from that first publication in vernacular form” (Art. 11.7.2).
We have interpreted names proposed in the following languages to be vernacular: all German names with the suffix “–en”, all Spanish names with the suffix “–os” or “–as” and all French names with the suffix “–iens”. The most important issue was to determine the correct status of names originally proposed with the suffix “–es.” We have used the principle that all family-group names proposed by non-French writers with the suffix “–es” were in fact latinized names with an ending that differs from those regulated by the Commission (these are mostly older names proposed before rules of zoological nomenclature became well-established). For each name with the suffix “–es” first proposed by French workers (e.g., Latreille, Lacordaire, Lameere) we went through the entire work containing those names and established if these authors consistently used either vernacular or latinized names in their work. French vernacular names often have accents in them (“é” or “è”) while latinized names do not. We have found that determining whether an author used vernacular or latinized names in a particular work was fairly straightforward (
As in
Article 29.2 includes a list of suffixes to be used for groups at the superfamily (-oidea), family (-idae), subfamily (-inae), tribe (-ini) and subtribe (-ina) names. The suffix of other categories is not regulated by the Code. The rank of supertribe (-itae) was used here for a small number of groups in which recent changes in classification required an additional rank between subfamily and tribe (e.g., Staphylinoidea).
The correct spelling of family-group names depends on the stem of its type genus (Art. 29.3). As pointed out by
Table 1. List of common Coleoptera generic suffixes with their associated family-group name endings (based partly on
Generic ending | Meaning | Derived family-group name ending | Type genus example | Family-group name example |
---|---|---|---|---|
-apion | pear (Greek) | -api- | Aspidapion | Aspidapiina |
-arthron | joint (Greek) | -arthr- | Decarthron | Decarthrina |
-aspis | shield (Greek) | -aspid- | Anaspis | Anaspidinae |
-baris | flat-bottomed boat (Greek) | -barid- | Baris | Baridini |
-chlamys | mantle (Greek) | -clamyd- | Spodochlamys | Spodochlamydini |
-celis | spot (Greek) | -celid- | Xiphoscelis | Xiphoscelidini |
-ceras | horn (Greek) | -cerat- | Megaceras | Megaceratini |
-cnema | shin or tibia (Greek) | -cnem- | Pachycnema | Pachycnemina |
-cupes | dainty (Latin) | -cuped- | Cupes | Cupedidae |
-dacne | bite (Greek) | -dacn- | Dacne | Dacnini |
-deres | neck, throat (Greek) | -der- | Aglycyderes | Aglycyderini |
-derma | skin (Greek) | -dermat- | Cryptoderma | Cryptodermatinae |
-dytes | diver (Greek) | -dyt- | Aspidytes | Aspidytidae |
-genys | jaw (Greek) | -geny- | Chaetogenys | Chaetogenyini |
-hospes | guest (Latin) | -hospit- | Termitohospes | Termitohospitini |
-ides | similar to, derived from (Greek) | -id- | Anaides | Anaidinae |
-ifer | carrier of (Latin) | -ifer- | Undulifer | Unduliferinae |
-iger | carrier of (Latin) | -iger- | Apoderiger | Apoderigerina |
-ites | like (Latin, Greek) | -it- | Aegialites | Aegialitinae |
-loma | edge or fringe (Greek) | -lomat- | Discoloma | Discolomatinae |
-macer | thin (Latin) | -macr- | Rhynchitomacer | Rhynchitomacrini |
-mycter | nose (Greek) | -mycter- | Eurymycter | Eurymycterini |
-odes | similar to (Greek) | -od- | Agyrtodes | Agyrtodini |
-odon | tooth (Greek) | -odont- | Pentodon | Pentodontini |
-oides | like (Greek) | -oid- | Acmaeoderoides | Acmaeoderoidina |
-omma | eye (Greek) | -ommat- | Omma | Ommatidae |
-onyx | yellow gem stone (Greek) | -onych- | Trichonyx | Trichonychini |
-ops | eye (Greek) | -op- | Achaenops | Achaenopina |
-opsis | appearance (Greek) | -opse- | Brachyceropsis | Brachyceropseini |
-otes | quality, nature (Greek) | -ot- | Agriotes | Agriotini |
-pholis | horny scale (Greek) | -pholid- | Trachypholis | Trachypholidini |
-pteryx | wing (Greek) | -pteryg- | Trichopteryx | Trichopterygini |
-pus | foot (Greek) | -pod- | Baripus | Baripodina |
-rhinus | snout (Greek) | -rhin- | Platyrhinus | Platyrhinini |
-rhynchus | snout (Greek) | -rhynch- | Doydirhynchus | Doydirhynchini |
-rhipis | fan (Greek) | -rhipid- | Xenorhipis | Xenorhipidini |
-soma | body (Greek) | -somat- | Platysoma | Platysomatini |
-stoma | mouth (Greek) | -stomat- | Stenostoma | Stenostomatini |
-teles | perfect (Greek) | -tel- | Abroteles | Abrotelina |
-termes | wood-worm (Latin) | -termit- | Philotermes | Philotermitini |
-thorax | chest (Greek) | -thorac- | Mecyclothorax | Mecyclothoracini |
-trox | gnawer (Greek) | -trog- | Trox | Trogidae |
-trupes | borer (Greek) | -trup- | Ceratotrupes | Ceratotrupini |
-typus | shape (Greek) | -typ- | Amarotypus | Amarotypini |
It should be noted that if a family-group name was not formed in accordance with Art. 29.3 but its original spelling is in prevailing usage then the current spelling is to be maintained (Art. 29.5). We have conserved the spelling of several family-group names currently used as valid however we did not do so for names that are listed as synonyms. For names based on incorrect stems proposed after 1999, we have considered that prevailing usage cannot be used to conserve the original spellings because too few references using these names could be found. We have therefore corrected the stems of such names unless the name of the type genus was an arbitrary combination of letters (Art. 29.4).
As stated in Art. 35.4.1 “A family-group name based upon an unjustified emendation … or an incorrect spelling of the name of the type genus must be corrected, unless it is preserved under Article 29.5 or unless the spelling of the genus-group name used to form the family-group name is preserved under Articles 33.2.3.1 or 33.3.1.” When an unjustified emendation or an incorrect subsequent spelling of the type genus is in prevailing usage and is attributed to the original author and date (Art. 33.2.3.1; 33.3.1), the correct spelling of the type genus is that in current usage.
In the glossary of the Code, a name in prevailing usage is defined as a “name which is adopted by at least a substantial majority of the most recent authors concerned with the relevant taxon, irrespective of how long ago their work was published”. The unfortunate subjectivity in this definition, as pointed out by
The Principle of Coordination (Art. 36.1) is, unfortunately, still overlooked by some authors. In some instances authors who propose new ranks for previously established suprageneric names are sometimes treated as the author of those names when in fact only a change of rank was presented. Based on the Code “A name established at any rank in the family-group is deemed to have been simultaneously established for nominal taxa at all ranks in the family-group; all these taxa have the same type genus, and their names are formed from the stem of the name of the type genus [Art. 29.3] with appropriate change of suffix [Art. 34.1]. The name has the same authorship and date at every rank.”
Principle of PriorityAs for species- and genus-group names, the oldest available name for a family-group taxon should be considered as valid (Art. 23). However four important exceptions need further discussion. Firstly, when a little-known family-group name was discovered to be older than a name currently used as valid for a particular taxon, we used the Reversal of Precedence to conserve usage of the younger name if the conditions of Art. 23.9.2 could be fulfilled. Younger names conserved using Reversal of Precedence are listed in Appendix 1. In some instances we could not fulfill all conditions of Art. 23.9.2 although we considered that using the newly discovered older name as valid would threaten stability or cause confusion. In those cases we maintained usage of the younger name as valid and either submitted an application to the Commission to conserve the younger name or made a recommendation that such submission should be submitted in the near future (Art. 23.9.3).
According to Art. 35.5 “If after 1999 a name in use for a family-group taxon (e.g., for a subfamily) is found to be older than a name in prevailing usage for a taxon at higher rank in the same family-group taxon (e.g., for the family within which the older name is the name of a subfamily) the older name is not to displace the younger name.” Indeed, we encountered a small number of cases in which the replacement of a name at the higher rank (e.g., Lymexyloidea Fleming, 1821) by the discovery of an older name for a taxon at a lower rank (e.g., Hylecoetoidea Germar, 1818) would not have served the stability of well-established names. In such cases usage of the younger name at the higher rank was conserved.
In cases where a family-group name was replaced before 1961 because of the synonymy of its type genus (e.g., Lepiceridae Hinton, 1936 instead of Cyathoceridae Sharp, 1882), the substitute name is to be maintained if it is in prevailing usage (Art. 40.2). In such cases, the valid family-group name retains its own author but takes the priority of the replaced name. Based on Recommendation 40A, we have cited those names with their original author and date, followed by the date of its priority enclosed in parentheses, as determined by Art. 40.2.1 (e.g., Lepiceridae Hinton, 1936 (1882)).
Lastly, if the stability and continuity of the meaning of a family-group name is threatened by the discovery that the type genus was originally misidentified, or that the type genus was based on a misidentified type species, or that an older type species of the type genus had been overlooked, then the case is to be referred to the Commission for a ruling (Art. 65.2.1). In cases where the oldest available name for a family-group taxon is based on misidentified type genus, or an altered concept of the type genus (e.g., see Scydmaenini Reitter, 1882), we have preferred to consider this name as invalid and to use instead the family-group name which is in prevailing usage (e.g., Cyrtoscydmini L. W. Schaufuss, 1889) until an application is submitted and a ruling is rendered by the Commission.
A list of problem cases based on the Principle of Priority, with comments on implementation of solutions or necessary actions to be taken in the future, is given in Appendix 2.
Principle of HomonymyBased on the Principle of Homonymy “when two or more names are homonyms, only the senior, as determined by the Principle of Priority…, may be used as the valid name…” (Art. 55.2). Here we report several instances in which family-group names in Coleoptera are identical to other family-group names in zoological nomenclature. These names are either based on identical type genera or on type genera that are similar but not identical. In the first instance the family-group name based on the preoccupied type genus is permanently invalid (Art. 39) but available.
When family-group names are homonyms because their type genera are similar but not identical, the case must be referred to the Commission for a ruling to remove homonymy (Art. 55.3.1). Such is the case with Adeliini Kirby, 1825 (type genus Adelium Kirby, 1819) and the hymenopteran name Adeliini Viereck, 1918 (type genus Adelius Haliday, 1833) which are both correctly formed from the stem of their type genus. More than thirty such cases were encountered during our research on Coleoptera family-group names.
Junior homonyms conserved using Reversal of Precedence are listed in Appendix 1. A list of problem cases based on the Principle of Homonymy, with comments on implementation of solutions or necessary actions to be taken in the future, is given in Appendix 3.
Submissions to the CommissionAs mentioned above, we have encountered several cases which require an application to the Commission because of problems with priority and/or homonymy. Some of these cases have been submitted recently (
Authorship of the three volumes on Coleoptera of the series Encyclopédie d’histoire naturelle is debated. The title pages on the volumes list Jean Charles Chenu as the author of the Encyclopédie d’histoire naturelle with the assistance of Eugène Desmarest for the Coleoptera section. However, the Société Entomologique de France recorded the livraisons received for 1851 in their Bulletin (p. cxxv) as “Encyclopédie d’histoire naturelle, ou traité complet de cette science, sous la direction de M. le docteur Chenu. Coléoptères, par M.E. Desmarest.” This was likely the title on the wrapper and clearly suggests that Desmarest was responsible alone for the three volume series on Coleoptera. This is also substained by Desmarest himself who in the Bulletin de la Société Entomologique de France for 1860 (p. lxiii) speaks of the three volumes series as being his work, published under the direction of Chenu. Based on the above fact, we credit the three-volume series to Desmarest alone (
The articles presented by Mulsant (and Rey) in the “Opuscules Entomologiques” are considered here to be reprints (as a compilation) of original publications in various Annales, mostly printed at Lyon, for the following reasons: In the dedication to the first volume Mulsant wrote “ces Opuscules, publiés déjà ça et là dans nos Recueils académiques”. The dates of the dedications in the Opuscules are always later than the dates on which they were presented to the various Sociétés, frequently in the following year. Although published at Paris, the Opuscules (see verso of title pages) were printed at Lyon by F. Dumoulin and others, the same printers which published the Mémoires de l’Académie... des Sciences, Belles-Lettres et Arts de Lyon, etc. Rey himself, in later publications such as the continuation of the “Histoire Naturelle des Coléoptères de France” (e.g., Annales de la Société Linnéenne de Lyon (N. S.) 32[1885]: 1-186 + [4], pls. 1-2 [see p. 106]) cited the Annales de la Société Linnéenne de Lyon article before the Opuscules version. E. A.
However, the first fascicles of the Histoire Naturelle series up to the “Pectinipèdes” were not reprinted in journals. It seems that most of the succeeding fascicles were submitted to the various Sociétés, were printed by Dumoulin, Barret, Pinier, etc. at Lyon, the dedications were then added and they were sent to Paris for separate publication with the same typesetting, and with changes only to the pagination and some titles. Since the issuing of the journal volumes was frequently delayed by the compilation of many papers (usually in the following year), the separate edition often appeared first, as shown by records of other journals and, in particular, the Bibliographie de la France. Therefore we have given date priority to the Histoire Naturelle versions in most instances, only those page numbers are cited with the scientific names in the catalogue unless evidence was found to the contrary, and the alternate versions are cited only in the bibliography, following each reference.
Format of the catalogue and conventions usedThis paper is organized into two main parts: 1) a synoptic classification containing only the names which we consider as valid herein and 2) a catalogue of all family-group names organized in the same order as in the synoptic classification. It should be noted that our publication is first and foremost a nomenclatural treatment of family-group names. We want to emphasize that the classification used here is not based on newly generated phylogenetic data. Furthermore, we do not necessarily endorse all parts of the classification presented, particularly those which were not based on phylogenetic approaches. For the most part, we follow currently accepted concepts in recent taxon-specific catalogues as well as comprehensive syntheses such as the Handbook of Zoology (
Only family-group names that are based on genera are included in our catalogue. For each family-group name the original spelling, author, year of publication, page number, correct stem and type genus are given. Complete data and comments regarding a particular family-group name are presented with the lowest-rank name when the same stem is used in more than one rank, since the same criteria apply in accordance with the Principle of Coordination (Art. 36.1). The correct stem to be used, which is given in square brackets (e.g., [stem: Uralocole-]), is especially important for synonyms, since the status of a name may be changed from synonym to valid in the future. Subsequent alternative spellings of family-group names (i.e., simple changes in suffix depending on change in rank) are not given here. Both available and unavailable names are listed together in the catalogue in order to enhance information retrieval. Unavailable names are preceded by an asterisk “*” and are identified as such in the “Comments” section along with the reason for this status. As is common practice in publications of the International Commission on Zoological Nomenclature, we use small capital letters for family-group names. Names in the catalogue are organized in a “phylogenetic” framework, as accepted in recent publications on the group, down to subfamily level. Valid tribes and subtribes are listed in alphabetical order within each subfamily. Synonyms are listed in chronological order under each valid taxon.
Known works of Johannes Gistel (who also published under the name Johannes Gistl and as G. Tilesius; see
Because of the importance of the Principle of Priority, we have tried to find the most accurate date of publication (given in square brackets in the References section) for works cited in the manuscript. The date of publication was determined either from the original publication itself, from the date of “stamps” when received in libraries of natural history institutions or from secondary literature sources. A list of natural history institutions and secondary literature sources is given at the beginning of the References section. Data on dates of publication were provided by AFN, PB, AED, MAAZ, CHCL, ABTS and YB. References for type genera are not included here to conserve space and because they are for the most part available in other recent publications (e.g.,
Names on the Official List of Family-Group Names in Zoology and type genera on the Official List of Generic Names in Zoology are included in our catalogue and are summarized in Appendix 4 and 5 respectively. Cases involving family-group names and/or their type genera that are awaiting a ruling by the Commission are summarized in Appendix 6.
Results Number of family-group namesA total of 4887 family-group names are included in our catalogue (see Figure 1). Of the names recorded, 4492 are available, of which 183 are permanently invalid because they are based on a preoccupied type genus or a genus which has been suppressed by the Commission. The majority of names were proposed in the suborder Polyphaga (4314) followed by Adephaga (531), Myxophaga (16), Archostemata (17) and Protocoleoptera (9). Within Adephaga the family Carabidae contains the highest number of names by far (441) followed by Dytiscidae (47). The five superfamilies with the highest number of names in Polyphaga are the Curculionoidea (862), Chrysomeloidea (794), Staphylinoidea (594), Tenebrionoidea (579) and Scarabaeoidea (436). Overall, the five families with the highest number of family-group names proposed are Curculionidae (555), Staphylinidae (493), Cerambycidae (468), Carabidae (441) and Tenebrionidae (323).
The number of Coleoptera family-group names that appeared in the 19th and 20th Centuries are almost identical (2331 and 2556 respectively) while the last decade saw an increase of 5% (246 names) in the total number of names in the literature (Figure 2). A large number of names proposed in the middle of the 19th Century are unavailable. Those names were generally proposed in vernacular form and were not made available subsequently (Art. 11.7.2). Unavailable names proposed after 1930 generally lacked a description or bibliographic reference to such a description (Arts 13, 15). Several names proposed in the last decade were either not explicitly introduced as new taxa or did not include necessary information about the type genus and are therefore unavailable (Art. 16).
Figure 2. Number of family-group names proposed in Coleoptera by decade. White bars = available names. Black bars = unavailable names.
A summary of the most significant contributions, in terms of the total number of names proposed by author, is shown in Table 2. Eighteen authors proposed 50 or more family-group names in Coleoptera. Lacordaire was the most prolific with 353 new names. Although the majority of Lacordaire’s names were proposed in vernacular form, a large proportion of them (90%) were subsequently made available and many are used as valid today.
Table 2. Summary of number of family-group names in Coleoptera by author. Only authors that have proposed fifty or more names are included. Authors listed in decreasing order by total number of names proposed. For each author we give the total number of name proposed, the percentage of those names that are available, the range of years in which those names were proposed and the number of families in which they proposed new family-group names in.
Rank | Author | Country of origin | Total family-group names | Percent available | Publication (year range) | Number of families |
---|---|---|---|---|---|---|
1 | Lacordaire, Jean Théodore | France | 353 | 90 | 1848–1872 | 34 |
2 | Gistel, Johannes von Nepomuk Franz Xaver | Germany | 135 | 99 | 1848–1856 | 42 |
3 | LeConte, John Lawrence | USA | 131 | 100 | 1847–1883 | 45 |
4 | Thomson, James | USA | 124 | 99 | 1857–1877 | 6 |
5 | Mulsant, Étienne | France | 122 | 76 | 1839–1880 | 25 |
6 | Chapuis, Félicien | France | 120 | 72 | 1869–1876 | 7 |
7 | Jeannel, René | France | 109 | 99 | 1910–1967 | 6 |
8 | Reitter, Edmund | Germany | 87 | 100 | 1875–1926 | 24 |
9 | Legalov, Andrei Aleksandrovich | Russia | 73 | 100 | 2001–2009 | 9 |
10 | Voss, Eduard | Germany | 66 | 97 | 1922–1972 | 6 |
11 | Burmeister, Hermann Carl Conrad | Germany | 66 | 98 | 1840–1873 | 2 |
12 | Casey, Thomas Lincoln | USA | 64 | 100 | 1884–1922 | 14 |
13 | Horn, George Henry | USA | 63 | 100 | 1867–1893 | 20 |
14 | Latreille, Pierre André | France | 62 | 98 | 1802–1834 | 30 |
15 | Rey, Claudius | France | 57 | 53 | 1853–1886 | 12 |
16 | Thomson, Carl Gustaf | Sweden | 55 | 100 | 1857–1867 | 27 |
17 | Blanchard, Charles Émile | France | 55 | 71 | 1845–1853 | 20 |
18 | Laporte, François Louis Nompar de Caumont (Comte Castelnau) | France | 51 | 100 | 1834–1840 | 21 |
The authors with the lowest percentage of available family-group names (Mulsant, Chapuis, Rey and Blanchard) originally proposed their names in vernacular form and a significant proportion of those names were not made available subsequently. All names proposed by LeConte, Reitter, Casey, Horn, Legalov, C. G. Thomson and Laporte are considered available.
Thirteen of the authors in Table 2 were active exclusively during the 19th Century while two authors, Reitter and Casey, published their works containing new family-group names in both the 19th and 20th Centuries. Two authors published family-group names exclusively during the 20th Century (Jeannel and Voss). Legalov is our only contemporary colleague in this list; he proposed more than 50 new family-group names in the last decade. It should be noted that the recent proliferation of new scientific names proposed by Legalov is treated by some authors as “extreme splitting” (
The broad taxonomic interests and expertise of Lacordaire, Gistel, LeConte, Mulsant, Latreille and C. G. Thomson are exemplified by the fact that they introduced new family-group names in 25 or more families. This was done either in catalogues (e.g., Gistel, C. G. Thomson) and/or in large scale taxonomic treatments (e.g., Lacordaire, LeConte, Latreille, Mulsant). On the other hand, the taxonomic expertise of J. Thomson, Chapuis, Jeannel, Voss, Burmeister and Legalov is much more targeted towards a small number of families (fewer than 10).
New nomenclatural actsThe following synonymies are recorded here for the first time: Agronomina Gistel, 1848 syn. nov. of Amarina Zimmermann, 1832 (Carabidae), Hylepnigalioini Gistel, 1856 syn. nov. of Melandryini Leach, 1815 (Melandryidae), Polycystophoridae Gistel, 1856 syn. nov. of Malachiinae Fleming, 1821 (Melyridae), Sclerasteinae Gistel, 1856 syn. nov. of Ptilininae Shuckard, 1839 (Ptinidae), Phloeonomini Ádám, 2001 syn. nov. of Omaliini MacLeay, 1825 (Staphylinidae), Sepedophilini Ádám, 2001 syn. nov. of Tachyporini MacLeay, 1825 (Staphylinidae), Phibalini Gistel, 1856 syn. nov. of Cteniopodini Solier, 1835 (Tenebrionidae); Agronoma Gistel 1848 (type species Carabus familiaris Duftschmid, 1812, designated herein) syn. nov. of Amara Bonelli, 1810 (Carabidae), Hylepnigalio Gistel, 1856 (type species Chrysomela caraboides Linnaeus, 1760, by monotypy) syn. nov. of Melandrya Fabricius, 1801 (Melandryidae), Polycystophorus Gistel, 1856 (type species Cantharis aeneus Linnaeus, 1758, designated herein) syn. nov. of Malachius Fabricius, 1775 (Melyridae), Sclerastes Gistel, 1856 (type species Ptilinus costatus Gyllenhal, 1827, designated herein) syn. nov. of Ptilinus Geoffroy, 1762 (Ptinidae), Paniscus Gistel, 1848 (type species Scarabaeus fasciatus Linnaeus, 1758, designated herein) syn. nov. of Trichius Fabricius, 1775 (Scarabaeidae), Phibalus Gistel, 1856 (type species Chrysomela pubescens Linnaeus, 1758, by monotypy) syn. nov. of Omophlus Dejean, 1834 (Tenebrionidae). The following replacement name is used here for the first time: Gompeliina Bouchard, 2011 nom. nov. for Olotelina Báguena Corella, 1948 (Aderidae).
Reversal of Precedence (Article 23.9) is used to conserve usage of the following names (family-group names followed by genus-group names): Perigonini Horn, 1881 nom. protectum over Trechicini Bates, 1873 nom. oblitum (Carabidae), Anisodactylina Lacordaire, 1854 nom. protectum over Eurytrichina LeConte, 1848 nom. oblitum (Carabidae), Smicronychini Seidlitz, 1891 nom. protectum over Desmorini LeConte, 1876 nom. oblitum (Curculionidae), Bagoinae Thomson, 1859 nom. protectum over Lyprinae Gistel 1848 nom. oblitum (Curculionidae), Aterpina Lacordaire, 1863 nom. protectum over Heliomenina Gistel, 1848 nom. oblitum (Curculionidae), Naupactini Gistel, 1848 nom. protectum over Iphiini Schönherr, 1823 nom. oblitum (Curculionidae), Cleonini Schönherr, 1826 nom. protectum over Geomorini Schönherr, 1823 nom. oblitum (Curculionidae), Magdalidini Pascoe, 1870 nom. protectum over Scardamyctini Gistel, 1848 nom. oblitum (Curculionidae), Agrypninae/-ini Candèze, 1857 nom. protecta over Adelocerinae/-ini Gistel, 1848 nom. oblita and Pangaurinae/-ini Gistel, 1856 nom. oblita (Elateridae), Prosternini Gistel, 1856 nom. protectum over Diacanthini Gistel, 1848 nom. oblitum (Elateridae), Calopodinae Costa, 1852 nom. protectum over Sparedrinae Gistel, 1848 nom. oblitum (Oedemeridae), Adesmiini Lacordaire, 1859 nom. protectum over Macropodini Agassiz, 1846 nom. oblitum (Tenebrionidae), Bolitophagini Kirby, 1837 nom. protectum over Eledonini Billberg, 1820 nom. oblitum (Tenebrionidae), Throscidae Laporte, 1840 nom. protectum over Stereolidae Rafinesque, 1815 nom. oblitum (Throscidae) and Lophocaterini Crowson, 1964 over Lycoptini Casey, 1890 nom. oblitum (Trogossitidae); Monotoma Herbst, 1799 nom. protectum over Monotoma Panzer, 1792 nom. oblitum (Monotomidae); Pediacus Shuckard, 1839 nom. protectum over Biophloeus Dejean, 1835 nom. oblitum (Cucujidae), Pachypus Dejean, 1821 nom. protectum over Pachypus Billberg, 1820 nom. oblitum (Scarabaeidae), Sparrmannia Laporte, 1840 nom. protectum over Leocaeta Dejean, 1833 nom. oblitum and Cephalotrichia Hope, 1837 nom. oblitum (Scarabaeidae).
Synoptic classification of the world ColeopteraOrder COLEOPTERA
†Suborder PROTOCOLEOPTERA
†Superfamily Tshekardocoleoidea Rohdendorf, 1944
†Family Tshekardocoleidae Rohdendorf, 1944
†Family Labradorocoleidae Ponomarenko, 1969
†Family Oborocoleidae Kukalová, 1969
†Superfamily Permocupedoidea Martynov, 1933
†Family Permocupedidae Martynov, 1933
†Family Taldycupedidae Rohdendorf, 1961
†Superfamily Permosynoidea Tillyard, 1924
†Family Ademosynidae Ponomarenko, 1968
†Family Permosynidae Tillyard, 1924
Suborder ARCHOSTEMATA
Family Crowsoniellidae Iablokoff-Khnzorian, 1983
Family Cupedidae Laporte, 1836
Subfamily Priacminae Crowson, 1962
†Subfamily Mesocupedinae Ponomarenko, 1969
Subfamily Cupedinae Laporte, 1836
Family Micromalthidae Barber, 1913
Family Ommatidae Sharp and Muir, 1912
†Subfamily Brochocoleinae Hong, 1982
Subfamily Tetraphalerinae Crowson, 1962
Subfamily Ommatinae Sharp and Muir, 1912
†Tribe Lithocupedini Ponomarenko, 1969
†Tribe Notocupedini Ponomarenko, 1966
Tribe Ommatini Sharp and Muir, 1912
Family Jurodidae Ponomarenko, 1985
†Family Triadocupedidae Ponomarenko, 1966
†Family Magnocoleidae Hong, 1998
†Family Obrieniidae Zherikhin and Gratshev, 1994
†Subfamily Kararhynchinae Zherikhin and Gratshev, 1994
†Tribe Kararhynchini Zherikhin and Gratshev, 1994
†Tribe Kenderlykaini Legalov, 2009
†Subfamily Obrieniinae Zherikhin and Gratshev, 1994
Suborder MYXOPHAGA
†Superfamily Asiocoleoidea Rohdendorf, 1961
†Family Asiocoleidae Rohdendorf, 1961
†Family Tricoleidae Ponomarenko, 1969
†Superfamily Rhombocoleoidea Rohdendorf, 1961
†Family Rhombocoleidae Rohdendorf, 1961
†Superfamily Schizophoroidea Ponomarenko, 1968
†Family Schizophoridae Ponomarenko, 1968
†Family Catiniidae Ponomarenko, 1968
†Family Schizocoleidae Rohdendorf, 1961
Superfamily Lepiceroidea Hinton, 1936 (1882)
Family Lepiceridae Hinton, 1936 (1882)
Superfamily Sphaeriusoidea Erichson, 1845
Family Torridincolidae Steffan, 1964
Subfamily Torridincolinae Steffan, 1964
Subfamily Deleveinae Endrödy-Younga, 1997
Family Hydroscaphidae LeConte, 1874
Family Sphaeriusidae Erichson, 1845
Suborder ADEPHAGA
†Family Tritarsidae Hong, 2002
Family Gyrinidae Latreille, 1810
Subfamily Spanglerogyrinae Folkerts, 1979
Subfamily Gyrininae Latreille, 1810
Tribe Enhydrini Régimbart, 1882
Subtribe Dineutina Desmarest, 1851
Subtribe Enhydrina Régimbart, 1882
Tribe Gyrinini Latreille, 1810
Subtribe Gyrinina Latreille, 1810
Subtribe Heterogyrina Brinck, 1956
Tribe Orectochilini Régimbart, 1882
Family Trachypachidae Thomson, 1857
†Subfamily Eodromeinae Ponomarenko, 1977
Subfamily Trachypachinae Thomson, 1857
Family Rhysodidae Laporte, 1840
Tribe Leoglymmiini Bell and Bell, 1978
Tribe Dhysorini Bell and Bell, 1978
Tribe Medisorini Bell and Bell, 1987
Tribe Rhysodini Laporte, 1840
Tribe Clinidiini Bell and Bell, 1978
Tribe Omoglymmiini Bell and Bell, 1978
Tribe Sloanoglymmiini Bell and Bell, 1991
Family Carabidae Latreille, 1802
†Subfamily Protorabinae Ponomarenko, 1977
†Subfamily Conjunctiinae Ponomarenko, 1977
Subfamily Nebriinae Laporte, 1834
Tribe Nebriini Laporte, 1834
Tribe Notiokasiini Kavanaugh and Nègre, 1983
Tribe Notiophilini Motschulsky, 1850
Tribe Opisthiini Dupuis, 1912
Tribe Pelophilini Kavanaugh, 1996
Subfamily Cicindinae Csiki, 1927
Subfamily Cicindelinae Latreille, 1802
Tribe Amblycheilini Csiki, 1903
Tribe Cicindelini Latreille, 1802
Subtribe Apteroessina Rivalier, 1971
Subtribe Cicindelina Latreille, 1802
Subtribe Dromicina Thomson, 1859
Subtribe Iresiina Rivalier, 1971
Subtribe Theratina Horn, 1893
Tribe Collyridini Brullé, 1834
Subtribe Collyridina Brullé, 1834
Subtribe Tricondylina Naviaux, 1991
Tribe Ctenostomatini Laporte, 1834
Tribe Manticorini Laporte, 1834
Tribe Megacephalini Laporte, 1834
Subfamily Carabinae Latreille, 1802
Tribe Carabini Latreille, 1802
Tribe Ceroglossini Lapouge, 1927
Tribe Cychrini Perty, 1830
Tribe Pamborini Hope, 1838
Subfamily Loricerinae Bonelli, 1810
Subfamily Omophroninae Bonelli, 1810
Subfamily Elaphrinae Latreille, 1802
Subfamily Migadopinae Chaudoir, 1861
Tribe Amarotypini Erwin, 1985
Tribe Migadopini Chaudoir, 1861
Subtribe Aquilicina Moret, 2005
Subtribe Migadopina Chaudoir, 1861
Subfamily Hiletinae Schiødte, 1848
Subfamily Scaritinae Bonelli, 1810
Tribe Carenini MacLeay, 1887
Tribe Clivinini Rafinesque, 1815
Subtribe Ardistomina Putzeys, 1867
Subtribe Clivinina Rafinesque, 1815
Subtribe Forcipatorina Bänninger, 1938
Tribe Dalyatini Mateu, 2002
Tribe Dyschiriini Kolbe, 1880
†Tribe Palaeoaxinidiini McKay, 1991
Tribe Pasimachini Putzeys, 1867
Tribe Promecognathini LeConte, 1853
Tribe Salcediini Alluaud, 1930 (1929)
Subtribe Androzelmina Bell, 1998
Subtribe Salcediina Alluaud, 1930 (1929)
Subtribe Solenogenyina Bell, 1998
Tribe Scaritini Bonelli, 1810
Subtribe Acanthoscelina Csiki, 1927
Subtribe Corintascarina Basilewsky, 1973
Subtribe Dyscherina Basilewsky, 1973
Subtribe Ochyropina Basilewsky, 1973
Subtribe Oxylobina Andrewes, 1929
Subtribe Scapterina Putzeys, 1867
Subtribe Scaritina Bonelli, 1810
Subtribe Storthodontina Jeannel, 1946
Subfamily Broscinae Hope, 1838
Tribe Broscini Hope, 1838
Subtribe Axonyina Roig-Juñent, 2000
Subtribe Baripodina Jeannel, 1941
Subtribe Broscina Hope, 1838
Subtribe Creobiina Jeannel, 1941
Subtribe Nothobroscina Roig-Juñent, 2000
Subfamily Apotominae LeConte, 1853
Subfamily Siagoninae Bonelli, 1813
Tribe Enceladini Horn, 1881
Tribe Lupercini Lecordier, 1977
Tribe Siagonini Bonelli, 1813
Subfamily Melaeninae Csiki, 1933
Subfamily Gehringiinae Darlington, 1933
Tribe Gehringiini Darlington, 1933
Subtribe Gehringiina Darlington, 1933
Subtribe Helenaeina Deuve, 2007
Subfamily Trechinae Bonelli, 1810
Tribe Bembidiini Stephens, 1827
Subtribe Anillina Jeannel, 1937
Subtribe Bembidiina Stephens, 1827
Subtribe Tachyina Motschulsky, 1862
Subtribe Xystosomina Erwin, 1994
Tribe Horologionini Jeannel, 1949
Tribe Pogonini Laporte, 1834
Tribe Trechini Bonelli, 1810
Subtribe Aepina Fowler, 1887
Subtribe Cnidina Jeannel, 1958
Subtribe Perileptina Sloane, 1903
Subtribe Plocamotrechina Jeannel, 1960
Subtribe Trechina Bonelli, 1810
Subtribe Trechodina Jeannel, 1926
Tribe Zolini Sharp, 1886
Subtribe Chalteniina Roig-Juñent and Cicchino, 2001
Subtribe Sinozolina Deuve, 1997
Subtribe Zolina Sharp, 1886
Subfamily Patrobinae Kirby, 1837
Tribe Lissopogonini Zamotajlov, 2000
Tribe Patrobini Kirby, 1837
Subtribe Deltomerina Chaudoir, 1871
Subtribe Deltomerodina Zamotajlov, 2002
Subtribe Patrobina Kirby, 1837
Subtribe Platidiolina Zamotajlov and Lafer, 2001
Subfamily Psydrinae LeConte, 1853
Tribe Amblytelini Blackburn, 1892
Tribe Mecyclothoracini Jeannel, 1940
Tribe Meonini Sloane, 1898
Tribe Moriomorphini Sloane, 1890
Tribe Psydrini LeConte, 1853
Tribe Tropopterini Sloane, 1898
Subfamily Nototylinae Bänninger, 1927
Subfamily Paussinae Latreille, 1806
Tribe Metriini LeConte, 1853
Tribe Mystropomini Horn, 1881
Tribe Ozaenini Hope, 1838
Tribe Paussini Latreille, 1806
†Subtribe Arthropteritina Luna de Carvalho, 1961
Subtribe Carabidomemnina Wasmann, 1928
Subtribe Cerapterina Billberg, 1820
†Subtribe Eopaussina Luna de Carvalho, 1951
Subtribe Heteropaussina Janssens, 1950
Subtribe Homopterina Wasmann, 1920
Subtribe Paussina Latreille, 1806
Subtribe Pentaplatarthrina Jeannel, 1946
Tribe Protopaussini Gestro, 1892
Subfamily Brachininae Bonelli, 1810
Tribe Brachinini Bonelli, 1810
Subtribe Aptinina Gistel, 1848
Subtribe Brachinina Bonelli, 1810
Subtribe Mastacina Erwin, 1970
Subtribe Pheropsophina Jeannel, 1949
Tribe Crepidogastrini Jeannel, 1949
Subfamily Harpalinae Bonelli, 1810
Tribe Abacetini Chaudoir, 1873
Tribe Amorphomerini Sloane, 1923
Tribe Anthiini Bonelli, 1813
Tribe Atranini Horn, 1881
Tribe Bascanini Basilewsky, 1953
Tribe Calophaenini Jeannel, 1948
Tribe Catapieseini Bates, 1882
Tribe Chaetodactylini Tschitschérine, 1903
Tribe Chaetogenyini Emden, 1958
Tribe Chlaeniini Brullé, 1834
Subtribe Callistina Laporte, 1834
Subtribe Chlaeniina Brullé, 1834
Tribe Cnemalobini Germain, 1911
Tribe Cratocerini Lacordaire, 1854
Tribe Ctenodactylini Laporte, 1834
Tribe Cuneipectini Sloane, 1907
Tribe Cyclosomini Laporte, 1834
Subtribe Cyclosomina Laporte, 1834
Subtribe Masoreina Chaudoir, 1871
Tribe Dercylini Sloane, 1923
Tribe Drimostomatini Chaudoir, 1872
Tribe Dryptini Bonelli, 1810
Tribe Enoicini Basilewsky, 1985
Tribe Galeritini Kirby, 1825
Subtribe Galeritina Kirby, 1825
Subtribe Planetina Jedlička, 1941
Tribe Geobaenini Péringuey, 1896
Tribe Ginemini Ball and Shpeley, 2002
Tribe Glyptini Horn, 1881
Tribe Graphipterini Latreille, 1802
Tribe Harpalini Bonelli, 1810
Subtribe Anisodactylina Lacordaire, 1854 nomen protectum
Subtribe Harpalina Bonelli, 1810
Subtribe Pelmatellina Bates, 1882
Subtribe Stenolophina Kirby, 1837
Tribe Helluonini Hope, 1838
Subtribe Helluonina Hope, 1838
Subtribe Omphrina Jedlička, 1941
Tribe Hexagoniini Horn, 1881 (1834)
Tribe Idiomorphini Bates, 1891
Tribe Lachnophorini LeConte, 1853
Subtribe Lachnophorina LeConte, 1853
Subtribe Selinina Jeannel, 1948
Tribe Lebiini Bonelli, 1810
Subtribe Actenonycina Bates, 1871
Subtribe Agrina Kirby, 1837
Subtribe Apenina Ball, 1983
Subtribe Calleidina Chaudoir, 1873
Subtribe Celaenephina Habu, 1982
Subtribe Cymindidina Laporte, 1834
Subtribe Demetriadina Bates, 1886
Subtribe Dromiusina Bonelli, 1810
Subtribe Gallerucidiina Chaudoir, 1872
Subtribe Lebiina Bonelli, 1810
Subtribe Metallicina Basilewsky, 1984
Subtribe Nemotarsina Bates, 1883
Subtribe Pericalina Hope, 1838
Subtribe Pseudotrechina Basilewsky, 1984
Subtribe Sugimotoina Habu, 1975
Subtribe Trichina Basilewsky, 1984
Tribe Licinini Bonelli, 1810
Subtribe Dicaelina Laporte, 1834
Subtribe Dicrochilina Ball, 1992
Subtribe Lestignathina Ball, 1992
Subtribe Licinina Bonelli, 1810
Tribe Melanchitonini Jeannel, 1948
Tribe Microcheilini Jeannel, 1948
Tribe Morionini Brullé, 1835
Tribe Odacanthini Laporte, 1834
Tribe Omphreini Ganglbauer, 1891
Tribe Oodini LaFerté-Sénectère, 1851
Tribe Orthogoniini Schaum, 1857
Tribe Panagaeini Bonelli, 1810
Subtribe Brachygnathina Basilewsky, 1946
Subtribe Panagaeina Bonelli, 1810
Subtribe Tefflina Basilewsky, 1946
Tribe Peleciini Chaudoir, 1880
Subtribe Agonicina Sloane, 1920
Subtribe Peleciina Chaudoir, 1880
Tribe Pentagonicini Bates, 1873
Tribe Perigonini Horn, 1881 nomen protectum
Tribe Physocrotaphini Chaudoir, 1863
Tribe Platynini Bonelli, 1810
Tribe Pseudomorphini Hope, 1838
Tribe Pterostichini Bonelli, 1810
Subtribe Abacomorphina Tschitschérine, 1902
Subtribe Euchroina Chaudoir, 1874
Subtribe Metiina Straneo, 1951
Subtribe Microcephalina Tschitschérine, 1898
Subtribe Pterostichina Bonelli, 1810
Tribe Sphodrini Laporte, 1834
Subtribe Atranopsina Baehr, 1982
Subtribe Calathina Laporte, 1834
Subtribe Dolichina Brullé, 1834
Subtribe Pristosiina Lindroth, 1956
Subtribe Sphodrina Laporte, 1834
Subtribe Synuchina Lindroth, 1956
Tribe Xenaroswellianini Erwin, 2007
Tribe Zabrini Bonelli, 1810
Subtribe Amarina Zimmermann, 1832
Subtribe Zabrina Bonelli, 1810
Tribe Zuphiini Bonelli, 1810
Subtribe Dicrodontina Machado, 1992
Subtribe Leleupidiina Basilewsky, 1951
Subtribe Metazuphiina Mateu, 1992
Subtribe Mischocephalina Mateu, 1992
Subtribe Patriziina Basilewsky, 1953
Subtribe Zuphiina Bonelli, 1810
Family Haliplidae Aubé, 1836
†Family Triaplidae Ponomarenko, 1977
†Family Colymbotethidae Ponomarenko, 1994
†Family Parahygrobiidae Ponomarenko, 1977
†Family Coptoclavidae Ponomarenko, 1961
†Subfamily Necronectinae Ponomarenko, 1977
†Subfamily Charonoscaphinae Ponomarenko, 1977
†Subfamily Coptoclavinae Ponomarenko, 1961
†Subfamily Coptoclaviscinae Soriano, Ponomarenko and Delclos, 2007
†Subfamily Hispanoclavinae Soriano, Ponomarenko and Delclos, 2007
†Family Liadytidae Ponomarenko, 1977
Family Meruidae Spangler and Steiner, 2005
Family Noteridae Thomson, 1860
Subfamily Noterinae Thomson, 1860
Tribe Neohydrocoptini Zalat, Saleh, Angus and Kaschef, 2000
Tribe Noterini Thomson, 1860
Tribe Pronoterini Nilsson, 2005
Tribe Tonerini Miller, 2009
Subfamily Notomicrinae Zimmermann, 1919
Subfamily Phreatodytinae Uéno, 1957
Family Amphizoidae LeConte, 1853
Family Aspidytidae Ribera, Beutel, Balke and Vogler, 2002
Family Hygrobiidae Régimbart, 1879 (1837)
Family Dytiscidae Leach, 1815
Subfamily Agabinae Thomson, 1867
Subfamily Colymbetinae Erichson, 1837
Tribe Anisomeriini Brinck, 1948
Tribe Carabdytini Pederzani, 1995
Tribe Colymbetini Erichson, 1837
Subfamily Copelatinae Branden, 1885
Subfamily Coptotominae Branden, 1885
Subfamily Dytiscinae Leach, 1815
Tribe Aciliini Thomson, 1867
Tribe Aubehydrini Guignot, 1942
Tribe Cybisterini Sharp, 1880
Tribe Dytiscini Leach, 1815
Tribe Eretini Crotch, 1873
Tribe Hydaticini Sharp, 1880
Tribe Hyderodini Miller, 2000
Subfamily Hydrodytinae Miller, 2001
Subfamily Hydroporinae Aubé, 1836
Tribe Bidessini Sharp, 1880
Tribe Carabhydrini Watts, 1978
Tribe Hydroporini Aubé, 1836
Tribe Hydrovatini Sharp, 1880
Tribe Hygrotini Portevin, 1929
Tribe Hyphydrini Gistel, 1848
Tribe Laccornini Wolfe and Roughley, 1990
Tribe Methlini Branden, 1885
†Tribe Schistomerini Palmer, 1957
Tribe Vatellini Sharp, 1880
Subfamily Laccophilinae Gistel, 1848
Tribe Agabetini Branden, 1885
Tribe Laccophilini Gistel, 1848
Subfamily Lancetinae Branden, 1885
Subfamily Matinae Branden, 1885
†Subfamily Palaeogyrininae Schlechtendal, 1894
†Subfamily Liadytiscinae Prokin and Ren, 2010
Suborder POLYPHAGA
Series STAPHYLINIFORMIA
Superfamily Hydrophiloidea Latreille, 1802
Family Hydrophilidae Latreille, 1802
Subfamily Helophorinae Leach, 1815
Subfamily Epimetopinae Zaitzev, 1908
Subfamily Georissinae Laporte, 1840
Subfamily Hydrochinae Thomson, 1859
Subfamily Spercheinae Erichson, 1837
Subfamily Horelophinae Hansen, 1991
Subfamily Horelophopsinae Hansen, 1997
Subfamily Hydrophilinae Latreille, 1802
Tribe Anacaenini Hansen, 1991
Tribe Berosini Mulsant, 1844
Tribe Chaetarthriini Bedel, 1881
Tribe Hydrophilini Latreille, 1802
Subtribe Acidocerina Zaitzev, 1908
Subtribe Globuloseina García, 2001
Subtribe Hydrobiusina Mulsant, 1844
Subtribe Hydrophilina Latreille, 1802
Tribe Laccobiini Houlbert, 1922
Tribe Sperchopsini Hansen, 1991
Subfamily Sphaeridiinae Latreille, 1802
Tribe Andotypini Hansen, 1991
Tribe Borborophorini Hansen, 1991
Tribe Coelostomatini Heyden, 1891 (1890)
Tribe Megasternini Mulsant, 1844
Tribe Omicrini Smetana, 1975
Tribe Protosternini Hansen, 1991
Tribe Rygmodini Orchymont, 1916
Tribe Sphaeridiini Latreille, 1802
Tribe Tormissini Hansen, 1991
Family Sphaeritidae Shuckard, 1839
Family Synteliidae Lewis, 1882
Family Histeridae Gyllenhal, 1808
Subfamily Niponiinae Fowler, 1912
Subfamily Abraeinae MacLeay, 1819
Tribe Abraeini MacLeay, 1819
Tribe Acritini Wenzel, 1944
Tribe Acritomorphini Wenzel, 1944
Tribe Plegaderini Portevin, 1929
Tribe Teretriini Bickhardt, 1914
Subfamily Trypeticinae Bickhardt, 1913
Subfamily Trypanaeinae Marseul, 1857
Subfamily Saprininae Blanchard, 1845
Subfamily Dendrophilinae Reitter, 1909
Tribe Anapleini Olexa, 1982
Tribe Bacaniini Kryzhanovskij, 1976
Tribe Dendrophilini Reitter, 1909
Tribe Paromalini Reitter, 1909
Subfamily Onthophilinae MacLeay, 1819
Subfamily Tribalinae Bickhardt, 1914
Subfamily Histerinae Gyllenhal, 1808
Tribe Exosternini Bickhardt, 1914
Tribe Histerini Gyllenhal, 1808
Tribe Hololeptini Hope, 1840
Tribe Omalodini Kryzhanovskij, 1972
Tribe Platysomatini Bickhardt, 1914
Subfamily Haeteriinae Marseul, 1857
Tribe Haeteriini Marseul, 1857
Tribe Nymphistrini Tishechkin, 2007
Tribe Synoditulini Tishechkin, 2007
Subfamily Chlamydopsinae Bickhardt, 1914
Superfamily Staphylinoidea Latreille, 1802
Family Hydraenidae Mulsant, 1844
Subfamily Orchymontiinae Perkins, 1997
Subfamily Prosthetopinae Perkins, 1994
Tribe Coelometoponini Perkins, 2005
Tribe Nucleotopini Perkins, 1994
Tribe Parasthetopini Perkins, 1994
Tribe Prosthetopini Perkins, 1994
Tribe Protosthetopini Perkins, 1994
Tribe Pterosthetopini Perkins, 1994
Subfamily Hydraeninae Mulsant, 1844
Tribe Hydraenidini Perkins, 1980
Tribe Hydraenini Mulsant, 1844
Tribe Limnebiini Mulsant, 1844
Tribe Madagastrini Perkins, 1997
Tribe Parhydraenini Perkins, 1997
Subfamily Ochthebiinae Thomson, 1859
Tribe Ochthebiini Thomson, 1859
Subtribe Enicocerina Perkins, 1997
Subtribe Meropathina Perkins, 1997
Subtribe Neochthebiina Perkins, 1997
Subtribe Ochthebiina Thomson, 1859
Subtribe Protochthebiina Perkins, 1997
Tribe Ochtheosini Perkins, 1997
Family Ptiliidae Erichson, 1845
Subfamily Ptiliinae Erichson, 1845
Tribe Discheramocephalini Grebennikov, 2009
Tribe Nanosellini Barber, 1924
Tribe Ptenidiini Flach, 1889
Tribe Ptiliini Erichson, 1845
Tribe Ptinellini Reitter, 1906 (1891)
Subfamily Cephaloplectinae Sharp, 1883
Subfamily Acrotrichinae Reitter, 1909 (1856)
Family Agyrtidae Thomson, 1859
Subfamily Agyrtinae Thomson, 1859
Subfamily Necrophilinae Newton, 1997
Subfamily Pterolomatinae Thomson, 1862
Family Leiodidae Fleming, 1821
Subfamily Camiarinae Jeannel, 1911
Tribe Agyrtodini Jeannel, 1936
Tribe Camiarini Jeannel, 1911
Tribe Neopelatopini Jeannel, 1962
Subfamily Catopocerinae Hatch, 1927 (1880)
Tribe Catopocerini Hatch, 1927 (1880)
Tribe Glacicavicolini Westcott, 1968
Subfamily Leiodinae Fleming, 1821
Tribe Agathidiini Westwood, 1838
Tribe Estadiini Portevin, 1914
Tribe Leiodini Fleming, 1821
Tribe Pseudoliodini Portevin, 1926
Tribe Scotocryptini Reitter, 1884
Tribe Sogdini Lopatin, 1961
Subfamily Coloninae Horn, 1880 (1859)
Subfamily Cholevinae Kirby, 1837
Tribe Anemadini Hatch, 1928
Subtribe Anemadina Hatch, 1928
Subtribe Eocatopina Jeannel, 1936
Subtribe Eunemadina Newton, 1998
Subtribe Nemadina Jeannel, 1936
Subtribe Paracatopina Jeannel, 1936
Tribe Cholevini Kirby, 1837
Subtribe Catopina Chaudoir, 1845
Subtribe Cholevina Kirby, 1837
Tribe Eucatopini Jeannel, 1921
Tribe Leptodirini Lacordaire, 1854 (1849)
Subtribe Anthroherponina Jeannel, 1910
Subtribe Bathysciina Horn, 1880
Subtribe Bathysciotina Guéorguiev, 1974
Subtribe Leptodirina Lacordaire, 1854 (1849)
Subtribe Pholeuina Reitter, 1886
Subtribe Platycholeina Horn, 1880
Subtribe Spelaeobatina Guéorguiev, 1974
Tribe Oritocatopini Jeannel, 1936
Tribe Ptomaphagini Jeannel, 1911
Subtribe Baryodirina Perreau, 2000
Subtribe Ptomaphagina Jeannel, 1911
Subtribe Ptomaphaginina Szymczakowski, 1964
Tribe Sciaphyini Perreau, 2000
Subfamily Platypsyllinae Ritsema, 1869
Family Silphidae Latreille, 1806
Subfamily Silphinae Latreille, 1806
Subfamily Nicrophorinae Kirby, 1837
Family Staphylinidae Latreille, 1802
Subfamily Glypholomatinae Jeannel, 1962
Subfamily Microsilphinae Crowson, 1950
Subfamily Omaliinae MacLeay, 1825
Tribe Anthophagini Thomson, 1859
Tribe Aphaenostemmini Peyerimhoff, 1914
Tribe Corneolabiini Steel, 1950
Tribe Coryphiini Jakobson, 1908
Subtribe Boreaphilina Zerche, 1990
Subtribe Coryphiina Jakobson, 1908
Tribe Eusphalerini Hatch, 1957
Tribe Hadrognathini Portevin, 1929
Tribe Omaliini MacLeay, 1825
Subfamily Empelinae Newton and Thayer, 1992
Subfamily Proteininae Erichson, 1839
Tribe Anepiini Steel, 1966
Tribe Austrorhysini Newton and Thayer, 1995
Tribe Nesoneini Steel, 1966
Tribe Proteinini Erichson, 1839
Tribe Silphotelini Newton and Thayer, 1995
Subfamily Micropeplinae Leach, 1815
Subfamily Neophoninae Fauvel, 1905
Subfamily Dasycerinae Reitter, 1887
Subfamily Protopselaphinae Newton and Thayer, 1995
Subfamily Pselaphinae Latreille, 1802
Supertribe Batrisitae Reitter, 1882
Tribe Amauropini Jeannel, 1948
Tribe Batrisini Reitter, 1882
Subtribe Ambicocerina Leleup, 1970
Subtribe Batrisina Reitter, 1882
Subtribe Leupeliina Jeannel, 1954
Subtribe Stilipalpina Jeannel, 1954
Tribe Thaumastocephalini Poggi, Nonveiller, Colla, Pavićević and Rada, 2001
Supertribe Clavigeritae Leach, 1815
Tribe Clavigerini Leach, 1815
Subtribe Apoderigerina Jeannel, 1954
Subtribe Clavigerina Leach, 1815
Subtribe Clavigerodina Schaufuss, 1882
Subtribe Disarthricerina Jeannel, 1949
Subtribe Hoplitoxenina Célis, 1969
Subtribe Lunillina Célis, 1969
Subtribe Mastigerina Jeannel, 1954
Subtribe Miroclavigerina Jeannel, 1949
Subtribe Neocerina Jeannel, 1954
Subtribe Radamina Jeannel, 1954
Subtribe Thysdariina Jeannel, 1954
Tribe Colilodionini Besuchet, 1991
Tribe Tiracerini Besuchet, 1986
Supertribe Euplectitae Streubel, 1839
Tribe Bythinoplectini Schaufuss, 1890
Subtribe Bythinoplectina Schaufuss, 1890
Subtribe Pyxidicerina Raffray, 1904
Tribe Dimerini Raffray, 1908
Tribe Euplectini Streubel, 1839
Tribe Jubini Raffray, 1904
Tribe Mayetiini Winkler, 1925
Tribe Metopiasini Raffray, 1904
Subtribe Metopiasina Raffray, 1904
Subtribe Rhinoscepsina Bowman, 1934
Tribe Trichonychini Reitter, 1882
Subtribe Bibloporina Park, 1951
Subtribe Panaphantina Jeannel, 1950
Subtribe Trichonychina Reitter, 1882
Subtribe Trimiina Bowman, 1934
Tribe Trogastrini Jeannel, 1949
Subtribe Phtegnomina Park, 1951
Subtribe Rhexiina Park, 1951
Subtribe Trogastrina Jeannel, 1949
Supertribe Faronitae Reitter, 1882
Supertribe Goniaceritae Reitter, 1882 (1872)
Tribe Arnylliini Jeannel, 1952
Tribe Barrosellini Leleup, 1973
Tribe Brachyglutini Raffray, 1904
Subtribe Baradina Park, 1951
Subtribe Brachyglutina Raffray, 1904
Subtribe Decarthrina Park, 1951
Subtribe Eupseniina Park, 1951
Tribe Bythinini Raffray, 1890
Subtribe Bythinina Raffray, 1890
Subtribe Machaeritina Jeannel, 1950
Subtribe Xenobythina Jeannel, 1950
Tribe Cyathigerini Schaufuss, 1872
Tribe Goniacerini Reitter, 1882 (1872)
Tribe Imirini Jeannel, 1949
Tribe Iniocyphini Park, 1951
Subtribe Iniocyphina Park, 1951
Subtribe Natypleurina Newton and Thayer, 1992
Tribe Machadoini Jeannel, 1951
Tribe Proterini Jeannel, 1949
Tribe Pygoxyini Reitter, 1909
Tribe Speleobamini Park, 1951
Tribe Tychini Raffray, 1904
Tribe Valdini Park, 1953
Supertribe Pselaphitae Latreille, 1802
Tribe Arhytodini Raffray, 1890
Tribe Attapseniini Bruch, 1933
Tribe Ctenistini Blanchard, 1845
Tribe Hybocephalini Raffray, 1890
Tribe Odontalgini Jeannel, 1949
Tribe Pachygastrodini Leleup, 1969
Tribe Phalepsini Jeannel, 1949
Tribe Pselaphini Latreille, 1802
Tribe Schistodactylini Raffray, 1890
Tribe Tmesiphorini Jeannel, 1949
Tribe Tyrini Reitter, 1882
Subtribe Centrophthalmina Jeannel, 1949
Subtribe Janusculina Cerruti, 1970
Subtribe Somatipionina Jeannel, 1949
Subtribe Tyrina Reitter, 1882
Subfamily Phloeocharinae Erichson, 1839
Subfamily Olisthaerinae Thomson, 1858
Subfamily Tachyporinae MacLeay, 1825
Tribe Deropini Smetana, 1983
Tribe Megarthropsini Cameron, 1919
Tribe Mycetoporini Thomson, 1859
Tribe Tachyporini MacLeay, 1825
Tribe Vatesini Seevers, 1958
Subfamily Trichophyinae Thomson, 1858
Subfamily Habrocerinae Mulsant and Rey, 1876
Subfamily Aleocharinae Fleming, 1821
Tribe Actocharini Bernhauer and Schubert, 1911
Tribe Aenictoteratini Kistner, 1993
Tribe Akatastopsisini Pace, 2000
Tribe Aleocharini Fleming, 1821
Subtribe Aleocharina Fleming, 1821
Subtribe Compactopediina Kistner, 1970
Subtribe Hodoxenina Kistner, 1970
Tribe Athetini Casey, 1910
Subtribe Athetina Casey, 1910
Subtribe Coptotermoeciina Kistner and Pasteels, 1970
Subtribe Microceroxenina Kistner, 1970
Subtribe Nasutiphilina Kistner, 1970
Subtribe Schistogeniina Fenyes, 1918
Subtribe Taxicerina Lohse, 1989
Subtribe Termitotelina Kistner, 1970
Subtribe Thamiaraeina Fenyes, 1921
Tribe Autaliini Thomson, 1859
Tribe Cordobanini Bernhauer, 1910
Tribe Corotocini Fenyes, 1918
Subtribe Abrotelina Seevers, 1957
Subtribe Corotocina Fenyes, 1918
Subtribe Eburniogastrina Jacobson, Kistner and Pasteels, 1986
Subtribe Nasutitellina Jacobson, Kistner and Pasteels, 1986
Subtribe Sphuridaethina Pace, 1988
Subtribe Termitocharina Seevers, 1957
Subtribe Termitocupidina Jacobson, Kistner and Pasteels, 1986
Subtribe Termitogastrina Bernhauer and Scheerpeltz, 1926
Subtribe Termitoiceina Jacobson, Kistner and Pasteels, 1986
Subtribe Termitopithina Jacobson, Kistner and Pasteels, 1986
Subtribe Termitoptochina Fenyes, 1921
Subtribe Timeparthenina Fenyes, 1921
Tribe Crematoxenini Mann, 1921
Tribe Cryptonotopseini Pace, 2003
Tribe Deinopsini Sharp, 1883
Tribe Diestotini Mulsant and Rey, 1871
Tribe Diglottini Jakobson, 1909
Tribe Digrammini Fauvel, 1900
Tribe Dorylogastrini Wasmann, 1916
Tribe Dorylomimini Wasmann, 1916
Tribe Drepanoxenini Kistner and Watson, 1972
Tribe Ecitocharini Seevers, 1965
Tribe Ecitogastrini Fenyes, 1918
Tribe Eusteniamorphini Bernhauer and Scheerpeltz, 1926
Tribe Falagriini Mulsant and Rey, 1873
Tribe Feldini Kistner, 1972
Tribe Gymnusini Heer, 1839
Tribe Himalusini Klimaszewski, Pace and Center, 2010
Tribe Homalotini Heer, 1839
Subtribe Bolitocharina Thomson, 1859
Subtribe Dinardopsina Bernhauer and Scheerpeltz, 1926
Subtribe Gyrophaenina Kraatz, 1856
Subtribe Homalotina Heer, 1839
Subtribe Silusina Fenyes, 1918
Tribe Hoplandriini Casey, 1910
Subtribe Hoplandriina Casey, 1910
Subtribe Platandriina Hanley, 2002
Subtribe Pseudoplandriina Hanley, 2002
Tribe Hygronomini Thomson, 1859
Subtribe Hygronomina Thomson, 1859
Subtribe Saphoglossina Bernhauer and Scheerpeltz, 1926
Tribe Hypocyphtini Laporte, 1835
Tribe Leucocraspedini Fenyes, 1921
Tribe Liparocephalini Fenyes, 1918
Tribe Lomechusini Fleming, 1821
Subtribe Aenictobiina Kistner, 1997
Subtribe Lomechusina Fleming, 1821
Subtribe Myrmedoniina Thomson, 1867
Subtribe Termitozyrina Seevers, 1957
Tribe Masuriini Cameron, 1939
Tribe Mesoporini Cameron, 1959
Tribe Mimanommatini Wasmann, 1912
Subtribe Dorylophilina Fenyes, 1921
Subtribe Mimanommatina Wasmann, 1912
Tribe Mimecitini Wasmann, 1917
Subtribe Labidopullina Jacobson and Kistner, 1991
Subtribe Leptanillophilina Fenyes, 1918
Subtribe Mimecitina Wasmann, 1917
Subtribe Mimonillina Bernhauer and Scheerpeltz, 1926
Tribe Myllaenini Ganglbauer, 1895
Tribe Oxypodini Thomson, 1859
Subtribe Aphytopodina Bernhauer and Scheerpeltz, 1926
Subtribe Blepharhymenina Klimaszewski and Peck, 1986
Subtribe Dinardina Mulsant and Rey, 1873
Subtribe Meoticina Seevers, 1978
Subtribe Oxypodina Thomson, 1859
Subtribe Tachyusina Thomson, 1859
Tribe Oxypodinini Fenyes, 1921
Tribe Paglini Newton and Thayer, 1992
Tribe Paradoxenusini Bruch, 1937
Tribe Pediculotini Ádám, 1987
Tribe Philotermitini Seevers, 1957
Tribe Phyllodinardini Wasmann, 1916
Tribe Phytosini Thomson, 1867
Tribe Placusini Mulsant and Rey, 1871
Tribe Pronomaeini Mulsant and Rey, 1873
Tribe Pseudoperinthini Cameron, 1939
Tribe Pygostenini Fauvel, 1899
Tribe Sahlbergiini Kistner, 1993
Tribe Sceptobiini Seevers, 1978
Tribe Skatitoxenini Kistner and Pasteels, 1969
Tribe Termitodiscini Wasmann, 1904
Subtribe Athexeniina Pace, 2000
Subtribe Termitodiscina Wasmann, 1904
Tribe Termitohospitini Seevers, 1941
Subtribe Hetairotermitina Seevers, 1957
Subtribe Termitohospitina Seevers, 1941
Tribe Termitonannini Fenyes, 1918
Subtribe Perinthina Bernhauer and Scheerpeltz, 1926
Subtribe Termitonannina Fenyes, 1918
Tribe Termitopaediini Seevers, 1957
Tribe Termitusini Fenyes, 1918
Subtribe Termitospectrina Seevers, 1957
Subtribe Termitusina Fenyes, 1918
Tribe Trichopseniini LeConte and Horn, 1883
Tribe Trilobitideini Fauvel, 1899
Subfamily Trigonurinae Reiche, 1866
Subfamily Apateticinae Fauvel, 1895
Subfamily Scaphidiinae Latreille, 1806
Tribe Cypariini Achard, 1924
Tribe Scaphidiini Latreille, 1806
Tribe Scaphiini Achard, 1924
Tribe Scaphisomatini Casey, 1893
Subfamily Piestinae Erichson, 1839
Subfamily Osoriinae Erichson, 1839
Tribe Eleusinini Sharp, 1887
Tribe Leptochirini Sharp, 1887
Tribe Osoriini Erichson, 1839
Subtribe Osoriina Erichson, 1839
Subtribe Parosoriina Bernhauer and Schubert, 1911
Tribe Thoracophorini Reitter, 1909
Subtribe Clavilispinina Newton and Thayer, 1992
Subtribe Glyptomina Newton and Thayer, 1992
Subtribe Lispinina Bernhauer and Schubert, 1910
Subtribe Thoracophorina Reitter, 1909
Subfamily Oxytelinae Fleming, 1821
Tribe Blediini Ádám, 2001
Tribe Coprophilini Heer, 1839
Tribe EuphaniiniReitter, 1909
Tribe Oxytelini Fleming, 1821
Tribe Planeustomini Jacquelin du Val, 1857
Subfamily Oxyporinae Fleming, 1821
Subfamily Megalopsidiinae Leng, 1920
Subfamily Scydmaeninae Leach, 1815
†Supertribe Hapsomelitae Poinar and Brown, 2004
Supertribe Mastigitae Fleming, 1821
Tribe Clidicini Casey, 1897
Tribe Leptomastacini Casey, 1897
Tribe Mastigini Fleming, 1821
Supertribe Scydmaenitae Leach, 1815
Tribe Cephenniini Reitter, 1882
Tribe Chevrolatiini Reitter, 1882
Tribe Cyrtoscydmini Schaufuss, 1889
Tribe Eutheiini Casey, 1897
Tribe Leptoscydmini Casey, 1897
Tribe Plaumanniolini Costa Lima, 1962
Tribe Scydmaenini Leach, 1815
Subfamily Steninae MacLeay, 1825
Subfamily Euaesthetinae Thomson, 1859
Tribe Alzadaesthetini Scheerpeltz, 1974
Tribe Austroesthetini Cameron, 1944
Tribe Euaesthetini Thomson, 1859
Tribe Fenderiini Scheerpeltz, 1974
Tribe Nordenskioldiini Bernhauer and Schubert, 1911
Tribe Stenaesthetini Bernhauer and Schubert, 1911
Subfamily Solieriinae Newton and Thayer, 1992
Subfamily Leptotyphlinae Fauvel, 1874
Tribe Cephalotyphlini Coiffait, 1963
Tribe Entomoculiini Coiffait, 1957
Tribe Leptotyphlini Fauvel, 1874
Tribe Metrotyphlini Coiffait, 1963
Tribe Neotyphlini Coiffait, 1963
Subfamily Pseudopsinae Ganglbauer, 1895
Subfamily Paederinae Fleming, 1821
Tribe Paederini Fleming, 1821
Subtribe Astenina Hatch, 1957
Subtribe Cryptobiina Casey, 1905
Subtribe Cylindroxystina Bierig, 1943
Subtribe Dolicaonina Casey, 1905
Subtribe Echiasterina Casey, 1905
Subtribe Lathrobiina Laporte, 1835
Subtribe Lithocharina Casey, 1905
Subtribe Medonina Casey, 1905
Subtribe Paederina Fleming, 1821
Subtribe Scopaeina Mulsant and Rey, 1878
Subtribe Stilicina Casey, 1905
Subtribe Stilicopsina Casey, 1905
Tribe Pinophilini Nordmann, 1837
Subtribe Pinophilina Nordmann, 1837
Subtribe Procirrina Bernhauer and Schubert, 1912
Subfamily Staphylininae Latreille, 1802
Tribe Arrowinini Solodovnikov and Newton, 2005
Tribe Diochini Casey, 1906
Tribe Maorothiini Assing, 2000
Tribe Othiini Thomson, 1859
Tribe Platyprosopini Lynch Arribálzaga, 1884
Tribe Staphylinini Latreille, 1802
Subtribe Amblyopinina Seevers, 1944
Subtribe Anisolinina Hayashi, 1993
Subtribe Eucibdelina Sharp, 1889
Subtribe Hyptiomina Casey, 1906
Subtribe Philonthina Kirby, 1837
Subtribe Quediina Kraatz, 1857
Subtribe Staphylinina Latreille, 1802
Subtribe Tanygnathinina Reitter, 1909
Subtribe Xanthopygina Sharp, 1884
Tribe Xantholinini Erichson, 1839
†Subfamily Protactinae Heer, 1847
Series SCARABAEIFORMIA
Superfamily Scarabaeoidea Latreille, 1802
Family Pleocomidae LeConte, 1861
Subfamily Pleocominae LeConte, 1861
†Subfamily Cretocominae Nikolajev, 2002
†Subfamily Archescarabaeinae Nikolajev, 2010
Family Geotrupidae Latreille, 1802
Subfamily Taurocerastinae Germain, 1897
Subfamily Bolboceratinae Mulsant, 1842
Tribe Athyreini Lynch Arribálzaga, 1878
Tribe Bolbelasmini Nikolajev, 1996
Tribe Bolboceratini Mulsant, 1842
Tribe Bolbochromini Nikolajev, 1970
Tribe Eubolbitini Nikolajev, 1970
Tribe Eucanthini Nikolajev, 2003
Tribe Gilletinini Nikolajev, 1990
Tribe Odonteini Shokhin, 2007
Tribe Stenaspidiini Nikolajev, 2003
Subfamily Geotrupinae Latreille, 1802
Tribe Ceratotrupini Zunino, 1984
Tribe Enoplotrupini Paulian, 1945
†Tribe Cretogeotrupini Nikolajev, 1996
Tribe Geotrupini Latreille, 1802
Tribe Lethrini Oken, 1843
Family Belohinidae Paulian, 1959
Family Passalidae Leach, 1815
Subfamily Aulacocyclinae Kaup, 1868
Tribe Aulacocyclini Kaup, 1868
Tribe Ceracupedini Boucher, 2006
Subfamily Passalinae Leach, 1815
Tribe Leptaulacini Kaup, 1871
Tribe Macrolinini Kaup, 1871
Tribe Passalini Leach, 1815
Tribe Proculini Kaup, 1868
Tribe Solenocyclini Kaup, 1871
Family Trogidae MacLeay, 1819
†Subfamily Avitortorinae Nikolajev, 2007
Subfamily Troginae MacLeay, 1819
Subfamily Omorginae Nikolajev, 2005
Family Glaresidae Kolbe, 1905
Family Diphyllostomatidae Holloway, 1972
Family Lucanidae Latreille, 1804
†Subfamily Protolucaninae Nikolajev, 2007
Subfamily Aesalinae MacLeay, 1819
Tribe Aesalini MacLeay, 1819
Tribe Ceratognathini Sharp, 1899
Tribe Nicagini LeConte, 1861
†Subfamily Ceruchitinae Nikolajev, 2006
Subfamily Syndesinae MacLeay, 1819
Subfamily Lampriminae MacLeay, 1819
Tribe Lamprimini MacLeay, 1819
Tribe Streptocerini Kikuta, 1986
Subfamily Lucaninae Latreille, 1804
Tribe Chiasognathini Burmeister, 1847
Tribe Lucanini Latreille, 1804
Tribe Platycerini Mulsant, 1842
Tribe Platyceroidini Paulsen and Hawks, 2008
†Subfamily Paralucaninae Nikolajev, 2000
Family Ochodaeidae Mulsant and Rey, 1871
†Subfamily Cretochodaeinae Nikolajev, 1995
Subfamily Ochodaeinae Mulsant and Rey, 1871
Tribe Enodognathini Scholtz, 1988
Tribe Ochodaeini Mulsant and Rey, 1871
Subfamily Chaetocanthinae Scholtz, 1988
Tribe Chaetocanthini Scholtz, 1988
Tribe Pseudochodaeini Scholtz, 1988
Tribe Synochodaeini Scholtz, 1988
Family Hybosoridae Erichson, 1847
†Subfamily Mimaphodiinae Nikolajev, 2007
Subfamily Anaidinae Nikolajev, 1996
Subfamily Ceratocanthinae Martínez, 1968
Tribe Ceratocanthini Martínez, 1968
Tribe Ivieolini Howden and Gill, 2000
Tribe Scarabatermitini Nikolajev, 1999
Subfamily Hybosorinae Erichson, 1847
Subfamily Liparochrinae Ocampo, 2006
Subfamily Pachyplectrinae Ocampo, 2006
Family Glaphyridae MacLeay, 1819
Subfamily Glaphyrinae MacLeay, 1819
Subfamily Amphicominae Blanchard, 1845
†Subfamily Cretoglaphyrinae Nikolajev, 2005
Family Scarabaeidae Latreille, 1802
†Subfamily Lithoscarabaeinae Nikolajev, 1992
Subfamily Chironinae Blanchard, 1845
Subfamily Aegialiinae Laporte, 1840
Subfamily Eremazinae Iablokoff-Khnzorian, 1977
Subfamily Aphodiinae Leach, 1815
Tribe Aphodiini Leach, 1815
Subtribe Aphodiina Leach, 1815
Subtribe Didactyliina Pittino, 1985
Subtribe Proctophanina Stebnicka and Howden, 1995
Tribe Corythoderini Schmidt, 1910
Tribe Eupariini Schmidt, 1910 nomen protectum
Tribe Odontolochini Stebnicka and Howden, 1996
Tribe Odochilini Rakovič, 1987
Tribe Psammodiini Mulsant, 1842
Subtribe Phycocina Landin, 1960
Subtribe Psammodiina Mulsant, 1842
Subtribe Rhyssemina Pittino and Mariani, 1986
Tribe Rhyparini Schmidt, 1910
Tribe Stereomerini Howden and Storey, 1992
Tribe Termitoderini Tangelder and Krikken, 1982
Subfamily Aulonocneminae Janssens, 1946
Subfamily Termitotroginae Wasmann, 1918
Subfamily Scarabaeinae Latreille, 1802
Tribe Ateuchini Perty, 1830
Subtribe Ateuchina Perty, 1830
Subtribe Scatimina Vaz-de-Mello, 2008
Tribe Coprini Leach, 1815
Tribe Deltochilini Lacordaire, 1856
Tribe Eucraniini Burmeister, 1873
Tribe Gymnopleurini Lacordaire, 1856
Tribe Oniticellini Kolbe, 1905
Subtribe Drepanocerina van Lansberge, 1875
Subtribe Eurysternina Vulcano, Martínez and Pereira, 1961
Subtribe Helictopleurina Janssens, 1946
Subtribe Oniticellina Kolbe, 1905
Tribe Onitini Laporte, 1840
Tribe Onthophagini Burmeister, 1846
Tribe Phanaeini Hope, 1838
Tribe Scarabaeini Latreille, 1802
Tribe Sisyphini Mulsant, 1842
†Subfamily Prototroginae Nikolajev, 2000
†Subfamily Cretoscarabaeinae Nikolajev, 1995
Subfamily Dynamopodinae Arrow, 1911
Tribe Dynamopodini Arrow, 1911
Tribe Thinorycterini Semenov and Reichardt, 1925
Subfamily Phaenomeridinae Erichson, 1847
Subfamily Orphninae Erichson, 1847
Tribe Aegidiini Paulian, 1984
Tribe Orphnini Erichson, 1847
Subfamily Allidiostomatinae Arrow, 1940
Subfamily Aclopinae Blanchard, 1850
Tribe Aclopini Blanchard, 1850
†Tribe Holcorobeini Nikolajev, 1992
Tribe Phaenognathini Iablokoff-Khnzorian, 1977
Subfamily Melolonthinae Leach, 1819
Tribe Ablaberini Blanchard, 1850
Tribe Automoliini Britton, 1978
Tribe Chasmatopterini Lacordaire, 1856
Tribe Colymbomorphini Blanchard, 1850
Tribe Comophorinini Britton, 1957
†Tribe Cretomelolonthini Nikolajev, 1998
Tribe Dichelonychini Burmeister, 1855
Tribe Diphucephalini Laporte, 1840
Tribe Diphycerini Medvedev, 1952
Tribe Diplotaxini Kirby, 1837
Tribe Euchirini Hope, 1840
Tribe Heteronychini Lacordaire, 1856
Tribe Hopliini Latreille, 1829
Subtribe Hopliina Latreille, 1829
Subtribe Pachycnemina Laporte, 1840
Tribe Lichniini Burmeister, 1844
Tribe Liparetrini Burmeister, 1855
Tribe Macrodactylini Kirby, 1837
Tribe Maechidiini Burmeister, 1855
Tribe Melolonthini Leach, 1819
Subtribe Enariina Dewailly, 1950
Subtribe Heptophyllina Medvedev, 1951
Subtribe Leucopholina Burmeister, 1855
Subtribe Melolonthina Leach, 1819
Subtribe Pegylina Lacroix, 1989
Subtribe Rhizotrogina Burmeister, 1855
Subtribe Schizonychina Burmeister, 1855
Tribe Oncerini LeConte, 1861
Tribe Pachypodini Erichson, 1840
Tribe Pachytrichini Burmeister, 1855
Tribe Phyllotocidiini Britton, 1957
Tribe Podolasiini Howden, 1997
Tribe Scitalini Britton, 1957
Tribe Sericini Kirby, 1837
Subtribe Phyllotocina Burmeister, 1855
Subtribe Sericina Kirby, 1837
Subtribe Trochalina Brenske, 1898
Tribe Sericoidini Erichson, 1847
Tribe Systellopini Sharp, 1877
Tribe Tanyproctini Erichson, 1847
Subtribe Macrophyllina Burmeister, 1855
Subtribe Tanyproctina Erichson, 1847
Subfamily Rutelinae MacLeay, 1819
Tribe Adoretini Burmeister, 1844
Subtribe Adoretina Burmeister, 1844
Subtribe Adorrhinyptiina Arrow, 1917
Subtribe Pachyrhinadoretina Ohaus, 1912
Subtribe Prodoretina Ohaus, 1912
Subtribe Trigonostomusina Ohaus, 1912
Tribe Alvarengiini Frey, 1975
Tribe Anatistini Lacordaire, 1856
Tribe Anomalini Streubel, 1839 nomen protectum
Subtribe Anisopliina Burmeister, 1844
Subtribe Anomalina Streubel, 1839 nomen protectum
Subtribe Isopliina Péringuey, 1902
Subtribe Leptohopliina Potts, 1974
Subtribe Popilliina Ohaus, 1918
Tribe Anoplognathini MacLeay, 1819
Subtribe Anoplognathina MacLeay, 1819
Subtribe Brachysternina Burmeister, 1844
Subtribe Phalangogoniina Ohaus, 1918
Subtribe Platycoeliina Burmeister, 1844
Subtribe Schizognathina Ohaus, 1918
Tribe Geniatini Burmeister, 1844
Tribe Rutelini MacLeay, 1819
Subtribe Areodina Burmeister, 1844
Subtribe Desmonychina Arrow, 1917
Subtribe Didrepanephorina Ohaus, 1918
Subtribe Heterosternina Bates, 1888 nomen protectum
Subtribe Lasiocalina Ohaus, 1918
Subtribe Oryctomorphina Burmeister, 1847
Subtribe Parastasiina Burmeister, 1844
Subtribe Rutelina MacLeay, 1819
Subfamily Dynastinae MacLeay, 1819
Tribe Agaocephalini Burmeister, 1847
Tribe Cyclocephalini Laporte, 1840
Tribe Dynastini MacLeay, 1819
Tribe Hexodontini Lacordaire, 1856
Tribe Oryctini Mulsant, 1842
Tribe Oryctoderini Endrödi, 1966
Tribe Pentodontini Mulsant, 1842
Subtribe Cheiroplatina Carne, 1957
Subtribe Dipelicina Carne, 1957
Subtribe Pentodontina Mulsant, 1842
Subtribe Pseudoryctina Carne, 1957
Tribe Phileurini Burmeister, 1847
Subtribe Cryptodina Burmeister and Schaum, 1840
Subtribe Phileurina Burmeister, 1847
Subfamily Cetoniinae Leach, 1815
Tribe Cetoniini Leach, 1815
Subtribe Cetoniina Leach, 1815
Subtribe Euphoriina Horn, 1880
Subtribe Leucocelina Kraatz, 1882
Tribe Cremastocheilini Burmeister and Schaum, 1841
Subtribe Aspilina Krikken, 1984
Subtribe Coenochilina Burmeister, 1842
Subtribe Cremastocheilina Burmeister and Schaum, 1841
Subtribe Cymophorina Krikken, 1984
Subtribe Genuchina Krikken, 1984
Subtribe Goliathopsidina Krikken, 1984
Subtribe Heterogeniina Krikken, 1984
Subtribe Lissogeniina Krikken, 1984
Subtribe Macromina Burmeister and Schaum, 1840
Subtribe Nyassinina Krikken, 1984
Subtribe Oplostomina Krikken, 1984
Subtribe Pilinurgina Krikken, 1984
Subtribe Spilophorina Krikken, 1984
Subtribe Telochilina Krikken, 1984
Subtribe Trichoplina Krikken, 1984
Subtribe Trogodina Krikken, 1984
Tribe Diplognathini Burmeister, 1842
Tribe Goliathini Latreille, 1829
Subtribe Coryphocerina Burmeister, 1842
Subtribe Dicronocephalina Krikken, 1984
Subtribe Goliathina Latreille, 1829
Subtribe Ichnestomatina Burmeister, 1842
Tribe Gymnetini Kirby, 1827
Subtribe Blaesiina Schoch, 1895
Subtribe Gymnetina Kirby, 1827
Tribe Phaedimini Schoch, 1894
Tribe Schizorhinini Burmeister, 1842
Subtribe Lomapterina Burmeister, 1842
Subtribe Schizorhinina Burmeister, 1842
Tribe Stenotarsiini Kraatz, 1880
Subtribe Anochiliina Krikken, 1984
Subtribe Coptomiina Schenkling, 1921
Subtribe Chromoptiliina Krikken, 1984
Subtribe Doryscelina Schenkling, 1921
Subtribe Euchroeina Paulian and Descarpentries, 1982
Subtribe Heterophanina Schoch, 1894
Subtribe Heterosomatina Krikken, 1984
Subtribe Pantoliina Krikken, 1984
Subtribe Parachiliina Krikken, 1984
Subtribe Stenotarsiina Kraatz, 1880
Tribe Taenioderini Mikšić, 1976
Subtribe Chalcotheina Mikšić, 1976
Subtribe Taenioderina Mikšić, 1976
Tribe Trichiini Fleming, 1821
Subtribe Cryptodontina Lacordaire, 1856
Subtribe Incina Burmeister, 1842
Subtribe Osmodermatina Schenkling, 1922
Subtribe Platygeniina Krikken, 1984
Subtribe Trichiina Fleming, 1821
Tribe Valgini Mulsant, 1842
Subtribe Microvalgina Kolbe, 1904
Subtribe Valgina Mulsant, 1842
Tribe Xiphoscelidini Burmeister, 1842
†Family Coprinisphaeridae Genise, 2004
†Family Pallichnidae Genise, 2004
Series ELATERIFORMIA
Superfamily Scirtoidea Fleming, 1821
Family Decliniidae Nikitsky, Lawrence, Kirejtshuk and Gratshev, 1994
Family Eucinetidae Lacordaire, 1857
Family Clambidae Fischer von Waldheim, 1821
Subfamily Calyptomerinae Crowson, 1955
Subfamily Acalyptomerinae Crowson, 1979
Subfamily Clambinae Fischer von Waldheim, 1821
Family Scirtidae Fleming, 1821
Subfamily Scirtinae Fleming, 1821
Subfamily Nipponocyphoninae Lawrence and Yoshitomi, 2007
Subfamily Stenocyphoninae Lawrence and Yoshitomi, 2007
†Family Elodophthalmidae Kirejtshuk and Azar, 2008
†Family Mesocinetida Kirejtshuk and Ponomarenko, 2010
Superfamily Dascilloidea Guérin-Méneville, 1843 (1834)
Family Dascillidae Guérin-Méneville, 1843 (1834)
Subfamily Dascillinae Guérin-Méneville, 1843 (1834)
Tribe Cinnabariini Pic, 1914
Tribe Dascillini Guérin-Méneville, 1843 (1834)
Subfamily Karumiinae Escalera, 1913
Tribe Emmitini Escalera, 1914
Tribe Escalerinini Paulus, 1972
Tribe Genecerini Pic, 1914
Tribe Karumiini Escalera, 1913
Family Rhipiceridae Latreille, 1834
Superfamily Buprestoidea Leach, 1815
Family Schizopodidae LeConte, 1859
Subfamily Schizopodinae LeConte, 1859
Tribe Dystaxiini Théry, 1929
†Tribe Electrapatini Iablokoff-Khnzorian, 1962
Tribe Schizopodini LeConte, 1859
Family Buprestidae Leach, 1815
Subfamily Julodinae Lacordaire, 1857
Subfamily Polycestinae Lacordaire, 1857
Tribe Acmaeoderini Kerremans, 1893
Subtribe Acmaeoderina Kerremans, 1893
Subtribe Acmaeoderoidina Cobos, 1955
Subtribe Nothomorphina Cobos, 1955
Tribe Astraeini Cobos, 1980
Tribe Bulini Bellamy, 1995
Tribe Haplostethini LeConte, 1861
Tribe Paratracheini Cobos, 1980
Tribe Perucolini Cobos, 1980
Tribe Polycestini Lacordaire, 1857
Subtribe Polycestina Lacordaire, 1857
Subtribe Xenopseina Volkovitsh, 2008
Tribe Polyctesini Cobos, 1955
Tribe Prospherini Cobos, 1980
Tribe Ptosimini Kerremans, 1903
Tribe Thrincopygini LeConte, 1861
Tribe Tyndaridini Cobos, 1955
Subtribe Mimicoclytrinina Bellamy, 2003
Subtribe Pseudacherusiina Cobos, 1980
Subtribe Tylaucheniina Cobos, 1959
Subtribe Tyndaridina Cobos, 1955
Tribe Xyroscelidini Cobos, 1955
Subfamily Galbellinae Reitter, 1911
Subfamily Chrysochroinae Laporte, 1835
Tribe Chrysochroini Laporte, 1835
Subtribe Chalcophorina Lacordaire, 1857 (1848)
Subtribe Chrysochroina Laporte, 1835
Subtribe Eucallopistina Bellamy, 2003
Tribe Dicercini Gistel, 1848
Subtribe Dicercina Gistel, 1848
Subtribe Haplotrinchina Holyński, 1993
Subtribe Hippomelanina Holyński, 1993
Subtribe Pseudoperotina Tôyama, 1987
Tribe Evidini Tôyama, 1987
Tribe Paraleptodemini Cobos, 1975
Subtribe Euchromatina Holyński, 1993
Subtribe Euplectaleciina Holyński, 1993
Subtribe Hypoprasina Holyński, 1993
Subtribe Paraleptodemina Cobos, 1975
Subtribe Pristipterina Holyński, 1993
Tribe Paratassini Bílý and Volkovitsh, 1996
Tribe Poecilonotini Jakobson, 1913
Subtribe Poecilonotina Jakobson, 1913
Subtribe Nesotrinchina Bílý, Kubáň and Volkovitsh, 2009
Tribe Sphenopterini Lacordaire, 1857
Tribe Vadonaxiini Descarpentries, 1970
Subfamily Buprestinae Leach, 1815
Tribe Actenodini Gistel, 1848
Tribe Anthaxiini Gory and Laporte, 1839
Tribe Bubastini Obenberger, 1920
Tribe Buprestini Leach, 1815
Subtribe Agaeocerina Nelson, 1982
Subtribe Buprestina Leach, 1815
Subtribe Lamprocheilina Holyński, 1993
Subtribe Trachykelina Holyński, 1988
Tribe Chrysobothrini Gory and Laporte, 1836
Tribe Coomaniellini Bílý, 1974
Tribe Curidini Holyński, 1988
Subtribe Anilarina Bílý, 2000
Subtribe Curidina Holyński, 1988
Subtribe Neocuridina Holyński, 1988
Tribe Epistomentini Levey, 1978
Tribe Exagistini Tôyama, 1987
Tribe Julodimorphini Kerremans, 1903
Tribe Kisanthobiini Richter, 1949
Tribe Maoraxiini Holyński, 1984
Tribe Melanophilini Bedel, 1921
Tribe Melobaseini Bílý, 2000
Tribe Mendizabaliini Cobos, 1968
Tribe Nascionini Holyński, 1988
Tribe Phrixiini Cobos, 1975
Tribe Pterobothrini Volkovitsh, 2001
Tribe Stigmoderini Lacordaire, 1857
Tribe Thomassetiini Bellamy, 1987
Subtribe Philanthaxiina Holyński, 1988
Subtribe Thomassetiina Bellamy, 1987
Tribe Trigonogeniini Cobos, 1956
Tribe Xenorhipidini Cobos, 1986
Subtribe Trichinorhipidina Bellamy, 2006
Subtribe Xenorhipidina Cobos, 1986
Subfamily Agrilinae Laporte, 1835
Tribe Agrilini Laporte, 1835
Subtribe Agrilina Laporte, 1835
Subtribe Amorphosternina Cobos, 1974
Subtribe Amyiina Holyński, 1993
Subtribe Rhaeboscelidina Cobos, 1976
Tribe Aphanisticini Jacquelin du Val, 1859
Subtribe Anthaxomorphina Holyński, 1993
Subtribe Aphanisticina Jacquelin du Val, 1859
Subtribe Cylindromorphina Portevin, 1931
Subtribe Cylindromorphoidina Cobos, 1979
Subtribe Germaricina Cobos, 1979
Tribe Coraebini Bedel, 1921
Subtribe Amorphosomatina Majer, 2000
Subtribe Cisseina Majer, 2000
Subtribe Clematina Majer, 2000
Subtribe Coraebina Bedel, 1921
Subtribe Dismorphina Cobos, 1990
Subtribe Ethoniina Majer, 2000
Subtribe Geraliina Cobos, 1988
Subtribe Meliboeina Majer, 2000
Subtribe Synechocerina Majer, 2000
Subtribe Toxoscelina Majer, 2000
Tribe Tracheini Laporte, 1835
Subtribe Brachina LeConte, 1861
Subtribe Leiopleurina Holyński, 1993
Subtribe Pachyschelina Böving and Craighead, 1931
Subtribe Tracheina Laporte, 1835
†Subfamily Parathyreinae Alexeev, 1994
Superfamily Byrrhoidea Latreille, 1804
Family Byrrhidae Latreille, 1804
Subfamily Byrrhinae Latreille, 1804
Tribe Byrrhini Latreille, 1804
Tribe Exomellini Casey, 1914
Tribe Morychini El Moursy, 1961
Tribe Pedilophorini Casey, 1912
Tribe Simplocariini Mulsant and Rey, 1869
Subfamily Syncalyptinae Mulsant and Rey, 1869
Tribe Microchaetini Paulus, 1973
Tribe Syncalyptini Mulsant and Rey, 1869
Subfamily Amphicyrtinae LeConte, 1861
Family Elmidae Curtis, 1830
Subfamily Larainae LeConte, 1861
Tribe Laraini LeConte, 1861
Tribe Potamophilini Mulsant and Rey, 1872
Subfamily Elminae Curtis, 1830
Tribe Ancyronychini Ganglbauer, 1904
Tribe Elmini Curtis, 1830
Subtribe Elmina Curtis, 1830
Subtribe Stenelmina Mulsant and Rey, 1872
Tribe Macronychini Gistel, 1848
Family Dryopidae Billberg, 1820 (1817)
Family Lutrochidae Kasap and Crowson, 1975
Family Limnichidae Erichson, 1846
Subfamily Hyphalinae Britton, 1971
Subfamily Limnichinae Erichson, 1846
Tribe Limnichini Erichson, 1846
Tribe Wooldridgeini Spangler, 1999
Subfamily Cephalobyrrhinae Champion, 1925
Subfamily Thaumastodinae Champion, 1924
Family Heteroceridae MacLeay, 1825
Subfamily Elythomerinae Pacheco, 1964
Subfamily Heterocerinae MacLeay, 1825
Tribe Augylini Pacheco, 1964
Tribe Heterocerini MacLeay, 1825
Tribe Micilini Pacheco, 1964
Tribe Tropicini Pacheco, 1964
Family Psephenidae Lacordaire, 1854
Subfamily Afroeubriinae Lee, Satô, Shepard and Jäch, 2007
Subfamily Eubriinae Lacordaire, 1857
Subfamily Eubrianacinae Jakobson, 1913
Subfamily Psephenoidinae Bollow, 1938
Subfamily Psepheninae Lacordaire, 1854
Family Cneoglossidae Champion, 1897
Family Ptilodactylidae Laporte, 1836
Subfamily Anchytarsinae Champion, 1897
Subfamily Cladotominae Pic, 1914
Subfamily Aploglossinae Champion, 1897
Subfamily Araeopidiinae Lawrence, 1991
Subfamily Ptilodactylinae Laporte, 1836
Family Podabrocephalidae Pic, 1930
Family Chelonariidae Blanchard, 1845
Family Eulichadidae Crowson, 1973
Family Callirhipidae Emden, 1924
Superfamily Elateroidea Leach, 1815
Family Rhinorhipidae Lawrence, 1988
Family Artematopodidae Lacordaire, 1857
Subfamily Electribiinae Crowson, 1975
Subfamily Allopogoniinae Crowson, 1973
Subfamily Artematopodinae Lacordaire, 1857
Tribe Artematopodini Lacordaire, 1857
Tribe Ctesibiini Crowson, 1973
Tribe Macropogonini LeConte, 1861
Family Brachypsectridae LeConte and Horn, 1883
Family Cerophytidae Latreille, 1834
Family Eucnemidae Eschscholtz, 1829
Subfamily Perothopinae Lacordaire, 1857
Subfamily Phyllocerinae Reitter, 1905
Tribe Anelastini Reitter, 1911
Tribe Phyllocerini Reitter, 1905
Subfamily Pseudomeninae Muona, 1993
Tribe Pseudomenini Muona, 1993
Tribe Schizophilini Muona, 1993
Subfamily Palaeoxeninae Muona, 1993
Subfamily Phlegoninae Muona, 1993
Subfamily Anischiinae Fleutiaux, 1936
Subfamily Melasinae Fleming, 1821
Tribe Calyptocerini Muona, 1993
Tribe Ceballosmelasini Muona, 1993
Tribe Dirhagini Reitter, 1911
Tribe Epiphanini Muona, 1993
Tribe Hylocharini Jacquelin du Val, 1859
Tribe Melasini Fleming, 1821
Subtribe Compsocnemina Muona, 1993
Subtribe Melasina Fleming, 1821
Tribe Neocharini Muona, 1993
Tribe Xylobiini Reitter, 1911
Subfamily Eucneminae Eschscholtz, 1829
Tribe Dendrocharini Fleutiaux, 1920
Tribe Dyscharachthini Muona, 1993
Tribe Entomosatopini Muona, 1993
Tribe Eucnemini Eschscholtz, 1829
Tribe Galbitini Muona, 1991
Tribe Mesogenini Muona, 1993
Tribe Muonajini Özdikmen, 2008
Tribe Perrotiini Muona, 1993
Tribe Phaenocerini Muona, 1993
Tribe Proutianini Muona, 1993
Subfamily Macraulacinae Fleutiaux, 1923
Tribe Anelastidini Muona, 1993
Tribe Echthrogasterini Cobos, 1965
Tribe Euryptychini Mamaev, 1976
Tribe Jenibuntorini Muona, 1993
Tribe Macraulacini Fleutiaux, 1923
Tribe Nematodini Leiler, 1976
Tribe Oisocerini Muona, 1993
Tribe Orodotini Muona, 1993
†Tribe Throscogeniini Iablokoff-Khnzorian, 1962
Family Throscidae Laporte, 1840 nomen protectum
†Family Praelateriidae Dolin, 1973
Family Elateridae Leach, 1815
Subfamily Cebrioninae Latreille, 1802
Subfamily Agrypninae Candèze, 1857 nomen protectum
Tribe Agrypnini Candèze, 1857 nomen protectum
Tribe Anaissini Golbach, 1984
†Tribe Cryptocardiini Dolin, 1980
Tribe Euplinthini Costa, 1975
Subtribe Cleidecostina Johnson, 2002
Subtribe Compsoplinthina Costa, 1975
Subtribe Euplinthina Costa, 1975
Tribe Hemirhipini Candèze, 1857
Tribe Oophorini Gistel, 1848
Tribe Platycrepidiini Costa and Casari-Chen, 1993
Tribe Pseudomelanactini Arnett, 1967
Tribe Pyrophorini Candèze, 1863
Subtribe Hapsodrilina Costa, 1975
Subtribe Nyctophyxina Costa, 1975
Subtribe Pyrophorina Candèze, 1863
Tribe Tetralobini Laporte, 1840
Subfamily Thylacosterninae Fleutiaux, 1920
Subfamily Lissominae Laporte, 1835
Subfamily Semiotinae Jakobson, 1913
Subfamily Campyloxeninae Costa, 1975
Subfamily Pityobiinae Hyslop, 1917
Subfamily Oxynopterinae Candèze, 1857
Subfamily Dendrometrinae Gistel, 1848
Tribe Crepidomenini Candèze, 1863
Tribe Dendrometrini Gistel, 1848
Subtribe Dendrometrina Gistel, 1848
Subtribe Denticollina Stein and Weise, 1877 (1848)
Subtribe Hemicrepidiina Champion, 1896
Tribe Dimini Candèze, 1863
Tribe Hypnoidini Schwarz, 1906 (1860)
Tribe Pleonomini Semenov and Pjatakova, 1936
Tribe Prosternini Gistel, 1856 nomen protectum
Tribe Senodoniini Schenkling, 1927
Subfamily Negastriinae Nakane and Kishii, 1956
Tribe Negastriini Nakane and Kishii, 1956
Tribe Quasimusini Schimmel and Tarnawski, 2009
Subtribe Loebliquasimusina Schimmel and Tarnawski, 2009
Subtribe Quasimusina Schimmel and Tarnawski, 2009
Subtribe Striatoquasimusina Schimmel and Tarnawski, 2009
Subtribe Wittmeroquasimusina Schimmel and Tarnawski, 2009
Subfamily Elaterinae Leach, 1815
Tribe Agriotini Laporte, 1840
Subtribe Agriotina Laporte, 1840
Subtribe Cardiorhinina Candèze, 1863
Tribe Ampedini Gistel, 1848
Tribe Dicrepidiini Thomson, 1858
Tribe Elaterini Leach, 1815
Tribe Megapenthini Gurjeva, 1973
Tribe Melanotini Candèze, 1859 (1848)
Tribe Odontonychini Girard, 1973
Tribe Physorhinini Candèze, 1859
Tribe Pomachiliini Candèze, 1859
Tribe Synaptini Gistel, 1856
Subfamily Cardiophorinae Candèze, 1859
Subfamily Hemiopinae Fleutiaux, 1941
Subfamily Physodactylinae Lacordaire, 1857
Subfamily Eudicronychinae Girard, 1971
Subfamily Subprotelaterinae Fleutiaux, 1920
Subfamily Morostomatinae Dolin, 2000
†Subfamily Protagrypninae Dolin, 1973
†Tribe Desmatini Dolin, 1975
†Tribe Hypnomorphini Dolin, 1975
†Tribe Protagrypnini Dolin, 1973
Family Plastoceridae Crowson, 1972
Family Drilidae Blanchard, 1845
Subfamily Drilinae Blanchard, 1845
Subfamily Thilmaninae Kazantsev, 2004
Tribe Euanomini Kazantsev, 2010
Tribe Thilmanini Kazantsev, 2004
Family Omalisidae Lacordaire, 1857
†Family Berendtimiridae Winkler, 1987
Family Lycidae Laporte, 1836
Subfamily Libnetinae Bocák and Bocáková, 1990
Subfamily Dictyopterinae Houlbert, 1922
Tribe Dictyopterini Houlbert, 1922
Tribe Lycoprogenthini Bocák and Bocáková, 2008
Tribe Taphini Bocák and Bocáková, 1990
Subfamily Lyropaeinae Bocák and Bocáková, 1989
Tribe Alyculini Bocák and Bocáková, 2008
Tribe Antennolycini Bocák and Bocáková, 2008
Tribe Lyropaeini Bocák and Bocáková, 1989
Tribe Miniduliticolini Kazantsev, 2003
Tribe Platerodrilini Kazantsev, 2004
Subfamily Ateliinae Kleine, 1929
Tribe Ateliini Kleine, 1929
Tribe Dilophotini Kleine, 1929
Subfamily Lycinae Laporte, 1836
Tribe Calochromini Lacordaire, 1857
Tribe Calopterini Green, 1949
Subtribe Acroleptina Bocáková, 2005
Subtribe Calopterina Green, 1949
Tribe Conderini Bocák and Bocáková, 1990
Tribe Dihammatini Bocák and Bocáková, 2008
Tribe Erotini LeConte, 1881
Tribe Eurrhacini Bocáková, 2005
Tribe Leptolycini Leng and Mutchler, 1922
Tribe Lycini Laporte, 1836
Tribe Lyponiini Bocák and Bocáková, 1990
Tribe Macrolycini Kleine, 1929
Tribe Melanerotini Kazantsev, 2010
Tribe Metriorrhynchini Kleine, 1926
Subtribe Hemiconderinina Bocák and Bocáková, 1990
Subtribe Metriorrhynchina Kleine, 1926
Subtribe Trichalina Kleine, 1929
Tribe Platerodini Kleine, 1929
Tribe Slipinskiini Bocák and Bocáková, 1992
Tribe Thonalmini Kleine, 1933
Subfamily Dexorinae Bocák and Bocáková, 1989
Family Telegeusidae Leng, 1920
Family Phengodidae LeConte, 1861
Subfamily Phengodinae LeConte, 1861
Subfamily Mastinocerinae LeConte, 1881
Subfamily Penicillophorinae Paulus, 1975
Family Rhagophthalmidae Olivier, 1907
Family Lampyridae Rafinesque, 1815
Subfamily Psilocladinae McDermott, 1964
Subfamily Amydetinae Olivier, 1907
Tribe Amydetini Olivier, 1907
Tribe Vestini McDermott, 1964
Subfamily Lampyrinae Rafinesque, 1815
Tribe Cratomorphini Green, 1948
Tribe Lamprocerini Olivier, 1907
Tribe Lamprohizini Kazantsev, 2010
Tribe Lampyrini Rafinesque, 1815
Tribe Lucidotini Lacordaire, 1857
Subtribe Dadophorina Olivier, 1907
Subtribe Lamprigerina McDermott, 1964
Subtribe Lucidotina Lacordaire, 1857
Subtribe Photinina LeConte, 1881
Tribe Pleotomini Summers, 1874
Subfamily Luciolinae Lacordaire, 1857
Tribe Curtosini McDermott, 1964
Tribe Luciolini Lacordaire, 1857
Subfamily Photurinae Lacordaire, 1857
Family Omethidae LeConte, 1861
Subfamily Omethinae LeConte, 1861
Subfamily Matheteinae LeConte, 1881
Subfamily Driloniinae Crowson, 1972
Family Cantharidae Imhoff, 1856 (1815)
Subfamily Cantharinae Imhoff, 1856 (1815)
Tribe Cantharini Imhoff, 1856 (1815)
Tribe Podabrini Gistel, 1856
Subfamily Silinae Mulsant, 1862
Tribe Silini Mulsant, 1862
Tribe Tytthonyxini Arnett, 1962
Subfamily Dysmorphocerinae Brancucci, 1980
Subfamily Malthininae Kiesenwetter, 1852
Tribe Malchinini Brancucci, 1980
Tribe Malthinini Kiesenwetter, 1852
Tribe Malthodini Böving and Craighead, 1931
Subfamily Chauliognathinae LeConte, 1861
Tribe Chauliognathini LeConte, 1861
Tribe Ichthyurini Champion, 1915
Subfamily Cydistinae Paulus, 1972
Subfamily Pterotinae LeConte, 1861
Subfamily Ototretinae McDermott, 1964
Subfamily Ototretadrilinae Crowson, 1972
†Subfamily Lasiosynidae Kirejtshuk, Chang, Ren and Kun, 2010
Series DERODONTIFORMIA
Superfamily Derodontoidea LeConte, 1861
Family Derodontidae LeConte, 1861
Subfamily Peltasticinae LeConte, 1861
Subfamily Derodontinae LeConte, 1861
Subfamily Laricobiinae Mulsant and Rey, 1864
Family Nosodendridae Erichson, 1846
Family Jacobsoniidae Heller, 1926
Series BOSTRICHIFORMIA
Superfamily Bostrichoidea Latreille, 1802
Family Dermestidae Latreille, 1804
Subfamily Dermestinae Latreille, 1804
Tribe Dermestini Latreille, 1804
Tribe Marioutini Jakobson, 1913
Subfamily Thorictinae Agassiz, 1846
Tribe Thaumaphrastini Anderson, 1949
Tribe Thorictini Agassiz, 1846
Subfamily Orphilinae LeConte, 1861
Subfamily Trinodinae Casey, 1900
†Tribe Cretonodini Kirejtshuk and Azar, 2009
Tribe Thylodriini Semenov, 1909
Tribe Trinodini Casey, 1900
Tribe Trinoparvini Háva, 2010
Subfamily Attageninae Laporte, 1840
Tribe Attagenini Laporte, 1840
Tribe Egidyellini Semenov, 1914
Subfamily Megatominae Leach, 1815
Tribe Anthrenini Gistel, 1848
Tribe Megatomini Leach, 1815
Family Endecatomidae LeConte, 1861
Family Bostrichidae Latreille, 1802
Subfamily Dysidinae Lesne, 1921
Subfamily Polycaoninae Lesne, 1896
Subfamily Bostrichinae Latreille, 1802
Tribe Apatini Billberg, 1820
Tribe Bostrichini Latreille, 1802
Tribe Dinapatini Lesne, 1910
Tribe Sinoxylini Marseul, 1857
Tribe Xyloperthini Lesne, 1921
Subfamily Psoinae Blanchard, 1851
Subfamily Dinoderinae Thomson, 1863
Subfamily Lyctinae Billberg, 1820
Tribe Lyctini Billberg, 1820
Tribe Trogoxylini Lesne, 1921
Subfamily Euderiinae Lesne, 1934
Family Ptinidae Latreille, 1802
Subfamily Eucradinae LeConte, 1861
Tribe Eucradini LeConte, 1861
Tribe Hedobiini Mulsant and Rey, 1868
Subfamily Ptininae Latreille, 1802
Tribe Gibbiini Jacquelin du Val, 1860
Tribe Meziini Bellés, 1985
Tribe Ptinini Latreille, 1802
Tribe Sphaericini Portevin, 1931
Subfamily Dryophilinae Gistel, 1848
Tribe Dryophilini Gistel, 1848
Tribe Ptilineurini Böving, 1927
Subfamily Ernobiinae Pic, 1912
Subfamily Anobiinae Fleming, 1821
Subfamily Ptilininae Shuckard, 1839
Subfamily Alvarenganiellinae Viana and Martínez, 1971
Subfamily Xyletininae Gistel, 1848
Tribe Lasiodermini Böving, 1927
Tribe Metholcini Zahradník, 2009
Tribe Xyletinini Gistel, 1848
Subfamily Dorcatominae Thomson, 1859
Subfamily Mesocoelopodinae Mulsant and Rey, 1864
Tribe Tricorynini White, 1971
Tribe Mesocoelopodini Mulsant and Rey, 1864
Series CUCUJIFORMIA
Superfamily Lymexyloidea Fleming, 1821
Family Lymexylidae Fleming, 1821
Subfamily Hylecoetinae Germar, 1818
Subfamily Lymexylinae Fleming, 1821
Subfamily Atractocerinae Laporte, 1840
Subfamily Melittommatinae Wheeler, 1986
Superfamily Cleroidea Latreille, 1802
Family Phloiophilidae Kiesenwetter, 1863
Family Trogossitidae Latreille, 1802
Subfamily Peltinae Latreille, 1806
Tribe Ancyronini Kolibáč, 2006
Tribe Colydiopeltini Kolibáč, 2006
Tribe Decamerini Crowson, 1964
Tribe Lophocaterini Crowson, 1964 nomen protectum
Tribe Peltini Latreille, 1806
Tribe Thymalini Léveillé, 1888
Subfamily Trogossitinae Latreille, 1802
Tribe Calityini Reitter, 1922
Tribe Egoliini Lacordaire, 1854
Tribe Gymnochilini Lacordaire, 1854
Tribe Larinotini Ślipiński, 1992
†Tribe Lithostomatini Kolibáč and Huang, 2008
Tribe Trogossitini Latreille, 1802
Family Chaetosomatidae Crowson, 1952
Family Metaxinidae Kolibáč, 2004
Family Thanerocleridae Chapin, 1924
Subfamily Zenodosinae Kolibáč, 1992
Subfamily Thaneroclerinae Chapin, 1924
Tribe Isoclerini Kolibáč, 1992
Tribe Thaneroclerini Chapin, 1924
Tribe Viticlerini Winkler, 1982
Family Cleridae Latreille, 1802
Subfamily Tillinae Fischer von Waldheim, 1813
Subfamily Hydnocerinae Spinola, 1844
Tribe Callimerini Kolibáč, 1998
Tribe Hydnocerini Spinola, 1844
Tribe Lemidiini Kolibáč, 1998
Subfamily Clerinae Latreille, 1802
Subfamily Korynetinae Laporte, 1836
Family Acanthocnemidae Crowson, 1964
Family Phycosecidae Crowson, 1952
Family Prionoceridae Lacordaire, 1857
Tribe Lobonychini Majer, 1987
Tribe Prionocerini Lacordaire, 1857
Family Mauroniscidae Majer, 1995
Family Melyridae Leach, 1815
Subfamily Rhadalinae LeConte, 1861
Subfamily Melyrinae Leach, 1815
Tribe Arthrobrachini Majer, 1987
Tribe Astylini Pic, 1929
Tribe Cerallini Pic, 1929
Tribe Melyrini Leach, 1815
Subfamily Dasytinae Laporte, 1840
Tribe Chaetomalachiini Majer, 1987
Tribe Danaceini Thomson, 1859
Tribe Dasytini Laporte, 1840
Tribe Gietellini Constantin and Menier, 1987
Tribe Listrini Majer, 1990
Subfamily Malachiinae Fleming, 1821
Tribe Amalthocini Majer, 2002
Tribe Attalomimini Majer, 1995
Tribe Carphurini Champion, 1923
Tribe Lemphini Wittmer, 1976
Tribe Malachiini Fleming, 1821
Tribe Pagurodactylini Constantin, 2001
Superfamily Cucujoidea Latreille, 1802
†Family Parandrexidae Kirejtshuk, 1994
†Family Sinisilvanidae Hong, 2002
Family Boganiidae Sen Gupta and Crowson, 1966
Subfamily Paracucujinae Endrödy-Younga and Crowson, 1986
Subfamily Boganiinae Sen Gupta and Crowson, 1966
Family Byturidae Gistel, 1848
Subfamily Platydascillinae Pic, 1914
Subfamily Byturinae Gistel, 1848
Family Helotidae Chapuis, 1876
Family Protocucujidae Crowson, 1954
Family Sphindidae Jacquelin du Val, 1860
Subfamily Protosphindinae Sen Gupta and Crowson, 1979
Subfamily Odontosphindinae Sen Gupta and Crowson, 1979
Subfamily Sphindiphorinae Sen Gupta and Crowson, 1979
Subfamily Sphindinae Jacquelin du Val, 1860
Family Biphyllidae LeConte, 1861
Family Erotylidae Latreille, 1802
Subfamily Xenoscelinae Ganglbauer, 1899
Subfamily Pharaxonothinae Crowson, 1952
Subfamily Loberinae Bruce, 1951
Subfamily Languriinae Hope, 1840
Tribe Hapalipini Leschen, 2003
Tribe Languriini Hope, 1840
Tribe Thallisellini Sen Gupta, 1968
Subfamily Cryptophilinae Casey, 1900
Tribe Cryptophilini Casey, 1900
Tribe Empocryptini Leschen, 2003
Tribe Toramini Sen Gupta, 1967
Subfamily Erotylinae Latreille, 1802
Tribe Dacnini Gistel, 1848
Tribe Encaustini Crotch, 1876
Tribe Erotylini Latreille, 1802
Tribe Megalodacnini Sen Gupta, 1970
Tribe Tritomini Curtis, 1834
Family Monotomidae Laporte, 1840
Subfamily Rhizophaginae Redtenbacher, 1845
Subfamily Monotominae Laporte, 1840
Tribe Europini Sen Gupta, 1988
Tribe Lenacini Crowson, 1952
Tribe Monotomini Laporte, 1840
†Tribe Rhizophtomini Kirejtshuk and Azar, 2009
Tribe Thionini Crowson, 1952
Family Hobartiidae Sen Gupta and Crowson, 1966
Family Cryptophagidae Kirby, 1826
Subfamily Cryptophaginae Kirby, 1826
Tribe Caenoscelini Casey, 1900
Tribe Cryptophagini Kirby, 1826
Tribe Picrotini Crowson, 1980
Subfamily Atomariinae LeConte, 1861
Tribe Atomariini LeConte, 1861
Tribe Cryptafricini Leschen, 1996
Tribe Hypocoprini Reitter, 1879
Family Agapythidae Sen Gupta and Crowson, 1969
Family Priasilphidae Crowson, 1973
Family Phloeostichidae Reitter, 1911
Family Silvanidae Kirby, 1837
Subfamily Brontinae Blanchard, 1845
Tribe Brontini Blanchard, 1845
Tribe Telephanini LeConte, 1861
Subfamily Silvaninae Kirby, 1837
Family Cucujidae Latreille, 1802
Family Myraboliidae Lawrence and Britton, 1991
Family Cavognathidae Sen Gupta and Crowson, 1966
Family Lamingtoniidae Sen Gupta and Crowson, 1969
Family Passandridae Blanchard, 1845
Family Phalacridae Leach, 1815
Subfamily Phaenocephalinae Matthews, 1899
Subfamily Phalacrinae Leach, 1815
Family Propalticidae Crowson, 1952
Family Laemophloeidae Ganglbauer, 1899
Family Tasmosalpingidae Lawrence and Britton, 1991
Family Cyclaxyridae Gimmel, Leschen and Ślipiński, 2009
Family Kateretida Kirby, 1837
Family Nitidulidae Latreille, 1802
Subfamily Calonecrinae Kirejtshuk, 1982
Subfamily Maynipeplinae Kirejtshuk, 1998
Subfamily Epuraeinae Kirejtshuk, 1986
Tribe Epuraeini Kirejtshuk, 1986
Tribe Taenioncini Kirejtshuk, 1998
Subfamily Carpophilinae Erichson, 1842
Subfamily Amphicrossinae Kirejtshuk, 1986
Subfamily Meligethinae Thomson, 1859
Subfamily Nitidulinae Latreille, 1802
Tribe Cychramini Gistel, 1848
Tribe Cychramptodini Kirejtshuk and Lawrence, 1992
Tribe Cyllodini Everts, 1898
Tribe Lawrencerosini Kirejtshuk, 1991
Tribe Mystropini Murray, 1864
Tribe Nitidulini Latreille, 1802
Subfamily Cillaeinae Kirejtshuk and Audisio, 1986
Subfamily Cryptarchinae Thomson, 1859
Tribe Arhinini Kirejtshuk, 1987
Tribe Cryptarchini Thomson, 1859
Tribe Eucalosphaerini Kirejtshuk, 1987
Tribe Platyarchini Kirejtshuk, 1998
Subfamily Cybocephalinae Jacquelin du Val, 1858
Family Smicripidae Horn, 1880
Family Bothrideridae Erichson, 1845
Subfamily Teredinae Seidlitz, 1888
Tribe Sosylopsini Dajoz, 1980
Tribe Sysolini Ślipiński and Pal, 1985
Tribe Teredini Seidlitz, 1888
Subfamily Xylariophilinae Pal and Lawrence, 1986
Subfamily Anommatinae Ganglbauer, 1899
Subfamily Bothriderinae Erichson, 1845
Family Cerylonidae Billberg, 1820
Subfamily Euxestinae Grouvelle, 1908
Subfamily Loeblioryloninae Ślipiński, 1990
Subfamily Ostomopsinae Sen Gupta and Crowson, 1973
Subfamily Murmidiinae Jacquelin du Val, 1858
Subfamily Ceryloninae Billberg, 1820
Family Alexiidae Imhoff, 1856
Family Discolomatidae Horn, 1878
Subfamily Notiophyginae Jakobson, 1915
Tribe Dystheamonini John, 1954
Tribe Notiophygini Jakobson, 1915
Tribe Pachyplacini John, 1954
Subfamily Discolomatinae Horn, 1878
Subfamily Aphanocephalinae Jakobson, 1904
Subfamily Cephalophaninae John, 1954
Subfamily Pondonatinae John, 1954
Family Endomychidae Leach, 1815
Subfamily Merophysiinae Seidlitz, 1872
Subfamily Pleganophorinae Jacquelin du Val, 1858
Subfamily Anamorphinae Strohecker, 1953
Subfamily Leiestinae Thomson, 1863
Subfamily Mycetaeinae Jacquelin du Val, 1857
Subfamily Eupsilobiinae Casey, 1895
Subfamily Xenomycetinae Strohecker, 1962
Subfamily Danascelinae Tomaszewska, 2000
Subfamily Endomychinae Leach, 1815
Subfamily Epipocinae Gorham, 1873
Subfamily Stenotarsinae Chapuis, 1876
Subfamily Lycoperdininae Bromhead, 1838
Family Coccinellidae Latreille, 1807
Subfamily Microweiseinae Leng, 1920
Tribe Microweiseini Leng, 1920
Tribe Serangiini Pope, 1962
Tribe Sukunahikonini Kamiya, 1960
Subfamily Coccinellinae Latreille, 1807
Tribe Argentipilosini Gordon and de Almeida, 1991
Tribe Aspidimerini Mulsant, 1850
Tribe Azyini Mulsant, 1850
Tribe Brachiacanthini Mulsant, 1850
Tribe Carinodulini Gordon, Pakaluk and Ślipiński, 1989
Tribe Cephaloscymnini Gordon, 1985
Tribe Chilocorini Mulsant, 1846
Tribe Chnoodini Mulsant, 1850
Tribe Coccidulini Mulsant, 1846
Tribe Coccinellini Latreille, 1807
Tribe Cranophorini Mulsant, 1850
Tribe Cryptognathini Mulsant, 1850
Tribe Cynegetini Thomson, 1866
Tribe Diomini Gordon, 1999
Tribe Epilachnini Mulsant, 1846
Tribe Epivertini Pang and Mao, 1979
Tribe Eremochilini Gordon and Vanderberg, 1987
Tribe Hyperaspidini Mulsant, 1846
Tribe Limnichopharini Miyatake, 1994
Tribe Monocorynini Miyatake, 1988
Tribe Noviini Mulsant, 1846
Tribe Ortaliini Mulsant, 1850
Tribe Oryssomini Gordon, 1974
Tribe Platynaspini Mulsant, 1846
Tribe Plotinini Miyatake, 1994
Tribe Poriini Mulsant, 1850
Tribe Scymnillini Casey, 1899
Tribe Scymnini Mulsant, 1846
Tribe Selvadiini Gordon, 1985
Tribe Shirozuellini Sasaji, 1967
Tribe Stethorini Dobzhansky, 1924
Tribe Sticholotidini Weise, 1901
Tribe Telsimiini Casey, 1899
Tribe Tetrabrachini Kapur, 1948
Family Corylophidae LeConte, 1852
Subfamily Periptyctinae Ślipiński, Lawrence and Tomaszewska, 2001
Subfamily Corylophinae LeConte, 1852
Tribe Aenigmaticini Casey, 1900
Tribe Cleidostethini Bowestead, Booth, Ślipiński and Lawrence, 2001
Tribe Corylophini LeConte, 1852
Tribe Foadiini Ślipiński, Tomaszewska and Lawrence, 2009
Tribe Orthoperini Jacquelin du Val, 1857
Tribe Parmulini Poey, 1854
Tribe Peltinodini Paulian, 1950
Tribe Rypobiini Paulian, 1950
Tribe Sericoderini Matthews, 1888
Tribe Teplinini Pakaluk, Ślipiński and Lawrence, 1994
Family Akalyptoischiidae Lord, Hartley, Lawrence, McHugh and Miller, 2010
Family Latridiidae Erichson, 1842
Subfamily Latridiinae Erichson, 1842
Subfamily Corticariinae Curtis, 1829
†Subfamily Tetrameropseinae Kirejtshuk and Azar, 2008
Superfamily Tenebrionoidea Latreille, 1802
Family Mycetophagidae Leach, 1815
Subfamily Esarcinae Reitter, 1882
Subfamily Mycetophaginae Leach, 1815
Tribe Mycetophagini Leach, 1815
Tribe Typhaeini Thomson, 1863
Subfamily Bergininae Leng, 1920
Family Archeocrypticidae Kaszab, 1964
Family Pterogeniidae Crowson, 1953
Family Ciidae Leach, 1819
Subfamily Sphindociinae Lawrence, 1974
Subfamily Ciinae Leach, 1819
Tribe Ciini Leach, 1819
Tribe Orophiini Thomson, 1863
Tribe Xylographellini Kawanabe and Miyatake, 1996
Subtribe Syncosmetina Lopes-Andrade, 2008
Subtribe Xylographellina Kawanabe and Miyatake, 1996
Family Tetratomidae Billberg, 1820
Subfamily Tetratominae Billberg, 1820
Subfamily Piseninae Miyatake, 1960
Subfamily Penthinae Lacordaire, 1859
Subfamily Hallomeninae Gistel, 1848
Subfamily Eustrophinae Gistel, 1848
Tribe Eustrophini Gistel, 1848
Tribe Holostrophini Nikitsky, 1998
Family Melandryidae Leach, 1815
Subfamily Melandryinae Leach, 1815
Tribe Anisoxiellini Nikitsky, 2007
Tribe Dircaeini Kirby, 1837
Tribe Hypulini Gistel, 1848
Tribe Melandryini Leach, 1815
Tribe Orchesiini Mulsant, 1856
Tribe Serropalpini Latreille, 1829
Tribe Xylitini Thomson, 1864
Tribe Zilorini Desbrochers des Loges, 1900
Subfamily Osphyinae Mulsant, 1856 (1839)
Family Mordellidae Latreille, 1802
Subfamily Ctenidiinae Franciscolo, 1951
Subfamily Mordellinae Latreille, 1802
Tribe Conaliini Ermisch, 1956
Tribe Mordellini Latreille, 1802
Tribe Mordellistenini Ermisch, 1941
Tribe Reynoldsiellini Franciscolo, 1957
Tribe Stenaliini Franciscolo, 1955
Family Ripiphoridae Gemminger, 1870 (1855)
Subfamily Ptilophorinae Gerstaecker, 1855
Subfamily Pelecotominae Seidlitz, 1875
Subfamily Hemirhipidiinae Heller, 1921
Subfamily Ripidiinae Gerstaecker, 1855
Tribe Eorhipidiini Iablokoff-Khnzorian, 1986
Tribe Ripidiini Gerstaecker, 1855
Subfamily Ripiphorinae Gemminger, 1870 (1855)
Tribe Macrosiagonini Heyden, 1908
Tribe Ripiphorini Gemminger, 1870 (1855)
Family Zopheridae Solier, 1834
Subfamily Colydiinae Billberg, 1820
Tribe Acropini Sharp, 1894
Tribe Adimerini Sharp, 1894
Tribe Colydiini Billberg, 1820
Tribe Gempylodini Sharp, 1893
Tribe Nematidiini Horn, 1878
Tribe Orthocerini Blanchard, 1845
Tribe Rhagoderini Horn, 1878
Tribe Rhopalocerini Reitter, 1911
Tribe Synchitini Erichson, 1845
Subfamily Zopherinae Solier, 1834
Tribe Latometini Ślipiński and Lawrence, 1999
Tribe Monommatini Blanchard, 1845
Tribe Phellopsini Ślipiński and Lawrence, 1999
Tribe Pycnomerini Erichson, 1845
Tribe Usechini Horn, 1867
Tribe Zopherini Solier, 1834
Family Ulodidae Pascoe, 1869
Family Promecheilidae Lacordaire, 1859
Family Chalcodryidae Watt, 1974
Family Trachelostenidae Lacordaire, 1859
Family Tenebrionidae Latreille, 1802
Subfamily Zolodininae Watt, 1975
Subfamily Lagriinae Latreille, 1825 (1820)
Tribe Adeliini Kirby, 1828
Tribe Belopini Reitter, 1917
Tribe Chaerodini Doyen, Matthews and Lawrence, 1990
Tribe Cossyphini Latreille, 1802
Tribe Goniaderini Lacordaire, 1859
Tribe Laenini Seidlitz, 1895
Tribe Lagriini Latreille, 1825 (1820)
Subtribe Lagriina Latreille, 1825 (1820)
Subtribe Statirina Blanchard, 1845
Tribe Lupropini Ardoin, 1958
Tribe Pycnocerini Lacordaire, 1859 nomen protectum
Subfamily Nilioninae Oken, 1843
Subfamily Phrenapatinae Solier, 1834
Tribe Archaeoglenini Watt, 1975
Tribe Penetini Lacordaire, 1859
Tribe Phrenapatini Solier, 1834
Subfamily Pimeliinae Latreille, 1802
Tribe Adelostomini Solier, 1834
Tribe Adesmiini Lacordaire, 1859 nomen protectum
Tribe Akidini Billberg, 1820
Tribe Anepsiini LeConte, 1862
Tribe Asidini Fleming, 1821
Tribe Boromorphini Skopin, 1978
Tribe Branchini LeConte, 1862
Tribe Caenocrypticini Koch, 1958
Tribe Ceratanisini Gebien, 1937
Tribe Cnemeplatiini Jacquelin du Val, 1861
Subtribe Actizetina Watt, 1992
Subtribe Cnemeplatiina Jacquelin du Val, 1861
Subtribe Rondoniellina Ferrer and Moragues, 2000
Subtribe Thorictosomatina Watt, 1992
Tribe Cnemodinini Gebien, 1910
Tribe Coniontini Waterhouse, 1858
Tribe Cossyphodini Wasmann, 1899
Subtribe Cossyphodina Wasmann, 1899
Subtribe Cossyphoditina Basilewsky, 1950
Subtribe Esemephina Steiner, 1980
Subtribe Paramellonina Andreae, 1961
Tribe Cryptochilini Solier, 1841
Subtribe Calognathina Lacordaire, 1859
Subtribe Cryptochilina Solier, 1841
Subtribe Homebiina Endrödy-Younga, 1989
Subtribe Horatomina Koch, 1955
Subtribe Vansoniina Koch, 1955
Tribe Cryptoglossini LeConte, 1862 nomen protectum
Tribe Edrotini Lacordaire, 1859
Tribe Elenophorini Solier, 1837
Tribe Epitragini Blanchard, 1845 nomen protectum
Tribe Erodiini Billberg, 1820 nomen protectum
Tribe Evaniosomini Lacordaire, 1859
Tribe Falsomycterini Gebien, 1910
Tribe Idisiini Medvedev, 1973
Tribe Klewariini Gebien, 1910
Tribe Kuhitangiini Medvedev, 1962
Tribe Lachnogyini Seidlitz, 1894
Subtribe Lachnodactylina Reitter, 1904
Subtribe Lachnogyina Seidlitz, 1894
Subtribe Netuschiliina Ferrer and Yvinec, 2004
Tribe Leptodini Lacordaire, 1859
Tribe Nycteliini Solier, 1834
Tribe Nyctoporini Lacordaire, 1859
Tribe Phrynocarenini Gebien, 1928
Tribe Physogasterini Lacordaire, 1859
Tribe Pimeliini Latreille, 1802
Tribe Praociini Eschscholtz, 1829
Tribe Sepidiini Eschscholtz, 1829
Subtribe Hypomelina Koch, 1955
Subtribe Molurina Solier, 1834
Subtribe Oxurina Koch, 1955
Subtribe Phanerotomeina Koch, 1958
Subtribe Sepidiina Eschscholtz, 1829
Subtribe Trachynotina Koch, 1955
Tribe Stenosini Schaum, 1859 (1834)
Tribe Tentyriini Eschscholtz, 1831
Tribe Thinobatini Lacordaire, 1859
Tribe Trilobocarini Lacordaire, 1859
Tribe Vacronini Gebien, 1910
Tribe Zophosini Solier, 1834
Subfamily Tenebrioninae Latreille, 1802
Tribe Acropteronini Doyen, 1989
Tribe Alphitobiini Reitter, 1917
Tribe Amarygmini Gistel, 1848
Tribe Amphidorini LeConte, 1862
Tribe Apocryphini Lacordaire, 1859
Tribe Blaptini Leach, 1815
Subtribe Blaptina Leach, 1815
Subtribe Gnaptorina Medvedev, 2001
Subtribe Gnaptorinina Medvedev, 2001
Subtribe Prosodina Skopin, 1960
Subtribe Remipedellina Semenov, 1907
Tribe Bolitophagini Kirby, 1837 nomen protectum
Tribe Centronopini Doyen, 1989
Tribe Cerenopini Horn, 1870
Tribe Dissonomini Medvedev, 1968
Tribe Eulabini Horn, 1870
Tribe Falsocossyphini Ferrer, 2006
Tribe Heleini Fleming, 1821
Subtribe Asphalina Matthews and Lawrence, 2005
Subtribe Cyphaleina Lacordaire, 1859
Subtribe Heleina Fleming, 1821
Tribe Helopini Latreille, 1802
Subtribe Helopina Latreille, 1802
Subtribe Cylindrinotina Español, 1956
Tribe Helopinini Lacordaire, 1859
Subtribe Aptilina Koch, 1958
Subtribe Helopinina Lacordaire, 1859
Subtribe Micrantereina Reitter, 1917
Subtribe Oncosomina Koch, 1958
Tribe Melanimonini Seidlitz, 1894 (1854)
Tribe Opatrini Brullé, 1832
Subtribe Heterocheirina Koch, 1956
Subtribe Heterotarsina Blanchard, 1845
Subtribe Opatrina Brullé, 1832
Subtribe Neopachypterina Bouchard, Löbl and Merkl, 2007
Tribe Palorini Matthews, 2003
Tribe Pedinini Eschscholtz, 1829
Subtribe Dendarina Mulsant and Rey, 1854
Subtribe Eurynotina Mulsant and Rey, 1854
Subtribe Leichenina Mulsant, 1854
Subtribe Loensina Koch, 1956
Subtribe Melambiina Mulsant and Rey, 1854
Subtribe Pedinina Eschscholtz, 1829
Subtribe Platynotina Mulsant and Rey, 1853
Subtribe Pythiopina Koch, 1953
Tribe Platyscelidini Lacordaire, 1859
Tribe Praeugenini De Moor, 1970
Tribe Rhysopaussini Wasmann, 1896
Tribe Scaurini Billberg, 1820
Tribe Scotobiini Solier, 1838
Tribe Tenebrionini Latreille, 1802
Tribe Titaenini Fauvel, 1905
Tribe Toxicini Oken, 1843
Subtribe Eudysantina Bouchard, Lawrence, Davies and Newton, 2005
Subtribe Nycteropina Lacordaire, 1859
Subtribe Toxicina Oken, 1843
Tribe Triboliini Gistel, 1848
Tribe Ulomini Blanchard, 1845
Subfamily Alleculinae Laporte, 1840
Tribe Alleculini Laporte, 1840
Subtribe Alleculina Laporte, 1840
Subtribe Gonoderina Seidlitz, 1896
Subtribe Mycetocharina Gistel, 1848
Subtribe Xystropodina Solier, 1835
Tribe Cteniopodini Solier, 1835
Subfamily Diaperinae Latreille, 1802
Tribe Crypticini Brullé, 1832
Tribe Diaperini Latreille, 1802
Subtribe Adelinina LeConte, 1862
Subtribe Diaperina Latreille, 1802
Tribe Ectychini Doyen, Matthews and Lawrence, 1990
Tribe Gnathidiini Gebien, 1921
Subtribe Anopidiina Jeannel and Paulian, 1945
Subtribe Gnathidiina Gebien, 1921
Tribe Hyociini Medvedev and Lawrence, 1982
Subtribe Brittonina Medvedev and Lawrence, 1986
Subtribe Hyociina Medvedev and Lawrence, 1982
Subtribe Uptonina Medvedev and Lawrence, 1986
Tribe Hypophlaeini Billberg, 1820
Tribe Leiochrinini Lewis, 1894
Tribe Myrmechixenini Jacquelin du Val, 1858
Tribe Phaleriini Blanchard, 1845
Tribe Scaphidemini Reitter, 1922
Tribe Trachyscelini Blanchard, 1845
Subfamily Stenochiinae Kirby, 1837
Tribe Cnodalonini Oken, 1843
Tribe Stenochiini Kirby, 1837
Tribe Talanini Champion, 1887 (1883)
Family Prostomidae Thomson, 1859
Family Synchroidae Lacordaire, 1859
Family Stenotrachelidae Thomson, 1859
Subfamily Stenotrachelinae Thomson, 1859
Subfamily Cephaloinae LeConte, 1862
Subfamily Nematoplinae LeConte, 1862
Subfamily Stoliinae Nikitsky, 1985
Family Oedemeridae Latreille, 1810
Subfamily Polypriinae Lawrence, 2005
Subfamily Calopodinae Costa, 1852 nomen protectum
Subfamily Oedemerinae Latreille, 1810
Tribe Asclerini Gistel, 1848
Tribe Ditylini Mulsant, 1858
Tribe Nacerdini Mulsant, 1858
Tribe Oedemerini Latreille, 1810
Tribe Stenostomatini Mulsant, 1858
Family Meloidae Gyllenhal, 1810
Subfamily Eleticinae Wellman, 1910
Tribe Derideini Wellman, 1910
Tribe Eleticini Wellman, 1910
Subtribe Eleticina Wellman, 1910
Subtribe Eospastina Selander, 1966
Tribe Ertlianini Selander, 1966
Tribe Spasticini Kaszab, 1959
Subtribe Anthicoxenina Selander, 1966
Subtribe Protomeloina Abdullah, 1965
Subtribe Spasticina Kaszab, 1959
Subtribe Xenospastina Selander, 1966
Subfamily Meloinae Gyllenhal, 1810
Tribe Cerocomini Leach, 1815
Tribe Epicautini Parker and Böving, 1924
Tribe Eupomphini LeConte, 1862
Tribe Lyttini Solier, 1851
Tribe Meloini Gyllenhal, 1810
Tribe Mylabrini Rafinesque, 1815
Tribe Pyrotini MacSwain, 1956
Subfamily Tetraonycinae Böving and Craighead, 1931
Subfamily Nemognathinae Laporte, 1840
Tribe Horiini Latreille, 1802
Tribe Nemognathini Laporte, 1840
Subtribe Nemognathina Laporte, 1840
Subtribe Zonitidina Mulsant, 1857
Subtribe Sitarina Mulsant, 1857
Tribe Stenoderini Selander, 1991
Family Mycteridae Oken, 1843
Subfamily Mycterinae Oken, 1843
Subfamily Eurypinae Thomson, 1860
Subfamily Hemipeplinae Lacordaire, 1854
Family Boridae Thomson, 1859
Subfamily Borinae Thomson, 1859
Subfamily Synercticinae Lawrence and Pollock, 1994
Family Trictenotomidae Blanchard, 1845
Family Pythidae Solier, 1834
Family Pyrochroidae Latreille, 1806
Subfamily Tydessinae Nikitsky, 1986
Subfamily Pilipalpinae Abdullah, 1964
Subfamily Pedilinae Lacordaire, 1859
Subfamily Pyrochroinae Latreille, 1806
Subfamily Agnathinae Lacordaire, 1859
Family Salpingidae Leach, 1815
Subfamily Othniinae LeConte, 1861
Subfamily Prostominiinae Grouvelle, 1914
Subfamily Agleninae Horn, 1878
Subfamily Inopeplinae Grouvelle, 1908
Subfamily Dacoderinae LeConte, 1862
Subfamily Aegialitinae LeConte, 1862
Subfamily Salpinginae Leach, 1815
Family Anthicidae Latreille, 1819
Subfamily Eurygeniinae LeConte, 1862
Tribe Eurygeniini LeConte, 1862
Tribe Ictistygnini Borchmann, 1936
Tribe Mitraelabrini Abdullah, 1969
Subfamily Macratriinae LeConte, 1862
Tribe Macratriini LeConte, 1862
†Tribe Camelomorphini Kirejtshuk and Azar, 2008
Subfamily Steropinae Jacquelin du Val, 1863
Subfamily Copobaeninae Abdullah, 1969
Subfamily Lemodinae Lawrence and Britton, 1991
Subfamily Tomoderinae Bonadona, 1961
Subfamily Anthicinae Latreille, 1819
Tribe Anthicini Latreille, 1819
Tribe Endomiini Kaszab, 1956
Tribe Formicomini Bonadona, 1974
Tribe Microhoriini Bonadona, 1974
Subfamily Notoxinae Stephens, 1829
Family Aderidae Csiki, 1909
Tribe Aderini Csiki, 1909
Subtribe Aderina Csiki, 1909
Subtribe Cnopina Báguena Corella, 1948
Subtribe Gompeliina Bouchard, 2011
Subtribe Syzetoninina Báguena Corella, 1948
Tribe Emelinini Báguena Corella, 1948
Tribe Euglenesini Seidlitz, 1875
Subtribe Euglenesina Seidlitz, 1875
Subtribe Pseudolotelina Báguena Corella, 1948
Tribe Phytobaenini Báguena Corella, 1948
Family Scraptiidae Gistel, 1848
Subfamily Scraptiinae Gistel, 1848
Tribe Allopodini Franciscolo, 1964
Tribe Scraptiini Gistel, 1848
Subfamily Anaspidinae Mulsant, 1856
Tribe Anaspidini Mulsant, 1856
Tribe Anaspimordini Franciscolo, 1954
Tribe Menuthianaspidini Franciscolo, 1972
Tribe Pentariini Franciscolo, 1954
Subfamily Lagrioidinae Abdullah and Abdullah, 1968
Subfamily Afreminae Levey, 1985
Subfamily Ischaliinae Blair, 1920
Superfamily Chrysomeloidea Latreille, 1802
Family Oxypeltidae Lacordaire, 1868
Family Vesperidae Mulsant, 1839
Subfamily Philinae Thomson, 1861
Subfamily Vesperinae Mulsant, 1839
Subfamily Anoplodermatinae Guérin-Méneville, 1840
Tribe Anoplodermatini Guérin-Méneville, 1840
Tribe Hypocephalini Blanchard, 1845
Tribe Mysteriini Prosen, 1960
Family Disteniidae Thomson, 1861
Tribe Cyrtonopini Gressitt, 1940
Tribe Disteniini Thomson, 1861
Tribe Dynamostini Lacordaire, 1868
Tribe Heteropalpini Villiers, 1961
Family Cerambycidae Latreille, 1802
Subfamily Parandrinae Blanchard, 1845
Tribe Erichsoniini Thomson, 1861
Tribe Parandrini Blanchard, 1845
Subfamily Prioninae Latreille, 1802
Tribe Acanthophorini Thomson, 1864
Tribe Aegosomatini Thomson, 1861
Tribe Anacolini Thomson, 1857
Tribe Cacoscelini Thomson, 1861
Tribe Callipogonini Thomson, 1861
Tribe Calocomini Galileo and Martins, 1993
Tribe Cantharocnemini Thomson, 1861
Tribe Ergatini Fairmaire, 1864
Tribe Eurypodini Gahan, 1906 (1868)
Tribe Hopliderini Thomson, 1864
Tribe Macrodontiini Thomson, 1861
Tribe Macrotomini Thomson, 1861
Subtribe Archetypina Lameere, 1912
Subtribe Basitoxina Lameere, 1912
Subtribe Macrotomina Thomson, 1861
Subtribe Platygnathina Gilmour, 1954
Subtribe Xixuthrina Lameere, 1912
Tribe Mallaspini Thomson, 1861
Tribe Mallodonini Thomson, 1861
Tribe Meroscelisini Thomson, 1861
Tribe Prionini Latreille, 1802
Tribe Remphanini Lacordaire, 1868
Tribe Solenopterini Lacordaire, 1868
Tribe Tereticini Lameere, 1913
Tribe Vesperoctenini Vives, 2005
Subfamily Lepturinae Latreille, 1802
Tribe Desmocerini Blanchard, 1845
Tribe Encyclopini LeConte, 1873
Tribe Lepturini Latreille, 1802
Tribe Oxymirini Danilevsky, 1997
Tribe Rhagiini Kirby, 1837
Tribe Rhamnusiini Sama, 2009
Tribe Teledapini Pascoe, 1871
Tribe Sachalinobiini Danilevsky, 2010
Tribe Xylosteini Reitter, 1913
Subfamily Spondylidinae Audinet-Serville, 1832
Tribe Anisarthrini Mamaev and Danilevsky, 1973
Tribe Asemini Thomson, 1861
Tribe Atimiini LeConte, 1873
Tribe Saphanini Gistel, 1848
Tribe Spondylidini Audinet-Serville, 1832
Subfamily Necydalinae Latreille, 1825
Subfamily Dorcasominae Lacordaire, 1868
Subfamily Apatophyseinae Lacordaire, 1869
Subfamily Cerambycinae Latreille, 1802
Tribe Acangassuini Galileo and Martins, 2001
Tribe Achrysonini Lacordaire, 1868
Tribe Agallissini LeConte, 1873
Tribe Alanizini Di Iorio, 2003
Tribe Anaglyptini Lacordaire, 1868 nomen protectum
Tribe Aphanasiini Lacordaire, 1868
Tribe Aphneopini Lacordaire, 1868
Tribe Auxesini Lepesme and Breuning, 1952
Tribe Basipterini Fragoso, Monné and Campos Seabra, 1987
Tribe Bimiini Lacordaire, 1868
Tribe Bothriospilini Lane, 1950
Tribe Brachypteromatini Sama, 2008
Tribe Callichromatini Swainson, 1840
Tribe Callidiini Kirby, 1837
Tribe Callidiopini Lacordaire, 1868
Tribe Cerambycini Latreille, 1802
Subtribe Cerambycina Latreille, 1802
Subtribe Sphallotrichina Martins and Monné, 2005
Tribe Certallini Fairmaire, 1864
Tribe Chlidonini Waterhouse, 1879
Tribe Cleomenini Lacordaire, 1868
Tribe Clytini Mulsant, 1839
Tribe Compsocerini Thomson, 1864
Tribe Coptommatini Lacordaire, 1869
Tribe Curiini LeConte, 1873
Tribe Deilini Fairmaire, 1864
Tribe Dejanirini Lacordaire, 1868
Tribe Diorini Lane, 1950
Tribe Distichocerini Pascoe, 1867
Tribe Dodecosini Aurivillius, 1912
Tribe Dryobiini Arnett, 1962 nomen protectum
Tribe Eburiini Blanchard, 1845
Tribe Ectenessini Martins, 1998
Tribe Elaphidiini Thomson, 1864
Tribe Eligmodermini Lacordaire, 1868
Tribe Erlandiini Aurivillius, 1912
Tribe Eroschemini Lacordaire, 1868
Tribe Eumichthini Linsley, 1940
Tribe Gahaniini Quentin and Villiers, 1969
Tribe Glaucytini Lacordaire, 1868
Tribe Graciliini Mulsant, 1839
Tribe Hesperophanini Mulsant, 1839
Subtribe Daramina Sama, 2008
Subtribe Hesperophanina Mulsant, 1839
Tribe Hesthesini Pascoe, 1867
Tribe Heteropsini Lacordaire, 1868 nomen protectum
Tribe Hexoplini Martins, 2006
Tribe Holopleurini Chemsak and Linsley, 1974
Tribe Holopterini Lacordaire, 1868
Tribe Hyboderini Linsley, 1940
Tribe Hylotrupini Zagajkevich, 1991
Tribe Ibidionini Thomson, 1861
Subtribe Compsina Martins and Galileo, 2007
Subtribe Ibidionina Thomson, 1861
Subtribe Tropidina Martins and Galileo, 2007
Tribe Ideratini Martins and Napp, 2009
Tribe Lissonotini Swainson, 1840
Tribe Luscosmodicini Martins, 2003
Tribe Lygrini Sama, 2008
Tribe Macronini Lacordaire, 1868
Tribe Megacoelini Quentin and Villiers, 1969
Tribe Methiini Thomson, 1860
Tribe Molorchini Gistel, 1848
Tribe Mythodini Lacordaire, 1868
Tribe Necydalopsini Lacordaire, 1868
Tribe Neocorini Martins, 2005
Tribe Neostenini Lacordaire, 1868
Tribe Obriini Mulsant, 1839
Tribe Ochyrini Pascoe, 1871
Tribe Oedenoderini Aurivillius, 1912
Tribe Oemini Lacordaire, 1868
Subtribe Methioidina Martins, 1997
Subtribe Oemina Lacordaire, 1868
Tribe Opsimini LeConte, 1873
Tribe Oxycoleini Martins and Galileo, 2003
Tribe Paraholopterini Martins, 1997
Tribe Phalotini Lacordaire, 1868
Tribe Phlyctaenodini Lacordaire, 1868
Tribe Phoracanthini Newman, 1840
Tribe Phyllarthriini Lepesme and Breuning, 1956
Tribe Piesarthriini McKeown, 1947
Tribe Piezocerini Lacordaire, 1868
Subtribe Haruspicina Martins, 1976
Subtribe Piezocerina Lacordaire, 1868
Tribe Platyarthrini Bates, 1870
Tribe Plectogasterini Quentin and Villiers, 1969
Tribe Plectromerini Nearns and Braham, 2008
Tribe Pleiarthrocerini Lane, 1950
Tribe Protaxini Gahan, 1906
Tribe Prothemini Lacordaire, 1868
Tribe Psebiini Lacordaire, 1868
Tribe Pseudocephalini Aurivillius, 1912 (1861)
Tribe Pseudolepturini Thomson, 1861
Tribe Psilomorphini Lacordaire, 1868
Tribe Pteroplatini Thomson, 1861
Tribe Pyrestini Lacordaire, 1868
Tribe Rhagiomorphini Newman, 1841
Tribe Rhinotragini Thomson, 1861
Tribe Rhopalophorini Blanchard, 1845
Tribe Rosaliini Fairmaire, 1864
Tribe Sestyrini Lacordaire, 1868
Tribe Smodicini Lacordaire, 1868
Tribe Spintheriini Lacordaire, 1869
Tribe Stenhomalini Miroshnikov, 1989
Tribe Stenoderini Pascoe, 1867
Tribe Stenopterini Gistel, 1848
Tribe Strongylurini Lacordaire, 1868
Tribe Tessarommatini Lacordaire, 1868
Tribe Thraniini Gahan, 1906
Tribe Thyrsiini Marinoni and Napp, 1984
Tribe Tillomorphini Lacordaire, 1868
Tribe Torneutini Thomson, 1861
Tribe Trachyderini Dupont, 1836
Subtribe Ancylocerina Thomson, 1864
Subtribe Trachyderina Dupont, 1836
Tribe Tragocerini Pascoe, 1867
Tribe Trichomesiini Aurivillius, 1912
Tribe Tropocalymmatini Lacordaire, 1868
Tribe Typhocesini Lacordaire, 1868
Tribe Unxiini Napp, 2007
Tribe Uracanthini Blanchard, 1853
Tribe Vesperellini Sama, 2008
Tribe Xystrocerini Blanchard, 1845
Subfamily Lamiinae Latreille, 1825
Tribe Acanthocinini Blanchard, 1845
Tribe Acanthoderini Thomson, 1860
Tribe Acmocerini Thomson, 1864
Tribe Acridocephalini Dillon and Dillon, 1959
Tribe Acrocinini Swainson, 1840
Tribe Aderpasini Breuning and Teocchi, 1978
Tribe Aerenicini Lacordaire, 1872
Tribe Agapanthiini Mulsant, 1839
Tribe Amphoecini Breuning, 1951
Tribe Ancitini Aurivillius, 1917
Tribe Ancylonotini Lacordaire, 1869
Tribe Anisocerini Thomson, 1860
Tribe Apomecynini Thomson, 1860
Tribe Astathini Thomson, 1864
Tribe Batocerini Thomson, 1864
Tribe Calliini Thomson, 1864
Tribe Ceroplesini Thomson, 1860
Subtribe Ceroplesina Thomson, 1860
Subtribe Crossotina Thomson, 1864
Tribe Cloniocerini Lacordaire, 1872
Tribe Colobotheini Thomson, 1860
Tribe Compsosomatini Thomson, 1857
Tribe Cyrtinini Thomson, 1864
Tribe Desmiphorini Thomson, 1860
Tribe Dorcadionini Swainson, 1840
Tribe Dorcaschematini Thomson, 1860
Tribe Elytracanthinini Bousquet, 2009
Tribe Enicodini Thomson, 1864
Tribe Eupromerini Galileo and Martins, 1995
Tribe Forsteriini Tippmann, 1960
Tribe Gnomini Thomson, 1860
Tribe Gyaritini Breuning, 1950
Tribe Heliolini Breuning, 1951
Tribe Hemilophini Thomson, 1868 nomen protectum
Tribe Homonoeini Thomson, 1864
Tribe Hyborhabdini Aurivillius, 1911
Tribe Lamiini Latreille, 1825
Tribe Laticraniini Lane, 1959
Tribe Mauesiini Lane, 1956
Tribe Megabasini Thomson, 1860
Tribe Mesosini Mulsant, 1839
Tribe Microcymaturini Breuning and Teocchi, 1985
Tribe Moneilemini Thomson, 1864
Tribe Monochamini Gistel, 1848
Tribe Morimonellini Lobanov, Danilevsky and Murzin, 1981
Tribe Morimopsini Lacordaire, 1869
Tribe Nyctimeniini Gressitt, 1951
Tribe Obereini Thomson, 1864
Tribe Oculariini Breuning, 1950
Tribe Onciderini Thomson, 1860
Tribe Oncideropsidini Aurivillius, 1922
Tribe Onocephalini Thomson, 1860
Tribe Onychogleneini Aurivillius, 1923
Tribe Parmenini Mulsant, 1839
Tribe Petrognathini Blanchard, 1845
Tribe Phacellini Lacordaire, 1872
Tribe Phantasini Kolbe, 1897
Tribe Phrynetini Thomson, 1864
Tribe Phymasternini Teocchi, 1989
Tribe Phytoeciini Mulsant, 1839
Tribe Pogonocherini Mulsant, 1839
Tribe Polyrhaphidini Thomson, 1860
Tribe Pretiliini Martins and Galileo, 1990
Tribe Proctocerini Aurivillius, 1922
Tribe Prosopocerini Thomson, 1864
Tribe Pteropliini Thomson, 1860
Tribe Rhodopinini Gressitt, 1951
Tribe Saperdini Mulsant, 1839
Tribe Stenobiini Breuning, 1950
Tribe Sternotomini Thomson, 1860
Tribe Tapeinini Thomson, 1857
Tribe Tetraopini Thomson, 1860
Tribe Tetraulaxini Breuning and Teocchi, 1977
Tribe Tetropini Portevin, 1927
Tribe Theocrini Lacordaire, 1872
Tribe Tmesisternini Blanchard, 1853
Tribe Tragocephalini Thomson, 1857
Tribe Xenicotelini Matsushita, 1933
Tribe Xenofreini Aurivillius, 1923
Tribe Xenoleini Lacordaire, 1872
Tribe Xylorhizini Lacordaire, 1872
Tribe Zygocerini Thomson, 1864
Family Megalopodidae Latreille, 1802
Subfamily Megalopodinae Latreille, 1802
Subfamily Palophaginae Kuschel and May, 1990
Subfamily Zeugophorinae Böving and Craighead, 1931
Family Orsodacnidae Thomson, 1859
Subfamily Orsodacninae Thomson, 1859
Subfamily Aulacoscelidinae Chapuis, 1874
Family Chrysomelidae Latreille, 1802
Subfamily Sagrinae Leach, 1815
Tribe Carpophagini Chapuis, 1874
Tribe Diaphanopsidini Monrós, 1958
Tribe Megamerini Chapuis, 1874
Tribe Sagrini Leach, 1815
Subfamily Bruchinae Latreille, 1802
Tribe Amblycerini Bridwell, 1932
Subtribe Amblycerina Bridwell, 1932
Subtribe Spermophagina Borowiec, 1987
Tribe Bruchini Latreille, 1802
Subtribe Acanthoscelidina Bridwell, 1946
Subtribe Bruchina Latreille, 1802
Subtribe Megacerina Bridwell, 1946
Tribe Eubaptini Bridwell, 1932
Tribe Kytorhinini Bridwell, 1932
Tribe Pachymerini Bridwell, 1929
Subtribe Caryedontina Bridwell, 1929
Subtribe Caryopemina Bridwell, 1929
Subtribe Pachymerina Bridwell, 1929
Tribe Rhaebini Blanchard, 1845
Subfamily Donaciinae Kirby, 1837
Tribe Donaciini Kirby, 1837
Tribe Haemoniini Chen, 1941
Tribe Plateumarini Böving, 1922
Subfamily Criocerinae Latreille, 1804
Tribe Criocerini Latreille, 1804
Tribe Lemini Gyllenhal, 1813
Tribe Pseudocriocerini Heinze, 1962
Subfamily Cassidinae Gyllenhal, 1813
Tribe Alurnini Chapuis, 1875
Tribe Anisoderini Chapuis, 1875
Tribe Aproidini Weise, 1911
Tribe Arescini Chapuis, 1875
Tribe Aspidimorphini Chapuis, 1875
Tribe Basiprionotini Gressitt, 1952 (1929)
Tribe Botryonopini Chapuis, 1875
Tribe Callispini Chapuis, 1875
Tribe Callohispini Uhmann, 1960
Tribe Cassidini Gyllenhal, 1813
Tribe Cephaloleiini Chapuis, 1875
Tribe Chalepini Weise, 1910
Tribe Coelaenomenoderini Weise, 1911
Tribe Cryptonychini Chapuis, 1875
Tribe Delocraniini Spaeth, 1929
Tribe Dorynotini Monrós and Viana, 1949 (1923)
Tribe Eugenysini Hincks, 1952
Tribe Eurispini Chapuis, 1875
Tribe Exothispini Weise, 1911
Tribe Goniocheniini Spaeth, 1942
Tribe Gonophorini Chapuis, 1875
Tribe Hemisphaerotini Monrós and Viana, 1951 (1929)
Tribe Hispini Gyllenhal, 1813
Tribe Hispoleptini Chapuis, 1875
Tribe Hybosispini Weise, 1910
Tribe Imatidiini Hope, 1840
Tribe Ischyrosonychini Chapuis, 1875
Tribe Leptispini Fairmaire, 1868
Tribe Mesomphaliini Hope, 1840
Tribe Nothosacanthini Gressitt, 1952 (1929)
Tribe Oediopalpini Monrós and Viana, 1947 (1910)
Tribe Omocerini Hincks, 1952 (1923)
Tribe Oncocephalini Chapuis, 1875
Tribe Promecothecini Chapuis, 1875
Tribe Prosopodontini Weise, 1910
Tribe Sceloenoplini Uhmann, 1930
Tribe Spilophorini Chapuis, 1875
Tribe Uroplatini Weise, 1910
Subfamily Chrysomelinae Latreille, 1802
Tribe Chrysomelini Latreille, 1802
Tribe Timarchini Motschulsky, 1860
Subfamily Galerucinae Latreille, 1802
Tribe Alticini Newman, 1834
Tribe Decarthrocerini Laboissière, 1937
Tribe Galerucini Latreille, 1802
Tribe Hylaspini Chapuis, 1875
Tribe Luperini Gistel, 1848
Tribe Metacyclini Chapuis, 1875
Tribe Oidini Laboissière, 1921 (1875)
Subfamily Lamprosomatinae Lacordaire, 1848
Tribe Lamprosomatini Lacordaire, 1848
Tribe Neochlamysini Monrós, 1959
Tribe Sphaerocharini Chapuis, 1874
Subfamily Cryptocephalinae Gyllenhal, 1813
Tribe Clytrini Kirby, 1837
Tribe Cryptocephalini Gyllenhal, 1813
Subtribe Achaenopina Chapuis, 1874
Subtribe Cryptocephalina Gyllenhal, 1813
Subtribe Monachulina Leng, 1920
Subtribe Pachybrachina Chapuis, 1874
Subtribe Stylosomina Chapuis, 1874
Tribe Fulcidacini Jakobson, 1924
Subfamily Eumolpinae Hope, 1840
Tribe Bromiini Baly, 1865 (1863)
Tribe Caryonodini Bechyné, 1951
Tribe Cubispini Monrós, 1954
Tribe Eumolpini Hope, 1840
Tribe Euryopini Chapuis, 1874
Tribe Habrophorini Bechyné and Špringlová de Bechyné, 1969
Tribe Hemydacnini Bechyné, 1951
Tribe Megascelidini Chapuis, 1874
Tribe Merodini Chapuis, 1874
Tribe Pygomolpini Bechyné, 1949
Tribe Rosiroiini Bechyné, 1950
Tribe Typophorini Baly, 1865
Subfamily Spilopyrinae Chapuis, 1874
Subfamily Synetinae LeConte and Horn, 1883
†Subfamily Protoscelidinae Medvedev, 1968
Superfamily Curculionoidea Latreille, 1802
Family Nemonychidae Bedel, 1882
Subfamily Nemonychinae Bedel, 1882
Subfamily Cimberidinae Gozis, 1882
Tribe Cimberidini Gozis, 1882
Tribe Doydirhynchini Pierce, 1916
†Tribe Kuschelomacrini Riedel, 2010
Subfamily Rhinorhynchinae Voss, 1922
Tribe Mecomacerini Kuschel, 1994
Subtribe Brarina Legalov, 2009
Subtribe Mecomacerina Kuschel, 1994
Tribe Rhinorhynchini Voss, 1922
†Subfamily Slonikinae Zherikhin, 1977
†Tribe Slonikini Zherikhin, 1977
†Tribe Ulyaniscini Legalov, 2009
†Subfamily Eccoptarthrinae Arnoldi, 1977
†Subfamily Brenthorrhininae Arnoldi, 1977
†Tribe Brenthorrhinini Arnoldi, 1977
†Tribe Brenthorrhinoidini Legalov, 2003
†Subfamily Distenorrhininae Arnoldi, 1977
†Subfamily Eobelinae Arnoldi, 1977
†Tribe Eobelini Arnoldi, 1977
†Subtribe Eobelina Arnoldi, 1977
†Subtribe Procurculionina Arnoldi, 1977
†Tribe Karataucarini Legalov, 2009
†Tribe Nanophydini Arnoldi, 1977
†Tribe Oxycorynoidini Arnoldi, 1977
†Tribe Probelini Legalov, 2009
†Subfamily Paleocartinae Legalov, 2003
†Tribe Nebrenthorrhinini Legalov, 2007
†Tribe Paleocartini Legalov, 2003
†Subfamily Metrioxenoidinae Legalov, 2009
†Subfamily Cretonemonychinae Gratshev and Legalov, 2009
†Subfamily Selengarhynchinae Gratshev and Legalov, 2009
Family Anthribidae Billberg, 1820
Subfamily Anthribinae Billberg, 1820
Tribe Anthribini Billberg, 1820
Tribe Basitropini Lacordaire, 1865
Tribe Corrhecerini Lacordaire, 1865
Tribe Cratoparini LeConte, 1876
†Tribe Cretanthribini Legalov, 2009
Tribe Decataphanini Lacordaire, 1865
Tribe Discotenini Lacordaire, 1865
Tribe Ecelonerini Lacordaire, 1865
Tribe Ischnocerini Lacordaire, 1865
Tribe Gymnognathini Valentine, 1960
Tribe Jordanthribini Morimoto, 1980
Tribe Mauiini Valentine, 1990
Tribe Mecocerini Lacordaire, 1865
Tribe Mycteini Morimoto, 1972
Tribe Ozotomerini Morimoto, 1972
Tribe Piesocorynini Valentine, 1960
Tribe Platyrhinini Bedel, 1882
Tribe Platystomini Pierce, 1916
Tribe Proscoporhinini Lacordaire, 1865
Tribe Ptychoderini Jekel, 1855
Tribe Sintorini Lacordaire, 1865
Tribe Stenocerini Kolbe, 1895
Tribe Tophoderini Lacordaire, 1865
Tribe Trigonorhinini Valentine, 1999
Tribe Tropiderini Lacordaire, 1865
Tribe Xenocerini Lacordaire, 1865
Tribe Xylinadini Lacordaire, 1865
Tribe Zygaenodini Lacordaire, 1865
Subfamily Choraginae Kirby, 1819
Tribe Apolectini Lacordaire, 1865
Tribe Araecerini Lacordaire, 1865
Tribe Cisanthribini Zimmerman, 1994
Tribe Choragini Kirby, 1819
Tribe Valenfriesiini Alonso-Zarazaga and Lyal, 1999
Tribe Xenorchestini Lacordaire, 1865
Subfamily Urodontinae Thomson, 1859
†Family Ulyanidae Zherikhin, 1993
Family Belidae Schönherr, 1826
Subfamily Belinae Schönherr, 1826
Tribe Agnesiotidini Zimmerman, 1994
Tribe Belini Schönherr, 1826
Subtribe Belina Schönherr, 1826
Subtribe Homalocerina Legalov, 2009
Tribe Pachyurini Kuschel, 1959
Subfamily Oxycoryninae Schönherr, 1840
Tribe Aglycyderini Wollaston, 1864
Tribe Alloxycorynini Legalov, 2009
Tribe Distenorrhinoidini Legalov, 2009
Tribe Metrioxenini Voss, 1953
Subtribe Afrocorynina Voss, 1957
Subtribe Metrioxenina Voss, 1953
Subtribe Zherichinixenina Legalov, 2009
Tribe Oxycorynini Schönherr, 1840
Subtribe Allocorynina Sharp, 1890
Subtribe Oxycorynina Schönherr, 1840
Subtribe Oxycraspedina Marvaldi and Oberprieler, 2006
Family Caridae Thompson, 1992
Subfamily Carinae Thompson, 1992
Subfamily Chilecarinae Legalov, 2009
Tribe Carodini Legalov, 2009
Tribe Chilecarini Legalov, 2009
†Subfamily Baissorhynchinae Zherikhin, 1993
Family Attelabidae Billberg, 1820
Subfamily Attelabinae Billberg, 1820
Tribe Attelabini Billberg, 1820
Subtribe Attelabina Billberg, 1820
Subtribe Euscelina Voss, 1925
Subtribe Euscelophilina Voss, 1925
Subtribe Henicolabina Legalov, 2007
Subtribe Himatolabina Legalov, 2003
Subtribe Hybolabina Voss, 1925
Subtribe Isolabina Legalov, 2007
Subtribe Lagenoderina Voss, 1925
Subtribe Lamprolabina Voss, 1925
Subtribe Metocalolabina Legalov, 2003
Subtribe Omolabina Legalov, 2003
Subtribe Paramecolabina Legalov, 2003
Subtribe Phialodina Legalov, 2003
Subtribe Phymatolabina Voss, 1925
Subtribe Phymatopsinina Legalov, 2003
Subtribe Pleurolabina Legalov, 2003
Tribe Euopini Voss, 1925
Tribe Pilolabini Voss, 1925
Subfamily Apoderinae Jekel, 1860
Tribe Apoderini Jekel, 1860
Tribe Clitostylini Voss, 1929
Subtribe Allapoderina Legalov, 2003
Subtribe Clitostylina Voss, 1929
Subtribe Pseudophrysina Legalov, 2003
Tribe Hoplapoderini Voss, 1926
Subtribe Afroapoderina Legalov, 2003
Subtribe Hoplapoderina Voss, 1926
Subtribe Paratomapoderina Legalov, 2003
Tribe Trachelophorini Voss, 1926
Subfamily Rhynchitinae Gistel, 1848
Tribe Auletini Desbrochers des Loges, 1908
Subtribe Auletina Desbrochers des Loges, 1908
Subtribe Auletobiina Legalov, 2001
Subtribe Guineauletina Legalov, 2003
Subtribe Mandelschtamiina Legalov, 2003
Subtribe Pseudauletina Voss, 1933
Subtribe Pseudomesauletina Legalov, 2003
Tribe Auletorhinini Voss, 1935
Tribe Byctiscini Voss, 1923
Subtribe Byctiscina Voss, 1923
Subtribe Listrobyctiscina Legalov, 2003
Subtribe Svetlanaebyctiscina Legalov, 2003
Tribe Cesauletini Legalov, 2003
Tribe Deporaini Voss, 1929
Subtribe Chonostropheina Morimoto, 1962
Subtribe Deporaina Voss, 1929
Tribe Minurini Legalov, 2003
Tribe Rhinocartini Voss, 1931
Tribe Rhynchitini Gistel, 1848
Subtribe Acritorrhynchitina Legalov, 2007
Subtribe Anisomerinina Legalov, 2003
Subtribe Eugnamptina Voss, 1930
Subtribe Lasiorhynchitina Legalov, 2003
Subtribe Perrhynchitina Legalov, 2003
Subtribe Rhynchitallina Legalov, 2003
Subtribe Rhynchitina Gistel, 1848
Subtribe Temnocerina Legalov, 2003
Subfamily Isotheinae Scudder, 1893
Tribe Isotheini Scudder, 1893
Subtribe Depasophilina Legalov, 2003
†Subtribe Isotheina Scudder, 1893
†Tribe Toxorhynchini Scudder, 1893
Subfamily Pterocolinae Lacordaire, 1865
Family Brentidae Billberg, 1820
Subfamily Brentinae Billberg, 1820
Tribe Brentini Billberg, 1820
Subtribe Arrhenodina Lacordaire, 1865
Subtribe Brentina Billberg, 1820
Subtribe Eremoxenina Semenov, 1892
Tribe Cyladini Schönherr, 1823
Tribe Cyphagogini Kolbe, 1892
Subtribe Atopobrentina Damoiseau, 1965
Subtribe Cyphagogina Kolbe, 1892
†Subtribe Dominibrentina Poinar, 2009
Subtribe Hoplopisthiina Senna and Calabresi, 1919
Subtribe Stereodermina Sharp, 1895
Tribe Pholidochlamydini Damoiseau, 1962
Tribe Taphroderini Lacordaire, 1865
Tribe Trachelizini Lacordaire, 1865
Subtribe Acratina Alonso-Zarazaga, Lyal, Bartolozzi and Sforzi, 1999
Subtribe Ithystenina Lacordaire, 1865
Subtribe Microtrachelizina Zimmerman, 1994
Subtribe Pseudoceocephalina Kleine, 1922
Subtribe Rhyticephalina Kleine, 1922
Subtribe Trachelizina Lacordaire, 1865
Subtribe Tychaeina Schoenfeldt, 1908
Tribe Ulocerini Schönherr, 1823
Subfamily Eurhynchinae Lacordaire, 1863
†Tribe Axelrodiellini Legalov, 2009
Tribe Eurhynchini Lacordaire, 1863
Subfamily Apioninae Schönherr, 1823
Supertribe Apionitae Schönherr, 1823
Tribe Apionini Schönherr, 1823
Subtribe Apionina Schönherr, 1823
Subtribe Aplemonina Kissinger, 1968
Subtribe Aspidapiina Alonso-Zarazaga, 1990
Subtribe Catapiina Alonso-Zarazaga, 1990
Subtribe Ceratapiina Alonso-Zarazaga, 1990
Subtribe Exapiina Alonso-Zarazaga, 1990
Subtribe Ixapiina Alonso-Zarazaga, 1990
Subtribe Kalcapiina Alonso-Zarazaga, 1990
Subtribe Malvapiina Alonso-Zarazaga, 1990
Subtribe Metapiina Alonso-Zarazaga, 1990
Subtribe Oxystomatina Alonso-Zarazaga, 1990
Subtribe Piezotrachelina Voss, 1959
Subtribe Prototrichapiina Wanat, 1995
Subtribe Synapiina Alonso-Zarazaga, 1990
Subtribe Trichapiina Alonso-Zarazaga, 1990
Tribe Chilapiini Wanat, 2001
Tribe Noterapiini Kissinger, 2004
Tribe Podapiini Wanat, 2001
Tribe Rhinorhynchidiini Zimmerman, 1994
Supertribe Antliarhinitae Schönherr, 1823
Supertribe Cybebitae Lacordaire, 1863
Supertribe Mecolenitae Wanat, 2001
Supertribe Myrmacicelitae Zimmerman, 1994
Tribe Lispotheriini Wanat, 2001
Tribe Myrmacicelini Zimmerman, 1994
Supertribe Rhadinocybitae Alonso-Zarazaga, 1992
Tribe Notapionini Zimmerman, 1994
Tribe Rhadinocybini Alonso-Zarazaga, 1992
Supertribe Tanaitae Schönherr, 1839
Subfamily Ithycerinae Schönherr, 1823
Subfamily Microcerinae Lacordaire, 1863
Subfamily Nanophyinae Gistel, 1848
Tribe Corimaliini Alonso-Zarazaga, 1989
Tribe Nanophyini Gistel, 1848
Family Dryophthoridae Schönherr, 1825
Subfamily Dryophthorinae Schönherr, 1825
Subfamily Cryptodermatinae Bovie, 1908
Subfamily Orthognathinae Lacordaire, 1865
Tribe Orthognathini Lacordaire, 1865
Tribe Rhinostomini LeConte, 1874
Subfamily Rhynchophorinae Schönherr, 1833
Tribe Diocalandrini Zimmerman, 1993
Tribe Litosomini Lacordaire, 1865
Tribe Ommatolampini Lacordaire, 1865
Tribe Polytini Zimmerman, 1993
Tribe Rhynchophorini Schönherr, 1833
Tribe Sphenophorini Lacordaire, 1865
Subfamily Stromboscerinae Lacordaire, 1865
Family Brachyceridae Billberg, 1820
Subfamily Brachycerinae Billberg, 1820
Tribe Brachycerini Billberg, 1820
Tribe Byrsopini Germar, 1829
Subfamily Cryptolarynginae Schalkwyk, 1966
Subfamily Erirhininae Schönherr, 1825
Tribe Aonychini Zimmerman, 1993
Tribe Arthrostenini Reitter, 1913
†Tribe Cretuliini Legalov, 2009
Tribe Erirhinini Schönherr, 1825
Tribe Himasthlophallini Zherikhin, 1991
Tribe Stenopelmini LeConte, 1876
Tribe Tadiini Zimmerman, 1993
Tribe Tanysphyrini Gistel, 1848
Subfamily Ocladiinae Lacordaire, 1865
Tribe Desmidophorini Morimoto, 1962
Tribe Ocladiini Lacordaire, 1865
Subfamily Raymondionyminae Reitter, 1913
Tribe Myrtonymini Kuschel, 1990
Tribe Raymondionymini Reitter, 1913
Family Curculionidae Latreille, 1802
Subfamily Curculioninae Latreille, 1802
Tribe Acalyptini Thomson, 1859
Subtribe Acalyptina Thomson, 1859
Subtribe Derelomina Lacordaire, 1865
Subtribe Notolomina Franz, 2006
Subtribe Phyllotrogina Franz, 2006
Subtribe Staminodeina Franz, 2006
Tribe Acentrusini Alonso-Zarazaga, 2005
Tribe Ancylocnemidini Voss, 1962
Tribe Anthonomini Thomson, 1859
Tribe Camarotini Schönherr, 1833
Subtribe Camarotina<