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The type status is described of 57 taxa from the superfamily Orthalicoidea in the collection of the Brussels museum. Two new species are described: Stenostylus perturbatus sp. n., and Suniellus adriani sp. n. New lectotypes are designated for Bulimulus (Naesiotus) amastroides Ancey, 1887; Bulimulus blanfordianus Ancey, 1903; Bulimulus montivagus chacoensis Ancey, 1897; Bulimus coloratus Nyst, 1845; Plecochilus dalmasi Dautzenberg, 1900; Placostylus porphyrostomus elata Dautzenberg, 1923; Bulimulus ephippium Ancey, 1904; Bulimus fulminans Nyst, 1843; Bulimus funckii Nyst, 1843; Orphnus thompsoni lutea Cousin, 1887; Bulimus melanocheilus Nyst, 1845; Orphnus thompsoni nigricans Cousin, 1887; Orphnus thompsoni olivacea Cousin, 1887; Bulimulus pollonerae Ancey, 1897; Orphnus thompsoni zebra Cousin, 1887. New combinations are: Bostryx borellii (Ancey, 1897); Bostryx carandaitiensis (Preston, 1907); Protoglyptus mazei (Crosse, 1874); Kuschelenia (Vermiculatus) sanborni (Haas, 1947). New synonymies are established for the following nominal taxa: Orphnus thompsoni var. lutea Cousin, 1887 = Kara thompsonii (Pfeiffer, 1845); Orphnus thompsoni var. nigricans Cousin, 1887 = Kara thompsonii (Pfeiffer, 1845); Thaumastus nystianus var. nigricans Cousin, 1887 = Drymaeus (Drymaeus) nystianus (Pfeiffer, 1853); Orphnus thompsoni var. olivacea Cousin, 1887 = Kara thompsonii (Pfeiffer, 1845); Orphnus thompsoni var. zebra Cousin, 1887 = Kara thompsonii (Pfeiffer, 1845).
Bolivia, Amphibulimidae, Bulimulidae, Bothriembryontidae, Megaspiridae, Orthalicidae, Simpulopsidae, types, biohistory
The Orthalicoidea is a dominant faunal element in the Neotropics (
One of the interesting parts of the RBINS malacological
collection is the former collection of Philippe Dautzenberg (1849–1935;
see
Excerpts of autographs from the Dautzenberg archive. A C.F. Ancey. B H. Crosse. C H.C. Fulton.
Two other authors need mentioning, who were not referred to by
A–B Excerpts of autographs from the Dautzenberg archive. A A. Morelet. B H.B. Preston. C Cover of the inventory of the collection of A. Cousin, in Dautzenberg’s handwriting.
The second author is Pierre-Henri Nyst, a Belgian
paleontologist who in his youth described several species of non-marine
molluscs from South America, supplied by Belgian explorers (
The following criteria were applied to assess potential type material: (a) the locality fits with the original description (taking into account changes in geographical names due to political or administrative reasons); (b) alleged type material is in accordance with the established understanding of the taxon. In order to fulfill the requirements of article 74 of the International Code of Zoological Nomenclature (ICZN), any lectotype designations herein are to be understood as to have the sole purpose of fixing the status of these specimens as the sole name-bearing types of the nominal taxa, to ensure the name’s proper and consistent application, even if this is not explicitly done in every single case but abbreviated as “lectotype designation”. Lectotypes are designated herein using the following criteria, in order of preference: (1) the relevant specimen was figured in the original description, or in subsequent revisionary works; (2) if no original figure was published, a specimen was selected that matches as closely as possible the measurements given in the original description.
For each taxon, the original publication—in which the
taxon was proposed—is mentioned, as well as papers in which reference
is made to the type material. The type locality is quoted from the
original publication in the original wording and language, with
clarifying notes between square brackets. The name of the collector, if
given in the original paper, is only mentioned (in italics) if it
might give a clue about the type status of material present in the
collection. The text of the original, or oldest, label is quoted,
together with information from subsequent labels if containing
information necessary for a correct interpretation. All labels have been
photographed and are figured for future historic reference. The
dimensions of the type are quoted, as given in the original paper.
Dimensions of the type specimens have been taken with a digital caliper,
using the methods figured by
Abbreviations used for depositories of material are: ANSP, Academy of Natural Sciences, Philadelphia, U.S.A.; FMNH, Field Museum of Natural History, Chicago, U.S.A.; MHNG, Muséum d’Histoire Naturelle, Genève, Switzerland; MNHN, Museum national d’Histoire naturelle, Paris, France; NHM, Natural History Museum, London, U.K.; RMNH, Nederlands Centrum voor Biodiversiteit Naturalis (formerly Rijksmuseum van Natuurlijke Historie), Leiden, Netherlands; SMF, Natur-Museum Senckenberg, Frankfurt am Main, Germany; UF, Florida State Museum, Gainesville, U.S.A.; ZMB, Zoologisches Museum, Humboldt Universität, Berlin, Germany. Other abbreviations used are: /, end of line in cited text; coll., collection; D, shell diameter; H, shell height; leg., legit, collected; MT, type collection of RBINS Malacology Section; W, number of whorls.
Systematics Systematic list of nominal taxa arranged in generic orderThis systematic list follows the arrangement of families as proposed by
At the family level the new scheme of
Family Amphibulimidae P. Fischer, 1873
Dryptus Albers, 1860
Dryptus funckii Nyst, 1843
Plekocheilus (Eurytus) Albers, 1850
Plekocheilus (Eurytus) coloratus Nyst, 1845; Plekocheilus (Eurytus) dalmasi Dautzenberg, 1900
Plekocheilus (Plekocheilus) Guilding, 1828
Plekocheilus (Plekocheilus) fulminans Nyst, 1845
Family Orthalicidae Martens, 1860
Kara Strebel, 1910
Kara lutea Cousin, 1887; Kara nigricans Cousin, 1887; Kara olivacea Cousin, 1887; Kara zebra Cousin, 1887
Family Megaspiridae Pilsbry, 1904
Thaumastus Albers, 1860
Thaumastus blanfordianus Ancey, 1903
Family Bothriembryontidae Iredale, 1937
Aspastus Albers, 1850
Aspastus porphyrochila Dautzenberg and Bernier, 1901
Placostylus Beck, 1837
Placostylus auriculatus Dautzenberg and Bouge, 1923; Placostylus elatus Dautzenberg, 1923; Placostylus paraspiritus A.W.B. Powell, 1951; Placostylus whareana A.W.B. Powell, 1951
Family Simpulopsidae Schileyko, 1999
Rhinus Albers, 1860
Rhinus argentinus Ancey, 1901
Simpulopsis (Eudioptus) Albers, 1860
Simpulopsis (Eudioptus) ephippium Ancey, 1904
Family Bulimulidae Tryon, 1867
Bostryx Troschel, 1847
Bostryx alausiensis Cousin, 1887; Bostryx albicolor Morelet, 1863; Bostryx baeri Dautzenberg, 1901; Bostryx bonneti Ancey, 1902; Bostryx borellii Ancey, 1887; Bostryx carandaitiensis Preston, 1907; Bostryx emaciatus Morelet, 1863; Bostryx huayaboensis Dautzenberg, 1901; Bostryx iocosensis Dautzenberg, 1901; Bostryx juana Cousin, 1887; Bostryx metagyra Pilsbry and Olsson, 1949; Bostryx moniezi Dautzenberg, 1901; Bostryx perforatus Haas, 1951; Bostryx spiculatus Morelet, 1860; Bostryx veruculum Morelet, 1860
Drymaeus (Drymaeus) Albers, 1850
Drymaeus (Drymaeus) abruptus Rolle, 1904; Drymaeus (Drymaeus) colimensis Rolle, 1895; Drymaeus (Drymaeus) icterica Ancey, 1892; Drymaeus (Drymaeus) jousseaumei Dautzenberg, 1901; Drymaeus (Drymaeus) nigricans Cousin, 1887; Drymaeus (Drymaeus) scolioides Dautzenberg, 1901; Drymaeus (Drymaeus) solidus Preston, 1907
Drymaeus (Mesembrinus) Albers, 1850
Drymaeus (Mesembrinus) interruptus Preston, 1909; Drymaeus (Mesembrinus) pallidus Preston, 1909
Naesiotus Albers, 1850
Naesiotus albemarlensis Dall, 1917; Naesiotus amastroides Ancey, 1887; Naesiotus bizonalis Ancey, 1887; Naesiotus chacoensis Ancey, 1897; Naesiotus cucullinus Dall, 1917; Naesiotus duncanus Dall, 1893; Naesiotus gilderoyi Van Mol, 1972; Naesiotus lycodus Dall, 1917; Naesiotus pollonerae Ancey, 1897
Protoglyptus Pilsbry, 1897
Protoglyptus dejectus Fulton, 1901; Protoglyptus mazei Crosse, 1874
Rabdotus Albers, 1850
Rabdotus hesperius Pilsbry and Ferriss, 1924
Kuschelenia (Vermiculatus) Breure, 1978
Kuschelenia (Vermiculatus) sanborni Haas, 1947
Alphabetic list of taxa by species namehttp://species-id.net/wiki/Bulimulus_(Drymaeus)_abruptus
Figs 12A, 12i“Huancabamba in Peru”; see remarks.
“Huancabamba, / Peru 1904”; in Rolle’s handwriting. Another label, in Dautzenberg’s handwriting, indicating that he obtained it on 16.vii.1907 from Rolle.
“Alt. 44, diam. max. 24 (..) mm”; figured specimen H 36.6, D 16.9, W 6.5.
RBINS/MT2332, two syntypes (Dautzenberg coll.).
Rolle did not state on how many specimens his
description was based upon. However, there is another specimen in the
NHM collection which will be designated lectotype (Breure and Ablett,
unpublished data). The locality is ambiguous, as there are several
places called “Huancabamba” in Peru. There were, however, several
other species described by Rolle from the same locality, among them Columbinia huancabambensis, which is regarded by
Bulimulidae, Drymaeus (Drymaeus) abruptus (Rolle, 1904).
http://species-id.net/wiki/Thaumastus_alausiensis
Figs 10A–B, 10i[Ecuador] “sur le versant du mont Hacu, entre Achapallas et la rivière Sula, sur le territoire Alausi, province de Chimborazo”.
“Racu, descente de Achupalla / au Rio Sula”, in Cousin’s handwriting.
“long. 25 mm; diam. 9 à 10 mm”; figured specimen H 25.2, D 11.0, W 8.2.
RBINS/MT2333, paralectotype; MT2334, 12 paralectotypes (partly juvenile), Cousin leg. (Dautzenberg coll.).
The material which Breure (1975) used to select a lectotype, came from the Jousseaume collection and originated from Cousin. However, the locality was imprecise (“Équateur” [Ecuador]). At that time, no other material from Cousin was known. In RBINS, material originating from Cousin and exactly corresponding to the type locality has been found. Therefore it may now be questioned if the Jousseaume material was validly regarded as the prime type material. However, in the inventory of Cousin’s collection, a total of 30 specimens is mentioned and it cannot be excluded that Jousseaume’s material originated from the Cousin collection.
Bulimulidae, Bostryx alausiensis (Cousin, 1887).
http://species-id.net/wiki/Bulimulus_(Naesiotus)_albemarlensis
[Ecuador, Galápagos, Isla Isabela] “near [Puerto] Villamil at 2300 to 3300 feet elevation”.
“Near Villamil / Albemarle Isl., Galapagos” on label of Stanford University.
“Length of shell 15, (...) diameter 9 mm”; largest specimen H 13.6, D 9.0, W 5.6.
RBINS/MT1911, two paratypes, Ochsner leg.
The material was donated by H.G. Schenk. For data
on the role of Schenck and the relationship between Stanford University
and RBINS, see
Bulimulidae, Naesiotus albemarlensis (Dall, 1917).
http://species-id.net/wiki/Bulimus_albicolor
Figs 11I, 11ix[Peru, Dept. Ayacucho] “Huanta et de la vallée de l’Apurimac”.
“Pérou”; taxon name in Morelet’s handwriting, locality data in Dautzenberg’s handwriting (“ex auctore”).
“Longit 28, diam. 9 mm”; figured specimen H 22.2, D 9.76, W 5+.
RBINS/MT2335, one syntype, ex Morelet (Dautzenberg coll.).
Additional syntype material is present in the MHNG and NHM collections.
Bulimulidae, Bostryx orophilus (Morelet, 1860).
http://species-id.net/wiki/Bulimulus_(Naesiotus)_amastroides
Figs 14B, 14vi[Ecuador] “Îles Galapagos”.
“I. Galapagos”, in Ancey’s handwriting.
“Long., 9 1/2; diam., 4 2/3 mill.”; figured specimen H 9.25, D 4.5, W 7.0.
RBINS/MT1866, lectotype (design. n.), ex Géret ex Ancey (Dautzenberg coll.).
Ancey writes that this species may only be compared to Naesiotus calvus (Sowerby, 1833), which was described from Isla San Salvador. However,
Bulimulidae, Naesiotus amastroides (Ancey, 1887).
http://species-id.net/wiki/Bulimulus_(Rhinus)_argentinus
Figs 16A–B, 16i“Gualeguaychu, province d’Entrerios, République Argentine”.
“Gualeguaychu / Prov. d’Entrerios / (Rép. Argentine)”, marked “types” in Ancey’s handwriting.
“Long. 19–21, diam. 12 1/2–14 mill.”; figured specimen H 19.7, D 13.1, W 6.3.
RBINS/MT1867, three syntypes, ex Géret ex Ancey (Dautzenberg coll.).
Simpulopsidae, Rhinus argentinus (Ancey, 1901).
http://species-id.net/wiki/Placostylus_caledonicus_auriculata
Figs 9B, 9ii[New Caledonia] “Forêt du Mt. Ignambi, au dessus d’Oubatche 500 m. altit.”.
“N. Cal.” in Dautzenberg’s handwriting.
Not given. Figured specimen H 81.7, D 42.0, W 5.7.
RBINS/MT2339. One possible syntype (Dautzenberg coll.).
The specimen has a label glued onto the dorsal side “Bulimus / P[seu?]do-Caledonicus”.
Bothriembryontidae, Placostylus caledonicus (Petit, 1845).
http://species-id.net/wiki/Peronaeus_baeri
Figs 11C, 11iii“Iocos (Peruvia) Baer legit. 1900”; see remarks.
“Iocos Pérou / Baer legit.”, in Dautzenberg’s handwriting.
“Altit. 14 1/2, latit. 3 1/2 millim.”; figured specimen H 13.6, D 3.1, W 10.1.
RBINS/MT2336, three paralectotypes, Baer leg. (Dautzenberg coll.).
Bulimulidae, Bostryx baeri (Dautzenberg, 1901).
http://species-id.net/wiki/Bulimulus_eschariferus_bizonalis
Figs 14C, 14v[Ecuador] “îles Galapagos”.
“I Galapagos”, also stating “type de var. bizonalis Anc.”, in Ancey’s handwriting.
Not given. Figured specimen H 16.0, D 6.3, W 7.5.
RBINS/MT2337, one syntype, ex Géret ex Ancey ex Deshayes (Dautzenberg coll.).
Ancey did not state on how many specimens his description was based. Therefore, the specimen is considered a syntype.
Bulimulidae, Naesiotus eschariferus (Sowerby, 1833).
http://species-id.net/wiki/Bulimulus_blanfordianus
Figs 4C–D, 4ii“Iquico, Bolivia, 3500 m. above the sea (fide Fulton)”; see remarks.
“Iquico / Bolivia / 3500 mtr.”, in Fulton’s handwriting. Taxon label in Dautzenberg’s handwriting.
“Long. 55, lat. 25 1/2 mill.”; figured specimen H 52.5, D 25.1, W 6.1.
RBINS/MT1865, lectotype (design. n.), ex Géret ex Ancey (Dautzenberg coll.).
Ancey did not state on how many specimens his description was based. Despite the fact that the specimen is slightly smaller than published by Ancey, there is enough evidence to consider this shell as from the original type series. It is now designated lectotype.
The type locality probably refers to Dept. La Paz, Ikiko [16°34'S, 67°44'W], where elevations around 3500 m are found.
Megaspiridae, Thaumastus (Thaumastus) blanfordianus (Ancey, 1903).
http://species-id.net/wiki/Bulimulus_bonneti
Figs 11A, 11i“Bolivie (teste A. Bonnet)”.
“Bolivie”; see remarks.
“Long. 23 1/2, lat. 13 mill.”; figured specimen H 21.9, D 14.7, W 5+.
RBINS/MT2338, one paralectotype, ex Géret ex Ancey (Dautzenberg coll.).
Bulimulidae, Bostryx bonneti (Ancey, 1902).
http://species-id.net/wiki/Bulimulus_borellii
Figs 10F–G, 10iv[Bolivia, Dept. Tarija] “Mission de San Francisco, sur le haut-Pilcomayo en Bolivie”.
“Mission / de San Francisco / Haut-Pilcomayo, Bolivie”, in Ancey’s handwriting.
[Measurements of three specimens given] “a) Long 34, lat. 11 mill.—b) Long 31, lat. 9 mill.—c) Long. 32, lat. 10 1/2 mill.”; largest figured specimen H 28.8, D 10.3, W 8.8.
RBINS/MT2340, two syntypes, ex Géret ex Ancey, Borelli leg. (Dautzenberg coll.).
The specimens are smaller than the measurements given by Ancey, although they are from the type locality. This taxon was hitherto classified as Drymaeus. However, the sculpture of the protoconch shows very fine spiral lines and dispersed granules, more or less axially arranged. It thus belongs to Bostryx (s.l.).
Bulimulidae, Bostryx borellii (Ancey, 1897) (comb. n.).
http://species-id.net/wiki/Bulimulus_(Drymaeus)_carandaitiensis
Figs 10H, 10v[Bolivia, Dept. Chuquisaca] “Carandaiti, province of Cordillera, Bolivia, 1000 metres”.
“Carandaiti / Prov. of Cordillera 1000 metres / Bolivia (co-type)”, in Preston’s handwriting.
“Alt. 35, diam. maj. 14 mill.”; figured specimen H 32.1, D 12.9, W 9.3.
RBINS/MT2341, one syntype, ex Preston (Dautzenberg coll.).
This taxon was hitherto regarded a Drymaeus species, but the protoconch is smooth (not worn) in both specimens examined. It is here tentatively placed in Bostryx, despite the fact that the colour pattern of the specimen in RBINS is unlike the ones normally found in this genus.
Bulimulidae, Bostryx carandaitiensis (Preston, 1907) (comb. n.).
http://species-id.net/wiki/Bulimulus_montivagus_chacoensis
Figs 14D–E, 14i[Bolivia, Dept. Tarija] “Caiza, Gran Chaco (Bolivie)”.
“Caiza, Chaco de Bolivie”, in Ancey’s handwriting.
[Measurements of three specimens given] “a) Long. 22, lat. 7 1/2 mill—b) Long. 22, lat. 8 mill.—c) Long. 18, lat. 7 1/2 mill.”; figured specimen H 20.9, D 7.7, W 8.5.
RBINS/MT2342, lectotype (design. n.), ex Géret ex Ancey, Borelli leg. (Dautzenberg coll.).
The RBINS material corresponds with the original
measurements and is here designated lectotype. The shell shape,
especially the flaring basal lip, and the number of whorls (8+) are not
typical for Naesiotus, and this species is only tentatively placed here.
Bulimulidae, Naesiotus montivagus (d’Orbigny, 1835).
http://species-id.net/wiki/Bulimulus_(Drymaeus)_chacoensis
Figs 10E, 10iii“To the north of the Rio Pilcomayo, Chaco, Bolivia”.
“N of riv. Pilcomayo / Chaco Bolivie 600 m / alt. (co-type)”; label in Dautzenberg’s handwriting, see remarks.
“Alt. 30, diam. maj. 9.5 mm”; figured specimen H 30.1, D 10.1, W 8.1.
RBINS/MT2343, one syntype, ex Preston (Dautzenberg coll.).
The specimen is not accompanied by an original
Preston label. As Dautzenberg always accurately documented on his labels
the source and date of his acquisition (in this case “Preston
14.xi.07”), there is hardly any doubt that this is an original type
specimen. The protoconch is smooth, confirming that this taxon should
be classified within Bostryx (cf.
Bulimulidae, Bostryx chacoensis (Preston, 1907).
http://species-id.net/wiki/Otostomus_colimensis
Figs 12C, 12iii[Mexico] “Colima”.
“Colima Mexico”; label in Dautzenberg’s handwriting, see remarks.
“Alt. 31, diam. 15 mm.”; figured specimen H 29.1, D 14.1, W 6.3.
RBINS/MT2344, two paralectotypes, ex Rolle (Dautzenberg coll.).
This material is not accompanied by an original
Rolle label, but Dautzenberg documented that he received the shells
from him on 16.vii.1907. The fact that Rolle was a dealer may account
for the delay between the time of publication and the acquisitions by
Dautzenberg. The material is from the type locality and is here
considered as syntypes.
Bulimulidae, Drymaeus (Drymaeus) colimensis (Rolle, 1904).
http://species-id.net/wiki/Bulimus_coloratus
Figs 3A–B, 3i“la province de Cumana, dans la Colombie [sic, Venezuela]”.
“Colombie / Cumana”, indicating “Type (Nyst)” in Nyst’s handwriting.
“49 millimètres de longueur sur 30 de largeur [H 49 D 30]”; lectotype H 47.3, D 29.2, W 4.7.
RBINS/MT2345, lectotype (design. n.); MT2346, paralectotype, ex Nyst.
The type material was found in the RBINS collection and is now figured for the first time since the original publication. Of the two specimens present, one shows the ‘yellow shadow’ which is characteristic for this species, and is here designated lectotype. The taxon is only known from confirmed localities in northern Colombia; the original locality, which is in Venezuela, Edo. Sucre, seems erroneous. The type locality is now restricted to Sierra Nevada de Santa Marta.
Amphibulimidae, Plekocheilus (Eurytus) coloratus (Nyst, 1845).
http://species-id.net/wiki/Bulimulus_(Naesiotus)_cucullinus
Figs 14A, 14ii[Ecuador, Galápagos, Isla Española] “Hood Island, between 200 and 600 feet [61–183 m]”.
“Hood Is. 380 ft. / under stones, Galapagos” on a label of Stanford University.
“Length of shell 19, diameter 9.5 mm.”; figured specimen H 17.7, D 8.6, W 6.8.
RBINS/MT1833, two paratypes, Ochsner leg.
The material was donated by H.G. Schenk.
Bulimulidae, Naesiotus cucullinus (Dall, 1917).
http://species-id.net/wiki/Plecochilus_dalmasi
Figs 3C–E, 3iiNot given.
[Colombia] “Sierra de Santa Marta”, in Dautzenberg’s handwriting.
“Longit.: 26 millim., latit.: 16 millim.”; H 26.3, D 15.3, W 4.0.
RBINS/MT668, lectotype (design. n.); MT2347, two paralectotypes (Dautzenberg coll.).
Dautzenberg did not state on how many specimens
his description was based, but one of the specimens corresponds to the
original dimensions and agrees with his figure. This shell is here
designated lectotype. The label states that it was collected on
15.iii.1896, which corresponds with the date of excursion to Sierra de
Santa Marta mentioned in the station list (
Amphibulimidae, Plekocheilus (Eurytus) dalmasi (Dautzenberg, 1900).
http://species-id.net/wiki/Bulimulus_(Protoglyptus)_dejectus
Figs 15B, 15iii[Brazil] “Santa Catharina (fide Linnaea Institute label)”.
“St. Catharina”; see remarks.
“Maj. diam. 10, alt. 29 mm.”; figured specimen H 28.0, D 10.0, W 7.6.
RBINS/MT2348, one syntype, ex Sowerby and Fulton (Dautzenberg coll.).
During a recent visit to the NHM, another
specimen was found which will be designated lectotype (Breure and
Ablett, unpublished data). The Brussels specimen thus will become a
paralectotype. Although the specimen is not accompanied by an original
Fulton label, Dautzenberg has noted on his label that he purchased this
specimen from Sowerby and Fulton on 27.ii.1907. The generic placement
of this taxon is somewhat puzzling. Three taxa may be considered, viz. Protoglyptus, Naesiotus, and Rhinus.
The protoconch sculpture consists of axial wrinkles, partly broken
into granules, which is a sculpture not characteristic for Protoglyptus nor Naesiotus. However, there is considerable variation in protoconch sculpture within these two taxa (
Bulimulidae, Protoglyptus dejectus (Fulton, 1907).
http://species-id.net/wiki/Placostylus_porphyrostomus_elata
Figs 9A, 9i“Nouvelle-Calédonie, St. Vincent (Coll. D. ex Rossiter)”.
“St. Vincent”, in Dautzenberg’s handwriting.
“hauteur 88 millim. Diam. max. 36 millim.”; lectotype H 86.3, D 37.8, W 7.3.
RBINS/MT702, lectotype (design. n.), Rossiter leg. (Dautzenberg coll.).
The specimen was marked by Dautzenberg as “type”.
As he did not state how many specimens he had seen, the shell is here
designated lectotype.
Bothriembryontidae, Placostylus porphyrostomus (Pfeiffer, 1851).
http://species-id.net/wiki/Bulimus_emaciatus
Figs 11B, 11ii[Peru] “dans les vallées et sur les plateaux de l’interieur de la Sierra, depuis Ayacucho jusqu’au Cuzco”.
“Pérou”. Locality and taxon label in Morelet’s handwriting.
“Longit. 22; diam. 5 1/2 mill.”; figured specimen H 18.5, D 5.4, W 9.2.
RBINS/MT2349, two syntypes, ex Morelet (Dautzenberg coll.).
Dautzenberg documented that these specimens
originate from the Morelet collection. It is not clear if Dautzenberg
had personal information from Morelet, when he added “Ayacucho” to his
own label. Since a lectotype was designated by
Bulimulidae, Bostryx emaciatus (Morelet, 1863).
http://species-id.net/wiki/_Bulimulus_ephippium
Figs 16C, 16ii“Bahia, Brazil (teste H. Fulton)”.
“Bahia” in Fulton’s handwriting; taxon name in Ancey’s handwriting.
“Longit. 20, diam. 12 mill.”; lectotype H 19.5, D 12.5, W 5.5.
RBINS/MT2350, lectotype (design. n.), ex Géret ex Ancey (Dautzenberg coll.).
Simpulopsidae, Simpulopsis (Eudioptus) ephippium (Ancey, 1904).
http://species-id.net/wiki/Bulimus_fulminans
Figs 6A–C, 6i“la Colombie [sic, Venezuela], dans la province de Cumana [Edo. Sucre]”.
“Venezula (sic) / Cumana”, in Nyst’s handwriting.
“60 millimètres de longeur sur 28 de largeur”; lectotype H 59.2, D 32.4, W 4.8.
RBINS/MT2351, lectotype (design. n.), ex Nyst.
The specimen corresponds to the original measurements given by Nyst, whose label is dated 1874 and indicated “type”. As there is no evidence that the description of Nyst was based on one specimen, it is here designated lectotype.
Amphibulimidae, Plekocheilus (Plekocheilus) fulminans (Nyst, 1843).
http://species-id.net/wiki/Bulimus_funckii
Figs 5A–B, 5i“la Colombie [sic, Venezuela], dans la province de Cumana [Edo. Sucre]”.
“Venezuela / Cumana”, in Nyst’s handwriting.
“90 millimètres de longeur sur 40 de largeur ”; lectotype H 86.3, D 44.5, W 5.5.
RBINS/MT2352, lectotype (design. n.), ex Nyst.
Amphibulimidae, Dryptus funckii (Nyst, 1843).
http://species-id.net/wiki/Bulimulus_gilderoyi
Figs 14G, 14iii[Ecuador] “Galápagos, Santa Cruz, à proximité du Cerro Coralon”.
“Galapagos, Santa Cruz, (...) Cerro Coralon”, in Van Mol’s handwriting.
“Hauteur 25.3 Largeur max. 15.5 (mm)”; holotype H 25.4, D 15.6 W 6.6.
RBINS/MT106, holotype; MT107, one paratype; MT108, 19 paratypes, all Van Mol leg., 27.x.1970.
This taxon was published on 10.ix.1972; A.G. Smith published a paper in which he described the same species as Naesiotus cavagnaroi on 21.i.1972.
Bulimulidae, Naesiotus cavagnaroi A.G. Smith, 1972.
http://species-id.net/wiki/Bulimulus_alternatus_hesperius
Figs 15C, 15iv[U.S.A., Texas] “east side of the Pecos [river] at the High Bridge”.
“near Pecos river / Texas”; see remarks.
“34 mm. long, 17.4 mm. diameter”; figured specimen H 34.2, D 17.2, W 7.7.
RBINS/MT2353, two probable paratypes, ex Eyerdam, J.H. Ferriss leg., 1924 (Dautzenberg coll.).
Pilsbry and Ferriss (in
Bulimulidae, Rabdotus alternatus (Say, 1829).
http://species-id.net/wiki/Bulimulus_(Ataxus)_huayaboensis
Figs 11H, 11viii“Huayabo (Marañon) Pérou, à 2000 m. d’altitude”.
“Huayabo Marañon / Pérou 2000m alt.”, in Dautzenberg’s handwriting.
“Long. 24, diam. maj. 10 millim.”; figured specimen H 22.9, D 8.5, W 7.2.
RBINS/MT2354, five paralectotypes, Baer leg. (Dautzenberg coll.).
The holotype designation of a specimen in the MNHN collection (
Bulimulidae, Bostryx huyaboensis (Dautzenberg, 1901).
“Province of Matto-Grosso, Brazil (Germain)”.
“Matto-Grosso”, in Ancey’s handwriting.
“Long. 29, alt. (obl.) 13 mill.”; figured specimen 28.1, D 13.4, W 6.6.
RBINS/MT1881, one syntype, ex Géret ex Ancey, P. Germain leg. (Dautzenberg coll.).
Bulimulidae, Drymaeus (Drymaeus) poecilus ictericus (Ancey, 1892).
http://species-id.net/wiki/Bulimulus_(Drymaeus)_interruptus
Figs 12E, 12v“Merida, Venezuela”.
“Merida Venezuela”; see remarks.
“Alt. 25, diam. maj. 10.5 mm”; figured specimen H 23.6, D 10.6, W 6.1
RBINS/MT2257, one syntype, ex Preston (Dautzenberg coll.).
The specimen is not accompanied by the original Preston label, but Dautzenberg documented that the shell was acquired on 19.xii.1907 from Preston as a “co-type”. It is therefore considered a syntype. See also pallidus Preston, 1909.
Bulimulidae, Drymaeus (Mesembrinus) granadensis (Pfeiffer, 1848).
http://species-id.net/wiki/Bulimulus_(Drymaeus)_interruptus
Figs 11D, 11iv“Iocos (Peruvia) Baer legit. 1900”; see remarks.
“Iocos Pérou / Baer legit.”, in Dautzenberg’s handwriting.
“Altit. 13, latit. 3 millim.”; figured specimen H 11.3, D 2.8, W 10.3.
RBINS/MT2355, two paralectotypes, Bear leg., 1900 (Dautzenberg coll.).
Bulimulidae, Bostryx iocosensis (Dautzenberg, 1901).
http://species-id.net/wiki/Drymaeus_jousseaumei
Figs 13A, 13i“Rio Mixiollo, Province Huallaga, Pérou” [Dept. San Martin].
“Rio Mixiolla / prov. Huallaga Pérou”, in Dautzenberg’s handwriting.
“Long. 50, diam. maj. 22 mill.”; figured specimen H 47.6, D 21.6, W 6.9.
RBINS/MT2356, two paralectotypes, Baer leg. (Dautzenberg coll.).
Dautzenberg (1901c: 309) reports to have seen three specimens. The holotype designation of a specimen in the MNHN collection (
Bulimulidae, Drymaeus (Drymaeus) jousseaumei Dautzenberg, 1901.
http://species-id.net/wiki/Thaumastus_juana
Figs 10C–D, 10ii[Ecuador, Prov. Azuay] “Gualacco [sic, Gualaceo], province de Cuença”.
“chemin à Gualacco, rives du Paute, avant le pont”, in Cousin’s handwriting.
“long., 20 à 23 mm; diam. 10 à 12 mm”; figured specimen H 19.1, D 8.76, W 7.7.
RBINS/MT2357, seven paralectotypes, Cousin leg. (Dautzenberg coll.).
The lectotype is in the MNHN collection (
Bulimulidae, Bostryx juana (Cousin, 1887).
http://species-id.net/wiki/Orphnus_thompsoni_lutea
Figs 7A, 7i[Ecuador, Prov. Azuay] “Cuença”.
“Cuenca Ecuador”, label in Dautzenberg’s handwriting.
Not given. Lectotype H 77.6, D 37.2, W 6.7.
RBINS/MT2358, lectotype (design. n.); MT2359, five paralectotypes, ex Cousin (Dautzenberg coll.).
In the inventory of Cousin’s collection, a total of 11 specimens is mentioned for this variety. From among the specimens in RBINS, a lectotype is here designated and the taxon is now considered a junior subjective synonym of Bulimus thompsonii Pfeiffer, 1845 (syn. n.).
Orthalicidae, Kara thompsonii (Pfeiffer, 1845) (comb. n.).
http://species-id.net/wiki/Bulimulus_(Naesiotus)_lycodus
Figs 14F, 14iv[Ecuador, Galápagos, Isla Santa Cruz] “Indefatigable Island at 450 to 550 feet elevation [137–167 m]”.
“Indefatigable Island” on label of Stanford University.
“Length of shell 11, diameter 8 mm.”; figured specimen H 11.4, D 7.1, W 6.2.
RBINS/MT1834, one paratype, Ochsner leg.
The material is accompanied by a label from Stanford University and was donated by H.G. Schenck.
Bulimulidae, Naesiotus lycodus (Dall, 1917).
http://species-id.net/wiki/Bulimulus_mazei
Figs 15A, 15ii[Martinique] “Massif des Pitons, ad 730 met. altitudinem supra mare (Mazé)”.
“Martinique”, “Massif des Pitons”, in handwriting of Crosse.
“Long. 19 mill., diam. maj. 7”; figured specimen H 14.5, D 5.9, W 6.4.
RBINS/MT2360, one paralectotype, ex Crosse, Mazé leg. (Dautzenberg coll.).
The holotype designation of a specimen in the MNHN collection (
Bulimulidae, Protoglyptus mazei (Crosse, 1874) (comb. n.).
http://species-id.net/wiki/Bulimus_melanocheilus
Figs 4A–B, 4i“l’Amérique mériodionale, au Pampas”.
“Amerique mériodionale / (Pampas) / Pampaijacu”, in Nyst’s handwriting.
“longeur de 80 millimètres sur 35 de largeur”; lectotype H 78.5, D 36.6, W 6.2.
RBINS/MT2361, lectotype (design. n.), ex Nyst.
The specimen is accompanied by a label of Nyst, dated 1874, and marked “type Nyst”.
Megaspiridae, Thaumastus (Thaumastus) melanocheilus (Nyst, 1845).
http://species-id.net/wiki/Bostryx_metagyra
“Peru”.
“Peru”.
RBINS/MT2362, seven paratypes.
The specimens were exchanged with FMNH and
originate from the original series as deposited in the Museo de Historia
Natural in Lima; see also
Bulimulidae, Bostryx metagyra Pilsbry and Olsson, 1949.
http://species-id.net/wiki/Bulimulus_(Bostryx)_moniezi
Figs 11G, 11vii“le Haut-Pérou”.
“Andes / Pérou” (handwriting of Dupuis?).
“Longit. 14 millim., latit. 6 1/2 millim.”; figured specimen H 13.9, D 6.2, W 9.4.
RBINS/MT1830, one syntype, ex P. Dupuis.
The specimen is labelled “co-type” but fits the
original measurements. It is not accompanied by a Dautzenberg label;
however, its type status is not questioned here as Paul Dupuis is known
to have been in close contact with Dautzenberg (
Bulimulidae, Bostryx moniezi (Dautzenberg, 1896).
http://species-id.net/wiki/Orphnus_thompsoni_nigricans
Figs 7B, 7ii[Ecuador, Prov. Azuay] “Cuença”.
“Cuenca, Ecuador”; see remarks.
Not given. Lectotype H 62.8, D 30.3, W 6.1.
RBINS/MT2363, lectotype (design. n.); MT2364, three paralectotypes, ex Cousin (Dautzenberg coll.).
The original Cousin label mentions the taxon name only; the locality has been added by Dautzenberg, probably on the basis of Cousin’s paper. One specimen was found with the Cousin label stuffed inside the aperture and is here designated lectotype. The taxon is considered a junior subjective synonym of Bulimus thompsonii Pfeiffer, 1845 (syn. n.).
Orthalicidae, Kara thompsonii (Pfeiffer, 1845) (comb. n.).
http://species-id.net/wiki/Thaumastus_nystianus_nigricans
Figs 12B, 12ii[Ecuador, Prov. Pichincha] “les bords du chemin qui conduit de Pomasqui à Chilguiltina”.
“chemin de Pomasqui à Chilguiltina” in Dautzenberg’s handwriting; see remarks.
Not given. Figured specimen H 37.3, D 17.7, W 6.3.
RBINS/MT2365, 13 possible syntypes, ex Cousin (Dautzenberg coll.).
The material is not accompanied by an original Cousin label, but it originates from Cousin; the specimens are treated herein as possible syntypes. This taxon is now considered a junior subjective synonym of Bulimus nystianus Pfeiffer, 1853 (syn. n.).
Bulimulidae, Drymaeus (Drymaeus) nystianus (Pfeiffer, 1853).
http://species-id.net/wiki/Orphnus_thompsoni_olivacea
Figs 7C, 7iii[Ecuador, Prov. Azuay] “Cuença”.
“Cuenca, Ecuador”; see remarks.
Not given. Lectotype H 64.5, D 32.9, W 6.2.
RBINS/MT2366, lectotype (design. n.); MT2367, three paralectotypes, ex Cousin (Dautzenberg coll.).
The original Cousin label mentions the taxon name only; the locality has been added by Dautzenberg, probably on the basis of Cousin’s paper. One specimen was found with the Cousin label stuffed inside the aperture and is here designated lectotype. The taxon is considered a junior subjective synonym of Bulimus thompsonii Pfeiffer, 1845 (syn. n.).
Orthalicidae, Kara thompsonii (Pfeiffer, 1845) (comb. n.).
http://species-id.net/wiki/Bulimulus_(Drymaeus)_interruptus_pallidus
Figs 12F, 12vi“Merida, Venezuela”.
“Merida (Venezuela)”; see remarks.
Not given. Figured specimen H 24.0, D 11.0, W 6.2.
RBINS/MT2258, one syntype, ex Preston (Dautzenberg coll.).
The specimen is not accompanied by a Preston label, but Dautzenberg documented that he acquired the specimen from Preston on 19.xii.1907; the type status of the specimen is here not contested and it is regarded as a syntype.
Bulimulidae, Drymaeus (Mesembrinus) granadensis (Pfeiffer, 1848).
http://species-id.net/wiki/Placostylus_(Maoristylus)_ambagiosus_paraspiritus
Figs 8B, 8ii[New Zealand, North Island] “one mile south of Cape Maria van Diemen”.
“Headland, 1 mile S. of Cape Maria van / Diemen.”.
“Height 72.25 mm. Diameter 32.0 mm.”; figured specimen H 82.2, D 39.2, W 5.8.
RBINS/MT1969, one paratype, ex L. Germain (Dautzenberg coll.).
The specimen is labelled “paratypes”, despite the fact that it is only one specimen.
Bothriembryontidae, Placostylus ambagiosus (Suter, 1906).
http://species-id.net/wiki/Bulimulus_(Ataxus)_perforatus
“Ninabamba on the Pampas River, an affluent of the Apurimac River, Peru. Altitude 2, 000 m”.
“Ninabamba, Rio Pampas, 2.000 m. / alt., Peru” typewritten by Weyrauch.
“Height 19.5 mm, width 8.2 mm.”; largest specimen H 20.4, D 7.53, W 9.4.
RBINS/MT2368, two paratypes, W. Weyrauch leg.
The material was exchanged on the basis of material from the Dautzenberg collection (see Introduction) with FMNH, where the holotype of this taxon is kept.
Bulimulidae, Bostryx perforatus (Haas, 1951).
http://species-id.net/wiki/Bulimulus_pollonerae
Figs 14H–I, 14vii“San Lorenzo, province de Jujuy, République de Argentine”.
“San Lorenzo / prov. de Jujuy / Rep. Argentine”, in Ancey’s handwriting.
“Long. 15 1/2, lat. 6 1/2 mm.”; lectotype H 15.3, D 6.33, W 7.7.
RBINS/MT2369, lectotype (design. n.); MT2370, one paralectotype, ex Géret ex Ancey, Borelli leg. (Dautzenberg coll.).
The type material consists of a subadult and an adult specimen; the latter is here designated lectotype.
Bulimulidae, Naesiotus pollonerae (Ancey, 1897).
http://species-id.net/wiki/Leucocharis_porphyrochila
Figs 8A, 8i“Nouvelle-Calédonie”.
“Houaïlou N. Caléd.”, in Dautzenberg’s handwriting.
“Altit. 43 millim., latit. 22 millim.”; figured specimen H 41.9, D 22.0, W 5.9.
RBINS/MT2371, two syntypes, ex Bernier (Dautzenberg coll.).
Two additional syntypes are in the MNHN-collection (V. Héros, pers. commun.).
Bothriembryontidae, Aspastus (Leucocharis) porphyrochila (Dautzenberg and Bernier, 1901).
http://species-id.net/wiki/Bulimulus_(Scutalus)_sanborni
Figs 15D, 15i“Carhuamayo, basin of Lake Junín, Department of Loreto [sic, Junín], 15, 000–18, 000 feet [4572–5486 m]”.
“Carhuamayo above Lake Junin / Dept. Loreto, Peru” typewritten (by FMNH technician?).
“Height 10.6 mm, width 6 mm”; figured specimen H 10.0, D 5.5 W 4.5.
RBINS/MT2372, three paratypes, C.C. Sanborn leg., 1946.
The material was acquired by exchange with FMNH on the basis of material from the Dautzenberg collection (see Introduction).
Bulimulidae, Kuschelenia (Vermiculatus) sanborni (Haas, 1947) (comb. n.).
http://species-id.net/wiki/Drymaeus_scoliodes
Figs 13B, 13ii“Rio Mixiolla, province Huallaga, Pérou”.
“Rio Mixiolla prov. / Huallaga Pérou”, in Dautzenberg’s handwriting.
“Long. 63, diam. maj. 22 millim.”; figured specimen H 40.7, D 21.3, W 6.8.
RBINS/MT2377, one paralectotype, Baer leg. (Dautzenberg coll.).
The specimen is not marked as type, but
corresponds to the data as given by Dautzenberg (1901). The holotype
designation of a specimen in the MNHN collection (
Bulimulidae, Drymaeus (Drymaeus) scoliodes (Dautzenberg, 1901).
http://species-id.net/wiki/Bulimulus_(Drymaeus)_solidus
Figs 13C, 13iii“Bogota, United States of Colombia”.
“Bogota / U.S. Colombia”, in Preston’s handwriting.
“Alt. 32.5, diam. maj. 15 mm.”; figured specimen H 33.8, D 17.3, W 6.2.
RBINS/MT2259, one syntype, ex Preston (Dautzenberg coll.).
The specimen is marked “co-type” on the original label.
Bulimulidae, Drymaeus (Drymaeus) solidus (Preston, 1907).
http://species-id.net/wiki/Bulimus_spiculatus
Figs 11E, 11vNo type locality given [interior of Peru].
“Pérou”; see remarks.
“Longit. 20; diam. 4 mill.”; figured specimen H 21.8, D 4.26, W 11.4.
RBINS/MT2373, four probable syntypes, ex Morelet (Dautzenberg coll.).
Dautzenberg documented that he received the
specimens from the Morelet collection (“ex auctore”). The material is
regarded as probable syntypes. It may be noted that
Bulimulidae, Bostryx spiculatus (Morelet, 1860).
http://species-id.net/wiki/Bulimulus_(Protoglyptus)_subcostatus
“Jaën, Department of Cajamarca, Peru. Altitude 1, 500–2, 100 feet [457–604 m]”.
“Jaen, Camarca [sic], Peru, 550–700 m / alt. W. Weyrauch leg.” typewritten (by FMNH technician?).
“Height 12.4 mm, width 5 mm”.
RBINS/MT2379, two paratypes, W. Weyrauch leg.
Haas (1948) did not mention how many paratypes he had in his material. It is thus possible that these specimens formed part of the original series, as they are labelled “paratypes”. The species was adequately figured by Haas. The material was received in exchange from FMNH where the holotype is kept.
Bulimulidae, Naesiotus subcostatus (Haas, 1948).
http://species-id.net/wiki/Bulimus_veruculum
Figs 11F, 11viNo type locality given [interior of Peru].
“Balsa de Cocharcas / Pérou”, see remarks.
“Longit. 24; diam. 4 1/2 mill.”; figured specimen H 21.9, D 3.85, W 16.7.
RBINS/MT2374, four syntypes, ex Morelet (Dautzenberg coll.).
Dautzenberg documented that he received the specimens from the Morelet collection. The material is regarded as syntypes.
Bulimulidae, Bostryx veruculum (Morelet, 1860).
http://species-id.net/wiki/Placostylus_(Maoristylus)_whareana
Figs 8C, 8iii“Whareana, east coast between Waikuku Beach and Parengarenga”.
“Whareana Valley, S. of North Cape”.
“Height 79.5 mm. Diameter 35.0 mm.”; figured specimen H 65.9, D 30.7, W 7.0.
RBINS/MT376, one paratype, ex L. Germain.
The label reads “paratypes”, but only one specimen was found.
Bothriembryontidae, Placostylus ambagiosus (Suter, 1906).
http://species-id.net/wiki/Orphnus_thompsoni_zebra
Figs 7D, 7iv[Ecuador, Prov. Cañar] “près Azagues [sic, Azogues], sur la pente, à environs 2400m alt.”.
“près Azagues sur la / pente à envir. 2400m alt.”; see remarks.
Not given. Lectotype H 46.4, D 25.4, W 5.4.
RBINS/MT2375, lectotype (design. n.); MT2376, nine paralectotypes, ex Cousin (Dautzenberg coll.).
The original Cousin label mentions the taxon name only; the locality has been added by Dautzenberg, probably on the basis of Cousin’s paper. Several specimens are juvenile or damaged. One specimen is here designated lectotype. The taxon is considered a junior subjective synonym of Kara thompsonii (Pfeiffer, 1851) (syn. n.).
Orthalicidae, Kara thompsonii (Pfeiffer, 1851) (comb. n.).
A–B, i Plekocheilus (Eurytus) coloratus (Nyst, 1845), lectotype RBINS/MT2345 (H=47.3). C–E, ii Plekocheilus (Eurytus) dalmasi (Dautzenberg, 1900), lectotype RBINS/MT668 (H=26.3).
A–B, i Thaumastus (Thaumastus) melanocheilus (Nyst, 1845), lectotype RBINS/MT2361 (H=78.5). C–D, ii Thaumastus (Thaumastus) blanfordianus (Ancey, 1903), lectotype RBINS/MT1865 (H=52.5).
A–B, i Dryptus funckii (Nyst, 1843), lectotype RBINS/MT2352 (H=86.3).
A–C, i Plekocheilus (Plekocheilus) fulminans (Nyst, 1845), lectotype RBINS/MT2351 (H=59.2).
A–D Kara thompsonii (Pfeiffer, 1851); A, i lectotype var. lutea Cousin, 1887 RBINS/MT2358 (H=77.6); B, ii lectotype of var. nigricans Cousin, 1887 RBINS/MT2363 (H=62.8); C, iii lectotype of var. olivacea Cousin, 1887 RBINS/MT2366 (H=64.5); D, iv lectotype of var. zebra Cousin, 1887 RBINS/MT2375 (H=46.4).
A, i Aspastus porphyrochilus (Dautzenberg and Bernier, 1901), syntype RBINS/MT2371 (H=41.9). B–C Placostylus ambagiosus (Suter, 1906) B, ii paratype of Placostylus ambagiosus paraspiritus Powell, 1951 RBINS/MT1969 (H=82.2). C, iii paratype of Placostylus ambagiosus whareana Powell, 1951 RBINS/MT376 (H=65.9).
A–B Placostylus A, i lectotype of Placostylus elata Dautzenberg, 1923 RBINS/MT702 (H=86.3) B, ii Placostylus auriculata Dautzenberg, 1923 RBINS/MT2339 (H=81.7).
Bostryx species. A–B, i Bostryx alausiensis (Cousin, 1887), paralectotype RBINS/MT2333 (H=25.2). C–D, ii Bostryx juana (Cousin, 1887), paralectotype RBINS/MT2357 (H= 19.1). E, iii Bostryx chacoensis (Preston, 1907), syntype RBINS/MT2343 (H=30.1) F–G, iv Bostryx borellii (Ancey, 1897), syntype RBINS/MT/xx (H=28.8). H, v Bostryx carandaitiensis (Preston, 1907), syntype RBINS/MT2341 (H=32.1).
Bostryx species. A, i Bostryx bonneti (Ancey, 1902), paralectotype RBINS/MT2338 (H=21.9). B, ii Bostryx emaciatus (Morelet, 1863), paralectotype RBINS/MT2349 (H=18.3). C, iii Bostryx baeri (Dautzenberg, 1901), paralectotype RBINS/MT2336 (H=13.6). D, iv Bostryx iocosensis (Dautzenberg, 1901), paralectotype RBINS/MT2355 (H=11.3). E, v Bostryx spiculatus (Morelet, 1860), probable syntype RBINS/MT2373 (H=21.3). F, vi Bostryx veruculum (Morelet, 1860), probable syntype RBINS/MT2374 (H=21.9). G, vii Bostryx moniezi (Dautzenberg, 1901), syntype RBINS/MT1830 (H=13.9). H, viii Bostryx huayaboensis (Dautzenberg, 1901), paralectotype RBINS/MT2354 (H=22.9). I, ix Bostryx orophilus (Morelet, 1860), syntype of albicolor Morelet, 1863 RBINS/MT2335 (H=22.2).
Drymaeus species. A, i Drymaeus (Drymaeus) abruptus (Rolle, 1904), paralectotype RBINS/MT2332 (H=36.6). B, ii Drymaeus (Drymaeus) nystianus (Pfeiffer, 1853), lectotype of nigricans Cousin, 1887 RBINS/MT2365 (H=37.3). C, iii Drymaeus (Drymaeus) colimensis (Rolle, 1904), paralectotype RBINS/MT2344 (H=29.1). D, iv Drymaeus (Drymaeus) poecilus ictericus (Ancey, 1892), syntype RBINS/MT1881 (H=28.1). E–F Drymaeus (Mesembrinus) granadensis (Pfeiffer, 1848) E, v syntype of interruptus Preston, 1909 RBINS/MT2257 (H=22.6). F, vi syntype of pallidus Preston, 1909 RBINS/MT2258 (H=24.0).
Drymaeus species. A, i Drymaeus (Drymaeus) jousseaumei (Dautzenberg, 1901), paralectotype RBINS/MT2356 (H=47.6). B, ii Drymaeus (Drymaeus) scolioides (Dautzenberg, 1901), paralectotype RBINS/MT2377 (H=40.7). C, iii Drymaeus (Drymaeus) solidus (Preston, 1907), syntype RBINS/MT2259 (H=33.8).
Naesiotus species. A, ii Naesiotus cucullinus (Dall, 1917), paratype RBINS/MT1833 (H=17.7). B, vi Naesiotus amastroides (Ancey, 1887), lectotype RBINS/MT1866 (H=9.25). C, v Naesiotus eschariferus (Sowerby, 1833), syntype of Bulimulus eschariferus bizonalis Ancey, 1887 RBINS/MT2337 (H=16.0). D–E, i Naesiotus montivagus (d’Orbigny, 1835), lectotype of Bulimulus montivagus chacoensis Ancey, 1897 RBINS/MT2342 (H=20.9). F, iv Naesiotus lycodus (Dall, 1917), paratype RBINS/MT1834 (H=11.4). G, iii Naesiotus cavagnaroi A.G. Smith, 1972, holotype of Bulimulus gilderoyi Van Mol, 1972 RBINS/MT106 (H=25.4). H–I, vii Naesiotus pollonerae (Ancey, 1897), lectotype RBINS/MT2369 (H=15.3).
A, ii Protoglyptus mazei (Crosse, 1874), paralectotype RBINS/MT2360 (H=14.5). B, iii Protoglyptus dejectus (Fulton, 1907), paralectotype RBINS/MT2348 (H=28.0). C, iv Rabdotus dealbatus (Say, 1821), probable paratype of Bulimulus alternatus hesperius Pilsbry and Ferriss, 1924 RBINS/MT2353 (H=34.2). D, i Kuschelenia (Vermiculatus) sanborni (Haas, 1947), paratype RBINS/MT2372 (H=10.0).
A–B, i Rhinus argentinus (Ancey, 1901), syntype RBINS/MT1867 (H=19.7). C, ii Simpulopsis (Eudioptus) ephippium (Ancey, 1904), lectotype RBINS/MT2350 (H=19.5).
Familiy Bulimulidae Tryon, 1867
Stenostylus is a genus of high-altitude species, known to occur above 3000 m (
1 | Surface with strong, thickened growth striae and traces of spiral impressions, shell height above 50 mm | 2 |
– | Surface of shell relatively smooth, shell height up to 50 mm | 3 |
2(1) | Last whorl relatively stout, aperture ovate; Peru, western slopes of Andes | Stenostylus zilchi Weyrauch, 1956 |
– | Last whorl relatively elongated, aperture elongate-ovate; Peru, eastern slopes of Andes | Stenostylus perturbatus sp. n. |
3(1) | Spire pointed, whorls relatively flat; Peru, western slopes of Andes | Stenostylus meleagris (Pfeiffer, 1854) |
– | Spire obtuse, whorls rounded | 4 |
4(3) | Shell thin, smooth, shell height up to 20 mm; (?Colombia), Ecuador, eastern slopes of Andes | Stenostylus colmeiroi (Hidalgo, 1872) |
– | Shell rather solid, with thickened growth striae, shell height more than 20 mm; Colombia, Cordillera Oriental | Stenostylus nigrolimbatus (Pfeiffer, 1854) |
urn:lsid:zoobank.org:act:D4D62ABB-D38C-458C-99CD-506CC09E3BB3
http://species-id.net/wiki/Stenostylus_perturbatus
Figs 17A–BA large species of Stenostylus, with a very elongated spire, a roughly sculptured surface, and the aperture relatively small, and narrow, compared to other species of the genus.
Shell 56.5 mm, 2.0 times as long as wide, elongate, with straight sides, imperforate; rather thin. Colour dark-brown to blackish, with axially oriented, yellowish lines, varying from small to somewhat broader, in some places forming patches, but always irregular, never from suture to suture; upper whorls light-brown, the first ones denuded of the periostracum. Surface rather shining, with thickened growth striae where the colour is yellowish; some traces of spiral impressions, especially visible at the upper side of the last whorl. Protoconch eroded. Whorls 6.2, slight convex, the last 0.75 total shell height; suture well impressed. Aperture elongate-ovate, 0.45 times shell height, 1.49 times as long as wide, somewhat shining inside, whitish. Peristome thin, hardly expanded; columellar margin thin, curved, transitioning into the parietal wall, which has a very thin, whitish callus.
A–B Stenostylus perturbatus spec. nov., holotype RBINS/MT698 (H=56.5). C–E Suniellus adriani spec.nov., holotype RBINS/MT2378 (H=32.5).
Holotype H 56.5, D 27.7, HA 25.5, WA 17.1.
Peru, Dept. Pasco, Huancabamba.
RBINS/MT698, holotype (ex Preston).
This new species is placed in Stenostylus,
despite its protoconch sculpture being unknown, on account of its
general shape, the thickened growth striae, and the traces of a pearly
lustre inside the aperture (cf.
The single specimen known was found under the same manuscript name of Preston, from whom Dautzenberg obtained the shell on 30.xii.1909. The lip is damaged, especially at the columellar and basal side, and has partly been broken off.
(L.), perturbatus, unquiet; referring to the distinctive colour pattern of the shell. The epithet is used as an adjective.
Hitherto this taxon was regarded a subgenus of Scutalus Albers, 1860. The latter genus appears to be polyphyletic (Breure, unpublished data); Suniellus is now interpreted as the high-altitude sister-group of Kuschelenia Hylton Scott, 1951 (cf.
1 | Last whorl saccate, aperture elongate-ovate | 2 |
– | Last whorl regularly rounded, aperture ovate | 3 |
2(1) | Relatively small (up to 16 mm); Bolivia | Suniellus chillu Breure, 1978 |
– | Relatively large (more than 25 mm); Ecuador | Suniellus adriani sp. n. |
3(1) | Shell height more than 25 mm; Colombia | Suniellus goudoti (Petit, 1843) |
– | Shell height less than 25 mm; Peru | Suniellus troscheli (Philippi, 1867) |
urn:lsid:zoobank.org:act:B2AF1109-5F1F-41CE-A02C-4A7308AFF2D7
http://species-id.net/wiki/Suniellus_adriani
Figs 17C–EA relatively large, and elongate species of Suniellus, with the last whorl a little saccate.
Shell 32.5 mm, 1.86 times as long as wide, elongate-ovate, with hardly convex sides, imperforate; very thin. Colour yellowish-olivaceus with brownish streaks, especially on last whorl. Upper whorls lighter in colour. Surface rather shining, with irregularly spaced, thickened growth striae, in between with weaker incrassations and, especially at upper part of last whorl, some oblong granules. Protoconch eroded, but on the dorsal side traces visible of axial wrinkles, partly anastomosing. Whorls 5.7, hardly convex, the last 0.81 times total shell height, a little saccate; suture well impressed. Aperture elongate-ovate, 0.54 times shell height, approx. 1.4 times as long as wide, with a pearly lustre inside. Peristome thin and simple, columella thread-like, somewhat dilated above and appressed at the transition to the parietal wall, which bears a hardly noticeable callus.
Holotype H 32.5, D 17.4, HA 17.4, WA 12.2.
Ecuador, Prov. Pichincha, “San Diego Cuchu” (see remarks).
RBINS/MT2378, holotype, ex Cousin (Dautzenberg coll.).
This novelty closely resembles Suniellus chillu Breure, 1978 from Bolivia in shell shape, but is much larger.
The lip is damaged at the upper palatal side and the insertion to the shell has been torn off, leaving some tears at the upper part of the last whorl, just behind the lip. The type locality could not be found in modern gazetteers; it is a mixture of Spanish and Quechua words, and may not be officially known under this name. It is likely to be in the páramo area, as Cousin on a second label has written “pie de la nieve / Frutillas por arriba” [at the foot of the snow / Strawberries above]; snow in the 19th century probably occurred at lower elevations than today, but an elevation for this locality of above 2750 m may be a safe guess (González, pers. commun.). There is only one specimen known.
The epithet is a patronym in honour of Adrián González—a Cuban malacologist now living in Quito, Ecuador—in recognition for his contributions to Neotropical malacology by his photographic work, his books aimed at a general public, and his dedicated fieldwork.
First of all I want to express my most sincere gratitude to Thierry Backeljau and Rose Sablon (RBINS) for their kind hospitality and all their support during my stay in Brussels. Also the support of Jan van Tol (NCB Naturalis) is gratefully acknowledged. Jackie Van Goethem (RBINS) kindly informed me about the status of some type specimens included in the Dautzenberg collection. I am very much indebted to Jonathan Ablett (London) and Virginie Héros (Paris) for providing photographs for taxa comparison, and to Jennifer Gallichan (Cardiff) for discussion on some Ancey types. Robert Herschler (Washington) and Henk Mienis (Jerusalem) kindly provided information on Preston material in the collections in their charge. The comments of Thierry Backeljau (Brussels), Francisco Borrero (Cincinnati), and two reviewers greatly improved a previous draft of the manuscript, and is gratefully acknowledged.