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The Agrilus occipitalis species–group is redefined and diagnosed. Two species from this group, Agrilus auriventris Saunders, 1873 and Agrilus occipitalis (Eschscholtz, 1822), are known as the serious pests of cultivated Citrus trees. Overall twenty-three taxa are included in the Agrilus occipitalis species–group. A complete list of references, type material, species examined and distribution is given for each taxon. The host plants, adult occurrence and altitude range is cited for most taxa. Habitus of all taxa and aedeagi of available males are pictured. Images of primary type specimens are provided. A character state matrix table for diagnostic characters is given for all taxa to facilitate their determination.
The following new taxonomic and nomenclatural acts are proposed. New species:eight new species are described: Agrilus mucidus sp. n., Agrilus nebulosus sp. n., Agrilus picturatus sp. n., Agrilus pluvius sp. n., Agrilus pseudoambiguus sp. n., Agrilus tesselatus sp. n., Agrilus trepanatus sp. n. and Agrilus umrongso sp. n. Proposed synonyms: eight synonyms are proposed: celebicola Obenberger, 1924, syn. n. (synonym of occipitalis Eschscholtz, 1822); connexus Kerremans, 1900, syn. n. (synonym of occipitalis Eschscholtz, 1822); cupricauda Saunders, 1867 syn. n. (synonym of occipitalis Eschscholtz, 1822); evinadus Gory & Laporte, 1839, syn. n. (synonym of occipitalis Eschscholtz, 1822); nirius Obenberger, 1924 syn. reconfirmed (synonym of occipitalis Eschscholtz, 1822); oblatus Kerremans, 1900, syn. n. (synonym of occipitalis Eschscholtz, 1822); samoensis Blair, 1928, syn. n. (synonym of auriventris Saunders, 1873); tebinganus Obenberger, 1924, syn. n. (synonym of occipitalis Eschscholtz, 1822). New lectotype designations: six lectotypes are designated: Agrilus celebicola Obenberger, 1924; Agrilus cupricauda Saunders, 1867; Agrilus korenskyi Obenberger, 1923; Agrilus kurandae Obenberger, 1923; Agrilus nirius Obenberger, 1924; Agrilus nitidus Kerremans, 1898.
Buprestidae, Agrilini, Agrilus, new species, synonymy, lectotype designation, Citrus, pest
This publication presents the first comprehensive revision of the Agrilus taxa of the occipitalis species–group. The group was established and defined by
The state of species taxonomy has remained unrevised despite the fact that the two most serious pests in the citrus orchards Agrilus occipitalis (Eschscholtz, 1822) and Agrilus auriventris Saunders, 1873 belong to this group. Most of the species from the Agrilus occipitalis species–group are distributed in South and Eastern Asia but some spread well beyond this area: Agrilus diversornatus Jendek, 2011 to Russian Far East; Agrilus occipitalis and Agrilus biakanus Curletti, 2006 to Australasia. The occurrence of the chronic Citrus pest Agrilus auriventris in Polynesia and Agrilus occipitalis in Micronesia is most likely an introduction.
The format of the taxonomic part, style of the new species descriptions and morphological terms follow those used in
According to Article 74.7.3 of the
Abbreviations for collections
BMNH The Natural History Museum, London, United Kingdom
EJCB Collection of E. Jendek, Bratislava, Slovak Republic [currently in Ottawa, Canada]
IZAS Institute of Zoology, Academia Sinica, Beijing, China
MNHN Muséum national d’Histoire Naturelle, Paris, France
NHMB Naturhistorisches Museum, Basel, Switzerland
NMPC National Museum (Natural History), Prague, Czech Republic
NSMT National Science Museum (Natural History), Tokyo, Japan
USNM National Museum of Natural History, Washington D. C. , USA
ZIN Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia
Diagnosis. Medium (>5 mm) or large species (>10 mm), body shape cuneiform or parallel. Head usually impressed medially with curved sculpture at both sides; eyes small or moderate. Pronotum with anteromedial, posteromedial and lateral impressions; rarely without apparent impressions. Prehumerus carinal or filamentary; rarely tubercular or obsolete. Elytra monochromatic or with elytral apices carmine; apices separately arcuate, rarely separately subtruncate; elytral pubescence fasciate or ornamental, rarely absent. Prosternal process flat, sides subparallel or dilated, rarely narrowed. Sexual dimorphism not obvious except for the longer ventral pubescence in male.
Distribution.East, South and Southeast Asia, Indonesia, Australasia, Oceania.
Host plants. Adoxaceae (Sambucus), Rutaceae (Citrus, Euodia, Fagara, Zanthoxylum), Rosacae (Sorbaria).
Within the Agrilus occipitalis species–group seven subgroups can be recognized (Appendix):
A: elytral apices separately subtruncate
Agrilus tesselatus-subgroup
Species: Agrilus tesselatus sp. n.
Diagnostic characters: antennae very short, serrate from antennomere 5; pronotum obviously convex, without apparent impressions, maximum width of pronotum at posterior margin; groove on apex of last ventrite deeply sinuate; ovipositor square.
B: elytral apices separately subarcuate
B1: species with elytral apex pubescent and prosternal lobe predominantly emarginate.
Agrilus perroti-subgroup
Species: Agrilus perroti Descarpentries & Villiers, 1963, Agrilus umrongso sp. n. and Agrilus zanthoxylumi Li Meng Lou, 1989.
Diagnostic characters: body large or medium sized; head with deep medial longitudinal impressions; maximum width of pronotum at posterior margin, rarely at middle; elytral pubescence at least partly fasciate; apex of elytra pubescent and concolor with elytral disk; ovipositor prolonged.
Agrilus auroapicalis-subgroup
Species: Agrilus auroapicalis Kurosawa, 1957, Agrilus diversornatus Jendek, 2011, Agrilus ishigakianus Tôyama, 1985.
Diagnostic characters: body medium sized; head with deep medial longitudinal impressions; anterior pronotal lobe obvious; pronotum widest at posterior margin or at middle; prehumerus long, reaching to anterior pronotal angles, often joined with pronotal marginal carina; elytral pubescence at least partly fasciate; apex of elytra with strikingly different color than disk; prosternal lobe emarginate; apex of pygidium arcuate; groove on apex of last ventrite arcuate; ovipositor prolonged.
B2: species with prosternal lobe subtruncate or arcuate, elytral apex glabrous (except for Agrilus yamawakii) and prosternal process often dilated.
Agrilus inamoenus-subgroup
Species: Agrilus inamoenus Kerremans, 1892, Agrilus mucidus sp. n., Agrilus pluvius sp. n., Agrilus tonkineus Kerremans, 1895
Diagnostic characters: body medium or large, robust; head with deep medial longitudinal impressions; pronotum widest at posterior margin or at middle; elytral pubescence mosaic or at least partly mosaic; apex of pygidium angulate or with short protrusion, rarely arcuate; apex of last ventrite smooth, without medial carinula; ovipositor square.
Agrilus ambiguus-subgroup
Species: Agrilus ambiguus Kerremans, 1895, Agrilus picturatus sp. n., Agrilus pseudoambiguus sp. n.
Diagnostic characters: body medium sized, slender; head with medium or deep medial impressions; antennae long; pronotum widest in middle, rarely at anterior margin; elytral pubescence mosaic or at least partly mosaic; pygidium angulate, with protrusion or long spine; ovipositor prolonged.
Agrilus auriventris-subgroup
Species: Agrilus alesi Obenberger, 1935, Agrilus auriventris Saunders, 1873, Agrilus nebulosus sp. n., Agrilus trepanatus sp. n., Agrilus yamawakii Kurosawa, 1957.
Diagnostic characters: body medium or large, robust; head with or without medial impression; pronotum widest at anterior margin or in middle, rarely at posterior margin; elytral pubescence fasciate or ornamental, rarely missing; elytral apex concolor with elytral disk, rarely indistinctly carmine; ovipositor prolonged.
Agrilus occipitalis-subgroup
Species: Agrilus biakanus Curletti, 2006, Agrilus horniellus Obenberger, 1935, Agrilus occipitalis (Eschscholtz, 1822), Agrilus sordidulus Obenberger, 1916.
Diagnostic characters: body medium, robust; head rarely without medial impression; pronotum widest in middle; elytral pubescence fasciate or ornamental, rarely missing; pygidium arcuate or with short protrusion, never with long spine; ovipositor prolonged.
http://species-id.net/wiki/Agrilus_alesi
Fig. 11 (habitus) ; Fig. 31 (aedeagus) ; Fig. 41 (pygidium)Agrilus alesi Obenberger, 1935. Type locality. Loo-choo: Okinawa. Lectotype designated by
Agrilus sacchari Obenberger, 1940. Type locality. Insulae Loo-Choo; Okinawa. Lectotype designated by
Agrilus aritai Tôyama, 1985. Type locality. Hirano, Ishigakijima Is. Holotype exa-mined by
Size 7.2–8.5 mm. Agrilus alesi can be distinguished from closely related Agrilus auriventris by the smaller size; by the more transverse pronotum with the maximum width in the middle; by the obvious anterior pronotal lobe and by the pygidium with the long spine on apical margin (Fig. 41). See also Appendix.
CHINA: 2 ♂, 1 ♀ (USNM): “CB [citrus borer?] adults, Chengtu, China, v.28.-vi.5. ‘[19]39, Kovlieu [Note: Chengtu may refer to many po- pulated places in several provinces of China”. JAPAN: Kyushu: 23 (USNM): “Kagoshima, Kyushu, 5-v-1940, F. Yano”; 2 ♂, 1 ♀ (USNM): “Kagoshima: Kyushu, Japan, 5-v-1940 (F. Yano)”. For further records see
5–6–7–9.
Citrus:
CHINA[provincial level unknown]. JAPAN: Kyushu; Ryukyu islands (Okinawa incl.).
http://species-id.net/wiki/Agrilus_ambiguus
Fig. 8 (habitus) ; Fig. 28 (aedeagus)Agrilus ambiguus Kerremans, 1895. Type locality. Sikkim: Kurseong. Lectotype designated by
Size 6.7–10 mm. Agrilus ambiguus differs from the closely related Agrilus picturatus sp. n. and Agrilus pseudoambiguus sp. n. by having the head only feebly impressed medially; by the black vertex and by having elytral apices concolor with elytral disk (See also Appendix).
INDIA: Assam: 1 ♀ (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso, 700m, 25°27'N, 92°43'E, 17.-25.v., Dembický & Pacholátko leg.”. Meghalaya: 7 (EJCB): “NE India, Meghalaya state, West Garo Hills, Nokrek Nat.Park, 9–17. V. 1996 alt.1100+150m, GPS N25°29.6', E90°19.5' (WGS 84), E. Jendek & O. Šauša leg.”; 7 (EJCB): “NE India, Meghalaya, 1400 m, Nokrek N. P. , 3 km S Daribokgiri, 25°27'N, 90°19'E, 26.iv.1999, Dembický & Pacholátko leg.”. 1 ♂ (EJCB): “NE India, Meghalaya, 1400 m, Nokrek N. P. , 3 km S Daribokgiri, 25°27'N, 90°19'E, 26.iv.1999, J. Rolčík leg.”. West Bengal: 2 (NMPC): “Darjeeling [labeled as pescheti Obnb.; nomen nudum]”. LAOS: Louang Namtha: 1 ♂, 2 ♀ (EJCB): “Laos, Louang Namtha pr., 21°09'N, 101°19'E, Namtha–Muang Sing, 5-31.v.1997, 900-1200 m, Vit Kubáň leg.”. Phongsali: 1 ♂ (EJCB): “Lao-N, Phongsaly prov., 21°41'-2'N, 102°06'–08'E, 28.v.-20.vi.2003 Phongsaly env., ~1500m, Vít Kubáň leg.”; 1 ♂ (EJCB): “Lao, Phongsaly prov. 21°41'N, 102°06'E Phongsaly env. 6–17.v.2004, 1500 m, P. Pacholátko leg.”.
4–5–6. Altitude range: 700–1500 m.
Unknown.
INDIA: Assam; Meghalaya; Sikkim; West Bengal. LAOS: Louang Namtha; Phongsali.
http://species-id.net/wiki/Agrilus_auriventris
Fig. 12 (habitus) ; Fig. 32 (aedeagus); Fig. 42 (pygidium)Agrilus auriventris Saunders, 1873. Type locality. Japan. Lectotype designated by
Agrilus graptelytrus Obenberger, 1914. Type locality. China, Tonkin: Phu-long-thuan. Lectotype designated by
Agrilus fleutiauxi Bourgoin, 1922. Type locality. not given [Note: Indo-Chine française is cited in the title of the publication]. Lectotype designated by
Agrilus pidjinus Obenberger, 1924. Type locality. China: Hong-Kong. Lectotype designated by
Agrilus samoensis Blair, 1928. Type locality. Upolu: Apia; Malololelei. Holotype (Fig. 61), ♀, (BMNH): “Type H. T. [p] [round label with red border] \ Samoan Is. [p] Upolu Malololelei 2000 ft, 28.xi.1924 [h] P. A. Buxton and G. H. Hopkins [p] \ Agrilus Samoensis Type Blr. [h] frt. K. G. Blair [p]”. Paratypes: 1 paratype (MNHN); 1 paratype (BPBM). Described from 4 specimens (holotype, paratypes).
Size: 5.0–8.8 mm. Agrilus auriventris can be distinguished from the most close Agrilus nebulosus sp. n. by the pygidium which is angulate or armed with a short protrusion (Fig. 42). See also Appendix.
CHINA: Guangxi: 1 (IZAS): “Guangxi, Longzhou, 140m, 1.v.1963, Y. Wang leg.”. VIETNAM: 1 (IZAS): “Tonkin, Hoa Binh, vii.1931, A. De Cooman leg.”. For further records see
4–5–6–7–8–10. Altitude range: 140–700 m.
Citrussp. For the detailed bibliography see
CHINA: Fujian; Guangdong; Guangxi; Hong Kong; Hubei; Hunan; Jiangxi; Sichuan; Taiwan; Zhejiang. JAPAN: Honshu; Kyushu. LAOS. MYANMAR. SAMOA. VIETNAM.
http://species-id.net/wiki/Agrilus_auroapicalis
Fig. 5 (habitus) ; Fig. 46 (Holotype)Agrilus auroapicalis Kurosawa, 1957. Type locality. Mt. Nanjin-zan, Formosa. Holotype (Fig. 46) examined by
Agrilus laurenconi Descarpentries & Villiers, 1963. Type locality. Tonkin: Hoa-Binh. Holotype (Fig. 55) examined by
Habitus of Agrilus: 1 Agrilus tesselatus sp. n – Holotype 2 Agrilus perroti Descarpentries & Villiers, 1963 3 Agrilus umrongso sp. n – Holotype 4 Agrilus zanthoxylumi Li Meng Lou, 1989 5 Agrilus auroapicalis Kurosawa, 1957 6 Agrilus auroapicalis ishigakianus Tôyama, 1985 7 Agrilus diversornatus Jendek, 2011 – Holotype 8 Agrilus ambiguus Kerremans, 1895 9 Agrilus picturatus sp. n – Holotype 10 Agrilus pseudoambiguus sp. n – Holotype 11 Agrilus alesi Obenberger, 1935 12 Agrilus auriventris Saunders, 1873.
Size 5.8–8.1 mm. Agrilus auroapicalis differs from the similar Agrilus diversornatus mainly by the larger eyes, by pronotum widest in the middle and by pronotal lobe obviously arcuate. See also Appendix.
CHINA: Guizhou: 1 ♀ (MNHN): “Kouy-Tchéou, R. P. J. R. Chaffanjon 1903”. Taiwan: 1 (EJCB): “Meiyuan, Nantou Hsien, Taiwan, 5.v.1993, Luo Chinchi leg.”. LAOS: Louangphrabang: 1 ♀ (EJCB): “Laos-N, 23.iv.1999, Louangphrabang prov. 20°42'N, 102°54'E, 25 km E Muang Ngoy, 1000 m, Vít Kubáň leg.”. VIETNAM: Gia Lai: 1 (EJCB): “S Vietnam, Gia Lai-Kon Tum pr., 5 km N Ankhé, 19.x.1979”; 1 (EJCB): “Vietnam, Gialai, Contum Tram Cap, 20.4.1995, Gorochov”.
4–5–10. Altitude range: 1000 m.
Unknown.
CHINA: Guizhou; Taiwan. LAOS: Louangphrabang. VIETNAM: Gia Lai; Hoa Binh.
http://species-id.net/wiki/Agrilus_auroapicalis_ishigakianus
Fig. 6 (habitus) ; Fig. 27 (aedeagus)Agrilus auroapicalis ishigakianus Tôyama, 1985. Type locality. Mt. Omotodake, Ishigaki-jima Isl. Types not examined. See Remarks. Described from 3 specimens (holotype, alotype, paratype).
Size 6.1 mm; it can be distinguished from Agrilus auroapicalis auroapicalis by having the body smaller, less produced apically; by the golden-brown dorsal color; by the ornamental elytral pubescence more extensive mostly along the suture and by having the color of elytral apices less contrasting to that of the nominal subspecies. See also Appendix.
JAPAN: Ryukyu islands: 1 ♂ (EJCB): “1996.4.19, Ishigaki Is., Okinawa, Ryukyu, K. Takahashi leg.”.
Euodia meliifolia:
JAPAN: Ryukyu islands (Okinawa incl.).
The type specimens of this taxon were not studied; they should be preserved in NSMT as stated by
http://species-id.net/wiki/Agrilus_biakanus
Fig. 47 (habitus of holotype)Agrilus biakanus Curletti, 2006. Type locality. Mokmer, Biak Isl., N. G. Type specimens were not examined. Image of the holotype (Fig. 47) was adopted from
Size: 6.9–9.2 mm. This species is distinctive by the glabrous elytra (
Unknown.
INDONESIA: Irian Jaya.
No specimens of this species were available for this study. The assignment of this taxon to Agrilus occipitalis species–group is based on the original description and on the image of aedeagus which is very similar to that of Agrilus occipitalis.
http://species-id.net/wiki/Agrilus_diversornatus
Fig. 7 (habitus of holotype)Agrilus diversornatus Jendek, 2011. Type locality. Eastern Russia, south Primorskiy kray, Lazovskii zapovednik, kordon Korpad’, 43°15'17"N, 134°07'59"E. Holotype (Fig. 7) examined by
Size 6.8–7.1 mm. From the similar Agrilus auroapicalis it differs mostly by the smaller eyes; by pronotum widest at the posterior margin and by the pronotal lobe being obviously angulate. See also Appendix.
Known only from type specimens.
Adult occurrence: 7–8.
Sorbaria:
RUSSIA: Primorskiy kray.
http://species-id.net/wiki/Agrilus_horniellus
Fig. 16 (habitus) ; Fig. 35 (aedeagus)Agrilus horniellus Obenberger, 1935. Type locality. See: Agrilus horni Théry, 1904. See: Agrilus horni Théry, 1904.
Agrilus horni Théry, 1904. Type locality. Nalanda. Lectotype designated by
Size: 6.5–7.3 mm. Agrilus horniellus differs from the very similar Agrilus occipitalis by having the apex of pygidium arcuate and the apex of elytra concolor with the elytral disk. See also Appendix.
SRILANKA:1 ♂ (EJCB): “Sri Lanka: Anu Distr., 6 miles south of Tantirimalai, 2000ft, 31 Oct 1976”; 1 ♀ (EJCB): “Ceylon, E. Prov., Pottuvil, 1-12/vii.-1983 Ole Mehl. leg.”; 1 ♀ (EJCB): “Ceylon, N. C. Prov., Anuradhapura, 22-26/vi.-1985, Ole Mehl. leg.”.
6–7–10. Altitude range: 610 m.
Unknown.
SRILANKA.
Agrilus horniellus may be conspecific with Agrilus occipitalis, but its original taxonomic concept was tentatively retained due to limited specimens available for examination.
http://species-id.net/wiki/Agrilus_inamoenus
Fig. 21 (habitus) ; Fig. 38 (aedeagus)Agrilus inamoenus Kerremans, 1892. Type locality. Carin Cheba, 900–1100 m. Lectotype designated by
Size 7.8–9.8 mm. Within the subgroup, Agrilus inamoenus is distinctive by the short antennae, by the prosternal lobe absent or vague and by the tubercular prehumerus. See also Appendix.
CHINA: 1 (IZAS): “[China] Shuangjiang, vi.1953 [in Chinese]”. Yunnan: 1 (IZAS): “Yunnan Xishuangbanna Xiaomengyang, 850m, 9.vii.1957, S. Y. Wang leg. [in Chinese]”; 3 (IZAS): “Yunnan Xishuangbanna Menghun, 750m, 2–7.vi.1958, S. Y. Wang leg. [in Chinese]”; 1 (IZAS): “Yunnan Xishuangbanna Damenglong, 650m, 5.v.1958, X. W. Meng leg. [in Chinese]”. THAILAND: Chaiyaphum: 1 (EJCB): “Thailand, Chaiyaphum Tat Tone NP, near stream, 15°58.771'N, 102°02.397'E, Malaise trap, 5–12.vii.2006, T. Jaruphan and O. Budsawong leg.”. Chiang Rai: 1 ♂ (USNM): “Khun Tan Mts, N Siam 3000 ft, HM. Smith May [19]33”. VIETNAM: Lam Dong: 1 (MNHN): “Djiring, Annam, H. Perrot”. For further re- cords see also
4–5–6–7. Altitude range: 420–1600 m.
Citrus:
CHINA: Fujian; Yunnan. LAOS: Borikhamxai; Khammouan; Louang Namtha; Louangphrabang; Savannakhet; Xaignabouri; Xiangkhoang. MYA-NMAR: Karen State. THAILAND: Chaiyaphum; Chiang Mai, Chiang Rai. VIETNAM: Binh Dinh; Gia Lai; Hoa Binh; Lam Dong; Son La.
urn:lsid:zoobank.org:act:1C67DFF8-5BF1-416C-9F7F-CD0F18B685C5
http://species-id.net/wiki/Agrilus_mucidus
Fig. 22 (habitus of holotype) ; Fig. 39 (aedeagus)BODY. Size: 9.6–10.7 mm (Holotype 10 mm).Shape: subparallel, Build: robust, Posterior tapering part: short with broad apex, Color (dorsally): unicolored, Sexual modifications in male: not apparent. HEAD.Size: very large, Medial impression: deep, Epistoma: raised above frons. Vertex: Shape: markedly convex, Sculpture elements: rugae, Sculpture shape: semispherical, Sculpture density: dense, Sculpture intensity: rough. Eyes: Size: small, Shape: markedly protruding head outline, Lower margin: in line or below with antennal socket, Medial orbit: converging ventrally or subparallel. Antennae: Length: long, Width: slender, Serration: from antennomere 4, Antennomere 7–10 (shape): with obvious collum. PRONOTUM.Shape: transverse, Sides: markedly arcuate or slightly arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe: Size: mode-rate, Shape: arcuate or subangulate, Position: at level with anterior angles. Posterior angles: Shape: acute or obtuse or rectangular, Apex: blunt or sharp. Disk: Conve-xity: flat, Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (intensity): shallow, Lateral impression (size): wide. Prehumerus: Development: filamentary, rarely carinal, Shape: bisinuate, Extent: to 1/3 of pronotal length, rarely to 1/2 of pronotal length, Anterior end: distant from lateral carina, Posterior end: joined with posterior angle or margin, Arc: moderate or obvious. Lateral carinae: Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of marginal carina. ELYTRA.Color: monochromatic, Humeral carina: absent. Apices: Arrangement: separate, Shape: arcuate. Pubescence: Color: monochromatic, Character: homogenous or with patches or spots of denser pubescence, Extent: entire ornamental with indication of fasciae or entire ornamental. Tomentum: Spots (pattern): postmedial only. STERNUM. Prosternal lobe: Size: large, Distal margin: arcuate. Prosternal process: Shape: subparallel, Sides: arcuate, Angles: obtuse, Angles (tips): blunt, Disc: flat. ABDOMEN.Tomentum: absent or present. Pygidium: Apical margin: angulate. Sternal groove: Extent: on three apical ventrites, Shape on the apex of last ventrite: arcuate. LEGS. Metatarsus: Size to metatibia: distinctly shorter than metatibia. Tarsomere 1: Size to following tarsomeres: longer than 2-3 but shorter than 2-4. GENITALIA. Aedeagus (Fig. 39): Symmetry: symmetric, Shape: widest in basal part, rarely subparallel, Modifications: apex of medial lobe sharply pointed. Ovipositor: Shape: square (uritiform).
From the very close Agrilus tonkineus, it can be distinguished by the flat pronotum; by the bi-sinuate prehumerus and by the presence of transverse tomentose strip at apical third of elytra. See also Appendix.
China, Hainan, Baihualing, 19.018, 109.836, altitude 300 m.
Holotype (Fig. 22), ♂, (EJCB): “China, Hainan, Baihualing, 19.018, 109.836, alt. 300 m, vi. 2008 [p]”. Paratypes: 1 paratype (EJCB), 3 paratypes (IZAS) from the same locality as holotype; 1 paratype (EJCB): “[China] Hainan Qiongzhong Baihuashan Mt, 27.v.1997, P. Y. Yu leg. [transcription from Chinese]”. 1 paratype (USNM), 1 paratype (EJCB): “Taichow, China, 1933”.
Habitus of Agrilus: 13 Agrilus nebulosus sp. n – Holotype 14 Agrilus trepanatus sp. n – Holotype 15 Agrilus yamawakii Kurosawa, 1957 16 Agrilus horniellus Obenberger, 1935 17 Agrilus occipitalis (Eschscholtz, 1822) – Laos, Vientiane, Ban Phabat 18 Agrilus occipitalis (Eschscholtz, 1822) – Papua New Guinea, Sideia island 19 Agrilus occipitalis (Eschscholtz, 1822) – Philippines, Palawan 20 Agrilus sordidulus Obenberger, 1916 21 Agrilus inamoenus Kerremans, 1892 22 Agrilus mucidus sp. n – Holotype 23 Agrilus pluvius sp. n – Holotype 24 Agrilus sordidulus Obenberger, 1916.
5–6 . Altitude range: 300 m.
Unknown.
CHINA:Hainan; Zhejiang.
The specific name is Latin adjective mucidus (moldy). It refers to the elytral tomentum of the species.
urn:lsid:zoobank.org:act:2DF8A3BE-19D6-40D4-B183-D3EE333BF5F5
http://species-id.net/wiki/Agrilus_nebulosus
Fig. 13 (habitus of holotype) ; Fig. 33 (aedeagus)BODY. Size: 6.4–7.5 mm (Holotype 6.5 mm).Shape: cuneiform, Posterior tapering part: long with narrow apex, Color (dorsally): unicolored, Sexual modifications in male: not apparent. HEAD. Medial impression: deep, rarely shallow, Epistoma: raised above frons, Vertex: Sculpture elements: rugae, Sculpture density: dense, Eyes: Size: moderate, Lower margin: in line or below with antennal socket, Medial orbit: converging ventrally, Antennae: Length: long, Width: slender, Serration: from antennomere 4. PRONOTUM. Shape: transverse, Sides: slightly arcuate, Maximal width: at anterior margin, Anterior margin: wider than posterior, Anterior lobe: Size: moderate, Shape: arcuate, Position: at level with anterior angles, Posterior angles: Shape: obtuse or rectangular, Apex: sharp, Disk: Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Prehumerus: Development: carinal, Shape: arcuate, Extent: to 1/3 of pronotal length, Anterior end: distant from lateral carina, Posterior end: joined with posterior angle or margin, Arc: moderate or weak, Lateral carinae: Convergence: moderately convergent, Junction: present, Narrowest point: at posterior 1/5-1/4 of marginal carina. ELYTRA. Color: monochromatic, Humeral carina: absent, Apices: Arrangement: separate, Shape: arcuate, Pubescence: Color: monochromatic, Extent: entire ornamental, rarely entire ornamental with indication of stripes. STERNUM. Prosternal lobe: Size: large, Distal margin: arcuate, Prosternal process: Size: wide, Shape: dilated, rarely subparallel, Sides: straight, Angles: acute, Angles (tips): blunt, Disc: flat, Projection (extend): distinctly beyond angles, Mesosternum: Mesosternal projection: flat. ABDOMEN. Tomentum: absent, Pygidium: Apical margin: arcuate, rarely angulate, Sternal groove: Extent: on all ventrites or on three apical ventrites, Shape on the apex of last ventrite: arcuate, rarely arcuately sinuate, Emargination (deepness): very shallow. LEGS. Metatarsus: Size to metatibia: distinctly shorter than metatibia, Tarsomere 1: Size to following tarsomeres: longer than 2-3 but shorter than 2-4 or subequal or longer than 2-4. GENITALIA. Aedeagus (Fig. 33): Symmetry: symmetric, Shape: widest in basal part, rarely subparallel, Modifications: apex of medial lobe sharply pointed, Ovipositor: Shape: markedly elongate.
Aedeagus of Agrilus: 25 Agrilus perroti Descarpentries & Villiers, 1963 26 Agrilus zanthoxylumi Li Meng Lou, 1989 27 Agrilus auroapicalis ishigakianus Tôyama, 1985 28 Agrilus ambiguus Kerremans, 1895 29 Agrilus picturatus sp. n – Holotype 30 Agrilus pseudoambiguus sp. n – Holotype 31 Agrilus alesi Obenberger, 1935 32 Agrilus auriventris Saunders, 1873 33 Agrilus nebulosus sp. n – Holotype 34 Agrilus yamawakii Kurosawa, 1957 35 Agrilus horniellus Obenberger, 1935 36 Agrilus occipitalis (Eschscholtz, 1822) – Philippines, Palawan 37 Agrilus sordidulus Obenberger, 1916 38 Agrilus inamoenus Kerremans, 1892 39 Agrilus mucidus sp. n – Holotype 40 Agrilus tonkineus Kerremans, 1895.Pygidium of Agrilus 41 Agrilus alesi Obenberger, 1935 42 Agrilus auriventris Saunders, 1873 43 Agrilus occipitalis (Eschscholtz, 1822) – Laos, Vientiane, Ban Phabat 44 Agrilus occipitalis (Eschscholtz, 1822) – Papua New Guinea, Sideia island 45 Agrilus occipitalis (Eschscholtz, 1822) – Thailand, Mae Hong Son, Pai.
From the close Agrilus auriventris, it can be distinguished by the generally smaller size and slender body; by the pygidium arcuate apically and by the the groove on the apex of last ventrite arcuate (rarely sinuate). See also Appendix.
North Vietnam, 21°35N, 106°30E, 52 km southwest of Lang Son, altitude 370 m.
Holotype (Fig. 13), ♂, (EJCB): “N Vietnam, 21°35N, 106°30E, 52 km SW of Lang Son, 27.iv.-6.v.1996, 370 m, Dembický and Pacholátko leg.”. Paratypes:1 ♂ paratype, 7 ♀ paratypes (EJCB) from the same locality as holotype.
5–6 . Altitude range: 300 m.
Unknown.
VIETNAM:Lang Son.
The specific name is Latin adjective nebulosus (misty, hazy, indefinite, obscure). It refers to the faint ornamental elytral pubescence of the species.
http://species-id.net/wiki/Agrilus_occipitalis
Figs 17–19 (habitus) ; Fig. 36 (Aedeagus); Figs 43–45 (pygidium) ; Fig. 60 (Lectotype)Buprestis occipitalis Eschscholtz, 1822. Type locality. Auf der Inßel [= Insel] Luzon, bei Manilla. Lectotype (Fig. 60) designated by
Agrilus evinadus Gory & Laporte, 1839. Type locality. Java. Lectotype (Fig. 51) designated by
Agrilus occipitalis Gory, 1841. Type locality. Iles Philippines. Lectotype designated by
Agrilus marmoreus Deyrolle, 1864. Type locality. I. Mysole et Batchian. Lectotype (Fig. 56) designated by
Agrilus cupricauda Saunders, 1867. Type locality. Penang. Lectotype by present designation (Fig. 50), ♀, (BMNH): “Type H. T. [p] [round label with red border] \ Penang [h] [oval blue label] \ Penang (Lamb.) Pascoe Coll. [p] \ Agrilus cupricauda Typ ES [h]”. Described from unknown number of syntypes.
Agrilus nitidus Kerremans, 1898. Type locality. Australie: Cocktown. Lectotype by present designation (Fig. 57), ♀, (BMNH): “Syn-Type [p] [round label with blue border] \ Cocktown Stauding [h] \ nitidus Kerr. Type [h] \ A. nitidus Kerrem. Australie [h] \
Agrilus connexus Kerremans, 1900. Type locality. not given [Sumatra, Hindrapoera is cited in the title and introductory text]. Holotype by monotypy (Fig. 49), ♂, (BMNH): “Type [p] [round label with red border] \ Sumatra Weyers [h] \ connexus Kerr. Type [h] \
Agrilus oblatus Kerremans, 1900. Type locality. Sumatra. Lectotype (Fig. 59) designated by
Agrilus korenskyi Obenberger, 1923. Type locality. Australia. Lectotype by present designation (Fig. 53), ♂, (NMPC): “Australia [h] \ Typus [p] [red label] \ A. Kořenskyi m. Type [h] Det. Dr. Obenberger [p]”. Described from unknown number of syntypes.
Agrilus kurandae Obenberger, 1923. Type locality. Kuranda (Quensland). Lectotype by present designation (Fig. 54), ♀, (NMPC): “Kuranda Queensland [h] \ Typus [p] [red label] \ Agrilus kurandae m. Type [h] Det. Dr. Obenberger [p]”. Described from unknown number of syntypes.
Agrilus celebicola Obenberger, 1924. Type locality. Celebes. Lectotype by present designation (Fig. 48), ♀, (NMPC): “Drs. Sarasin S. O. Celebes Kolaka [p] [yellow label] \ TYPUS [p] [red label] \ Agrilus celebicola m. Type [h] Det. Dr. Obenberger [p]”. Described from unknown number of syntypes.
Agrilus nirius Obenberger, 1924 Type locality. Java; Ins. Batoe; Tanah Masa. Lectotype by present designation (Fig. 58), ♀, (NMPC): “Java Buitenzorg [h] \ Typus [p] [red label] \ Agrilus Nirius m. Type [h] Det. Dr. Obenberger [p]”. Secondary: 6 paralectotypes (NMPC); 1 paralectotype (RMNH). Described from unknown number of syntypes.
Agrilus tebinganus Obenberger, 1924. Type locality. Ostsumatra. Lectotype by present designation (Fig. 63), ♀, (NMPC): “Sumatra [h] \ Typus [p] [red label] \ tebinganus Kerr. n. sp. type \ Agrilus tebinganus m. Type [h] [Obenberger’s MS] Det. Dr. Obenberger [p]”. Secondary: 1 paralectotype (MNHN). Described from unknown number of syntypes.
Types of Agrilus: 46 Agrilus auroapicalis Kurosawa, 1957 – Holotype 47 Agrilus biakanus Curletti, 2006 – Holotype (source: Curletti, 2006) 48 Agrilus celebicola Obenberger, 1924 – Lectotype49 Agrilus connexus Kerremans, 1900 – Holotype by monotypy 50 Agrilus cupricauda Saunders, 1867 – Lectotype 51 Agrilus evinadus Gory & Laporte, 1839 – Lectotype 52 Agrilus horni Théry, 1904 – Lectotype 53 Agrilus korenskyi Obenberger, 1923 – Lectotype 54 Agrilus kurandae Obenberger, 1923 – Lectotype 55 Agrilus laurenconi Descarpentries & Villiers, 1963 – Holotype 56 Agrilus marmoreus Deyrolle, 1864 – Lectotype 57 Agrilus nitidus Kerremans, 1898 – Lectotype 58 Agrilus nirius Obenberger, 1924 – Lectotype 59 Agrilus oblatus Kerremans, 1900 – Lectotype 60 Agrilus occipitalis (Eschscholtz, 1822) – Lectotype 61 Agrilus samoensis Blair, 1928 – Holotype 62 Agrilus sordidulus Obenberger, 1916 – Holotype by monotypy 63 Agrilus tebinganus Obenberger, 1924 – Lectotype.
Size: 5.5–9.3 mm. Agrilus occipitalis is very variable in the size, color and the shape of body parts. The pygidium varies from arcuate to subangulate (Figs 43-45). From the close Agrilus horniellus and Agrilus sordidulus, it differs by the characters cited in diagnosis at Agrilus horniellus and Agrilus sordidulus. See also characters in Appendix.
INDONESIA: Java: 1 ♂ (EJCB): “F. H. Doesburg, Java, Samarang”; 1 ♂ (EJCB): “Java Samarang”; 1 ♂ (EJCB): “Java Malang”; 1 (USNM): “L. G. E. Kalshoven, Java 250m, Buitenzorg, ix.1924, NS 176”; 1 (USNM): “Dr. L. J. Toxopeus, Preanger, Java, Bandoeng, 27.xi.31, Djeroek”; 1 (USNM): “Dr. L. J. Toxopeus, Preanger, Java, Bandoeng, xi.1932”; 1 ♂ (EJCB): “Indonesia, Java, Bandung, iv.1993 on Citrus trees”; 8 (EJCB): “Indonesia, Java isl, East Java prov., 6 km SE of Lasem, Celering Mt., 23. I. 1998, St. Jákl leg.”; 1 ♂, 1 ♀ (EJCB): “Indonesia, Java cent., Lasem env. - 4 km E of, Gunung Celering 140 m, 23–24. I. 1998, R. Červenka lgt.”. Kalimantan: 3 (EJCB): “Borneo occ. Pontianak 1901”. Lesser Sunda: 2 ♂, 2 ♀ (EJCB): “Sumbawa Colffs.”; 3 ♂ (EJCB): “W Timor, 350 m, Buraen, 50 km S Kupang, 26.i.-9.ii.2006, S. Jákl leg.”. Sulawesi: 1 (EJCB): “Celebes”; 2 ♂, 1 ♀ (USNM): “Celebes NEJ, Watampone vi 1935, leg. L. E. C. Veen”.Sumatra: 1 ♂ (EJCB): “W Sumatra, 1991”; 1 ♀ (EJCB): “W Sumatra, Solok, Jul 1995”; 1 ♂ (EJCB): “Sumatra, Harau valley, April 1996”; 1 ♂, 4 ♀ (EJCB): “W Sumatra, Harau Valley, 700 m, iv.2004, S. Jákl leg.”; 1 ♂ (EJCB): “Indonesia, W Sumatra, Harau Valley, 500-800 m, ca 20 km N of Payakumbuh, iv-v.2006, S. Jákl leg.”; 3 ♂ (EJCB): “Indonesia, W Sumatra, Harau Valley, 500–800 m, ca 20 km N of Payakumbuh, 5-28.ii.2006, S. Jákl leg.”; 1 ♀ (EJCB): “Indonesia, W Sumatra, Harau Valley, 500-800 m, N of Payakumbuh, iv-v.2006”; 1 ♂ (EJCB): “Indonesia, W Sumatra, Harau Valley, 500-800 m, ca 20 km N of Payakumbuh, iv-v.2006, S. Jákl leg.”; 1 ♂ (EJCB): “Indonesia, W Sumatra, Harau Valley, 500–800 m, ca 20 km N of Payakumbuh, ii.2006, S. Jákl leg.”; 8 (EJCB): “Indonesia, W Sumatra, Mt. Tandikat, 400-600m, ca 25 km N Pariaman, i. 2007 S. Jákl leg.”; 1 ♂ (EJCB): “Indonesia, W Sumatra, Harau Valley, 500–800 m, ca 20 km N of Payakumbuh, v-vi.2007, S. Jákl leg.”; 8 (EJCB): “Indonesia, W Sumatra, Harau Valley, 500-800 m, ca 20 km N of Payakumbuh, v-vi.2007, S. Jákl leg.”; 2 ♂, 2 ♀ (EJCB): “Indonesia, W Sumatra, Harau Valley, 500-800 m, ca 20 km N of Payakumbuh, viii.2009, S. Jákl leg.”. LAOS: Vientiane: cca 55 (EJCB): “Laos centr., 27. IV. -1. V. 1997, 70 km NE Vientiane, Ban Phabat env., 150 m, 18°16.1N, 103°10.9E, E. Jendek & O. Šauša leg.”; 1 ♂ (EJCB): “C Laos, Viang Chan prov. Lao Pako resort, 100 m, 50km NE Vientiane, 2002, M. Štrba leg. 28–30. V. ”; 1 ♂, 1 ♀ (EJCB): “LAOS, Vientiane prov., Lao Pako env. 200 m, 55 km NE Vientiane, 1–4.v.2004, F. & L. Kantner leg.”; 1 ♀ (EJCB): “Laos centr., Viang Chan pr., Ban PA Kho resort, ca.50 km, NE of Vientiane, ~90 m, 18°10'N, 102°52'E, 9.–14.vi.2007, M. Štrba leg.”. MALAYSIA: Johor: 1 (EJCB): “Malaysia: Pahang, Tioman island, 2 km N Ayer Batang, 18.7.1993, leg. Schuh”. Pahang: 1 ♀ (EJCB): “Malaysia-W, Pahang pr., 30km E Ipoh, 1500m, Cameron Highlands, Tanah Rata, 20.ii.-3.iii.1998, P. Čechovský leg.”; 1 ♀ (EJCB): “Malaysia-W, Pahang pr., 30km E Ipoh, 1500m, Cameron Highlands, Tanah Rata, 21-24.vi.2001, P. Čechovský leg.”; Perak: 1 ♀ (EJCB): “Malaysia-W, Perak 900m, 40km SE Ipoh, 4°25'N, 101°23'E, Cameron Highlands, Ringlet, M.Říha leg. 25.iv.–5.v.2001”. NORTHERN MARIANA ISLANDS: 13 (USNM): “Mariana Isls.: Saipan Island, Aug. 20, 1944, David G. Hall”; 38 (USNM): “Lake Hagoya, Tinian Is., VI-10-46 \ Oakley 526 on Citrus leaves”; 2 (USNM): “Chalon [?]avlav, Saipan, vi-19-46, Oakley 734, on orange leaves”; 1 (USNM): “Rota Rota, vi-23-46, Townes”; 1 (USNM): “Vlig Bay, Guam, 22-i-48, Mechler \ with Annona recticulata 48-2448”; 5 (USNM): “Chalon Lavlau, Saipan, vi-19-46, Oakley 734, on orange leaves”; 6 (USNM): “Marpo Valley, Tinian Isl., vi-9-46, Oakley 520, on sour orange leaves”. PAPUA NEW GUINEA: 1 ♂ (CBCS): “New Guinea, Sideia island, Sideia Mission, 28 Dec. 1988, Leg. G. Hangay”. PHILIPPINES: Luzon isl. group: 22 (USNM): “Manilla PI, in citrus branches, CollnRC McGregor”; 3 (USNM): “Malinao, Tayabas, Baker”; 1 (USNM): “Manila, PI, CollnRC McGregor”; 1 (USNM): “Mt. Makiling, Luzon, Baker, ”; 1 ♂ (EJCB): “Mindoro”; 6 (USNM): “Santo Tomas, Batangas, Luzon”; 8 (USNM): “Manile, Philippines”; 1 (USNM): “Luzon, P. I. , Montalban”; 1 (USNM): “Los Banos, Philippine Is., vi-vii-17”; 3 (USNM): “Manila P. I. , VI-24, R. C. Mc. Gregor”; 5 (USNM): “Manila P. I. , April 24, R. C. Mc. Gregor”; 1 (USNM): “Manila P. I. , V-24, R. C. Mc. Gregor”; 22 (USNM): “in Citrus wood, Manila, P. I. , 8.17.25, FC Brosius”; 1 (USNM): “Los Banos, Luzon PI, X.1945, Bmalkin”; 1 ♀ (EJCB): “Mt. Maquiling Philippines, elev. 50m, 28-II-48, R. Afenir”; 2 ♂, 1 ♀ (EJCB): “Lipa, Batangas, elev. 100m, 10-VIII-1948, Bigornia, A.”;Mindanao isl. group: 2 (USNM): “Zamboanga, Mindanao, Baker”; 1 (USNM): “Zamboanga, Mindanao, Baker”; 3 (USNM): “Iligan, Mindanao”; 2 (USNM): “Surigao, Mindanao, Baker”; 3 (USNM): “Dapitan, Mindanao, Baker”; 1 ♂, 1 ♀ (NMPC): “Dapitan Mindanao Baker”; 6 (USNM): “Davao, Mindanao, Baker”; 1 (USNM): “Diklom, Bukidnon, Mindanao”; 2 ♀ (EJCB): “Philippinen Mindanao”; 15 (USNM): “Butuan, Mindanao, Baker”; 1 ♀ (EJCB): “Mindanao Phillippines 28.vii.1977 M. Sato leg.”; 3 (CBCS): “Philippines, S. E. Mindanao, ix.2009, local collector”. Palawan isl. group: 11 (EJCB): “Philippines, Palawan, 1–21. II. 2000, 800 m, 9°42'N, 118°31E, Salakot waterfalls, E. Jendek leg.”. Visayas islands: 1 (USNM): “Isl. Biliran, Philippines, Baker”; 4 (USNM): “Victoria, occ. Negros, in gardin”; 5 (USNM): “Island Samar, Baker”; 1 ♀ (EJCB): “Victorias, Occ. Negros, 10/26/[19]29”; 6 (USNM): “Calicoan isl., P. I. , x-15-45 FF Bibby 601”; 1 (EJCB): “Masbate P. I. VIII.28 1952 Henry Townes”; 1 ♂ (EJCB): “Malubog, Toledo City, Cebu Is., 12.vi.1986, Hawkeswood T. J. , on steam of Citrus”; 1 (CBCS): “Philippines, Leyte Isl., Mt. Balocaue, vi.2009, local collector”. THAILAND: 1 ♂ (EJCB): “S Thailand Covaz 2.6.1995”, Chiang Mai: 1 ♂ (EJCB): “Thailand 1. VI. 1990 Sansai, Chiang Mai, S. Steinke leg.”. Mae Hong Son: 2 ♂ (EJCB): “Thailand bor., prov. Mae Hong Son, Pai, 24–30. IV. 1997, R. Šigut leg.”. Nakhon Ratchasima: 1 ♂ (MHCB): “Thailand Corat 12.vii.1995, leg. Lehman & Steinke”; 1 ♂, 1 ♀ (EJCB): “Thailand Corat VI.1997”.Yala: 1 ♀ (EJCB): “S Thailand 7-8. V. 1992 Betong, L. Dembický leg.”; 1 ♀ (EJCB): “S Thailand Betong, Gunung Cang dun vill., Yala dist. 25.3.–22.4.93, J. Horák leg.”.VIETNAM: 2 ♀ (EJCB): “Cochinchine”.
1–2–3–4–5–6–7–8–9–10–11–12. Altitude range: 50–1500 m.
Citrus sp.:
AUSTRALIA: Queensland. CAMBODIA: Phnum Penh. CHINA: Taiwan. INDONESIA: Irian Jaya; Java; Kalimantan; Lesser Sunda (incl. West Timor); Maluku; Sulawesi; Sumatra. LAOS: Vientiane. MALAYSIA: Johor; Malaysia Peninsular; Pahang; Perak. NORTHERN MARIANA ISLANDS. PAPUA NEW GUINEA. PHILIPPINES: Luzon isl. group; Mindanao isl. group; Palawan isl. group; Visayas islands. THAILAND: Chiang Mai; Mae Hong Son; Nakhon Ratchasima; Yala. VIE-TNAM: Ba Ria-Vung Tau.
Remarks.
http://species-id.net/wiki/Agrilus_perroti
Fig. 2 (habitus) ; Fig. 25 (aedeagus)Agrilus perroti Descarpentries & Villiers, 1963. Type locality. Tonkin: Thanh-Moï. Holotype examined by
See
INDIA: West Bengal: 1 ♂, 1 ♀ (MNHN): “British Bootang, Maria Basti, Durel [leg.]”. For further records see:
4. Altitude range: 400 m.
Unknown.
CHINA: Fujian; Guangxi; Yunnan. INDIA: West Bengal. VIE-TNAM.
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http://species-id.net/wiki/Agrilus_picturatus
Fig. 9 (habitus of holotype) ; Fig. 29 (aedeagus)BODY. Size: 8.4 mm (Holotype).Shape: cuneiform, Build: slender, Color (dorsally): bicolored. HEAD. Medial impression: deep, Frons: Shape: flat, Vertex: Sculpture elements: rugae, Sculpture shape: semispherical or subparallel, Sculpture density: dense, Sculpture intensity: rough, Eyes: Size: moderate, Lower margin: in line or below with antennal socket, Medial orbit: converging ventrally, Antennae: Length: long, Width: slender, Serration: from antennomere 4, Antennomere 7-10 (shape): with obvious collum. PRONOTUM. Shape: visually square, Sides: slightly arcuate or straight, Maximal width: at middle, Anterior margin: subequal to posterior, Anterior lobe: Size: obvious, Shape: arcuate, Position: projecting beyond anterior angles, Posterior angles: Shape: rectangular, Apex: sharp, Disk: Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (intensity): deep, Prehumerus: Development: carinal, Shape: arcuate, Extent: to 1/3 of pronotal length, Anterior end: distant from lateral carina, Posterior end: joined with posterior angle or margin, Arc: moderate, Lateral carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent or present, Narrowest point: at posterior angles. ELYTRA. Color: dichromatic, Alternative color: apical portion, Humeral carina: absent, Apices: Arrangement: separate, Shape: arcuate, Modifications: margin obviously denticulate, Pubescence: Color: monochromatic, Extent: entire ornamental, rarely entire ornamental with indication of stripes. STERNUM. Prosternal lobe: Distal margin: arcuate, Prosternal process: Shape: dilated, rarely subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: flat, Projection (extend): distinctly beyond angles, Mesosternum: Mesosternal projection: flat. ABDOMEN. Pygidium: Apical margin: with shortly projecting carina, Last ventrite: Disk: with medial carinula, Sternal groove: Extent: on all ventrites or on three apical ventrites, Shape on the apex of last ventrite: arcuate. LEGS. Metatarsus: Size to metatibia: about as long or longer than metatibia, Tarsomere 1: Size to following tarsomeres: subequal or longer than 2-4. GENITALIA. Aedeagus (Fig. 29): Symmetry: symmetric, Shape: widest in basal part, Modifications: apex of medial lobe sharply pointed.
From the very similar Agrilus pseudoambiguus sp. n., it differs by having the pronotum more elongate with sides almost straight; by having an obvious pronotal lobe and by the elytral apices being distinctly denticulate. See also Appendix.
Thailand, Sakon Nakhon province, Phu Phane National Park, 17°07'30"N, 104°01'E, altitude 350 m.
Holotype (Fig. 9), ♂, (EJCB): “Thai, Sakon Nakhon, Phu Phane Nat.Park, 17°07'30"N, 104°01'E, 350m, ix.2000, local collector”.
9. Altitude range: 350 m.
Unknown.
THAILAND: Sakhon Nakhon.
The specific name is the Latin adjective picturatus (painted). It refers to the elytral pubescence of the species.
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http://species-id.net/wiki/Agrilus_pluvius
Fig. 23 (habitus of holotype)BODY. Size: 12.6 mm (Holotype).Shape: cuneiform, Build: robust, Color (dorsally): unicolored. HEAD. Medial impression: deep, Epistoma: raised above frons, Vertex: Sculpture elements: rugae, Sculpture shape: semispherical, Sculpture density: dense, Eyes: Size: small, Lower margin: in line or below with antennal socket, Medial orbit: subparallel, Antennae: Serration: from antennomere 4. PRONOTUM. Shape: transverse, Sides: slightly arcuate, Maximal width: at middle, Anterior margin: subequal to posterior, Anterior lobe: Size: moderate, Shape: arcuate, Width: wide, Position: at level with anterior angles or not reaching level of anterior angles, Posterior angles: Shape: obtuse, Apex: sharp, Disk: Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (intensity): deep, Prehumerus: Development: carinal, Shape: bisinuate, Extent: to 1/3 of pronotal length, Anterior end: distant from lateral carina, Posterior end: joined with posterior angle or margin, Arc: moderate, Lateral carinae: Interspace: narrow, Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles. ELYTRA. Color: monochromatic, Humeral carina: absent, Apices: Arrangement: separate, Shape: arcuate, Pubescence: Color: monochromatic, Density: dense, Extent: entire ornamental. STERNUM. Prosternal lobe: Size: large, Distal margin: arcuate or subtruncate, Prosternal process: Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: flat, Projection (extend): distinctly beyond angles, Mesosternum: Mesosternal projection: flat. ABDOMEN. Tomentum: present, Pygidium: Apical margin: arcuate, Sternal groove: Extent: on three apical ventrites, Shape on the apex of last ventrite: arcuately sinuate, Emargination (deepness): very shallow. LEGS. Metatarsus: Size to metatibia: about as long or longer than metatibia, Tarsomere 1: Size to following tarsomeres: subequal or longer than 2-4. GENITALIA. Ovipositor: Shape: square (uritiform).
Agrilus pluvius sp. n. is very distinctive by the large size and it differs from all members of the subgroup by having the apical half of elytra more elongate; by the arcuate apical margin of pygidium and by the distinctly sinuate sternal groove on the apex of last ventrite. See also Appendix.
Northeastern India, Meghalaya, southwest of Cherrapunjee, 25°13'–14'N 91°40'E, altitude 900 m.
Holotype (Fig. 23), ♀, (EJCB): “NE India, Meghalaya, SW of Cherrapunjee, 25°13'–14'N 91°40'E, 900m, 5.–24.v.2005, P. Pacholátko leg.”.
5. Altitude range: 900 m.
Unknown.
INDIA: Meghalaya.
The specific name is Latin adjective pluvius (rainy). It refers to the type locality which is known for the highest precipitation in the world.
urn:lsid:zoobank.org:act:9A413050-3A7B-480A-9990-AE8CBA32019D
http://species-id.net/wiki/Agrilus_pseudoambiguus
Fig. 10 (habitus of holotype) ; Fig. 30 (aedeagus)BODY. Size: 5.9–8.1 mm (Holotype 8.1 mm).Shape: cuneiform, Color (dorsally): bicolored, rarely unicolored, Sexual modifications in male: not apparent. HEAD. Medial impression: deep, Epistoma: raised above frons, Vertex: Shape: markedly convex, Sculpture elements: rugae, Sculpture shape: semispherical or subparallel, Sculpture density: dense, Eyes: Size: small, Lower margin: in line or below with antennal socket, Medial orbit: converging ventrally, Antennae: Length: long, Width: slender, Serration: from antennomere 4, Antennomere 7-10 (shape): with obvious collum. PRONOTUM. Shape: visually square, Sides: slightly arcuate, rarely subangulate, Maximal width: at middle, Anterior margin: subequal to posterior or wider than posterior, Anterior lobe: Size: absent or vague, rarely moderate, Shape: arcuate, Position: at level with anterior angles, Posterior angles: Shape: rectangular, rarely acute, rarely obtuse, Apex: sharp, Disk: Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (intensity): deep, Lateral impression (size): narrow, Prehumerus: Development: carinal, Shape: arcuate, Extent: to 1/3 of pronotal length, Anterior end: distant from lateral carina, Posterior end: joined with posterior angle or margin, Arc: moderate or weak, Lateral carinae: Convergence: moderately convergent, Junction: absent or present, Narrowest point: at posterior angles. ELYTRA. Color: dichromatic, Alternative color: apical portion, Apices: Arrangement: separate, Shape: arcuate, Pubescence: Color: monochromatic, Extent: entire ornamental, rarely entire ornamental with indication of stripes. STERNUM. Prosternal lobe: Distal margin: arcuate, Prosternal process: Shape: dilated or subparallel, Sides: straight, Angles: obtuse, Disc: flat, Projection (extend): distinctly beyond angles, Mesosternum: Mesosternal projection: flat. ABDOMEN. Pygidium: Apical margin: angulate, Sternal groove: Extent: on all ventrites or on three apical ventrites, Shape on the apex of last ventrite: arcuate, rarely arcuately sinuate, Emargination (deepness): very shallow. LEGS. Metatarsus: Size to metatibia: somewhat shorter as metatibia, Tarsomere 1: Size to following tarsomeres: subequal or longer than 2-4. GENITALIA. Aedeagus (Fig. 30): Symmetry: symmetric, Shape: subparallel, Modifications: apex of medial lobe sharply pointed, Ovipositor: Shape: markedly elongate.
From very similar Agrilus picturatus sp. n., this new species differs by having the pronotum more transverse with sides slightly arcuate; by absent or vague pronotal lobe and by the smooth or very finely denticulate elytral apices. See also Appendix.
Laos, Louang Namtha pr., 21°09'N, 101°19'E, Namtha - Muang Sing, 900–1200 m.
Holotype (Fig. 10), ♂, (EJCB): “Laos, Louang Namtha pr., 21°09'N, 101°19'E, Namtha - Muang Sing, 5–31.v.1997, 900–1200 m, Vit Kubáň leg.”. Paratypes: 2 paratypes (EJCB) with the same data as holotype. 1 paratype (EJCB): “Laos NE, Hua Phan prov., 20°19'N, 104°25'E, 25 km SE Vieng Xai (by road), Ban Kangpabong env., 14–18.v.2001, D. Hauck leg.”
5. Altitude range: 900–1200 m.
Unknown.
LAOS:Houaphan; Louang Namtha.
The specific name is derived from Greek prefix pseudo- (having the appearance of) and the specific name ambiguus; it refers to the similarity of the species to Agrilus ambiguus.
http://species-id.net/wiki/Agrilus_sordidulus
Fig. 20 (habitus) ; Fig. 37 (aedeagus) ; Fig. 62 (Holotype by monotypy)Agrilus sordidulus Obenberger, 1916. Type locality. Ostindien: Trichinopoli. Holotype by monotypy (Fig. 62), (NHMB): “Typus [p] [red label with black border] \ 1220 [h] [blue label] \ Koll. Dr. A. Frh. v. Hoschek [p] Trichinopolis Ind. or. [h] \ Agrilus sordidulus Typ! [h] Det. Obenberger [p]”. Described from 1 specimen.
Size: 6.1–8.4 mm. Agrilus sordidulus can distinguished from the similar Agrilus occipitalis and Agrilus horniellus by the more robust body with the narrowing apical part of elytra shorter; by the head deeply, medially impressed and by the very deep medial pronotal impressions. See also Appendix.
INDIA: Karnataka: 1 (CBCS): “S. Coorg, S. India, Ammatti, 3100 ft, II.1952”. Kerala: 2 ♂ (EJCB): “S India, Kerala, Thekkady Periyar Lake, 9.34 N, 77.10 E, 900–1000 m, 19-27. IV. 1997, Dembický & Pacholátko leg.”. Tamil Nadu: 1 ♂, 8 ♀ (EJCB): “S India, Tamil Nadu, Nilgiri Hills, 15 km SE Kotagiri, near Kunchappanai, alt. 900 m, 13-20. V. 1994, 11°22'N, 76°56'E, Z. Kejval lgt.”; 1 ♂, 3 ♀ (EJCB): “India S, Tamil Nadu, Nilgiris, 15 km SE of Kotagiri, Kunchappanai, 900 m, 11.22 N 76.56 E, 7–22.v.2000, leg. P. Pacholátko”; 1 ♂, 1 ♀ (EJCB): “India S, Tamil Nadu, Nilgiri Hills, 11 km SE of Kotagiri, 1100±100 m, 11.24 N 76.56 E, Kunchappanai, leg. L. Dembický, 3-15.v.2002”.
2–4–5. Altitude range: 900–1200 m.
Unknown.
INDIA: Karnataka; Kerala; Tamil Nadu.
urn:lsid:zoobank.org:act:4298A27E-4E89-42F4-A989-AF7CC76DB545
http://species-id.net/wiki/Agrilus_tesselatus
Fig. 1 (habitus of holotype)BODY. Size: 9.9 mm (Holotype).Shape: subparallel, Build: robust, Posterior tapering part: short with broad apex, Color (dorsally): unicolored. HEAD. Medial impression: deep, Epistoma: raised above frons, Frons: Shape: markedly convex, Vertex: Shape: markedly convex, Sculpture elements: rugae, Sculpture shape: semispherical, Eyes: Size: small, Shape: markedly protruding head outline, Lower margin: in line or below with antennal socket, Medial orbit: converging ventrally, Antennae: Length: short, Width: solid, Serration: from antennomere 5, Antennomere 7-10 (shape): without collum, Antennomere 7-10 (length): markedly wider than long. PRONOTUM. Shape: transverse, Sides: straight, Maximal width: at posterior margin, Anterior margin: narrower than posterior or subequal to posterior, Anterior lobe: Size: obvious, Position: projecting beyond anterior angles, Posterior angles: Shape: obtuse, Apex: blunt, Disk: Convexity: strongly convex, Impressions: absent or medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (intensity): shallow, Lateral impression (size): narrow, Prehumerus: Development: carinal, Shape: straight, Anterior end: distant from lateral carina, Posterior end: distant from angles and margin, Lateral carinae: Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles, Scutellum: Disc: impressed, Scutellar carina: obsolete or absent. ELYTRA. Color: monochromatic, Humeral carina: absent, Apices: Arrangement: separate, Shape: subtruncate, Truncation: transverse, Pubescence: Color: monochromatic, Extent: entire ornamental. STERNUM. Prosternal lobe: Distal margin: angulately emarginate, Delimitation: angulate, Emargination (width): wide, Prosternal process: Shape: subparallel, Sides: straight, Angles: rectangular, Angles (tips): blunt, Disc: flat, Projection (extend): distinctly beyond angles, Mesosternum: Mesosternal projection: flat. ABDOMEN. Tomentum: absent, Pygidium: Apical margin: arcuate, Sternal groove: Extent: on apical ventrite, Shape on the apex of last ventrite: angulately sinuate, Emargination (width): markedly wide. LEGS. Metatarsus: Size to metatibia: distinctly shorter than metatibia, Tarsomere 1: Size to following tarsomeres: longer than 2-3 but shorter than 2-4. GENITALIA. Ovipositor: Shape: square (uritiform).
The very distinctive species which differs from all other members of Agrilus occipitalis species-group mainly by characters given for the subgroup definition. See also Appendix.
NorthVietnam, Tonkin, Ninh Binh province, Cuc-Phuong national park, 20°18'N, 105°39'00"E.
Holotype (Fig. 1), ♀, (EJCB): “Vietnam N, Tonkin, Cuc-Phuong nat. park, 2–12. V. 1991, E. Jendek leg.”.
5.
Unknown.
VIETNAM: Ninh Binh.
The specific name is the Latin adjective tesselatus (checkered). It refers to the elytral pubescence of the species.
http://species-id.net/wiki/Agrilus_tonkineus
Fig. 24 (habitus) ; Fig. 40 (aedeagus)Agrilus tonkineus Kerremans, 1895. Type locality. Hanoï. Lectotype designated by
Agrilus blatteiceps Bourgoin, 1925. Type locality. Laos: Vien Poukha. Holotype by monotypy examined by
Size: 7.5–9.5 mm. Very close to Agrilus mucidus sp. n. from which it can be distinguished by the pronotum more convex; by the arcuate prehumerus and by the absence of transverse tomentose strip at apical third of elytra. See also Appendix.
CHINA: Fujian: 1 ♂ (EJCB): “Shunchang Fujian, 27.iv.1979, Shicheng Ji leg”. Hainan: 1 ♀ (USNM): “Hainan Is, Woh Hau Chuen, E of Nodoa, Jul 3, 1929”; 1 ♀ (MNHN): “Hainan, Hu...[illegible], G. Ros leg. 23.v.[19]36”. Yunnan: 1 (IZAS): “Yunnan Xishuangbanna Menghun, 1200-1400m, 28.iv.1958, C. P. Hong leg. [in Chinese]”; 1 (IZAS): “Yunnan Xishuangbanna Mengsong, 1600m, 28.iv.1958, S. Y. Wang leg. [in Chinese]”; 1 (IZAS): “Yunnan Xishuangbanna Damenglong, 650m, 11.iv.1958, L. Y. Zheng leg. [in Chinese]”. For further records see
4–5–6–7. Altitude range: 420–1600 m.
Unknown.
CHINA: Fujian; Hainan; Yunnan. LAOS: Borikhamxai; Louang Namtha; Vientiane; Xaignabouri; Xiangkhoang. VIETNAM: Ha Noi; Ha Tay; Hoa Binh.
urn:lsid:zoobank.org:act:46090B44-156C-4387-BADF-35E4F03F81E4
http://species-id.net/wiki/Agrilus_trepanatus
Fig. 14 (habitus of holotype)BODY. Size: 10.2–12.7 mm (Holotype 12.7 mm).Shape: subparallel, Build: robust, Posterior tapering part: short with broad apex, Color (dorsally): bicolored. HEAD. Medial impression: deep, Epistoma: raised above frons, Vertex: Sculpture elements: rugae, Sculpture shape: semispherical, Sculpture density: dense, Eyes: Size: small, Lower margin: in line or below with antennal socket, Medial orbit: subparallel, Antennae: Length: long, Width: slender, Serration: from antennomere 4, Antennomere 7-10 (shape): with obvious collum. PRONOTUM. Shape: visually square, Sides: slightly arcuate, Maximal width: at middle, Anterior margin: subequal to posterior, Anterior lobe: Size: moderate or obvious, Shape: arcuate, Position: at level with anterior angles, Posterior angles: Shape: obtuse or rectangular, Apex: sharp, Disk: Convexity: strongly convex, Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Prehumerus: Development: carinal, Shape: arcuate or bisinuate, Extent: to 1/2 of pronotal length or to 1/3 of pronotal length, Modifications: with rudiment at anterior angle, Anterior end: distant from lateral carina, Posterior end: joined with posterior angle or margin, Arc: moderate, Lateral carinae: Interspace: narrow, Convergence: moderately convergent, Junction: absent, rarely present, Narrowest point: at posterior 1/5-1/4 of marginal carina, Modifications: submarginal carina posteriorly obliterate. ELYTRA. Color: monochromatic, Humeral carina: absent, Apices: Arrangement: separate, Shape: arcuate, Pubescence: Color: monochromatic, Extent: entire ornamental with indication of stripes. STERNUM. Prosternal lobe: Size: large, Distal margin: arcuate, Prosternal process: Shape: dilated or subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: flat, Projection (extend): distinctly beyond angles, Mesosternum: Mesosternal projection: flat. ABDOMEN. Pygidium: Apical margin: arcuate, Sternal groove: Extent: on three apical ventrites, Shape on the apex of last ventrite: arcuate or arcuately sinuate, Emargination (deepness): very shallow. LEGS. Metatarsus: Size to metatibia: distinctly shorter than metatibia, Tarsomere 1: Size to following tarsomeres: subequal or longer than 2–4. GENITALIA. Ovipositor: Shape: markedly elongate.
Agrilus trepanatus sp. n. can be distinguished from all species of the group by the large body; by the strikingly bicolor dorsal side and by the head obviously deeply impressed medially. See also Appendix.
South India, Karnataka state, Coorg district, northeastern Virajpet, 75°50'E, 12°13'N, altitude 500 m.
Holotype (Fig. 14), ♀, (EJCB): “S-India, Karnataka state, Coorg distr., NE Virajpet, 75°50'E, 12°13'N, ca 500m, 4–8.vi.1999, Z. Kejval & M. Trýzna leg.”. Paratypes: 1 paratype, ♀ (EJCB): “India, Karnataka, 12 km SW Yellapur, 7.vii-14.viii.84, B. Gill FIT 500 m”.
6–7–8. Altitude range: 500 m.
Unknown.
INDIA: Karnataka.
The specific name is an adjective derived from the Greek verb trepao (drill, bore) in Latinized form trepano. It refers to the conspicuously impressed head of this species.
urn:lsid:zoobank.org:act:A12BB28B-30FB-4AC7-8D0A-99F504EA8221
http://species-id.net/wiki/Agrilus_umrongso
Fig. 3 (habitus of holotype)BODY. Size: 12 mm (Holotype).Shape: cuneiform, Posterior tapering part: long with narrow apex, Color (dorsally): unicolored. HEAD. Medial impression: deep, Epistoma: raised above frons, Vertex: Shape: markedly convex, Sculpture elements: rugae, Sculpture shape: semispherical, Sculpture density: dense, Eyes: Size: small, Lower margin: in line or below with antennal socket, Medial orbit: converging ventrally, Antennae: Serration: from antennomere 4. PRONOTUM. Shape: transverse, Sides: slightly arcuate, Maximal width: at middle or at posterior margin, Anterior margin: subequal to posterior, Anterior lobe: Size: moderate, Shape: arcuate, Position: at level with anterior angles or projecting beyond anterior angles, Posterior angles: Shape: obtuse, Apex: sharp, Disk: Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Prehumerus: Development: carinal, Shape: bisinuate, Extent: to 1/3 of pronotal length, Anterior end: distant from lateral carina, Posterior end: distant from angles and margin, Arc: weak, Lateral carinae: Convergence: moderately convergent, Junction: present, Narrowest point: at posterior angles. ELYTRA. Color: monochromatic, Humeral carina: absent, Apices: Arrangement: separate, Shape: arcuate, Pubescence: Color: monochromatic, Density: dense, Extent: entire ornamental with indication of stripes. STERNUM. Prosternal lobe: Size: large, Distal margin: arcuately emarginate, Emargination (width): narrow, Prosternal process: Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: flat, Projection (extend): distinctly beyond angles, Mesosternum: Mesosternal projection: flat. ABDOMEN. Tomentum: present, Pygidium: Apical margin: arcuate, Sternal groove: Extent: on three apical ventrites, Shape on the apex of last ventrite: arcuate. LEGS. Metatarsus: Size to metatibia: distinctly shorter than metatibia, Tarsomere 1: Size to following tarsomeres: longer than 2-3 but shorter than 2-4. GENITALIA. Ovipositor: Shape: markedly elongate.
Agrilus umrongso sp. n. can be distinguished from the very close Agrilus perroti by the head much more deeply impressed medially; by the apically arcuate pygidium and by the distinctly emarginate prosternal lobe. See also Appendix.
Northeastern India, Assam, 5 km north of Umrongso, altitude 700 m, 25°27'N, 92°43'E.
Holotype (Fig. 3), ♀, (EJCB): “NE India, Assam, 1999, 5 km N of Umrongso, 700m, 25°27'N, 92°43'E, 17.–25.v., Dembický & Pacholátko leg.”.
5. Altitude range: 700 m.
Unknown.
INDIA: Assam.
The specific name is a noun in apposition. It refers to the Umrongso, the type locality of the species.
http://species-id.net/wiki/Agrilus_yamawakii
Fig. 15 (habitus) ; Fig. 34 (aedeagus)Agrilus yamawakii Kurosawa, 1957. Type locality. Mt. Fukuchiyama, Fukuoka Pref., Kyűshű, Japan. Holotype examined by
Size: 6.9–11.7 mm. Agrilus yamawakii differs from other member of the group by the following combination of characters: the antennae moderately long; the elytral pubescence obsolete at least in proximal part; apical-most part of elytral apices pubescent. See also Appendix.
KOREA SOUTH:1 ♂ (MNHN): “Corée, Mirinai, Chass. indigènes”. For further records see
5–6–7.
Fagara (=Zanthoxylum) ailanthoides; Fagara mantchurica:
CHINA: Liaoning; Nei Mongol; Taiwan. JAPAN: Honshu; Kyushu; Ryukyu isl. (Okinawa incl.); Shikoku; Tsushima. KOREA NORTH. KOREA SOUTH.
http://species-id.net/wiki/Agrilus_zanthoxylumi
Fig. 4 (habitus) ; Fig. 26 (aedeagus)Agrilus zanthoxylumi Li Meng Lou, 1989. Type locality. Shaanxi (Baoji, Weinan). Type specimens not found. Described from unknown number of specimens. See also Remarks.
See
See
5–6.
Zanthoxylum: Li Meng Lou (1989); Zanthoxylum bungeanum:
CHINA: Gansu; Hubei; Shaanxi; Shandong; Yunnan; Zhejiang.
Remarks.The authorship of the name zanthoxylumi had changed several times.
All collection curators are sincerely acknowledged for their help in accessing the specimens. For consultation and comments to the early version of the manuscript I thank Aleš Smetana & Vasily Grebennikov (both Ottawa, Canada).
Character state matrix for species from Agrilus occipitalis species-group with the most important diagnostic characters. (doi: 10.3897/zookeys.256.4272.app) File format: Adobe PDF file (pdf).