ZooKeys 392: 1–367, doi: 10.3897/zookeys.392.6328
Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London
Abraham S.H. Breure 1, Jonathan D. Ablett 2
1 Naturalis Biodiversity Center, P.O. Box 9517, Leiden, the Netherlands
2 Natural History Museum, Division of Higher Invertebrates, London, SW7 5BD, UK

Corresponding author: Abraham S.H. Breure (ashbreure@gmail.com)

Academic editor: A. Schileyko

received 28 September 2013 | accepted 24 January 2014 | Published 21 March 2014
(C) 2014 Abraham S.H. Breure. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
For reference, use of the paginated PDF or printed version of this article is recommended.

Citation: Breure ASH, Ablett JD (2014) Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London. ZooKeys 392: 1–367. doi: 10.3897/zookeys.392.6328

Abstract

The type status is described of 404 taxa classified within the family Bulimulidae (superfamily Orthalicoidea) and kept in the London museum. Lectotypes are designated for Bulimus aurifluus Pfeiffer, 1857; Otostomus bartletti H. Adams, 1867; Helix cactorum d’Orbigny, 1835; Bulimus caliginosus Reeve, 1849; Bulimus chemnitzioides Forbes, 1850; Bulimus cinereus Reeve, 1849; Helix cora d’Orbigny, 1835; Bulimus fallax Pfeiffer, 1853; Bulimus felix Pfeiffer, 1862; Bulimus fontainii d’Orbigny, 1838; Bulimus fourmiersi d’Orbigny, 1837; Bulimus (Mesembrinus) gealei H. Adams, 1867; Bulimus gruneri Pfeiffer, 1846; Bulimus humboldtii Reeve, 1849; Helix hygrohylaea d’Orbigny, 1835; Bulimus jussieui Pfeiffer, 1846; Bulimulus (Drymaeus) binominis lascellianus E.A. Smith, 1895; Helix lichnorum d’Orbigny, 1835; Bulimulus (Drymaeus) lucidus da Costa, 1898; Bulimus luridus Pfeiffer, 1863; Bulimus meleagris Pfeiffer, 1853; Bulimus monachus Pfeiffer, 1857; Bulimus montagnei d’Orbigny, 1837; Helix montivaga d’Orbigny, 1835; Bulimus muliebris Reeve, 1849; Bulimus nigrofasciatus Pfeiffer in Philippi 1846; Bulimus nitelinus Reeve, 1849; Helix oreades d’Orbigny, 1835; Helix polymorpha d’Orbigny, 1835; Bulimus praetextus Reeve, 1849; Bulinus proteus Broderip, 1832; Bulimus rusticellus Morelet, 1860; Helix sporadica d’Orbigny, 1835; Bulimus sulphureus Pfeiffer, 1857; Helix thamnoica var. marmorata d’Orbigny, 1835; Bulinus translucens Broderip in Broderip and Sowerby I 1832; Helix trichoda d’Orbigny, 1835; Bulinus ustulatus Sowerby I, 1833; Bulimus voithianus Pfeiffer, 1847; Bulimus yungasensis d’Orbigny, 1837.

The type status of the following taxa is changed to lectotype in accordance with Art. 74.6 ICZN: Bulimulus (Drymaeus) caucaensis da Costa, 1898; Drymaeus exoticus da Costa, 1901; Bulimulus (Drymaeus) hidalgoi da Costa, 1898; Bulimulus (Drymaeus) interruptus Preston, 1909; Bulimulus (Drymaeus) inusitatus Fulton, 1900; Bulimulus latecolumellaris Preston, 1909; Bulimus (Otostomus) napo Angas, 1878; Drymaeus notabilis da Costa, 1906; Drymaeus notatus da Costa, 1906; Bulimulus (Drymaeus) nubilus Preston, 1903; Drymaeus obliquistriatus da Costa, 1901; Bulimus (Drymaeus) ochrocheilus E.A. Smith, 1877; Bulimus (Drymaeus) orthostoma E.A. Smith, 1877; Drymaeus expansus perenensis da Costa, 1901; Bulimulus pergracilis Rolle, 1904; Bulimulus (Drymaeus) plicatoliratus da Costa, 1898; Drymaeus prestoni da Costa, 1906; Drymaeus punctatus da Costa, 1907; Bulimus (Leptomerus) sanctaeluciae E.A. Smith, 1889; Bulimulus (Drymaeus) selli Preston, 1909; Drymaeus subventricosus da Costa, 1901; Bulimulus (Drymaeus) tigrinus da Costa, 1898; Drymaeus volsus Fulton, 1907; Drymaeus wintlei Finch, 1929; Bulimus zhorquinensis Angas, 1879; Bulimulus (Drymaeus) ziczac da Costa, 1898.

The following junior subjective synonyms are established: Bulimus antioquensis Pfeiffer, 1855 = Bulimus baranguillanus Pfeiffer, 1853; Drymaeus bellus da Costa, 1906 = Drymaeus blandi Pilsbry, 1897; Bulimus hachensis Reeve 1850 = Bulimus gruneri Pfeiffer, 1846 = Bulimus columbianus Lea, 1838; Bulimus (Otostomus) lamas Higgins 1868 = Bulimus trujillensis Philippi, 1867; Bulimulus (Drymaeus) binominis lascellianus E.A. Smith, 1895 = Bulimulus (Drymaeus) binominis E.A. Smith, 1895; Drymaeus multispira da Costa, 1904 = Helix torallyi d’Orbigny, 1835; Bulimulus (Drymaeus) plicatoliratus Da Costa, 1898 = Bulimus convexus Pfeiffer, 1855; Bulimus sugillatus Pfeiffer, 1857 = Bulimus rivasii d’Orbigny, 1837; Bulimus meridionalis Reeve 1848 [June] = Bulimus voithianus Pfeiffer, 1847.

New combinations are: Bostryx montagnei (d’Orbigny, 1837); Bostryx obliquiportus (da Costa, 1901); Bulimulus heloicus (d’Orbigny, 1835); Drymaeus (Drymaeus) lusorius (Pfeiffer, 1855); Drymaeus (Drymaeus) trigonostomus (Jonas, 1844); Drymaeus (Drymaeus) wintlei Finch, 1929; Drymaeus (Mesembrinus) conicus da Costa, 1907; Kuschelenia (Kuschelenia) culminea culminea (d’Orbigny, 1835); Kuschelenia (Kuschelenia) culmineus edwardsi (Morelet, 1863); Kuschelenia (K.) gayi (Pfeiffer, 1857); Kuschelenia (Kuschelenia) tupacii (d’Orbigny, 1835); Kuschelenia (Vermiculatus) anthisanensis (Pfeiffer, 1853); Kuschelenia (Vermiculatus) aquilus (Reeve, 1848); Kuschelenia (Vermiculatus) bicolor (Sowerby I, 1835); Kuschelenia (Vermiculatus) caliginosus (Reeve, 1849); Kuschelenia (Vermiculatus) cotopaxiensis (Pfeiffer, 1853); Kuschelenia (Vermiculatus) filaris (Pfeiffer, 1853); Kuschelenia (Vermiculatus) ochracea (Morelet, 1863); Kuschelenia (Vermiculatus) petiti (Pfeiffer, 1846); Kuschelenia (Vermiculatus) purpuratus (Reeve, 1849); Kuschelenia (Vermiculatus) quechuarum (Crawford, 1939); Naesiotus cinereus (Reeve, 1849); Naesiotus dentritis (Morelet, 1863); Naesiotus fontainii (d’Orbigny, 1838); Naesiotus orbignyi (Pfeiffer, 1846); Protoglyptus pilosus (Guppy, 1871); Protoglyptus sanctaeluciae (E.A. Smith, 1889).

Type material of the following taxa is figured herein for the first time: Bulimus cinereus Reeve, 1849; Bulimus coriaceus Pfeiffer, 1857; Bulimulus laxostylus Rolle, 1904; Bulimus pliculatus Pfeiffer, 1857; Bulimus simpliculus Pfeiffer, 1855.

Keywords

Bulimulidae, types

Introduction and methods

This is the third paper on the types of Orthalicoidea in the Natural History Museum, London. Earlier papers (Breure and Ablett 2011, 2012) have presented the context of the collection, the criteria used for the selection of lectotypes, some biohistorical notes, and a list of type specimens belonging to the Amphibulimidae, Bothriembryontidae, and Odontostomidae. The aim of this paper is to supplement the earlier papers with data of the type specimens classified with the Bulimulidae (sensu Breure and Romero 2012).

References are given to the original publication, plus those of following papers where type material has been mentioned or is (re-)figured. Dates of publication are in accordance with Coan et al. (2013a, 2013b) and Duncan (1937). Abbreviations used for depositories of material are: ANSP, Academy of Natural Sciences, Philadelphia, U.S.A.; MHNG, Muséum d’Histoire Naturelle, Genève, Switserland; MNHN, Muséum d’Histoire Naturelle, Paris, France; NHMUK, Natural History Museum, London, U.K.; RBINS, Royal Belgian Institute of Natural Sciences, Brussels, Belgium; RMNH, Naturalis Biodiversity Center, Leiden, the Netherlands (formerly the collections of Rijksmuseum van Natuurlijke Historie, Leiden, and Zoölogisch Museum, Amsterdam); SMF, Senckenberg Natur-Museum, Frankfurt am Main, Germany; ZMB, Zoologisches Museum, Humboldt Universität, Berlin, Germany. Other abbreviations used are: / end of line in cited text; coll., collection; D, diameter; H, shell height; leg., legit, collected; W, number of whorls. See Breure and Whisson 2012: fig. 1 for the way measurements on the shell have been taken. Label styles in the Cuming collection (”M.C. label style”) are explained in Breure and Ablett (2011: 7–8). Although most figures have been composed with the shells enlarged, its sizes retained relatively, one need to check the actual shell height in the legend.

Systematics
Systematic list of taxa arranged in generic order

This systematic list follows Breure (1979); the family classification is amended as proposed by Breure and Romero (2012). The latter authors conserved the well-known name Bulimulidae under Art. 35.5 ICZN, although Peltellinae Gray, 1855 falls within the same group. The generic classification has been adapted from Breure (1979) and unpublished data from the senior author. It may be noted that ongoing phylogenetic research may alter the classification. Within the family, genus and species level taxa are presented in alphabetical order.

Family Bulimulidae Tryon, 1867

Auris Spix, 1827

Auris swainsoni Pfeiffer, 1845.


Bostryx Troschel, 1847 sensu Breure 1979 (see also Breure 2012b)

Bostryx acalles Pfeiffer, 1853; Bostryx affinis Broderip in Broderip and Sowerby I 1832; Bostryx agueroi Weyrauch, 1960; Bostryx aileenae Breure, 1978; Bostryx albicans Broderip in Broderip and Sowerby I 1832; Bostryx albicolor Morelet, 1863; Bostryx albus Sowerby I, 1833; Bostryx andoicus Morelet, 1863; Bostryx apodemeta d’Orbigny, 1835; Bostryx atacamensis Pfeiffer, 1856; Bostryx balsanus Morelet, 1863; Bostryx cactorum d’Orbigny, 1835; Bostryx ceratacme Pfeiffer, 1855; Bulimus cercicola Morelet 1863; Bostryx compactus Fulton, 1902; Bostryx conspersus Sowerby I, 1833; Bostryx costatus Weyrauch, 1960; Bostryx costifer Weyrauch, 1960; Bostryx delumbis Reeve, 1849; Bostryx denickei J.E. Gray, 1852; Bostryx dentritis Morelet, 1863; Bostryx depstus Reeve, 1849; Bostryx derelictus Broderip in Broderip and Sowerby I 1832; Bostryx devians Dohrn, 1863; Bostryx emaciatus Morelet, 1863; Bostryx erosus Broderip in Broderip and Sowerby I 1832; Bostryx ferrugineus Reeve, 1849; Bostryx glomeratus Weyrauch, 1960; Bostryx guttatus Broderip in Broderip and Sowerby I 1832; Bostryx hamiltoni Reeve, 1849; Bostryx holostoma Pfeiffer, 1846; Bostryx huascensis Reeve, 1848; Bostryx infundibulum Pfeiffer, 1853; Bostryx kathiae Breure, 1978; Bostryx lactifluus Pfeiffer, 1857; Bostryx lesueureanus Morelet, 1860; Bostryx lichnorum d’Orbigny, 1835; Bostryx limensis Reeve, 1849; Bostryx limonoica d’Orbigny, 1835; Bostryx longinquus Morelet, 1863; Bostryx luridus Pfeiffer, 1863; Bostryx mejillonensis Pfeiffer, 1857; Bostryx metagyra Pilsbry and Olsson 1949; Bostryx minor Weyrauch, 1960; Bostryx modestus Broderip in Broderip and Sowerby I 1832; Bostryx moniezi Dautzenberg, 1896; Bostryx montagnei d’Orbigny, 1837; Bostryx mordani Breure, 1978; Bostryx multispira da Costa, 1904; Bostryx nanus Reeve, 1849; Bostryx nigrolimbatus Pfeiffer, 1853; Bostryx obliquistriatus da Costa, 1901; Bostryx orophilus Morelet, 1860; Bostryx papillatus Morelet, 1860; Bostryx paposensis Pfeiffer, 1856; Bostryx paucicostatus Breure, 1978; Bostryx philippii Pfeiffer, 1842; Bostryx pictus Pfeiffer, 1855; Bostryx pruinosus Sowerby I, 1833; Bostryx pupiformis Broderip in Broderip and Sowerby I 1832; Bostryx pustulosus Broderip in Broderip and Sowerby I 1832; Bostryx radiatus Morelet, 1863; Bostryx reconditus Reeve, 1849; Bostryx rehderi Weyrauch, 1960; Bostryx rhodolarynx Reeve, 1849; Bostryx rodriguezae Weyrauch, 1967; Bostryx rusticellus Morelet, 1860; Bostryx scabiosus Sowerby I, 1833; Bostryx scalaricosta Morelet, 1860; Bostryx scalariformis Broderip in Broderip and Sowerby I 1832; Bostryx serotinus Morelet, 1860; Bostryx simpliculus Pfeiffer, 1855; Bostryx spiculatus Morelet, 1860; Bostryx stenacme Pfeiffer, 1857; Bostryx terebralis Pfeiffer, 1842; Bostryx torallyi d’Orbigny, 1835; Bostryx tricinctus Reeve, 1848; Bostryx tumidulus Pfeiffer, 1842; Bostryx turritus Broderip in Broderip and Sowerby I 1832; Bostryx veruculum Morelet, 1860; Bostryx vilchezi Weyrauch, 1960; Bostryx virgultorum Morelet, 1863; Bostryx voithianus Pfeiffer, 1847; Bostryx woodwardi Pfeiffer, 1857.


Bulimulus Leach, 1814

Bulimulus barbadensis Pfeiffer, 1853; Bulimulus cacticolus Reeve, 1849; Bulimulus dysoni Pfeiffer, 1846; Bulimulus effeminatus Reeve, 1848; Bulimulus erectus Reeve 1849; Bulimulus haplochrous Pfeiffer, 1855; Bulimulus heloica d’Orbigny, 1835; Bulimulus ignavus Reeve, 1849; Bulimulus inutilis Reeve, 1850; Bulimulus istapensis Crosse & Fischer, 1873; Bulimulus juvenilis Pfeiffer, 1855; Bulimulus mollicellus Reeve, 1849; Bulimulus monachus Pfeiffer, 1857; Bulimulus montevidensis Pfeiffer, 1846; Bulimulus nubeculatus Pfeiffer, 1853; Bulimulus pervius Pfeiffer, 1853; Bulimulus pessulatus Reeve, 1848; Bulimulus petenensis Morelet, 1851; Bulimulus pliculatus Pfeiffer, 1857; Bulimulus rubrifasciatus Reeve, 1848; Bulimulus sporadica d’Orbigny, 1835; Bulimulus transparens Reeve, 1849; Bulimulus turritella d’Orbigny, 1835; Bulimulus vesicalis Pfeiffer, 1853.


Drymaeus (Drymaeus) Albers, 1850

Drymaeus (Drymaeus) abruptus Rolle, 1904; Drymaeus (Drymaeus) abscissus Pfeiffer, 1855; Drymaeus (Drymaeus) abyssorum d’Orbigny, 1835; Drymaeus (Drymaeus) aequatorianus E.A. Smith, 1877; Drymaeus (Drymaeus) acervatus Pfeiffer, 1857; Drymaeus (Drymaeus) acuminatus da Costa, 1906; Drymaeus (Drymaeus) alabastrinus da Costa, 1906; Drymaeus (Drymaeus) albolabiatus E.A. Smith, 1877; Drymaeus (Drymaeus) ambustus Reeve, 1849; Drymaeus (Drymaeus) angustus da Costa, 1906; Drymaeus (Drymaeus) antioquiensis Pfeiffer, 1855; Drymaeus (Drymaeus) arcuatostriatus Pfeiffer, 1855; Drymaeus (Drymaeus) auris Pfeiffer, 1866; Drymaeus (Drymaeus) baranguillanus Pfeiffer, 1853; Drymaeus (Drymaeus) bartletti H. Adams, 1867; Drymaeus (Drymaeus) bellus da Costa, 1906; Drymaeus (Drymaeus) bogotensis Pfeiffer, 1855; Drymaeus (Drymaeus) bolivarii d’Orbigny, 1835; Drymaeus (Drymaeus) bolivianus Pfeiffer, 1846; Drymaeus (Drymaeus) boucardi da Costa, 1907; Drymaeus (Drymaeus) bourcieri Pfeiffer, 1853; Drymaeus (Drymaeus) brachysoma d’Orbigny, 1835; Drymaeus (Drymaeus) buckleyi Sowerby III, 1895; Drymaeus (Drymaeus) canaliculatus Pfeiffer, 1845; Drymaeus (Drymaeus) castaneostrigatus da Costa, 1906; Drymaeus (Drymaeus) caucaensis da Costa, 1898; Drymaeus (Drymaeus) chamaeleon Pfeiffer, 1855; Drymaeus (Drymaeus) chimborasensis Reeve, 1848; Drymaeus (Drymaeus) chiriquensis da Costa, 1901; Drymaeus (Drymaeus) clathratus Pfeiffer, 1858; Drymaeus (Drymaeus) coarctatus Pfeiffer, 1845; Drymaeus (Drymaeus) confluens Pfeiffer, 1855; Drymaeus (Drymaeus) convexus Pfeiffer, 1855; Drymaeus (Drymaeus) cuticula Pfeiffer, 1855; Drymaeus (Drymaeus) cuzcoensis Reeve, 1849; Drymaeus (Drymaeus) cylindricus da Costa, 1901; Drymaeus (Drymaeus) dacostae Sowerby III, 1892; Drymaeus (Drymaeus) dombeyanus Pfeiffer, 1846; Drymaeus (Drymaeus) dunkeri Pfeiffer in Philippi 1846; Drymaeus (Drymaeus) elsteri da Costa, 1901; Drymaeus (Drymaeus) exoticus da Costa, 1901; Drymaeus (Drymaeus) expatriatus Preston, 1909; Drymaeus (Drymaeus) fabrefactus Reeve, 1848; Drymaeus (Drymaeus) fallax Pfeiffer, 1853; Drymaeus (Drymaeus) farrisi Pfeiffer, 1858; Drymaeus (Drymaeus) felix Pfeiffer, 1862; Drymaeus (Drymaeus) fenestratus Pfeiffer, 1846; Drymaeus (Drymaeus) flexilabris Pfeiffer, 1853; Drymaeus (Drymaeus) flexuosus Pfeiffer, 1853; Drymaeus (Drymaeus) fucatus Reeve, 1849; Drymaeus (Drymaeus) fusoides d’Orbigny, 1835; Drymaeus (Drymaeus) gealei H. Adams, 1867; Drymaeus (Drymaeus) geometricus Pfeiffer, 1846; Drymaeus (Drymaeus) hidalgoi da Costa, 1898; Drymaeus (Drymaeus) humboldtii Reeve, 1849; Drymaeus (Drymaeus) hygrohylea d’Orbigny, 1835; Drymaeus (Drymaeus) inclinatus Pfeiffer, 1862; Drymaeus (Drymaeus) incognitus da Costa, 1907; Drymaeus (Drymaeus) knorri Pfeiffer in Philippi 1846; Drymaeus (Drymaeus) lattrei Pfeiffer in Philippi 1846; Drymaeus (Drymaeus) linostoma d’Orbigny, 1835; Drymaeus (Drymaeus) lophoica d’Orbigny, 1835; Drymaeus (Drymaeus) lucidus da Costa, 1898; Drymaeus (Drymaeus) marmarina d’Orbigny, 1835; Drymaeus (Drymaeus) murrinus Reeve, 1848; Drymaeus (Drymaeus) musivus Pfeiffer, 1855; Drymaeus (Drymaeus) napo Angas, 1878; Drymaeus (Drymaeus) notabilis da Costa, 1906; Drymaeus (Drymaeus) notatus da Costa, 1906; Drymaeus (Drymaeus) nystianus Pfeiffer, 1853; Drymaeus (Drymaeus) ochrocheilus E.A. Smith, 1877; Drymaeus (Drymaeus) orthostoma E.A. Smith, 1877; Drymaeus (Drymaeus) phryne Pfeiffer, 1863; Drymaeus (Drymaeus) plicatoliratus da Costa, 1898; Drymaeus (Drymaeus) poecila d’Orbigny, 1835; Drymaeus (Drymaeus) praetextus Reeve, 1849; Drymaeus (Drymaeus) protractus Pfeiffer, 1855; Drymaeus (Drymaeus) pseudofusoides da Costa, 1906; Drymaeus (Drymaeus) pulcherrimus H. Adams, 1867; Drymaeus (Drymaeus) punctatus da Costa, 1907; Drymaeus (Drymaeus) quadrifasciatus Angas, 1878; Drymaeus (Drymaeus) recedens Pfeiffer, 1864; Drymaeus (Drymaeus) regularis Fulton, 1905; Drymaeus (Drymaeus) rosenbergi da Costa, 1900; Drymaeus (Drymaeus) saccatus Pfeiffer, 1855; Drymaeus (Drymaeus) scitulus Reeve, 1849; Drymaeus (Drymaeus) scitus H. Adams, 1867; Drymaeus (Drymaeus) selli Preston, 1909; Drymaeus (Drymaeus) serratus Pfeiffer, 1855; Drymaeus (Drymaeus) smithii da Costa, 1898; Drymaeus (Drymaeus) solidus Preston, 1907; Drymaeus (Drymaeus) spadiceus da Costa, 1906; Drymaeus (Drymaeus) spectatus Reeve, 1849; Drymaeus (Drymaeus) strigatus Sowerby I, 1833; Drymaeus (Drymaeus) subhybridus da Costa, 1906; Drymaeus (Drymaeus) subinterruptus Pfeiffer, 1853; Drymaeus (Drymaeus) subventricosus da Costa, 1901; Drymaeus (Drymaeus) sykesi da Costa, 1906; Drymaeus (Drymaeus) tigrinus da Costa, 1898; Drymaeus (Drymaeus) vespertinus Pfeiffer, 1858; Drymaeus (Drymaeus) volsus Fulton, 1907; Drymaeus (Drymaeus) xanthostoma d’Orbigny, 1835; Drymaeus (Drymaeus) yungasensis d’Orbigny, 1837; Drymaeus (Drymaeus) zhorquinensis Angas, 1879; Drymaeus (Drymaeus) ziczac da Costa, 1898; Drymaeus (Drymaeus) zoographica d’Orbigny, 1835.


Drymaeus (Mesembrinus) Albers, 1850

Drymaeus (Mesembrinus) aestivus Pfeiffer, 1857; Drymaeus (Mesembrinus) amandus Pfeiffer, 1855; Drymaeus (Mesembrinus) andicola Pfeiffer, 1847; Drymaeus (Mesembrinus) apicepunctata Preston, 1914; Drymaeus (Mesembrinus) apiculata J.E. Gray, 1834; Drymaeus (Mesembrinus) attenuatus Pfeiffer, 1853; Drymaeus (Mesembrinus) aureolus Guppy, 1866; Drymaeus (Mesembrinus) aurifluus Pfeiffer, 1857; Drymaeus (Mesembrinus) broadwayi E.A. Smith, 1896; Drymaeus (Mesembrinus) bugabensis Martens, 1893; Drymaeus (Mesembrinus) californicus Reeve, 1848; Drymaeus (Mesembrinus) cancellata da Costa, 1906; Drymaeus (Mesembrinus) castus Pfeiffer, 1847; Drymaeus (Mesembrinus) championi Martens, 1893; Drymaeus (Mesembrinus) citronellus Angas, 1879; Drymaeus (Mesembrinus) columbiensis Pfeiffer, 1856; Drymaeus (Mesembrinus) conicus da Costa, 1907; Drymaeus (Mesembrinus) demotus Reeve, 1850; Drymaeus (Mesembrinus) depictus Reeve, 1849; Drymaeus (Mesembrinus) deshayesi Pfeiffer, 1845; Drymaeus (Mesembrinus) discrepans Sowerby I, 1833; Drymaeus (Mesembrinus) dubius Pfeiffer, 1853; Drymaeus (Mesembrinus) dutaillyi Pfeiffer, 1857; Drymaeus (Mesembrinus) electrum Reeve, 1848; Drymaeus (Mesembrinus) erubescens Pfeiffer, 1847; Drymaeus (Mesembrinus) feriatus Reeve, 1850; Drymaeus (Mesembrinus) fidustus Reeve, 1849; Drymaeus (Mesembrinus) flavidulus E.A. Smith, 1877; Drymaeus (Mesembrinus) floridanus Pfeiffer, 1857; Drymaeus (Mesembrinus) fuscobasis E.A. Smith, 1877; Drymaeus (Mesembrinus) gabbi Angas, 1879; Drymaeus (Mesembrinus) gruneri Pfeiffer, 1846; Drymaeus (Mesembrinus) hachensis Reeve, 1850; Drymaeus (Mesembrinus) hepatostomus Pfeiffer, 1861; Drymaeus (Mesembrinus) hoffmanni Martens, 1893; Drymaeus (Mesembrinus) hondurasanus Pfeiffer, 1846; Drymaeus (Mesembrinus) hypozonus Martens, 1893; Drymaeus (Mesembrinus) immaculatus C.B. Adams in Reeve 1850; Drymaeus (Mesembrinus) incarnatus Pfeiffer, 1855; Drymaeus (Mesembrinus) inglorius Reeve, 1848; Drymaeus (Mesembrinus) interruptus Preston, 1909; Drymaeus (Mesembrinus) inusitatus Fulton, 1900; Drymaeus (Mesembrinus) jonasi Pferiffer in Philippi 1846; Drymaeus (Mesembrinus) keppelli Pfeiffer, 1853; Drymaeus (Mesembrinus) koppeli Sowerby III, 1892; Drymaeus (Mesembrinus) laetus Reeve, 1849; Drymaeus (Mesembrinus) lascellianus E.A. Smith, 1895; Drymaeus (Mesembrinus) lirinus Morelet, 1851; Drymaeus (Mesembrinus) lividus Reeve, 1850; Drymaeus (Mesembrinus) loxensis Pfeiffer, 1846; Drymaeus (Mesembrinus) lucidus Reeve, 1848; Drymaeus (Mesembrinus) lusorius Pfeiffer, 1855; Drymaeus (Mesembrinus) manupictus Reeve, 1848; Drymaeus (Mesembrinus) meridanus Pfeiffer, 1846; Drymaeus (Mesembrinus) monilifer Reeve, 1848; Drymaeus (Mesembrinus) moricandi Pfeiffer, 1847; Drymaeus (Mesembrinus) mossi E.A. Smith, 1896; Drymaeus (Mesembrinus) moussoni Pfeiffer, 1853; Drymaeus (Mesembrinus) muliebris Reeve, 1849; Drymaeus (Mesembrinus) nigrofasciatus Pfeiffer in Philippi 1846; Drymaeus (Mesembrinus) nitelinus Reeve, 1849; Drymaeus (Mesembrinus) nitidus Broderip in Broderip and Sowerby I 1832; Drymaeus (Mesembrinus) pallens Reeve, 1849; Drymaeus (Mesembrinus) panamensis Broderip in Broderip and Sowerby I 1832; Drymaeus (Mesembrinus) pervariabilis Pfeiffer, 1853; Drymaeus (Mesembrinus) prestoni da Costa, 1906; Drymaeus (Mesembrinus) primula Reeve, 1848; Drymaeus (Mesembrinus) puellaris Reeve, 1850; Drymaeus (Mesembrinus) rawsonis H. Adams, 1873; Drymaeus (Mesembrinus) rectilinearis Pfeiffer, 1855; Drymaeus (Mesembrinus) roseatus Reeve, 1848; Drymaeus (Mesembrinus) signifer Pfeiffer, 1855; Drymaeus (Mesembrinus) sisalensis Morelet, 1849; Drymaeus (Mesembrinus) sowerbyi Pfeiffer, 1847; Drymaeus (Mesembrinus) studeri Pfeiffer, 1847; Drymaeus (Mesembrinus) subpellucidus E.A. Smith, 1877; Drymaeus (Mesembrinus) sulcosus Pfeiffer, 1841; Drymaeus (Mesembrinus) sulphureus Pfeiffer, 1857; Drymaeus (Mesembrinus) tenuilabris Pfeiffer, 1866; Drymaeus (Mesembrinus) translucens Broderip in Broderip and Sowerby I 1832; Drymaeus (Mesembrinus) trimarianus Martens, 1893; Drymaeus (Mesembrinus) trinitarius E.A. Smith, 1986; Drymaeus (Mesembrinus) tristis Pfeiffer, 1855; Drymaeus (Mesembrinus) tropicalis Morelet, 1849; Drymaeus (Mesembrinus) umbraticus Reeve, 1850; Drymaeus (Mesembrinus) varicosus Pfeiffer, 1853; Drymaeus (Mesembrinus) vincentinus Pfeiffer, 1846; Drymaeus (Mesembrinus) virginalis Pfeiffer, 1856; Drymaeus (Mesembrinus) wintlei Finch, 1929.


Kuschelenia (Kuschelenia) Hylton Scott, 1951

Kuschelenia (Kuschelenia) confusus Reeve, 1848; Kuschelenia (Kuschelenia) culminea d’Orbigny, 1835; Kuschelenia (Kuschelenia) edwardsi Morelt, 1863; Kuschelenia (Kuschelenia) gayi Pfeiffer, 1857; Kuschelenia (Kuschelenia) jussieui Pfeiffer, 1846; Kuschelenia (Kuschelenia) lithoica d’Orbigny, 1835; Kuschelenia (Kuschelenia) thamnoica d’Orbigny, 1835; Kuschelenia (Kuschelenia) tupacii d’Orbigny, 1835.


Kuschelenia (Vermiculatus) Breure, 1978

Kuschelenia (Vermiculatus) anthisanensis Pfeiffer, 1853; Kuschelenia (Vermiculatus) aquilus Reeve, 1848; Kuschelenia (Vermiculatus) badius Sowerby I, 1835; Kuschelenia (Vermiculatus) bicolor Sowerby I, 1835; Kuschelenia (Vermiculatus) caliginosus Reeve, 1849; Kuschelenia (Vermiculatus) coagulatus Reeve, 1849; Kuschelenia (Vermiculatus) cotopaxiensis Pfeiffer, 1853; Kuschelenia (Vermiculatus) filaris Pfeiffer, 1853; Kuschelenia (Vermiculatus) nucinus Reeve, 1850; Kuschelenia (Vermiculatus) ochraceus Morelet, 1863; Kuschelenia (Vermiculatus) peaki Breure, 1978; Kuschelenia (Vermiculatus) petiti Pfeiffer, 1846; Kuschelenia (Vermiculatus) polymorpha d’Orbigny, 1835; Kuschelenia (Vermiculatus) purpuratus Reeve, 1849; Kuschelenia (Vermiculatus) quechuarum Crawford, 1939; Kuschelenia (Vermiculatus) subfasciatus Pfeiffer, 1853.


Naesiotus Albers, 1850 sensu Breure 1979

Naesiotus achatellinus Forbes, 1850; Naesiotus albemarlensis Dall, 1917; Naesiotus apertus Pfeiffer, 1855; Naesiotus catlowiae Pfeiffer, 1853; Naesiotus chamayensis Weyrauch, 1967; Naesiotus chemnitzioides Forbes, 1850; Naesiotus cinereus Reeve, 1849; Naesiotus crepundia d’Orbigny, 1835; Naesiotus curtus Reibisch, 1892; Naesiotus darwini Pfeiffer, 1846; Naesiotus exornatus Reeve, 1849; Naesiotus fernandezae Weyrauch, 1958; Naesiotus fontainii d’Orbigny, 1838; Naesiotus fourmiersi d’Orbigny, 1837; Naesiotus galapaganus Pfeiffer, 1855; Naesiotus irregularis Pfeiffer, 1848; Naesiotus jacobi Sowerby I, 1833; Naesiotus lycodus Dall, 1917; Naesiotus montivaga d’Orbigny, 1835; Naesiotus munsterii d’Orbigny, 1837; Naesiotus nucula Pfeiffer, 1853; Naesiotus nux Broderip, 1832; Naesiotus orbignyi Pfeiffer, 1846; Naesiotus pallidus Reibisch, 1892; Naesiotus paziana d’Orbigny, 1835; Naesiotus perspectivus Pfeiffer, 1846; Naesiotus phlegonis Dall & Ochsner, 1928; Naesiotus quitensis Pfeiffer, 1848; Naesiotus rimatus Pfeiffer, 1847; Naesiotus rivasii d’Orbigny, 1837; Naesiotus rocayana d’Orbigny, 1835; Naesiotus rugiferus Sowerby I, 1833; Naesiotus sugillatus Pfeiffer, 1857; Naesiotus terebra Reibisch, 1892; Naesiotus trichoda d’Orbigny, 1835; Naesiotus unifasciatus Sowerby I, 1833; Naesiotus ustulatus Sowerby I, 1833; Naesiotus ventrosus Reibisch, 1892; Naesiotus verrucosus Pfeiffer, 1855; Naesiotus wolfi Reibisch, 1892.


Neopetraeus Martens, 1885

Neopetraeus altoperuvianus Reeve, 1849; Neopetraeus atahualpa Dohrn, 1863; Neopetraeus binneyanus Pfeiffer, 1857; Neopetraeus cora d’Orbigny, 1835; Neopetraeus decussatus Reeve, 1849; Neopetraeus excoriatus Pfeiffer, 1855; Neopetraeus lobbii Reeve, 1849; Neopetraeus myristicus Reeve, 1849; Neopetraeus patasensis Pfeiffer, 1858; Neopetraeus platystomus Pfeiffer, 1858; Neopetraeus ptychostylus Pfeiffer, 1858.


Newboldius Pilsbry, 1932

Newboldius crichtoni Broderip, 1836; Newboldius illustris Rolle, 1905.


Protoglyptus Pilsbry, 1897

Protoglyptus martinicensis Pfeiffer, 1846; Protoglyptus pilosus Guppy, 1871; Protoglyptus sanctaeluciae E.A. Smith, 1889.


Rabdotus Albers, 1850

Rabdotus juarezi Pfeiffer, 1866; Rabdotus liquabilis Reeve, 1848; Rabdotus ragsdalei Pilsbry, 1890; Rabdotus schiedeanus Pfeiffer, 1841.


Scutalus Albers, 1850

Scutalus baroni (Helix) Fulton, 1896; Scutalus baroni (Bulimulus) Fulton, 1897; Scutalus chiletensis Weyrauch, 1967; Scutalus cretaceus Pfeiffer, 1855; Scutalus grandiventris Weyrauch, 1960; Scutalus latecolumellaris Preston, 1909; Scutalus proteus Broderip in Broderip and Sowerby I 1832; Scutalus versicolor Broderip in Broderip and Sowerby I 1832.


Stenostylus Pilsbry, 1898

Stenostylus meleagris Pfeiffer, 1853; Stenostylus nigrolimbatus Pfeiffer, 1853.


Nomina inquirenda

clarus Pfeiffer, 1857; dukinfieldi Melvill, 1900; gelidus Reeve, 1849; nivalis d’Orbigny, 1835; pallens Reeve, 1849; sowerbyi Pfeiffer, 1847.


Alphabetic list of taxa by species name
Bulimulus (Drymaeus) abruptus Rolle, 1904

http://species-id.net/wiki/Bulimulus_abruptus

Figs 26J–L, L1i
Bulimulus (Drymaeus) abruptus Rolle 1904: 35.
Type locality.

”Huancabamba in Peru”; see remarks.

Label.

”Huancabamba, / Peru 1904”; in Rolle’s handwriting. Another label in pencil indicating that it came from the Fulton collection.

Dimensions.

”Alt. 44, diam. max. 24 (..) mm”; figured specimen herein H 42.5, D 21.5 W 6.3.

Type material.

NHMUK 1947.2.10.1, syntype (ex Rolle).

Remarks.

Rolle did not state on how many specimens his description was based upon. However, the shell height is close to the original measurements and together with the indication ”type” on the label in Rolle’s handwriting, there is no doubt about the status of the specimen. The locality is ambiguous, as there are several places called ”Huancabamba” in Peru. There were, however, several other species described from the same locality, amoung them Columbinia huancabambensis, which is regarded by Loosjes and Loosjes-van Bemmel (1984: 33), as occurring in northern Peru. Another species described in the same paper, Systrophia moellendorffii, was said by Haas (1955: 367) to have been rediscovered in the Chanchamayo valley [Dept. Pasco]. Given the fact that a third species described by Rolle—Newboldius illustris—is known to occur in the same region, makes it likely that Huancabamba in Dept. Pasco is the locality where Rolle’s taxon occurs. This species was hitherto considered as published in 1905 (Richardson 1995: 95); however, on the copy in the Smithsonian Institution Libraries the title page of the issue in which Rolle’s paper appeared is marked ”Rec’d Mar. 2/04” (see http://www.biodiversitylibrary.org/item/53294#page/211/mode/1up); p. 48 of this issue mentions ”Ausgegeben 22. Februar”.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) abruptus (Rolle, 1904).

Bulimus abscissus Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_abscissus

Figs 26A–C, L1ii
Bulimus abscissus Pfeiffer 1855f: 116; Pfeiffer 1859b: 376; Breure 1979: 105 (lectotype designation).
Type locality.

”Province of Quito, Ecuador”.

Label.

”Ecuador”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 28, diam. 13 mill.”; lectotype H 27.8, D 14.3, W 5.5.

Type material.

NHMUK 1975497, lectotype; 1975498 five paralectotypes (Cuming coll.).

Remarks.

Pfeiffer did not state on how many specimens his description was based; no variety has been mentioned by him. Two sets of labels are with this lot, which is originating from the Cuming collection; one reading ”Ecuador” in the handwriting of one of Cuming’s secretaries. On both labels the taxon name is in Pfeiffer’s handwriting. One of the lots has been labelled ”abscissus var.” in a later handwriting.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) abcissus (Pfeiffer, 1855).

Helix abyssorum d’Orbigny, 1835

http://species-id.net/wiki/Helix_abyssorum

Figs 27A–C, L1iii
Helix abyssorum d’Orbigny 1835: 17.
Bulimus abyssorum; d’Orbigny 1837 [1834–1847]: 308, pl. 39 figs 7–8 [7 Aug. 1837; text 6 May 1838]; Gray 1854: 21.
Drymaeus abyssorum; Pilsbry 1898 [1897–1898]: 192, pl. 37 figs 3–4; Breure 1975b: 1149, pl. 7 fig. 2 (lectotype designation); Cuezzo et al. 2013: 153.
Drymaeus (Drymaeus) hygrohylaeus; Miquel 1989b: 77, fig. 1.
Type locality.

”provincia Lagunacensi (republica Boliviana)”; see Breure 1973: 113.

Label.

”pampa ruis, Pce Laguna, Bolivia”, in d’Orbigny’s handwriting.

Dimensions.

”Longit. 52 mil., latit. 25 millim.”; figured specimen herein H 43.7 D 23.3, W 6.2.

Type material.

NHMUK 1854.12.4.125, five paralectotypes (d’Orbigny coll.).

Remarks.

d’Orbigny did not state on how many specimens his description was based. The types series appears to have been split between the MNHN and NHMUK collections. None of the specimens in London match the original figures, while one of the specimens in Paris is close to the published dimensions. The current systematic position follows the synonymisation of this taxon with Helix hygrohylea d’Orbigny, 1835, by Miquel (1989b: 77); this author, however, overlooked that Helix abyssorum has page priority. Cuezzo et al. (2013) disageed with this synonymisation.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) abyssorum (d’Orbigny, 1835).

Bulimus acalles Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_acalles

Figs 6A, L1iv
Bulimus acalles Pfeiffer 1853b [7 Dec.]: 258; Pfeiffer 1853d [31 Dec.]: 410; Pfeiffer 1853 [31 Dec.] in Küster and Pfeiffer 1840–1865: 84, pl. 30 figs 27–28.
Bulimulus (Lissoacme) acalles; Pilsbry 1896 [1895–1896]: 160, pl. 50 figs 53–54.
Type locality.

”in Andibus Peruvianis”.

Label.

”Peru”. M.C. label style I.

Dimensions.

”Long. 10, diam. 6 mill.”; figured specimen herein H 13.2, D 7.7, W 5.3.

Type material.

NHMUK 20100651, two possible syntypes (Cuming coll.).

Remarks.

The specimens found are slightly larger than the original measurements; the largest has shell height 13.2 mm. The material—not accompanied by a taxon label in Pfeiffer’s handwriting—is from the Cuming collection, on which Pfeiffer based the description for this taxon; he did not state on how many specimens his description was based. The specimens are treated as possible syntypes.

Current systematic position.

Bulimulidae, Bostryx acalles (Pfeiffer, 1853).

Bulimus acervatus Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_acervatus

Figs 50A–C, L1v
Bulimus acervatus Pfeiffer 1857a: 157.
Drymaeus (Drymaeus) acervatus; Breure and Eskens 1981: 4, pl. 8 fig. 6 (lectotype designation).
Type locality.

”Brasilia. (Strain)”.

Label.

”Brazils. Mr Strain”; taxon name in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 41, diam. 21 mill.”; lectotype H 41.5, D 24.7, W 6.5.

Type material.

NHMUK 1975461, lectotype (Cuming coll.).

Remarks.

The current systematic position follows Richardson (1995: 96).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) acervatus (Pfeiffer, 1857).

Bulimus achatellinus Forbes, 1850

http://species-id.net/wiki/Bulimus_achatellinus

Figs 14J, L2i
Bulimus achatellinus Forbes 1850: 56, pl. 9 figs 5a–b; Pfeiffer 1853 in Küster and Pfeiffer 1840–1865: 93, pl. 31 figs 21-23.
Bulimulus achatellinus; Pilsbry 1897 [1897–1898]: 99, pl. 16 figs 28–29.
Naesiotus achatellinus; Breure and Borrero 2008: 10.
Type locality.

[Ecuador, Galápagos, Isla San Cristóbal] ”Chatham Island, Gelepagos”.

Label.

”Chatham Is Galapagos”, added on a different label with E.A. Smith’s handwriting. The taxon name is on a second label ”Bulimus achatenellinus [sic] Forbes, figured specimen, Proc Zool Soc (...) fig. 5”. M.C. label style III.

Dimensions.

”Long. 19, diam. 10 mill.”; holotype H 19, D 9.52, W 8.7.

Type material.

NHMUK 1855.4.5.25, holotype (Cuming coll.).

Remarks.

Forbes writes ”This shell (...)”, therefore the single specimen found, together with the indication on the label that it was figured, makes it justifiable to consider it a holotype.

Current systematic position.

Bulimulidae, Naesiotus achatellinus (Forbes, 1850).

Drymaeus acuminatus da Costa, 1906

http://species-id.net/wiki/Drymaeus_acuminatus

Figs 45A–C, L2ii
Drymaeus acuminatus da Costa 1906a: 8, pl. 1 fig. 4; Simone 2006: 135, fig. 440.
Drymaeus (Drymaeus) acuminatus; Breure 1979: 106.
Type locality.

”Matto Grosso, Brazil”.

Label.

”Matto Grosso / Brazil”; in da Costa’s handwriting.

Dimensions.

”Long. 33, diam. 14 mm.”; holotype H 33.4, D 14.1, W 7.4.

Type material.

NHMUK 1907.11.21.10, holotype (da Costa coll.).

Remarks.

da Costa wrote ”A rather peculiar shell, unlike any known to the writer, who obtained it at the dispersal of the collection of the late Mr. Miers”; given the singular form, it may be assumed that da Costa only had one specimen at hand.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) acuminatus da Costa, 1906.

Bulimus (Drymaeus) aequatorianus E.A. Smith, 1877

http://species-id.net/wiki/Bulimus_aequatorianus

Figs 28E–G, L2iii
Bulimus (Drymaeus) aequatorianus E.A. Smith 1877: 363, pl. 30 fig. 7.
Drymaeus (Drymaeus) aequatorianus; Breure 1979: 106 (lectotype designation); Breure and Eskens 1981: 4; Breure and Borrero 2008: 19.
Type locality.

”Ecuador”.

Label.

”Ecuador”. M.C. label style III.

Dimensions.

”Long. 26 1/2 mill., diam. 11”; lectotype H 26.6, D 13.4, W 6.1.

Type material.

NHMUK 1975137, lectotype. NHMUK 1975138, three paralectotypes (Cuming coll.).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) aequatorianus (E.A. Smith, 1877).

Bulimus aequatorius Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_aequatorius

Figs 69A–B, L2v–vi
Bulimus aequatorius Pfeiffer 1853d: 420; Pfeiffer 1854 in Küster and Pfeiffer 1840–1865: 101, pl. 13 figs 1–4; Pfeiffer 1854c: 155; Breure 1979: 85 (lectotype designation).
Bulimulus aequatorius; Pilsbry 1897 [1897–1898]: 30, pl. 7 figs 1–5.
Scutalus (Vermiculatus) aequatorius; Breure and Borrero 2008: 18.
Type locality.

”in republica Aequatorius, in monte Schinchulagua Equador”.

Label.

”Woods on the / mountain / Schinchulaqua / Equador”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 34, diam. 17 mill.”. Figured specimen H 37.8, D 18.2, W 6.5.

Type material.

NHMUK 1975377, lectotype; 1975378, two paralectotypes (Cuming coll.).

Remarks.

Pfeiffer did no indicate on how many specimens his description was based. He described two varieties from ”Chimborazo (Bourcier)”. The second label in the lot indicates ”var.” and ”Chimborazo, Equador / Mons. Bourcier / Consul General”. The systematic position at the species level follows Richardson (1995: 345).

Current systematic position.

Bulimulidae, Kuschelenia (Vermiculatus) aequatorius (Pfeiffer, 1853) (comb. n.).

Bulimus aestivus Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_aestivus

Figs 19A–B, L2iv
Bulimus aestivus Pfeiffer 1857c: 331.
Drymaeus (Drymaeus) aestivus; Breure 1979: 106.
Type locality.

”Meobamba, Peru (Mr. Gueinzius)”.

Label.

”Ecuador”; taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 17, diam. 7 1/3 mill.”; syntype H 16.9 D 7.9, W 6.4.

Type material.

NHMUK 1975462, syntype (Cuming coll.).

Remarks.

Pfeiffer described this taxon from Cuming’s collection, but did not state on how many specimens his description was based. The specimen found corresponds to the shell height given by Pfeiffer. This is a case where the locality information provided by Cuming to the describer (viz. Pfeiffer) has been lost; the label ”Ecuador” on the frontside of the wooden tablet has been added in a later handwriting and may be erroneous.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) aesitvus (Pfeiffer, 1857) (comb.n.).

Bulinus affinis Broderip in Broderip and Sowerby I 1832

http://species-id.net/wiki/Bulinus_affinis

Figs 8A, L3i–ii
Bulinus affinis Broderip in Broderip and Sowerby I 1832b: 106; Sowerby I 1833 [Sowerby I and II 1832–1841]: fig. 30.
Bulimus affinis; Reeve 1848 [1848–1850]: pl. 23 fig. 154.
Type locality.

”in Peruviâ (Mexillones, desert of Atacama)”.

Label.

”Peru”; taxon name in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”long 1, lat. 1/2 poll. [H 25.3, D 12.7 mm]”; figured specimen herein H 27.5, D 12.6, W 8.3.

Type material.

NHMUK 20100610, five possible syntypes (Cuming coll.).

Remarks.

Broderip described this taxon from the Cuming collection, but did not indicate on how many specimens his description was based. These specimens are herein regarded as possible syntypes; one is corresponding to Reeve’s figure. The presence of a label in Pfeiffer’s handwriting together with specimens apparently seen by Reeve as evidenced by the figured specimen, corroborates the assumption (Breure and Ablett 2011: 5) that—at least in some cases—material from the Cuming collection was either returned after having been sent to Pfeiffer or Pfeiffer made the identification during a visit to London.

Current systematic position.

Bulimulidae, Bostryx affinis (Broderip in Broderip and Sowerby I 1832).

Bostryx (Peroneus) agueroi Weyrauch, 1960

http://species-id.net/wiki/Bostryx_agueroi

Fig. 4E
Bostryx (Peroneus) agueroi Weyrauch 1960b: 126, pl. 12 figs 39–41; Neubert and Janssen 2004: 198, pl. 9 fig. 96.
Bostryx agueroi Breure 1979: 51.
Type locality.

”Mittel-Peru, Fundo Yacca, auf der rechten Seite des Río Cañete, an der Autostraße von Cañete nach Yauyos, 2300 m”.

Label.

” WW 1462-A/ 2 Paratypes” ”C-Peru: Yacca, 2300m, Rio Canete, leg. W. Weyrauch”; printed label.

Dimensions.

”H. 22, 8 D. 8, 5”; figured specimen herein H 21.1, D 7.4, W 7.3.

Type material.

NHMUK 1975333, two paratypes (ex Weyrauch).

Current systematic position.

Bulimulidae, Bostryx agueroi Weyrauch, 1960.

Bostryx aileenae Breure, 1978

http://species-id.net/wiki/Bostryx_aileenae

Fig. 5A
Bostryx aileenae Breure 1978: 49, pl. 1 fig. 6; Breure 1979: 51.
Type locality.

”Peru, Dept. Lima, Río Cañete valley, 1 km above Puente Auco, 2070 m”.

Label.

”Peru, Dept. Lima, Río Cañete valley, 1 km above Puente Auco, 2070 m. Sta. 79”.

Dimensions.

”shell height 14.9 mm, diameter 7.6 mm”; figured specimen herein H 13.7, D 7.0, W 5.9.

Type material.

NHMUK 1975229, three paratypes (Breure leg.).

Current systematic position.

Bulimulidae, Bostryx aileenae Breure, 1978.

Drymaeus alabastrinus da Costa, 1906

http://species-id.net/wiki/Drymaeus_alabastrinus

Figs 43J–L, L3iii
Drymaeus alabastrinus da Costa 1906b: 98, pl. 11 fig. 4.
Drymaeus (Drymaeus) alabastrinus; Breure 1979: 106.
Type locality.

”Honda, Colombia”.

Label.

”Honda, Colombia”; in da Costa’s handwriting.

Dimensions.

”Long. 33, diam. 15 mill.”; figured specimen herein H 33.0, D 15.3, W 7.6.

Type material.

NHMUK 1907.11.21.16, holotype (da Costa coll.).

Remarks.

da Costa did not clearly state on how many specimens his description was based. However, since he mentioned ”a shell” it may be safely assumed that he had only one specimen at hand.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) alabastrinus da Costa, 1906.

Bulimulus (Naesiotus) albemarlensis Dall, 1917

http://species-id.net/wiki/Bulimulus_albemarlensis

Figs 13F, L3iv
Bulimulus (Naesiotus) albemarlensis Dall 1917b: 377.
Naesiotus albemarlensis; Breure 1979: 67; Breure and Borrero 2008: 10.
Type locality.

[Ecuador, Galápagos, Isla Isabela] ”near Villamil, at 2300 to 3300 feet elevation”.

Label.

”Galapagos Isl. / Albemarle Isl.”.

Dimensions.

”Length of shell 15; diameter 9 mm”; figured specimen herein H 14.7, D 10.0, W 5.8.

Type material.

NHMUK 1937.6.18.13-16, three possible paratypes, California Academy of Sciences Expedition 1905–1906 leg. (ex Schlesch).

Remarks.

The type status of these specimens is not fully resolved and is here viewed with much caution, as it remains unclear how Schlesch obtained specimens from the original series.

Current systematic position.

Bulimulidae, Naesiotus albemarlensis (Dall, 1917).

Bulinus albicans Broderip in Broderip and Sowerby I 1832

http://species-id.net/wiki/Bulinus_albicans

Figs 7I, L3v–vi
Bulinus albicans Broderip in Broderip and Sowerby I 1832b: 105; Sowerby I 1833 [Sowerby I and II 1832–1841]: figs 22–22*.
Bulimus albicans; Reeve 1848 [1848–1850]: pl. 22 fig. 141.
Type locality.

”Copiapo, Chili”.

Label.

”Chili”; taxon name in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”long. 9/12, lat. 7/12 poll. [H 19, D 14.8 mm]”; figured specimen herein H 22.1, D 13.1, W 5.7.

Type material.

NHMUK 20100611, five possible syntypes (Cuming coll.).

Remarks.

Broderip described this taxon from the Cuming collection, but did not indicate on how many specimens his description was based. These specimens are herein regarded as possible syntypes; one is corresponding to Reeve’s figure. This is another case of material that has been studied by both Reeve and Pfeiffer. See also affinis Broderip, 1832.

Current systematic position.

Bulimulidae, Bostryx albicans (Broderip in Broderip and Sowerby I 1832).

Bulimus albicolor Morelet, 1863

http://species-id.net/wiki/Bulimus_albicolor

Figs 4F, L3vii
Bulimus albicolor Morelet 1863: 199, pl. 11 fig. 9.
Bulimulus (Peronaeus) albicolor; Pilsbry 1896 [1895–1896]: 148, pl. 46 figs 49–50.
Bostryx albicolor; Breure 1979: 51.
Type locality.

[Peru, Dept. Ayacucho] ”Huanta et de la vallée de l’Apurimac”.

Label.

”Pérou / Apurimac, Huanta”; in Morelet’s handwriting.

Dimensions.

”Longit 28, diam. 9 mm”; figured specimen herein H 28.2, D 10.92, W 8.3.

Type material.

NHMUK 1893.2.4.169–170, two syntypes (Morelet coll.).

Remarks.

Further syntype material is present in the MHNG and RBINS collections. See also Breure (2011). The current systematic position follows Richardson (1995: 36).

Current systematic position.

Bulimulidae, Bostryx orophilus (Morelet, 1860).

Bulimus albolabiatus E.A. Smith, 1877

http://species-id.net/wiki/Bulimus_albolabiatus

Figs 29G–I, L3viii
Bulimus (Drymaeus) albolabiatus E.A. Smith 1877: 363, pl. 39 fig. 4.
Drymaeus albolabiatus; Pilsbry 1898 [1897–1898]: 201, pl. 36 fig. 37.
Drymaeus (Drymaeus) albolabiatus; Breure 1979: 106; Breure and Borrero 2008:
Type locality.

”Malacatos, South Ecuador” [Prov. Loja].

Label.

”Malacatos, S. Ecuador”; both locality and taxon name in Smith’s handwriting.

Dimensions.

Not given; figured specimen herein H 34.0, D 14.2, W 6.3.

Type material.

NHMUK 1877.3.28.3, holotype.

Remarks.

According to the label, a paratype has been ”smashed [in] 1896”. The current systematic position follows Richardson (1995: 96).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) albolabiatus (E.A. Smith, 1877).

Bulinus albus Sowerby I, 1833

http://species-id.net/wiki/Bulinus_albus

Figs 7J, L3ix–x
Bulinus albus Broderip [sic]; Sowerby I 1833 [Sowerby I and II 1832–1841]: fig. 51 [19 July]; Sowerby I 1833b: 73 [20 Sept.] (see remarks).
Type locality.

[Chile] ”Copiapo”.

Label.

”Chili”. M.C. label style I.

Dimensions.

”long. 0.8, lat. 0.5 poll. [H 20.3, D 12.7 mm]”; figured specimen herein H 15.0, D 10.7, W 4.7.

Type material.

NHMUK 1842.5.10.149–151, three possible syntypes (Cuming coll.).

Remarks.

The name was introduced in Sowerby I 1833 [Sowerby I and II 1832–1841] as ”B. albus Brod.”. However, as can be seen in Sowerby I (1833 in Sowerby I and II 1832–1841: 73), the authorship should be credited to G.B. Sowerby I (”G.B.S.”). It is not known if this species was described from the Cuming collection, but this cannot be excluded a priori (see also Breure and Ablett 2011: 3). One specimen is closely resembling Sowerby’s figure. The material is herein considered as possible syntypes. The registration numbers suggest that six specimens were originally present; only three of these could be found.

As mentioned before, the publication dates of Sowerby’s publications are according to Coan et al. (2013a) and Duncan (1937). Interestingly, d’Orbigny 1838 [1834–1847]: 280 mentioned this taxon as ”Bulimus albus Broderip”, with reference to Sowerby I (1833 in Sowerby I and II 1832–1841), and said it had publication date 11 June 1833 [contra Duncan 1937: 78, who reports 20 September 1833]. At the same time he noticed that Duclos had published Bulimus olorinus on 15 June 1833 (Duclos 1833: pl. 24). The text pertaining to this plate describes this taxon from ”Chili”, based on a specimen from Duclos’ collection and collected by Gay. He signed the text with ”15 juin 1833”; however, it seems likely from the context that this was the date Duclos finished his description and not the date the text appeared in print. Without additional bibliographical research it seems premature to give priority to Duclos’ taxon. The current systematic position follows Richardson (1995: 15).

Current systematic position.

Bulimulidae, Bostryx albus (Sowerby I, 1833).

Bulimus altoperuvianus Reeve, 1849

http://species-id.net/wiki/Bulimus_altoperuvianus

Figs 56A–C, L4i
Bulimus altoperuvianus Reeve 1849 [1848–1850]: pl. 72 figs 521a–b; Breure 1979: 100.
Neopetraeus altoperuvianus; Pilsbry 1898 [1897–1898]: 173, pl. 32 figs 30–31; Breure 1978: 209 (lectotype designation).
Type locality.

[Peru, Dept. Amazonas] ”Chachapoyas, Alto-Peru”.

Label.

”Province of Caxamarca [Dept. Cajamarca], Peru”. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 46.5, D 25.4, W 7.2.

Type material.

NHMUK 1975437, lectotype; 1975438, one paralectotype (Cuming coll.).

Remarks.

The lectotype corresponds to Reeve’s fig. 521a, while fig. 521b corresponds to the paralectotype.

Current systematic position.

Bulimuludae, Neopetraeus altoperuvianus (Reeve, 1849).

Bulimus amandus Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_amandus

Figs 19C, L4ii
Bulimus amandus Pfeiffer 1855d: 96, pl. 31 fig. 4; Breure 1979: 117 (lectotype designation).
Drymaeus (Mesembrinus) amandus; Breure and Eskens 1981: 50.
Type locality.

”Venezuela”.

Label.

No locality given. No taxon label in Pfeiffer’s handwriting.

Dimensions.

”Long. 30, diam. 11 1/2 mill.”; lectotype H 29, D 13.1, W 6.3.

Type material.

NHMUK 1997457, lectotype (Cuming coll.).

Remarks.

The lectotype corresponds to Pfeiffer’s figure, although the colours have faded. The current systematic position corresponds to Richardson (1995: 97).

Current systematic position.

Drymaeus (Mesembrinus) amandus (Pfeiffer, 1855).

Bulimus ambustus Reeve, 1849

http://species-id.net/wiki/Bulimus_ambustus

Figs 40J–L, L4iii
Bulimus ambustus Reeve 1849 [1848–1850]: pl. 74 fig. 535.
Drymaeus ambustus; Pilsbry 1898 [1897–1898]: 264, pl. 46 figs 66–67.
Drymaeus (Drymaeus) ambustus; Breure and Eskens 1981: 5.
Type locality.

”—?”.

Label.

[Ecuador] ”Between Jacunga [Prov. Cotapaxi, Latacunga] and / [Prov. Tungurahua] Ambato”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 29.0, D 13.1, W 6.3.

Type material.

NHMUK 1975441/1–3, lectotype and two paralectotypes, Bourcier leg. (Cuming coll.).

Remarks.

Although the label given is more precise than the locality given in Reeve’s publication and the specimens bear a label in Pfeiffer’s handwriting, one of the shells does corresponds to the specimen figured by Reeve. The current systematic position follows Richardson (1995: 97).

Current systematic position.

Drymaeus (Drymaeus) ambustus (Reeve, 1849).

Bulimus andicola Pfeiffer, 1847

http://species-id.net/wiki/Bulimus_andicola

Figs 23A, L4iv
Bulimus andicola Pfeiffer 1847: 115; Reeve 1848 [1848–1850]: pl. 55 fig. 364; Breure 1979: 117 (lectotype designation).
Bulimulus (Lissoacme) andicola; Pilsbry 1896 [1895–1896]: 166, pl. 50 fig. 46.
Type locality.

”Columbian Andes”.

Label.

”Andes of Columbia”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.

Dimensions.

”Long. 24, diam. 11 mill.”; figured specimen herein H 24.1, D 10.7, W 7.1.

Type material.

NHMUK 1975315, lectotype; NHMUK 1975316, two paralectotypes (Cuming coll.).

Current systematic position.

Drymaeus (Mesembrinus) andicola (Pfeiffer, 1847).

Bulimus andoicus Morelet, 1863

http://species-id.net/wiki/Bulimus_andoicus

Figs 10G, L4v
Bulimus andoicus Morelet 1863: 198, pl. 11 fig. 13; Breure 1979: 50.
Bulimulus (Lissoacme) andoicus; Pilsbry 1896 [1895–1896]: 147, pl. 46 figs 42–44.
Bostryx andoicus; Breure 1978: 50 (lectotype designation).
Type locality.

[Peru] ”vallées (..) d’Ayacucho”.

Label.

”Pérou, Pomacoche”, in Morelet’s handwriting.

Dimensions.

”Long. 26–30; diam. 9–10 mill.”; figured specimen herein H 29.5, D 12.12, W 7+.

Type material.

NHMUK 1893.2.4.171, lectotype (Morelet coll.).

Remarks.

The type locality was broadly defined by Morelet and probably also covers adjacent parts of Dept. Apurimac. There are two places called ”Pomacocha” in the valley of Río Apurimac, one in Dept. Ayacucho and the other in Dept. Apurimac. Given the range in dimensions given by Morelet, it is clear that he had a larger type series at hand. The top of the lectotype is slightly damaged. There are additional lots in the MHNG collection, as described by Breure 1978. The current systematic position corresponds to Richardson (1995: 16).

Current systematic position.

Bulimulidae, Bostryx andoicus (Morelet, 1863).

Drymaeus angustus da Costa, 1906

http://species-id.net/wiki/Drymaeus_angustus

Figs 37L–N, L4vi
Drymaeus angustus da Costa 1906a: 9, pl. 1 figs 7–8; Linares and Vera 2012: 181.
Drymaeus (Drymaeus) angusta [sic]; Breure 1979: 106.
Type locality.

[Colombia] Bogotá.

Label.

”Bogota”; in da Costa’s handwriting.

Dimensions.

”Long. 31.5, diam. 10”; figured specimen herein H 31.5, D 10.4, W 6.5.

Type material.

NHMUK 1907.11.21.14, holotype (da Costa coll.).

Remarks.

da Costa wrote ”this unique specimen”, which makes the specimen found the holotype. The taxon name on the label reads ”angusta”, but the name ”angustus” was published. The top of the holotype is slightly damaged.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) angustus angustus da Costa, 1906.

Bulimus anthisanensis Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_anthisanensis

Figs 69E–F, 69I, L5i
Bulimus anthisanensis Pfeiffer 1853d: 406; Pfeiffer 1854 in Küster and Pfeiffer 1840–1865: 104, pl. 33 figs 20–21; Breure 1979: 85.
Bulimulus anthisanensis; Pilsbry 1897 [1897–1898]: 32, pl. 4 figs 41–42.
Scutalus (Vermiculatus) anthisanensis; Breure 1978: 170, pl. 8 fig. 8 (lectotype designation); Breure and Borrero 2008: 18.
Type locality.

[Ecuador, Prov. Napo] ”monte Anthisana (44000’ [sic]) reipublicae Aequatoris (Bourcier)”.

Label.

”the Anthisana / 14000 feet high / Monsr Bourcier / Consul General”; taxon name in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 40, diam. 17 mill. (Mus. Cuming)”; figured specimen herein H 40.2, D 19.5, W 6.6.

Type material.

NHMUK 1975372, lectotype; 1975373, three paralectotypes (Cuming coll.).

Remarks.

The published altitude at which the type material was collected is clearly in error, as evidenced by the label information.

Current systematic position.

Bulimulidae, Kuschelenia (Vermiculatus) anthisanensis (Pfeiffer, 1853) (comb. n.).

Bulimus antioquensis Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_antioquensis

Figs 31D–F, L5ii
Bulimus antioquensis Pfeiffer 1855b: 291; Pfeiffer 1859b: 394; Breure 1979: 106 (lectotype designation).
Drymaeus (Drymaeus) antioquensis; Breure and Eskens 1981: 6.
Drymaeus antioquensis; Linares and Vera 2012: 181.
Type locality.

[Colombia] ”Province of Antioquia, New Granada. (Schlim)”.

Label.

”Province of Antioquia / Monsr Schlim”; taxon name in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 30, diam. 13 mill.”; figured specimen herein H 29.6, D 16.4, W 6.1.

Type material.

NHMUK 1975450, lectotype, Schlim leg. (Cuming coll.).

Remarks.

This taxon is now considered a junior subjective synonym of Bulimus baranguillanus Pfeiffer, 1853 (syn. n.). The current systematic position is based on a revision of Colombian Drymaeus species (Breure and Borrero, unpublished data).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) baranguillanus (Pfeiffer, 1853).

Bulimus apertus Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_apertus

Figs 15D, L5iii
Bulimus apertus Pfeiffer in Dunker et al. 1855: 107.
Naesiotus apertus Breure 1979: 68 (lectotype designation).
Type locality.

”?”.

Label.

”Andes of Peru”; taxon name in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 19, diam. 10 mill.”; figured specimen herein H 18.1 D 9.23 W 6.5.

Type material.

NHMUK 1975317, lectotype (Cuming coll.).

Remarks.

This is the first time a locality for this taxon is published; Bulimus apertus is now considered a junior subjective synonym of Bulimus orbignyi Pfeiffer, 1846 (syn. n.), whose type locality is ”Bolivia” (see under Bulimus orbignyi Pfeiffer). Any specific Peruvian locality for this taxon may likely be expected in one of the southern departments, close to the Bolivian border.

Current systematic position.

Bulimulidae, Naesiotus orbignyi (Pfeiffer, 1846).

Bulimulus apicepunctata Preston, 1914

http://species-id.net/wiki/Bulimulus_apicepunctata

Figs 23J–K, L5iv
Bulimulus apicepunctata Preston 1914: 523; Breure 1979: 117.
Drymaeus (Mesembrinus) apicepunctata; Breure and Eskens 1981: 52.
Type locality.

”Santa Rita, E. Peru”.

Label.

”Santa Rita, E. Peru”.

Dimensions.

”Alt. 17.5, diam. maj. 9, diam. min. 7 mm”; figured specimen herein H 17.5, D 8.3, W 6.6.

Type material.

NHMUK 1915.1.6.23, holotype (ex Preston).

Remarks.

Preston did not state on how many specimens his description was based; the reference of Breure (1979) ”HT 1915.1.6.23” has to be interpreted as lectotype designation under Art. 74.6 ICZN. This is the first time the type has been figured.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) apicepunctata (Preseton, 1914).

Bulimus apiculatus J.E. Gray, 1834

http://species-id.net/wiki/Bulimus_apiculatus

Figs 23B, L5v
Bulimus apiculatus J.E. Gray 1834: 66; Breure 1979: 117.
Type locality.

Not given.

Label.

No locality given.

Dimensions.

”Axis 10, diam. 4 1/4 lin.” [H 21.1, D 9.0 mm]; figured specimen herein H 20.2, D 9.5, W 6.2.

Type material.

NHMUK 1982297, holotype (see remarks).

Remarks.

Gray (1834) refers to the type material as ”This shell...”, which may imply that he had only one specimen at hand. It is a juvenile shell, with the lip damaged. The taxon was considered to be a junior subjective synonym of Helix elongata Röding, 1789 by Pilsbry (1899: 25), with which we concur.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) elongatus (Röding, 1789).

Helix apodemeta d’Orbigny, 1835

http://species-id.net/wiki/Helix_apodemeta

Figs 9A–B, L5vi
Helix apodemeta d’Orbigny 1835: 10.
Bulimus apodemetes d’Orbigny 1837 [1834–1847]: 279, pl. 30 figs 5–8 [3 April 1837; text 23 April 1838]; Gray 1854: 16.
Bulimulus (Lissoacme) apodemetus; Pilsbry 1896 [1895–1896]: 187, pl. 51 fig. 1–3.
Bulimulus (Bulimulus) apodemetus; Breure 1975b: 1145.
Type locality.

”republica Argentina; republica Boliviana”; see Breure 1973: 114.

Label.

”Monte grande, chiquitos (Bolivia)” [.178, . 181]; ”Pampa ruis (laguna) Bolivia” [.179]; ”Vilna (Valle grande) Bolivia” [.180]; ”Sn Lorenzo Parana Rep. argentina” [.182], all in in d’Orbigny’s handwriting.

Dimensions.

”Assez variables; les plus along´s ont, de longueur, 28 millimètres sur 18 de largeur, tandis que les plus courts offerent 23 millimètres de longueur, sur 12 de largeur”; largest specimen H 28.3, shortest H 22.7, figured specimen H 23.0, D 12.6, W 6.2 [.182].

Type material.

NHMUK 1854.12.4.178–182, five lots with resp. eight, seven, seven, three and three syntypes (d’Orbigny coll.).

Remarks.

d’Orbigny (1935) did not state on how many specimens his description was based. In d’Orbigny (1838 [1834–1847]: 280) the localities were specified as ”sur les coteaux du Parna, province d’Entre-rios, près de Feliciano (...) près de San-Lorenzo, province de Santa-Fe. (...) la république de Bolivia (...) des provinces de Valle grande et de la Laguna (...); (...) des plaines de Santa-Cruz de la Sierra, (...) la province de Chiquitos”; see also Breure 1973. The shell of the largest specimen among the lots, in. 178, is broken on the ventral side. The protoconch of the type material proves to be smooth and the taxon is now being transferred from the genus Bulimulus Leach, 1814 to Bostryx Troschel, 1847 sensu Breure 1979 (but see Breure 2012b).

Current systematic position.

Bulimulidae, Bostryx apodemetus (d’Orbigny, 1835) (comb. n.).

Bulimus aquilus Reeve, 1848

http://species-id.net/wiki/Bulimus_aquilus

Figs 70A–B, 70K, L6i
Bulimus aquilus Reeve 1848 [1848–1850]: pl. 22 fig. 138; Breure 1979: 85.
Bulimulus aquilus; Pilsbry 1897 [1897–1898]: 17, pl. 5 figs 72–73.
Scutalus (Vermiculatus) aquilus (Reeve); Breure 1978: 170 (lectotype designation).
Type locality.

”Peru, Tacna”.

Label.

”Peru”. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 27.1, D 14.6, W 5.8.

Type material.

NHMUK 1975376, lectotype (Cuming coll.).

Remarks.

The current systematic position at the species level follows Richardson (1995: 347).

Current systematic position.

Bulimulidae, Kuschelenia (Vermiculatus) aquilus (Reeve, 1848) (comb. n.).

Bulimus arcuatostriatus Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_arcuatostriatus

Figs 41A–C, L6iii
Bulimus arcuatostriatus Pfeiffer 1855d: 95; 1859: 394; Breure 1979: 106 (lectotype designation).
Drymaeus (Drymaeus) arcuatostriatus; Breure and Eskens 1981: 6.
Type locality.

”Peru”.

Label.

”Peru”; taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 30, lat. 13 mill.”; figured specimen herein H 27.6, D 15.6, W 6.5.

Type material.

NHMUK 1975455, lectotype (Cuming coll.).

Remarks.

The basal lip of the lectotype is damaged. The current systematic position folows Richardson (1995: 99).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) arcuatostriatus (Pfeiffer, 1855).

Bulimus atacamensis Pfeiffer, 1856

http://species-id.net/wiki/Bulimus_atacamensis

Figs 2A–B, L6ii
Bulimus atacamensis Pfeiffer 1856c: 207; Pfeiffer 1859b: 486; Breure 1979: 51.
Bostryx atacamensis; Breure 1978: 53 (lectotype designation).
Type locality.

”Paposo in deserto Atacamensi reipublicae Chilensis. (R.A. Philippi)”.

Label.

”Desert of Atacama, / Chile // Philippi has sent it”, ”Bolivia” [in a later handwriting]; taxon label in Pfeiffer’s handwriting.

Dimensions.

”Long. 19, lat. 5 1/2 mill.”; figured specimen herein H 19.0, D 5.9, W 10.2.

Type material.

NHMUK 1975312, lectotype; NHMUK 1975128, two paralectotypes (Cuming coll.).

Remarks.

The current systematic position follows Richardson (1995: 17).

Current systematic position.

Bulimulidae, Bostryx atacamensis (Pfeiffer, 1856).

Bulimulus atahualpa Dohrn, 1863

http://species-id.net/wiki/Bulimulus_atahualpa

Figs 56D–F, L6iv–v
Bulimulus atahualpa Dohrn 1863: 153.
Neopetraeus atahualpa; Breure 1979: 100.
Type locality.

Not given.

Label.

”Huallaga”, ”Province of / Patas, Andes of Peru” [see remarks]. M.C. label style IV.

Dimensions.

”Long 43, lat 21 (..) mill.” (largest specimen); figured specimen herein H 38.5, D 26.4, W 6+.

Type material.

NHMUK 1975436, three possible syntypes (Cuming coll.).

Remarks.

Dohrn (1863) gives the measurements of five specimens, but it is unclear how many specimens his description was actually based upon. The specimens in NHMUK are accompanied by two sets of labels. One is in Dohrn’s handwriting (teste S.P. Dance, 1958) and has ”Huallaga” as the locality. The other label set originates from Farris and has ”Province of Patas, Andes of Peru”; the taxon name is in Pfeiffer’s handwriting, who identified it as Bulimus tessellatus Shuttleworth, 1852. The latter label may have been added to the material in a later stage. Due to the fact the other label has original handwriting, the specimens are considered possible syntypes. The current systematic position follows Richardson (1995: 239).

Current systematic position.

Bulimulidae, Neopetraeus atahualpa (Dohrn, 1863).

Bulimus attenuatus Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_attenuatus

Figs 17F–H, L6vi
Bulimus attenuatus Pfeiffer 1853b: 256; Pfeiffer 1853d: 336; Pfeiffer 1853 in Küster and Pfeiffer 1840–1865: 83, pl. 30 figs 9–10; Breure 1979: 106 (lectotype designation).
Drymaeus (Mesembrinus) attenuatus attenuatus; Thompson 2011: 114.
Type locality.

[Mexico] ”Vera Cruz”.

Label.

”Vera Cruz”; taxon name in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 34, lat. 13 mill.”; figured specimen herein H 34.0, D 14.4, W 6.1.

Type material.

NHMUK 1975458, lectotype; NHMUK 1975459, two paralectotypes (Cuming coll.).

Remarks.

The current systematic position follows Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) attenuatus (Pfeiffer, 1853).

Bulimus aureolus Guppy, 1866

http://species-id.net/wiki/Bulimus_aureolus

Figs 21A, L7i
Bulimus aureolus Guppy 1866: 49.
Drymaeus (Mesembrinus) aureolus; Breure 1979: 117.
Type locality.

[Trinidad].

Label.

”Trinidad”; in Guppy’s handwriting.

Dimensions.

”Height 0.6 inch, breadth 0.3 inch” [H 15.2, D 7.6 mm]; figured specimen H 23.2, D 10.71, W 6.2.

Type material.

NHMUK 1866.1.3.8, two syntypes (ex Guppy).

Remarks.

Guppy described this taxon from immature specimens: ”The specimens from which the species was described by me in the Annals and Magazine of Natural History (1866) were young and imperfect.” (Guppy 1871: 307). There is another shell (NHMUK 1875.2.8.1), also presented by Guppy, which could be from the series he is referring to in the second paper; it is therefore not considered a syntype since this specimen was later collected than the type series, but unfortunately all three specimens have been mixed and there is no way of separating the two lots.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) aureolus (Guppy, 1866).

Bulimus aurifluus Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_aurifluus

Figs 25A, L6vii
Bulimus aurifluus Pfeiffer 1857b: 319, pl. 35 fig. 10; Pfeiffer 1859b: 400.
Type locality.

[Mexico] ”Cordova”.

Label.

”Cordova (Mexico)”; taxon name in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 22, diam. 10 mill.”; figured specimen herein H 21.8, D 11.1, W 5.1.

Type material.

NHMUK 1975502, lectotype; 1975502/2, one paralectotype (Cuming coll.).

Remarks.

One of the two specimens found corresponds to the figure of Pfeiffer and is here selected as the lectotype (design. n.). The current systematic position follows Thompson (2011: 112).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) aurifluus (Pfeiffer, 1857).

Bulimus auris Pfeiffer, 1866

http://species-id.net/wiki/Bulimus_auris

Figs 37A–C, L7ii
Bulimus auris Pfeiffer 1866: 831; Pfeiffer 1868: 36; Breure 1979: 106 (lectotype designation).
Drymaeus (Drymaeus) auris; Breure and Eskens 1981: 8.
Type locality.

”Venezuela”.

Label.

”Venezuela”; taxon name in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 39, diam. 14 1/2 mill.”; figured specimen herein H 38.8, D 20.1, W 6+.

Type material.

NHMUK 1975499, lectotype (Cuming coll.).

Remarks.

Pfeiffer did not state on how many specimens his description was based. Only one specimen was found, which corresponds to Pfeiffer’s original measurements; the top is damaged.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) auris (Pfeiffer, 1866).

Bulinus badius Sowerby, 1835

http://species-id.net/wiki/Bulinus_badius

Figs 67J, L7iii
Bulinus badius Sowerby I 1835: 141.
Bulimus badius; Reeve 1849 [1848–1850]: pl. 39 fig. 235.
Bulimulus badius; Pilsbry 1897 [1897–1898]: 28, pl. 4 fig. 49.
Type locality.

”in provincia Peruviae Xagua” (see remarks).

Label.

”Prov. of Xagua, Peru”. M.C. label style I.

Dimensions.

”long. 1, lat. 0.6 poll. [H 25.3, D 15.2 mm]”; figured specimen herein H 25.8, D 15.5, W 5.9.

Type material.

NHMUK 20100631, three possible syntypes (Cuming coll.).

Remarks.

The specimen figured by Reeve was found in the collection and, together with two other specimens, are considered possible syntypes. The type locality may refer to Dept. Jauja in Central Peru (teste Crawford 1939: 330) or to Dept. Amazonas, Prov. Bagua; the latter seems slightly outside the range known for this subgenus. The current systematic position at species level follows Richardson (1995: 348).

Current systematic position.

Bulimulidae, Kuschelenia (Vermiculatus) bicolor (Sowerby I, 1835) (comb. n.).

Bulimus balsanus Morelet, 1863

http://species-id.net/wiki/Bulimus_balsanus

Figs 5B, L7iv
Bulimus balsanus Morelet, 1863: 192, pl. 9 fig. 8; Breure 1979: 51.
Bostryx balsanus; Breure 1978: 53 (lectotype designation).
Type locality.

[Peru] ”Balsa de Cocharcas”.

Label.

”Balsa de Cocharcas”, in Morelet’s handwriting.

Dimensions.

”Long. 19, diam. 8 1/2 mill.”; figured specimen herein H 17.9, D 9.0, W 6.4.

Type material.

NHMUK 1893.2.4.173, lectotype; 1893.2.4.174, paralectotype (Morelet coll.).

Remarks.

The current systematic position follows Richardson (1995: 35).

Current systematic position.

Bulimulidae, Bostryx nigropileatus (Reeve, 1849).

Bulimus baranguillanus Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_baranguillanus

Figs 31A–C, L7v
Bulimus baranguillanus Pfeiffer 1853d: 334; Pfeiffer 1853 in Küster and Pfeiffer 1840–1865: 246, pl. 66 figs 5–6; Pfeiffer 1854b: 136; Breure 1979: 106 (lectotype designation).
Drymaeus baranguillanus; Pilsbry 1898 [1897–1898]: 208, pl. 35 figs 21–22; Linares and Vera 2012: 181.
Drymaeus (Drymaeus) baranguillanus; Breure and Eskens 1981: 8.
Type locality.

”Baranguilla in Andibus Colombianus”.

Label.

”Baranguilla / Andes of Colombia”; taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 32, diam. 13 1/2 mill.”; figured specimen herein H 31.5, D 15.6, W 7.2.

Type material.

NHMUK 1975452, lectotype (Cuming coll., ex Bland).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) baranguillanus (Pfeiffer, 1853).

Bulimus barbadensis Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_barbadensis

Figs 60A–B, L8i
Bulimus barbadensis Pfeiffer 1853d: 435; Breure 1979: 62 (lectotype designation).
Bulimulus fuscus (Guilding, 1828); Breure 1974: 38, pl. 5 figs 1–4.
Type locality.

[West Indies] ”Barbados. (Mus. Cuming)”.

Label.

No locality given. The taxon label is in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 20, diam. 10 mill.”; figured specimen herein H 21.3, D 10.8 W 5.9.

Type material.

NHMUK 1974054, lectotype; 1974055, two paralectotypes (Cuming coll.).

Remarks.

The current systematic position follows the revison of Breure (1974).

Current systematic position.

Bulimulidae, Bulimulus fuscus (Guilding, 1828).

Helix (Xenothauma) baroni Fulton, 1896

http://species-id.net/wiki/Helix_baroni

Figs 65A–B, L8iii
Helix (Xenothauma) baroni Fulton 1896: 101, pl. 6 figs 7–7b; Breure 1979: 83; Neubert and Janssen 2004: 201, pl. 11 fig. 135.
Bulimulus baroni; Pilsbry 1898 [1897–1898]: 172, pl. 25 figs 64–66.
Scutalus (Scutalus) baroni; Breure 1978: 164 (lectotype designation); Köhler 2007: 147, fig. 61.
Type locality.

[Peru, Dept. Cajamarca] ”Rio Yonan, Peru, 4000 feet (C.T. Baron)”.

Label.

”Rio Yonan, Peru, 4000’ / C.T. Baron”, in Fulton’s handwriting.

Dimensions.

”Height 12 millim., maj. diam. 30 millim.”; figured specimen herein H 12.7, D 29.3, W 4.6.

Type material.

NHMUK 1896.6.23.1, lectotype; 1896.6.23.2, one paralectotype, Baron leg. (ex Fulton).

Remarks.

Fulton did not mention on how many specimens his description was based. From Neubert and Janssen (2004: 201) it may be concluded that the type series was distributed to various museums. ”Rio Yonan” is the local name of Rio Jequetepeque at the village of Yonan, near Tembladera (V. Mogollón, pers. commun.).

Current systematic position.

Bulimulidae, Scutalus baroni (Fulton, 1896).

Bulimulus (Drymaeus) baroni Fulton, 1897

http://species-id.net/wiki/Bulimulus_baroni

Figs 65E–F, L8ii
Bulimulus (Drymaeus) baroni Fulton 1897: 213, pl. 6 fig. 8–8b; Breure 1979: 83.
Scutalus (Scutalus) baroni; Breure 1978: 164 (lectotype designation).
Type locality.

”Rio Yonan, Peru, 4000 feet (C.T. Baron)”.

Label.

”4000 ft / Rio Yonan / Peru”, in Fulton’s handwriting.

Dimensions.

”Long. 31–36 millim., maj. diam. 19–22 millim.”; figured specimen herein H 34.3, D 23.2, W 6.3.

Type material.

NHMUK 1897.8.3.173, lectotype; 1897.8.3.179, one paralectotype, Baron leg. (ex Fulton).

Remarks.

Fulton states ”Type specimens (three) in British Museum”. Only two specimens have been found. This taxon is considered a junior subjective synonym of Scutalus cretaceus (Pfeiffer, 1855) by Breure (1978).

Current systematic position.

Bulimulidae, Scutalus cretaceus (Pfeiffer, 1855).

Otostomus bartletti H. Adams, 1867

http://species-id.net/wiki/Otostomus_bartletti

Figs 41G–I, L8iv
Otostomus bartletti H. Adams 1867a: 442, pl. 38 fig. 4; Breure 1979: 106.
Drymaeus (Drymaeus) bartletti; Pilsbry 1898 [1897–1898]: 224, pl. 32 fig. 44.
Type locality.

”[Upper Amazons, and on the River Ucayali, Eastern Peru]”.

Label.

”E. Peru”; ”Upper Amazons” added in a later handwriting. Taxon label in H. Adam’s handwriting.

Dimensions.

”Long. 25 mill., diam. maj. 26 mill.”; figured specimen herein H 28.5, D 26.7, W 4+.

Type material.

NHMUK 1867.5.18.4, lectotype, Bartlett leg.

Remarks.

The type locality was not specified for each species, but is taken from the title of Adam’s paper. ”Two examples, one in good condition, of this very finely marked and interesting species were obtained by Mr. Bartlett” (H. Adams 1867a: 442). Only one specimen is present, of which the index card in the collection states ”The specimen, although not registered, appears to be 1867.5.18.4 which was purchased from Bartlett along with other types described at the same time. No specimen can be found with this number. The writings on the board are the same as those on the other specimens puchased at the same time, including the original labels. Locality exactly correct”. The specimen, of which the top is damaged, is the figured specimen by Adam and is here designated lectotype (design. n.).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) bartletti (H. Adams, 1867).

Drymaeus bellus da Costa, 1906

http://species-id.net/wiki/Drymaeus_bellus

Figs 28H–J, L8v
Drymaeus bellus da Costa 1906a: 8, pl. 1 fig. 5; Breure 1979: 106; Linares and Vera 2012: 181.
Type locality.

”Colombia, San Martin”.

Label.

”San Martin, Colombia”, in da Costa’s handwriting.

Dimensions.

”Long. 33, diam. 16 mm.”; figured specimen herein H 33.2, D 16.6, W 6.3.

Type material.

NHMUK 1907.11.21.8, holotype; 1907.11.21.9, paratype (da Costa coll.).

Remarks.

The current systematic position follows the revision of Breure and Borrero (unpublished data) and da Costa’s taxon is now placed in the synonymy of Drymaeus blandi Pilsbry, 1897 (syn. n.).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) blandi Pilsbry, 1897.

Bulinus bicolor Sowerby I, 1835

http://species-id.net/wiki/Bulinus_bicolor

Figs 66O–P, L9i
Bulinus bicolor Sowerby I 1835: 141; Reeve 1848 [1848–1850]: pl. 44 fig. 276; Breure 1979: 85.
Scutalus (Vermiculatus) bicolor; Breure 1978: 171 (lectotype designation).
Type locality.

”provinciae Peruviae Xagua” (see remarks).

Label.

”Peru” on two labels with different handwriting, ”Zagua” written on board. M.C. label style I, V.

Dimensions.

”Long. 0.9, lat. 0.4 poll. [H 22.8, D 10.1 mm]”; figured specimen herein H 23.6, D 10.9, W 5.8.

Type material.

NHMUK 1975151 lectotype; 1975152, two paralectotypes (Cuming coll.).

Remarks.

Sowerby described this taxon from the Cuming collection. So far the publication date for this taxon has been cited as 1834 or 1835. According to Duncan (1937: 78) Sowerby’s paper was published on 3 April 1835. The type locality has not be found in any modern gazetteer. Weyrauch (1967: 386) simply lists this species as from Peru, but distinguished Scutalus bicolor badius (Sowerby I, 1835) from ”N-Peru”. According to Crawford (1939: 330) the name ”Xagua” could refer to Dept. Jauja, which is in Central Peru. The specimen figured by Reeve has been selected lectotype by Breure (1978). The current systematic position at the species level follows Richardson (1995: 348).

Current systematic position.

Bulimulidae, Kuschelenia (Vermiculatus) bicolor (Sowerby I, 1835) (comb. n.).

Bulimus binneyanus Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_binneyanus

Figs 58C–E, L9ii
Bulimus binneayanus Pfeiffer 1857e: 229; Pfeiffer 1858a: 257, pl. 42 fig. 4; Breure 1979: 100.
Neopetraeus binneyanus; Pilsbry 1898 [1897–1898]: 164, pl. 32 fig. 43; Breure 1978: 210 (lectotype designation).
Type locality.

”in Andibus prov. Patas, Peru (Dr. Farris)”.

Label.

”Province of Patas / Andes of Peru”; taxon name in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 26, diam. 19 mill.”; figured specimen herein H 25.2, D 20.8, W 5.2.

Type material.

NHMUK 1975426, lectotype; 1975427, two paralectotypes (Cuming coll.).

Remarks.

Pfeiffer did not state on how many specimens his description was based; from Pfeiffer 1858 it is clear this he described this taxon from the Cuming collection. The specimen originally figured by Pfeiffer (1858a: pl. 42 fig. 4) was selected lectotype by Breure (1978).

Current systematic position.

Bulimulidae, Neopetraeus binneyanus (Pfeiffer, 1857).

Bulimus bogotensis Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_bogotensis

Figs 42A–C, L9iii
Bulimus bogotensis Pfeiffer 1855d: 93; Pfeiffer 1859b: 390; Breure 1979: 106 (lectotype designation).
Drymaeus bogotensis; Linares and Vera 2012: 182 [partim].
Drymaeus (Drymaeus) bogotensis; Breure and Eskens 1981: 10.
Type locality.

[Colombia] ”Santa Fé de Bogota”.

Label.

”Santa Fe da Bogata [sic]”; taxon name in Pfeiffer’s handwriting. M.C. label style III.

Dimensions.

”Long. 38, diam. 14 mill.”; figured specimen herein H 37.4, D 17.5, W 6.9.

Type material.

NHMUK 1975191, lectotype (Cuming coll.).

Remarks.

Pfeiffer described this taxon from Cuming’s collection; he did not state on how many specimens his description was based. Only one specimen was found. On the original label, Pfeiffer has added ”(spectatus Reeve t. 81. / f. 601 b.)”. Reeve described Bulimus spectatus from specimens in ”Mus. Taylor”; his figure does not correspond to the specimen designated as lectotype and we consider Reeve’s taxon to be distinct.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) bogotensis (Pfeiffer, 1855).

Helix bolivarii d’Orbigny, 1835

http://species-id.net/wiki/Helix_bolivarii

Figs 52A–B
Helix bolivarii d’Orbigny 1835: 17.
Bulimus bolivarii d’Orbigny 1837 [1834–1847]: 309, pl. 39 figs 5–6 [7 Aug. 1837; text 6 May 1838].
Drymaeus bolivarii (d’Orbigny); Breure 1975b: 1150, pl. 4 fig. 2 (lectotype designation).
Type locality.

”provincia Cochabambacensi (republica Boliviana); see Breure 1973: 114.

Label.

Not found.

Dimensions.

”Longit. 54 millim., latit. 26 millim.”.

Type material.

Not found.

Remarks.

No material of this species was found. In the NHMUK-copy of the ”Voyage...” a note on p. 309 states ”No examples of this sp. could be traced in d’O. colln in Sept. 1958. S.P. Dance”. Also, in Gray (1854) there is no ”B.M.” following the name, so it may safely assumed that type material of this taxon never came to London. The only extant material is thus the material in the Paris museum. As this is an incomplete specimen, the original figures by d’Orbigny are here copied.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) bolivarii (d’Orbigny, 1835).

Bulimus bolivianus Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_bolivianus

Figs 42D–F, L9iv
Bulimus bolivianus Pfeiffer 1846a: 34; Pfeiffer 1848b: 105; Reeve 1849 [1848–1850]: pl. 81 fig. 599.
Drymaeus bolivianus; Pilsbry 1898 [1897–1898]: 244, pl. 43 fig. 66.
Drymaeus (Drymaeus) bolivianus; Breure 1979: 107 (lectotype designation); Breure and Eskens 1981: 12.
Type locality.

[Venezuela] ”Merida, Andes of Bolivia”.

Label.

”Merida”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 33, diam. 13 mill.”; figured specimen herein H 33, D 13.7, W 7.2.

Type material.

NHMUK 1975444, lectotype; 1975445, one paralectotype (Cuming coll.); see remarks.

Remarks.

Pfeiffer did not state on how many specimens his description was based; the lot found contains three specimens. The smallest shell of the lot is a juvenile specimen not belonging to this species, and is here excluded from the type material. The locality as originally stated by Pfeiffer seems in error as there is no place with that name in Bolivia; the taxon is now tentatively assigned to the Venezuelan malacofauna. Reeve 1848 [1848–1850]: pl. 44 fig. 281 figured a specimen from ”Mus. Dennison”, which he mistakenly held for Bulimus bolivianus. Later he corrected his mistake on pl. 81, stating ”The shell here represented is the original type of the species described by Dr. Pfeiffer. To that which I have mistaken for it at Pl. XLIV Sp. 281, the name B. annulatus may be given”. The current systematic position follows Richardson (1995: 105).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) bolivianus (Pfeiffer, 1846).

Drymaeus boucardi da Costa, 1907

http://species-id.net/wiki/Drymaeus_boucardi

Figs 36A–C, L9v
Drymaeus boucardi da Costa 1907: 305, pl. 26 figs 5–5a; Breure 1979: 107.
Type locality.

[Panama] ”Chiriqui”.

Label.

”Chiriqui”, in da Costa’s handwriting.

Dimensions.

”Long. 27, diam. 11 mm.”; figured specimen herein H 26.9, D 12.4, W 5.9.

Type material.

NHMUK 1907.11.21.26, holotype, Boucard leg. (da Costa coll.).

Remarks.

In his remarks, da Costa states ”this shell’, which might imply that he had only one specimen. The material is considered herein a holotype. The basal part of the lip is damaged. This taxon has not been mentioned by Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) boucardi da Costa, 1907.

Bulimus bourcieri Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_bourcieri

Figs 26G–I, L9vi
Bulimus bourcieri Pfeiffer 1853d: 314; Pfeiffer 1854 in Küster and Pfeiffer 1840–1865: 98, pl. 32 figs 3–4.
Drymaeus bourcieri; Pilsbry 1898 [1897–1898]: 241, pl. 33 fig. 45.
Drymaeus (Drymaeus) bourcieri; Breure 1979: 107 (lectotype designation); Breure and Eskens 1981: 12; Breure and Borrero 2008: 20.
Type locality.

”Pichincha reipublicae Aequatoris (Bourcier)”.

Label.

”Pichincha / Ecuador / Monsr Bourcier”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 25, diam. 11 mill.”; figured specimen herein H 23.9, D 13.0, W 5.5.

Type material.

NHMUK 1975446, lectotype; 1975447, two paralectotypes, Bourcier leg. (Cuming coll.).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) bourcieri (Pfeiffer, 1853).

Helix brachysoma d’Orbigny, 1835

http://species-id.net/wiki/Helix_brachysoma

Figs 52C–D
Helix brachysoma d’Orbigny 1835: 18.
Bulimus brachysoma d’Orbigny 1837 [1834–1847]: 309, pl. 39 figs 9–10 [7 Aug. 1837; text 6 May 1838].
Type locality.

”provincia Santa Cruz de la Sierra (republica Boliviana)”; see Breure 1973: 115.

Dimensions.

”Longit. 40 millim., latit. 19 millim.”.

Remarks.

In the NHMUK copy of the ”Voyage...”, there is a note in pencil in the margin of p. 309, which states ”No examples of this sp. could be traced in d’O. colln. in Sept. 1958. S.P. Dance”. Also, in Gray (1854) there is no ”B.M.” following the name, so it may safely assumed that type material of this taxon never came to London. As this species has not been traced in the Paris museum (Breure 1975b), the type material must be considered lost. The original figures of d’Orbigny are here reproduced.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) brachysoma (d’Orbigny 1835).

Bulimulus (Drymaeus) broadwayi E.A. Smith, 1896

http://species-id.net/wiki/Bulimulus_broadwayi

Figs 20I, L9vii
Bulimulus (Drymaeus) broadwayi E.A. Smith 1896: 243, pl. 8 fig. 9.
Drymaeus broadwayi; Pilsbry 1899: 22, pl. 12 fig. 5.
Drymaeus (Mesembrinus) broadwayi; Breure 1979: 117.
Type locality.

[Trinidad].

Label.

”Trinidad”.

Dimensions.

”Longit. 14 millim., diam. 8 1/2”; holotype H 11.9, D 7.59, W 5.4.

Type material.

NHMUK 1895.11.28.10, holotype, W.E. Broadway leg. (ex W. Moss).

Remarks.

Smith remarked ”Mr. Moss, who received from him [Broadway] a fine series (...), has liberally placed one of his three specimens of this species in the British Museum collection”. The specimen found is thus the holotype, despite the fact it is somewhat smaller than the dimensions given by Smith.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) broadwayi (E.A. Smith, 1896).

Bulimulus (Drymaeus) buckleyi Sowerby III, 1895

http://species-id.net/wiki/Bulimulus_buckleyi

Figs 36D–F, L10i
Bulimulus (Drymaeus) buckleyi Sowerby III 1895: 214, pl. 13 figs 3–4.
Drymaeus (Drymaeus) buckleyi; Breure 1979: 107 (lectotype designation); Breure and Eskens 1981: 12; Breure and Borrero 2008: 20.
Type locality.

”Ecuador”.

Label.

”Ecuador”. The label also mentions ”2 co-types”.

Dimensions.

”Long. 27, diam. 9 mm.”; figured specimen herein H 27.3, D 11.1, W 5.7.

Type material.

NHMUK1907.11.21.48, lectotype; 1907.11.21.49–50, two paralectotypes (da Costa coll.).

Remarks.

Sowerby (1895) stated: ”Types in the collection of Mr. S.I. Da Costa. The shells were collected by Mr. C. Buckley in 1872, but not mentioned in Mr. E.T. Higgins’ paper [Higgins 1872]”.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) buckleyi (Sowerby III, 1895).

Otostomus bugabensis Martens, 1893

http://species-id.net/wiki/Otostomus_bugabensis

Figs 25B, L10ii
Otostomus bugabensis Martens 1893 [1890–1901]: 218, pl. 13 figs 21–21a.
Drymaeus bugabensis; Pilsbry 1899: 64, pl. 13 figs 34–35.
Drymaeus (Mesembrinus) bugabensis; Breure 1979: 117 (lectotype designation); Breure and Eskens 1981: 54, pl. 6 fig. 4; Thompson 2011: 114.
Type locality.

”S. Panama, Bugaba, Department of Chiriqui, at an elevation of 1000 feet (Champion)”.

Label.

”Bugaba Panama” on two labels in Martens’ handwriting, one of which states ”Champion 166”.

Dimensions.

”Long. 27, diam. 11 millim.”; lectotype H 26.3, D 11.0, W 5.8 (see remarks).

Type material.

NHMUK 1901.6.22.958, lectotype; 1901.6.22.820, paralectotype, Champion leg. (Godman coll.).

Remarks.

The lectotype has a damaged palatal lip and could be only subadult. Both shells are discoloured by age. The current systematic position is according to Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) bugabensis (Martens, 1893).

Bulimus cacticolus Reeve, 1849

http://species-id.net/wiki/Bulimus_cacticolus

Figs 60E–F, L10iii
Bulimus cacticolus Reeve 1849 [1848–1850]: pl. 58 fig. 393.
Type locality.

”Curiana, Venezuela. (Mus. Dyson)”.

Label.

”Curiana, Venezuela”, in a later handwriting on board. M.C. label style I.

Dimensions.

Not given; figured specimen herein H 24.4, D 13.7, W 6.2.

Type material.

NHMUK 20100515, one syntype (Cuming coll.).

Remarks.

The specimen corresponds to Reeve’s figure, except that the spire is less pointed; therefore, considered a possible syntype. No other material is present in the collection that can be matched to Reeve’s figure or can be attributed to Dyson.

Current systematic position.

Bulimulidae, Bulimulus cacticolus (Reeve, 1849).

Helix cactorum d’Orbigny, 1835

http://species-id.net/wiki/Helix_cactorum

Figs 6D, L10iv
Helix cactorum d’Orbigny 1835: 10.
Bulimus hennahi Gray; d’Orbigny 1837 [1834–1847]: 283, pl. 30 figs 3–4 [3 April 1837; text 6 May 1838].
Type locality.

”provincia Tacnacensi (rep. Peruviana)”.

Label.

”Tacna, Pérou”, in d’Orbigny’s handwriting.

Dimensions.

”Longit. 25 millim., latit. 15 millim.”; figured specimen herein H 24.4, D 12.8, W 7.0.

Type material.

NHMUK 1854.12.4.189, lectotype and six paralectotypes (d’Orbigny coll.).

Remarks.

Seven specimens are present in the d’Orbigny collection, of which one corresponds to d’Orbigny 1837 [1834–1847]: pl. 30 fig. 4; it is here designated lectotype (design. n.). This taxon has been synonymized with Bostryx hennahi (J.E. Gray, 1830) by Richardson (1995: 27); the latter species was described from ”Arica”, which is presumably meant to be in (nowadays) Chile, while Tacna is in southern Peru; both localities are in the coastal area.

Current systematic position.

Bulimulidae, Bostryx hennahi (J.E. Gray, 1830).

Bulimus californicus Reeve, 1848

http://species-id.net/wiki/Bulimus_californicus

Figs 23C, L10v
Bulimus californicus Reeve 1848 [1848–1850]: pl. 56 fig. 378.
Type locality.

”California”.

Label.

”California”. M.C. label style III.

Dimensions.

Not given; figured specimen herein H 25.9, D 11.7, W. 6.3.

Type material.

NHMUK 1975136, three syntypes (Cuming coll.).

Remarks.

This taxon is considered as a junior subjective synonym of Drymaeus (Mesembrinus) ziegleri (Pfeiffer, 1847) by Richardson (1995: 198).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) ziegleri (Pfeiffer, 1847).

Bulimus caliginosus Reeve, 1849

http://species-id.net/wiki/Bulimus_caliginosus

Figs 69C–D, 69K, L10vi
Bulimus caliginosus Reeve 1849 [1848–1850]: pl. 82 fig. 609.
Type locality.

”—?”.

Label.

”?”. M.C. label style I.

Dimensions.

Not given; figured specimen herein H 35.4, D 17.9, W 6.1.

Type material.

NHMUK 20100518.1–3, lectotype and two paralectotypes (Cuming coll.).

Remarks.

This taxon was classified as a Naesiotus species by Breure (1979). The sculpture of the protoconch is, however, not straightly axially ribbed but with a waving sculpture that is typical for species classified with Vermiculatus Breure, 1978. The specimen figured by Reeve is here designated lectotype (design. n.) to define the taxon.

Current systematic position.

Bulimulidae, Kuschelenia (Vermiculatus) caliginosus (Reeve, 1849) (comb. n.).

Bulimus canaliculatus Pfeiffer, 1845

http://species-id.net/wiki/Bulimus_canaliculatus

Figs 38G–I, L11i
Bulimus canaliculatus Pfeiffer 1845a: 68; Reeve 1848 [1848–1850]: pl. 41 fig. 256.
Drymaeus (Drymaeus) canaliculatus; Breure 1979: 107 (lectotype designation); Breure and Eskens 1981: 12, pl. 5 fig. 6.
Type locality.

”Bolivia”.

Label.

”Bolivia”; taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 37, diam. 14 mill.”; figured specimen herein H 36.0, D 16.2, W 6.6.

Type material.

NHMUK 1975514, lectotype; 1975515, one paralectotype (Cuming coll.).

Remarks.

The lectotype has also been figured by Reeve (1848 [1848–1850]). The current systematic position follows Richardson (1995: 107).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) canaliculatus (Pfeiffer, 1845).

Drymaeus prestoni cancellata da Costa, 1906

http://species-id.net/wiki/Drymaeus_prestoni_cancellata

Figs 24K, L11ii
Drymaeus prestoni cancellata da Costa 1906a: 9, pl. 1 fig. 10.
Drymaeus (Mesembrinus) prestoni cancellata; Breure 1979: 117.
Drymaeus (Mesembrinus) tripictus hoffmanni (Martens, 1893); Thompson 2011: 117.
Type locality.

”Chiriqui, Panama”.

Label.

”Chiriqui”, in da Costa’s handwriting.

Dimensions.

Not given; figured specimen herein H 18.1, D 9.8, W 5.3.

Type material.

NHMUK 1907.11.21.13, holotype (da Costa coll.).

Remarks.

As da Costa speaks about ”this pretty shell”, it is implied that he had only one specimen at hand; thus the specimen found should be regarded holotype. The current systematic position follows the synonymisation by Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) tripictus hoffmanni (Martens, 1893).

Drymaeus castaneostrigatus da Costa, 1906

http://species-id.net/wiki/Drymaeus_castaneostrigatus

Figs 36G–I, L11iii
Drymaeus castaneostrigatus da Costa 1906b: 98, pl. 11 fig. 5; Breure 1979: 107.
Type locality.

”Pozuzo, Eastern Peru”.

Label.

”Pozuzo. E. Peru”, in da Costa’s handwriting.

Dimensions.

”Long. 19, diam. 8.5 mm”; figured specimen herein H 20.4, D 8.6, W 6.3.

Type material.

NHMUK 1907.11.21.19; holotype (da Costa coll.).

Remarks.

da Costa referred to ”This shell (..)”, which might imply that he had only seen one specimen. The NHMUK material is herein regarded the holotype.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) castaneostrigatus da Costa, 1906.

Bulimus castus Pfeiffer, 1847

http://species-id.net/wiki/Bulimus_castus

Figs 19D, L11iv
Bulimus castus Pfeiffer 1847: 112; Reeve 1848 [1848–1850]: pl. 40 figs 1–2.
Drymaeus (Mesembrinus) castus; Breure 1979: 117 (lectotype designation).
Type locality.

”Central America ?”.

Label.

”Central America?”. M.C. label style III.

Dimensions.

”Long. 19, diam. 9 mill.”; figured specimen herein H 19.0, D 9.4, W 5.1.

Type material.

NHMUK 1975197, lectotype; 1975198, two paralectotypes (Cuming coll.).

Remarks.

The specimen figured by Reeve (1848 [1848–1850]) was designated lectotype by Breure (1979). The current systematic position follows Thompson (2011: 109).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) castus castus (Pfeiffer, 1847).

Bulimus catlowiae Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_catlowiae

Figs 15C, L11v
Bulimus catlowiae Pfeiffer 1853d: 427; Breure and Coppois 1978: 182 (lectotype designation); Breure 1979: 68.
Type locality.

[Ecuador] ”prope Quito (Bourcier)”.

Label.

”near Quito / Monsr Bourcier / Consul General”; taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 25, diam. 11 mill.”; figured specimen herein H 25.1, D 12.8, W 6.7.

Type material.

NHMUK 1975414, lectotype; 1975415, two paralectotypes (Cuming coll.).

Remarks.

This taxon is considered a junior subjective synonym of Naesiotus quitensis (Pfeiffer, 1848) (Richardson 1995: 229).

Current systematic position.

Bulimulidae, Naesiotus quitensis (Pfeiffer, 1848).

Bulimulus (Drymaeus) caucaensis da Costa, 1898

http://species-id.net/wiki/Bulimulus_caucaensis

Figs 31G–I, L11vi
Bulimulus (Drymaeus) caucaensis da Costa 1898: 81, pl. 6 fig. 3.
Drymaeus (Drymaeus) caucaensis; Pilsbry 1898 [1897–1898]: 247, pl. 50 fig. 83; Linares and Vera 2012: 183.
Drymaeus (Drymaeus) caucaensis; Breure 1979: 107.
Type locality.

”Valley of the R. Cauca, Colombia”.

Label.

”Cauca, Colombia”, in da Costa’s handwriting.

Dimensions.

”Long. 35, diam. 16 mm.”; figured specimen herein H 34.8, D 16.6, W 6.3.

Type material.

NHMUK 1907.11.21.43, lectotype (da Costa coll.).

Remarks.

The specimen is labelled ”type”; however, since da Costa did not state on how many specimens his description was based upon, the reference of Breure (1979) to ”HT 1907.11.21.43” has to be interpreted as lectotype designation according to Art. 74.6 ICZN.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) caucaensis (da Costa, 1898).

Bulimus ceratacme Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_ceratacme

Figs 3F–G, L11vii
Bulimus ceratacme Pfeiffer 1855c: 8; Pfeiffer 1859b: 424.
Bostryx ceratacme; Breure 1978: 57 (lectotype designation); Breure 1979: 52.
Type locality.

”Peru?”.

Label.

”Peru”; taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 18 1/2, diam. 6 mill.”; figured specimen herein H 18.0, D 6.11, W 7.5.

Type material.

NHMUK 1975374, lectotype (Cuming coll.).

Remarks.

The current systematic position follows Richardson (1995: 19).

Current systematic position.

Bulimulidae, Bostryx ceratacme (Pfeiffer, 1855).

Bulimus cercicola Morelet, 1863

http://species-id.net/wiki/Bulimus_cercicola

Figs 4G, L12i
Bulimus cercicola Morelet 1863: 192, pl. 9 fig. 7.
Bulimulus (Lissoacme) cercicola; Pilsbry 1896 [1895–1896]: 184, pl. 46 fig. 63.
Bostryx cereicola [sic]; Breure 1978: 56; Breure 1979: 52.
Type locality.

[Peru] ”vallées chaudes d’Abancay et d’Acostambo, situées à l’ouest du Cuzco”.

Label.

”Abancay, Pérou”, in Morelet’s handwriting.

Dimensions.

”Long. 20; diam. 9 mill.”; figured specimen herein H 18.7, D 8.75, W 5+.

Type material.

NHMUK 1893.2.4.175–176, two syntypes (Morelet coll.).

Remarks.

One of the syntypes is severely damaged; the other has the top damaged. The current systematic position as given by Richardson (1995: 36).

Current systematic position.

Bulimulidae, Bostryx orophilus (Morelet, 1860).

Bulimus chamaeleon Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_chamaeleon

Figs 40A–C, L12ii
Bulimus chamaeleon Pfeiffer 1855f: 116; Breure 1979: 107.
Type locality.

[Ecuador] ”Quito (Mr. Bourcier)”.

Label.

”near Quito / Ecuador / Monsr Bourcier / Consul General”; taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long 27, diam. 11 1/2 mill.”; figured specimen herein H 27.6, D 12.2, W 6.2.

Type material.

NHMUK 1975448, three syntypes (Cuming coll.).

Remarks.

Richardson (1995: 98) listed this taxon as a nomen nudum in Pfeiffer 1853d: 422, and Pfeiffer 1854 in Küster and Pfeiffer 1840–1865: 105, pl. 33 figs 17–18. However, the name Bulimus chamaeleon seems to have been published for the first time in Pfeiffer 1855f: 116. The current systematic position follows Richardson (1995).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) ambustus (Reeve, 1849).

Naesiotus (Naesiotus) subcostatus chamayensis Weyrauch, 1967

http://species-id.net/wiki/Naesiotus_subcostatus_chamayensis

Fig. 15E
Naesiotus (Naesiotus) subcostatus chamayensis Weyrauch 1967a: 409, figs 97–99; Breure 1979: 68; Neubert and Janssen 2004: 203, pl. 10 fig. 121; Breure 2012a: 6.
Type locality.

”Norte interandino de Perú, cerca de pueblo Chamaya, sobre la margen izquierda del río Chamaya, afluente occidental del río Maranon, 550–660 m”.

Label.

”Above Chamaya, Río Chamaya, western affluent of Río Marañon, 600 m”; printed label.

Dimensions.

”A. 18.2, D. 7.3 [mm]”; figured specimen H 17.5 D 7.7 W 7.5.

Type material.

NHMUK 1975423, three paratypes (ex Weyrauch).

Current systematic position.

Bulimulidae, Naesiotus subcostatus chamayensis Weyrauch, 1967.

Otostomus championi Martens, 1893

http://species-id.net/wiki/Otostomus_championi

Figs 19E–F, L12iii
Otostomus championi Martens 1893 [1890–1901]: 222, pl. 14 fig. 5.
Drymaeus (Mesembrinus) championi (von Martens); Breure 1979: 107 (lectotype designation); Breure and Eskens 1981: 14; Thompson 2011: 117.
Type locality.

”W. Guatemala, Hacienda de Los Nubes, Cerro Zunil, Pacific slope, in the vicinity of the coffee-plantations, at an elevation of about 5000 feet (Champion)”.

Label.

”Cerro Zunil / Champion”.

Dimensions.

”Long. 27, diam. 14 1/2 mill.”; figured specimen herein H 27.0, D 13.8, W 6.2.

Type material.

NHMUK 1901.6.22.781, lectotype (Godman coll.).

Remarks.

The specimen is not accompanied by a taxon label in von Marten’s handwriting, but bears the name of the collector and corresponds to the original figure. The current systematic position follows Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) championi (Martens, 1893).

Bulimus chemnitzioides Forbes, 1850

http://species-id.net/wiki/Bulimus_chemnitzioides

Figs 12I, L12iv
Bulimus chemnitzioides Forbes 1850: 55, pl. 9 figs 6a–b; Pfeiffer 1854 in Küster and Pfeiffer 1840–1865: 94, pl. 31 figs 21–23.
Bulimulus chemnitzioides; Pilsbry 1897 [1897–1898]: 124, pl. 24 figs 45–46.
Type locality.

[Ecuador] ”Chatham Island, Gelepagos [sic, Galápagos]”.

Label.

”Chatham Island / Galapagos / Proc. Zool. Soc. / fig. 6”.

Dimensions.

”Long. 19, diam. 4 mil..”; figured specimen herein H 18.5, D 3.64, W 16.0.

Type material.

NHMUK 1855.4.5.23, lectotype; 1850.4.23.14–15, two paralectotypes (ex Kellett and Wood).

Remarks.

The specimen originally figured is here designated lectotype (design. n.). The two other specimens also originate from the voyage of Kellett and Wood (Forbes 1850, Seemann 1853), and are considered to be part of the original series.

Current systematic position.

Bulimulidae, Naesiotus chemnitzioides (Forbes, 1850).

Scutalus (Scutalus) chiletensis Weyrauch, 1967

http://species-id.net/wiki/Scutalus_chiletensis

Fig. 65G
Scutalus (Scutalus) chiletensis Weyrauch 1967a: 373, figs 24–30; Breure 1979: 83; Neubert and Janssen 2004: 204, pl. 12 fig. 138; Breure 2012a: 6, pl. 5 figs 46–47.
Type locality.

”Norte de Péru, vertiente occidental de la Cordillera Occidental, cerro encima del pueblo Chilete, sobre la margen izquierda del río Jequetepeque, 850–950 m”.

Label.

”N-Peru, mountain above Chilete, Rio Jequetepeque, 850-950m”; printed label.

Dimensions.

”A. 33.1, D. 21 [mm]”; figured specimen herein H 34.2 D 23.4 W 6.0.

Type material.

NHMUK 1975384, five paratypes (ex Weyrauch).

Current systematic position.

Bulimulidae, Scutalus chiletensis Weyrauch, 1967.

Bulimus chimborasensis Reeve, 1848

http://species-id.net/wiki/Bulimus_chimborasensis

Figs 38A–C, L12v
Bulimus chimborasensis Reeve 1848 [1848–1850]: pl. 44 fig. 275.
Drymaeus chimborasensis; Pilsbry 1898 [1897–1898]: 261, pl. 40 fig. 1; Linares and Vera 2012: 183.
Drymaeus (Drymaeus) chimborasensis; Breure 1979: 107; Breure and Borrero 2008: 20.
Type locality.

”Chimborazo, Columbia, New Granada”.

Label.

”Chimborazo”. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 39.7, D 15.0, W 7.0.

Type material.

NHMUK 1975460, three syntypes (Cuming coll.).

Remarks.

Reeve described this species from ”Mus. Dennison”, and said it was identical to Bulimus decoratus Lea, 1838 (not Férussac, 1821); he referred to Lea’s figure (pl. 23 fig. 8). The specimens found correspond to Reeve’s description, but none can be matched to the shell figured on his plate 44.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) chimborasensis (Reeve, 1848).

Drymaeus chiriquensis da Costa, 1901

http://species-id.net/wiki/Drymaeus_chiriquensis

Figs 42J–L, L12vi
Drymaeus chiriquiensis da Costa 1901: 238, pl. 24 fig. 1; Pilsbry 1901 [1901–1902]: 162, pl. 48 fig. 47.
Drymaeus (Drymaeus) chiriquiensis; Breure 1979: 107.
Drymaeus (Mesembrinus) chiriquiensis Thompson 2011: 114.
Type locality.

”Boqueti, Chiriquí, Panama”.

Label.

”Boqueti / Chiriqui / Panama”, in da Costa’s handwriting.

Dimensions.

”Long. 29, diam. 14 mm.”; figured specimen herein H 29.1, D 13.6, W 5.5.

Type material.

NHMUK 1907.11.21.119, holotype (da Costa coll.).

Remarks.

As da Costa (1901) writes ”A single specimen has been received of this pretty shell”, it is clear that the specimen has to be regarded as the holotype. The current systematic position follows Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) chiriquensis da Costa, 1901.

Bulimus cinereus Reeve, 1849

http://species-id.net/wiki/Bulimus_cinereus

Figs 15F–G, L12vii
Bulimus cinereus Reeve 1849 [1848–1850]: pl. 56 fig. 372.
Type locality.

”Bolivia”.

Label.

”Bolivia”. M.C. label style I, V.

Dimensions.

Not given; figured specimen herein H 15.5, D 6.8, W 7.5.

Type material.

NHMUK 20100519, lectotype (Cuming coll.).

Remarks.

The specimen figured by Reeve 1849 [1848–1850] was found and is now designated lectotype (design. n.). This hitherto unfigured taxon was regarded a nomen inquirendum by Breure (1979: 136) and Richardson (1995: 387). The protoconch is sculptured with axial riblets, typical of Naesiotus sensu Breure 1979.

Current systematic position.

Bulimulidae, Naesiotus cinereus (Reeve, 1849) (comb. n.).

Bulimus citronellus Angas, 1879

http://species-id.net/wiki/Bulimus_citronellus

Figs 20E, L13i
Bulimus citronellus Angas 1879: 479, pl. 40 fig. 5; Thompson 2011: 119.
Drymaeus (Mesembrinus) citronellus; Breure 1979: 117 (lectotype designation).
Type locality.

”Uren to Lipurio, Costa Rica (Gabb)”.

Label.

”Uren to Lipurio / low hills / Gabb / Costa rica”, in Angas’s handwriting.

Dimensions.

”Diam. 6, alt. 12 lin. [H 25.0, D 12.5 mm]”; figured specimen herein H 25.0, D 11.9, W 6.9.

Type material. NHMUK 1879.7.22.19–22, lectotype and three paralectotypes (ex Angas).

Remarks.

Angas (1879) writes that he has seen ”only two specimens”. The lot is marked ”type” and has been registered as having four specimens (1879.7.22.19-22). The current systematic position follows Richardson (1995: 183).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) sulphureus (Pfeiffer, 1857).

Bulimus clarus Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_clarus

Figs 73I–J, L13ii
Bulimus clarus Pfeiffer 1857c: 330.
Type locality.

”Meobamba, Peru (Mr. Gueinzius)”.

Label.

”Ecuador” [see remarks]; taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 16, diam. 7 2/3 mill.”. See remarks.

Type material.

NHMUK 1975487, two possible syntypes [see remarks] (Cuming coll.).

Remarks.

The type material consists of a mixed lot, one specimen belonging to Naesiotus sensu Breure 1979 (H 15.9, D 8.2, W 7.0), the other to Drymaeus (H 15.8, D 7.6, W 5.4). The latter corresponds more closely to the original measurements, but is subadult. However, as the locality differs from the type locality, and the collector is not mentioned on the label, there remains doubt whether this is the original lot. The taxon is now considered a nomen inquirendum.

Current systematic position.

Bulimulidae.

Bulimus clathratus Pfeiffer, 1858

http://species-id.net/wiki/Bulimus_clathratus

Figs 43D–F, L13iii
Bulimus clathratus Pfeiffer 1858a: 258; Pfeiffer 1859a: 47.
Drymaeus (Drymaeus) clathratus; Breure 1979: 107 (lectotype designation); Breure and Eskens 1981: 14.
Type locality.

”Province of Patas, Andes of Peru (Dr. Farris)”.

Label.

”Province of Patas, / Andes of Peru / Dr Farris”; taxon label in Pfeiffer’s handwriting.

Dimensions.

”Long. 30, diam. 11 mill.”; figured specimen herein H 30, D 12.2, W 7.8.

Type material.

NHMUK 1975449, lectotype (Cuming coll.).

Remarks.

This hitherto unfigured taxon is known from a single specimen. The current systematic position follows Richardson (1995: 110).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) clathratus (Pfeiffer, 1858).

Bulimus coagulatus Reeve, 1849

http://species-id.net/wiki/Bulimus_coagulatus

Figs 66E, L13iv
Bulimus coagulatus Reeve 1849 [1848–1850]: pl. 77 fig. 558.
Scutalus (Vermiculatus) coagulatus (Reeve); Breure 1978: 173 (lectotype designation); Breure 1979: 85.
Type locality.

”Peru”.

Label.

”Peru”. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 15.6, D 9.3, W 5.1.

Type material.

NHMUK 1975351, lectotype; 1975352 two paralectotypes (Cuming coll.).

Remarks.

This is a mixted lot of four specimens. The lectotype is an inmature specimen and is considered by Ramírez and Thomé (2005) to be synonymous with Scutalus versicolor (Broderip in Broderip and Sowerby 1832). They identified one of the other specimens as Bostryx conspersus (NHMUK 1975352); this specimen is no longer considered as type material.

Current systematic position.

Bulimulidae, Scutalus versicolor (Broderip in Broderip and Sowerby 1832).

Bulimus coarctatus Pfeiffer, 1845

http://species-id.net/wiki/Bulimus_coarctatus

Figs 37F–H, L13v
Bulimus coarctatus Pfeiffer 1845b: 73; Pfeiffer 1848b: 90; Pfeiffer 1856 [1854–1860]: 80, pl. 22 figs 22–23; Breure 1979: 108 (lectotype designation).
Drymaeus coarctatus; Pilsbry 1898 [1897–1898]: 195, pl. 28 figs 17–18; Simone 2006: 136, fig. 445.
Drymaeus (Drymaeus) coarctatus; Breure and Eskens 1981: 16, pl. 5 fig. 5.
Type locality.

”Locality unknown”.

Label.

”Brazil” (added in later handwriting); taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 34, diam. 17 mill.”; figured specimen herein 34.6, D 20.2, W 6.9.

Type material.

NHMUK 1975560, lectotype; 1975561, one paralectotype (Cuming coll.).

Remarks.

The specimen designated lectotype has the best match with the original measurements. The other specimen has been figured in Pfeiffer 1856 [1854–1860]: pl. 22 figs 22–23.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) coarctatus (Pfeiffer, 1845).

Bulimus columbiensis Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_columbiensis

Figs 19I–J, L25v
Bulimus columbiensis Pfeiffer 1855g: 124; Breure 1979: 118 (lectotype designation).
Bulimus gratus Pfeiffer 1856a: 159. New name for Bulimus columbiensis Pfeiffer non Lea.
Drymaeus (Mesembrinus) gratus; Breure and Eskens 1981: 71, pl. 7 fig. 9.
Drymaeus columbiensis; Linares and Vera 2012: 193.
Type locality.

”Columbia”.

Label.

”Columbia”; ”Venezuela” in a later handwriting. Labels for both taxon names in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 28, diam. 11 1/2 mill.”; figured specimen herein H 28.0, D 11.9, W 7.2.

Type material.

NHMUK 1975521, lectotype (Cuming coll.).

Remarks.

The original taxon label has been corrected to Bulimus gratus, in a different ink. However, also a label with this name in Pfeiffer’s handwriting has been glued to the board. It may be noted that Lea’s name (Bulimus columbianus) is sufficiently different to retain Pfeiffer’s original taxon name. On the front side, a later hand has written ”Venezuela”, which may be seen as a further precision to reflect the later political-administrative situation. Breure and Borrero (unpublished data) could not recognise this species in the Colombian material they studied.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) columbiensis (Pfeiffer, 1855).

Bulimulus compactus Fulton, 1902

http://species-id.net/wiki/Bulimulus_compactus

Figs 7H, L13vi
Bulimulus compactus Fulton 1902: 69.
Bostryx compactus; Breure 1979: 52.
Type locality.

”Chicani, Bolivia”.

Label.

”Chicani Bolivia”, in Fulton’s handwriting.

Dimensions.

”Alt. 16, maj. diam. 9 millim.”; figured specimen herein H 15.8, D 7.62, W 6.0.

Type material.

NHMUK 1902.5.28.1, syntype (ex Fulton).

Remarks.

Fulton did not state on how many specimens his description was based; the specimen is therefore regarded as a syntype. The current systematic position follows Richardson (1995: 20).

Current systematic position.

Bulimulidae, Bostryx compactus (Fulton, 1902).

Bulimus confluens Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_confluens

Figs 31J–L, L14i
Bulimus confluens Pfeiffer 1855f: 115; Pfeiffer 1859b: 443; Breure 1979: 108 (lectotype designation).
Drymaeus confluens; Linares and Vera 2012: 183.
Drymaeus (Drymaeus) confluens; Breure and Eskens 1981: 16, pl. 8 fig. 1.
Type locality.

[Colombia] ”Marmato, New Granada”.

Label.

”Venezuela” [see remarks]; taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 40, diam. 17 mill.”; figured specimen herein H 39.5, D 20.8, W 6.3.

Type material.

NHMUK 1975196, lectotype and one paralectotype (Cuming coll.).

Remarks.

According to Pfeiffer (1859: 443) the type is in ”Mus. Cuming”. The original specimen appears to be lost. The specimens are labelled ”Bulimus con-/fluens Pfr. var. / Venezuela”. There is no variety mentioned by Pfeiffer, nor in his original paper nor in subsequent publications, which is interpreted as him considering this a variable taxon; the specimens are thus regarded to be from the type series.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) confluens (Pfeiffer, 1855).

Bulimus confusus Reeve, 1848

http://species-id.net/wiki/Bulimus_confusus

Figs 66I–K, L14ii
Bulimus confusus Reeve 1848 [1848–1850]: pl. 48 fig. 316; Pfeiffer 1853d: 426; Breure 1979: 85.
Bulimulus confusus; Pilsbry 1898 [1897–1898]: 282, pl. 45 fig. 32.
Scutalus (Vermiculatus) confusus; Breure 1978: 173 (lectotype designation).
Type locality.

Not given.

Label.

”Peru”, corrected to ”?Peru” on a label in E.A. Smith’s handwriting. M.C. label style III, V.

Dimensions.

Not given; figured specimen herein H 29.0, D 13.1, W 6.0.

Type material.

NHMUK 1975194, lectotype; 1975195, two paralectotypes (Cuming coll.).

Remarks.

The protoconch sculpture is typically for Kuschelenia Hylton Scott, 1952. The placement of this taxon within Drymaeus by Richardson (1995) is thus erroneous. Breure (1978) mentioned ”New Grenada” as locality on the label; however, no mentioning of Peru which may be the original locality, if Crawford’s interpretation (Crawford 1939: 328) was correct to synonymize this taxon with Kuschelenia culminea (d’Orbigny, 1835).

Current systematic position.

Bulimulidae, Kuschelenia (Kuschelenia) culminea (d’Orbigny, 1835) (comb. n.).

Drymaeus conicus da Costa, 1907

http://species-id.net/wiki/Drymaeus_conicus

Figs 25E, L14iii
Drymaeus conicus da Costa 1907: 305, pl. 26 figs 7–7a; Breure 1979: 108 (lectotype designation).
Type locality.

”Oaxarca [sic], Mexico”.

Label.

”Oaxarca, Mexico”, in da Costa’s handwriting.

Dimensions.

”Long. 17.5, diam. 7 mm.”; lectotype H 17.5, D 7.5, W 6.9.

Type material.

NHMUK 1907.11.21.32, lectotype and one paralectotype (da Costa coll.).

Remarks.

da Costa did not state on how many specimens his description was based. A lectotype was designated by Breure (1979); the paralectotype is slightly damaged at the palatal lip. The current systematic position follows Richardson (1995: 111), but the taxon is now placed in the subgenus Mesembrinus Albers, 1850 (comb. n.).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) conicus da Costa, 1907.

Bulinus conspersus Sowerby I, 1833

http://species-id.net/wiki/Bulinus_conspersus

Figs 6C, L15i
Bulinus conspersus Sowerby I 1833b: 73.
Bulimus conspersus; Reeve 1848 [1848–1850]: pl. 22 fig. 137.
Type locality.

[Peru] ”collinis prope Lima” (see remarks).

Label.

”Peru”. M.C. label style I, V.

Dimensions.

”Long. 0.65, lat. 0.4 poll. [H 16.5, D 10.1 mm]”; figured specimen herein H 19.8, D 12.1, W 5.5.

Type material.

NHMUK 20100619, five probable syntypes (Cuming coll.).

Remarks.

The original material of Sowerby has not been traced; however, one of the specimens from this lot corresponds to Reeve (1848 [1848-1850]) and the specimens are regarded probable syntypes. Ramírez (2004) has restricted the type locality to Lomas de Atocongo, near Lima.

Current systematic position.

Bulimulidae, Bostryx conspersus (Sowerby I, 1833).

Bulimus convexus Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_convexus

Figs 39A–C, L14iv
Bulimus convexus Pfeiffer 1855f: 116; Pfeiffer 1859b: 444; Breure 1979: 108 (lectotype designation).
Drymaeus (Drymaeus) convexus; Breure and Eskens 1981: 16. pl. 8 fig. 7.
Type locality.

”New Granada”.

Label.

”Ecuador”, added in a later handwriting; taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 38, diam. 14 mill.”; figured specimen herein H 38.5, D 16.2, W 7.2.

Type material.

NHMUK 1975192, lectotype; 1975193, two paralectotypes (Cuming coll.).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) convexus (Pfeiffer, 1855).

Helix cora d’Orbigny, 1835

http://species-id.net/wiki/Helix_cora

Figs 58A–B, L14v
Helix cora d’Orbigny 1835: 15.
Bulimus cora d’Orbigny 1837 [1834–1847]: 307, pl. 34 figs 14–15 [3 April 1837; text 6 May 1838].
Type locality.

[Peru] ”republica Peruviana”.

Label.

”Perou”, in d’Orbigny’s handwriting.

Dimensions.

”Latit. 44 millim.; longit. 29 millim.”; figured specimen herein H 43.9, D 28.0, W 6.7.

Type material.

NHMUK 1854.12.4.124, lectotype and two paralectotypes (d’Orbigny coll.).

Remarks.

d’Orbigny 1838 [1834–1847] specifies the type locality as ”sur les montagnes des environs de Trujillo”, which is in Dept. La Libertad. The lot contains two juvenile, and one adult specimen; the latter corresponds to d’Orbigny’s pl. 34 fig. 15 and is here designated lectotype (design. n.). The arrangement by Richardson (1995: 356), referring d’Orbigny 1837–1838 to Bulinus proteus Broderip in Broderip and Sowerby I 1832, may be in error.

Current systematic position.

Bulimulidae, Neopetraeus cora (d’Orbigny, 1835).

Bulimus coriaceus Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_coriaceus

Fig. 60K–L, L67ii
Bulimus coriaceus Pfeiffer 1857b: 318.
Type locality.

[Mexico] ”Cordova”.

Label.

”Mexico”, taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 18, diam. 9 mill.”. Figured specimen H 17.1, D 9.5, W 5.8.

Type material.

NHMUK 1975409, three possible syntypes (Cuming coll.).

Remarks.

The type material was based on material collected by Sallé; however, the material is accompanied by a taxon label in Pfeiffer’s handwriting and is herein considered as possible syntypes. Pfeiffer did not state on how many specimens his description was based. The type material was up till now unfigured. The current systematic position follows Thompson (2011: 106).

Current systematic position.

Bulimulidae, Bulimulus coriaceus (Pfeiffer, 1857).

Bostryx (Bostryx) pygmaeus costatus Weyrauch, 1960

http://species-id.net/wiki/Bostryx_pygmaeus_costatus

Fig. 1I
Bostryx (Bostryx) pygmaeus costatus Weyrauch 1960b: 122, pl. 11 figs 12–16; Breure 1979: 53; Neubert and Janssen 2004: 206, pl. 4 fig. 47; Breure 2012a: 7.
Type locality.

[Peru] ”Bei Alis, auf der rechten Seite des Río Alis, Zufluss des Río Cañete, 3300 m”.

Label.

”C-Peru, Alis, Río Alis, affluent of Río Cañete, 3300 m”; printed label.

Dimensions.

”H. 8.7 D. 5.1”; figured specimen herein H 8.45, D 4.9, W 5.2.

Type material.

NHMUK 1975357, one paratype (ex Weyrauch, WW 318).

Current systematic position.

Bulimulidae, Bostryx pygmaeus Weyrauch, 1960.

Bostryx (Elatibostryx) imeldae costifer Weyrauch, 1960

http://species-id.net/wiki/Bostryx_imeldae_costifer

Fig. 1J
Bostryx (Elatibostryx) imeldae costifer Weyrauch 1960b: 129, pl. 12 figs 35–38; Breure 1979: 53; Neubert and Janssen 2004: 206, pl. 7 fig. 77; Breure 2012a: 7.
Type locality.

[Peru, Dept. Lima] ”Mittel-Peru, Quichao, 5 km von Laraos, am Fusspad nach Yauyos, auf der linken Seite des Río Mayo im Tal des Río Cañete, 3500 m”.

Label.

”C-Peru, Quichao, 5 kms from Laraos, valley of Río Cañete, 3500 m”; printed label.

Dimensions.

”H. 11.0 D. 5.0 [mm]”; figured specimen herein H 9.6, D 4.2, W 6.2.

Type material.

NHMUK 1975354, one paratype, ex Weyrauch (WW 3319).

Current systematic position.

Bulimulidae, Bostryx imeldae Weyrauch, 1958.

Bulimus cotopaxiensis Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_cotopaxiensis

Figs 69G–H, 69J, L15ii
Bulimus cotopaxiensis Pfeiffer 1853d: 419; Pfeiffer 1854c: 155; Pfeiffer 1854 in Küster and Pfeiffer 1840–1865: 103, pl. 33 figs 9–10; Breure 1979: 86; Breure and Borrero 2008: 18.
Bulimulus cotopaxiensis; Pilsbry 1897 [1897–1898]: 31, pl. 4 figs 50–51.
Scutalus (Vermiculatus) cotopaxiensis; Breure 1978: 175, pl. 9 fig. 9 (lectotype designation).
Type locality.

[Ecuador] ”reipublicae Aequatoris, montem Cotopaxi”.

Label.

”Cotopaxi, Equador / Monsr Bourcier / Consul General”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 34, diam. 16 mill.”; figured specimen herein H 33.9, D 17.1, W 6.2.

Type material.

NHMUK 1975370, lectotype; 1975371, two paralectotypes (Cuming coll.).

Remarks.

The original paper only quotes material collected by Bourcier as a variety from Cayembe. However, if the type was in the Pfeiffer collection it must now considered to be lost (Dance 1966). Both the locality, and measurements agree with Pfeiffer’s paper; the taxon label is from his hand, so there is no doubt that these specimens can be considered as type material. The shell height of the lectotype as quoted by Breure (1978) is in error. This species was listed as synonym under Scutalus anthisanensis (Pfeiffer, 1853) by Richardson (1995: 346); however, the shell shape of the two taxa is different, and provisionally the two taxa are kept separate awaiting further studies.

Current systematic position.

Bulimulidae, Kuschelenia (Vermiculatus) cotopaxiensis (Pfeiffer, 1853) (comb. n.).

Helix crepundia d’Orbigny, 1835

http://species-id.net/wiki/Helix_crepundia

Figs 15H–I, L14vi
Helix crepundia d’Orbigny 1835: 14.
Bulimus crepundia; d’Orbigny 1837 [1834–1847]: 275, pl. 33 figs 18–19 [19 June / 7 Aug 1837; text 23 April 1838].
Bulimulus crepundia; Pilsbry 1897 [1897–1898]: 90, pl. 11 figs 33–34.
Naesiotus crepundia; Breure 1975b: 1146, pl. 8 fig. 5.
Type locality.

”provincia Chiquitensi, republica Boliviana”; see Breure 1973: 116.

Label.

Only the taxon name is given in d’Orbigny’s handwriting.

Dimensions.

”Latit. 15 millim., long. 10 millim.”, corrected to ”Longueur totale, 25 millimètres” in d’Orbigny 1838 [1834–1847]; figured specimen herein H 26.6, D 11.1, W 7.9.

Type material.

NHMUK 1854.12.4.173, six paralectotypes; 1854.12.4.174, one paralectotype (d’Orbigny coll.).

Remarks.

One specimen was marked with ‘x’ on the original board, which suggests a special selection; however, it is smaller than the type (H 23.7) and has the lip damaged. Another specimen, which shows the full lip as in d’Orbigny’s figure, is here figured. The lectotype is in MNHN (Breure 1975b). In the collation of Coan et al. (2013a) the plates as issued in ‘Livraison 25’ and ‘Livraison 26’ appear to be incorrect; plate 38 is mentioned twice, while plate 33 is missing. Since no taxonomic implications seem involved, these dates have both been cited herein as it is not clear which is the correct date for each plate.

Current systematic position.

Bulimulidae, Naesiotus crepundia (d’Orbigny, 1835).

Bulimus cretaceus Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_cretaceus

Figs 65C–F, L15iii
Bulimus cretaceus Pfeiffer 1855g: 123; Breure 1979: 83.
Scutalus (Scutalus) cretaceus (Pfeiffer); Breure 1978: 166, figs 277–278, pl. 8 fig. 7 (lectotype designation).
Type locality.

”Eastern Islands [sic, see remarks] (Captain Keppell)”.

Label.

”No locality but I / had the from Cap. Keppell / who got them in the / Eastern Islands but don’t / recollect”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 36, diam. 18 mill.”; figured specimen herein H 35.2, D 23.2, W 6.2.

Type material.

NHMUK 1975388, lectotype; 1975389, one paralectotype, ex Keppell (Cuming coll.).

Remarks.

Pilsbry (1902 [1901–1902]: 141) was the first to recognize that this species was from Peru. The type locality is here restricted to Dept. Cajamarca, Río Jequetepeque, near Tembladera (restr. n.).

Current systematic position.

Bulimulidae, Scutalus cretaceus (Pfeiffer, 1855).

Bulinus crichtoni Broderip, 1836

http://species-id.net/wiki/Bulinus_crichtoni

Figs 59A–C, L15iv
Bulinus crichtoni Broderip 1836: 44.
Newboldius crichtoni; Breure 1979: 99.
Type locality.

[Peru] ”ad Ambo juxta Huanuco Peruviae”.

Label.

”Peru, Ambo, Sir A Crichton”, added in pencil. Taxon label probably in Broderip’s handwriting.

Dimensions.

”Long. 3 (circiter), latit. 1 1/8 poll. [H 76.0, D 28.5 mm]”; holotype H 71.3, D 33.5, W 4+.

Type material.

NHMUK 1958.9.3.4, holotype (Broderip coll.).

Remarks.

The top whorl of the specimen is missing. Broderip refers to ”the apex of the shell under description, the only specimen I ever saw, is broken, and its actual length is 2 inches and 7/8 [72.8 mm]”; we consider this specimen the holotype.

Current systematic position.

Bulimulidae, Newboldius crichtoni (Broderip, 1836).

Helix culminea d’Orbigny, 1835

http://species-id.net/wiki/Helix_culminea

Figs 66F–H, L15v
Helix culminea d’Orbigny 1835: 13; Breure 1979: 88.
Bulimus culmineus; d’Orbigny 1837 [1834–1847]: 288, pl. 33 figs 8–9 [19 June / 7 Aug 1837; text 6 May 1838].
Bulimulus culmineus; Pilsbry 1897 [1897–1898]: 25, pl. 5 figs 74–75.
Scutalus culmineus culmineus; Breure 1975b: 1143, pl. 1 fig. 3.
Type locality.

”culminibus Andesensibus, republica Boliviana” (see remarks).

Label.

”Titicaca, Bolivia”, in d’Orbigny’s handwriting.

Dimensions.

”Latit. 17 millim., longit. 13 millim.”; figured specimen herein H 31.2, D 14.1, W 6.3.

Type material.

NHMUK 1854.12.4.198, six paralectotypes (d’Orbigny coll.).

Remarks.

d’Orbigny (1838 [1834–1847]: 289) specified the type locality as ”les montagnes de la province de Carangas, à l’ouest d’Oruro, principalement sur celle du ‘Pucara’, à cinq lieues du bourg de Totora”. Breure (1973: 130, fig. 3) located ‘Pucara’ at approximately 17°58’S, 068°21’W. This place could not be found using modern gazetteers, but Totora is located at 17°47’S, 068°08’W. The old French distance ‘lieu’ equals 4.44 kms, thus the Pucara mountain is about 22.2 km from Totora; however, d’Orbigny did not state in which direction he travelled. Breure (1975) restricted the type locality to this area, contrary to Weyrauch (1967a: 393) who selected the peninsula Capachica, in the Peruvian part of Lake Titicaca, as type locality; he referred to Crawford (1939) who had studied the types of d’Orbigny in NHMUK. However, Weyrauch may have overlooked that d’Orbigny (1838 [1834–1847]: 289) specified a second locality ”sur toutes les îles et sur toutes les montagnes du lac de Titicaca”. The specimens from the latter locality are those in the NHMUK collection, while those from ”Carangas” are in the MNHN collection (Breure 1975b); the lectotype has been selected from the latter collection, and the type locality is thus in Bolivia.

Current systematic position.

Bulimulidae, Kuschelenia (Kuschelenia) culminea (d’Orbigny, 1835).

Bulimulus (Naesiotus) curtus Reibisch, 1892

http://species-id.net/wiki/Bulimulus_curtus

Figs 13G–H, L16i
Bulimulus (Naesiotus) curtus Reibisch 1892: 21, pl. 1 fig. 14.
Naesiotus curtus; Breure 1979: 68.
Type locality.

[Ecuador, Galápagos, Isla San Cristóbal] ”Chatam-Island [sic] (Wolf)”.

Label.

”Chatham Is. Galapagos”.

Dimensions.

”Long. 10, 25, diam. maj. 5 mm”; figured specimen herein H 9.6, D 4.62, W 6.9.

Type material.

NHMUK 1894.6.8.8–9, two syntypes, T. Wolf leg.

Remarks.

Reibisch did not state on how many specimens his description was based. The current systematic position follows Richardson (1995: 217).

Current systematic position.

Bulimulidae, Naesiotus curtus (Reibisch, 1892).

Bulimus cuticula Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_cuticula

Figs 42G–I, L16ii
Bulimus cuticula Pfeiffer 1855d: 95; Breure 1979: 108 (lectotype designation).
Drymaeus (Drymaeus) cuticulus; Breure and Eskens 1981: 20; Simone 2006: 136, fig. 446.
Type locality.

”Rio Janeiro”.

Label.

”Rio”, taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 28, diam. 13 mill.”; figured specimen herein H 27.5, D 13.75, W 4.7.

Type material.

NHMUK 1975451, lectotype (Cuming coll.).

Remarks.

There is only one specimen, of which the lip has been broken, and the last whorl is badly damaged. The specimen is probably subadult.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) cuticula (Pfeiffer, 1855).

Bulimus cuzcoensis Reeve, 1849

http://species-id.net/wiki/Bulimus_cuzcoensis

Figs 43A–C, L16iii
Bulimus cuzcoensis Reeve 1849 [1848–1850]: pl. 71 fig. 514; Reeve 1850: 98; Pfeiffer 1853d: 344; Breure 1979: 108 (lectotype designation).
Type locality.

[Peru] ”Cuzco, Bolivia [sic]”.

Label.

”Peru”, added in a later handwriting. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 36.6, D 15.7, W 7.2.

Type material.

NHMUK 1975453, lectotype; 1975454, two paralectotypes (Cuming coll.).

Remarks.

The current systematic position follows Richardson (1995: 112).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) cuzcoensis (Reeve, 1849).

Bulimulus dacostae Sowerby III, 1892

http://species-id.net/wiki/Bulimulus_dacostae

Figs 33M–O, L16iv
Bulimulus dacostae Sowerby III 1892: 297, pl. 23 figs 15–16.
Drymaeus dacostae; Pilsbry 1898 [1897–1898]: 214, pl. 43 figs 77–78; Linares and Vera 2012: 184.
Drymaeus (Drymaeus) dacostae; Breure 1979: 108.
Type locality.

[Colombia] ”Bogota (Mus. Da Costa)”.

Label.

”Bogota”, in da Costa’s handwriting.

Dimensions.

”Long. 26, diam. maj. 11 millim.”; figured specimen herein H 26.3, D 11.0, W 6.3.

Type material.

NHMUK 1907.11.21.51, holotype (da Costa coll.).

Remarks.

Sowerby writes ”I have seen as yet only one specimen of this species”. Therefore the specimen is considered as holotype.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) dacostae (Sowerby III, 1892).

Bulimus darwini Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_darwini

Figs 13J, L66vii
Bulimus darwini Pfeiffer 1846a: 29; Pfeiffer 1848b: 199; Reeve 1848 [1848–1850]: pl. 21 fig. 136.
Type locality.

[Ecuador] ”Gallapagos [sic] Islands”.

Label.

”Galapagos Is.”, in E.A. Smith’s handwriting. M.C. label style III, V.

Dimensions.

”Long. 17, diam. 19 mill.”; figured specimen herein H 8.64, D 6.87 W 4.9.

Type material.

NHMUK 20100523, two possible syntypes, C. Darwin leg. (Cuming coll.).

Remarks.

The two juvenile specimens were labelled ”ustulatus Sow.” by E.A. Smith as coming from the Cuming museum, and were probably collected by Darwin during his visit to the archipelago with the ‘Beagle’. They are considered as possible syntypes. The current systematic position follows Richardson (1995: 217).

Current systematic position.

Bulimulidae, Naesiotus darwini (Pfeiffer, 1846).

Bulimus decussatus Reeve, 1849

http://species-id.net/wiki/Bulimus_decussatus

Figs 56G–I, L16vi
Bulimus decussatus Reeve 1849 [1848–1850]: pl. 72 fig. 519; Reeve 1850: 99; Pfeiffer 1853d: 431; Breure 1979: 100.
Neopetraeus decussatus; Breure 1978: 215 (lectotype designation).
Type locality.

”Andes of Caxamarca [sic, Cajamarca], Peru”.

Label.

”Andes of Caxamarca, Peru”; in unknown handwriting, obviously added in a later stage (see remarks).

Dimensions.

Not given; figured specimen herein H 37.7, D 16.5, W 7.7.

Type material.

NHMUK 1975180, lectotype (Cuming coll.).

Remarks.

The printed label, that is typical of specimens figured by Reeve, has been lost. The label, probably in the handwriting of a NHMUK employee during the latter half of the 19th century, refers to Reeve’s species number (519), stating that it is ”the type”. In the lower left corner, ”one on right” has been added (probably by E.A. Smith), which seems to indicate that the specimens originally were glued on board. There is a second note ”now marked x”, which apparently refers to the type specimen. See also Bulimus myristicus Reeve, 1849, which is a junior synonym (Breure 1978).

Current systematic position.

Bulimulidae, Neopetraeus decussatus (Reeve, 1849).

Bulimus delumbis Reeve, 1849

http://species-id.net/wiki/Bulimus_delumbis

Figs 9C, L17i
Bulimus delumbis Reeve 1849 [1848–1850]: pl. 76 fig. 555; Pfeiffer 1853d: 418; Breure 1979: 53.
Bulimulus delumbis; Pilsbry 1901 [1901–1902]: 138, pl. 25 fig. 15.
Bostryx delumbis; Breure 1978: 71 (lectotype designation).
Type locality.

”—?”.

Label.

”Peru + Bolivia”, added in a later handwriting. M.C. label style I, V.

Dimensions.

Not given; figured specimen herein H 21.2, D 14.0, W 6.0.

Type material.

NHMUK 1975124, lectotype; 1975558, one paralectotype (Cuming coll.).

Remarks.

The current systematic position follows Richardson (1995: 21).

Current systematic position.

Bulimulidae, Bostryx delumbis (Reeve, 1849).

Bulimus demotus Reeve, 1850

http://species-id.net/wiki/Bulimus_demotus

Figs 25F, L17iii
Bulimus feriatus Reeve 1848 [1848–1850]: pl. 54 fig. 354; Breure 1979: 118 (lectotype designation).
Bulimus demotus Reeve 1850 [1848–1850]: ii. New name for Bulimus feriatus Reeve, 1848 (Dec.), not Reeve, 1848 (Nov.); Pfeiffer 1853d: 340.
Drymaeus demotus; Pilsbry 1898 [1897–1898]: 306, pl. 43 fig. 81.
Type locality.

”Venezuela”.

Label.

”Venezuela”. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 31.8, D 15.3, W 6.2.

Type material.

NHMUK 1975504, lectotype; 1975505, two paralectotypes (Cuming coll.).

Remarks.

The lectotype corresponds to Reeve’s figure. The current systematic position follows Richardson (1995: 131).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) granadensis (Pfeiffer, 1848).

Bulimus denickei J.E. Gray, 1852

http://species-id.net/wiki/Bulimus_denickei

Figs 7G, L17iv
Bulimus denickei J.E. Gray 1852: 92.
Bostryx denickei; Breure 1979: 53.
Type locality.

[Peru, Dept. Arequipa] ”Chala, near Callao”.

Label.

”Chala, near Callao”. M.C. label style IV.

Dimensions.

Not given; figured specimen herein H 26.4, D 17.5, W 5.9.

Type material.

NHMUK 1851.5.15.1–3, three syntypes, E. Denicke leg. (Cuming coll.).

Remarks.

The current systematic position follows Richardson (1995: 40).

Current systematic position.

Bulimulidae, Bostryx reentsi (Philippi, 1851).

Bulimus dentritis Morelet, 1863

http://species-id.net/wiki/Bulimus_dentritis

Figs 16G, L17ii
Bulimus dentritis Morelet, 1863: 206, pl. 9 fig. 5; Breure 1979: 62.
Bulimulus dentritis; Breure 1978: 72 (lectotype designation).
Type locality.

[Peru, Dept. Cuzco] ”près de Huiro, vallée de Santa-Anna”.

Label.

”Miota, Vallée de Sta Anna”, in Morelet’s handwriting.

Dimensions.

”Longit. 20, diam. 8 mill.”; figured specimen herein H 19.9, D 7.42, W.

Type material.

NHMUK 1893.2.4.237, paralectotype (Morelet coll.).

Remarks.

The species is hitherto classified as Bulimulus. However, the protoconch sculpture consists of axial riblets; it is here re-classified under Naesiotus sensu Breure 1979. The NHMUK specimen fits the measurements given by Morelet, but is not from the type locality. Other type material is present in MHNG from various other localities, of which one specimen from the type locality was designated lectotype (Breure 1978).

Current systematic position.

Bulimulidae, Naesiotus dentritis (Morelet, 1863) (comb.n.).

Bulimus depictus Reeve, 1849

http://species-id.net/wiki/Bulimus_depictus

Figs 22C, L17v
Bulimus depictus Reeve 1849 [1848–1850]: pl. 74 fig. 529; Pfeiffer 1853d: 337; Breure 1979: 118 (lectotype designation).
Drymaeus depictus; Pilsbry 1898 [1897–1898]: 290, pl. 45 fig. 17.
Drymaeus (Mesembrinus) depictus; Breure and Eskens 1981: 55, pl. 6 fig. 5.
Type locality.

”New Granada”.

Label.

”Venezuela”, added in a later handwriting. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 27.9, D 12.1, W 5+.

Type material.

NHMUK 1975529, lectotype; 1975530, two paralectotypes (Cuming coll.).

Remarks.

The top of the lectotype is damaged. The difference between the published type locality and the locality on the label is explained by political changes around 1850.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) depictus (Reeve, 1849).

Bulimus depstus Reeve, 1849

http://species-id.net/wiki/Bulimus_depstus

Figs 6B, L17vi
Bulimus depstus Reeve 1849 [1848–1850]: pl. 73 fig. 525 [text species 524]; Reeve 1850: 97; Pfeiffer 1853d: 428; Breure 1979: 53.
Bostryx depstus; Breure 1978: 73 (lectotype designation).
Type locality.

”Chachapoyas, Alto-Peru”.

Label.

No label found (see remarks).

Dimensions.

Not given; figured specimen herein H 18.5, D 9.71, W 7.1.

Type material.

NHMUK 1975318, lectotype; 1975319, two paralectotypes (Cuming coll.).

Remarks.

Breure (1978) stated he found a label with the text ”Peru”; however, this label may have been lost.

Current systematic position.

Bulimulidae, Bostryx depstus (Reeve, 1849).

Bulinus derelictus Broderip, 1832

http://species-id.net/wiki/Bulinus_derelictus

Figs 10H, L17vii
Bulinus derelictus Broderip in Broderip and Sowerby I 1832b: 107; Reeve 1848 [1848–1850]: pl. 23 fig. 151; Neubert and Janssen 2004: 208, pl. 6 fig. 65.
Type locality.

”Cobijam Bolivia [now Chile] (Puerto del Mar)”.

Label.

”Bolivia S. America”, with a taxon label in Pfeiffer’s handwriting. M.C. label style I, V.

Dimensions.

”Long. 6/12, lat. 5/8 poll. [H 12.7, D 15.8 mm, see remarks]”; figured specimen herein H 30.3, D 18.8, W 6.6.

Type material.

NHMUK 20100609, four probable syntypes (Cuming coll.).

Remarks.

Broderip described this taxon from the Cuming collection, but did not state on how many specimens his description was based. The original measurments seem to be erroneous, as these indicate a height/diameter ratio uncommon for this genus. Neubert and Janssen (2004) reported three specimens, with a maximum height of 23.1 and diameter of 15.3 mm, labelled ”Peru” (ex Cuming), which they consider syntypes. The specimen corresponding to Reeve’s figure has been found in the NHMUK collection, together with three others; they are considered probable syntypes.

Current systematic position.

Bulimulidae, Bostryx derelictus (Broderip in Broderip and Sowerby I 1832).

Bulimus deshayesi Pfeiffer, 1845

http://species-id.net/wiki/Bulimus_deshayesi

Figs 24D, L18i
Bulimus deshayesi Pfeiffer 1845c: 73; Pfeiffer 1848b: 200; Reeve 1848 [1848–1850]: pl. 40 fig. 250; Breure 1979: 118 (lectotype designation).
Drymaeus deshayesi; Pilsbry 1898 [1897–1898]: 303, pl. 45 fig. 27; Linares and Vera 2012: 194.
Drymaeus (Mesembrinus) deshayesi; Breure and Eskens 1981: 55, pl 7 fig. 1.
Type locality.

”Locality unknown”.

Label.

”New Granada”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.

Dimensions.

”Long. 45, diam. 15 mill.”; figured specimen herein H 44.5, D 16.8, W 8.1.

Type material.

NHMUK 1975526, lectotype; 1975527, one paralectotype (Cuming coll.).

Remarks.

The current systematic position follows the revision of Breure and Borrero (unpublished data).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) deshayesi (Pfeiffer, 1845).

Bulimus devians Dohrn, 1863

http://species-id.net/wiki/Bulimus_devians

Figs 4C, L18ii
Bulimus devians Dohrn 1863: 155.
Type locality.

No type locality given [Peru].

Label.

”[Peru]”, added in a later handwriting. M.C. label style III.

Dimensions.

”Long. 15, lat. 6 mill.”; figured specimen herein H 14.9, D 6.78, W 5.1.

Type material.

NHMUK 1975339, one possible syntype (Cuming coll.).

Remarks.

Dohrn’s collection is known to be completely destroyed in the 1939–1945 war (Dance 1966, Köhler 2007). The specimen fits the original measurement, although it is unknown on how many specimens Dohrn based his description. Moreover, the lack of original data on the label prevents to ascertain the status of this specimen, and the specimen is only treated as possible syntype.

Current systematic position.

Bulimulidae, Bostryx devians (Dohrn, 1863).

Bulinus discrepans Sowerby I, 1833

http://species-id.net/wiki/Bulinus_discrepans

Figs 23D, L18iii
Bulinus discrepans Sowerby I 1833b: 72; Reeve 1848 [1848–1850]: pl. 23 fig. 145.
Type locality.

”Americâ Centrali” (see remarks).

Label.

”Conchagua, C. America”. M.C. label style IV, V.

Dimensions.

”Long. 0.7, lat. 0.33 poll. [H 17.7, D 8.4 mm]”; figured specimen herein H 20.7, D 9.3, W 6.5.

Type material.

NHMUK 1975181, four syntypes (Cuming coll.).

Remarks.

Sowerby added is his remarks ”This was found at Conchagua” [El Salvador, dept. La Union], which is thus the type locality. Of the four specimens in the lot, two are juveniles and one is subadult; the largest specimen is marked ”59”. The current systematic position follows Thompson (2011: 119).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) discrepans (Sowerby I, 1833).

Bulimus dombeyanus ‘Férussac’ Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_dombeyanus

Figs 37I–K, L18v
Helix dombeyana ‘F[érussac]’ Pfeiffer 1842: 76 (= Bulimus) [nomen nudum].
Bulimus dombeyanus ‘Fér.’ Pfeiffer 1842: 114 [nomen nudum]; ‘Fér.’ Pfeiffer 1846c: 83.
Type locality.

”Peru (Mus. Britt.)”.

Label.

”Peru”. M.C. label style IV.

Dimensions.

”Long. 56, diam. 26 mill.”; figured specimen herein H 56.8, D 31.7, W 7.1.

Type material.

NHMUK 1844.10.2.32, syntype (ex Férussac, ”purchd of Turner”).

Remarks.

This taxon is generally regarded as a Mexican species (Pilsbry 1899, Solem 1957, Thompson 2011); the locality thus could be a case of mislabeling.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) dombeyanus (Pfeiffer, 1846).

Bulimus dubius Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_dubius

Figs 25C, L18iv
Bulimus dubius Pfeiffer 1853b: 257; Breure 1979: 118 (lectotype designation).
Drymaeus (Mesembrinus) dubius; Breure and Eskens 1981: 57, pl. 7 fig. 10.
Drymaeus dubius; Linares and Vera 2012: 194.
Type locality.

”in Andibus Novae Granadae”.

Label.

”Andes, N. Granada”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 28, diam. 10 mill.”; figured specimen herein H 27.1, D 11.0, W 6.2+.

Type material.

NHMUK 1975519, lectotype; 1975520, two paralectotypes (Cuming coll.).

Remarks.

Of the lectotype the lip and the apex are damaged. Of the paralectotypes, one shell is broken.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) dubius (Pfeiffer, 1853).

Bulimulus (Drymaeus) dukinfieldi Melvill, 1900

http://species-id.net/wiki/Bulimulus_dukinfieldi

Figs 60C–D, L19i
Bulimulus (Drymaeus) dukinfieldi Melvill 1900: 116, fig.
?Bulimulus dukinfieldi; Breure 1979: 62.
Bulimulus dukenfieldi; Simone 2006: 118, fig. 359.
Thaumastus dukinfieldi; Simone 2006: 152, fig. 518.
Type locality.

”Salto Grande do Rio dos Patos (upper waters of Rio Ivahy), Paraná, Brazil”.

Label.

”Salto Grande do Rio / dos Patos. (Upper waters of River Ivahy). / Paraná. Brazil.”.

Dimensions.

”Long. 28, lat. 13 mm”; figured specimen herein H 28.1, D 14.8, W 5.9.

Type material.

NHMUK 1900.9.27.2, lectotype (ex Melvill).

Remarks.

Melvill wrote ”Two specimens, precisely similar, were obligingly handed me by Mr. R. Dukinfield Darbishire”; ”the type” was placed in the NHMUK collection, the whereabouts of the second specimen are unknown. The reference to ”HT 1900.9.27.2” in Breure 1979 has to be interpreted as lectotype designation under Art. 74.6 ICZN. This shell seems to have been damaged several times during life-time, which may have influenced the shape of the shell and aperture. This is probably the reason why this taxon has been referred to different genera (Simone 2006: 118, 152); the classification of Breure (1979) is here tentatively retained until further studies have clarified the systematic position of this nomen inquirendum.

Current systematic position.

Bulimulus (?) dukinfieldi Melvill, 1900. Nomen inquirendum.

Bulimus dunkeri Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_dunkeri

Figs 50D–F, L19ii
Bulimus dunkeri Pfeiffer in Philippi 1846 [1845–1847]: 112, pl. 4 fig. 10; Breure 1979: 108 (lectotype designation).
Drymaeus (Drymaeus) dunkeri; Breure and Eskens 1981: 20; Thompson 2011: 110.
Type locality.

”respublica mexicana, prope Michoacan”.

Label.

”Michoacan”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 18, diam. 8’’’ [H 39.2, D 17.4 mm]”; figured specimen herein H 34.8, D 18.5, W 6.0.

Type material.

NHMUK 1975512, lectotype; 1975513, two paralectotypes (Cuming coll.).

Remarks.

The original measurements are interpreted as German lines (2.18 mm), and do not correspond to those of the specimen in the NHMUK. However, Pfeiffer’s collection is known to have been destroyed (Dance 1966), and type material of this taxon is neither in the SMF (Neubert and Janssen 2004) nor ZMB (Köhler 2007) collections. The current systematic position follows Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) dunkeri dunkeri (Pfeiffer, 1846).

Bulimulus (Peronaeus) durangoanus Martens, 1893

http://species-id.net/wiki/Bulimulus_durangoanus

Figs 73F–H, L19iii
Bulimulus (Peronaeus) durangoanus Martens 1893 [1890–1901]: 246, pl. 15 figs 11–11a.
Bulimulus durangoanus; Pilsbry 1897 [1897–1898]: 127, pl. 18 figs 32–33.
Type locality.

”N. Mexico, Villa Lerdo, State of Durango”.

Label.

”Mexico”, taxon label in Martens’ handwriting.

Dimensions.

”Long. 15, diam. 6 (...) millim.”. Figured specimen H 15.0, D 6.4, W 6.3.

Type material.

NHMUK 1901.6.22.871, holotype (ex Godman).

Remarks.

Martens wrote ”I have seen only one specimen of this species”; the specimen found is thus the holotype. The current systematic position follows Thompson (2011: 124).

Current systematic position.

Bulimulidae, Naesiotus durangoanus (Martens, 1893).

Bulimus dutaillyi Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_dutaillyi

Figs 22G, L19iv
Bulimus dutaillyi Pfeiffer 1857d: 390; Breure 1979: 118 (lectotype designation).
Drymaeus (Mesembrinus) dutaillyi; Breure and Eskens 1981: 57, pl. 7 fig. 2; Simone 2006: 145, fig. 485.
Type locality.

”Brazils”.

Label.

”Brazils Mon. Dutailly”, taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 31, diam. 12 mill.”; figured specimen herein H 30.8, D 12.5, W 7.1.

Type material.

NHMUK 1975516, lectotype, Dutailly leg. (Cuming coll.).

Remarks.

Simone (2006) erroneously had the shell height as 33.0 mm.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) dutaillyi (Pfeiffer, 1857).

Bulimus dysoni Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_dysoni

Figs 62K–L, L19v
Bulimus dysoni Pfeiffer 1846b: 39; Pfeiffer 1848b: 183; Reeve 1849 [1848–1850]: pl. 62 fig. 425; Breure 1979: 62.
Bulimulus dysoni; Pilsbry 1897 [1897–1898]: 56, pl. 10 fig. 83.
Bulimulus (Bulimulus) dysoni; Breure 1974: 47, pl. 4 figs 4–5.
Type locality.

”Honduras”.

Label.

”Honduras”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.

Dimensions.

”Long. 20, diam. 9 1/2 mill.”; figured specimen herein H 19.2, D 9.5, W 7.4.

Type material.

NHMUK 1975453, three syntypes (Cuming coll.).

Remarks.

The specimen figured by Reeve is damaged at the penultimate whorl.

Current systematic position.

Bulimulidae, Bulimulus dysoni (Pfeiffer, 1846).

Bulimus edwardsi Morelet, 1863

http://species-id.net/wiki/Bulimus_edwardsi

Figs 67C–D, 67L, L19vi
Bulimus edwardsi Morelet 1863: 182, pl. 9 fig. 1; Breure 1979: 88.
Bulimulus edwardsi; Pilsbry 1897 [1897–1898]: 27, pl. 7 figs 11–13.
Type locality.

[Peru, Dept. Huancavelica] ”le type (...) Huancabelica; la variété, (...) vallée de Huanta” [Dept. Ayacucho].

Label.

”Pérou Huanta”, taxon label (”var.”) in Morelet’s handwriting.

Dimensions.

”Longit. 29; diam. 12 mill.”; figured specimen herein H 25.4, D 13.3, W 5.7.

Type material.

NHMUK 1893.2.4.177–178, two syntypes (Morelet coll.).

Remarks.

Morelet considered this to be a variable taxon and distinguished explicitly a smaller variety ”Testa minor, (...)”. Crawford (1939: 329) compared the two specimens in NHMUK to Scutalus bicolor (Sowerby, 1835), and expressed doubts whether the variety is identical to typical edwardsi; the typical form is considered by him as a synonym of Scutalus culmineus (d’Orbigny, 1835). With the current material at hand we prefer to treat Morelet’s material as a geographical form. Weyrauch (1967a: 386) considers Morelet’s taxon as a subspecies of Scutalus culmineus. This species is now placed in the genus Kuschelenia Hylton Scott, 1951.

Current systematic position.

Bulimulidae, Kuschelenia (Kuschelenia) culmineus edwardsi (Morelet, 1863) (comb. n.).

Bulimus effeminatus Reeve, 1848

http://species-id.net/wiki/Bulimus_effeminatus

Figs 62A–B, L20i
Bulimus effeminatus Reeve 1848 [1848–1850]: pl. 51 fig. 338; Breure 1979: 62.
Bulimulus effeminatus; Breure 1978: 143, pl. 9 figs 10–11 (lectotype designation).
Type locality.

”—?”.

Label.

”Valley of the / Madeleine [= Río Magdalena, Colombia] / N. Grenada”. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 31.1, D 14.2, W 7.9.

Type material.

NHMUK 1975508, lectotype; 1975509, two paralectotypes (Cuming coll.).

Remarks.

This taxon was erroneously placed under Drymaeus by Richardson (1995: 119).

Current systematic position.

Bulimulidae, Bulimulus effeminatus (Reeve, 1848).

Bulimus electrum Reeve, 1848

http://species-id.net/wiki/Bulimus_electrum

Figs 22E, L20ii
Bulimus electrum Reeve 1848 [1848–1850]: pl. 56 fig. 373; Breure 1979: 118 (lectotype designation).
Drymaeus electrum; Pilsbry 1898 [1897–1898]: 310, pl. 41 fig. 36.
Drymaeus (Mesembrinus) electrum; Breure and Eskens 1981: 57, pl. 7 fig. 3.
Type locality.

”Venezuela”.

Label.

”Venezuela”. M.C. label style IV, V.

Dimensions.

Not given; figured specimen herein H 29.6, D 14.1, W 5.8.

Type material.

NHMUK 1975510, lectotype; 1975511, two paralectotypes (Cuming coll.).

Remarks.

The lectotype corresponds to Reeve’s original figure. The current systematic position is in accordance with Richardson (1995).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) electrum (Reeve, 1848).

Drymaeus elsteri da Costa, 1901

http://species-id.net/wiki/Drymaeus_elsteri

Figs 33G–I, L20iii
Drymaeus elsteri da Costa 1901: 238, pl. 24 fig. 6; Pilsbry 1901 [1901–1902]: 156, pl. 48 fig. 53; Breure 1979: 108 (lectotype designation).
Type locality.

”Chachapoyas, prov. Amazonas, Peru”.

Label.

”Chacapoyas [sic]”, in da Costa’s handwriting.

Dimensions.

”Long. 34, diam. 15 mm.”; figured specimen herein H 33.9, D 14.9, W 6.0.

Type material.

NHMUK 1907.11.21.34, lectotype; 1907.11.21.35–36, two paralectotypes (da Costa coll.).

Remarks.

da Costa indicated that he had several specimens at hand when describing this taxon. The lectotype chosen is closely matching the original measurements.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) elsteri da Costa, 1901.

Bulimus emaciatus Morelet, 1863

http://species-id.net/wiki/Bulimus_emaciatus

Figs 3E, L20iv
Bulimus emaciatus Morelet 1863: 201, pl. 11 fig. 10; Breure 1979: 53.
Bulimulus (Peronaeus) emaciatus; Pilsbry 1896 [1895–1896]: 143, pl. 45 figs 27–28.
Bostryx emaciatus; Breure 1978: 74, fig. 101 (lectotype designation).
Type locality.

[Peru] ”les vallées et sur les plateaux de l’intérieur de la Sierra, depuis Ayacucho jusqu’au Cuzco” (see remarks).

Label.

”Pérou”, taxon label in Morelet’s handwriting.

Dimensions.

”Longit. 22, diam. 5 1/2 mill.”; figured specimen herein H 21.7, D 5.43, W 10.6.

Type material.

NHMUK 1893.2.4.248–250, three paralectotypes (Morelet coll.).

Remarks.

Breure (1978) upon selecting a lectotype from the MHNG collection, also fixated the type locality to Acobamba according to the label information of the specimen. Although there are several localities with that name in southern Peru, it is now assumed that it should be in Dept. Huancavelica [S -12.866667, W -74.566667], given the context of the other localities mentioned in Breure (1978). The current systematic position follows Richardson (1995: 23).

Current systematic position.

Bulimulidae, Bostryx emaciatus (Morelet, 1863).

Bulimus erectus Reeve 1849

http://species-id.net/wiki/Bulimus_erectus

Figs 60G–H, L20v
Bulimus erectus Reeve 1849 [1848–1850]: pl. 58 fig. 392; Pfeiffer 1853d: 439.
Bulimulus erectus; Pilsbry 1897 [1897–1898]: 60, pl. 10 fig. 99.
Type locality.

”Curiana, Venezuela”.

Label.

”Curiana, Venezuela”. M.C. label style I, V.

Dimensions.

Not given; figured specimen herein H 22.0 D 12.24 W 6.8.

Type material.

NHMUK 20100516, three syntypes, Dyson leg. (Cuming coll.).

Remarks.

Although none of the specimens exactly fits Reeve’s figure, they are regarded as possibly syntypes as they are originating from Dyson. See also Bulimus cacticolus Reeve, 1849.

Current systematic position.

Bulimulidae, Bulimulus erectus (Reeve, 1849).

Bulimus erosus Broderip in Broderip and Sowerby I 1832

http://species-id.net/wiki/Bulimus_erosus

Figs 7A, L20vi
Bulimus erosus Broderip in Broderip and Sowerby I 1832b: 106; Reeve 1848 [1848–1850]: pl. 22 fig. 140.
Bulimulus (Lissoacme) erosus; Pilsbry 1896 [1895–1896]: 160, pl. 49 fig. 34.
Type locality.

”Peruviâ, Huantajaya near Iquiqui”.

Label.

”Peru”, taxon label in Pfeiffer’s handwriting. M.C. label style I, V.

Dimensions.

”long. 11/12, lat. 6/12 poll. [H 23.2, D 12.66 mm]”; figured specimen herein H 22.9, D 13.32, W 6.3.

Type material.

NHMUK 20100612, five syntypes (Cuming coll.).

Remarks.

Breure (1979: 136), not having seen any type material, considered this taxon doubtfully a Bostryx and listed it as nomen inquirendum. The specimen now found in the NHMUK collection confirms this classification and agrees with Richardson (1995: 24).

Current systematic position.

Bulimulidae, Bostryx erosus (Broderip in Broderip and Sowerby I 1832).

Bulimus erubescens Pfeiffer, 1847

http://species-id.net/wiki/Bulimus_erubescens

Figs 25D, L21i
Bulimus erubescens Pfeiffer 1847: 112; Reeve 1848 [1848–1850]: pl. 57 fig. 381; Breure 1979: 118 (lectotype designation).
Drymaeus erubescens; Pilsbry 1899: 9, pl. 13 fig. 89.
Type locality.

”Locality unknown”.

Label.

”Jamaica”, added in a later handwriting on the front of the cardboard. M.C. label style I, V.

Dimensions.

”Long. 24, diam. 10 mill.”; figured specimen herein H 24.2, D 9.74 W 6.3.

Type material.

NHMUK 1975562, lectotype; 1975563, paralectotype (Cuming coll.).

Remarks.

The lectotype corresponds to Pfeiffer’s original measurements and is figured in Reeve (1848 [1848–1850]). The locality is probably erroneous; the species is not listed in Rosenberg and Muratov (2005). This taxon is considered by Richardson (1995: 172) to be a synonym of Drymaeus (Mesembrinus) rufescens (J.E. Gray, 1825), described from Jamaica but with very poor diagnostic features. The locality of the latter taxon may also be in error; the type material of Bulimus rufescens Gray, 1825 has not been located and may be lost.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) (?) rufescens (J.E. Gray, 1825).

Bulinus eschariferus Sowerby I, 1838

http://species-id.net/wiki/Bulinus_eschariferus

Figs 74A, L21ii
Bulinus eschariferus Sowerby I 1838 [Sowerby I and II 1832–1841]: fig. 85.
Bulimus eschariferus; Reeve 1848 [1848–1850]: pl. 20 fig. 121.
Bulimulus eschariferus; Pilsbry 1897 [1897–1898]: 108, pl. 22 fig. 9.
Type locality.

”Galapagos”.

Label.

”Chatham Is. / Galapagos”, in E.A. Smith’s handwriting. M.C. label style III.

Dimensions.

Not given. Figured specimen H 18.4, D 7.1, W 7.5.

Type material.

NHMUK 1975153, one possible syntype (Cuming coll.).

Remarks.

Sowerby did not state on how many specimens he based himself. The specimen found was figured by Reeve and the label is evidently based on the information given by him. The material is considered as possible syntype; see Breure and Ablett 2011: 2–12 for historical details on the Cuming collection and the relationship with Sowerby’s and Reeve’s taxa.

Current systematic position.

Bulimulidae, Naesiotus eschariferus (Sowerby I, 1833).

Bulimus excoriatus Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_excoriatus

Figs 58F–G, L21iii
Bulimus excoriatus Pfeiffer 1855g: 123; Breure 1979: 100.
Neopetraeus excoriatus; Breure 1978: 215, pl. 3 fig. 2 (lectotype designation).
Type locality.

”Andes of Peru (Captain Keppell)”.

Label.

”Andes of Peru”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 39, diam. 19 mill.”; figured specimen herein H 38.4, D 25.2, W 6.2.

Type material.

NHMUK 1975500, lectotype, Keppell leg. (Cuming coll.).

Remarks.

The specimen is subadult and aged; it is possibly a synonym of Neopetraeus catamarcanus (Pfeiffer, 1858), in which case Neopetraeus excoriatus would have priority.

Current systematic position.

Bulimulidae, Neopetraeus excoriatus (Pfeiffer, 1855).

Bulimus exornatus Reeve, 1849

http://species-id.net/wiki/Bulimus_exornatus

Fig. L21iv
Bulimus exornatus Reeve 1849 [1848–1850]: pl. 77 fig. 560; Breure 1979: 69 (lectotype designation).
Bulimulus (Lissoacme) exornatus; Pilsbry 1896 [1895–1896]: 171, pl. 50 fig. 55.
Type locality.

”Chilon, Bolivia” [Dept. Santa Cruz].

Label.

”Bolivia”. M.C. label style I, V.

Dimensions.

Not given; lectotype H 13.1, D —, W 5.5.

Type material.

NHMUK 1975331, lectotype (Cuming coll.).

Remarks.

The lectotype was damaged when it was returned from a loan in 1988; the remaining shell is too fragile to be figured herein. The lot consisted of a second specimen (1975332), which appears to be a juvenile Drymaeus species; this specimen is now excluded as type material.

Current systematic position.

Bulimulidae, Naesiotus exornatus (Reeve, 1849).

Drymaeus exoticus da Costa, 1901

http://species-id.net/wiki/Drymaeus_exoticus

Figs 33D–F, L21v
Drymaeus exoticus da Costa 1901: 239, pl. 24 fig. 10; Pilsbry 1901 [1901–1902]: 156, pl. 48 fig. 52; Breure 1979: 109; Linares and Vera 2012: 185.
Type locality.

”The hot country, Upper Magdalena River, Colombia”.

Label.

”hot country / Columbia”, taxon label in da Costa’s handwriting.

Dimensions.

”Long. 23.5, diam. 11 mm.”; figured specimen herein H 24.8, D 12.2, W 6.1.

Type material.

NHMUK 1907.11.21.38, lectotype (da Costa coll.).

Remarks.

This material was considered as holotype by Breure (1979). However, da Costa did not state on how many specimens his description was based. Breure (unpublished data) found another specimen in the Dautzenberg collection with a label in da Costa’s handwriting, but without enough evidence to consider it a type specimen. However, it cannot be excluded that da Costa had more than one specimen at hand; according to Art. 74.6 ICZN, the NHMUK specimen should be considered as a lectotype.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) exoticus (da Costa, 1901).

Bulimulus (Drymaeus) expatriatus Preston, 1909

http://species-id.net/wiki/Bulimulus_expatriatus

Figs 51A–C, L21vi
Bulimulus (Drymaeus) expatriatus Preston 1909: 510, pl. 10 fig. 4; Breure 1979: 109 (lectotype designation).
Drymaeus (Drymaeus) expatriatus; Breure and Eskens 1981: 21.
Type locality.

”E. Bolivia”.

Label.

”E. Bolivia”, taxon label in Preston’s handwriting.

Dimensions.

”Alt. 28, diam. maj. 11.5 mm”; figured specimen herein H 28.1, D 11.5, W 7.0.

Type material.

NHMUK 1975201, lectotype; 1915.1.6.43, one paralectotype (ex Preston).

Remarks.

The paralectotype is a juvenile shell.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) expatriatus (Preston, 1909).

Bulimus fabrefactus Reeve, 1848

http://species-id.net/wiki/Bulimus_fabrefactus

Figs 38D–F, L22i
Bulimus fabrefactus Reeve 1848 [1848–1850]: pl. 49 fig. 319; Breure 1979: 109 (lectotype designation).
Drymaeus fabrefactus; Pilsbry 1898 [1897–1898]: 260, pl. 40 fig. 5; Linares and Vera 2012: 185.
Drymaeus (Drymaeus) fabrefactus; Breure and Eskens 1981: 22, pl. 5 fig. 4.
Type locality.

[Venezuela] ”Province of Merida, New Granada”.

Label.

”New Granada”. M.C. label style I, V.

Dimensions.

Not given; figured specimen herein H 38.5, D 16.5, W 7.2.

Type material.

NHMUK 1975531, lectotype (Cuming coll.).

Remarks.

Linares and Vera (2012) have a number of specific localities for this taxon within Colombia.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) fabrefactus (Reeve, 1848).

Bulimus fallax Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_fallax

Figs 26D–F, L21vii
Bulimus fallax Pfeiffer 1853d: 375; Pfeiffer 1854 in Küster and Pfeiffer 1840–1865: 98, pl. 32 figs 5–6.
Drymaeus fallax; Pilsbry 1898 [1897–1898]: 239, pl. 33 figs 43–44.
Type locality.

[Ecuador] ”Tunguragua reipublicae Aequatoris (Bourcier)”.

Label.

”Tunguragua / Equador / Monsr Boissier [sic] / Consul General”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 24, diam. 12 mill.”; figured specimen herein H 22.4, D 13.8, W 5.7.

Type material.

NHMUK 1969142, lectotype and one paralectotype, Bourcier leg. (Cuming coll.).

Remarks.

Given the label, there is no doubt that this material was part of the type series. One specimen is now designated lectotype (design. n.) to define this poorly understood taxon.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) fallax (Pfeiffer, 1853).

Bulimus farrisi Pfeiffer, 1858

http://species-id.net/wiki/Bulimus_farrisi

Figs 45J–L, L22ii
Bulimus farrisi Pfeiffer 1858a: 258, pl. 42 fig. 8; Breure 1979: 109 (lectotype designation).
Drymaeus farrisi; Pilsbry 1898 [1897–1898]: 268, pl. 47 fig. 6.
Drymaeus (Drymaeus) farrisi; Breure and Eskens 1981: 22.
Type locality.

”Province of Patas, Andes of Peru (Dr. Farris)”.

Label.

”Province of Patas, / Andes of Peru / Dr. Farris”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 47, diam. 16 mill.”; figured specimen herein H 46.8, D 18.4, W 7.2.

Type material.

NHMUK 1975506, lectotype; 1975507, three paralectotypes, Farris leg. (Cuming coll.).

Remarks.

Breure and Eskens (1981) stated they had seen four paralectotypes. Lot NHMUK 1975509 seems to be missing.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) farrisi (Pfeiffer, 1858).

Bulimus felix Pfeiffer, 1862

http://species-id.net/wiki/Bulimus_felix

Figs 45D–F, L22iii
Bulimus felix Pfeiffer 1862: 387, pl. 37 fig. 2.
Drymaeus felix; Pilsbry 1898 [1897–1898]: 211, pl. 35 fig. 20; Linares and Vera 2012: 185 [partim].
Type locality.

”New Granada”.

Label.

”New Granada”, taxon label in Pfeiffer’s handwriting. M.C. label style III.

Dimensions.

”Long. 33, diam. 13 mill.”; lectotype H 34.0, D 14.8, W 6.2.

Type material.

NHMUK 1975206, lectotype and three paralectotypes (Cuming coll.).

Remarks.

Pfeiffer did not state on how many specimens his description was based. A lot with four specimens was found, which is considered as type material. The specimen marked with ‘x’ has been selected as lectotype (design. n.) to define this often confused taxon.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) felix (Pfeiffer, 1862).

Bulimus fenestratus Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_fenestratus

Figs 51J–L, L22iv
Bulimus fenestratus Pfeiffer 1846a: 29; Pfeiffer 1848b: 101; Reeve 1848 [1848–1850]: pl. 36 fig. 214; Breure 1979: 109 (lectotype designation).
Drymaeus (Drymaeus) fenestratus; Pilsbry 1899: 34, pl. 7 fig. 11.
Drymaeus (Drymaeus) fenestratus; Breure and Eskens 1981: 22, pl. 5 fig. 2; Thompson 2011: 110.
Type locality.

”Mexico”.

Label.

”Mexico”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.

Dimensions.

”Long. 45, diam. 18 mill.”; figured specimen herein H 44.0, D 20.3, W 6.6.

Type material.

NHMUK 1975525, lectotype (Cuming coll.).

Remarks.

The current systematic position follows Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) fenestratus (Pfeiffer, 1846).

Bulimus feriatus Reeve, 1848

http://species-id.net/wiki/Bulimus_feriatus

Figs 17I–K, L22vi
Bulimus feriatus Reeve 1848 [1848–1850]: pl. 54 fig. 354; Pfeiffer 1853d: 323; Breure 1979: 118 (lectotype designation).
Drymaeus feriatus; Pilsbry 1898 [1897–1898]: 203, pl. 34 fig. 10.
Drymaeus (Mesembrinus) feriatus; Breure and Eskens 1981: 26.
Type locality.

”Venezuela”.

Label.

No locality label. M.C. label style III, V.

Dimensions.

Not given; figured specimen herein H 29.4, D 12.1, W 6.0.

Type material.

NHMUK 1975204, lectotype; 1975205, two paralectotypes (Cuming coll.).

Remarks.

The current systematic position follows Richardson (1995: 126).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) feriatus (Reeve, 1848).

Naesiotus (Maranhoniellus) fernandezae Weyrauch, 1958

http://species-id.net/wiki/Naesiotus_fernandezae

Fig. 12H
Naesiotus (Maranhoniellus) fernandezae Weyrauch 1958: 122, pl. 9 figs 45–46; Breure 1979: 69; Neubert and Janssen 2004: 209, pl. 10 fig. 111; Breure 2012a: 8.
Type locality.

[Peru, Dept. Amazonas] ”Interandines Nord-Peru, auf rechter Seite des Río Marañon, bei Balsas, 850 m”.

Label.

”N-Peru: Balsas, Rio Marañon, 850 m, leg. B. Fernandez”, printed label.

Dimensions.

”H. 16.0 D. 3.4”; figured specimen herein H 15.1, D 3.3, W 8.1.

Type material.

NHMUK 19831, one paratype (ex Weyrauch).

Current systematic position.

Bulimulidae, Naesiotus fernandezae Weyrauch, 1958.

Bulimus ferrugineus Reeve, 1849

http://species-id.net/wiki/Bulimus_ferrugineus

Figs 9D, L22v
Bulimus ferrugineus Reeve 1849 [1848–1850]: pl. 62 fig. 424; Pfeiffer 1853d: 416; Breure 1979: 53.
Bulimulus ferrugineus; Pilsbry 1897 [1897–1898]: 29, pl. 9 fig. 37.
Bostryx ferrugineus; Breure 1978: 77 (lectotype designation).
Type locality.

”Peru”.

Label.

”Peru”. M.C. label style V.

Dimensions.

Not given; figured specimen herein H 19.0, D 10.7, W 5.4.

Type material.

NHMUK 1975380, lectotype; 1975381, two paralectotypes (Cuming coll.).

Remarks.

All type material is subadult; the two paralectypes are relatively bigger, with the largest having a shell height of 26.4 mm. The current systematic position follows Richardson (1995: 25).

Current systematic position.

Bulimulidae, Bostryx ferrugineus (Reeve, 1849).

Bulimus fidustus Reeve, 1849

http://species-id.net/wiki/Bulimus_fidustus

Figs 23G–H, L23i
Bulimus fidustus Reeve 1849 [1848–1850]: pl. 76 fig. 557; Breure 1979: 119 (lectotype designation).
Drymaeus fidustus; Pilsbry 1898 [1897–1898]: 308, pl. 50 fig. 95; Linares and Vera 2012: 194.
Drymaeus (Mesembrinus) fidustus; Breure and Eskens 1981: 69.
Type locality.

[Colombia, Dept. Putumayo / Ecuador, Prov. Napo] ”Sebundoi, New Granada”.

Label.

”Sebundoi / New Granada”. M.C. label style I, V.

Dimensions.

Not given; figured specimen herein H 22.5, D 11.4, W 6.8.

Type material.

NHMUK 1975517, lectotype; 1975518, one paralectotype (Cuming coll.).

Remarks.

The lip of the lectotype is slightly damaged.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) fidustus (Reeve, 1849).

Bulimus filaris Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_filaris

Figs 70C–D, 70L, L23ii
Bulimus filaris Pfeiffer 1853d: 653; Pfeiffer 1854e: 50; Breure 1979: 86.
Type locality.

Not given.

Label.

No locality given, taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long 26, diam. 12 mill.”; figured specimen herein H 25.1, D 12.3, W 5.6.

Type material.

NHMUK 1975569, two syntypes (Cuming coll.).

Remarks.

Pfeiffer did not state on how many specimens his description was based. On the label a note is added by a later hand ”same as one of / Morelet”; it is unclear to us to what taxon this refers. The current systematic position at the species level follows Richardson (1995).

Current systematic position.

Bulimulidae, Kuschelenia (Vermiculatus) filaris (Pfeiffer, 1853) (comb. n.).

Bulimus (Liostracus) flavidulus E.A. Smith, 1877

http://species-id.net/wiki/Bulimus_flavidulus

Figs 24A, L23iii
Bulimus (Liostracus) flavidulus E.A. Smith 1877: 364, pl. 39 fig. 3; Breure 1979: 119 (lectotype designation).
Drymaeus (Mesembrinus) flavidulus; Breure and Eskens 1981: 70; Breure and Borrero 2008: 24.
Type locality.

[Ecuador, Prov. El Oro] ”Zaruma, South Ecuador”.

Label.

”Zaruma, S. Ecuador”, taxon label in Smith’s handwriting.

Dimensions.

”Long. 21 mill, diam. 9”; figured specimen herein H 21.4, D 9.4, W 7.1.

Type material.

NHMUK 1975134, lectotype; 1877.3.28.5, two paralectotypes.

Remarks.

The lip of the lectotype is slightly damaged.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) flavidulus (E.A. Smith, 1877).

Bulimus flexilabris Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_flexilabris

Figs 49J–L, L23iv
Bulimus flexilabris Pfeiffer 1853d: 652; Pfeiffer 1854e: 50; Breure 1979: 109 (lectotype designation).
Drymaeus flexilabris; Simone 2006: 137, fig. 451.
Drymaeus (Drymaeus) bivittatus (Sowerby I, 1833); Breure and Eskens 1981: 10.
Type locality.

”in Brasilia”.

Label.

”Brazils”, taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 28, diam. 12 1/2 mill.”; figured specimen herein H 27.1, D 13.8, W 7.1.

Type material.

NHMUK 1975559, lectotype (Cuming coll.).

Remarks.

This taxon is a junior subjective synonym of Drymaeus bivittatus (Sowerby I, 1833), of which the type material has not been traced in the NHMUK collection.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) bivittatus (Sowerby I, 1833).

Bulimus flexuosus Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_flexuosus

Figs 34A–C, L23v
Bulimus flexuosus Pfeiffer 1853d: 329; Pfeiffer 1854b: 136; Pfeiffer 1854 in Küster and Pfeiffer 1840–1865: 244, pl. 65 figs 6–7; Breure 1979: 109 (lectotype designation).
Drymaeus flexuosus; Linares and Vera 2012: 186 [partim].
Drymaeus (Drymaeus) flexuosus; Breure and Eskens 1981: 26, pl. 5 fig. 3.
Type locality.

[Colombia, Dept. Caldas] ”Marinato [sic, Marmato] Novae Granadae”.

Label.

”Marinata / New Grenada”, taxon label in Pfeiffer’s handwriting. M.C. label style III.

Dimensions.

”Long. 40, diam. 14 mill.”; figured specimen herein H 43.0, D 20.0, W 7.2.

Type material.

NHMUK 1975202, lectotype; 1975203, two paralectotypes (Cuming coll.).

Remarks.

This taxon is restricted to the Central Cordillera in Colombia and has partly been confused with other Drymaeus species (Breure and Borrero, unpublished data).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) flexuosus (Pfeiffer, 1853).

Bulimus floridanus Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_floridanus

Figs 19H, L24i
Bulimus floridanus Pfeiffer 1857c: 330; Breure 1979: 119 (lectotype designation).
Drymaeus (Mesembrinus) dominicus (Reeve)[sic]; Breure and Eskens 1981: 55.
Type locality.

[U.S.A.] ”Florida”.

Label.

”Florida”, taxon label in Pfeiffer’s handwriting. M.C. label style III.

Dimensions.

”Long. 15 2/3–17, diam. 7 1/2 mill.”; figured specimen herein H 16.7, D 8.0, W 6.2.

Type material.

NHMUK 1975199, lectotype; 1975200, one paralectotype (Cuming coll.).

Remarks.

This taxon was considered by Richardson (1995: 190) as a junior subjective synonym of Drymaeus umbraticus (Reeve, 1850), of which the type material also is present in the NHMUK collection (see page 201); this classification is here followed.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) umbraticus (Reeve, 1850).

Bulimus fontainii d’Orbigny, 1838

http://species-id.net/wiki/Bulimus_fontainii

Figs 16H, L24ii
Bulimus fontainii d’Orbigny 1838 [1834–1847]: 273.
Bulimulus fontainii; Breure and Borrero 2008: 10.
Type locality.

[Ecuador, Prov. Guayas] ”environs de Guayaquil”.

Label.

”Guayaquil”, label in d’Orbigny’s handwriting.

Dimensions.

”Longueur totale, 13 millimètres; largeur 6 millimètres”; figured specimen herein H 13.2, D 6.4, W 8.5.

Type material.

NHMUK 1854.12.4.165, lectotype (d’Orbigny coll.).

Remarks.

d’Orbigny did not state on how many specimens his description was based. The single specimen found in his collection matches the dimensions given, and is here designated lectotype (design. n.). The weak axial sculpture of the protoconch makes us considering this taxon to belong to Naesiotus sensu Breure 1979, awaiting further studies of this group.

Current systematic position.

Bulimulidae, Naesiotus fontainii (d’Orbigny, 1838) (comb. n.).

Bulimus fourmiersi d’Orbigny, 1837

http://species-id.net/wiki/Bulimus_fourmiersi

Figs 16I, L24iii
Bulimus fourmiersi d’Orbigny 1837 [1834–1847]: 273, pl. 30 figs 12–14 [3 April 1837; text 23 April 1838].
Type locality.

[Argentina] ”la province de Corrientes, non loin du Rio de Santa-Lucia, au lieu nommé Pasto reito”; see Breure 1973: 133.

Label.

”corrientes Rep. argentina”, in d’Orbigny’s handwriting.

Dimensions.

”Long. 11 millim., lat. 7 millim.”; see remarks.

Type material.

NHMUK 1854.12.4.210, lectotype (d’Orbigny coll.).

Remarks.

The single specimen is severely damaged, and only the upper whorls are retained. The original figures of d’Orbigny, which are heirein designated as lectotype (design. n.), are figured instead of the type specimen. The protoconch scultpture suggests this taxon to belong to Naesiotus; more detailed studies are needed to corraborate this classification. According to the collation data the figures were published in 1837, the text in 1838 (Coan et al. 2013a). Miquel (1989a: 2) considered it as ”sp. inquirenda”; Cuezzo et al. (2013: 151) classified it with Bulimulus Leach, 1814.

Current systematic position.

Bulimulidae, Naesiotus fourmiersi (d’Orbigny, 1837).

Bulimus fucatus Reeve, 1849

http://species-id.net/wiki/Bulimus_fucatus

Figs 45G–I, L24v
Bulimus fucatus Reeve 1849 [1848–1850]: pl. 83 fig. 615; Breure 1979: 109 (lectotype designation).
Drymaeus fucatus; Pilsbry 1898 [1897–1898]: 234, pl. 42 fig. 63; Breure and Borrero 2008: 21; Linares and Vera 2012: 187.
Type locality.

[Colombia] ”Sebundoi, New Granada”.

Label.

”New Granada”.

Dimensions.

Not given; figured specimen herein H 23.5, D 11.1, W 6.2.

Type material.

NHMUK 1874.12.11.224, lectotype, ex Mrs. T. Lombe-Taylor.

Remarks.

Reeve did not state on how many specimens his description was based. The distribution of this species in Colombia needs further evidence (Breure and Borrero, unpublished data).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) fucatus (Reeve, 1849).

Bulimus (Liostracus) fuscobasis E.A. Smith, 1877

http://species-id.net/wiki/Bulimus_fuscobasis

Figs 24B, L24vi
Bulimus (Liostracus) fuscobasis E.A. Smith 1877: 365, pl. 39 fig. 6; Breure 1979: 119 (lectotype designation).
Drymaeus (Mesembrinus) fuscobasis (E.A. Smith); Breure and Eskens 1981: 70.
Type locality.

”Tarapoto, Andes of Peru, Mr. Spruce (Mus. Cuming)”.

Label.

”Tarapoto, Andes of Peru / Mr. Spruce”; taxon label in Smith’s handwriting. M.C. label style III.

Dimensions.

”Longit. 29 mill., diam. 12”; figured specimen herein H 28.7, D 12.3, W 8.2.

Type material.

NHMUK 1975139, lectotype; 1975140, one paralectotype, Spruce leg. (Cuming coll.).

Remarks.

The specimen selected lectotype by Breure (1979) is marked with a red dot. The specimens were accompanied by a label ”holotype”; however, the original label clearly states ”types” and it is not clear whom has attached the (subsequent?) label ”holotype”. Smith described the species from ”two specimens”.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) fuscobasis (E.A. Smith, 1877).

Helix fusoides d’Orbigny, 1835

http://species-id.net/wiki/Helix_fusoides

Figs 29D–F, L24iv
Helix fusoides d’Orbigny 1835: 19.
Bulimus fusoides d’Orbigny 1837 [1834–1847]: 315, pl. 40 figs 12–13 [18 Sept. 1837; text 6 May 1838].
Drymaeus fusoides; Pilsbry 1898 [1897–1898]: 201, pl. 38 figs 17–18.
Type locality.

”provincia Yungacensi (republica Boliviana)”

Label.

”Yunga..... olivia”; in d’Orbigny’s handwriting.

Dimensions.

”Longit. 4 centim., latit. 1 centim. 4 millim.” (corrected to ”Long. 40 millim., lat. 15 millim.” in d’Orbigny 1837); figured specimen herein H 24.6, D —, W 5.4.

Type material.

NHMUK 1854.12.4.133, one paralectotype.

Remarks.

d’Orbigny did not state on how many specimens his description was based. The type locality was specified in d’Orbigny 1838 [1834-1847]: 316 as ”près du lieu dit Yunga de la Palma” (see Breure 1973: 117, 132). Breure (1975: 1151) has selected a lectotype from the material found in MNHN. The specimen in London has lost the lower part of the last whorl and moreover does not correspond to d’Orbigny 1837 [1834-1847]: pl. 40 figs 12-13. The current systematic position follows Richardson (1995: 128).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) fusoides (d’Orbigny, 1835).

Bulimus gabbi Angas, 1879

http://species-id.net/wiki/Bulimus_gabbi

Figs 24L, L24vii
Bulimus gabbi Angas 1879: 477, pl. 40 fig. 3; Breure 1979: 119 (lectotype designation).
Drymaeus gabbi; Pilsbry 1899: 70, pl. 6 figs 7–9, 11.
Drymaeus (Mesembrinus) gabbi; Thompson 2011: 117.
Type locality.

[Costa Rica, Prov. Limón, Cerro Kámuk] ”flanks of Pico Blanco, at an altitude from 3000 to 6000 feet [914–1828 m]”.

Label.

”flanks / of Pico Blanco / 3–6000 ft / Costa Rica”.

Dimensions.

”Diam. 7, alt. 10 1/2 lin. [H 22.2, D 14.8 mm]”; figured specimen herein H 20.6, D 12.06, W 4.7.

Type material.

NHMUK 1879.7.22.23, lectotype; 1879.7.22.24–26, three paralectotypes, Gabb leg. (ex Angas).

Remarks.

Angas did not state on how many specimens his description was based; he distinguished three varieties, which differ in their colour pattern. The lectotype corresponds to Angas’ right-hand sided figure. The current systematic position follows Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) gabbi (Angas, 1879).

Bulimus galapaganus Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_galapaganus

Figs 14H, L24viii
Bulimus galapaganus Pfeiffer 1855a: 58; Pfeiffer 1859b: 503; Breure 1979: 69 (lectotype designation).
Type locality.

[Ecuador] ”Galapagos Islands”.

Label.

”Galapagos Islds”, taxon label in Pfeiffer’s handwriting. M.C. label style III.

Dimensions.

”Long. 15 1/2, diam. 6 mill.”; figured specimen herein H 15.1, D 6.28, W 7.6.

Type material.

NHMUK 1975146, lectotype; 1975147, one paralectotype (Cuming coll.).

Remarks.

The current systematic position follows Richardson (1995: 220).

Current systematic position.

Bulimulidae, Naesiotus galapaganus (Pfeiffer, 1855).

Bulimus gayi Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_gayi

Figs 67E–F, 67M, L25i
Bulimus gayi Pfeiffer 1857d: 389; Breure 1979: 88.
Scutalus (Kuschelenia) gayi; Breure 1978: 177 (lectotype designation).
Type locality.

”Bolivia”.

Label.

”Bolivia”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 27, diam. 16 mill.”; figured specimen herein H 27.7, D 16.2, W 5.8.

Type material.

NHMUK 1975382, lectotype; 1975383, one paralectotype (Cuming coll.).

Remarks.

This taxon is now transferred to the genus Kuschelenia Hylton Scott, 1952.

Current systematic position.

Bulimulidae, Kuschelenia (Kuschelenia) gayi (Pfeiffer, 1857) (comb. n.).

Bulimus (Mesembrinus) gealei H. Adams, 1867

http://species-id.net/wiki/Bulimus_gealei

Figs 51G–I, L25iii
Bulimus (Mesembrinus) gealei H. Adams 1867b: 309, pl. 19 fig. 21.
Type locality.

”Mexico. Collected by Mr. Boucard”.

Label.

”Mexico”.

Dimensions.

”Long. 29, diam. 14 mill.”; figured specimen herein H 26.4, D 13.4, W 6+.

Type material.

NHMUK 1867.1.9.20, lectotype and two paralectotypes; 1867.1.9.21, three paralectotypes, ex Geale.

Remarks.

One of the specimens corresponds to Adams 1867b: pl. 19 fig. 21 and is here selected lectotype (design. n.) to fixate this taxon; it has the top whorls slightly damaged. According to the register the specimens were purchased from Mr R. Geale; the label does not mention the name Boucard. Köhler (2007: 145, fig. 90) selected a lectotype (ZMB 4457a) for the senior subjective synonym Bulimulus (Scutalus) fenestrellus Martens, 1864.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) fenestrellus (Martens, 1864).

Bulimus gelidus Reeve, 1849

http://species-id.net/wiki/Bulimus_gelidus

Figs 62C–D, L25ii
Bulimus gelidus Reeve 1849 [1848–1850]: pl. 76 fig. 555; Breure 1978: 142; Breure 1979: 62 (lectotype designation); Miquel 1991: 98, fig. 25.
Bulimulus gelidus; Breure 2008: 245, figs 6–7 [not 8]; see remarks.
Type locality.

”Central America”.

Label.

”C. America”. M.C. label style I, V.

Dimensions.

Not given; figured specimen herein H 32.5, D 15.0, W 7.7.

Type material.

NHMUK 1975402, lectotype (Cuming coll.).

Remarks.

On the back of the cardboard a handwriting in pencil has noted ”seems to be the same as monte-vidensis Pfr.”. Miquel (1991) placed this taxon is the synonymy of Syphalomphix bonariensis Rafinesque, 1833, a species from Argentina. However, it must be noted that most Bulimulus species are very similar and often hard to tell apart. Reconsidering the identification made in Breure (2008), we now disagree with him, as the Ecuadorian shells have different dimensions, and the last whorl is less globose than in the lectotype of Bulimus gelidus. The locality on the label may be erroneous, and Reeve’s taxon is herein considered as nomen inquirendum.

Current systematic position.

Bulimulidae, Bulimulus. Nomen inquirendum.

Bulimus geometricus Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_geometricus

Figs 46J–L, L25iv
Bulimus geometricus Pfeiffer 1846c: 84; Pfeiffer 1848b: 59; Reeve 1848 [1848–1850]: pl. 44 fig. 278; Breure 1979: 109 (lectotype designation).
Drymaeus geometricus; Linares and Vera 2012: 187.
Drymaeus (Drymaeus) geometricus; Breure and Eskens 1981: 27, pl. 8 fig. 2.
Type locality.

[Colombia] ”Vallis Magdalenae Novae Granadae”.

Label.

”Valley of the Madelein / New Granada”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.

Dimensions.

”Long. 35, diam. 13 mill.”; figured specimen herein H 34.6, D 18.0, W 6.9.

Type material.

NHMUK 1975564, lectotype; 1975565, two paralectotypes (Cuming coll.).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) geometricus (Pfeiffer, 1846).

Bostryx (Bostryx) zilchi glomeratus Weyrauch, 1960

http://species-id.net/wiki/Bostryx_zilchi_glomeratus

Fig. 3I
Bostryx (Bostryx) zilchi glomeratus Weyrauch 1960b: 124, pl. 12 figs 29–34; Neubert and Janssen 2004: 210, pl. 3 fig. 35; Breure 2012a: 8.
Type locality.

”Mittel-Peru: Quichao, 5 km von Laraos, am Fusspfad nach Yauyos, im Tale des Río Cañete und auf der linken Seite des Río Mayo, 3500 m”.

Label.

”C-Peru, Quichao, 5 kms. from Laraos, valley of Río Cañete, 3500 m”; printed label.

Dimensions.

”H. 19.5, D. 6.5”; figured specimen herein H 18.6, D 6.1, W 7.7.

Type material.

NHMUK 1975154, five paratypes, ex Weyrauch (WW 3320).

Current systematic position.

Bulimulidae, Bostryx zilchi Weyrauch, 1958.

Scutalus grandiventris Weyrauch, 1960

http://species-id.net/wiki/Scutalus_grandiventris

Fig. 65H
Scutalus grandiventris Weyrauch, 1960a: 42, pl. 5 figs 27–33; Neubert and Janssen 2004: 211, pl. 12 fig. 140; Breure 2012a: 8.
Type locality.

”N-Peru, am Westhang der westlichen Anden, oberhalb Cascas ± 100 km nö. Trujillo und Cajamarca (1400 m)”.

Label.

”N-Peru: Cascas, 1400 m”; printed label.

Dimensions.

”H. 53.0, D. 29.7”; figured specimen herein H 54.2 D 29.1 W 7.0.

Type material.

NHMUK 1975386, two paratypes, ex Weyrauch (WW 1364).

Current systematic position.

Bulimulidae, Scutalus grandiventris Weyrauch, 1960.

Bulimus gruneri Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_gruneri

Figs 24E, L25vii
Bulimus gruneri Pfeiffer 1846a: 30; Pfeiffer 1848b: 213; Reeve 1849 [1848–1850]: pl. 51 fig. 332.
Type locality.

”Mexico”.

Label.

”Mexico”, added in a later handwriting. M.C. label style I, V.

Dimensions.

”Long. 28, diam. 10 mill.”; figured specimen herein H 27.8, D 10.5, W 7.3.

Type material.

NHMUK 20100563/1, lectotype; NHMUK 20100563/2–3, two paralectotypes; NHMUK 20100562, three paralectotypes (Cuming coll.).

Remarks.

The quotation by Richardson (1995: 196) of 1842 as the year of publication of Bulimus gruneri appears erroneous, as Pfeiffer does not mention this taxon until his 1846 paper. Two lots have been traced, one of which contains a specimen closest matching the original dimensions; this specimen is here designated lectotype (design. n.) to define the taxon. The other lot contains one specimen resembling Reeve 1848 [1848–1850]: pl. 51 fig. 332. The locality ”Mexico” seems in error, as this taxon has been recognised as a Colombian species (Breure and Borrero, unpublished data) and is now considered as a junior subjective synonym of Bulimus columbianus Lea, 1838 (syn. n.).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) columbianus (Lea, 1838).

Bulimus gueinzii Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_gueinzii

Figs 30M–O, L25vi
Bulimus gueinzii Pfeiffer 1857c: 330; Breure 1979: 109 (lectotype designation).
Drymaeus (Drymaeus) gueinzii; Breure and Eskens 1981: 27, pl. 6 fig. 8.
Type locality.

”Meobamba, Peru (Mr Gueinzius)”.

Label.

”Meobamba”, taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

”Long. 23, diam. 10 mill.”; figured specimen herein H 23.2, D 10.9, W 6.0.

Type material.

NHMUK 1975539, lectotype, Gueinzius leg. (Cuming coll.).

Remarks.

The current systematic position follows Richardson (1995: 134).

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) gueinzii (Pfeiffer, 1857).

Bulinus guttatus Broderip, 1832

http://species-id.net/wiki/Bulinus_guttatus

Figs 7E, L26i
Bulinus guttatus Broderip in Broderip and Sowerby I 1832a: 31.
Bulimus guttatus; Reeve 1848 [1848–1850]: pl. 22 fig. 144.
Type locality.

[Chile] ”Peruviâ. (Cobija or Puerto De la Mar)”.

Label.

”Bolivia, S. America”, added in a later handwriting. M.C. label style III, V.

Dimensions.

”long. 7/8, lat. 3/8 poll. [H 22.1, D 9.5 mm]”; figured specimen herein H 22.0, D —, W 6.2.

Type material.

NHMUK 20110175, five possible syntypes (Cuming coll.).

Remarks.

The type locality is now part of Chile, but during the 19th century has also been part of Bolivia and Peru. This taxon has been treated as nomen inquirendum by Breure (1979). Of the five specimens, two are subadult or juvenile. One specimen corresponds to Reeve 1848 [1848–1850]: pl. 22 fig. 144, and is damaged both at the lip and on the dorsal side of the last whorl.

Current systematic position.

Bulimulidae, Bostryx guttatus (Broderip in Broderip and Sowerby I 1832).

Bulimus hachensis Reeve, 1850

http://species-id.net/wiki/Bulimus_hachensis

Figs 19K–L, L26ii
Bulimus hachensis Reeve 1850 [1848–1850]: pl. 85 fig. 627; Pfeiffer 1853d: 421; Breure 1979: 119 (lectotype designation).
Drymaeus hachensis; Pilsbry 1899: 90, pl. 12 fig. 20; Linares and Vera 2012: 195.
Drymaeus (Mesembrinus) hachensis; Breure and Eskens 1981: 71.
Type locality.

”Guatemala, banks of the Rio Hacha”.

Label.

”Rio Hacha, Guatemala”. M.C. label style I, V.

Dimensions.

Not given; figured specimen herein H 26.4, D 11.3, W 7.7.

Type material.

NHMUK 1975392, lectotype (Cuming coll.).

Remarks.

Reeve did not state on how many specimens his description was based. We consider this taxon to be a junior subjective synonym of Bulimus columbianus Lea, 1838 (syn. n.), which is partly based on unpublished data of Breure and Borrero.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) columbianus (Lea, 1838).

Bulimus hamiltoni Reeve, 1849

http://species-id.net/wiki/Bulimus_hamiltoni

Figs 1F, L26iii
Bulimus hamiltoni Reeve 1849 [1848–1850]: pl. 83 fig. 610; Pfeiffer 1853d: 429.
Bulimulus (Peronaeus) hamiltoni; Pilsbry 1896 [1895–1896]: 149, pl. 46 fig. 51.
Bostryx hamiltoni; Breure 1978: 80 (lectotype designation).
Type locality.

”Near the Lake of Titicaca, Bolivia”.

Label.

”Bolivia”. A second label indicates ”appartient [?] à Mr Pentlan[d]” [belongs to Mr. Pentland] and refers to the figure published in Reeve 1848–1850.

Dimensions.

Not given; figured specimen herein H 17.0, D 6.55, W 8.0.

Type material.

NHMUK 1849.5.14.53, lectotype; 1849.5.14.54–57, four paralectotypes.

Remarks.

According to Reeve the material was collected by Mr. Pentland and belonged to ”Mus. Hamilton”; the lectotype corresponds to Reeve 1849 [1848–1850]: pl. 83 fig. 610. Breure (1978) erroneously mentioned the lectotype as 1975329 and the paralectotypes as 1975330. The current systematic position follows Richardson (1995: 26).

Current systematic position.

Bulimulidae, Bostryx hamiltoni (Reeve, 1849).

Bulimus haplochrous Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_haplochrous

Figs 61G–H, L26iv
Bulimus haplochrous Pfeiffer 1855g: 125; Pfeiffer 1859b: 502; Breure 1979: 63.
Bulimulus haplochrous; Breure 1978: 144 (lectotype designation).
Type locality.

”—?”.

Label.

”New Granada”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 30, diam. 12 mill.”; figured specimen herein H 28.6, D 13.3, W 5.3.

Type material.

NHMUK 1975405, lectotype; 1975406, two paralectotypes (Cuming coll.).

Remarks.

Pfeiffer did not state on how many specimens his description was based. The current systematic position follows Richardson (1995: 76).

Current systematic position.

Bulimulidae, Bulimulus haplochrous (Pfeiffer, 1855).

Helix heloica d’Orbigny, 1835

http://species-id.net/wiki/Helix_heloica

Figs 62E–F, L26v
Helix heloica d’Orbigny 1835: 11.
Bulimus heloicus; d’Orbigny 1837 [1834–1847]: 272, pl. 30 figs 9–11 [3 April 1837; text 23 April 1838].
Bulimulus (Lissoacme) heloicus; Pilsbry 1896 [1895–1896]: 193, pl. 51 figs 12–13.
Type locality.

[Bolivia] ”provincia Chiquitensi, republica Boliviana”.

Label.

”Sn Xavier de Chiquitos Bolivia”, label in d’Orbigny’s handwriting.

Dimensions.

”Longit. 28 millim., diam. 6 millim.”; figured specimen herein H 25.6, D 12.0, W 6.8.

Type material.

NHMUK 1854.12.4.164, five syntypes (d’Orbigny coll.).

Remarks.

d’Orbigny (1835) did not state on how many specimens his description was based; the type locality was specified in d’Orbigny 1838 [1834–1847]: 273 as ”près de la Mission de Bibosi, province de Santa-Cruz de la Sierra, et dans la partie orientaler de l’immense fôret (Monte grande) qui sépare Santa-Cruz de la province de Chiquitos, non loin du lieu nommé Potrero de la Cruz”. The locality on the label probably refers to San Javier, Dept. Santa Cruz in Bolivia; see Breure 1973: 119, 127. The protoconch sculpture suggests that this taxon belongs to Bulimulus Leach, 1814.

Current systematic position.

Bulimulidae, Bulimulus heloicus (d’Orbigny, 1835) (comb. n.).

Bulimus hepatostomus Pfeiffer, 1861

http://species-id.net/wiki/Bulimus_hepatostomus

Figs 17D–E, L26vii
Bulimus hepatostomus Pfeiffer 1861: 23, pl. 3 fig. 4; Breure 1979: 110 (lectotype designation).
Drymaeus (Drymaeus) hepatostomus; Breure and Eskens 1981: 28, pl. 8 fig. 10.
Drymaeus (Mesembrinus) hepatostomus; Thompson 2011: 115.
Type locality.

”Mexico (Mr Boucard)”.

Label.

”Tepenistlahuaca / Mexico. Mr Boucard”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

”Long. 32, diam. 13 mill.”; figured specimen herein H 30.1, D 15.0, W 6.1.

Type material.

NHMUK 1975571, lectotype; 1975221, one paralectotype, Boucard leg. (Cuming coll.).

Remarks.

The current systematic position is according to Thompson (2011).

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) hepatostomus (Pfeiffer, 1861).

Bulimulus (Drymaeus) hidalgoi da Costa, 1898

http://species-id.net/wiki/Bulimulus_hidalgoi

Figs 32A–C, L26vi
Bulimulus (Drymaeus) hidalgoi da Costa, 1898: 80, pl. 6 fig. 2; Breure 1979: 110.
Drymaeus (Drymaeus) hidalgoi; Breure and Borrero 2008: 21.
Type locality.

”Ecuador”.

Label.

”Equador”, label in da Costa’s handwriting.

Dimensions.

”Long. 39, diam. 13 mm.”; figured specimen herein H 39.0, D 17.0, W 6.6.

Type material.

NHMUK 1907.11.21.28, lectotype; 1907.11.21.29–30, two paralectotypes, Buckley leg. (da Costa coll.).

Remarks.

da Costa did not state on how many specimens his description was based, only that he had ”examples collected by the late Mr. Buckley”. The reference of Breure (1979) to ”HT 1907.11.21.28” has to be interpreted as a lectotype designation under Art. 74.6 ICZN.

Current systematic position.

Bulimulidae, Drymaeus (Drymaeus) hidalgoi (da Costa, 1898).

Bulimus holostoma Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_holostoma

Figs 2O–P, L26viii
Bulimus holostoma Pfeiffer 1846a: 28; Pfeiffer 1848b: 161; Reeve 1848 [1848–1850]: pl. 69 fig. 490; Breure 1979: 54 (lectotype designation).
Type locality.

[Chile] ”Cobija, Bolivia”.

Label.

”Cobija Bolivia Hills under Bushes”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.

Dimensions.

”Long. 9, diam. 2 2/3 mill.”; figured specimen herein H 7.84, D 2.43, W 5.9.

Type material.

NHMUK 1975345, lectotype; 1975346, one paralectotype (Cuming coll.).

Remarks.

Pfeiffer described this species from the Cuming collection, but did not state on how many specimens his description was based. The current systematic position follows Richardson (1995: 27).

Current systematic position.

Bulimulidae, Bostryx holostoma (Pfeiffer, 1846).

Bulimus hondurasanus Pfeiffer, 1846

http://species-id.net/wiki/Bulimus_hondurasanus

Figs 24I, L27i
Bulimus hondurasanus Pfeiffer 1846a: 29; Reeve 1849 [1848–1850]: pl. 59 fig. 400; Breure 1979: 119 (lectotype designation).
Drymaeus hondarusanus; Rehder 1966: 286, figs 5–6.
Drymaeus (Mesembrinus) hondurasanus; Thompson (2011: 121).
Type locality.

”Honduras”.

Label.

”Honduras on leaves / of Bushes Dyson”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.

Dimensions.

”Long. 18 1/2, diam. 10 mill.”; figured specimen herein H 28.6, D 13.7, W 6.6.

Type material.

NHMUK 1975265, lectotype; 1975117, one paralectotype, Dyson leg. (Cuming coll.).

Remarks.

Pfeiffer (1846a: 30) mentioned that his material originated from Dyson, which is confirmed by the information on the label. Therefore there is no doubt about the type status of the specimens. However, Pfeiffer’s dimensions (H/D ration 1.85) do not fit the specimens found in the NHMUK collection (H/D ratio 2.08). The figure as presented by Reeve 1849 [1848–1850] confirms the latter ratio, which makes it likely that Pfeiffer’s measurements are either erroneous or that he had more material in his own collection. As the Pfeiffer collection must considered to be lost (Dance 1966), the London specimens are the sole reference. We consider it probable that the published dimensions of Pfeiffer are in error.

Current systematic position.

Bulimulidae, Drymaeus (Mesembrinus) hondurasanus (Pfeiffer, 1846).

Bulimus