Citation: Kamiński MJ (2013) A cladistically based reinterpretation of the taxonomy of two Afrotropical tenebrionid genera Ectateus Koch, 1956 and Selinus Mulsant & Rey, 1853 (Coleoptera, Tenebrionidae, Platynotina). In: Bouchard P, Smith AD (Eds) Proceedings of the Third International Tenebrionoidea Symposium, Arizona, USA, 2013. ZooKeys 415: 81–132. doi: 10.3897/zookeys.415.6406
On the basis of a newly performed cladistic analysis a new classification of the representatives of two Afrotropical tenebrionid genera, Ectateus Koch, 1956 and Selinus Mulsant & Rey, 1853 sensu Iwan 2002a, is provided. Eleoselinus is described as a new genus. The genus Monodius, previously synonymized with Selinus by Iwan (2002), is redescribed and considered as a separate genus. Following new combinations are proposed: Ectateus calcaripes (Gebien, 1904), Monodius laevistriatus (Fairmaire, 1897), Monodius lamottei (Gridelli, 1954), Monodius plicicollis (Fairmaire, 1897), Eleoselinus villiersi (Ardoin, 1965) and Eleoselinus ursynowiensis (Kamiński, 2011). Neotype for Ectateus calcaripes and lectotypes for E. crenatus (Fairmaire, 1897), E. ghesquierei Koch, 1956 and Monodius malaisei malaisei Koch, 1956 are designated to fix the taxonomic status of these taxa. The following synonymies are proposed: Selinus monardi Kaszab, 1951 and Ectateus latipennis Koch, 1956 with E. crenatus (Fairmaire, 1897). Identification keys are provided to all known species of Ectateus sensu novum, Eleoselinus, Monodius and Selinus sensu novum.
Africa, ecoregions, cladistics, identification key, new genus, taxonomy, Pedinini
Pursuant to the classification of the family Tenebrionidae presented by
According to the results of a cladystic analysis performed by
The current taxonomic concept of the genus Ectateus was proposed by
According to the results of a cladistic analysis performed by
The aim of this paper was to test the monophyly of Ectateus and Selinus and propose a stable classification for the representatives of these genera.
Morphological studies. The descriptive sequence used in this study is in accordance with
Measurements, taken using a filar micrometer, were as follows: width of anterior elytral margin (from humeral angle to scutellum); body length (from anterior margin of labrum to elytral apex); body width (maximum elytral width).
For examination of internal structures, specimens were dissected and whole abdomens were cleared in 10% cold potassium hydroxide overnight (
Images were taken using a Canon 1000D body with accordion bellows and Industar 61L/3 MC 50 mm f/2.8 lens, and with a Hitachi S-3400N SEM in MIIZ. Chosen SEM photographs were colored using Photoshop CS5.
Entomological material. This study was based on the material from the following collections:
BMNH Natural History Museum, London, Great Britain
HNHM Hungarian Natural History Museum, Budapest, Hungary
JFCS Julio Ferrer Collection, Haninge, Sweden
MHNG Muséum d’histoire naturelle de la Ville de Genève, Geneva, Switzerland
MIIZ Museum and Institute of Zoology, Polish Academy of Sciences, Warsaw, Poland
MNB Museum für Naturkunde, Germany, Berlin
MNHN Muséum National d’Histoire naturelle, Paris, France
MHNL Centre de Conservation des Collections, Muséum d’Histoire Naturelle, Lyon, France
MRAC Royal Museum for Central Africa, Tervuren, Belgium
RBINS Royal Belgian Institute of Natural Sciences, Brussels, Belgium
TMNH Ditsong National Museum of Natural History, Pretoria, Republic of South Africa
SMNS Staatliches Museum für Naturkunde, Stuttgart, Germany
ZMAS Zoological Museum, Academy of Sciences, Sankt Petersburg, Russia
Phylogenetic analysis. Based on the results of a comparative analysis of the morphology of available material, including the type specimens, I propose a following synonymy: Selinus monardi Kaszab, 1951 and Ectateus latipennis Koch, 1956 with Ectateus crenatus (Fairmaire, 1897). Also, I disagree with the synonymy of Selinus calcaripes Gebien, 1904 with Ectateus curtulus (Fairmaire, 1893) proposed by
The operational taxonomic units (OTUs) representing the genus Ectateus consists of all (8) known species (considering above mentioned nomenclatural acts): Ectateus crenatus (Fairmaire, 1897), Ectateus curtulus (Fairmaire, 1893), Ectateus ghesquierei Koch, 1956, Ectateus laevistriatus (Fairmaire, 1897), Ectateus lamottei (Gridelli, 1954), Ectateus modestus (Fairmaire, 1887), Ectateus ursynowiensis Kamiński, 2011 and Ectateus villiersi Ardoin, 1965. Also, all (7) known species of Selinus were included in the phylogenetic analysis: Selinus convexipennis Gebien, 1904, Selinus gravis Koch, 1956, Selinus malaisei Koch, 1956, Selinus medius Fairmaire, 1897, Selinus planus (Fabricius, 1792), Selinus plicicollis Fairmaire, 1897 and Selinus striatus (Fabricius, 1794). The above mentioned taxa form the ingroup.
Zidalus latipes (Sahlberg, 1823) was used as the most distant outgroup on which the character polarization process was performed. According to
Lechius abacoides (Fairmaire, 1902), Pseudoselinus punctatostriatus (Gerstaecker, 1854), Upembarus upembaensis Koch, 1956 were used to test the monophyly of the clade Ectateus+Selinus. According to the results of
The data matrix originated in Mesquite (
Species distribution. The distribution of species was illustrated using DIVA-GIS version 7.5 (
Character matrix. A matrix of 40 characters was constructed for 20 operational taxonomic units (Table 1). Characters used for phylogenetic analyses have been treated as unordered. The missing data for Ectateus curtulus are caused by the fact that this species is only known from one specimen (holotype, female). The character states are presented in this section.
Character matrix for the cladistic analysis of the species of Ectateus and Selinus (sensu
Taxon / character | 1 | 6 | 11 | 16 | 21 | 26 | 31 | 36 |
---|---|---|---|---|---|---|---|---|
Zidalus latipes | 00000 | 00000 | 01000 | 10000 | 00000 | 00100 | 00000 | 00000 |
Lechius abacoides | 00000 | 10000 | 01000 | 00000 | 00000 | 00000 | 00000 | 00000 |
Pseudoselinus punctatostriatus | 00000 | 01000 | 01000 | 00101 | 00000 | 01100 | 00000 | 00000 |
Upembarus upembaensis | 00000 | 01000 | 01000 | 00100 | 00000 | 00100 | 00000 | 00000 |
Ectateus crenatus | 00111 | 00110 | 10211 | 10011 | 11100 | 00001 | 00100 | 10001 |
Ectateus curtulus | 0?010 | 00100 | 10211 | 00011 | 10100 | 00??? | ????? | ??000 |
Ectateus ghesquierei | 00111 | 00110 | 10211 | 00011 | 11100 | 00001 | 00100 | 10001 |
Ectateus laevistriatus | 01001 | 10000 | 00101 | 10001 | 00000 | 10110 | 01011 | 01100 |
Ectateus lamottei | 01001 | 10000 | 00101 | 00001 | 00000 | 10110 | 01011 | 01100 |
Ectateus modestus | 01111 | 00110 | 10211 | 10011 | 10100 | 00001 | 00100 | 10000 |
Ectateus ursynowiensis | 10000 | 00000 | 00100 | 01000 | 00101 | 00001 | 00100 | 00000 |
Ectateus villiersi | 10000 | 00001 | 00100 | 01000 | 00001 | 00001 | 00100 | 00000 |
Selinus calcaripes | 01111 | 00110 | 10211 | 10011 | 10000 | 00001 | 00100 | 10000 |
Selinus convexipennis | 01001 | 10000 | 00101 | 00001 | 00000 | 10110 | 01010 | 01100 |
Selinus gravis | 01000 | 10000 | 00100 | 00001 | 00000 | 10110 | 01010 | 01100 |
Selinus malaisei | 01000 | 10000 | 00100 | 00000 | 00010 | 10110 | 11010 | 02100 |
Selinus medius | 01000 | 10000 | 00100 | 00000 | 00010 | 10110 | 11010 | 02100 |
Selinus planus | 01000 | 10001 | 00100 | 00001 | 00000 | 01100 | 00010 | 00010 |
Selinus plicicollis | 01000 | 10000 | 00100 | 00000 | 00010 | 10110 | 11010 | 02100 |
Selinus striatus | 01000 | 10001 | 00100 | 00001 | 00000 | 01100 | 00010 | 00010 |
Head (characters 1–7)
1. Anntenna: (0) slender, longer than pronotum; (1) robust, shorter than pronotum.
2. Antennomeres from 7 to 11: (0) widened, their width greater than the length; (1) elongated, their length greater than the width (Fig. 3).
Head, dorsal view (1, 2), ventral view (4); antenna (3); pronotal disc (5, 6); mesotibia (7). Ectateus calcaripes (3), Ectateus crenatus (2), Ectateus ghesquierei (5), Monodius medius (7), Monodius plicicollis (4), Selinus planus (1, 6).
3. Circular depressions on the lateral sides of clypeus and genae: (0) absent (Fig. 1); (1) present (Fig. 2).
4. Fronto-clypeal suture: (0) fine (Fig. 1); (1) coarse, clearly visible (Fig. 2).
5. Indentation between frons and clypeus on the lateral edge: (0) shallow (Fig. 1); (1) deep (Fig. 2).
6. Anterior tentorial pit: (0) shallow; (1) deep, clearly visible (Fig. 4).
7. Anterior part of mentum: (0) not elongated; (1) elongated.
Prothorax (characters 8–18)
8. Anterior pronotal angles: (0) straight; (1) curved outwards (Fig. 5).
9. Lateral pronotal sides: (0) rounded; (1) sinusoidal (Fig. 5).
10. Pronotum: (0) widest at the middle (Fig. 5); (1) widest at the base (Fig. 6).
11. Pronotal margins: (0) not erected upwards; (1) strongly erected upwards (
12. Ratio of prothorax width (tw) and pronotal disc height (dh): (0) < 5; (1) > 6 (
13. Apophyseal depressions: (0) absent; (1) trapezoidal (Fig. 6); (2) rounded (Fig. 5).
14. Pronotal base: (0) the same width as elytral base; (1) narrower than elytral base.
15. Posterior pronotal angles: (0) not protruding towards elytra; (1) strongly protruding towards elytra.
16. Punctures on pronotal disc: (0) fine, the intervals between the punctures are greater than the 2 diameters of the puncture; (1) coarse, the intervals between the punctures are smaller than the diameter of the puncture.
17. Intercoxal process of prosternum: (0) flat or dented (Fig. 12); (1) bellied (
18. Intercoxal process of prosternum: (0) not widened at the apex; (1) strongly widened at the apex.
Elytral disc (8, 9); 5th abdominal ventrite (10, 11), intercoxal process of prosternum (12), male protarsi (13). Ectateus crenatus (8, 12), Monodius convexipennis (10), Monodius malaisei (9), Monodius plicicollis (13), Selinus striatus (11).
Mesothorax (characters 19–24)
19. Scutellum: (0) situated at the level of elytra; (1) impressed.
20. Elytral surface: (0) dull; (1) shiny.
21. Elytral intervals with transverse sculpture: (0) no; (1) yes (Fig. 8, 27).
22. Elytral intervals: flat (0); strongly convex (1).
23. Elytral striae: (0) impressed on whole length, with fine punctures (Fig. 9); (1) impressed mainly near conspicuous punctures (Fig. 8).
24. Margins of elytra in basal part: (0) rounded; (1) subparallel (elytral humeri slightly protruding outwards).
Metathorax (character 25)
25. Metaventrite: (0) without a coarse longitudinal depression; (1) with a coarse longitudinal depression.
Abdomen (characters 26–27)
26. 5th abdominal ventrite: (0) relatively narrow; (1) strongly widened (Fig. 10).
27. 5th abdominal ventrite: (0) unbordered (Fig. 10); (1) bordered (Fig. 11).
Legs (character 28–31)
28. Male protarsi widened: (0) no; (1) yes (Fig. 13).
29. Female protarsi widened: (0) no; (1) yes.
30. Male profemora (0) relatively wide (length/width = 3.2-3.6); (1) relatively slender (length/width = 4.0-5.6).
31. Denticle at the apex of the inner face of male mesotibia: (0) small, sometimes absent; (1) large (Fig. 7).
Male and female genitalia (character 32–39)
32. Penis wide: (0) no (Figs 20–21); (1) yes, at least 4 times wider than clavae (Figs 14–19).
Aedeagal tegmina (dorsal and vental views). Monodius gravis (14), Monodius plicicollis (15), Monodius convexipennis (16), Monodius malaisei (17), Monodius lamottei (18), Monodius laevistriatus (19), Ectateus calcaripes (20), Selinus striatus (21).
33. Clavae: (0) straight (Figs 14–19, 21); (1) curved, hook-shaped (Fig. 20).
34. Clavae: (0) short, their length less than half of the length of parameres; long, their length more than half of the length of parameres (1).
35. Parameres strongly extended apically: (0) no; (1) yes (Fig. 18).
36. Parameres narrowest in the half of their length (0) no; (1) yes (Fig. 20).
37. Apex of parameres: (0) not fused (Fig. 20); (1) fused, not emarginated at apex (Figs 14, 16); (2) fused, emarginated at apex (Figs 15, 17).
38. Bursa copulatrix: (0) without additional sacs; (1) with 2 additional sacs (Fig. 23).
Female genitalia. Ovipositor of Monodius plicicollis (22); bursa copultrix of Monodius medius (23), spermatheca of Monodius gravis (24).
39. Paraproct longer than coxites: (0) no (Fig. 22); (1) yes.
Other (character 40)
40. Body size: (0) more than 10.0 mm; (1) less than 9.0 mm.
Phylogenetic analysis. The cladistic analysis yield a single most parsimonious cladogram (Fig. 25) with a length of 57 steps, a consistency index (CI) of 74 and a retention index (RI) of 90. According to the obtained cladogram the genera Ectateus and Selinus, in their current interpretations, are paraphyletic (Fig. 25). The Ectateus clade is supported by two synapomorphies: male protibia slender (length/width = 4.0-5.6) (char. 30:1) and clavae curved, hook-shaped (char. 33:1). Also one homoplasy was recovered for this clade – male protarsi relatively narrow (char. 28:0). The Selinus clade is supported by single synapomorphy – clavae long, their length more than half of the length of parameres (char. 34:1) – and two homoplasies: antennomeres from 7 to 11 elongated (their length greater than the width), anterior tentorial pit deep, clearly visible (char. 2:1, 6:1). The monophyly of the Ectateus+Selinus clade was supported during the analysis by the following two synapomorphies: ratio of prothorax width (tw) and pronotal disc height (dh) greater than 5 (char. 12: 0) and apophyseal depressions on pronotal disc present (char. 13:1).
Phylogeny of the species of Ectateus and Selinus sensu
Two main species groups were recovered within the Ectateus clade – modestus group and villiersi group. The branch support reported for these groups was relatively high (Fig. 25). The phylogenetic relationships within the modestus group were supported unequally. Relatively low Jackknife values were reported within the clade composed of Ectateus modestus, Selinus calcaripes, Ectateus ghesquierei and Ectateus crenatus.
According to the results of a cladistic analysis the modestus group is characterized by following synapomorphies: fronto-clypeal suture coarse, clearly visible (char. 4:1), anterior pronotal angles curved outwards (char. 8:1), pronotal margins strongly erected upwards (char. 11:1), apophyseal depressions rounded (char. 13:2), pronotal base narrower than elytral base (char. 14:1), scutellum impressed (char. 19:1) and elytral intervals with transverse sculpture (char. 21:1). Additionally, one homoplasy was recovered: posterior pronotal angles strongly protruding towards elytra (char. 15:1).
Despite the fact that the species aggregated in the modestus group (Fig. 25) are homogeneous in their morphology the cladistic analysis revealed some species groups. According to the results Ectateus curtulus is a sister taxon to all other modestus group species. This relationship is supported highly supported (Jackknife support = 91; char. 3:1, 6:1, 9:1). Unfortunately, Ectateus curtulus is known only form a single specimen (holotype, female), therefore the above mentioned phylogenetic hypothesis should be reconsidered once the male specimen will be found.
The four remaining species occurred in two separate clades (Fig. 25). The first clade which consists of Ectateus ghesquierei and Ectateus crenatus is defined by the following synapomorphies: convex elytral intervals (char. 22:1) and a small body size (char. 40:1). The other clade composed of Ectateus modestus and Ectateus calcaripes comb. n. is only supported by a single homoplasy - antennomeres from 7 to 11 elongated (char. 2:1). However, these two species are very similar in general morphology – the females are almost impossible to separate or distinguish (Figs 45, 49).
According to the results of a cladistic analysis the villiersi group is characterized by following synapomorphies: anntenna robust, shorter than pronotum (char. 1:1), intercoxal process of prosternum bellied (char. 17:1) and metaventrite with coarse longitudinal depression (char. 25: 1).
Taking into consideration other significant morphological differences between modestus and villiersi groups (char. 1, 4, 8, 11, 13–15, 17, 19, 21, 25) it is reasonable to treat them as two separate genera.
Two main highly supported species group were recovered within the Selinus clade – convexipennis group and planus group (Fig. 25). The first group contains the type species (convexipennis) of Monodius Koch, 1956 (genus synonimized with Selinus by Iwan in
According to the results of a cladistic analysis the convexipennis group is characterized by following synapomorphies: 5th abdominal ventrite strongly widened (char. 26:1), female protarsi widened (char. 29:1), penis wide, at least 4 times wider than clavae (char. 32:1), apex of parameres fused (char. 37:1) and bursa copulatrix with 2 additional sacs (char. 38:1).
Selinus gravis occurred as a sister taxon to all other convexipennis group species, however this relationship is not highly supported (Fig. 25). The remaining species of the above mentioned group were divided into two separate clades (Fig. 25). The first one which consists of Selinus malaisei, Selinus medius and Selinus plicicollis is defined by the following synapomorphies: margins of elytra in basal part subparallel (elytral humeri slightly protruding outwards) (char. 24:1), denticle at the apex of the inner face of male mesotibia large (char. 31:1), apex of parameres fused and emarginated at apex (char. 37:2). This clade is also supported by a single homoplasy – elytral surface shiny (char. 20:0). The second clade (Selinus convexipennis, Selinus laevistriatus and Selinus lamottei) is defined by two homoplasies: indentation between frons and clypeus on the lateral edge deep (char. 5:1) and posterior pronotal angles strongly protruding towards elytra (char. 15:1).
According to the results of a cladistic analysis the planus group is characterized by a following synapomorphy: paraproct longer than coxites (char. 39:1). Additionally, two homoplasies were recovered: pronotum widest at the base (char. 10:1) and 5th abdominal ventrite bordered (char. 27:1).
Because of significant morphological differences between convexipennis group and planus group, especially the ones concerning the male (char. 32, 37) and female genitalia (char. 38, 39), I propose to consider them as two separate genera.
On the basis of the aforementioned results I propose to classify the analyzed ingroup species in four genera: Ectateus (based on modestus group), Monodius stat. r. (based on convexipennis group), Eleoselinus gen. n. (based on villiersi group) and Selinus (based on planus group).
A new classification and diagnostic characters of the analyzed ingroups species are presented below.
http://species-id.net/wiki/Ectateus
Anchophthalmus modestus Fairmaire, 1887; by original designation.
The following character combination is unique for Ectateus within the whole subtribe Platynotina: (1) fronto-clypeal suture coarse, clearly visible, (2) anterior pronotal angles curved outwards, (3) pronotal margins strongly erected upwards, (4) apophyseal depressions rounded, (5) pronotal base narrower than elytral base, (6) posterior pronotal angles strongly protruding towards elytra, (7) scutellum impressed, (8) elytral intervals with transverse sculpture (9) male protarsi relatively narrow, (10) male protibia slender (length/width = 4.0–5.6) and (11) clavae curved, hook-shaped.
Ectateus specimens have been collected in the following ecoregions of Central Africa (Cameroon, Central African Republic, Democratic Republic of the Congo, Equatorial Guinea, Gabonese Republic, Republic of Rwanda, Republic of the Congo, South Sudan): Albertine Rift montane forests, Angolan Miombo woodlands, Atlantic Equatorial coastal forests, East Sudanian savanna, Mount Cameroon and Bioko montane forests, Northeastern Congolian lowland forests, Northwestern Congolian lowland forests, Northern Congolian forest-savanna mosaic, Southern Congolian forest-savanna mosaic, Western Congolian forest-savanna mosaic (Fig. 41).
Ectateus calcaripes (Gebien, 1904), comb. n., Ectateus crenatus (Fairmaire, 1897), Ectateus curtulus (Fairmaire, 1893), Ectateus ghesquierei Koch, 1956 and Ectateus modestus (Fairmaire, 1887).
1 | Clypeus and genae without depressions. Pronotal margins rounded. Elytral intervals with conspicuous punctures | Ectateus curtulus |
– | Circular depressions on the lateral sides of clypeus and genae (Fig. 2). Pronotal margins sinusoidal (Fig. 5). Elytral intervals without punctures or punctures very fine (Fig. 8, 27) | 2 |
2 | Body size: 7.0–9.0 mm. Antennomeres form 7 to 11 transverse. Elytral striae with deep punctures; intervals convex (Fig. 8) | 3 |
– | Body size: 11.5–14.0 mm. Antennomeres form 7 to 11 elongated. Elytral striae with superficial punctures; intervals flat (Fig. 27) | 4 |
3 | Pronotal disc with a longitudinal groove in the middle (Fig. 5). Male protibiae as in Fig. 33; mesofemorae simple | Ectateus ghesquierei |
– | Pronotal disc without longitudinal groove. Male protibiae as in Fig. 34; mesofemorae with a large denticle on the posterior face (Fig. 35) | Ectateus crenatus |
4 | Intercoxal process protruding towards mesoventrite; peaked at the apex. Male protibiae as in Fig. 31 | Ectateus calcaripes |
– | Intercoxal process not protruding towards mesoventrite; rounded at the apex. Male protibiae as in Fig. 32 | Ectateus modestus |
Elytral disc (26, 27, 28); male mesotibia (29); male protibia (30). Monodius gravis (26, 29, 30), Ectateus calcaripes (27), Ectateus lamottei (28).
Male protibiae (31–34); male mesofemora (35); male mesotibia (36). Ectateus calcaripes (31), Ectateus modestus (32), Ectateus ghesquierei (33), Ectateus crenatus (34–36).
http://species-id.net/wiki/Ectateus_calcaripes
Figs 3, 20, 27, 31, 41, 45The types of Selinus calcaripes seems to be lost. According to the information provided by
Based only on the original species descriptions
The results of a cladistic analysis confirmed the aforementioned assumption that Ectateus curtulus and Selinus calcaripes represent two distinct morphological forms (Fig. 25). They can be easily distinguished by the structure of head (char. 3, 5), pronotum (char. 9) and elytra (elytral intervals with conspicuous punctures in Ectateus curtulus). Additionally, the results shows that Selinus calcaripes is very closely related to Ectateus modestus – which is consistent with
Taking into consideration the difficulties associated with Selinus calcaripes I propose to designate a neotype to clarify the taxonomic status of this species. Additionally, on the basis of the results of a cladistic analysis I propose to treat this taxon as a independent species – not as a synonym of Ectateus curtulus.
Neotype designated here, male (MNHN): “Rep. Centrafric., La Maboke, 2.IX.1967, L. Matile rec.”. Other material: male (MNHN): “Musée du Congo, Haut-Uele: Yebo moto, VII-1926, L. Burgeon”, male (MRAC): “Musée du Congo, Haut-Uele: Yebo moto, V-1926, L. Burgeon”, male (MRAC): “Musée du Congo, Bambesa, 15-X-1933, J. V. Leroy”, male and 3 females (MRAC): “Coll. Mus. Tervuren, Oubanghi-Chari: Bangui I/III.1968, ex. coll. Breuning”, male and female (RBINS): “R.I.Sc.Nat. Belg., I.G. 16.364”, 4 females (MNHN): “Calcaripes, Geb., det. Dr. Kaszab”, male and female (MNHN): “Boukoko, Rep. Centr Afric.”, male and female (MNHN): “La Maboke, Rep. Centr Afric.”, 2 males and female (MNHN): “Lamaboké, 10.XI.1965”, male and female (MNHN): “Lamaboké, 20-21.XI.1966”, male (MNHN): “Lamaboké, 18.XI.1965”, 2 females (HNHM): “Cameroon, Dja Reserve, Ekom, 21-26.XII. 1996, G. Hácz & G. Köszegi”, 3 males (MNB): “Kamerun Buea, 1-7.III.1912, v. Rothkirch S.G.”, female (MNB): “Kamerun, Brea 1000m, 2. - 7. III- 1912; v. Rothkirch S.G.”, female (MNB): “Kamerun, Soppo 25. II. 1912, v. Rothkirch S.G.”, 2 males (MNHN): “Lamaboké, Rep. Centrafric., V-1964, J. Carayon”, female (MNHN): “1968-69, La Maboké, Rép. Centre Afr., P. Teochhi leg.”, male (MNHN): “21.VIII.1969, La Maboké, Rép. Centre Afr., P. Teochhi leg.”, female (RBINS): “Coll. R. I. Sc. N. B., congo belge Beni, crottes d’ éléphants, 18-I-1952 Ch. Verbeke”, female (MNHN): “Muséum Paris, La Maboké, Rép. Centrafric.”, male (MRAC): “Musée du Congo, Région de Sassa, 1895-96, Colmant”, male and female (MNHN): “Cameroun, Dr. G. Nonveiller”, female (MNHN): “Muséum Paris, Congo Moyen, Rég. de M`Baiki, (D Fidao), Pitard 1919”, male (MNHN): “Uganda Prot., Mabira Forest, Chagwe., 3, 500-3, 800 ft.”, female (MNHN): “Uganda, Nimuli to, Murchison Falls”, male (MNB): “Neu-Kamerun, No. 3360-71, Tessmann S.G.”
Habitus as in Fig. 45. Body length = 11.5–14.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.4–2.6).
Dorsal side of head dull, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture coarse. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 8.0–8.6). Mentum with median part narrow. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.0–1.1). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.1–1.2). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–3.0).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.4–0.5); dull, with coarse punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and protruding outwards. Lateral margins of pronotal disc sinusoidal. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions rounded. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures; intervals non-convex, with transverse sculpture (Fig. 27). Elytral base slightly rounded. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; situated in a depression.
Intercoxal process protruding towards mesoventrite; peaked at the apex. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.2). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly narrow. Protibiae as in Fig. 31. Mesotibiae and mesofemorae with large denticle. Metafemorae with an hair fringe. Female legs simple.
Male genitalia. Parameres narrowest in the half of their length; length equal to the 0.2 of the rest of aedeagal tegmen (Fig. 20). Clavae hook-shaped (Fig. 20). Female genitalia. Paraproct equal to coxites. Bursa copulatrix with a sclerite in the distal part. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of Central Africa (Cameroon, Central African Republic, Democratic Republic of the Congo, South Sudan): Atlantic Equatorial coastal forests, East Sudanian savanna, Mount Cameroon and Bioko montane forests, Northeastern Congolian lowland forests, Northwestern Congolian lowland forests (Fig. 41).
http://species-id.net/wiki/Ectateus_crenatus
Figs 2, 8, 12, 34–36, 41, 46While describing Ectateus latipennis, Koch has noted that types of Ectateus crenatus were unknown to him. The characters used by Koch to separate those two species (body size, pronotum structure) were based only on the
The examination of the type material representing Selinus monardi resulted in similar conclusions – there are no consistent morphological characters to separate it from Ectateus crenatus.
Three specimens with “type” labels are available.
Habitus as in Fig. 46. Body length = 7.0–9.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.4–2.6).
Dorsal side of head shiny, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture coarse. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 5.7–6.5). Mentum with median part narrow. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.0–1.2). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.1–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–3.0).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6); shiny, with coarse punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and protruding outwards. Lateral margins of pronotal disc sinusoidal. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions rounded. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with coarse punctures (Fig. 8); intervals convex, with transverse sculpture and fine puncturation (Fig. 8). Elytral base slightly rounded. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; situated in a depression.
Intercoxal process protruding towards mesoventrite, peaked at the apex, slightly saddle-like. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.2). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly narrow. Protibiae as in Fig. 34. Mesotibiae and mesofemorae with large denticle (Figs 35–36). Metafemorae with an hair fringe. Female legs simple.
Male genitalia. Parameres narrowest in the half of their length; length equal to 0.2 of the rest of aedeagal tegmen. Clavae hook-shaped. Female genitalia. Paraproct equal to coxites. Bursa copulatrix without sclerites. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of Central Africa (Cameroon, Democratic Republic of the Congo, Equatorial Guinea, Gabonese Republic, Republic of the Congo): Atlantic Equatorial coastal forests, Northern Congolian forest-savanna mosaic, Northwestern Congolian lowland forests, Southern Congolian forest-savanna mosaic, Western Congolian forest-savanna mosaic (Fig. 41).
http://species-id.net/wiki/Ectateus_curtulus
Fig. 47Holotype (Fig. 47), female (MNHN): „l’Oubanghi”.
Habitus as in Fig. 47. Body length ca. 12.5 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum ca. 1.2; length ratio elytra / the middle of pronotum ca. 2.6).
Dorsal side of head dull, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture coarse. Clypeal emargination relatively deep (clypeal emargination width / depth ratio ca. 8.1). Mentum with median part wide. Submentum with short base. 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd ca. 3.0).
Pronotal disc transverse (middle of pronotum length / width ratio ca. 0.5); dull, with coarse punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and protruding outwards. Lateral margins of pronotal disc rounded. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions rounded. Pronotal hypomera dull, without punctures.
Elytra oblong (elytra length / width ratio ca. 1.2). Elytral striae with conspicuous punctures; intervals non-convex, with transverse sculpture and conspicuous punctuation (2 diameters apart). Elytral base slightly rounded. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; situated in a depression.
Intercoxal process protruding towards mesoventrite, peaked at the apex, slightly saddle-like. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite ca. 2.1). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Female legs simple.
Female genitalia. Paraproct equal to coxites. Bursa copulatrix without sclerites. Spermatheca unknown.
The only known specimen was collected in the Oubanghi (Central Africa). Because of the general character of the geographical reference it can not be translated into ecoregions.
http://species-id.net/wiki/Ectateus_ghesquierei
Figs 5, 33, 41, 48Six syntypes of Ectateus ghesquierei are available. Lectotype designation is needed to fix the taxonomic status of the genus and the species. Lectotype designated here, (MRAC): “Musée du Congo Belge, Kasai: Kondué, E. Luja”; Paralectotypes, 3 females (MRAC): same data, 2 females (MRAC): “Coll. Mus. Congo, Mayidi, 1943, Rév. P. Van Eyen”. Other material: female (MNHN): “Voka Congo, X.1977”, male and female (MNHN): “2.11.1963., No 78, sifted litter, leg. Endrõy-Younga” “Soil-Zoological Exp., Congo-Brazzaville, Kindamba, Méya, Louolo river”.
Habitus as in Fig. 48. Body length = 8.0–9.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.5–2.6).
Dorsal side of head dull, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture coarse. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 9.0–9.3). Mentum with median part narrow. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.0–1.1). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.1–1.2). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.7–3.0).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.4–0.5); dull, with coarse punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and protruding outwards. Lateral margins of pronotum sinusoidal. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions rounded (Fig. 5). Pronotal hypomera shiny; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.3). Elytral striae with conspicuous punctures; intervals convex, with transverse sculpture. Elytral base slightly rounded. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular, situated in a depression.
Intercoxal process protruding towards mesoventrite, peaked at the apex, slightly saddle-shaped. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.0–2.2). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly narrow. Protibiae as in Fig. 33. Mesotibiae with large denticle. Metafemorae without fringle of hairs. Female legs simple.
Male genitalia. Parameres narrowest in the half of their length; length equal to the 0.2 of the rest of aedeagal tegmen. Clavae hook-shaped. Female genitalia. Paraproct equal to coxites. Bursa copulatrix without sclerites. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of Central Africa (Democratic Republic of the Congo, Republic of the Congo): Southern Congolian forest-savanna mosaic, Western Congolian forest-savanna mosaic (Fig. 41).
http://species-id.net/wiki/Ectateus_modestus
Figs 32, 41, 49Lectotype (designated by Kamiński in
Habitus as in Fig. 49. Body length = 12.0–14.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.4–2.6).
Dorsal side of head dull, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture coarse. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 8.0–8.4). Mentum with median part narrow. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.0–1.1). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.7–3.0).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.4–0.5); dull, with coarse punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and protruding outwards. Lateral margins of pronotum sinusoidal. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions rounded. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures; intervals non-convex, with transverse sculpture. Elytral base slightly rounded. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; situated in a depression.
Intercoxal process not protruding towards mesoventrite, rounded at the apex. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite ca. 2.0. 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly narrow. Protibiae as in Fig. 32. Mesotibiae and mesofemorae with large denticle. Metafemorae with an hair fringe. Female legs simple.
Male genitalia. Parameres narrowest in the half of their length; length equal to 0.2 of the rest of aedeagal tegmen. Clavae hook-shaped. Female genitalia. Paraproct equal to coxites. Bursa copulatrix with a sclerite in distal part. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of Central Africa (Angola, Democratic Republic of the Congo, Republic of Rwanda, Republic of the Congo): Albertine Rift montane forests, Angolan Miombo woodlands, Atlantic Equatorial coastal forests, Western Congolian forest-savanna mosaic (Fig. 41).
Ectateus villiersi Ardoin, 1965; here designated.
The presence of the basal depressions on the pronotal disc place Eleoselinus near to (
Non-dimorphic legs distinguish Eleoselinus from: Anchophthalmops, Anchophthalmus, Ectateus, Microselinus, Monodius, Phymatoplata, Platykochius, Platymedvedevia and Selinus. From Kochogaster it can be easily distinguished by a triangular submentum and lack of sclerites in bursa copulatrix (
The following character combination is unique for Eleoselinus within the whole subtribe Platynotina: (1) anntenna robust, shorter than pronotum, (2) shallow anterior tentorial pits, (3) presence of apophyseal and basal depressions on pronotal disc, (4) intercoxal process of prosternum bellied, (5) metaventrite with coarse longitudinal depression, (6) 5th abdominal ventrite unbordered, (7) non dimorphic legs and maillary palps, (8) elytral intervals with fine punctures, (9) curved, hook-shaped clavae and (10) longitudinal coxites of ovipositor.
Body length = 10.5–13.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.4–2.9).
Dorsal side of head dull, with fine punctures (the intervals between the punctures are greater than the 2 diameters of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 5.5–5.7). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.0–1.2). Length of antennae slightly greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle ca. 0.9). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 3.2–3.5).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6); dull, with fine punctures (the intervals between the punctures are greater than the 3 diameters of the puncture). Lateral margins of pronotum narrowing towards apex. Apophyseal and basal depressions on pronotal disc present. Pronotal hypomera dull, without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.3). Elytral striae with fine punctures (the intervals between the punctures are greater than the 2 diameters of the puncture). Elytral intervals dull, non-convex, without punctures of with very fine punctuation. Elytral base slightly sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular.
Intercoxal process of prosternum bellied. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2), with longitudinal depression. In both sexes abdominal process without tubercles, relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.2). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Legs. Protarsi narrow. Other leg segments simple.
Male genitalia. Parameres narrowing towards apex; length equal to the 0.2 of the rest of aedeagal tegmen. Clavae hook-shaped. Female genitalia. Paraproct equal to coxites. Coxites narrow and long. Bursa copulatrix without sclerites.
The name is derived from the combination of Eleo (prefix indicating the genus Eleodes Eschscholtz, 1829 – a poster beetle genus of the Third International Tenebrionoidea Symposium in Tempe, Arizona) and Selinus. This genus is named to thank the Steering Committee of the Third International Tenebrionoidea Symposium: Aaron Smith (lead organizer), Rolf Aalbu, Patrice Bouchard, Kojun Kanda, Nico Franz, Warren Steiner and Quentin Wheeler.
Eleoselinus gen. n. specimens have been collected in the following ecoregion of Central Africa (Republic of the Congo): Western Congolian forest-savanna mosaic (Fig. 42).
Eleoselinus villiersi (Ardoin, 1965), comb. n. and Eleoselinus ursynowiensis (Kamiński, 2011), comb. n.
1 | Pronotal sides evenly narrowing towards apex. Elytral striae impressed on whole length (see |
Eleoselinus villiersi |
– | Pronotal sides rounded. Elytral striae impressed only near the punctures (see |
Eleoselinus ursynowiensis |
http://species-id.net/wiki/Eleoselinus_villiersi
Fig. 58Holotype, male (MNHN): “Brazzaville, Congo, V-1963”, “Muséum Paris, Mission A. Descarpentries et A. Villers, 1963-1964”. Other specimens, female (MNHN): “Allotype” same data as holotype, 7 males and 9 females (MNHN): “Juil 1959, Brazzaville, Congo. L. Vincent”, male “Kimpoko, Kongo”, female (MNHN): “Muséum Paris, Congo, Brazzaville, Mission Chari-Tchad, Dr J. Decorse 1904”.
Habitus as in Fig. 58. Body length = 10.5–12.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.7–2.9).
Dorsal side of head dull, with fine punctures (the intervals between the punctures are greater than the 4 diameters of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 5.5–5.7). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.0–1.2). Length of antennae slightly greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle ca. 0.9). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 3.2–3.5).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6), dull, with fine punctures (the intervals between the punctures are greater than the 3 diameters of the puncture). Anterior pronotal angles rounded and slightly protruding towards apex. Lateral margins of pronotal disc narrowing towards apex. Apophyseal and basal depressions on pronotal disc present. Pronotal hypomera dull, without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.3). Elytral striae with fine punctures (the intervals between the punctures are greater than the 4 diameters of the puncture). Elytral intervals dull, non-convex, without punctures. Elytral base slightly sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular.
Intercoxal process of prosternum bellied. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2); with longitudinal depression. In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.2). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Legs. Protarsi slightly narrow. Legs simple.
Male genitalia. Parameres narrowing towards apex; length equal to the 0.2 of the rest of aedeagal tegmen. Clavae hook-shaped. Female genitalia. Paraproct equal to coxites. Coxites narrow and long. Bursa copulatrix without sclerites.
This species has been collected in the following ecoregion Central Africa (Republic of the Congo).
http://species-id.net/wiki/Eleoselinus_ursynowiensis
Fig. 59Holotype, male (MNHN): “Juil 1959, Brazzaville, Congo. L. Vincent”, male “Kimpoko, Kongo”; Paratypes, 10 males and 9 females (MNHN and MIIZ): same data as holotype; female (MNHN): “Muséum Paris, Congo et Oubanghi, Decaux 1896”; male (MNHN): “Muséum Paris, Congo Franc., Env. De Brazzaville, E. Roubaud et A. Weiss, 1907”.
Because the original description (
This species has been collected in the following ecoregion Central Africa (Republic of the Congo).
Selinus convexipennis Gebien, 1904; by original designation.
The following character combination is unique for Monodius within the whole subtribe Platynotina: (1) antennomeres from 7 to 11 elongated (their length greater than the width), (2) anterior tentorial pit deep, clearly visible, (3) 5th abdominal ventrite strongly widened, (4) female protarsi widened, (5) penis wide, at least 4 times wider than clavae, (6) clavae long, their length more than half of the length of parameres, (7) apex of parameres fused and (8) bursa copulatrix with 2 additional sacs.
Monodius specimens have been collected in the following ecoregions of West and Central Africa (Burkina Faso, Cameroon, Federal Republic of Nigeria, Ivory Coast, Republic of Benin, Republic of Ghana, Republic of Liberia, Republic of Niger, Sierra Leone, Togolese Republic): Cross-Sanaga-Bioko coastal forests, Atlantic Equatorial coastal forests, Central African mangroves, Eastern Guinean forests, Guinean forest-savanna mosaic, Mount Cameroon and Bioko montane forests, Northern Congolian forest-savanna mosaic, Northwestern Congolian lowland forests, West Sudanian savanna, Western Guinean lowland forests (Figs 42–43).
Monodius convexipennis (Gebien, 1904), Monodius gravis Koch, 1956, Monodius laevistriatus (Fairmaire, 1897), comb. n., Monodius lamottei (Gridelli, 1954), comb. n., Monodius malaisei Koch, 1956, Monodius medius (Fairmaire, 1897), Monodius plicicollis (Fairmaire, 1897), comb. n.
1 | Elytral surface dull. Margins of elytra in the basal part subparallel (elytral humeri slightly protruding outwards). Denticle at the apex of the inner face of male mesotibia large (Fig. 7). Apex of parameres emarginated at the apex (Figs 15, 17) | 2 |
– | Elytral surface shiny. Margins of elytra in the basal part rounded. Denticle at the apex of the inner face of male mesotibia small or absent (e.g. Fig. 29). Apex of parameres connected (Figs 14, 16) | 4 |
2 | Pronotal apophyseal depressions coarse. Male mesofemorae with a denticle (similar to the one in Ectateus laevistriatus, Fig. 40). Parameres strongly emarginated at the apex (Fig. 15) | Monodius plicicollis |
– | Pronotal apophyseal depressions fine. Male mesofemorae wihout denticles. Parameres slightly emarginated at the apex (Fig. 17) | 3 |
3 | Male protibiae with median dilatation on the inner face (similar to the one in Monodius convexipennis, Fig. 37) | Monodius medius |
– | Male protibiae almost straight (Fig. 38) | Monodius malaisei |
4 | Body size: 16.0–19.0 mm. Elytral intervals with fine punctures (Fig. 26). Male protibiae as in Fig. 30 | Monodius gravis |
– | Body size: 12.0-14.5 mm. Elytral intervals with conspicuous punctures. Male protibiae as in Fig. 37 | 5 |
5 | Pronotal disc with two circular depressions in the middle. Aedeagal tegmen as in Fig. 19 | Monodius laevistriatus |
– | Pronotal disc wtihout circular depressions. Aedeagal tegmen as in Fig. 16, 18 | 6 |
6 | Elytral intervals with conspicuous punctures. Elytral apex as in Fig. 51. Aedeagal tegmen as in Fig. 16 | Monodius convexipennis |
– | Elytral intervals with very coarse punctures (Fig. 28). Elytral apex rounded. Aedeagal tegmen as in Fig. 18 | Monodius lamottei |
Male protibiae (37–39); male mesofemora (40). Monodius convexipennis (37), Monodius malaisei (38), Selinus planus (39), Ectateus laevistriatus (40).
http://species-id.net/wiki/Monodius_convexipennis
Figs 10, 16, 37, 42, 50, 5145 males and 41 females (MIIZ): “Mus. Zool. Polonicum Warszawa 12/45”, “Kamerun, Barombi, Conradt”, “Selinus convexipennis H. Gebien 1939”, 20 males and 12 females (MNHN): “Muséum Paris”, “Kamerun, L. Conradt, 1898-1899”, 3 males and female (MRAC): “Coll. Mus. Tervuren Cameroun, Mt Balmayo (M. Barga), XII 1972, Ex. Coll. Breuning”, female (MRAC): “Coll. Mus. Tervuren, Oubanghi-Chari”, “ex. Coll. Breuning”, 4 females (MNHN): “Muséum Paris, 1952, coll. R. Oberthur”, “Afriq Occid, Johann-Albrechts Höhe, Station Kamerun, L.Conradt, 1896”, male (MNHN): “Muséum Paris, coll. P. Ardoin, 1978”, “Cameroun Yaoundé Vadon!”, 3 males and 4 females (MNB): “Kamerun, John.Albreschtshöhe, 8.11.1898-10.3.1899, leg Conradt”, 6 males and 4 females (MNB): “N.W.Kamerun, Moliwe b. Victoria, 10-20.12.07, Frfr. v. Maltzan G.”, male and 6 females (MNB): “N.-Kamerun, Joh.-Albrechtshöhe, III.96, L.Conradt S. ”, 2 males (MNB): “S.-Kamerun, Bipindi, IX.1898, Zenker S. V.”, 2 females (MNB): “Kamerun, Bascho, Adamatz S.G.”, male and females (MNB): “Kamerun, Nokundange, 16-30 VI 05, G. Leßmann S. G.”, “1908-09.”, male (MNB): “Neu-Kamerun, Dengdeng Station, 26.III.1914, Mildbraed S.G.”, male (MNB): “Kamerun, Duala, Schäfer S.G.”, 3 males (MNHN): “Muséum Paris ex. Coll. P. Ardoin 1978”, “Duala Bothkiych”, 3 males and female (SMNS): “Malimba Pahl 91”.
Habitus as in Fig. 50. Body length = 13.0–14.5 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.4–2.6).
Dorsal side of head dull, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 7.0–7.5). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.1–1.2). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–2.9).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6), dull, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and strongly protruding towards front. Lateral margins of pronotal disc subparallel at their basal half. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions trapezoidal. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures, impressed on the whole length. Elytral intervals shiny, non-convex, with conspicuous punctures (the intervals between the punctures are smaller than the diameter of the puncture). Elytral base slightly sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum rounded.
Intercoxal process slightly protruding towards mesoventrite. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles, relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.3). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly widened. Protibiae as in Fig. 37. Mesofemorae with a small denticle at the apex. Metatibiae and Metafemorae with an hair fringe. Female legs. Protarsi slightly widened. Other leg parts simple.
Male genitalia. Parameres narrowing towards apex; length equal to the 0.3 of the rest of aedeagal tegmen (Fig. 16). Clavae straight (Fig. 16). Female genitalia. Paraproct equal to coxites. Bursa copulatrix with two sacs. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of Central Africa (Cameroon): Atlantic Equatorial coastal forests, Central African mangroves, Cross-Sanaga-Bioko coastal forests, Mount Cameroon and Bioko montane forests, Northern Congolian forest-savanna mosaic, Northwestern Congolian lowland forests (Fig. 42).
Distribution of the species of Ectateus sensu novum. The division of Afrotropical Realm into ecoregions was adopted after
Distribution of the species of Monodius convexipennis, Monodius gravis, Monodius laevistriatus, Monodius lamottei and Eleoselinus gen. n. The division of Afrotropical Realm into ecoregions was adopted after
http://species-id.net/wiki/Monodius_gravis
Figs 14, 24, 26, 29, 30, 42, 52Female (MNHN): “Schantung, Tsingtau, Miss. Mus. Steyl”, 3 males and 2 females (MNHN): “Muséum Paris, Bas Dahomey, Torricada E. Poisson 1902”, “Novembre”, female (MNB): “Süd Togo zw. Haho u. Shioftuss Laadschaften Gamme u. Gaohe Smend S.”, 2 females (MIIZ): “Museum Zool. Polonicum, Warszawa, 12/45”, “Selinus n.sp. H. Gebien det. 1939”, “Guinea”, male (MNHN): “Wydah R. P. Ménager”.
Habitus as in Fig. 52. Body length = 16.0–19.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / middle of pronotum = 2.2–2.4).
Dorsal side of head dull, with fine punctures (the intervals between the punctures are greater than the 2 diameters of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 7.1–7.5). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.1–1.2). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–2.9).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6), dull, with fine punctures (the intervals between the punctures are greater than the 3 diameters of the puncture). Anterior pronotal angles sharp and strongly protruding toward the front. Lateral margins of pronotal disc rounded. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions trapezoidal. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures; impressed on the whole length. Elytral intervals shiny, non-convex, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Elytral base slightly sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum rounded.
Intercoxal process slightly protruding towards mesoventrite. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.3). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi widened. Protibiae as in Fig. 30. Mesofemorae with a small denticle at the apex. Metafemorae with an hair fringe. Female legs. Protarsi slightly widened. Other leg parts simple.
Male genitalia. Parameres narrowing towards apex; length equal to the 0.3 of the rest of aedeagal tegmen (Fig. 14). Clavae straight (Fig. 14). Female genitalia. Paraproct equal to coxites. Bursa copulatrix with two sacs. Spermatheca with narrow ducts (Fig. 24).
This species has been collected in the following ecoregions of West Africa (Republic of Benin, Togolese Republic): Guinean forest-savanna mosaic (Fig. 42).
http://species-id.net/wiki/Monodius_laevistriatus
Figs 19, 40, 42, 53Holotype, female (MNHN): “TYPE”, “Sierra-Leone”, “Selinus laevistriatus Frm [unreadable] Type”, “Muséum Paris, 1906, Coll. L. Fairmaire”. Other material: males and 2 female (MNHN): “Fort Camp, 1070m, 26-V-1963”, “Mission ENS-IFAN, aux Monts LOMA, Sierra Leone”, “Muséum Paris, coll. P. Ardoin, 1978”, “Monodius laevistriatus, Fairm., P. Ardoin det. 1966”.
Habitus as in Fig. 53. Body length = 12.0–14.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.2–2.4).
Dorsal side of head dull, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 7.1–7.5). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.1–1.2). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–2.9).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6); dull, with fine punctures (the intervals between the punctures are greater than the 2 diameters of the puncture); with two circular depressions in the middle. Anterior pronotal angles sharp and strongly protruding towards front. Lateral margins of pronotal disc rounded. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions trapezoidal. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures; impressed on the whole length. Elytral intervals shiny, non-convex, with conspicuous punctures (the intervals between the punctures are smaller than the diameter of the puncture). Elytral base slightly sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum rounded.
Intercoxal process not protruding towards mesoventrite. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles, relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.3). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly widened. Protibiae as in Monodius convexipennis. Mesofemorae with a large denticle at the apex, mesotibia with a small denticle at the apex. Metafemorae with an hair fringe. Female legs. Protarsi slightly widened. Other leg parts simple.
Male genitalia. Parameres strongly narrowed toward apex; length equal to the 0.5 of the rest of aedeagal tegmen (Fig. 19). Clavae straight (Fig. 19). Female genitalia. Paraproct equal to coxites. Bursa copulatrix with two sacs. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of West Africa (Sierra Leone): Western Guinean lowland forests (Fig. 42).
http://species-id.net/wiki/Monodius_lamottei
Figs 18, 28, 42, 54Holotype, male (MNHN): “Muséum Paris, Nimba (Guinée), M. Lamotte II. VI. 42”, “Typus”, “Keoulenta”; Paratypes, male and female (MNHN): same data as holotype. Other material: 3 females (MNHN): “Muséum Paris, Côte d’Ivoire, Réserve du Banco, R. Paulian & C. Delamare”, 2 males and 2 females (MNHN): “Coll Mus. Tervuren, Côte d’Ivoire: Bingervillie, X.1961, J. Decelle”, “Muséum Paris Coll. P. Ardoin, 1978”, female (MRAC): “Coll Mus. Tervuren, Côte d’Ivoire: Korea, S. de Daloa, J. Decelle, VII / IX.1962”, “Ectateus laevistriatus, det. Ardoin 1965”, female (MRAC): “Coll Mus. Tervuren, Côte d’Ivoire: Adlapoté, 80 km. W. Abidjan, J. Decelle, II-1962”, “Ectateus laevistriatus, det. Ardoin 1965”, males (TMNH): “Adiopodoumé, B. Côte d’Ivoire, Ledoux”, female (MNHN): “Côte d’Ivoire”, “Muséum Paris Coll. P. Ardoin, 1978”.
Habitus as in Fig. 54. Body length = 12.0–14.5 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.2–2.4).
Dorsal side of head dull, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively shallow (clypeal emargination width / depth ratio = 10.0–11.5). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.1–1.2). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–2.9).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6), dull, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and strongly protruding towards front. Lateral margins of pronotal disc rounded. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions trapezoidal. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures, impressed on the whole length. Elytral intervals shiny, non-convex, with coarse punctures (the intervals between the punctures are smaller than the diameter of the puncture). Elytral base slightly sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum rounded.
Intercoxal process protruding towards mesoventrite. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles, relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.3). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly widened. Protibiae as in Monodius convexipennis. Mesotibiae with a small denticle at the apex. Metafemorae with an hair fringe. Female legs. Protarsi slightly widened. Other leg parts simple.
Male genitalia. Parameres extended towards apex; length equal to the 0.5 of the rest of aedeagal tegmen (Fig. 18). Clavae straight (Fig. 18). Female genitalia. Paraproct equal to coxites. Bursa copulatrix with two sacs. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of West Africa (Ivory Coast, Republic of Liberia): Eastern Guinean forests, Western Guinean lowland forests (Fig. 42).
http://species-id.net/wiki/Monodius_malaisei
Figs 9, 17, 38, 43, 55Twelve syntypes of Monodius malaisei malaisei are available. Lectotype designation is needed to fix the taxonomic status of the species and the subspecies. Lectotype designated here, male (TMNH): “Paratype No: 3224, Monodius malai-, sei Koch”, “Ob. Volta, Pundu, Olsufiew”; Paralectotypes, male (TMNH): same data as holotype, except the number referring to paratype (3226, 3227). Other material: 12 males and 15 females (MNHN): “Muséum Paris, Haute Volta, Gaoua, H. Labouret 1924”, 2 males and 2 females (ZMAS): “Poundou, Hante, Volta, Afr. Occ. Fr., Oлϲуфъев 927”, male and female (ZMAS): “Ouagadougou, Afrique Occ. Fr. Oлϲуфъев VII. VIII 927”, 3 males and 4 females (MRAC): “Coll. Mus. Tervuren, Haute Volta: Bobo-Dioulasso, 10.V.1964, R. Siffointe”, 2 males (MNHN): “Oyo Yoruba P. François”, “Muséum Paris, Coll. L. Fairmaire”, 2 males (MNHN): “Abétifi, Côte d’Ivoire”, “Muséum Paris”, 2 males (MNHN): “Tortiya, Cte Ivoire, II.59, R. Villemain”, “Muséum Paris”, 2 males (MNB): “Togo, Station Ho, Schroder S.”, male (MNHN): “Muséum Paris Soudan Franc Région Volta Sikasso- Bobo-San A. Chevalier 1900”, “Mai-Juin”, 2 males (MNHN): “Muséum Paris coll. P.Ardoin 1978”, “Monodius Selinus malaisei Koch ssp. nigeriensis Koch P. Ardoin det.1972”, “Niamey-Niger Leg. Loups”, 2 males and 2 females (MNHN): “Muséum Paris, Coll. P. Ardoin, 1978”, “IX.1971, Pabré, Haute Volta, R.P. Fernandez”.
Habitus as in Fig. 55. Body length = 13.0-14.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.2–1.3; length ratio elytra / the middle of pronotum = 2.2–2.4).
Dorsal side of head dull, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively shallow (clypeal emargination width / depth ratio = 10.0–11.5). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.1–1.2). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–2.9).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6), dull, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and strongly protruding towards front. Lateral margins of pronotal disc rounded. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions trapezoidal. Pronotal hypomera dull, without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures; impressed on the whole length. Elytral intervals dull, non-convex, with fine punctures (the intervals between the punctures are greater than the 4 diameters of the puncture). Elytral base slightly sinusoidal. Elytral humeri slightly protruding laterad. Wings absent. Scutellum rounded.
Intercoxal process protruding towards mesoventrite. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles, relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.3). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly widened. Protibiae as in Fig. 38. Mesotibiae with a large denticle at the apex. Metafemorae with an hair fringe. Female legs. Protarsi slightly widened. Other leg parts simple.
Male genitalia. Parameres extended towards apex; length equal to the 0.2 of the rest of aedeagal tegmen (Fig. 17). Clavae straight (Fig. 17). Female genitalia. Paraproct equal to coxites. Bursa copulatrix with two sacs. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of West Africa (Republic of Ghana, Ivory Coast, Burkina Faso, Federal Republic of Nigeria, Republic of Niger): Eastern Guinean forests, Guinean forest-savanna mosaic, West Sudanian savanna (Fig. 43).
Distribution of the species of Monodius malaisei, Monodius medius and Monodius plicicollis. The division of Afrotropical Realm into ecoregions was adopted after
1 | Male metatibiae curved and slightly dilated on distal half | Monodius malaisei malaisei Koch, 1956 |
– | Male metatibiae straight | Monodius malaisei nigeriensis Koch, 1956 |
http://species-id.net/wiki/Monodius_medius
Figs 7, 23, 43, 56Holotype, male (MNHN): “Type”, “Muséum Paris, Paris, Collection Leon Fairmaire, 1906”, “Selinus medius Fm guinea, Acut. det.”. Other material: 6 males and 2 females (RBINS): “Selinus, angulatipes, det. H. Gebien 1913, n. sp.”, “Type”, “Asenté Akem, (Ashanti), Guineé” (Syntypes of Selinus angulatipes Gebien, 1921), 11 males and 6 females (MNHN): “Lamto, Pacobo, V. 1968”, “Côte d’Ivoire, CL. Girard Col.”, male (MRAC): “Coll. Mus. Tervuren, Côte d’Ivoire, Kossou 18.2.1975, R. Jocqué”, 2 males (MRAC): “Coll. Mus. Tervuren, Côte d’Ivoire: Bouaké, VII-, 1977, P. M. Elsen”, male and female (MRAC): “Coll. Mus. Tervuren, Togo: Missahoué 650 m., VI.1963, Mme Y. Schach”.
Habitus as in Fig. 56. Body length = 12.0–15.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.2–1.3; length ratio elytra / the middle of pronotum = 2.2–2.4).
Dorsal side of head dull, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively shallow (clypeal emargination width / depth ratio = 10.0–11.5). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.1–1.2). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–2.9).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6); dull, with fine punctures (the intervals between the punctures are greater than the 2 diameters of the puncture). Anterior pronotal angles sharp and strongly protruding towards front. Lateral margins of pronotal disc rounded. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions trapezoidal. Pronotal hypomera dull, without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures, impressed on the whole length. Elytral intervals dull, non-convex, with fine punctures (the intervals between the punctures are greater than the 4 diameters of the puncture). Elytral base slightly sinusoidal. Elytral humeri slightly protruding laterad. Wings absent. Scutellum rounded.
Intercoxal process not protruding towards mesoventrite. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles, relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.3). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 3 diameters of the puncture).
Male legs. Protarsi slightly widened. Protibiae as in Monodius convexipennis. Mesotibiae with a large denticle at the apex. Metafemorae with an hair fringe. Female legs. Protarsi slightly widened. Other leg parts simple.
Male genitalia. Similar as in Monodius malaisei. Female genitalia. Paraproct equal to coxites. Bursa copulatrix with two sacs (Fig. 23). Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of West Africa (Republic of Ghana, Ivory Coast): Eastern Guinean forests, Guinean forest-savanna mosaic (Fig. 43).
http://species-id.net/wiki/Monodius_plicicollis
Figs 4, 13, 15, 22, 43, 57Holotype, male (MNHN): “Selinus plicicollis, Fairm, Togo”, “Togoland L. Conradt 1892-1893”, “Muséum Paris, 1906, Coll. L. Fairmaire”; Paratype, male (MNHN): same data as holotype. 11 males and 10 females (MRAC): “Coll. Mus. Tervuren Togo: Niamtougou, 21/24-VII-1969, F. Puylaert”, female (MIIZ): “Mus. Zool. Polonicum Warszawa 12/45”, “Selinus plicicollis H.Gebien det.1939”, “Dahomey”, male and female (MNHN): “Museum Paris, Moyen-Dahomey, Plateau de Zaguanado, Saison des Drages te. Des Tornades P. Ducorps 1910”, female (MNB): “Togo Bismarckburg, 12-14.IV.93, L. Conradt S.”, male and female (MNHN): “Abboekuta, P. Francois”, “Museum Paris, Coll. Ch. Alluaud, coll. L.Fairmaire 1906”, female (TMNH): “Oyo Yoruba P. François”, “Selinus plicicollis”, 7 males and 10 females (MNHN): “Togoland L. Conradt 1892-1893”, “Muséum Paris, 1906, Coll. L. Fairmaire”, 2 females (MNHN): “26.XII.1975, Akoumapé, Togo, J-Cl. Martin”, “Museum Paris, coll. P. Ardoin, 1978”, 6 males and 7 females (MNHN): “Museum Paris Dahomey env. De Porto-Novo, Waterlot 1910”.
Habitus as in Fig. 57. Body length = 13.0–18.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.2–1.3; length ratio elytra / the middle of pronotum = 2.2–2.4).
Dorsal side of head dull, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively shallow (clypeal emargination width / depth ratio = 10.0–11.5). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.1–1.2). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–2.9).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.5–0.6); dull, with fine punctures (the intervals between the punctures are greater than the 2 diameters of the puncture). Anterior pronotal angles sharp and strongly protruding towards front. Lateral margins of pronotal disc rounded. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions trapezoidal; very coarse. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures; impressed on the whole length. Elytral intervals dull, non-convex, with fine punctures (the intervals between the punctures are greater than the 4 diameters of the puncture). Elytral base slightly sinusoidal. Elytral humeri slightly protruding laterad. Wings absent. Scutellum rounded.
Intercoxal process not protruding towards mesoventrite. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.3). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 3 diameters of the puncture).
Male legs. Protarsi slightly widened. Protibiae straight. Mesotibiae and mesofemorae with a large denticle at the apex. Metafemorae with an hair fringe. Female legs. Protarsi slightly widened. Other leg parts simple.
Male genitalia. Parameres extended towards apex; length equal to the 0.2 of the rest of aedeagal tegmen (Fig. 15). Clavae straight (Fig. 15). Female genitalia. Paraproct equal to coxites (Fig. 22). Bursa copulatrix with two sacs. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of West Africa (Togolese Republic, Republic of Benin, Federal Republic of Nigeria): Eastern Guinean forests, Guinean forest-savanna mosaic, West Sudanian savanna (Fig. 43).
http://species-id.net/wiki/Selinus
Opatrum planum Fabricius, 1792; designated by
The following character combination is unique for Selinus within the whole subtribe Platynotina: (1) anterior tentorial pit deep, clearly visible, (2) antennomeres from 7 to 11 elongated (their length greater than the width), (3) pronotum widest at the base, (4) 5th abdominal ventrite bordered, (5) paraproct longer than coxites, (6) clavae long, their length more than half of the length of parameres.
Specimens of this genus have been collected in the following ecoregions of West Africa (Ivory Coast, Republic of Benin, Republic of Ghana, Republic of Guinea, Republic of Mali, Togolese Republic): Eastern Guinean forests, Guinean forest-savanna mosaic, West Sudanian savanna, Western Guinean lowland forests (Fig. 44).
Distribution of the species of Selinus sensu novum. The division of Afrotropical Realm into ecoregions was adopted after
Selinus planus (Fabricius, 1792) and Selinus striatus (Fabricius, 1794).
1 | Body size: 12.0–14.0 mm. Pronotal sides evenly narrowing towards apex. Pronotal and elytral surface with fine punctures (the intervals between the punctures are greater than the 3 diameters of the puncture). Male protibiae as in Fig. 39 | Selinus planus |
– | Body size: 10.0–11.0 mm. Pronotal sides parallel in their basal half. Pronotal and elytral surface with conspicuous punctures (the intervals between the punctures are smaller than the diameter of the puncture). Male protibiae with very shallow dilatation near the middle | Selinus striatus |
http://species-id.net/wiki/Selinus_planus
Figs 1, 6, 39, 44, 60Paratype, female (MNHN): “Guinée, coll. R. Oberthür, ex coll. Deyrolle”. Other material: 10 males and 9 females (MNHN): “Muséum Paris, Bas Dahomey, Torricada E. Poisson 1902”, “Novembre”, 6 males and 5 females (MNHN): “Muséum Paris Dahomey env. De Porto-Novo, Waterlot 1910”, 3 males (MNHN): “Muséum Paris coll. P. Ardoin 1978”, “15/20.X.1967, Takoradi, Ghana, Cl. Besnard leg.”, “♂”, female (MNHN): “Ashanti”, “Muséum Paris, (Coll. Ch. Alluaud), coll. L.Fairmaire 1906”, 2 males and 3 females (MNB): “Togo Amedzowe”, 2 males and female (MRAC): “Coll. Mus Tervuren, Togo: Missahoué 650 m., VI.1963, Mme Y. Schach”, female (MNHN): “Under log, river bank”, “Akuse, Gold Coast, 23-II-29”, “Museum Paris”, male (MNHN): “Abétifi, Côte d’Ivoire”, “Muséum Paris”, 2 females (MRAC): “Coll. Mus. Tervuren, Ghana: Volta River, 12.VII.1964, G. Marlier”, 2 males (MNHN): “Togo, Palime, Forêt de Klouto, 20-24-IV-74 S. Vit”, female (MRAC): “Coll. Mus. Tervuren, Côte d’Ivoire: Eremankono, S. de Divo, J. Decelle VII-1962”, male and female (MNHN): “Goldküste, Ostertag”, 2 males (MNHN): “Addah.”, “Muséum Paris”, male (MNHN): “Gold Coast”, “Koumassi”, 3 males and female (MNHN): “Dahomey, Athieme, J. M. Renou 1898”, female (BMNH): “Aburi, Gold Coast, W. H. Patterson., 1914-29”, female (MNHN): “Talanzoa”, “Muséum Paris, Nimba (Guinée), M. Lamotte II. VI. 42”.
Habitus as in Fig. 60. Body length = 12.0–14.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.7–3.0).
Dorsal side of head dull, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 4.0–4.5). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.0–1.1). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–3.0).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.4–0.5); dull, with fine punctures (the intervals between the punctures are greater than the 3 diameters of the puncture). Anterior pronotal angles sharp and slightly protruding towards apex. Lateral margins of pronotum narrowing towards apex. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions trapezoidal. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures (sometimes absent). Elytral intervals shiny, non-convex; with conspicuous punctures (the intervals between the punctures are greater than the 3 diameters of the puncture). Elytral base slightly sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular.
Intercoxal process protruding towards mesoventrite. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.2). 5th abdominal ventrite with complete bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly widened. Protibiae as in Fig. 39. Metafemorae with an hair fringe. Female legs. Protarsi slightly widened. Other leg parts simple.
Male genitalia. Parameres narrowing towards apex; length equal to the 0.2 of the rest of aedeagal tegmen. Clavae straight. Female genitalia. Paraproct longer than coxites. Spermatheca with narrow ducts.
This species has been collected in the following ecoregions of West Africa (Ivory Coast, Republic of Benin, Republic of Ghana, Republic of Guinea, Togolese Republic): Eastern Guinean forests, Guinean forest-savanna mosaic, West Sudanian savanna (Fig. 44).
http://species-id.net/wiki/Selinus_striatus
Figs 11, 21, 44, 613 females (MNHN): “Muséum Paris, Côte d’Ivoire, Reserve du Banco, R. Paulian & C. Delamare”, 2 males and female (MNHN): “Muséum Paris (coll. Ch. Alluaud) coll. L. Fairmaire 1906”, “Assinie Côte occid. Afrique Ch. Alluaud 1886”, male and female (MNHN): “Coll Mus. Tervuren, Côte d’Ivoire: Bingervillie, X.1961, J. Decelle”, “Muséum Paris Coll. P. Ardoin, 1978”, female (MRAC): “Cerole de Sassandra, Cote d’Ivoire 4.1962, J.Hamon Orstom Rec.”, “Selinus striatus Fab. P. Ardoin det.1963”, “♀”, female (MRAC): “Coll. Mus. Tervuren Côte d’Ivoire: Amanikro, 50 km. N.W. Abengourou J.Decelle V/VI.1961, “Récolté sur cacaoyer”, “Selinus striatus Fab. P.Ardoin det.1965”, male and female (MNHN): “Muséum Paris (coll. Ch.Alluaud) coll. L. Fairmaire 1906”, “Rhobomp, Sierra Leone”, male (MNHN): “Muséum Paris, Iles se Los, Tamara, J. Serand 1913”, male (MHNL): “Dimbokru, Côte d’Ivoire, G. Skibiski, Lyon”, male (MNHN): “Muséum Paris coll. P. Ardoin 1978”, “3.V.1967, Takoradi, Ghana, Cl.Besnard leg.”, “♂”.
Habitus as in Fig. 61. Body length = 10.0–11.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.7–2.9).
Dorsal side of head dull, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture fine. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 4.0–4.4). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.1–1.3). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–3.0).
Pronotal disc transverse (middle of pronotum length / width ratio = 0.4–0.5); dull, with fine punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and slightly protruding towards apex. Lateral margins of pronotal disc narrowing towards apex. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions trapezoidal. Pronotal hypomera dull; without punctures.
Elytra oblong (elytra length / width ratio = 1.1–1.2). Elytral striae with fine punctures (sometimes absent). Elytral intervals shiny, non-convex, with conspicuous punctures (the intervals between the punctures are smaller than the diameter of the puncture). Elytral base slightly sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular.
Intercoxal process protruding towards mesoventrite. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1–2.2). 5th abdominal ventrite with complete bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
Male legs. Protarsi slightly slightly widened. Male protibiae with very shallow dilatation near the mddle. Metafemorae with an hair fringe. Female legs. Protarsi slightly widened. Other leg parts simple.
Male genitalia. Parameres narrowing towards apex; length equal to the 0.2 of the rest of aedeagal tegmen (Fig. 21). Clavae straight (Fig. 21). Female genitalia. Paraproct longer than coxites. Spermatheca and bursa copulatrix as in Selinus planus.
This species has been collected in the following ecoregions of West Africa (Ivory Coast, Republic of Ghana, Republic of Guinea, Republic of Mali): Eastern Guinean forests, Guinean forest-savanna mosaic, West Sudanian savanna, Western Guinean lowland forests (Fig. 44).
Body habitus: Ectateus calcaripes (45), Ectateus crenatus (46), Ectateus curtulus (47), Ectateus ghesquierei (48) and Ectateus modestus (49).
Body habitus: Monodius convexipennis (50), Monodius gravis (52), Monodius laevistriatus (53) and Monodius lamottei (54). Apex of elytra of Monodius convexipennis (51).
Body habitus: Monodius malaisei (55), Monodius medius (56), Monodius plicicollis (57), Eleoselinus villiersi (58) and Eleoselinus ursynowiensis (59).
Body habitus: Selinus planus (60) and Selinus striatus (61).
I am grateful to Laurent Soldati (France), Patrice Bouchard (Kanada) and the anonymous reviewer and for the valuable comments to the previous versions of this manuscript.