ZooKeys 176: 1–11, doi: 10.3897/zookeys.176.2372
Cartographic analysis of woodlice fauna of the former USSR
Daria M. Kuznetsova 1, Konstantin B. Gongalsky 1
1 A.N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia

Corresponding author: Konstantin B. Gongalsky (gongalsky@gmail.com)

Academic editor: S. Taiti

received 15 November 2011 | accepted 31 January 2012 | Published 20 March 2012

(C) 2012 Daria M. Kuznetsova. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

For reference, use of the paginated PDF or printed version of this article is recommended.


An inventory of the woodlice fauna of the former USSR yielded 190 species, 64 of them were recorded from the territory of Russia. According to the cartographic analysis, the limits of distribution of epigean terrestrial isopods over the area, excluding mountains, is explained by temperature. No woodlice records were found outside the isocline of 120 days a year with the mean daily air temperature >10°C. The highest species diversity was found between the isoclines of 180 and 210 days. These areas correspond to forest-steppe and steppe zones.


Woodlice, mean annual airtemperature, database, Russia


Studies of spatial differentiation of various taxa are among the most important frontiers of modern biogeography. For some well-studied groups, mainly, vertebrates and plants, such trends are already discovered (Loiselle et al. 2003; Guisan and Thuiller 2005; Grenouillet et al. 2011), but for soil-dwelling invertebrates they are only at the stage of species inventory. However, there are certain groups of invertebrates for which analysis of spatial differentiation is already possible due to the large number of records from different geographical localities. Woodlice are among such groups.

There is no faunistic list of terrestrial isopods for the territory of the former USSR until now, as well as of the territory of Russia. However, there are extensive regional lists (Borutzky 1948, 1953; Zalesskaya and Rybalov 1982; Khisametdinova 2007; Gongalsky and Kuznetsova 2011), and numerous records scattered in the literature devoted to soil macrofauna. At the same time, there are only a few ecological studies about factors affecting woodlice distribution over regions of the former USSR (Gongalsky et al. 2005; Khisametdinova 2009).

The aim of the study is to determine the factors affecting woodlice distribution over the plain area of the former Soviet Union. To achieve this, an inventory of species distribution across the study area was made. The task was to create a database indicating locations with woodlice presence/absence overlaid with several environmental variables values distribution.

Material and methods Database

The first step was to compile a list of species for the study area. We made a database of isopod presence or absence in the locations across the whole territory of the former USSR (both plains and mountains). For each record the database includes information about date, data source, geographical coordinates, location, isopod species list or information about woodlice absence in the soil fauna list, biotope, and natural zone.

Three types of information sources of terrestrial isopod locations were used: i) available literature on soil fauna surveys; ii) collections of the Zoological Museum of Moscow State University (Moscow, Russia) and the Zoological Institute of the Russian Academy of Sciences (St.-Petersburg, Russia); and iii) authors’ personal collections. Here we provide a list of woodlice from the territory of the former USSR since some species and localities were not included in the list of Schmalfuss (2003), although it covered the majority of species. To work with regional databases, a specific list would be useful. Since such a list for this area did not exist, the proposed compilation would be a start to be completed in the future. We used the taxonomic system proposed by Schmalfuss (2003) for species naming. Isopod absence was recorded only in extensively surveyed locations.

For cartographic analysis, 259 locations were chosen, 44 of which with woodlice absence. Due to the difficulty of tracing ecological trends in the mountains, only plain territories were involved into the analysis. Some species were excluded from the analysis: i) synanthropic species and ii) species inhabiting azonal locations, such as sea coasts, caves and anthills.

Then database records with isopod presence or absence locations were laid on the geographic maps to perform cartographic analysis.

Cartographic analysis

The map of woodlice distribution was visually compared with the maps of environmental factors (mean annual temperature; the period with temperature above 10°C; mean precipitation; permafrost distribution; soil pH and soil type; vegetation type; natural zones) found in the Agricultural Atlas of the USSR (Tulupnikov 1960) and the Geographical Atlas of the USSR (Kolosova 1980). The data were verified using the WorldClim database (Hijmans et al. 2005).

The database is maintained in MS Excel. Cartographic analysis is done in MapInfo 8.5.

Results and discussion Limits of isopod distribution

Woodlice have not been recorded northwards the isocline of 120 days a year with temperature >10°C (Fig. 1). The northern border of woodlice distribution matches the distribution of this parameter. Other parameters did not coincide with isopod distribution as well as with this isocline (data not shown).

Figure 1.

Map of woodlice presence or absence over the plain territory of the former USSR. The duration of period with temperature >10°C is adapted from Geographical Atlas of the USSR (Kolosova 1980).

Species diversity

In total, 190 species were recorded from the territory of the former USSR (Appendix 1). Among them, 64 were recorded from the territory of Russia. Northernmost natural zone with woodlice records is southern taiga. No woodlice records were in tundra, northern and middle taiga. The species diversity increases southwards, but decreases in the deserts. However, this may be due to the low number of locations extensively studied to reveal local faunas.

Distribution of isopods is known to be limited by natural factors, such as temperature and moisture (Harding and Sutton 1985, Hopkin 1991). In our study, the limiting factor of woodlice distribution towards the north turned out to be the length of the warm period, expressed as number of days when the temperature was above 10°C. The highest species diversity was observed between isoclines of 180 and 210 days with temperature >10°C. Colder conditions slow down their physiological processes (Hopkin 1991) and limit their distribution. For a better understanding of distribution of woodlice, a Species Distribution Modeling (Elith and Leathwick 2009, Franklin 2009) should be applied, which is a next step in the analysis of the database of Russian isopods.


The authors are grateful to Dr H. Schmalfuss and Dr Ch. Schmidt for the help with isopod identifications, and to Dr K.G. Mikhailov and Dr B.V. Mezhov for allowing working with crustacean material kept at Zoological Museum of Moscow State University.

The study is supported by Russian Foundation for Basic Research (grant 11-04-00245) and the Program “Biodiversity” of the Presidium of the Russian Academy of Sciences.

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Appendix 1

List of woodlice species from the territory of the former USSR. Abbreviations: Ab – Abkhazia, Ar – Armenia, Az – Azerbaijan, Bl – Belarus, Ge – Georgia, Kz – Kazakhstan, Kg – Kyrgyzstan, Lt – Lithuania, Md – Moldova, Ru – Russia, Td – Tajikistan, Tu – Turkmenistan, Ua – Ukraine, Uz – Uzbekistan; S, N, W, E – south, north, west, east. References to authorships of the species can be found in Schmalfuss and Wolf-Schwenninger (2002).

1 Acaeroplastes kosswigi Verhoeff, 1941 Az: Nabran’
2 Agabiformius orientalis (Dollfus, 1905) Ru: Volgograd region
3 Armadillidium azerbaidzhanum Schmalfuss, 1990 Ar: Khastarak; Az: Baku, Adjikend, Drmbon, Lenkoran, Salyany, Sheki; Ge: Kakhetia, Vashlovan; Ru: Rostov, Stavropol regions, N Osetia
4 Armadillidium granulatum Brandt, 1833 Ru: Krasnodar region; Ua: S Crimea
5 Armadillidium nasatum Budde-Lund, 1885 Ru: Moscow; Ab: Sukhum
6 Armadillidium opacum (C.Koch, 1841) Ua: Kiev
7 Armadillidium pallasii Brandt, 1833 Ab: Sukhum; Ua: Crimea, Odessa; Ru: Krasnodar region
8 Armadillidium pictum Brandt, 1833 Ua: Crimea
9 Armadillidium pulchellum (Zenker, 1798) Lt: Vilnius
10 Armadillidium traiani Demianowicz, 1932 Md
11 Armadillidium versicolor Stein, 1859 Ru: Penza, Saratov, Tula regions
12 Armadillidium vulgare Latreille, 1804 Ab: Sukhum; Az; Ge: Adygeni, Tbilisi; Ru: Dagestan, Krasnodar, Kaluga, Volgograd, Rostov regions; Ua: Crimea, Kiev, Odessa
13 Armadillidium zenckeri Brandt, 1833 Ua: Crimea, Zakaspyisk region
14 Armadillo alievi Schmalfuss, 1990 Az: Baku, Kobustan
15 Armadillo officinalis Dumeril, 1816 Ru: Krasnodar region; Ua: S Crimea, Odessa
16 Armadilloniscus ellipticus (Harger, 1878) Ru: Krasnodar region; Ab: Gagry
17 Borutzkyella revasi (Borutzky, 1973) Ab: Gudauty region
18 Buddelundiella cataractae Verhoeff, 1930 Ge: Tskhaltubo
19 Caucasocyphonethes cavaticus Borutzky, 1948 Ru: Krasnodar region
20 Caucasoligidium cavernicola Borutzky, 1950 Ab: Gudauty, Sukhum; Ge: Gogolety
21 Caucasonethes borutzkyi Verhoeff, 1932 Ge: Tskhaltubo
22 Chaetophiloscia cellaria Dollfus, 1884 Ru: Rostov region
23 Chaetophiloscia elongata Dollfus, 1884 Ua: Crimea
24 Chaetophiloscia hastata Verhoeff, 1929 Ru: Krasnodar region
25 Colchidoniscus kutaissianus Borutzky, 1974 Ge: Tskhaltubo
26 Cylisticoides angulatus Schmalfuss, 2003 Az: Istisu, Lenkoran, Yardymly; Kz: Astana; Ru: Krasnodar region
27 Cylisticus albomaculatus Borutzky, 1957 Ru: Rostov, Voronezh, Volgograd regions
28 Cylisticus arnoldii Borutzky, 1961 Ua: Kharkov, Zmiev
29 Cylisticus birsteini Borutzky, 1961 Ru: Krasnodar region
30 Cylisticus caucasius Verhoeff, 1917 Ab: Gudauty, Gagry, Kelassuri, Sukhum; Ge: Kutaisi, Tkibuli, Tskhaltubo, Shovi; Ru: Krasnodar region
31 Cylisticus convexus De Geer, 1778 Ru: Chelyabinsk, Kaluga, Moscow, Rostov, Voronezh regions, Ua: S Crimea, Kiev
32 Cylisticus cretaceus Borutzky, 1957 Ru: Rostov region; Ua: Lugansk region
33 Cylisticus desertorum Borutzky, 1957 Ru: Rostov, Stavropol regions; Ua
34 Cylisticus giljarovi Borutzky, 1977 Ru: Adygea, Stavropol, Krasnodar regions, N Osetia
35 Cylisticus iners Budde-Lund, 1880 Ar: Azizbekov, Leninakan, Tshakhkavan; Az: Airidja, Mardakert, Zakatalinsk regions; Ru: Chechnya; Ge: Manglisi, Shuahevi
36 Cylisticus lencoranensis Borutzky, 1977 Az: Prishib
37 Cylisticus mitis Budde-Lund, 1885 Ge: Kutaisi
38 Cylisticus orientalis Borutzky, 1939 Ru: Orenburg region
39 Cylisticus rotabilis Budde-Lund, 1885 Ua: S Crimea
40 Cylisticus sarmaticus Borutzky, 1977 Ru: Rostov region; Ua: Zaporozhye region
41 Cylisticus silvestris Borutzky, 1957 Ru: Moscow, Rostov, Stavropol regions
42 Cylisticus strouhali Borutzky, 1977 Ar: W Vanadzor, Spitaki
43 Desertoniscus birsteini Borutzky, 1945 Tu
44 Desertoniscus bulbifrons Borutzky, 1945 Tu
45 Desertoniscus elongatus Borutzky, 1945 Tu
46 Desertoniscus kirghizicus Borutzky, 1978 Kg
47 Desertoniscus reductus Borutzky, 1978 Td
48 Desertoniscus subterraneus Verhoeff, 1930 Kg; Tu: (Kizil-arvat); Td: Samgar massif
49 Desertoniscus tekinus Borutzky, 1945 Tu
50 Desertoniscus zhelochovtzevi Borutzky, 1945 Uz
51 Detonella papillicornis (Richardson, 1904) Ru: Kamchatka, Sakhalin regions
52 Halophiloscia couchii (Kinahan, 1858) Ru: Krasnodar region; Ua: Crimea
53 Haplophthalmus danicus Budde-Lund, 1880 Ru: Krasnodar, Rostov regions
54 Hemilepistoides messerianus Borutzky, 1945 Tu
55 Hemilepistus buddelundi Borutzky, 1945 Tu
56 Hemilepistus communis Borutzky, 1945 Td: Samgar massif
57 Hemilepistus crenulatus (Pallas, 1771) Td: Samgar massif; Tu: Central Karakum, Kyzyl-Arvat; Uz: Zakaspiysk, Fergana regions; Kg
58 Hemilepistus cristatus Budde-Lund, 1885 Tu: Kyzyl-Arvat
59 Hemilepistus elongatus Budde-Lund, 1885 Ru: Rostov, Stavropol regions; Tu: SW part
60 Hemilepistus fedtschenkoi (Uljanin, 1875) Kz: Semipalatinsk;Tu: Krasnovodsk; Uz: Bukhara, Samarkand, Syrdaryinsk, Zakaspyisk regions
61 Hemilepistus heptneri Borutzky, 1945 Tu
62 Hemilepistus klugii (Brandt, 1833) Az: Baku
63 Hemilepistus magnus Borutzky, 1945 Uz
64 Hemilepistus nodosus Budde-Lund, 1885 Tu; Kz
65 Hemilepistus pavlovskii Borutzky, 1954 Kz
66 Hemilepistus reductus Borutzky, 1945 Uz: Bukhara, Samarkand, Syrdaryinsk regions
67 Hemilepistus rhinoceros Borutzky, 1958 Kz
68 Hemilepistus ruderalis (Pallas, 1771) Ru: Volgograd region; Kz: Djanybek
69 Hemilepistus russonovae Borutzky, 1951 Az: Baku
70 Hemilepistus zachvatkini Verhoeff, 1930 Td: Samgar massif
71 Hyloniscus riparius C. Koch, 1838 Ru: Moscow, Penza, Pskov, Rostov, Tula regions; Ua: Kiev region
72 Leptotrichus panzerii (Audonin, 1826) Ua: Crimea
73 Leptotrichus tauricus Budde-Lund, 1885 Ua: Crimea
74 Ligia cinerascens Budde-Lund, 1885 Ru: Kurily islands
75 Ligia italica Fabricius, 1798 Ua: Crimea
76 Ligia pallasii Brandt, 1833 Kadakh?
77 Ligidium birsteini Borutzky, 1950 Ab: Gagry
78 Ligidium cavaticum Borutzky, 1950 Ru: Krasnodar region
79 Ligidium fragile Budde-Lund, 1885 Ab: Sukhum
80 Ligidium germanicum Verhoeff, 1901 Md
81 Ligidium hypnorum Cuvier, 1792 Ab: Sukhum; Bl: Belovezha National Park; Ru: Tver, Kaluga, Moscow regions; Ua: Crimea, Kiev
82 Ligidium margaritae Borutzky, 1955 Kz: Alma-Ata
83 Ligidium nodulosum Verhoeff, 1918 Ab: Gagry
84 Ligidium shadini Borutzky, 1948 Td
85 Ligidium tauricum Verhoeff, 1930 Ua: Crimea
86 Ligidium zaitzevi Borutzky, 1950 Ab: Sukhum
87 Ligidium zernovi Borutzky, 1948 Kg
88 Mingrelloniscus inchhuricus Borutzky, 1974 Ge: Megrelia
89 Nagurus matekini Borutzky, 1959 Kg
90 Oniscus asellus Linne, 1758 Lt: Vilnius; Ru: Pskov region; Ua: Kiev
91 Parcylisticus armenicus Borutzky, 1970 Ar: Daralagez
92 Parcylisticus dentifrons Budde-Lund, 1885 Az: Kutkashen; Ge: Manglisi; Ru: Astrakhan, Stavropol regions, Chechnya, Dagestan, Kabardino-Balkaria, N Osetia; Ua: Crimea
93 Parcylisticus georgianus Schmalfuss, 2003 Ge: Adigeni, Batumi, Kutaisi, Mestia
94 Parcylisticus golovatchi Schmalfuss, 2003 Az: Shikahokh
95 Parcylisticus mrovdaghicus (Borutzky, 1970) Az: Avash, Dashsalty, Kelbadjar, Kirovobad, Lenkoran, Zuvand
96 Parcylisticus urartuensis Borutzky, 1970 Ar
97 Parcylisticus zangezuricus Borutzky, 1970 Ar
98 Platyarthrus armenicus Borutzky, 1976 Ar: Megri
99 Platyarthrus hoffmannseggii Brandt, 1833 Ru: Krasnodar region
100 Platyarthrus luppovae Borutzky, 1953 Td
101 Platyarthrus mesasiaticus Borutzky, 1976 Tu
102 Platyarthrus ocellatus Borutzky, 1953 Td
103 Platyarthrus schoblii Budde-Lund, 1885 Ua: Crimea
104 Porcellio bistriatus Budde-Lund, 1885 Ab: Sukhum; Ru: Krasnodar region
105 Porcellio crassicornis C. Koch, 1841 Bl: Minsk
106 Porcellio dilatatus Brandt, 1833 Ar: Sevan
107 Porcellio laevis Latreille, 1804 Ab: Sukhum; Ru: Altay, Kalmykia, Moscow, Primorie, Rostov regions; Ua: S Crimea, Odessa; Uz: Bukhara region
108 Porcellio lamellatus Budde-Lund, 1885 Ua: Crimea
109 Porcellio obsoletus Budde-Lund, 1885 Ua: S Crimea
110 Porcellio scaber Latreille, 1804 Bl: Belovezha, Berezinsky reserves; Lt: Vilnius; Ru: Belgorod, Kaluga, Moscow, Nizhni Novgorod, Primorie, Rostov regions, Kamchatka, S Kuruly islands, Sakhalin; Ua: Kiev, Kremenetz, Vinnickaya region (Yampol’)
111 Porcellio spinicornis Say, 1818 Bl: Minsk; Lt: Vilnius; Md; Ru: Kaluga, Leningrad, Moscow Pskov regions; Ua: Kiev
112 Porcellio uljanini Budde-Lund, 1885 Ua: Crimea
113 Porcellio variabilis Lucas, 1849 Ua: Crimea
114 Porcellionides approximatus Budde-Lund, 1885 Md; Ru: Stavropol region; Ua: Crimea
115 Porcellionides linearis (Budde-Lund, 1885) Uz: Nukus
116 Porcellionides pruinosus Brandt, 1833 Ab: Sukhum; Ar: Shorzha; Az: Baku, Khachmas, Nabran; Ru: Baikal, Volgograd, Krasnodar region, Moscow, Rostov, Saratov regions; Ua: Crimea
117 Porcellionides rectifrons (Budde-Lund, 1885) Ua: Crimea
118 Porcellium collicola (Verhoeff, 1907) Md
119 Porcellium conspersum C. Koch, 1841 Bl: Belovezha Reserve; Ua
120 Protracheoniscus abricossovi Borutzky, 1945 Tu
121 Protracheoniscus alabashensis Borutzky, 1959 Kg
122 Protracheoniscus almaatinus Borutzky, 1975 Kz: Alma-Ata
123 Protracheoniscus anatolii Borutzky, 1959 Kg
124 Protracheoniscus armenicus Borutzky, 1975 Ge: Megri
125 Protracheoniscus asiaticus (Uljanin, 1875) Ru: Moscow, Nizhni Novgorod, Rostov, Ryazan’ regions; Td: Smagar massif
126 Protracheoniscus atrecicus Borutzky, 1945 Tu: Bugdaily
127 Protracheoniscus bugdajliensis Borutzky, 1975 Tu: Bugdaily
128 Protracheoniscus cristatus Borutzky, 1945 Az: Lenkoran, Sara isl.; Tu
129 Protracheoniscus darevskii Borutzky, 1975 Ar: Megri
130 Protracheoniscus delilensis Borutzky, 1945 Tu
131 Protracheoniscus desertorum Verhoeff, 1930 Turkestan?
132 Protracheoniscus digitifer Borutzky, 1945 Tu
133 Protracheoniscus fossuliger Verhoff, 1901 Ru: Krasnodar, Rostov regions
134 Protracheoniscus giljarovi Borutzky, 1957 Ru: Rostov region; Ua: Lugansk region
135 Protracheoniscus gissarensis Borutzky, 1975 Td: Dushanbe
136 Protracheoniscus hirsutulus Verhoff, 1930 Uz: Tashkent
137 Protracheoniscus kopetdagicus Borutzky, 1945 Tu
138 Protracheoniscus kryszanovskii Borutzky, 1957 Ru: Volgograd region, Kalmykia
139 Protracheoniscus latus (Uljanin, 1875) Td: Zeravshvan valley
140 Protracheoniscus litoralis (Budde-Lund, 1885) Ua: Crimea
141 Protracheoniscus major (Dollfus, 1903) Ru: Rostov region; Ua: Kiev
142 Protracheoniscus maracandicus (Uljanin, 1875) Td: Smagar massif; Uz: Bukhara, Samarkand, Syrdaryinsk regions
143 Protracheoniscus marginatus (Uljanin, 1875) Ua: Crimea
144 Protracheoniscus nogaicus Demianowitz, 1931 Md; Ru: Rostov region
145 Protracheoniscus orientalis (Uljanin, 1875) Az: Baku, Mardakert; Bl: Berezinsky reserve; Kz: Mangyshlak; Tu: Bugdaily; Ru: Moscow, Orenburg, Primorie regions; Ua: Kremenetz, Odessa regions, Uz: Nukus; Zakaspyisk region
146 Protracheoniscus panphilovi Borutzky, 1959 Kg
147 Protracheoniscus politus (C. Koch, 1841) Ru: Moscow region
148 Protracheoniscus scythicus Demianowicz, 1932 Md
149 Protracheoniscus steinbergi Borutzky, 1961 TU: SW part
150 Protracheoniscus taschkentensis Verhoeff, 1930 Uz: Tashkent; Td
151 Protracheoniscus tashausicus Borutzky, 1976 Tu; Ru: Rostov region
152 Protracheoniscus topczievi Borutzky, 1975 Ru: Krasnodar, Rostov regions; Ua: Zaporozhye region
153 Protracheoniscus tuberculatus (Borutzky, 1945) Tu
154 Protracheoniscus turcomanicus Borutzky, 1945 Tu
155 Protracheoniscus tzvetkovi Borutzky, 1975 Kz: Alma-Ata, Uzun-Agach; Ru: Moscow region?
156 Protracheoniscus uljanini Borutzky, 1953 Td
157 Protracheoniscus verhoeffi Strouhal, 1929 Ge: Tbilisi
158 Protracheoniscus zenkevitschi (Borutzky, 1945) Tu
159 Psachonethes czerkessicus Borutzky, 1969 Ru: Krasnodar region
160 Pseudobuddelundiella hostensis Borutzky, 1967 Ru: Krasnodar region
161 Pseudobuddelundiella ljovuschkini Borutzky, 1967 Ru: Krasnodar region
162 Schizidium davidi (Dollfus, 1887) Az: Divichi
163 Schizidium golovatchi Schmalfuss, 1988 Ar: Shikalyukh; Az: Baku; Ge: Batumi
164 Schizidium reinoehli Schmalfuss, 1988 Ru: Rostov region
165 Tadzhikoniscus coecus Borutzky, 1976 Td
166 Tauroligidium stygium Borutzky, 1950 Ua: Crimea
167 Tauronethes lebedinskyi Borutzky, 1949 Ua: Crimea
168 Titanethes albus (C. Koch, 1841) Ua: Crimea
169 Trachelipus azerbaidzhanus Schmalfuss, 1986 Az: E part
170 Trachelipus caucasius (Verhoeff, 1918) Ab: Gagry; Ru: Krasnodar region
171 Trachelipus difficilis Radu, 1950 Bl: Belovezha, Berezinsky reserves; Ua: S Crimea
172 Trachelipus ensiculorum Verhoeff, 1949 Ar: Yerevan
173 Trachelipus gagriensis (Verhoeff, 1918) Ab: Gagry
174 Trachelipus kervillei (Arcangeli, 1938) Ru: Rostov region
175 Trachelipus lignaui (Verhoeff, 1918) Ab: Gagry; Ru: Rostov region
176 Trachelipus longipennis Budde-Lund, 1885 Ab; Ua: S Crimea
177 Trachelipus lutschnikii (Verhoeff, 1933) Ru: Krasnodar region
178 Trachelipus rathkii Brandt, 1833 Ab: Sukhum; Az: Airidja; Ar: Chaldyr; Ge: Kutaisi; Bl: Belovezha pusha; Lt: Vilnius; Md; Ru: Belgorod, Kursk, Tver, Maryi-El, Kaluga, Mordovia, Moscow, Penza, Rostov, Leningrad, Saratov, Tula regions; Ua: Crimea, Kiev
179 Trachelipus razzautii (Arcangeli, 1913) Ru: Krasnodar region
180 Trachelipus sarculatus (Budde-Lund, 1896) Ua: Crimea
181 Trachelipus trachealis Budde-Lund, 1885 Md
182 Trichoniscus aphonicus Borutzky, 1977 Ab
183 Trichoniscus gudauticus Borutzky, 1977 Ab
184 Trichoniscus pusillus Brandt, 1833 Ua: Crimea, Kiev
185 Trichoniscus pygmaeus Sars, 1898 Ru: Krasnodar region
186 Turanoniscus anacanthotermitis Borutzky, 1969 Uz: Tashkent
187 Tylos granuliferus Budde-Lund, 1885 Ru: Primorie region, S Kuril Islands
188 Tylos ponticus Grebnicki, 1874 Ua: Crimea, Odessa
189 Typhloligidium coecum (Carl, 1904) Ua: Crimea
190 Typhloligidium karabijajlae Borutzky, 1962 Ua: Crimea