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Citation: Nardi G, Háva J (2013) Italian Dermestidae: notes on some species and an updated checklist (Coleoptera). ZooKeys 360: 45–81. doi: 10.3897/zookeys.360.6023
An up-to-date checklist of the Italian Dermestidae is provided. The presence of 95 species in Italy is confirmed, while further 5 species (Dermestes (Dermestes) vorax Motschulsky, 1860, Thorictuspilosus Peyron, 1857, T. wasmanni Reitter, 1895, Attagenus (Attagenus) simonis Reitter, 1881 and Globicornis (G.) breviclavis (Reitter, 1878)) and 1 subspecies (A. (A.) tigrinus pulcher Faldermann, 1835) are excluded from the Italian fauna.
Attagenus (Attagenus) calabricus Reitter, 1881 and A. (A.) lobatus Rosenhauer, 1856 are for the first time recorded from Abruzzi and Tuscany respectively; A. (A.) silvaticus Zhantiev, 1976 is recorded for the first time from mainland Italy (Apulia); Anthrenus (Anthrenus) angustefasciatus Ganglbauer, 1904 is new to northern Italy (Friuli-Venezia Giulia), central Italy (Tuscany), Apulia and Basilicata; A. (A.) munroi Hinton, 1943 is new to central Italy (Elba Island); A. (A.) delicatus Kiesenwetter, 1851 is for the first time recorded from Apulia; Globicornis (Globicornis) fasciata (Fairmaire & Brisout de Barneville, 1859) is new to southern Italy (Basilicata); G. (Hadrotoma) sulcata (C.N.F. Brisout de Barneville, 1866) is for the first time recorded from central Italy (Abruzzi), Campania and Sicily, whileTrogoderma inclusum LeConte, 1854 is new to Apulia.
Seven species (Dermestes (Dermestes) peruvianus Laporte de Castelnau, 1840, D. (Dermestinus) carnivorus Fabricius, 1775, D. (Dermestinus) hankae Háva, 1999, D. (Dermestinus) intermedius intermedius Kalík, 1951, D. (Dermestinus) szekessyi Kalík, 1950, Anthrenus (Anthrenops) coloratus Reitter, 1881 and Trogodermaangustum (Solier, 1849)) recently recorded from Italy (without further details) are discussed.
The lectotype and a paralectotype are designated forAttagenus (A.) calabricus Reitter, 1881 from Calabria.
Attagenus pellio (Linnaeus, 1758) var. pilosissimus Roubal, 1932 is removed from synonymy with A. (A.) pellio and recognized as a valid species (stat. prom.); it is known from Lombardy, Apulia and Calabria.
Dermestidae, lectotype designation, new status, taxonomy, distribution, new records, chorotypes, checklist, Italy
The main purpose of the present paper is to enhance knowledge of the Dermestidae of Italy. It is chiefly based on material collected by the staff of the Centro Nazionale per lo Studio e la Conservazione della Biodiversità Forestale “Bosco Fontana” di Verona (Italy), during entomological surveys in various Italian Protected Areas (e.g.
This paper is organized in three sections: “Notes on some species”, “Literature records” and “A new checklist of the Dermestidae of Italy”.
The classification of the subfamilies, tribes, genera and subgenera follows that proposed by
The first section is based chiefly on study of new material. With very few exceptions, identifications were made by J. Háva.
The second section summarizes the literature data updating the Italian distribution of the species provided by
In both sections, nomenclatural combinations and possible synonyms (listed chronologically-alphabetically) found in the literature that includes Italian records are provided below the valid name of each species or subspecies.
The first section concerns 25 species. As far as possible, the following data are provided for each record: region, province, general area and/or commune, locality of collection, altitude, UTM coordinates, date of collection, collector/s, collecting method, possible research project, number of specimens, and, in parentheses, abbreviation of depository. The labels of the examined specimens are generally written in Italian; hereunder, the regions and the collecting methods were translated into English. The label data of type specimens are cited verbatim between quotation marks, a forward slash separating the different lines on a label, and a comma separating subsequent labels. Comments and interpretations are given in square brackets.
The mainland Italian regions are listed from north to south, and from west to east, all toponyms are listed alphabetically. Chorotypes were assigned according to
In the second section, the literature records updating the checklist of Dermestidae of Italy provided by
Finally, a new checklist of the Italian Dermestidae is provided (Table 1). The total number of species for each geographic division includes only those recorded with certain, so generic records for “Italy” and records followed by a question mark are excluded.
Acronyms for chorotypes follow
Checklist of the Dermestidae of Italy.
Taxa | Distribution | Chorotype | |||
---|---|---|---|---|---|
Dermestinae Latreille, 1804 | |||||
Dermestini Latreille, 1804 | |||||
Genus Dermestes Linnaeus, 1758 | |||||
Subgenus Dermestes Linnaeus, 1758 | |||||
1. Dermestes (Dermestes) ater DeGeer, 1774 | N | COS | |||
2. Dermestes (Dermestes) bicolor Fabricius, 1781 | PAL | ||||
Dermestes (Dermestes) bicolor bicolor Fabricius, 1781 | N | S | Sa | ||
3. Dermestes (Dermestes) haemorrhoidalis Küster, 1852 | N | COS | |||
4. Dermestes (Dermestes) hispanicus Kalík, 1952 | Sa | WME | |||
5. Dermestes (Dermestes) lardarius Linnaeus, 1758 | N | S | Si | Sa | COS |
6. Dermestes (Dermestes) peruvianus Laporte de Castelnau, 1840 | N? | S | COS | ||
Subgenus Dermestinus Zhantiev, 1967 | |||||
7. Dermestes (Dermestinus) aurichalceus Küster, 1846 | N | S? | Sa | MED | |
8. Dermestes (Dermestinus) carnivorus Fabricius, 1775 | S | COS | |||
9. Dermestes (Dermestinus) erichsoni Ganglbauer, 1904 | N | S | Si | Sa | EUR |
10. Dermestes (Dermestinus) frischii Kugelann, 1792 | N | S | Si | Sa | COS |
11. Dermestes (Dermestinus) gyllenhalii Laporte de Castelnau, 1840 | EUR | ||||
Dermestes (Dermestinus) gyllenhalii gyllenhalii Laporte de Castelnau, 1840 | N | S | Si? | ||
12. Dermestes (Dermestinus) hankae Háva, 1999 | S | WME | |||
13. Dermestes (Dermestinus) intermedius Kalík, 1951 | EUR | ||||
Dermestes (Dermestinus) intermedius intermedius Kalík, 1951 | IT | IT | |||
14. Dermestes (Dermestinus) laniarius Illiger, 1801 | PAL | ||||
Dermestes (Dermestinus) laniarius laniarius Illiger, 1801 | N | S | Si | Sa | |
15. Dermestes (Dermestinus) maculatus DeGeer, 1774 | N | S | Si | Sa | COS |
16. Dermestes (Dermestinus) murinus Linnaeus, 1758 | PAL | ||||
Dermestes (Dermestinus) murinus murinus Linnaeus, 1758 | N | S | Si | Sa | |
17. Dermestes (Dermestinus) mustelinus Erichson, 1846 | N | S | Si | Sa | EUR |
18. Dermestes (Dermestinus) pardalis Billberg, 1808 | N? | IT | Sa | WME | |
19. Dermestes (Dermestinus) sardous Küster, 1846 | MED | ||||
Dermestes (Dermestinus) sardous sardous Küster, 1846 | N | S | Sa | ||
20. Dermestes (Dermestinus) szekessyi Kalík, 1950 | IT | IT | SIE | ||
21. Dermestes (Dermestinus) undulatus Brahm, 1790 | N | S | Si | Sa | OLA |
Subgenus Montandonia Jacquet, 1886 | |||||
22. Dermestes (Montandonia) fuliginosus P. Rossi, 1792 | N | S | Si | EUR | |
23. Dermestes (Montandonia) hirticollis Fabricius, 1792 | Sa | WME | |||
24. Dermestes (Montandonia) olivieri Lepesme, 1939 | N | S | Si | Sa | TEM |
Thorictinae Agassiz, 1846 | |||||
Thaumaphrastini Anderson, 1949 | |||||
Genus Thorictodes Reitter, 1875 | |||||
25. Thorictodes heydeni Reitter, 1875 | N | S | COS | ||
Thorictini Agassiz, 1846 | |||||
Genus Thorictus Germar, 1834 | |||||
26. Thorictus grandicollis Germar, 1842 | TUM | ||||
Thorictus grandicollis grandicollis Germar, 1842 | N | S | Si | Sa | |
27. Thorictus mauritanicus Lucas, 1846 | Si | Sa | MED | ||
28. Thorictus stricticollis Kraatz, 1859 | MED | ||||
Thorictus stricticollis stricticollis Kraatz, 1859 | Sa | ||||
29. Thorictus studti John, 1971 | Si | SICI | |||
Orphilinae LeConte, 1900 | |||||
Genus Orphilus Erichson, 1848 | |||||
30. Orphilus niger (P. Rossi, 1790) | N | S | Si | Sa | CEM |
Trinodinae T.L. Casey, 1900 | |||||
Thylodriini Semenov, 1909 | |||||
Genus Thylodrias Motschulsky, 1839 | |||||
31. Thylodrias contractus Motschulsky, 1839 | N | COS | |||
Trinodini T.L. Casey, 1900 | |||||
Genus Trinodes Dejean, 1821 | |||||
32. Trinodes hirtus (Fabricius, 1781) | N | S | Si | Sa | TUE |
Attageninae Laporte, 1840 | |||||
Attagenini Laporte, 1840 | |||||
Genus Attagenus Latreille, 1802 | |||||
Subgenus Attagenus Latreille, 1802 | |||||
33. Attagenus (Attagenus) bifasciatus (A.G. Olivier, 1790) | S | Si | Sa | TUM | |
34. Attagenus (Attagenus) brunneus Faldermann, 1835 | N | S | Si | Sa | OLA |
35. Attagenus (Attagenus) calabricus Reitter, 1881 | S | Si | APPE | ||
36. Attagenus (Attagenus) cyphonoides Reitter, 1881 | S | SCO | |||
37. Attagenus (Attagenus) fallax Gené, 1839 | S | Si | Sa | MED | |
38. Attagenus (Attagenus) fasciatus (Thunberg, 1795) | N | S | COS | ||
39. Attagenus (Attagenus) lobatus Rosenhauer, 1856 | S | Si | Sa | CEM | |
40. Attagenus (Attagenus) maritimus (Gené, 1839) | Si | Sa | WME | ||
41. Attagenus (Attagenus) obtusus (Gyllenhal, 1808) | Sa | TEM | |||
42. Attagenus (Attagenus) pantherinus (Ahrens, 1814) | N | Sa | EUR | ||
43. Attagenus (Attagenus) pellio (Linnaeus, 1758) | N | S | Si | Sa | COS |
44. Attagenus (Attagenus) pilosissimus Roubal, 1932 | N | S | ALAP | ||
45. Attagenus (Attagenus) punctatus (Scopoli, 1772) | N | S | Si | EUR | |
46. Attagenus (Attagenus) rossii Ganglbauer, 1904 | S | Si | Sa | MED | |
47. Attagenus (Attagenus) schaefferi (Herbst, 1792) | OLA | ||||
Attagenus (Attagenus) schaefferi schaefferi (Herbst, 1792) | N | S | Si | Sa | |
48. Attagenus (Attagenus) silvaticus Zhantiev, 1976 | S | Si | SIE | ||
49. Attagenus (Attagenus) simplex Reitter, 1881 | S | Si | Sa | MED | |
50. Attagenus (Attagenus) tigrinus (Fabricius, 1792) | |||||
Attagenus (Attagenus) tigrinus tigrinus (Fabricius, 1792) | N | S | Si | Sa | MED |
51. Attagenus (Attagenus) trifasciatus (Fabricius, 1787) | N | Si? | MED | ||
52. Attagenus (Attagenus) unicolor (Brahm, 1790) | COS | ||||
Attagenus (Attagenus) unicolor unicolor (Brahm, 1790) | N | S | Si | Sa | |
53. Attagenus (Attagenus) uniformis Fairmaire, 1860 | Si | NAF | |||
Megatominae Leach, 1815 | |||||
Anthrenini Gistel, 1848 | |||||
Genus Anthrenus Geoffroy, 1762 | |||||
Subgenus Anthrenops Reitter, 1881 | |||||
54. Anthrenus (Anthrenops) coloratus Reitter, 1881 | S | OLA+AFR | |||
Subgenus Anthrenus Geoffroy, 1762 | |||||
55. Anthrenus (Anthrenus) angustefasciatus Ganglbauer, 1904 | N | S | EUR | ||
56. Anthrenus (Anthrenus) delicatus Kiesenwetter, 1851 | N | S | Si | Sa | MED |
57. Anthrenus (Anthrenus) festivus Erichson, 1846 | S? | Si | Sa | MED | |
58. Anthrenus (Anthrenus) flavipes LeConte, 1854 | COS | ||||
Anthrenus (Anthrenus) flavipes flavipes LeConte, 1854 | S | Sa | |||
59. Anthrenus (Anthrenus) goliath Saulcy, 1868 | N | S | Si | Sa | MED |
60. Anthrenus (Anthrenus) munroi Hinton, 1943 | S | Sa | MED | ||
61. Anthrenus (Anthrenus) mroczkowskii Kalík, 1954 | S | SEU | |||
62. Anthrenus (Anthrenus) pimpinellae (Fabricius, 1775) | COS | ||||
Anthrenus (Anthrenus) pimpinellae isabellinus Küster, 1848 | N | S | Si | Sa | |
Anthrenus (Anthrenus) pimpinellae pimpinellae (Fabricius, 1775) | N | S | Si | ||
63. Anthrenus (Anthrenus) scrophulariae (Linnaeus, 1758) | COS | ||||
Anthrenus (Anthrenus) scrophulariae scrophulariae (Linnaeus, 1758) | N | S | Si | Sa | |
Subgenus Florilinus Mulsant & Rey, 1868 | |||||
64. Anthrenus (Florilinus) museorum (Linnaeus, 1761) | N | S | Si | Sa | COS |
65. Anthrenus (Florilinus) oberthueri Reitter, 1881 | N | Si | WME | ||
Subgenus Helocerus Mulsant & Rey, 1868 | |||||
66. Anthrenus (Helocerus) fuscus A.G. Olivier, 1790 | N | S | Si | Sa | SCO |
67. Anthrenus (Helocerus) minutus Erichson, 1846 | S | Si | Sa | MED | |
Subgenus Nathrenus Casey, 1900 | |||||
68. Anthrenus (Nathrenus) biskrensis Reitter, 1887 | Si | NAF | |||
69. Anthrenus (Nathrenus) molitor Aubé, 1850 | N | Sa | MED | ||
70. Anthrenus (Nathrenus) signatus Erichson, 1846 | N | S | Si | EUR | |
71. Anthrenus (Nathrenus) verbasci (Linnaeus, 1767) | N | S | Si | Sa | COS |
72. Anthrenus (Nathrenus) versicolor Reitter, 1887 | Si | Sa | MED | ||
Megatomini Leach, 1815 | |||||
Genus Anthrenocerus Arrow, 1915 | |||||
73. Anthrenocerus australis (Hope, 1843) | N | AUST (i) | |||
Genus Ctesias Stephens, 1830 | |||||
Subgenus Ctesias Stephens, 1830 | |||||
74. Ctesias (Ctesias) serra (Fabricius, 1792) | N | S | Si | Sa | EUR |
Genus Globicornis Latreille, 1829 | |||||
Subgenus Globicornis Latreille, 1829 | |||||
75. Globicornis (Globicornis) bifasciata (Perris, 1866) | Si | Sa | WME | ||
76. Globicornis (Globicornis) fasciata (Fairmaire & Brisout de Barneville, 1859) | N | S | Si? | Sa | CEU |
77. Globicornis (Globicornis) luckowi Herrmann, Háva & Kadej, 2011 | N | ALPC | |||
78. Globicornis (Globicornis) nigripes (Fabricius, 1792) | N | S | EUR | ||
79. Globicornis (Globicornis) picta (Küster, 1851) | S | Si | Sa | EUR | |
80. Globicornis (Globicornis) tristis (Reitter, 1881) | S | EME | |||
81. Globicornis (Globicornis) variegata (Küster, 1851) | N | S | Si | Sa | SEU |
Subgenus Hadrotoma Erichson, 1848 | |||||
82. Globicornis (Hadrotoma) corticalis (Eichhoff, 1863) | N | S | Si | EUR | |
83. Globicornis (Hadrotoma) emarginata (Gyllenhal, 1808) | N | S | Si | Sa | EUR |
84. Globicornis (Hadrotoma) sulcata (C.N.F. Brisout de Barneville, 1866) | S | Si | WME | ||
Genus Megatoma Herbst, 1791 | |||||
Subgenus Megatoma Herbst, 1791 | |||||
85. Megatoma (Megatoma) ruficornis Aubé, 1866 | S | SEU | |||
86. Megatoma (Megatoma) undata (Linnaeus, 1758) | N | S | EUR | ||
Genus Phradonoma Jacquelin du Val, 1859 | |||||
87. Phradonoma villosulum (C. Duftschmid, 1825) | N | CAE | |||
Genus Reesa Beal, 1967 | |||||
88. Reesa vespulae (Milliron, 1939) | N | SCO | |||
Genus Trogoderma Dejean, 1821 | |||||
89. Trogoderma angustum Solier, 1849 | IT | IT | SCO | ||
90. Trogoderma glabrum (Herbst, 1783) | N | S | Si? | SCO | |
91. Trogoderma granarium Everts, 1898 | N | S | Si | COS | |
92. Trogoderma inclusum LeConte, 1854 | N | S | Si | Sa | COS |
93. Trogoderma megatomoides Reitter, 1881 | N | OLA | |||
94. Trogoderma variabile Ballion, 1878 | N | Sa | COS | ||
95. Trogoderma versicolor (Creutzer, 1799) | N | S | Si | Sa | COS |
Total species | 61 | 67 | 57 | 59 |
The following abbreviations are used in the text: dint. = dintorni = environs; env. = environs; ex = specimen/s; FA = F. Angelini leg.; FEI = Forum Entomologi Italiani (http://www.entomologiitaliani.net); Fraz. = Frazione = hamlet; GC = G. Scaglioni leg.; GNP = Gargano National Park; leg. = legit or legerunt; PC = P. Cornacchia leg.; Prog. = Progetto = Project; prov. = province; sn = sweep net.
AHG Andreas Herrmann private collection, Stade, Germany
CNBFVR Centro Nazionale per lo Studio e la Conservazione della Biodiversità Forestale “Bosco Fontana” di Verona, Sede di Bosco Fontana. Marmirolo (Mantua), Italy
GNAC Gianluca Nardi private collection, Cisterna di Latina (Latina), Italy
HNHM Hungarian Natural History Museum, Budapest, Hungary
JHAC Private Entomological Laboratory and Collection, Jiří Háva, Prague-west, Czech Republic
MCZR Museo Civico di Zoologia, Rome, Italy
MZUF Museo di Storia Naturale, Sezione di Zoologia, Università di Firenze, Florence, Italy
PCOP Paolo Cornacchia private collection, Porto Mantovano (Mantua), Italy
http://species-id.net/wiki/Dermestes_peruvianus
[Latium: Roma province, ] Roma env., [no other data, ] 1 ex (JHAC).
Subcosmopolitan (
Italy? (
Species described from Peru (cf.
http://species-id.net/wiki/Dermestes_carnivorus
Tuscany: Firenze [= Florence], V.1948, [no collector, ] 1 ♀ (JHAC).
Cosmopolitan (cf.
Italy (without further details) (
http://species-id.net/wiki/Dermestes_vorax
None.
Asiatic (
Italy (without further details) (
http://species-id.net/wiki/Dermestes_hankae
Italy mer. [= southern Italy], [no other data], 1 ♂ (JHAC).
W-Mediterranean: Spain, France, Italy and Algeria (
Italy (without further details) (
Species described from southern France (
The specific name (hankae) is correct (
This species is very similar to Dermestes (Dermestinus) pardalis Billberg, 1808 (
http://species-id.net/wiki/Dermestes_intermedius_intermedius
None.
Turano-European reaching only as far as (Dermestes (Dermestinus) intermedius iranicus Háva & Kalík, 1999) Iran, Iraq and Syria eastward (cf.
Italy (without further details) (
The above generic Italian records by
http://species-id.net/wiki/Dermestes_szekessyi
None.
Sibero-European (cf.
Italy (without further details) (
The above generic Italian record of
http://species-id.net/wiki/Dermestes_olivieri
Apulia: Foggia prov., [GNP, ] Mattinata, strada per M.te Sacro [= road to Mount Sacro], Masseria Vaira dint., 613 m, 33T 584160 4821825, 24.V.2010, I. Toni leg., sn on Ferula communis [(Apiaceae)], Prog. Foresta Umbra, 1 ex (CNBFVR); Taranto prov., Ginosa Marina, VIII.1985, P. Abbazzi leg., 2 ex (MZUF). Tuscany: Arezzo [prov.], Cavriglia, [Fraz.] Montegonzi, 16.V.1991, Lisa leg., 1 ex (MZUF); [Arezzo prov., ] Ris. Nat. [= Riserva Naturale = Nature Reserve] “Valle Inferno-Bandella”, Casa Giardino, 8.V.1998, L. Bartolozzi, B. Cecchi A. & Sforzi leg., 1 ex (MZUF); [Firenze prov., ] Firenze [= Florence], V.1896, ex-coll. Beccari, 1 ex (MZUF); Firenze prov., Greve [in Chianti], [Fraz.] Strada in Chianti, 20.I.1957, A. Bandinelli leg., 1 ex (MZUF); [Grosseto prov.], Parco Naturale della [= Natural Park of] Maremma, Mti [= Monti = Mounts] dell’Uccellina, S. Rabano, X.1988, L. Bartolozzi leg., 4 ex (MZUF).
Turano-European with extension to Tunisia (cf.
All of Italy, Sicily and Sardinia (cf.
New collecting sites from Tuscany and Apulia. In Tuscany, the species was formally recorded from a sole locality (
http://species-id.net/wiki/Thorictus_pilosus
None.
Turanian-E-Mediterranean: Cyprus, Egypt, Greece, Iraq, Israel, Lebanon, Libya, Syria, Tunisia, Turkey and Uzbekistan (
“S?” [= peninsular Italy?] (
The above doubtful record for peninsular Italy has already been ignored by other authors (
http://species-id.net/wiki/Thorictus_wasmanni
None.
Turanian: endemic of Uzbekistan (
“Si?” [= Sicily?] (
The above doubtful Sicilian record has already been ignored by other authors (
http://species-id.net/wiki/Attagenus_calabricus
Abruzzi: Pescara prov., Maiella, Valle Orfento, Caramanico, 23.VI.1988, G. Osella leg., 1 ex (JHAC). Apulia: Brindisi prov., Francavilla Fontana, 30.V.1998, C. Esposito leg., 1 ex (GNAC); [Foggia prov., GNP, ] Mattinata, 22.VI.1991, on flowers on the beach, M. Mantic leg., 4 ex (JHAC); [Foggia prov., GNP, ] Gargano, Mattinata, 8–10.V.1995, Z. Svec leg., 1 ex (JHAC); [Foggia prov., GNP, Vico del Gargano, Fraz.] San Menaio, V.1996, M. Fikacek leg., 2 ex (JHAC); Taranto prov., Circ. [= Circum] Mar Piccolo, 24.V.1992, Montemurro leg., 1 ex (JHAC); Taranto prov., Grottaglie, VI.1369 [sic!], FA, 1 ex (MCSV). Basilicata: [Matera prov., ] Marina di Nova Siri, 4.VI.2008, W. Apfel leg., 1 ex (JHAC); [Matera prov., ] Matera, 4.VI.1989, FA, 1 ex (AHG); [Matera prov., ] Matera, 350 m, 4.VI.1989, FA, 11 ex (MCSV); Potenza prov., Francavilla sul Sinni, 320 m, 5.VI.1988, querceta [= oak wood], FA, 1 ex (MCSV); [Potenza or Matera prov., ] Pollino, 19.VI.1994, FA, 1 ex (AHG). Calabria: “Calabria / Baudi [leg.]” [handwritten by unknown], “Holotypus 1881 / Attagenus / Calabricus / Reitter” [red-framed; handwritten by unknown], “Calabricus m. / Calab. Baudi [leg.]” [handwritten probably by Reitter], “coll. Reitter” [printed], “Lectotype / Attagenus calabricus Reitter / J. Háva des. 2012”, 1 ex (HNHM); “Calabria / Baudi [leg.]” [handwritten by unknown], “Paratypus 1881 / Attagenus / Calabricus / Reitter” [red-framed; handwritten by unknown], “coll. Reitter [printed]”, “Paralectotype / Attagenus calabricus Reitter / J. Háva des. 2012”, 1 ex (HNHM); Calabria [without further data], 1 ex (JHAC); [Cosenza prov., Cassano all‘Ionio, Fraz.] Marina di Sibari env., 3 m, sandy dunes, 14.6.2009, P. Kresl leg., 6 ex (JHAC); [Cosenza prov., ] Roseto Capo Spulico, 27.V.–3.VI.2006, V. Hron leg., 1 ex (JHAC); [Cosenza prov., ] Sila, Mte. [= Mount] Oliveto, 2.VII.1929, C. Confalonieri leg., 1 ex (JHAC); [Cosenza prov., ] Sila, Sud [= South of] S. Giovanni [in] Fiore, 1100 m, 4.VII.[19]87, FA, 2 ex (MCSV); Reggio Calabria prov., Asprom. [= Aspromonte], str. [= strada = road] Cimina–Zomaro, 600–800 m, 6.VI.1994, FA, 6 ex (JHAC).
Apenninic endemic (
Campania (
The generic records for Italy (
First record from the Abruzzi region; this new site (cf.
http://species-id.net/wiki/Attagenus_lobatus
Tuscany: Firenze [= Florence], 28.IX.1988, L. Chelazzi leg., 1 ♂ (MZUF).
Centralasiatic-Mediterranean; this species was introduced to the USA (cf.
Latium, Campania, Apulia, Calabria, Sicily and Sardinia (cf.
First record for Tuscany. The new locality is the northernmost in Italy (cf.
Apulia: Bari prov., Gioia del Colle, 22.IV.1993, De Marzo leg., 1 ♂ (JHAC); Foggia prov., [GNP, ] Foresta Umbra, Monte Sant’Angelo, caserme forestali [= barracks of the Italian National Forestry Service], 792 m, 33T 582306 4630267, 27.IV.2010, PC, death on windowsill, Prog. Foresta Umbra, 1 ♀ (JHAC). Lombardy: Bergamo prov., Oltre il Colle, 7.VII.2005, FA, 1 ♂ (JHAC).
Alpino-Apenninic endemic (cf.
Apulia and Calabria (
Attagenus pellio var. pilosissimus from Apulia was listed as a synonym of Attagenus (Attagenus) pellio by
Attagenus (Attagenus) spp., males 1 Antennal club of Attagenus (Attagenus) pilosissimus from Italy (Apulia, Bari prov., Gioa del Colle) 2 Antennal club of Attagenus (Attagenus) pellio from Czech Republic (Central Bohemia, Prague) 3 Habitus of Attagenus (Attagenus) pilosissimus from Italy (Apulia, Bari prov., Gioia del Colle) (length 4.5 mm) 4 Abdomen in ventral view of Attagenus (Attagenus) pilosissimus from Italy (Apulia, Bari prov., Gioa del Colle) 5 Aedeagus (setation omitted) of Attagenus (Attagenus) pilosissimus from Italy (Lombardy, Bergamo prov., Oltre il Colle) (length 0.5 mm) 6 Aedeagus (setation omitted) of Attagenus (Attagenus) pellio from Czech Republic (Central Bohemia, Prague) (length 0.5 mm) (photos and drawings by J. Háva).
http://species-id.net/wiki/Attagenus_silvaticus
Apulia: Foggia prov., [GNP, ] Foresta Umbra, Vico del Gargano, Bosco Sfilzi, 391 m, 33T 585633 4634653, 23.V.2010, D. Birtele & I. Toni leg., window flight trap, Prog. Foresta Umbra, 1 ♀ (CNBFVR).
European (Czech Republic, Hungary, northwestern Russia, Sicily, Slovakia, Ukraine) with extension eastward to “Caucasus”, Anatolia, Iran and West Siberia (
Sicily (
First record for mainland Italy. The above specimen was intercepted in a beech wood (D. Birtele, pers. comm., 2012).
http://species-id.net/wiki/Attagenus_simonis
None.
E-Mediterranean: Egypt, Israel, Jordan and Syria (
Latium (
This species was recently excluded from the Italian fauna (
http://species-id.net/wiki/Attagenus_tigrinus_pulcher
None.
Mediterranean (cf.
Sardinia (
This subspecies is excluded here from the Italian fauna.
http://species-id.net/wiki/Anthrenus_coloratus
Apulia: [Foggia prov., GNP, ] Gargano, V.1991, J. Háva leg., 12 ex (JHAC).
Turano-Europeo-Mediterranean with extension to Canary Islands, Tajikistan, “India” and Sudan; this species was introduced to and is widespread in the USA (
Italy (
The generic Italian record by
http://species-id.net/wiki/Anthrenus_angustefasciatus
Friuli-Venezia Giulia: Trieste prov., Carso, [Sgonico, Fraz.] Gabrovizza S. Primo, 12.VI.1988, sn, PC, 1 ♀ (PCOP). Tuscany: Grosseto prov., Pitigliano, 23.I.[20]02, L. Saltini leg., 1 ex (GNAC). Apulia: [Brindisi prov., ] Brindisi env., 18.V.1997, J. Hrdlicka leg., 4 ex (JHAC). Basilicata: Potenza prov., [Pollino National Park, ] Terranova di Pollino, 900 m, 27–30.VI.2005, GC, sn, 1 ♀ (PCOP).
European with extension southward to a part of the Maghreb: Algeria, Bosnia and Herzegovina, Croatia, Czech Republic, France mainland, Italy, Morocco, Portugal, Spain, Switzerland, Yugoslavia and European Turkey (cf.
The records for Corsica (
Calabria (
First records for northern Italy (Friuli-Venezia Giulia), for central Italy (Tuscany), for Apulia and Basilicata; the former record is expected since this species occurs in neighbouring foreign regions (
http://species-id.net/wiki/Anthrenus_delicatus
Apulia: Foggia prov., [GNP, ] Promontorio del Gargano, Vieste, SP 89 [= Strada Provinciale n. 89 = Provincial Road n. 89], Casa Cupari dint., 354 m, 33T 5920001 4630150, 29.V.2010, abandoned quarry, sn on grasses, PC, Prog. Foresta Umbra, 6 ex (CNBFVR; JHAC); Foggia prov., [GNP, ] Promontorio del Gargano, Peschici, Lampia del Principe dint., 161 m, 33T 587282 4639346, 28.IV.2010, maquis sn, PC, Prog. Foresta Umbra, 1 ♀ (CNBFVR). Latium: Roma prov., M.ti [= Monti = Mounts] Simbruini, Jenne dint., M.te [= Monte = Mount] Porcaro, 800 m, 31.V.2007, sn, PC GC, 1 ex (PCOP). Sicily: Palermo prov., Corleone, Bosco della Ficuzza, 700 m, 1.VI.2008, sn near a watering trough, PC GC, 1 ex (PCOP).
Mediterranean, reaching eastward as far as Caucasus and Iran (cf.
Trentino, Venezia Giulia (
Moreover,
First detailed records from Apulia. About Sicily,
http://species-id.net/wiki/Anthrenus_munroi
Tuscany: [Livorno prov.], Is. [= Isola = Island] d’Elba, dint. Marina di Campo, 15–17.VI.1999, Abbazzi, Bartolozzi, Lo Cascio & Sforzi leg., 1 ♂ (MZUF).
Mediterranean (eastwards as far as Ukraine, Turkey, Syria, Lebanon, Jordan, and Israel), with extension to Hungary and Portugal (cf.
Apulia, Basilicata, Calabria (
First record for central Italy; this new site is the northernmost in Italy. The species also occurs in the facing Corsica (cf.
Anthrenus (Anthrenus) pimpinellae pimpinellae (Fabricius, 1775) and Anthrenus (Nathrenus) verbasci (Linnaeus, 1767) were the sole congeneric species previously recorded from Elba Island (
http://species-id.net/wiki/Anthrenus_pimpinellae_pimpinellae
Apulia: Foggia prov., [GNP, ] Promontorio del Gargano, Vieste, SP 89 [= Strada Provinciale n. 89 = Provincial Road n. 89], Casa Cupari dint., 354 m, 33T 5920001 4630150, abandoned quarry, sn on grasses, 29.V.2010, PC, Prog. Foresta Umbra, 4 ex (CNBFVR).
Cosmopolitan (cf.
All of Italy and Sicily (
Species common and widespread in Italy but from the Gargano National Park (cf.
http://species-id.net/wiki/Anthrenus_signatus
Basilicata: Potenza prov., Pignola, Ris. [= Riserva = Reserve] WWF, L. [= Lago = Lake] Pignola, 700 m, 21.VI.1992, FA, 1 ♂ (JHAC).
European (cf.
South Tyrol, Venetia, Venezia Giulia, Latium?, Apulia, Basilicata and Sicily (cf.
This species was recorded in Basilicata region from the Pollino National Park (
Anthrenus signatus ab. bielawskii Mroczkowski, 1958 from Bulgaria (
http://species-id.net/wiki/Globicornis_breviclavis
None.
Endemic to the Near East: Armenia, Bulgaria, “Caucasus”, Georgia, European Turkey, Ukraine, and Southern European Russia (Adygeya, Krasnodar kraj) (
Piedmont: Graian and Pennine Alps (
This species described from Caucasus (
http://species-id.net/wiki/Globicornis_fasciata
Basilicata: Potenza prov., Parco Naz. del Pollino [= Pollino National Park], Casa del Conte dint., tra [= between] Acqua Tremola e [= and] Piani di S. Francesco, 1100–1500 m, 9.VII.2002, sn, GC, 3 ex (PCOP).
European (Belgium, Denmark, France, Germany, Italy, Spain, Sweden, Switzerland) with extension southward to Tunisia (cf.
Piedmont (
The records for northern and peninsular Italy (
First record from southern Italy. Five species of this genus are now recorded from the Pollino National Park (cf.
The records for Piedmont (
In the literature (see also above), the authorship of this species was often attributed only to Fairmaire.
http://species-id.net/wiki/Globicornis_sulcata
Abruzzi: Parco Naz. [= National Park], L‘Aquila prov., Pescasseroli, 1300 m, 1.IV.2003, FA, 1 ♀ (JHAC). Basilicata: Potenza prov., Moliterno, 500 m, 11.V.2003, FA, 1 ♀ (JHAC); Potenza prov., Ruoti, 29.IX.1996, Abies–Quercus, FA, 1 ♂ (JHAC). Campania: Avellino prov., Cilento, Novi Velia, M. [= Mount] Sacro, 1900 m, 7.XII.1996, Fagus, FA, 1 ♂ (JHAC). Sicily: Palermo prov., Ficuzza, 700 m, 1–4.V.2000, bosco leccio [= ilex wood], FA, 1 ♂ (JHAC); Palermo prov., Bosco della Ficuzza, 27.V.2006, M. Sarovec leg., 1 ♂, 4 ♀♀ (JHAC).
W-Mediterranean: France, Italy and Spain.
Basilicata (
First records from central Italy (Abruzzi), Campania and Sicily. In the Sicilian site, it was also collected Anthrenus (Anthrenus) delicatus (see above).
http://species-id.net/wiki/Trogoderma_angustum
None.
Subcosmopolitan (cf.
Italy (without further details) (
The above generic Italian record by
http://species-id.net/wiki/Trogoderma_inclusum
Apulia: Lecce prov., Salento, Torre S. Giovanni dint., 1–7.VII.2002, PC GC, sn, 1 ex (PCOP). Venetia: Verona prov., Verona, [Fraz.] Cancello, [about 500 m], 22.VI.1986, sn, PC, 1 ex (PCOP).
Cosmopolitan (cf.
Lombardy (
The first Italian record of this species was provided by
In Italy, this species and Trogoderma versicolor (Creutzer, 1799) have often been confused (cf.
The old Sicilian records of Attagenus (Attagenus) trifasciatus (Fabricius, 1787) and Trogoderma glabrum (Herbst, 1783) (
Thorictus mauritanicus Lucas, 1846 and Thorictus stricticollis Kraatz, 1859 were both recorded from “Italy” by
Thorictus sicilianus John, 1965 (= Thorictus sicilianus John, 1966) from Sicily and Morocco (
Dermestes (Dermestinus) gyllenhalii gyllenhalii Laporte de Castelnau, 1840
Dermestes ? atomarius Erichson, 1846:
Si? (
Dermestes (Dermestinus) laniarius laniarius Illiger, 1801
Dermestes laniarius Illig.:
Dermestes laniarius Illiger, 1801:
Si (
Thorictus grandicollis grandicollis Germar, 1842
Thorictus grandicollis Germ.:
N (
Attagenus (Attagenus) cyphonoides Reitter, 1881
Attagenus alfierii Pic[, 1910]:
Attagenus (Attagenus) cyphonoides Reitter, 1881:
Attagenus (Attagenus) cyphonoides Reitter, 1881:
S (
Attagenus (Attagenus) fasciatus (Thunberg, 1795)
Attagenus fasciatus (Thunberg, 1795):
Attagenus (Attagenus) fasciatus (Thunberg, 1795):
N (
Attagenus (Attagenus) pantherinus (Ahrens, 1814)
Attagenus pantherinus:
Attagenus (Attagenus) pantherinus (Ahrens, 1814):
Attagenus pantherinus (Ahrens, 1814):
N (
Attagenus (Attagenus) rossii Ganglbauer, 1904
Dermestes bifasciatus Rossi, 1794: 79.
Attagenus bifasciatus Rossi:
Attagenus bifasciatus:
Attagenus (Lanorus) bifasciatus Rossi:
Attagenus (Lanorus) bifasciatus Oliv. [= (A.G. Olivier, 1790)] a. Rossii Ganglb.:
Attagenus (Lanorus) bifasciatus Oliv. v. Rossii Ganglb.:
Attagenus bifasciatus Oliv. var. Rossii Ganglb.:
Attagenus bifasciatus (Olivier, 1790) var. Rossii Ganglbauer, 1904:
Attagenus (Attagenus) rossii Ganglbauer, 1904:
Attagenus rossii Ganglbauer, 1904:
Attagenus rossii Ganglbauer, 1804 [sic!]: FEI 2012.
S (
Attagenus (Attagenus) trifasciatus (Fabricius, 1787)
Attagenus trifasciatus Fabr.:
Attagenus ? trifasciatus (Fabricius, 1787):
Si? (
Anthrenocerus australis (Hope, 1843)
Anthrenocerus australis (Hope, 1843): Herrmann and
N (Herrmann and
Globicornis (Globicornis) luckowi Herrmann, Háva & Kadej, 2011
Globicornis (Globicornis) luckowi Herrmann, Háva & Kadej, 2011:
N (
Globicornis (Hadrotoma) corticalis (Eichhoff, 1863)
Globicornis corticalis (Eichhoff, 1863):
S (
Reesa vespulae (Milliron, 1939)
Reesa vespulae (Milliron, 1939):
N (
Trogoderma glabrum (Herbst, 1783)
Trogoderma elongatulum Fabr. [= (Fabricius, 1801)]:
Si? (
Trogoderma variabile Ballion, 1878
Trogoderma variabile Ballion, 1878:
Trogoderma variabile Ballion:
N (
The present-day dermestid fauna of Italy is summarized in the checklist below (Table 1).
The following abbreviations are used: AFR = Afrotropical; ALAP = Alpino-Apenninic endemic; ALPC = Central-Alpine endemic; APPE = Apenninic endemic; AUST = Australian; CAE = Centralasiatic-European; CEM = Centralasiatic-Europeo-Mediterranean; CEU = Central European; COS = Cosmopolitan; EME = E-Mediterranean; EUM = Europeo-Mediterranean; EUR = European; IT = Italy and/or Italian mainland without further details; (i) = introduced to Europe; MED = Mediterranean; N = northern Italy (Trentino-Alto Adige, Venetia, Friuli-Venezia Giulia, Lombardy, Piedmont, Aosta Valley, Liguria and Emilia-Romagna); NAF = N-African; OLA = Holarctic; PAL = Palearctic; S = peninsular Italy (the remainig of continental Italy); Sa = Sardinia and small circum-Sardinian islands; SCO = Subcosmopolitan; SEU = S-European; Si = Sicily and small circum-Sicilian islands; SICI = Sicilian endemic; SIE = Sibero-European; TEM = Turano-Europeo-Mediterranean; TUE = Turano-European; TUM = Turano-Mediterranean; WME = W-Mediterranean.
The updated checklist of Italian Dermestidae includes 95 species, moreover Anthrenus (Anthrenus) pimpinellae occurs with two subspecies, but the taxonomic status of Anthrenus (Anthrenus) pimpinellae isabellinus needs to be clarified (cf.
Some of the above records come from the following two protected areas of southern peninsular Italy: the Gargano National Park (Apulia) and the Pollino National Park (Basilicata and Calabria). 24 species of Dermestidae are now known from the former Park, while 23 species are now known from the Pollino National Park, but only 15 species are common to both (cf.
We are indebted to Claudio Angeloro (Corpo Forestale dello Stato - Ufficio Territoriale per la Biodiversità di Foresta Umbra, Italy) for the assistance in Apulia during the research of the CNBFVR; Daniele Birtele (CNBFVR), Paolo Cornacchia (Porto Mantovano, Italy), Carmine Esposito (Velletri, Italy), Gianluca Scaglioni (Porto Mantovano, Italy) and Ilaria Toni (CNBFVR) for the collection of specimens; to Luca Bartolozzi (Museo di Storia Naturale, Sezione di Zoologia, Università di Firenze, Florence, Italy) for the loan of specimens from the collections under his care; to Franco Mason (Verona, Italy) for his constructive comments on a previous version of this paper; to Ottó Merkl (Hungarian Natural History Museum, Budapest, Hungary) for the information on the types of Attagenus calabricus; to Miloslav Rakovič (Prague, Czech Republic) and to Jonathan Cooter (Oxford, England) for the linguistic revision of the manuscript. We thank Riaan Stals (Pretoria, South Africa) and an anonymous referee for their reviews.