Holotype, male (Fig. 1a). Length 1.74 mm. Color black; elytra and pronotum with greenish-bronze lustre; antenna and tarsi ferruginous. Body subovate; in dorsal aspect and in profile with weak constriction between pronotum and elytron. Rostrum with indistinct, irregular, longitudinal ridges. Pronotum with weak subapical constriction; densely punctate. Elytra with striae deeply impressed, intervals weakly costate, subglabrous. Meso- and metafemur ventrally weakly dentate. Metafemur subapically without stridulatory patch. Aedeagus (Fig. 1b) with apex extended, pointed; body in profile at middle with marked depression; transfer apparatus relatively complex, symmetrical; ductus ejaculatorius with bulbus. Intraspecific variation. Length 1.50–1.74 mm. Female rostrum dorsally subglabrous.

Holotype (MZB): ARC0530 (EMBL # FN429236), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, S02°31.912', E140°30.416', 785 m, 02-XII-2007, sifted. Paratypes (SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani: 3 exx, ARC0531 (EMBL # FN429237), same data as holotype; 2 exx, ARC0550 (EMBL # FN429256), S02°31.776', E140°30.215', 945 m, 21-XI-2007, sifted; 4 exx, ARC0564 (EMBL # FN429270), S02°31.912', E140°30.416', 785 m, 02-XII-2007, sifted.

Jayapura Reg. (Cyclops Mts). Elevation: 785–945 m.

Sifted from leaf litter in montane forest.

This epithet is based on a combination of the Latin adjective aeneus (of bronze) and the noun penna (wing, elytron) and refers to its coloration.

Trigonopterus aeneipennis Riedel, sp. n. was coded as “Trigonopterus sp. 50” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spax” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 2a). Length 1.91 mm. Color black with greenish-bronze lustre; antenna and tarsi ferruginous. Body subrhomboid; in dorsal aspect and in profile with distinct constriction between pronotum and elytron. Rostrum with weak median wrinkle; epistome simple. Eyes small. Pronotum with weak subapical constriction; disk sparsely punctate. Elytra with strial punctures distinct; intervals flat, subglabrous; interval 7 subapically costate, forming angulate ridge; sutural interval apically with knob. Meso- and metafemur ventrally weakly dentate. Metafemur subapically without stridulatory patch. Onychium ca. 1.8× longer than tarsomere 3. Aedeagus (Fig. 2b) with apex weakly asymmetrical; long median extension somewhat shifted to the left; transfer apparatus hook-shaped; ductus ejaculatorius with indistinct bulbus, torn off in holotype. Intraspecific variation. Length 1.63–1.91 mm. Female rostrum dorsally subglabrous, sparsely punctate.

Holotype (SMNK): ARC1089 (EMBL # HE615719), PAPUA NEW GUINEA, Simbu Prov., Karimui Dist., Haia, S06°43.948', E144°59.856', 915 m, 26-IX-2009. Paratypes (NAIC, SMNK, ZSM): PAPUA NEW GUINEA, Simbu Prov.: 12 exx, ARC1090 (EMBL # HE615720), ARC1091 (EMBL # HE615721), same data as holotype; 1 ex, Haia, S06°41.624', E145°00.728', 960 m, 25-IX-2009, sifted; 1 ex, ARC1105 (EMBL # HE615734, Haia, S06°41.018', E145°00.995', 1090 m, 04-X-2009.

Simbu Prov. (Haia). Elevation: 915–1090 m.

Sifted from leaf litter in primary forest.

This epithet is based on the Latin adjective aeneus (of bronze) and refers to its coloration.

Trigonopterus aeneus Riedel, sp. n. was coded as “Trigonopterus sp. 267” by Tänzler et al. (2012).

Holotype, male (Fig. 3a). Length 2.04 mm. Color black; legs and antenna ferruginous. Body subglobose; in dorsal aspect with weak constriction between pronotum and elytron; with more distinct constriction in profile. Rostrum in basal half with 3 ridges posteriorly continued to and uniting on forehead; apical half scabrous; epistome smooth, forming indistinct transverse ridge. Pronotum punctate-rugose, interspaces between punctures forming longitudinal wrinkles; with distinct subapical constriction. Elytra subglabrous, striae deeply impressed, intervals costate; apex extended ventrad, beak-shaped. Femora edentate. Metafemur with denticulate dorsoposterior edge, subapically without stridulatory patch. Abdominal venter steeply flexed dorsad, concealed in elytral capsule. Aedeagus (Fig. 3b) with apex medially pointed; body in apical half with broad depression visible in lateral aspect; in basal half with x-shaped sclerite; subglabrous, with sparse indistinct setae; transfer apparatus markedly flagelliform, longer than body, curled, pointing apicad. Intraspecific variation. Length 1.86–2.04 mm.

Holotype (MZB): ARC1688 (EMBL # HE615975), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Angkasa indah, S02°30.346', E140°42.087', 490 m, 28-VI-2010, sifted. Paratypes (SMNK): 4 exx, ARC1689 (EMBL # HE615976), same data as holotype.

Jayapura Reg. (Cyclops Mts). Elevation: 490 m.

Sifted from leaf litter in primary forest.

This epithet is based on the Greek noun agathis (ball, globe) in apposition and refers to the species' habitus.

Trigonopterus agathis Riedel, sp. n. was coded as “Trigonopterus sp. 109” by Tänzler et al. (2012).

Holotype, male (Fig. 4a). Length 2.55 mm. Color black with slight bluish lustre; legs deep ferruginous, antenna light ferruginous. Body ovate; without constriction between pronotum and elytron; in profile evenly convex. Rostrum dorsally in basal half with median ridge, coarsely punctate and with sparse cream-colored scales; apically smooth, with small punctures. Eyes large. Pronotum dorsally densely punctate with minute punctures; laterally with larger punctures, anteriorly above procoxa squamose with subtriangular cream-colored scales. Elytra dorsally subglabrous, punctures minute, striae hardly visible; laterally strial punctures deep, lateral 5 striae distinct. Femora elongate, edentate. Profemur converging from base to apex. Metafemur with simple dorsoposterior edge; subapically without stridulatory patch. Tibiae simple, without rows or brushes of long setae. Metathoracic and abdominal venter with sparse cream-colored scales. Aedeagus (Fig. 4b) symmetrical, apically pointed; ductus ejaculatorius without bulbus. Intraspecific variation. Length 2.54–2.84 mm. Female rostrum dorsally subglabrous, in apical half with minute punctures.

Holotype (MZB): ARC0488 (EMBL # FN429195), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°31.7', E140°30.3', 850–1000 m, 30-XI-2007, beaten. Paratypes (ARC, SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani: 4 exx, ARC0489 (EMBL # FN429196), ARC0490 (EMBL # FN429197), same data as holotype; 2 exx, S02°31.7', E140°30.3', 860–1150 m, 21-XI-2007, beaten; 3 exx, S02°31.6', E140°30.4', 900–1100 m, 28-XI-2007, beaten; 5 exx, S02°31.6', E140°30.4', 1000–1200 m, 30-XI-2007, beaten; 5 exx, 950–1450 m, 03-X-1992; 1 ex, 1100–1600 m, 05-X-1991; 1 ex, 700–1400 m, 23-XII-2004; 5 exx, 300–1400 m, 10-VIII-1991.

Jayapura Reg. (Cyclops Mts). Elevation: 700–1100 m.

Collected by beating foliage in montane forests.

This epithet is based on the Latin adjective agilis (quick) and refers to the behavior of this species and its close relatives.

Trigonopterus agilis Riedel, sp. n. was coded as “Trigonopterus sp. 14” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spm” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 5a). Length 2.26 mm. Color black; tarsi and antenna ferruginous. Body subrhomboid; in dorsal aspect with marked constriction between pronotum and elytron; in profile with shallow constriction. Rostrum dorsally dull, with 3 irregular ridges, with rows of erect setae; epistome forming angulate ridge and median denticle. Pronotum with distinct subapical constriction, sparsely punctate with small setiferous punctures, behind subapical constriction scales larger, subclavate, yellowish. Elytra subglabrous, striae weakly impressed, with minute punctures, each puncture with minute seta; apex bordered by curved ridge, median suture incised. Femora edentate. Meso- and metatibia in basal half widened, subapically narrowed. Metafemur with denticulate dorsoposterior edge, subapically without stridulatory patch. Aedeagus (Fig. 5b) widening towards subtruncate, medially pointed apex; transfer apparatus flagelliform, curved, shorter than body; ductus ejaculatorius basally swollen, without bulbus. Intraspecific variation. Length 2.00–2.28 mm. Color of elytra and legs black or ferruginous. Female rostrum dorsally punctate-rugose, in apical half without setae; epistome simple.

Holotype (MZB): ARC0556 (EMBL # FN429262), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°31.182', E140°30.542', 1510 m, 30-XI-2007, sifted. Paratypes (ARC, SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani: 34 exx, ARC0557 (EMBL # FN429263), ARC0558 (EMBL # FN429264), ARC0559 (EMBL # HE615318), same data as holotype; 14 exx, S02°31.281', E140°30.535', 1420 m, 30-XI-2007, sifted; 7 exx (1 marked ARC0094), 1320 m, 23-XII-2004, sifted; 5 exx, 300–1400 m, 10-VIII-1991.

Jayapura Reg. (Cyclops Mts). Elevation: 1320–1510 m.

Sifted from leaf litter in montane forest.

This epithet is based on a combination of the Latin adjective amplus (wide) and the noun penna (wing, elytron) and refers to the basally widened elytra.

Trigonopterus amplipennis Riedel, sp. n. was coded as “Trigonopterus sp. 40” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus span” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 6a). Length 2.49 mm. Color black; antenna and tarsi ferruginous. Body subovate; in dorsal aspect and in profile with distinct constriction between pronotum and elytron. Eyes large. Rostrum medially with two rows of course squamiferous punctures; pair of lateral furrows with row of larger overlapping almond-shaped white scales. Pronotum densely punctate; dorsally punctures containing inconspicuous setae, anterolaterally with scattered white scales. Elytra with striae distinct; intervals flat, subglabrous, each with one row of minute punctures. Femora ventrally with acute tooth. Mesofemur and metafemur dorsally sparsely squamose with white scales. Metafemur subapically with stridulatory patch. Metatibia at middle curved ventrad, subapically with brush of long setae; uncus hook-like extended, curved ventrobasad. Aedeagus (Fig. 6b) with apodemes 2.5 × as long as body; sides of body weakly bisinuate, converging; apex extended, pointed, markedly curved ventrad, sinuate in profile; transfer apparatus flagelliform, subequal to body of aedeagus; ductus ejaculatorius without bulbus.

Holotype (SMNK): ARC1115 (EMBL # HE615744), PAPUA NEW GUINEA, Simbu Prov., Karimui Dist., Haia, Supa, S06°40.078', E145°03.207' to S06°39.609', E145°03.012', 1220–1450 m, 02-X-2009.

Simbu Prov. (Haia). Elevation: ca. 1220–1450 m.

Collected by beating foliage in montane forest.

This epithet is based on a combination of the Latin nouns ancora (anchor) and uncus (hook; tibial uncus) in apposition and refers to the species´ remarkable metatibia.

Trigonopterus ancoruncus Riedel, sp. n. was coded as “Trigonopterus sp. 78” by Tänzler et al. (2012).

Holotype, male (Fig. 7a). Length 2.63 mm. Color black; antenna light ferruginous; tarsi and tibiae deep ferruginous. Body subovate-hexa-goniform; in dorsal aspect with shallow constriction between pronotum and elytron; in profile evenly convex. Rostrum in basal half with median and pair of submedian carinae; laterally somewhat flattened; with sparse suberect scales; epistome with transverse, angulate ridge. Pronotum with anterior margin curved dorsad, forming lateral angles; disk subglabrous, anteriorly densely coarsely punctate, towards sides forming edges; laterally in front and behind procoxa with cavity. Elytra subglabrous; intervals flat, with minute punctures; striae weakly incised on disk, towards glabrous sides forming edges; interval 7 subapically forming edge, apex angulate. Femora edentate. Metafemur dorsally squamose with indistinct suberect scales; in apical third without transverse row of setae, subapically with stridulatory patch. Aedeagus (Fig. 7b) apically subangulate, median tip truncate; with complex, symmetrical transfer apparatus; ductus ejaculatorius with indistinct bulbus. Intraspecific variation. Length 2.60–2.63 mm. No female specimen available.

Holotype (SMNK): ARC1088 (EMBL # HE615718), PAPUA NEW GUINEA, Simbu Prov., Karimui Dist., Haia, S06°41.216', E145°00.945', 965 m, 27-IX-2009, sifted. Paratype (NAIC): 1 ex, ARC1098 (EMBL # HE615727): PAPUA NEW GUINEA, Simbu Prov., Karimui Dist., Haia, S06°40.976', E145°00.979', 1135 m, 27-IX-2009, sifted.

Simbu Prov. (Haia). Elevation: 965–1135 m.

. Sifted from leaf litter in primary forest.

This epithet is based on the Latin adjective angulatus (with angles) and refers to the outline of its body in dorsal aspect.

Trigonopterus angulatus Riedel, sp. n. was coded as “Trigonopterus sp. 194” by Tänzler et al. (2012).

Holotype, male (Fig. 8a). Length 2.83 mm. Color black; legs and antenna ferruginous. Body elongate; in dorsal aspect and in profile with distinct constriction between pronotum and elytron. Rostrum basally with distinct median and pair of submedian carinae, in apical ¼ relatively smooth, basally sparsely squamose. Pronotum densely punctate with large punctures except small glabrous area at center; interspaces smaller than puncture´s diameter. Elytra dorsally punctate with deep punctures; near base and along suture more densely punctate, punctation confuse; basal margin near elytral suture somewhat swollen and glabrous; striae impressed as fine lines; laterally punctation relatively sparse, behind humerus with row of deep punctures. Femora edentate. Profemur in basal third posteriorly with callus. Metafemur subapically without stridulatory patch. Aedeagus (Fig. 8b) apically subangulate, medially truncate, with pair of stout setae; body flattened, sides subparallel; ductus ejaculatorius without bulbus. Intraspecific variation. Length 2.24–2.83 mm. Body of females more slender. Female rostrum dorsally subglabrous, punctate, basally sparsely squamose. Female abdominal ventrites 1–2 flat.

Holotype (MZB): ARC0626 (EMBL # FN429283), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°32.3', E140°30.4', 350–620 m, 19-XI-2007, beaten. Paratypes (ARC, SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts: 4 exx, Sentani, ARC0455 (EMBL # FN429166), ARC0456 (EMBL # FN429167), ARC0479 (EMBL # FN429186), ARC0631 (EMBL # FN429286), S02°31.3', E140°30.5', 1200–1420 m, 30.XI.2007; 2 exx, ARC0627 (EMBL # FN429284), ARC0630 (EMBL # FN429285), same data as holotype; 1 ex, Cyclops Mts, Angkasa indah ARC1693 (EMBL # HE615980), S02°30.355', E140°42.103' to S02°30.346', E140°42.087', 450–520 m, 28-VI-2010; 5 exx (1 marked as “ARC0046”), 950–1450 m, 03-X-1992; 3 exx, 400–800 m, 07-VIII-1992; 1 ex, 1100–1600 m, 05-X-1991; 2 exx, 300–1400 m, 10-VIII-1991; 6 exx, 1200–1400 m, 09-VIII-1992; 3 exx, Lereh, 500–1000 m, 26-I-1996.

Jayapura Reg. (Cyclops Mts; Lereh). Elevation: 520–1200 m.

Collected by beating foliage in montane forests.

This epithet is based on the Latin adjective angustus (narrow) and refers to its habitus.

Trigonopterus angustus Riedel, sp. n. was coded as “Trigonopterus sp. 12” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spl” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 9a). Length 4.90 mm. Color black except basal half of elytron bright orange; tarsi and antenna ferruginous. Body elongate; in dorsal aspect with distinct constriction between pronotum and elytron. Rostrum slender, basally scabrous with distinct median ridge and pair of irregular submedian ridges, in apical 1/3 punctate. Pronotum large, subquadrate, densely punctate. Elytra densely punctate with small irregular punctures; striae partly impressed as fine lines, partly indistinct. Femora dentate with acute tooth. Profemur enlarged, subovate, posteriorly concave and polished, in basal third with callus. Metafemur subapically with stridulatory patch. Thoracic and abdominal venter concave, densely punctate, sparsely setose with thin erect setae. Aedeagus (Fig. 9b) with apodemes 3.0 × as long as body; apex rounded, at middle subtruncate; transfer apparatus flagelliform, stout, more than 2 × as long as body; ductus ejaculatorius basally swollen, without bulbus. Intraspecific variation. Length 4.08–4.90 mm. Female rostrum dorsally subglabrous, sparsely punctate, at base coarsely punctate. Male pronotum larger, with sides posteriorly almost straight; female pronotum smaller, with sides rather convex. Male elytra narrower, converging apicad; female elytra wider, lateral contour convex. Female abdominal venter flat, subglabrous, punctate, with sparse short recumbent setae.

Holotype (SMNK): ARC1136 (EMBL # HE615765), PAPUA NEW GUINEA, Simbu Prov., Karimui Dist., Haia, Supa, S06°39.905', E145°03.880' to S06°39.796', E145°03.873', 1220–1320 m, 01-X-2009. Paratypes (NAIC, SMNK, ZSM): Simbu Prov., Karimui Dist., Haia, Supa: 2 exx, ARC1137 (EMBL # HE615766), ARC1138 (EMBL # HE615767), same data as holotype; 1 ex, Haia, Supa station, S06°40.047', E145°03.464' to S06°39.905', E145°03.880', 1075–1220 m, 01-X-2009, beaten; 1 ex, Haia, Supa station, S06°39.815', E145°03.169' to S06°39.609', E145°03.012', 1240–1450 m, 30-IX-2009, beaten.

Simbu Prov. (Haia). Elevation: 1220–1240 m.

Collected by beating foliage in montane forests.

This epithet is based on the Latin adjective apicalis (pertaining to the apex) and refers to the species´ contrasting elytral coloration.

Trigonopterus apicalis Riedel, sp. n. was coded as “Trigonopterus sp. 259” by Tänzler et al. (2012).

Holotype, male (Fig. 10a). Length 3.53 mm. Color black; tarsi and antenna ferruginous. Body ovate; almost without constriction between pronotum and elytron; in profile evenly convex. Rostrum in basal third swollen, dorsally coarsely punctate and with indistinct median carina; apically shining, punctures more shallow, with longitudinal furrows. Pronotum densely punctate except along impunctate midline. Elytra punctate with small punctures, along basal margin with transverse row of deeper and denser punctures; striae impressed as fine lines; lateral stria behind humeri simple, not deepened. Profemur and mesofemur with anteroventral ridge ending abruptly 1/3 before apex. Metafemur with anteroventral ridge ending with bluntly angled tooth 1/3 before apex; with denticulate dorsoposterior edge; subapically with stridulatory patch. Mesotibia ventrally with spine in subapical 1/3, apically with premucro. Metatibia apically with uncus and larger subtriangular premucro. Aedeagus (Fig. 10b) with distinct, symmetrical transfer-apparatus; ductus ejaculatorius with bulbus. Intraspecific variation. Length 2.63–3.59 mm. Female rostrum basally simple, not swollen. Female mesotibia in subapical 1/3 simple, apically without premucro. Female metatibia without premucro.

Holotype (MZB): ARC570 (EMBL#FN429273), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°32.0', E140°30.4', 700–900m, 02.XII.2007, beaten. Paratypes (ARC, SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani: 9 exx, ARC0464 (EMBL # FN429174), ARC0496 (EMBL # FN429203), ARC0498 (EMBL # FN429205), S02°31.8', E140°30.5', 600–900 m, 28.XI.2007; 12 exx, S02°31.6', E140°30.4', 900–1100 m, 28-XI-2007, beaten; 2 exx, S02°31.6', E140°30.4', 1000–1200 m, 30-XI-2007, beaten; 1 ex, S02°31.7' E140°30.3', 860–1150 m, 21-XI-2007, beaten; 2 exx, S02°32.0', E140°30.4', 700–900 m, 02-XII-2007, beaten; 4 exx, 950–1450 m, 03-X-1992; 1 ex, S02°31.794', E140°30.190', 800–860 m, 21-XI-2007, “Mim2”, beaten; 7 exx, 1100–1600 m, 05-X-1991; 1 ex, 1200–1400 m, 09-VIII-1992; 3 exx, 300–1400 m, 10-VIII-1991; 5 exx, 800–1000 m, 07-VIII-1992; 4 exx, 950–1450 m, 03-X-1992.

Jayapura Reg. (Cyclops Mts). Elevation: 860–1200 m.

Collected by beating foliage in montane forests.

This epithet is based on the Latin participle armatus (armed) and refers to the teeth of the male meso- and metatibia.

Trigonopterus armatus Riedel, sp. n. was coded as “Trigonopterus sp. 7” by Riedel et al. (2010), respectively “Trigonopterus spg” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 11a). Length 2.55 mm. Color ferruginous; dorsal surface of head and pronotum black. Body subovate; in dorsal aspect with distinct constriction between pronotum and elytron; in profile with weak constriction. Rostrum densely punctate-reticulate, without longitudinal furrows or ridges. Eyes large, approximate. Pronotum coarsely punctate-reticulate. Elytra densely striate-punctate; striae deeply impressed; intervals each with dense row of deeply impressed punctures, similar to striae; interspaces costate, subglabrous. Femora edentate. Metafemur subapically with stridulatory patch. Aedeagus (Fig. 11b) apically subangulate, subglabrous; transfer apparatus spiniform, long, subequal to body; ductus ejaculatorius without bulbus. Intraspecific variation. Length 2.52–2.64 mm. Female rostrum dorsally in apical half with punctures usually isolated.

Holotype (MZB): ARC1767 (EMBL # HE616044), WEST NEW GUINEA, Jayawijaya Reg., Poga, S03°47.575', E138°33.155' to S03°47.473', E138°33.163', 2620–2715 m, 15-VII-2010. Paratypes (SMNK, ZSM): 21 exx, ARC1768 (EMBL # HE616045), ARC1769 (EMBL # HE616046), same data as holotype.

Jayawijaya Reg. (Poga). Elevation: ca. 2620–2715 m.

Beaten from foliage of upper montane forests.

This epithet is based on the Latin participle ascendens (climbing up) and refers to its occurrence on higher elevations.

Trigonopterus ascendens Riedel, sp. n. was coded as “Trigonopterus sp. 169” by Tänzler et al. (2012).

Holotype, male (Fig. 12a). Length 4.24 mm. Color black; antenna partly ferruginous. Body ovate; in dorsal aspect and in profile with constriction between pronotum and elytron. Rostrum slender, dorsally with distinct median carina and sublateral ridges; furrows bordering ridges containing each one row of mesad directed white narrow scales; subapically shining, punctate, setose. Eyes large. Pronotum large, subglabrous, with minute punctures, in basal half sides separated by densely punctate edge. Elytra punctate with minute punctures; striae impressed as fine lines; basal margin bisinuate, bordered by row of large punctures continued behind humeri. Profemur large, anteriorly at middle with large tooth. Mesofemur and metafemur dorsally densely squamose with white scales; with anteroventral ridge at middle with inconspicuous tooth. Metafemur with smooth dorsoposterior edge; subapically without stridulatory patch. Aedeagus (Fig. 12b) apically weakly pointed; ductus ejaculatorius near insertion to transfer apparatus swollen, subapically with weak bulbus. Intraspecific variation. Length 3.28–4.16 mm. Female rostrum dorsally largely subglabrous, with submedian rows of minute punctures; in basal 1/5 punctate-rugose, with white recumbent scales.

Holotype (MZB): ARC0444 (EMBL # FN429155), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°31.685', E140°30.430', 1010m, 28-XI-2007, beaten. Paratypes (ARC, NHMB, SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani: 1 ex, ARC0445 (EMBL # FN429156), same data as holotype; 1 ex, ARC0658 (EMBL # FN429305), S02°31.794', E140°30.190', 800–860 m, 21-XI-2007, “Mim2”, beaten; 16 exx, ARC0666 (EMBL # FN429313), ARC0667 (EMBL # FN429314), S02°31.6', E140°30.4', 900–1100 m, 28-XI-2007, beaten; 3 exx, 1100–1600 m, 05-X-1991; 1 ex, 300–1400 m, 10-VIII-1991; 1 ex, 800–1000 m, 07-VIII-1992; 6 exx (1 marked as “ARC0130”), 950–1450 m, 03-X-1992; 2 exx, Lake Sentani, III-1992, 300 m.

Jayapura Reg. (Cyclops Mts). Elevation: 300–1200 m.

Collected by beating foliage in montane forest.

This epithet is based on the Latin noun augur in apposition and refers to the large eyes that help the species to see birds, presumably important predators.

Trigonopterus augur Riedel, sp. n. was coded as “Trigonopterus sp. 21” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spu” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 13a). Length 2.58 mm. Color orange-ferruginous; pronotum and parts of head black. Body fusiform, almost without constriction between pronotum and elytron; in profile dorsally flat, towards apex convex. Rostrum with median costa flat, pair of submedian furrows containing sparse rows of mesad-directed setae. Pronotum subglabrous, punctate with small to minute punctures. Elytra subglabrous, punctation confused with small to minute punctures. Femora with anteroventral ridge terminating with tooth in apical third. Anteroventral ridge of mesofemur high, at middle with subangulate incision. Metafemur with crenulate dorsoposterior edge; subapically with stridulatory patch; posteroventral ridge at middle with knob. Mesotibia with dorsal edge pushed forward, apex slightly curved ventrad. Abdominal ventrite 1 besides metacoxa with brush of erect scales. Aedeagus (Fig. 13b) with body subrectangular; apex with pair of long pointed brushes; transfer-apparatus complex, symmetrical; ductus ejaculatorius with bulbus. Intraspecific variation. Length 2.11–2.96 mm. Female mesofemur with evenly denticulate anteroventral ridge, without incision at middle. Posteroventral ridge of female metafemur simple. Female mesotibia straight. Abdominal ventrite 1 besides metacoxa simple.

Holotype (MZB): ARC0752 (EMBL # HE615435), WEST NEW GUINEA, Jayawijaya Reg., Jiwika, Kurulu, S03°57.043', E138°57.410', 1920–1950 m, “Mim 3”, 26-XI-2007. Paratypes (ARC, SMNK, NHMB, ZSM): WEST NEW GUINEA, Jayawijaya Reg.: 45 exx, ARC0753 (EMBL # HE615436), ARC0754 (EMBL # HE615437), same data as holotype; 34 exx, Jiwika, Kurulu, S03°56.146', E138°57.710', 2245–2290 m, “Mim 4”, 26-XI-2007; 35 exx, Jiwika, Kurulu, S03°56.5', E138°57.1', 1900–2245 m, 26-XI-2007; 45 exx, Jiwika, Kurulu, S03°57.161', E138°57.357', 1875 m, 11-VII-2010, sifted; 135 exx, Jiwika, Kurulu, S03°57.161', E138°57.357' to S03°56.977', E138°57.441', 1875–1990 m, 12-VII-2010; 2 exx, ARC1711 (EMBL # HE615991), ARC1712 (EMBL # HE615992), Jiwika, Kurulu, S03°57.161', E138°57.357' to S03°56.977', E138°57.441', 1875–1990 m, 12-VII-2010; 73 exx (1 marked ARC0034), Jiwika, Kurulu, 1800–2300 m, 31-V-1998; 20 exx, Jiwika, Kurulu, 1700–2300 m, 02-IX-1991; 9 exx, Jiwika, Kurulu, 1700–2300 m, 06-IX-1991; 15 exx, Jiwika, 1750–2100 m, 05-VII-1994; 18 exx, Jiwika, Kurulu, trail to Wandanku, 2240–2420 m, 28-IX-1996; 17 exx, Jiwika, Kurulu, trail to Wandanku, 1900–2150 m, 28-29-IX-1996; 19 exx, Jiwika, Kurulu, 1900–2300 m, 29-IX-1992; 6 exx, Jiwika, 1700–2100 m, 05-XII-1995; 14 exx, Jiwika, 1700–2000 m, 11-IX-1991; 4 exx, Jiwika, trail to Wandanku, 2240–2420 m, 28-IX-1996; 1 ex, Jiwika, 2300 m, 1992; 69 exx, Baliem-vall., ca. 1700 m, III-1992.

Jayawijaya Reg. (Jiwika). Elevation: 1875–2240 m.

Beaten from foliage of montane forests.

This epithet is based on the name of the Balim-river area which is close to the type locality.

Trigonopterus balimensis Riedel, sp. n. was coded as “Trigonopterus sp. 172” by Tänzler et al. (2012). At the time, it was lumped with a closely related but distinct species from Poga and Lake Habbema.

Holotype, male (Fig. 14a). Length 2.42 mm. Color black; base of elytra and legs ferruginous; antenna light ferruginous. Body subovate; in dorsal aspect with marked constriction between pronotum and elytron; in profile almost evenly convex. Rostrum dorsally scabrous, basally with indistinct median ridge; epistome forming angulate ridge. Pronotum moderately densely punctate. Elytra with striae marked by small punctures; interval 4 basally with cluster of few subovate, cream-colored, recumbent scales; interval 7 subapically forming indistinct ridge. Meso- and metafemur with anteroventral ridge weakly dentate. Meso- and metatibia in basal half widened, subapically narrowed; dorsal edge basally granulate; uncus large, peg-shaped. Metafemur with denticulate dorsoposterior edge, subapically without stridulatory patch. Aedeagus (Fig. 14b) with sides of body in apical third converging, apex rounded; orifice retracted; endophallus denticulate; transfer apparatus symmetrical; ductus ejaculatorius without bulbus. Intraspecific variation. Length 1.98–2.48 mm. Female rostrum dorsally subglabrous in apical half, with small punctures, epistome simple.

Holotype (MZB): ARC0523 (EMBL # FN429229), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°31.912', E140°30.416', 785 m, 02-XII-2007, sifted. Paratypes (SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts: 4 exx, ARC0524 (EMBL # FN429230), same data as holotype; 16 exx, ARC0532 (EMBL # FN429238), Doyo, S02°32.478', E140°28.835', 365 m, 27-XI-2007, sifted; 2 exx, Sentani, S02°32.031', E140°30.412', 710 m, 02-XII-2007, sifted; 5 exx (1 marked “ARC0105”), Sentani, 600 m, 22-XII-2004, sifted; 1 ex, Sentani, 700 m, 22-XII-2004, sifted; 17 exx, Sentani, S02°32.221', E140°30.526', 575 m, 19-XI-2007, sifted; 1 ex, Sentani, S02°32.166' E140°30.512', 620 m, 19-XI-2007, sifted; 3 exx, Sentani, S02°32.291', E140°30.505', 515 m, 19-XI-2007, sifted.

Jayapura Reg. (Cyclops Mts). Elevation: 365–785 m.

Sifted from leaf litter in primary forest.

This epithet is based on the Latin adjective basalis (characterized by the base) and refers to the elytral base differing in color and vestiture from the remainder.

Trigonopterus basalis Riedel, sp. n. was coded as “Trigonopterus sp. 42” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spap” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 15a). Length 3.34 mm. Color black, legs deep ferruginous, antenna lighter ferruginous. Body ovate; almost without constriction between pronotum and elytron; in profile evenly convex. Rostrum dorsally tricarinate, with distinct median and pair of lateral carinae. Pronotum densely punctate. Elytra densely punctate; strial punctures slightly larger than minute punctures on intervals; striae impressed as fine lines; lateral stria behind humeri simple, not deepened. Femora edentate. Metafemur with denticulate dorsoposterior edge; subapically with stridulatory patch. Mesotibia simple, in basal half dorsal contour denticulate, but without distinct angulation. Metatibia with minute premucro; without suprauncal projection. Aedeagus (Fig. 15b) apically subangulate; dorsum sublaterally sparsely setose; transfer apparatus dentiform, short; ductus ejaculatorius with bulbus. Intraspecific variation. Length 3.31–3.34 mm. No female specimen available.

Holotype (MZB): ARC1794 (EMBL # HE616071), WEST NEW GUINEA, Jayawijaya Reg., Bokondini, S03°41.787', E138°40.229' to S03°41.778', E138°40.129', 1705–1710 m, 17-VII-2010. Paratype (SMNK): 3 exx, ARC1795 (EMBL # HE616072), ARC2347 (EMBL # HF548203), ARC2348 (EMBL # HF548204) same data as holotype.

Jayawijaya Reg. (Bokondini). Elevation: ca. 1705–1710 m.

Beaten from foliage of montane forest.

This epithet is based on the Latin adjective conformis (like, similar) and refers to the similarity of this species, both to some closely related sibling species, and to others of only superficial resemblance.

Trigonopterus conformis Riedel, sp. n. was coded as “Trigonopterus sp. 86” by Tänzler et al. (2012).

Holotype, male (Fig. 16a). Length 2.55 mm. Color dark brown; antenna, tarsi, and elytra ferruginous. Body dull, microreticulate; subovate; in dorsal aspect with marked constriction between pronotum and elytron; in profile with shallow constriction. Rostrum sparsely punctate, with sublateral pair of furrows; epistome simple. Eyes large, divided into dorsal and ventral portions by marked incision of posterior margin. Pronotum with distinct subapical constriction; densely punctate with deep punctures; each puncture with one elongate-ovate, ochre scale. Elytra with striae deeply incised; intervals costate, each with one row of narrow scales. Metafemur subapically without stridulatory patch. Tibial apex with stout uncus and minute premucro. Onychium ca. 2.3 × longer than tarsomere 3. Aedeagus (Fig. 16b) with sides of apical half converging to apex, in profile markedly curved; transfer apparatus tubuliform; ductus ejaculatorius without bulbus. Intraspecific variation. Length 2.55–2.63 mm. Color with ferruginous elytra or entirely brown.

Holotype (MZB): ARC1731 (EMBL # HE616008), WEST NEW GUINEA, Jayawijaya Reg., Poga, S03°47.406', E138°35.507', 2410 m, 14-VII-2010, sifted. Paratypes (SMNK): WEST NEW GUINEA, Jayawijaya Reg., Poga: 1 ex, ARC1724 (EMBL # HE616004), S03°48.382', E138°34.780'; 2330 m, 13-VII-2010, sifted.

Jayawijaya Reg. (Poga). Elevation: 2330–2410 m.

Sifted from leaf litter in montane forest.

This epithet is based on the Latin participle constrictus (constricted) and refers both to the constriction of the eye and the body between pronotum and elytron.

Trigonopterus constrictus Riedel, sp. n. was coded as “Trigonopterus sp. 54” by Tänzler et al. (2012).

Holotype, male (Fig. 17a). Length 2.12 mm. Color black; antenna light ferruginous; legs deep ferruginous. Body subovate, with shallow constriction between pronotum and elytron; in profile evenly convex. Rostrum in basal half medially carinate, with pair of sublateral furrows posteriorly converging on forehead; weakly punctate, sparsely setose; epistome with transverse, angulate ridge. Pronotum with subapical constriction dorsally distinct, laterally indistinct; disk with longitudinal impressions, sparsely punctate, sparsely setose. Elytra with striae deeply incised, towards sides with deep interspersed punctures; intervals costate-carinate, subglabrous, with sparsely setose with minute recumbent setae; apex rounded. Femora edentate. Metafemur with simple dorsoposterior edge, in apical third with transverse row of small suberect setae, subapically with stridulatory patch. Abdominal ventrite 5 basally with pair of teeth. Aedeagus (Fig. 17b) apically subangulate, glabrous; with complex, symmetrical transfer apparatus; ductus ejaculatorius without bulbus. Intraspecific variation. Length 2.06–2.12 mm. Female rostrum in basal half with median carina less distinct.

Holotype (MZB): ARC0767 (EMBL # HE615450), WEST NEW GUINEA, Manokwari, Mt. Meja, S00°51.497', E134°04.949', 220 m, 05-XII-2007, sifted. Paratypes (SMNK, ZSM): WEST NEW GUINEA, Manokwari, Mt. Meja: 2 exx, ARC766 (EMBL # HE615449), ARC0768 (EMBL # HE615451), same data as holotype; 6 exx, S00°51.400', E134°04.918', 225 m, 06-XII-2007, sifted; 1 Ex, 200 m, 30-XII-2004.

Manokwari Reg. (Mt. Meja). Elevation: 220–225 m.

Sifted from leaf litter in lowland forest.

This epithet is based on the Latin adjective costatus (ribbed) and refers to the elytral sculpture.

Trigonopterus costatus Riedel, sp. n. was coded as “Trigonopterus sp. 202” by Tänzler et al. (2012).

Holotype, male (Fig. 18a). Length 2.80 mm. Color black; tarsi and antenna ferruginous. Body elongate; in dorsal aspect and in profile with distinct constriction between pronotum and elytron. Rostrum with median costa and pair of submedian costae, furrows between with sparse rows of suberect scales; epistome flat. Pronotum with marked subapical constriction, anteriorly densely punctate; disk sparsely punctate, deeply sculptured, with median ridge and pair of broad submedian ridges; behind constriction anteriorly with pair of lateral angular protrusions. Elytra subglabrous except basally striae 1–3 with each one fovea; remaining striae indistinct, marked by minute punctures; apex rounded. Femora edentate. Metafemur subapically with stridulatory patch. Tarsomere 3 small, hardly larger than preceding, onychium ca. 2.1 × longer than tarsomere 3. Aedeagus (Fig. 18b) apically angulate, sparsely setose; transfer-apparatus flagelliform; ductus ejaculatorius with indistinct bulbus.

Holotype (SMNK): ARC1094 (EMBL # HE615723), PAPUA NEW GUINEA, Simbu Prov., Karimui Dist., Haia, S06°43.948', E144°59.856', 915 m, 26-IX-2009, sifted.

Simbu Prov. (Haia). Elevation: 915 m.

Sifted from leaf litter in primary forest.

This epithet is a combination of the Latin nouns costa (rib, ridge) and collum (neck; pronotum) refers to the sculpture of its pronotum.

Trigonopterus costicollis Riedel, sp. n. was coded as “Trigonopterus sp. 166” by Tänzler et al. (2012).

Holotype, male (Fig. 19a). Length 2.39 mm. Color orange-ferruginous; pronotum and parts of head black. Body subrhomboid, almost without constriction between pronotum and elytron; in profile dorsally flat, towards apex convex. Rostrum without median ridge, basally swollen and densely punctate, sparsely setose. Antenna with funicle swollen, continuous with club. Pronotum subglabrous, sparsely punctate with small to minute punctures. Elytra subglabrous, punctation confused with small to minute punctures. Profemur with anteroventral ridge terminating with tooth in apical third. Mesofemur with anteroventral ridge irregularly serrate; posteroventral ridge in apical half, terminating with rectangular protrusion. Metafemur laterally flattened; dorsally sparsely squamose with silvery scales; anteroventral ridge distinct; dorsoposterior edge simple; subapically with stridulatory patch. Uncus of metatibia small. Aedeagus (Fig. 19b). Body with sides converging; apex medially weakly extended, sublaterally with pair of sparse setose brushes; transfer-apparatus complex, symmetrical; ductus ejaculatorius with bulbus. Intraspecific variation. Length 2.22–2.85 mm. Female rostrum dorsally subglabrous with pair of submedian row of small punctures. Female antennal funicle less swollen. Female mesofemur along posteroventral edge simple, without rectangular protrusion.

Holotype (MZB): ARC0756 (EMBL # HE615439), WEST NEW GUINEA, Jayawijaya Reg., Jiwika, Kurulu, S03°56.146', E138°57.710', 2245–2290 m, “Mim 4”, 26-XI-2007. Paratypes (ARC, SMNK, ZSM): WEST NEW GUINEA, Jayawijaya Reg.: 3 exx, ARC0757 (EMBL # HE615440), ARC0758 (EMBL # HE615441), same data as holotype; 72 exx, ARC1785 (EMBL # HE616062), ARC1786 (EMBL # HE616063), Bokondini, S03°40.345', E138°42.386' to S03°40.255', E138°42.189', 1655–1700 m, 18-VII-2010; 22 exx, ARC1788 (EMBL # HE616065), ARC1789 (EMBL # HE616066), Bokondini, S03°41.787', E138°40.229' to S03°41.778', E138°40.129', 1705–1710 m, 17-VII-2010; 2 exx (1 marked ARC00611), Ilugwa, Melanggama, 1900–2200 m, 09-12-IX-1990; 2 exx, Ilugwa, Melanggama, trail to Pass Valley, 2100–2300 m, 09-10-IX-1990; 3 exx, Jiwika, Kurulu, ca. 1700–2300 m, 06-IX-1991; 1 ex, Jiwika, 1800–2300 m, 31-V-1998.

Distribution. Jayawijaya Reg. (Jiwika, Ilugwa, Bokondini). Elevation: 1700–2245 m.

Biology. Beaten from foliage of montane forests.

Etymology. This epithet is based on a combination of the Latin adjective crassus (thick) and cornu (horn, antenna) and refers to the thickened antennal funicle.

Notes. Trigonopterus crassicornis Riedel, sp. n. was coded as “Trigonopterus sp. 171” by Tänzler et al. (2012).

Holotype, male (Fig. 20a). Length 3.00 mm. Color black. Body slender, subrhomboid; without constriction between pronotum and elytron; in profile evenly convex. Rostrum in apical third smooth; in basal 2/3 with broad median costa and pair of sublateral ridges; furrows containing sparse row of setae. Eyes with dorsal margin carinate. Head bordering eye with elongate impression. Pronotum subglabrous, sparsely punctate with minute punctures. Elytra subglabrous, punctation confused with minute punctures; striae hardly visible, impressed as very fine lines; lateral stria behind humeri with row of ca. 5 deep punctures. Femora subglabrous, including dorsum of metafemur without scales. Mesofemur on posterior surface with longitudinal ridge. Metafemur on posterior surface with two longitudinal furrows; dorsally with smooth ridge; subapically without stridulatory patch. Meso- and metatibia subapically simple, with uncus, without premucro. Aedeagus (Fig. 20b). Apex symmetrical, with median acute extension; body dorsally with two rows of sparse short setae; transfer apparatus small, dentiform; endophallus without distinct sclerites; ductus ejaculatorius without bulbus. Intraspecific variation. Length 1.95–3.00 mm.

Holotype (SMNK): ARC1839 (EMBL # HE616116), PAPUA NEW GUINEA, Eastern Highlands Prov., Aiyura, S06°21.033', E145°54.597', 2169 m, 06-II-2010. Paratypes (NAIC): PAPUA NEW GUINEA, Eastern Highlands Prov.: 1 ex, ARC1849 (EMBL # HE616126), Okapa, Kimiagomo village, Hamegoya, S06°25.727', E145°35.455', S06°25.117', E145°35.225', 1891–2131 m, 18-III-2010.

Eastern Highlands Prov. (Aiyura, Okapa). Elevation: 2131–2169 m.

Beaten from foliage of montane forests.

This epithet is based on a combination of the Latin nouns cuneus (wedge) and penna (wing, elytron) and refers to the shape of elytra.

Trigonopterus cuneipennis Riedel, sp. n. was coded as “Trigonopterus sp. 96” by Tänzler et al. (2012).

Holotype, male (Fig. 21a). Length 3.27 mm. Color black; legs deep ferruginous, antenna light ferruginous. Body ovate; with weak constriction between pronotum and elytron; in profile evenly convex. Rostrum dorsally densely rugose-punctate, in basal half with brown erect scales, in apical half sparsely setose. Eyes large. Pronotum densely punctate with subtriangular, setiferous punctures. Elytral striae distinct, marked by regular rows of small punctures; intervals with row of minute punctures; laterally behind humeri with ridge bordered by row of deep punctures of stria 8. Legs squamose with inconspicuous brownish scales. Femora with anteroventral ridge terminating in apical third. Metafemur with weakly denticulate dorsoposterior edge; subapically with stridulatory patch. Abdominal ventrite 5 at middle with shallow, subquadrate impression. Aedeagus (Fig. 21b) with apex subangulate; transfer-apparatus flagelliform, longer than body of aedeagus; ductus ejaculatorius without bulbus. Intraspecific variation. Length 2.67–3.27 mm. Female rostrum in apical half with relatively small, sparse punctures. Abdominal ventrite 5 simple.

Holotype (MZB): ARC478 (EMBL # FN429185), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°31.2', E140°30.5', 1420–1520 m, 30.XI.2007, beaten. Paratypes (ARC, SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani: 5 exx, same data as holotype; 4 exx, ARC0423 (EMBL # FN429134), ARC0483 (EMBL # FN429190), ARC0674 (EMBL # FN429319), ARC0675 (EMBL # FN429320), S02°31.3', E140°30.5', 1200–1420 m, 30.XI.2007, beaten; 2 exx (1 marked as “ARC0409”), 1100–1600 m, 05-X-1991.

Jayapura Reg. (Cyclops Mts). Elevation: ca. 1420 m.

Collected by beating foliage in montane forests.

This epithet is based on the type locality, the Cyclops Mountains.

Trigonopterus cyclopensis Riedel, sp. n. was coded as “Trigonopterus sp. 5” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spe” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 22a). Length 1.65 mm. Color black; antenna and tarsi ferruginous. Body subovate; in dorsal aspect without constriction between pronotum and elytron; in profile with distinct constriction. Rostrum at base with median ridge and pair of submedian ridges; at middle with anteriorly hollowed protuberance dorsally bearing pair of denticles; between protuberance and epistome relatively flat, with sparse erect scales; epistome at middle with dorsoposteriad directed horn. Pronotum punctate-rugose, each puncture containing one narrow transparent scale; medially with indistinct ridge. Elytra with striae deeply impressed; punctures large, each containing one downcurved seta; intervals weakly costate, subglabrous. Metafemur dorsoposteriorly simple, subapically without stridulatory patch. Aedeagus (Fig. 22b). Body widening to shortly before apex; medially weakly extended; endophallus denticulate; transfer apparatus spiniform, curved; ductus ejaculatorius with weak bulbus. Intraspecific variation. Length 1.65–1.92 mm. Female rostrum dorsally even, without teeth or cavities, medially subglabrous, sublaterally punctate.

Holotype (MZB): ARC0538 (EMBL # FN429244), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, S02°31.594', E140°30.407', 1065 m, 21-XI-2007, sifted. Paratypes (ARC, SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani: 2 exx, ARC0544 (EMBL # FN429250), S02°31.683', E140°30.281', 960 m, 21-XI-2007, sifted; 6 exx, ARC0545 (EMBL # FN429251), Sentani, S02°31.776', E140°30.215', 945 m, 21-XI-2007, sifted; 1 ex, Sentani, S02°31.603', E140°30.434', 1095 m, 28-XI-2007, sifted; 2 exx, Sentani, 950–1450 m, 03-X-1992; 6 exx (1 marked ARC0026), Sentani, 1000 m, 23-XII-2004, sifted; 1 ex, Sentani, 1100 m, 23-XII-2004, sifted; 3 exx, ARC1690 (EMBL # HE615977), ARC1691 (EMBL # HE615978), Angkasa indah, S02°30.346', E140°42.087', 490 m, 28-VI-2010, sifted.

Jayapura Reg. (Cyclops Mts). Elevation: 490–1095 m.

Sifted from leaf litter in montane forest.

This epithet is based on a combination of the Latin nouns dens (tooth) and rostrum (snout) and refers to the dorsal protrusions of the rostrum.

Trigonopterus dentirostris Riedel, sp. n. was coded as “Trigonopterus sp. 49” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spaw” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 23a). Length 2.20 mm. Color black; legs ferruginous, antenna light ferruginous. Body subglobose; in dorsal aspect with weak constriction between pronotum and elytron; in profile almost evenly convex. Rostrum with pair of sublateral furrows and pair of submedian row of punctures, each containing row of mesally directed setae; epistome simple. Pronotum with large disk and distinct lateral edges; moderately densely punctate with small setiferous punctures. Elytra converging to subangulate apex; striae distinct, marked by small punctures; intervals subglabrous, with row of minute punctures; some punctures with one minute recumbent seta. Metafemur with weakly denticulate dorsoposterior edge; subapically with stridulatory patch. Abdominal ventrites 1–2 deeply excavated. Aedeagus (Fig. 23b) with body flattened, sides subparallel, apex subangulate; apodemes ca. 3 X as long as body; transfer-apparatus spiniform; ductus ejaculatorius, without bulbus. Intraspecific variation. Length 1.68–2.43 mm. Female rostrum more slender than in males, dorsally punctures smaller.

Holotype (SMNK): ARC1175 (EMBL # HE615803), PAPUA NEW GUINEA, Morobe Prov., Huon peninsula, Mindik, S06°27.311', E147°24.073', 1570 m, 10-X-2009. Paratypes (ARC, NAIC, SMNK, ZSM): Morobe Prov., Huon peninsula, Mindik: 5 exx, same data as holotype; 12 exx, ARC1179 (EMBL # HE615807), ARC1180 (EMBL # HE615808), S06°27.221', E147°24.185', 1670 m, 10-X-2009; 5 exx, 1450 m, 26-IV-1998; 1 ex, 1400–1550 m, 27-IV-1998.

Morobe Prov. (Mindik). Elevation: 1450–1670 m.

Sifted from leaf litter in montane forest.

This epithet is based on the Latin adjective discoidalis (shaped like a disk) and refers to the species´ outline when viewed from above.

Trigonopterus discoidalis Riedel, sp. n. was coded as “Trigonopterus sp. 88” by Tänzler et al. (2012).

Holotype, male (Fig. 24a). Length 3.31 mm. Color black; tarsi and tibiae dark ferruginous; antenna light ferruginous. Body subrhomboid; in dorsal aspect with marked constriction between pronotum and elytron; in profile with moderate constriction. Rostrum medially punctate, with pair of sublateral furrows converging posteriorly on forehead; epistome forming indistinct, angulate ridge. Pronotum with marked subapical constriction, sides subparallel, behind subapical constriction projecting with marked angular protrusions; center of disk densely punctate, laterally with sparse punctures. Elytra of subtriangular shape; median suture carinate; striae moderately impressed; intervals weakly punctate; surface weakly microreticulate, with sparse clusters of white recumbent scales; interval 7 subapically forming indistinct ridge. Femora edentate. Meso- and metatibia in basal half markedly widened, dorsal edge curved, denticulate. Metafemur with denticulate dorsoposterior edge, subapically without stridulatory patch. Metathoracic and abdominal venter forming common concavity, subglabrous. Aedeagus (Fig. 24b) with body parallel-sided, markedly curved ventrad; apex medially extended; transfer apparatus spiniform; ductus ejaculatorius without bulbus. Intraspecific variation. Length 3.22–3.69 mm. Integument relatively smooth and with scattered white scales in specimens from Sentani, dull-coriaceous and almost nude in specimens from Angkasa indah. Aedeagus with median tip acute in specimens from Sentani, median tip more rounded in specimens from Angkasa indah.

Holotype (MZB): ARC0512 (EMBL # FN429218), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°32.031', E140°30.412', 710 m, 02-XII-2007, sifted. Paratypes (SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts: 1 ex, ARC0534 (EMBL # FN429240), Sentani, S02°31.516', E140°30.436', 1150 m, 21-XI-2007, sifted; 2 exx, ARC0549 (EMBL # FN429255), Sentani, S02°31.776', E140°30.215', 945 m, 21-XI-2007, sifted; 2 exx, Sentani, S02°31.603', E140°30.434', 1095 m, 28-XI-07, sifted; 1 ex, Sentani, S02°31.912', E140°30.416', 785 m, 02-XII-2007, sifted; 1 ex, ARC0092, Sentani, 1000 m, 23-XII-2004, sifted; 1 ex, S02°31.425', E140°30.474', 1265 m, 30-XI-2007, sifted; 2 exx, ARC1684 (EMBL # HE615971), ARC1685 (EMBL # HE615972), Angkasa indah, S02°30.346', E140°42.087', 490 m, 28-VI-2010, sifted.

Jayapura Reg. (Cyclops Mts). Elevation: 490–1265 m.

Sifted from leaf litter in primary forest.

This epithet is based on the name of the dromedary camel (Camelus dromedaries L.) and refers to the body shape.

Trigonopterus dromedarius Riedel, sp. n. was coded as “Trigonopterus sp. 41” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spao” in the EMBL/GenBank/DDBJ databases. Specimens from Angkasa indah and Sentani exhibit minor morphological differences (e.g. tip of aedeagus; surface of integument) and a high cox1 p-distance of 7.79%. These allopatric populations may be regarded “subspecies”.

Holotype, male (Fig. 25a). Length 3.69 mm. Color black; base of antennal scape ferruginous. Body subovate; with weak constriction between pronotum and elytron; in profile evenly convex. Rostrum punctate-rugose, with pair of sublateral furrows. Eyes dorsally bordered by furrow. Pronotum subglabrous, densely punctate with minute punctures; disk subquadrate, rounded towards sides. Elytra tapering apicad; striae indistinct, marked by rows of small punctures; intervals flat, with confused minute punctures; elytral base with row of foveae continued laterally behind humeri bordering ridge. Femora subglabrous, edentate. Metafemur dorsally partly covered with silvery scales; with weakly denticulate dorsoposterior edge; subapically with stridulatory patch. Mesotibia and metatibia with inconspicuous straight premucro below tarsal insertion. Aedeagus (Fig. 25b) with sides of body basally converging, in apical 1/3 with weak constriction; apex with median, broad angular extension, sparsely setose; transfer apparatus spiniform; ductus ejaculatorius without bulbus. Intraspecific variation. Length 3.38–4.64 mm. Female rostrum slender, dorsally subglabrous, sparsely punctate with small punctures. Pronotum of males anteriorly subangulate, in females sides curving more evenly towards apex. Elytra of females shorter and dorsally rather convex; elytra of males apically slightly extended and dorsally slightly flattened.

Holotype (MZB): ARC1790 (EMBL # HE616067), WEST NEW GUINEA, Jayawijaya Reg., Bokondini, S03°41.787', E138°40.229' to S03°41.778', E138°40.129', 1705–1710 m, 17-VII-2010. Paratypes (ARC, SMNK, ZSM): 91 exx, ARC1791 (EMBL # HE616068), ARC1792 (EMBL # HE616069), same data as holotype; 6 exx, ARC1793 (EMBL # HE616070), Bokondini, S03°40.345', E138°42.386' to S03°40.255', E138°42.189', 1655–1700 m, 18-VII-2010; 85 exx, Angguruk, 1600–1700 m, 21-IX-1991; 1 ex, Angguruk, 1200–1500 m, 23-IX-1992; 2 exx, Angguruk – Tanggeam, 1500–1800 m, 28-29-IX-1991.

Jayawijaya Reg. (Bokondini, Angguruk). Elevation: 1700–1710 m.

Beaten from foliage of montane forests.

This epithet is based on the Latin adjective durus (hard, tough) and refers to the physical properties of the species. The name would be equally fitting many other species of this genus and should not be seen as diagnostic.

Trigonopterus durus Riedel, sp. n. was coded as “Trigonopterus sp. 102” by Tänzler et al. (2012).

Holotype, male (Fig. 26a). Length 1.41 mm. Color black; antenna light ferruginous, club dark; legs and head dark ferruginous. Body subglobose; with distinct constriction between pronotum and elytron; in profile almost evenly rounded, with weak constriction between pronotum and elytron. Rostrum basally with indistinct, median ridge and pair of submedian ridges, sparsely setose; apical 1/3 subglabrous; epistome simple. Eyes small. Pronotum with weak subapical constriction; disk densely, coarsely punctate; each puncture with short, yellowish scale. Elytra with striae deeply incised; with suberect, yellowish scales; intervals subglabrous, markedly costate. Femora subapically ventrally constricted. Metafemur subapically with stridulatory patch. Protibia with long, hook-shaped uncus. Aedeagus (Fig. 26b) with apex rounded; transfer apparatus symmetrical, spiniform; ductus ejaculatorius with indistinct bulbus. Intraspecific variation. Length 1.28–1.54 mm. Female rostrum basally without ridges; apical half subglabrous. Suberect scales may be partly abraded.

Holotype (MZB): ARC0781 (EMBL # HE615464), WEST NEW GUINEA, Manokwari, Arfak Mts, S01°04.087', E133°54.268', 1520 m, 08-XII-2007, sifted. Paratypes (SMNK, ZSM): 21 exx, ARC0782 (EMBL # HE615465), ARC0783 (EMBL # HE615466), same data as holotype; Mokwam, Siyoubrig, S01°06.107', E133°54.888', 1530 m, 10–XII-2007.

Manokwari Reg. (Arfak Mts). Elevation: 1520 m.

Sifted from leaf litter in montane forest dominated by Nothofagus.

This epithet is based on the Latin noun echinus (hedgehog) in apposition and refers to the species´ general habitus and its suberect scales resembling spines.

Trigonopterus echinus Riedel, sp. n. was coded as “Trigonopterus sp. 230” by Tänzler et al. (2012).

Holotype, male (Fig. 27a). Length 1.48 mm. Color black; antenna light ferruginous; legs deep ferruginous. Body subglobose, in dorsal aspect with marked constriction between pronotum and elytron; dorsally flattened; anteriorly profile almost straight, convex at apex. Rostrum coarsely rugose-punctate; epistome forming distinct, transverse ridge. Pronotum coarsely punctate-reticulate. Elytra with striae deeply incised, with sparse rows of setae; intervals costate, subglabrous, with few minute punctures; interval 7 subapically forming ridge, projecting dentiform; apex subangulate. Legs. Anteroventral ridge of meso- and metafemur ending in apical third, forming indistinct tooth. Metafemur dorsoposteriorly simple, subapically with stridulatory patch. Aedeagus (Fig. 27b) with apex subtruncate, slightly angulate; transfer apparatus spiniform, shorter than body of aedeagus; ductus ejaculatorius without bulbus. Intraspecific variation. Length 1.32–2.02 mm. Body of females subovate. Female rostrum dorsally subglabrous, coarsely punctate; epistome simple. Female elytral apex laterally not dentiform.

Holotype (MZB): ARC0737 (EMBL # HE615420), WEST NEW GUINEA, Jayawijaya Reg., Jiwika, Kurulu, S03°57.161', E138°57.357', 1875 m, 24-XI-2007, sifted. Paratypes (ARC, SMNK, ZSM): WEST NEW GUINEA, Jayawijaya Reg., Jiwika, Kurulu: 27 exx, ARC0738 (EMBL # HE615421), ARC0739 (EMBL # HE615422), ARC0740 (EMBL # HE615423), ARC0741 (EMBL # HE615424), ARC0742 (EMBL # HE615425), same data as holotype; 64 exx, S03°57.161', E138°57.357', 1875 m, 11-VII-2010, sifted; 8 exx (1 marked ARC0074), 1900–2050 m, 24-X.1993, sifted;12 exx, 1900–2000 m, 23-IX-1992, sifted; 3 exx, 1900–2050 m, 24-IX-1992, sifted; 2 exx, ca. 1700–2300 m, 02-IX-1991, sifted; 7 exx, 1700–2000 m, 11-IX-1991, sifted.

Jayawijaya Reg. (Jiwika). Elevation: 1875–1900 m.

Sifted from leaf litter in montane forest.

This epithet is the latinized form of the Greek word edaphos (soil, ground) and treated as an adjective. It refers to the habit of this litter-dwelling species.

Trigonopterus edaphus Riedel, sp. n. was coded as “Trigonopterus sp. 224” by Tänzler et al. (2012).

Holotype, male (Fig. 28a). Length 3.66 mm. Color black; tarsi and antenna ferruginous. Body ovate; with weak constriction between pronotum and elytron; in profile evenly convex. Rostrum dorsally with distinct median carina and sparse rows of upcurved scales; submedian ridges indistinct, irregular. Pronotum densely punctate except along impunctate midline. Elytra punctate with minute punctures, especially on intervals; strial punctures slightly larger; striae impressed as fine lines; lateral stria behind humeri simple, not deepened. Femora edentate. Metafemur with denticulate dorsoposterior edge; subapically with fine stridulatory patch. Metatibia apically with uncus, without premucro. Abdominal ventrite 5 flat, densely setose with suberect setae. Aedeagus (Fig. 28b) slightly asymmetrical; with distinct, asymmetrical transfer-apparatus; ductus ejaculatorius with bulbus. Intraspecific variation. Length 3.38–3.72 mm. Female rostrum dorsally in basal half with median and pair of submedian ridges, apical half punctate. Female abdominal ventrite 5 flat, sparsely setose with recumbent setae.

Holotype (MZB): ARC426 (EMBL # FN429137), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°31.2', E140°30.5', 1420–1520 m, 30-XI-2007, beaten. Paratypes (ARC, SMNK, ZSM): WEST NEW GUI-NEA, Jayapura Reg., Cyclops Mts, Sentani: 9 exx, ARC0425 (EMBL # FN429136), ARC0427 (EMBL # FN429138), same data as holotype; 1 ex (marked as “ARC0411”), 1200–1400 m, 9-VIII-1992; 1 ex, 1100–1600 m, 5.X.1991.

Jayapura Reg. (Cyclops Mts). Elevation: 1400–1420 m.

Collected by beating foliage in montane forests.

This epithet is the latinized form of the Greek noun eremites (hermit) and refers to the species´ restricted occurrence in the montane forests of the Cyclops Mountains.

Trigonopterus eremitus Riedel, sp. n. was coded as “Trigonopterus sp. 6” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spf” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 29a). Length 1.70 mm. Color of antenna, tarsi, tibiae, and rostrum ferruginous; head and pronotum black; elytra black, with ferruginous patches near base of intervals 2–4, continued to apex with irregular patches on intervals 1–4 to apex. Body subovate; in dorsal aspect and in profile with distinct constriction between pronotum and elytron. Eyes large. Rostrum punctate, without distinct longitudinal ridges; laterally with rows of cream-colored scales. Pronotum densely punctate-reticulate; with few scattered cream-colored scales; each puncture containing one brownish seta. Elytra with striae deeply impressed, intervals costate, with scattered white scales. Femora ventrally edentate, subapically constricted. Tibial uncus simple, curved. Metafemur subapi- cally with stridulatory patch. Aedeagus (Fig. 29b) with sides of body sinuate, converging; apex extended, pointed, curved ventrad; transfer apparatus compact, symmetrical; ductus ejaculatorius basally swollen, without bulbus. Intraspecific variation. Length 1.63–1.86 mm. Ferruginous color of elytra more or less extensive. Female rostrum dorsally subglabrous, laterally punctate. Body of males rather slender, females slightly stouter.

Material examined. Holotype (MZB): ARC0784 (EMBL # HE615467), WEST NEW GUINEA, Manokwari Reg., Manokwari, Arfak Mts, S01°04.087', E133°54.268', 1520 m, 08-XII-2007. Paratypes (SMNK, ZSM): WEST NEW GUINEA, Manokwari Reg., Manokwari, Arfak Mts: 6 exx, same data as holotype; 4 exx, ARC0775 (EMBL # HE615458), ARC0776 (EMBL # HE615459), S01°03.723', E133°54.145', 1385 m, 08-XII-2007; 2 exx, Mokwam, Siyoubrig, S01°06.107', E133°54.888', 1530 m, 10-XII-2007.

Manokwari Reg. (Arfak Mts). Elevation: 1385–1530 m.

Sifted from leaf litter in montane forest.

This epithet is based on the Greek euops (well-sighted) and refers to the species´ relatively large eyes, at least by comparison with other edaphic species.

Trigonopterus euops Riedel, sp. n. was coded as “Trigonopterus sp. 53” by Tänzler et al. (2012).

Holotype, male (Fig. 30a). Length 2.73 mm. Color black; elytra orange-red, apically changing to black. Body with distinct constriction between pronotum and elytron; in profile almost evenly convex. Rostrum dorsally relatively flat, sparsely punctate, dorsolaterally with pair of furrows continuing along eye, furrows containing row of mesad directed setae. Pronotum densely punctate with punctures of subtriangular shape, interspaces larger than puncture´s width. Elytra with striae distinct, dorsally punctures small, along base and laterally punctures large; interspaces subglabrous. Femora edentate. Metafemur with indistinct, simple dorsoposterior edge; subapically with stridulatory patch. Thoracic and abdominal venter partly with dense erect setae. Aedeagus (Fig. 30b) apically subangulate, with pair of sublateral setose brushes; transfer apparatus flagelliform, longer than body; ductus ejaculatorius without bulbus. Intraspecific variation. Length 2.73–2.85 mm. Female venter subglabrous, sparsely setose with short recumbent setae.

Holotype (MZB): ARC0477 (EMBL # FN429184), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°31.2', E140°30.5', 1420–1520 m, 30-XI-2007, beaten. Paratypes (SMNK, ZSM): WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani: 9 exx, ARC0475 (EMBL # FN429182), ARC0476 (EMBL # FN429183), same data as holotype; 1 ex, S02°31.3', E140°30.5', 1200–1420 m, 30-XI-2007; 1 ex, S02°31.182', E140°30.542', 1510 m, 30-XI-2007, sifted.

Jayapura Reg. (Cyclops Mts). Elevation: 1420–1510 m.

Collected by beating foliage in montane crippled forests.

This epithet is based on the Latin adjective ferrugineus (rusty, red-brown) and refers to the species´ elytra.

Trigonopterus ferrugineus Riedel, sp. n. was coded as “Trigonopterus sp. 29” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spac” in the EMBL/GenBank/DDBJ databases.

Holotype, male (Fig. 31a). Length 2.63 mm. Color black, legs and antenna ferruginous. Body subrhomboid; almost without constriction between pronotum and elytron; in profile dorsally flat, towards apex convex. Rostrum in apical half dorsally flattened, slightly widened, rugose-punctate; basal half markedly swollen in profile, narrow in dorsal aspect; with indistinct median ridge, densely punctate; dorsolaterally with pair of wide furrows containing erect narrow scales. Pronotum subglabrous, punctate, punctures becoming larger and denser towards apex. Elytra densely punctate; strial punctures slightly larger than minute punctures on intervals; striae impressed as fine lines. Anteroventral ridge of femora terminating with tooth in apical third; tooth largest in metafemur. Metafemur laterally markedly flattened; dorsally with row of silvery scales; posteroventral ridge indistinct, at middle with denticle; dorsoposterior edge weakly denticulate; subapically with stridulatory patch. Aedeagus (Fig. 31b). Body with sides subparallel, extended dorsad; profile of body subtriangular; apex subtruncate, subglabrous; transfer-apparatus complex, symmetrical; ductus ejaculatorius with bulbus. Intraspecific variation. No female specimen available.

Holotype (MZB): ARC1807 (EMBL # HE616084), WEST NEW GUINEA, Jayawijaya Reg., Bokondini, S03°41.787', E138°40.229' to S03°41.778', E138°40.129', 1705–1710 m, 17-VII-2010. Paratype (SMNK): WEST NEW GUINEA, Jayawijaya Reg.: 1 ex, ARC1805 (EMBL # HE616082), Bokondini, S03°40.345', E138°42.386' to S03°40.255', E138°42.189´, 1655–1700 m, 18-VII-2010.

Jayawijaya Reg. (Bokondini). Elevation: 1700–1705 m.

Beaten from foliage of montane forests.

This epithet is a combination of the Latin noun fusus (spindle) and the suffix -formis (-shaped) and refers to the habitus of this species.

Trigonopterus fusiformis Riedel, sp. n. was coded as “Trigonopterus sp. 107” by Tänzler et al. (2012).

Holotype, male (Fig. 32a). Length 1.98 mm. Color black, antenna and tarsi ferruginous. Body laterally somewhat compressed; ovate; without constriction between pronotum and elytron; in profile almost evenly convex. Rostrum dorsally in basal half with pair of sublateral furrows; medially even, sparsely punctate. Eyes large. Pronotum subglabrous. Elytra subglabrous, laterally sparsely punctate with small punctures, striae indistinct. Femora with anteroventral ridge. Profemur subparallel. Meso- and metafemur with dorsoposterior edge subapically worn; metafemur subapically without stridulatory patch. Tibiae simple, without rows or brushes of long setae. Metaventrite laterally forming acute process over metacoxa, reaching tibial insertion. Metaventrite and abdominal ventrite 1 subglabrous, with sparse recumbent setae. Abdominal ventrite 2 fused to and forming common cavity with ventrite 1. Abdominal ventrite 5 anteriorly with pair of distinct longitudinal protrusions, indistinct median cavity posteriorly open. Aedeagus (Fig. 32b) apically with median extension and with pair of submedian teeth; body with two conspicuous pairs of endophallic sclerites; ductus ejaculatorius without bulbus. Intraspecific variation. Length 1.98–2.18 mm. Female rostrum subglabrous except in basal ¼ with ridges. Female abdominal ventrite 5 flat.

Holotype (SMNK): ARC0960 (EMBL # HE615593), PAPUA NEW GUINEA, Central Prov., Moroka area, Kailaki, Mt. Berogoro, S09°24.213', E147°33.870' to S09°23.647', E147°34.244', 500–600 m, 20-IX-2009. Paratypes (NAIC, SMNK, ZSM): PAPUA NEW GUINEA, Central Prov.: 1 ex, ARC0961 (EMBL # HE615594), same data as holotype; 8 exx, Moroka area, Kailaki, Mt. Berogoro, S09°24.213', E147°33.870' to S09°23.647', E147°34.244', 500–565 m, 26-X-2009; 4 exx, ARC0942 (EMBL # HE615575), ARC0943 (EMBL # HE615576), Varirata N.P., S09°26.150', E147°21.520' to S09°26.148', E147°21.361', 700–800 m, 19-IX-2009; 16 Ex, ARC0991 (EMBL # HE615624), Moroka area, Kailaki, Wariaga, S09°25.350', E147°31.047' to S09°25.403', E147°31.315', 650–820 m, 27-X-2009; 9 exx, Moroka area, Kailaki, Wariaga, S09°25.350', E147°31.047' to S09°25.683', E147°31.707', 650–920 m, 27-X-2009; 18 exx, Moroka area, Kailaki, Wariaga, S09°25.403', E147°31.315' to S09°25.683', E147°31.707', 820–920 m, 27-X-2009; 5 exx, Moroka area, Kailaki, Beremutana ridge, S09°25.515', E147°33.136' to S09°25.754', E147°33.485', 535–700 m, 28-X-2009, beaten; 5 exx, Moroka area, Kailaki, Beremutana ridge, S09°25.754', E147°33.485' to S09°25.940', E147°33.703', 700–845 m, 28-X-2009, beaten; 2 exx, Moroka area, Kailaki, Beremutana ridge, S09°25.515', E147°33.136' to S09°25.754', E147°33.485', 535–650 m, 28-X-2009, beaten.

Central Prov. (Varirata, Moroka). Elevation: 565–820 m.

Collected by beating foliage in primary forests.

This epithet is based on the Latin adjective glaber (hairless, bald) and refers to its smooth body.

Trigonopterus glaber Riedel, sp. n. was coded as “Trigonopterus sp. 164” by Tänzler et al. (2012).

Holotype, male (Fig. 33a). Length 2.90 mm. Color black. Body subovate; almost without constriction between pronotum and elytron; in profile evenly convex. Rostrum dorsally relatively flat, with two indistinct submedian rows of punctures, dorsolaterally with pair of furrows continuing along eye; surface weakly microreticulate. Pronotum sparsely punctate with double-punctures each consisting of two minute approximate punctures. Elytra with striae distinct, dorsally punctures small, laterally large; intervals flat, with row of minute punctures, subglabrous. Femora edentate. Metafemur with simple dorsoposterior edge; subapically with stridulatory patch. Tibial base dentiform, when leg extended tibial tooth overlapping femoral apex dorsally. Thoracic and abdominal venter with dense erect setae. Aedeagus (Fig. 33b) apically subangulate, subglabrous; transfer apparatus flagelliform, 1.5 X longer than body; endophallus with two pairs of sclerites; ductus ejaculatorius without bulbus. Intraspecific variation. Length 2.75–2.88 mm. Female venter subglabrous.

Holotype (MZB): ARC1776 (EMBL # HE616053), WEST NEW GUINEA, Jayawijaya Reg., Bokondini, S03°40.345', E138°42.386' to S03°40.255', E138°42.189', 1655–1700 m, 18-VII-2010. Paratypes (SMNK, ZSM): 81 exx, ARC1777 (EMBL # HE616054), ARC1778 (EMBL # HE616055), same data as holotype.

Jayawijaya Reg. (Bokondini). Elevation: ca. 1655–1700 m.

Beaten from foliage of montane forests.

This epithet is based on a combination of the Greek nouns gonatos (knee) and ceros (horn) in apposition and refers to the peculiar extensions of the tibial base.

Trigonopterus gonatoceros Riedel, sp. n. was coded as “Trigonopterus sp. 121” by Tänzler et al. (2012).

Holotype, male (Fig. 34a). Length 2.15 mm. Color black; legs and rostrum deep ferruginous; antenna light ferruginous. Body laterally somewhat compressed, ovate; without constriction between pronotum and elytron; in profile evenly convex. Rostrum dorsally in basal third with low median ridge and pair of submedian ridges; apically subglabrous. Eyes large. Pronotum densely punctate; dorsally punctures small, laterally larger, each with one minute seta; without scales. Elytra dorsally subglabrous, stria 1–2 hardly visible; laterally strial punctures large, relatively shallow. Femora with anteroventral ridge. Profemur converging from base to apex. Meso- and metafemur with dorsoposterior edge in apical third shortened; metafemur subapically without stridulatory patch. Tibiae simple, without rows or brushes of long setae; metatibia subapically with small suprauncal projection. Metaventrite laterally forming acute process over metacoxa, reaching tibial insertion. Metaventrite and abdominal ventrite 1 subglabrous, with sparse recumbent setae. Abdominal ventrite 2 similar to ventrites 3–4. Abdominal ventrite 5 with transverse depression, without distinct cavity; subapically with median denticle. Aedeagus (Fig. 34b) apically sinuate, with deep median incision; ductus ejaculatorius without bulbus. Intraspecific variation. Length 1.98–2.48 mm. Female abdominal ventrite 5 almost flat, densely punctate.

Holotype (MZB): ARC0441 (EMBL # FN429152), WEST NEW GUINEA, Jayapura Reg., Cyclops Mts, Sentani, S02°32.2', E140°30.4', 545–700 m, 02-XII-2007, beaten. Paratypes (ARC, SMNK, ZSM): WEST NEW GUI-NEA, Jayapura Reg., Cyclops Mts, Sentani: 38 exx, ARC0442 (EMBL # FN429153), ARC0443 (EMBL # FN429154), same data as holotype; 1 ex, ARC0653, S02°32.3', E140°30.4', 350–620 m, 19-XI-2007, beaten; 2 exx, S02°31.8', E140°30.5', 600–900 m, 28-XI-2007; 2 exx, S02°32.0', E140°30.4', 700–900 m, 02-XII-2007; 4 exx, S02°31.6', E140°30.4', 900–1100 m, 28-XI-2007, beaten; 11 exx, 950–1450 m, 03-X-1992; 21 exx, 600–1000 m, 05-X-1991; 2 exx, 800–1000 m, 07-VIII-1992; 6 exx, 300–1400 m, 10-VIII-1991; 5 exx, 400–800 m, 07-VIII-1992; 4 exx, 350–850 m, 16-X-1996; 1 ex, 300–550 m, 02-X-1992.

Jayapura Reg. (Cyclops Mts). Elevation: 620–950 m.

Collected by beating foliage in primary forests.

This epithet is based on the Latin noun granum (small kernel, seed) in apposition and refers to the general habitus.

Trigonopterus granum Riedel, sp. n. was coded as “Trigonopterus sp. 15” by Riedel et al. (2010) and Tänzler et al. (2012), respectively “Trigonopterus spo” in the EMBL/GenBank/DDBJ databases. It is closely related to Trigonopterus pseudogranum sp. n., Trigonopterus velaris sp. n., and Trigonopterus imitatus sp. n.; from the latter two it can be distinguished by its sparsely punctate body and the structure of its male abdominal ventrite 5. The externally very similar Trigonopterus pseudogranum sp. n. is best separated by the cox1-sequence which diverges 12.1 %.

Holotype, male (Fig. 35a). Length 3.14 mm. Color black; basal half of elytra with large orange spot. Body elongate-subovate; in dorsal aspect and in profile with distinct constriction between pronotum and elytron. Rostrum slender; dorsally with distinct median costa and pair of narrow submedian ridges; at base with erect white clavate scales, apically replaced by bristles. Eyes large, medially approximate. Pronotum densely punctate-reticulate; sides anteriorly with punctures containing small white scales. Elytra subglabrous, irregularly punctate with minute punctures; striae obsolete; basal margin with ridge extending behind humeri, bordered by row of small indistinct punctures. Femora with tooth in apical half. Mesofemur and metafemur dorsally sparsely squamose with white scales. Metafemur with smooth dorsoposterior edge; subapically without stridulatory patch. Abdominal ventrites 1–2 laterally swollen, medially concave; ventrite 2 posteriorly truncate, posterior face markedly projecting over ventrite 3. Aedeagus (Fig. 35b) subapically slightly widened; apex subangulate, asymmetrical, shifted to left; transfer apparatus dentiform, basally supported by crescent-shaped sclerite; ductus ejaculatorius subapically without bulbus. Intraspecific variation. Length 3.27–3.53 mm. Female rostrum dorsally subglabrous, with submedian rows of minute punctures, with lateral furrow containing row of sparse setae. Female abdominal ventrites 1–2 convex.

Holotype (SMNK): ARC1831 (EMBL # HE616108), PAPUA NEW GUINEA, Eastern Highlands Prov., Okapa, Kimiagomo village, Afiyaleto, S06°25.593', E145°34.862', S06°25.212', E145°35.498', 1911 m, 18-III-2010. Paratypes (NAIC, SMNK, ZSM): PAPUA NEW GUINEA, Eastern Highlands Prov.: 4 exx, ARC1830 (EMBL # HE616107), ARC1832 (EMBL # HE616109), same data as holotype; 1 ex, Okapa, Afiyaleto village, S06°25.593', E145°34.862', 1940 m, 18-III-2010, beaten.

Eastern Highlands Prov. (Okapa). Elevation: 1911–1940 m.

Beaten from foliage of montane forest.

This epithet is based on the Greek noun helios (sun) in apposition and refers to the bright orange spot on the elytra.

Trigonopterus helios Riedel, sp. n. was coded as “Trigonopterus sp. 136” by Tänzler et al. (2012).

Holotype, male (Fig. 36a). Length 2.64 mm. Color black; tarsi and antenna ferruginous. Body subovate; with weak constriction between pronotum and elytron; in profile evenly convex. Rostrum sparsely squamose with suberect scales; in basal half with median carina; in front of eyes with flat lateral extensions; subapically weakly scabrous; epistome smooth, posteriorly forming transverse, angulate ridge, with weak median denticle. Pronotum without subapical constriction, densely coarsely punctate; laterally above procoxa with fovea. Elytra with striae moderately incised on disk; intervals flat, with rows of small punctures; laterally subglabrous; apex subangulate, with pair of sublateral knobs; extended ventrad, slightly beak-shaped. Femora edentate. Metafemur dorsally sparsely squamose, with weakly denticulate dorsoposterior edge; subapically with stridulatory patch. Abdominal ventrite 2 projecting dentiform over elytral edge in profile. Aedeagus (Fig. 36b) apically rounded, sparsely setose; with complex, symmetrical transfer apparatus; ductus ejaculatorius with bulbus. Intraspecific variation. Length 2.28–2.64 mm. Female rostrum with median carina and transverse ridge of epistome less distinct.

Holotype (SMNK): ARC1172 (EMBL # HE615800), PAPUA NEW GUINEA, Morobe Prov., Huon peninsula, Mindik, S06°27.311', E147°24.073', 1570 m, 10-X-2009. Paratypes (NAIC): Morobe Prov., Huon peninsula, Mindik: 1 ex, ARC1176 (EMBL # HE615804), S06°27.221', E147°24.185', 1670 m, 10-X-2009.

Morobe Prov. (Mindik). Elevation: 1570–1670 m.

Sifted from leaf litter in montane forest.

This species is dedicated to the people of Papua New Guinea. The epithet is based on the family name Hitolo, found on page 221 of the Papua New Guinea Telephone Directory of 2010 and treated in genitive plural.

Trigonopterus hitoloorum Riedel, sp. n. was coded as “Trigonopterus sp. 195” by Tänzler et al. (2012).

Holotype, male (Fig. 37a). Length 2.58 mm. Color black; legs and rostrum deep ferruginous; antenna light ferruginous. Body laterally somewhat compressed, ovate, without constriction between pronotum and elytron; in profile evenly convex. Rostrum dorsally in basal third punctate-rugose; apically subglabrous. Eyes large. Pronotum densely punctate, punctures dorsally small, laterally becoming larger, bearing each one minute seta; without scales. Elytra dorsally subglabrous, stria 1–2 hardly visible; laterally strial punctures large, relatively shallow. Profemur converging from base to apex. Meso- and metafemur with in apical 1/3 with anteroventral ridge terminating as tooth. Metafemur subapically without stridulatory patch. Tibiae simple, without rows or brushes of long setae. Metaventrite laterally forming acute process over metacoxa, reaching tibial insertion. Metaventrite and abdominal ventrite 1 with long erect setae, especially near mesocoxa. Abdominal ventrite 2 similar to ventrites 3–4. Abdominal ventrite 5 with deep, transversely ovate cavity; basal third simple, swollen; laterally setose. Aedeagus (Fig. 37b) apically sinuate, with deep narrow median incision; ductus ejaculatorius without bulbus. Intraspecific variation. Length 1.98–2.58 mm. Female rostrum subglabrous except in basal ¼ with ridges. Female venter sparsely setose, abdominal ventrite 5 flat.

Holotype (MZB): ARC1697 (EMBL # HE615984), WEST NEW GUINEA, Biak Reg., Supiori Isl., Korido, S00°49.715', E135°35.055', 50–100 m, 9-VII-2010, beaten. Paratypes (SMNK, ZSM): 4 exx, ARC1698 (EMBL # HE615985), ARC1699 (EMBL # HE615986), same data as holotype.

Biak Reg. (Supiori Isl.). Elevation: ca. 50–100 m.

Collected by beating foliage in primary forests.

This epithet is based on the Latin participle imitatus (imitated) and refers to its morphological similarity with sibling species.

Trigonopterus imitatus Riedel, sp. n. was coded as “Trigonopterus sp. 273” by Tänzler et al. (2012). It is closely related to Trigonopterus granum sp. n., Trigonopterus pseudogranum sp. n., and Trigonopterus velaris sp. n. from which it can be distinguished by the male venter with long setae. Despite its close morphological similarity its cox1-sequence diverges 9.9–12.3 % from the other species.

Holotype, male (Fig. 38a). Length 2.90 mm. Color black, elytra with greenish-bronze lustre; antenna light ferruginous, legs dark ferruginous. Body subovate; in dorsal aspect and in profile almost without constriction between pronotum and elytron. Rostrum with distinct median and pair of submedian carinae, in apical third scabrous; epistome simple. Pronotum relatively small, with weak subapical constriction; disk densely punctate, longitudinally rugose; each puncture with seta, a few with scale. Elytra with striae deeply impressed; with sparse, yellowish scales; intervals subglabrous. Meso- and metafemur edentate. Metafemur with dorsoposterior edge serrate, subapically without stridulatory patch. Metatibia apically with blunt premucro. Onychium ca. 1.1× longer than tarsomere 3. Aedeagus (Fig. 38b) with apex asymmetrical, right side forming biramose extension; basal orifice ventrally with rim; transfer apparatus spiniform, curved, directed basad; ductus ejaculatorius without bulbus. Intraspecific variation. Length 2.45–2.90 mm. Female rostrum dorsally subglabrous, with dense small punctures, in basal third with longitudinal ridges. Female metatibia apically without premucro.

Holotype (MZB): ARC0858 (EMBL # HE615540), WEST NEW GUINEA, Biak Isl., Korim, Nernu, S00°55.784', E136°01.530', 165 m, 15-XII-2007, sifted. Paratypes (ARC, SMNK, ZSM): WEST NEW GUINEA, Biak Isl.: 28 exx, ARC0859 (EMBL # HE615541), same data as holotype; 1 ex, ARC0883 (EMBL # HE615565), Korim, Nernu, S00°55.888', E136°01.671', 180 m, 15-XII-2007, sifted; 3 exx (1 marked ARC0093), Korim, Nernu, 150 m, 20-I-2005, sifted.

Biak Isl.. Elevation: 150–180 m.

Sifted from leaf litter in lowland forest.

This epithet is based on the Latin participle inflatus (swollen) and refers to the shape of the elytra that appear somewhat disproportionate in relation to the pronotum.

Trigonopterus inflatus Riedel, sp. n. was coded as “Trigonopterus sp. 63” by Tänzler et al. (2012).

Holotype, male (Fig. 39a). Length 3.41 mm. Color black; antenna and tarsi ferruginous. Body subovate; without constriction between pronotum and elytron; in profile evenly convex. Rostrum in basal half with distinct median carina and pair of indistinct sublateral ridges; with sparse rows of white scales; subapical third subglabrous, weakly punctate; in front of antennal insertion with weak constriction. Eyes large, medially approximate. Pronotum densely punctate with small punctures; sides separated by indistinct edge bearing dense row of punctures; laterally above coxa sparsely squamose with white scales. Elytral striae distinct with small punctures; intervals flat, subglabrous, sparsely punctate with minute punctures; basal margin straight, simple. Femora with anteroventral ridge distinct, at base abruptly ending and forming markedly projecting blunt angle; edentate. Mesofemur and metafemur dorsally densely squamose with white scales but partly abraded. Metafemur with smooth dorsoposterior edge; subapically without stridulatory patch. Aedeagus (Fig. 39b) with apodemes ca. 3× as long as body; sides of body in basal half subparallel, markedly sclerotized, mid-portion of body weakly sclerotized; apex subangulate; transfer apparatus flagelliform, longer than body; ductus ejaculatorius subapically with bulbus. Intraspecific variation. Length 3.38–3.41 mm. No female specimen available.

Holotype (MZB): ARC1694 (EMBL # HE615981), WEST NEW GUINEA, Biak Reg., Supiori Isl., Korido, S00°49.715´, E135°35.055´, 50–100 m, 09-VII-2010. Paratype (SMNK): 1 ex, ARC1695 (EMBL # HE615982), same data as holotype.

Biak Reg. (Supiori Isl.). Elevation: ca. 50–100 m.

Beaten from foliage of lowland forest.

This epithet is based on the Latin adjective insularis (of an island) and refers to the type locality, Supiori Island.

Trigonopterus insularis Riedel, sp. n. was coded as “Trigonopterus sp. 57” by Tänzler et al. (2012).