Modified from Rasnitsyn and Brothers (2009): Head hypognathous, without ocular carina, without subantennal groove accommodating antennal base; vertex with or without longitudinal median sulcus; antenna filiform, with scape short to moderate, never elongate, flagellomeres variable in number (14–19), most often 16, without apparent sexual dimorphism; mandibles often asymmetrical, with 3–4 teeth. Pronotum short medially; mesoscutum with notauli and median sulcus; axillae meeting anterior to mesoscutellum or separated by scutoscutellar sulcus; propodeum areolate. Forewing with costal space moderate to wide; C and pterostigma present; basal sections of Rs and M subequal in length, not continuously aligned (i.e., not forming smooth basal vein); cell [2R1] (i.e., marginal cell) closed, wide (not triangular), moderately short to moderately long; cells [1Rs] and/or [2Rs] closed; cell [1M] closed, small, distant from [1Rs]; cell [2M] open or delimited by spectral or nebulous 2m-cu; 1cu-a antefurcal. Hind wing with no posterobasal lobe; 4–5 distal hamuli; basal cell [R] closed; free apex of Rs present, short to long; free apices of M and Cu short or absent, that of A absent. Legs with trochantelli; tibial spur formula 1-2-2; tarsi pentamerous; tarsal plantulae present in some females; pretarsal claws with preapical tooth. Metasoma rather short, compact, not much sculptured, attached low on propodeum, first segment longest [articulatory ring referred to by Rasnitsyn and Brothers (2009) is not evident and apparently an incorrect interpretation], apical sternum of female elongate, nearly reaching metasomal apex; ovipositor external but not very long, sheaths at most as long as metasoma, not fitting tightly to ovipositor (often preserved detached).

Andyrossia Rasnitsyn and Jarzembowski, 2000 (Wealden, England, Barremian; a replacement name for Arossia Rasnitsyn and Jarzembowski in Rasnitsyn et al. 1998); Ahiromaimetsha Perrichot, Azar, Nel, and Engel gen n. (Lebanese amber, Neocomian); Iberomaimetsha Ortega-Blanco, Perrichot, and Engel gen. n. (Spanish amber, Lower Albian); Guyotemaimetsha Perrichot et al., 2004a (French amber, latest Albian/earliest Cenomanian); Afrapia Rasnitsyn and Brothers, 2009 (Orapa Mine, Botswana, Turonian); Afromaimetsha Rasnitsyn and Brothers, 2009 (Orapa Mine, Botswana, Turonian); Maimetshorapia Rasnitsyn and Brothers, 2009 (Orapa Mine, Botswana, Turonian); Maimetsha Rasnitsyn, 1975 (Taimyr amber, Santonian); and Ahstemiam McKellar and Engel gen. n. (Canadian amber, Campanian). Table 1 summarizes the species diversity of the family.

Male: total body length (excluding mandibles and parameres) 1.7 to 2.9 mm; compound eyes oval and bulging, large, 0.66× height of head; vertex without longitudinal medial line; fine occipital carina present; mandibles asymmetrical, right with four teeth and left with three teeth; scape flattened laterally, short, its maximal width 0.9× its length; pedicel nearly as broad as long, 0.75× length of scape; 14 flagellar articles cylindrical in shape and progressively shortening and slightly narrowing; maxillary palpus apparently with five palpomeres; labial palpus apparently with three palpomeres, all short; forewing with pterostigma moderately long, shorter than [2R1]; r-rs originating within apical one-third of pterostigmal length; 2R present as long stub, about half length of 1R; [1M] small, nearly rhombic, with first abscissa of Rs+M one-half length of second abscissa (2Rs+M), so that apical corner (junction of Rs+M and 1m-cu crossvein) positioned anterior to pterostigma; [1Rs] triangular, with Rs and 1rs-m crossvein both fade to become nebulous posteriorly; cross-veins 2rs-m and 2m-cu at most spectral, generally absent; hind wing with cells [R] and [Cu] enclosed by nebulous veins, with free apex of Cu.

Female: very similar to male but larger, total body length (excluding mandibles and ovipositor) 3.7 to 4.1 mm; pedicel more elongate, 0.75× as broad as long, 0.85× length of scape; legs with tarsal plantulae; ovipositor apparently simple, not serrate or toothed.

Guyotemaimetsha is most similar to Maimetsha in the absence of cell [2Rs]. It differs in cell [1Rs] being anteriorly petiolate instead of sessile, the hind wing with a free apex of Cu, the absence of a medial line on the vertex, and the presence of tarsal plantulae in females.

Holotype MNHN.F.A30174 (♂); paratype MNHN.F.A30175 (♂); in amber collection of the Museum National d’Histoire Naturelle, Paris, France.

Specimens IGR.ARC-1 (♀, missing anterior part of the head and with cuticle somewhat altered by taphonomy), IGR.ARC-309.11 (sex unknown, missing apices of right forewing and metasoma), IGR.ARC-370.7 (= scan ESRF-A-018g, reconstructed, ♀, in opaque amber), IGR.ARC-378.1 (♂, missing head, most metasoma and apices of all legs), IGR.ARC-378.2 (scan ERSF A-401-378, not reconstructed; ♂), IGR.ARC-393.4 (= radiograph ESRF-A-053d, not reconstructed; ♂), IGR.CDL-2.5 (♂, with forewing fragment in IGR.CDL-2.16, missing apical flagellomeres of left antenna; ex collection Arnaud), IGR.CDL-2.33 (♂, missing left antenna and apices of right antenna, right forewing, and all legs; ex collection Arnaud). All specimens deposited in the amber collection of the Department of Geosciences of the University Rennes I, France.

IGR.ARC-171 (♂, missing head, anterodorsal part of metasoma, and forelegs); IGR.ARC-191 (♂?, missing part of the head and mostly obscured, preserved in very dark amber); IGR.ARC-275 (♀, missing parts of mesosoma, left legs, and with metasoma mostly obscured by a bacterial film). These specimens agree in most respects with the definition of Guyotemaimetsha enigmatica but little is preserved and attribution to this species cannot be asserted. All specimens deposited in the amber collection of the Department of Geosciences of the University Rennes I, France.

Font-de-Benon quarry, 1 km east of Archingeay, Charente-Maritime, France.

Lithological subunit A1sl-A, Uppermost Albian–Lowermost Cenomanian, middle Cretaceous.

As for the genus (see above).

Male: Integument reddish-brown in color, with dense, minute punctures and fine, decumbent pubescence. Head transverse, with length approximately 0.8x width; ocelli separated from each other by one ocellar diameter; antennae inserted between compound eyes, closer to each other than to compound eye margin or posterior clypeal margin; scape globular in lateral view, laterally compressed in frontal view, with faintly concave apex; pedicel with sides convex; flagellum with dense coat of short, inclined setae, with first article longest, twice length of pedicel; penultimate article as broad as long; apical article with rounded apex; clypeus transverse, about twice as broad as long, with anterior margin rounded and posterior margin straight; mandibles large, with outer margin convex, overlapping apically when closed; right mandible with basal tooth large, second and third teeth equally smaller, apical tooth largest; left mandible with basal tooth large, median tooth smallest, apical tooth largest. Mesosoma compact, 0.7x as high as long; pronotum in dorsal view reduced to a short neck; mesoscutum broader than long, about 0.4x mesosomal length, abruptely truncate anteriorly to form a flat vertical surface, bordered by a transverse carina at anterolateral corners, with low dorsal convexity; median mesoscutal sulcus crenulate, notauli deeply impressed, slightly diverging anteriorly; mesoscutellum with low dorsal convexity, small triangular axillae separated anteriorly by large foveate groove; propodeum areolate, gradually sloping to pronounced posterior lip; meso- and metapleuron with anterior margin foveolate. Forewing hyaline, with costal cell distinctly enlarged around midlength; Rs junction with Sc+R forming an acute angle; cell [1M] with vein M slightly arched, roughly aligned with Rs, vein Cu shortest, M and Rs+M subequal in length; cells [1M] and [1Rs] well distant from each other, separated by long vein 2Rs+M; 2Cu significantly longer than 1Cu and 1m-cu; M and Cu nebulous at apex, reaching wing margin; 2A fading to nebulous vein posterior of [2Cu]. Hind wing with Sc+R, R, M+Cu, Cu, and A tubular, other veins nebulous; Sc+R and R more heavily sclerotized than M+Cu, Cu, and A; Cu fading to nebulous vein apically; free apex of Rs and Cu short. Metacoxa approximately 1.4x length of pro-and mesocoxae, all with broad bases and distinct apical constriction, in lateral view inserted posteroventrally on mesosoma; metafemur distinctly enlarged around midlength; all tibiae approximately subequal to femora in length, with basitarsus length slightly less than combined length of all subsequent tarsomeres. Metasoma compact, as long as mesosoma; parameres trapezoidal, narrowed toward apex, with outer surface slightly convex, with short erect setae around apex; cerci small, spatulate, inserted just anterior to parameres.

Female: Very similar to male but with pedicel cylindrical and each leg with a small plantar lobe (or tarsal plantula) on ventral apex of tarsomeres I–IV; ovipositor approximately as long as metasomal length, apex acute. Integument sculpturing and pubescence not visible, altered on specimen IGR.ARC.1 and not reconstructed in microtomographic scan of specimen IGR.ARC-370.7.

Iberomaimetsha rasnitsyni Ortega-Blanco, Perrichot & Engel, sp. n.

Antennae with 16 articles; pedicel straight; notauli parallel; forewing costal cell thinner than pterostigma width; prestigma incrassate, wider than base of R, and about the length of 1Rs (distinctly separated from pterostigma); pterostigma longer than distance from base of Rs to base of 2r-rs; cell [2R1] short, about 2.4x longer than wide; cells [1Rs] and [2Rs] present but with 2rs-m slightly sclerotized; extremely light nebulous 2m-cu (only visible playing with light incidence angle); protibial spur biseriate apically.

The new genus-group name is a combination of Iberia, referring to the Iberian Peninsula, and Maimetsha, type genus of the family. The name is feminine.

Iberomaimetsha is well distinguished from Maimetsha and Guyotemaimetsha in the simultaneous presence of cells [1Rs] and [2Rs] (rm cells sensu Rasnitsyn and Brothers 2009); it differs from Afromaimetsha by its parallel notauli (instead of diverging anteriorly); the prestigma is incrassate, clearly wider than basalmost R, not as in Afrapia; the origin of Rs in Iberomaimetsha is well separated from pterostigma, not as close as in Maimetshorapia; the pedicel is straight and not “comma-shaped” as in Ahstemiam (see below); Andyrossia was described from just a forewing but Iberomaimetsha differs clearly in several details, such as the length of cell [2R1] (around 2.4 times longer than wide versus 3.6 in Andyrossia), and the width of cell [C] (narrower than pterostigmal width in Iberomaimetsha versus wider in Andyrossia).

Holotype MCNA 9928 (♀); paratype MCNA 8765 (♂). All specimens deposited in the Museo de Ciencias Naturales de Álava, Vitoria-Gasteiz, Spain.

Peñacerrada I, Moraza, Burgos Province, Basque-Cantabrian Basin, Spain.

Escucha Formation, Lower Albian, Lower Cretaceous.

Mandibles moderately large, each with three teeth, with almost straight margins and teeth (when mandibles opened, teeth facing forward, not curved inward); all flagellomeres of similar length (gradual slight reduction of length); protibial distal margin without comb of stiff setae; tarsal plantulae present on female; cell [1M] about as large as cell [2Cu], both almost square or rhomboidal; 2Rs+M much shorter than 1m-cu and shorter than cell [1Rs]; apicalmost sector of Cu arising at midlength of cell [2Cu]; metasoma elongate, about as long as remainder of body; female ovipositor with apex of second valvula 4-toothed.

Female (holotype): Body length 2.8 mm. Head rounded in anterior view, length 0.4 mm, width 0.85 mm; compound eyes not especially bulging, oval, large, well separated by 0.43 mm dorsally; basal part of palps not visible but maxillary palps with at least five palpomeres; labial palps with at least three palpomeres; slight, triangular, supra-antennal elevation above torulus, intertorular area flattened; toruli closer to each other than to inner margin of compound eyes; antennae 2.4 mm in length, inserted on frons below midpoint of compound eyes; scape and pedicel short and globular, flagellomeres cylindrical; lengths of antennomeres as follows (all in mm): scape 0.15, pedicel 0.10, flagellomeres 0.22, 0.20, 0.20, 0.20, 0.18, 0.15, 0.14, 0.14, 0.12, 0.12, 0.12, 0.12, 0.12, 0.15; malar space short, 0.15 mm in length. Mesosoma only laterally visible, 1.15 mm in length; pronotum 0.20 mm in length; mesoscutum 0.35 mm in length; mesoscutellum strongly convex, 0.30 mm in length; propodeum slightly rugose, 0.30 mm in length. Only left mid and hind legs are complete, but foreleg can be reconstructed by mixing right and left parts; no visible division of metatrochantellus; tarsi pentamerous, tarsomeres I–IV each with apicoventral plantar lobe (well visible on left metatarsus: Figs 5, 6D); pretarsal claws with small preapical tooth; leg measurements (all in mm): profemur (with protrochantellus) 0.85, protibia 0.50, protarsomeres I–V 0.30/0.15/0.12/?/?; mesofemur 0.75, mesotibia 0.72, mesotarsomeres 0.32/0.15/0.10/0.07/0.12; metafemur 0.95, metatibia 0.90, metatarsomeres 0.38/0.15/0.10/0.08/0.14. Forewing hyaline, length 2.4 mm, maximum width 0.95 mm, densely covered by microtrichia; all veins tubular except for nebulous crossveins 2rs-m and 2m-cu, thus only cells [2Rs] and [2M] pseudo-opened apically (Fig. 7A); 1rs-m not aligned with r-rs; 2m-cu curved at each distal end; veins M and Cu reaching wing margin, with distal end of Cu strongly bent posteriorly; cell [1R1] longer than both cells [1Rs] + [2Rs]; cell [1Rs] about half the surface of cells [1M] and [2Rs]; vein r-rs emerging at two-thirds from base of pterostigma; cell [2R1] short, with vein Rs slightly anteriorly bent distad 2rs-m. Hind wing hyaline and densely covered by microtrichia, length 1.65 mm, maximum width 0.45 mm; five strong hamuli present on costal vein in distal half of wing (Fig. 7B); distal sections of veins Rs and Cu reaching nebulous wing margin; vein A lost beyond cu-a. Metasoma pedunculate, elongate and globoid, length 1.3 mm, greatest height 0.5 mm; sterna hard, convex; no apparent metasomal armature on sternum II or III; cerci very short; ovipositor moderately exserted, first valvula 0.65 mm long (as preserved), second valvula 0.35 mm long, apically with four dorsal teeth, with sheaths covered by very short, sparse setae (Figs. 4, 5A, 5C).

Male: Very similar to female except in following minor differences: Body length 3.63 mm; tarsi without plantar lobes; metasoma slightly longer than mesosoma.

The specific epithet is a patronym honoring Prof. Alexandr P. Rasnitsyn for his numerous important contributions to the study of Hymenoptera and his generous friendship with the authors.

Holotype MCNA 13049 (?); paratypes MCNA 8758(♀?), MCNA 8790 (♂), MCNA 9373 (?), MCNA 9918 (♀), and MCNA 10732 (?). All specimens deposited in the Museo de Ciencias Naturales de Álava, Vitoria-Gasteiz, Spain.

Peñacerrada I, Moraza, Burgos Province, Basque-Cantabrian Basin, Spain.

Escucha Formation, Lower Cretaceous, Lower Albian.

Mandibles distinctly large, exserted, asymmetrical, right 4-toothed, left 3-toothed, arched inwards (teeth pointing toward teeth of opposite mandible); basal flagellomeres (from F1 to F3–4) distinctly larger than apical flagellomeres (F1–F4 about 2–1.5× the length of F7–F14); protibial distal margin with a comb of stiff setae; cell [1M] square or rhomboidal, distinctly smaller than [2Cu] which has a short but distinct connection between 1M+Cu and M (then, [2Cu] composed by five aristae); 2Rs+M longer than 1m-cu and longer than cell [1Rs]; apicalmost sector of Cu arising from almost apical posterior margin of [2Cu]; metasoma moderately short, as long as mesosoma.

Body length (from paratypes, as incomplete metasoma in holotype) around 2.55–2.61 mm. Head rounded in anterior view, length 0.41 mm, width 0.68 mm with a short occipital carina and apparently some polygonal dorsal sculpture; compound eyes not bulging, slightly oval, separated dorsally by 0.39 mm; toruli about at same distance to inner margin of compound eyes and each other; antennae 2.54 mm in length, inserted on frons below midpoint of compound eyes; scape and pedicel short and globular, flagellomeres cylindrical; lengths of antennomeres as follows (all in mm): scape 0.12, pedicel 0.09, flagellomeres 0.29, 0.28, 0.27, 0.23, 0.20, 0.18, 0.15, 0.14, 0.12, 0.12, 0.11, 0.11, 0.10, 0.12; malar space short, length approximately 0.11 mm. Mesosoma (dorsal view) 0.95 mm in length; pronotum not distinct dorsally; mesoscutal length 0.27 mm, with parallel notauli and longitudinal medial line, all three present as linear series of grooves more than well-delimited lines (Fig. 9); axillae wide and long, well distinct, separating completely mesoscutellum and mesoscutum; mesoscutellum distinctly convex, length 0.22 mm; propodeum 0.29 mm in length, areolate (seen in paratype MCNA 8790). Legs distinctly elongate; division of trochantellus not visible; tarsi with no plantar lobes visible; leg measurements (all in mm): profemur 0.82, protibia 0.46, protarsomeres I–V 0.34/0.15/0.10/0.09/0.10; mid-legs very poorly preserved but apparently with mesofemur 0.68, mesotibia ?, mesotarsomeres 0.24/0.12/0.10/?/?; metafemur 1.0, metatibia 0.83, metatarsomeres 0.46/0.20/0.15/0.09/0.12. Forewing hyaline, length 2.1 mm, maximum width 0.80 mm, densely covered by microtrichia; all veins tubular except for nebulous 2rs-m and 2m-cu, thus, only cells [2Rs] and [2M] pseudo-opened apically (Figs 9, 10); 1rs-m not aligned with r-rs; 2m-cu curved S-like; veins M and Cu reaching wing margin (although M somewhat nebulous), with apicalmost sector of Cu slightly angled posteriorly; cell [1R1] longer than both cells [1Rs]+[2Rs]; cell [1Rs] almost ¾ the surface of cell [1M] and almost half of [2Rs]; vein r-rs emerging at two-thirds from base of pterostigma; cell [2R1] short, with vein Rs slightly anteriorly bent distad 2rs-m. Hind wing difficult to observe, hyaline and densely covered by microtrichia, length approximately 1.22 mm, maximum width approximately 0.41 mm; three strong hamuli present on costal vein in distal half of wing. Metasoma (checked on paratype MCNA 8790) pedunculate, short and slightly flattened, length 0.98 mm, greatest height 0.41 mm; sterna hard, convex; no apparent metasomal armature on sternum II or III.

Male: Parameres spatulate, with apparently three short stiff setae apically (Fig. 10).

The specific epithet is a noun in apposition and is the Anglo-Saxon word for nightmare, in reference to the terrifying appearance of the species in frontal view and with its enlarged mandibles.

Ahiromaimetsha najlae Perrichot, Azar, Nel & Engel, sp. n.

Antennae apparently with 16 articles; pedicel straight; forewing costal cell apically about as wide as pterostigmal width; prestigma not incrassate, not wider than base of R, about as long as 1Rs (distinctly separated from pterostigma); cell [2R1] long, nearly 4x longer than wide; cells [1Rs] and [2Rs] present, former greatly enlarged; 2Rs+M absent owing to confluence of 1m-cu with 2Rs; 1rs-m exceptionally minute (Figs 11, 12D); 2rs-m completely sclerotized; protibial spur bifurcate apically; female tarsi with apicoventral plantar lobes.

The new genus-group name is a combination of Ahirom, Phoenician king of Byblos (ca. 1000 BC) whose sarcophagus bears the oldest inscription in the Phoenician alphabet, and Maimetsha, type genus of the family. The name is considered to be feminine.

Ahiromaimetsha can be distinguished most easily from other genera by the effective absence of 2Rs+M owing to the confluence of 1m-cu with the second free abscissa of Rs. Like Iberomaimetsha rasnitsyni and Andyrossia joyceae, Ahiromaimetsha najlae has a large cell [1Rs] but even more so than in the aforementioned species and, unlike Iberomaimetsha rasnitsyni, the prestigma is not incrassate.

Holotype MKN-1A (♀), collection Azar, provisionally deposited in the Muséum National d’Histoire Naturelle, Paris, France.

Maknouniyyeh, Caza Jezzine, Mouhafazit Loubnan Al-Janoubi (South Lebanon district), Lebanon.

Neocomian (Valanginian–Lowermost Aptian), Lower Cretaceous.

As for the genus (see above).

Female: Total body length ca. 5.2 mm. Head nearly rounded in anterior view, length 0.97 mm, width 1.35 mm, finely setose; compound eyes somewhat bulging, oval, large, length 0.78 mm, width 0.62 mm; ocelli positioned on swelling; mandibles strong, asymmetrical, right with four teeth and left with three teeth both with apical tooth largest; maxillary palps well developed, elongate, distinctly longer than mandibles, with six palpomeres; labial palps short, with four palpomeres; antennae partly damaged but complete, with apparently 16 segments (some segments have secondary constrictions owing to compression), scape and pedicel short and globulous, flagellomeres cylindrical, length of antennomeres as follows (all in mm): scape 0.3, pedicel 0.16, flagellomeres 0.47, 0.52, 0.45, 0.34, 0.32, 0.32, 0.22, 0.17, 0.17, 0.16, 0.16, 0.14, 0.11, 0.11; antenna with fine setae and sparse sensilla. Mesosomal length 2.21 mm, height 1.09 mm, finely setose but without visible foveae or sculpturing; mesoscutal length 0.71 mm, with strong anterior median depression; mesoscutellum not clearly visible, obscured by wings. Metacoxa with an outer basal extension; metatrochantellus diagonally divided and appearing two-segmented; protibial calcar bifurcate (Fig. 12B), with defined velum and trunk, velum smooth, trunk with spine-like microsculpture, basitarsal comb composed of blunt, flattened pegs; pretarsal claws cleft; leg measurements (all in mm, given for those observable): procoxa 0.21, protrochanter 0.42, profemur 0.82, protibia 0.60, probasitarsus 0.61, second protarsomere 0.16, third protarsomere 0.11, mesocoxa 0.39, mesotrochanter 0.55, mesofemur 0.81, metacoxa 0.48, metatrochanter 0.16, metafemur 1.10. Wings hyaline, forewing length 4.81 mm, width 1.54 mm; hind wing length 3.12 mm, width 1.15 mm; venation as in figure 12D; five distal hamuli on hind wing. Metasomal length 2.11 mm; height 0.81 mm; metasomal sterna well sclerotized, convex; ovipositor long, length 2.16 mm, with clearly visible denticles at apex (Fig. 12C).

Male: Unknown.

The specific epithet is a matronym honoring Dr Najla Zeidane-Gèze, wife of the collector of the holotype.

Ahstemiam cellula McKellar & Engel, sp. n.

Total body length near 1.2 mm; compound eyes globular and protuberant; fine occipital carina present; pedicel ‘comma-shaped’ in lateral view, 0.54x length of scape and inserted deeply into scape’s apex; 14 flagellomeres, cylindrical in shape and progressively shortening; maxillary palpus with at least three palpomeres, ultimate palpomere with three apical stiff setae, penultimate palpomere with one apical and two mid-body stiff setae; labial palpus apparently with three palpomeres, all short and bearing numerous stiff setae; junction between Sc+R and Rs very close to pterostigma, 2Sc+R shorter than Rs; pterostigma elongate, longer than [2R1], with gradual apical taper; r-rs originating within apical one-third of pterostigmal length; 2R present as small stub; [1M] relatively small, apical corner (junction of Rs+M and 1m-cu crossvein) positioned posterior to pterostigma; [1Rs] triangular, bounded by very thin M posteriorly, 2Rs and 1rs-m forming anterior and apical margins of cell, both fade to become nebulous throughout most of cell’s length; first apparent metasomal segment much shorter than cell [2R1].

The new genus-group name is the inverse of the type genus, Maimetsha, and is considered a meaningless euphonious combination of letters. The name is designated to be feminine in gender.

Ahstemiam is most similar to Maimetshorapia. The new genus differs in the presence of a prestigma (abscissa of R between 1Rs and pterostigma) that is only slightly inflated (as opposed to distinctly incrassate) and faded apically, as well as a first apparent metasomal tergum that is much shorter than cell [2R1]. The nebulous veins bordering the posterior edges of cell [1Rs], as well as the small size of the cell itself, are distinct among known members of the family Maimetshidae.

Holotype CNC-CAS 1038 (♂) (Fig. 13). Deposited in Canadian National Collection of Insects and Arthropods, Ottawa, Ontario, Canada.

Grassy Lake, Alberta, Canada.

Uppermost Foremost Formation, Campanian, Upper Cretaceous.

As for the genus (see above).

Male (holotype): Body color apparently dark brown, with paler brown antennae and femora, and yellow legs distal to femora (Fig. 13). Head slightly distorted in type specimen, appearing broad (transverse) and foreshortened (exsagittal), with length 0.57× width; ventral margin of gena with prominent notches, more so adjacent to mandibles; frons deeply impressed, antennal bases inserted in shallow depression between compound eyes (Fig. 13); vertex with sparse, short, inclined setae; scape taphonomically distorted, in lateral view, elongate and narrow basally with pronounced apical flare, and concave apex (Fig. 14); pedicel with row of midlength stiff setae inclined upon ventral margin; apical flagellar article slightly wider than preceding articles, and tapering to bluntly-rounded apex; flagellum with dense coat of short, curled setae in various orientations. Mesoscutum with low dorsal convexity, apparently near one-half of mesosomal length; mesoscutellum with low dorsal convexity, overhanging metanotum slightly, bearing numerous short, inclined setae; metanotum with prominent ridge along posterior margin which projects posterodorsally over propodeum; propodeum gradually sloping to pronounced posterior lip. Forewing at least 0.96 mm in length and 0.42 mm in width, hyaline with microtrichial vestiture and pterostigma pale brown, like tubular veins in wing; Sc+R slightly thicker than other veins, narrowing basally but expanding apical to junction with Rs; fusion of Rs and M angled, veins not parallel to each other; 2rs-m apparently nebulous throughout its course, but preserved in only one wing, which appears taphonomically affected in this region; apex of M not preserved in type specimen; 2Cu significantly longer than 1m-cu; 3Cu appears relatively short, terminating posterior to 1rs-m; 2A fading to nebulous vein at midlength of cell [2Cu]. Hind wings poorly visible; apparent length near 0.64 mm; only anterior veins partly visible; broad, open costal cell apparent near apex of Sc+R; R appears to curve anteriorly and rapidly fade along anterior margin of wing; 2Rs and putative 1rs-m both fade distally, and are removed from 1R by relatively long 1Rs; faint curved vein within posterior portion of wing base assumed to represent A; marginal setae visible along posterior margin of wing are fine, elongate and relatively sparse, with greater density in apical positions. Legs with pervasive coat of short, stiff setae densely inclined upon most surfaces; pro- and mesocoxae with elongate setae inclined upon ventral surface; profemur with two rows of short, stiff setae suberect upon ventral surface, protibia bearing row of at least three stout spines approximately one-half of calcar’s length on posterior surface; calcar gently curved and robust (as wide as probasitarsus), deeply inserted into protibial apex, extending only slightly beyond apex; fine setae inclined upon both sides of calcar, and matching longitudinal comb of erect spicules on basal half of probasitarsus; probasitarsus length slightly less than all subsequent tarsomeres combined, with approximately four irregularly distributed spines upon apical half of plantar surface; protarsomeres III and IV much shorter than others, one-half to two-thirds length of protarsomere V; all tarsi with serrate pretarsal claws and large arolium, with two apical spines on all tarsomeres (potentially absent on protarsomeres, but orientation obscures observation); meso- and metacoxae both approximately 1.5× length of procoxa, with broad bases and apical constriction; mesofemur with two rows of short, stiff setae inclined upon ventral surface; mesotibia with patch of fine, short spines along posterior surface near apex; two mesotibial spurs short and stout, slightly longer than apical width of mesotibia, and bearing minute setae along both margins; metafemur with significant midlength inflation, laterally compressed, with two rows of short, stiff setae suberect along ventral surface; metatibia with moderate apical expansion, bearing two apical spurs slightly longer than metatibial apical width, as well as comb-like row of six spicules each approximately two-thirds of spurs’ length; metatarsi not preserved. Metasoma with six apparent terga; terga appear to bear frills of elongate setae along posterior and lateral margins, terga more pilose within posterior of metasoma; parameres spatulate, each with four elongate setae erect on apex.

Female: Unknown.

The specific epithet is the Latin diminutive noun in apposition, cellula (meaning, “small room”) and refers to the very small size of cell [1Rs].