The genus Lasinus can be readily separated from the other genera of the Pselaphodes complex (Hlaváč 2003) by a combination of the following characters: 1) head with well-defined setose frontal and vertexal foveae, 2) maxillary palpi small, with palpomeres III–IV symmetrical, neither roundly expanded nor projecting laterally, 3) antennal club three-segmented, with antennomeres VIII–IX often modified in males, 4) pronotal lateral and median foveae well-defined, 5) pronotum lacking antebasal sulcus connecting foveae, 6) pronotal longitudinal sulcus present, well- to weakly-defined, 7) tarsal segments II linear, segments III inserted at the apex of the II, 8) basal carinae on the first visible tergite (IV) present, short, 9) median setose metaventral fovea absent, 10) metaventral horny processes reduced to short, stout protuberances.

Length 2.5–3.8 mm. Head lacking dorsally visible postgenae, with well-defined setose frontal and vertexal foveae, maxillary palpi small, symmetric. Antennae with scapes distinctly longer than pedicels, club three-segmented, antennomeres VIII–IX often modified in males, species characteristic. Pronotum with well-defined median and lateral foveae, longitudinal sulcus sometimes weakly-defined but always present, antebasal transverse sulcus absent. Elytra with two basal foveae and two striae, lacking carinae, punctate, covered by short, golden pubescence. Metaventrite with two stout, short protuberances instead of long horn-like processes, metaventral apex sharp, with shallow excavation, surface punctate and pubescent, median metaventral fovea absent. Legs long and slender, roughly punctate and pubescent, protrochanters, mesotrochanters, profemora and mesofemora with spines of various shape and length, metatrochanters and metafemora lacking spines, tarsomeres II linear, tarsomeres III inserted at the apex of II.

Females of all species bear on mesotrochanters one or two more spines than males, females antennomeres VII–IX simple, without modification.

Due to the simple second tarsal segments and the absence of the median metaventral foveae, Lasinus is most closely related to the genera Paralasinus Hlaváč & Nomura, 2001, Pselaphodes Westwood, 1870 and Dayao Yin, Li & Zhao, 2011. From the latter two genera Lasinus can be separated by the completely symmetric palpomeres II–IV. In Pselaphodes and Dayao, the maxillary palpi have at least some palpomeres II–IV asymmetric, roundly expanded, or slightly to distinctly projecting laterally. Lasinus differs from Paralasinus by the pronotum lacking an antebasal sulcus, which is present in Paralasinus.

Members of the genus are usually collected by sifting leaf-litter in forested areas.

Distribution. China, Vietnam, Japan, Russia (Kuril Islands).

Tonkin (Ha Noi, Vietnam).

(4 ♀♀). LECTOTYPE, ♀, here designated: (h) Tonkin / red label (p) TYPE / (h) Lasinus mandarinus (p) A. Raffray det. / (p) MUSÉUM PARIS, 1917, COLL. A. RAFFRAY / red label (p) LECTOTYPE Lasinus mandarinus Raffray, Bekchiev, Hlaváč & Nomura des., 2013. (MNHN). PARALECTOTYPES, 3 ♀♀: same data as lectotype, bearing the following red label: (p) PARALECTOTYPE Lasinus mandarinus Raffray, Bekchiev, Hlaváč & Nomura des., 2013. (MNHN).

Redescription of this species given below is based on four females deposited in MNHN having a status of syntypes. Raffray (1890) also mentioned four females in his original description. One female is here designated as lectotype, and three remaining females paralectotypes, in order to ensure the stability of nomenclature and provide a unique name-bearing type for Lasinus mandarinus.

Body (Fig. 1) unicoloured, dark brown, maxillary palpi yellow, length 3.30–3.60 mm.

Head elongate, about 1.15 times longer than wide, slightly longer than pronotum; median sulcus well-defined along whole length of head. Genae simple, without protuberance.

Antennae very long, about 2.3–2.4 mm; scapes about 5 times longer than pedicels, pedicels short, as long as wide; antennomeres III–VII subequal in length, slightly shorter than V; antennomeres VI–VIII about as long as pedicels; IX slender, 2.25 times longer than wide, 1.10 times as long as X; late about 1.6 times longer than wide; terminal segments 1.2 times as long as X and 1.6 times longer than wide.

Pronotum as long as wide, gibbose, with prominent lateral swellings before lateral foveae; lateral and median setose foveae well-defined; median sulcus originates in median fovea and almost reaching anterior margin of pronotum.

Legs long and slender; protrochanters with small apical spine; profemora with long spine before middle; mesotrochanters at apex with two spines, outer one fairly bigger; mesofemora with minuscule spine at basal third.

First visible abdominal tergite (IV) glabrous, slightly more than two times longer than second (V); short basal carinae well-defined, distance between carinae 0.4 of maximal tergal width.

Lasinus mandarinus is close to Lasinus sinicus by the pronotum with prominent lateral swellings before lateral foveae, but it differs from the latter by the proportion of antennomere X which is 1.6 times as long as wide.

Vietnam (Tonkin, Ha Noi).

Nagasaki, Suwo-sama (=Suwa shrine).

LECTOTYPE, ♂, here designated: (h) Lasinus spinosus. Type D.S. Japan. Lewis. [label where the type specimen was originaly mounted] / (h) Lasinus spinosus ♂ (p) TYPE (h) D. S. / (p) Japan. G. Lewis / round label with red margin (p) TYPE / round label with blue margin (p) SYNTYPE / (p) Sharp Coll. 1905-313. / red label (p) LECTOTYPE Lasinus spinosus Sharp, Bekchiev, Hlaváč & Nomura, des., 2013 (BMNH). PARALECTOTYPE, 1 ♀, here designated: (h) Lasinus spinosus ♀ (p) TYPE (h) D. S. / (p) Japan. G. Lewis / round label with blue margin (p) SYNTYPE / red label (p) PARALECTOTYPE Lasinus spinosus Sharp, Bekchiev, Hlaváč & Nomura, des., 2013 (BMNH).

(8 ♂♂, 9 ♀♀). (1 ♂) Japan, Saga Pref., Kashima City, Mt. Kyogatake., 19.X.1986, S. Nomura leg.; (1 ♂) Japan, Fukuoka Pref., Hiko-san Mts., 3.V.1983, S. Nomura leg.; (1 ♂) Japan, Kumamoto Pref., Ueki-cho, 10.IV.1981, S. Naomi leg.; (1 ♂, 1 ♀) Japan, Kyushu, Nagasaki Pref., Isahaya-shi, Jôyama, Atagoyama, 18.III.1998, S. Nomura leg.; (1 ♂) Japan, Kyushu, Oita Pref., Shonai-machi, Nishi-Ohara, 20.VI.1998, K. Ôtsuka leg.; (1 ♂, 1 ♀) Japan, Kyushu, Miyazaki Pref., Tano-cho, Aoidake, 6.IX.1993, S. Nomura leg.; (1 ♂, 1 ♀) Japan, Fukuoka, Hikosan Mts., 27.XII.1982, S. Nomura leg.; (1 ♂, 1 ♀) Japan, Nagasaki City, Suwa Shrine, 2.V.1985, S. Nomura leg.; (1 ♀) Japan, Miyazaki Pref., Kiyotake-cho, Kaeda vall., 27.IV.1993, S. Nomura leg.; (2 ♀♀) Japan, Miyazaki Pref., Aya-Minami, 9.V.1985, S. Nomura leg.; (2 ♀♀) Japan, Kyushu, Oita Pref., Kujû Mts., Makinoto pass., 14.X.1991, S. Nomura leg. (NSMT, PCPH, NMNH, PCSK).

Redescription of this species given below is based on one male and one female deposited in BMNH having a status of syntypes. Sharp (1874) mentioned three specimens in his original description. One male is here designated as lectotype, another female is paralectotype, in order to ensure the stability of nomenclature and provide a unique name-bearing type for Lasinus spinosus.

Body (Fig. 2) unicoloured, reddish-brown, maxillary palpi yellow, length 2.90–3.10 mm.

Head elongate, about 1.15 times longer than wide, slightly longer than pronotum; median sulcus visible on rostrum and on vertex reaching level of vertexal foveae. Genae with weak protuberance, covered with erected, dense golden setae.

Antennae about 2.02 mm long (Fig. 11); scapes long, about 3.7 times longer than pedicels; pedicels about 1.18 times shorter than antennomeres III; antennomeres IV and V as long as wide; antennomeres VI 1.22 times shorter than pedicels; antennomeres VII 1.25 times shorter than VI; antennomeres VIII about 1.27 times longer than and distinctly wider than VII; IX about 1.5 times longer than wide and about same length as terminal antennomeres, IX in male with small and shallow discoidal plate on apical half bearing small pore-like structure with one long seta, in female unmodified; antennomeres X quadrate, 1.5 times shorter than IX; terminal antennomeres 1.6 times longer than X and about 1.5 times longer than wide.

Pronotum about as wide as long, wrinkly, evenly rounded before lateral foveae; lateral and median setose foveae well-defined; median longitudinal sulcus very thin, but distinct.

Legs long and slender; protrochanters with large apical spine; profemora with long spine in middle of its length; mesotrochanters at apex with one small (male) or two (female) spines, in some cases one spine is slightly stronger; mesofemora with minuscule spine at basal third.

Abdomen slightly wider than elytra, first visible abdominal tergite (IV) finely punctate with dense and short golden setae, about 4 times as long as next tergite, basal carinae very short, distance between carinae 0.4 of maximal tergal width. Aedeagus (Fig. 20) 0.64 mm long; median lobe weakly narrowed apically, with short and large apical lobe; endophallus with two spines and one lamella; dorsal spine very big, enlarged at apex, forming large plate; ventral spine long, acute at apex; lamella small finely dentate in apical part; parameres short and slender, not overlapping apical lobe.

Lasinus spinosus shares with Lasinus monticola, Lasinus mikado, Lasinus inexpectatus, Lasinus saoriae and Lasinus yamamotoi the evenly rounded pronotal lateral margins, but differs from all of these species by the shape of the antennae and aedeagus.

Japan (Kyushu).

Hiko (900 m), Fukuoka, Kyushu.

(21 ♂♂, 19 ♀♀). (3 ♂♂, 1♀): Japan, Shikoku, Ehime Pref., Oda-cho, Mt. Odamiyama, Buna st., 2.IX.1993, E. Yamamoto leg.; (1 ♂) Nara, Nara Park, 8.VIII.1980, I. Löbl leg.; (1 ♀) Japan: Honshu, Kanagawa Pref., Aikawa-chô Mt., Hasuge-san, 7.I.2006, T. Lackner leg.; (1 ♂) Japan, Fukushima pref, Okutadami, Alizu, Mt. Asakusadake, 22.VII.1987, S. Nomura leg.; (5 ♂♂) Japan, Shimane Pref., Kanagi-machi, Atoyama, 9.V.1991, T. Nakamura leg.; (2 ♂♂) Japan, Ehime Pref., Narukawa-keikoku 600–700 m, 1.II.1997, M. Sakai leg.; (1 ♂) Japan, Ehime Pref, Komi, Yanadani, 2.X.1994, M. Sakai leg.; (1 ♂, 7 ♀♀) Japan, Kyushu, Kagoshima Pref., Kirishima, Kurinodake Spa, 8.III.1999, H. Hoshina leg.; (1 ♂) Japan, Kyushu, Miyazaki Pref., Tsuno-chô, Mt. Osuzuyama, 700 m, 8.IX.1994, S. Nomura leg.; (2 ♂♂) Japan, Kyushu, Miyazaki Pref., Takachiho-chô, Onino-iwaya, 3.XII.1994, S. Nomura leg.; (1 ♂) Japan, Kyushu, Miyazaki Pref., Wanizukayama Mts., 6.IX.1993, S. Nomura leg.; (1 ♂, 1 ♀) Japan, Kyushu, Miyazaki Pref., Aya-chô, 10.II.1994, S. Nomura leg.; (1 ♂) Japan, Kyushu, Kagoshima Pref., Aira-chô, 30.I.1985, T. Tanabe leg.; (1 ♀) Japan, Honshu, Tokyo Pref., Fussa-shi, Tamagawa Riverside, Mutsumi-bashi, 12.II.2007, S. Nomura leg.; (1 ♀) Japan, Honshu, Tokyo Pref., Okutama, Nippara, Ogawadani, 4.IV.2006, S. Nomura leg.; (4 ♀♀) Japan, Kyushu, Nagasaki Pref., Unzen Mt. Kinugasayama, 16.III.2007, S. Nomura leg.; (1 ♀) Japan, Kyushu, Kagoshima Pref., Osumi Mt., Hoyoshidake, 19.III.1994, S. Nomura leg.; (1 ♂) Japan, Kioto, 11.VI.1881, G. Lewis leg.; (2 ♀♀) Japan, Miyanoshita, 11.VI.1881, G. Lewis leg. (BMNH, NSMT, PCPH, NMNH, PCSK).

Body bicoloured (Fig. 3), darker brown with more reddish elytra, maxillary palpi yellow, length 2.80–3.30 mm.

Head elongate, about 1.06 times longer than wide and as long as pronotum; median sulcus weakly defined along whole length of head. Genae with triangular, prominent protuberance, covered with erect, dense golden setae (Fig. 6).

Antennae about 2.23 mm long (Fig. 12); scapes long, about 4 times longer than pedicels; pedicels shortest, quadrate and as long as IV and V each; antennomeres III about 1.25 times longer than pedicels; antennomeres VI–VII subequal in length; VIII slightly shorter than VII; IX about 1.5 times longer than wide, with apical, nail-shaped protuberance on ventral side in male, in female unmodified; antennomeres X quadrate, 1.25 times shorter than IX; terminal antennomeres 1.5 times longer than X and about 1.5 times longer than wide.

Pronotum slightly longer than wide, wrinkly, evenly rounded before lateral foveae (Fig. 8); lateral and median setose foveae well-defined; median sulcus weakly-defined.

Legs long and slender; protrochanters with small apical spine; profemora with long spine before middle; mesotrochanters at apex with two (males) minuscule spines or three (females) spines, median one minuscule; mesofemora with minuscule spine at basal third.

Abdomen slightly wider than elytra, first visible tergite (IV) about 3 times as long as second (V), finely punctate with dense, very short golden setae; basal carinae well-defined but very short, distance between carinae 0.5 of the maximal tergal width. Aedeagus (Fig. 21) 0.66 mm long; median lobe weakly narrowed apically, with short and large apical lobe, curved downwards in middle; endophallus with two spines and one large lamella; dorsal spine large, enlarged in middle to form broad plate, acutely angled at apex, with one small tooth in middle; ventral spine short, acute at apex; lamella large; parameres long, overlapping apical lobe, enlarged apically.

Lasinus monticola is close to Lasinus mikado and Lasinus yakushimanus by the presence of nail-shaped protuberance on the antennomeres IX, butit differs from Lasinus yakushimanus by the shape of the genal region of the head and the pronotum, and it differs from Lasinus mikado by the absence of a deep concavity on antennomeres IX. Lasinus monticola can be readily separated from both species also by the shape of aedeagus.

Japan (Honshu, Shikoku, Kyushu).

(5 ♂♂, 3 ♀♀). HOLOTYPE, ♂, labelled as follows: (p) [Guanxi, CHINA], “locality in Chinese characters”, 25.V.1996, S. Uéno leg. / (p) ab. Liangshui 1710 m, Mt. Miao’ershan, Xing’an Xian. / red label (p) HOLOTYPE Lasinus sinicus sp. n., Bekchiev, Hlaváč & Nomura det., 2013. (NSMT). PARATYPES: 1♂: China, Vill, 86, Gansu: Mrijishan (h), 1000 m, Rougemont: 1 ♂, 2 ♀♀: same data as holotype, but specimens collected on 26.V.1996; 1 ♂, 1 ♀: China, Shaanxi, Nanwutaishan, 4.04.03, leg. Rougemont; 1 ♂: China, W Hubei, 21.VI–13.VII, Guanmenshan-1500 m, pit fall traps, 31.45N, 110.4E, leg. Jaroslav Turna, 2003. All paratypes bear the following red label: (p) PARATYPE Lasinus sinicus sp. n. Bekchiev, Hlaváč & Nomura det., 2013. (NSMT, MCSN, PCPH).

Description. Body (Fig. 4) bicolored, head, pronotum and abdomen almost black, elytra reddish-brown, maxillary palpi yellow, length 3.25–3.50 mm.

Head elongate, about 1.15 times longer than wide, slightly longer than pronotum; median sulcus absent along whole length of head. Genae simple, lacking protuberances.

Antennae long about 2.10 mm; scapes long, about 4 times longer than pedicels; pedicels shortest, 1.25 times shorter than antennomeres III; antennomeres III, IV, VII and VIII subequal in length, slightly shorter than V and VI; antennomeres XI 0.71 times longer than wide only slightly enlarged on the apex of antennomeres in male, in female unmodified; X 0.85 times longer than wide; XI 0.66 times longer than wide.

Pronotum about as long as wide, wrinkly, gibbose, with prominent lateral swellings before lateral foveae (Fig. 9); lateral and median setose foveae well-defined; median sulcus present only on disc, not originates from median fovea, very short and fine, largely separated from anterior margin of pronotum.

Legs long and slender; protrochanters with small apical spine; profemora with small spine in middle; mesotrochanters at apex with small median spine (male) or two (female) spines; mesofemora with small spine at basal third.

First visible abdominal tergite (IV) glabrous, very long, about 3.50 times longer than second (V); basal carinae very short, distance between carinae about 0.5 of maximal tergal width. Aedeagus (Fig. 22) 0.66 mm long; median lobe weakly narrowed apically, with short and very large apical lobe; endophallus with one large, bifid spine and one lamella; lamella large, with dentation on inner left part; parameres short and slender, not overlapping apical lobe.

Lasinus sinicus is close to Lasinus mandarinus by the similar shape of the pronotum with prominent lateral swellings before the lateral foveae. They can be separated from it by the proportion of antennomere X which is almost as long as wide in Lasinus sinicus.

The specific name is derived from China, where the species was discovered.

China (Guangxi, Gansu, Shaanxi, Hubei)

(14 ♂♂, 17 ♀♀). HOLOTYPE, ♂, labelled as follows: (p) [Japan, Miyanoshita, Lewis], red label (p) HOLOTYPE Lasinus mikado sp. n., Bekchiev, Hlaváč & Nomura det., 2013. (BMNH); PARATYPES (2 ♂♂, 1♀): Japan, Nanatsukahara, Shobara City, Hiroshima Pref., 10.X.1987, I. Okamoto leg.; (1 ♂) Japan, Honshu, Chiyoda-ku, Tokyo Pref., Fukiage Gyoen, Imperial palace, 19.I.2001, S. Nomura leg.; (1 ♂) same data, 8.V.2001; (1 ♀) 18.XII.2003; (2 ♀♀) 2.II.2004; (3 ♀♀) 12.II.2007; (1 ♂) Japan, Honshu, Fussa-shi, Tokyo Pref., Tamagawa riverside, Mutsumi-bashi, 12.II.2007, S. Nomura leg.; (1 ♂) Japan, Honshu, Akiu-machi, Miyagi Pref., Futakuchi Valley, 27.VII.1990, S. Nomura leg.; (3 ♂♂, 1 ♀) Japan, Honshu, Saitama Pref., Ranzan-machi, Kagamata, 5.IV.1996, K. Toyoda leg.; (1 ♂) Japan, Honshu, Chiba Pref., Kôzaki-jinja, Kôzaki-machi, 14.X.2001, S. Nomura leg.; (1 ♂, 2♀♀) Japan, Honshu, Niiharu-mura, Gunma Pref., Hôshi-onsen, 600 m, 20.X.2001, S. Nomura leg.; (2 ♀♀) Japan, Niigata Pref., Kuroiwa, Shibata, 22.XI.1990, H. Koike leg.; (1 ♂) Japan, Nara, 27–31.VII.1980, C. Besuchet leg.; (1 ♂) Japan, Shikoku, Ishizuchi Mts., Omogo Valley, 700 m, 18-25.VIII.1980, J. Peck leg.; (1 ♂) Japan, Ôhira, Shimamaki-mura, Hokkaido, 4-18. VI. 1994, S. Hori leg.; (1 ♂) Japan, Teshio-gawa, Teshio-chô, Hokkaido, 22. VII. 1992, S. Hori leg.; (4 ♂♂, 2 ♀♀) Russia, Kunashir Island, Tretiakovo VIII., 20.VII.1990, S. Kurbatov leg.; (1 ♀) same data, 18.VII.1990; (1 ♂) Japan, 1890, Schönfeldt leg.; (7 ♂♂, 10 ♀♀) Japan, Kanagawa, Sauter leg., (BMNH, NSMT, NHMW, NMNH, MHNG, PCPH, PCSK).

Body (Fig. 5) unicoloured, reddish-brown, maxillary palpi light brown, length 2.87–3.1 mm.

Head elongate, about 1.10 times longer than wide, slightly longer than pronotum; median sulcus visible on rostrum, on vertex reaching level of vertexal foveae. Genae with weak protuberance, covered with erected, dense golden setae.

Antennae about 2.15 mm long (Fig. 13); scapes long, about 3.4 times longer than pedicels; pedicels about 1.18 times shorter than antennomeres III, antennomeres V and VI as long as pedicels; VII 1.25 times shorter than VI; antennomeres VIII about 1.27 times longer and distinctly wider than VII; antennomeres IX about 1.5 times longer than wide, about the same length as terminal antennomeres, in male with deep ventral concavity on apical half terminating with nail-shaped protuberance, in female unmodified; antennomere X quadrate, 1.5 times shorter than IX; terminal antennomeres 1.6 times as long as X and about 1.5 times longer than wide.

Pronotum about as wide as long, wrinkly, evenly rounded before lateral foveae; lateral and median setose foveae well-defined; median sulcus thin.

Legs long and slender; protrochanters with large apical spine; profemora with long spine in middle of its length; mesotrochanters at apex with two minuscule (male) or two strong (female) and one minuscule spine; mesofemora with minuscule spine at basal third.

Abdomen slightly wider than elytra; first visible abdominal tergite (IV) finely punctate, with dense and long golden setae, about 4 times longer than second visible tergite (V); basal carinae very short, distance between carinae 0.53 of maximal tergal width. Aedeagus (Fig. 23) 0.61 mm long; median lobe weakly narrowed apically, with long and relatively narrow apical lobe; endophallus with one bifid spine and two small lamellas; lamellas finely dentate on the apical part; parameres long, overlapping apical lobe, enlarged at apex.

Lasinus mikado is close to Lasinus monticola and Lasinus yakushimanus by the presence of a nail-shaped protuberance on antennomeres IX, it differs from both by the presence of a deep concavity on antennomeres IX and by the shape of aedeagus.

The name is derived from the Japanese word – „mikado“, meaning the Emperor of Japan.

Japan (Hokkaido, Honshu, Shikoku), Russia (Kuril Islands).

The original type series of Lasinus spinosus in the Sharp collection is in fact a mix of Lasinus spinosus and Lasinus mikado sp. n.

(4 ♂♂, 1♀).HOLOTYPE, ♂, labelled as follows: (p) [Japan, Ehime, Nomura Dam, Nomura-cho, 27.V.1994, M. Sakai leg.] red label (p) HOLOTYPE Lasinus yamamotoi sp. n., Bekchiev, Hlaváč & Nomura det., 2013. (NSMT). PARATYPES: (2 ♂♂, 1♀); (1 ♂) same data with holotype; (1 ♀) same data with holotype but in 23.VII.1994; (1 ♂) Japan, Shikoku, Ehime Pref., Uchiko-chô, Shiromawari, 9.VII.1995, E. Yamamoto leg. (NSMT, PCPH, NMNH).

Body unicoloured, head, pronotum and abdomen reddish-brown, elytra slightly lighter, maxillary palpi yellow, length 2.85–2.95 mm.

Head elongate, 1.08 times longer than wide and as long as pronotum; median sulcus visible on rostrum, relatively shallow at level of vertexal foveae. Genae with weak protuberance, covered with erected, dense golden setae.

Antennae (Fig. 14) about 2.02 mm long; scapes long, about 2.85 times longer than pedicels; pedicels short, 1.42 times shorter than each of antennomeres III–IV; antennomeres V and VI of same length; antennomeres VII 1.22 times longer than wide; VIII longer than wide; XI longer than wide, simple, only slightly oblique on ventral side in male, in female unmodified; X slightly longer than wide; XI longer than wide.

Pronotum slightly longer than wide, surface evenly wrinkly, evenly rounded before lateral foveae; lateral foveae well-defined; median setose fovea small; median sulcus very thin.

Legs long and slender; protrochanters with long apical spine; profemora with short, strong spine in middle; mesotrochanters with one small (male) or two (female) spines; mesofemora with small spine at basal third.

First visible abdominal tergite (IV) very long, about four times longer than second visible tergite (V), with fine punctation in anterior part, disc glabrous, surrounded with short golden pubescence on sides; carinae short, distance between carinae 0.4 of maximal tergal width. Aedeagus (Fig. 24) 0.57 mm long; median lobe weakly narrowed apically, with large and relatively short apical lobe; endophallus with two spines and small lamella; ventral spine very big, forming large plate; dorsal spine big, curved, acute at apex; lamella small, finely dentate in apical part; parameres short and slender, reaching apical lobe, enlarged at apex.

Lasinus yamamotoi resembles Lasinus inexpectatus due to the unmodified antennal clubs in both sexes, but it can be distinguished from the latter by the proportion of the antennomeres VII and the shape of the aedeagus.

Patronimic, dedicated to Mr. Eiji Yamamoto (Japan).

Japan (Shikoku).

(3 ♂♂, 1♀). HOLOTYPE, ♂, labelled as follows: (p) Japan, Kyushu, Miyazaki Pref., Aya-chô, Ôtsuribashi, 25.IV.1993, leg. S. Nomura. red label (p) HOLOTYPE, Lasinus inexpectatus sp. n., Bekchiev, Hlaváč & Nomura det. 2013 (NSMT). PARATYPES, (2 ♂♂, 1♀): 1 ♂, 1 ♀, same data as holotype, 1 ♂: (p) Japan, Kyushu, Miyazaki Pref., Nangoh-choh, Ohshima (3), leg. Atsushi Nagai (NSMT, NMNH, PCPH).

Body unicoloured, reddish-brown, elytra slightly lighter, maxillary palpi light brown, length 2.80–3.20 mm.

Head about 1.05 longer than wide, and as long as pronotum; median sulcus very weakly-defined, slightly visible only on rostrum, absent on vertex. Genae with weak protuberance, covered with erected, dense golden setae.

Antennae 2.20–2.24 mm long (Fig. 15); scapes long, about 3.25 times longer than pedicels; pedicels slightly shorter than antennomeres III–VII combined, which are subequal; VIII slightly longer and about 1.30 times wider than VII; antennomeres IX rectangular, simple, in male slightly shorter (0.18 mm) than in the female (0.24 mm); X about as wide as long; terminal antennomeres 1.60 times longer than X.

Pronotum 1.05 times longer than wide, wrinkly, evenly rounded before lateral foveae; lateral and median setose foveae well-defined; median sulcus thin, present only on disc.

Legs long and slender; protrochanters with large apical spine; profemora with long spine in middle of its length; mesotrochanters at apex with one minuscule (male) or two strong (female) spines; mesofemora with minuscule spine at basal third.

Abdomen slightly wider than elytra; first visible abdominal tergite (IV) finely punctate with dense and long golden setae on sides, disc glabrous, about 3 times longer than second visible tergite (V); basal carinae very short, almost invisible in males. Aedeagus (Fig. 25) 0.63 mm long; median lobe weakly narrowed apically, with long and narrow apical lobe; endophallus with three spines and small lamella; ventral spine large, forming large plate, acute and curved at apex; dorsal spine large, curved, acute at apex; lateral spine very slender and small, acute at the apex; lamella small, finely dentate on apical part; parameres short, unequal in length, not overlapping apical lobe, enlarged at apex.

Lasinus inexpectatus is close to Lasinus sinicus due to the simple antennomeres IX lacking any sexual character. The two species can be separated by the absence of swellings before lateral foveae on the pronotum in Lasinus inexpectatus. From all other Japanese species, Lasinus inexpectatus also differs by the shapes of the aedeagus and the antennae.

The name is realated with the „unexpected” discovery of this species.

Japan (Kyushu).

(8 ♂♂, 7 ♀♀). HOLOTYPE, ♂, labelled as follows: (p) [Japan, Kyushu, Kagoshima Pref., Yakushima, Hananoegô, 17.III.2001, H. Hoshina leg.] red label (p) HOLOTYPE Lasinus yakushimanus sp. n., Bekchiev, Hlaváč & Nomura det., 2013 (NSMT). PARATYPES: (6 ♂♂, 3 ♀♀) same data as holotype; (1 ♂, 3♀♀) Japan, Kagoshima Pref., Yakushima, Mt. Nonkidake, 1350 m, 4.IX.2006, S. Nomura leg.; (1♀) Japan, Kagoshima Pref., Yakushima, Mt. Aikodake, 200 m, 5.IX.2006, S. Nomura leg. (NSMT, PCPH, NMNH, PCSK).

Body unicoloured, reddish-brown, maxillary palpi yellow dark, length 2.84–2.96 mm.

Head elongate, about 1.07–1.11 times longer than wide and about as long as pronotum; median sulcus visible on rostrum and on vertex reaching level of vertexal foveae. Genae with weak protuberance, covered with erected, dense golden setae (Fig. 7).

Antennae about 1.94 mm long (Fig. 16); scapes long, about 3.0 times longer than pedicels; pedicels as long as antennomeres III; antennomeres IV and V slightly longer than wide; antennomeres VI 1.62 times longer than wide; antennomeres VII as long as wide; antennomeres VIII as long as and slightly wider than VII; IX about 1.20 times longer than wide, in male with shallow ventral excavation on apical half terminating with small nail-shaped protuberance, in female unmodified; antennomeres X 1.21 longer than wide; terminal antennomeres about 1.27 times longer than wide.

Pronotum about as wide as long, wrinkly, with weak lateral swellings before lateral foveae (Fig. 10); lateral and median setose foveae well-defined; median sulcus thin.

Legs long and slender; protrochanters with large apical spine; profemora with longer spine in middle of its length; mesotrochanters at apex with two minuscule (male) or two strong and one minuscule (female) spines; mesofemora with minuscule spine at basal third.

Abdomen slightly wider than elytra; first visible abdominal tergite (IV) finely punctate, with dense, very short golden setae; carinae short, distance between carinae 0.49 of maximal tergal width. Aedeagus (Fig. 26) 0.56 mm long; median lobe weakly narrowed apically, with short and large apical lobe; endophallus with two spines; ventral spine large, forming narrow curved plate; dorsal spine slender, evenly curved, acute at apex; parameres long, overlapping apical lobe, enlarged at apex.

Lasinus yakushimanus is close to Lasinus okinawanus by sharing the presence of weak lateral swelling on the pronotum, and to Lasinus monticola by the presence of a nail-shaped protuberance on antennomeres IX, it differs from both by the shapes of the antennae and the aedeagus.

The name is associated with the name of the locality, Yakushima, where the speciments was found.

Japan (Yaku-shima Island).

(11 ♂♂, 9 ♀♀).HOLOTYPE, ♂, labelled as follows: (p) [Japan, Ryukyus, Kagoshima Pref., Amami-ôshima Is., Mt. Yuidake, 10.VIII.1984, S. Nomura leg., ], red label (p) HOLOTYPE Lasinus amamianus sp. n., Bekchiev, Hlaváč & Nomura det., 2013. (NSMT). PARATYPES: (4 ♂♂, 4 ♀♀) same data as holotype; (1 ♂, 1 ♀) Japan, Ryukyus, Kagoshima Pref., Amami-ôshima Is., Mt. Yuidake, 15.V.1983, S. Nomura leg.; (2 ♂♂, 1 ♀) Japan, Ryukyus, Kagoshima Pref., Amami-ôshima Is., Mt. Yuidake, 8.V.1987, S. Nomura leg.; (1 ♂) Japan, Kagoshima Pref., Tokunoshima Is., Mt. Inutabudake, 3.V.1988, S. Nomura leg.; (3 ♂♂, 3 ♀♀) Japan, Kagoshima Pref., Tokunoshima Is., Yonama, 4.V.1988, S. Nomura leg. (NSMT, PCPH, NMNH, PCSK).

Body unicoloured, reddish-brown, elytra slightly brighter, maxillary palpi yellow dark, length 2.80–3.00 mm.

Head elongate, about 1.10 times longer than wide, as long as pronotum; median sulcus shallow, reaching level of vertexal foveae. Genae with weak protuberance, covered with erected, dense golden setae.

Antennae about 1.96 mm long (Fig. 17); scapes about 3.40 times longer than pedicels; pedicels 1.40 times shorter than antennomeres III; III slightly longer than wide; IV as long as III; antennomeres V slightly longer than wide; VI about 1.20 times longer than wide; antennomeres VII 1.2 times longer than wide; VIII 1.16 times longer than wide; IX about 1.10 times longer than wide, in male with well-developed tubercles in apical ventral part, in female unmodified; antennomeres X as wide as long; terminal antennomeres about 1.47 times longer than wide.

Pronotum about as wide as long, wrinkly, with weak lateral swellings before lateral foveae; lateral and median setose foveae well-defined; median sulcus thin and deep.

Legs long and slender; protrochanters with large apical spine; profemora with long spine in middle; mesotrochanters at apex with one (male) or two (female) spines; mesofemora with minuscule spine at basal third.

Abdomen slightly wider than elytra, first visible abdominal tergite (IV) finely punctate with dense and long, golden setae; carinae short, distance between them 0.47 of maximal tergal width. Aedeagus (Fig. 27) 0.59 mm long; median lobe weakly narrowed apically, with long and narrow apical lobe; endophallus with two spines and one small lamella; ventral spine large, enlarged, forming large plate, acute at left apex; dorsal spine very big, acute at apex; lamella large, finely dentate on apical part; parameres short and slender, reaching apical lobe.

Lasinus amamianus and Lasinus saoriae differ from all other species of the genus by the shape of antennae, especially by the presence of tubercles on antennomeres IX. Lasinus amamianus can be readily separated from Lasinus saoriae by the proportion of antennomeres VII and VIII and by the shape of aedeagus.

The species name is associated with the name of the locality, Amami-ôshima Island, where most of the specimens were found.

Japan (Amami-ôshima, Tokunoshima Islands).

(6 ♂♂, 6 ♀♀). HOLOTYPE, ♂, labelled as follows: (p) [Japan, Ryukyus, Okinawa Is., Kunigami-son, Mt. Yonahadake, 22.III.2005, S. Nomura leg.], red label (p) HOLOTYPE Lasinus saoriae sp. n., Bekchiev, Hlaváč & Nomura det., 2013. (NSMT). PARATYPES: (2 ♂♂, 5 ♀♀) same data as holotype; (2 ♂♂, 1 ♀) Japan, Okinawa, Yona-Kunigami, 15.III.1985, leg. S. Nomura; (1 ♂) Japan, Okinawa, Kunigami, 16.III.1985, leg. S. Nomura (NSMT, PCPH, NMNH).

Body unicoloured, reddish-brown, maxillary palpi yellow dark, length 2.70–3.10 mm.

Head elongate, about 1.08 longer than wide, slightly shorter than pronotum; median sulcus shallow, reaching level of vertexal foveae. Genae with weak protuberance, covered with erected, dense golden setae.

Antennae about 1.8 mm long (Fig. 18); scapes long, about twice longer than pedicels; pedicels 1.22 times longer than antennomeres III; antennomeres IV as long as wide; antennomeres V slightly longer than wide; antennomeres VI about 1.33 times longer than wide; antennomeres VII and VIII slightly longer than wide; VII 0.88 times longer than wide; VIII 0.9 times longer than wide; IX about 1.26 times longer than wide, in male with short tubercles in apical ventral part, in female unmodified; antennomeres X about 1.33 longer than wide; terminal antennomeres about as long as wide.

Pronotum slightly longer than wide, wrinkly, evenly rounded before lateral foveae; lateral and median setose foveae well-defined; median sulcus thin and deep.

Legs long and slender; protrochanters with large apical spine; profemora with long spine in middle of its length; mesotrochanters at apex with one (male) or three (female) spines; mesofemora with minuscule spine at basal third.

Abdomen slightly wider than elytra; first visible abdominal tergite (IV) finely punctate, with sparse golden setae; carinae very small, distance between them 0.48 of maximal tergal width. Aedeagus (Fig. 28) 0.61 mm long; median lobe weakly narrowed apically; endophallus with two spines and two lamellas; ventral spine large, enlarged in middle, acute at apex; dorsal spine slender, acute at apex; dorsal lamella small, finely dentate on apical part; ventral lamella large; parameres very short and slender, not reaching apical lobe.

Lasinus saoriae strongly resembles Lasinus amamianus from which it differs essentially by the shapes of the antennae and the aedeagus.

The specific epithet is dedicated to Saori Takeuchi (Japan), a family friend of the first author.

Japan (Okinawajima Island).

(4 ♂♂, 4 ♀♀). HOLOTYPE, ♂, labelled as follows: (p) [Japan, Ryukyus, Okinawa Is., Okinawajima, Mt. Oppadake, Nakijin-son, 26.VI.1998, S. Nomura leg.], red label (p) HOLOTYPE Lasinus okinawanus sp. n., Bekchiev, Hlaváč & Nomura det., 2013 (NSMT). PARATYPES: (2 ♂♂, 3 ♀♀) same data as holotype; (1 ♂, 1 ♀) Japan, Ryukyus, Okinawa Pref., Nago-shi, Mt. Nagodake., 2.IX.2006, S. Nomura leg. (NSMT, PCPH, NMNH).

Body unicoloured, light reddish-brown, maxillary palpi yellow, length 3.0–3.2 mm.

Head elongate, about 1.03 longer than wide, and as long as pronotum; median sulcus visible on rostrum, on vertex reaching level of vertexal foveae. Genae with weak protuberance, covered with erected, dense golden setae.

Antennae long about 1.94 mm (Fig. 19); scapes 1.16 times longer than wide, 1.40 times as long as pedicels; pedicels as long as wide and as long as antennomeres III; antennomeres IV 1.25 times shorter than III; antennomeres V 1.20 times longer than wide; antennomeres VI 1.60 times longer than wide; antennomeres VII as long as wide; antennomeres VIII about 1.40 times longer and distinctly wider than VII; antennomeres IX as long as wide and about same length as terminal antennomeres, in male with large, shallow, in apical half highly inclined discoidal plate, in female unmodified; antennomeres X quadrate; terminal antennomeres 1.60 times as long as X and about 1.37 times longer than wide.

Pronotum about as wide as long, wrinkly, with weak lateral swellings before lateral foveae; lateral and median setose foveae well-defined; median longitudinal sulcus present.

Legs long and slender; protrochanters with large apical spine; profemora with longer spine in middle of its length; mesotrochanters at apex with one (male) or three (female) spines; mesofemora with minuscule spine at basal third.

Abdomen slightly wider than elytra, first visible abdominal tergite (IV) finely punctate with sparse, short golden setae; carinae short, distance between them 0.39 of maximal tergal width. Aedeagus (Fig. 29) 0.60 mm long; median lobe weakly narrowed apically, with long and narrow apical lobe; endophallus with two spines and two lamellas; ventral spine very large, short, curved downwards in middle, acute rightward at apex; dorsal spine very strong, acute at apex; dorsal lamella small finely dentate on apical part; ventral lamella large; parameres long and slender, different in length, left one overlapping apical lobe, right one shorter, .

Lasinus okinawanus is most closely related to Lasinus spinosus by the antennomeres IX with a shallow cavity, and the lack of an apical nail-shaped protuberance. It differs from latter by antennomeres IX having highly inclined shallow discoidal plate in apical half, and by the shape of the aedeagus.

The specific name is associated with the name of the type locality, Okinawa Island, where the type specimens were found.

Japan (Okinawajima Island).