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The work treats 52 species–group names related to genus Agrilus incorrectly cited in the world catalogue of Bellamy (2008). The name dimorphus Théry, 1941 from the genus Aphanisticus and mulleri Théry, 1925 from the genus Australodraco are also treated. Four primary or secondary homonyms are replaced by substitute names. Most of the proposed changes refer to the availability, validity, spelling and authorship of the names. The following new nomenclatural acts are proposed: Four new substitute names for homonyms: gola Jendek for filiformis Gory & Laporte, 1839 not Herbst, 1801; lukesi Obenberger, 1936 for modicus Kerremans, 1892 not Solier, 1833; thomsoni Jendek for impressipennis Thomson not Uhler, 1855; walkerianus Jendek (Aphanisticus) for sulcicollis Walker not Lacordaire, 1835. New synonyms: turei Curletti, 2002 is an objective synonym of thurei Curletti, 1996. Lectotype designations: A lectotype is designated for Agrilus dualaecola Obenberger, 1923.
Coleoptera, Buprestidae, Agrilus, Aphanisticus, Australodraco, new synonyms, new substitute names, lectotype designation
In 2008–2009, a monumental catalogue on world buprestid beetles was published by Chuck L. Bellamy. The major part of the Volume 4 (
This work is aimed to clarify and correct some taxonomic and nomenclatural problems published in the catalogue. The term Agrilus related species–group names means names originally or subsequently assigned to the genus Agrilus.
All data were checked from original publications rather than to rely on data published in the Zoological Records. The vast majority of treated taxa belongs to Agrilus, but some taxa of other genera are also influenced (see Abstract).
In the course of work, several Agrilus species–group names were found primary or se-condary homonyms. These names are replaced by substitute names in the separate section.
Treated names are cited alphabetically. Each name has the corrected status in parenthesis with its generic assignment if other than Agrilus. Pages where and how a taxon is listed in
Agrilus acutipennis Horn (unavailable name)
Pages 2216, 2258 – cited as an available synonym of olivaceoniger Fisher, 1928 on page 2216 and quadriimpressus on page 2258.
Remark.
Agrilus aenescens Kerremans, 1903 (valid name)
Page 1953 – cited as a synonym of aenescentellus Obenberger, 1936c.
Remark. The name aenescentellus Obenberger, 1936c was proposed as a replacement name for aenescens Kerremans, 1903 not Shilsky, 1888. Because the name aenescens Shilsky is unavailable (see
Agrilus agadiensis Théry, 1910 (valid name)
Page 2050 – cited as a synonym of cupriventris Gory & Laporte, 1839
Remark. The name agadiensis Théry, 1910 was cited by
Agrilus atriplicis Escalera (unavailable synonym of rabaticus Théry, 1930)
Page 2261 – cited as available synonym of rabaticus
Remark.
Agrilus bandanus Obenberger, 1933 (unavailable synonym of sulphurifer Burmeister, 1872)
Page 2318 – cited as available synonym of sulphurifer
Remark. The name bandanus Obenberger, 1933 was proposed for an aberration, which is an unavailable name (ICZN, Article 45.6.2).
Agrilus bayeri Obenberger, 1935d (synonym of sinensis splendidicollis Fairmaire, 1889)
Page 2294 – cited as beyeri; synonym of sinensis splendidicollis
Remark. Original spelling of this patronymic name is bayeri (dedicated to Prof. E. Bayer) not beyeri which is incorrect subsequent spelling.
Agrilus bolamanus Kerremans (unavailable synonym of buraicus Obenberger, 1928)
Page 2010 – cited as available synonym of buraicus
Remark.
Agrilus borrei Kerremans (unavailable synonym of deborrei Dugès, 1891)
Page 2059 – cited as available synonym of deborrei
Remark.
Agrilus caliginulus Obenberger, 1935e (valid name)
Page 2012 – cited as calliginulus
Remark. Original spelling of the name is caliginulus not calliginulus which is incorrect subsequent spelling.
Agrilus cameroni Bellamy, 1999 (unavailable name)
Page 2012 – cited as valid name for species
Remark. The name cameroni Bellamy, 1999 was proposed as a new replacement name for dimorphus Théry, 1941, which was supposed to be preoccupied by dimorphus Obenberger, 1923.
Agrilus camerounensis Obenberger (unavailable name)
Page 2012 – cited as cameroonensis and valid name for species
Remark. The name camerounensis Obenberger, 1931 is listed as a valid name of species but I failed to find original publication neither by Obenberger or anybody else. The name camerounensis Obenberger is an unavailable name introduced by
Agrilus capicola Kerremans, 1898 (valid name)
Page 2013 – cited as capicolus
Remark. Original spelling of the name is capicola not capicolus. The name is to be treated as a noun in apposition to the name of its genus as ruled be ICZN (Article 31.2.2). The name capicolus is an incorrect subsequent spelling.
coeruleoniger Fisher, 1929 (valid name) Page 2032 – cited as coeruleonigra
Remark. Original spelling of the name is coeruleonigra. The ending has to be changed to coeruleoniger to agree with the genus gender. This is a mandatory change ruled by ICZN (Article 34.2).
Agrilus confusulan Obenberger, 1935a (unavailable synonym of grandis Gory & Laporte, 1839)
Page 2115 – cited as confusulus and unavailable synonym of grandis
Remark. Original spelling of this unavailable name proposed for an aberration is confusulan.
Agrilus coronifrons Obenberger, 1931a (valid name)
Page 2227 – cited as a synonym of patricius
Remark. I failed to trace any author who proposed the synonymisation of the names patricius Obenberger, 1931a and coronifrons Obenberger, 1931a. The last use of both names was that of
Agrilus cyaneus Olivier (unavailable synonym of viridis Linné, 1758) Page 2354 – cited as available synonym of viridis
Remark.
Agrilus dicalis Kerremans (unavailable synonym of dualis Kerremans, 1903)
Page 2069 – cited as valid name for species.
Remark. Kerremans never proposed the name dicalis. This name was introduced by Obenberger (1936) who misspelled the name dualis Kerremans, 1903. This lapse was repeated by
Agrilus dimorphus Théry, 1941 (Aphanisticus; valid name for species)
Page 2012, 2395 – cited as a synonym of cameroni Bellamy, 1999 in Agrilus on page 2012 and as a valid name for species in Aphanisticus on page 2395.
Remark. The name dimorphus Théry, 1941 is a valid name for species in Aphanisticus. See also remarks at the name cameroni Bellamy, 1989.
Agrilus dualaecola Obenberger, 1923 (synonym of roscidinus Obenberger, 1923)
Page 2268, 2310 – cited as a synonym of roscidinus on page 2268 and synonym of subcurtulus on page 2310.
Remark.
Agrilus cyanophilus Schaefer, 1946 (unavailable synonym of suvorovi Obenberger, 1935)
Page 2320–2321, 2358 – cited as unavailable synonym of suvorovi on Page 2320 and as cyanophila an unavailable synonym of viridis on Page 2358
Remark. The name cyanophila Schaefer, 1946 was proposed for an aberration. The name with the correct ending cyanophilus is an unavailable synonym of suvorovi Obenberger, 1935 (see also
Agrilus enriquei Murria Beltrán & Murria Beltrán, 2007 (valid name) Page 2083 – cited as enriguei
Remark. Original spelling of this patronymic name is enriquei (dedicated to Enrique Murria Beltrán) not enriguei which is incorrect subsequent spelling.
Agrilus escalerai Obenberger, 1921 (valid name) Page 2084 – cited as escaleri
Remark. Original spelling of this patronymic name is escalerai (dedicated to Escalera) not escaleri which is incorrect subsequent spelling.
Agrilus ferrugineoguttatus Gory & Laporte (unavailable synonym of discolor Fåhraeus, 1851) Page 2072 – cited as available synonym of discolor
Remark.
Agrilus fulgidicollellus Obenberger, 1935 (unavailable name)
Page 2328 – cited as available synonym of thoracicus
Remark.
Agrilus goryi Saunders, 1871 (valid name)
Page 2118 – cited as valid name proposed as a replacement name
Remark. The name goryi Saunders, 1871 is a name proposed by indication to the bibliographic reference of Laporte and Gory (1839) (see ICZN Article 12.2) and not a proposal of a new replacement name.
guerryi Obenberger, 1933 (valid name)
Page 2118 – cited as guercyi
Remark. Original spelling of this patronymic name is guerryi (dedicated to Guerry) not guerrcyi which is an incorrect subsequent spelling.
Agrilus ingnoratus Obenberger, 1924 (unavailable synonym of ribbei Kiesenwetter, 1879) Page 2266 – cited as available synonym of ribbei
Remark.
Agrilus insulicola Kerremans, 1912 (valid name)
Page 2141 – cited as insulicolus
Remark. Original spelling of this name is insulicola not insulicolus. The name is to be treated as a noun in apposition to the name of its genus as ruled be ICZN (Article 31.2.2). The name insulicolus is an incorrect subsequent spelling.
Agrilus javicola Fisher, 1935 (valid name)
Page 2146 – cited as javicolus
Remark. Original spelling of this name is javicola not javicolus. The name is to be treated as a noun in apposition to the name of its genus as ruled be ICZN (Article 31.2.2). The name javicolus is an incorrect subsequent spelling.
Agrilus kachovskii Obenberger, 1935e (valid name)
Page 2148 – cited as kackovskii
Remark. Original spelling of this patronymic name is kachovskii (dedicated to Kachovski) not kackovskii which is incorrect subsequent spelling.
Agrilus linearis Panzer (unavailable synonym of viridis Linné, 1758)
Page 2354 – cited as available synonym of viridis
Remark.
Agrilus mucronatus Gory & Laporte (unavailable synonym of goryi Saunders, 1871)
Page 2110 – cited as available synonym of goryi
Remark.
Agrilus muscarinus Baudon (unavailable synonym of muscarius Kerremans, 1895)
Page 2196 – cited as available synonym of muscarius
Remark.
Agrilus mulleri Théry, 1925 (Australodraco; valid name)
Page 2369 – cited as muelleri
Remark. Original spelling of this patronymic name is mulleri (dedicated to Franklin Müller) not mülleri which would authorized the mandatory change of ü to ue (ICZN, Article 32.5.2). The name muelleri is an incorrect subsequent spelling.
Agrilus novicus Westhoff (unavailable synonym of viridis Linné, 1758)
Page 2356 – cited as available synonym of viridis
Remark.
Agrilus otini Théry, 1934 (synonym of lukuledianus Kerremans, 1907)
Page 2172 – cited as subspecies of lukuledianus
Remark. The act of
Agrilus parapupala Curletti, 1998a (valid name)
Page 2225 – cited as parapupalus
Remark.
Agrilus patrizii Théry, 1927 (valid name)
Page 2227 – cited as patrizzii
Remark. Original spelling of this name is patrizii not patrizzii which is an incorrect subsequent spelling.
Agrilus politus Weiss (unavailable synonym of cuprescens Ménétriés, 1832)
Pages 2048, 2356 – cited as unavailable synonym of cuprescens on page 2048 and as unavailable synonym of viridis on page 2356
The unavailable name politus cited by
Agrilus pseudroberti Fleischer, 1934 (unavailable synonym of cyanescens Ratzeburg, 1837)
Page 2055 – cited as pseudoroberti; unavailable synonym of cyanescens
Remark. The original spelling of this unavailable name proposed by
pupala Obenberger, 1935e (valid name)
Page 2255 – cited as pupalus
Remark. Original spelling of the name is pupala not pupalus. Following the ICZN, Article 31.2.2., the name is to be treated as a noun in apposition and original spelling is to be retained. The pupalus is incorrect subsequent spelling. See also parapupala.
Agrilus pygaera Obenberger (unavailable name)
Page 2257 – cited as valid name of species
Remark.
Agrilus pyri Blanchard, 1845 (available synonym of sinuatus Olivier, 1790b)
Page 2296 – cited as unavailable synonym of sinuatus
Remark.
Agrilus rugiplumbeus Cobos, 1964 (valid name)
Page 2274 – cited as rugiplumbus
Remark. Original spelling of this name is rugiplumbeus not rugiplumbus that is incorrect subsequent spelling.
Agrilus shamyl Obenberger, 1922 (subspecies of lineola Kiesenwetter, 1857)
Page 2167 – cited as shamyi; subspecies of lineola
Remark. Original spelling of the name is shamyl not shamyi which is an incorrect subsequent spelling.
Agrilus sibiricola Obenberger, 1924 (synonym of laticornis Illiger, 1803)
Page 2291 – cited as sibiricolus
Remark. Original spelling of this name is sibiricola not sibiricolus. The name is to be treated as a noun in apposition to the name of its genus as ruled be ICZN (Article 31.2.2). The name sibiricolus is an incorrect subsequent spelling.
Agrilus spiniger Gory & Laporte, 1839 (available synonym of spinamajor Chevrolat, 1838) Page 2302 – cited as unavailable synonym of spinamajor
Remark.
Agrilus sulcifer Bétis (unavailable synonym of hyperici Creutzer, 1799)
Page 2130 – cited as available synonym of hyperici
Remark.
Agrilus thurei Curletti, 1996 (valid name) correct original spelling
Page 2337 – cited as a synonym of turei
Remark. The name thurei Curletti, 1996 was originally dedicated to A. Thure which follows from the derivatio nominis as well as from the name of collector cited on the labels of type specimens. The name was changed by
Following the ICZN (Article 32.4): the original spelling of a name is the “correct original spelling”, unless it is demonstrably incorrect as provided in Article 32.5. The Article 32.5 requires clear evidence of an inadvertent error in the original publication itself without recourse to any external source of information. The publication of
The best solution for the stability of the nomenclature is the reversal to original state by following the ICZN Article 32.5. The name thurei Curletti, 1996 is a correct original spelling and the name turei Curletti, 2002 an unjustified emendation and available synonym of thurei.
Agrilus turei Curletti, 2002 (available synonym of thurei Curletti, 1996) unjustified emendation
Page 2337 – cited as valid name
Remark. See comments at thurei above.
Agrilus viridis Seidlitz (unavailable synonym of cuprescens Ménétriés, 1832)
Pages 2047, 2356 – cited as unavailable synonym of cuprescens on page 2047 and unavailable synonym of viridis on page 2356
Remark.
Agrilus zanthoxylumi Li Meng Lou, 1989 (valid name)
Page 2367 – Zhang and Wang are cited as authors.
Remark. The authorship of the name has been changed several times. Jendek & Grebennikov (2011) stated that Li Meng Lou (1989) is the first who established the name by presenting characters (ICZN, Article 13.1.1).
Agrilus lukesi Obenberger, 1936b new substitute name
Remark. The name modicus Kerremans, 1892b is replaced by its synonym lukesi Obenberger, 1936b due to primary homonymy with modicus Solier, 1833 (recently in Paragrilus).
Agrilus thomsoni Jendek new replacement name
Remark. The name is impressipennis Thomson, 1879 is replaced by a new replacement name thomsoni Jendek due to primary homonymy with impressipennis Uhler, 1855 (currently synonym of fallax Say, 1833).
Etymology: The name is patronymic and dedicated to Thomson.
Agrilus walkerianus Jendek (Aphanisticus) new replacement name
Remark. The name sulcicollis Walker, 1858 (recently in Aphanisticus) is replaced by a new replacement name walkerianus Jendek due to primary homonymy with sulcicollis Lacordaire, 1835.
Etymology: The name is patronymic and dedicated to Walker.
Agrilus gola Jendek new replacement name
Remark. The name filiformis Gory & Laporte, 1839 (originally proposed in Agrilus) is replaced by a new replacement name gola Jendek due to primary homonymy with filiformis Herbst, 1801 (originally proposed in Buprestis; currently synonym of viridis Linné, 1758).
Etymology: The name is derived from the first two letters of the names Gory and Laporte.
Aleš Smetana and Yves Bousquet (both Ottawa, Canada) are sincerely acknowledged for consultation on several nomenclatural problems.