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ZooKeys 182: 1–70, doi: 10.3897/zookeys.182.2720
Annotated type catalogue of the Bothriembryontidae and Odontostomidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London
Abraham S.H. Breure 1, Jonathan D. Ablett 2
1 Netherlands Centre for Biodiversity Naturalis, P.O. Box 9517, Leiden, the Netherlands
2 Natural History Museum, Division of Higher Invertebrates, London, SW7 5BD, UK

Corresponding author: Abraham S.H. Breure (ashbreure@gmail.com)

Academic editor: E. Neubert

received 20 January 2011 | accepted 26 March 2012 | Published 10 April 2012


(C) 2012 Abraham S.H. Breure. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.


For reference, use of the paginated PDF or printed version of this article is recommended.

Abstract

The type status is described for specimens of 84 taxa classified within the families Bothriembryontidae and Odontostomidae (superfamily Orthalicoidea) and kept in the Natural History Museum, London. Lectotypes are designated for Bulimus (Liparus) brazieri Angas, 1871; Bulimus broderipii Sowerby I, 1832; Bulimus fuligineus Pfeiffer, 1853; Helix guarani d’Orbigny, 1835; Bulimus (Tomigerus) ramagei E.A. Smith, 1890; Helix rhodinostoma d’Orbigny, 1835; Bulimus (Bulimulus) ridleyi E.A. Smith, 1890. The type status of the following taxa is changed to lectotype in accordance with Art. 74.6 ICZN: Placostylus (Euplacostylus) cylindricus Fulton, 1907; Bulimus pyrostomus Pfeiffer, 1860; Bulimus turneri Pfeiffer, 1860. The following taxon is synonymised: Bulimus oblitus Reeve, 1848 = Bahiensis neglectus (Pfeiffer, 1847).

Keywords

Bothriembryontidae, Odontostomidae, types

Introduction and methods

This is the second paper on the types of Orthalicoidea in the Natural History Museum, London. An earlier paper (Breure and Ablett 2011) has presented the context of the collection, the criteria used for the selection of lectotypes, some biohistorical notes, and a list of type specimens belonging to the Amphibulimidae. The aim of this publication is to supplement the earlier paper with data on the type specimens classified with the Bothriembryontidae and Odontostomidae (sensu Breure and Romero in press).

Abbreviations used for depositories of material are: ANSP, Academy of Natural Sciences, Philadelphia, U.S.A.; MHNG, Muséum d’Histoire Naturelle, Geneve, Switserland; MNHN, Muséum National d’Histoire Naturelle, Paris, France; MNHNM, Museo Nacional de Historia Natural, Montevideo, Uruguay; NHMUK, Natural History Museum, London, U.K.; RBINS, Royal Belgian Institute of Natural Sciences, Brussels, Belgium; RMNH, Netherlands Centre for Biodiversity Naturalis, Leiden, the Netherlands; SMF, Senckenberg Natur-Museum, Frankfurt am Main, Germany; UF, Florida Museum of Natural History, Gainesville, U.S.A.; ZMB, Museum für Naturkunde, Berlin, Germany. Other abbreviations used are: / end of line in cited text; coll., collection; D, diameter; H, shell height; leg., legit, collected; W, number of whorls. See Fig. 1 for the way measurements of the shell have been taken. Label styles in the Cuming collection (M.C.) are explained in Breure and Ablett (2011: 7–8, fig. 6).

Systematics Systematic list of taxa arranged in generic order

This systematic list follows Breure (1979); the family classification amended as proposed by Breure and Romero (in press). The generic classification has been adapted from Simone (2006) for the Odontostomidae (but see Remarks below); the classification of Bothriembryontidae is based on unpublished data from the senior author. Within each family, genus and at species level, taxa are presented in alphabetical order.

Family Bothriembryontidae Iredale, 1937

Remarks. The Gondwanan group within the Orthalicoidea (Placostylidae sensu Neubert et al. 2009, Bothriembryon, Prestonella, Plectostylus, and Discoleus) appears to be monophyletic (Breure and Romero in press). Thus the oldest family name, i.e. Bothriembryontidae, is used herein for this group.


Aspastus Albers, 1850

Aspastus miltocheilus (Reeve, 1848); Aspastus sellersi (Cox, 1871).

Bothriembryon Pilsbry, 1894

Bothriembryon angasianus (Pfeiffer, 1864); Bothriembryon brazieri(Angas, 1871); Bothriembryon dux (Pfeiffer, 1861); Bothriembryon kingii (J.E. Gray, 1825); Bothriembryon leeuwinensis (E.A. Smith, 1894); Bothriembryon physoides (Reeve, 1849); Bothriembryon rhodostomus (J.E. Gray, 1834); Bothriembryon spenceri (Tate, 1894); Bothriembryon tasmanicus (Pfeiffer, 1853).

Callistocharis Pilsbry, 1900

Callistocharis crassilabrum (Garrett, 1872); Callistocharis ochrostoma (Garrett, 1872); Callistocharis subroseus Fulton, 1915.

Eumecostylus Martens in Albers, 1860

Eumecostylus backhuysi Delsaerdt, 2010; Eumecostylus calus (E.A. Smith, 1891); Eumecostylus cylindricus Fulton, 1907; Eumecostylus foxi Clench, 1941; Eumecostylus hargravesi (Cox, 1871); Eumecostylus sanchristovalensis (Cox, 1870).

Euplacostylus Crosse, 1875

Euplacostylus koroensis(Garrett, 1872).

Placocharis Pilsbry, 1900

Placocharis guppyi E.A. Smith, 1891; Placocharis kreftii (Cox, 1872); Placocharis macgillivrayi (Pfeiffer, 1855); Placocharis strangei (Pfeiffer, 1853).

Placostylus Beck, 1837

Placostylus alba (Crosse, 1874); Placostylus asopeus (Gassies, 1871); Placostylus bairdii (Reeve, 1848); Placostylus bulbulus (Gassies, 1871); Placostylus corpulentus (Gassies, 1871); Placostylus cuninculinsulae (Cox, 1872); Placostylus eddystonensis (Pfeiffer, 1855); Placostylus edwardsianus (Gassies, 1863); Placostylus falcicula (Gassies, 1871); Placostylus gatopensis (Crosse, 1870); Placostylus grammica (Crosse, 1870); Placostylus infundibulum (Gassies, 1871); Placostylus necouensis (Gassies, 1871); Placostylus ouensis (Gassies, 1870); Placostylus pinicola (Gassies, 1870); Placostylus priscus Powell, 1938; Placostylus pyrostomus (Pfeiffer, 1860); Placostylus salomonis (Pfeiffer, 1853); Placostylus savesi (Crosse, 1886); Placostylus singularis (Morelet, 1857); Placostylus sinistrorsa (Crosse, 1884); Placostylus souvillei (Morelet, 1857); Placostylus superfasciatus (Gassies, 1871).

Plectostylus Beck, 1837

Plectostylus broderipii (Sowerby I, 1832); Plectostylus coturnix (Sowerby I, 1832); Plectostylus elegans (Pfeiffer, 1842); Plectostylus elongata (d’Orbigny, 1837); Plectostylus punctulifer (Sowerby I, 1833); Plectostylus reflexus (Pfeiffer, 1842); Plectostylus variegata (Pfeiffer, 1842).

Poecilocharis Kobelt, 1891

Poecilocharis turneri (Pfeiffer, 1860).

Prestonella Connolly, 1929

Prestonella bowkeri (Sowerby III, 1890); Prestonella nuptialis (Melvill and Ponsonby, 1894); Prestonella quadingensis Connolly, 1929.

Santacharis Iredale, 1927

Santacharis fuligineus (Pfeiffer, 1853).

Family Odontostomidae Pilsbry and Vanatta, 1894

Remarks. We follow here the classification of Simone (2006), with exceptions mentioned below. However, it must be emphasized that a revision of this family, combining morphological and molecular data, is much overdue.


Anostoma Fischer von Waldheim, 1807

Anostoma brunneum Verdcourt, 1990; Anostoma carinatum Pfeiffer, 1853.

Bahiensis Jousseaume, 1877

Bahiensis fuscagula d’Orbigny, 1837; Bahiensis guarani (d’Orbigny, 1835); Bahiensis longulus (Pfeiffer, 1859); Bahiensis neglecta (Pfeiffer, 1847); Bahiensis oblita (Reeve, 1848); Bahiensis occultus (Reeve, 1849); Bahiensis paralellus (Pfeiffer, 1857); Bahiensis rhodinostoma (d’Orbigny, 1835).

Biotocus Salgado and Leme, 1990

Biotocus cumingii (Pfeiffer, 1849); Biotocus turbinatus (Pfeiffer, 1845).

Bonnanius Jousseaume, 1900

Bonnanius ramagei (E.A. Smith, 1890).

Clessinia Döring, 1874

Clessinia costatus (Pfeiffer, 1849).


Remarks. Molecular evidence appear to suggest that taxa hitherto placed in Spixia and Clessinia belong to the same group (Breure and Romero in press); in case further studies should confirm the synonymy of these taxa, Clessinia has priority over Spixia Pilsbry & Vanatta, 1894.


Cyclodontina Beck, 1837

Cyclodontina catharinae (Pfeiffer, 1857); Cyclodontina charpentieri (Pfeiffer, 1850); Cyclodontina corderoi Klappenbach, 1958; Cyclodontina minor (d’Orbigny, 1837).

Hyperaulax Pilsbry, 1897

Hyperaulax ridleyi (E.A. Smith, 1890).

Moricandia Pilsbry & Vanatta, 1898

Moricandia tolerata (Fulton, 1903).

Odontostomus Beck, 1837

Odontostomus grayanus Pfeiffer, 1845; Odontostomus odontostomus (Sowerby I, 1824).


Remarks. Simone (2006) seems to have misinterpreted this genus, by considering Plagiodontes daedaleus (Deshayes, 1851) as a member of Odontostomus, whilst placing the type species of this genus in Macrodontes Swainson, 1840 without further discussion.


Plagiodontes Döring, 1876

Plagiodontes dentata (Wood, 1828); Plagiodontes patagonica (d’Orbigny, 1835).

Spixia Pilsbry & Vanatta, 1894

Spixia major (d’Orbigny, 1837).

Alphabetic list of taxa by species name
Bulimus ouveanus alba Crosse, 1874

http://species-id.net/wiki/Bulimus_ouveanus_alba

Figs 8G–H
Bulimus ouveanus var. γ alba Crosse 1874: 109; Breure and Schouten 1985: 66.
Placostylus fibratus ouveanus (Dotzauser in Mousson); Neubert et al. 2009: 74, fig. 96
Type locality.

“in loco Siandé dicto, Novae Caledoniae (E. Marie)”.

Label.

“Siandé”; label in Crosse’s handwriting glued to the shell.

Dimensions.

“Long. 48, diam. maj. 23 1/2 mill.”; holotype H 48.1, D 23.8, W 6.1.

Type material.

NHMUK 1929.9.14.6, holotype (Marie coll.).

Remarks.

The shell is accompanied by another label in Crosse’s handwriting, stating that on 30 July 1884 he received a message from E. Marie that this shell had to be considered a monstrosity. Neubert et al. (2009) ascribe this taxon to Pfeiffer, but it is unclear if Pfeiffer ever saw this specimen.

Current systematic position.

Bothriembryontidae, Placostylus fibratus ouveanus (Dotzauer in Mousson, 1869).

Bulimus angasianus Pfeiffer, 1864

http://species-id.net/wiki/Bulimus_angasianus

Figs 2A–B, iii
Bulimus angasianus Pfeiffer 1864: 528; B.J. Smith 1992: 101.
Bothriembryon (Bothriembryon) angasianus (Pfeiffer); Breure 1979: 93.
Type locality.

“Port Lincoln, South Australia”.

Label.

“Port Lincoln / South Australia”; presumably in Angas’ handwriting.

Dimensions.

“Long. 25 1/2, diam. 14 mill.”; syntype H 20.6 D 14.6, W 4.5.

Type material.

NHMUK 1870.10.26.176, one syntype (Angas coll.).

Remarks.

The label bears “type” in the original handwriting, but the specimen does not match the dimensions given by Pfeiffer. There are three additional syntypes in the Angas collection in the Hancock Museun [now part of the Great North Museum in Newcastle-upon-Tyne] (D. Gordon, pers. comm.)

Current systematic position.

Bothriembryontidae, Bothriembryon angasianus (Pfeiffer, 1864).

Bulimus asopeus Gassies, 1871

http://species-id.net/wiki/Bulimus_asopeus

Figs 7E, 7v
Bulimus asopeus Gassies 1871: 87; Breure and Schouten 1985: 66; Neubert et al. 2009: 51, 54, fig. 18.
Type locality.

[New Caledonia] “Ile des Pins, sud de la Nouvelle-Calédonie”.

Label.

No information given. The specimen is marked by Gassies with “T” on the ventral side of the last whorl.

Dimensions.

“Long. 50 mill.; diam. maj. 34”; holotype H 49.7, D 35.1, W 6.5.

Type material.

NHMUK 1883.11.10.1173, holotype, R.P. Lambert leg. (ex Gassies).

Remarks.

The specimen is a monstrosity and was classified as Placostylus fibratus fibratus (Martyn, 1784) in a recent revision by Neubert et al. (2009).

Current systematic position.

Bothriembryontidae, Placostylus fibratus fibratus (Martyn, 1784).

Eumecostylus vicinus backhuysi Delsaerdt, 2010

http://species-id.net/wiki/Eumecostylus_vicinus_backhuysi

Eumecostylus vicinus backhuysi Delsaerdt 2010: 43, pl. 6 figs 10–13.
Type locality.

“Marau area, SE Guadalcanal, Solomon Islands”.

Label.

“Marau area, SE Guadalcanal, Solomon Islands”.

Dimensions.

“68.4 x 29.3 mm”; holotype H 68.4, D 29.3, W 4+.

Type material.

NHM 20100568, holotype, R. Moylan leg. (ex Delsaerdt ex Backhuys).

Remarks.

The apex of the specimen is missing.

Current systematic position.

Bothriembryontidae, Eumecostylus vicinus backhuysi Delsaerdt, 2010.

Bulimus bairdii Reeve, 1848

http://species-id.net/wiki/Bulimus_bairdii

Figs 7A, 7i
Bulimus bairdii Reeve 1848 [1848–1850]: pl. 43 fig. 272; Breure and Schouten 1985: 67 (lectotype designation); Neubert et al. 2009: 51, 53, fig. 8.
Type locality.

“—?”.

Label.

No locality given. “Type specimen” added in a later handwriting (E.A. Smith?)

Dimensions.

Not given. Specimen figured herein H 78.1, D 36.2, W 7.3.

Type material.

NHMUK 1975223, lectotype.

Remarks.

Reeve indicated that his specimen was in “Mus. Brit.”. His description leaves room for the possibility that Reeve passed additional specimens to other collectors/institutions; therefore, we concur with Breure and Schouten (1985) to designate this specimen as lectotype. Neubert et al. (2009), who consider this specimen a syntype, treat this taxon as a synonym of Placostylus fibratus fibratus (Martyn, 1784).

Current systematic position.

Bothriembryontidae, Placostylus fibratus fibratus (Martyn, 1784).

Bulimus (Mesembrinus?) bowkeri Sowerby III, 1890

http://species-id.net/wiki/Bulimus_bowkeri

Figs 6A–C, 6i
Bulimus (Mesembrinus?) bowkeri Sowerby III 1890: 581, pl. 56 fig. 5.
Prestonella bowkeri (Sowerby); Herbert 2007: 6, figs 2–7, 9–11, 19, 25 (lectotype designation).
Type locality.

[South Africa] “Somerset (East), Cape Colony. Forest about 3, 000 ft above sea level (Col. Bowker)”.

Label.

“Somerset (East), Cape / Colony. Forest about / 3, 000 ft above sea level / Col. Bowker”.

Dimensions.

“Long. 20, maj. diam. 11 mill.”; lectotype H 19.6, D 10.3, W 4.7.

Type material.

NHM 1889.11.4.9, lectotype.

Current systematic position.

Bothriembryontidae, Prestonella bowkeri (Sowerby III, 1890).

Bulimus (Liparus) brazieri Angas, 1871

http://species-id.net/wiki/Bulimus_brazieri

Figs 2C–E, 2ii
Bulimus (Liparus) brazieri Angas 1871: 19, pl. 1 fig. 28; B.J. Smith 1992: 102.
Bothriembryon (Bothriembryon) brazieri (Angas); Breure 1979: 93.
Type locality.

[Western Australia] “Sinclair’s Range, King George’s Sound”; see Remarks.

Label.

“ Stirling Range, King George’s Sound”.

Dimensions.

“Length 8 lines, breadth 4 lines”; lectotype H 14.3, D 8.7, W 4.9.

Type material.

NHMUK 1870.11.5.8, lectotype, and one paralectotype (Angas coll.).

Remarks.

According to Iredale (1939) the published locality was in error and should read “Stirling Range”. The taxon forms part of an ill-defined species complex. The material corresponds to the original figures of Angas. The shell pictured in the right-hand view is here selected lectotype (design. n.). The specimens are not full-grown.

Current systematic position.

Bothriembryontidae, Bothriembryon brazieri (Angas, 1871).

Bulinus broderipii Sowerby I, 1832

http://species-id.net/wiki/Bulinus_broderipii

Figs 4A–B, 4i
Bulinus broderipii Sowerby I in Broderip and Sowerby 1832a: 30; Sowerby 1833 [1833–1838]: figs 1–1*.
Type locality.

“prope Copiapo Chilensium”.

Label.

“Chili” (in a later handwriting on board). M.C. label style I.

Dimensions.

“long. 1 6/8, lat. 1 1/8 poll. [44.3, 28.5 mm]”; lectotype H 44.3, D 26.1, W 5.9.

Type material.

NHMUK 20100655, lectotype, and three paralectotypes (Cuming coll.).

Remarks.

This taxon may be confused with several other Plectostylus species; a revision of this group is pending (F. Cádiz, pers. commun.). One of the specimens matches the original shell height and corresponds to the figures in Sowerby 1833–1838; this shell is here selected as lectotype (design. n.).

Current systematic position.

Bothriembryontidae, Plectostylus broderipii (Sowerby I, 1832).

Anostoma octodentatum brunneum Verdcourt, 1991

http://species-id.net/wiki/Anostoma_octodentatum_brunneum

Figs 16D–F
Anostoma octodentatum brunneum Verdcourt 1991: 81, fig. 1.
Anostoma brunneum Verdcourt; Simone 2006: 174, fig. 617.
Type locality.

“Brazil, Goias State, about 20 km SE. Dianapolis, near Jardim”.

Label.

“Brazil, Goiás, near Jardim, c. 20 km SE Dianapolis”.

Dimensions.

“Alt. 16 mm, greater diam. 33–34 mm”; holotype H 16.3, D 32.6, W 4.3.

Type material.

NHMUK 1990125, holotype and NHMUK 1990126, (juvenile) paratype (R.M. Hartley leg., 12.ii.1987); in ethanol.

Remarks.

A paratype in the MZSP collection was figured by Simone (2006).

Current systematic position.

Odontostomidae, Anostoma octodentatum brunneum Verdcourt, 1991.

Bulimus bulbulus Gassies, 1871

http://species-id.net/wiki/Bulimus_bulbulus

Figs 7F, 7vi
Bulimus bulbulus Gassies 1871: 193.
Type locality.

[New Caledonia] “les environs de Nouméa, de Concepcion, Baie du Sud, etc., fossile à l’île des Pins”.

Label.

No type locality. Taxon label in Gassies’ handwriting.

Dimensions.

“Diam. maj. 4 mill., alt. 5 mill.”; specimen figured herein H 4.13, D 3.56, W 2.1.

Type material.

NHMUK 1883.11.10.223-229, seven syntypes (Gassies coll.).

Remarks. Three specimens are still glued on board, four others separate in capsule. These specimens are juveniles, probably of Placostylus fibratus (Martyn, 1784).

Current systematic position.

Bothriembryontidae, Placostylus cf. fibratus (Martyn, 1784).

Placostylus calus E.A. Smith, 1891

http://species-id.net/wiki/Placostylus_calus

Figs 11A, 11i
Placostylus calus E.A. Smith 1891: 489, pl. 40 fig. 7.
Placostylus (Placocharis) calus (E.A. Smith) [sic, no parenthesis required]; Breure and Schouten 1985: 63 (lectotype designation).
Eumecostylus calus (E.A. Smith); Delsaerdt 2010: 44, pl. 7 figs 1–4.
Type locality.

“Solomon Islands”.

Label.

“Solomon Is.”, in Smith’s handwriting.

Dimensions.

“Longit. 78 millim., diam. maj. 33”; lectotype H 78, D 33.7, W 5+.

Type material.

NHMUK 1891.11.3.4, lectotype; NHMUK1891.11.3.5-6, two paralectotypes (ex Brazier).

Remarks.

The apex of the lectotype is damaged.

Current systematic position.

Bothriembryontidae, Eumecostylus calus (E.A. Smith, 1891).

Anostoma carinatum Pfeiffer, 1853

http://species-id.net/wiki/Anostoma_carinatum

Figs 16A–C, 16i
Anostoma carinatum Pfeiffer 1853a: 57.
Ringicella carinata (Pfeiffer); Simone 2006: 175, fig. 622.
Type locality.

“Brasilia (Mus. Dennison.)”.

Label.

“Brazils”.

Dimensions.

“Diam. maj. 24 1/2, (..) alt. 13 mill.”; specimen figured herein H 13.3, D 24.3, W 4.9.

Type material.

NHMUK 20100509, three syntypes (Cuming coll.).

Remarks. Although the taxon label is not in Pfeiffer’s handwriting, the specimens fit the original measurements and are considered probable syntypes.

Current systematic position.

Odontostomidae, Anostoma carinatum Pfeiffer, 1853.

Bulimus catharinae Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_catharinae

Figs 23A–B, 23i
Bulimus catharinae Pfeiffer 1857: 389; Breure 1974: 113.
Cyclodontina (Cyclodontina) fusiformis (Menke); Breure and Schouten 1985: 6, pl. 1 figs 1–2 (lectotype designation).
Cyclodontina catharinae (Pfeiffer); Simone 2006: 165, fig. 567.
Type locality.

“St. Catherine’s, Brazil”.

Label.

“St Catherines Brazil”; taxon label in Pfeiffer’s handwriting.

Dimensions.

“23–25, 7 1/2–8 mill.”; specimen figured herein H 24.4, D 7.5, W 8.1.

Type material.

NHMUK 1975261, lectotype and two paralectotypes (Cuming coll.).

Remarks.

Simone revalidated this taxon without providing further discussion; however, we follow here the synonymization of Breure and Schouten (1985).

Current systematic position.

Odontostomidae, Cyclodontina fusiformis (Menke, 1828).

Bulimus charpentieri Pfeiffer, 1850

http://species-id.net/wiki/Bulimus_charpentieri

Figs 21G–I, 21ii
Bulimus charpentieri “Grateloup” Pfeiffer 1850: 14; Pfeiffer in Küster and Pfeiffer 1854 [1840–1865]: 143, pl. 45 figs 14–15; Breure 1974: 113.
Type locality.

“Cardova [sic, Córdoba] reipubl. Argentinae”.

Label.

“Brazils”. M.C. label style III.

Dimensions.

“Long. 20, diam. 5 2/3 mill.”; specimen figured herein H 17.9, D 5.15, W 9.2.

Type material.

NHMUK 20110386, syntype (Cuming coll.).

Current systematic position.

Odontostomidae, Cyclodontina charpentieri (Pfeiffer, 1850).

Cyclodontina (Spixia) corderoi Klappenbach, 1958

http://species-id.net/wiki/Cyclodontina_corderoi

Figs 21J–K, 21iii
Cyclodontina (Spixia) corderoi Klappenbach 1958: 2, pls 1–2; Breure 1974: 114.
Type locality.

[Uruguay] “Pozo Hondo, próxima a la población Tambores en el departemento de Tacuarembó”.

Label.

“Pozo Hondo, Tambores, Depto Tacuarembó, Rep. del Uruguay”.

Dimensions.

“Largo total 17.5, Ancho 5 (en millimetros)”; specimen figured herein H 16.1, D 5.44, W 7+.

Type material.

NHMUK 1961679, three paratypes, M.A. Klappenbach & P. San Martin leg., 12.x.1957, ex MNHNM 0716.

Remarks.

The apex of the shell is slightly damaged.

Current systematic position.

Odontostomidae, Cyclodontina corderoi Klappenbach, 1958.

Bulimus corpulentus Gassies, 1871

http://species-id.net/wiki/Bulimus_corpulentus

Figs 10A, 10i
Bulimus corpulentus Gassies 1871: 183; Breure and Schouten 1985: 67.
Placostylus corpulentus (Gassies); Neubert et al. 2009: 112, fig. 184.
Type locality.

“Fossile à l’île des Pins et à l’îlot Koutoumo”.

Label.

“New Caledonia”.

Dimensions.

“Long. 100 mill.; diam. maj. 55 mill.”; specimen figured herein H 94.0, D 64.0, W 6.7.

Type material.

NHMUK 1883.11.10.1178, syntype, R.P. Lambert leg. (ex Gassies).

Remarks.

The surface of the specimen is covered by various encrustations.

Current systematic position.

Bothriembryontidae, Placostylus corpulentus (Gassies, 1871).

Bulimus costatus Pfeiffer, 1848

http://species-id.net/wiki/Bulimus_costatus

Figs 22F–G, 22ii
Bulimus costatus Pfeiffer 1848: 114; Pfeiffer 1849a: 111; Reeve 1849 [1848–1850]: pl. 65 fig. 450.
Clessinia costata (Pfeiffer); Breure and Schouten 1985: 10, pl. 1 fig. 4 (lectotype designation); Simone 2006: 164, fig. 563.
Type locality.

“Brasilia” [Brazil].

Label.

“Brazil”. M.C. label style III. Another label indicates that this specimen was also figured in Reeve’s Conchologica Iconica.

Dimensions.

“Long. 18, diam. 5 1/2 mill.”; lectotype H 17.8, D 5.6, W 8.8.

Type material.

NHMUK 1975250, lectotype and two paralectotypes (Cuming coll.).

Remarks.

Although the first publication was in Pfeiffer (1848), he refers to Pfeiffer (1849) as the original description; this paper was describing new species from the Cuming collection.

Current systematic position.

Odontostomidae, Clessinia costata (Pfeiffer, 1848).

Bulinus coturnix Sowerby I, 1832

http://species-id.net/wiki/Bulinus_coturnix

Figs 4C–D, 4ii
Bulinus coturnix Sowerby I in Broderip and Sowerby 1832a: 30.
Type locality.

[Chile] “Huasco”.

Label.

“Huasco”. M.C. label style I.

Dimensions.

“Long 1 1/4, lat. 7/8 poll. [H 31.7, D 22.2 mm]”; specimen figured herein H 32.4, D 21.0, W 5.0.

Type material.

NHMUK 20100620, five possible syntypes (Cuming coll.).

Remarks.

Of the five specimens, only one is adult; although it does not correspond to the original measurements, it originates from the type locality. The material is considered as possible syntypes.

Current systematic position.

Bothriembryontidae, Plectostylus coturnix (Sowerby I, 1832).

Bulimus crassilabrum Garrett, 1872

http://species-id.net/wiki/Bulimus_crassilabrum

Figs 13A, 13i
Bulimus crassilabrum Garrett 1872: 233, pl. 18 fig. 5.
Type locality.

[Fiji] “Vanna Levu Id., Viti Isles”.

Label.

“Vanua Levu”.

Dimensions.

“Length 41 mill., diam. 21 mill.”; specimen figured herein H 44.0, D 22.1, W 5.1.

Type material.

NHMUK 20030801, one syntype, ex Kennard ex Garrett coll. (“cotype ex auct.”).

Remarks.

Richardson (1995: 286) placed this taxon in the synonymy of Aspastus rugatus (Garrett, 1872), without further comments. Judging by the descriptions and figures of both species, there is every reason to consider these two taxa identical. Based on Garrett’s 1872 paper, Bulimus crassilabrum has page priority.

Current systematic position.

Bothriembryontidae, Callistocharis crassilabrum (Garrett, 1872).

Tomigerus cumingii ‘Newcomb’ Pfeiffer, 1849

http://species-id.net/wiki/Tomigerus_cumingii

Figs 19A–C, 19i
Tomigerus cumingii ‘Newcomb’ Pfeiffer 1849b: 67.
Biotocus cumingii (Pfeiffer); Simone 2006: 177, fig. 633.
Type locality.

“Para[, ] Brasiliae”.

Label.

“Para”. M.C. label style III.

Dimensions.

“Diam. maj. 7 1/2, min. 5, alt. 6 mill.”; specimen figured herein H 6.45, D 6.72, W 4.8.

Type material.

NHMUK 1975587, three syntypes, ex Newcomb (Cuming coll.).

Current systematic position.

Odontostomidae, Biotocus cumingii (‘Newcomb’ Pfeiffer, 1849).

Bulimus cuninculinsulae Cox, 1872

http://species-id.net/wiki/Bulimus_cuninculinsulae

Figs 12C, 12iii
Bulimus cuninculinsulae Cox 1872b: 19, 233, pl. 18 fig. 5; Breure and Schouten 1985: 68 (lectotype designation).
Type locality.

[Australia] “Rabbit Island, near Lord Howe’s Island, Pacific Ocean”.

Label.

“Rabbit Is. / near Lord / Howe’s Is. / Pacific ocean”.

Dimensions.

“Length 1.65, breadth 0.75 of an inch [H 41.9, D 19.1 mm]”; lectotype H 41.6, D 20.5, W 5.6.

Type material.

NHMUK 1880.12.11.47-48, lectotype and one paralectotype (ex Cox).

Remarks.

The specimen figured by Cox (1872) has been chosen by Breure and Schouten (1985) as the lectotype; the paralectotype matches the original measurements (H 41.9).

Current systematic position.

Bothriembryontidae, Placostylus cuninculinsulae (Cox, 1872).

Placostylus (Euplacostylus) cylindricus Fulton, 1907

http://species-id.net/wiki/Placostylus_cylindricus

Figs 11C, 11iii
Placostylus (Euplacostylus) cylindricus Fulton 1907: 154, pl. 10 fig. 3.
Placostylus (?Proaspastus) cylindricus (Fulton); Breure and Schouten 1985: 64.
Eumecostylus cylindricus (Fulton); Delsaerdt 2010: 39, pl. 5 figs 1–5.
Type locality.

“Isabel Island, Solomons” (see remarks).

Label.

“Isabel Id. / Solomon Ids.”, in Fulton’s handwriting.

Dimensions.

“Maj. diam. 23, alt. 71 mm.”; lectotype H 71.3, D 27.4, W 6.7.

Type material.

NHMUK 1907.5.3.160, lectotype (ex Fulton).

Remarks.

Fulton (1907) mentioned that he had three specimens, of which the NHMUK specimen corresponds to his figure. Contrary to Breure and Schouten (1985), it should not be considered as holotype, but is here designated as the lectotype (Art. 74.6 ICZN). The specimen has the apex damaged. Clench (1941) questioned the type locality and suggested it was erroneous. Also Delsaerdt (2010) has been unable to locate this species on Santa Isabel; it is only known from Guadalcanal.

Current systematic position.

Bothriembryontidae, Eumecostylus cylindricus (Fulton, 1907).

Pupa dentata Wood, 1828

http://species-id.net/wiki/Pupa_dentata

Figs 25A–D, 25i
Pupa dentata Wood 1828: 24, 50, pl. 8 fig. 71.
Type locality.

Not given.

Label.

“Montevideo”.

Dimensions.

Not given. Specimen figured herein H 14.16, D 7.17, W 7.9.

Type material.

NHMUK 1840.9.12.50, two syntypes, ex Mrs. Mawe.

Remarks.

Wood (1828: 24) referred to material from “Br.M. [British Museum]”. This supports the claim expressed on the label that this material may be considered as probable syntypes. It is not clear, however, on what evidence the locality “Montevideo” is based, as Wood did not state it in his publication. One specimen is damaged.

Current systematic position.

Odontostomidae, Plagiodontes dentata (Wood, 1828).

Bulimus dux Pfeiffer, 1861

http://species-id.net/wiki/Bulimus_dux

Figs 3A–B, 3i
Bulimus dux Pfeiffer 1861: 24; B.J. Smith 1992: 103.
Bothriembryon (Bothriembryon) dux (Pfeiffer); Breure 1978: 203 (lectotype designation); Breure 1979: 94.
Type locality.

“King George’s Sound, [Western] Australia”.

Label.

“King Georges Sound / W. Australia”; taxon label damaged, possibly in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

“Long. 51, diam. 26 mill.”; lectotype H 48.5, D 28, W 6.0.

Type material. NHMUK 19598, lectotype; 19599, two paralectotypes (Cuming coll.).

Remarks.

These specimens were identified by E.A. Smith as type material.

Current systematic position.

Bothriembryontidae, Bothriembryon dux (Pfeiffer, 1861).

Bulimus eddystonensis Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_eddystonensis

Figs 9D, 9iv
Bulimus eddystonensis Pfeiffer 1855a: 8.
Placostylus (Placostylus) eddystonensis (Pfeiffer); Breure and Schouten 1985: 59 (lectotype designation); Neubert et al. 2009: 94, fig. 149.
Type locality.

“Eddystone Island, Australian Seas” [Solomon Islands; see remarks].

Label.

“Eddystone Isld / Australian sea”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

“Long. 74, diam. 34 mill.”; lectotype H 73.3, D 36.3, W 6.2.

Type material.

NHMUK 1975230, lectotype and two paralectotypes (Cuming coll.).

Remarks. The type locality was in error and was subsequently corrected to New Caledonia, Hienghène (Gassies 1863: 260); see Neubert et al. 2009: fig. 156 for distributional data.

Current systematic position.

Bothriembryontidae, Placostylus eddystonensis (Pfeiffer, 1855).

Bulimus edwardsianus Gassies, 1863

http://species-id.net/wiki/Bulimus_edwardsianus

Figs 9F, 9vi
Bulimus edwardsianus Gassies 1863: 40, pl. 4 fig. 2; Breure and Schouten 1985: 60.
Placostylus caledonicus (Petit); Neubert et al. 2009: 104, fig. 171.
Type locality.

[New Caledonia] “Nékété, Kanala, l’île Nu, l’île Pôt”.

Label.

“Nékété”, taxon label in Gassies’ handwriting, marked “T”.

Dimensions.

“Long. 80 mill.—Diam. maj. 34 mill.”; specimen figured herein H 72.0, D 36.5, W 6.7.

Type material.

NHMUK 1883.11.10.1172, one possible syntype (ex Gassies ex Marie).

Remarks.

Neubert et al. (2009) consider this specimen a possible syntype; it is not the originally figured specimen.

Current systematic position.

Bothriembryontidae, Placostylus caledonicus (Petit de la Saussaye, 1845).

Succinea elegans Pfeiffer, 1842

http://species-id.net/wiki/Succinea_elegans

Figs 4G–H, 4iv
Succinea elegans Pfeiffer 1842: 56; Pfeiffer 1843: 187; Pfeiffer 1855, in Küster and Pfeiffer 1840–1865: pl. 60 fig. 7. Not Succinea elegans Risso, 1826.
Plectostylus coquimbensis perelegans (Pilsbry); Breure 1978: 201, pl. 9 fig. 14 (lectotype designation).
Type locality.

“Huasco, Chile (Bridges)”.

Label.

“Questa de Arenas / Huasco, Chili”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

“Long. 36, diam. 17 mill.”; lectotype H 30.0, D 16.2, W 5.1.

Type material.

NHMUK 1975360, lectotype; 1975361, three paralectotypes (Cuming coll.).

Remarks.

The lectotype corresponds to the specimen figured by Pfeiffer (1855 [1840–1865]). The original shell height may have been in error. This taxon is now considered a junior subjective synonym of Bulinus broderipii Sowerby I, 1832 (F. Cádiz, unpublished data).

Current systematic position.

Bothriembryontidae, Plectostylus broderipii (Sowerby I, 1832) (synon. n.).

Bulimus broderipi elongata d’Orbigny, 1837

http://species-id.net/wiki/Bulimus_broderipi_elongata

Figs 4E–F, 4iii
Bulimus broderipi var. elongata d’Orbigny 1837 [1834–1847]: 266.
Type locality.

“Cobija, Bolivia” [Chile].

Label.

“Cobija”; in d’Orbigny’s handwriting.

Dimensions.

“Long. 30 millim.; lat. 17 millim.”; lectotype H 30.5, D 16.4, W 5.2.

Type material.

NHMUK 1854.12.4.102, lectotype and six paralectotypes (d’Orbigny coll.).

Remarks.

No material of this taxon has been found in the MNHN. As the species was not figured by d’Orbigny, the specimen that most closely matches the given dimensions has been chosen as the lectotype. This taxon is now considered a junior subjective synonym of Bulinus broderipii Sowerby I, 1832 (F. Cádiz, unpublished data).

Current systematic position.

Bothriembryontidae, Plectostylus broderipii (Sowerby I, 1832) (synon. n.).

Bulimus falcicula Gassies, 1871

http://species-id.net/wiki/Bulimus_falcicula

Figs 7B, 7ii
Bulimus falcicula Gassies 1871: 190.
Placostylus (Placostylus) falcicula (Gassies); Breure and Schouten 1985: 59.
Placostylus (Placostylus) fibratus fibratus (Martyn); Neubert et al. 2009: 50, fig. 15.
Type locality.

[New Caledonia] “Baie du Sud (Nouvelle-Calédonie)”.

Label.

No locality label; taxon label in Gassies’ handwriting.

Dimensions.

“Long. 80 mill.; diam. maj. 40 mill.”; specimen figured herein H 74.5, D 35.9, W 6.5.

Type material.

NHMUK 1883.11.10.1174, syntype, R.P. Lambert leg. (ex Gassies).

Remarks.

The taxon label is marked “T”, and the shell is marked with “245” on the last whorl.

Current systematic position.

Bothriembryontidae, Placostylus fibratus fibratus (Martyn, 1784).

Placostylus (Eumecostylus) foxi Clench, 1950

http://species-id.net/wiki/Placostylus_foxi

Figs 11B, 11ii
Placostylus (Eumecostylus) foxi Clench 1950: 3, figs 1–2; Breure and Schouten 1985: 68.
Eumecostylus foxi Clench; Delsaerdt 2010: 34, pl. 3 figs 9–15.
Type locality.

“Heuru western end of San Cristobal Island, Solomon Islands”.

Label.

“Brit. Solomon Isl. / Heuru, western Cristobal Isl.”.

Dimensions.

“Length 86.8, Gt. width 32 [mm]”; specimen figured herein H 82.3, D 31.5, W 5.9.

Type material.

NHMUK 1953.2.4.1, paratype (ex Schlesch).

Remarks.

Delsaerdt (2010) corrected the coordinates of the type locality as given by Clench. The specimen in NHMUK was obtained “Through Dr Quick” from the Schlesch collection and has a label “paratype” glued within the aperture. It may be noted that Clench (1950) stated “Paratypes (...) in the Museum of Comparative Zoology, the B.P. Bishop Museum and the collections of A.G. Stevenson, Reverend Fox and Walter J. Eyerdam”. It is not clear how the specimen reached the Schlesch collection or if it was genuinely part of the original series.

Current systematic position.

Bothriembryontidae, Eumecostylus foxi (Clench, 1950).

Bulimus fuligineus Pfeiffer, 1853

http://species-id.net/wiki/Bulimus_fuligineus

Figs 15A, 15i
Bulimus fuligineus Pfeiffer 1853b: 301; Pfeiffer 1854, in Küster and Pfeiffer 1840–1865: 157, pl. 48 figs 5–6.
Bulimus fuligineus var. β Pfeiffer 1859: 363.
Placostylus (Santacharis) fuligineus (Pfeiffer); Solem 1959: 134, pl. 8 figs 3, 6.
Type locality.

[Vanuatu] “Novis Hebridis”.

Label.

“New Hebrides”. “Isle of Aneiteum”; taxon labels in Pfeiffer’s handwriting. M.C. label style IV.

Dimensions.

“Long. 38, diam. 16 mill.”; lectotype H 39.4, D 19.2, W 4.6.

Type material.

NHMUK 1975226, lectotype and two paralectotypes; 20100650, three paralectotypes of the varietal form, all Macgillivray leg. (Cuming coll.).

Remarks.

Two lots, each consisting of three specimens proved to be present, one corresponding to the variety recognized by Pfeiffer (1859). All material was taken at Vanuatu, Prov. Tafea, Anatom (of which Anaiteum and Aneityum are a few of the variant names), according to the label glued inside the aperture of one of the specimens and the label of the variety lot. The specimen figured by Pfeiffer (1854: pl. 48 figs 5–6) is now seclected lectotype (design. n.).

Current systematic position.

Bothriembryontidae, Santacharis fuligineus (Pfeiffer, 1853).

Bulimus fuscagula d’Orbigny, 1837

http://species-id.net/wiki/Bulimus_fuscagula

Figs 23E–F, 23iii
Bulimus fuscagula d’Orbigny 1837 [1834–1847]: 318, pl. 39 figs 1–2; Gray 1854: 22.
Bahiensis miliola (d’Orbigny); Simone 2006: 169, fig. 591b.
Type locality.

[Brazil] “Rio de Janeiro”.

Label.

“In Bresil”

Dimensions.

“Long. 30 millim.; lat. 8 millim.”; lectotype H 28.6, D 7.6, W 9.4.

Type material.

NHMUK 1854.12.4.141, lectotype (d’Orbigny coll.).

Remarks.

This specimen was thought by d’Orbigny to represent Auricula fuscagula Lea, 1837 [= Cyclodontina punctatissima (Lesson, 1830)], which is, however, a distinct species. The type lot consisted of a second specimen (NHMUK 1854.12.4.142) which indeed belongs to this species described by Lesson. Therefore, to solve the potential confusion, specimen NHMUK 1854.12.4.141 is here designated as lectotype (design. n.) of Bulimus fuscagula d’Orbigny.

Current systematic position.

Odontostomidae, Bahiensis fuscagula (d’Orbigny, 1837).

Bulimus souvillei gatopensis Crosse, 1870

http://species-id.net/wiki/Bulimus_souvillei_gatopensis

Figs 8B–C, 8iii
Bulimus souvillei var. δ gatopensis Crosse 1870: 242; Breure and Schouten 1985: 69; Neubert et al. 2009: 61.
Type locality.

“Gatope et in loco ‘Ferme modèle’ dicto, Novae Caledoniae. (E. Marie)”.

Label.

No locality given. Taxon label in Crosse’s handwriting (?).

Dimensions.

“Long. 88–95, diam. maj. 47–56 mill.”; specimen figured herein H 94.5, D 55.5, W 6.5.

Type material.

NHMUK 1929.9.14.4, syntype, ex Marie coll.

Remarks.

A label (from Crosse?) with the note “type” is glued on the dorsal side of the last whorl. Neubert et al. (2009) have commented on the type locality of this material, noting it to be erroneous or based on introduced material. A further syntype is in the MNHN collection.

Current systematic position. Bothriembryontidae, Placostylus fibratus guestieri (Gassies, 1869).

Bulimus fibratus grammica Crosse, 1870

http://species-id.net/wiki/Bulimus_fibratus_grammica

Figs 9A, 9i
Bulimus fibratus grammica Crosse 1870: 242; Neubert et al. 2009: 62, fig. 54.
Type locality.

“Nouvelle-Calédonie”.

Label.

“New Caledonia” added in a later handwriting. “type de l’auteur” glued on shell.

Dimensions.

“Long. 111, diam. maj. 47 mill.”; specimen figured herein H 111.3, D 52.5, W 7.4.

Type material.

NHMUK 1929.9.14.5, syntype, ex Marie coll.

Remarks.

In the recent revision of New Caledonian Placostylus (Neubert et al. 2009) an argument is made for the current systematic position of this taxon.

Current systematic position.

Bothriembryontidae, Placostylus fibratus goroensis (Souverbie, 1870).

Bulimus grayanus Pfeiffer, 1845

http://species-id.net/wiki/Bulimus_grayanus

Figs 24E–H, 24ii
Bulimus grayanus Pfeiffer 1845b: 73.
Macrodontes grayanus (Pfeiffer); Simone 2006: 162, fig. 556.
Type locality.

“Brazils”.

Label.

“Brazil”. M.C. label style III.

Dimensions.

“Long. 35, diam. 11 mill.”; specimen figured herein H 34.1, D 12.8, W 6.0.

Type material.

NHMUK 20110387, syntype (Cuming coll.).

Current systematic position.

Odontostomidae, Odontostomus grayanus (Pfeiffer, 1845).

Helix guarani d’Orbigny, 1835

http://species-id.net/wiki/Helix_guarani

Figs 18A–B, 18iii
Helix guarani d’Orbigny 1835: 21.
Bulimus guarani d’Orbigny 1837 [1834–1847]: 518, pl. 41bis fig. 1.
Type locality.

“provincia Corrientesensi (republica Argentina)”.

Label.

“corrientes. rep. Argentina”, label in d’Orbigny’s handwriting.

Dimensions.

“Longit. 23 millim., latit. 8 1/2”; lectotype H 20.5, D 7.4, H 8.1.

Type material.

NHMUK 1854.12.4.140, lectotype and one paralectotype (d’Orbigny coll.).

Remarks.

The specimen corresponding to d’Orbigny 1837 [1834–1847]: pl. 41 bis fig. 1 has been selected lectotype (design. n.); the apex of the specimen is slightly damaged and its current shell height is somewhat smaller than in the original measurements. This species is tentatively classified with Bahiensis, awaiting a generic level revision of this group.

Current systematic position.

Odontostomidae, Bahiensis guarani (d’Orbigny, 1835).

Placostylus guppyi E.A. Smith, 1891

http://species-id.net/wiki/Placostylus_guppyi

Figs 14C, 14iii
Placostylus guppyi E.A. Smith 1891: 489, pl. 40 fig. 6; Breure and Schouten 1985: 69 (lectotype designation).
Placocharis guppyi (Smith); Delsaerdt 2010: 60, pl. 11 figs 12–13.
Type locality.

“Solomon Islands”; restricted to West Guadalcanal by Delsaerdt (2010).

Label.

“Solomon Is.”, taxon label in Smith’s handwriting.

Dimensions.

“Longit. 80 millim., diam. maj. 37”; lectotype H 80.1, D 40.1, W 5.5.

Type material.

NHMUK 1891.11.6.110, lectotype; 1891.11.6.111, paralectotype, Brazier leg.

Current systematic position.

Bothriembryontidae, Placocharis guppyi (E.A. Smith, 1891).

Bulimus hargravesi Cox, 1871

http://species-id.net/wiki/Bulimus_hargravesi

Figs 12A, 12i
Bulimus hargravesi Cox 1871: 323, pl. 34 fig. 3; Breure and Schouten 1985: 69.
Eumecostylus hargravesi (Cox); Delsaerdt 2010: 47, pl. 8 figs 1–6.
Type locality.

“Treasury Island, Solomon Islands”, corrected to Malaita, Solomon Islands (Delsaerdt 2010).

Label.

“Solomon Isls”.

Dimensions.

“Diam. 1.05, long. 2.60 unc. [H 64.3, D 25.9 mm]”; lectotype H 65.0, D 30.9, W 5.3.

Type material.

NHMUK 1880.12.11.43, lectotype (ex Cox).

Remarks.

The holotype designation (Breure and Schouten 1985) has to be interpreted as lectotype designation (Art. 74.6 ICZN); the claim of Delsaerdt (2010: unnumbered page opposite contents [not repeated in main text]) of a lectotype designation is thus erroneous. Delsaerdt interpreted Cox’ abbreviation “unc.” equalling 25.4 mm given his dimensions of the type specimens as 26.67 x 66.04 mm. Cox’ abbreviation can also be read as the Roman uncia, equal to 2.473 cm (Rowlett 2004), but according to Woodward (1851 [1851–1856]: 61) the uncia equals the inch (= 25.4 mm).

Current systematic position.

Bothriembryontidae, Eumecostylus hargravesi (Cox, 1871).

Bulimus infundibulum Gassies, 1871

http://species-id.net/wiki/Bulimus_infundibulum

Figs 8A, 8i
Bulimus infundibulum Gassies 1871: 86; Breure and Schouten 1985: 69; Neubert et al. 2009: 55, fig. 23.
Type locality.

[New Caledonia] “Ilôt Koutoumo, sud de la Nouvelle-Calédonie”.

Label.

No locality label. Taxon label in Gassies’ handwriting and marked “T”.

Dimensions.

“Long. 75 mill.; diam. maj. 40”; holotype H 75.6, D 46, W 6.5.

Type material.

NHMUK 1883.11.10.1163, holotype, ex Gassies.

Remarks.

Gassies (1871) remarked that he had seen only one specimen. The single specimen known is a monstrosity; it is marked “241” on the last whorl.

Current systematic position.

Bothriembryontidae, Placostylus fibratus guestieri (Gassies, 1869)

Bulimus kingii J.E. Gray, 1825

http://species-id.net/wiki/Bulimus_kingii

Figs 2F–G, 2i
Bulimus kingii J.E. Gray 1825: 414; B.J. Smith 1992: 104.
Bothriembryon (Bothriembryon) kingii (Gray); Breure 1978: 205 (lectotype designation); Breure 1979: 94.
Type locality.

[Australia] “New Holland, Capt. King”.

Label.

“Australia”.

Dimensions.

“Long. 1, diam. 1/2 unc.”; lectotype H 24.5, D 10.6, W 5.5.

Type material.

NHMUK 195910, lectotype; 195911, one paralectotype, ex P.P. King R.N.

Remarks.

According to Pilsbry (1900) and Iredale (1939) the type locality is Western Australia, King George Sound, near Bald Head. Captain King visited this locality on 20 Juanuary 1818 and stated “An abundance of shells of the helix tribe (Helix bulimus) was found on the top and sides of the hill” (King 1827: 12).

Current systematic position.

Bothriembryontidae, Bothriembryon kingii (J.E. Gray, 1825).

Bulimus koroensis Garrett, 1872

http://species-id.net/wiki/Bulimus_koroensis

Figs 13E, 13iv
Bulimus koroensis Garrett 1872: 236, pl. 18 fig. 9.
Type locality.

“Koro Isl., Viti Isles”.

Label.

“Koro, Viti”.

Dimensions.

“Length 53 mill.; diam. 18 mill.”; specimen figured herein H 51.2, D 18.0, W 5.1.

Type material.

NHMUK 20030802, two probable syntypes, ex Taylor, ex Garrett.

Current systematic position.

Bothriembryontidae, Euplacostylus koroensis (Garrett, 1872).

Bulimus (Charis) kreftii Cox, 1872

http://species-id.net/wiki/Bulimus_kreftii

Figs 14A, 14i–ii
Bulimus (Charis) kreftii Cox 1872b: 19, pl. 4 fig. 4.
Placocharis kreftii (Cox); Delsaerdt 2010: 67, pl. 11 figs 14–17 (lectotype designation).
Type locality.

“Solomon Islands”, restricted to Florida Islands by Delsaerdt (2010).

Label.

“Solomon Isles”, in Cox’ handwriting (see remarks).

Dimensions.

“Length 2.10, breadth 0.92 of an inch [H 53.3, D 23.3]”; specimen figured herein H 54.9, D 24.3, W 5.2.

Type material.

NHMUK 20070068, lectotype and three paralectotypes, ex Kennard ex Cox; 1880.12.11.46, paralectotype, ex Cox.

Remarks.

Both series are accompanied by a label in Cox’ handwriting. The specimen registered 1880.12.11.46 was presented by J.C. Cox, has a taxon label “Bulimus kreftii” and corresponds to pl.4 fig. 4 of Cox (1872). The other lot (ex A.S. Kennard collection) has a taxon label “Placostylus krefftii” [sic, misspelled] and is “(Ex Auct.)”; the locality on Kennard’s label “New Caledonia” is clearly in error. It is therefore more likely that the specimen presented by Cox was truly part of the type series; those from the Kennard collection may have been collected at a later stage, for which the genus name Placostylus on the taxon label written by Cox (Fig. 14ii) is an indication.

Current systematic position.

Bothriembryontidae, Placocharis kreftii (Cox, 1872).

Bulimus leeuwinensis E.A. Smith, 1894

http://species-id.net/wiki/Bulimus_leeuwinensis

Figs 3C–E, 3ii
Bulimus leeuwinensis E.A. Smith 1894: 94, pl. 7 fig. 27; B.J. Smith 1992: 105.
Bothriembryon (Bothriembryon) leeuwinensis (Smith); Breure 1978: 206 (lectotype designation); Breure 1979: 94.
Type locality.

“Cape Leeuwin, S.W. Australia”.

Label.

“Cape Leeuwin, S.W. Australia”.

Dimensions.

“Longit. 27 mm.; diam. 12.5 mm.”; lectotype H 27, D 13.7, W 5.4.

Type material.

NHMUK 1891.11.21.128, lectotype; 1891.11.21.129–132, four paralectotypes, J.J. Walker leg.

Current systematic position.

Bothriembryontidae, Bothriembryon leeuwinensis (E.A. Smith, 1894).

Bulimus longulus ‘Behn’ Pfeiffer, 1859

http://species-id.net/wiki/Bulimus_longulus

Figs 17A–B, 17i
Bulimus longulus ‘Behn’ Pfeiffer 1859a: 44.
Bahiensis longulus (Pfeiffer); Simone 2006: 169, fig. 590.
Type locality.

[Brazil] “Chicatas Brasiliae”.

Label.

“Brazils”. M.C. label style III.

Dimensions.

“Long. 30, diam. 7 1/2 mill.”; specimen figured herein H 29.8, D 7.23, W 11.2.

Type material.

NHMUK 20110388, syntype (Cuming coll.).

Remarks.

Pfeiffer (1859) stated that the material was in “Mus. Cuming”. The specimens found are not accompanied by a taxon label in Pfeiffer’s handwriting, but their type status is not disputed. Simone (2006) listed the specimen as lectotype, but we are unaware of a previous lectotype designation for this taxon. His listing does not fulfill the requirements of ICZN Art. 74.7, and therefore the specimen is regarded as a syntype. The registration number quoted by him was incorrect. The type locality has not been found using modern gazetteers.

Current systematic position.

Odontostomidae, Bahiensis longulus (Pfeiffer, 1859).

Bulimus macgillivrayi Pfeiffer, 1855

http://species-id.net/wiki/Bulimus_macgillivrayi

Figs 14B, 14v
Bulimus macgillivrayi Pfeiffer 1855b: 108, pl. 32 fig. 2.
Placostylus (Placocharis) macgillivrayi (Pfeiffer); Breure and Schouten 1985: 63 (lectotype designation).
Placocharis macgillivrayi (Pfeiffer); Delsaerdt 2010: 61, pl. 10 figs 9–11.
Type locality.

“Wanderer Bay, Guadalcanar, Salomon’s Islands (Macgillivray)”.

Label.

“Wanderer Bay, Guadal- / canar, Solomon’s Islds (Macgillivray)”. M.C. label style III.

Dimensions.

“Long. 59, diam. 22 mill.”; lectotype H 58.5, D 24.2, W 5.4.

Type material.

NHMUK 1975232, lectotype, and two paralectotypes, Macgillivray leg. (Cuming coll.).

Current systematic position.

Bothriembryontidae, Placocharis macgillivrayi (Pfeiffer, 1855).

Pupa spixii major d’Orbigny, 1837

http://species-id.net/wiki/Pupa_spixii_major

Figs 22A–E, 22i
Helix spixii var. major d’Orbigny 1835: 21 [nomen nudum].
Pupa spixii var. α major d’Orbigny 1837 [1834–1847]: 320.
Pupa spixii (d’Orbigny); Gray 1854: 23 [partim].
Spixia striata (Spix); Breure 1976: 1158 [partim].
Type locality.

“...pays habité par les Guarayos, au sein des forêts humides des frontiers nord de la province de Chiquitos (république de Bolivia), et dans la province de Corrientes (république Argentine), en un bois voisin de la rivière de Santa-Lucia, au lieu dit “Pasto reito””; see Breure 1973: 122.

Label.

Two labels, “guarayos” and “corrientes, Rep. Argentina”, both in d’Orbigny’s handwriting.

Dimensions.

“Long. var. A, 35 millim., (...) Lat. var. A, 12 millim.”; lectotype H 34.8, D 11, W 10.6.

Type material.

NHMUK 1854.12.4.232, lectotype, matching best the original shell height; NHMUK 1854.12.4.232, six paralectotypes (d’Orbigny coll.).

Remarks.

In d’Orbigny (1835) only the name of this taxon is mentioned, together with a locality, thus it was invalidly published. The first valid publication is in d’Orbigny 1837 [1834–1847], where only a brief description was given but no figures of the ventral view of the shell (only a side view of a living snail). There is no explicit mentioning of “var. major” on the label, as d’Orbigny has done with varieties of other taxa; the only link are the localities quoted in his original paper. This taxon is here figured for the first time. As the lots referring to Pupa spixii refer to two taxa, a lectotype is here designated for each (design. n.); see also minor d’Orbigny.

Current systematic position.

Odontotomidae, Spixia striata (Spix, 1827).

Bulimus miltocheilus Reeve, 1848

http://species-id.net/wiki/Bulimus_miltocheilus

Figs 10C, 10iii
Bulimus miltocheilus Reeve 1848 [1848–1850]: pl. 49 fig. 322; Breure and Schouten 1985: 70.
Aspastus miltocheilus (Reeve); Delsaerdt 2010: 21 (lectotype designation).
Type locality.

[Solomon Islands] “San Christoval, southeastern island of Solomon’s Group, northeast coast of New Holland”. Restricted to [Makira] “Kirakira (Wainoni-Bay)” by Rensch and Rensch (1935: 72).

Label.

“San Cristoval, Solomon Is.”. M.C. label style IV.

Dimensions.

Not given. Specimen figured herein H 68.3, D 25.0, W 6.0.

Type material.

NHMUK 20070072, two paralectotypes (Cuming coll.).

Remarks.

Delsaerdt (2010) has selected Reeve’s original figure as the lectotype.

Current systematic position.

Bothriembryontidae, Aspastus miltocheilus (Reeve, 1848).

Pupa spixii minor d’Orbigny, 1837

http://species-id.net/wiki/Pupa_spixii_minor

Figs 21A–F, 21i
Helix spixii var. minor d’Orbigny 1835: 21 [nomen nudum].
Pupa spixii var. β minor d’Orbigny 1837 [1834–1847]: 320.
Pupa spixii (d’Orbigny); Gray 1854: 23 [partim].
Spixia striata (Spix); Breure 1976: 1158 [partim].
Type locality.

[Bolivia] “province de Chiquitos, entre Santo-Corazon et San-Juan”; see Breure 1973: 123.

Label.

“chiquitos, Bolivia”; in d’Orbigny’s handwriting.

Dimensions.

“Long. (...) var. B, 30 millim. Lat. (...) var. B, 7 millim.”; lectotype H 29.2, D 7.46, W 11.1.

Type material.

NHMUK 1854.12.4.231, lectotype and seven paralectotypes.

Remarks.

In d’Orbigny (1835) only the name of this taxon is mentioned, together with a locality, thus it was invalidly published. One specimen, which most closely matches the original shell height, is here designated as the lectotype. As the lots referring to Pupa spixii refer to two taxa, a lectotype is here designated for each (design. n.); see also major d’Orbigny. This taxon is here figured for the first time.

Current systematic position.

Odontostomidae, Cyclodontina minor (d’Orbigny, 1837).

Bulimus necouensis Gassies, 1871

http://species-id.net/wiki/Bulimus_necouensis

Figs 8I, 8iv
Bulimus necouensis Gassies 1871: 191; Neubert et al. 2009: 68, fig. 75.
Type locality.

[New Caledonia] “Nécoué, Baie Lebris (Nouvelle-Calédonie)”.

Label.

No locality information on the label, which gives the taxon name as “Bulimus Nekouensis”. A second label states “New Caledonia” but this is likely to have been added after the receipt of the material.

Dimensions.

“Long. 77 mill.; diam. maj. 33 mill.”; holotype H 73.9, D 33.6, W 6.5.

Type material.

NHMUK 1883.11.10.1175, holotype, E. Marie leg. (ex Gassies).

Remarks.

The specimen is marked “T” above the aperture on the last whorl. Neubert et al. (2009) have clarified the type status of this specimen and the taxonomic status of the taxon.

Current systematic position.

Bothriembryontidae, Placostylus fibratus alexander (Crosse, 1855).

Bulimus neglectus Pfeiffer, 1847

http://species-id.net/wiki/Bulimus_neglectus

Figs 18H–I, 18i
Bulimus neglectus Pfeiffer 1847: 67.
Clessinia neglecta (Pfeiffer); Breure and Schouten 1985: 10 (lectotype designation); Simone 2006: 164, fig. 564.
Type locality.

[Brazil] “Brasilia”.

Label.

“Brazils”, taxon label in Pfeiffer’s handwriting. M.C. label style I.

Dimensions.

“Long 19, diam. 6 mill.”; lectotype H 16.8, D 5.18, W 8.2.

Type material.

NHMUK 1975263, lectotype and one paralectotype (Cuming coll.).

Remarks.

A note accompanies the material stating “types / var. designation not Pfeiffer’s”. However, the label on the back of the cardbox has Pfeiffer’s handwriting, and the type status of these specimens is not disputed here. Simone (2006) incorrectly gave the size of the lectotype as 21 mm height. This species is tentatively classified with Bahiensis, a group awaiting a generic level revision.

Current systematic position.

Odontostomidae, Bahiensis neglecta (Pfeiffer, 1847).

Buliminus nuptialis Melvill and Ponsonby, 1894

http://species-id.net/wiki/Buliminus_nuptialis

Figs 6D–F, 6iii
Buliminus nuptialis Melvill and Ponsonby 1894: 92, pl. 1 fig. 5.
Prestonella nuptialis (Melvill and Ponsonby); Herbert 2007: 11, fig. 12–13.
Type locality.

[South Africa, Eastern Cape] “Craigie Burn, Somerset East (Mrs. Mary Layard Barber)”.

Label.

“Craigie Burn, Somerset / East, S. Africa”.

Dimensions.

“Long. 15, lat. 8.50 mill.”; holotype H 14.6, D 9.82, W 4.4.

Type material.

NHMUK 1905.1.26.29, holotype, Mrs. Mary Layard Barber (née Bowker) (ex E.L. Layard ex Ponsonby).

Remarks.

The data on the collector and the way this specimen has found its way to the NHMUK collection have been taken from Herbert (2007). The size of the specimen was erroneously given as “length 11.5 mm”.

Current systematic position.

Bothriembryontidae, Prestonella nuptialis (Melvill and Ponsonby, 1894).

Bulimus oblitus Reeve, 1848

http://species-id.net/wiki/Bulimus_oblitus

Figs 18F–G, 18ii
Bulimus oblitus Reeve 1848 [1848–1850]: pl. 56 fig. 376.
Clessinia oblita (Reeve); Breure and Schouten 1985: 10 (lectotype designation); Simone 2006: 164, fig. 565.
Type locality.

“Brazil”.

Label.

“Brazil”. M.C. label style III.

Dimensions.

Not given. Lectotype H 20.9, D 6.27, W 9.2+.

Type material.

NHMUK 1975254, lectotype and two paralectotypes (Cuming coll.).

Remarks.

This species is tentatively classified with Bahiensis, awaiting a revision at generic level of this group. Bulimus oblitus is now considered a junior subjective synonym of Bahiensis neglecta (Pfeiffer, 1847) (synon.n.).

Current systematic position.

Odontostomidae, Bahiensis neglecta (Pfeiffer, 1847).

Bulimus occultus Reeve, 1849

http://species-id.net/wiki/Bulimus_occultus

Figs 17C–D, 17ii
Bulimus occultus Reeve 1849 [1848–1850]: pl. 83 fig. 617.
Cyclodontina (Bahiensis) occultus (Reeve); Breure and Schouten 1985: 7, pl. 1 fig. 3 (lectotype designation).
Bahiensis occultus (Reeve); Simone 2006: 169, fig. 592.
Type locality.

“Brazil”.

Label.

“Brazil”. M.C. label style III.

Dimensions.

Not given. Lectotype H 23.6, D 5.8, W 9.0.

Type material.

NHMUK 1975258, lectotype (Cuming coll.).

Remarks.

See also the discussion under Bulimus parallelus Pfeiffer, 1857.

Current systematic position.

Odontostomidae, Bahiensis occultus (Reeve, 1849).

Bulimus ochrostoma Garrett, 1872

http://species-id.net/wiki/Bulimus_ochrostoma

Figs 13B, 13iii
Bulimus ochrostoma Garrett 1872: 232, pl. 18 fig. 3.
Type locality.

[Fiji, Taviuni Island] “Tavinni [sic] Island, Viti Islands”.

Label.

“Taviuni, Viti”.

Dimensions.

“Length 29 mill., diam. 16 mill.”; specimen figured herein H 35.8, D 18.3, W 4.7.

Type material.

NHMUK 20030803, two probable syntypes (ex Taylor ex Garrett).

Remarks.

Garrett’s original description mentioned that types were in the author’s collection and in the ANSP. The specimens originate from Garrett and were probably part of the original series.

Current systematic position.

Bothriembryontidae, Callistocharis ochrostoma (Garrett, 1872).

Bulimus odontostoma Sowerby I, 1824

http://species-id.net/wiki/Bulimus_odontostoma

Figs 24A–D, 24i
Bulimus odontostoma Sowerby I 1824: 59, pl. 5 fig. 3.
Type locality.

[Brazil] “Rio de Janeiro”.

Label.

“Brazil”, in a later hand. M.C. label style IV.

Dimensions.

“Axis 1 4/10, diam. 4/10 unc. [H 35.6, D 10.16 mm]”; specimen figured herein H 37.1, D 12.8, W 6.2.

Type material.

NHMUK 20100632, two possible syntypes (Cuming coll.).

Remarks.

According to Sowerby, he had seen “A few specimens...”. Since it is not unlikely that Sowerby made use of the Cuming collection to describe species (Petit 2009), the specimens are considered possible syntpes.

Current systematic position.

Odontostomidae, Odontostomus gargantua (Férussac, 1821).

Bulimus ouensis Gassies, 1870

http://species-id.net/wiki/Bulimus_ouensis

Figs 7D, 7iv
Bulimus ouensis Gassies 1870: 142.
Placostylus (Placostylus) fibratus (Martyn); Breure and Schouten 1985: 60.
Placostylus fibratus fibratus (Martyn); Neubert et al. 2009: 50, fig. 10.
Type locality.

“Insula Ouen, Novae Caledoniae”.

Label.

Only a taxon label is present, with the name written in Gassies’ handwriting and marked “T”.

Dimensions.

“Long. 86, diam. maj. 41 mill.”; holotype H 83.7, D 40.7, W 7.3.

Type material.

NHMUK 1883.11.10.1176, holotype (ex Gassies).

Remarks.

The specimen is marked “T” on the last whorl beside the aperture. Gassies (1870) mentioned that he had seen only one specimen.

Current systematic position.

Bothriembryontidae, Placostylus fibratus fibratus (Martyn, 1784).

Bulimus parallelus Pfeiffer, 1857

http://species-id.net/wiki/Bulimus_parallelus

Figs 17E–F, 17iii
Bulimus parallelus Pfeiffer 1857: 389; Breure and Schouten 1985: 7 (lectotype designation).
Moricandia parallela (Pfeiffer); Simone 2006: 172, fig. 606.
Type locality.

“St. Catherine’s, Brazil”.

Label.

“St. Catherines Brazil”, taxon label in Pfeiffer’s handwriting. M.C. label style III.

Dimensions.

“Long. 22, diam. 7 mill.”; lectotype H 22.5, D 7.17, W 7.3.

Type material.

NHMUK 1975256, lectotype and one paralectotype (Cuming coll.).

Remarks.

Simone (2006) has revalidated this taxon as a separate species, removing it from the synonymy of Bahiensis occultus (Reeve, 1849) (See also Pisbry 1901 [1901–1902]: 48), and placing it in the genus Moricandia Pilsbry & Vanatta, 1898. Pfeiffer’s type is somewhat stouter, but has the same characteristic brown blotch behind the lip. Tentatively, the synonymy with Bulimus occultus Reeve, 1849 is here retained.

Current systematic position.

Odontostomidae, Bahiensis occultus (Reeve, 1849).

Helix patagonica d’Orbigny, 1835

http://species-id.net/wiki/Helix_patagonica

Figs 25E–G, 25ii
Helix patagonica d’Orbigny 1835: 22.
Bulimus patagonicus d’Orbigny 1837 [1834–1847]: 321, pl. 41 figs 17–18.
Plagiodontes patagonicus (d’Orbigny); Breure 1976: 1159, pl. 5 fig. 4, pl. 10 fig. 4 (lectotype designation).
Type locality.

[Argentina] “Patagonia”.

Label. “baie blanche [Bahia Blanca], Patagonia”, in d’Orbigny’s handwriting.

Dimensions.

“Longit. 22 1/2 millim., latit. 11 millim.”; specimen figured herein H 22.3, D 11.2, W 7.5.

Type material.

NHMUK 1854.12.4.235, five paralectotypes (d’Orbigny coll.).

Remarks.

The lectotype is in the MNHN collection (Breure 1976).

Current systematic position.

Odontostomidae, Plagiodontes patagonica (d’Orbigny, 1835).

Bulimus physoides Reeve, 1849

http://species-id.net/wiki/Bulimus_physoides

Figs 2H–I, 2iv
Bulimus physoides Reeve 1849 [1848–1850]: pl. 70 fig. 507.
Bulimus (Liparus) physoides Reeve; E.A. Smith 1894: 95, pl. 7 fig. 30.
Type locality.

“—?”.

Label.

“W. Australia”, taxon label in Pfeiffer’s handwriting. M.C. label style III.

Dimensions.

Not given. Specimen figured herein H 19.2, D 11.22, W 5.0.

Type material.

NHMUK 1975224–1975225, three possible syntypes (Cuming coll.).

Remarks.

Reeve (legend to pl. 70: “Mus. Pfeiffer”) evidently based his description on material from Pfeiffer, who referred to “physodes (Menke mss.)”; this species was published as such by Menke (1843: 7), and Reeve’s name is therefore probably an emendation of Menke’s name. However, as Menke’s material is presumably lost (B.J. Smith 1992: 105), these specimens are interesting, since they may have originated via Pfeiffer from Menke’s collection.

Current systematic position.

Bothriembryontidae, Bothriembryon melo (Quoy and Gaimard, 1832).

Bulimus pinicola Gassies, 1870

http://species-id.net/wiki/Bulimus_pinicola

Figs 8D, 8ii
Bulimus pinicola Gassies 1870: 142.
Placostylus (Placostylus) fibratus (Martyn); Breure and Schouten 1985: 60.
Placostylus fibratus guestieri (Gassies); Neubert et al. 2009: 55, fig. 21.
Type locality.

“Insula Pinorum, Novae Caledoniae”.

Label.

Only a taxon label “Bulimus pinicula” in Gassies handwriting, marked “T”.

Dimensions.

“Long. 90, diam. maj. 44 mill.”; specimen figured herein H 92.8, D 44.3, W 6.7.

Type material.

NHMUK 1883.11.10.1164, one syntype (ex Gassies).

Remarks.

Marked “T” on last whorl besides aperture. Gassies (1870) stated that he had seen three specimens.

Current systematic position.

Bothriembryontidae, Placostylus fibratus guestieri (Gassies, 1869).

Placostylus ambagiosus priscus Powell, 1938

http://species-id.net/wiki/Placostylus_ambagiosus_priscus

Fig. 12D
Placostylus ambagiosus priscus Powell 1938: 149, pl. 34 figs 9–10.
Type locality.

[New Zealand, North Island] “about three quarters of a mile east of the extreme north-western headland [Cape Maria van Diemen]”.

Label.

“Cape Maria van Diemen (mainland)”

Dimensions.

“Height, 77 mm.; breadth, 31 mm.”; specimen figured herein H 70.3, D 28.5, W 6.5.

Type material.

NHMUK 1939.3.17.18, one paratype (ex Powell).

Remarks.

The holotype of this subspecies, which is supposedly of Pleistocene age, is in the Auckland Museum. Buckley et al. (2011) have revised the New Zealand Placostylus and showed that this taxon falls within the overall variation of Placostylus ambagiosus Suter, 1906.

Current systematic position.

Bothriembryontidae, Placostylus ambagiosus Suter, 1906.

Bulinus