Indonesia, Java (Xyleborus hirtipennis, lectotype, USNM); Sarawak, Malaysia (Xyleborus puberulus, holotype, BMNH).

Lectotype of Arixyleborus hirtipennis bears all essential features of Arixyleborus puberulus, only the declivital rugosities more organized into rows, shining area of elytra smaller, less clearly distinguished from rugose area. These are exceptionally plastic in Arixyleborus puberulus, Arixyleborus hirtipennis represents small deviation in the large range of variation of declivital surface in Arixyleborus puberulus.

Indonesia, Java, Buitenzorg (Diuncus justus, holotype, NHMW); Philippines, Luzon, (Xyleborus marginicollis, holotype, NHMW).

1.5 mm.

Type specimen of Xyleborus marginicollis Schedl represents one end of a continuum of variation in Diuncus justus: short (1.5 mm) but robust (most representatives of Diuncus justus slightly longer and more slender). Diagnostic characters identical: surface of declivity devoid of vestiture, no elytral denticles, smooth impression across interstriae 2 and 3 (very shallow).

Malaysia, Sabah, Danum Valley (2, R.A. Beaver det., MSUC); Sumatra (USNM).

Prolonged large representative of Debus. Elytral declivity deeply excavated, edge of declivity with two pairs of long teeth, but only few tubercles. Declivital surface smooth.

Debus amphicranoides (Hagedorn) possibly senior synonym of the following (NHMW): Debus birmanus (Eggers), Debus cyclopus (Schedl), Debus interponens (Schedl), Debus robustipennins (Schedl). Debus birmanus identical except slightly larger, with slightly longer declivital posterolateral processes, much smaller upper tooth on declivity. Debus interponens similar except lacks constricted declivity and has shorter posterolateral declivital processes. Schedl (1954) considered Debus (as Xyleborus) interponens possible altitudinal variation of Xyleborus robustipennins; the two essentially identical, originated from different elevations. Debus cyclopus similar except narrower elytral apical emargination. Debus robustipennis larger. Schedl (1954) mentioned that it only differed from Debus amphicranoides in minor differences in declivital teeth shape.

Malaysia, Burma (2 indiv., BMNH).

Very similar to Debus amphicranoides, possibly a synonym. Holotype at FRI not available.

Malaysia, Sarawak (holotype, BMNH); Malaysia, Sabah, Danum Valley (9 indiv., Hulcr det., MSUC.).

Elytral declivity slightly with much higher number of declivital tubercles than other Debus. Declivity flat, not excavated, not emarginate at apex. Depth of emargination varies. Similar to Debus pumilus, but uniformly brown, with more and larger tubercles on the declivity. Significant intraspecific size variation.

Elytral declivity superficially different from other Debus spp, but its structure homologous. Few small or large tubercles in the interstriae 1 (usually 3 pairs), displaced by broadened interstriae 1 and positioned on first striae or on interstriae 2. Strial punctures greatly reduced on declivity, difficult to follow as interstria 1 broad, displacing other striae. No tubercles originating on second striae. Smaller tubercles on striae 3 and beyond, creating tuberculated area surrounding declivity. Other characters shared with Debus spp.: extended pronotal disc, triangular protibiae with large and long but sparse denticles (<7), inflated prosternal posterocoxal process, antennal club shape.

Xyleborus persimilis (Eggers) and Debus dolosus (Blandford) probably synonyms. Xyleborus persimilis (lectotype, USNM) with slightly broader, more excavated declivity. Browne (1961) suggested that Xyleborus subdolosus is only a local variety of Debus dolosus.

Indonesia, Kalimantan (2, holotype & allotype, NSMT); Indonesia, Kalimantan (2 paratypes, NHMW).

Elytral apex not emarginate, but all other diagnostic characters of Debus present: elongated pronotal disc, broad antennal club type 2, triangular protibiae, flat elytral declivity with tubercles on elevated lateral sulcus (appears as if formed by interstriae 2 through 4).

Xyleborus shoreae: India, Kumaon, (2), Beeson det., BMNH); Malaysia, Kedah, (two labels: Xyleborus shoreae, Browne det., Xyleborus fallax, Schedl det., BMNH); Thailand, Pong Yaeng N. P., (Beaver det.); Borneo (Schedl det., BMNH); Debus fallax: Malaysia, Sabah, Danum Valley (Hulcr det.); Malaysia, Sabah, Danum Valley (51, Hulcr det.); New Guinea, Morobe Province, Bulolo (Jordal det.); New Guinea (BBM, 5 indiv.); Sulawesi (Browne det., BMNH); Thailand, Pong Yaeng N. P. (2, Hulcr det.); PNG, Madang Prov. (36), Oro Prov. (66), West Sepik (123) (Hulcr coll, det.); Philippines, Luzon, Mt. Makiling (Xyleborus amphicranulus Egg. holotype, Xyleborus fallax syn., SMTD).

Holotype of Xyleborus shoreae in FRI, inaccessible, non-type specimens identified by several authorities available. Location of Xyleborus fallax holotype unclear. Wood and Bright (1992) indicated IRSNB as holotype depository, however museum personell reports that holotype has never been deposited there. Xyleborus shoreae is a variant of Debus fallax (Eichhoff), declivital emargination shallower than in most Debus fallax. All other characters identical to Debus fallax: color uniformly brown to bicolored (light brown to orange pronotum), elytral denticles all small except the denticle in the middle of declivital face, which is slightly longer than others; declivity surface shining, most specimens with remnants of strial punctures, size 2.6 - 3.0 mm. Declivity emargination depth intermediate between Debus fallax and Debus emarginatus, most other characters (size, coloration, flat posterolateral processes) shared with Debus fallax. Maiti and Saha (2004) had access to holotype, redescription and illustration fits Debus fallax. Stebbing not consistent in distinguishing Xyleborus shoreae from Xyleborus fallax, assigned similar specimens to either species (Maiti and Saha 2004).

Reported from India and Thailand, and by Browne (1983) as imported from PNG to Japan. Despite the name “shoreae”, the species is a broad generalist (Wood and Bright 1992).

Malaysia, Sarawak, Mt. Penrissen, 4500 ft. (lectotype, NHMW).

All diagnostic features of genus Debus present, including antennal club form, prolonged pronotum, emarginate declivity. Similar to Debus amphicranoides (Hagedorn), but with less constricted declivity and longer posterolateral declivital processes. Schedl (1954) considered Debus interponens altitudinal variant of Debus robustipennins, the two are allegedly identical, only differing by their origins from different elevations.

Indonesia, Borneo (lectotype, NHMW).

All diagnostic features of Debus present, including antennal club form, prolonged pronotum, emarginate declivity.

Lectotype of Xyleborus robustipennis Schedl very similar to non-type specimens of Debus amphicranoides (Hagedorn) in USNM, only slightly larger. Schedl (1954) indicated that Xyleborus robustipennis differs from Xyleborus amphicranoides very little, merely by shallower and wider declivital emargination, having the lateral declivital costa between teeth 1 and 2 more elevated, and lateral declivital process shorter. Type of Debus amphicranoides not available, thus synonymy could not be confirmed.

Malaysia (BMNH); Malaysia, Sabah, Danum Valley (3, Hulcr det.).

An “elegant” form of Debus fallax. Longer, smooth declivity, shallowly emarginate, no tubercles or granules on declivital sides except two pairs of slender teeth, one long, one short. Declivity shagreen when dry.

Xyleborus terminaliae: Philippines, Pagbilao (holotype, USNM). Xyleborus exesus: Japan, (holotype, BMNH). Debus emarginatus: Indonesia, Sumatra, Bandar Baroe (homotype, compared to type by Eggers, NHMW); Indonesia, Java, Bandjar (homotype, compared to type by Eggers, NHMW, 2 indiv.); Philippines, Laguna, Pangil (homotype, NHMW); Malaysia, Sabah, Danum Valley (17 indiv., Hulcr det., MSUC); New Guinea (BBM, 20 indiv.); New Guinea, Ambunti (4, BBM); New Guinea (FICB); New Guinea, Gulf Province, Ivimka (UCD); Thailand, Pong Yaeng N. P. (Hulcr det., MSUC); PNG, Madang Prov. (79, Hulcr coll.).

Holotypes of Xyleborus exesus Blandford, Xyleborus palmeri Hopkins, and Xyleborus terminaliae Hopkins share all diagnostic characters with homotype and large series of non-types of Debus emarginatus (Eichhoff). Xyleborus exesus: declivity with slightly less steep slope, less pronounced lateral tubercles (granules), dominant tubercle in middle of lateral sulcus slightly longer. Schedl (1973e) suggested synonymy of non-New Guinean Xyleborus emarginatus Schedl with Xyleborus exesus Blandford, based on shared shallow declivital emargination. Holotype of Xyleborus exesus damaged, missing elytron, fits range of Debus emarginatus variation. Xyleborus palmeri Hopkins is larger variant of typical Debus emarginatus.

India, Sunderbans Div. (holotype, BMNH).

All diagnostic features of Microperus present: small size, elytral punctures aligned in striae, and prolonged body shape (Hulcr and Cognato in press). Similar to Microperus pometianus, but slightly longer, with distinctly elevated and long declivital costa.

Microperus corporaali: Indonesia, Kotangan an der Ostkusgte (lectotype, USNM); Xyleborus fulvulus: Indonesia, Sumatra (paratype, USNM).

Xyleborus fulvulus identical to Microperus corporaali (identical antennae, body shape, declivital surface and shape, posterolateral declivital costa, declivital vestiture as one row of erect setae per intrestria, backward-bent setae in strial punctures). Paratype not mentioned by Anderson and Anderson (1971).

Australia, Queensland, Geagana (lectotype, NHMW).

All diagnostic features of Microperus present (elytral mycangia, absence of scutellum, small size, prolonged body shape, abundant vestiture). Similar to Microperus intermedius, but substantially longer, elytra often bicolored, usually without convexity on elytral disc.

Malaysia, Selangor (lectotype, BMNH); Malaysia, Selangor (Schedl det., BMNH); Malaysia, Sabah, Danum Valley, (13, Hulcr det., MSUC).

Very similar to Microperus diversicolor (e.g., antennal club type 3), except pronotum bright yellow with brown patch, elytra black, declivity commencing closer to elytral base, declivital interstriae covered with many small sharp hooks (similar as in Microperus parva, but larger). Characteristic elytral disc: anterior portion inflated, convex, boundary between elytral disc and declivity slightly concave, impressed.

Schedl (1979) designated lectotype in NHMW, another unspecified “type” resides in BMNH. Possibly synonymous with Coptodryas undulata (Sampson) (as Xyleborus leprosulus Schedl, syn. Wood 1989) (Schedl 1939).

Creates irregularly branching galleries with transverse brood chambers (Beaver and Browne 1978).

Xyleborus artelaevis: Malaysia, Perak, (holotype, NHMW); New Guinea, Gulf Province, Ivimka, (R.A. Beaver det., UCD); Indonesia, Sulawesi (R. A. Beaver det., BMNH). Planiculus bicolor: Nagasaki, Japan (syntype, BMNH); Fiji, Namosi (Xyleborus rameus (syn. Planiculus bicolor) Schedl det., BMNH).

Holotype of Xyleborus artelaevis virtually identical to Planiculus bicolor (Blandford), except first segment of antennal club more convex. All other characters identical, including uniform granules in declivital interstriae 1, 2, and 3 (same size granules in interstriae 1–3 characteristic for Planiculus bicolor). Xyleborus artelaevis holotype deteriorated, missing or damaged body parts including antenne.

Malaysia, Peninsula (holotype, NHMW).

Diagnostic characters of Pseudowebbia: regular type of pronotum (not extremely prolonged and flat as in Webbia), circular antennal club (not broadened), triangular to broadly rounded protibia (not thin and sickle-like as in Webbia). Elytral declivity deeply excavated, surrounded by highly elevated circumdeclivital costa with no teeth.

Type species of Taphrodasus Wood, 1980. Morphological limits of Taphrodasus never specified. Characters listed by Wood (1980) are either autapomorphic to Taphrodasus percorthylus, or present in other genera, mostly Webbia. Taphrodasus not a valid genus, see below.

Indonesia, Java, Buitenzorg (holotype, NHMW).

Xyleborus superbus Schedl (1951) preoccupied by Xyleborus superbus Schedl (1942c). Renamed Xyleborus superbulus (Schedl, 1958a). Xyleborus superbus Schedl (1942c) later placed in Coptoborus (Wood and Bright 1992). Replacement name unnecessary, original name Streptocranus superbus (Schedl, 1951) restored.

Truncaudum agnatum: New Guinea, Hatam (cotype, MCG). Xyleborus polyodon: Philippines, Luzon, Mt. Makiling; (unspecified “type”, SMTD).

Type of Xyleborus polyodon similar to Truncaudum agnatum, except tubercles on and around declivity larger, pointed. Tubercles in homologous position. Eggers’s unspecified “type” in SMTD from the same collection series as lectotype at USNM (Anderson & Andreson, 1971).

Uganda (Premnobius circumcinctus, holotype, NHMW).

The only known African Truncaudum. Truncaudum synapomorphies: impressed submentum, antennal club type 1, curved outer edge of protibiae, complete denticulated circumdeclivital costa. Very similar to Truncaudum impexus [(Schedl): Hulcr and Cognato (in press)], except declivity slightly convex (mostly flat in Truncaudum spp.), antenna type 1, several adjacent denticles on each stria on the upper edge of circumdeclivital costa (mostly a single flat tubercle in Truncaudum spp.). Otherwise remarkably similar to Asian relatives. Length: 2.8 mm.

Described as Premnobius by Schedl (1941), later treated as Xyleborus (Schedl, 1962b), but combination never offically published.

Malaysia, Perak (holotype, BMNH).

Transferred to Taphrodasus (Wood & Bright, 1992) without discussion of characters. Webbia synapomorphies: dorsal aspect of pronotum long and quadrangular, pronotal disc long and flat, frontal slope of pronotum short, scutellum suppressed, costate and setose elytral bases. Differs from most Webbia spp. by densely pubescent and excavated declivity and elongated body shape. Length: 2.4 mm. Characters shared with Pseudowebbia percorthylus [(Schedl, 1935): Hulcr and Cognato, this volume] (type species of Taphrodasus) limited to excavated declivity with dense setae, genus-level characters different.

Malaysia, N.S. Triang (homotype, NHMW); Malaysia, Perak (BMNH); Malaysia, Borneo (BMNH).

Type in Hamburg museum lost (Wood and Bright 1992).Most features characteristic of Webbia (elongated pronotum, suppressed scutellum), unrelated to type species of Taphrodasus: Pseudowebbia percorthylus ((Schedl, 1935): Hulcr & Cognato, this volume). Similar to Webbia divisus, except declivity with long, dense, erect setae, not scales.

Xyleborinus perminutissimus: Indonesia, Java, Mt. Gede (lectotype, NHMW). Xyleborinus perpusillus: Indonesia, Sumatra (holotype, USNM); Malaysia, Sarawak, Gunung Buda (R.A. Beaver det., MSUC); Malaysia, Sabah, Danum Vallery (29 indiv., Hulcr coll.); New Guinea, Oro Province, Kanga (5 indiv, Hulcr coll.).

Lectotype of Xyleborinus perminutissimus (Schedl, 1934d) virtually identical to holotype of Xyleborinus perpusillus (Eggers). Slightly smaller tubercles in some declivital interstriae, but pattern identical: tubercles missing from interstriae 2.