Research Article |
Jiang-Li Tan‡, James M. Carpenter§, Cornelis van Achterberg‡
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Corresponding author: Jiang-Li Tan ( tanjl@nwu.edu.cn ) Academic editor: Andreas Köhler
© 2018 Jiang-Li Tan, James M. Carpenter, Cornelis van Achterberg.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Tan J-L, Carpenter JM, van Achterberg C (2018) An illustrated key to the genera of Eumeninae from China, with a checklist of species (Hymenoptera, Vespidae). ZooKeys 740: 109-149. https://doi.org/10.3897/zookeys.740.22654
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Abstract
An illustrated key to the currently recognized genera of the subfamily Eumeninae (Vespidae) from China is presented together with a list of 267 species and subspecies, belonging to 51 genera. Nortozumia van der Vecht, 1937 is reported for the first time from China. Two replacement names are proposed for junior primary homonyms: Ancistrocerus rufofrustius Tan & Carpenter, nom. n. replacing Ancistrocerus rufopictus (Kostylev) and Orientalicesa confasciatus Tan & Carpenter, nom. n. replacing Orientalicesa unifasciatus (von Schulthess, 1934).
Keywords
homonym, illustrated key, new record, Oriental, Palaearctic
Introduction
Eumeninae or potter wasps are the largest subfamily of the Vespidae with 3773 valid species in 205 genera (
The Chinese Eumeninae were first catalogued by Liu (1936–1937) resulting in a list of 57 species divided among nine genera. Unfortunately, his research stopped after his only revision (Pareumenes;
Material and methods
Specimens were collected by hand net or with Malaise traps. The studied specimens are deposited in the collections of College of Life Sciences, Northwest University, Xi’an (
Morphological terminology follows
Key to genera of Eumeninae from China
| 1 | Tergum I more or less petiolate, slender and its apical half parallel-sided or slightly narrowed posteriorly (1a); tergum I usually at least twice as long as wide (but in e.g., Pseudozumia slightly longer than wide); tergum II twice maximum width of tergum I in dorsal view, but approx. 1.5 × in Pseudozumia | 2 |
| – | Tergum I robust and its apical half widened posteriorly (1aa); tergum I much less than twice as long as wide; tergum II at most twice as wide as tergum I | 18 |
| 2 | Middle tibia with two spurs (2a) | 3 |
| – | Middle tibia with one spur (2aa) | 5 |
| 3 | Propodeum with valvula distinctly protruding, elongate and rectangular (3a), orifice acutely pointed dorsally (3b) | Zethus Fabricius |
| – | Propodeum with valvula not protruding (3aa), orifice rounded dorsally (3bb) | 4 |
| 4 | Maxillary palpus 3-segmented (4a); tergum I more than twice as long as wide, longitudinally striate (4b); tegula posteriorly acute (4c) | Calligaster de Saussure |
| – | Maxillary palpus 4-segmented (4aa); tergum I less than twice as long as wide, punctate (4bb); tegula posteriorly truncate (4cc) | Discoelius Latreille |
| 5 | Propodeal valvula relatively short, rounded, submarginal carina distinctly protruding (5a); frons distinctly longer than clypeus; second submarginal cell narrow anteriorly, vein 1-M and vein 2-M meeting almost at right angle (5b) | 6 |
| – | Propodeal valvula elongate, rectangular, submarginal carina not protruding (4aa); frons shorter than clypeus; second submarginal cell wide anteriorly, vein 1-M and vein 2-M meeting at obtuse angle (5bb) | 8 |
| 6 | Female with fovea anterior to mid-ocellus (6a); metanotum monodentiform mesally (6d); metasomal petiole abruptly swollen apically in lateral view (6c); tegula not exceeding parategula posteriorly (6b); anterior face of pronotum smooth (6e) | Labus de Saussure |
| – | Female without fovea on vertex (6aa); metanotum obtuse mesally (6dd); metasomal petiole not conspicuously swollen in lateral view (6cc); tegula more or less exceeding parategula posteriorly (6bb); anterior face of pronotum distinctly punctuate laterally (6ee) | 7 |
| 7 | Mesoscutum distinctly longer than wide (7a), apical margin of tergum I wider than half width of TII (7b); parategula short, almost absent, tegula distinctly exceeding parategula (7c) | Leptomicrodynerus Giordani Soika |
| – | Mesoscutum distinctly wider than long (7aa); apical margin of tergum I narrower than half width of tergum II (7bb); parategula normal, tegula slightly exceeding parategula (7cc) | Cyrtolabulus van der Vecht |
| 8 | Propodeum dorsally with elongate fovea from which a carina runs to orifice, usually with dentiform projections above valvulae (8a); axillary fossa narrower than long, slit-like (8b); tegula with narrow posterior lobe which about equals parategula posteriorly (8c) | 9 |
| – | Propodeum without fovea or dentiform projections (8aa); axillary fossa oval, broader than long (8bb); tegula short, convex and not equalling parategula posteriorly (8cc), or long, with narrow posterior lobe which surpasses parategula posteriorly | 14 |
| 9 | Mesepisternum with epicnemial carina present (9a) | 10 |
| – | Mesepisternum with epicnemial carina absent (9aa) | 13 |
| 10 | Metasomal petiole with transverse carina basally (10a, a’) | 11 |
| – | Metasomal petiole not carinate (10aa, aa’) | 12 |
| 11 | Tergum I with its lateral margins fused ventrally; sternum I reduced to posterior crescentic sclerite (11a); [mesonotum and propodeum smooth between fine punctures] | Ectopioglossa Perkins |
| – | Tergum I with lateral margins not meeting ventrally, sternum I visible along entire petiole length (11aa) | Nortozumia van der Vecht (new record) |
| 12 | Mesoscutum with a pair of prescutellar longitudinal groove (12a); forewing with parastigma longer than pterostigma (12b); sternum I irregularly rugose posteriorly, with rugae running in longitudinal direction (12c) | Pseudozumia de Saussure |
| – | Mesoscutum without prescutellar longitudinal groove (12aa); forewing with parastigma shorter than pterostigma (12bb); sternum I smooth basally, its posterior two thirds transversely striate or smooth (12cc) | Coeleumenes van der Vecht |
| 13 | Forewing with parastigma longer than pterostigma (13a); sternum I gradually widened backwards with regular transverse striae (13b); female with cephalic fovea (13c); hind tibia with number of short spines on its outside (13d) | Pareumenes de Saussure |
| – | Forewing with parastigma shorter than pterostigma (13aa); sternum I narrower basally, more or less fused with tergum I, posteriorly short, triangular and without ruga (13bb); female without cephalic fovea (13cc); hind tibia without spines on its outside (13dd) | Pseumenes Giordani Soika |
| 14 | Tergum I impunctate or with only a few small punctures (14c); propodeum inclining posteriorly into a slope (14a); tergum II without lamella separated by preapical thickening, sometimes with pale border (14b) | 15 |
| – | Tergum I with dense, coarse punctation (14cc); propodeum less inclined posteriorly (14aa); tergum II with apical lamella more or less separated from disc by preapical thickening (14bb) | 17 |
| 15 | Clypeus apically rounded (15a); temple in dorsal view as long as eye (15b) | Katamenes Meade-Waldo |
| – | Clypeus apically truncate or emarginated (15aa); temple in dorsal view shorter than eye (15bb) | 16 |
| 16 | Tergum I slightly longer than mesosoma, with section after spiracles shorter than section before spiracles (16a); male: terminal sternum with a longitudinal groove (16b) | Delta de Saussure |
| – | Tergum I much longer than mesosoma, with section after spiracles longer than section before spiracles (16aa); male: terminal sternum without groove (16bb) | Phimenes Giordani Soika |
| 17 | Pronotum with pretegular carina absent (17b); parastigma of forewing shorter than half of pterostigma (17a); female: apical margin of clypeus emarginated (17c); propodeum lateral margin rounded, without distinct border with posterior face; male: apical antennal segment medium-sized and curved (17d) | Eumenes Latreille |
| – | Pronotum with pretegular carina present (17bb); parastigma of forewing longer than half of pterostigma (17aa); female: apical margin of clypeus truncated (17cc); propodeum lateral side distinctly separated from its posterior face; male: apical antennal segment small and straight (17dd) | Oreumenes Bequaert |
| 18 | Tegula evenly rounded posteriorly and usually not reaching apex of parategula (18a); male antenna apically spiralled (18b) | 19 |
| – | Tegula protruding posteriorly, emarginate or truncate adjoining parategula (18aa); male antenna apically hooked or simple (18bb) | 22 |
| 19 | Labial palpi 3-segmented, in female segment II and III both broadly flattened, fringed with setae, forming a psammophore (19a); sterna II-V in male usually with central apical brush (19b) | 20 |
| – | Labial palpus 4-segmented, in female cylindrical, without psammophore (19aa); sterna II-V in male usually without central apical brush (19bb) | 21 |
| 20 | Maxillary palpus 5- (20a) or 6- (20a’) segmented; female: labial palpus segment II thick basally, segment II, III not curved (20b); male: mandible with penultimate tooth often without deep excision, and axis of penultimate tooth is at an oblique angle relative to main axis of mandible and approximately parallel to axis of apical tooth (20c); body <9 mm | Pterocheilus Klug |
| – | Maxillary palpus 6-segmented (20aa); female: labial palpus segment II and III slender, flat and curved (20bb); male: mandible with distance between second and third tooth broad (20cc) or deeply and broadly excised, and axis of penultimate tooth is at approximately a right angle relative to the main axis of the mandible and relative to the apical tooth (20cc’); body >10 mm | Onychopterocheilus Blüthgen |
| 21 | Propodeum with lateral carinae well developed (21a); female without cephalic foveae (21b); vertex weakly longitudinally rugose posteriorly (21b); temples and mesosoma with very fine pubescence; male sterna II-VI with conspicuous fringe of setae (21c) | Tropidodynerus Blüthgen |
| – | Propodeum with lateral carinae weak or absent (21aa); female with cephalic foveae; vertex not rugose; temples and mesosoma with long setae (21bb); male sterna without fringe of setae (21cc) | Odynerus Latreille |
| 22 | Anterior face of pronotum with two close and deeply impressed pits, which may be approximated, or with series of elongate foveae (22a); tegula broad, wider than long, not surpassing parategula (except Jucancistrocerus) (22b); propodeum valvula bilamellate (with submarginal carina produced into pointed lamella apically and valvula enlarged and free posteriorly from submarginal carina) (22c) | 23 |
| – | Anterior face of pronotum without deep pits or foveae (except punctures) (22aa, aa’); submarginal carina, propodeum valvula and tegula variable | 28 |
| 23 | Tergum I with two transverse carinae (23a) or with one (23a’) | 24 |
| – | Tergum I without transverse carinae (23aa) | 27 |
| 24 | Tegula densely punctate, sieve-like, surpassing parategula posteriorly (24a); propodeal dorsum without extending horizontal area (24b); pretegular carina absent (24c); [carina of tergum I indistinct in some species] | Jucancistrocerus Blüthgen |
| – | Tegula usually finely punctate (24aa); propodeal dorsum extending horizontally, forming shelf-like area behind metanotum (24bb); pretegular carina present (24cc) | 25 |
| 25 | Tergum I with two carinae, tergum wider than long in dorsal view, both carinae relatively close near each other (25a) | Subancistrocerus de Saussure |
| – | Tergum I with one (25aa) or two carinae (25aa’); if with two carinae, then tergum I longer than wide in dorsal view, more or less petiole-like and distance between both carinae relatively large | 26 |
| 26 | Anterior face of pronotum with foveae separated (26a); tergum II usually smooth basally, forming an acarinarium (26b); metasoma sessile, tergum I nearly as wide as tergum II (26b’) | Parancistrocerus Bequaert |
| – | Anterior face of pronotum with contiguous foveae (26aa); tergum II ridged basally, not forming an acarinarium; tergum I in dorsal view longer than wide, tergum II much wider than tergum I (26bb) | Pseudonortonia Giordani Soika |
| 27 | Tergum I relatively short, gradually widened, with its lateral sides divergent in dorsal view (27a); vertex strongly depressed, forming an acute triangle with face (27b, arrowing part); in female, apical margin of clypeus truncated and with two longitudinal carinae (27c); anterior face of pronotum with foveae coalesced (27b) | Paraleptomenes Giordani Soika |
| – | Tergum I relatively long, roughly parallel-sided in dorsal view (27aa); vertex normal, not forming an acute triangle with face (27bb); apical margin of clypeus emarginate and without carina (27cc); anterior face of pronotum with foveae separated (22a). [Note: if tergum I of Parancistrocerus spp. has an indistinct transverse carina, then it is difficult to separate them from Stenodynerus; Parancistrocerus spp. usually have an acarinarium on tergum II basally and tergum I more or less lengthened medially in dorsal view, while Stenodynerus spp. have tergum II ridged basally and tergum I medio-dorsally without elongation] | Stenodynerus de Saussure |
| 28 | Tergum I transversely carinate (28a) or horizontal and vertical faces of tergum clearly separated (Pararrhynchium, 28a’) | 29 |
| – | Tergum I evenly curved, without transverse carina (28aa) | 34 |
| 29 | Tergum I with broad medio-longitudinal furrow posterior to transverse carina (29a); notauli clearly indicated (29b); male antenna simple apically (29c) | Symmorphus Wesmael |
| – | Tergum I without medio-longitudinal furrow (29aa); notauli absent or nearly so (29bb); male antenna hooked apically (29cc) | 30 |
| 30 | Parastigma of forewing with more than half the length of pterostigma, measured along posterior part, often nearly equal (30a) | 31 |
| – | Parastigma half the length of pterostigma or less, measured along posterior part (30aa) | 33 |
| 31 | Tergum II with well-developed apical lamella (31a) | Lissodynerus Giordani Soika |
| – | Tergum II lacking an apical lamella (31aa) | 32 |
| 32 | Clypeus wide ventrally and slightly emarginated medio-ventrally (32a); male: terminal sternum without teeth basally (32b) | Orancistrocerus van der Vecht |
| – | Clypeus narrower ventrally and deeply emarginated medio-ventrally (32aa); male: terminal sternum with 2–3 teeth basally (32bb) | Archancistrocerus Giordani Soika |
| 33 | Axillary fossa narrower than long, slit-like (33b); humeri (“shoulder”) rounded (33a); propodeal dorsum strongly extending horizontally, forming shelf-like area behind metanotum (33c) | Pararrhynchium de Saussure |
| – | Axillary fossa oval, broader than long (33bb); humeri angular or pointed (33aa); propodeal dorsum slightly extending behind metanotum, below level of metanotum (33cc) | Ancistrocerus Wesmael |
| 34 | Metanotum with serrate bilobed ridge (34a, a’) | 35 |
| – | Metanotum low toothed (34aa), including blunt or rounded off ridge or flat | 36 |
| 35 | Clypeus higher than wide (35a); metanotum with truncate teeth; mid-anterior face of pronotum smooth and with short transverse rugae (35b); tergum I distinctly narrower than tergum II (35c); male terminal antennal segment small (35d) | Apodynerus Giordani Soika |
| – | Clypeus wider than high (35aa); mid-anterior face of pronotum usually densely punctate and with an upper trace of transverse carina (35bb); tergum I slightly narrower than tergum II (35cc); male terminal antennal segment relatively large (35dd) | Antepipona de Saussure |
| 36 | Tergum II with lamella behind transverse band (36a) | 37 |
| – | Tergum II without lamella behind transverse band, at most with narrow border (36aa). Note: few Euodynerus spp. may possess a distinct lamella on tergum II (see 45a), they can be separated from Leptochilus by having tergum I not depressed subapically (depressed in Leptochilus); and differs from Gribodia, Stenodynerellus and Epsilon by having oval axillary fossa, broader than long (slit-like, narrower than long in Gribodia, Stenodynerellus and Epsilon) | 40 |
| 37 | Tergum I depressed subapically, gradually widened with lateral sides divergent in dorsal view (37 aa); propodeum with submarginal carina projecting as rounded lobe above valvula, bilamellate (37bb); epicnemial carina absent (37cc); axillary fossa oval, broader than long (37dd) | Leptochilus de Saussure |
| – | Tergum I not depressed subapically, usually with lateral sides roughly parallel in dorsal view (37aa); propodeum with submarginal carina not differentiated from valvula, mono-lamellate (37bb; except Epsilon); epicnemial carina present (37cc); axillary fossa narrower than long, slit-like (37dd) | 38 |
| 38 | Palpal formula 5:3 (38a); male vertex sometimes with large and deep depression (38b); propodeum without shelf-like protruding part and with lateral carinae well developed (38c); [metanotum angulated, second submarginal cell with second recurrent vein nearly or completely interstitial with third submarginal cell; terga I-V each with apical lamella] | Gribodia Zavattari |
| – | Palpal formula 6:4 (38aa); male vertex without large and deep depression (38bb); propodeum with shelf-like protruding part (38cc) or absent (38cc’) | 39 |
| 39 | Clypeus longer than wide, apical margin usually truncate (39a); propodeum with submarginal carina not differentiated from valvula (39b); propodeum usually with dorsal surface at about same level as metanotum, lateral margin rounded; metanotum usually smoothly convex (39c); terga I-V each with apical lamella (39d); second submarginal cell with second recurrent vein variable | Stenodyneriellus Giordani Soika |
| – | Clypeus longer than wide, apical margin emarginate (39aa); propodeum with submarginal carina projecting as rounded lobe above valvula (39bb); propodeum without raised shelf-like part, lateral margin carinate; metanotum angulated (39cc); only tergum II with lamella (39dd); second submarginal cell with second recurrent vein nearly or completely interstitial with third submarginal cell (39ee) | Epsilon de Saussure |
| 40 | Axillary fossa in dorsal view much narrower than long, often slit-like (40a); tegula short, not exceeding parategula | 41 |
| – | Axillary fossa in dorsal view not slit-like, at least as wide as long, oval (40aa); tegula usually equal to or exceeding parategula | 45 |
| 41 | Propodeum with raised shelf-like part nearly at level of metanotum, metanotum flat (41a) | 42 |
| – | Propodeum below level of metanotum, metanotum various (41aa) | 43 |
| 42 | Forewing with third submarginal cell separated from apex of marginal cell by about half its length (42a); male sternum VII with basal truncate process (42b) | Allorhynchium van der Vecht |
| – | Forewing with third submarginal cell separated from apex of marginal cell by about its own length (42aa); male sternum VII without process (42bb) | Orientalicesa Koçak and Kemal |
| 43 | Parastigma of forewing more than half length of pterostigma, measured along posterior part or nearly equal (43a); palpal formula 6:4 (38aa) | 44 |
| – | Parastigma shorter than half length of pterostigma, measured along posterior part (43aa); palpal formula 5:3 (38a) | Okinawepipona Yamane |
| 44 | Mesoscutum posteriorly and scutellum smooth, very sparsely and finely punctate (44a); metanotum depressed medially (44b); male middle femur basally emarginated (44c) | Rhynchium Spinola |
| – | Mesoscutum and scutellum richly punctate throughout (44aa); metanotum not depressed medially (44bb); male middle femur not basally emarginate (44cc) | Anterhynchium de Saussure |
| 45 | Tergum I behind apical band with well-developed lamella (45a) | 46 |
| – | Tergum I behind apical band without well-developed lamella (45aa) | 47 |
| 46 | Metanotum between horizontal and vertical area with hemi-circular carina (46a) | Antodynerus de Saussure |
| – | Metanotum between areas without hemi-circular shaped carina (46aa) | Euodynerus Dalla Torre |
| 47 | Pretegular carina absent (47a) | 48 |
| – | Pretegular carina present, at least posteriorly (47aa) | 49 |
| 48 | Tegula sparsely punctate, slightly exceeding parategula and much smaller than scutellum (48a); mid-anterior face of pronotum smooth (48b); metanotum with lateral lamellae (48c); female with single cephalic fovea situated near occipital carina (48d) | Parodontodynerus Blüthgen |
| – | Tegula densely punctate, distinctly exceeding parategula, almost as large as scutellum (48aa); mid-anterior face of pronotum densely punctate (48bb); metanotum without lateral lamellae (48cc); female with single cephalic fovea situated halfway posterior ocelli and occipital carina (48dd) | Brachyodynerus Blüthgen |
| 49 | Tegula narrower and longer, surpassing parategula posteriorly (49a). [female vertex with reniform fovea, about as wide as ocellar triangle; hind coxa with ventral lobes] | Allodynerus Blüthgen |
| – | Tegula broad, equal to parategula posteriorly (49aa) | 50 |
| 50 | Propodeal valvula mono-lamellate (50a), with transverse carina, and with a dentate ridge laterally (50b); [metanotum ridge roughly bidentate-shaped] | Pseudepipona de Saussure |
| – | Propodeal valvula bilamellate (50aa), without transverse carina, and without dentate ridge laterally; [small species (6 mm body length); anterior face of pronotum smooth; vertex with very small pits] | Asiodynerus Kurzenko |
Head and pronotum (a, e, aa, ee), part of mesonotum in dorsal view (b, bb, dd), metanotum magnified (d), and metasomal segment I in lateral view (c, cc): a–e Labus spiniger (de Saussure) aa, ee, lower cc Leptomicrodynerus tieshengi Giordani Soika bb, dd, upper cc Cyrtolabulus suavis van der Vecht.
Forewing (a, aa), metasomal sternum I (SI) (b, bb), head in dorsal view (c, cc) and hind tibia (d, dd). a–b Pareumenes quadrispinosus conjunctus Liu c, d Pareumeness. sansibaricus (von Schulthess) aa–cc Pseumenes d. depressus (de Saussure); dd. Pseumenes depressus annulatus van der Vecht.
Metasomal tergum I (a, aa) in dorsal view, propodeum in lateral view (b, bb), mesosoma in lateral view (c, cc) and metanotum in dorsal view (d, dd). a–d Leptochilus m. mauritanicus (Lepeletier) aa Gribodia confluenta (Smith) bb, cc Stenodyneriellus guttulatus (de Saussure) dd Stenodyneriellus sp.
Checklist of the species of the subfamily Eumeninae from China
Allodynerus Blüthgen, 1938
Allodynerus Blüthgen, 1938 (1937), Konowia 16: 280; Type species: “Lionotus floricola Sauss. 1852” [= Odynerus floricola de Saussure, 1853].
Allodynerus delphinalis delphinalis (Giraud, 1866)
Allodynerus mandschuricus Blüthgen, 1953
Allorhynchium van der Vecht, 1963
Allorhynchium van der Vecht, 1963, Zool. Verh., Leiden 60: 57 (key), 58, genus. Type species: Vespa argentata Fabricius, 1804.
Allorhynchium chinense (de Saussure, 1862)
Allorhynchium lugubrinum (Cameron, 1900)
Allorhynchium metallicum (de Saussure, 1852)
Ancistrocerus Wesmael, 1836
Ancistrocerus Wesmael, 1836, Bull. Acad. R. Belg. 3: 45. Type species: Vespa parietum Linnaeus, 1758.
Ancistrocerus arcanus Giordani Soika, 1993
Ancistrocerus aureovillosus Giordani Soika, 1977
Ancistrocerus deqinensis You and Li, 2013
Ancistrocerus frigidus Giordani Soika, 1977 (1976)
Ancistrocerus hirsutus hirsutus (Meade-Waldo, 1910)
Ancistrocerus hirsutus supiensis Giordani Soika, 1977 (1976)
Ancistrocerus hirsutus tinkiensis Giordani Soika, 1977 (1976)
Ancistrocerus khangmarensis Giordani Soika, 1966
Ancistrocerus krausei Giordani Soika, 1966
Ancistrocerus melanocerus (Dalla Torre, 1894)
Ancistrocerus melanurus Morawitz, 1889
Ancistrocerus montuosus Gusenleitner, 1993
Ancistrocerus nigricornis (Curtis, 1791)
Ancistrocerus parapoloi Giordani Soika, 1966
Ancistrocerus parietum (Linnaeus, 1758)
Ancistrocerus rufofrustius Tan & Carpenter, nom. n.
Replacement name for Ancistrocerus rufopictus (Kostylev, 1940) (junior primary homonym of Odynerus lineaticollis var. rufopictus Meade-Waldo, 1915).
Ancistrocerus sikhimensis (Bingham, 1897)
Ancistrocerus terayamai Yamane, 1993
Ancistrocerus tibetanus Giordani Soika, 1966
Ancistrocerus transpunctatus You & Li, 2013
Ancistrocerus trifasciatus shibuyai (Yasumatsu, 1938)
Ancistrocerus waltoni (Meade-Waldo, 1910)
Antepipona de Saussure, 1855
Antepipona de Saussure, 1855, Ét. Fam. Vesp. 3: 244. Type species: Odynerus silaos de Saussure, 1853.
Antepipona asiamontana Gusenleitner, 2004
Antepipona biguttata (Fabricius, 1787)
Antepipona bipustulata (de Saussure, 1855)
Antepipona brunneola Giordani Soika, 1986
Antepipona deflenda (Saunders, 1853)
Antepipona excelsa excelsa Giordani Soika, 1982
Antepipona ferruginea Kim & Yamane, 2003
Antepipona ferruginea Kim & Yamane, 2003
Antepipona guttata diffinis (de Saussure, 1855)
Antepipona menkei Giordani Soika, 1986
Antepipona ovalis (de Saussure, 1853)
Antepipona plurimacula Giordani Soika, 1971
Antepipona rufescens (Smith, 1857)
Antepipona shantungensis Giordani Soika, 1993
Antepipona sexfasciata Soika, 1986
Antepipona tytides (Cameron, 1904)
Anterhynchium de Saussure, 1863
Anterhynchium de Saussure, 1863, Mém. Soc. Phys. Hist. Nat. Genève 17: 205. Type species: Rygchium [= Rhynchium] synagroides de Saussure, 1852.
Anterhynchium (Anterhynchium) mellyi (de Saussure, 1852)
Anterhynchium (Dirhynchium) flavomarginatum flavomarginatum (Smith, 1852)
Anterhynchium (Dirhynchium) flavomarginatum formosicola (von Schulthess, 1934)
Anterhynchium (Dirhynchium) flavopunctatum flavopunctatum (Smith, 1852)
Anterhynchium (Dirhynchium) flavopunctatum opulentum (Giordani Soika, 1973)
Anterhynchium (Dirhynchium) inamurai (Sonan, 1937)
Anterhynchium (Dirhynchium) yunnanensis Giordani Soika, 1973
Antodynerus de Saussure, 1855
Antodynerus de Saussure, 1855, Ét. Fam. Vesp. 3: 287.Type species: “Odynerus punctum (Fabricius)” sensu de Saussure, 1853 [= Vespa flavescens Fabricius, 1775]
Antodynerus limbatus (de Saussure, 1852)
Apodynerus Giordani Soika, 1993
Apodynerus Giordani Soika, 1993, Boll. Mus. Civ. Stor. Nat. Venezia 42: 155. Type species: Odynerus troglodytes de Saussure, 1855.
Apodynerus formosensis continentalis Giordani Soika, 1994
Apodynerus formosensis formosensis (von Schulthess, 1934)
Apodynerus troglodytes troglodytes (de Saussure, 1855)
Apodynerus yayeyamensis yayeyamensis (Matsmura, 1926)
Archancistrocerus Giordani Soika, 1986
Archancistrocerus Giordani Soika, 1986, Boll. Mus. Civ. Stor. Nat. Venezia 35: 143, genus. Type species: Archancistrocerus diffinis Giordani Soika, 1986, monotypy.
Archancistrocerus diffinis Giordani Soika, 1986
Note: Archancistrocerus is a junior synonym and will be included in Allorhynchium (
Asiodynerus Kurzenko, 1977
Asiodynerus Kurzenko, 1977, Ins. Mongol. 5: 557. Type species: Odynerus lucifer Kostylev, 1937.
Asiodynerus lucifer (Kostylev, 1937 (1936))
Brachyodynerus Blüthgen, 1938
Brachyodynerus Blüthgen, 1938, Deutsch. Entomol. Zeitschr.: 450, 459, genus. Type species: Odynerus (Lionotus) magnificus Morawitz, 1867.
Brachyodynerus perarrus Kurzenko, 1977
Calligaster de Saussure, 1852
Calligaster de Saussure, 1852, Ét. fam. Vesp. 1: 22, genus. Type species: Calligaster cyanoptera de Saussure, 1852.
Calligaster himalayensis (Cameron, 1904)
Coeleumenes van der Vecht, 1963
Coeleumenes van der Vecht, 1963, Zool. Verh., Leiden 60: 45, genus. Type species: Montezumia impavida Bingham, 1897
Coeleumenes burmanicus (Bingham, 1897)
Coeleumenes thoracicus (Sonan, 1939)
Cyrtolabulus van der Vecht, 1969
Cyrtolabulus van der Vecht, 1969, Entomol. Ber., Amst. 29: 1, replacement name for Cyrtolabus van der Vecht, 1963, non Voss, 1925. Type species: Cyrtolabus suavis van der Vecht, 1963.
Cyrtolabulus exiguus (de Saussure, 1853)
Cyrtolabulus yunnanensis Lee, 1982
Delta de Saussure, 1855
Delta de Saussure, 1855, Ét. Fam. Vesp. 3: 130, 132, 143. Type species: Vespa maxillosa DeGeer, 1775 [= Vespa emarginata Linnaeus, 1758]
Delta campaniforme campaniforme (Fabricius, 1775)
Delta conoideum (Gmelin, 1790)
Delta esuriens okinawae Giordani Soika, 1986
Delta pyriforme pyriforme (Fabricius, 1775)
Discoelius Latreille, 1809
Discoelius Latreille, 1809, Gen. Crust. et Insect. 4: 140. Type species: Vespa zonalis Panzer, 1801, monotypy.
Discoelius dufourii dufourii (Lepeletier, 1841)
Discoelius dufourii manchurianus Yasumatsu, 1934
Discoelius emeishanensus Zhou and Li, 2013
Discoelius esakii Yasumatsu, 1934
Discoelius longinodus Yamane, 1996
Discoelius nigriclypeus Zhou & Li, 2013
Discoelius wangi Yamane, 1996
Discoelius zonalis (Panzer, 1801)
Ectopioglossa Perkins, 1912
Ectopioglossa Perkins, 1912, Ann. Mag. Nat. Hist. (8) 9: 118, genus. Type species: Ectopioglossa australensis Perkins, 1912, monotypy.
Ectopioglossa ovalis Giordani Soika, 1993
Ectopioglossa taiwana (Sonan, 1938)
Epsilon Saussure, 1855
Epsilon de Saussure, 1855, Ét. Fam. Vesp. 3: 229, 252. Type species: Odynerus dyscherus de Saussure, 1852.
Epsilon fujianense Lee, 1981
Eumenes Latreille, 1802
Eumenes Latreille, 1802, Hist. Nat. Crust. Ins. 3: 360, genus. Type species: “Eumenes coarctata, Fab.” [= Vespa coarctata Linnaeus, 1758].
Eumenes architectus Smith, 1859
Eumenes assamensis Meade-Waldo, 1910
Eumenes atrophicus (Fabricius, 1798)
Eumenes buddha Cameron, 1897
Eumenes coarctatus coarctatus (Linnaeus, 1758)
Eumenes coronatus coronatus (Panzer, 1799)
Eumenes ferrugiantennus Zhou, Chen & Li, 2012
Eumenes formosensis Giordani Soika, 1973
Eumenes fraterculus Dalla Torre, 1894
Eumenes fulvopilosellus Giordani Soika, 1965
Eumenes kangrae Dover, 1925
Eumenes kiangsuensis Giordani Soika, 1941
Eumenes labiatus flavoniger Giordani Soika, 1941
Eumenes labiatus labiatus Giordani Soika, 1941
Eumenes labiatus sinicus Giordani Soika, 1941
Eumenes mediterraneus manchurianus Giordani Soika, 1971
Eumenes micado Cameron, 1904 (Kim and Yoon (2001) state that the record is very doubtful)
Eumenes multipictus de Saussure, 1855
Eumenes nigriscutatus Zhou, Chen & Li, 2012
Eumenes pedunculatus pedunculatus (Panzer, 1799)
Eumenes pomiformis (Fabricius, 1781)
Eumenes punctatus de Saussure, 1852
Eumenes quadratus obsoletus Dover, 1926
Eumenes quadratus quadratus Smith, 1852
Eumenes quadratus urainus Sonan, 1939
Eumenes rubronotatus Pérez, 1905
Eumenes septentrionalis khangmarensis Giordani Soika, 1966
Eumenes septentrionalis septentrionalis Giordani Soika, 1940
Eumenes tosawae lofouensis Giordani Soika, 1973
Eumenes tosawae tosawae Giordani Soika, 1934
Eumenes transbaicalicus Kurzenko, 1984
Eumenes tripunctatus (Christ, 1791)
Eumenes variepunctatus Giordani Soika, 1941
Euodynerus Dalla Torre, 1904
Euodynerus Dalla Torre, 1904, Gen. Ins. 19: 38. Type species: Vespa dantici Rossi, 1790.
Euodynerus (Euodynerus) caspicus caspicus (Morawitz, 1873)
Euodynerus (Euodynerus) dantici brachytomus (Kostylev, 1940)
Euodynerus (Euodynerus) dantici violaceipennis Giordani Soika, 1973
Euodynerus (Euodynerus) fastidiosus (de Saussure, 1853)
Euodynerus (Euodynerus) rufinus rufinus Blüthgen, 1942
Euodynerus (Euodynerus) semisaecularis semisaecularis (Dalla Torre, 1889)
Euodynerus (Euodynerus) variegatus kruegeri (von Schulthess, 1928)
Euodynerus (Pareuodynerus) adiacens Giordani Soika, 1973
Euodynerus (Pareuodynerus) deqinensis Ma, Chen & Li, 2017
Euodynerus (Pareuodynerus) ferrugineus Ma, Chen & Li, 2017
Euodynerus (Pareuodynerus) nipanicus nipanicus (von Schulthess, 1908)
Euodynerus (Pareuodynerus) nipanicus ryukyuensis Tano, 1987
Euodynerus (Pareuodynerus) nipanicus tonkinensis Giordani Soika, 1973
Euodynerus (Pareuodynerus) notatus notatus (Jurine, 1807)
Euodynerus (Pareuodynerus) quadrifasciatus quadrifasciatus (Fabricius, 1793)
Euodynerus (Pareuodynerus) similinipanicus Ma, Chen & Li, 2017
Euodynerus (Pareuodynerus) strigatus (Radoszkowski, 1893)
Euodynerus (Pareuodynerus) trilobus (Fabricius, 1787)
Gribodia Zavattari, 1912
Gribodia Zavattari, 1912, Arch. Naturgesch. 78A (4): 161. Type species: Monobia cavifrons Gribodo, 1891 [= Odynerus confluenta Smith, 1857], monotypy.
Gribodia nigra Nguyen & Xu, 2015
Jucancistrocerus Blüthgen, 1938
Jucancistrocerus Blüthgen, 1938, Deutsch. Entomol. Zeitschr.: 442, 460.Type species: Odynerus (Ancistrocerus) jucundus Mocsáry, 1883, monotypy.
Jucancistrocerus (Jucancistrocerus) alashanicus Kurzenko, 1977
Jucancistrocerus (Jucancistrocerus) angustifrons (Kostylev, 1940)
Jucancistrocerus (Eremodynerus) atrofasciatus (Morawitz, 1885)
Jucancistrocerus (Eremodynerus) chotanensis (Blüthgen, 1942)
Katamenes Meade-Waldo, 1910
Katamenes Meade-Waldo, 1910, Ann. Mag. Nat. Hist. (8) 5: 46, genus. Type species: Katamenes watsoni Meade-Waldo, 1910, monotypy.
Katamene arbustorum arbustorum (Panzer, 1799)
Katamene indetonsus (Morawitz, 1895)
Katamene tauricus tauricus (de Saussure, 1855)
Labus de Saussure, 1867
Labus de Saussure, 1867, Reise Novara, Zool. 2 (1), Hym.: 3, genus. Type species: Labus spiniger de Saussure, 1867
Labus exiguus (de Saussure, 1855)
Labus lofuensis Giordani Soika, 1973
Leptochilus de Saussure, 1853
Leptochilus de Saussure, 1853, Ét. Fam. Vesp 1: 233. Type species: Pterocheilus mauritianus [!] [= Pterocheilus mauritanicus Lepeletier, 1841].
Leptochilus (Lionotulus) chinensis Gusenleitner, 2001
Leptochilus (Lionotulus) gobicus (Kostylev, 1940)
Leptochilus (Lionotulus) incertus (Kostylev, 1940)
Leptochilus (Neoleptochilus) tibetanus Giordani Soika, 1966
Lissodynerus Giordani Soika, 1993
Lissodynerus Giordani Soika, 1993, Boll. Mus. Civ. Stor. Nat. Venezia 42: 135. Type species: Odynerus septemfasciatus Smith, 1857.
Lissodynerus septemfasciatus feanus (Giordani Soika, 1941)
Leptomicrodynerus Giordani Soika, 1985
Leptomicrodynerus Giordani Soika, 1985, Lavori Soc. Ven. Sci. Nat. 10: 37. Type species: Leptomicrodynerus tieshengi Soika, 1985, monotypy.
Leptomicrodynerus tieshengi Giordani Soika, 1985
Note: The characteristics of this genus also fit well with Eumenidiopsis Giordani Soika, 1939, and further research is needed to solve the problem.
Nortozumia van der Vecht, 1937 (new record)
Nortozumia van der Vecht, 1937, Treubia 16: 263, genus. Type species: Zethus rufofemoratus Cameron, 1903.
Nortozumia sp.
Odynerus Latreille, 1802
Odynerus Latreille, 1802, Hist. Nat. Crust. Ins. 3: 362. Type species: Vespa spinipes Linnaeus, 1758.
Odynerus (Odynerus) tristis tianshanicus Kurzenko, 1977.
Okinawepipona Yamane, 1987
Okinawepipona Yamane, 1987, Mem. Kagoshima Univ. Res. Center S. Pacific 8: 52. Type species: Anterhynchium kogimai Giordani Soika, 1986, monotypy.
Okinawepipona curcipunctura Nguyen & Xu, 2014
Okinawepipona kogimai taiwana Yamane, 1987
Okinawepipona nigra Nguyen & Xu, 2014
Onychopterocheilus Blüthgen, 1955
Onychopterocheilus Blüthgen, 1955, Mitt. Münch. Entomol. Ges. 44/45: 406, 407. Type species: “Pterocheilus daw (Dusmet, 1909)” [= Odynerus (Hoplomerus) daw Dusmet, 1903], monotypy.
Onychopterocheilus (Asiapterocheilus) bensoni (Giordani Soika, 1941)
Onychopterocheilus (Asiapterocheilus) nigropilosus (Kostylev, 1940)
Onychopterocheilus (Asiapterocheilus) rongsharensis (Giordani Soika, 1977)
Onychopterocheilus (Asiapterocheilus) tibetanus (Meade-Waldo, 1913)
Onychopterocheilus (Asiapterocheilus) waltoni (Meade-Waldo, 1913)
Onychopterocheilus (Onychopterocheilus) chinensis Gusenleitner, 2005
Onychopterocheilus (Onychopterocheilus) dementievi (Kostylev, 1940)
Onychopterocheilus (Onychopterocheilus) eckloni (Morawitz, 1885)
Onychopterocheilus (Onychopterocheilus) wuhaiensis Gusenleitner, 2005
Orancistrocerus van der Vecht, 1963
Orancistrocerus van der Vecht, 1963, Zool. Verh., Leiden 60: 58 (key), 99, genus. Type species: Odynerus drewseni de Saussure, 1857
Orancistrocerus aterrimus aterrimus (de Saussure, 1852)
Orancistrocerus drewseni drewseni (de Saussure, 1857)
Orancistrocerus drewseni ingens (von Schulthess, 1934)
Orancistrocerus drewseni opulentissimus (Giordani Soika, 1941)
Orancistrocerus moelleri aulicus Giordani Soika, 1973
Oreumenes Bequaert, 1926
Oreumenes Bequaert, 1926, Ann. S. Afr. Mus. 23: 488. Type species: Eumenes harmandi Perez, 1905 [= Eumenes decoratus Smith, 1852], monotypy.
Oreumenes decoratus (Smith, 1852)
Orientalicesa Koçak & Kemal, 2010
Orientalicesa Koçak & Kemal, 2010, CESA Misc. Pap. 151: 4, replacement name for Kennethia Giordani Soika, 1994, non De Dekker, 1979. Type species: Odynerus unifasciatus von Schulthess, 1934.
Orientalicesa confasciatus Tan & Carpenter, nom. n.
Replacement name for Orientalicesa unifasciatus (von Schulthess, 1934) (junior primary homonym of Odynerus unifasciatus de Saussure, 1852).
Paraleptomenes Giordani Soika, 1970
Paraleptomenes Giordani Soika, 1970, Boll. Mus. Civ. Stor. Nat. Venezia 20/21: 79, genus. Type species: Paraleptomenes nurseanus Giordani Soika, 1970, monotypy.
Paraleptomenes kosempoensis (von Schulthess, 1934)
Paraleptomenes miniatus miniatus (de Saussure, 1855)
Parancistrocerus Bequaert, 1925
Parancistrocerus Bequaert, 1925, Trans. Am. Entomol. Soc. 51: 64. Type species: Odynerus fulvipes de Saussure, 1855 [= O. “flavipes Fabricius” sensu de Saussure, 1852, non Vespa flavipes Fabricius, 1775].
Parancistrocerus hongkongensis Gusenleitner, 2002
Parancistrocerus intermediatus (Sonan, 1939)
Parancistrocerus kuraruensis (Sonan, 1939)
Parancistrocerus nitobei (Sonan, 1939)
Parancistrocerus taihorinensis (von Schulthess, 1934)
Parancistrocerus taikonus (Sonan, 1939)
Parancistrocerus yachowensis konkunesis Giordani Soika, 1994
Parancistrocerus yachowensis yachowensis Giordani Soika, 1986
Parancistrocerus yamanei Gusenleitner, 2000
Pararrhynchium de Saussure, 1855
Pararrhynchium de Saussure, 1855, Ét. Fam. Vesp. 3: 173. Type species: Rhynchium ornatum Smith, 1852, monotypy.
Pararrhynchium ornatum bifasciatulum Giordani Soika, 1986
Pararrhynchium ornatum infrenis Giordani Soika, 1973
Pararrhynchium ornatum multifasciatum Giordani Soika, 1986
Pararrhynchium ornatum ornatum (Smith, 1852)
Pararrhynchium ornatum sauteri (von Schulthess, 1934)
Pararrhynchium paradoxum paradoxum (Gussakovskij, 1932)
Pararrhynchium sinense (von Schulthess, 1913)
Pararrhynchium smithii (de Saussure, 1855)
Pararrhynchium taiwanum Kim & Yamane, 2007
Pareumenes de Saussure, 1855
Pareumenes de Saussure, 1855, Ét. Fam. Vesp. 3: 133. Type species: Eumenes quadrispinosus de Saussure, 1855.
Pareumenes (Nortonia) taiwanus (Sonan, 1937)
Pareumenes (Pareumenes) chinensis Liu, 1941
Pareumenes (Pareumenes) obtusus Liu, 1941
Pareumenes (Pareumenes) quadrispinosus acutus Liu, 1941
Pareumenes quadrispinosus conjunctus Liu, 1941
Pareumenes (Pareumenes) quadrispinosus interruptus Liu, 1941
Pareumenes (Pareumenes) quadrispinosus quadrispinosus (de Saussure, 1855)
Pareumenes (Pareumenes) quadrispinosus transitorus Liu, 1941
Parodontodynerus Blüthgen, 1938
Parodontodynerus Blüthgen, 1938 (1937), Konowia 16: 280. Type species: Eumenes ephippium Klug, 1817.
Parodontodynerus laudatus (Kostylev, 1940)
Phimenes Giordani Soika, 1992
Phimenes Giordani Soika, 1992, Lavori Soc. Ven. Sci. Nat. 17: 41, 66, genus, replacement name for Phi de Saussure, 1855, non de Saussure, 1854. Type species: Vespa arcuata Fabricius, 1775.
Phimenes flavopictus flavopictus (Blanchard, 1804)
Phimenes flavopictus formosanus (Zimmermann, 1931)
Phimenes sparsipunctatus Gusenleitner, 2002
Pseudepipona de Saussure, 1856
Pseudepipona de Saussure, 1856, Ét. Fam. Vesp. 3: 309. Type species: Odynerus herrichii de Saussure, 1856, monotypy.
Pseudepipona (Pseudepipona) augusta (Morawitz, 1867)
Pseudepipona (Pseudepipona) herrichii herrichii (de Saussure, 1856)
Pseudepipona (Pseudepipona) lativentris rubricans Kurzenko, 1976
Pseudepipona przewalskyi (Morawitz, 1885)
Pseudonortonia Giordani Soika, 1936
Pseudonortonia Giordani Soika, 1936, Ann. Mus. Civ. Stor. Nat. Genova 59: 268, genus. Type species: Odynerus difformis de Saussure, 1853
Pseudonortonia abbreviaticornis Giordani Soika, 1941
Pseudozumia de Saussure, 1875
Pseudozumia de Saussure, 1875, Smithson. Misc. Coll. 254 (I): 128, division of genus Montezumia de Saussure. Type species: Montezumia indica de Saussure, 1855, monotypy.
Pseudozumia indica indica (de Saussure, 1855)
Pseudozumia indica paulonotata Giordani Soika, 1941
Pseudozumia indosinensis Giordani Soika, 1960
Pseumenes Giordani Soika, 1935
Pseumenes Giordani Soika, 1935, Ann. Mus. Civ. Stor. Nat. Genova 57: 145. Type species: Eumenes eximius Smith, 1861.
Pseumenes depressus depressus (de Saussure, 1855)
Pseumenes imperatrix (Smith, 1857)
Pterocheilus Klug, 1805
Pterocheilus Klug, 1805, Beitr. Naturk. 1: 143. Type species: Vespa phalerata Panzer, 1797.
Pterocheilus albofasciatus Smith, 1878
Pterocheilus napalkovi Kurzenko, 1977
Rhynchium Spinola, 1806
Rhynchium Spinola, 1806, Ins. Ligur. 1: 84. Type species: Rygchium europaeum Spinola, 1806 [= Vespa oculata Fabricius, 1781], monotypy.
Rhynchium brunneum brunneum (Fabricius, 1793)
Rhynchium quinquecinctum quinquecinctum (Fabricius, 1787)
Stenodyneriellus Giordani Soika, 1962
Stenodyneriellus Giordani Soika, 1962 (1961), Boll. Mus. Civ. Stor. Nat. Venezia 14: 65, 71. Type species: Stenodyneriellus turneriellus Giordani Soika, 1962.
Stenodyneriellus depressus Li & Chen, 2016
Stenodyneriellus guttulatus (de Saussure, 1862)
Stenodyneriellus maolanensis Li & Chen, 2016
Stenodyneriellus similiguttulatus Li & Chen, 2016
Stenodynerus de Saussure, 1863
Stenodynerus de Saussure, 1863, Mém. Soc. Phys. Hist. Nat. Genève 17: 228. Type species: Odynerus chinensis de Saussure, 1863.
Stenodynerus baronii Giordani Soika, 1975
Stenodynerus bluethgeni van der Vecht, 1971
Stenodynerus chinensis chinensis (de Saussure, 1863)
Stenodynerus clyppeopictus (Kostylev, 1940)
Stenodynerus copiosus Gusenleitner, 2012
Stenodynerus frauenfeldi (de Saussure, 1867)
Stenodynerus funebris (André, 1884)
Stenodynerus incurvitus Gusenleitner, 2003
Stenodynerus morawitzi Kurzenko, 1977
Stenodynerus morbillosus Giordani Soika, 1979
Stenodynerus nepalensis Giordani Soika, 1985
Stenodynerus ninglangensis Ma & Li, 2016
Stenodynerus nudus (Morawitz, 1889)
Stenodynerus pappi luteifasciatus Kim & Yamane, 2004
Stenodynerus pappi pappi Giordani Soika, 1976
Stenodynerus pullus Gusenleitner, 1981
Stenodynerus reflexus Ma & Li, 2016
Stenodynerus similibaronii Ma & Li, 2016
Stenodynerus taiwanus Kim & Yamane, 2004
Stenodynerus tenuilamellatus Ma & Li, 2016
Stenodynerus tergitus Kim, 1999
Subancistrocerus de Saussure, 1855
Subancistrocerus de Saussure, 1855, Ét. Fam. Vesp. 3: 206. Type species: Odynerus sichelii de Saussure, 1855.
Subancistrocerus camicrus (Cameron, 1904)
Subancistrocerus compressus Li & Chen, 2014
Subancistrocerus jinghongensis Li & Chen, 2014
Subancistrocerus kankauensis (von Schulthess, 1934)
Subancistrocerus sichelii (de Saussure, 1855)
Symmorphus Wesmael, 1836
Symmorphus Wesmael, 1836, Bull. Acad. R. Belg. 3: 45. Type species: Odynerus elegans Wesmael, 1833.
Symmorphus ambotretus Cumming, 1989
Symmorphus angustatus (Zetterstedt, 1838)
Symmorphus apiciornatus (Cameron, 1911)
Symmorphus aurantiopictus Giordani Soika, 1986
Symmorphus bifasciatus (Linnaeus, 1761)
Symmorphus cavatus Li and Chen, 2014
Symmorphus foveolatus Gussakovskij, 1933
Symmorphus fuscipes (Herrich-Schaeffer, 1838)
Symmorphus hoozanensis (von Schulthess, 1934)
Symmorphus lucens (Kostylev, 1938)
Symmorphus mizuhonis Tsuneki, 1977
Symmorphus nigriclypeus Li & Chen, 2014
Symmorphus ornatus Gusenleitner, 2000
Symmorphus sichuanensis Lee, 1981
Symmorphus sublaevis (Kostylev, 1940)
Symmorphus tianchiensis Li & Chen, 2014
Symmorphus violaceipennis Giordani Soika, 1966
Symmorphus yananensis Gusenleitner, 2002
Symmorphus yunnanensis Gusenleitner, 2002
Tropidodynerus Blüthgen, 1939
Tropidodynerus Blüthgen, 1939, Veröff. Deutsch. Kolon. Übersee Mus. Bremen 2: 259, 260. Type species: “Hoplomerus interruptus (Brullé, 1832) = H. mandibularis Morawitz, 1885” [= Polistes interrupta Brullé, 1832].
Tropidodynerus concavus Li & Chen, 2015
Tropidodynerus liupanshanensis Li & Chen, 2015
Zethus Fabricius, 1804
Zethus Fabricius, 1804, Syst. Piez.: xii, 282. Type species: “Zethus coeruleo-pennis Fab.” [= Vespa coeruleopennis Fabricius, 1798].
Zethus dolosus Bingham, 1897
Zethus malayanus Gusenleitner, 2010
Zethus nanlingensis Nguyen & Xu, 2017
Zethus taiwanus Yeh & Lu, 2017
Zehtus velamellatus Tan, 2018
Zethus nigerrimus Gusenleitner, 2001
Acknowledgements
We wish to thank Mr. Christophe Barthélémy (Hong Kong) and two anonymous reviewers for their comments. The first author wishes to thank Dr. Claire Villemant (
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