♂ (CUMZ), Thailand, Chiang Mai Province, Chom Thong District, Doi Inthanon National Park, 1, 700 m a.s.l., 18°31'55"N, 98°29'30"E, 20.12.2013, leg. N. Likhitrakarn & S. Chaiwong.

2 ♂, 6 ♀, 2 juveniles (CUMZ), 1 ♂ (NHMW), same locality, together with holotype. 1 ♂, 1 ♀ (CUMZ), same locality, 25.01.2013, leg. N. Likhitrakarn.

To emphasize the clearly wrinkled postcollum metaterga.

Differs from congeners mainly in the very strongly developed paraterga with evident oblong ridges. Gonopod process h prominent, hook-shape, much longer than solenophore.

Length 15.5–18.2 (♂) or 16.5–21.0 mm (♀), width of midbody pro- and metazonae 1.50–1.70 and 1.95–2.15 mm (♂) or 1.55–2.0 and 2.0–2.5 mm (♀), respectively.

Coloration of live animals blackish-brown (Fig. 1A) with a pattern of contrasting light brown paraterga and posterior halves of midbody metaterga and epiproct, dark brown to light brown head, legs and antennae; coloration in alcohol faded after two months of preservation, paraterga, legs and epiproct being light brown to whitish; head to metazonae 3 blackish, thereafter metazonae with a light brown to whitish cross (Fig. 1B, D, F); venter and a few basal podomeres light brown to yellow-brown, legs increasingly darker brown distally (Fig. 1B–J).

Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae moderately long (Fig. 1A–C), reaching body segment 3 (♂) or 2 (♀) when stretched dorsally. In width, head < segment 3 = 4 < collum < 2 < 5–17 (♂, ♀); thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4+4 anterior, 3+3 intermediate and 5+5 posterior; a small incision laterally in posterior half; caudal corner of paraterga rounded, slightly declined ventrad, produced behind rear tergal margin (Fig. 1B, C).

Tegument leathery and shining, prozonae very finely shagreened, metaterga leathery, finely microgranulate and delicately rugulose; surface below paraterga finely microgranulate. Postcollum metaterga with two transverse rows of setae on evident oblong ridges: 2+2 in anterior (pre-sulcus) and 3+3 in posterior (post-sulcus) row, caudal row being more strongly developed than anterior one (Fig. 1B–F, H); behind segment 10, metaterga with: 2+2 in anterior and 3(4)+3(4) in posterior row. Tergal setae long, strong, slender, about 1/3 of metatergal length. Axial line visible on metaterga. Paraterga very strongly developed (Fig. 1B–F, H), especially well so in ♂, set at about 1/4 midbody height, mostly upturned, all lying high, but always below dorsum; shoulders well-developed, mostly rounded; caudal corner almost completely to fully pointed, extending increasingly beyond tergal margin, posterior edge mostly oblique, especially strongly so on segments 16–19 (Fig. 1F, H); paraterga very thin blunt blades in lateral view, a little thicker only on pore-bearing segments. Calluses on paraterga delimited by a sulcus only dorsally. Paraterga 2 broad, anterior edge angular, lateral edge with two evident incisions in anterior half; posterior edge slightly concave (Fig. 1B, C). Lateral edge of paraterga with evident incisions, one in anterior 1/3, the other at midway, caudal incision being smaller in pore-bearing segments. Ozopores evident, lateral, lying in an ovoid groove at about 1/3 in front of caudal corner. Transverse sulcus usually distinct (Fig. 1B, D, F), slightly incomplete on segment 19, complete on metaterga 4–18, deep, reaching bases of paraterga, clearly beaded at bottom. Stricture between pro- and metazonae wide, clearly ribbed at bottom down to base of paraterga (Fig. 1B–E). Pleurosternal carinae complete crests with a sharp caudal tooth on segment 7 (♂, ♀), a small, caudal, mostly sharp tooth until segment 17 (♂) or 16 (♀), thereafter missing (Fig. 1C, E, H). Epiproct (Fig. 1F, G) conical, flattened dorsoventrally, with two strong apical papillae; tip subtruncate; pre-apical papillae evident, lying close to tip. Hypoproct roundly subtrapeziform (Fig. 1G), setiferous knobs at caudal edge well-separated and evident.

Sterna very densely setose, with a small cone caudally near each coxa, rear cones being a bit better developed than front ones (Fig. 2D); a deeply notched sternal lobe between ♂ coxae 4 (Fig. 1I, J). Legs moderately long and slender, midbody ones ca 1.0–1.2 (♂) or 0.9–1.1 times (♀) as long as body height, ♂ legs of segments 5–16 with an evident adenostyle (tubercle) on femur, postfemur, tibia and tarsus (Fig. 2D–F); tarsal brushes present until ♂ legs 7.

Gonopods (Figs 2A–C, 3) simple; coxa a little curved caudad, sparsely setose distoventrally. Prefemur densely setose, about 1/3 as long as femorite + “postfemoral” part. Femorite rather stout, expanded distad, slightly curved, showing a mesal groove; lobe l simple; process m apicoventral and spiniform; solenophore long and slender, typically coiled, tip subtruncate; process h strongly developed, curved and acute, longer than solenophore.

♂ (CUMZ), Thailand, Loei Province, Phuluang District, Phuluang Wildlife Sanctuary, 1, 486 m a.s.l., 17°16'44.9"N, 101°31'10.2"E, 20.07.2011, leg. Sira Noommeechai.

1 ♀ (CUMZ), same data, together with holotype.

To emphasize the close resemblance to Tylopus hilaroides Golovatch, 1984.

Very similar to Tylopus hilaroides, especially as regards its gonopod conformation, but differs in the presence of two rows of setae on metaterga 3–18 (an anterior transverse row of 2+2 setae and a posterior row of 4+4 insertion points versus solely an anterior transverse row of 2+2 setae), by the transverse sulcus visible starting already from metatergum 4 (versus metatergum 5), as well as in gonopod process z with two evident spines along distal margin (versus three spines) and process h being smaller (versus stouter).

Length 34 (♂) or 33 mm (♀), width of midbody pro- and metazonae 3.1 and 4.3 mm (♂) or 3.2 and 4.1 mm (♀), respectively.

Coloration of live animals dark castaneous brown (Fig. 4A); legs red-brown, venter and a few basal podomeres light brown to yellow-brown; coloration of alcohol material after a half year preservation faded to dark brown; antennae and epiproct light brown to pallid, venter and a few basal podomeres light brown to pallid (Fig. 4B–H).

Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae moderately long (Fig. 4A), reaching behind body segment 3 (♂, ♀) when stretched dorsally. In width, head < segment 3 < 4 < 5 < collum < segment 2 < 6–17 (♂) or head < segment 3 < 4 < collum < segment 2 < 5–17 (♀); thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4+4 anterior, 3+3 intermediate, and 4+4 posterior; a setigerous incision laterally in posterior 1/3; caudal corner of paraterga very narrowly rounded, not drawn behind rear tergal margin (Fig. 4B, C).

Tegument rather smooth and shining, prozonae finely shagreened, metaterga often rugose (Fig. 4A–F); surface below paraterga finely microgranular (Fig. 4C, E, H). Postcollum metaterga with an anterior transverse row of 2+2 setae visible at least as insertion points, and a posterior row of 4+4 insertion points. Tergal setae long, strong, slender, about 1/3 of metatergal length. Axial line clearly visible both on pro- and metazonae. Paraterga strongly developed (Fig. 4B–H), especially so in ♂, lying rather high (at 1/3 of midbody height), slightly upturned, but lying below dorsum; anterior edge rounded, caudal corner very narrowly rounded, starting from segment 13 extending increasingly beyond rear tergal margin, pointed, on segments 15–19 tips strongly curved mesad (Fig. 4F, H); lateral edge on poreless segments with two evident (anterior larger, posterior one smaller) setigerous incisions in anterior 1/3, but with only one strong (anterior) incision on pore-bearing segments (Fig. 4B, D, F); posterior edge oblique. Calluses on paraterga narrow, delimited by a sulcus only dorsally in segments 2–3, but both dorsally and ventrally in following segments. Paraterga 2 broad, posterior edge clearly oblique. Paraterga 2 and 3 broadly angular anteriorly, following segments with rounded anterior edges (Fig. 4B). Ozopores evident, lateral, lying in an ovoid groove at about 1/3 in front of posterior edge of metaterga. Transverse sulcus usually distinct (Fig. 4B, D, H), slightly incomplete on segments 4 and 19, complete on metaterga 5–18 (♂, ♀), narrow, wavy, rather deep, not reaching bases of paraterga, at most faintly ribbed at bottom. Stricture between pro- and metazonae broad and deep, beaded at bottom down to well below base of paraterga (Fig. 4B, D, H). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2–4, thereafter split into a sharp front and a sharp caudal tooth, the former gradually turning into a bulge, the latter tooth gradually reduced until segment 17 (♂, ♀). Epiproct (Fig. 4F, G) conical, flattened dorsoventrally, with two evident apical papillae; tip subtruncate; pre-apical papillae small, lying rather close to tip. Hypoproct roundly subtrapeziform, setiferous knobs at caudal edge small and well-separated (Fig. 4G).

Sterna densely setose, without modifications; a single, linguiform, sternal lobe between ♂ coxae 4 (Fig. 4I, J). Legs rather long and slender, midbody ones ca 1.2–1.3 (♂) or 1.1–1.4 times (♀) as long as body height; appressed setation ventrally on coxa, prefemur and femur, but tarsal brushes absent; ♂ prefemora distinctly bulged laterally (Fig. 5C), ♂ postfemora and tibiae on segments 7–17 with an evident adenostyle at midway on ventral side (Fig. 5C).

Gonopods (Fig. 5A, B) simple; coxa a little curved caudad, sparsely setose distoventrally. Prefemur densely setose, about 1/3 as long as femorite + “postfemoral” part. Femorite rather stout, expanded distad, slightly curved, showing a mesal groove; lobe l simple; process z with two evident spines along dorsal margin; process h short and slender, curved, with an acute tip; solenophore long and slender, typically coiled, tip subtruncate.

♂ (CUMZ), Thailand, Tak Province, Umphang District, Pa Wai Waterfall, 804 m a.s.l., 16°34'29.6"N, 98°50'3.2"E, 20.01.2011, leg. C. Sutcharit & N. Likhitrakarn.

1 ♂ (CUMZ), same data, together with holotype. 3 ♀ (CUMZ), same district, Thee Lor Sue Waterfall, 591 m a.s.l., 15°55'38.1"N, 98°45'12.8"E, 19.01.2011, leg. N. Likhitrakarn.

To emphasize the light brown triangle on terga; noun in apposition.

This new species shows a peculiar colour pattern, much like that observed in Tylopus schawalleri Golovatch, 2013, but differs in gonopod process h being rather short and coiled (versus high and strongly twisted), as well as by the presence of a process m (versus absent).

Length 21.2–27.8 (♂) or 22.1–24.0 mm (♀), width of midbody pro- and metazona 1.97–1.83 and 2.65–2.78 mm (♂) or 2.43–2.58 and 3.05–3.14 mm (♀), respectively.

Coloration of live animals light brown (Fig. 6A); paraterga, legs and epiproct light brown, head and collum blackish, following terga with a light brown triangle and blackish collar covering both pro- and metazonae; coloration of alcohol material after three years of preservation faded to whitish with a pattern of a contrasting dark brown inverted triangle at anterior edge of metazonae and a light brown triangle at posterior edge of prozonae (Fig. 6B–H).

Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae very short (Fig. 6A, B), reaching only behind body segment 2 (♂) or collum (♀) when stretched dorsally. In width, head < segment 3 < 4 < collum < segment 2 < 5–17 (♂, ♀); thereafter body gently and gradually tapering. Collum with three transverse rows of strong setae: 4+4 anterior, 2+2 intermediate, and 4+4 posterior; a rounded incision laterally in posterior half; caudal corner of paraterga rounded, slightly declined ventrad, produced behind rear tergal margin (Fig. 6B, C).

Tegument rather smooth and shining, prozonae very finely shagreened, metaterga smooth and finely rugulose, leathery; surface below paraterga finely microgranulate (Fig. 6B–H). Postcollum metaterga with two transverse rows of setae on small knobs to oblong ridges: 2+2 in anterior (pre-sulcus), 3+3 in posterior (post-sulcus) row, caudal row more strongly developed than anterior one, starting from metaterga 11 with 2+2 in anterior and 4(3)+4(3) in posterior row. Tergal setae long, strong, slender, about 1/3 of metatergal length. Axial line visible. Paraterga very strongly developed (Fig. 6B–F, H), especially well in ♂, set high, at about 1/3 of midbody height, slightly upturned, always lying high, but below dorsum; shoulders well-developed, mostly regularly rounded and narrowly bordered, fused to callus; caudal corner narrowly rounded to fully pointed, extending increasingly beyond rear tergal margin, posterior edge clearly oblique (Fig. 6B, D, F); paraterga very thin blunt blades in lateral view, a little thicker only on pore-bearing segments. Calluses on paraterga delimited by a sulcus both dorsally and ventrally. Paraterga 2 broad, anterior edge angular, lateral edge with three evident incisions in anterior half (Fig. 6B, C). Lateral edge of following paraterga with two clear incisions, one in anterior 1/3, the other at midway, front one being particularly evident. Ozopores evident, lateral, lying in an ovoid groove at about 1/3 in front of caudal corner. Transverse sulcus usually distinct (Fig. 6B, D, H), slightly incomplete on segments 4 and 19, complete on metaterga 5–18, deep, reaching bases of paraterga, clearly ribbed at bottom. Stricture between pro- and metazonae very wide, clearly beaded at bottom down to base of paraterga (Fig. 6B, D, H). Pleurosternal carinae complete crests with a sharp caudal tooth on segment 12 (♂) or 7 (♀), thereafter increasingly well reduced in size and sharpness until segment 17 (♂) or 14 (♀), onward missing (Fig. 6C, E, H). Epiproct (Fig. 6F–H) conical, flattened dorsoventrally, subtruncate, with two evident apical papillae directed caudally, both pointed at tip; pre-apical papillae evident, lying close to tip. Hypoproct roundly subtrapeziform (Fig. 6G), small setiferous knobs at caudal edge well-separated and evident.

Sterna very densely setose, with a small cone caudally near each coxa; a single, linguiform, deeply medially notched sternal lobe between ♂ coxae 4 (Fig. 9I, J). Legs moderately long and slender, midbody ones ca 1.1–1.2 (♂) or 1.0–1.1 times (♀) as long as body height, ♂ femora with 2–4 small adenostyles on ventral side (Fig. 7C); tarsal brushes present until ♂ leg 7.

Gonopods (Fig. 7A, B) very simple; coxa a little curved caudad, sparsely setose distoventrally. Prefemur densely setose, about 1/3 as long as femorite + “postfemoral” part. Femorite rather slender, expanded distad, slightly curved, showing a mesal groove; lobe l simple; solenophore long and slender, typically coiled, tip subtruncate; process m evident, but not spiniform; process h prominent, coiled, acute at tip.

The ♂ paratype on metatergum 19 shows 3+3 and 6+6 setae without knobs in the anterior and posterior rows, respectively.

♂ (here designated) of Agnesia nodulipes (NHMW-3986), Laos, Luang Prabang, 1938–1939, leg. C. Dawydoff.

Lectotype designation proposed herewith is necessary to ensure the species is based on a complete ♂ coming from a certain locality, because (1) Attems (1953) provided no information on the number and sex of syntypes, and (2) he stated their provenance to have been both from Luang Prabang, Laos and Mount Fan-Si-Pan, Lao Cai Province, Vietnam. No paralectotype material could be traced in the Vienna Museum.

Lectotype ca 24 mm long, width of midbody pro- and metazonae 2.1 and 2.9 mm (vs 3.0 in width, as given in the available description (Attems 1953)). Coloration of alcohol material after long preservation rather uniformly light reddish brown (Fig. 8A–G) with light yellow antennae, paraterga, epiproct and legs (versus dark maroon with light yellowish brown mid-dorsal parts of prozonae, paraterga a little lighter, antennae light chestnut brown and legs yellow brown, as given in the original description (Attems 1953)).

Clypeolabral region densely setose; vertex rather smooth, only faintly rugulose; epicranial suture distinct. Antennae rather long and slender (Fig. 8A, B), reaching behind body segment 3 when stretched dorsally. In width, head < segments 3 and 4 < collum < 2 < 5–16, gently and gradually tapering thereafter. Collum smooth, with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 4+4 posterior; caudal corner of paraterga subrectangular, narrowly rounded (Fig. 8A, B), drawn behind rear tergal margin.

Tegument smooth and shining; metaterga rugulose, prozonae finely shagreened, surface below paraterga finely microgranulate. Metaterga 2–17 with two transverse rows of setae: 2+2 in anterior (pre-sulcus) row and 3(2)+3(2) in posterior (post-sulcus) one, setae being borne on very small tubercles growing a little larger laterally, metaterga 18 and 19 with two transverse rows of setae: 2+2 in anterior and 4+4 in posterior row. Tergal setae long, strong, slender, about 1/3 of metatergal length. Axial line visible. Paraterga very strongly developed (Fig. 8A–G), subhorizontal, lying below dorsum, thin blunt blades in lateral view, a little thicker only on pore-bearing segments, on postcollum segments extending increasingly beyond rear tergal margin, nearly pointed to pointed, caudal tip on paraterga 18–19 clearly curved mesad. Calluses delimited by a sulcus only dorsally, rather narrow. Paraterga 2 broad, slightly upturned, anterior edge rounded, lateral edge with three small incisions in anterior half; posterior edge oblique (Fig. 8A, B). Anterior edge of postcollum metaterga broadly rounded, bordered and fused to callus, lateral edge with two small incisions in anterior half on poreless segments, with only one incision near front 1/3 on pore-bearing ones. Ozopores evident, lateral, lying inside an ovoid groove at about 1/3 of metazonital length. Transverse sulcus complete on metaterga 5–18, incomplete on metatergum 19, rather wide, reaching bases of paraterga, faintly beaded at bottom (Fig. 8A, C, F). Stricture between pro- and metazonae very wide, shallow, faintly beaded at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segment 2 (Fig. 8B), with anteriorly bulged crests and a sharp denticle caudally on segments 3–8, thereafter only a small sharp caudal tooth on segments 9–15, onward missing (Fig. 8B & D). Epiproct (Fig. 8F, G) conical, flattened dorsoventrally, apical papillae evident; tip subtruncate; pre-apical papillae rather large, lying close to tip. Hypoproct (Fig. 8G) roundly subtrapeziform, setigerous knobs at caudal margin evident and well-separated.

Sterna sparsely setose, starting from segment 6 with a small cone caudally near each coxa, rear cones being a little larger than front ones; a rather large, linguiform, densely setose, sternal lobe between ♂ coxae 4 (Fig. 8H, I). Legs moderately long and slender, midbody ones ca 1.2–1.3 times as long as body height, legs of segments 8–18 with an evident adenostyle on each prefemur, postfemur and tibia, with two adenostyles on each femur (Fig. 9C); tarsal brushes present only until ♂ legs 4.

Gonopods (Fig. 9A, B) rather simple; prefemur densely setose, about 1/3 as long as femorite + “postfemoral” part. Femorite stout, expanded distad, slightly curved, showing a mesal groove; lobe l simple; solenophore long and slender, typically coiled, tip subtruncate; process h high, strongly twisted, tip bifid; process m rather long and spiniform, process z knife-shaped.

This is the type species of Tylopus Jeekel, 1968, originally recorded from two localities: Luang Prabang Province, Laos and Mount Fan-Si-Pan, Lao Cai Province, Vietnam (Attems 1953). Golovatch (1984) redescribed and illustrated only a gonopod, but the locality remained unclear. So the lectotype is herewith selected for the sole type specimen still kept in the Vienna Museum.

This species has recently been reported from Nam Xay Commune (22°05’N, 104°05’E), 1, 000 m a.s.l., Van Ban District, Lao Cai Province; Son Tay Commune, 600 m a.s.l., Huong Son District, Ha Tinh Province; and Chem Waterfall, 430 m a.s.l., Pu Mat National Park (18°46'–19°12'N, 104°01'–104°56'E), Nghe An Province, Vietnam (Nguyen 2012).

♂ of Anoplodesmus hilaris (NHMW-4248), Vietnam, Danang Prov., Mount Bana, 1, 500 m, 28.09.1931, leg. C. Dawydoff.

Length ca 38 mm, width of midbody pro- and metazonae 3.7 and 5.1 mm, respectively (vs 3.4 and 5.0 mm in width, as given in the available descriptions (Attems 1937, 1938)). Coloration of alcohol material after long preservation brown (Fig. 10A–G) with light yellow antennae, paraterga, epiproct and legs (versus dark brown with prozonae and posterior halves of metazonae blackish brown; edge of paraterga, antennae and legs yellowish brown, as given in the descriptions (Attems 1937, 1938)).

Clypeolabral region densely setose, vertex smooth, epicranial suture distinct. Antennae rather long (Fig. 10A, B), reaching behind body segment 3 when stretched dorsally. In width, head < segments 3 and 4 < collum < 2 < 5–16, gently and gradually tapering thereafter. Collum smooth, with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 3+3 posterior; caudal corner of paraterga subrectangular, narrowly rounded (Fig. 10A, B). Tegument smooth and shining; metaterga faintly rugulose, prozonae finely shagreened, surface below paraterga finely microgranulate. Postcollum metaterga with an anterior transverse row (pre-sulcus) of 2+2, mostly abraded setae; caudal (post-sulcus) row barely traceable as 3+3 insertion points. Tergal setae short, simple, slender, about 1/5 metatergal length. Axial line barely visible, starting from collum. Paraterga very strongly developed (Fig. 10A–F), all subhorizontal and lying below dorsum, thin blunt blades in lateral view, a little thicker only on pore-bearing segments, on postcollum segments extending increasingly beyond rear tergal margin, nearly pointed to pointed, caudal tips on paraterga 17–19 evidently curved mesad. Calluses delimited by a sulcus only dorsally, rather narrow. Paraterga 2 broad, anterior edge rounded, lateral edge with three small incisions in anterior half; posterior edge concave (Fig. 10A). Anterior edge of postcollum segments broadly rounded, bordered and fused to callus, lateral edge with two small incisions in anterior half on poreless segments, with only one incision near front 1/3 on pore-bearing ones; posterior edge oblique. Ozopores evident, lateral, lying inside an ovoid groove at about 1/4 metazonital length in front of caudal corner. Transverse sulcus complete on metaterga 5–18, incomplete on metaterga 4 and 19, rather deep, wide, line-shaped, reaching bases of paraterga, ribbed at bottom (Fig. 10A, C, F). Stricture between pro- and metazonae shallow, broad, beaded at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segment 2 (Fig. 10B), with a sharp tooth caudally on segments 3–7, only a small sharp caudal tooth on segments 8–16, onward missing (Fig. 10A, D, E). Epiproct (Fig. 10E–G) conical, flattened dorsoventrally, apical papillae very small; tip subtruncate; pre-apical papillae very small, lying close to tip. Hypoproct (Fig. 10G) roundly subtriangular, setiferous knobs at caudal margin small and well-separated.

Sterna sparsely setose, until segment 6 with an evident cone caudally near coxae; on segment 4 with an evident, central cone between coxae; on segment 5 with a small central lobe with a paramedian pair of evident, sparsely setose, apical cones between coxae (Fig. 10H, I). Legs long and slender, midbody ones ca 1.2–1.3 (♂) or 0.8–0.9 (♀) times as long as midbody height, all legs until segment 17 with an evident adenostyle on each postfemur and tibia (Fig. 11C); tarsal brushes absent.

Gonopods (Fig. 11A, B) simple; coxa a little curved caudad, sparsely setose distoventrally. Prefemur densely setose, about 1/3 as long as femorite + “postfemoral” part. Femorite rather stout, expanded distad, slightly curved, showing a mesal groove; lobe l simple; process z with two small spines along ventral margin; process h short and stout, curved, tip acute; solenophore long and slender, typically coiled, tip subtruncate.

Endemic to Vietnam, Tylopus hilaris is currently known from Mount Bana, 1, 500 m a.s.l., Danang Province (Attems, 1937); Bach Ma National Park (16°05'–16°05'N, 107°43'–107°53'E), Thua Thein Hue Province; Mount Ngoc Linh (15°00'–15°18'N, 107°41'–108°01'E), Kon Tum Province, central Vietnam (Nguyen 2012).

(here designated) of Sundanina sigma (NHMW-3987), Vietnam, Lao Cai Prov., Sapa (= Chapa), 1938–1939, leg. C. Dawydoff.

2 ♂ of Sundanina sigma (NHMW-3987), same data, together with lectotype.

Lectotype designation proposed herewith is necessary to ensure the species is based on a complete ♂, because Attems (1953) provided no information on the number and sex of syntypes.

Length 18–19 mm (♂), width of midbody pro- and metazonae 1.0–1.1 and 1.5–1.7 mm (♂), respectively (versus length 25 mm and width of midbody metazonae 1.6 mm, as given in the original description (Attems 1953)). Lectotype 18 mm long, 1.0 and 1.5 mm wide on midbody pro- and metazonae, respectively. Coloration of alcohol material after long preservation rather uniformly brown (Fig. 12A–G) (versus blackish brown with edges of paraterga and legs yellowish brown, as given in the description (Attems 1953)).

Clypeolabral region sparsely setose, vertex smooth, epicranial suture distinct. Antennae long and slender (Fig. 12B), extending behind body segment 5 (♂) dorsally. In width, head < segments 2–4 < collum < 5–17 (♂), gently and gradually tapering thereafter. Collum smooth, with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 3+3 posterior; caudal corner of paraterga subrectangular, narrowly rounded (Fig. 12A). Tegument smooth and shining; metaterga and prozonae finely shagreened; surface below paraterga finely microgranulate. Postcollum with an anterior (pre-sulcus) transverse row of 2+2, mostly abraded setae; caudal (post-sulcus) row barely traceable as 3+3 insertion points. Tergal setae short, simple, slender, about 1/3 of metatergal length. Axial line barely visible, starting from collum. Paraterga very strongly developed (Fig. 12A, C, F, G), mostly subhorizontal and lying below dorsum, thin blunt blades in lateral view, a little thicker only on pore-bearing segments, posterior edge concave, caudal tip narrowly rounded. Calluses delimited by a sulcus only dorsally, rather narrow. Paraterga 2 broad, slightly upturned, anterior edge nearly straight, lateral edge with three more or less evident incisions; posterior edge clearly concave (Fig. 12A, B). Anterior edge of postcollum segments oblique, bordered and fused to callus, lateral edge with a strong incision near front 1/3; posterior edge oblique. Ozopores evident, lateral, lying inside an ovoid groove at about 1/4 metazonital length before caudal corner. Transverse sulcus complete on metaterga 5–18, shallow, not reaching bases of paraterga, faintly beaded at bottom (Fig. 12A, C, F). Stricture between pro- and metazonae broad, shallow, ribbed at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segment 2, with a small sharp caudal tooth on segments 3–7, onward missing (Fig. 12B, D, E). Epiproct (Fig. 12E–G) conical, flattened dorsoventrally, apical papillae evident and large; tip subtruncate; pre-apical papillae small, but visible, lying close to tip. Hypoproct (Fig. 12G) roundly subtrapeziform, setiferous knobs at caudal margin small and well-separated.

Sterna sparsely setose, with a large central cone between coxae 3 and a small central lobe with a paramedian pair of evident, sparsely setose, apical cones between coxae 4 (♂) (Fig. 12H, I); segments 8–16 with a strong cone caudally near each coxa. Legs long and slender, midbody ones ca 1.4–1.5 (♂) as long as body height, legs on segments 8–18 with a small adenostyle on each prefemur, femur and postfemur (Fig. 13C); tarsal brushes absent.

Gonopods (Fig. 13A, B) very simple; coxa a little curved caudad, sparsely setose distoventrally. Prefemur densely setose, about 1/3 as long as femorite + “postfemoral” part. Femorite rather slender, expanded distad, slightly curved, showing a mesal groove; lobe l simple; solenophore long and slender, typically coiled, tip subtruncate; process h short, rather curved, tip acute.

Endemic to Vietnam, Tylopus sigma is only known from Sapa (= Chapa), Lao Cai Province, Vietnam (Attems 1953).

♂ of Anoplodesmus mutilatus (NHMW-4245), locality unknown; a slide with mounted gonopod.

Gonopod (Fig. 14) rather simple. Coxa long and slender, with several setae distodorsally. Prefemur densely setose, nearly 1/3 as long as femorite + “postfemoral” part. Femorite stout, slightly curved, slightly enlarged distad, showing a mesal groove, “postfemoral” part demarcated by an oblique lateral sulcus; lobe l simple; process h long, rather simple, slightly curved, tip small and bifid; process z high, slightly curved, tip acute; solenophore long and slender, typically coiled, tip microdenticulate.

This species was described both from Luang Prabang, Xieng Kuang, Laos and Pic de Langbiang (Mount Langbian), Lamdong Province, Vietnam (Attems 1953). Golovatch (1984) redescribed and illustrated only a gonopod, but the locality remained unclear. As all our attempts at locating a torso of Tylopus sigma in the collection of the Naturhistorisches Museum Wien, Austria had failed, we could only revise the very same right gonopod mounted on a slide. Fortunately, the gonopod is easily distinguished from congeners.