This genus is very similar to Pangia Wang & Ren, 2012 but it can be distinguished from the latter as follows: mandible reduced, apical tooth erect (Fig. 4), penis guide usually simple (Figs 25, 33, 41), ovipositor elongate-oval and without styli, spermatheca composed of two or three globular parts (Figs 10–11). In Pangia, the mandible is normal, apical tooth is bent, penis guide is strongly asymmetrical and complex, ovipositor is triangularly elongate and usually bearing short styli, and spermatheca has a large part and a small process.

This genus is also similar to Catanella Miyatake, 1961, but it can be distinguished from the latter as follows: mandible reduced with erect apical tooth (Fig. 4), antenna with 9 antennomeres, with antennomere 3 strongly triangular (Fig. 5). In Catanella, the mandible is normal, apical tooth is bent, antennae with 8 antennomeres, with antennomere 3 elongate and not expanded.

Body minute, hemispherical with head in repose drawn into prothorax and closely fitting ventrally against prominent prosternal lobe (Fig. 1); dorsum glabrous, pronotum and elytral outer margins with sparse long setation (Figs 12–20). Head transverse, ventrally flattened with clypeal region prominent anteriorly (Fig. 2); frontoclypeus deeply emarginated around exposed antennal insertions. Mandible reduced, apical tooth erect (Fig. 4). Antenna with 9 antennomeres; antennomere 1 stout, antennomere 2 globular and smaller than 1, antennomere 3 strongly triangular; club, oval and flat with apex angulate (Fig. 5). Terminal maxillary palpomere always longer than wide, barrel-shaped, truncate at apex (Fig. 6).

Pronotum strongly transverse, anterior corner rounded. Scutellum relatively large, triangular. Elytra usually smooth without visible punctures. Wings with greatly reduced venation. Prosternum strongly prominent medially forming a broad lobe concealing mouthparts from below; prosternal process subtruncate apically, broad, without carinae (Fig. 3). Mesoventrite very short and broad. Metaventrite large and broad, surface shining and glabrous (Fig. 1). Epipleuron moderately narrow, incomplete, reaching 2/3 of elytral length, with clearly delimited cavities to accommodate apices of meso- and metafemora. Abdomen with 5 ventrites (Fig. 7), ventrite 1 and 5 much longer than 2–4; hind margin of terminal ventrite rounded and smooth. Abdominal postcoxal lines incomplete, reaching lateral margin of ventrite, without associated pits or pores. Femora, especially profemur, broad, flat, closely fitting into depressions on ventral surface, protecting tibiae and tarsi from below; meso- and metatibiae conspicuously protuberant externally beyond middle, usually triangular; tarsus with 3 (Figs 8–9), rarely 4 tarsomeres.

Male genitalia: tegmen strongly asymmetrical, parameres extremely short or distinctly reduced sparsely setose apically (Figs 24–25). Female genitalia: ovipositor oval, without styli; spermatheca small and well sclerotised (Figs 10–11).

China, India, Japan, Mayotte Island (Indian Ocean), Sri Lanka, Vietnam.

This species can be distinguished from other Microserangium species by its yellow or burgundy elytral disk with dark brown margins and its penis guide acutely triangular (Figs 12, 25).

TL: 1.32–1.42mm, TW: 1.20–1.30mm, TH: 0.76–0.86mm, TL/TW: 1.09–1.10; PL/PW: 0.38–0.40; EL/EW: 0.88–0.89; HW/TW: 0.46; PW/TW: 0.73.

Body shiny and glabrous (Fig. 12). Head brown, frons yellowish brown. Pronotum brown to black, scutellum dark brown. Elytra with disk yellow to burgundy and margins dark brown. Underside dark red. Legs yellowish brown, tibiae and tarsi yellow.

Head transverse and ventrally flattened; frontal punctures large and sparsely distributed, with short sparsely distributed setae on frons; eyes round, large and coarsely faceted, widest interocular distance 0.39× width of head. Antennal club oval and flat, apex rounded (Fig. 27).

Pronotum transverse, anterior corners rounded. Pronotal disk with large and fine punctures both associated with long sparsely distributed setae, the large punctures distinctly larger than those on frons, 0.5–3.0 diameters apart. Elytra smooth and shiny with sparsely distributed long setae along margins, punctures inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat, weakly furrowed. Metaventrite shiny and glabrous, punctures fine and sparsely distributed, 1.0–2.0 diameters apart. Meso- and metatibiae angulate externally beyond middle, almost triangular; tarsus with 3 tarsomeres (Fig. 28).

Male genitalia. Penis strongly curved along entire length, apex narrowed and acicular, penis capsule with short outer arm and indistinct inner one (Figs 22–23). Tegmen slender and strongly asymmetrical. Penis guide in lateral view slender, almost straight, pointed apically (Fig. 24); in ventral view acutely triangular (Fig. 25). Parameres distinct, small, about half as long as penis guide.

Holotype ♂: China, Yunnan: Menglun, Xishuangbanna, 21°55.27'N, 101°16.64'E, ca 550m, 21.viii.2005, Wang XM leg. Paratypes (8): Yunnan: 1♀, Mengxing, Mengla, Xishuangbanna, 21°52.63'N, 101°27.07'E, ca 690m, 3.v.2008, Wang XM leg.; 1♀, Xishuangbanna Plant Park, Xishuangbanna, 21°56.05'N, 101°15.55'E, ca 550m, 22.viii.2005, Wang XM leg.; 1♀, Nuozadu, Simao, 22°34.0'N, 100°33.39'E, ca 750m, 12.v.2008, Wang XM leg.; 2♂♂, Dadugang, Puer, 22°22.35'N, 100°56.68'E, ca 950m, 5.v.2009, Ren SX & Wang XM leg.; 1♂1♀, Yaoqu, Mengla, 21°46.98'N, 101°29.34'E, ca 700m, 7.v.2009, Wang XM leg.; 1♂, Daheishan, Jiangcheng, 22°33.74'N, 101°50.87'E, ca 1300m, 17.v.2009, Wang XM leg.

China (Yunnan).

The specific epithet is formed from the Latin adjective erythrinus red colored, referring to the elytral disk being of yellow to burgundy color.

This species can be distinguished from other Microserangium species by its relatively small body, dark brown dorsum (Fig. 13), fine and sparse pronotal punctation, rather small penis capsule (Fig. 30), and rather wide penis guide (Fig. 33).

TL: 1.15–1.29mm, TW: 1.02–1.15mm, TH: 0.63–0.69mm, TL/TW: 1.11–1.13; PL/PW: 0.34–0.44; EL/EW: 0.89–0.90; HW/TW: 0.45; PW/TW: 0.77.

Body shiny and glabrous (Fig. 13). Dorsum uniformly dark brown. Head brown. Underside reddish brown, legs yellowish brown, tibiae and tarsi yellow.

Head transverse and ventrally flattened; frons with four to six medium punctures and many fine punctures, and several long widely separated setae; eyes round, moderately large and coarsely faceted, widest interocular distance 0.57× width of head. Antennal club oval and flat, apex blunt (Fig. 35).

Pronotum transverse, anterior corners inconspicuous and blunt, glabrous, punctures fine and sparsely distributed, associated with long sparsely distributed setae. Elytra smooth and shiny, with sparse row of long setae along margins, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat, weakly furrowed. Metaventrite shiny and glabrous, punctures in center fine, 1.0–3.0 diameters apart. Meso- and metatibiae angulate externally beyond middle, almost triangular; tarsus with 3 tarsomeres (Fig. 36).

Male genitalia. Penis strongly curved along entire length, apex narrowed and pointed, penis capsule small (Figs 30–31). Tegmen slender and strongly asymmetrical. Penis guide in lateral view slender, almost straight, apex pointed (Fig. 32), in ventral view flat and rather wide (Fig. 33). Parameres small and short, less than 1/3 length of penis guide.

Holotype ♂: China, Hainan: Wushi, 19°8.99'N, 109°53.84'E, ca 320m, 14.vii.1999, Peng ZQ leg.; Paratypes (2): 1♀, same data as holotype; 1♂, Nanbin, 18°21.37'N, 109°11.0'E, ca 10m, 24.iii.1998, Peng ZQleg.

China (Hainan).

The specific epithet is formed from the Latin adjective fuscus, referring to the elytral disk being uniformly dark brown.

This species can be distinguished from other Microserangium species by its male genitalia with relatively long parameres and tongue-shaped penis guide (Figs 40–41).

TL: 1.38–1.62mm, TW: 1.19–1.48mm, TH: 0.76–0.82mm, TL/TW: 1.09–1.17; PL/PW: 0.42–0.46; EL/EW: 0.89–1.00; HW/TW: 0.42; PW/TW: 0.73.

Body shiny and glabrous (Fig. 14). Dorsum uniformly dark brown to black. Head yellowish brown. Underside reddish brown, legs yellowish brown, tibiae and tarsi yellow.

Head transverse and ventrally flattened; frontal punctures medium-sized and densely distributed, 1.0–1.5 diameter apart, associated with several long sparsely distributed setae; eyes round, large and coarsely faceted, widest interocular distance 0.42× width of head. Antennal club oval and flat, apex angular (Fig. 43).

Pronotum transverse, anterior corners inconspicuous and blunt, Pronotal disk densley covered with large and fine punctures both associated with long sparsely distributed setae. Elytra smooth and shiny, with sparse row of long setae along margin, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat, weakly furrowed. Metaventrite shiny and glabrous, punctures at center fine but conspicuous, 2.0–4.0 diameters apart. Meso- and metatibiae angulate externally beyond middle, almost triangular; tarsus with 3 tarsomeres (Fig. 44).

Male genitalia. Penis moderately curved, apex strongly narrowed and acicular, penis capsule with short outer arm and inconspicuous inner one (Figs 38–39). Tegmen slender and asymmetrical. Penis guide in lateral view slender, almost straight, apex pointed (Fig. 40), in ventral view flat and tongue-shape, apex slightly pointed (Fig. 41). Parameres moderately long, slightly less than ½ length of penis guide.

Holotype ♂: China, Hainan: Bawangling, 19°05.65'N, 109°6.73'E, ca 330m, 21.iii.1996, Peng ZQ leg. Paratypes (11): Hainan: 5♂♂, same data to holotype; 2♂♂, Wuzhishan, 18°47.0'N, 109°31.98'E, ca 650m, viii.1995, Peng ZQleg.; 1♂, Limushan, 19°16.08'N, 109°47.32'E, ca 280m, 21.iv. 1996, Peng ZQleg.; 1♂, Shijing, Diaoluoshan, 18°56.15'N, 109°56.90'E, ca 200m, ix. 1995, Peng ZQleg.; 1♀, Wuzhishan, 18°47'N, 109°31.98'E, ca 650m, 22.xi.1991, Peng ZQ leg.; 1♂, Limushan, 19°16.08'N, 109°47.32'E, ca 280m, 22.vii.2006, Wang XMleg.

China (Hainan).

The specific epithet is formed from the Latin adjective glossoides, referring to the tongue-shaped penis guide.

This species is similar to Microserangium glossoides, from which it differs in having a relatively large outer arm of the penis capsule, small parameres, and narrow and curved penis guide (Figs 46–49). In Microserangium glossoides, the penis capsule is inconspicuous, the parameres are distinctly longer than in Microserangium shennongensis, and the penis guide is tongue-shaped (Figs 38–41).

TL: 1.25–1.35mm, TW: 1.09–1.15mm, TH: 0.66–0.69mm, TL/TW: 1.15–1.17; PL/PW: 0.42–0.44; EL/EW: 0.94–1.11; HW/TW: 0.42; PW/TW: 0.73.

Body shiny and glabrous (Fig. 15). Dorsum uniformly black. Head reddish brown. Underside dark brown, legs brown, tibiae and tarsi yellow.

Head transverse and ventrally flattened; frontal punctures large, conspicuous and densely distributed, 0.5–1.0 diameter apart, associated with long sparsely distributed setae; eyes round, large and coarsely faceted, widest interocular distance 0.50× width of head. Antennal club oval and flat, apex truncated (Fig. 51).

Pronotum short and strongly transverse, anterior corners inconspicuous and blunt. Pronotal disk with densely distributed large and fine punctures both associated with long sparsely distributed setae. Elytra smooth and shiny, with sparse row of long setae along margins, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat, weakly furrowed. Metaventrite shiny and glabrous, punctures fine and densely distributed, 1.5–2.0 diameters apart. Meso- and metatibiae weakly angulate externally beyond middle; tarsus with 3 tarsomeres (Fig. 52).

Male genitalia. Penis strongly curved along entire length, apex narrowed and acicular, penis capsule with short but distinct outer arm and small inner one (Figs 46–47). Tegmen rather slender and strongly asymmetrical. Penis guide in lateral view slender, outer margin arcuate, apex pointed (Fig. 48), in ventral view slender and moderately curved, apex pointed (Fig. 49). Parameres inconspicuous, small.

Holotype ♂: China, Hunan: Shennong Valley National Forest Park, Yanling, 26°29.95'N, 114°0.18.98'E, ca 800m, 9.x.2010, Wang XM leg. Paratypes (4): Hunan: 1♂3♀♀, same data as the holotype.

China (Hunan).

The specific epithet refers to the Shennong Valley National Forest Park, the type locality of this ladybird.

The male genitalia of this species are similar to Microserangium erythrinum, but this species can be distinguished from the latter by its uniformly dark elytra and semilunate penis guide (Figs 16, 56–57). In Microserangium erythrinum, theelytral disk is yellow or burgundy with dark brown margins and the penis guide is acutely triangular (Figs 12, 24–25).

TL: 1.42–1.55mm, TW: 1.24–1.35mm, TH: 0.76–0.86mm, TL/TW: 1.14–1.15; PL/PW: 0.40–0.44; EL/EW: 0.93–0.98; HW/TW: 0.44; PW/TW: 0.73.

Body shiny and glabrous (Fig. 16). Dorsum uniformly black. Head black, clypeus brown. Underside black, legs dark brown, tibiae and tarsi yellow.

Head transverse and ventrally flattened; frontal punctures large and densely distributed, 0.3–1.0 diameter apart, associated with long sparsely distributed setae; eyes round, large and coarsely faceted, widest interocular distance 0.50× width of head. Antennal club oval and flat, apex blunt (Fig. 59).

Pronotum short and strongly transverse, anterior corner inconspicuous and blunt, mat and shagreened. Pronotal disk with densely distributed large and fine shallow punctures both associated with long sparsely distributed setae. Elytra smooth and shiny, with sparse row of long setae along margins, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat weakly furrowed. Metaventrite shiny and glabrous, punctures fine and sparsely distributed, 2.0–3.0 diameters apart. Meso- and metatibiae strongly angulate externally beyond middle; tarsus with 3 tarsomeres (Fig. 60).

Male genitalia. Penis strongly curved along entire length, apex narrowed and acicular, penis capsule with large outer arm and inconspicuous inner one (Figs 54–55). Tegmen rather slender and strongly asymmetrical. Penis guide in lateral view small, almost straight, apex pointed (Fig. 56), in ventral view small and semilunate (Fig. 57). Parameres conspicuous.

Holotype ♂: China, Hainan: Bawangling, 19°05.65'N, 109°6.73'E, ca 330m, 21.iii.1996, Peng ZQ leg. Paratypes (12): Hainan: 4♂♂1♀, same data to holotype; 1♂1♀, Bawangling, 19°05.65'N, 109°6.73'E, ca 330m, 3.ix.1998, Peng ZQleg.; 1♂, Bawangling, 19°05.65'N, 109°6.73'E, ca 330m, 20.iv.2000, Peng ZQleg.; 1♂1♀, Diaoluoshan, 18°56.15'N, 109°56.90'E, ca 200m, 26.vii.2006, Wang XM leg.; 1♂, Diaoluoshan, 18°56.15'N, 109°56.90'E, ca 200m, 7.v.2005, Wang XM leg.

China (Hainan).

The specific epithet is formed from the the Latin adjective semilunatus, referring to the semilunate penis guide.

This species can be distinguished as follows: frontal punctures fine and sparsely distributed, pronotum densely covered with large punctures associated with long sparsely distributed setae, and punctures in central part of metaventrite moderately large and densely distributed. The penis guide of the male genitalia is also unique (Fig. 65).

TL: 1.52–1.62mm, TW: 1.35–1.45mm, TH: 0.86–0.89mm, TL/TW: 1.11–1.12; PL/PW: 0.44–0.45; EL/EW: 0.90–0.91; HW/TW: 0.43; PW/TW: 0.70.

Body shiny and glabrous (Fig. 17). Dorsum uniformly black. Head dark brown, frons brown. Underside dark brown, legs reddish brown, tibiae and tarsi yellow.

Head transverse and ventrally flattened; frontal punctures fine and sparsely distributed, 1.0–2.0 diameters apart, associated with long sparsely distriuted setae, eyes round, large and coarsely faceted, widest interocular distance 0.47× width of head. Antennal club oval and flat, apex blunt (Fig. 67).

Pronotum strongly transverse, anterior corners inconspicuous and blunt, mat and shagreened. Pronotal disk with densely distributed large punctures associated with long sparsely distributed setae. Elytra smooth and shiny, with long sparsely distributed setae along margins, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat weakly furrowed. Metaventrite shiny and glabrous, with densely distributed medium size punctures at center, 1.0–2.0 diameters apart. Meso- and metatibiae angulate externally beyond middle; tarsus with 3 tarsomeres (Fig. 68).

Male genitalia. Penis strongly curved, apex narrowed and pointed, penis capsule with conspicuous outer arm (Figs 62–63). Tegmen slender and strongly asymmetrical. Penis guide in lateral view slender, straight, apex pointed (Fig. 64), in ventral view acute triangular, apex pointed (Fig. 65). Parameres conspicuous, small, short, rectangular with penis guide (Fig. 64).

China, Hainan: 2♂♂, Tianchi, Jingfengling, 18°44.42'N, 108°51.80'E, ca 820m, viii.1995, Peng ZQ leg. 2♂♂, Xinan, Diaoluoshan, 18°56.15'N, 109°56.90'E, 18.ix.1995, Peng ZQ leg.; 1♂, Bawangling, 19°05.65'N, 109°6.73'E, ca 330m, 5.v.2005, Wang XM leg.

China (Hainan).

This species is similar to Microserangium okinawense, but can be separated from the latter by the inconspicuous inner arm of the penis capsule and triangular penis guide (Figs 70–73). In Microserangium okinawense, the inner arm of the penis capsule is small but distinct, and the penis guide is almost triangular with inner and outer margins arcuate (Figs 78–81).

TL: 1.19–1.35mm, TW: 1.02–1.15mm, TH: 0.66–0.76mm, TL/TW: 1.16–1.17; PL/PW: 0.38–0.44; EL/EW: 0.87–1.00; HW/TW: 0.43; PW/TW: 0.71.

Body shiny and glabrous (Fig. 18). Dorsum uniformly black, anterior corner of pronotum yellowish brown. Head dark brown, except frons medium brown. Underside dark brown, legs yellowish brown, tibiae and tarsi yellow.

Head transverse and ventrally flattened; frontal punctures fine and sparsely distributed, 1.0–2.0 diameters apart, with long sparsely distributed setae; eyes round, large and coarsely faceted, widest interocular distance 0.40× width of head. Antennal club oval and flat, apex blunt (Fig. 75).

Pronotum strongly transverse, anterior corners inconspicuous and blunt, mat and shagreened. Pronotal disk with densely distributed large and fine punctures both associated with long sparsely distributed setae, the finer punctures slightly deeper than the large ones. Elytra smooth and shiny, with long sparsely distributed setae along margins, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat, weakly furrowed. Metaventrite shiny and glabrous, punctures fine and sparsely distributed, 1.5–3.0 diameters apart. Meso- and metatibiae distinctly angulate externally beyond middle, almost triangular; tarsus with 3 tarsomeres (Fig. 76).

Male genitalia. Penis strongly curved along entire length, apex narrowed and pointed, penis capsule with large outer arm and inconspicuous inner one (Figs 70–71). Tegmen slender and asymmetrical. Penis guide in lateral view slender and straight, apex pointed (Fig. 72), in ventral view flat and triangular, apex pointed (Fig. 73). Parameres conspicuous, small (Fig. 72).

Holotype ♂: China, Guangxi: Pinglongshan, Fulong, Shangsi, 21°49.88'N, 107°56.79'E, ca 160m, 29.vii.2005, Wang XM leg. Paratypes (28): Guangxi: 1♂4♀♀, same data to holotype; 6♂♂3♀♀, Fulong, Shiwandashan, Shangsi, 21°49.88'N, 107°56.79'E, ca 160m, 7.xi.2004, Wang XM et al.leg.; 2♂♂11♀♀, Hongqilinchang, Shiwandashan, Shangsi, 21°52.79'N, 107°55.05'E, ca 900m, 7.xi.2004, Wang XM et al.leg.; Guizhou: 1♂, Xianheping, Anlong, 24°58.66'N, 105°36.45'E, ca 1500m, 12.ix.2007, Wang XM leg.

China (Guangxi, Guizhou).

The specific epithet is formed from the Latin adjective deltoides, referring to the triangular penis guide.

This species is close to Microserangium deltoides. The differences between these two are discussed in the diagnosis of Microserangium deltoides.

TL: 1.35–1.42mm, TW: 1.15–1.25mm, TH: 0.76–0.79mm, TL/TW: 1.13–1.17; PL/PW: 0.40–0.42; EL/EW: 0.95–1.00; HW/TW: 0.42; PW/TW: 0.68.

Body shiny and glabrous (Fig. 19). Dorsum uniformly dark brown to black. Head reddish brown. Underside reddish brown, legs yellowish brown, tibiae and tarsi yellow.

Head transverse and ventrally flattened; frontal punctures fine and inconspicuous, with long sparsely distributed setae; eyes round, large and coarsely faceted, widest interocular distance 0.44× width of head. Antennal club oval and flat, apex blunt (Fig. 83).

Pronotum short and strongly transverse, anterior corners inconspicuous and blunt, mat and shagreened. Pronotal disk with densely distributed large and fine shallow punctures both associated with long sparsely distributed setae. Elytra smooth and shiny, with sparse row of long setae along margins, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat weakly furrowed. Metaventrite shiny and glabrous, punctures in center fine and conspicuous, 1.0–2.0 diameters apart. Meso- and metatibiae protuberant externally beyond middle, almost triangular; tarsus with 3 tarsomeres (Fig. 84).

Male genitalia. Penis strongly curved along entire length, apex narrowed and pointed, penis capsule conspicuous (Figs 78–79). Tegmen rather slender and strongly asymmetrical. Penis guide in lateral view slender, straight, widest in apical 1/3, narrowed at base, apex pointed (Fig. 80); in ventral view flat and shovel-shaped, apex slightly blunt (Fig. 81). Parameres conspicuous, small (Fig. 80).

China, Taiwan: 2♂♂1♀, Pingdong, Shuangliu, 22°13.07'N, 120°47.77'E, 200–400m, 21. X. 2012, S. Ren SX et al. leg.; 1♂1♀, Jiaxian & Tengzhi, 23°4.38'N, 120°36.94'E, 370m-1450m, 25. X. 2012, S. Ren SX et al. leg.; Hainan: 4♂♂, Wufenchang, Limushan, 19°16.23'N, 109°47.48'E, ca 280m, ix.1996, Peng ZQ leg. 2♂♂3♀♀, Bawangling, 19°05.65'N, 109°6.73'E, ca 330m, 5.v.2005, Wang XMleg.; 1♂, Yinggeling, 19°02.25'N, 109°33.85'E, ca 830m, 23.xi.1997, Peng ZQleg.; 1♀, Yinggeling, 19°02.25'N, 109°33.85'E, ca 830m, 8.v.2005, Wang XM leg.; 2♀♀, Limushan, 19°16.08'N, 109°47.32'E, ca 280m, 22.vii.2006, Wang XM leg.

China (Hainan, Taiwan).

This species is similar to Microserangium bacthaiensis Hoàng, 1978 in morphological characters and male genitalia, but it can be distinguished by the different shape of the penis guide.

TL: 1.25–1.38mm, TW: 1.12–1.22mm, TH: 0.76–0.82mm, TL/TW: 1.12–1.14; PL/PW: 0.42–0.44; EL/EW: 0.94–0.95; HW/TW: 0.43; PW/TW: 0.76.

Body shiny and glabrous (Fig. 20). Dorsum uniformly black. Head brown, except frons yellowish brown. Underside dark brown, except prosternum brown. Legs yellowish brown, tibiae and tarsi yellow (Fig. 1).

Head transverse and ventrally flattened; frontal punctures inconspicuous and sparsely distributed, with long sparsely distributed setae (Fig. 2); eyes round, large and coarsely faceted, widest interocular distance 0.44× width of head (Fig. 2). Antennal club, oval and flat, apex angular (Fig. 5).

Pronotum short and strongly transverse, with anterior corners inconspicuous and blunt, mat and shagreened. Pronotal disk with densely distributed large and fine punctures both associated with long sparsely distributed setae, punctures inconspicuous. Elytra smooth and shiny, with several long setae along margins, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat, weakly furrowed (Fig. 3). Metaventrite shiny and glabrous, punctures fine and sparsely distributed, 3.0–4.0 diameters apart. Meso- and metatibiae protuberant externally beyond middle; tarsus with 3 tarsomeres (Fig. 9).

Male genitalia. Penis strongly curved along entire length, apex strongly narrowed and acicular, penis capsule with large outer arm and inconspicuous inner one (Figs 86–87). Tegmen rather slender and strongly asymmetrical. Penis guide in lateral view slender, straight, apex pointed (Fig. 88), in ventral view flat and triangular, apex finger-shaped (Fig. 89). Parameres conspicuous, small (Fig. 88).

Holotype ♂: China, Yunnan: Dadugang, Puer, 22°22.35'N, 100°56.68'E, ca 950m, 26.iv.2008, Wang XM leg. Paratypes (20): Yunnan: 3♂♂2♀♀, same data to holotype; 2♂♂2♀♀, Tongbiguan, Nabang, Yingjiang, 24°37.86'N, 97°34.75'E, ca 1000m, 22–23.v.5.2008, Wang XM et al.leg.; 2♂♂5♀♀, Ganlongjing, Lianhuatan, Hekou, 22°56.59'N, 103°31.68'E, ca 710m, 20.v.2009, Wang XM et al. leg.; 1♀, Yaoqu, Mengla, 700m, 7.v.2009, Wang XM leg.; 1♂, Ainiguzhai, Jinghong, 21°54.67'N, 101°10.31'E, ca 660m, 11.v.2009, Wang XM leg.; 1♂, Lianhuatan, Hekou, 22°53.86'N, 103°34.04'E, ca 900m, 22.iv.2008, Wang XM leg.; 1♂, Daheishan, Jiangcheng, 22°33.62'N, 101°50.16'E, ca 1240m, 17.v.2009, Wang XM leg.

China (Yunnan).

The specific epithet is formed from the Latin adjective dactylicus, referring to the finger-shaped apex of the penis guide.