Catalogue |
Corresponding author: Marcin J. Kamiński ( kaminskientomo@gmail.com ) Academic editor: Wolfgang Schawaller
© 2019 Marcin J. Kamiński, Kojun Kanda, Ryan Lumen, Jonah M. Ulmer, Christopher C. Wirth, Patrice Bouchard, Rolf Aalbu, Noël Mal, Aaron D. Smith.
This is an open access article distributed under the terms of the CC0 Public Domain Dedication.
Citation:
Kamiński MJ, Kanda K, Lumen R, Ulmer JM, Wirth CC, Bouchard P, Aalbu R, Mal N, Smith AD (2019) A catalogue of the tribe Sepidiini Eschscholtz, 1829 (Tenebrionidae, Pimeliinae) of the world. ZooKeys 844: 1-121. https://doi.org/10.3897/zookeys.844.34241
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This catalogue includes all valid family-group (six subtribes), genus-group (55 genera, 33 subgenera), and species-group names (1009 species and subspecies) of Sepidiini darkling beetles (Coleoptera: Tenebrionidae: Pimeliinae), and their available synonyms. For each name, the author, year, and page number of the description are provided, with additional information (e.g., type species for genus-group names, author of synonymies for invalid taxa, notes) depending on the taxon rank. Verified distributional records (loci typici and data acquired from revisionary publications) for all the species are gathered. Distribution of the subtribes is illustrated and discussed.
Several new nomenclatural acts are included. The generic names Phanerotomea Koch, 1958 [= Ocnodes Fåhraeus, 1870] and Parmularia Koch, 1955 [= Psammodes Kirby, 1819] are new synonyms (valid names in square brackets).
The following new combinations are proposed: Ocnodes acuductus acuductus (Ancey, 1883), O. acuductus ufipanus (Koch, 1952), O. adamantinus (Koch, 1952), O. argenteofasciatus (Koch, 1953), O. arnoldi arnoldi (Koch, 1952), O. arnoldi sabianus (Koch, 1952), O. barbosai (Koch, 1952), O. basilewskyi (Koch, 1952), O. bellmarleyi (Koch, 1952), O. benguelensis (Koch, 1952), O. bertolonii (Guérin-Méneville, 1844), O. blandus (Koch, 1952), O. brevicornis (Haag-Rutenberg, 1875), O. brunnescens brunnescens (Haag-Rutenberg, 1871), O. brunnescens molestus (Haag-Rutenberg, 1875), O. buccinator (Koch, 1952), O. bushmanicus (Koch, 1952), O. carbonarius (Gerstaecker, 1854), O. cardiopterus (Fairmaire, 1888), O. cataractus (Koch, 1952), O. cinerarius (Koch, 1952), O. complanatus (Koch, 1952), O. confertus (Koch, 1952), O. congruens (Péringuey, 1899), O. cordiventris (Haag-Rutenberg, 1871), O. crocodilinus (Koch, 1952), O. dimorphus (Koch, 1952), O. distinctus (Haag-Rutenberg, 1871), O. dolosus (Péringuey, 1899), O. dorsocostatus (Gebien, 1910), O. dubiosus (Péringuey, 1899), O. ejectus (Koch, 1952), O. epronoticus (Koch, 1952), O. erichsoni (Haag-Rutenberg, 1871), O. ferreirae ferreirae (Koch, 1952), O. ferreirae zulu (Koch, 1952), O. fettingi (Haag-Rutenberg, 1875), O. fistucans (Koch, 1952), O. fraternus (Haag-Rutenberg, 1875), O. freyi (Koch, 1952), O. freudei (Koch, 1952), O. fulgidus (Koch, 1952), O. funestus (Haag-Rutenberg, 1871), O. gemmeulus (Koch, 1952), O. gibberosulus (Péringuey, 1908), O. gibbus (Haag-Rutenberg, 1879), O. globosus (Haag-Rutenberg, 1871), O. granisterna (Koch, 1952), O. granulosicollis (Haag-Rutenberg, 1871), O. gridellii (Koch, 1960), O. guerini guerini (Haag-Rutenberg, 1871), O. guerini lawrencii (Koch, 1954), O. guerini mancus (Koch 1954), O. haemorrhoidalis haemorrhoidalis (Koch, 1952), O. haemorrhoidalis salubris (Koch, 1952), O. heydeni (Haag-Rutenberg, 1871), O. humeralis (Haag-Rutenberg, 1871), O. humerangula (Koch, 1952), O. imbricatus (Koch, 1952), O. imitator imitator (Péringuey, 1899), O. imitator invadens (Koch, 1952), O. inflatus (Koch, 1952), O. janssensi (Koch, 1952), O. javeti (Haag-Rutenberg, 1871), O. junodi (Péringuey, 1899), O. kulzeri (Koch, 1952), O. lacustris (Koch, 1952), O. laevigatus (Olivier, 1795), O. lanceolatus (Koch, 1953), O. licitus (Peringey, 1899), O. luctuosus (Haag-Rutenberg, 1871), O. luxurosus (Koch, 1952), O. maputoensis (Koch, 1952), O. marginicollis (Koch, 1952), O. martinsi (Koch, 1952), O. melleus (Koch, 1952), O. mendicus estermanni (Koch, 1952), O. mendicus mendicus (Péringuey, 1899), O. miles (Péringuey, 1908), O. mimeticus (Koch, 1952), O. misolampoides (Fairmaire, 1888), O. mixtus (Haag-Rutenberg, 1871), O. monacha (Koch, 1952), O. montanus (Koch, 1952), O. mozambicus (Koch, 1952), O. muliebris curtus (Koch, 1952), O. muliebris muliebris (Koch, 1952), O. muliebris silvestris (Koch, 1952), O. nervosus (Haag-Rutenberg, 1871), O. notatum (Thunberg, 1787), O. notaticollis (Koch, 1952), O. odorans (Koch, 1952), O. opacus (Solier, 1843), O. osbecki (Billberg, 1815), O. overlaeti (Koch, 1952), O. ovulus (Haag-Rutenberg, 1871), O. pachysoma ornata (Koch, 1952), O. pachysoma pachysoma (Péringuey, 1892), O. papillosus (Koch, 1952), O. pedator (Fairmaire, 1888), O. perlucidus (Koch, 1952), O. planus (Koch, 1952), O. pretorianus (Koch, 1952), O. procursus (Péringuey, 1899), O. protectus (Koch, 1952), O. punctatissimus (Koch, 1952), O. puncticollis (Koch, 1952), O. punctipennis planisculptus (Koch, 1952), O. punctipennis punctipennis (Harold, 1878), O. punctipleura (Koch, 1952), O. rhodesianus (Koch, 1952), O. roriferus (Koch, 1952), O. rufipes (Harold, 1878), O. saltuarius (Koch, 1952), O. scabricollis (Gerstaecker, 1854), O. scopulipes (Koch, 1952), O. scrobicollis griqua (Koch, 1952), O. scrobicollis simulans (Koch, 1952), O. semirasus (Koch, 1952), O. semiscabrum (Haag-Rutenberg, 1871), O. sericicollis (Koch, 1952), O. similis (Péringuey, 1899), O. sjoestedti (Gebien, 1910), O. spatulipes (Koch, 1952), O. specularis (Péringuey, 1899), O. spinigerus (Koch, 1952), O. stevensoni (Koch, 1952), O. tarsocnoides (Koch, 1952), O. temulentus (Koch, 1952), O. tenebrosus melanarius (Haag-Rutenberg, 1871), O. tenebrosus tenebrosus (Erichson, 1843), O. tibialis (Haag-Rutenberg, 1871), O. torosus (Koch, 1952), O. transversicollis (Haag-Rutenberg, 1879), O. tumidus (Haag-Rutenberg, 1871), O. umvumanus (Koch, 1952), O. vagus (Péringuey, 1899), O. vaticinus (Péringuey, 1899), O. verecundus (Péringuey, 1899), O. vetustus (Koch, 1952), O. vexator (Péringuey, 1899), O. virago (Koch, 1952), O. warmeloi (Koch, 1953), O. zanzibaricus (Haag-Rutenberg, 1875), Psammophanes antinorii (Gridelli, 1939), and P. mirei (Pierre, 1979).
The type species [placed in square brackets] of the following genus-group taxa are designated for the first time, Ocnodes Fåhraeus, 1870 [Ocnodes scrobicollis Fåhraeus, 1870], Psammodophysis Péringuey, 1899 [Psammodophysis probes Péringuey, 1899], and Trachynotidus Péringuey, 1899 [Psammodes thoreyi Haag-Rutenberg, 1871].
A lectotype is designated for Histrionotus omercooperi Koch, 1955 in order to fix its taxonomic status. Ulamus Kamiński is introduced here as a replacement name for Echinotus Marwick, 1935 [Type species. Avicula echinata Smith, 1817] (Mollusca: Pteriidae) to avoid homonymy with Echinotus Solier, 1843 (Coleoptera: Tenebrionidae).
Africa, distribution, Molurini, new synonyms, new combinations, nomen novum, nomenclature, type species
The Sepidiini Eschscholtz, 1829 are a diverse tribe of ground-dwelling darkling beetles (Tenebrionidae) of the subfamily Pimeliinae Latreille, 1802 (Figs
Morphology of representative species of the different subtribes of Sepidiini 1 Arturium dallastai (Molurina) 2 Psammophanes jockli (Molurina) 3 Phrynocolus dentatus (Molurina) 4 Euphrynus sexdentatus (Molurina) 5 Psammophanes raffrayi (Molurina) 6 Distretus mormocyle (Molurina) 7 Psammophanes poccilator (Molurina) 8 Euphrynus sexdentatus (Molurina) 9 Moluris nitida (Molurina) 10 Dichtha inflata (Molurina) 11 Psammorhyssus titanus (Phanerotomeina) 12 Dichtha cubica (Molurina) 13 Brachyphrynus petrosus (Molurina) 14 Echinotus spinicollis (Sepidiina) 15 Psammodes attenuata (Phanerotomeina) 16 Vieta muscosa (Sepidiina) 17 Sepidium bidentatum (Sepidiina) 18 Psammodes ponderosus (Phanerotomeina) 19 Sepidium cristatum (Sepidiina) 20 Ocnodes scabricollis (Phanerotomeina) 21 Sepidium hystryx (Sepidiina) 22 Vieta spiculosa (Sepidiina) 23 Amiantus picteti (Molurina) 24 Sepidiostenus compressus (Sepidiina) 25 Psammodes vialis (Phanerotomeina) 26 Dimoniacis jacksoni (Sepidiina) 27 Physophrynus haroldi (Molurina) 28 Tarsocnodes molossa (Phanerotomeina) 29 Tarsocnodes nielseni (Phanerotomeina) 30 Physophrynus bredoi (Molurina) 31 Psammodes fartus (Phanerotomeina) 32 Vietomorpha foveipennis (Sepidiina) 33 Ocnodes sp. (Phanerotomeina) 34 Physophrynus burdoi (Molurina) 35 Oxura setosa (Oxurina) 36 Ocnodes sp. (Phanerotomeina) 37 Somaticus aeneus (Trachynotina) 38 Trachynotus omercooperi (Trachynotina) 39 Ocnodes procrustes (Phanerotomeina) 40 Peringueyia dispar (Sepidiina) 41 Amiantus gibbosus (Molurina) 42 Sepidium crassicaudatum (Sepidiina) 43 Sepidiostenus ruspolii (Sepidiina) 44 Psammophanes sp. (Molurina) 45 Ocnodes guerini (Phanerotomeina).
From the morphological perspective, Sepidiini are defined by the following combination of characters (
The last comprehensive checklist of the species currently classified within Sepidiini was published by
From the strictly formal point of view, the validity of many names introduced after 1937 remained questionable (e.g., Histrionotus omercooperi Koch, 1955), since their unusual “descriptions” were incorporated in remarks concerning other taxa. Furthermore, the taxonomic affiliation of many genera and species is uncertain because of ambiguous remarks made by the contributors, see notes in the catalogue below.
The main aim of this work is to synthesise available nomenclatural, taxonomic, and distributional information concerning Sepidiini.
Characters proposed to define Sepidiini. 46 Ventral portion of head (Dichtha inflata) 47 11-segmented antenna (Ocnodes similis) 48 connection of pronotum and elytra (Ocnodes procrustes) 49 mesocoxa (Trachynotidus sp.). Abbreviations: cr - cardo; cx - coxa; gu - gula; me - mentum; mp - maxillary palp; pg - postgena; sc - scutellum; st - stipes; tn - trochantin; tr - trochanter.
All nomenclaturally available family-, genus-, and species-group names are included. The author, year, and page of the original description are provided for each scientific name. The type genus for each family-group name and the type species and type fixation for each genus-group name are included. Type species fixed by original designation were only accepted when an explicit statement (e.g., “Type species.”) was used in the original publication (see
Type deposition data and the status of the name bearing types was primarily taken from the original publications. However, when authors provided the name of the entomological collection for the type deposition without referring to any public institutions the following publications were used in order to locate those collections:
Allard, Bates, Crotch, DeGeer, Desbrochers des Loges, Dohrn, Dupont, Gory, Hope, Klug, Pallas, Pascoe, Sahlberg, and Westwood collections –
Banks collection –
Fabricius collection –
Gestro collection –
Haag-Rutenberg collection –
Kocher collection –
Kolbe collection –
Spinola collection –
Type deposition information provided in this catalogue was verified against the databases originating from the Basel (
The following format for deposition information was used throughout the catalogue:
Agricultural Institute Agricultural Institute in Pretoria, Pretoria, South Africa
Barcelona Museum Museu de Ciències Naturals de Barcelona, Barcelona, Spain
Basel Museum Naturhistorisches Museum Basel, Basel, Switzerland
Berlin Museum Museum für Naturkunde, Berlin, Germany
Bloemfontein Museum Bloemfontein National Museum, Bloemfontein, South Africa
Bologna Museum Bologna Zoological Museum, Bologna, Italy
Bremen Museum Übersee Museum, Bremen, Germany
British Museum The Natural History Museum, London, United Kingdom
Brussels Museum Muséum des sciences naturelles de Belgique, Brussels, Belgium
Budapest Museum Magyar Természettudományi Múzeum, Budapest, Hungary
California Academy Museum of the California Academy of Sciences, San Francisco, USA
Cambridge Museum Harvard Museum of Natural History, Cambridge, USA
Cape Museum Iziko South African Museum, Cape Town, South Africa
Companhia Diamantes Companhia de Diamantes de Angola, Luanda, Angola
Copenhagen Museum Statens Naturhistoriske Museum, University of Copenhagen, Copenhagen, Denmark
Cornell University Cornell University, Ithaca, USA
Ditsong Museum Ditsong National Museum of Natural History, Pretoria, South Africa
Dundo Museum Dundo Museum, Dundo, Angola
Durban Museum Durban Natural Science Museum, Durban, South Africa
Florence Museum Museo di Storia Naturale di Firenze, Florence, Italy
Frankfurt Museum Naturmuseum Senckenberg, Frankfurt, Germany
Geneva Museum Muséum d’Histoire Naturelle, Geneva, Switzerland
Genoa Museum Civic Museum of Natural History Giacomo Doria, Genoa, Italy
Glasgow Museum Hunterian Zoology Museum, Glasgow, Scotland
Hamburg University Universität Hamburg, Hamburg, Germany
Humboldt University Humboldt-Universität, Berlin, Germany
Kenya Museum National Museums of Kenya, Nairobi, Kenia
Kiel Museum Zoologischen Museum Kiel, Kiel, Germany
Leiden Museum Naturalis Museum, Leiden, Holland
Lund University Zoological Museum, Lund University, Lund, Sweden
Madrid Museum Museo Nacional de Ciencias Naturales, Madrid, Spain
Maputo Museum Centro de Investigação Científica Algodoeira, Maputo, Mozambique
Marseille Museum Muséum d’histoire naturelle de Marseille, Marseille, France
McGregor Museum McGregor Museum, Kimberley, South Africa
Milan Museum Museo Civico di Storia Naturale, Milano, Italy
Monaco Museum Nouveau Musée National de Monaco, Monaco, Monaco
Munich Museum Bayerisches Nationalmuseum, Munich, Germany
Museo Civico Filangieri Museo Civico Filangieri, Naples, Italy
National Congo nstitute of the National Parks of Belgian Congo (temporarily preserved in Tervuren Museum)
Naturhistoriska riksmuseet Naturhistoriska riksmuseet, Stockholm, Sweden
New York Museum American Museum New York, New York, USA
Ohio State Ohio State University, Columbus, USA
Oxford University Oxford University Museum of Natural History, Oxford, United Kingdom
Paris Museum Muséum National d’Histoire Naturelle, Paris, France
Prague Museum Národní muzeum, Prague, Czech Republic
Pretoria University University of Pretoria, Pretoria, South Africa
Rabat Institute d’Entomologie de l’Institut Scientifque Chérifen, Rabat, Morocco
Rhodes University Rhodes University, Grahamstown, South Africa
Rhodesia Museum National Museum of Southern Rhodesia, Bulawayo, Zimbabwe
South African National
Collection South African National Collection of Insects, Pretoria, South Africa
Stellenbosch University Stellenbosch University, Stellenbosch, South Africa
Stuttgart Museum Staatliches Museum für Naturkunde Stuttgart, Stuttgart, Germany
Tervuren Museum Musée royal de l’Afrique centrale, Tervuren, Belgium
Torino Museum Museo Regionale di Scienze Naturali di Torino, Turin, Italy
Trieste Museum Museo Civico di Storia Naturale, Trieste, Italy
Uppsala University Uppsala universitet Evolutionsmuseet, Uppsala, Sweden
Vienna Museum Naturhistorisches Museum Wien, Wien, Austria
Warsaw Museum Muzeum i Instytut Zoologii, Polska Akademia Nauk, Warsaw, Poland
Windhoek Museum National Museum of Namibia, Windhoek, Namibia
The distribution of all subtribes was illustrated using Quantum GIS (QGIS) v. 2.4. All vector layers were downloaded from the Natural Earth webpage (http://www.naturalearthdata.com). The list of localities was built by consulting available literature and is available as an Suppl. material to this publication (Suppl. material
A total of 1009 valid species and subspecies divided over 55 genera (33 subgenera) and six subtribes is listed in this catalogue. The subtribe Molurina is the most diverse with 382 valid species-group taxa, followed by the Trachynotina (218), Phanerotomeina (177), Sepidiina (124), Oxurina (63), and Hypomelina (45). In most cases, the species and subspecies diversity is not equally divided over the available genera (Fig.
Although this publication focuses on the nomenclature and classification of Sepidiini, the examined references enabled to reveal the most urgent taxonomic problems within the tribe. According to
A database containing 2523 distributional records (857 not georeferenced) was created during this study (Fig.
According to the acquired data, Sepidiini are widely distributed throughout the Afrotropical Realm, except its northwestern parts (Fig.
Type genus. Sepidium Fabricius, 1775
Subtribes included. Hypomelina, Molurina, Oxurina, Phanerotomeina, Sepidiina, Trachynotina.
Distribution. Western Palaearctic (mainly Mediterranean Basin) and Afrotropical Realm (with exception of western Africa) (Fig.
Type genus. Hypomelus Solier, 1843
Taxonomic diversity. (9 gen., 45 spp.): Argenticrinis (1 sp.), Bombocnodulus (6), Brinckia (8), Hypomelus (14), Iugidorsum (4), Sulcipectus (2), Trachynotidus (8), Triangulipenna (1), Uniungulum (1).
Distribution. Southern part of the Afrotropical Realm. Majority of species were described from the Namibian coast. A single genus, Bombocnodulus, reaching north to Central Africa (Fig.
Type species. Argenticrinis haackei Louw, 1979 (by original designation); syn. of Psammodes lossowi Koch, 1952)
lossowi (Koch, 1952: 339) Psammodes [Louw, 1980: 216]
Type data. Holotype (Ditsong Museum)
= Argenticrinis haackei Louw, 1979: 101 [syn. by
Type data. Holotype (Windhoek Museum) and paratypes (Ditsong Museum, Windhoek Museum)
Type species. Psammodes crinicollis Haag-Rutenberg, 1879 (by monotypy)
Notes. Although the description is unusual, it meets the criteria of Art. 11 and 13
Genus originally described under Phanerotomeina, and subsequently transferred to Hypomelina by
crinicollis crinicollis (Haag-Rutenberg, 1879: 293) Psammodes [Koch, 1955: 36]
Type data. Holotype (British Museum)
= Psammodes inquinatus Péringuey, 1899: 292 [syn. by
Type data. Holotype (Cape Museum)
crinicollis fortuitus (Péringuey, 1899: 292) Psammodes [Penrith, 1986: 61]
Type data. Syntypes (Cape Museum)
Notes.
dollmani Penrith, 1986: 64
Type data. Holotype (British Museum) and paratype (Ditsong Museum)
longantennatus Penrith, 1986: 62
Type data. Holotype (Ditsong Museum) and paratype (Windhoek Museum)
torridus Penrith, 1986: 62
Type data. Holotype (British Museum) and paratypes (British Museum, Ditsong Museum)
wittei Penrith, 1986: 66
Type data. Holotype (Brussels Museum) and paratypes (Brussels Museum, Ditsong Museum)
Type species. Psammodes debilis Péringuey, 1899 (by original designation)
australis Penrith, 1986: 81
Type data. Holotype (Ditsong Museum) and paratypes (Ditsong Museum, Windhoek Museum)
debilis (Péringuey, 1899: 277) Psammodes [Koch, 1962b: 121]
Type data. Lectotype, designated by
delicata Penrith, 1986: 77
Type data. Holotype (Windhoek Museum) and paratypes (Ditsong Museum, Windhoek Museum)
insularis (Péringuey, 1908: 410) Trachynotidus [Koch, 1962b: 121]
Type data. Lectotype, designated by
oblonga (Thunberg, 1787: 37) Sepidium [Ferrer, 2009: 114]
Type data. Holotype (Uppsala University)
oograbiensis Koch, 1962b: 118
Type data. Holotype (Ditsong Museum) and paratype (Cape Museum)
serratina Koch, 1962b: 119
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Ditsong Museum)
vaga (Péringuey, 1908: 411) Trachynotidus [Koch, 1962b: 122]
Type data. Lectotype, designated by
Type species. Hypomelus bicolor Solier, 1843 (by original designation); syn. of Helops peronatus Germar, 1823
Notes. Interpreted as a subgenus of Psammodes for a long time (e.g.,
The species composition mostly follows that in Gebien’s catalogue (1937a).
basalis (Haag-Rutenberg, 1871a: 70, in key) Psammodes [Gebien, 1937a: 772]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Note. A detailed morphological description was provided by
inaequalis Solier, 1843: 98
Type data. Holotype (Torino Museum – Spinola coll.)
= Hypomelus flagrans Péringuey, 1899: 273 [homonym of Psammodes flagrans Péringuey, 1899: 295 published on the same date;
= Psammodes dentipennis
Type data. Holotype (Cape Museum)
interstitialis (Haag-Rutenberg, 1871a: 70, in key) Psammodes [Gebien, 1937a: 772]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Note. A detailed morphological description was provided by
obliquatus Solier, 1843: 97
Type data. Holotype (Geneva Museum – Gory collection)
= Hypomelus sabulosus Solier, 1843: 308 [syn. by
Type data. Syntypes (Paris Museum)
obliteratus Solier, 1843: 96
Type data. Holotype (Geneva Museum – Gory collection)
peringueyi (
= Hypomelus plausibilis Péringuey, 1899: 295 [homonym of Psammodes plausibilis Péringuey, 1899: 271 published on the same date;
Type data. Syntypes (Cape Museum)
peronatus (Germar, 1823: 149) Helops [Gebien, 1937a: 771]
Type data. Syntypes (Cape Museum, Paris Museum)
= Oxura psammodioides Guérin-Méneville, 1834: 20 [syn. by
Type data. Syntypes (Paris Museum)
= Hypomelus bicolor Solier, 1843: 100 [syn. by
Type data. Syntypes (Paris Museum)
profugus (Péringuey, 1899: 277) Psammodes [
Type data. Syntypes (Cape Museum)
reflexicollis (Haag-Rutenberg, 1871a: 70, in key) Psammodes [Gebien, 1937a: 772]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
reflexus (Haag-Rutenberg, 1871a: 70, in key) Psammodes [Gebien, 1937a: 772]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
servus Péringuey, 1899: 294
Type data. Syntypes (Cape Museum)
setosocostatus (Haag-Rutenberg, 1871a: 70, in key) Psammodes [Gebien, 1937a: 772]
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
villosocostatus Solier, 1843: 98
Type data. Holotype (Torino Museum – Spinola coll.)
vulpinus (Haag-Rutenberg, 1873: 45, replacement name) Psammodes [Gebien, 1937a: 772]
= Psammodes hirtipennis Haag-Rutenberg, 1871b: 92 [junior primary homonym of Psammodes hirtipennis Haag-Rutenberg, 1871a: 105]
Type data. Holotype (Munich Museum)
Type species. Iugidorsum cumstriis Louw, 1979 (by original designation)
cumstriis cumstriis Louw, 1979: 106
Type data. Holotype (Windhoek Museum) and paratypes (Windhoek Museum and Ditsong Museum)
cumstriis magnum Louw, 1979: 106
Type data. Holotype (Ditsong Museum) and paratype (Windhoek Museum)
cumstriis prominens Louw, 1979: 106
Type data. Holotype and paratypes (Ditsong Museum)
sinestriis Louw, 1979: 107
Type data. Holotype and paratype (Windhoek Museum)
Type species. Sulcipectus levis Louw, 1979 (by original designation)
cumcavus Louw, 1979: 113
Type data. Holotype and paratypes (Windhoek Museum)
levis Louw, 1979: 110
Type data. Holotype and paratypes (Windhoek Museum)
Type species. Psammodes thoreyi Haag-Rutenberg, 1871 (here designated)
Notes. In 1904, Péringuey described a new species named damarinus under “Gen. Trachynotideus Péring”. The spelling “Trachynotideus” was generally treated as an incorrect subsequent spelling of Trachynotidus by subsequent authors (e.g.,
alstoni (Péringuey, 1885: 116) Clinocranion [Péringuey, 1899: 297]
Type data. Holotype (Cape Museum)
angulicollis (Haag-Rutenberg, 1871a: 69, in key) Psammodes [Péringuey, 1904: 234]
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. A detailed morphological description was provided by
cognatus Péringuey, 1899: 297
Type data. Holotype (Cape Museum)
cruentus Péringuey, 1908: 411
Type data. Syntypes (Cape Museum)
eximius Péringuey, 1899: 298
Type data. Syntypes (British Museum, Cape Museum)
gravis (
= Hypomelus grandis Solier, 1843: 101 [homonym of Phanerotoma grande Solier, 1843: 90 published on the same date;
Type data. Holotype (Geneva Museum – Gory collection)
rufozonatus (Fairmaire, 1888a: 194) Trachynotus [Gebien, 1937a: 771]
Type data. Holotype (Paris Museum)
= Trachynotidus manifestus Péringuey, 1899: 297 [syn. by
Type data. Syntypes (Cape Museum)
thoreyi (Haag-Rutenberg, 1871b: 104) Psammodes [Gebien, 1937a: 771]
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
Type species. Triangulipenna lacuna Louw, 1979 (by original designation)
lacuna Louw, 1979: 115
Type data. Holotype (Cape Museum) and paratypes (Cape Museum, Windhoek Museum)
Type species. Uniungulum hoeschi Koch, 1962 (by original designation)
hoeschi Koch, 1962b: 114
Type data. Holotype (Ditsong Museum) and paratypes (Budapest Museum, Ditsong Museum)
Type genus. Moluris Latreille, 1802
Taxonomic diversity. (15 gen., 382 spp.): Amiantus (15 sp.), Arturium (16), Brachyphrynus (11), Dichtha (5), Distretus (22), Euphrynus (4), Glyptophrynus (8), Melanolophus (11), Moluris (15), Phrynocolus (14), Phrynophanes (13), Physophrynus (8), Psammodes (169), Psammophanes (63), Psammotyria (8).
Distribution. With exception of western Africa, widely distributed in the Afrotropical Realm. Glyptophrynus is the only Malagasy representative of the whole tribe (Fig.
Type species. Amiantus gibbosus Fåhraeus, 1870 (by subsequent designation by
browni Haag-Rutenberg, 1871a: 47
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
connexus Haag-Rutenberg, 1871a: 49
Type data. Syntypes (British Museum)
costipennis Kolbe, 1886: 292
Type data. Holotype (Berlin Museum)
decemcostatus (Gebien, 1910a: 156) Phrynocolus [
Type data. Holotype (Hamburg University)
gibbosus Fåhraeus, 1870: 280
Type data. Syntypes (Naturhistoriska riksmuseet)
globulipennis Péringuey, 1896: 167
Type data. Holotype (Cape Museum)
= Amiantus multicostatus Fairmaire, 1899a: 181 [syn. by
Type data. Syntypes (Basel Museum, Paris Museum)
lobicollis Kolbe, 1886: 291
Type data. Holotype (Berlin Museum)
mechowi (Quedenfeldt, 1885: 6) Distretus [
Type data. Syntypes (Berlin Museum)
octocostatus Péringuey, 1896: 167
Type data. Holotype (Cape Museum)
octocristatus Fairmaire, 1899: 181
Type data. Holotype (Paris Museum)
opacus Haag-Rutenberg, 1871a: 49
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
pusillus Péringuey, 1904: 235
Type data. Syntypes (Cape Museum)
rusticus Fåhraeus, 1870: 280
Type data. Syntypes (Naturhistoriska riksmuseet)
scrobipennis Haag-Rutenberg, 1875: 68
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
undosus Distant, 1892: 199
Type data. Holotype (Ditsong Museum)
Type species. Melanolophus ater Waterhouse, 1885 (by original designation)
absciri Koch, 1959: 7
Type data. Holotype (Munich Museum)
ater (Waterhouse, 1885: 234) Melanolophus [Koch, 1951: 83]
Type data. Syntypes (British Museum)
Notes.
auriculatus (Gebien, 1910a: 155) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Basel Museum)
benanum (Wilke, 1921: 167) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Basel Museum)
crispatus (Fairmaire, 1887: 184) Phrynocolus [Koch, 1951: 83]
Type data. Syntypes (Paris Museum)
= Phrynocolus undatocostatus Kolbe, 1891: 30 [syn. by
Type data. Holotype (Berlin Museum)
dallastai Ardoin, 1977: 811
Type data. Holotype and paratype (Tervuren Museum)
fiomicum (Wilke, 1921: 167) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Berlin Museum)
fulleborni (Wilke, 1921: 167) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Berlin Museum)
gebieni (Wilke, 1921: 168) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Basel Museum)
glauningi (Wilke, 1921: 167) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Berlin Museum)
methneri (Wilke, 1921: 166) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Basel Museum)
parvulus (Gestro, 1895: 132) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Genoa Museum)
pretiosum (Wilke, 1921: 167) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Basel Museum)
tenuecostatus (Gebien, 1910a: 155) Phrynocolus [Koch, 1951: 83]
Type data. Syntypes (Basel Museum, Berlin Museum)
undaticostis (Fairmaire, 1887: 183) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Paris Museum)
wembericum (Wilke, 1921: 168) Phrynocolus [Koch, 1951: 83]
Type data. Holotype (Berlin Museum)
Type species. Brachyphrynus spissicornis Fairmaire, 1882 (by monotypy)
abyssinicus abyssinicus (Haag-Rutenberg, 1871a: 39) Phrynocolus [Koch, 1951: 85]
Type data. Syntypes (Basel Museum, British Museum)
Notes. Treated as a synonym of Psammophanes catenatus (Reiche, 1850) by
abyssinicus breuningi Kaszab, 1963: 348
Type data. Holotype (Tervuren Museum) and paratypes (Budapest Museum, Tervuren Museum)
gallanus (Wilke, 1921: 163) Phrynocolus [Koch, 1951: 85]
Type data. Holotype (Berlin Museum)
kuntzeni (Wikle, 1921: 163) Phrynocolus [Koch, 1951: 85]
Type data. Syntypes (Basel Museum)
petrosus erlangeri (Wilke, 1921: 164) Phrynocolus [Koch, 1951: 85]
Type data. Syntypes (Basel Museum)
petrosus petrosus (Gerstaecker, 1871: 59) Phrynocolus [Koch, 1951: 85]
Type data. Holotype (Berlin Museum)
= Phrynocolus ikutanus Fairmaire, 1897: 113 [syn. by
Type data. Syntypes (Paris Museum)
placidus (Kolbe, 1885: 112) Phrynocolus [Koch, 1951: 85]
Type data. Holotype (Geneva Museum)
somalicus (Wilke, 1921: 164) Phrynocolus [Koch, 1951: 85]
Type data. Syntypes (Basel Museum)
spissicornis Fairmaire, 1882a: 72
Type data. Holotype (Paris Museum)
subnodosus (Gebien, 1937b: 48) Phrynocolus [Koch, 1951: 85]
Type data. Holotype (Trieste Museum) and paratype (Basel Museum)
wachei (Wilke, 1921: 163) Phrynocolus [Koch, 1951: 85]
Type data. Syntypes (Basel Museum)
Type species. Cryptogenius inflatus Gerstacker, 1854 (by original designation)
Notes. “Dichtha incantatoris / incantatoria Koch, 1952” is considered here as a nomen nudum, since no published record of this species-group name was found during the present work.
cubica (Guérin-Méneville, 1845: 285) Moluris [Haag-Rutenberg, 1871a: 41]
Type data. Holotype (Paris Museum)
inflata (Gerstaecker, 1854: 532) Cryptogenius [Haag-Rutenberg, 1871a: 41]
Type data. Syntypes (Berlin Museum)
modesta Robiche, 2013: 159
Type data. Holotype (Paris Museum) and paratypes (Paris Museum, Gérard Robiche collection)
transvalica Brancsik, 1914: 65
Type data. Syntypes (Budapest Museum)
quedenfeldti Kolbe, 1886: 293
Type data. Syntypes (Berlin Museum)
Type species. Moluris amplipennis Fåhraeus, 1870 (by subsequent designation by
Type species. Moluris amplipennis Fåhraeus, 1870 (by subsequent designation by
amplipennis (Fåhraeus, 1870: 262) Moluris [Haag-Rutenberg, 1871a: 43]
Type data. Syntypes (Naturhistoriska riksmuseet)
dissociatus (Péringuey, 1899: 274) Psammodes [Gebien, 1937a: 757]
Type data. Holotype (Cape Museum)
fahraei Haag-Rutenberg, 1871a: 43
Type data. Holotype (Naturhistoriska riksmuseet)
inaequalis Fairmaire, 1894: 320
Type data. Holotype (Basel Museum)
mashunus (Péringuey, 1896: 167) Amiantus [Gebien, 1937a: 757]
Type data. Holotype (Cape Museum)
undosus Kolbe, 1886: 291
Type data. Syntypes (Berlin Museum)
undatus (Haag-Rutenberg, 1875: 69) Amiantus [Gebien, 1910b: 153]
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
variabilis Gebien, 1910a: 153
Type data. Syntypes (Basel Museum, Tervuren Musuem)
variolosus (Guérin-Méneville, 1854: 245) Moluris [Haag-Rutenberg, 1871a: 44]
Type data. Holotype (Warsaw Museum – Dohrn coll.)
= Moluris pilicornis Fåhraeus, 1870: 263 [syn. by
Type data. Syntypes (Naturhistoriska riksmuseet)
vietus (Péringuey, 1899: 273) Psammodes [Gebien, 1937a: 757]
Type data. Holotype (Cape Museum)
Type species. Distretus (Perdistretus) vilhenai Koch, 1953 (by original designation)
acutecostatus (Fairmaire, 1888b: 260) Dichtha [Koch, 1953: 65]
Type data. Syntypes (Basel Museum, Leiden Museum)
angolanus Koch, 1953b: 72
Type data. Holotype (Cape Museum)
angustipennis Péringuey, 1892: 52
Type data. Holotype (Cape Museum)
Notes. Considered as a synonym of Perdistretus acutecostatus Fairmaire, 1888b by
Originally described in combination with the generic name “Dichtrethus”, which is treated as an incorrect subsequent spelling of Distretus, not in prevailing usage.
auritus Koch, 1953b: 73
Type data. Holotype (Munich Museum)
duartei Koch, 1953b: 70
Type data. Holotype (Munich Museum) and paratypes (Basel Museum, Ditsong Museum, Munich Museum)
gracilis Gebien, 1910a: 152
Type data. Syntypes (Tervuren Museum)
mormolyce Koch, 1953b: 68
Type data. Holotype (Munich Museum) and paratype (Basel Museum, Ditsong Museum, Munich Museum)
seminitidus Quedenfeldt, 1888: 184
Type data. Holotype (Berlin Museum)
strioliceps Koch, 1953b: 71
Type data. Holotype (Munich Museum)
schoutedeni Koch, 1954a: 435
Type data. Holotype (Tervuren Museum) and paratypes (Ditsong Museum, Tervuren Museum)
upembensis Koch, 1954a: 437
Type data. Holotype (National Congo) and paratypes (Ditsong Museum, National Congo)
vilhenai Koch, 1953b: 65
Type data. Holotype (Dundo Museum) and paratype (Ditsong Museum, Dundo Museum)
Type species. Euphrynus spinithorax Fairmaire, 1897 (by monotypy)
carinatus (Fåhraeus, 1870: 281) Amiantus [Koch, 1952: 345]
Type data. Holotype (Naturhistoriska riksmuseet)
= Amiantus costatus Péringuey, 1896: 168 [syn. by
Type data. Syntypes (Cape Museum)
jansei Koch, 1952: 343
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Ditsong Museum)
sexdentatus Koch, 1952: 344
Type data. Holotype (Ditsong Museum) and paratype (Basel Museum)
spinithorax Fairmaire, 1897: 114
Type data. Holotype (Geneva Museum)
Type species. Glyptophrynus tenuesculptus Fairmaire, 1899 (by monotypy)
Notes. Treated as a synonym of Phrynocolus by several authors (e.g.,
cordipennis Koch, 1962a: 12
Type data. Holotype and paratypes (Ditsong Museum)
madecassus madecassus (Fairmaire, 1901: 183) Phrynocolus [Wilke, 1921: 174]
Type data. Syntypes (Berlin Museum, British Museum)
madecassus pauliani Koch, 1962a: 15
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Budapest Museum, Ditsong Museum)
ovipennis ovipennis (Fairmaire, 1899b: 533) Phrynocolus [Wilke, 1921: 174]
Type data. Syntypes (Paris Museum)
ovipennis serricostatus Koch, 1962a: 17
Type data. Holotype (Ditsong Museum)
tenuesculptus crassigranulatus Wilke, 1921: 174
Type data. Syntypes (Berlin Museum)
tenuesculptus tenuesculptus Fairmaire, 1899b: 532
Type data. Syntypes (Basel Museum, Paris Museum)
voeltzkowi Wilke, 1921: 174
Type data. Syntypes (Berlin Museum)
Type species. Melanolophus septemcostatus Fairmaire, 1882 (by monotypy)
Notes. Treated as a synonym of Amiantus by several authors (e.g.,
gridellii Koch, 1956: 170
Type data. Holotype (Trieste Museum) and paratypes (Ditsong Museum, Trieste Museum)
lomianus Koch, 1956: 173
Type data. Holotype (Trieste Museum) and paratypes (Basel Museum, Ditsong Museum, Trieste Museum)
picteti picteti (Haag-Rutenberg, 1871a: 46) Amiantus [Koch, 1960: 258]
Type data. Holotype (Geneva Museum)
picteti septemcostatus Fairmaire, 1882a: 70
Type data. Holotype (Basel Museum)
Notes. Synonymised with the nominotypical form by
picteti splendidus
Type data. Holotype (Ditsong Museum)
praeplanatus Koch, 1960: 261
Type data. Holotype (Ditsong Museum)
sexcostatus benardellii Koch, 1960: 258
Type data. Holotype (Ditsong Museum) and paratypes (Geneva Museum, Munich Museum)
sexcostatus gibbithorax Koch, 1956: 175
Type data. Holotype (Trieste Museum) and paratypes (Ditsong Museum, Trieste Museum)
sexcostatus hellardi Koch, 1960: 258
Type data. Holotype and paratype (Ditsong Museum)
sexcostatus sexcostatus (Gahan, 1900: 28) Amiantus [Koch, 1956: 175]
Type data. Holotype (British Museum)
sexcostatus tuberculatus Koch, 1960: 258
Type data. Holotype (Ditsong Museum) and paratypes (Berlin Museum, Munich Museum, Milan Museum)
Type species. Tenebrio gibbus Pallas, 1781 (by monotypy)
= Physodera Solier, 1843: 78 [junior subjective synonym proposed by
Type species. Pimelia gibba Fabricius, 1787 (by original designation)
chevrolati Haag-Rutenberg, 1871a: 52
Type data. Holotype (Paris Museum)
discoidea Guérin-Méneville, 1845: 286
Type data. Holotype (Paris Museum)
Notes. According to
ferrari Haag-Rutenberg, 1871a: 55
Type data. Holotype (Vienna Museum)
gibba (Pallas, 1781: 46) Tenebrio [Latreille, 1802: 169]
Type data. Syntypes (Humboldt University – Pallas collection)
= Pimelia planata Thunberg, 1787: 49 [syn. by
Type data. Syntypes (Uppsala University)
= Pimelia gibba Fabricius, 1787: 24 [junior secondary homonym of Tenebrio gibba Pallas, 1781: 46]
Type data. Syntypes (Copenhagen Museum, Glasgow Museum, Kiel Museum)
= Pimelia bistriata Herbst, 1799: 50 [syn. by
Type data. Syntypes (Berlin Museum)
gibbicollis Haag-Rutenberg, 1871a: 107
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
gibbosa (Thunberg, 1787: 49) Pimelia [Solier, 1843: 79]
Type data. Syntypes (Uppsala University)
= Opatrum gibbosum Thunberg, 1821: 33 [syn. by
Type data. Holotype (Uppsala University)
gibbosa globulicollis Solier, 1843: 80
Type data. Holotype (Torino Museum – Spinola coll.)
herbsti Haag-Rutenberg, 1871a: 54, replacement name
= Pimelia gibba Herbst, 1799: 48 [junior secondary homonym of Tenebrio gibba Pallas, 1781: 46]
Type data. Holotype (Berlin Museum)
nitida Haag-Rutenberg, 1871a: 52
Type data. Syntypes (Brussels Museum)
pseudonitida Péringuey, 1908: 406
Type data. Holotype (Cape Museum)
redtenbacheri Haag-Rutenberg, 1871a: 56
Type data. Holotype (Vienna Museum)
rustica Haag-Rutenberg, 1871a: 54
Type data. Holotype (Naturhistoriska riksmuseet)
semiscabra Solier, 1843: 81
Type data. Holotype (Torino Museum – Spinola coll.)
strigosa (Herbst, 1799: 49) Pimelia [Haag-Rutenberg, 1871a: 55]
Type data. Syntypes (Berlin Museum)
= Moluris rouleti Solier, 1843: 80 [syn. by
Type data. Holotype (Geneva Museum – Gory collection)
tuberculata Haag-Rutenberg, 1871a: 107
Type data. Syntypes (Paris Museum)
Type species. Cryptogenius dentatus Solier, 1843 (by original designation)
Type species. Phrynocolus frondosus Gerstacker, 1871 (original designation)
denhardti denhardti Wilke, 1921: 165 [Koch, 1951: 93]
Type data. Holotype (Berlin Museum)
denhardti fractus Koch, 1969: 13
Type data. Holotype (Munich Museum)
denhardti humeralis Koch, 1969: 12
Type data. Holotype (Geneva Museum)
desaegeri Koch, 1969: 15
Type data. Holotype (Brussels Museum) and paratypes (Brussels Museum, Tervuren Museum)
frondosus Gerstaecker, 1871: 59 [Koch, 1951: 93]
Type data. Holotype (Cape Museum)
transversus Fairmaire, 1887: 183 [Koch, 1951: 93]
Type data. Holotype (Paris Museum)
subfrondosus Wilke, 1921: 166 [Koch, 1951: 93]
Type data. Syntypes (Basel Museum, Berlin Museum)
= Cryptogenius Solier, 1843: 37 [junior homonym of Cryptogenius Westwood, 1842 (Coleoptera: Hybosoridae)]
Type species. Cryptogenius dentatus Solier, 1843 (by original designation)
dentatus (Solier, 1843: 38) Cryptogenius [Lacordaire, 1859: 201]
Type data. Syntypes (Cape Museum, Paris Museum)
felinus Koch, 1951: 89
Type data. Holotype (Paris Museum) and paratypes (Basel Museum)
spinolai spinolai (Solier, 1843: 39) Cryptogenius [Lacordaire, 1859: 201]
Type data. Holotype (Warsaw Museum – Dupont collection)
Notes.
= Phrynocolus niloticus Haag-Rutenberg, 1871a: 38 [syn. by
Type data. Syntypes (Basel Museum, British Museum)
= Phrynocolus cultratus Fairmaire, 1891a: 249 [syn. by
Type data. Holotype (Paris Museum)
spinolai wilkei Koch, 1951: 88
Type data. Holotype and paratypes (Basel Museum)
theryi Koch, 1951: 89
Type data. Holotype (Basel Museum) and paratypes (Basel Museum, Budapest Museum, Ditsong Museum)
Type species. Phrynocolus spinipennis Gebien, 1910 (by original designation)
spinipennis Gebien, 1910a: 154
Type data. Syntypes (Basel Museum)
incertae sedis
menghallensis Wilke, 1922: 381
Type data. Holotype (Berlin Museum)
Notes. This species and
Type species. Moluris gredleri Haag-Rutenberg, 1877 (by original designation)
Notes. Originally described as a subgenus of Phrynocolus. Elevated to generic level by
citernii (Gridelli, 1939b: 229) Psammodes [Koch, 1969: 21]
Type data. Holotype (Geneva Museum)
cryptisculptus Koch, 1969: 4
Type data. Holotype (Munich Museum)
discoideus (Fairmaire, 1891b: CCXCIV) Phrynocolus [Koch, 1969: 17]
Type data. Holotype (Paris Museum)
gredleri (Haag-Rutenberg, 1877: 515) Moluris [Koch, 1951: 93]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
= Phrynocolus unicarinatus Wilke, 1921: 170 [syn. by
Type data. Holotype (Basel Museum)
humilis (Wilke, 1921: 170) Phrynocolus [Koch, 1951: 93]
Type data. Holotype (Berlin Museum)
lateritius (Wilke, 1921: 169) Phrynocolus [Koch, 1951: 93]
Type data. Syntypes (Basel Museum, Berlin Museum)
neumanni (Wilke, 1921: 169) Phrynocolus [Koch, 1951: 93]
Type data. Holotype (Basel Museum)
reticulatus Wilke, 1921: 169
Type data. Syntypes (Basel Museum, Berlin Museum)
schereri Koch, 1969: 20
Type data. Holotype (Munich Museum)
scortecii Koch, 1969: 19
Type data. Holotype (Geneva Museum)
schoutedeni (Koch, 1951: 92) Phrynocolus [Koch, 1969: 24]
Type data. Holotype (Brussels Museum) and paratypes (Basel Museum, Brussels Museum)
squamifer gridellianus Koch, 1960: 262
Type data. Holotype and paratype (Ditsong Museum)
squamifer squamifer (Gridelli, 1939b: 228) Psammodes [Koch, 1960: 262]
Type data. Syntypes (Geneva Museum, Trieste Museum)
Type species. Physophrynus burdoi Fairmaire, 1882 (by monotypy)
bufo (Haag-Rutenberg, 1871a: 48) Amiantus [Koch, 1953a: 177]
Type data. Holotype (Warsaw Museum – Dohrn coll.)
= Amiantus reichardi Kolbe, 1886: 228 [syn. by
Type data. Holotype (Berlin Museum)
burdoi Fairmaire, 1882b: L
Type data. Holotype (Paris Museum)
bredoi Mal, 2005: 9
Type data. Holotype and paratypes (Brussels Museum)
bufocrenatocostatus (Fairmaire, 1887: 181) Amiantus [Koch, 1953a: 177]
Type data. Holotype (Fairmaire collection)
haroldi (Haag-Rutenberg, 1871a: 47) Amiantus [Koch, 1953a: 177]
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
kaszabi Koch, 1953a: 176
Type data. Holotype (Budapest Museum)
manicanus (Péringuey, 1899: 226) Amiantus [Koch, 1953a: 177]
Type data. Holotype (Cape Museum)
revoili Fairmaire, 1887: 182
Type data. Holotype (Paris Museum)
Type species. Psammodes longicornis Kirby, 1819 (by monotypy)
= Piesomera Solier, 1843: 77 [junior subjective synonym proposed by
Type species. Pimelia scabra Fabricius, 1775 (by monotypy)
= Psammodophysis Péringuey, 1899: 296 [junior subjective synonym proposed by
Type species. Psammodophysis probes Péringuey, 1899 (here designated)
= Parmularia Koch, 1955: 35, syn. n. [homonym of Parmularia Macgillivray, 1887 (Bryozoa: Cheilostomida)]
Type species. Psammodes caffra Fåhraeus, 1870 (by monotypy)
Notes. Originally described as a monotypic subgenus of Psammodes. Interpreted here as a synonym of the nominal form, as sustaining a weakly defined and monotypic subgenus within present Psammodes seems to be unjustified.
algoensis Péringuey, 1899: 275
Type data. Holotype (Cape Museum)
asperulipennis Fairmaire, 1888: 193
Type data. Holotype (Paris Museum)
atratus Haag-Rutenberg, 1871a: 73, in key
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
basuto Koch, 1953c: 7
Type data. Holotype (Ditsong Museum) and paratypes (Ditsong Museum, Lund University, Munich Museum)
barbatus Fåhraeus, 1870: 268
Type data. Holotype (Naturhistoriska riksmuseet)
= Psammodes praeliator Péringuey, 1899: 272 [syn. by
Type data. Holotype (Cape Museum)
batesi Haag-Rutenberg, 1871a: 77
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. Treated as a synonym of Psammodes ponderosus Fåhraeus, 1870 by
bennigseni Kraatz, 1897: 46
Type data. Holotype (Berlin Museum)
blapsoides Haag-Rutenberg, 1871a: 63, in key
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
brunneus brunneus (Olivier, 1795: 14) Pimelia [Haag-Rutenberg, 1871b: 42]
Type data. Syntypes (Paris Museum)
brunneus rufocastaneus Haag-Rutenberg, 1871b: 42
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
brunnipes Haag-Rutenberg, 1871a: 72, in key
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
caelatus Péringuey, 1899: 281
Type data. Syntypes (Cape Museum)
caffra Fåhraeus, 1870: 265
Type data. Holotype (Naturhistoriska riksmuseet)
caraboides Haag-Rutenberg, 1871a: 69, in key
Type data. Syntypes (British Museum, Cape Museum, Geneva Museum)
Notes. A detailed morphological description was provided by
carinatus Haag-Rutenberg, 1871a: 103
Type data. Syntypes (Berlin Museum)
clarus Haag-Rutenberg, 1873: 76
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
collaris Haag-Rutenberg, 1871b: 101
Type data. Holotype (Berlin Museum)
coloratus Haag-Rutenberg, 1871a: 71, in key
Type data. Holotype (Berlin Museum)
Notes. A detailed morphological description was provided by
comatus Haag-Rutenberg, 1871a: 106
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
comptus Haag-Rutenberg, 1871a: 109
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
convexus (Solier, 1843: 91) Phanerotoma [Haag-Rutenberg, 1871b: 61]
Type data. Holotype (Paris Museum)
coriaceus (Gerstaecker, 1854: 532) Phanerotoma [Haag-Rutenberg, 1871b: 68]
Type data. Syntypes (Berlin Museum)
= Psammodes manifestus
Type data. Holotype (Cape Museum)
costalis Haag-Rutenberg, 1871a: 97
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll., Royal Museum)
dejeani (Solier, 1843: 71) Moluris [Haag-Rutenberg, 1871a: 92]
Type data. Holotype (Paris Museum)
depressicollis Haag-Rutenberg, 1871a: 72, in key
Type data. Syntypes (British Museum)
Notes. A detailed morphological description was provided by
devexus Fåhraeus, 1870: 266
Type data. Holotype (Cape Museum)
diabolicus diabolicus Koch, 1952: 335
Type data. Holotype (Ditsong Museum) and paratype (Basel Museum, Budapest Museum, Ditsong Museum)
diabolicus tactilis Koch, 1962b: 123
Type data. Holotype and paratype (Ditsong Museum)
difficilis Haag-Rutenberg, 1871a: 73, in key
Type data. Syntypes (British Museum, Geneva Museum)
Notes. A detailed morphological description was provided by
dilutus Haag-Rutenberg, 1871a: 64, in key
Type data. Holotype (Warsaw Museum – Dohrn coll.)
Notes. A detailed morphological description was provided by
dimidiatus Haag-Rutenberg, 1871a: 71, in key
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
discrepans Péringuey, 1904: 230
Type data. Holotype (Cape Museum)
dohrni Haag-Rutenberg, 1871a: 67, in key
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
eberlanzi Koch, 1952: 337
Type data. Holotype (Ditsong Museum)and paratypes (British Museum, Budapest Museum, Ditsong Museum)
egregius Haag-Rutenberg, 1871a: 74
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
ethologus Koch, 1953c: 10
Type data. Holotype (Ditsong Museum) and paratype (Durban Museum)
expletus Quedenfeldt, 1885: 4
Type data. Syntypes (Berlin Museum)
Notes. Type specimens of this species were unknown to
fartus Péringuey, 1904: 232
Type data. Holotype (Cape Museum)
= Psammodes illotus Péringuey, 1904: 233 [syn. by
Type data. Syntypes (Cape Museum)
Psammodes ferrugineus Haag-Rutenberg, 1871a: 79
Type data. Syntypes (Cape Museum, Munich Museum – Haag-Rutenberg coll.)
flagrans Péringuey, 1899: 273
Type data. Holotype (Cape Museum)
fragilis Haag-Rutenberg, 1871a: 68, in key
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
fritschi Haag-Rutenberg, 1871a: 103
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
funestus Haag-Rutenberg, 1871a: 72, in key
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
gariesus Péringuey, 1899: 282
Type data. Holotype (Cape Museum)
gerstaeckeri Haag-Rutenberg, 1871b: 100
Type data. Holotype (Berlin Museum)
gibbus coelata (Solier, 1843: 64) Moluris [Haag-Rutenberg, 1871a: 82]
Type data. Holotype (Paris Museum)
gibbus gibbus (Linnaeus, 1760: 226) Tenebrio [Ferrer & Holson, 2009: 30]
Type data. Lectotype, designated by Ferrer and Holson (2009) (Naturhistoriska riksmuseet)
= Pimelia striata Fabricius, 1775: 251 [syn. by Ferrer and Holson (2009: 34)]
Type data. Syntypes (Copenhagen Museum, Kiel Museum)
= Tenebrio glandiformis Pallas, 1781: 45 [syn. by Haag-Rutenberg, 1871a: 82]
Type data. Syntypes (Humboldt University – Pallas collection)
gibbus gravidus (Solier, 1843: 69) Moluris [Haag-Rutenberg, 1871a: 85]
Type data. Syntypes (Paris Museum)
gibbus hemisphaericus (Solier, 1843: 68) Moluris [Haag-Rutenberg, 1871a: 85]
Type data. Holotype (Paris Museum)
gibbus nigrocostatus Haag-Rutenberg, 1871a: 85
Type data. Syntypes (Munich Museum)
gibbus solieri Gebien, 1910b: 161, replacement name
= Moluris unicolor Solier, 1843: 64 [junior secondary homonym of Pimelia unicolor Fabricius, 1787: 316].
Type data. (Warsaw Museum – Dupont collection)
gibbus unicolor (Fabricius, 1787: 316) Pimelia [Haag-Rutenberg, 1871a: 82]
Type data. Syntypes (British Museum, Kiel Museum, Naturhistoriska riksmuseet)
glaber Koch, 1953c: 10
Type data. Holotype (Lund University) and paratypes (Ditsong Musuem, Lund University)
glabratus bienus Koch, 1953b: 77
Type data. Holotype (Munich Museum)
glabratus glabratus Harold, 1878: 106
Type data. Holotype (Berlin Museum)
grandis (Solier, 1843: 90) Phanerotoma [Gemminger, 1870: 122]
Type data. Holotype (Paris Museum)
= Psammodes lugubris Fåhraeus, 1870: 269 [syn. by
Type data. Holotype (Naturhistoriska riksmuseet)
granulatus (Solier, 1843: 87) Phanerotomea [Haag-Rutenberg, 1871b: 53]
Type data. Holotype (Paris Museum)
granulifer Haag-Rutenberg, 1871b: 54
Type data. Syntypes (Geneva Museum)
guillarmodi Koch, 1952: 340
Type data. Holotype (Ditsong Museum)
haagi Gebien, 1910b: 156, replacement name
= Psammodes obliteratus Haag-Rutenberg, 1871a: 103 [junior secondary homonym of Hypomelus obliteratus Solier, 1843: 97]
Type data. Holotype (Munich Museum)
hirtipennis Haag-Rutenberg, 1871a: 105
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
hirtipes (Laporte, 1840: 198) Moluris [Gebien, 1910b: 159]
Type data. Holotype (Paris Museum)
= Moluris reichei Solier, 1843: 67 [syn. by
Type data. Holotype (Paris Museum)
hirtus (Bertoloni, 1849: 399) Moluris [Gerstaecker, 1854: 532]
Type data. Holotype (Bologna Museum)
herculeanus Haag-Rutenberg, 1871a: 68, in key
Type data. Syntypes (Naturhistoriska riksmuseet)
Notes. A detailed morphological description was provided by
herero Péringuey, 1908: 409
Type data. Holotype (Cape Museum)
hottentottus Péringuey, 1899: 267
Type data. Holotype (Cape Museum)
incongruens Péringuey, 1899: 281
Type data. Syntypes (Cape Museum)
infernalis Harold, 1878: 106
Type data. Syntypes (Munich Museum)
intermedius Péringuey, 1899: 272
Type data. Holotype (Cape Museum)
janitor Koch, 1953c: 11
Type data. Holotype (Ditsong Museum) and paratypes (Cape Museum, Ditsong Museum, Lund University, Rhodes University)
kamagasus Péringuey, 1908: 409
Type data. Holotype (Cape Museum)
kirschi Haag-Rutenberg, 1871b: 102
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
kubub Péringuey, 1908: 408
Type data. Syntypes (Cape Museum)
kuisip Péringuey, 1908: 504
Type data. Holotype (Cape Museum)
Notes. Originally described under the name Psammodes tuberculifer (intended redescription; page: 407). However, in erratum (page: 504), renamed kuisip.
lanuginosus Haag-Rutenberg, 1871a: 105
Type data. Holotype (Warsaw Museum – Dohrn coll.)
lethargicus Péringuey, 1899: 284
Type data. Holotype (Cape Museum)
laevicollis (Solier, 1843: 65) Moluris [Haag-Rutenberg, 1871a: 78]
Type data. Holotype (Paris Museum)
longicornis Kirby, 1819: 480
Type data. Syntypes (British Museum)
= Phanerotoma ruficore Solier, 1843: 86 [syn. by
Type data. Syntypes (Paris Museum)
longipes Haag-Rutenberg, 1871a: 108
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
lucidus Fåhraeus, 1870: 267
Type data. Holotype (Naturhistoriska riksmuseet)
mashunus Péringuey, 1899: 269
Type data. Syntypes (Cape Museum)
memnonius Haag-Rutenberg, 1871a: 50, in key
Type data. Holotype (British Museum)
Notes. A detailed morphological description was provided by
mimipinguis Koch, 1953c: 9
Type data. Holotype (Lund University) and paratypes (Ditsong Museum, Munich Museum)
moschleri Haag-Rutenberg, 1875: 73
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
muata Harold, 1878: 106
Type data. Syntypes (Munich Museum, Warsaw Museum)
mulleri Péringuey, 1899: 269
Type data. Syntypes (Cape Museum)
nigrisaxicola Koch, 1953b: 78
Type data. Holotype (British Museum)
nitens Fåhraeus, 1870: 267
Type data. Holotype (Naturhistoriska riksmuseet)
nitidicollis Haag-Rutenberg, 1871a: 91
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
nitidipennis (Fairmaire, 1897: 114) Amiantus [Gebien, 1937a: 763]
Type data. Holotype (Paris Museum)
nitidissimus Haag-Rutenberg, 1871a: 92
Type data. Holotype (Warsaw Museum – Dohrn coll.)
obsulcatus Haag-Rutenberg, 1871a: 72, in key
Type data. Holotype (Geneva Museum)
Notes. A detailed morphological description was provided by
ovatus (Solier, 1843: 90) Phanerotoma [Haag-Rutenberg, 1871b: 62]
Type data. Holotype (Paris Museum)
ovipennis Haag-Rutenberg, 1871a: 102
Type data. Holotype (Warsaw Museum – Dohrn coll.)
perfidus Péringuey, 1899: 283
Type data. Holotype (Cape Museum)
piceus Haag-Rutenberg, 1871a: 67, in key
Type data. Syntype (Geneva Museum)
Notes. A detailed morphological description was provided by
pilifer Haag-Rutenberg, 1871a: 69, in key
Type data. Holotype (British Museum – Bates coll.)
Notes. A detailed morphological description was provided by
pilosellus Haag-Rutenberg, 1875: 71
Type data. Syntypes (British Museum – Bates coll.)
pilosipennis Haag-Rutenberg, 1871a: 89
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
pilosus (Thunberg, 1787: 49) Pimelia [Haag-Rutenberg, 1871a: 104]
Type data. Syntypes (Uppsala University)
pinguis (Solier, 1843: 70) Moluris [Haag-Rutenberg, 1871a: 86]
Type data. Holotype (Paris Museum)
= Psammodes rotundipennis Péringuey, 1899: 268 [syn. by
Type data. Holotype (Cape Museum)
placidus Péringuey, 1899: 280
Type data. Syntypes (Cape Museum)
plicatus (Solier, 1844: 72) Moluris [Haag-Rutenberg, 1871a: 95]
Type data. Holotype (Marseille Museum)
plicipennis Gemminger, 1870: 1899, replacement name
= Phanerotoma plicatus Solier, 1844: 299 [homonym of Moluris plicatus Solier, 1844: 284 published on the same date;
Type data. Holotype (Paris Museum)
ponderosus Fåhraeus, 1870: 264
Type data. Syntypes (Cape Museum, Naturhistoriska riksmuseet)
probes (Péringuey, 1899: 296) Psammodophysis [Gebien, 1910b: 159]
Type data. Holotype (Cape Museum)
procerus (Fåhraeus, 1870: 271) Hypomelus [Gebien, 1910b: 159]
Type data. Holotype (Naturhistoriska riksmuseet)
procustes (Westwood, 1875: 224) Moluris [Gebien, 1910b: 159]
Type data. Holotype (Oxford University – Westwood coll.)
= Psammodes giganteus Haag-Rutenberg, 1879: 290 [syn. by
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
productus Haag-Rutenberg, 1871b: 101
Type data. Holotype (Berlin Museum)
profanus Péringuey, 1899: 271
Type data. Holotype (Cape Museum)
propinquus Quedenfeldt, 1885: 5
Type data. Syntypes (Berlin Museum)
protensus Haag-Rutenberg, 1871a: 73, in key
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
pubescens (Solier, 1843: 85) Phanerotomea [Haag-Rutenberg, 1871b: 37]
Type data. Holotype (Paris Museum)
pustulifer Haag-Rutenberg, 1871a: 71, in key
Type data. Syntypes (Naturhistoriska riksmuseet)
Notes. A detailed morphological description was provided by
quadricostatus (Fåhraeus, 1870: 272) Hypomelus [Gebien, 1910b: 159]
Type data. Syntypes (Naturhistoriska riksmuseet)
raucus Haag-Rutenberg, 1875: 159
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
refleximargo (Gebien, 1920: 90) Trachynotidus [Gebien, 1937a: 771]
Type data. Holotype (Hamburg University – Michaelsen coll.)
retrospinosus Haag-Rutenberg, 1871a: 61, in key
Type data. Syntypes (Geneva Museum, Warsaw Museum)
Notes. A detailed morphological description was provided by
rotundicollis Haag-Rutenberg, 1871b: 69
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
rufofasciatus Haag-Rutenberg, 1871a: 96
Type data. Syntypes (Cape Museum, Warsaw Museum)
rufonervosus Haag-Rutenberg, 1871a: 96
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
rufostriatus Haag-Rutenberg, 1875: 70
Type data. Syntypes (British Museum, Cape Museum, Munich Museum – Haag-Rutenberg coll.)
rugulosipennis Haag-Rutenberg, 1871a: 98
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
rugulosus (Solier, 1843: 93) Phanerotomea [Haag-Rutenberg, 1871b: 49]
Type data. Holotype (Paris Museum)
= Psammodes exilis Péringuey, 1899: 280 [syn. by
Type data. Syntypes (Cape Museum)
Notes. Interpreted as a synonym of Psammodes caraboides Haag-Rutenberg, 1871 by
rusticus Péringuey, 1899: 270
Type data. Syntypes (Cape Museum)
scaber (Fabricius, 1775: 251) Pimelia [Haag-Rutenberg, 1871a: 109]
Type data. Holotype (British Museum)
scabratus scabratus (Solier, 1843: 74) Moluris [Haag-Rutenberg, 1871a: 110]
Type data. Holotype (Warsaw Museum – Dupont collection)
scabratus gariepinus Koch, 1953c: 5
Type data. Holotype (Ditsong Museum) and paratypes (Cape Museum, Ditsong Museum)
scabriusculus Haag-Rutenberg, 1871a: 98
Type data. Syntypes (Brussels Museum)
schultzei Peinguey, 1908: 408
Type data. Holotype (Cape Museum)
segnis Haag-Rutenberg, 1871a: 71, in key
Type data. Syntypes (Vienna Museum)
Notes. A detailed morphological description was provided by
sellatus sellatus Haag-Rutenberg, 1875: 72
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
sellatus uriai Koch, 1953b: 75
Type data. Holotype (Ditsong Museum)
semipilosus Haag-Rutenberg, 1871a: 80
Type data. Syntypes (Geneva Museum)
= Psammodes approximans Péringuey, 1899: 270 [syn. by
Type data. Holotype (Cape Museum)
semivillosus Haag-Rutenberg, 1871a: 80
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
setipennis Haag-Rutenberg, 1871a: 107
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
solitarius Péringuey, 1899: 273
Type data. Holotype (Cape Museum)
spiculosus Haag-Rutenberg, 1871a: 111
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
= Psammodes karrooensis Péringuey, 1899: 267 [syn. by
Type data. Syntypes (Cape Museum)
spinosus Haag-Rutenberg, 1871a: 62, in key
Type data. Holotype (Cape Museum)
Notes. A detailed morphological description was provided by
splendens Haag-Rutenberg, 1871a: 73, in key
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
steinhelli Haag-Rutenberg, 1878: 91
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
striatopilosus Haag-Rutenberg, 1871a: 90
Type data. Syntypes (Geneva Museum)
subaeneus Harold, 1878: 106
Type data. Holotype (Munich Museum)
subcostatus (Solier, 1843: 88) Phanerotomea [Haag-Rutenberg, 1871b: 46]
Type data. Holotype (British Museum)
subgranulatus Haag-Rutenberg, 1871a: 78
Type data. Syntypes (Cape Museum, Munich Museum – Haag-Rutenberg coll.)
tenuipes (Fåhraeus, 1870: 273) Hypomelus [Haag-Rutenberg, 1871b: 47]
Type data. Holotype (Cape Museum)
timarchoides Haag-Rutenberg, 1871a: 79
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
togatus Koch, 1953c: 10
Type data. Holotype (Ditsong Museum) and paratypes (Ditsong Museum, Lund University)
tomentosus (Solier, 1843: 73) Moluris [Haag-Rutenberg, 1871a: 93]
Type data. Holotype (Paris Museum)
trachysceloides Haag-Rutenberg, 1871a: 72, in key
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
transvaalensis Haag-Rutenberg, 1875: 81
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
=Psammodes laetulus Péringuey, 1899: 278 [syn. by
Type data. Syntypes (Cape Museum)
tricostatus (Fåhraeus, 1870: 273) Hypomelus [Haag-Rutenberg, 1871b: 48]
Type data. Syntypes (Naturhistoriska riksmuseet)
= Psammodes mendax Péringuey, 1899: 283 [syn. by
Type data. Holotype (Cape Museum)
= Psammodes praestans Péringuey, 1899: 282 [syn. by
Type data. Syntypes (Cape Museum)
Notes. Interpreted as a synonym of Hypomelus tenuipes Fåhraeus, 1870 by
tristis Fåhraeus, 1870: 269
Type data. Syntypes (Naturhistoriska riksmuseet)
tuberculipennis Haag-Rutenberg, 1871a: 60, in key
Type data. Holotype (British Museum)
Notes. A detailed morphological description was provided by
= Psammodes interventor Péringuey, 1908: 410 [syn. by
Type data. Holotype (Cape Museum)
Notes.
tumidipennis Haag-Rutenberg, 1871a: 88
Type data. Syntypes (Geneva Museum, Warsaw Museum – Dohrn coll.)
undulatus Haag-Rutenberg, 1871a: 102
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
uniformis uniformis Haag-Rutenberg, 1871a: 88
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
uniformis uniformis Koch, 1953c: 7
Type data. Holotype (Cape Museum) and paratypes (Ditsong Museum, Lund University, McGregor Museum, Rhodes University)
uniformis rugigaster Koch, 1953c: 7
Type data. Holotype (Cape Museum) and paratypes (Cape Museum, Ditsong Museum, Stellenbosch University, Lund University)
validus Kratz, 1897: 48
Type data. Holotype (Berlin Museum)
velutinus Haag-Rutenberg, 1871a: 87
Type data. Syntypes (Vienna Museum)
ventricosus Fåhraeus, 1870: 264
Type data. Syntypes (Naturhistoriska riksmuseet)
vialis tuberculifer Haag-Rutenberg, 1871a: 60, in key
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
vialis vialis (Burchell, 1822: 305) Moluris [Gebien, 1937a: 760]
Type data. Syntypes (Oxford University – Burchell coll.)
= Moluris pierreti Amyot, 1835: 129 [syn. by
Type data. Syntypes (Paris Museum)
villosostriatus Haag-Rutenberg, 1871a: 87
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
villosulus Haag-Rutenberg, 1871a: 81
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
vittatus (Solier, 1843: 113) Trachynotus [Haag-Rutenberg, 1871a: 85]
Type data. Holotype (Paris Museum)
volvulus Haag-Rutenberg, 1871a: 68, in key
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. A detailed morphological description was provided by
=Psammodes adventitus Péringuey, 1899: 299 [syn. by Péringuey, (1904: 297)]
Type data. Holotype (Cape Museum)
zschokkei Koch, 1953b: 84
Type data. Holotype (Ditsong Museum) and paratypes (Cape Museum, Ditsong Museum)
Type species. Moluris catenata Reiche, 1850 (by original designation)
Notes. Originally described as a subgenus of Psammodes, and elevated to the generic level by
Type species. Psammodes acuticosta Fairmaire, 1884 (by original designation)
acuticosta (Fairmaire, 1884: LXXIV) Psammodes [Koch, 1953a: 155]
Type data. Syntypes (Basel Museum, Cape Museum)
lomii (Gridelli, 1939b: 230) Psammodes [Koch, 1953a: 155]
Type data. Syntypes (Trieste Museum)
Type species. Moluris catenata Reiche, 1850 (by original designation)
angulicauda (Lesne, 1922: 691) Psammodes [Lesne, 1922: 691]
Type data. Syntypes (Paris Museum)
antinorii (Gridelli, 1939b: 234) Psammodes comb. n.
Type data. Syntypes (Trieste Museum)
Notes. This species was unknown to
beccarii beccarii (Gridelli, 1939a: 105) Psammodes [Gridelli, 1939a: 105]
Type data. Holotype (Vienna Museum) and paratype (Monaco Museum)
beccarii sudanicus Koch, 1953a: 169
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Ditsong Museum, Munich Museum)
borosi Koch, 1953a: 173
Type data. Holotype (Brussels Museum) and paratypes (Basel Museum, Brussels Museum, Budapest Museum, Ditsong Museum)
castanopterus (Haag-Rutenberg, 1875: 69) Amiantus [
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
catenatus (Reiche, 1850: 366) Moluris [Lesne, 1922: 692]
Type data. Holotype (Paris Museum)
= Psammodes abyssinicus Haag-Rutenberg, 1871b: 32 [syn. by
Type data. Syntypes (Munich Museum, Warsaw Museum)
camiadei Robiche, 2013: 157
Type data. Holotype (Paris Museum) and paratypes (Paris Museum, Gérard Robiche collection)
densepunctatus Koch, 1953a: 158
Type data. Holotype (Munich Museum) and paratypes (Ditsong Museum, Munich Museum)
duodecimcostatus (Lesne, 1922: 693) Psammodes [Lesne, 1922: 693]
Type data. Syntypes (Paris Museum)
Notes. Originally described as a subspecies of catenatus; status elevated by
granuliger Koch, 1953a: 160
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Ditsong Museum)
gridellii gridellii Koch, 1953a: 164
Type data. Holotype (Munich Museum)
gridellii microsetosus Koch, 1953a: 165
Type data. Holotype (Munich Museum)
gurannicus (Lesne, 1922: 694) Psammodes [Lesne, 1922: 694]
Type data. Holotype (Paris Museum)
impressiventris (Fairmaire, 1897: 115) Psammodes [Gebien, 1937a: 762]
Type data. Holotype (Basel Museum)
kilimandjarus Koch, 1953a: 174
Type data. Holotype (Paris Museum) and paratypes (Munich Museum)
leakeyi Koch, 1953a: 171
Type data. Holotype (Kenya Museum) and paratypes (Ditsong Museum)
mirei (Pierre, 1979: 7) Psammodes comb. n.
Notes. Originally described as Psammodes (Psammophanes) mirei. According to the original description, this species is allied to Psammophanes (Psammophanes) naivashanus (Lesne, 1922). Based on this information, P. mirei is hereby included within subgenus Psammophanes.
Type data. Holotype and paratypes (Paris Museum)
nairobiensis Koch, 1953a: 173
Type data. Holotype (Royan Brussels) and paratypes (Basel Museum, Brussels Museum, Budapest Museum, Ditsong Museum, Munich Museum, Kenya Museum)
naivashanus (Lesne, 1922: 690) Psammodes [Lesne, 1922: 690]
Type data. Syntypes (Budapest Museum, Paris Museum)
pilosiusculus ecostatus (Lesne, 1922: 691) Psammodes [Lesne, 1922: 691]
Type data. Syntypes (Paris Museum)
pilosiusculus pilosiusculus (Gebien, 1913: 60) Psammodes [Gebien, 1937a: 763]
Type data. Holotype (Munich Museum) and paratypes (Basel Museum, Budapest Museum, Munich Museum, Tervuren Museum)
pilosiusculus ruandanus Koch, 1953a: 168
Type data. Holotype (Tervuren Museum)
plicatoides Koch, 1953a: 164
Type data. Holotype and paratype (Kenya Museum)
plicatus aethiopicus Koch, 1953a: 162
Type data. Holotype (Munich Museum)
plicatus multilineatus Koch, 1953a: 161
Type data. Holotype (Kenya Museum) and paratypes (Basel Museum, Budapest Museum, Munich Museum, Kenya Museum)
plicatus plicatus (Gerstaecker, 1871: 59) Phrynocolus [Koch, 1953a: 160]
Type data. Syntypes (Berlin Museum)
plicatus sulcatus
(
= Psammodes plicipennis Fairmaire, 1891b: CCXCIII [junior primary homonym of Psammodes plicipennis Gemminger, 1870: 1899]
Type data. Holotype (Paris Museum)
praetenuis praetenuis Koch, 1953a: 163
Type data. Holotype (Munich Museum) and paratypes (Basel Museum, Budapest Museum, Ditsong Museum, Munich Museum)
praetenuis subtomentosus Koch, 1953a: 163
Type data. Holotype (Munich Museum) and paratypes (Budapest Museum, Munich Museum)
pyriformis (Gridelli, 1939b) Psammodes [Koch, 1953a: 171]
Type data. Syntypes (Trieste Museum)
raffrayi pseudocatenatus Koch, 1953a: 167
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Budapest Museum, Ditsong Museum, Munich Museum)
raffrayi raffrayi (Lesne, 1922: 694) Psammodes [Lesne, 1922: 694]
Type data. Syntypes (Paris Museum)
rubrolineatus (Lesne, 1922: 693) Psammodes [Lesne, 1922: 693]
Type data. Syntypes (Basel Museum)
sexcostatus (Gerstäcker, 1884: 54) Phrynocolus [Lesne, 1922: 691]
Type data. Holotype (Berlin Museum)
somalicus Koch, 1953a: 158
Type data. Holotype (Munich Museum) and paratype (Disong Museum)
terrenus crassecostatus Koch, 1953a: 175
Type data. Holotype (Tervuren Museum) and paratypes (Budapest Museum, Ditsong Museum)
terrenus rugilineatus Koch, 1953a: 176
Type data. Holotype (Tervuren Museum) and paratypes (National Museums of Kenya)
terrenus terrenus (Lesne, 1922: 694) Psammodes [Lesne, 1922: 694]
Type data. Holotype (Paris Museum)
vagecostatus (Fairmaire, 1882b: L) Psammodes [
Type data. Holotype (Paris Museum)
Notes. Treated as a synonym of Amiantus castanopterus Haag-Rutenberg, 1875 by
Type species. Phrynocolus frommi Wilke, 1921 (by original designation)
frommi (Wilke, 1921: 169) Phrynocolus [Koch, 1953a: 157]
Type data. Syntypes (Berlin Museum)
Type species. Psammophanes (Psammophrynus) jokli Koch, 1953 (by original designation)
jokli Koch, 1953a: 152
Type data. Holotype (Brussels Museum) and paratypes (Budapest Museum, Brussels Museum, Ditsong Museum, Munich Museum)
penicillatus penicillatus Koch, 1953a: 153
Type data. Holotype (Tervuren Museum)
penicillatus piacatus Koch, 1953a: 154
Type data. Holotype (Tervuren Museum)
poccilator Koch, 1953a: 153
Type data. Holotype (Tervuren Museum)
Type species. Psammodes bisbicostatus Gebien, 1910 (by original designation)
bisbicostatus bisbicostatus (Gebien, 1910a: 153) Psammodes [Koch, 1953a: 147]
Type data. Syntypes (Basel Museum, Tervuren Museum)
bisbicostatus leleupi Koch, 1953a: 148
Type data. Holotype (Brussels Museum) and paratypes (Basel Museum, Budapest Museum, Ditsong Museum, Tervuren Museum)
circumscriptus Koch, 1953a: 151
Type data. Holotype (Cape Museum) and paratypes (Basel Museum, Brussels Museum, Budapest Museum, Ditsong Museum)
erectepilosus erectepilosus Koch, 1953a: 150
Type data. Holotype (Tervuren Museum) and paratypes (Ditsong Museum, Tervuren Museum)
erectepilosus tanganyikanus Koch, 1953a: 150
Type data. Holotype (Munchen Museum)
maculicollis Koch, 1953a: 150
Type data. Holotype (Brussels Museum) and paratypes (Basel Museum, Budapest Museum, Ditsong Museum, Tervuren Museum)
neavei (Gebien, 1910a: 153) Psammodes [Koch, 1953a: 149]
Type data. Syntypes (Basel Museum, Tervuren Museum)
punctipilus Koch, 1953a: 149
Type data. Holotype (Brussels Museum) and paratype (Tervuren Museum)
prosodoides (Gebien, 1910a: 153) Psammodes [Koch, 1953a: 151]
Type data. Syntypes (Basel Museum, Cape Museum, Tervuren Museum)
Type species. Psammophanes (Psammotyriopsis) bredoi Koch, 1953 (by original designation)
bredoi Koch, 1953a: 144
Type data. Holotype (Budapest Museum) and paratype (Budapest Museum, Royal Brusseles)
Type species. Psammodes gracilentus Fairmaire, 1882 (by original designation)
ahlmedoensis Koch, 1969: 31
Type data. Holotype and paratypes (Munich Museum)
arabicus (Gebien, 1938: 58, in Schuster & Gebien, 1938) Psammodes [Koch, 1953a: 155]
Type data. Syntypes (Hamburg University)
benardellii Koch, 1965: 126
Type data. Holotype (Milan Museum)
gracilentus (Fairmaire, 1882a: 69) Psammodes [Koch, 1953a: 156]
Type data. Syntypes (Paris Museum)
hemmingi Koch, 1969: 25
Type data. Holotype (Munich Museum) and paratypes (Ditsong Museum)
nogalus Koch, 1962c: 242
Type data. Holotype (Milan Museum)
Type species. Psammodes ertli Kolbe, 1904 (by original designation)
Notes. Originally described as a subgenus of Psammodes. Elevated to generic level by
attenuatus attenuatus (Fairmaire, 1887: 180) Moluris [Koch, 1953a: 142]
Type data. Holotype (Paris Museum)
= Moluris tentyrioides Fairmaire, 1891a: 249 [syn. by
Type data. Holotype (Paris Museum)
attenuatus magnophthalmus (Koch, 1953a: 143) Psammodes [Koch, 1953a: 143]
Type data. Syntypes (Basel Museum, Ditsong Museum, Munich Museum)
ertli ertli (Kolbe, 1904: 301) Psammodes [Koch, 1955]
Type data. Syntypes (Basel Museum, Berlin Museum, Tervuren Museum)
ertli punctativentris (Koch, 1953a: 143) Psammodes [Koch, 1953a: 143]
Type data. Holotype (Budapest Museum)
ertli spinosocostatus (Kolbe, 1904: 302) Psammodes [Koch, 1955]
Type data. Syntypes (Berlin Museum)
lateridens lateridens (Fairmaire, 1887: 179) Moluris [Koch, 1953a: 143]
Type data. Holotype (Paris Museum)
lateridens nyassicus (Koch, 1953a: 143) Psammodes [Koch, 1953a: 143]
Type data. Holotype (Budapest Museum)
quadriplicatus (Gebien, 1910b: 159, replacement name) Psammodes [Koch, 1953a: 141]
= Psammodes quadricostatus Fairmaire, 1891b: CCXCIII [junior secondary homonym of Hypomelus quadricostatus Fåhraeus, 1870: 272]
Type data. Holotype (Paris Museum)
Type genus. Oxura Kirby, 1819
Taxonomic diversity. (8 gen., 63 spp.): Decoriplus (11 ssp.), Miripronotum (1), Namibomodes (4), Oxura (9), Palpomodes (4), Pterostichula (17), Stenethmus (11), Synhimba (6).
Distribution. The majority of species were described from Namibia. A small number of species of Stenethmus were described from the northern part of Tanzania, while some species of Decoriplus from Central Africa (Fig.
Type species. Psammodes pictus Haag-Rutenberg, 1871 (by original designation)
aequabilis Louw, 1979: 125
Type data. Holotype (Ditsong Museum) and paratypes (Ditsong Museum and Windhoek Museum)
clavus Louw, 1979: 126
Type data. Holotype and paratypes (Ditsong Museum)
convexus Louw, 1979: 127
Type data. Holotype (Ditsong Museum)
costimargo Louw, 1979: 128
Type data. Holotype (Windhoek Museum) and paratypes (Budapest Museum, Cape Museum, Ditsong Museum, Windhoek Museum)
discicollis Louw, 1979: 130
Type data. Holotype and paratype (Ditsong Museum)
granulimargo Louw, 1979: 131
Type data. Holotype (Ditsong Museum) and paratypes (Ditsong Museum, Windhoek Museum)
hamatus Louw, 1979: 133
Type data. Holotype (British Museum) and paratypes (British Museum, New York Museum)
aequabilis hieroglyphicus (Haag-Rutenberg, 1871a: 69, in key) Psammodes [Louw, 1979: 134]
Type data. Lectotype, designated by
Notes.A detailed morphological description was provided by
humerus Louw, 1979: 136
Type data. Holotype (Ditsong Museum) and paratypes (British Museum, Ditsong Museum, Paris Museum, Windhoek Museum)
pictus (Haag-Rutenberg, 1871a: 69, in key) Psammodes [Louw, 1979: 122]
Notes. A detailed morphological description was provided by
Type data. Holotype (Naturhistoriska riksmuseet)
striatulus Louw, 1979: 138
Type data. Holotype (Ditsong Museum) and paratypes (Budapest Museum, Ditsong Museum, Paris Museum, Windhoek Museum)
Type species. Miripronotum prominoculatum Louw, 1979 (by original designation)
prominoculatum Louw, 1979: 119
Type data. Holotype and paratypes (Windhoek Museum)
Type species. Psammodes serrimargo Gebien, 1938 (by original designation)
maculicollis Koch, 1962b: 111
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Budapest Museum, Ditsong Museum)
rubra Koch, 1962b: 112
Type data. Holotype and paratype (Ditsong Museum)
serrimargo (Gebien, 1938a: 86) Psammodes [Koch, 1952: 221]
Type data. Syntypes (Basel Museum, Bremen Museum, Cape Museum)
zarcoi Koch, 1962b: 110
Type data. Holotype (Ditsong Museum) and paratypes (Budapest Museum, Ditsong Museum)
Type species. Oxura setosa Kirby, 1819 (by monotypy)
= Oxyura Agassiz, 1846: 267 [junior homonym of Oxyura Bonaparte, 1831 (Aves: Anatidae)]
Type species. Oxura setosa Kirby, 1819 (by monotypy)
Notes. Unjustified emendation of Oxura Kirby, 1819.
connexa (Haag-Rutenberg, 1871a: 46, in key) Psammodes [Louw, 1979: 159]
Type data. Holotype (Paris Museum) and paratypes (Cape Museum, Paris Museum)
Notes. A detailed morphological description was provided by
femoralis Haag-Rutenberg, 1871b: 111
Type data. Lectotype, designated by
margoabsoluta margoabsoluta Louw, 1979: 164
Type data. Holotype (Ditsong Museum) and paratypes (Ditsong Museum, Windhoek)
margoabsoluta puncticollis Louw, 1979: 165
Type data. Holotype (Windhoek Museum) and paratypes (Pretoria University, Windhoek Museum)
punctipennis Haag-Rutenberg, 1871b: 111
Type data. Lectotype, designated by
rufotibiata rufotibiata Louw, 1979: 167
Type data. Holotype (Windhoek Museum) and paratypes (Ditsong Museum and Windhoek Museum)
rufotibiata planipennata Louw, 1979: 168
Type data. Holotype and paratypes (Windhoek Museum)
setosa Kirby, 1819: 414
Type data. Lectotype, designated by
vestita Solier, 1843: 119
Type data. Holotype (Torino Museum)
Type species. Psammodes physopterus Gebien, 1920 (by monotypy)
Notes. Originally described as a subgenus of Namibomodes. Elevated to the generic level by
Type species. Psammodes physopterus Gebien, 1920 (by monotypy)
halophilus (Koch, 1958:57) Namibomodes [Koch, 1958: 58]
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
physopterus angolensis (Koch, 1958: 58) (Namibomodes) [Koch, 1958: 58]
Type data. Holotype (Ditsong Museum)
physopterus physopterus (Gebien, 1920) Psammodes [Koch, 1952: 223]
Type data. Holotype (Hamburg University – Michaelsen coll.)
Type species. Namibomodes rudebecki Koch, 1952 (by monotypy)
Notes. Originally described as a subgenus of Namibomodes.
rudebecki (Koch, 1952: 223) Namibomodes [Koch, 1952: 223]
Type data. Holotype (Lund University)
Type species. Pterostichula (Pterostichula) calathoides Koch, 1952 (by original designation)
Type species. Pterostichula (Pterostichula) calathoides Koch, 1952 (by original designation)
aridipaludis Louw, 1979: 146
Type data. Holotype (Cape Museum) and paratype (Cape Museum, Windhoek)
broomoides Koch, 1952: 228
Type data. Holotype (Ditsong Museum) and paratypes (Cape Museum, Ditsong Museum)
calathoides Koch, 1952: 227
Type data. Holotype (Cape Museum) and paratypes (Agricultural Institute, Basel Museum, Budapest Museum, Cape Museum, Ditsong Museum)
diaphana Louw, 1979: 145
Type data. Holotype (Ditsong Museum) and paratypes (Budapest Museum, Ditsong Museum, Windhoek Museum)
dubia Louw, 1979: 141
Type data. Holotype (Windhoek Museum) and paratype (Cape Museum, Windhoek Museum)
ellamariae Koch, 1952: 229
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Budapest Museum, Cape Museum, Ditsong Museum, Munich Museum)
infuscata Koch, 1952: 227
Type data. Holotype (Ditsong Museum)
kung Koch, 1952: 229
Type data. Holotype and paratype (Ditsong Museum)
namaqua Koch, 1952: 228
Type data. Holotype (Ditsong Museum)
quarzophila Koch, 1952: 227
Type data. Holotype (Ditsong Museum) and paratypes (Ditsong Museum, Munich Museum)
solitudo Louw, 1979: 147
Type data. Holotype (Windhoek Museum) and paratype (Cape Museum, Windhoek Museum)
Type species. Pterostichula (Ripicolodes) misanthropa Koch, 1952 (by original designation)
arenicola Koch, 1952: 232
Type data. Holotype (Ditsong Museum)
frontalis Koch, 1952: 232
Type data. Holotype (Ditsong Museum)
misanthropa misanthropa Koch, 1952: 231
Type data. Holotype (Ditsong Museum) and paratypes (Cape Museum, Ditsong Museum, Munich Museum)
misanthropa kunenensis Koch, 1952: 231
Type data. Holotype and paratype (Ditsong Museum)
omurambestris Koch, 1952: 230
Type data. Holotype (Ditsong Museum)
parvicollis Louw, 1979: 153
Type data. Holotype (Ditsong Museum) and paratypes (Cape Museum, Ditsong Museum, Windhoek Museum)
Type species. Psammodes tentyriiniformis Hesse, 1935 (by original designation)
Notes. Classified within Tentyriini by
borealis Kaszab, 1972: 231
Type data. Holotype (Budapest Museum)
impuncticollis Gebien, 1937b: 42
Type data. Holotype and paratype (Basel Museum)
massaicus Kaszab, 1972: 231
Type data. Holotype (Budapest Museum)
orientalis Kaszab, 1972: 232
Type data. Holotype (Budapest Museum)
poggii Ferrer, 2004b: 513
Type data. Holotype (Geneva Museum)
punctipleuris Kaszab, 1972: 233
Type data. Holotype and paratype (Budapest Museum)
punctiventris Genien, 1937b: 43
Type data. Syntypes (Basel Museum)
rhodesianus Kaszab, 1972: 232
Type data. Holotype (Budapest Museum)
szunyoghyi Kaszab, 1972: 230
Type data. Holotype and paratype (Budapest Museum)
tentyriiniformis tentyriiniformis (Hesse, 1935: 546) Psammodes [Gebien, 1937b: 41]
Type data. Holotype (Ditsong Museum) and paratypes (Cape Museum, Ditsong Museum)
tentyriiniformis septentrionalis Gebien, 1937b: 44
Type data. Syntypes (Basel Museum, Budapest Museum)
Type species. Psammodes cordiformis Haag-Rutenberg, 1871 (by original designation)
cordiforme (Haag-Rutenberg, 1871a: 62, in key) Psammodes [Koch, 1952: 219]
Type data. Syntypes (Naturhistoriska riksmuseet)
Notes. A detailed morphological description was provided by
hyalinum hyalinum Koch, 1952: 220
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Budapest Museum, California Academy, Ditsong Museum, Munich Museum)
hyalinum ovambo Koch, 1952: 220
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, California Academy, Ditsong Museum, Munich Museum)
melancholica (Haag-Rutenberg, 1871a: 69, in key) Psammodes [Koch, 1952: 218]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
pruinosum Koch, 1952: 219
Type data. Holotype (Ditsong Museum) and paratypes (Barcelona Museum, Basel Museum, British Museum, Budapest Museum, Ditsong Museum, Munich Museum)
sculpturatum (Haag-Rutenberg, 1871a: 62, in key) Psammodes [Koch, 1952: 219]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
Type genus. Phanerotomea Koch, 1958 [junior objective synonym proposed of Ocnodes]
Taxonomic diversity. (5 gen., 177 spp.): Huilamus (1 sp.), Ocnodes (149), Psammoryssus (1), Stridulomus (1), Tarsocnodes (25).
Distribution. Widely distributed in the southern part of the Afrotropical Realm. Only two species, Ocnodes gridellii (Koch, 1960) and O.
humerangula (Koch, 1952), were described north from the equator. None of the known species were reported from the Eastern Cape (Fig.
Type species. Huilamus welwitschi Koch, 1953 (by original designation)
welwitschi Koch, 1953b: 80
Type data. Holotype (Ditsong Museum) and paratypes (British Museum, Ditsong Museum, Munich Museum)
Type species. Ocnodes scrobicollis Fåhraeus, 1870 (here designated)
Type species. Moluris (Phanerotoma) bertolonii Guérin-Méneville, 1844 (by original designation)
arnoldi arnoldi (Koch, 1952: 313) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Ditsong Museum, Durban Museum, Rhodesia Museum)
arnoldi sabianus (Koch, 1952: 314) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
bertolonii (Guérin-Méneville, 1844: 148) Moluris [Koch, 1952: 314], comb. n.
Type data. Syntypes (Paris Museum)
Notes. While describing this species,
freyi (Koch, 1952: 316) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
guerini guerini (Haag-Rutenberg, 1871a: 71, in key) Psammodes [Koch, 1952: 314], comb. n.
Type data. Syntypes (Geneva Museum)
Notes. A detailed morphological description was provided by
guerini lawrencii (Koch, 1954b: 265) Phanerotomea comb. n.
Type data. Holotype (Cape Museum) and paratypes (Cape Museum)
guerini mancus (
Type data. Holotype (Maputo Museum)
junodi (Péringuey, 1899: 275) Psammodes [Koch, 1952: 314], comb. n.
Type data. Holotype (Cape Museum)
= Psammodes junodi Fairmaire, 1899a: 179 [syn. by
Type data. Holotype (Paris Museum)
Type species. Ocnodes scrobicollis Fåhraeus, 1870 (here designated)
Notes. This genus-group name was treated as a synonym of Psammodes (see
= Phanerotoma Solier, 1843: 82 [junior homonym of Phanerotoma Wesmael 1838: 695 (Insecta: Hymenoptera)]
= Phanerotomea Koch, 1958: 58, syn. n., replacement name
Type species. Phanerotoma elongatum Solier, 1843 (by original designation)
acuductus acuductus (Ancey, 1883: 118) Psammodes comb. n.
Type data. Syntypes (Paris Museum)
acuductus ufipanus (Koch, 1952: 301) Phanerotomea comb. n.
Type data. Holotype (Munich Museum) and paratypes (Ditsong Museum, Munich Museum, Tervuren Museum)
adamantinus (Koch, 1952: 245) Phanerotomea comb. n.
Type data. Holotype (Cape Museum)
argenteofasciatus (Koch, 1953b: 82) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
barbosai (Koch, 1952: 317), comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Cape Museum, Maputo Museum)
basilewskyi (Koch, 1952: 308), comb. n.
Type data. Holotype (Tervuren Museum)
bellmarleyi (Koch, 1952: 305), comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, British Museum, Cape Museum, Ditsong Museum, Durban Museum)
benguelensis (Koch, 1952: 276) Phanerotomea comb. n.
Type data. Holotype (Cape Museum) and paratype (Basel Museum, Cape Museum)
blandus (Koch, 1952: 291) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum)
brevicornis (Haag-Rutenberg, 1875: 79) Psammodes comb. n.
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
= Psammodes rugicollis Kolbe, 1883: 23 [syn. by
Type data. Syntypes (Berlin Museum)
brunnescens brunnescens (Haag-Rutenberg, 1871a: 72, in key) Psammodes comb. n.
Type data. Syntypes (Ditsong Museum, Munich Museum, Warsaw Museum, former Dohrn coll.)
Notes. A detailed morphological description was provided by
Type deposition information after
brunnescens molestus (Haag-Rutenberg, 1875: 75) Psammodes comb. n.
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
buccinator (Koch, 1952: 295) Phanerotomea comb. n.
Type data. Holotype (Brussels Museum) and paratypes (Basel Museum, Ditsong Museum, Brussels Museum, Tervuren Museum)
bushmanicus (Koch, 1952: 257) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
carbonarius (Gerstaecker, 1854: 532) Phanerotomea comb. n.
Type data. Syntypes (Berlin Museum)
cardiopterus (Fairmaire, 1888b: 259) Psammodes comb. n.
Type data. Syntypes (Leiden Museum)
cataractus (Koch, 1952: 290) Phanerotomea comb. n.
Type data. Holotype (Rhodesia Museum)
cinerarius (Koch, 1952: 272) Phanerotomea comb. n.
Type data. Holotype (Basel Museum) and paratype (Basel Museum, Ditsong Museum)
complanatus (Koch, 1952: 299) Phanerotomea comb. n.
Type data. Holotype (Royan Brussels) and paratypes (Basel Museum, Ditsong Museum, Brussels Museum)
concinnus Fåhraeus, 1870
Type data. Syntypes (Geneva Museum)
Notes. Type deposition information after Haag-Rutenberg (1781b).
confertus (Koch, 1952: 275) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum, Royal Museum)
congruens (Péringuey, 1899: 281) Psammodes comb. n.
Type data. Holotype (Cape Museum)
cordiventris (Haag-Rutenberg, 1871a: 74, in key) Psammodes comb. n.
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
crocodilinus (Koch, 1952: 311) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
dimorphus (Koch, 1952: 297) Phanerotomea comb. n.
Type data. Holotype (Basel Museum) and paratypes (Basel Museum, Ditsong Museum)
distinctus (Haag-Rutenberg, 1871a: 62, in key) Psammodes comb. n.
Type data. Syntypes (Basel Museum, British Museum)
Notes. A detailed morphological description was provided by
dolosus (Péringuey, 1899: 291) Psammodes comb. n.
Type data. Holotype (Cape Museum)
dorsocostatus (Gebien, 1910a: 153) Psammodes comb. n.
Type data. Syntypes (Basel Museum, Tervuren Museum)
dubiosus (Péringuey, 1899: 287) Psammodes comb. n.
Type data. Holotype (Cape Museum)
ejectus (Koch, 1952: 239) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Cornell University, Ditsong Museum)
epronoticus (Koch, 1952: 246) Phanerotomea comb. n.
Type data. Holotype (Cape Museum)
erichsoni (Haag-Rutenberg, 1871b: 63) Psammodes comb. n.
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
ferreirae ferreirae (Koch, 1952: 238) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Agricultural Institute, Basel Museum, British Museum, California Academy, Cape Museum, Ditsong Museum)
ferreirae zulu (Koch, 1952: 239) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Durban Museum, Munich Museum)
fettingi (Haag-Rutenberg, 1875: 77) Psammodes comb. n.
Type data. Syntypes (Basel Museum)
Notes. Type deposition information after
fistucans (Koch, 1952: 260) Phanerotomea comb. n.
Type data. Holotype and paratypes (Munich Museum – Haag-Rutenberg coll.)
fraternus (Haag-Rutenberg, 1875: 80) Psammodes comb. n.
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
freudei (Koch, 1952: 303) Phanerotomea comb. n.
Type data. Holotype (Basel Museum) and paratypes (Basel Museum, Ditsong Museum)
fulgidus (Koch, 1952: 294) Phanerotomea comb. n.
Type data. Holotype (Brussels Museum) and paratype (Basel Museum, Brussels Museum, Ditsong Museum)
funestus (Haag-Rutenberg, 1871a: 72, in key) Psammodes comb. n.
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
gemmeulus (Koch, 1952: 287) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Ditsong Museum)
gibberosulus (Péringuey, 1908: 407) Psammodes comb. n.
Type data. Syntypes (Cape Museum)
gibbus (Haag-Rutenberg, 1879: 292) Psammodes comb. n.
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
= Psammodes integer
Type data. Holotype (Cape Museum)
globosus (Haag-Rutenberg, 1871a: 74, in key) Psammodes comb. n.
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
= Psammodes myrmidon Péringuey, 1899: 286 [syn. by
Type data. Holotype (Cape Museum)
granisterna (Koch, 1952: 266) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum)
granulosicollis (Haag-Rutenberg, 1871a: 74, in key) Psammodes comb. n.
Type data. Holotype (British Museum)
Notes. A detailed morphological description was provided by
gridellii (Koch, 1960: 263) Phanerotomea comb. n.
Type data. Holotype and paratypes (Ditsong Museum)
haemorrhoidalis haemorrhoidalis (Koch, 1952: 310) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Munich Museum)
haemorrhoidalis salubris (Koch, 1952: 311) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Munich Museum, Tervuren Museum)
heydeni (Haag-Rutenberg, 1871a: 62, in key) Psammodes comb. n.
Type data. Syntypes (Basel Museum)
Notes. A detailed morphological description was provided by
humeralis (Haag-Rutenberg, 1871a: 62, in key) Psammodes comb. n.
Type data. Syntypes (Naturhistoriska riksmuseet)
Notes. A detailed morphological description was provided by
humerangula (Koch, 1952: 332) Tarsocnodes comb. n.
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
imbricatus (Koch, 1952: 256) Phanerotomea comb. n.
Type data. Holotype (Basel Museum)
imitator imitator (Péringuey, 1899: 289) Psammodes comb. n.
Type data. Syntypes (Cape Museum)
Notes.
imitator invadens (Koch, 1952: 241) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Ditsong Museum)
inflatus (Koch, 1952: 275) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Brussels Museum, Ditsong Museum, Tervuren Museum)
janssensi (Koch, 1952: 292) Phanerotomea comb. n.
Type data. Holotype (Brussels Museum) and paratypes (Basel Museum, Brussels Museum, Ditsong Museum)
javeti (Haag-Rutenberg, 1871a: 74, in key) Psammodes comb. n.
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
kulzeri (Koch, 1952: 273) Phanerotomea comb. n.
Type data. Holotype (Cape Museum)
lacustris (Koch, 1952: 291) Phanerotomea comb. n.
Type data. Holotype (Brussels Museum) and paratypes (Ditsong Museum, Tervuren Museum)
laevigatus (Olivier, 1795: 15) Pimelia comb. n.
Type data. Syntypes (Paris Museum)
= Pimelia marginata Herbst, 1799: 54 [syn. by Haag-Rutenberg, 1871b: 59]
Type data. Syntypes (Berlin Museum)
= Phanerotoma elongatum Solier, 1843: 89 [syn. by Haag-Rutenberg, 1871b: 59]
Type data. Syntypes (Paris Museum)
lanceolatus (Koch, 1953a: 177) Phanerotomea comb. n.
Type data. Holotype (Museum Budapest)
licitus (Peringuey, 1899: 290) Psammodes comb. n.
Type data. Syntypes (Cape Museum)
luctuosus (Haag-Rutenberg, 1871a: 72, in key) Psammodes comb. n.
Type data. Holotype (Munich Museum)
Notes. A detailed morphological description was provided by
luxurosus (Koch, 1952: 276) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum), and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum)
maputoensis (Koch, 1952: 244) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Tervuren Museum)
marginicollis (Koch, 1952: 282) Phanerotomea comb. n.
Type data. Holotype (Cape Museum) and parartype (Ditsong Museum)
martinsi (Koch, 1952: 259) Phanerotomea comb. n.
Type data. Holotype (Munich Museum) and paratypes (Ditsong Museum, Munich Museum)
melleus (Koch, 1952: 248) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Ditsong Museum, Munich Museum)
mendicus estermanni (Koch, 1952: 259) Phanerotomea comb. n.
Type data. Holotype (Basel Museum) and paratypes (Basel Museum, Ditsong Museum)
mendicus mendicus (Péringuey, 1899: 299) Psammodes comb. n.
Type data. Syntypes (Cape Museum)
miles (Péringuey, 1908: 408) Psammodes comb. n.
Type data. Holotype (Cape Museum)
mimeticus (Koch, 1952: 320) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Ditsong Museum, Munich Museum)
misolampoides (Fairmaire, 1888b: 258) Psammodes comb. n.
Type data. Holotype (Leiden Museum)
mixtus (Haag-Rutenberg, 1871a: 74, in key) Psammodes comb. n.
Type data. Syntypes (British Museum)
Notes. A detailed morphological description was provided by
monacha (Koch, 1952: 267) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum)
montanus (Koch, 1952: 283) Phanerotomea comb. n.
Type data. Holotype and paratype (Tervuren Museum)
mozambicus (Koch, 1952: 251) Phanerotomea comb. n.
Type data. Holotype (Basel Museum)
muliebris curtus (Koch, 1952: 286) Phanerotomea comb. n.
Type data. Holotype (Cape Museum) and paratypes (Basel Museum, Cape Museum, Ditsong Museum)
muliebris muliebris (Koch, 1952: 285) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, British Museum, Cape Museum, Ditsong Museum, Munich Museum)
muliebris silvestris (Koch, 1952: 286) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
nervosus (Haag-Rutenberg, 1871a: 74, in key) Psammodes comb. n.
Type data. Holotype (British Museum – Bates coll.)
Notes. A detailed morphological description was provided by
notatum (Thunberg, 1787: 48) Sepidium comb. n.
Type data. Holotype (Uppsala University)
notaticollis (Koch, 1952: 263) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum)
odorans (Koch, 1952: 321) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Budapest Museum, British Museum, California Academy, Ditsong Museum, Munich Museum)
opacus (Solier, 1843: 91) Phanerotomea comb. n.
Type data. Holotype (Warsaw Museum – Dupont collection)
osbecki (Billberg, 1815: 281) Moluris comb. n.
Type data. Lectotype, designated by
= Phanerotoma suturalis Solier, 1843: 92 [syn. by
Type data. Holotype (Paris Museum)
overlaeti (Koch, 1952: 274) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratype (Basel Museum, Ditsong Museum, Tervuren Museum)
ovulus (Haag-Rutenberg, 1871a: 73, in key) Psammodes comb. n.
Type data. Holotype (Geneva Museum)
Notes. A detailed morphological description was provided by
= Psammodes probus Péringuey, 1899: 283 [syn. by
Type data. Holotype (Cape Museum)
= Psammodes consors Péringuey, 1899: 288 [syn. by
Type data. Holotype (Cape Museum)
pachysoma ornata (Koch, 1952: 261) Phanerotomea comb. n.
Type data. Holotypes (Munich Museum)
pachysoma pachysoma (Péringuey, 1892: 52) Psammodes comb. n.
Type data. Syntypes (Cape Museum)
papillosus (Koch, 1952: 302) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
pedator (Fairmaire, 1888b: 257), comb. nov. Psammodes
Type data. Syntypes (Leiden Museum)
perlucidus (Koch, 1952: 246) Phanerotomea comb. n.
Type data. Holotype (Cape Museum) and paratypes (Basel Museum, Cape Museum, Ditsong Museum)
planus (Koch, 1952: 307) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum)
pretorianus (Koch, 1952: 279) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
procursus (Péringuey, 1899: 279) Psammodes comb. n.
Type data. Syntypes (Cape Museum)
procrustes (Westwood, 1875: 224) Moluris [Westwood, 1875: 224]
Type data. Holotype (Oxford University – Westwood coll.)
protectus (Koch, 1952: 323) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, California Academy, Cape Museum, Ditsong Museum, Dundo Museum, Munich Museum)
punctatissimus (Koch, 1952: 304) Phanerotomea comb. n.
Type data. Holotype (Brussels Museum) and paratypes (Basel Museum, Brussels Museum, Ditsong Museum)
puncticollis (Koch, 1952: 264) Phanerotomea comb. n.
Type data. Holotype (Brussels Museum) and paratypes (Basel Museum, Brussels Museum, Ditsong Museum, Tervuren Museum)
punctipennis planisculptus (Koch, 1952: 265) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum)
punctipennis punctipennis (Harold, 1878: 106) Psammodes comb. n.
Type data. Holotype (Munich Museum)
punctipleura (Koch, 1952: 306) Phanerotomea comb. n.
Type data. Holotype (Munich Museum) and paratype (Ditsong Museum)
rhodesianus (Koch, 1952: 292) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratype (Basel Museum, Cape Museum)
roriferus (Koch, 1952: 253) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, California Academy, Cape Museum, Ditsong Museum, Munich Museum)
rowleianus (Westwood, 1864: 8979) Moluris [Westwood, 1864: 8979]
Type data. Syntypes (British Museum)
= Psammodes zoutpansbergianus Péringuey, 1904: 231 [syn.
Type data. Holotype (Cape Museum)
rufipes (Harold, 1878: 106) Psammodes comb. n.
Type data. Syntypes (Munich Museum)
= Psammodes congoanus Gebien, 1920b: 7 [syn. by
Type data. Syntypes (Munich Museum)
saltuarius (Koch, 1952: 250) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
scabricollis (Gerstaecker, 1854: 532) Phanerotomea comb. n.
Type data. Holotype (Berlin Museum)
scopulipes (Koch, 1952: 319) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Ditsong Museum, Munich Museum)
scrobicollis griqua (Koch, 1952: 234) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Cape Museum, Ditsong Museum)
scrobicollis scrobicollis Fåhraeus, 1870: 270
Type data. Holotype (Naturhistoriska riksmuseet) and paratypes (Naturhistoriska riksmuseet, Warsaw Museum)
scrobicollis simulans (Koch, 1952: 235) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Agricultural Institute, Basel Museum, Cape Museum, Ditsong Museum, Durban Museum, Munchen Museum, Pretoria University)
semirasus (Koch, 1952: 296) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum)
semiscabrum (Haag-Rutenberg, 1871a: 73, in key) Psammodes comb. n.
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. Type deposition information after
= Psammodes sperabilis Peinguey, 1899: 289 [syn. by
Type data. Holotypes (Cape Museum)
sericicollis (Koch, 1952: 254) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Ditsong Museum, Munich Museum)
similis (Péringuey, 1899: 291) Psammodes comb. n.
Type data. Syntypes (Cape Museum)
sjoestedti (Gebien, 1910b: 372) Psammodes comb. n.
Type data. Holotype (Munich Museum)
spatulipes (Koch, 1952: 318) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum)
specularis (Péringuey, 1899: 286) Psammodes comb. n.
Type data. Syntypes (Cape Museum, Munich Museum)
spinigerus (Koch, 1952: 269) Phanerotomea comb. n.
Type data. Holotype (California Academy) and paratypes (Basel Museum, California Academy, Ditsong Museum)
stevensoni (Koch, 1952: 284) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratype (Basel Museum, Ditsong Museum)
tarsocnoides (Koch, 1952: 267) Phanerotomea comb. n.
Type data. Holotype (Basel Museum) and paratypes (Basel Museum, Ditsong Museum)
temulentus (Koch, 1952: 284) Phanerotomea comb. n.
Type data. Holotype (Basel Museum) and paratype (Basel Museum, Cape Museum)
tenebrosus melanarius (Haag-Rutenberg, 1871a: 71, in key) Psammodes comb. n.
Type data. Syntypes (Geneva Museum)
Notes. A detailed morphological description was provided by
tenebrosus tenebrosus (Erichson, 1843: 242) Moluris comb. n.
Type data. Syntypes (Humboldt University)
tibialis (Haag-Rutenberg, 1871a: 63, in key) Psammodes comb. n.
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
torosus (Koch, 1952: 235) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Agricultural Institute, Basel Museum, Cape Museum, Ditsong Museum, Munich Museum, Pretoria University)
transversicollis (Haag-Rutenberg, 1879: 291) Psammodes comb. n.
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
= Psammodes cinctipennis Fairmaire, 1899: 180 [syn. with P. valens by
Type data. Syntypes (Paris Museum)
= Psammodes valens Péringuey, 1899: 276 [syn. by
Type data. Syntypes (Cape Museum)
tumidus (Haag-Rutenberg, 1871a: 73, in key) Psammodes comb. n.
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
Notes. A detailed morphological description was provided by
umvumanus (Koch, 1952: 280) Phanerotomea comb. n.
Type data. Holotype (Cape Museum)
vagus (Péringuey, 1899: 288) Psammodes comb. n.
Type data. Syntypes (Cape Museum)
vaticinus (Péringuey, 1899: 279) Psammodes comb. n.
Type data. Holotype (Cape Museum)
verecundus (Péringuey, 1899: 286) Psammodes comb. n.
Type data. Syntypes (Cape Museum)
vetustus (Koch, 1952: 263) Phanerotomea comb. n.
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Tervuren Museum)
vexator (Péringuey, 1899: 287) Psammodes comb. n.
Type data. Holotype (Cape Museum)
virago (Koch, 1952: 236) Phanerotomea comb. n.
Type data. Holotype (Ditsong Museum) and paratypes (Agricultural Institute, Basel Museum, California Academy of Sciences, Cape Museum, Cornell University, Ditsong Museum, Munich Museum, Pretoria University, Rhodesia Museum)
warmeloi (Koch, 1953b: 86) Phanerotomea comb. n.
Type data. Holotype (Cape Museum) and paratypes (Basel Museum, Cape Museum, Ditsong Museum)
zanzibaricus (Haag-Rutenberg, 1875: 78) Psammodes comb. n.
Type data. Holotype (Cape Museum)
Type species. Psammoryssus titanus Kolbe, 1886 (by monotypy)
titanus Kolbe, 1886: 290
Type data. Syntypes (Berlin Museum)
Type species. Psammodes sulcicollis Péringuey, 1885 (by monotypy)
sulcicollis (Péringuey, 1885: 110) Psammodes [Koch, 1955: 37]
Type data. Syntypes (Cape Museum)
= Psammodes rehbocki Kolbe, 1904: 299 [syn. by
Type data. Syntypes (Berlin Museum)
Type species. Psammodes molossa Haag-Rutenberg, 1871 (by original designation)
albarenarum Penrith, 1987: 252
Type data. Holotype (Bloemfontein Museum) and paratypes (British Museum, Ditsong Museum, Pretoria University)
aquamontis Penrith, 1987: 249
Type data. Holotype (Ditsong Museum) and paratypes (British Museum, Ditsong Museum, Windhoek Museum)
brendelli Penrith, 1987: 240
Type data. Holotype (British Museum) and paratypes (British Museum, Ditsong Museum)
albarenarumcompressitarsis (Müller, 1887: 299) Psammodes [Gebien, 1920: 83]
Type data. Lectotype, designated by
albarenarumdilaticollis (Muller, 1887: 298) Psammodes [Gebien, 1937a: 759]
Type data. Lectotype, designated by
ephialtes Koch, 1952: 334
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, British Museum, Cape Museum, Ditsong Museum, Munich Museum)
errans (Péringuey, 1892: 53) Psammodes [Gebien, 1920: 83]
Type data. Syntypes (Cape Museum)
Notes. In his original description,
= Tarsocnodes spectabilis Gebien, 1920: 84 [syn. by
Type data. Syntypes (Hamburg University)
Notes. Type deposition information after
finitima Koch, 1952: 331
Type data. Holotype (Ditsong Museum)
gracilipes Koch, 1952: 332
Type data. Holotype (Ditsong Museum)
Notes. Type deposition information after
granulicauda Penrith, 1987: 242
Type data. Holotype (Ditsong Museum) and Paratypes (British Museum, Ditsong Museum, Windhoek Museum)
iflundi Koch, 1952: 335
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Cape Museum, Ditsong Museum, California Academy)
ephialteslaevipennis (Haag-Rutenberg, 1879: 291) Psammodes [Koch, 1962d: 334]
Type data. Lectotype, designated by
madida Koch, 1952: 326
Type data. Holotype (Ditsong Museum)
michaelis Penrith, 1986: 247
Type data. Holotype (Windhoek Museum) and parartypes (Ditsong Museum, Windhoek Museum)
molossa (Haag-Rutenberg, 1871a: 83) Psammodes [Gebien, 1920: 83]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
= Moluris (Phanerotoma) gravida (Westwood, 1875: 223) [syn. by.
Type data. Syntypes (British Museum)
monasterialis Koch, 1952: 329
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Companhia Diamantes, Ditsong Museum, Dundo Museum)
= Tarsocnodes variabilis dissoluta Koch, 1952: 330 [syn. by
Type data. Holotype (Tervuren Museums) and paratypes (Ditsong Museum)
= Tarsocnodes variabilis variabilis Koch, 1952: 330 [syn. by
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Ditsong Museum, Munich Museum, Tervuren Museum)
nielseni Ferrer, Evanno & Evanno, 2010: 195
Type data. Holotype (Naturhistoriska riksmuseet) and paratype (Ferrer collection)
praegrandis horribillis Koch, 1952: 328
Type data. Holotype (Ditsong Museum)
praegrandis praegrandis Koch, 1952: 327
Type data. Holotype (Tervuren Museum) and paratypes (Basel Museum, Brussels Museum, Companhia Diamantes, Dundo Museum, Munich Museum, Tervuren Museum)
prozeskyorum Penrith, 1986: 254
Type data. Holotype (Ditsong Museum) and paratypes (British Museum, Cape Museum, Ditsong Museum)
rugicollis Gebien, 1920: 85
Type data. Holotype and paratypes (Munich Museum)
Notes. Type deposition information after
tarsalis (Haag-Rutenberg, 1871a: 52) Psammodes [Gebien, 1920: 83]
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. Type deposition information after
variolata Koch, 1952: 329
Type data. Holotype (Tervuren Museum)
vernayi Koch, 1952: 250
Type data. Holotype (Ditsong Museum)
whiteheadi Penrith, 1986: 239
Type data. Holotype and paratypes (Windhoek Museum)
Type genus. Sepidium Fabricius, 1775
Taxonomic diversity. (8 gen., 124 spp.): Dimoniacis (3 ssp.), Echinotus (2), Peringueyia (1), Sepidiopsis (3), Sepidiostenus (7), Sepidium (51), Vieta (52), Vietomorpha (5).
Distribution. Widely distributed throughout the Mediterranean area and Sub-Saharan Africa, except its western part. Majority of the species were described form from the Horn of Africa. Vieta representatives are the only species, which were described from the area south from equator, while only Sepidium species have loci tipici north from Tropic of Cancer (Fig.
Type species. Dimoniacis jacksoni Koch, 1958 (by original designation)
jacksoni Koch, 1958: 44
Type data. Holotype (Ditsong Museum)
lavranosi Ardoin, 1979: 60
Type data. Holotype and paratypes (Paris Museum)
puccionii Ferrer, 1995: 27
Type data. Holotype (Florence Museum)
Type species. Sepidium spinicollis Laporte, 1840 (by original designation)
Notes. During the compilation of this catalog, a junior homonym of Echinotus Solier, 1843 was found: Echinotus Marwick, 1935: 301 (Type species. Avicula echinata Smith, 1817; Mollusca: Pteriidae). Ulamus Kaminski, nom. nov. is introduced here as a replacement name for the above-mentioned pteriid genus. This newly introduced name honours Stanisław Marcin Ulam, Polish-American scientist, inventor of the Monte Carlo method of computation.
natalensis Chevrolat, 1874: 331
Type data. Holotype (Paris Museum)
spinicollis (Laporte, 1840: 197) Sepidium [Solier, 1843: 31]
Type data. Syntypes (Paris Museum)
Type species. Echinotus dispar Péringuey, 1899 (by monotypy)
dispar (Péringuey, 1899: 302) Echinotus [Koch, 1958: 44]
Type data. Holotype (Cape Museum)
Type species. Sepidiopsis cornigera Gestro, 1892 (by original designation)
ardoini Ferrer, 1995: 26
Type data. Holotype (Florence Museum)
cornigera Gestro, 1892: 772
Type data. Holotype (Genoa Museum)
villosa Gestro, 1892: 773
Type data. Holotype (Genoa Museum)
Type species. Sepidiostenus erinaceus Fairmaire, 1884 (by monotypy)
= Sepidiacis Fairmaire, 1884: CXLVI [junior subjective synonym proposed by
Type species. Sepidiacis compressa Fairmaire, 1884 (subsequent designation by
Notes. This taxon was redescribed as new by
compressus (Fairmaire, 1884: CXLVI) Sepidiacis
Type data. Holotype (Paris Museum)
dolichopus Gestro, 1898: 517
Type data. Holotype (Genoa Museum)
erinaceus Fairmaire, 1887: 185
Type data. Holotype (Paris Museum)
fairmairei Gestro, 1898: 512
Type data. Holotype (Genoa Museum)
longipennis Gestro, 1898: 516
Type data. Syntypes (Genoa Museum)
pradieri (Guérin-Méneville, 1858: 128) Sepidium [Gestro, 1892: 776]
Type data. Syntypes (Paris Museum)
ruspolii Gestro, 1898: 514
Type data. Holotype (Genoa Museum)
Type species. Sepidium tricuspidatum Fabricius, 1775 (by subsequent designation by Latreille (1810: 429))
= Espidium Rafinesque, 1815: 113
Notes. Unnecessary replacement name for Sepidium Fabricius, 1775
aitagiae Escalera, 1913: 41
Type data. Holotype (Madrid Museum)
aliferum Erichson, 1841: 178
Type data. Holotype (British Museum)
= Sepidium douei Solier, 1843: 18 [syn. by
Type data. Syntypes (Paris Museum)
aper Fairmaire, 1884: LXXV
Type data. Holotype (Paris Museum)
barbarum Solier, 1843: 23
Type data. Syntypes (Warsaw Museum – Dupont collection)
= Sepidium servillei Solier, 1843: 24 [syn. by
Type data. Holotype (Paris Museum)
= Sepidium pallens Allard, 1874: 137 [syn. by
Type data. Holotype (Paris Museum)
= Sepidium barbarum solieri Desbrochers des Loges, 1881: 101 [syn. by
Type data. Syntypes (Paris Museum)
bicaudatum Fairmaire, 1871: 388
Type data. Holotype (Paris Museum)
bidentatum Solier, 1843: 15
Type data. Syntypes (Paris Museum)
bilobatum Gahan, 1900: 30
Type data. Holotype (British Museum)
boranum Mal, 1986b: 151
Type data. Holotype (Trieste Museum) and paratypes (Berlin Museum, British Museum, Budapest Museum, Tervuren Museum, Trieste Museum)
bulbiferum Gerstaecker, 1884: 55
Type data. Syntypes (Berlin Museum)
brevicaudatum Fairmaire, 1882b: LI
Type data. Holotype (Paris Museum)
capricorne Desbrochers des Loges, 1881: 96
Type data. Holotype (Paris Museum)
Notes. Species concept after
crassicaudatum Gestro, 1878: 320
Type data. Holotype (Genoa Museum)
cristatum Fabricius, 1775: 250
Type data. Syntypes (Copenhagen Museum, Kiel Museum)
= Tenebrio notoceros Pallas, 1781: 59 [syn. by
Type data. Syntypes (Humboldt University – Pallas collection)
cylindrigerum Fairmaire, 1882a: 75
Type data. Syntypes (Paris Museum)
cyrenaicum Schuster, 1928: 122
Type data. Syntypes (Basel Museum)
dathan Crotch, 1872: 268
Type data. Holotype (Cambridge Museum)
= Sepidium abiram Crotch, 1872: 268 [syn. by
Type data. Holotype (Cambridge Museum)
= Sepidium vietaeformis Reitter, 1914: 385 [syn. by
Type data. Syntypes (Basel Museum, Budapest Museum, Vienna Museum)
fusiforme Kwieton, 1980: 17
Type data. Holotype and paratype (Budapest Museum)
gypsicola Escalera, 1913: 42
Type data. Syntypes (Madrid Museum)
hoseini Escalera, 1911: 301
Type data. Holotype (Madrid Museum)
hystryx desertica Espanol, 1944: 12
Type data. Holotype (Barcelona Museum)
hystryx hystryx Antoine, 1932: 185
Type data. Holotype (Paris Museum)
hystryx ifniensis Escalera, 1940: 5
Type data. Syntypes (Madrid Museum)
Notes.
inaequale Reitter, 1914: 386
Type data. Syntyptes (Budapest Museum)
kaszabi Mal, 1990: 64
Type data. Holotype and paratype (British Museum)
kelleri Fairmaire, 1893: 151
Type data. Holotype (Paris Museum)
lusitanicum Kaszab & Pinheiro, 1972
Type data. Holotype (Budapest Museum) and paratype (Madrid Museum)
magnum Gahan, 1900: 29
Type data. Holotype and paratypes (British Museum)
mali Ferrer & Martínez, 2012, replacement name
= Sepidium elongatum Mal, 1984: 200 [junior primary homonym of Sepidium elongatus Olivier, 1795: 8]
Type data. Holotype (Brussels Museum) and paratypes (British Museum, Brussels Museum, Julio Ferrer collection)
marraquense Escalera, 1911: 302
Type data. Syntypes (Basel Museum, Madrid Museum)
mesopotamicum Reitter, 1914: 386
Type data. Syntypes (Basel Museum, Budapest Museum, Vienna Museum)
mskalicum Escalera, 1914: 307
Type data. Syntypes (Madrid Museum)
obtusangulum Fairmaire, 1882a: 73
Type data. Holotype (Paris Museum)
pagesii Fairmaire, 1894: 321
Type data. Syntypes (Basel Museum, British Museum, Budapest Museum)
penicilligerum Karsch, 1881: 49
Type data. Syntypes (Berlin Museum)
perforatum Allard, 1874: 130
Type data. Holotype (Paris Museum)
Notes. Species concept after
peyerimhoffi Antoine, 1932: 183
Type data. Syntypes (Basel Museum, Budapest Museum, Paris Museum)
reichei Allard, 1870: 49
Type data. Syntypes (Paris Museum)
= Sepidium reichei bispinicollis Reitter, 1914: 389 [syn. by
Type data. Syntypes (Budapest Museum)
requieni Solier, 1843: 29
Type data. Holotype (Paris Museum)
ruspolii ruspolii Fairmaire, 1893: 150
Type data. Holotype (Paris Museum) and paratype (Basel Museum)
ruspolii spectabile Kulzer, 1960: 309
Type data. Holotype (Berlin Museum) and paratypes (Basel Museum)
scebelianum Mal, 1986b: 153
Type data. Holotype (Trieste Museum) and paratypes (Berlin Museum, British Museum, Budapest Museum, Tervuren Museum, Trieste Museum)
siculum Solier, 1843: 19
Type data. Syntypes (Paris Museum)
= Sepidium genei Solier, 1843: 20 [syn. by
Type data. Syntypes (Paris Museum)
tricuspidatum korah Crotch, 1872: 268
Type data. Holotype (Cambridge Museum)
tricuspidatum mogadoricum Escalera, 1914: 306
Type data. Syntypes (Madrid Museum)
= Sepidium immundum Reitter, 1914: 388 [syn. by
Type data. Syntypes (Basel Museum, Budapest Museum)
= Sepidium mogadoricum guisseri Kocher, 1958: 111 [syn. by
Type data. Syntypes (Rabat Institute)
= Sepidium mogadoricum schrammi Kocher, 1958: 110 [syn. by
Type data. Syntypes (Rabat Institute)
tricuspidatum multispinosum Solier, 1843: 26
Type data. Holotype (Marseille Museum)
= Sepidium laghoatense Baudi, 1875: 695 [syn. by
Type data. Syntypes (Basel Museum)
tricuspidatum tomentosum Erichson, 1841: 179
Type data. Syntypes (Humboldt University)
= Sepidium barthelemyi Solier, 1843: 24 [syn. by
Type data. Syntypes (Marseille Museum)
= Sepidium maillei Solier, 1843: 27 [syn. by
Type data. Syntypes (Marseille Museum)
= Sepidium serratum Solier, 1843: 28 [syn. by
Type data. Holotype (Paris Museum)
= Sepidium serratum remotum Sahlberg, 1903: 50 [syn. By
Type data. Syntypes (Helsinki University)
tricuspidatum tricuspidatum Fabricius, 1775: 250
Type data. Syntypes (Copenhagen Museum, Kiel Museum)
= Tenebrio alexandrinus Forskål, 1775: 80 [
Type data. Syntypes (Copenhagen Museum)
= Sepidium flexuosum Solier, 1843: 25 [syn. by
Type data. Holotype (Paris Museum)
= Sepidium tricuspidatum cerisyi Solier, 1843: 26 [syn. by
Type data. Syntypes (Marseille Museum)
Notes. Originally described as a variety of the nominotypical form.
tualensis Escalera, 1940: 9
Type data. Syntypes (Madrid Museum)
uncinatum Erichson, 1841: 178
Type data. Holotype (Humboldt University)
= Sepidium mittrei Solier, 1843: 16 [syn. by
Type data. Syntypes (Paris Museum)
= Sepidium mittrei bicorne Solier, 1843: 17 [syn. by
Type data. Syntypes (Paris Museum)
Notes. Originally described as a variety of the nominotypical form.
= Sepidium subcornutum Escalera, 1925: 376 [syn. by
Type data. Syntypes (Madrid Museum)
= Sepidium sefrense Rotrou, 1943: 235 [syn. by
Type data. Holotype (Paris Museum)
variegatum (Fabricius, 1792: 112) Tenebrio [
Type data. Syntypes (British Museum, Kiel Museum)
= Sepidium variegatum angustatum Solier, 1843: 21 [syn. by Reitter (1916: 383)]
Type data. Syntypes (Paris Museum)
= Sepidium dufouri Solier, 1843: 21 [syn. by
Type data. Holotype (Paris Museum)
= Sepidium laterale Allard, 1874: 133 [syn. by
Type data. Syntypes (Paris Museum)
= Sepidium variegatum dispar Desbrochers des Loges, 1881: 101 [syn. by Reitter (1916: 383)]
Type data. Syntypes (Paris Museum)
= Sepidium variegatum integrum Desbrochers des Loges, 1881: 100 [syn. by
Type data. Syntypes (Paris Museum)
= Sepidium variegatum subfurcatum Desbrochers des Loges, 1881: 100 [syn. by
Type data. Syntypes (Paris Museum)
wagneri Erichson, 1841: 179
Type data. Syntypes (Humboldt University)
= Sepidium wagneri confusum Allard, 1874: 140 [syn. by
Type data. Syntypes (Paris Museum)
= Sepidium wagneri macrotys Antoine, 1951a: 94 [syn. by
Type data. Syntypes (Paris Museum)
Type species. Sepidium vestitum Guérin-Méneville, 1831 (designated by Hope (1841: 116))
= Dymonus Solier, 1843: 7 [junior subjective synonym proposed by
Type species. Sepidium vestitum Guérin-Méneville, 1831 (by original designation)
= Divieta Reitter, 1914: 390 [junior subjective synonym proposed by
Type species. Vieta costata Allard, 1874 (by subsequent designation by
algeriana Allard, 1870: 50
Type data. Syntypes (Paris Museum)
angolensis angolensis (Quedenfeldt, 1885: 7) Sepidium [Koch, 1958: 53]
Type data. Syntypes (Humboldt University, Paris Museum)
angolensis biena Koch, 1958: 53
Type data. Holotype and paratypes (California Academy)
angolensis eduardi Koch, 1958: 54
Type data. Holotype and paratypes (California Academy)
angolensis transversa (Fairmaire, 1888b: 262) Sepidium [Koch, 1958: 53]
Type data. Holotype (Leiden Museum)
= Sepidium ovampoense Péringuey, 1892: 55 [syn. by
Type data. Holotype (Cape Museum)
angustula Lesne, 1922: 697
Type data. Holotype (Paris Museum)
aper Fairmaire, 1887: 278
Type data. Holotype (Paris Museum)
apicorne (Fairemaire, 1882: 75) Sepidium [Koch, 1958: 42]
Type data. Holotype (Paris Museum)
borana Gridelli, 1939a: 114
Type data. Syntypes (Trieste Museum)
bulbifera Fairmaire, 1897: 116
Type data. Holotype (Paris Museum)
clypeata Gahan, 1896: 454
Type data. Holotype (British Museum)
cornutipennis Gebien, 1937b: 37
Type data. Holotype (Trieste Museum) and paratype (Basel Museum, Trieste Museum)
costata Allard, 1874: 149
Type data. Holotype (Basel Museum)
crinita Allard, 1882: LXXXVII
Type data. Syntypes (Paris Museum)
= Sepidium zambezianum Péringuey, 1892: 123 [syn. by
Type data. Syntypes (British Museum and Paris Museum)
deckerti Ferrer, 2004a: 217
Type data. Holotype (Berlin Museum)
erecticollis Ancey, 1881b: 461
Type data. Syntypes (Paris Museum)
erosa Allard, 1882: LXXXVII
Type data. Holotype (Basel Museum)
furcifera (Gerstaecker, 1884: 55) Sepidium [Gebien, 1937b: 56]
Type data. Syntypes (Berlin Museum)
gracilenta Ancey, 1881a: 397
Type data. Syntypes (Paris Museum)
grisea Gridelli, 1939a: 421
Type data. Syntypes (Trieste Museum)
grixonii Gestro, 1895: 375
Type data. Holotype (Genoa Museum)
hamaticollis (Fairmaire, 1887: 277) Sepidium [Koch, 1958: 43]
Type data. Holotype (Paris Museum)
holdhausi Reitter, 1914: 391
Type data. Holotype (Vienna Museum)
lacunosa Fairmaire, 1894: CCLII
Type data. Holotype (Basel Museum)
longehirta (Fairmaire, 1887: 277) Sepidium [Koch, 1958: 43]
Type data. Holotype (Paris Museum)
longepilosa Fairmaire, 1891b: CCXCIV
Type data. Holotype (Paris Museum)
luctuosa Fairmaire, 1894: 392
Type data. Holotype (Paris Museum)
lutulenta Gestro, 1895: 143
Type data. Holotype (Genoa Museum)
luxorii Allard, 1874: 150
Type data. Holotype (Paris Museum)
millingenii Kirchsberg, 1877: 203
Type data. Syntypes (Basel Museum)
montana Fairmaire, 1894: 392
Type data. Holotype (Paris Museum)
muscosa (Gerstaecker, 1871: 60) Sepidium [Koch, 1958: 43]
Type data. Syntypes (Berlin Museum)
ovalis Allard, 1874: 149
Type data. Syntypes (Basel Museum, British Museum)
pallidicornis Koch, 1958: 219
Type data. Holotype (Cape Museum) and paratypes (Cape Museum, Ditsong Museum)
protensa Fairmaire, 1891b: CCXCV
Type data. Syntypes (Paris Museum)
punctipennis Reitter, 1914: 390
Type data. Holotype (Vienna Museum)
ramosipilus Koch, 1958: 219
Type data. Syntypes (Cape Museum, Ditsong Museum)
rendiliana Lesne, 1922: 696
Type data. Holotype (Paris Museum)
robusta Lesne, 1922: 697
Type data. Holotype (Paris Museum)
russoi Gebien, 1937b: 38
Type data. Holotype (Trieste Museum) and paratype (Basel Museum, Museo Civico Filangieri)
senegalensis dongolensis Laporte, 1840: 197
Type data. Holotype (Paris Museum)
Notes. Taxonomic concept after
= Dymonus dufossei Solier, 1843: 10 [syn. by
Type data. Holotype (Paris Museum)
senegalensis senegalensis (Klug, 1835: 40)
Type data. Syntypes (Humboldt University)
= Sepidium vestitum Guérin-Méneville, 1831: 114 [syn. by
Type data. Syntypes (Paris Museum)
sexcornuta (Fairmaire, 1897: 115) Sepidium [Koch, 1958: 42]
Type data. Holotype (Paris Museum)
speculifera Gebien, 1910a: 158
Type data. Syntypes (Basel Museum, Tervuren Museum)
spiculosa (Gerstaecker, 1884: 55) Sepidium [Gebien, 1937b: 56]
Type data. Holotype (Berlin Museum)
subcaudata Lesne, 1922: 696
Type data. Holotype (Paris Museum)
tuberculata (Solier, 1844: 11) Dymonus [Koch, 1958: 43]
Type data. Holotype (Torino Museum – Spinola coll.)
= Dymonus gibbicollis Solier, 1843: 12 [syn. by
Type data. Holotype (Warsaw Museum – Dupont collection)
tuberosa Fairmaire, 1882a: 76
Type data. Holotype (Paris Museum)
uncigera Ancey, 1881b: 461
Type data. Holotype (Paris Museum)
vietomorphoides Kwieton, 1978: 11
Type data. Holotype (Prague Museum)
villosula (Fairmaire, 1882a: 74) Sepidium [Koch, 1958: 42]
Type data. Holotype (Paris Museum)
zavatartii Gridelli, 1939a: 110
Type data. Syntypes (Trieste Museum)
Type species. Vietomorpha foveipennis Fairmaire, 1887 (by monotypy)
abyssinica Mal, 1986a: 17
Type data. Holotype (Brussels Museum) and paratypes (British Museum, Budapest Museum, Ditsong Museum, Ohio State, Tervuren Museum, Paris Museum)
bartolozzii Mal, 1986a: 19
Type data. Holotype and paratype (Florence Museum)
crassipes Mal, 1986a: 16
Type data. Holotype (Ditsong Museum) and paratypes (British Museum, Budapest Museum, Ditsong Museum, Paris Museum)
foveipennis Fairmaire, 1887: 186
Type data. Holotype (Basel Museum)
= Vietomorpha arabica Schuster & Gebien, 1938: 59 [syn. by
Type data. Syntypes (Basel Museum)
tuberosa (Fairmaire, 1882a: 76) Vieta [Mal, 1986a: 21]
Type data. Holotype (Paris Museum)
= Vieta senegalensis somalica Gebien, 1937b: 41 [syn. by
Type data. Syntypes (Genoa Museum)
Type genus. Trachynotus Latreille, 1828
Taxonomic diversity. (10 gen., 218 spp.): Cyrtoderes (5 ssp.), Epairopsis (4), Ethmus (30), Histrionotus (2), Microphligra (1), Ossiporis (11), Oxycerus (1), Somaticus (148), Trachynotus (15), Trichethmus (1).
Distribution. Widely distributed in the southern part of the Afrotropical Realm (south from equator) (Fig.
Type species. Sepidium lacunosum Thunberg, 1784 (by subsequent designation by Bousquet & Bouchard (2013: 45); syn of Tenebrio cristatus DeGeer, 1778)
= Phligra Laporte, 1840: 197 [junior objective synonym; see Bousquet & Bouchard (2013: 45)]
Type species. Phligra degeeri Laporte, 1840 (by monotypy); syn of Tenebrio cristatus DeGeer, 1778.
cristatus cristatus (DeGeer, 1778: 653) Tenebrio [Bousquet & Bouchard, 2013: 45]
Type data. Syntypes (Naturhistoriska riksmuseet)
= Sepidium lacunosum Thunberg, 1787: 48 [syn. by
Type data. Syntypes (Uppsala University)
= Brachycerus cristatus Fabricius, 1798: 161 [syn. by Alonso-Zarazaga (2014: 73)]
Type data. Holotype (Lund University)
= Brachycerus areolatus Thunberg, 1799: 31 [syn. by
Type data. Syntypes (Uppsala University)
= Phligra degeeri Laporte, 1840: 197 [syn. by
Type data. Holotype (Paris Museum)
= Cyrtoderes curculioides Solier, 1843: 36 [syn. by
Type data. Holotype (Paris Museum)
cristatus nigritus Solier, 1843: 35
Type data. Holotype (Paris Museum)
cristatus sinuosus Solier, 1843: 34
Type data. Holotype (Paris Museum)
crucifera (Haag-Rutenberg, 1871a: 35) Phligra [Bousquet & Bouchard, 2013: 45]
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
hamaticollis (Péringuey, 1904: 235) Phligra [Bousquet & Bouchard, 2013: 45]
Type data. Syntypes (Cape Museum)
Type species. Trachynotus frontalis Haag-Rutenberg, 1873 (by original designation)
deckerti Ferrer, 2004: 193
Type data. Holotype (Berlin Museum)
frontalis frontalis (Haag-Rutenberg, 1873: 43) Trachynotus [Koch, 1955: 47]
Type data. Syntypes (Naturhistoriska riksmuseet, Warsaw Museum)
= Epairops laevigata Péringuey, 1892: 55 [syn. by
Type data. Syntypes (Cape Museum)
frontalis variegata (Péringuey, 1892: 55) Epairops [
Type data. Holotype (Cape Museum)
superbus
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
Type species. Ethmus maculatus Haag-Rutenberg, 1873 (by subsequent designation by
= Tynthlobia
Type species. Tynthlobia quadricostata Fairmaire, 1888 (by monotypy)
Type species. Ethmus (Ethmomerus) subcylindricus Koch, 1954 (by original designation)
brevis Koch, 1954b: 261
Type data. Holotype (Tervuren Museum) and paratypes (Budapest Museum, Ditsong Museum, Tervuren Museum)
subcylindricus Koch, 1954b: 259
Type data. Holotype (British Museum) and paratypes (British Museum, Ditsong Museum)
Type species. Ethmus latus Haag-Rutenberg, 1873 (by original designation)
herero herero Koch, 1958: 32
Type data. Holotype and paratypes (Ditsong Museum)
herero pronamibensis Koch, 1958: 32
Type data. Holotype and paratypes (Ditsong Museum)
latus bisbicostatus Koch, 1958: 30
Type data. Holotype (Ditsong Museum) and paratypes (Budapest Museum, Ditsong Museum)
latus kaokoanus Koch, 1958: 30
Type data. Holotype (Ditsong Museum) and paratypes (Budapest Museum, Ditsong Museum)
latus latus Haag-Rutenberg, 1873: 45 [
Type data. Holotype (Naturhistoriska riksmuseet)
paradisiacus angolanus Koch, 1958: 33
Type data. Holotype and paratypes (Ditsong Museum)
paradisiacus formosus Koch, 1958: 33
Type data. Holotype (Ditsong Museum)
paradisiacus paradisiacus Koch, 1958: 28
Type data. Holotype (Ditsong Museum) and paratypes (Budapest Museum, Ditsong Museum)
vernayi vernayi Koch, 1958: 26
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Budapest Museum, Ditsong Museum)
vernayi reductus Koch, 1958: 28
Type data. Holotypes and paratypes (Ditsong Museum)
vernayi marginatus Koch, 1958: 28
Type data. Holotype (Ditsong Museum)
Type species. Ethmus maculatus Haag-Rutenberg, 1873 (by subsequent designation by
acinopoides Koch, 1954b: 247
Type data. Holotype (Tervuren Museum) paratypes (Ditsong Museum, Tervuren Museum)
analis Gebien, 1910a: 150
Type data. Holotype (Basel Museum)
barbosai Koch, 1954b: 241
Type data. Holotype (Maputo Museum)
borgesi Koch, 1958: 21
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
cinereosparsus Gebien, 1910a: 151
Type data. Syntypes (Basel Museum)
dollmani Koch, 1954b: 248
Type data. Holotype (British Museum) and paratypes (British Museum, Ditsong Museum)
gedyei Koch, 1954b: 252
Type data. Holotype (British Museum) and paratypes (British Museum, Ditsong Museum)
maculatus Haag-Rutenberg, 1873: 45
Type data. Syntypes (Paris Museum, Munich Museum – Haag-Rutenberg coll.)
nyassicus Koch, 1954b: 249
Type data. Holotype and paratypes (Munich Museum)
plicatus Müller, 1887: 300
Type data. Holotype (Leiden Museum)
pygidialis Koch, 1954b: 253
Type data. Holotype (Ditsong Museum)
pustulatus Koch, 1954b: 257
Type data. Holotype (National Congo) and paratypes (Budapest Museum, Ditsong Museum, National Congo)
quadricostatus (Fairmaire, 1888b: 261) Tynthlobia [
Type data. Holotype (Leyden Museum)
sculptus Koch, 1954b: 255
Type data. Holotype (Munich Museum)
silvanus Koch, 1958: 22
Type data. Holotype (Ditsong Museum) and paratypes (Basel Museum, Budapest Museum, Ditsong Museum)
tessellatus Koch, 1958: 24
Type data. Holotype (Ditsong Museum)
incostatus Gebien, 1910a: 150
Type data. Syntypes (Basel Museum, Tervuren Museum)
Notes. Species unknown to Koch (1954,
Type species. Trachynotus lightfooti Péringuey, 1892 (by original designation)
lightfooti (Péringuey, 1892: 122) Trachynotus [Koch, 1955: 44]
Type data. Syntypes (Cape Museum)
omercooperi Koch, 1955: 43
Type data.
Notes. Although
Type species. Phligra minuta Péringuey, 1904: (by original designation)
Notes. In caption of plate 9 in his paper from 1955 Koch introduced the second representative of Microphligra: “Microphligra (Paraphligra) succulentium (subg. nov. in litt. sp. nov. in litt.)”. A habitus of this beetle is presented in the preceding page. However, Koch did not specify any characters separating this new entity from the Microphligra minuta, therefore this cannot be treated as a valid description according to the regulations of the
minuta (
Type data. Holotype (Cape Museum)
Type species. Ossiporis terrena Pascoe, 1866 (by monotypy)
= Epairops Fåhraeus, 1870: 282 [syn. by
Type species. Epairops fragilis Fåhraeus, 1870 (by monotypy)
angolensis Koch, 1953b: 86
Type data. Holotype (British Museum) and paratype (Ditsong Museum)
capeneri Koch, 1953d: 7
Type data. Holotype (Ditsong Museum) and paratypes (British Museum)
crenulimargo Koch, 1953d: 26
Type data. Holotype (British Museum) and paratypes (British Museum, Brussels Museum, Ditsong Museum)
marshalli Koch, 1953d: 24
Type data. Holotype (British Museum)
taterae Koch, 1953d: 2
Type data. Holotype (Tervuren Museum)
terrena chubbi Koch, 1953d: 25
Type data. Holotype (Ditsong Museum) and paratypes (Durban Museum)
terrena fragilis (Fåhraeus, 1870: 282) Epairops [Gebien, 1937b: 37]
Type data. Lectotype, designated by
terrena terrena Pascoe, 1866: 452
Type data. Holotype (Basel Museum, British Museum)
terrena rhodesiana Koch, 1953d: 8
Type data. Holotype (Ditsong Museum) and paratypes (British Museum, Ditsong Museum, Rhodesia Museum)
terrena zoutpansbergensis Koch, 1953d: 28
Type data. Holotype and paratypes (Ditsong Museum)
undulicostis Koch, 1953d: 6
Type data. Holotype (Rhodesia Museum)
Type species. Trachynotus resolutus Péringuey, 1904 (by original designation)
resolutus (Péringuey, 1904: 234) Trachynotus [Koch, 1955: 46]
Type data. Holotype (Ditsong Museum)
Type species. Sepidium rugosum Fabricius, 1781 (by original designation)
Type species. Sepidium acuminatum Quensel, 1806 (by original designation); syn. of Sepidium striatum Thunberg, 1787
adventitus (Péringuey, 1899: 229) Trachynotus [Koch, 1955: 169]
Type data. Syntypes (Cape Museum)
albanyensis Koch, 1955: 164
Type data. Holotype (British Museum)
algoensis Koch, 1955: 153
Type data. Syntypes (Budapest Museum, Ditsong Museum)
bisinuatus Koch, 1955: 155
Type data. Syntypes (Ditsong Museum)
caviventris Koch, 1955: 158
Type data. Holotype (Ditsong Museum)
cohaerens Koch, 1955: 166
Type data. Syntypes (Budapest Museum, Ditsong Museum)
corallipes Koch, 1955: 157
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
dimorphus Koch, 1955: 172
Type data. Syntypes (Basel Museum, Budapest Museum, Cape Museum)
georgensis Koch, 1955: 166
Type data. Holotype (Cape Museum)
hirundo Koch, 1955: 159
Type data. Holotype (Cape Museum) and paratypes (Cape Museum, Ditsong Museum)
Notes.
karrooensis Koch, 1955: 164
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
licinoides (Haag-Rutenberg, 1873: 7) Trachynotus [Koch, 1955: 176]
Type data. Syntypes (Munich Museum)
= Trachynotus rusticus Péringuey, 1899: 300 [syn. by
Type data. Syntypes (Cape Museum)
malaisei Koch, 1955: 172
Type data. Syntypes (Basel Museum, British Museum, Budapest Museum, Ditsong Museum)
marlothi Koch, 1955: 17
Type data. Holotype (Ditsong Museum)
moerens (Haag-Rutenberg, 1879: 294) Trachynotus [Koch, 1955: 162]
Type data. Holotype (British Museum)
Notes. Type deposition information after
namaquensis Koch, 1955: 170
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
nitens (Péringuey, 1899: 298) Trachynotus [Koch, 1955: 171]
Type data. Syntypes (Cape Museum)
nollothensis Koch, 1955: 158
Type data. Holotype (Cape Museum)
praephallatus frigidorae Koch, 1955: 162
Type data. Syntypes (Ditsong Museum)
praephallatus praephallatus Koch, 1955: 161
Type data. Holotype and paratype (Ditsong Museum)
punctiger (Haag-Rutenberg, 1873: 15) Trachynotus [Koch, 1955: 156]
Type data. Holotype (Museum Berlin)
purcelli Koch, 1955: 151
Type data. Syntypes (Cape Museum)
saxicola Koch, 1955: 177
Type data. Holotype (Ditsong Museum)
scaber (Haag-Rutenberg, 1873: 15) Trachynotus [Koch, 1955: 175]
Type data. Holotype (Munich Museum)
striatus (Thunberg, 1787: 48) SepidiumSepidium [Ferrer, 2009: 114]
Type data. Holotype (Uppsala University)
= Sepidium acuminatus Quensel, 1806: 130 [syn. by
Type data. Syntypes (Naturhistoriska riksmuseet)
suturalifer Koch, 1955: 151
Type data. Holotype (Cape Museum)
transmontanus Koch, 1955: 167
Type data. Holotype (Cape Museum)
vansonianus Koch, 1955: 154
Type data. Syntypes (Ditsong Museum)
Type species. Trachynotus brucki Haag-Rutenberg, 1873 (by original designation)
Trachynotus bohemani bohemani (Haag-Rutenberg, 1873: 11) Trachynotus [Koch, 1955: 94]
Type data. Holotype (Munich Museum) and paratype (Naturhistoriska riksmuseet)
Notes. Type deposition information after
bohemani gaerdesi Koch, 1955: 95
Type data. Holotype (Ditsong Museum)
bohemani scherzi Koch, 1955: 96
Type data. Syntypes (Ditsong Museum)
brucki brucki (Haag-Rutenberg, 1873: 13) Trachynotus [Koch, 1955: 93]
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. Type deposition information after
brucki ovamboanus Koch, 1955: 104
Type data. Holotype (Ditsong Museum)
brucki poweri (Hesse, 1935: 554) Trachynotus [Koch, 1955: 104]
Type data. Holotype (Cape Museum)
Notes. Type deposition information after
cinctus (Haag-Rutenberg, 1873: 12) Trachynotus [Koch, 1955: 97]
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. Type deposition information after
geniculatus geniculatus (Haag-Rutenberg, 1873: 21) Trachynotus [Koch, 1955: 99]
Type data. Syntypes (British Museum, Munich Museum)
geniculatus hessei Koch, 1955: 101
Type data. Syntypes (Budapest Museum, Ditsong Museum)
geniculatus pluricostatus Koch, 1955: 100
Type data. Syntypes (Ditsong Museum, Munich Museum)
hereroensis Koch, 1955: 99
Type data. Holotype (University Lund)
rugulosicollis (Hesse, 1935: 554) Trachynotus [Koch, 1955: 102]
Type data. Syntypes (Cape Museum, Ditsong Museum)
Type species. Trachynotus badeni Haag-Rutenberg, 1873 (by original designation)
badeni (Haag-Rutenberg, 1873: 10) Trachynotus [Koch, 1955: 87]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
= Trachynotus scrobiculatus Péringuey, 1885: 110 [syn. by
Type data. Holotype (Cape Museum)
seminitidus Koch, 1955: 89
Type data. Holotype (Ditsong Museum) and paratypes (Budapest Museum, Cape Museum, Naturhistoriska riksmuseum)
Trachynotus strangulatus arborarius Koch, 1955: 92
Type data. Syntypes (Budapest Museum, Ditsong Museum)
Trachynotus strangulatus auobensis Koch, 1955: 93
Type data. Holotype and paratypes (Ditsong Museum)
Trachynotus strangulatus patzelti Koch, 1955: 92
Type data. Holotype (Ditsong Museum) and paratype (California Academy of Science)
Trachynotus strangulatus rehobothensis Koch, 1955: 92
Type data. Holotype (Ditsong Museum) and paratypes (Cape Museum)
Trachynotus strangulatus strangulatus (Gebien, 1920: 92) Trachynotus [Koch, 1955: 90]
Type data. Holotype (Basel Museum)
Trachynotus wahlbergi ausensis Koch, 1955: 90
Type data. Holotype (Cape Museum) and paratype (Ditsong Museum)
Trachynotus wahlbergi wahlbergi (Haag-Rutenberg, 1873: 10) Trachynotus [Koch, 1955: 89]
Type data. Holotype (Naturhistoriska riksmuseum)
Notes. Type deposition information after
Notes. While describing this genus
Treated as a subgenus of Trachynotus (e.g.,
Type species. Clinocranion spinosum Solier, 1843 (by subsequent designation by
planatus drukeri Koch, 1955: 73
Type data. Syntypes (Budapest Museum, Ditsong Museum, Tervuren Museum)
planatus planatus (Solier, 1843: 116) Clinocranion [Koch, 1955: 72]
Type data. Holotype (Geneva Museum – Gory collection)
Notes. Type deposition information after
planatus subdamarensis Koch, 1955: 73
Type data. Holotypes and paratypes (Ditsong Museum)
spinosus (Solier, 1843: 115) Clinocranion [Koch, 1955: 71]
Type data. Holotype (Geneva Museum)
Type species. Trachynotus regalis Haag-Rutenberg, 1875 (original designation)
angustus (Péringuey, 1886: 125) Trachynotus [Koch, 1955: 106]
Type data. Syntypes (Cape Museum)
distinctus (Péringuey, 1892: 54) Trachynotus [Koch, 1955: 105]
Type data. Syntypes (Cape Museum)
regalis (Haag-Rutenberg, 1875: 82) Trachynotus [Koch, 1955: 106]
Type data. Syntypes (British Museum, Munich Museum – Haag-Rutenberg coll.)
Notes. Although the correct original spelling of this genus group name is Somaticum (Hope, 1840: 117 and errata), to our knowledge all subsequent authors have used the incorrect subsequent spelling Somaticus and that this incorrect subsequent spelling is in prevailing usage and attributed to the publication of the original spelling; therefore this genus-group name is to be preserved and deemed to be the correct original spelling (
Type species. Sepidium rugosum Fabricius, 1781 (by original designation)
= Gonopterus Solier, 1843: 101 [junior subjective synonym proposed by Gebien, 1910b: 163]
Type species. Sepidium rugosum Fabricius, 1781 (by monotypy)
aeneus (Solier, 1843: 111) Trachynotus [Koch, 1955: 82]
Type data. Syntypes (Geneva Museum)
Notes. Type deposition information after
bisbicostatus (Gebien, 1920: 93) Trachynotus [Koch, 1955: 86]
Type data. Neotype (Basel Museum), designated by
Notes. According to
decoratipes cisfluminis Koch, 1955: 81
Type data. Syntypes (Budapest Museum, Ditsong Museum)
decoratipes decoratipes Koch, 1955: 79
Type data. Syntypes (Basel Museum, Budapest Museum, Cape Museum, Ditsong Museum)
glabriventris Koch, 1955: 78
Type data. Syntypes (Basel Museum, Budapest Museum, Paris Museum)
rugosus rugosissimus Koch, 1955: 78
Type data. Syntypes (Cape Museum)
rugosus rugosus (Fabricius, 1781: 315) Sepidium [Koch, 1955: 76]
Type data. Holotype Copenhagen Museum) and paratypes (British Museum)
Notes. Type deposition information after
= Pimelia leucophrys Herbst, 1799: 115 [syn. by Haag-Rutenberg, 1873: 8]
Type data. Holotype (Berlin Museum)
rugosus testaceipes Koch, 1955: 77
Type data. Syntypes (Ditsong Museum)
stali (Haag-Rutenberg, 1873: 18) Trachynotus [Koch, 1955: 85]
Type data. Holotype (Naturhistoriska riksmuseet)
= Clinocranion latemarginatum Péringuey, 1885: 115 [syn. by
Type data. Holotype (Cape Museum)
straminicornis Koch, 1955: 81
Type data. Holotype (Ditsong Museum)
stridulatus Koch, 1955: 78
Type data. Syntypes (Budapest Museum, California Academy, Ditsong Museum, Stellenbosch University)
welwitschi Koch, 1955: 84
Type data. Syntypes (British Museum)
Type species. Tracheloeum laticolle Hope, 1840 (by original designation)
carinatus cancellatus Koch, 1955: 200
Type data. Holotype (Ditsong Museum)
carinatus carinatus (Solier, 1843: 109) Trachynotus [Koch, 1955: 199]
Type data. Holotype (Geneva Museum)
Notes. Type deposition information after Haag-Rutenberg (1871).
carinatus chevrolati (Haag-Rutenberg, 1873: 35) Trachynotus [Koch, 1955: 54]
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
contractus (Haag-Rutenberg, 1873: 27) Trachynotus [Koch, 1955: 189]
Type data. Holotype (Museum Berlin)
Notes. Type deposition information after
dilatatus (Haag-Rutenberg, 1873: 25) Trachynotus [Koch, 1955: 182]
Type data. Holotype (Munich Museum)
fahraeusi Koch, 1955: 185
Type data. Syntypes (Cape Museum)
Notes.
giganteus Koch, 1955: 194
Type data. Syntypes (Budapest Museum, Ditsong Museum)
hoffmanni (Haag-Rutenberg, 1878: 93) Trachynotus [Koch, 1955: 201]
Type data. Holotype (Stuttgart Museum)
impressicollis (Péringuey, 1885: 111) Trachynotus [Koch, 1955: 200]
Type data. Syntypes (Cape Museum)
intermedius (Haag-Rutenberg, 1878: 92) Trachynotus [Koch, 1955: 195]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
laticollis (Hope, 1840: 116) Tracheloeum [Koch, 1955: 179]
Type data. Holotype (Oxford University – Hope coll.)
Notes. Type deposition information after
maculosus (Fåhraeus, 1870: 276) Trachynotus [Koch, 1955: 184]
Type data. Syntypes (Naturhistoriska riksmuseet)
Notes. Type deposition information after
marginatus (Thunberg, 1787) Sepidium [
Type data. Holotype (Uppsala University)
= Trachynotus laevis Fåhraeus, 1870: 275 [syn. by Ferrer, 2009: 114]
Type data. Syntypes (Naturhistoriska riksmuseet, Warsaw Museum)
Notes. Type deposition information after
= Trachnotus glaber Fåhraeus, 1870: 275 [syn. by
Type data. Syntypes (Naturhistoriska riksmuseet)
pretorianus bushveldeus Koch, 1955: 183
Type data. Holotype (Ditsong Museum) and paratype (Cape Museum)
pretorianus pretorianus Koch, 1955: 182
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
silphoides metallescens Koch, 1955: 194
Type data. Holotype (South African National Collection)
silphoides nigronitens Koch, 1955: 193
Type data. Syntypes (Cape Museum, Ditsong Museum)
silphoides peringueyi Koch, 1955: 191, replacement name
= Trachynotus plicipennis Péringuey, 1899: 300 [junior primary homonym of Trachynotus plicipennis Haag-Rutenberg, 1873: 38]
Type data. Syntypes (Cape Museum)
silphoides silphoides (Fåhraeus, 1870: 274) Trachynotus [Koch, 1955: 191]
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. Type deposition information after
silphoides swazicola Koch, 1955: 193
Type data. Syntypes (Munich Museum)
similis (Haag-Rutenberg, 1873: 35) Trachynotus [Koch, 1955: 196]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
transvaalensis Koch, 1955: 197
Type data. Syntypes (Cape Museum, Ditsong Museum)
vittiger (Haag-Rutenberg, 1873: 22) Trachynotus [Koch, 1955: 187]
Type data. Holotype (British Museum)
Notes. Type deposition information after
Type species. Trachynotus bipunctatus Haag-Rutenberg, 1873 (by original designation)
albomaculatus (Haag-Rutenberg, 1873: 41) Trachynotus [Koch, 1955: 142]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
= Trachynotus terrenus Péringuey, 1885: 114 [syn. by Gebien, 1937a: 773]
Type data. Syntypes (Cape Museum)
barnardi Koch, 1955: 137
Type data. Holotype (Cape Museum)
bipunctatus bipunctatus (Haag-Rutenberg, 1873: 20) Trachynotus [Koch, 1955: 113]
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
bipunctatus pilosus (Péringuey, 1885: 112) Trachynotus [Koch, 1955: 121]
Type data. Syntypes (Cape Museum)
braunsi Koch, 1955: 135
Type data. Syntypes (Basel Museum, British Museum, Budapest Museum, Ditsong Museum, Tervuren Museum)
cordipennis Koch, 1955: 136
Type data. Holotype (Ditsong Museum)
dubius (Péringuey, 1885: 114) Trachynotus [Koch, 1955: 130]
Type data. Holotype (Cape Museum)
= Trachynotus sericeus Péringuey, 1886: 124 [syn. by Gebien, 1937a: 773]
Type data. Syntypes (Cape Museum)
= Trachynotus dubius maculipennis Gebien, 1920: 99 [syn. by
Type data. Syntypes (Hamburg University)
dutoiti Koch, 1955: 118
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
eremicola Koch, 1955: 129
Type data. Syntypes (Cape Museum, Ditsong Museum)
fitzsimonsi Koch, 1955: 126
Type data. Holotype (Ditsong Museum) and paratypes (Budapest Museum, Ditsong Museum)
goryi (Solier, 1843: 112) Trachynotus [Koch, 1955: 117]
Type data. Holotype (Geneva Museum)
Notes. Type deposition information after
gracilipes (Haag-Rutenberg, 1873: 19) Trachynotus [Koch, 1955: 128]
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
= Trachynotus attenuatus
Type data. Syntypes (Cape Museum)
gunvorae amnigenus Koch, 1955: 123
Type data. Holotype (Cape Museum)
gunvorae cylindricollis Koch, 1955: 122
Type data. Holotype (Cape Museum)
gunvorae gunvorae (Koch, 1953c: 12) Trachynotus [Koch, 1955: 121]
Type data. Holotype (Lund University) and paratypes (Ditsong Museum, Lund University)
haagi haagi (Péringuey, 1899: 299) Trachynotus [Koch, 1955: 114]
Type data. Holotype (Cape Museum)
haagi pilipeplus Koch, 1955: 116
Type data. Holotype (Ditsong Museum) and paratype (Naturhistoriska riksmuseet)
incostatus (Gebien, 1920: 99) Trachynotus [Koch, 1955: 138]
Type data. Syntypes (Basel Museum)
kungorum Koch, 1955: 140
Type data. Syntypes (Ditsong Museum)
plutus Koch, 1955: 134
Type data. Holotype (Ditsong Museum) and paratypes (California Academy, Ditsong Museum, Munich Museum)
pygmaeus (Fåhraeus, 1870: 279) Trachynotus [Koch, 1955: 141]
Type data. Holotype (Naturhistoriska riksmuseet)
= Trachynotus tantillus Péringuey, 1899: 301 [syn. by
Type data. Holotype (Cape Museum)
ratus Koch, 1955: 125
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
tentyrioides (Haag-Rutenberg, 1873: 24) Trachynotus [Koch, 1955: 124]
Type data. Holotype (Munich Museum)
= Trachynotus acuticostis
Type data. Syntypes (Basel Museum)
tibialis (Haag-Rutenberg, 1873: 20) Trachynotus [Koch, 1955: 132]
Type data. Holotype (Munich Museum)
Notes.
zinni Koch, 1955: 133
Type data. Syntypes (Cape Museum)
Type species. Trachynotus sordidus Gerstaecker, 1854 (by original designation)
angulatus (Fåhraeus, 1870: 277) Trachynotus [Koch, 1955: 206]
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. Type deposition information after
darlingtoni Koch, 1955: 212
Type data. Syntypes (Budapest Museum, Ditsong Museum)
funestus (Fåhraeus, 1870: 278) Trachynotus [Koch, 1955: 227]
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. Type deposition information after
griseus (Fåhraeus, 1870: 277) Trachynotus [Koch, 1955: 214]
Type data. Holotype (Naturhistoriska riksmuseet)
Notes. Type deposition information after
hispidus (Hesse, 1935: 555) Trachynotus [Koch, 1955: 225]
Type data. Syntype (Cape Museum, Ditsong Museum)
histrio Koch, 1955: 229
Type data. Holotype (Cape Museum)
lutulentus lutulentus (Péringuey, 1899: 301) Trachynotus [Koch, 1955: 222]
Type data. Holotype (Cape Museum)
lutulentus montisdraconis Koch, 1955: 223
Type data. Syntypes (Durban Museum)
metropolis Koch, 1955: 227
Type data. Syntypes (Basel Museum, British Museum, Budapest Museum, Ditsong Museum)
newtoni Koch, 1955: 223
Type data. Holotype (Ditsong Museum)
obscurus Koch, 1955: 208
Type data. Syntypes (Cape Museum)
schalkwykae Koch, 1955: 213
Type data. Syntypes (Pretoria University)
sinuatus Koch, 1955: 208
Type data. Holotype (Ditsong Museum)
sordidus (Gerstaecker, 1854: 532) Trachynotus [Koch, 1955: 205]
Type data. Syntypes (Berlin Museum)
terricola setulosus (Haag-Rutenberg, 1873: 31) Trachynotus [Koch, 1955: 216]
Type data. Syntypes (Munich Museum, Warsaw Museum)
Notes. Type deposition information after
terricola terricola (Fåhraeus, 1870: 278) Trachynotus [Koch, 1955: 215]
Type data. Syntypes (Naturhistoriska riksmuseet)
Notes. Type deposition information after
testudo Koch, 1955: 220
Type data. Holotype (Cape Museum)
varicollis brachythorax Koch, 1955: 211
Type data. Syntypes (Budapest Museum, Ditsong Museum)
varicollis disconnectus Koch, 1955: 212
Type data. Holotype (Cape Museum) and paratypes (Cape Museum, Ditsong Museum)
varicollis varicollis Koch, 1955: 209
Type data. Syntypes (Budapest Museum, Ditsong Museum)
vestitus (Haag-Rutenberg, 1873: 30) Trachynotus [Koch, 1955: 218]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
zumpti rhodesianus Koch, 1955: 220
Type data. Holotype (Ditsong Museum) and paratype (Cape Museum)
zumpti zumpti Koch, 1955: 219
Type data. Syntypes (Budapest Museum, Ditsong Museum)
Type species. Trachynotus crinitus Haag-Rutenberg, 1873 (by original designation)
crinitus (Haag-Rutenberg, 1873: 32) Trachynotus [Koch, 1955: 109]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
kraatzi kraatzi (Haag-Rutenberg, 1873: 33) Trachynotus [Koch, 1955: 110]
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
kraatzi fulvohirtus Koch, 1955: 111
Type data. Syntypes (Basel Museum, Budapest Museum, Ditsong Museum)
kraatzi orientalis Koch, 1955: 112
Type data. Holotype (Ditsong Museum) and paratypes (Stellenbosch University)
serratus (Péringuey, 1885: 112) Trachynotus [Koch, 1955: 112]
Type data. Holotype (Cape Museum)
Type species. Trachynotus peregrinator Koch, 1953a (by original designation)
peregrinator (Koch, 1953a: 179) Trachynotus [Koch, 1955: 230]
Type data. Holotype (British Museum) and paratypes (British Museum, Budapest Museum, Ditsong Museum)
tropicalis Koch, 1955: 231
Type data. Holotype (Frankfurt Museum)
incertae sedis
damarinus Péringuey, 1904: 233 [
Type data. Holotype (Naturhistoriska riksmuseet)
Notes.
scutelliformis (Laporte, 1840: 197) Sepidium
Type data. Holotype (Munich Museum)
Notes. See
Type species. Sepidium vittatus Fabricius, 1781 (here designated)
Notes. Trachynotus used to be attributed to Latreille (1829: 14). In that publication, Sepidium reticulatum was one of the originally included species and this species was later selected as the type species (see
= Hipomelus Dejean, 1834: 181 [junior objective synonym; see Bousquet & Bouchard (2013: 47)]
Type species. Sepidium vittatum Fabricius, 1781 (by subsequent designation by
albulus Péringuey, 1886: 127
Type data. Syntypes (Cape Museum)
elongatus (Olivier, 1795: 8) Sepidium [Haag-Rutenberg, 1873: 36]
Type data. Syntypes (Paris Museum)
leucographus Solier, 1843: 107
Type data. Syntypes (Paris Museum)
lutosus Péringuey, 1885: 113
Type data. Syntypes (Cape Museum)
meracus Péringuey, 1899: 302
Type data. Holotype (Cape Museum)
ornatus Haag-Rutenberg, 1873: 40
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
plicipennis Haag-Rutenberg, 1873: 38
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
proximus Laporte, 1840: 197
Type data. Holotype (Paris Museum)
recurvus Haag-Rutenberg, 1873: 38
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
reticulatus (De Geer, 1778: 651) Tenebrio [Haag-Rutenberg, 1873: 2]
Type data. Holotype (Naturhistoriska riksmuseet)
= Sepidium reticulatum Thunberg, 1791: 23 [syn. by
Type data. Syntypes (Uppsala University)
sctulosus Haag-Rutenberg, 1873: 31
Type data. Syntypes (Munich Museum – Haag-Rutenberg coll.)
tricostatus Haag-Rutenberg, 1873: 23
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
variegatus Haag-Rutenberg, 1878: 94
Type data. Holotype (Munich Museum – Haag-Rutenberg coll.)
albulusvicinus (Haag-Rutenberg, 1871b: 51) Psammodes [Koch, 1955: 46]
Type data. Syntypes (Geneva Museum, Warsaw Museum)
vittatus (Fabricius, 1781: 315) Sepidium [Haag-Rutenberg, 1873: 5]
Type data. Syntypes (British Museum, Copenhagen Museum)
= Sepidium vittatum Thunberg, 1791: 24 [syn. by
Type data. Syntypes (Uppsala University)
= Sepidium plicatus Wiedemann, 1823: 39 [syn. by
Type data. Holotype (Humboldt University)
= Trachynotus lacunosus Solier, 1843: 110 [syn. by
Type data. Syntypes (Paris Museum)
Type species. Trichethmus lobicolis Gebien, 1937b: 45 (by monotypy)
lobicolis Gebien, 1937b: 46
Type data. Holotype (Basel Museum)
Funding for this study was provided by the NSF ARTS Program (DEB #1754630). Publication costs were covered by the funding from the Species File Group of the Illinois Natural History Survey. The authors would like to thank Yves Bousquet (Gatineau, Quebec, Canada) and Ottó Merkl (Budapest, Hungary) for providing useful nomenclatural information.