Distinguished from other genera of Tenthredininae by deep furrow on ventral side of antennal flagellum and the very large, protruding and isolated supra-antennal tubercles.

Body robust (Figs 1, 8). Clypeus transversely subconvex at basal third, anterior margin sharp, quarter-circularly incised with acute lateral teeth; apical third of labrum deflexed, and labrum thus truncate at apex in front view (Figs 5, 11); mandibles strongly bent at apical third, asymmetric (Figs 2, 3), right one with 1 basal tooth, 3 inner teeth; one of them placed on the dorsal side (Fig. 3); left mandible with 1 basal tooth and 2 inner teeth, without dorsal tooth (Fig. 2); malar space longer than diameter of median ocellus; inner margins of eyes very feebly S-curved and converging downwards, distance between eyes much longer than height of eye (Figs 5, 11); supra-antennal tubercles large, highly elevated and quite free-standing (Figs 4, 10, 12); head elongated behind eyes, occipital carina low but distinct; interocellar furrow broad and deep, postocellar furrow broad and shallow. Antennae stout and uniformly thick; 2^nd antennomere about as long as broad, flagellum with a very deep and sharp longitudinal furrow along the outer-side from apex of the 3^rd antennomere to 9^th antennomere, 3^rd antennomere distinctly longer than 4^th (Figs 6, 9). Anterior lobe of pronotum broad, without marginal carina, with broadest part about 3 × diameter of an ocellus; ventral margins of propleura narrowly but distinctly meeting; mesoscutellum flattened, mesoscutellar appendage with an obtuse middle ridge. anterior basin of metascutellum narrow, furrow like; dorsal lobe of metepimeron long and linear; mesosternal thorns wanting; posterior corner of metepimeron round, without appendage. Basal plates (first tergite) of abdomen contiguous on meson, without large membranous blotch (Fig. 8). Inner tibial spur of fore leg bifurcate at apex; hind coxa small, hind femur not reaching apex of abdomen, distinctly shorter than hind tibia, hind tibial spur shorter than half length of metabasitarsus; metabasitarsus about as long as or slightly longer than following 3 tarsomeres together; claw cleft, inner tooth shorter than outer tooth. Venation similar to Tenthredo, anal cell of forewing with a short and erect cross vein at basal 2/5; hind wing with 2 closed middle cells, anal cell sessile or shortly petiolate (Figs 1, 8).

Himalaya: China (Tibet), India (Jammu & Kashmir, Uttaranchal, Himachal Pradesh), Nepal, Pakistan (Islamabad).

Holotype ♀ (Fig. 1).

Body length 11 mm. Greenish yellow, following parts black: apex of each mandible; antennae entirely; supra-antennal tubercle, frons and adjacent area of upper inner orbit except a stripe on frontal ridge, postocellar area, a broad band from upper hind corner of each eye to postocellar area (Figs 1, 4); a medial band on pronotum; meso- and metanotum, except mesoscutellum and 2 small triangular spots on mesoscutal lateral lobe and 2 linear spots on mesoscutal median lobe; abdomen above, except for tergites 8–10 and a broad band on posterior margin of tergites 3–4 (in the middle covering about two thirds, laterally about one third of the length of the tergite); depressed upper corner of mesepimeron and a minute and roundish dot in lower posterior corner of mesepisternum. Legs greenish yellow, following parts black: a small dot on extreme apex of each femur; a narrow dorsal stripe on each tibia and tarsus. Wings hyaline with very faint yellow tinge, vein C and stigma yellow brown, other veins pale brown. Body hairs silver.

Body shiny; labrum without large punctures; central area of clypeus, outer sides of mandibles with some large punctures, head above antenna including supra-antennal tubercle with large and deep punctures, interspaces between punctures about as large as a puncture, surface smooth and shiny; punctures on postorbit sparser but distinct; punctures on mesoscutal median lobe and mesoscutal lateral lobe smaller than punctures on head, interspaces shiny; posterior margin of mesoscutellum densely punctured, dorsal side of mesoscutellum hardly punctured; mesoscutellar appendage impunctate, parapsis of mesothorax microsculptured; metascutellum with some minute punctures; mesepisternum densely punctured, narrow interspaces smooth, strongly shiny; metepisternum densely punctured, mesepimeron and metepimeron polished, with scattered punctures. Abdominal tergites 1–2 strongly shiny, tergite 3 weakly microsculptured in basal half, tergites 4–7 distinctly microsculptured and punctured in basal half.

Clypeus and labrum as Fig. 5; left mandible as Fig. 2, right mandible as Fig. 3; ED = 1.7; supra-antennal tubercle larger than scape (Fig. 4), clearly above top of ocelli in lateral view; a distinct tubercle present on anterior margin of middle fovea and almost in line with anterior margins of supra-antennal tubercles; frontal walls broad and flat, parallel to each other; postocellar area 1.2 × as broad as long; lateral furrows deep, distinctly divergent backwards; POL : OOL : OCL = 1 : 2.5 : 2.5; head behind eyes about as long as eyes in dorsal view, lateral edge roundly curved (Fig. 4); occipital carina distinct in entire length. Antennae stout, weakly compressed, 0.9 × length of head and thorax combined, antennomere 3 0.9 × length of antennomeres 4 and 5 combined, antennomeres 6–8 1.7 × as long as broad (Fig. 6). Anal cell of hind wing sessile. Ovipositor sheath 1.1 × length of fore tibia. Lancet with 21 serrulae, serrulae 5–6 as Fig. 7.

Variability (females). Body length 10–11 mm. Punctation on upper head varies from sparse to dense; supra-antennal tubercle sometimes only with few punctures and / or smaller than scape; tergites 1–2 smooth to rather densely punctured laterally and on basal half; POL : OOL : OCL = 1 : 2.0–2.7 : 2.0–2.6; postocellar area 1.2–1.4 × as broad as long; head behind eyes 0.8–1.0 × as long as eyes in dorsal view. Ratio of length and breadth of antennomeres 6–8: 1.5–1.9. Black spot on lower mesepisternum may be indistinct or missing; pale color on tergite 4 may be reduced to a triangular spot; greenish color may alter (in dried specimens) to yellow.

Male. (Figs 8–10) Body length 8–9 mm. Color and structure similar to female. Generally darker than the female: pale stripe on upper inner orbit very narrow and only exceptionally nearly extending toward postocellar area; frontal ridges black or pale marked; hind tibia and tarsus entirely black; pale macula on mesoscutellum more or less reduced; mesoscutal median lobe usually entirely black, without pale lateral stripe; tegula black, basally pale; lower mesepisternum usually with broad black stripe, sometimes anteriorly reduced to a large spot; tergites 3-4 dorsally usually only with faintly indicated pale hind margin, sometimes completely black; subgenital plate apical margin rounded. Penis valve Fig. 10.

Pictures of several paratypes are given in original resolution at https://doi.org/10.6084/m9.figshare.781286

The specific epithet, an adjective, refers to the distinctly punctured head.

China (Tibet), Nepal (Karnali).

♀, China: Tibet, Yadong, Naiduilashan, 27°24'51"N, 88°56'08"E, 3100m, 2003.VIII.29, Wei Meicai leg. Coll. CSCS.

(7 ♀♀, 10 ♂♂, DEI-GISHym 17227–17243, in coll. CSCS, DEI, NHRS, NKME). Nepal, Prov. Karnali: 1 ♀, Umg. Churta 3000–3400 m, 31.V.2007 (DEI-GISHym 17228, BOLD:ACG2198); 1 ♀, Umg. Churta E Hochtal 3500–4000 m, 02.VI.–04.VI.2007; 4 ♂♂, Umg. Churta E Hochtal 3500–3800 m, 02.VI.2007; 1 ♀, Gothichaur, 29°12'10"N, 82°18'56"E, 2850 m, 09.VI.1997; 1 ♀, Gothichaur, Thymian-Wiese, 3100 m, 26.V.2007; 1 ♀ 2 ♂♂, Gothichaur, 29°11'54"N, 82°18'36"E, 2850 m, Sumpfwiese, 26.V.–05.VI.2007; 4 ♂♂, Gothichaur, 29°11'54"N, 82°18'36"E, 2850 m, Umg. Lager, 26.V.–05.VI.2007; all specimens F. Creutzburg leg.; 1 ♀, Gothichaur 2 km W, 2700 m; 20.-21.V.1995, M. Hartmann leg.; 1 ♀, Gothichaur, Wald, 29°12'10"N, 82°18'56"E, 2850 m, 08.VI.1997, M. Hartmann leg.

Lectotype of Allantus dorsatus ♀ (Fig. 11), additions based on other specimens are given in brackets [ ].

Body length 11 mm. Greenish yellow, following parts black: apex of each mandible; antennae entirely (Fig. 14); supra-antennal tubercle, frons and adjacent area of upper inner orbit except a short stripe on frontal ridge, postocellar area, a broad band from upper hind corner of each eye to postocellar area (Figs 12, 13); a medial band on pronotum; meso- and metanotum, except mesoscutellum and 2 small triangular spots on mesoscutal lateral lobe and 2 linear spots on mesoscutal median lobe (the latter sometimes absent); abdomen above, except for tergites 8–10 and posterior 2/5 of tergites 3–4; depressed upper corner of mesepimeron and a small and roundish spot in lower posterior corner of mesepisternum. Legs greenish yellow, following parts black: a small dot on extreme apex of each femur; a narrow dorsal stripe on fore and middle tibiae, dorsal side of hind tibia; a narrow stripe on fore and middle tarsi above (sometimes interrupted), hind tarsus entirely. Wings hyaline with faint yellowish tinge, veins and stigma pale brown, base of stigma slightly darkened. Body hairs silver.

Body shiny; labrum and outer side of mandible with some large punctures, head otherwise almost impunctate, frontal basin feebly microsculptured; mesoscutal median lobe and mesoscutal lateral lobe sparsely punctured, interspaces smooth and shiny; posterior margin of mesoscutellum densely punctured, dorsal side of mesoscutellum hardly punctured; mesoscutellar appendage and parapsis impunctate; metascutellum with some minute punctures; mesepisternum shallowly but distinctly punctured, surface smooth, strongly shiny; metepisternum densely punctured, mesepimeron and metepimeron polished, with scattered punctures. Abdominal tergites 1–2 strongly shiny, tergite 3 weakly microsculptured in basal half, tergites 4–7 distinctly microsculptured and punctured in basal half.

Clypeus, labrum, mandibles as Fig. 11; ED = 1.3; supra-antennal tubercle distinctly smaller than scape (Figs 12, 13), clearly lower than top of ocelli in lateral view; a small tubercle present in bottom of middle fovea and in line with posterior margins of supra-antennal tubercles; frontal walls broad and flat, weakly divergent forwards; postocellar area broader than long as 7 : 5; lateral furrows deep, weakly divergent backwards; POL : OOL : OCL = 15 : 29 : 27; head behind eyes about 0.7 × length of eyes in dorsal view, indistinctly narrowed (Fig. 12); occipital carina distinct in lower 2/3 and rather weak in dorsal 1/3. Antennae weakly compressed, as long as head and thorax combined, antennomere 3 0.9 × length of antennomeres 4 and 5 together, [antennomeres 6–8 1.5–2.4 × as long as broad (Fig. 14)]. Anal cell of hind wing with a very short petiole. Ovipositor sheath as long as fore tibia. [Lancet with 20 serrulae, serrulae 5–6 as Fig. 15].

Male. Body length 8 mm. Similar to female except head distinctly narrowed behind eyes in dorsal view, malar space slightly longer than diameter of median ocellus; pale stripe from upper inner orbit to postocellar area missing; subgenital plate rounded at apex; penis valve simple, valviceps weakly bent; harpe about 2 × as long as broad (see figs 678 and 680 in Saini 2007).

India: Himachal Pradesh, Jammu & Kashmir, Uttaranchal (Saini et al. 2006); Pakistan: Islamabad (Togashi 1989). It is ambiguous whether these records actually base on Tyloceridius dorsatus or on the new Tyloceridius stictocephalus. We studied material from Himachal Pradesh, Jammu & Kasmir and Uttaranchal. Referring to Muche (1983), Saini (2007) recorded Tyloceridius dorsatus also from Nepal and Bhutan, but Muche mentioned only 1 ♀ (!) from India (“Chaurengi” located in former Uttarpradesh).

Lectotype (Fig. 11): 1♀, “Himalaya, Plasow”; “Allantus dorsatus Mocs., India Oriental”; “Lectotypus Allantus dorsatus Mocs., design. A. Taeger, 1988” [red]; “Tyloceridius dorsatus (Mocs.), det. A. Taeger, 88”. (HNHM). Left antenna, apical 3 antennomeres of right antenna, right foreleg below femur, left middle tarsus, right hind tarsus and left leg below femur are missing.

Paralectotype. 1 ♀, “Himalaya, Plasow”; “Typus Allantus dorsatus Mocs.”; “Syntype [sic!] Allantus dorsatus Mocsáry, 1883. teste A. Taeger, 2011” [red]; “Tylocerus [sic!] gen. n. bituberculatus Cam., Malaise det. 1935”; “DEI-GISHym 10877” (HNHM, figs see https://doi.org/10.6084/m9.figshare.781292).

1 ♀ (Figs 12–15), India, Uttaranchal (former N Uttar Pradesh), 5. 7. 1989, Rishikesh [30.117°N, 78.317°E], A. Riedel leg. (SDEI); 1♀ same data (see https://doi.org/10.6084/m9.figshare.903712); 2 ♀♀, India, Kalatop, (H.P.), 8200’, 11.7.83, M.S. Saini (USNM); 1♀1♂, India, Himachal Pradesh, Kalatop, 2400m, July 1983, M.S. Saini collector, Tyloceridius dorsatus Malaise (!) (USNM); 1 ♀ 2 ♂♂, kept in NHRS, data unrecorded; 1 ♀ 1 ♂ India, Uttarakhand, Joshimath, 14.6.1983, leg. Balbir (NHRS, photo documentation by H. Vårdal).

Saini (2007) described both sexes and illustrated the lancet, penis valve and gonoforceps. This species is widely distributed at higher altitudes in Uttaranchal & Himachal Pradesh (Saini 2007).