ZooKeys 360: 45–81, doi: 10.3897/zookeys.360.6023
Italian Dermestidae: notes on some species and an updated checklist (Coleoptera)
Gianluca Nardi 1, Jiří Háva 2
1 MiPAAF, Corpo Forestale dello Stato, Centro Nazionale per lo Studio e la Conservazione della Biodiversità Forestale “Bosco Fontana” di Verona, Sede di Bosco Fontana, Strada Mantova 29, I-46045 Marmirolo (MN), Italy
2 Department of Forest Protection and Entomology, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamýcká 1176, CZ-165 21, Prague 6 - Suchdol, Czech Republic

Corresponding author: Gianluca Nardi (l_nardi@hotmail.com)

Academic editor: C. Majka

received 20 July 2013 | accepted 29 November 2013 | Published 6 December 2013


(C) 2013 Gianluca Nardi. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.


For reference, use of the paginated PDF or printed version of this article is recommended.

Citation: Nardi G, Háva J (2013) Italian Dermestidae: notes on some species and an updated checklist (Coleoptera). ZooKeys 360: 45–81. doi: 10.3897/zookeys.360.6023

Abstract

An up-to-date checklist of the Italian Dermestidae is provided. The presence of 95 species in Italy is confirmed, while further 5 species (Dermestes (Dermestes) vorax Motschulsky, 1860, Thorictuspilosus Peyron, 1857, T. wasmanni Reitter, 1895, Attagenus (Attagenus) simonis Reitter, 1881 and Globicornis (G.) breviclavis (Reitter, 1878)) and 1 subspecies (A. (A.) tigrinus pulcher Faldermann, 1835) are excluded from the Italian fauna.

Attagenus (Attagenus) calabricus Reitter, 1881 and A. (A.) lobatus Rosenhauer, 1856 are for the first time recorded from Abruzzi and Tuscany respectively; A. (A.) silvaticus Zhantiev, 1976 is recorded for the first time from mainland Italy (Apulia); Anthrenus (Anthrenus) angustefasciatus Ganglbauer, 1904 is new to northern Italy (Friuli-Venezia Giulia), central Italy (Tuscany), Apulia and Basilicata; A. (A.) munroi Hinton, 1943 is new to central Italy (Elba Island); A. (A.) delicatus Kiesenwetter, 1851 is for the first time recorded from Apulia; Globicornis (Globicornis) fasciata (Fairmaire & Brisout de Barneville, 1859) is new to southern Italy (Basilicata); G. (Hadrotoma) sulcata (C.N.F. Brisout de Barneville, 1866) is for the first time recorded from central Italy (Abruzzi), Campania and Sicily, whileTrogoderma inclusum LeConte, 1854 is new to Apulia.

Seven species (Dermestes (Dermestes) peruvianus Laporte de Castelnau, 1840, D. (Dermestinus) carnivorus Fabricius, 1775, D. (Dermestinus) hankae Háva, 1999, D. (Dermestinus) intermedius intermedius Kalík, 1951, D. (Dermestinus) szekessyi Kalík, 1950, Anthrenus (Anthrenops) coloratus Reitter, 1881 and Trogodermaangustum (Solier, 1849)) recently recorded from Italy (without further details) are discussed.

The lectotype and a paralectotype are designated forAttagenus (A.) calabricus Reitter, 1881 from Calabria.

Attagenus pellio (Linnaeus, 1758) var. pilosissimus Roubal, 1932 is removed from synonymy with A. (A.) pellio and recognized as a valid species (stat. prom.); it is known from Lombardy, Apulia and Calabria.

Keywords

Dermestidae, lectotype designation, new status, taxonomy, distribution, new records, chorotypes, checklist, Italy

Introduction

The main purpose of the present paper is to enhance knowledge of the Dermestidae of Italy. It is chiefly based on material collected by the staff of the Centro Nazionale per lo Studio e la Conservazione della Biodiversità Forestale “Bosco Fontana” di Verona (Italy), during entomological surveys in various Italian Protected Areas (e.g. Cerretti et al. 2003, Mason et al. 2006, Nardi and Vomero 2007, Cerretti et al. 2009, Nardi et al. 2011). Moreover, this paper gives an opportunity to provide a new checklist of the Dermestidae of Italy: it updates the nomenclature, systematics and the faunistic of the previous (Audisio et al. 1995) chiefly according to recent capital works (Háva 2007, Zhantiev 2011a, Háva 2011a).

Material and methods

This paper is organized in three sections: “Notes on some species”, “Literature records” and “A new checklist of the Dermestidae of Italy”.

The classification of the subfamilies, tribes, genera and subgenera follows that proposed by Háva (2004, 2007); species are listed alphabetically according to the nomenclature of Háva (2007), Kadej et al. (2007), Háva (2008), Zhantiev (2009) and Háva (2011a).

The first section is based chiefly on study of new material. With very few exceptions, identifications were made by J. Háva.

The second section summarizes the literature data updating the Italian distribution of the species provided by Audisio et al. (1995) or by Háva and Nardi (2011).

In both sections, nomenclatural combinations and possible synonyms (listed chronologically-alphabetically) found in the literature that includes Italian records are provided below the valid name of each species or subspecies.

The first section concerns 25 species. As far as possible, the following data are provided for each record: region, province, general area and/or commune, locality of collection, altitude, UTM coordinates, date of collection, collector/s, collecting method, possible research project, number of specimens, and, in parentheses, abbreviation of depository. The labels of the examined specimens are generally written in Italian; hereunder, the regions and the collecting methods were translated into English. The label data of type specimens are cited verbatim between quotation marks, a forward slash separating the different lines on a label, and a comma separating subsequent labels. Comments and interpretations are given in square brackets.

The mainland Italian regions are listed from north to south, and from west to east, all toponyms are listed alphabetically. Chorotypes were assigned according to Vigna Taglianti et al. (1993, 1999) and based on the distributions provided by Háva (2003, 2007, 2011b), Zhantiev (2009, 2011a, 2011b) and Háva et al. (2013), unless otherwise stated.

In the second section, the literature records updating the checklist of Dermestidae of Italy provided by Audisio et al. (1995) or by Háva and Nardi (2011) are detailed. These new faunistic data are based on papers subsequent to these two papers but also include some previously overlooked publications. These records concern 14 species. The data on distribution are listed according to the geographic divisions by Audisio et al. (1995): northern Italy (N), peninsular Italy (S), Sardinia and small circum-Sardinian islands (Sa) and Sicily and small circum-Sicilian islands (Si).

Finally, a new checklist of the Italian Dermestidae is provided (Table 1). The total number of species for each geographic division includes only those recorded with certain, so generic records for “Italy” and records followed by a question mark are excluded.

Acronyms for chorotypes follow Vigna Taglianti et al. (1993, 1999), while those for the geographic divisions of Italy follow Audisio et al. (1995); both are included in the foreword of the checklist.

Table 1.

Checklist of the Dermestidae of Italy.

Taxa Distribution Chorotype
Dermestinae Latreille, 1804
Dermestini Latreille, 1804
Genus Dermestes Linnaeus, 1758
Subgenus Dermestes Linnaeus, 1758
1. Dermestes (Dermestes) ater DeGeer, 1774 N COS
2. Dermestes (Dermestes) bicolor Fabricius, 1781 PAL
Dermestes (Dermestes) bicolor bicolor Fabricius, 1781 N S Sa
3. Dermestes (Dermestes) haemorrhoidalis Küster, 1852 N COS
4. Dermestes (Dermestes) hispanicus Kalík, 1952 Sa WME
5. Dermestes (Dermestes) lardarius Linnaeus, 1758 N S Si Sa COS
6. Dermestes (Dermestes) peruvianus Laporte de Castelnau, 1840 N? S COS
Subgenus Dermestinus Zhantiev, 1967
7. Dermestes (Dermestinus) aurichalceus Küster, 1846 N S? Sa MED
8. Dermestes (Dermestinus) carnivorus Fabricius, 1775 S COS
9. Dermestes (Dermestinus) erichsoni Ganglbauer, 1904 N S Si Sa EUR
10. Dermestes (Dermestinus) frischii Kugelann, 1792 N S Si Sa COS
11. Dermestes (Dermestinus) gyllenhalii Laporte de Castelnau, 1840 EUR
Dermestes (Dermestinus) gyllenhalii gyllenhalii Laporte de Castelnau, 1840 N S Si?
12. Dermestes (Dermestinus) hankae Háva, 1999 S WME
13. Dermestes (Dermestinus) intermedius Kalík, 1951 EUR
Dermestes (Dermestinus) intermedius intermedius Kalík, 1951 IT IT
14. Dermestes (Dermestinus) laniarius Illiger, 1801 PAL
Dermestes (Dermestinus) laniarius laniarius Illiger, 1801 N S Si Sa
15. Dermestes (Dermestinus) maculatus DeGeer, 1774 N S Si Sa COS
16. Dermestes (Dermestinus) murinus Linnaeus, 1758 PAL
Dermestes (Dermestinus) murinus murinus Linnaeus, 1758 N S Si Sa
17. Dermestes (Dermestinus) mustelinus Erichson, 1846 N S Si Sa EUR
18. Dermestes (Dermestinus) pardalis Billberg, 1808 N? IT Sa WME
19. Dermestes (Dermestinus) sardous Küster, 1846 MED
Dermestes (Dermestinus) sardous sardous Küster, 1846 N S Sa
20. Dermestes (Dermestinus) szekessyi Kalík, 1950 IT IT SIE
21. Dermestes (Dermestinus) undulatus Brahm, 1790 N S Si Sa OLA
Subgenus Montandonia Jacquet, 1886
22. Dermestes (Montandonia) fuliginosus P. Rossi, 1792 N S Si EUR
23. Dermestes (Montandonia) hirticollis Fabricius, 1792 Sa WME
24. Dermestes (Montandonia) olivieri Lepesme, 1939 N S Si Sa TEM
Thorictinae Agassiz, 1846
Thaumaphrastini Anderson, 1949
Genus Thorictodes Reitter, 1875
25. Thorictodes heydeni Reitter, 1875 N S COS
Thorictini Agassiz, 1846
Genus Thorictus Germar, 1834
26. Thorictus grandicollis Germar, 1842 TUM
Thorictus grandicollis grandicollis Germar, 1842 N S Si Sa
27. Thorictus mauritanicus Lucas, 1846 Si Sa MED
28. Thorictus stricticollis Kraatz, 1859 MED
Thorictus stricticollis stricticollis Kraatz, 1859 Sa
29. Thorictus studti John, 1971 Si SICI
Orphilinae LeConte, 1900
Genus Orphilus Erichson, 1848
30. Orphilus niger (P. Rossi, 1790) N S Si Sa CEM
Trinodinae T.L. Casey, 1900
Thylodriini Semenov, 1909
Genus Thylodrias Motschulsky, 1839
31. Thylodrias contractus Motschulsky, 1839 N COS
Trinodini T.L. Casey, 1900
Genus Trinodes Dejean, 1821
32. Trinodes hirtus (Fabricius, 1781) N S Si Sa TUE
Attageninae Laporte, 1840
Attagenini Laporte, 1840
Genus Attagenus Latreille, 1802
Subgenus Attagenus Latreille, 1802
33. Attagenus (Attagenus) bifasciatus (A.G. Olivier, 1790) S Si Sa TUM
34. Attagenus (Attagenus) brunneus Faldermann, 1835 N S Si Sa OLA
35. Attagenus (Attagenus) calabricus Reitter, 1881 S Si APPE
36. Attagenus (Attagenus) cyphonoides Reitter, 1881 S SCO
37. Attagenus (Attagenus) fallax Gené, 1839 S Si Sa MED
38. Attagenus (Attagenus) fasciatus (Thunberg, 1795) N S COS
39. Attagenus (Attagenus) lobatus Rosenhauer, 1856 S Si Sa CEM
40. Attagenus (Attagenus) maritimus (Gené, 1839) Si Sa WME
41. Attagenus (Attagenus) obtusus (Gyllenhal, 1808) Sa TEM
42. Attagenus (Attagenus) pantherinus (Ahrens, 1814) N Sa EUR
43. Attagenus (Attagenus) pellio (Linnaeus, 1758) N S Si Sa COS
44. Attagenus (Attagenus) pilosissimus Roubal, 1932 N S ALAP
45. Attagenus (Attagenus) punctatus (Scopoli, 1772) N S Si EUR
46. Attagenus (Attagenus) rossii Ganglbauer, 1904 S Si Sa MED
47. Attagenus (Attagenus) schaefferi (Herbst, 1792) OLA
Attagenus (Attagenus) schaefferi schaefferi (Herbst, 1792) N S Si Sa
48. Attagenus (Attagenus) silvaticus Zhantiev, 1976 S Si SIE
49. Attagenus (Attagenus) simplex Reitter, 1881 S Si Sa MED
50. Attagenus (Attagenus) tigrinus (Fabricius, 1792)
Attagenus (Attagenus) tigrinus tigrinus (Fabricius, 1792) N S Si Sa MED
51. Attagenus (Attagenus) trifasciatus (Fabricius, 1787) N Si? MED
52. Attagenus (Attagenus) unicolor (Brahm, 1790) COS
Attagenus (Attagenus) unicolor unicolor (Brahm, 1790) N S Si Sa
53. Attagenus (Attagenus) uniformis Fairmaire, 1860 Si NAF
Megatominae Leach, 1815
Anthrenini Gistel, 1848
Genus Anthrenus Geoffroy, 1762
Subgenus Anthrenops Reitter, 1881
54. Anthrenus (Anthrenops) coloratus Reitter, 1881 S OLA+AFR
Subgenus Anthrenus Geoffroy, 1762
55. Anthrenus (Anthrenus) angustefasciatus Ganglbauer, 1904 N S EUR
56. Anthrenus (Anthrenus) delicatus Kiesenwetter, 1851 N S Si Sa MED
57. Anthrenus (Anthrenus) festivus Erichson, 1846 S? Si Sa MED
58. Anthrenus (Anthrenus) flavipes LeConte, 1854 COS
Anthrenus (Anthrenus) flavipes flavipes LeConte, 1854 S Sa
59. Anthrenus (Anthrenus) goliath Saulcy, 1868 N S Si Sa MED
60. Anthrenus (Anthrenus) munroi Hinton, 1943 S Sa MED
61. Anthrenus (Anthrenus) mroczkowskii Kalík, 1954 S SEU
62. Anthrenus (Anthrenus) pimpinellae (Fabricius, 1775) COS
Anthrenus (Anthrenus) pimpinellae isabellinus Küster, 1848 N S Si Sa
Anthrenus (Anthrenus) pimpinellae pimpinellae (Fabricius, 1775) N S Si
63. Anthrenus (Anthrenus) scrophulariae (Linnaeus, 1758) COS
Anthrenus (Anthrenus) scrophulariae scrophulariae (Linnaeus, 1758) N S Si Sa
Subgenus Florilinus Mulsant & Rey, 1868
64. Anthrenus (Florilinus) museorum (Linnaeus, 1761) N S Si Sa COS
65. Anthrenus (Florilinus) oberthueri Reitter, 1881 N Si WME
Subgenus Helocerus Mulsant & Rey, 1868
66. Anthrenus (Helocerus) fuscus A.G. Olivier, 1790 N S Si Sa SCO
67. Anthrenus (Helocerus) minutus Erichson, 1846 S Si Sa MED
Subgenus Nathrenus Casey, 1900
68. Anthrenus (Nathrenus) biskrensis Reitter, 1887 Si NAF
69. Anthrenus (Nathrenus) molitor Aubé, 1850 N Sa MED
70. Anthrenus (Nathrenus) signatus Erichson, 1846 N S Si EUR
71. Anthrenus (Nathrenus) verbasci (Linnaeus, 1767) N S Si Sa COS
72. Anthrenus (Nathrenus) versicolor Reitter, 1887 Si Sa MED
Megatomini Leach, 1815
Genus Anthrenocerus Arrow, 1915
73. Anthrenocerus australis (Hope, 1843) N AUST (i)
Genus Ctesias Stephens, 1830
Subgenus Ctesias Stephens, 1830
74. Ctesias (Ctesias) serra (Fabricius, 1792) N S Si Sa EUR
Genus Globicornis Latreille, 1829
Subgenus Globicornis Latreille, 1829
75. Globicornis (Globicornis) bifasciata (Perris, 1866) Si Sa WME
76. Globicornis (Globicornis) fasciata (Fairmaire & Brisout de Barneville, 1859) N S Si? Sa CEU
77. Globicornis (Globicornis) luckowi Herrmann, Háva & Kadej, 2011 N ALPC
78. Globicornis (Globicornis) nigripes (Fabricius, 1792) N S EUR
79. Globicornis (Globicornis) picta (Küster, 1851) S Si Sa EUR
80. Globicornis (Globicornis) tristis (Reitter, 1881) S EME
81. Globicornis (Globicornis) variegata (Küster, 1851) N S Si Sa SEU
Subgenus Hadrotoma Erichson, 1848
82. Globicornis (Hadrotoma) corticalis (Eichhoff, 1863) N S Si EUR
83. Globicornis (Hadrotoma) emarginata (Gyllenhal, 1808) N S Si Sa EUR
84. Globicornis (Hadrotoma) sulcata (C.N.F. Brisout de Barneville, 1866) S Si WME
Genus Megatoma Herbst, 1791
Subgenus Megatoma Herbst, 1791
85. Megatoma (Megatoma) ruficornis Aubé, 1866 S SEU
86. Megatoma (Megatoma) undata (Linnaeus, 1758) N S EUR
Genus Phradonoma Jacquelin du Val, 1859
87. Phradonoma villosulum (C. Duftschmid, 1825) N CAE
Genus Reesa Beal, 1967
88. Reesa vespulae (Milliron, 1939) N SCO
Genus Trogoderma Dejean, 1821
89. Trogoderma angustum Solier, 1849 IT IT SCO
90. Trogoderma glabrum (Herbst, 1783) N S Si? SCO
91. Trogoderma granarium Everts, 1898 N S Si COS
92. Trogoderma inclusum LeConte, 1854 N S Si Sa COS
93. Trogoderma megatomoides Reitter, 1881 N OLA
94. Trogoderma variabile Ballion, 1878 N Sa COS
95. Trogoderma versicolor (Creutzer, 1799) N S Si Sa COS
Total species 61 67 57 59

The following abbreviations are used in the text: dint. = dintorni = environs; env. = environs; ex = specimen/s; FA = F. Angelini leg.; FEI = Forum Entomologi Italiani (http://www.entomologiitaliani.net); Fraz. = Frazione = hamlet; GC = G. Scaglioni leg.; GNP = Gargano National Park; leg. = legit or legerunt; PC = P. Cornacchia leg.; Prog. = Progetto = Project; prov. = province; sn = sweep net.

Acronyms of specimen depositories

AHG Andreas Herrmann private collection, Stade, Germany

CNBFVR Centro Nazionale per lo Studio e la Conservazione della Biodiversità Forestale “Bosco Fontana” di Verona, Sede di Bosco Fontana. Marmirolo (Mantua), Italy

GNAC Gianluca Nardi private collection, Cisterna di Latina (Latina), Italy

HNHM Hungarian Natural History Museum, Budapest, Hungary

JHAC Private Entomological Laboratory and Collection, Jiří Háva, Prague-west, Czech Republic

MCZR Museo Civico di Zoologia, Rome, Italy

MZUF Museo di Storia Naturale, Sezione di Zoologia, Università di Firenze, Florence, Italy

PCOP Paolo Cornacchia private collection, Porto Mantovano (Mantua), Italy

Notes on some species
Dermestes (Dermestes) peruvianus Laporte de Castelnau, 1840

http://species-id.net/wiki/Dermestes_peruvianus

Dermestes peruvianus Laport.: Bertolini 1904: 58.
Dermestes peruvianus Casteln.: Luigioni 1929: 1024; Porta 1929: 301.
Dermestes (Dermestes) peruvianus Castelnau, 1840: Plata-Negrache 1972: 150; Zhantiev 2011a.
Dermestes (Dermestes) peruvianus Laporte de Castelnau, 1840: Audisio et al. 1995: 12, 16; Háva 2007: 300.
Dermestes peruvianus Laporte de Castelnau: Pollini 1998: 909.
Material examined.

[Latium: Roma province, ] Roma env., [no other data, ] 1 ex (JHAC).

Chorotype.

Subcosmopolitan (Háva 2007). This species is also recorded from Portugal (Grosso-Silva and Serrano 2000) and Switzerland (Kenis 2005, Zhantiev 2011a). Both countries must be added to the European distribution summarized by Háva (2007), while the former country must also be added to those provided by Zhantiev (2011a).

Italian distribution.

Italy? (Bertolini 1904, Luigioni 1929, Porta 1929). Italian mainland? (Audisio et al. 1995). Italy (without further details) (Plata-Negrache 1972, Háva 2007). Italian mainland (Zhantiev 2011a).

Remarks.

Species described from Peru (cf. Háva and Kalík 2005); the first precise record from Italy is shown above.

Domenichini et al. (1993: 272) wrote that this species is common in food factories and was reared experimentally in an insectarium in Piacenza University (northern Italy), but the place of origin of this captive population is not mentioned. This species, and Dermestes (Dermestinus) carnivorus Fabricius, 1775 (see below), are also considered in an Italian manual of applied entomology (Pollini 1998), but no records from Italy are listed therein.

Dermestes (Dermestinus) carnivorus Fabricius, 1775

http://species-id.net/wiki/Dermestes_carnivorus

Dermestes carnivorus Fabricius: Pollini 1998: 906.
Dermestes (Dermestinus) carnivorus Fabricius, 1775: Háva 2007: 300; Zhantiev 2011a.
Material examined.

Tuscany: Firenze [= Florence], V.1948, [no collector, ] 1 ♀ (JHAC).

Chorotype.

Cosmopolitan (cf. Háva 2007).

Italian distribution.

Italy (without further details) (Háva 2007). Italian mainland (Zhantiev 2011a).

Remarks.

Pollini (1998: 906) wrote about this species that si è acclimatato in Europa e in India[= it is naturalized in Europe and India], but no Italian locality was mentioned. The above generic Italian record by Háva (2007), was based on the above specimen.

Dermestes (Dermestes) vorax Motschulsky, 1860

http://species-id.net/wiki/Dermestes_vorax

Dermestes vorax Motschulsky, 1860: Denux and Zagatti 2010: 357.
Dermestes (Dermestes) vorax Motschulsky, 1860: Háva and Nardi 2011: 408.
Material examined.

None.

Chorotype.

Asiatic (Háva 2003, 2007). This species was introduced to the coastal Croatia (Depoli 1928, Luigioni 1929, Porta 1934, all as Dermestes vorax Motsch.), but this record was later ignored (Mroczkowski 1968, Háva 2003, 2007, Zhantiev 2011a).

Italian distribution.

Italy (without further details) (Denux and Zagatti 2010).

Remarks.

Denux and Zagatti (2010) wrote that an Italian record of this species was provided by Lohse (1979) and Háva (2003), but in these papers this species is not recorded from this country. Moreover, a recent record from “Italy” (http://www.dermestidae.wz.cz/main.html; http://www.dermestidae.com/index.html) is erroneous since concerns a dark specimen of Dermestes (Dermestes) lardarius Linnaeus, 1758 (Hava, unpublished data), so it was ignored by Háva and Nardi (2011).

Dermestes (Dermestinus) hankae Háva, 1999

http://species-id.net/wiki/Dermestes_hankae

Dermestes (Dermestinus) hankae Háva, 1999: Háva 2007: 301.
Dermestes (Dermestinus) hankai [sic!] Háva, 1999: Zhantiev 2011a.
Material examined.

Italy mer. [= southern Italy], [no other data], 1 ♂ (JHAC).

Chorotype.

W-Mediterranean: Spain, France, Italy and Algeria (Háva 2007, Prieto and Háva 2013).

Italian distribution.

Italy (without further details) (Háva 2007). Italian mainland (Zhantiev 2011a).

Remarks.

Species described from southern France (Háva 1999: 143); the generic record for Italy (Háva 2007) was based on the above specimen.

The specific name (hankae) is correct (ICZN 1999, art. 31.1.2), so the name hankai is only a subsequent incorrect spelling.

This species is very similar to Dermestes (Dermestinus) pardalis Billberg, 1808 (Háva 1999, Herrmann and Bahillo de la Puebla 2003, Prieto and Háva 2013); the re-examination of specimens attributed to it could provide additional records of Dermestes (Dermestinus) hankae. In Italy, Dermestes (Dermestinus) pardalis is recorded from Sardinia (cf. Háva and Nardi 2011), generically from mainland Italy (Zhantiev 2011a), and from a non-specified locality of the northern mainland (Giachino 1982: 360, as Dermestes thoracicus Dejean[, 1821 (nomen nudum)]), but this record, based on very old specimens, should be confirmed.

Dermestes (Dermestinus) intermedius intermedius Kalík, 1951

http://species-id.net/wiki/Dermestes_intermedius_intermedius

Dermestes (Dermestinus) intermedius intermedius Kalík, 1951: Háva 2003: 18; Háva 2007: 301.
Dermestes (Dermestinus) intermedius Kalík, 1951: Zhantiev 2011a.
Material examined.

None.

Chorotype.

Turano-European reaching only as far as (Dermestes (Dermestinus) intermedius iranicus Háva & Kalík, 1999) Iran, Iraq and Syria eastward (cf. Háva 2007, Háva et al. 2011).

Italian distribution.

Italy (without further details) (Háva 2003, 2007). Italian mainland (Zhantiev 2011a).

Remarks.

The above generic Italian records by Háva (2003, 2007) were based on a personal communication of V. Kalík, who identified an Italian specimen; unfortunately, no further details are available (Háva, unpublished data).

Dermestes (Dermestinus) szekessyi Kalík, 1950

http://species-id.net/wiki/Dermestes_szekessyi

Dermestes (Dermestinus) szekessyi Kalík, 1950: Háva 2007: 302; Zhantiev 2011a.
Material examined.

None.

Chorotype.

Sibero-European (cf. Háva 2003, 2007, Háva et al. 2011, Zhantiev 2011a).

Italian distribution.

Italy (without further details) (Háva 2007). Italian mainland (Zhantiev 2011a).

Remarks.

The above generic Italian record of Háva (2007) was based on a personal communication of V. Kalík, who identified an Italian specimen; unfortunately, no further details are available (Háva, unpublished data).

Dermestes (Montandonia) olivieri Lepesme, 1939

http://species-id.net/wiki/Dermestes_olivieri

Dermestes ater Oliv. [= Olivier, 1790]: Ghiliani 1842: 33; Ghiliani 1847: 93; Reiche 1860: 720; Ghiliani 1887: 310; Bertolini 1890: 196; Iehl 1909: 26; Luigioni 1929: 537; Porta 1929: 301; Gridelli 1949: 176; Horion 1955: 197; Faggioli 1956: 174; Gulli 1961: 4.
Dermestes ater Ol.: Villa and Villa 1844: 38 [444]; Rottenberg 1870: 238; Steck 1887: 181; Ragusa 1892: 203; Bertolini 1904: 58; Vitale 1904: 75; Holdhaus 1911: 445; Panganetti-Hummler 1918: 87; Zangheri 1969: 1334.
Dermestes ater Olivier: Giachino 1982: 361.
Dermestes olivieri Lep.: Angelini 1986: 74; Angelini and Montemurro 1986: 573; Angelini 1987: 38; Angelini 1991: 206.
Dermestes (Dermestes) olivieri Lepesme, 1939: Audisio et al. 1995: 12; Bordoni et al. 2006: 60; FEI 2011.
Dermestes ater Degeer, 1774 [sic!]: Sapuppo 2002: 189.
Dermestes (Montandonia) olivieri Lepesme, 1939: Háva and Nardi 2011: 416.
Material examined.

Apulia: Foggia prov., [GNP, ] Mattinata, strada per M.te Sacro [= road to Mount Sacro], Masseria Vaira dint., 613 m, 33T 584160 4821825, 24.V.2010, I. Toni leg., sn on Ferula communis [(Apiaceae)], Prog. Foresta Umbra, 1 ex (CNBFVR); Taranto prov., Ginosa Marina, VIII.1985, P. Abbazzi leg., 2 ex (MZUF). Tuscany: Arezzo [prov.], Cavriglia, [Fraz.] Montegonzi, 16.V.1991, Lisa leg., 1 ex (MZUF); [Arezzo prov., ] Ris. Nat. [= Riserva Naturale = Nature Reserve] “Valle Inferno-Bandella”, Casa Giardino, 8.V.1998, L. Bartolozzi, B. Cecchi A. & Sforzi leg., 1 ex (MZUF); [Firenze prov., ] Firenze [= Florence], V.1896, ex-coll. Beccari, 1 ex (MZUF); Firenze prov., Greve [in Chianti], [Fraz.] Strada in Chianti, 20.I.1957, A. Bandinelli leg., 1 ex (MZUF); [Grosseto prov.], Parco Naturale della [= Natural Park of] Maremma, Mti [= Monti = Mounts] dell’Uccellina, S. Rabano, X.1988, L. Bartolozzi leg., 4 ex (MZUF).

Chorotype.

Turano-European with extension to Tunisia (cf. Háva 2007, Háva and Herrmann 2009, 2011, Háva et al. 2011, Háva and Nardi 2011).

Italian distribution.

All of Italy, Sicily and Sardinia (cf. Bertolini 1904, Luigioni 1929, Horion 1955, Audisio et al. 1995, Háva and Nardi 2011). These generic distributional data are most likely correct; nevertheless, precise records are known only from the following regions: Trentino (Bertolini 1890), Lombardy (Villa and Villa 1844, Giachino 1982), Piedmont (Ghiliani 1847, 1887), Aosta Valley (Iehl 1909), Romagna (Zangheri 1969), Tuscany (Bordoni et al. 2006), Latium (Horion 1955, FEI 2011), Abruzzi (Horion 1955), Apulia (Holdhaus 1911, Panganetti-Hummler 1918, Gridelli 1949, Horion 1955, Faggioli 1956, Angelini 1987), Basilicata (Angelini 1986, Angelini and Montemurro 1986), Calabria (Angelini 1991), Sicily (Ghiliani 1842, Rottenberg 1870, Steck 1887, Ragusa 1892, Vitale 1904, Luigioni 1929, Horion 1955, Audisio et al. 1995), and Sardinia (cf. Háva and Nardi 2011).

Remarks.

New collecting sites from Tuscany and Apulia. In Tuscany, the species was formally recorded from a sole locality (Bordoni et al. 2006); it is also for the first time recorded here from a nature reserve, of which the beetle-fauna is well known (cf. Zinetti and Terzani 2009). The species was already known from Apulia, from various localities, including the Gargano National Park (see the references above).

Sapuppo (2002) recorded Dermestes ater DeGeer, 1774 from two Sicilian localities based on specimens collected by A. Gulli and identified by A. Porta. The data from a locality are exactly the same as those of “Dermestes ater Oliv.” provided by Gulli (1961), and thus, both sites are attributed here to Dermestes (Montandonia) olivieri; the homonymy (Dermestes ater DeGeer and Dermestes ater Oliver) was very probably the cause of the error of Sapuppo (2002). Dermestes (Dermestes) ater DeGeer is a cosmopolitan species, but in Italy it is known from some northern regions only (Luigioni 1929: 537, as Dermestes (Dermestes) cadaverinus Fabr. [= Fabricius, 1775], Minelli and Negrisolo 1993: 92, as Dermestes ater Degeer, Audisio et al. 1995: 12, as Dermestes (Dermestes) ater Degeer).

Thorictus pilosus Peyron, 1857

http://species-id.net/wiki/Thorictus_pilosus

Thorictus pilosus Peyron, 1857: Audisio et al. 1995: 12.
Material examined.

None.

Chorotype.

Turanian-E-Mediterranean: Cyprus, Egypt, Greece, Iraq, Israel, Lebanon, Libya, Syria, Tunisia, Turkey and Uzbekistan (John 1963, Löbl 2007).

Italian distribution.

“S?” [= peninsular Italy?] (Audisio et al. 1995).

Remarks.

The above doubtful record for peninsular Italy has already been ignored by other authors (Löbl 2007, Zhantiev 2011a); this species is excluded from Italy here, since when considering its geographical distribution, its occurrence in this country is improbable. A doubtful record from Corsica (cf. Bertolini 1874: 96, as Thorictus piliger Schaum[, 1858], Luigioni 1929: 1022, as Thorictus pilosus Peyr., Porta 1929: 204, as Thorictus pilosus Peyron) has also never been confirmed (Löbl 2007, Zhantiev 2011a), and must be referred to Thorictus grandicollis grandicollis Germar, 1842 (Háva, unpublished data).

Thorictus wasmanni Reitter, 1895

http://species-id.net/wiki/Thorictus_wasmanni

Thorictus wasmanni Reitter, 1895: Audisio et al. 1995: 12.
Material examined.

None.

Chorotype.

Turanian: endemic of Uzbekistan (John 1963, Löbl 2007).

Italian distribution.

“Si?” [= Sicily?] (Audisio et al. 1995).

Remarks.

The above doubtful Sicilian record has already been ignored by other authors (Löbl 2007, Zhantiev 2011a); this species must be excluded from the Italian fauna, since its occurrence in Sicily is incompatible with its geonemy. It is known from the type specimens only (Zhantiev 2011b).

Attagenus (Attagenus) calabricus Reitter, 1881

http://species-id.net/wiki/Attagenus_calabricus

Attagenus calabricus Reitter, 1881: 37.
Attagenus (Lanorus) calabricus Reitt.: Reitter 1883: 89; Reitter 1891: 337; Ragusa 1892: 204; Reitter 1906: 378; Dalla Torre 1911: 52; Winkler 1926: 676; Luigioni 1929: 538; Porta 1929: 302.
Megatoma calabrica Rttr.: Reitter 1887: 48.
Attagenus calabricus Reitt.: Bertolini 1904: 58; Panganetti-Hummler 1918: 87; Schatzmayr 1941: 73; Angelini 1986: 74; Angelini and Montemurro 1986: 573; Angelini 1987: 38; Angelini 1991: 206.
Attagenus calabricus Rtt.: Liebmann 1962: 4.
Attagenus calabricus Reitter, 1881: Mroczkowski 1968: 82; Sparacio 1997: 73.
Attagenus (Lanorus) calabricus Reitter, 1881: Audisio et al. 1995: 13.
Attagenus (Attagenus) calabricus Reitter, 1881: Háva 2003: 59; Háva 2007: 308; Zhantiev 2011a.
Material examined.

Abruzzi: Pescara prov., Maiella, Valle Orfento, Caramanico, 23.VI.1988, G. Osella leg., 1 ex (JHAC). Apulia: Brindisi prov., Francavilla Fontana, 30.V.1998, C. Esposito leg., 1 ex (GNAC); [Foggia prov., GNP, ] Mattinata, 22.VI.1991, on flowers on the beach, M. Mantic leg., 4 ex (JHAC); [Foggia prov., GNP, ] Gargano, Mattinata, 8–10.V.1995, Z. Svec leg., 1 ex (JHAC); [Foggia prov., GNP, Vico del Gargano, Fraz.] San Menaio, V.1996, M. Fikacek leg., 2 ex (JHAC); Taranto prov., Circ. [= Circum] Mar Piccolo, 24.V.1992, Montemurro leg., 1 ex (JHAC); Taranto prov., Grottaglie, VI.1369 [sic!], FA, 1 ex (MCSV). Basilicata: [Matera prov., ] Marina di Nova Siri, 4.VI.2008, W. Apfel leg., 1 ex (JHAC); [Matera prov., ] Matera, 4.VI.1989, FA, 1 ex (AHG); [Matera prov., ] Matera, 350 m, 4.VI.1989, FA, 11 ex (MCSV); Potenza prov., Francavilla sul Sinni, 320 m, 5.VI.1988, querceta [= oak wood], FA, 1 ex (MCSV); [Potenza or Matera prov., ] Pollino, 19.VI.1994, FA, 1 ex (AHG). Calabria: “Calabria / Baudi [leg.]” [handwritten by unknown], “Holotypus 1881 / Attagenus / Calabricus / Reitter” [red-framed; handwritten by unknown], “Calabricus m. / Calab. Baudi [leg.]” [handwritten probably by Reitter], “coll. Reitter” [printed], “Lectotype / Attagenus calabricus Reitter / J. Háva des. 2012”, 1 ex (HNHM); “Calabria / Baudi [leg.]” [handwritten by unknown], “Paratypus 1881 / Attagenus / Calabricus / Reitter” [red-framed; handwritten by unknown], “coll. Reitter [printed]”, “Paralectotype / Attagenus calabricus Reitter / J. Háva des. 2012”, 1 ex (HNHM); Calabria [without further data], 1 ex (JHAC); [Cosenza prov., Cassano all‘Ionio, Fraz.] Marina di Sibari env., 3 m, sandy dunes, 14.6.2009, P. Kresl leg., 6 ex (JHAC); [Cosenza prov., ] Roseto Capo Spulico, 27.V.–3.VI.2006, V. Hron leg., 1 ex (JHAC); [Cosenza prov., ] Sila, Mte. [= Mount] Oliveto, 2.VII.1929, C. Confalonieri leg., 1 ex (JHAC); [Cosenza prov., ] Sila, Sud [= South of] S. Giovanni [in] Fiore, 1100 m, 4.VII.[19]87, FA, 2 ex (MCSV); Reggio Calabria prov., Asprom. [= Aspromonte], str. [= strada = road] Cimina–Zomaro, 600–800 m, 6.VI.1994, FA, 6 ex (JHAC).

Chorotype.

Apenninic endemic (Háva 2007).

Italian distribution.

Campania (Luigioni 1929), Apulia (Panganetti-Hummler 1918, Luigioni 1929, Angelini 1987), Basilicata (Angelini 1986, Angelini and Montemurro 1986), Calabria (Bertolini 1904, Luigioni 1929, Porta 1929, Schatzmayr 1941, Angelini 1991) and Sicily (Ragusa 1892, Bertolini 1904, Winkler 1926, Luigioni 1929, Porta 1929, Liebmann 1962, Mroczkowski 1968, Audisio et al. 1995, Sparacio 1997, Háva 2003, 2007, Zhantiev 2011a).

The generic records for Italy (Reitter 1883, 1891, 1906, Dalla Torre 1911, Winkler 1926, Mroczkowski 1968, Háva 2003, 2007), peninsular Italy (Audisio et al. 1995), southern Italy (Sparacio 1997) and mainland Italy (Zhantiev 2011a) refer to the above listed regions.

Remarks.

First record from the Abruzzi region; this new site (cf. Osella 1988) is at the northernmost limit of the species distribution, while the above record from the Brindisi province is at the easternmost limit. The southernmost site is the Pantelleria Island in the Sicilian Channel (Liebmann 1962).

Reitter (1881) described this species without giving any locality of collecting (see also Reitter 1887), but based on the specific name, the type locality is apparently in the Calabria region (southern peninsular Italy). The species was described based on an unspecified number of specimens without any type designation (Reitter 1881). The two type specimens above are thus syntypes and are designated here as lectotype and paralectotype, respectively; in this way “Calabria” becomes the type locality of the species (ICZN 1999, art. 76.2).

Attagenus (Attagenus) lobatus Rosenhauer, 1856

http://species-id.net/wiki/Attagenus_lobatus

Attagenus (Attagenus) lobatus Rosenhauer, 1856: Háva and Nardi 2011: 420.
Material examined.

Tuscany: Firenze [= Florence], 28.IX.1988, L. Chelazzi leg., 1 ♂ (MZUF).

Chorotype.

Centralasiatic-Mediterranean; this species was introduced to the USA (cf. Háva and Nardi 2011; Háva et al. 2013).

Italian distribution.

Latium, Campania, Apulia, Calabria, Sicily and Sardinia (cf. Háva and Nardi 2011).

Remarks.

First record for Tuscany. The new locality is the northernmost in Italy (cf. Háva and Nardi 2011).

Attagenus (Attagenus) pilosissimus Roubal, 1932, stat. prom.
Attagenus pellio L. var. pilosissimus Roubal, 1932: 66.
Attagenus pellio Lin. v. pilosissimus Roubal: Porta 1934: 171.
Attagenus pellio (Linnaeus, 1758) var. pilosissimus Roubal, 1932: Mroczkowski 1968: 89.
Attagenus pellio var. pilosissimus Roub.: Angelini 1986: 74.
Attagenus pellio var. pilosissimus Roubal: Angelini 1991: 206.
Attagenus (Attagenus) pellio (Linné, 1758) = Attagenus pilosissimus Roubal, 1932: Audisio et al. 1995: 13.
Attagenus (Attagenus) pellio (Linnaeus, 1758) = Attagenus pellio var. pilosissimus Roubal, 1932: Háva 2003: 66.
Attagenus (Attagenus) pellio (Linnaeus, 1758) = Attagenus pilosissimus Roubal, 1932: Háva 2007: 57.
Attagenus (Attagenus) pellio (Linnaeus, 1758) = Attagenus pellio var. pilosissimus Roubal, 1932: Háva and Nardi 2007: 122.
Material examined.

Apulia: Bari prov., Gioia del Colle, 22.IV.1993, De Marzo leg., 1 ♂ (JHAC); Foggia prov., [GNP, ] Foresta Umbra, Monte Sant’Angelo, caserme forestali [= barracks of the Italian National Forestry Service], 792 m, 33T 582306 4630267, 27.IV.2010, PC, death on windowsill, Prog. Foresta Umbra, 1 ♀ (JHAC). Lombardy: Bergamo prov., Oltre il Colle, 7.VII.2005, FA, 1 ♂ (JHAC).

Chorotype.

Alpino-Apenninic endemic (cf. Háva and Nardi 2007).

Italian distribution.

Apulia and Calabria (Roubal 1932, Porta 1934, Angelini 1986, 1991, Háva and Nardi 2007).

Remarks.

Attagenus pellio var. pilosissimus from Apulia was listed as a synonym of Attagenus (Attagenus) pellio by Audisio et al. (1995), because Mroczkowski (1968) listed pilosissimus as a “variety”. This means that this name was implicitly considered as a junior synonym as Mroczkowski also treated subspecific taxa in his paper. This opinion was confirmed by Háva and Nardi (2007: 122) through a study of the holotype, a female specimen which is very worn. A photo of this holotype is published at http://www.dermestidae.com/Attagenuspelliopilosissimus.html. Nevertheless, based on the study of the above new material, this variety is resurrected as a valid species (stat. prom.). Attagenus (Attagenus) pilosissimus and Attagenus (Attagenus) pellio show differences in the shape of the antennal club (Figs 1–2) and in the male genitalia (Figs 4–5): the median lobe of the aedeagus of the former species (Fig. 5) is longer and narrower, and parameres are parallel for most of their length. Moreover, Attagenus (Attagenus) pilosissimus also differs by the pubescence of the abdomen (Fig. 4) and pronotum (Fig. 3) and by the presence of only one small circular spot with whitish pubescence on each elytron (Fig. 3). Attagenus (Attagenus) pilosissimus is known only from mainland Italy (Lombardy, Apulia and Calabria), where, in sympatry, Attagenus (Attagenus) pellio (Linnaeus, 1758) also occurs (cf. Holdhaus 1911, as Attagenus pellio L., Háva and Nardi 2007).

Figures 1–6.

Attagenus (Attagenus) spp., males 1 Antennal club of Attagenus (Attagenus) pilosissimus from Italy (Apulia, Bari prov., Gioa del Colle) 2 Antennal club of Attagenus (Attagenus) pellio from Czech Republic (Central Bohemia, Prague) 3 Habitus of Attagenus (Attagenus) pilosissimus from Italy (Apulia, Bari prov., Gioia del Colle) (length 4.5 mm) 4 Abdomen in ventral view of Attagenus (Attagenus) pilosissimus from Italy (Apulia, Bari prov., Gioa del Colle) 5 Aedeagus (setation omitted) of Attagenus (Attagenus) pilosissimus from Italy (Lombardy, Bergamo prov., Oltre il Colle) (length 0.5 mm) 6 Aedeagus (setation omitted) of Attagenus (Attagenus) pellio from Czech Republic (Central Bohemia, Prague) (length 0.5 mm) (photos and drawings by J. Háva).

Attagenus (Attagenus) silvaticus Zhantiev, 1976

http://species-id.net/wiki/Attagenus_silvaticus

Attagenus (Attagenus) silvaticus Zhantiev, 1976: Háva and Nardi 2007: 122.
Attagenus silvaticus Zhantiev, 1976: Zhantiev 2011a.
Material examined.

Apulia: Foggia prov., [GNP, ] Foresta Umbra, Vico del Gargano, Bosco Sfilzi, 391 m, 33T 585633 4634653, 23.V.2010, D. Birtele & I. Toni leg., window flight trap, Prog. Foresta Umbra, 1 ♀ (CNBFVR).

Chorotype.

European (Czech Republic, Hungary, northwestern Russia, Sicily, Slovakia, Ukraine) with extension eastward to “Caucasus”, Anatolia, Iran and West Siberia (Háva 2007, Háva and Nardi 2007, Háva et al. 2010, as Attagenus silvaticus, Zhantiev 2011a).

Italian distribution.

Sicily (Háva and Nardi 2007, Zhantiev 2011a).

Remarks.

First record for mainland Italy. The above specimen was intercepted in a beech wood (D. Birtele, pers. comm., 2012).

Attagenus (Attagenus) simonis Reitter, 1881

http://species-id.net/wiki/Attagenus_simonis

Attagenus tigrinus F. [= (Fabricius, 1792)] v. Simoni [sic!] Reit.: Bertolini 1904: 58.
Attagenus bifasciatus Oliv. [= (Olivier, 1790)] v. Simoni [sic!] Reitt.: Luigioni 1923b: 4.
Attagenus (Lanorus) bifasciatus Oliv. a Simoni [sic!] Reitt.: Luigioni 1929: 538; Porta 1929: 302.
Attagenus bifasciatus (Olivier, 1790) var. simoni [sic!] Reitter, 1881: Sapuppo 2002: 187–188.
Attagenus (Attagenus) simonis Reitter, 1881: Háva and Nardi 2007: 418–419.
Material examined.

None.

Chorotype.

E-Mediterranean: Egypt, Israel, Jordan and Syria (Háva 2007).

Italian distribution.

Latium (Luigioni 1929), Campania (Bertolini 1904, Luigioni 1929, Porta 1929), Capri Island (Luigioni 1923b, 1929, Porta 1929) and Sicily (Luigioni 1929, Sapuppo 2002).

Remarks.

This species was recently excluded from the Italian fauna (Háva and Nardi 2007, 2011); also according to Mroczkowski (1968: 81, as Attagenus bifasciatus var. simonis), it occurs only in some East-Mediterranean countries. Nevertheless, an escatomediterranean distributional pattern (Zilli 2000) could explain the occurrence of this species in Italy.

Gulli (1961: 4, as Attagenus bifasciatus Oliv. var. Rossii Ganglb. [= Ganglbauer, 1904] and Attagenus bifasciatus var. Simoni [sic!]) recorded Sicilian specimens of Attagenus (Attagenus) rossii (identified by A. Porta) with a colour similar to Attagenus (Attagenus) simonis; these specimens are also listed by Sapuppo (2002), but unfortunately their re-examination has so far been impossible.

Attagenus (Attagenus) tigrinus pulcher Faldermann, 1835

http://species-id.net/wiki/Attagenus_tigrinus_pulcher

Attagenus pulcher Fald.: Bargagli 1872: 102, Bertolini 1874: 99.
Attagenus tigrinus F. [= (Fabricius, 1792)] var. pulcher Falder.: Bertolini 1904: 58.
Material examined.

None.

Chorotype.

Mediterranean (cf. Háva and Nardi 2011); Attagenus (Attagenus) tigrinus pulcher is a Caucasian endemic (Zhantiev 2009).

Italian distribution.

Sardinia (Bargagli 1872, Bertolini 1874, 1904).

Remarks.

This subspecies is excluded here from the Italian fauna. Zhantiev (2009) recognized this taxon as a valid subspecies. This paper was overlooked by Háva and Nardi (2011), so these authors, according to Háva (2007), listed it as a synonym of Attagenus (Attagenus) bifasciatus (A.G. Olivier, 1790). Nevertheless, the status of this subspecies needs to be revised (Háva, unpublished data).

Anthrenus (Anthrenops) coloratus Reitter, 1881

http://species-id.net/wiki/Anthrenus_coloratus

Anthrenus (Anthrenops) coloratus Reitter, 1881: Háva 2007: 311.
Anthrenus (Florilinus) coloratus Reitter, 1881: Zhantiev 2011a.
Material examined.

Apulia: [Foggia prov., GNP, ] Gargano, V.1991, J. Háva leg., 12 ex (JHAC).

Chorotype.

Turano-Europeo-Mediterranean with extension to Canary Islands, Tajikistan, “India” and Sudan; this species was introduced to and is widespread in the USA (Beal 1998, as Anthrenus coloratus, Háva 2003, 2007).

Italian distribution.

Italy (Háva 2007). Italian mainland (Zhantiev 2011a).

Remarks.

The generic Italian record by Háva (2007) was based on the above specimens.

Anthrenus (Anthrenus) angustefasciatus Ganglbauer, 1904

http://species-id.net/wiki/Anthrenus_angustefasciatus

Anthrenus (Anthrenus) pimpinellae Fabr. [= (Fabricius, 1775)] a. angustefasciatus Ganglb.: Luigioni 1929: 540.
Anthrenus pimpinellae Fabr. ab. angustefasciatus Ganglb.: Porta 1934: 171.
Anthrenus (Anthrenus) angustefasciatus Ganglbauer, 1904: Háva 2003: 79; Háva 2007: 311; Kadej et al. 2007: 728; Háva 2011b: 78; Zhantiev 2011a; Háva et al. 2013: 137.
Material examined.

Friuli-Venezia Giulia: Trieste prov., Carso, [Sgonico, Fraz.] Gabrovizza S. Primo, 12.VI.1988, sn, PC, 1 ♀ (PCOP). Tuscany: Grosseto prov., Pitigliano, 23.I.[20]02, L. Saltini leg., 1 ex (GNAC). Apulia: [Brindisi prov., ] Brindisi env., 18.V.1997, J. Hrdlicka leg., 4 ex (JHAC). Basilicata: Potenza prov., [Pollino National Park, ] Terranova di Pollino, 900 m, 27–30.VI.2005, GC, sn, 1 ♀ (PCOP).

Chorotype.

European with extension southward to a part of the Maghreb: Algeria, Bosnia and Herzegovina, Croatia, Czech Republic, France mainland, Italy, Morocco, Portugal, Spain, Switzerland, Yugoslavia and European Turkey (cf. Háva 2011b, Zhantiev 2011a, Háva et al. 2013).

The records for Corsica (Luigioni 1929, Porta 1934) were later ignored (Sainte-Claire Deville 1937, Mroczkowski 1968, as Anthrenus (Anthrenus) pimpinellae var. angustefasciatus, Zhantiev 2011a, Háva et al. 2013).

Italian distribution.

Calabria (Luigioni 1929, Porta 1934), Sardinia (Háva et al. 2013). Italy (without further details) (Háva 2003, 2007, Kadej et al. 2007, Háva 2011b). Italian mainland (Zhantiev 2011a).

Remarks.

Ganglbauer (1904: 42) described Anthrenus pimpinellae var. angustefasciatus from “Dalmatien [= Dalmatia]” that was only recently raised to species level (Háva 2003: 79); its diagnostic features were provided by different authors (Kadej 2005, Kadej et al. 2007, Háva 2011b, Háva and Zahradník 2011).

First records for northern Italy (Friuli-Venezia Giulia), for central Italy (Tuscany), for Apulia and Basilicata; the former record is expected since this species occurs in neighbouring foreign regions (Depoli 1928, as Anthrenus pimpinellae f. angustefasciatus Ganglb., Novak 1952, as Anthrenus pimpinellae a. angustefasciatus Gglb.). The generic records for Italy (Háva 2003, 2007, Kadej et al. 2007, Háva 2011b) and, probably also those generic for mainland Italy (Zhantiev 2011a), refer to previous literature records and, in part, to the above material.

Anthrenus (Anthrenus) delicatus Kiesenwetter, 1851

http://species-id.net/wiki/Anthrenus_delicatus

Anthrenus pimpinellae Fabr. [= (Fabricius, 1775)] var. delicatulus [sic!] Kiesw.: Ghiliani 1887: 311.
Anthrenus pimpinellae F. var. delicatus Kiesw.: Steck 1887: 181; Ragusa 1892: 206.
Anthrenus pimpinellae Fabr. Var. delicatulus [sic!] Kiesw.: Baudi 1890: 102.
Anthrenus pimpinellae F. [var.] Isabellinae Muls. [= Mulsant & Rey, 1868]: Ragusa 1892: 206.
Anthrenus pimpinellae F. var. delicatus K.: Bertolini 1904: 59.
Anthrenus pimpinellae F. ab. delicatus Kiesw.: Luigioni 1923a: 135.
Anthrenus (Anthrenus) pimpinellae F. a. delicatus Kiesw.: = Isabellinae Muls. Rey: Luigioni 1929: 540.
Anthrenus (Anthrenus) pimpinellae Fabr. V. delicatus Kiesw.: Porta 1929: 306.
Anthrenus pimpinellae v. delicatus Kiesw.: Liebmann 1962: 4.
Anthrenus (Anthrenus) delicatus Kiesenwetter, 1851: Audisio et al. 1995: 14; Kadej 2005: 728; Lo Cascio et al. 2006: 322; Háva and Nardi 2011: 423; Zhantiev 2011a.
Anthrenus (Anthrenus) delicatus delicatus Kiesenwetter, 1852 [sic!]: Háva 2007: 312.
Material examined.

Apulia: Foggia prov., [GNP, ] Promontorio del Gargano, Vieste, SP 89 [= Strada Provinciale n. 89 = Provincial Road n. 89], Casa Cupari dint., 354 m, 33T 5920001 4630150, 29.V.2010, abandoned quarry, sn on grasses, PC, Prog. Foresta Umbra, 6 ex (CNBFVR; JHAC); Foggia prov., [GNP, ] Promontorio del Gargano, Peschici, Lampia del Principe dint., 161 m, 33T 587282 4639346, 28.IV.2010, maquis sn, PC, Prog. Foresta Umbra, 1 ♀ (CNBFVR). Latium: Roma prov., M.ti [= Monti = Mounts] Simbruini, Jenne dint., M.te [= Monte = Mount] Porcaro, 800 m, 31.V.2007, sn, PC GC, 1 ex (PCOP). Sicily: Palermo prov., Corleone, Bosco della Ficuzza, 700 m, 1.VI.2008, sn near a watering trough, PC GC, 1 ex (PCOP).

Chorotype.

Mediterranean, reaching eastward as far as Caucasus and Iran (cf. Kadej 2005, Háva and Nardi 2011, Zhantiev 2011b). This species is also recorded from Cyprus (Zhantiev 2011a, Alziar and Lemaire 2012: 67, as Anthrenus delicatus), and this country must be added to the distribution summarized by Háva and Nardi (2011).

Italian distribution.

Trentino, Venezia Giulia (Luigioni 1929), Piedmont (Ghiliani 1887, Baudi 1890, Bertolini 1904, Luigioni 1929), Lombardy (Háva and Nardi 2004), Liguria (Luigioni 1929), Latium, Campania (Luigioni 1923a, Porta 1929), Sicily (Ragusa 1892, Bertolini 1904, Luigioni 1929, Porta 1929, Liebmann 1962, Audisio et al. 1995, Lo Cascio et al. 2006) and Sardinia (Bertolini 1904, Luigioni 1929, Porta 1929, Audisio et al. 1995, Háva and Nardi 2011, Zhantiev 2011a).

Moreover, Luigioni (1929) recorded this species generically for central and southern Italy; generic records for northern and peninsular Italy (Audisio et al. 1995) and for “Italy” (Kadej 2005, Háva 2007) refer to the above regions, the same is probably valid for a generic record for mainland Italy (Zhantiev 2011a).

Remarks.

First detailed records from Apulia. About Sicily, Ragusa (1892) wrote Ne posseggo vari esemplari [= I have several specimens] without giving the collecting sites, so the sole known precise Sicilian localities were the Pantelleria and Lipari Islands (Liebmann 1962, Lo Cascio et al. 2006); the above new site is located in one of the most important forested areas of western Sicily, a zone of great value for environmental conservation (cf. Sabella and Sparacio 2004, Mason et al. 2006, La Mantia et al. 2010, Falci et al. 2012). Luigioni (1923a) also recorded Anthrenus (Anthrenus) delicatus from the environs of Rome; this record was overlooked by Nardi (1997). As stated by Audisio et al. (1995: 16), the Italian distribution of this species should be verified, because it was considered for a long time as a variety or aberration of Anthrenus (Anthrenus) pimpinellae. Moreover, the colour of elytral fasciae in four of the above Apulian specimens are visually very similar to those of Anthrenus (Anthrenus) mroczkowskii Kalík, 1954, nevertheless their identification was established by the examination of the structure of antennae, 9th sternite and male genitalia (cf. Kadej 2005, Kadej et al. 2007). In this framework, it will be better to revise the specimens of Anthrenus (Anthrenus) mroczkowskii recorded so far from Italy (Angelini 1986, as Anthrenus pimpinellae mroczkowskii, Angelini and Montemurro 1986, as Anthrenus pimpinellae mroczkowskii). The distribution of Anthrenus (Anthrenus) mroczkowskii also includes the following countries: Albania, Bosnia and Herzegovina, Bulgaria, Corsica, Crete, Croatia, Greece, Slovenia, European Turkey and Algeria (introduced) (Háva 2007, as Anthrenus (Anthrenus) pimpinellae mroczkowskii, Kadej et al. 2007).

Anthrenus (Anthrenus) munroi Hinton, 1943

http://species-id.net/wiki/Anthrenus_munroi

Anthrenus munroi Hint.: Angelini 1986: 73; Angelini and Montemurro 1986: 573; Angelini 1987: 37.
Anthrenus (Anthrenus) munroi Hinton, 1943: Háva and Nardi 2011: 425.
Material examined.

Tuscany: [Livorno prov.], Is. [= Isola = Island] d’Elba, dint. Marina di Campo, 15–17.VI.1999, Abbazzi, Bartolozzi, Lo Cascio & Sforzi leg., 1 ♂ (MZUF).

Chorotype.

Mediterranean (eastwards as far as Ukraine, Turkey, Syria, Lebanon, Jordan, and Israel), with extension to Hungary and Portugal (cf. Kadej et al. 2007, Háva and Nardi 2011).

Italian distribution.

Apulia, Basilicata, Calabria (Angelini 1986, Angelini and Montemurro 1986, Angelini 1987) and Sardinia (Háva and Nardi 2011).

Remarks.

First record for central Italy; this new site is the northernmost in Italy. The species also occurs in the facing Corsica (cf. Háva and Nardi 2011).

Anthrenus (Anthrenus) pimpinellae pimpinellae (Fabricius, 1775) and Anthrenus (Nathrenus) verbasci (Linnaeus, 1767) were the sole congeneric species previously recorded from Elba Island (Holdhaus 1923, as Anthrenus pimpinellae F. and Anthrenus verbasci L.; Luigioni 1929, Porta 1929, both as Anthrenus (Anthrenus) pimpinellae Fabr. and Anthrenus (Nathrenus) verbasci Lin.)

Anthrenus (Anthrenus) pimpinellae pimpinellae (Fabricius, 1775)

http://species-id.net/wiki/Anthrenus_pimpinellae_pimpinellae

Anthrenus pimpinellae F.: Ragusa 1892: 206; Bertolini 1904: 59; Holdhaus 1911: 445; Panganetti-Hummler 1918: 87; Holdhaus 1923: 101; Luigioni 1923b: 4; Marcuzzi 1985: 8.
Anthrenus (Anthrenus) pimpinellae Fabr.: Luigioni 1929: 540; Porta 1929: 306.
Anthrenus (Anthrenus) pimpinellae pimpinellae (Fabricius, 1775): Háva and Nardi 2004: 120; Háva and Nardi 2011: 426.
Anthrenus (Anthrenus) pimpinellae (Fabricius, 1775) s.l.: Lo Cascio et al. 2006: 322.
Material examined.

Apulia: Foggia prov., [GNP, ] Promontorio del Gargano, Vieste, SP 89 [= Strada Provinciale n. 89 = Provincial Road n. 89], Casa Cupari dint., 354 m, 33T 5920001 4630150, abandoned quarry, sn on grasses, 29.V.2010, PC, Prog. Foresta Umbra, 4 ex (CNBFVR).

Chorotype.

Cosmopolitan (cf. Háva and Nardi 2011).

Italian distribution.

All of Italy and Sicily (Bertolini 1904, Luigioni 1929, Porta 1929; Háva and Nardi 2011). This species is also recorded from some minor islands: Elba (Holdhaus 1923), Capri (Luigioni 1923b, 1929, Porta 1929), Lampedusa (Ragusa 1892, Luigioni 1929) and Vulcano (Lo Cascio et al. 2006).

Remarks.

Species common and widespread in Italy but from the Gargano National Park (cf. Angelini 1987) was recorded only by Holdhaus (1911); it is also known from other Apulian localities (Panganetti-Hummler 1918, Marcuzzi 1985).

Anthrenus (Nathrenus) signatus Erichson, 1846

http://species-id.net/wiki/Anthrenus_signatus

Anthrenus (Nathrenus) signatus Erichson, 1846: Háva and Nardi 2011: 428.
Material examined.

Basilicata: Potenza prov., Pignola, Ris. [= Riserva = Reserve] WWF, L. [= Lago = Lake] Pignola, 700 m, 21.VI.1992, FA, 1 ♂ (JHAC).

Chorotype.

European (cf. Háva and Nardi 2011).

Italian distribution.

South Tyrol, Venetia, Venezia Giulia, Latium?, Apulia, Basilicata and Sicily (cf. Háva and Nardi 2011).

Remarks.

This species was recorded in Basilicata region from the Pollino National Park (Angelini 1986) only; the above specimen comes from a nature reserve, from which a sole congeneric species, Anthrenus (Nathrenus) verbasci, was known so far (Angelini 1996a, 1998, both as Anthrenus verbasci).

Anthrenus signatus ab. bielawskii Mroczkowski, 1958 from Bulgaria (Mroczkowski 1958: 6, 1968: 145) was recently ignored (Háva 2007, 2008, 2011a), since according to the ICZN (1999, art. 45.5 and 45.6.2), it is an infrasubspecific and unavailable name.

Globicornis (Globicornis) breviclavis (Reitter, 1878)

http://species-id.net/wiki/Globicornis_breviclavis

Hadrotoma breviclavis Reitter: Baudi 1890: 101.
Hadrotoma breviclavis Reitt.: Bertolini 1904: 59.
Globicornis (Globicornis) breviclavis Reitt.: Ganglbauer 1904: 31; Reitter 1906: 379; Dalla Torre 1911: 63; Winkler 1926: 679; Luigioni 1929: 539; Porta 1929: 304.
Globicornis (Globicornis) breviclavis (Reitter, 1878): Audisio et al. 1995: 13, 16.
Material examined.

None.

Chorotype.

Endemic to the Near East: Armenia, Bulgaria, “Caucasus”, Georgia, European Turkey, Ukraine, and Southern European Russia (Adygeya, Krasnodar kraj) (Háva 2007, Háva and Legalov 2010, Zhantiev 2011a).

Italian distribution.

Piedmont: Graian and Pennine Alps (Baudi 1890, Luigioni 1929, Porta 1929); Piedmont (without further details) (Bertolini 1904, Reitter 1906); Piedmont? (Dalla Torre 1911, Winkler 1926, Audisio et al. 1995). Liguria: Ligurian Apennines (Luigioni 1929); Liguria? (Audisio et al. 1995).

Remarks.

This species described from Caucasus (Reitter 1878, as Hadrotoma breviclavis) is excluded here from the Italian fauna. Ganglbauer (1904) listed the paper by Baudi (1890), while Dalla Torre (1911) referred to Ganglbauer (1904). The Italian records were ignored by Mroczkowski (1968: 114), who recorded this species only from the Caucasus. No specimens of this species are present in the P. Luigioni collection (MCZR) (A. Zilli, pers. comm. 2013). The above Italian records probably concern the similar Globicornis (Globicornis) luckowi Herrmann, Háva & Kadej, 2011 which is known only from southern Switzerland, Piedmont and Liguria (Herrmann et al. 2011, Háva et al. 2013). Nevertheless, as discussed for Attagenus (Attagenus) simonis, an escatomediterranean distributional pattern (Zilli 2000) could explain the occurrence of Globicornis (Globicornis) breviclavis in Italy.

Globicornis (Globicornis) fasciata (Fairmaire & Brisout de Barneville, 1859)

http://species-id.net/wiki/Globicornis_fasciata

Hadrotoma fasciata Fairm.: Bertolini 1904: 59; Luigioni 1923a: 135.
Globicornis fasciata Fair.: Vitale 1904: 75.
Globicornis (Globicornis) fasciata Frm. [18]59: Winkler 1926: 679.
Globicornis (Globicornis) fasciata Fairm.: Luigioni 1929: 539; Porta 1929: 304.
Globicornis fasciatus [sic!] Fairm.: Horion 1955: 208.
Globicornis (Globicornis) fasciata (Fairmaire, 1859): Mroczkowski 1968: 114; Audisio et al. 1995: 13, 16; Zhantiev 2011a.
Globicornis (Globicornis) fasciata (Fairmaire & Brisout, 1859): Háva 2003: 109; Háva and Nardi 2011: 429.
Globicornis (Globicornis) fasciata (Fairmaire & Brisout de Barneville, 1859): Háva 2007: 315.
Material examined.

Basilicata: Potenza prov., Parco Naz. del Pollino [= Pollino National Park], Casa del Conte dint., tra [= between] Acqua Tremola e [= and] Piani di S. Francesco, 1100–1500 m, 9.VII.2002, sn, GC, 3 ex (PCOP).

Chorotype.

European (Belgium, Denmark, France, Germany, Italy, Spain, Sweden, Switzerland) with extension southward to Tunisia (cf. Háva et al. 2010, Háva and Nardi 2011).

Italian distribution.

Piedmont (Porta 1929, Horion 1955), Tuscany (Luigioni 1929, Horion 1955), Latium (Luigioni 1923a, 1929, Porta 1929, Horion 1955), Sicily (Vitale 1904) and Sardinia (Bertolini 1904, Winkler 1926, Luigioni 1929, Porta 1929, Horion 1955, Mroczkowski 1968, Audisio et al. 1995, Háva 2003, 2007, Zhantiev 2011a).

The records for northern and peninsular Italy (Audisio et al. 1995) as those generic for Italy (Háva 2003) refers to above regions; the same is probably valid for a generic record for mainland Italy (Zhantiev 2011a).

Remarks.

First record from southern Italy. Five species of this genus are now recorded from the Pollino National Park (cf. Angelini 1986).

The records for Piedmont (Porta 1929, Horion 1955) and those generic for northern Italy (Audisio et al. 1995: 13) refers to the Maritime Alps (Audisio et al. 1995: 16). The species also occurs in a neighbouring French department (Sainte-Claire Deville 1937: 252, as Globicornis fasciata Fairm.). The record from Sicily (Vitale 1904) was omitted by subsequent authors (cf. Luigioni 1929, Audisio et al. 1995, Zhantiev 2011a), and thus, it should be confirmed.

In the literature (see also above), the authorship of this species was often attributed only to Fairmaire.

Globicornis (Hadrotoma) sulcata (C.N.F. Brisout de Barneville, 1866)

http://species-id.net/wiki/Globicornis_sulcata

Globicornis ? sulcata (C. Brisout de Barneville, 1866): Angelini 1986: 74.
Globicornis (Hadrotoma) ? sulcata (C. Brisout de Barneville, 1866): Audisio et al. 1995: 16.
Globicornis (Hadrotoma) sulcata (C. Brisout de Barneville, 1866): Audisio et al. 1995: 13.
Material examined.

Abruzzi: Parco Naz. [= National Park], L‘Aquila prov., Pescasseroli, 1300 m, 1.IV.2003, FA, 1 ♀ (JHAC). Basilicata: Potenza prov., Moliterno, 500 m, 11.V.2003, FA, 1 ♀ (JHAC); Potenza prov., Ruoti, 29.IX.1996, AbiesQuercus, FA, 1 ♂ (JHAC). Campania: Avellino prov., Cilento, Novi Velia, M. [= Mount] Sacro, 1900 m, 7.XII.1996, Fagus, FA, 1 ♂ (JHAC). Sicily: Palermo prov., Ficuzza, 700 m, 1–4.V.2000, bosco leccio [= ilex wood], FA, 1 ♂ (JHAC); Palermo prov., Bosco della Ficuzza, 27.V.2006, M. Sarovec leg., 1 ♂, 4 ♀♀ (JHAC).

Chorotype.

W-Mediterranean: France, Italy and Spain.

Italian distribution.

Basilicata (Angelini 1987, Audisio et al. 1995).

Remarks.

First records from central Italy (Abruzzi), Campania and Sicily. In the Sicilian site, it was also collected Anthrenus (Anthrenus) delicatus (see above).

Trogoderma angustum (Solier, 1849)

http://species-id.net/wiki/Trogoderma_angustum

Trogoderma angustum (Solier, 1849): Háva 2007: 319; Zhantiev 2011a.
Material examined.

None.

Chorotype.

Subcosmopolitan (cf. Háva 2007). This species is also recorded from Switzerland (Kenis 2005); this country must be added to the European distributions recently summarized by Háva (2007) and by Zhantiev (2011a).

Italian distribution.

Italy (without further details) (Háva 2007). Italian mainland (Zhantiev 2011a).

Remarks.

The above generic Italian record by Háva (2007) was based on a personal communication from V. Kalík, who identified Italian material; unfortunately, no further details are available (Háva, unpublished data).

Trogoderma inclusum LeConte, 1854

http://species-id.net/wiki/Trogoderma_inclusum

Trogoderma meridionalis Kraatz, 1858: 146.
Trogoderma meridionalis Kraatz: Mulsant and Rey 1868: 132.
Trogoderma meridionale [sic!] Kraatz: Bertolini 1874: 99.
Trogoderma versicolor Creutz. [= (Creutzer, 1799)] var. meridionale [sic!] Kr.: Ragusa 1892: 205; Porta 1929: 305.
Trogoderma versicolor Creut. v. meridionale [sic!] K.: Bertolini 1904: 59.
Trogoderma versicolor Creutz. a. meridionale [sic!] Kr.: Luigioni 1929: 540.
Trogoderma versicolor meridionale [sic!] Kraatz, 1858: Mroczkowski 1968: 107.
Trogoderma inclusum Le Conte: Dal Monte 1972: 102.
Trogoderma ? inclusum Lec.: Angelini and Montemurro 1986: 573.
Trogoderma inclusum: Nardi R. et al. 1993: 623, fig. 9.
Trogoderma [?] inclusum J.L. Leconte, 1854: Audisio et al. 1995: 13, 16.
Trogoderma inclusum Leconte: Nicoli Aldini 1999: 8.
Trogoderma inclusum LeConte, 1854: Háva 2007: 319; Háva and Nardi 2011: 432; Zhantiev 2011a.
Material examined.

Apulia: Lecce prov., Salento, Torre S. Giovanni dint., 1–7.VII.2002, PC GC, sn, 1 ex (PCOP). Venetia: Verona prov., Verona, [Fraz.] Cancello, [about 500 m], 22.VI.1986, sn, PC, 1 ex (PCOP).

Chorotype.

Cosmopolitan (cf. Háva and Herrmann 2011, Háva and Nardi 2011, Háva et al. 2011).

Italian distribution.

Lombardy (Nicoli Aldini 1999, Háva and Nardi 2011), Liguria (Luigioni 1929), Emilia-Romagna (Nicoli Aldini 1999), Latium (Nardi R. et al. 1993, Nicoli Aldini 1999), Basilicata [?] (Angelini and Montemurro 1986, Audisio et al. 1995), Sicily (Kraatz 1858, Mulsant and Rey 1868, Ragusa 1892, Bertolini 1904, Porta 1929, Mroczkowski 1968) and Sardinia (cf. Háva and Nardi 2011).

The first Italian record of this species was provided by Kraatz (1858) who recorded it from Sicily (without a precise locality), while some authors (Bertolini 1874, Dal Monte 1972, Háva 2007) and Zhantiev (2011a) recorded it from Italy and mainland Italy, respectively, without further details. Moreover, Nicoli Aldini (1999: 8) also recorded it from alcune città del Nord-Italia [= some towns of Northern Italy] but without mentioning the regions.

Remarks.

In Italy, this species and Trogoderma versicolor (Creutzer, 1799) have often been confused (cf. Nicoli Aldini 2003b, Háva and Nardi 2011); old Italian records in the literature must be thus considered with caution. Trogoderma inclusum is recorded here for the first time from Apulia and Venetia. This species is a stored product pest; it also develops in nests of social aculeate Hymenoptera, and is a predator of egg masses of Lymantria dispar (Linnaeus, 1758) (Lepidoptera, Erebidae, Lymantriinae) (cf. Háva and Nardi 2011).

Literature records

The old Sicilian records of Attagenus (Attagenus) trifasciatus (Fabricius, 1787) and Trogoderma glabrum (Herbst, 1783) (Ghiliani 1842) were ignored (or overlooked?) by subsequent authors and are thus considered as doubtful here (Table 1).

Thorictus mauritanicus Lucas, 1846 and Thorictus stricticollis Kraatz, 1859 were both recorded from “Italy” by Háva (2003, 2007) and from “Italian mainland” by Zhantiev (2011a). Háva’s (2003, 2007) records of Thorictus mauritanicus refer only to Sardinia and Sicily, while those of Thorictus stricticollis (Háva 2003, 2007) refer only to Sardinia (Háva, unpublished data). In this framework, the records of both species for mainland Italy (Zhantiev 2011a) are considered erroneous.

Thorictus sicilianus John, 1965 (= Thorictus sicilianus John, 1966) from Sicily and Morocco (John 1965, 1966, Audisio et al. 1995, Löbl 2007) was recently recognized as a junior synonym of Thorictus grandicollis grandicollis Germar, 1842 (Háva 2013); Attagenus picipennis Pic, 1894 from Sicily (Audisio et al. 1995) is a junior synonym of Attagenus (Attagenus) brunneus Faldermann, 1835 (Háva 2007: 57); Anthrenus (Attagenus) scrophulariae suecius Palm, 1940 that was recorded from southern Italy and Sicily (Audisio et al. 1995), was recently listed by Háva (2007: 313) as a junior synonym of Anthrenus (Anthrenus) scrophulariae scrophulariae (Linnaeus, 1758), but for an error this new synonym was not formally established (cf. Háva 2007: 57). These three taxa are thus excluded from the new checklist.

Dermestes (Dermestinus) gyllenhalii gyllenhalii Laporte de Castelnau, 1840

Dermestes ? atomarius Erichson, 1846: Sapuppo 2002: 189.

Si? (Sapuppo 2002).

Dermestes (Dermestinus) laniarius laniarius Illiger, 1801

Dermestes laniarius Illig.: Gulli 1961: 4.

Dermestes laniarius Illiger, 1801: Sapuppo 2002: 190.

Si (Gulli 1961, Sapuppo 2002).

Thorictus grandicollis grandicollis Germar, 1842

Thorictus grandicollis Germ.: Cerruti 1954: 111.

N (Cerruti 1954).

Attagenus (Attagenus) cyphonoides Reitter, 1881

Attagenus alfierii Pic[, 1910]: Trematerra 1987: 16.

Attagenus (Attagenus) cyphonoides Reitter, 1881: Háva and Nardi 2004: 121, Háva 2007: 308.

Attagenus (Attagenus) cyphonoides Reitter, 1881: Zhantiev 2011a.

S (Háva and Nardi 2004). Italy (Trematerra 1987, Háva 2007). Italian mainland (Zhantiev 2011a).

Attagenus (Attagenus) fasciatus (Thunberg, 1795)

Attagenus fasciatus (Thunberg, 1795): Nicoli Aldini 2003a 155–156, Nicoli Aldini 2003b: 123–125, Nicoli Aldini 2004: 983, 985, Zhantiev 2011a.

Attagenus (Attagenus) fasciatus (Thunberg, 1795): Háva and Nardi 2004: 121, Háva 2007: 308.

N (Nicoli Aldini 2003a, 2003b, 2004), S (Háva and Nardi 2004). Italy (Háva 2007). Italian mainland (Zhantiev 2011a).

Attagenus (Attagenus) pantherinus (Ahrens, 1814)

Attagenus pantherinus: Henry 1938: 66.

Attagenus (Attagenus) pantherinus (Ahrens, 1814): Háva 2007: 309, Háva and Nardi 2007: 121.

Attagenus pantherinus (Ahrens, 1814): Zhantiev 2011a.

N (Henry 1938, Háva and Nardi 2007), Sa (Zhantiev 2011a). Italy (Háva 2007).

Attagenus (Attagenus) rossii Ganglbauer, 1904

Dermestes bifasciatus Rossi, 1794: 79.

Attagenus bifasciatus Rossi: Rottenberg 1870: 238, Bertolini 1874: 99, Steck 1887: 181, Bertolini 1904: 58.

Attagenus bifasciatus: Failla-Tedaldi 1887: 158.

Attagenus (Lanorus) bifasciatus Rossi: Ragusa 1892: 204.

Attagenus (Lanorus) bifasciatus Oliv. [= (A.G. Olivier, 1790)] a. Rossii Ganglb.: Luigioni 1929: 538.

Attagenus (Lanorus) bifasciatus Oliv. v. Rossii Ganglb.: Porta 1929: 302.

Attagenus bifasciatus Oliv. var. Rossii Ganglb.: Gulli 1961: 4.

Attagenus bifasciatus (Olivier, 1790) var. Rossii Ganglbauer, 1904: Sapuppo 2002: 187.

Attagenus (Attagenus) rossii Ganglbauer, 1904: Háva 2007: 309, Háva and Nardi 2007: 123, Háva and Nardi 2011: 422.

Attagenus rossii Ganglbauer, 1904: Zhantiev 2011a.

Attagenus rossii Ganglbauer, 1804 [sic!]: FEI 2012.

S (Rossi 1794, Luigioni 1929, Porta 1929, Háva and Nardi 2007, 2011), Si (Rottenberg 1870, Bertolini 1874, Failla-Tedaldi 1887, Steck 1887, Ragusa 1892, Bertolini 1904, Luigioni 1929, Porta 1929, Gulli 1961, Sapuppo 2002, Háva and Nardi 2007, Zhantiev 2011a, FEI 2012), Sa (cf. Háva and Nardi 2011). Italy (Háva 2007). Italian mainland (Zhantiev 2011a).

Attagenus (Attagenus) trifasciatus (Fabricius, 1787)

Attagenus trifasciatus Fabr.: Ghiliani 1842: 33.

Attagenus ? trifasciatus (Fabricius, 1787): Sapuppo 2002: 189.

Si? (Ghiliani 1842, Sapuppo 2002).

Anthrenocerus australis (Hope, 1843)

Anthrenocerus australis (Hope, 1843): Herrmann and Háva 2007: 5, Háva 2007: 315, Zhantiev 2011a.

N (Herrmann and Háva 2007). Italy (Háva 2007). Italian mainland (Zhantiev 2011a).

Globicornis (Globicornis) luckowi Herrmann, Háva & Kadej, 2011

Globicornis (Globicornis) luckowi Herrmann, Háva & Kadej, 2011: Háva et al. 2013: 138.

N (Háva et al. 2013).

Globicornis (Hadrotoma) corticalis (Eichhoff, 1863)

Globicornis corticalis (Eichhoff, 1863): Biscaccianti et al. 2008: 50.

S (Biscaccianti et al. 2008).

Reesa vespulae (Milliron, 1939)

Reesa vespulae (Milliron, 1939): Nicoli Aldini 2003a: 156, 159, Nicoli Aldini 2003b: 123, 127, Nicoli Aldini 2004: 983, 987.

N (Nicoli Aldini 2003a, 2003b, 2004).

Trogoderma glabrum (Herbst, 1783)

Trogoderma elongatulum Fabr. [= (Fabricius, 1801)]: Ghiliani 1842: 33.

Si? (Ghiliani 1842).

Trogoderma variabile Ballion, 1878

Trogoderma variabile Ballion, 1878: Nicoli Aldini 1998: 115, Háva 2003: 141, Háva 2007: 320.

Trogoderma variabile Ballion: Nicoli Aldini 2003a: 156, Nicoli Aldini 2003b: 126–127.

N (Nicoli Aldini 1998, 2003a, 2003b), Sa (Zhantiev 2011a). Italy (Háva 2003, 2007). Italian mainland (Zhantiev 2011a).

A new checklist of the Dermestidae of Italy

The present-day dermestid fauna of Italy is summarized in the checklist below (Table 1).

The following abbreviations are used: AFR = Afrotropical; ALAP = Alpino-Apenninic endemic; ALPC = Central-Alpine endemic; APPE = Apenninic endemic; AUST = Australian; CAE = Centralasiatic-European; CEM = Centralasiatic-Europeo-Mediterranean; CEU = Central European; COS = Cosmopolitan; EME = E-Mediterranean; EUM = Europeo-Mediterranean; EUR = European; IT = Italy and/or Italian mainland without further details; (i) = introduced to Europe; MED = Mediterranean; N = northern Italy (Trentino-Alto Adige, Venetia, Friuli-Venezia Giulia, Lombardy, Piedmont, Aosta Valley, Liguria and Emilia-Romagna); NAF = N-African; OLA = Holarctic; PAL = Palearctic; S = peninsular Italy (the remainig of continental Italy); Sa = Sardinia and small circum-Sardinian islands; SCO = Subcosmopolitan; SEU = S-European; Si = Sicily and small circum-Sicilian islands; SICI = Sicilian endemic; SIE = Sibero-European; TEM = Turano-Europeo-Mediterranean; TUE = Turano-European; TUM = Turano-Mediterranean; WME = W-Mediterranean.

Discussion

The updated checklist of Italian Dermestidae includes 95 species, moreover Anthrenus (Anthrenus) pimpinellae occurs with two subspecies, but the taxonomic status of Anthrenus (Anthrenus) pimpinellae isabellinus needs to be clarified (cf. Háva and Nardi 2004). This total number is very probably still provisional since this family has been little studied in Italy, and the occurrence of further species recently described (e.g. Zhantiev 2001) or resurrected is possible. The previous checklist (Audisio et al. 1995), included only 81 species. Considering the four adopted geographic divisions, the relatively high species richness in the peninsular region (67) is unsurprising (cf. Minelli et al. 2002).

Some of the above records come from the following two protected areas of southern peninsular Italy: the Gargano National Park (Apulia) and the Pollino National Park (Basilicata and Calabria). 24 species of Dermestidae are now known from the former Park, while 23 species are now known from the Pollino National Park, but only 15 species are common to both (cf. Holdhaus 1911, Panganetti-Hummler 1918, Angelini 1987, 1996b, Háva and Nardi 2004). 24 and 23 species represent 37% and 34% of those currently recorded with certainty from peninsular Italy. These values confirm the importance of these Parks for insects (cf. Holdhaus 1911, Palm 1939, Grandi 1956, Angelini 1986, 1987, Rusek and Stumpp 1988, Agostini and Scali 1989, Rivosecchi 1989, Merz 1993, Angelini 1996b, Ruffo and Stoch 2006) and for biodiversity conservation.

Acknowledgments

We are indebted to Claudio Angeloro (Corpo Forestale dello Stato - Ufficio Territoriale per la Biodiversità di Foresta Umbra, Italy) for the assistance in Apulia during the research of the CNBFVR; Daniele Birtele (CNBFVR), Paolo Cornacchia (Porto Mantovano, Italy), Carmine Esposito (Velletri, Italy), Gianluca Scaglioni (Porto Mantovano, Italy) and Ilaria Toni (CNBFVR) for the collection of specimens; to Luca Bartolozzi (Museo di Storia Naturale, Sezione di Zoologia, Università di Firenze, Florence, Italy) for the loan of specimens from the collections under his care; to Franco Mason (Verona, Italy) for his constructive comments on a previous version of this paper; to Ottó Merkl (Hungarian Natural History Museum, Budapest, Hungary) for the information on the types of Attagenus calabricus; to Miloslav Rakovič (Prague, Czech Republic) and to Jonathan Cooter (Oxford, England) for the linguistic revision of the manuscript. We thank Riaan Stals (Pretoria, South Africa) and an anonymous referee for their reviews.

References
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