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The main objective of this work is to contribute to the taxonomic knowledge of the species of Capitellidae reported for the Eastern Tropical Pacific. This catalogue includes the original name of each species, new names, synonymies, type localities, the museum or institution where the type material is deposited, revision of the material reported for the region by different authors, new examined material, previous reports from other regions of the world, and comments on systematics and distributions. The catalogue lists 43 species in 19 genera. Of these, 6 species were erroneously recorded for the region (Decamastus gracilis Hartman, 1963; Decamastus nudus Thomassin, 1970; Mastobranchus variabilis Edwing, 1984; Notomastus aberans Day, 1957; Notomastus americanus Day, 1973; Notomastus latericeus Sars, 1851) and 5 species are found here to be questionable records for the Eastern Tropical Pacific (Capitella capitata (Fabricius, 1780); Dasybranchus glabrus Moore, 1909; Decamastus lumbricoides Grube, 1878; Notomastus lineatus Claparède, 1870 and Notomastus tenuis Moore, 1909).
Capitellidae, Catalogue, taxonomy, Eastern Tropical Pacific
Capitellids are typical inhabitants of marine soft bottom sediments. They are important in the energy budgets of the environments due to their feeding habits and usually high abundances. Their morphology is very simple; most of the species are similar morphologically to earthworms. Generally they live buried in sand or mud and feed by assimilating the organic matter adhering to sediments and are thus regarded as non-selective deposit feeders (
The taxonomy of the Capitellidae is based almost entirely on the distribution, orientation, and structure of chaetae for identification at all levels (Doyle 1991). The recognition of the genera in this family has been based traditionally on the chaetal formula, in other words, the number of segments with capillary chaetae and, if present, the mixture of capillary chaetae and hooded hooks (
The capitellids of the Eastern Tropical Pacific have been studied by numerous authors included in the checklist of
In first instance the list of species was taken from a checklist of polychaetes of the Tropical Oriental Pacific (
Abbreviations of the institutions cited in the catalogue:
BMNH The Natural History Museum, London, UK
CAS-IZ California Academy of Sciences-Invertebrate Zoology, San Francisco, USA
ECOSUR El Colegio de la Frontera Sur Chetumal, México
CP ICMyL UNAM Polychaete Collection of the Instituto de Ciencias del Mar y Limnología, Universidad Nacional Autónoma de México, México
LACM-AHF Natural History Museum of Los Angeles County, Los Angeles, USA
MNHN Muséum National d'Histoire Naturelle, Paris, France
UANL Universidad Autónoma de Nuevo León, México
USNM National Museum of Natural History, Washington, DC, USA
HZM Zoologisches Institut und Zoologisches Museum der Universität, Hamburg
ZMUC Zoological Museum, University of Copenhagen
Results TaxonomyClass Polychaeta Grube, 1850
Family Capitellidae Grube, 1862
Amastigos acutusPiltz, 1977
http://species-id.net/wiki/Amastigos_acutus
Near Carpinteria Hope Ranch, CaliforniaUSA, Stn. 11A, April 5 1971, fine grained, sandy beach.
Holotype (LACM-AHF POLY 1239) and 1 Paratype (LACM-AHF POLY 1240).
(4 specimens) Baja California Sur: La Paz Bay, Ensenada de La Paz (UANL-6457), Stn. 1 [24°06'50.3"N, 110°25'12.0"W] (1 specimen), August 17 2005; (UANL-6459), Stn. 7 [24°09'00.0"N, 110°21'38.0"W] (1 specimen) November 27 2005; (UANL-6456), Stn. 11 [24°08'28.9"N, 110°25'41.9"W] (1 specimen), November 27 2005, 0.20 m, Coll. Daniel Hernández-Valdez (DHV), Guerrero: Petacalco Bay (UANL-6458), Stn. E-2 [17°58'26"N, 102°03'45"W] (1 specimen), december 9 1992, 7 m, Coll. Edgar Amador-Silva.
USA: California (
Upon examining the holotype, we noticed some morphological characteristics that were omitted in the original description: a) the presence of palpodes on the anterior end of the prostomium, and b) the structure described as the pygidium is, in fact, the rupture of an abdominal segment. The rest of the description agrees well with the features of the type material
Eunotomastus gordiodes Moore, 1909
http://species-id.net/wiki/Anotomastus_gordiodes
San Diego, California.
Syntypes (LACM-AHF POLY 1450-42), (LACM-AHF POLY 1451-42), (LACM-AHF POLY 1505-43), (LACM-AHF POLY 1487-38), (LACM-AHF POLY 2275).
(50 specimens) Baja California: Todos Santos Bay (ECOSUR-CAP-1), Stn. IV-2 [28°56'32.3"N, 113°33'15.4"W] (1 specimen), 13 m, October 8 1979, Coll. Sergio Salazar-Vallejo (SSV); Los Angeles Bay, Municipal Beach (UANL-6463) [28°56'32.3"N, 113°33'15.4"W] (26 specimens), 1 m, June 27 2005; Baja California Sur: Magdalena Bay (UANL-6245), Stn. E-9 [25°36'8"N, 112°0'8"W] (1 specimen), December 6 1996; (UANL-6246), Stn. E-2 [24°34'9"N, 111°58'9"W] (2 specimens), December 5 1996; 1 m, Coll. Victoria Díaz-Castañeda (VDC); Sonora: Puerto Peñasco, La Choya Bay (UANL-6462) [31°20'37.8"N, 113°38'01.7"W] (16 specimens) 1 m, June 29 2005; (UANL-6464) (4 specimens), 1 m, August 7 2006, Coll. Jesús Angel de León-González (JALG) and María Elena García-Garza (MEGG).
USA: California (
Lumbricus capitatus Fabricius, 1780:279.
http://species-id.net/wiki/Capitella_capitata
Greenland, NW Atlantic, Arctic Ocean
Neotype ZMUC- POL-1967 (not seen).
(181 specimens) Baja California Sur: La Paz Bay, Magdalena Bay [24°38'09"N, 112°80'73"W] (139 specimens), December 6 1996, Coll. JALG; Sonora: Guaymas Bay (UANL 5238), Stn. 2 [27°53'28"N, 110°54'14"W] (18 specimens), May 6 1999; (UANL 5241), Stn. 6 [27°55'09"N, 110°52'39"W] (4 specimens), May 4 1999; (UANL 5242), Stn. 3 [27°54'13"N, 110°53'59"W] (2 specimens), May 6 1999; (UANL 5244), Stn. 1 [27°55'21"N, 110°53'14"W] (1 specimen), May 6 1999; Río Escondido [27°54'04.2"N, 110°54'08.2"W] (17 specimens), August 9 2006, 0.50 m, Coll. JALG.
This species has been reported to be broadly distributed around the world: Greenland (
The specimens revised in this work from Western México are similar morphologically to the neotype of Capitella capitata redescribed by
Dasybranchethus fauveli Monro, 1931
http://species-id.net/wiki/Dasybranchethus_pacifica
Baja California Sur, Concepción Bay, El Quemadito Beach [26°45'33.1"N, 111°52'36.5"W], June 26 2005; Santispac mangrove [26°45'43.2"N, 111°53'31.0"W], 1 m, June 26 2005, Coll. MEGG and JALG. Silt with a mixture of thick and fine sand, intertidal.
Holotype (UANL-6336) and 1 Paratype (UANL-6337)
México: Baja California Sur, Concepción Bay.
Dasybranchus caducus (Grube, 1846)
http://species-id.net/wiki/Dasybranchus_caducus
Mediterranean Sea.
Not seen
Not seen
México, Baja California Sur, Gulf of California, La Paz Bay, El Mogote Beach (
The material examined by
http://species-id.net/wiki/Dasybranchus_glabrus
California, San Diego Bay
Not seen
(2 specimens)Baja California Sur: La Paz Bay, Enfermería Beach, Stn. E-19 [24°13'54.4"N, 110°18'22.3"W] (2 specimens), December 1985, mixed silt, sands and fragmented shells.
USA: California (
Dasybranchus glabrus was described from California, and recorded by
http://species-id.net/wiki/Dasybranchus_lumbricoides
Philippine Islands
Not seen
Baja California Sur: 3.4 miles S W of Punta Arena, Carmen Island, Stn. 1746 [25°46'00"N, 111°15'00"W, to 25°49'40"N, 111°15'30"W] (1 specimen), 29-35 m, March 18 1949, sand, mud, pebbles. Coll. Allan Hancock Foundation Velero IV (AHF); California, Corona del Mar, New Port, Harbor, Stn. 1450-42 [33°36'04"N, 117°52'48"W] (1 specimen), 1.51 m; Stn. 1451-42, [33°36'04"N, 117°52'48"W] (1 specimen), 1.2 m.
California (
Dasybranchus lumbricoides was described from the Philippine Islands and
http://species-id.net/wiki/Dasybranchus_parplatyceps
Sonora, Puerto Peñasco, Stn. Beach [31°16'40"N, 113°30'W]. Intertidal, in silt under basalt boulders (Holotype, January 21 1972, Paratype, April 16 1972), Coll. J.D. Kudenov.
Holotype (LACM-AHFPOLY 1111) and 1 Paratype (LACM-AHF POLY 1112).
(120 specimens) Baja California: Los Angeles Bay, Moradas Beach [28°57'10.8"N, 113°33'27.0"W] (1 specimen), May 25 1986; (2 specimens), August 23 1987 Coll. JALG and SSV; La Gringa Beach (UANL-6431) [29°02'17.0"N, 113°32'57.2"W] (2 specimens), 1 m, June 27 2005; (UANL-6429) (2 specimens), 2 m, August 4 2006, Col. JALG and MEGG; Baja California Sur: La Paz Bay, Caimancito Beach, Stn. 1 [24°12'37"N, 110°18'19"W] (5 specimens), 1 m, March 19 1989; Stn. C-33 [24°13'55.6"N, 110°18'23.7"W] (2 specimens), 1.5 m, November 29 1986; Stn. C-37 [24°11'35.6"N, 110°18'00.7"W] (9 specimens), 1.5 m, November 29 1986; Enfermería mangrove, Stn. E-19 [24°13'54.4"N, 110°18'22.3"W] (2 specimens), 1 m March 1986, Coll. JALG; Cabo Pulmo-Los Frailes, Stn. CP-988-2 [23°24'27"N, 109°24'26"W] (5 specimens), 4-7 m, September 18 1988, Coll. JRB-Z; Enfermería Beach [24°13'55.6"N, 110°18'23.7"W] (2 specimens), 1m, June 15 1987; El Saladito Beach (2 specimens), 1m, May 15 1987; El Presidente Beach (2 specimens), October 8 1987; (2 specimens ), 1m, October 10 1987, Coll. JALG & SISV; Concepción Bay, El Requesón Beach (UANL-6430) [26°38'17.7"N, 111°49'55.5"W] (1 specimen), July 19 1985; (UANL-6438) (9 specimens) 1m, August 20 1985, Coll. JALG; El Quemadito Beach (UANL-6433) [26°45'33.1"N, 111°52'36.5"W) (1 specimen), 1m, June 25 2005; Los Cocos Beach (UANL-6437) [25°44'39.1"N, 111°53'55.4"W] (3 specimens), 1 m, June 25 2005, Coll. JALG and MEGG; Magdalena Bay Estero Rancho Nuevo, Santa Marina Bay, (UANL-6428) [24°19'15"N, 111°25'05"W] (6 specimens), 3 m, June 15 1998; Western coast of Baja California Peninsula, San Juaníco Bay (UANL-6439) [26°02'12"N, 112°51'12"W] (2 specimens), January 21 1999, 30 m, Coll. JALG; Sonora: Guaymas, Varadero Beach (UANL-6427) [27°54'04.3"N, 110°52'07.7"W] (11 specimens), 1 m, July 1 2005, Coll. JALG and MEGG; Nayarit: María Madre Islands (UANL-6463) [21°38'14"N, 116°32'13.7"W] (1 specimen), 3 m, November 2 1979, Coll. SS-V.
México: Sonora, Station Beach, Puerto Peñasco (
Upon careful examination the specimens reported by Bastida-Zavala (1993 and 1995) as Notomastus (Clistomastus) lineatus, Dasybranchus glabrus, Decamastus gracilis and Leiocapitella glabra were all found, to correspond to Dasybranchus parplatyceps Kudenov, 1975.
http://species-id.net/wiki/Dasybranchus_platyceps
Punta Cholla, Sonora, Gulf of California, May 9 1941, intertidal.
Holotype(LACM-AHFPOLY 0431) and 2 Paratypes (LACM-AHFPOLY 0432).
México: Punta Cholla, Sonora, Gulf of California (
Genus Decamastus Hartman, 1963
http://species-id.net/wiki/Decamastus_gracilis
California, Redondo Canyon, south wall, 1.65 miles WSW of end of Redondo Bach Pier [33°49'42"N, 118°25'18"W], December 5 1952, Coll. Allan Hancock Foundation R/V Velero IV, 232 m, green mud, fine sand
Holotype(LACM-AHFPOLY 000435) and 1 Paratype (LACM-AHFPOLY 000436).
Baja California Sur, La Paz Bay, Sta.989-2 (1 specimen), September 18, 1989, 4-17 m, Coll. Rolando Bastida-Zavala.
USA: California (
Decamastus gracilis was described from Southern California (USA) and
http://species-id.net/wiki/Decamastus_nudus
S.W. of Madagascar, in Tuléar region.
Not seen
(21 specimens) Baja California: Rocks Consag, Stn. 39 [30°59.4'N, 114°04.1'W] (1 specimen), 106.4 m, silty sand, March 16 1985; Sonora: North Tiburón Island, Sta. 26 [29°23.3'N, 112°30.7'W] (7 specimens), 71.9 m, fine sand, March 14 1985; Cabo San Miguel, Sta. 20 [28°08.0'N, 112°45.8'W] (10 specimens), 54.1 m, fine sand, March 13 1985; Santa María Bay, Stn. 3 [25°02.4'N, 108°31.7'W] (3specimens), 32 m, March 10 1985; Punta San Marcial, Sta. 10 [25°58.6'N, 111°06.9'W] (7 specimens), 39 m, March 11 1985, Coll. PH-A & VS-W
Madagascar (
Decamastus nudus, described from Madagascar, was reported by Hernández-Alcánatra and Solís-Weiss (1993, 1998, 1999) for the Gulf of California, but these specimens differ from the original description of
Capitella filiformis Claparède, 1864 by subsequent designation by
http://species-id.net/wiki/Heteromastus_filiformis
Port-Vendres France, Mediterranean Sea.
Not found
(19 specimens). Mazatlán Sinaloa, Estero de Urías (EMU-418) [23°12'N, 106°23'W] (19 specimens), 0 m, January 1979, Coll. Agnes Rutgers (AR).
Italy (
Leiocapitella glabra Hartman, 1947
http://species-id.net/wiki/Leiocapitella_glabra
Holotype, 1 mile northwest of San Gabriel Bay, Espíritu Santo Island, Gulf of California, México, Stn. 1107–40 [24°26'39"N, 110°22'53"W to 24°26'19"N, 110°22'45"W], 8–10 m, February 13 1940, Allan Hancock Pacific Expedition of 1940, Velero III; Paratype, 2 miles southwest of Cedros Islands light, Stn. 1265–41 [28°20'33"N, 115°10'10"W to 28°20'45'N, 115°09'45"W], 16 m, February 28 1941, Allan Hancock Pacific Expedition of 1941, Velero III, in soft bottom.
Holotype (LACM-AHF POLY 0425) and 1 Paratype (LACM-AHF POLY 0205).
(8 specimens)Baja California Sur:Cabo Pulmo-Los Frailes, Stn. CP-988 [23°24'27"N, 109°24'26"W] (8 specimens), 4-7 m May 7–8 1989, Coll. JRB-Z.
México:Baja California Sur, Espiritu Santo Island and Cedros Island (
After examination of the specimens reported by
Leiochrides australis Augener, 1914
http://species-id.net/wiki/Leiochrides_hemipodus
7.5 miles north of west end of Santa Catalina Island, California, bearing 002, Sta. 2798–54 [33°36'00"N, 118°36'03"W] 117 m, May 25 1954, Allan Hancock Pacific Expedition of 1940. Velero IV, blue green mud.
Holotype (LACM-AHF POLY 0429) and 1 Paratype (LACM-AHF POLY 0430).
(3 specimens)Baja California Sur: Cabo Falso, Stn. 13775 [22°34'15"N, 109°32'30"W] (1 specimen), 441.96 m, January 22 1970; Jalisco: Cabo Corrientes, Stn. 13753 [19°41'15"N, 105°53'00"W] (1 specimen), 381 m, January 18, 1970; Stn. 13756 [19°51'00"N, 105°57'30"W] (1 specimen), 426.72 m, January 18 1970, Coll. Allan Hancock Pacific Expedition of 1940. Velero IV.
USA: California (
http://species-id.net/wiki/Leiochrides_pallidor
Monterey Bay, California.
Not seen.
California (
Leiochrides pallidor it is a poorly known species and, until now, all descriptions have been incomplete. Chamberlin in his description only mentions that the species has 12 thoracic chaetigers, others features are not useful to characterize that species.
Mastobranchus trinchesii Eisig, 1887
http://species-id.net/wiki/Mastobranchus_variabilis
Holotype (USNM), Florida, Tampa Bay, Gulf of México [27°36.5'N, 82°55.8'W], 12 m, clean sand, January 1980; 1 Paratype (USNM), Off Florida, St. Lucie Country, Hutchinson Island, Sta. 5 [27°22'22.08"N, 80°13'46"W], 10.3 m, Coll. D. Beaumariage, P. Camp and R. Gallagher, May 10 1972; and 1 Paratype (LACM-AHF), Sta. 723 [27°37.1'N, 82°55.1'W], 10 m, January 1980.
Holotype (USNM 81993); 2 Paratypes (USNM 81994); 3 Paratypes (LACM- AHF 001369).
(6 specimens) Sinaloa: El Fuerte River, Stn. 51 [25°39.9'N, 109°30.6'W] (2 specimens), 49.5 m, March 20 1985, silty sands; Nayarit: María Madre Island Stn. 62C [21°38.2'N, 106°31.9'W] (4 specimens), 29.7 m, March 22 1985, fine sands Coll. PH-A.
USA: Alabama, Florida and North Carolina (
Mastobranchus variabilis was described from the Gulf of México and later reported as Mastobranchus? variabilis by
Mediomastus californiensis Hartman, 1944
http://species-id.net/wiki/Mediomastus_ambiseta
Newport Harbour Bay, Corona del Mar, Californa, USA [33°36'04"N, 117°52'48"W], June 2 1942, intertidal.
Holotype (LACM-AHF POLY 1451-42, Poly 0449).
(41 specimens) Baja California: Todos Santos Bay, Stn. V-1 [116°39.4'N, 31°46"W] (5 specimens) 6 m, October 9 1979; [116°39.7'N, 31°45"W] (3 specimens), October 15 1989; San Quintín Bay, Stn. 13 [30°27'48"N, 115°57'46"W] (8 specimens), November 18 1982; Sinaloa: Off Ohuira Bay, Stn. B-2 [25°35'54"N, 109°20'12"W] (25 specimens), September 17 1989; Sinaloa: Topolobampo Bay (UANL-1613) [17°56'53"N, 102°5'9"W] (2 specimens), September 17 1989, Coll. JALG.
USA:California (
http://species-id.net/wiki/Mediomastus_californiensis
California, Tomales Bay, USA [28°25'N, 123°W], June 8 1941, intertidal.
Holotype (LACM-AHF POLY 0428).
(408 specimens) Baja California Sur: La Paz Bay, Ensenada de La Paz, Stn. 1 [24°06'50.3"N, 110°25'12.0"W] (10 specimens); Stn. 3 [24°07'09.9"N, 110°21'18.2"W] (1 specimen); Stn. 4 [24°08'29.9"N, 110°21'08.7"W] (20 specimens); Stn. 8 [24°09'50.7"N, 110°21'38.0"W] (8 specimens); Stn. 10 [24°09'55.1"N, 110°25'26.2"W] (42 specimens); Stn. 11 [24°08'28.9"N, 110°25'41.9"W] (7 specimens); Stn. 12 [24°07'37.3"N, 110°25'10.6"W] (4 specimens), 17 August 2005, 0.15 m; Stn. 1 (6 specimens); Stn. 2 [24°06'30.6"N, 110°24'05.1"W] (23 specimens); Stn. 3 (16 specimens); Stn. 6 [24°09'59.8"N, 110°19'37.5"W] (2 specimens); Stn. 8 (13 specimens); Stn. 11 (6 specimens) 5-7 m, 27 November 2005, Coll. DHV; Stn. 1 (17); Stn. 2 (59 specimens); Stn. 3 (51 specimens); Stn. 4 (2 specimens); Stn. 7 [24°09'00.0"N, 110°21'38.0"W] (1 specimen); Stn. 8 (6 specimens); Stn. 9 [24°10'13.9"N, 110°24'26.2"W] (3 specimens); Stn. 10 (9 specimens); Stn. 11 (1 specimen); Stn. 12 (1specimen) 30 m, 6 March 2006, Stn. 1 (5 specimens), Stn. 2 (17 specimens); Stn. 3 (3 specimens); Stn. 4 (4 specimens); Stn. 5 [24°08'53.4"N, 110°20'17.3"W] (1 specimen); Stn. 8 (7 specimens), Stn. 9 (4 specimens), Stn. 10 (15 specimens); Stn. 11 (4 specimens); Stn. 12 (1 specimen) 0.15 m, June 1 2006, Coll. DHV; Manglar Zacatecas (UANL-6425) [24°9'56.2"N, 110°25'55.6"W] (2 specimens), 24 June 2005; (UANL 6422) (15 specimens), 1 m, 1 August 2006; Enfermería Beach (UANL-6424) [24°13'55.6"N, 110°18'23.7"W] (2 specimens), 0.15 m, June 24 2005; El Conchalito Beach (UANL-6423) [24°08'23.4"N, 110°21'05.8"W] (20 specimens), 1 m, August 1 2006, Coll. JALG and MEGG.
USA: Oregon to California (
http://species-id.net/wiki/Mediomastus_setosus
El Salvador, La Herradura, Estero Jaltepeque, [13°17'N, 89°02'W to 13°13'N, 88°54'W], February, 1955.
Holotype (HZM P-19159). Not seen.
El Salvador, Jaltepeque Estuary (Hartmann-Schöder 1959,
Neoheteromastus lineus Hartman, 1960
http://species-id.net/wiki/Neoheteromastus_lineus
San Nicolas, California, USA, 1609 m.
Holotype (LACM-AHF POLY 0421).
(3 specimens) Baja California Sur:Cabo Pulmo-Los Frailes (1 specimen), Stn. CP-589-1 [23°24'27"N, 109°24'26"W], 4–7 m, May 7-8 1989, Coll. JRB-Z. South East of Gulf of California, Stn. Talud IV T-3–20 [24°27'N, 108°35'W] (1 specimen), 1, 200-1, 274 m, August 26 2000; Stn. Talud IV T-4–26 [24°56.3'N, 109°11.8'W] (1 specimen), 1, 200–1, 274 m, August 26 2000, Coll. Nuria Méndez-Ubach.
USA: California in deep slope and abyssal depths (
Genus Neomediomastus Hartman, 1969
Mediomastus glabrus Hartman, 1960
http://species-id.net/wiki/Neomediomastus_glabrus
Southern California, Santa Catalina basin, Stn. 2850 [33°30'N, 118°35'W].
Holotype (LACM-AHF POLY 0426).
USA: Basins off Southern California (
Neonotomastus glabrus Fauchald, 1972
http://species-id.net/wiki/Neonotomastus_glabrus
Baja California Sur, Colorado Point, San Jose Island Stn. 11792 [25°20'00"N, 109°58'30"W], 405 m, November 24 1967.
Holotype (LACM AHF POLY 1027).
México:Baja California Sur and Jalisco in abyssal depths (
Brazil, Stn. 49 [11°34'S, 37°22'W] 26 m, November 23 1961 intertidal in soft bottoms.
Syntype (MNHN POLY TYPE 1301).
(3 specimens) Sonora: Puerto Peñasco, La Cholla Bay (UANL- 6519) [31°20'37.8"N, 113°38'01.7"W] (1 specimen), 1 m, August 7 2006, Coll. MEGG. Sinaloa, El Fuerte River, Stn. 51 [25°42.1'N, 109°30.6'W] (1 specimen), 49.5 m, March 20 1985; Nayarit, María Madre Island, Stn. 62C [21°38.2'N, 106°31.9'W] (1 specimen), March 22 1985, Coll. PHA.
Brazil (
The material of
Notodasus magnus Fauchald, 1972.
http://species-id.net/wiki/Notodasus_dexterae
Panamá, Naos Island [8°53'N, 79°33'W], July 1969.
Holotype (LACM-AHF POLY-2190) and 29 paratypes.
Panamá, Naos Island (Fauchald 1973;
http://species-id.net/wiki/Notodasus_harrisae
México: Baja California Sur, La Paz Bay, El Tesoro beach, [24°15'16.1"N, 110°18'55.4"W], 1 m, August 1, 2006, Coll. JALG and MEGG.
Holotype (UANL-6510), 1 Paratype (LACM-AHF POLY 2211) , and 1 Paratype (MNHN 1506).
(46 specimens). Baja California: Los Angeles Bay, Municipal beach (UANL-6512) [28°57'10.8"N, 113°33'27.0"W] (1 specimen), 1 m, June 27 2005, Coll. JALG and MEGG; Baja California Sur: El Mogote beach, Stn. 89 (UANL-6505) [24°10'39"N, 110°22'75"W] (4 specimens), 0.50 m, October 2003, Coll. A. Chávez; Ensenada de La Paz, Stn. 2 (UANL-6506) [24°06'30.6"N, 110°24'05.1"W] (1 specimen), 0.15, cm November 27 2005; Stn. 5 (UANL-6507) [24°08'53.4"N, 110°20'17.3"W] (2 specimens), 0.15 cm, June 5 2006, Coll. DHV; El Requesón beach (UANL-6508) [26°44'39.1"N, 111°49'55.5"W] (1 specimen), 1 m, June 25 2005; El Quemadito beach, (UANL-6509) [26°45'33.1"N, 111°52'36.5"W] (1 specimen), 1 m June 26 2005, Colls. JALG and MEGG; Balandra beach (UANL-6511) [24°19'18.1"N, 110°19'29.4"W] (35 specimens), 1 m, August 1 2006, Coll. JALG and MEGG; Sinaloa: Mazatlán, Estero de Urías (ICMyL EMU-420) [23°12'N, 106°23'W] (1 specimen), 0 m, January 1979, Coll. A.R.
México: Baja California, Baja California Sur, Sinaloa (
http://species-id.net/wiki/Notodasus–hartmanae
México:Paredón, Mar Muerto, Chiapas [16°03'26"N, 93°52'34"W], 0.5 m, April 14 2008, Coll. JALG, in compact mud, among numerous tubes of Diopatra rhizophorae Grube, with high content of organic matter.
Holotype (UANL-6513), 6 Paratypes (UANL-6514), 3 Paratypes (LACM-AHF POLY 2212) and 3 Paratypes (MNHN1507).
México: Chiapas (
http://species-id.net/wiki/Notodasus_kristiani
Sonora, Guaymas, Varadero beach [27°54'04.3"N, 110°52'07.7"W] 1 m, July 01 2005, Coll. MEGG and JALG.
Holotype (UANL-6515), 20 Paratypes (UANL-6517), 2 Paratypes (LACM-AHF POLY 2213) and 1 Paratype (MNNH 1508).
(2 specimens) Baja California: Los Angeles Bay, Municipal beach (ECOSUR) [28°56'32.3"N, 113°33'57.2"W] (1 specimen), 1 m, May 24 1986, Coll. P. Sánchez and E. Espinosa; Baja California Sur: Santa Marina Bay, Estero Rancho Nuevo (UANL-6515) [24°19'15"N, 111°25'05"W] (1 specimen), 3 m, June 21 1998, Coll. JALG.
México:Baja California, in mud pockets between Mytilus edulis beds, Baja California Sur, in soft sediments retained into Nextier boxes, Sonora, in mud with high content of organic matter (
http://species-id.net/wiki/Notodasus_magnus
Isla Carmen, Punta Arena, Gulf of California [25°46'00"N, 111°15'00"W; 25°49'40"N, 111°15'30"W], 29-35 m, March 18 1949, in silts with sand, mud and cobbles.
Holotype (LACM-AHF POLY 1031).
México: Baja California Sur, Isla Carmen (
Notomastus latericeus Sars, 1851.
http://species-id.net/wiki/Notomastus_aberans
South Africa, Kosi Bay
Holotype (ANO3 1961.16. 75–76)
(1 specimen) Western coast of Baja California Sur, Stn. A-3 (UANL-0038) [24°11.4'N, 111°23.5'W] (1 specimen), 74 m, June 17 1987.
South Africa (
Notomastus aberans was described from South Africa and
http://species-id.net/wiki/Notomastus_abyssalis
Baja California Sur: Punta Colorado, San José Island, Stn.11788 [25°21'00"N, 110°05'00"W], November 24 1967, 393m depth, only known for deep waters, from 375 to 481m, in silts with fine sand.
Holotype (LACM-AHF POLY 1012).
(10 specimens)Baja California Sur: Cabo Falso, Stn. 13774 [22°35'00"N, 109°35'00"W] (1 specimen), 438 m, January 22 1970; Stn. 13775 [22°34'00"N, 109°35'45"W] (1 specimen), 441 m, January 22 1970; Punta Colorado, San Jose Island, Stn. 11788 [25°21'00"N, 110°05'00"W] (1 specimen), 393 m, November 23 1967; Stn. 11792 [25°20'00"N, 109°58'30"W] (1 specimen), 405 m, November 23 1967; Stn. 11793 [25°20'00"N, 109°59'00"W] (2 specimens) 408 m November 23 1967; Sinaloa: Mazatlán, Crestón Island, Stn. 11761 [23°05'00"N, 107°59'20"W] (2 specimens) 405 m, November 14 1967; Jalisco: Cabo Corrientes Stn. 13752 [19°41'08"N, 105°53'30"W] (1 specimen) 375 m, January 18 1967; Nayarit: San Juanito Island, Tres Marías Island Stn. 13765 [21°37'00"N, 106°57'30"W] (1 specimen), 481 m, January 20 1970.
México:Baja California Sur, Southern and central part of the Gulf of California, and Nayarit, Tres Marías Islands (
http://species-id.net/wiki/Notomastus_americanus
North Carolina, Beaufort, June 4 1965.
Holtype (USNM 43118) and 14 Paratypes (USNM–43119).
(57 specimens), Baja California: Punta Willard, Stn. 34 [30 11.5N, 114 31.7'W] (1 specimen), 32.2 m, 15 March 1985; Rocks Consag, Stn. 39 [30°59.4"N, 114°04.1"W] (12 specimens), 106.4 m, March 16 1985, silty sand; Baja California Sur: Santa Inés Bay, Stn. 49 [26°59.6'N, 111°50.4'W] (3 specimens), 100 m, March 19 198; Sonora: Punta Arboleada, Stn. 14 [26°46.6'N, 110°06.7'W] (8 specimens), 92 m, March 12 1985; Stn. 15 [26°51.1'N, 110°06.5'W] (13 specimens), 48.8 m, March 12 1985; Cabo Tepoca, Stn. 44 [30°02.4'N, 112°55.4'W] (9 specimens), 100 m, March 17 1985, Sinaloa: Santa María Bay, Sta. 3 [25°02.4'N, 108°31.7'W] (1 specimen), 32 m, March 10 1985; Stn. 4 [24°56.9'N, 108°31.7'W] (4 specimens), 79m March 10 1985; Río Fuerte, Stn. 50, [25°46.8'N, 109°35.4'W] (4 specimens), 97 m March 20 1985; Stn. 51 [25°42.1'N, 109°30.6'W] (2 specimens), 49.5 m March 20 1985, Coll. PHA.
USA: North Carolina (
Notomastus americanus
http://species-id.net/wiki/Notomastus_angelicae
Sinaloa, West Río Fuerte [25°39'54"N, 109°28'36"W], 28.6 m, March 1985.Silt with fine sand.
Holotype (USNM 180697), 5 Paratypes (USNM 180698) and 5 Paratypes (LACM-AHF-POLY-1902).
(104 specimens) Baja Clifornia: Punta San Marcial Stn. 10 [25°58.6'N, 111°06.9'W] (4 specimens), 39 m, March 11 1985; Punta Willard, Stn. 30 [30°11.59'N, 114°31.7'W] (1 specimen), 32.2 m, 15 March 1985, ; Baja California Sur: western coast, Stn. E-14 [25°38'12"N, 112°23'18"W] (2 specimens), 80 m, Coll. JALG; Santa Inés Bay, Stn. 49 [26°59.6'N, 111°50.4'W] (3 specimens), 100 m, 19 March 1985; Stn. 49B [26°59.4' N, 111°53.5'W] 68 m, 19 March 1985; Sonora: Punta Arboleada, Stn. 14 [26°46.6'N, 110°06.7'W] (1 specimen), 92 m, March 12 1985; Cabo Tepoca, Stn. 44 [30°02.4'N, 112°55.4'W] (9 specimens), 100 m, March 17 1985; Sinaloa: Santa María Bay, Stn. 4 [24°56.9'N, 108°31.7'W] (4 specimens), 79 m, March 10 1985; Stn. 5 [24°54.6'N, 108°45.3'W] (2 specimens), 120 m, 10 March 1985; Río Fuerte, Stn. 50 [25°46.8'N, 109°35.4'W], (9 specimens), 97m; Stn. 51 [25°42.1'N, 109°30.6'W] (1 specimen), 49.5 m, March 20 1985; Nayarit: Islas Marías Stn. 62C, [21°38.2'N, 106°31.9'W] (56 specimens), 29.7 m, March 22 1985, Coll. PHA.
México:Baja California Sur, Sonora, Sinaloa (
http://species-id.net/wiki/Notomastus–cinctus
Baja California Sur, Cabo Falso [22°34'00"N, 109°32'30"W], 1, 450 m January 22 1970.
Holotype (LACM AHF POLY 1026).
.México: Baja California Sur, Cabo Falso; Guerrero, Zihuatanejo Bay, Nayarit, in 1, 200 to 1, 450 m (
http://species-id.net/wiki/Notomastus_hemipodus
North Carolina, Beaufort, June 15-18 1940, intertidal fine sands and mud.
Holotype (LACM-AHF POLY 0414), 1 Paratype (LACM-AHF POLY 1697), 1 Paratype (LACM-AHF POLY 0415), 2 Paratypes (LACM-AHF POLY 1701) and 5 Paratypes (LACM-AHF POLY 1709).
(57 specimens) Baja California: Consag Rocks, Stn. 38 [31°08.3N, 114°13.3"W] (4 specimens), 71.9 m, 16 March 1985; Stn. 39 [30°59.4"N, 114°04.1"W] (5 specimens), 106.4 m, March 16 1985; Santa Inés Bay, Stn. 49C [26°59.2"N, 111°58.3"W] (5 specimens), Coll. PHA; Baja California Sur: West Coast (UANL- 6486), Stn. 25 BIP II (1 specimen), October 6 1998, Colls. Eduardo Balart and EAS; (UANL 6485) (2 specimens), Rancho Bueno, Santa Marina Bay [24°08'23.4"N, 110°21'05.8"W], 1 m, January 21 1999, Coll. JALG; La Paz Bay, Ensenada de La Paz (UANL-6482), Stn. 12 [24°07'37.3"N, 110°25'10.6"W] (1 specimen), 0.15 m, November 27 2005, Coll. DHV; (UANL-6484), Stn. 10-A [24°09'55.1"N, 110°25'39.6"W] (1 specimen), 1 m, November 27 2005, Coll. DVH; El Conchalito beach (UANL-6481) [24°08'23.4 N, 110°21'05.8"W] (2 specimens), 1 m, June 24 2005; Concepción Bay, Los Cocos beach (UANL-6483) [26°44'39.1 N, 111°53'55.4"W] (2 specimens), 1 m, June 25 2005; Santispac mangrove (UANL-6487) [26°45'43.2N, 111°53'31.0"W] (6 specimens), 1 m, August 3 2006, Colls. JALG and MEGG; Sonora: Punta Arboleda Stn. 15 [26°51.1"N, 110°06.5"W] (13 specimens), 49.8 m, March 12 1985; Estero Tastiota Stn. 48 [28°16.4"N, 111°36.6"W] (5 specimen), 36.9 m, 18 March 1985; Sinaloa: Santa María Bay, Stn.3 [25°02.4"N, 108°31.7"W] (1 specimen), 32 m, March 10 1985; Stn. 5 [24°54.6"N, 108°45.3"W] (5 specimens), 79 m, March 10 1985; El Fuerte, River Stn. 52 [25°39.9"N, 109°28.6"W] (5 specimens), 28.6 m, March 20 1985, Coll. PHA.
USA: Beaufort, North Carolina (
http://species-id.net/wiki/Notomastus_latericeus
Norway (Atlantic Ocean)
Not seen
(15 specimens) Baja California: Stn. 30 [11.59'N, 114°31.7'W] (3 specimens), 32.2 m, March 1985; Punta Willard, Stn. 34 [30°11.5'N, 114°31.7'W] (1 specimen), 32.4 m, March 15 1985; Baja California Sur: Santa Inés Bay, Stn. 49B [26°59.4'N, 111°53.5'W] (1 specimen), 68 m, March 19 1985; Sonora: Punta Arboleada, Stn. 14 [26°46.6'N, 110°06.7'W] (1 specimen), 92 m; Cabo Tepoca, Stn. 44 [30°02.4'N, 112°55.4'W] (5 specimens), 104.1 m, March 17 1985; Sinaloa: Santa María Bay, Stn.4 [24°56.9'N, 108°31.7'W] (4 specimens), 32 m, Coll. PHA.
Norway (
Notomastus latericeus Sars, 1851 was described from Norway and has been reported widely around the world.
http://species-id.net/wiki/Notomastus_lineatus
Gulf of Naples (Mediterranean Sea).
Not seen
(7 specimens), Baja California: Cabo San Miguel, Sta. 19 [28°10.4'N, 112°48.1'W] (1 specimen), 30.4 m March 13 1985; Baja California Sur: Banco Gorda, Sta. 55 [23°08.7'N, 119°28.3'W] (1 specimen), 32.5m, March 21 1985; Santa Inés Bay, Sta. 49C [26°59.2'N, 111°58.3'W] (1 specimen), 28.9 m, March 19 1985; Punta San Marcial, Sta. 8 [25°33.4'N, 110°59.8'W] (1 specimen), 52 m, March 11 1985; Sta. 9 [25°47.8'N, 110°03.8'W] (1 specimen), 77.5 m, March 11 1985, Coll. PHA; La Paz Bay, Caimancito beach, Stn. C37 (2 specimens), November 29 1986, Coll. JRB-Z
Gulf of Naples, Mediterranean Sea (
The specimens reported by
http://species-id.net/wiki/Notomastus_magnus
USA: California, Tomales Bay.
(1)Paratype (LACM-POLY 0413) and 1 Paratype (LACM-POLY 2217).
(6 specimens) Baja California Sur: Magdalena Bay [25°36'8"N, 112°0'8"W] (1 specimen), 1.5 m, December 6 1996, Coll. JALG; Sonora: Puerto Peñasco, La Cholla Bay [31°20'37.8"N, 113°38'01.7"W] (5 specimens), 1.5 m, August 7 2006, Coll. JALG and MEGG.
USA: California (
http://species-id.net/wiki/Notomastus_polyodon
South Viet Nam, Nha Trang Bay, Stn. 26411. Soft bottoms, intertidal zone.
Holotype (LACM-AHF POLY 0301)
(59 specimens) Baja California Sur: west coast, Stn.A-3 (UANL-0038) [24°11.4'N, 111°23.5'W] (1 specimen), 74 m, June 17 1987, Coll. JALG.Baja California Sur: La Paz Bay, Caleritas beach (1 specimen), Stn. Ca-29 N-8, November 29 1986, Coll. JRB-Z; Ensenada de La Paz (UANL-6521), Stn. 1 [24°06'50.3"N, 110°25'12.0"W] (1 specimen); (UANL-6522), Stn. 4 [24°08'29.9"N, 110°21'08.7"W] (2 specimen); (UANL-6523), Stn. 6 [24°09'59.8"N, 110°19'37.5"W] (1 specimen); (UANL 6524), Stn. 7 [24°09'00.0"N, 110°21'38.0"W) (1 specimen); (UANL-6525), Stn. 8 [24°09'50.7"N, 110°23'35.9"W] (3 specimens); (UANL-6526), Stn. 9 [24°10'13.9"N, 110°24'26.2"W] (3 specimens); (UANL-6527), Stn. 10 [24°09'55.1"N, 110°25'39.6"W] (7 specimens); (UANL-6528), Stn. 11 [24°08'28.9"N, 110°25'41.9"W] (2 specimens), 0.15 m, Coll. DHV; Balandra beach (UANL-6529) [24°19'18.1"N, 110°19'29.4"W] (3 specimens), August 1 2006; Concepción Bay, Santispac beach (UANL 6530) [26°45'45.3"N, 111°53'30.0"W] (1 specimen), 1 m, June 26 2005, Coll. MEGG and JALG; El Quemadito beach, (UANL-6531) [26°45'33.1"N, 111°52'36.5"W] (14 specimens), 1 m, June 26 2005, Coll. MEGG and JALG; Sonora: Puerto Peñasco, La Cholla Bay (UANL-6532) [31°20'37.8"N, 113°38'01.7"W] (2 specimens), 1 m, June 29 2005. Sinaloa: Mazatlán, Punta Cerritos beach (UANL -6533) [23°18'28.6"N, 106°27'21.3"W] (21 specimens), 0.50 m, May 22 2006, Coll. JALG and MEGG; Estero de Urías (UANL-6534)[23°12'00"N, 106°23'00"W] (13 specimens), 0.50 m, January 1979, Coll. A.R.
South Viet Nam (
Notomastus polyodon was previously known only from the western Pacific, this is the first formal record from the eastern Pacific.
http://species-id.net/wiki/Notomastus_precocis
USA: Santa Catalina Gulf, 1400-2000m.
Holotype (LACM-AHF POLY 0416).
(6 specimens)Baja California Sur: Natividad Island Stn. 7358 [27°35'45"N, 115°08'30"W to 27°32'15"N, 115°05'00"W] (3 specimens), 182–201 m, April 21 1961; Jalisco: Cabo Corrientes, Stn. 13754 [19°41'15"N, 105°53'00"W] (2 specimens), 365 m, January 18, 1970; Nayarit: Punta Piedras, Stn. 13767 [21°54'30"N, 106°50'00"W] (1specimen), 246 m January 20 1970.
USA: California (
http://species-id.net/wiki/Notomastus_sonorae
Sonora, La Cholla Bay, Puerto Peñasco, march 28 1971. Intertidal, in sediments with fine sand.
Holotype (LACM-AHF POLY 1113).
(11 specimens) Sonora: Puerto Peñasco, La Cholla Bay (UANL- 6536) [31°20'37.8"N, 113°38'01.7"W], 1 m, June 29 2005, Coll. JALG and MEGG.
México: Sonora, (
http://species-id.net/wiki/Notomastus_tenuis
San Diego, California, USA, 1902-1903, intertidal. Coll. E. C. Starks.
Holotype (CAS-019718).
(80 specimens)California: Anaheim Slough Collections: (LACM-AHF), Stn. 903-38 (7 specimens), December 5 1938; (5 specimens), Stn. 905–38, December 7 1938; (2 specimens), Mission Bay, Stn. 1211–38; Baja California: Las Animas Island (LACM-AHF), Stn. F3893-11831 [28°41'30"N, 113°02'00"W] (1 specimen), 207 m, December 1 1967, Coll. VELERO IV; Baja California Sur: (LACM-AHF), Cabo Falso, Stn. F4285-13774 [22°35'00"N, 109°35'00"W] (1 specimen), 426.72 m, January 22 1970, Coll. VELERO IV; Sonora, Punta Arboleda (ICMyL-UNAM), Stn. 15 [26°51.1'N, 110°06.5'W] (13 specimens), 49.8 m, March 12 1985, Coll. PHA; Estero Tastiota (ICMyL-UNAM), Stn. 48 [28°16.4'N, 111°36.6'W] (5 specimens), 60.2 m, March 18 1985, Coll. PHA; Sinaloa: El Fuerte River (ICMyL-UNAM), Stn. 52 [25°39.9'N, 109°28.6'W] (5 specimens), 28.6 m, March 20 1985, Coll. PHA; Santa María Bay (ICMyL-UNAM), Stn. 3 [25°02.4'N, 108°31.7'W] (1 specimen), 32 m March 10 1985 ; Stn. 05 [24°54.6'N, 108°45.3'W] (5 specimens), 120 m, March 10 1985; Jalisco: Cabo Corrientes (LACM-AHF), Stn. F4205-13756 [19°51'30"N, 105°58'00"W] (1 specimen), 426.72 m January 18 1970, Coll. VELERO IV. Nayarit: Punta Piedras, San Juanito Island (LACM-AHF), Stn. F4296-13768 [21°53'00"N, 106°50'00"W] (1 specimen), 256 m January 20 1970, Coll. VELERO IV; North Rock Consag (ICMyL-UNAM), Stn. 38 [31°08.3'N, 114°13.3'W] (4 specimens), 71.9 m March 16 1985; Stn. 39 [30°59.4'N, 114°04.1'W] (12 specimens), 106.4 m, March 16 1985, Coll. PHA.
USA: California (
Notomastus tenuis was described from California by
Rashgua rubrocincta Wesenberg-Lund, 1949
http://species-id.net/wiki/Rashgua_lobatus
Baja California, Espíritu Santo Island, México, Stn. 1107-40, 6–10 m, intertidal in soft bottoms.
Holotype (LACM-AHF POLY 1107–40), 1 Paratype (LACM-AHF POLY 0411) and 1 Paratype (LACM-AHF POLY 1069-40).
(173 specimens) Baja California Sur: La Paz Bay, Caimancito beach, Stn. C-37 [24°08'23.4"N, 110°18'00"W] (8 specimens), 1.5 m, February 29 1986, Coll. Héctor Salaices-Polanco; Ensenada de la Paz (UANL-6554), Stn. 3 [24°07'09.9"N, 110°21'18.2"W] (12 specimens), 0.20 m, November 2005; (UANL-6555), Stn. 2 [24°06'30.6"N, 110°24'05.1"W] (52 specimens), 0.20 m, March 6 2006; (UANL-6556), Stn. 2 (106 specimens), 27 m, 1 June 2006, Coll. DHV; Sonora: Guaymas, Varadero beach (UANL-6557) [27°54'04.3"N, 110°52'07.7"W] (3 specimens), 1 m, July 1 2005, Coll. JALG and MEGG.
USA: Gulf of México, Mobile Bay, Mississippi, North Carolina (
Scyphoproctus djiboutiensis Gravier, 1904.
http://species-id.net/wiki/Scyphoproctus_oculatus
California, New Port Bay, intertidal, in soft bottoms with fine sand and shells fragments.
Holotype (LACM-AHF POLY 11108, POLY 0481) and 1 Paratype (BMNH AN03 1959.12.18.1).
(13 specimens) Baja California Sur: La Paz Bay, San Gabriel (UANL -6559) [24°18'44.3"N, 110°20'10.7"W] (2 specimens), August 31 2004; Concepción Bay, Santispac beach (UANL-6561) [26°45'45.3"N, 111°53'30.0"W] (3 specimens), 0.50 m, June 25 2005, Colls. JALG and MEGG.
USA: Southern California (
In this study we first found 43 species of Capitellidae from Eastern Tropical Pacific belonging to 19 genera. Following analysis of type and non-type material we conclude that this list should be reduced to 32 valid species in 17 genera. Of these, 14 species were originally described from the Eastern Tropical Pacific.
The species Capitella capitata
Amastigos acutus (
We thank Leslie Harris (LACM AHF), Kristian Fauchald (USNM), Alexander Muir (BMNH) and Tarik Meziane (MNHN) for their assistance and hospitality during our visits to these museums. Likewise we thank Vivianne Solís-Weiss (ICMyL-UNAM), Rolando Bastida-Zavala (UMAR), Sergio I. Salazar-Vallejo (ECOSUR) who allowed us to revise the material in their collections, and Pablo Hernández Alcántara for his help in the ICML revisions. We also wish to thank Harlan Dean (UM-B), Vivianne Solís-Weiss (ICMyL-UNAM) and Alexandra Rizzo (UERJ) for the critical reading of the manuscript. This study was partially financed by grants FOSEMARNAT 2004-CO1-245, CONACyT 61609, and H-029 from the Comisión Nacional para el Conocimiento y Uso de la Biodiversidad (CONABIO).