The genus Filipinolotis can be distinguished from other genera of the tribe Sticholotidini by the following characters: body rounded, glabrous, dorsum strongly convex (Figs 1a–c, 2a–c); antennae with 11 antennomeres (Figs 1e, 2e); hind wings atrophied; elytral epipleuron unevenly broadened; abdomen with five ventrites, abdominal postcoxal lines incomplete (Figs 1n, s, 2m).

Figure 1. 

Morphological characters of the genus Filipinolotis. a–t F. latefasciata Miyatake. a dorsal habitus b lateral habitus c frontal habitus d head, ventral e antenna f mandible g maxilla h labrum i labium j front leg k tarsal claws l prothorax, ventral m mesoventrite and metaventrite n male abdomen o penis p apex of penis q tegmen, lateral view r tegmen, ventral view s female abdomen t ovipositor.

Body rounded, dorsum strongly convex and glabrous (Figs 1a–c, 2a–c). Head smooth, with sparse short hairs (Figs 1c, 2c). Antennae with 11 antennomeres, scape and pedicel robust, pedicel shorter than scape; flagellum 9-segmented, gradually broadening towards apex, last-three antennomeres forming a fusiform club (Figs 1d–e, 2d–e).

Figure 2. 

Filipinolotis purpuratorotunda sp. n. a dorsal habitus b lateral habitus c frontal habitus d head, ventral e antenna f mandible g maxilla h labrum i labium j front leg k prothorax, ventral l mesoventrite and metaventrite m abdomen n penis o apex of penis p tegmen, ventral view q tegmen, lateral view.

Clypeus protruded with anterior margin slightly emarginate at middle (Figs 1c, 2c). Labrum transverse, narrowly rounded and covered with dense, long setae (Figs 1h, 2h). Mandible with two apical teeth, sharp and smooth without additional denticles (Figs 1d, f, 2d, f). Maxillary palp 4-segmented with terminal segment cut obliquely at apex (Figs 1d, g, 2d, g). Labial palpomere 3-segmented, with robust 2^ndsegment and slender terminal segment, ligula membranous and plump, insertion of labial palps is anterior on the prementum (Figs 1i, 2i).

Pronotum transverse, strongly convex, emarginate at anterior margin, hind margin bent into a curved shape; lateral margin slightly oblique, with fine groove continuing towards basal corner from anterior corner, anterior angles rounded and smooth, hind angles obtuse and smooth (Figs 1a–c, 2 a–c). Scutellum minute, triangular (Figs 1a, c; 2a, c). Elytra without humeral angles, strongly convex, almost as wide at anterior margin as hind margin of pronotum, lateral margins extremely narrow; lateral margins with narrow rim, invisible from above (Figs 1a–c, 2a–c). Hind wings atrophied.

Prosternum T-shaped. Prosternal process narrowed, less than 0.6 times width of coxal diameter and not prominent anteriorly between coxae, prosternal carinae present and fan-shaped (Figs 1l, 2k). Mesoventrite trapezoid-shaped, coarsely keeled on the anterior edge, metaventrite as long as mesoventrite (Figs 1m, 2l). Elytral epipleuron wide and uneven, gradually narrowed to apex. Legs with pubescence rather dense, coxae square-shaped, obvious robust; trochanter triangular, robust; femora thick, deeply grooved beneath for tibiae when retracted; length of tibia subequal to femur, slender, and widening but not angulate outwardly; tarsi 4-segmented; claws simple and slender, without teeth (Figs 1j, k, 2j). Abdomen with five ventrites in both sexes, ventrite 1 length measured below coxal cavity, almost as long as ventrites 2^nd to 4^th combined, abdominal postcoxal lines incomplete, not recurved but meeting hind margin of ventrite 1 (Figs 1n, s, 2m).

Male genitalia: tegmen stout, basal piece with distinct strut and additional, dorsal strut-like projection, parameres well developed and setose apically. Penis uniformly tubular, curved (Figs 1o–r, 2n–q).

Female genitalia: coxites long triangular with interior margin slightly emarginate, setose apically; styli conspicuous (Figure 1t).

Philippines.

This species can be recognized by the following combination of characters: pronotum reddish yellow, elytra blackish shiny with a broad transverse reddish yellow band in the middle (Figure 1a–c); penis guide of tegmen in ventral view broad and flat, with a rounded apex, parameres narrow and almost straight with dense setae at apices (Figure 1q–r); penis uniformly tubular, curved, gradually narrowing to apex (Figure 1o–r).

TL: 1.92–2.3 mm, TW: 1.86 mm, TH: 1.35–1.38 mm, TL/TW: 1.03–1.24; PL/PW: 0.38–0.42; EL/EW: 0.89–1.08; HW/TW: 0.39; PW/TW:0.67–0.74.

Head reddish yellow (Figure 1c, d). Pronotum reddish yellow (Figure 1a–c). Scutellum blackish shiny (Figure 1a, c). Elytra blackish and shiny, with a broad transverse reddish yellow band at middle, covering almost half of elytra (Figure 1a–c). Underside blackish brown, except claws yellowish.

Body rounded, dorsum strongly convex and glabrous (Figure 1a–c). Head large, with sparse short golden pubescence (Figure 1d). Pronotum glabrous with uniformly minute and dense punctures (Figure 1a–c). Elytra convex, entire surface uniformly punctate (Figure 1a–c). Elytral epipleuron with sparse golden pubescence. Prosternum with sparse golden pubescence and inconspicuous punctures (Figure 1l). Mesoventrite and metaventrite with sparse golden pubescence and inconspicuous punctures, mesoventrite with narrow intercoxal process, chin piece formed on the anterior margin of mesoventrite; metaventrite short, as wide as mesoventrite (Figure 1m). Legs with dense golden pubescence (Figure 1j, k). Abdomen with sparse golden hairs and uniform punctures (Figure 1n, s).

Male genitalia (Figure 1o–r). Tegmen stout and symmetrical, penis guide in lateral view wide at base and gradually narrowing to pointed apex, the basal 2/3 with membranous prominence; in ventral view broad and flat, rounded apically; parameres narrow and almost straight with dense setae at apices (Figure 1q–r), phallobase membranous. Penis simple tubular, curved, gradually narrowing along apical third to pointed apex, penis capsule membranous, without inner arm (Figure 1o–p).

Philippines: 1 male, Mt. Polis, Lepanto, Luzon Prov., Böttcherleg (MNB); 1 female, Mt. Data, Mountain, Luzon Prov., 7500 ft, 23.iv.1946; 1 male Philippines, Luzon (ANIC).

Philippines (Luzon).

This species is similar to F. latefasciata Miyatake but it can be distinguished from the latter as follows: body uniformly black with purple shine, without spots (Figure 2a–c); penis guide of tegmen in ventral view narrow with a nipple-shaped apex (Figure 2p, q); apex of the penis truncate, membranous (Figure 2o). In F. latefasciata, pronotum reddish yellow, elytra blackish and shiny, with a broad transverse reddish yellow band in the middle (Figure 1a–c); penis guide of tegmen in ventral view broad and flat, with rounded apex (Figure 1r); apex of the penis pointed (Figure 1o, p).

TL: 1.99–2.63 mm, TW: 1.84–2.55 mm, TH: 1.28–1.8 mm, TL/TW: 1.03–1.08; PL/PW: 0.40–0.49; EL/EW: 0.93; HW/TW: 0.34–0.39; PW/TW: 0.59–0.60.

Color: Head, pronotum, scutellum, elytra purplish shiny black, without spots (Figure 2a–c). Underside blackish brown, except yellowish claws.

Body rounded, dorsum strongly convex and glabrous (Figure 2a–c). Head large with scattered short golden pubescence (Figure 2c, d). Pronotum glabrous, with uniformly distributed punctures (Figure 2a–c). Scutellum of isosceles triangle shape (Figure 2a, c). Elytral surface uniformly punctate (Figure 2a–c). Prosternum, mesoventrite, and metaventrite with scattered golden pubescence and inconspicuous punctures (Figure 2k, l). Elytral epipleuron with uniform golden pubescence. Legs with dense golden pubescence (Figure 2j, l). Abdomen with sparse golden pubescence and uniformly distributed punctures (Figure 2m).

Male genitalia. Tegemen stout and symmetrical, tegminal strut stout and straight; penis guide in lateral view almost straight and gradually narrowing along apical third to pointed apex, with membranous prominence at basal half; in ventral view narrowest at base and gradually broadening to apex, then strongly narrowed to form a nipple-shaped apex; parameres with dense long setae apically, longer than penis guide; phallobase membranous (Figure 2p, q). Penis uniformly tubular, curved, basal capsule membranous, without distinct arms; apex of penis subtruncate with membranous appendage (Figure 2n, o).

Holotype: Philippines: 1 male, La Trinidad, Luzon Prov., 1300m, 4–5.IV.1968, Benqae (ANIC); Paratypes: Philippines: 1 male, Island of Basilan, Baker (ANIC); 1 male, Disimungal, Madela, Quirino, Eastern Luzon, XII.2014 (SCAU).

Philippines (Luzon).

The name purpuratorotunda is composed of the word purpuratus, which refers to the purplish color of the body and rotundus, referring to the rounded body shape.