Holotype: 1 ♂ (NHMB): [h]“Sichuan, 2500m \ Emei Shan \ 29°35'N/103°11'E”, [h]“CHINA, \ 22/24.VI.1990”, [h] “Cantharis \ thibetanomima \ Wittm. \ det. W. Wittmer”, [p]“HOLOTYPUS”, [p]“CANTHARIDAE \ CANTH00004087”. Paratypes: 1 ♂ (NHMB): same data to the holotype, [p]“CANTHARIDAE \ CANTH00003067”; 1 ♀ (NHMB): same data, [p]“CANTHARIDAE \ CANTH00003209”; 1 ♀ (NHMB): same data, [p] “CANTHARIDAE \ CANTH00003565”; 1 ♂ (NHMB): [p]“CHINA / Sichuan \ 103.20el/29.30nw \ Mt. Emei 500-1200m \ 4.-18.V.1989 \ S. &J. Kolíáč leg.”, [h]“thibetanomima”, [p]“CANTHARIDAE \ CANTH00003497”.

CHINA: Sichuan: 1 ♂, 1 ♀ (IZAS): Emei Shan, 2100–3100m, 25.VI.1955, leg. B.R. Ou.

Male (Fig. 1). Body black, except mandibles dark brown, antennomeres I–II brown on ventral sides, pronotum yellow, with a large black marking extending from anterior to posterior margin, elytra green, with strongly metallic shine.

Head rounded, surface matt on frons, densely punctate on vertex, eyes slightly protruding, head width acrossing eyes distinctly narrower than pronotum; terminal maxillary palpomeres nearly long-triangular, widest at basal one-third; antennae filiform, extending to elytral middle length, antennomeres II about 3 times longer than wide, III slightly longer than II, IV–XI each with a narrow, smooth longitudinal to oval groove nearly in middle of outer margin.

Pronotum wider than long, widest at anterior one-third, anterior margin straight, lateral margins sinuate, posterior margin bisinuate and narrowly bordered, anterior angles rounded, posterior angles slightly protruding, disc slightly convex at postero-lateral parts, surface lustrous, slightly largely and sparsely punctate.

Elytra nearly parallel-sided, about 3 times longer than width at humeri, about 4 times of length of pronotum, dorsum finely punctate, lustrous at anterior one-third parts, roughly but shallowly rugoluse-lacunose on the rest.

Legs: all outer tarsal claws each with a triangular lobe at base, inner claws simple.

Aedeagus (Figs 5–7): ventral process of each paramere narrow, distinctly shorter than conjoint dorsal plate; conjoint dorsal plate with apical margin widely emarginated in middle, lateroapical angles obtusely dentated, each side with a large tooth near apical margin on inner surface; laterophyse adhered to median lobe, with apex bent towards middle, the portion around the bending corner with upper margin slightly protuberant and bent dorsally.

Female. Similar to male, but eyes smaller, terminal maxillary palpomeres shorter, nearly widest in middle, antennae shorter, extending to elytral one-third length, antennomeres IV–XI without any groove, pronotum wider, elytra with lateral margins slightly diverging posteriorly, abdominal sternite VIII (Fig. 8) largely emarginated in middle and slightly emarginated on both sides of posterior margin, the portion between middle and each lateral emarginations rounded at apex.

Body length: 9.0–11.0 mm; width: 2.0–2.5 mm.

China (Sichuan). Excluded from Yunnan province at the moment.

In the study, the type series of this species were discovered to be plural and consist of two species. Except the true Taiwanocantharis thibetanomima, which was located in Sichuan, China, the paratypes from Yunnan, China designated in the original publication belong to another unknown species described below, Taiwanocantharis wittmeri sp. n. Besides, the photos of aedeagus provided by Wittmer (1997b) is not of Taiwanocantharis thibetanomima but Taiwanocantharis wittmeri sp. n. So it is necessary to redescribe and illustrate this species here.

Holotype♂ (NHMB): [p] “CHINA, Yunnan prov. \ 27°08'N 100°14'E, 2900 \ Yulongshan mts., -3500m \ Baishui 7.-12.VII.1990 \ Vít Kubáň leg.”, [p] “CANTHARIDAE \ CANTH00003678”. Paratypes: 1 ♀ (NHMB): same data to holotype, [p] “CANTHARIDAE \ CANTH00003695”; 1 ♀ (NHMB): same data, [p] “CANTHARIDAE \ CANTH00003057”; 1 ♀ (NHMB): same data, [p] “CANTHARIDAE \ CANTH00002059”; 1♂ (NHMB): [p] “YUNNAN, 23-24.JUN. \ Yulong Mts., 1993 \ 27.00N 100.12E \ Bolm lgt. 3200m”, [p] “CANTHARIDAE \ CANTH00003523”; 1 ♀ (NHMB): [p] “YUNNAN, 24-26 May \ Yulong mts., 1993 \ 27.01N 100.12E \ Bolm lgt. 3200m”, [p] “CANTHARIDAE \ CANTH00003665”; 1 ♀ (IZAS): [p] “YUNNAN 3300-3500m \ 27.07N 100.13E 1993\ YULONGSHAN mts. 20- \ Vít Kubáň leg. -21/6.”, [p] “IOZ(E) 217863”; 1 ♂ (IZAS): [p] “China N-YUNNAN \ 27°08'N 100°14'E \ Yulongshan mts. 2900- \ -3500m BAISHUI vill. \ lgt. D. Král 7-12/7’90”, [p] “IOZ(E) 217864”; 1 ♀ (NHMB): [p] “China Yunnan 1-19.VII \ HEISHUI 27.13N 100.19E \ 35km N of Lijiang \ legit S. Beĉváŕ 1992”, [p] “CANTHARIDAE \ CANTH00003632”; 1 ♂ (NHMB): [p] “CHINA: N. Yunnan \ 30km N of Lijiang \ 3000m, 3.VII.1990 \ L.& M. Bocák. lgt.”, [p] “CANTHARIDAE \ CANTH00004071”; 1 ♀ (NHMB): [p] “YUNNAN 2500-3100m \ 25.58N 100.21E 30/5 \ Jizushan mts. -3/6 \ Vit Kubáň leg. 1993”, [p] “CANTHARIDAE \ CANTH00003636”; 1 ♀ (NHMB): same data, [p] “CANTHARIDAE \ CANTH00003773”; 1 ♂ (NHMB): [p] “YUNNAN, 30 May-3 Jun \ JIZU MTS., 1993 \ 25.58N 100.21E \ Bolm lgt. 2800m”, [p] “CANTHARIDAE \ CANTH00003608”; 1 ♀ (NHMB): same data, [p] “CANTHARIDAE \ CANTH00003804”; 1 ♀ (NHMB): [p] “YUNNAN 2000-2800m \ 25.11N 100.24E \ Weibaoshan mts. \ W slope 25-28/6.92 \ David Král leg.”, [p] “CANTHARIDAE \ CANTH00003651”.

Male (Fig. 2). Body black, except mandibles dark brown, antennomeres I–II brown on ventral sides, pronotum yellow, with a large black marking extending from anterior to posterior margin, elytra mixed green and bronze, with strongly metallic shine.

Head rounded, surface matt on frons, densely punctate on vertex, eyes slightly protruding, head width acrossing eyes distinctly narrower than pronotum; terminal maxillary palpomeres nearly long-triangular, widest at basal one-third; antennae filiform, extending to elytral middle length, antennomeres II about 3 times longer than wide, III slightly longer than II, IV–XI each with a narrow, smooth longitudinal to oval groove nearly in middle of outer margin.

Pronotum wider than long, widest at anterior one-third, anterior margin straight, lateral margins sinuate, posterior margin bisinuate and narrowly bordered, anterior angles rounded, posterior angles nearly rectangular, not protruding, disc slightly convex at postero-lateral parts, surface lustrous, finely and sparsely punctate.

Elytra nearly parallel-sided, about 3 times longer than width at humeri, about 3.5 times of length of pronotum, dorsum finely punctate, lustrous at anterior one-third parts, roughly but shallowly rugoluse-lacunose on the rest.

Legs: all outer tarsal claws each with a triangular lobe at base, inner claws simple.

Aedeagus (Fig. 9–11): ventral process of each paramere narrow, nearly as long as conjoint dorsal plate; conjoint dorsal plate with apical margin triangularly emarginated in middle, lateroapical angles acutely dentated, each side with a small tooth near middle of apical margin on inner surface; laterophyse adhered to median lobe, with apex bent towards middle, the portion around the bending corner with upper margin slightly protuberant and bent dorsally.

Female. Similar to male, but eyes smaller, terminal maxillary palpomeres shorter, nearly widest in middle, antennae shorter, extending to elytral one-third length, antennomeres IV–XI without any groove, pronotum wider, elytra with lateral margins slightly diverging posteriorly, all tarsal claws simple, abdominal sternite VIII (Fig. 12) largely emarginated in middle and slightly emarginated on both sides of posterior margin, the portion between middle and each lateral emarginations tapered at apex.

Body length: 8.0–11.0 mm; width: 2.0–2.5 mm.

This species is similar to Taiwanocantharis thibetanomima, but can be distinguished from the latter by the following characters: all tarsal claws simple in female; aedeagus: ventral process of each paramere nearly as long as conjoint dorsal plate, conjoint dorsal plate with lateroapical angels acutely dentated, each side with a small tooth near middle of apical margin on inner surface; abdominal sternite VIII of female with the portions between middle and lateral emarginations tapered at apex.

China (Yunnan).

This species is named after the late distinguished taxonomist Dr. Walter Wittmer.

This new species was described based on a part of the paratypes, which are located in Yunnan, China, of Taiwanocantharis thibetanomima. What’s noted, two paratypes provided with the information as “Yulongshan Mts., Baishui, 2900–3500m, 27°08'N, 100°14'E, 7.–12.VII.1990, 22.–24.VI.1993”, according to the original publication by Wittmer (1997), can not found in the collection of NHMB. But four not three paratypes labeled as “Jizushan Mts., 2500–3100m, 25°58'N, 100°21'E, 30.V.–3.VI.1993” were found in NHMB, all of them were designated as the paratypes of Taiwanocantharis wittmeri sp. n.

Holotype♂ (MHBU): CHINA: Gansu: Wenxian, Huangtuling, 2350m, 9.VII.2003, leg. Y.B. Ba & Y. Yu. Paratypes: 1 ♀ (MHBU): same data to holotype; 1 ♀ (MHBU): same data, 8.VII.2003. 1 ♂ (NHMB): [Sichuan]: Da Tsien Lou [Kangding], 3.VI.(18)93 (collector, hand-writing, hardly readable).

Male (Fig. 3). Body black, except mandibles brown, antennomeres I–II brown on ventral sides, pronotum yellow, with a large black marking extending from anterior to posterior margin, elytra green, with strongly metallic shine.

Head rounded, surface matt on frons, densely punctate on vertex, eyes slightly protruding, head width acrossing eyes slightly narrower than anterior margin of pronotum; terminal maxillary palpomeres nearly long-triangular, widest at basal one-third; antennae filiform, extending to elytral middle length, antennomeres II about 3 times longer than wide, III slightly longer than II, IV–XI each with a narrow, smooth longitudinal to oval groove nearly in middle of outer margin.

Pronotum wider than long, widest at anterior one-third, anterior margin straight, lateral margins sinuate, posterior margin bisinuate and narrowly bordered, anterior angles rounded, posterior angles nearly rectangular, not protruding, disc slightly convex at postero-lateral parts, surface lustrous, finely and sparsely punctate.

Elytra nearly parallel-sided, about 3 times longer than width at humeri, about 4 times of length of pronotum, dorsum finely punctate, lustrous at anterior one-third parts, roughly but shallowly rugoluse-lacunose on the rest.

Legs: all outer tarsal claws each with a triangular lobe at base, inner claws simple.

Aedeagus (Figs 13–15): ventral process of each paramere narrow, slightly shorter than conjoint dorsal plate; conjoint dorsal plate with apical margin slightly emarginated in middle, lateroapical angles acutely dentated, without any tooth on inner surface; laterophyse adhered to median lobe, with apex bent towards middle, the portion around the bending corner with upper margin slightly protuberant and bent dorsally.

Female. Similar to male, but eyes smaller, terminal maxillary palpomeres shorter, nearly widest in middle, antennae shorter, extending to elytral one-third length, antennomeres IV–XI without any groove, pronotum wider, elytra with lateral margins slightly diverging posteriorly, abdominal sternite VIII (Fig. 16) largely emarginated in middle and slightly emarginated on both sides of posterior margin, the portion between middle and each lateral emarginations subrounded at apex.

Body length: 8.0–11.0 mm; width: 2.0–2.5 mm.

This species is similar to Taiwanocantharis thibetanomima, but can be distinguished from the latter by the aedeagus: conjoint dorsal plate of parameres with lateroapical angles acutely dentated, without any tooth on inner surface.

China (Gansu, Sichuan).

This specific name is derived from the Latin “a-” (none) + “dentatus” (toothed), referring to its conjoint dorsal plate of parameres without any tooth on inner surface.

Holotype♂ (MHBU): CHINA: Guangxi: Wuming, Damingshan, 1100m, 27.V.2011, leg. H.Y. Liu. Paratypes: 2 ♂♂, 1 ♀(MHBU): same data to holotype; 2♂♂(MHBU): same locality, 600–900 m, 25.V.2011, leg. H.Y. Liu; 1 ♂, 1 ♀(MHBU): same locality, 1230–1423m, 20.V.2011, leg. H.Y. Liu.

Male (Fig. 4). Head black, mouthparts yellow, slightly darkened at terminal maxillary and labial palpomeres, mandibles dark brown, antennae black, antennomeres I–II yellow on ventral sides, pronotum yellow, disc with two central and four prebasal small black markings, scutellum yellow, elytra green, with strongly metallic shine, thorax and abdomen yellow on ventral sides, abdominal sternites II–VIII each side with a small round black marking, legs yellow, tibiae black, with apical parts yellow on ventral sides, tarsi black.

Head rounded, surface lustrous, densely punctate on vertex, eyes strongly protruding, width acrossing eyes slightly narrower than pronotum; terminal maxillary palpomeres nearly long-triangular, widest at basal one-third; antennae filiform, extending to elytral middle length, antennomeres II about 3 times longer than wide, III slightly longer than II, IV–XI each with a narrow, smooth longitudinal to oval groove nearly in middle of outer margin.

Pronotum distinctly wider than long, widest at anterior one-third, anterior margin arcuate, lateral margins slightly sinuate, posterior margin arcuate and narrowly bordered, anterior angles rounded, posterior angles subrounded, not protruding, disc slightly convex at postero-lateral parts, surface lustrous, finely and sparsely punctate.

Elytra nearly parallel-sided, about 4 times longer than width at humeri, about 5 times of length of pronotum, dorsum finely punctate, lustrous at humeral parts, roughly but shallowly rugoluse-lacunose on the rest.

Legs: slender, all outer tarsal claws each with a triangular lobe at base, inner claws simple.

Aedeagus (Figs 17–19): ventral process of each paramere slightly widened, nearly as long as conjoint dorsal plate; conjoint dorsal plate slightly bent ventrally at apex in lateral view, with apical margin roundly emarginated in middle, each side of the emargination triangularly protuberant, the protuberance extending laterally into a short ridge on inner surface and bent ventrally, lateroapical angles blunt-coniformly dentated; laterophyses normal and separated on both sides of median lobe, with apices rounded.

Female. Similar to male, but eyes smaller, terminal maxillary palpomeres shorter, nearly widest in middle, antennae shorter, extending to elytral one-third length, antennomeres IV–XI without any groove, elytra with lateral margins slightly diverging posteriorly, abdominal sternites II–VII each side with a small round black marking, VIII (Fig. 20) moderately emarginated in middle and slightly emarginated on both sides of posterior margin, the portion between middle and each lateral emarginations rounded at apex.

Body length: 12.0–14.0 mm; width: 2.0–3.0 mm.

This new species is related to Taiwanocantharis satoi (Wittmer, 1997a), but can be distinguished from the latter by the following characters: pronotum with posterior angles not protruding, disc with a large central and four small prebasal black markings; legs yellow at coxae, trochanters and femora, black at tibiae and tarsi; abdominal sternite VIII of female moderately emarginated in middle and slightly emarginated on both sides of posterior margin.

China (Guangxi).

This specific name is derived from the Greek prefix “para-” (similar), referring to its close relationship to Taiwanocantharis satoi.

Cordicantharis drahuska: Holotype: 1 ♂ (NMPC): [p]“China: Shaanxi, 2.-4.7.1998 \ Qing Ling Shan mts., 3500m \ Hou Zen Zi-Tai Bai \ Jindra, Trýzna&Šafránek lgt.”, [p]“HOLOTYPUS \ Cordicantharis \ drahuska sp. n. \ V. Švihla det. 2003”.

Cantharis (s.str.) gansosichuana: Holotype: 1 ♂ (SKCR): [p]“CHINA: S Gansu \ Tepo [Tewo] 2500-2800m \ 26-28/vi/2001 \ S. Kazantsev leg.”, [p-h]“Cantharis \ gansosichuana \ sp. n. \ S. Kazantsev det. 2010”, [p]“HOLOTYPUS \ S. Kazantsev des.”.

CHINA: Sichuan: 2 ♂♂, 1 ♀ (NMPC): Erlangshan Mts., 2600m, E Luding, 14.–16.VI.2003, leg. S. Murzin. Gansu: 1 ♂, 6 ♀♀ (HBUM): Wenxian, Huangtuling, 2350m, 9.VII.2003, leg. Yi-Bin Ba & Yang Yu; 1 ♀ (HBUM): same data, 8.VII.2003. Ningxia: 1 ♂ (HBUM): Heshangpu Forestry center, 1.–2.VII.2008, leg. X.P. Wang & X.L. Liu; 1 ♀ (HBUM): same data, 2100m, 5.–6.VII.2008; 2 ♂♂, 13 ♀♀ (HBUM): Jingyuan, Erlonghe, 3.–4.VII.2009, leg. X.P. Wang & X.L. Zhao; 4 ♀♀ (HBUM): same locality, 23.VI.2008, leg. Hong-Fan Ran; 2 ♀♀ (HBUM): same locality, 6.VII.2009, leg. S.Y. Zhou & X.J. Meng; 1 ♀ (HBUM): same locality, 6.VII.2009, leg. H.F. Ran & S.S. Zhang; 4 ♀♀ (HBUM): same locality, 23.VI.2008, leg. H.F. Ran; 1 ♂, 4 ♀♀ (HBUM): Jingyuan, Xixia Forestry center, 9.–10.VII.2009, leg. X.P. Wang & X.L. Zhao; 1 ♂, 3 ♀♀ (HBUM): same locality, 27.–28.VII.2008, leg. X.M. Li, H.F. Ran & Q.Q. Wu; 3 ♂♂, 8 ♀♀ (HBUM): same data, 15.–16.VII.2008; 1 ♂ (HBUM): same locality, 27.VII.2008, leg. F. Yuan; 2 ♀♀ (HBUM): Wanghuanan Forestry center, 20.VI.2008, leg. H.F. Ran; 1 ♀ (HBUM): same locality, 3.–4.VII.2009, leg. G.D. Ren & Y.W. Ba; 2 ♀♀ (HBUM): Jingyuan, Dongshanpo, 8.VII.2009, leg. H.F. Ran & S.S. Zhang; 1 ♀ (HBUM): Hongxia Forestry center, 25.VI.2008, leg. H.F. Ran; 1 ♂ (HBUM): Longde, Fengtai Forestry center, 29.–30.VI.2008, leg. X.M. Li, H.F. Ran & Q.Q. Wu.

China (Sichuan, Gansu, Shaanxi, Ningxia). Newly recorded for Ningxia Hui Autonomous Region.

It had been noted by Švihla (2011) that Taiwanocantharis gansosichuana (Kazantsev, 2010) was possibly a junior synonym of Taiwanocantharis drahuska (Švihla, 2004) without examining the types of the former species. During our study, the holotypes of the both nominal species and a large series of additional material at our disposal were examined, and no difference between them was discovered, thereby we suggest Taiwanocantharis drahuska be synonymized with Taiwanocantharis gansosichuana.

Holotype:1 ♂ (NMPC): [p]“Laos: Hua Phan prov. \ Ban Saluei, Mt. Phu Phan \ 20.13°N, 103.95°E, 1300-2000m \ F. & L. Kantner lgt. \ 6-17.v.2004”, [p]“HOLOTYPUS \ Cantharis (s.str.) \ dedicata sp. n. \ V. Švihla det. 2005”.

CHINA: Yunnan: 2 ♂♂ (IZAS):Tengchong, Jietou town, Shaba, 1850m, 25.3926°N, 98.7035°E, 25.V.2006, leg. H.B. Liang & P. Hu.

China (new record: Yunnan); Laos.

Holotype: 1 ♂ (NHMB): [p]“N.E. BURMA \ Kambaiti, 7000ft \ 3/5 1934 \ R. Malaise”, [h]“Canthris \ malaisei \ Wittm. \ det. W. Wittmer”, [h]“HOLOTYPUS”, [p]“Naturhistorisches \ Museum Basel \ Coll. W. Wittmer”, [p]“CANTHARIDAE \ CANTH00003352”. Paratypes: 2 ♀♀ (NHMB): same data, 3. –7.V.1934; 1 ♀ (NHMB): same data, 12.IV.1934; 2 ♀♀ (NHMB): same data, 4.–8.VI.1934; 1 ♂, 1 ♀ (NHMB): same data, 2000m, 27.V.1934.

CHINA: Yunnan: 1 ♀ (NHMB): Gaoligong mts., 2200–2500m, 24°57'N, 98°45'E, 8.–16.V.1995, leg. Vit Kubáň; 1 ♀ (CAS): Lushui County, Pianma Township, Gangfang, 1675m, 26.12070°N/098.57830°E, 16.V.2005, attracted to uv and mv lights at night, Stop# HBL-05-11, H. B. Liang collector [CASENT 1036703].

China (new record: Yunnan); Myanmar.