Checklist |
Corresponding author: David C. Houghton ( david.houghton@hillsdale.edu ) Academic editor: Ana Previšić
© 2018 David C. Houghton, R. Edward DeWalt, Angelica J. Pytel, Constance M. Brandin, Sarah E. Rogers, David E. Ruiter, Ethan Bright, Patrick L. Hudson, Brian J. Armitage.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Houghton DC, DeWalt RE, Pytel AJ, Brandin CM, Rogers SE, Ruiter DE, Bright E, Hudson PL, Armitage BJ (2018) Updated checklist of the Michigan (USA) caddisflies, with regional and habitat affinities. ZooKeys 730: 57-74. https://doi.org/10.3897/zookeys.730.21776
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Based on examination of ~180,000 specimens from 695 collections of 443 localities collected from the 1930s to 2015 we report 295 species of caddisflies from Michigan. Of these, 41 are reported from the state for the first time. Another 18 species previously reported from Michigan are listed as doubtful. The 11 most abundant species collectively represented over half of all specimens collected. Conversely, 80 species were known from <10 specimens, and 27 species from a single specimen. The Michigan fauna is similar to those of Minnesota and Ohio, adjacent states with comparable recent collecting effort. Regional and habitat affinities for each Michigan species are reported herein. Due to the high level of species discovery over the last few years, despite a >80-year collecting history, it is likely that additional species remain undiscovered in the state.
Michigan, Trichoptera , caddisfly, checklist, species, diversity
Despite the ecological importance of caddisflies in aquatic ecosystems and their utility in biological monitoring (
The caddisflies of Michigan are known on a species level primarily from
Past faunal studies of Michigan caddisflies, with resulting numbers of specimens, reported species, and new state records. Nearly all of the specimens associated with these studies were examined during the current study.
Reference | Region | Specimens | Species | State records |
---|---|---|---|---|
|
statewide | ? | 101 | 101 |
|
statewide | ? | N/A | 7 |
|
statewide | 5,767 | 181 | 66 |
|
Houghton Creek | ? | 85 | 0 |
|
St. Clair/Detroit River | ? | 70 | 21 |
|
Manistee River watershed | 26,000+ | 134 | 11 |
|
Isle Royale National Park | 326 | 42 | 2 |
|
Black River Ranch | 38,248 | 117 | 3 |
Current paper | statewide | ~180,000 | 291 | 41 |
We have been collecting caddisflies in Michigan since the 1990s, primarily using ultraviolet light traps for adults. Other adult collecting methods have included malaise trapping, sweep netting, and aspirating from riparian rocks and vegetation. Larval collection methods included kick-netting, hand-picking, and Hess sampling. Most adult collecting took place during June and July, the peak emergence period of caddisflies in central Michigan (
Collecting sites were chosen to yield a geographically representative sample, paying particular attention to unique habitats, such as intermittent streams, waterfalls, and forested wetlands. We collected from sites that appeared to be the least disturbed of their general area. Unique areas, such as the Huron and Porcupine Mountains in the northeastern Upper Peninsula, the Black River Ranch in the northern Lower Peninsula, Sleeping Bear Dunes National Park in the northwestern Lower Peninsula, and Sarah Jane’s Natural Area in the southern Lower Peninsula were sampled more intensively. Most notably, Fairbanks Creek, a pristine small stream in the northern Lower Peninsula, was sampled every week from May to October 2010–2014.
Specimens were identified using
Approximately 180,000 specimens from 695 collections of 443 Michigan localities from the 1930s to 2015 were examined during this study (Figure
The 295 caddisfly species confirmed from Michigan. Taxa are arranged alphabetically by family and genus. The number of species within each family is listed after each respective family. Species reported from Michigan for the first time are in boldface type. Collection data for each newly-reported species are included in Suppl. material
Taxon | spcs | locs | Dep. | NG | NF | SA | Lk | SR | MR | LR |
---|---|---|---|---|---|---|---|---|---|---|
APATANIIDAE (1) | ||||||||||
Apatania zonella (Zetterstedt, 1840) | 49 | 4 | HCIC | 49 | 0 | 0 | 18 | 0 | 0 | 31 |
BRACHYCENTRIDAE (8) | ||||||||||
Brachycentrus americanus (Banks, 1899) | 3547 | 81 | HCIC | 1278 | 2218 | 51 | 6 | 1094 | 2343 | 60 |
B. fuliginosus Walker, 1852 | 25 | 6 |
|
0 | 25 | 0 | 0 | 0 | 25 | 0 |
B. incanus Hagen, 1861 | 1 | 1 | INHS | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
B. lateralis (Say, 1823) | 69 | 4 |
|
0 | 69 | 0 | 0 | 0 | 69 | 0 |
B. numerosus (Say, 1823) | 122 | 21 |
|
9 | 72 | 41 | 0 | 56 | 34 | 31 |
Micrasema charonis Banks, 1914 | 5 | 2 | INHS | 0 | 2 | 3 | 0 | 0 | 2 | 3 |
M. rusticum (Hagen, 1868) | 783 | 56 | HCIC | 24 | 722 | 37 | 4 | 97 | 669 | 13 |
M. wataga Ross, 1938 | 50 | 10 | HCIC | 40 | 10 | 0 | 0 | 3 | 44 | 3 |
DIPSEUDOPSIDAE (1) | ||||||||||
Phylocentropus placidus (Banks, 1905) | 274 | 24 | HCIC | 235 | 17 | 21 | 71 | 86 | 104 | 3 |
GLOSSOSOMATIDAE (8) | ||||||||||
Agapetus hessi Leonard & Leonard, 1949 | 102 | 3 | HCIC | 0 | 102 | 0 | 0 | 0 | 102 | 0 |
A. tomus Ross, 1941 | 63 | 8 | HCIC | 27 | 0 | 36 | 0 | 12 | 49 | 0 |
Glossosoma intermedium Klapálek, 1892 | 16 | 9 | HCIC | 18 | 0 | 0 | 2 | 0 | 14 | 0 |
G. lividum (Hagen, 1861) | 268 | 8 |
|
0 | 268 | 0 | 0 | 9 | 259 | 0 |
G. nigrior Banks, 1911 | 1796 | 68 | HCIC | 179 | 1575 | 42 | 22 | 1355 | 395 | 10 |
Protoptila erotica Ross, 1938 | 138 | 12 | HCIC | 4 | 130 | 4 | 1 | 0 | 63 | 74 |
P. maculata (Hagen, 1861) | 76 | 10 | HCIC | 2 | 5 | 69 | 1 | 4 | 38 | 22 |
P. tenebrosa (Walker, 1852) | 444 | 27 | HCIC | 223 | 220 | 1 | 6 | 197 | 223 | 18 |
GOERIDAE (1) | ||||||||||
Goera stylata Ross, 1938 | 2422 | 12 | HCIC | 30 | 2392 | 0 | 0 | 2361 | 58 | 3 |
HELICOPSYCHIDAE (1) | ||||||||||
Helicopsyche borealis (Hagen, 1861) | 6957 | 114 | HCIC | 281 | 6347 | 310 | 1186 | 1703 | 3161 | 800 |
HYDROPSYCHIDAE (35) | ||||||||||
Arctopsyche ladogensis (Kolenati, 1859) | 16 | 2 |
|
15 | 1 | 0 | 0 | 1 | 15 | 0 |
Cheumatopsyche analis (Banks, 1908) | 1137 | 99 | HCIC | 334 | 485 | 318 | 80 | 263 | 625 | 53 |
C. aphanta Ross, 1938 | 38 | 2 | HCIC | 0 | 37 | 1 | 0 | 0 | 38 | 0 |
C. campyla Ross, 1938 | 6683 | 65 | HCIC | 55 | 257 | 6371 | 56 | 13 | 248 | 6312 |
C. gracilis (Banks, 1899) | 1063 | 64 | HCIC | 248 | 804 | 11 | 2 | 33 | 912 | 114 |
C. minuscula (Banks, 1907) | 2 | 1 | HCIC | 2 | 0 | 0 | 0 | 0 | 0 | 2 |
C. oxa Ross, 1938 | 1609 | 58 | HCIC | 69 | 1425 | 112 | 3 | 1077 | 516 | 8 |
C. pasella Ross, 1941 | 44 | 9 | HCIC | 6 | 37 | 1 | 2 | 1 | 41 | 0 |
C. sordida (Hagen, 1861) | 7 | 4 | HCIC | 5 | 0 | 2 | 0 | 1 | 0 | 5 |
C. speciosa (Banks, 1904) | 61 | 2 | HCIC | 0 | 0 | 61 | 0 | 0 | 0 | 58 |
Diplectrona modesta Banks, 1908 | 1106 | 9 | HCIC | 5 | 1096 | 5 | 0 | 1106 | 0 | 0 |
Hydropsyche aerata Ross, 1938 | 5 | 2 | INHS | 0 | 0 | 4 | 0 | 0 | 0 | 0 |
H. alhedra (Ross, 1939) | 56 | 7 | HCIC | 15 | 40 | 1 | 0 | 5 | 49 | 2 |
H. alternans (Walker, 1852) | 118 | 11 | HCIC | 115 | 2 | 1 | 57 | 2 | 0 | 54 |
H. arinale Ross, 1938 | 1 | 1 | INHS | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
H. betteni Ross, 1938 | 1262 | 67 | HCIC | 74 | 1041 | 147 | 13 | 932 | 266 | 33 |
H. bronta (Ross, 1938) | 192 | 43 | HCIC | 37 | 96 | 59 | 1 | 17 | 141 | 33 |
H. cheilonis (Ross, 1938) | 17 | 8 | HCIC | 0 | 2 | 14 | 0 | 0 | 0 | 4 |
H. cuanis Ross, 1938 | 21 | 4 | INHS | 0 | 0 | 21 | 0 | 7 | 8 | 6 |
H. dicantha Ross, 1938 | 11 | 7 | HCIC | 1 | 6 | 2 | 0 | 2 | 6 | 1 |
H. frisoni Ross, 1938 | 73 | 11 | INHS | 1 | 36 | 32 | 0 | 0 | 67 | 3 |
H. incommoda Hagen, 1861 | 130 | 14 | HCIC | 1 | 12 | 116 | 1 | 2 | 13 | 74 |
H. leonardi Ross, 1938 | 2 | 1 | INHS | 0 | 2 | 0 | 0 | 0 | 2 | 0 |
H. morosa (Hagen, 1861) | 262 | 32 | HCIC | 62 | 162 | 39 | 10 | 18 | 165 | 61 |
H. phalerata Hagen, 1861 | 31 | 5 | HCIC | 0 | 4 | 27 | 0 | 0 | 0 | 23 |
H. placoda Ross, 1941 | 1 | 1 | HCIC | 0 | 0 | 1 | 0 | 0 | 0 | 1 |
H. scalaris Hagen, 1861 | 3 | 3 | INHS | 0 | 0 | 2 | 0 | 0 | 0 | 0 |
H. simulans Ross, 1938 | 26 | 4 | HCIC | 0 | 22 | 4 | 3 | 0 | 3 | 16 |
H. slossonae (Banks, 1905) | 1241 | 68 | HCIC | 137 | 1036 | 68 | 0 | 586 | 646 | 7 |
H. sparna (Ross, 1938) | 2712 | 113 | HCIC | 425 | 2018 | 261 | 12 | 1253 | 1330 | 88 |
H. vexa (Ross, 1938) | 12 | 6 | HCIC | 4 | 8 | 0 | 0 | 4 | 7 | 1 |
H. walkeri (Betten & Mosely, 1940) | 65 | 13 | HCIC | 42 | 22 | 1 | 1 | 1 | 24 | 39 |
Macrostemum zebratum (Hagen, 1861) | 533 | 15 | HCIC | 10 | 499 | 24 | 10 | 2 | 24 | 490 |
Parapsyche apicalis (Banks, 1908) | 220 | 19 | HCIC | 62 | 252 | 6 | 1 | 274 | 45 | 0 |
Potamyia flava (Hagen, 1861) | 119 | 16 | HCIC | 1 | 71 | 47 | 0 | 45 | 31 | 28 |
HYDROPTILIDAE (63) | ||||||||||
Agraylea multipunctata Curtis, 1834 | 4952 | 59 | HCIC | 127 | 927 | 3898 | 195 | 581 | 339 | 31 |
Hydroptila ajax Ross, 1938 | 27 | 3 | HCIC | 0 | 0 | 27 | 0 | 0 | 0 | 19 |
H. albicornis Hagen, 1861 | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
H. amoena Ross, 1938 | 8 | 4 | HCIC | 1 | 7 | 0 | 2 | 0 | 6 | 0 |
H. ampoda Ross, 1941 | 15 | 9 | HCIC | 15 | 0 | 0 | 0 | 0 | 15 | 0 |
H. angusta Ross, 1938 | 45 | 2 | HCIC | 0 | 0 | 45 | 0 | 0 | 0 | 45 |
H. antennopedia Sykora & Harris, 1994 | 111 | 9 | HCIC | 111 | 0 | 0 | 6 | 12 | 93 | 0 |
H. armata Ross, 1938 | 48 | 17 | HCIC | 6 | 13 | 29 | 2 | 2 | 40 | 1 |
H. berneri Ross, 1941 | 1 | 1 |
|
0 | 0 | 1 | 0 | 0 | 0 | 0 |
H. calia Denning, 1948 | 1 | 1 |
|
0 | 0 | 0 | 0 | 0 | 1 | 0 |
H. consimilis Morton, 1905 | 195 | 28 | HCIC | 31 | 140 | 24 | 0 | 62 | 129 | 4 |
H. delineata Morton, 1905 | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
H. grandiosa Ross, 1938 | 20 | 9 | HCIC | 0 | 9 | 11 | 0 | 1 | 9 | 1 |
H. hamata Morton, 1905 | 154 | 27 | HCIC | 46 | 100 | 8 | 62 | 5 | 75 | 4 |
H. jackmanni Blickle, 1963 | 477 | 37 | HCIC | 168 | 270 | 39 | 4 | 191 | 278 | 0 |
H. metoeca Blickle & Morse, 1954 | 166 | 16 | HCIC | 1 | 165 | 0 | 3 | 80 | 82 | 1 |
H. nicoli Ross, 1941 | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
H. novicola Blickle & Morse, 1954 | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
H. perdita Morton, 1905 | 11 | 6 | HCIC | 0 | 3 | 8 | 0 | 1 | 5 | 0 |
H. quinola Ross, 1947 | 3 | 2 | HCIC | 3 | 0 | 0 | 0 | 0 | 3 | 0 |
H. salmo Ross, 1941 | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
H. scolops Ross, 1938 | 18 | 2 |
|
0 | 16 | 2 | 0 | 0 | 16 | 0 |
H. spatulata Morton, 1905 | 9 | 5 | HCIC | 2 | 5 | 2 | 0 | 1 | 4 | 4 |
H. tortosa Ross, 1938 | 7 | 1 | HCIC | 0 | 7 | 0 | 0 | 0 | 7 | 0 |
H. tusculum Ross, 1947 | 1 | 1 | HCIC | 0 | 1 | 0 | 0 | 0 | 1 | 0 |
H. valhalla Denning, 1947 | 90 | 15 | HCIC | 88 | 2 | 0 | 0 | 2 | 87 | 1 |
H. waubesiana Betten, 1934 | 119 | 23 | HCIC | 7 | 36 | 76 | 8 | 17 | 51 | 3 |
H. wyomia Denning, 1948 | 23 | 5 | HCIC | 5 | 15 | 0 | 0 | 0 | 20 | 0 |
H. xera Ross, 1938 | 237 | 19 | HCIC | 189 | 51 | 0 | 2 | 1 | 235 | 2 |
Ithytrichia clavata Morton, 1905 | 222 | 6 | HCIC | 1 | 214 | 7 | 139 | 0 | 75 | 8 |
Leucotrichia pictipes (Banks, 1911) | 30 | 2 | HCIC | 15 | 15 | 0 | 0 | 0 | 21 | 9 |
Mayatrichia ayama Mosely, 1905 | 7 | 2 |
|
0 | 6 | 1 | 0 | 0 | 6 | 0 |
Neotrichia halia Denning, 1948 | 131 | 3 | HCIC | 131 | 0 | 0 | 0 | 0 | 5 | 126 |
N. minutisimella (Chambers, 1873) | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 0 | 1 |
N. okopa Ross, 1939 | 9 | 1 | INHS | 9 | 0 | 0 | 0 | 0 | 9 | 0 |
N. vibrans Ross, 1938 | 1 | 1 | HCIC | 0 | 1 | 0 | 0 | 0 | 0 | 1 |
Ochrotrichia arva (Ross, 1941) | 3 | 2 | HCIC | 0 | 3 | 0 | 0 | 2 | 1 | 0 |
O. spinosa (Ross, 1938) | 220 | 11 | HCIC | 209 | 4 | 7 | 7 | 46 | 161 | 6 |
O. riesi Ross, 1944 | 2 | 1 | INHS | 0 | 2 | 0 | 0 | 2 | 0 | 0 |
O. tarsalis (Hagen, 1861) | 2 | 2 | HCIC | 1 | 0 | 1 | 0 | 0 | 2 | 0 |
Orthotrichia aegerfasciella (Chambers, 1873) | 451 | 21 | HCIC | 1 | 64 | 386 | 38 | 5 | 54 | 5 |
O. balduffi Kingsolver & Ross, 1961 | 97 | 19 | HCIC | 11 | 42 | 44 | 25 | 2 | 36 | 4 |
O. cristata Morton, 1905 | 1813 | 49 | HCIC | 55 | 308 | 1450 | 163 | 71 | 187 | 0 |
O. curta Kingsolver & Ross, 1961 | 13 | 1 | HCIC | 13 | 0 | 0 | 13 | 0 | 0 | 0 |
Oxyethira aeola Ross, 1938 | 44 | 8 | HCIC | 0 | 44 | 0 | 0 | 5 | 39 | 0 |
O. anabola Blickle,1966 | 7 | 5 | HCIC | 5 | 2 | 0 | 0 | 1 | 4 | 2 |
O. araya Ross, 1941 | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
O. coercens Morton, 1905 | 115 | 19 | HCIC | 7 | 101 | 7 | 4 | 29 | 77 | 0 |
O. ecornuta Morton, 1893 | 73 | 3 | HCIC | 2 | 71 | 0 | 73 | 0 | 0 | 0 |
O. forcipata Mosely, 1934 | 170 | 18 | HCIC | 9 | 21 | 140 | 11 | 5 | 20 | 0 |
O. grisea Betten, 1934 | 180 | 5 |
|
0 | 106 | 74 | 50 | 0 | 56 | 0 |
O. itascae Monson & Holzenthal, 1993 | 4 | 4 | HCIC | 0 | 4 | 0 | 0 | 0 | 4 | 0 |
O. michiganensis Mosely, 1934 | 219 | 25 | HCIC | 120 | 99 | 0 | 3 | 32 | 183 | 1 |
O. novasota Ross, 1944 | 2 | 1 |
|
0 | 0 | 2 | 0 | 0 | 0 | 0 |
O. obtatus Denning, 1947 | 27 | 10 | HCIC | 4 | 15 | 8 | 12 | 0 | 5 | 3 |
O. pallida (Ross, 1904) | 757 | 10 | HCIC | 0 | 9 | 748 | 4 | 1 | 28 | 0 |
O. rivicola Blickle & Morse, 1954 | 85 | 7 | HCIC | 2 | 83 | 0 | 1 | 59 | 25 | 0 |
O. serrata Ross, 1938 | 366 | 17 | HCIC | 6 | 319 | 41 | 315 | 0 | 11 | 0 |
O. sida Blickle & Morse, 1954 | 10 | 5 | HCIC | 4 | 6 | 0 | 0 | 2 | 8 | 0 |
O. verna Ross, 1938 | 3 | 3 | HCIC | 1 | 1 | 1 | 0 | 1 | 0 | 0 |
O. zeronia Ross, 1941 | 73 | 12 | HCIC | 5 | 26 | 42 | 25 | 4 | 5 | 0 |
Stactobiella delira (Ross, 1938) | 27 | 4 | HCIC | 24 | 3 | 0 | 0 | 14 | 13 | 0 |
S. palmata (Ross, 1938) | 14 | 4 | HCIC | 5 | 9 | 0 | 0 | 0 | 5 | 9 |
LEPIDOSTOMATIDAE (10) | ||||||||||
Lepidostoma bryanti (Banks, 1908) | 7129 | 38 | HCIC | 157 | 6907 | 65 | 7 | 6822 | 300 | 0 |
L. carrolli Flint, 1958 | 1 | 1 | HCIC | 0 | 1 | 0 | 0 | 1 | 0 | 0 |
L. cinereum (Banks, 1914) | 154 | 5 |
|
8 | 146 | 0 | 1 | 134 | 19 | 0 |
L. costale (Banks, 1914) | 30 | 5 |
|
6 | 24 | 0 | 0 | 13 | 15 | 0 |
L. griseum (Banks, 1911) | 406 | 8 | HCIC | 0 | 406 | 0 | 0 | 405 | 1 | 0 |
L. liba Ross, 1941 | 1 | 1 | INHS | 0 | 1 | 0 | 0 | 1 | 0 | 0 |
L. sackeni (Banks, 1936) | 14 | 7 | HCIC | 2 | 12 | 0 | 0 | 11 | 3 | 0 |
L. togatum (Hagen, 1861) | 5623 | 87 | HCIC | 1167 | 4436 | 2 | 43 | 1022 | 4417 | 123 |
L. unicolor (Banks, 1911) | 4 | 2 | HCIC | 4 | 0 | 0 | 0 | 0 | 4 | 0 |
L. vernale (Banks, 1897) | 116 | 10 | HCIC | 1 | 115 | 0 | 0 | 112 | 1 | 3 |
LEPTOCERIDAE (46) | ||||||||||
Ceraclea alagma (Ross, 1938) | 1058 | 34 | HCIC | 28 | 867 | 163 | 841 | 5 | 52 | 2 |
C. albosticta (Hagen, 1861) | 2 | 1 | HCIC | 0 | 2 | 0 | 2 | 0 | 0 | 0 |
C. alces (Ross, 1941) | 1 | 1 | HCIC | 0 | 1 | 0 | 0 | 0 | 1 | 0 |
C. ancylus (Vorhies, 1909) | 11 | 5 | HCIC | 10 | 0 | 1 | 1 | 0 | 9 | 1 |
C. annulicornis (Martynov, 1910) | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 0 | 1 |
C. arielles (Denning, 1942) | 3637 | 13 | HCIC | 515 | 3122 | 0 | 1 | 28 | 3607 | 1 |
C. cancellata (Betten, 1942) | 163 | 31 | HCIC | 75 | 52 | 35 | 41 | 1 | 71 | 49 |
C. diluta (Hagen, 1861) | 29 | 10 | HCIC | 10 | 8 | 10 | 22 | 0 | 5 | 1 |
C. excisa (Morton, 1904) | 1 | 1 |
|
1 | 0 | 0 | 0 | 0 | 0 | 0 |
C. flava (Ross, 1904) | 39 | 1 | HCIC | 39 | 0 | 0 | 0 | 0 | 0 | 39 |
C. maculata (Banks, 1899) | 143 | 15 | HCIC | 39 | 68 | 36 | 11 | 1 | 100 | 31 |
C. mentiea (Walker, 1852) | 1 | 1 | INHS | 0 | 0 | 1 | 0 | 0 | 0 | 1 |
C. resurgens (Walker, 1852) | 7 | 4 | HCIC | 3 | 3 | 1 | 5 | 0 | 0 | 2 |
C. tarsipunctata (Vorhies, 1909) | 1532 | 18 | HCIC | 113 | 1250 | 126 | 321 | 42 | 533 | 628 |
C. transversa (Hagen, 1861) | 993 | 61 | HCIC | 311 | 666 | 14 | 53 | 17 | 866 | 53 |
C. wetzeli (Ross, 1941) | 30 | 3 | HCIC | 26 | 4 | 0 | 0 | 0 | 4 | 26 |
Leptocerus americanus (Banks, 1899) | 3037 | 85 | HCIC | 120 | 1010 | 1906 | 264 | 123 | 2139 | 365 |
Mystacides interjecta (Banks, 1914) | 1067 | 52 | HCIC | 233 | 806 | 28 | 965 | 25 | 53 | 0 |
M. sepulchralis (Walker, 1852) | 1774 | 102 | HCIC | 385 | 1299 | 89 | 1078 | 17 | 614 | 12 |
Nectopsyche albida (Walker, 1852) | 2572 | 63 | HCIC | 21 | 2347 | 201 | 774 | 111 | 356 | 1278 |
N. candida (Hagen, 1861) | 824 | 15 | HCIC | 4 | 92 | 728 | 90 | 0 | 350 | 383 |
N. diarina (Ross, 1944) | 76 | 16 | HCIC | 9 | 55 | 12 | 20 | 2 | 48 | 1 |
N. exquisita (Walker, 1852) | 226 | 19 | HCIC | 54 | 91 | 81 | 82 | 0 | 8 | 80 |
N. pavida (Hagen, 1861) | 170 | 18 | HCIC | 43 | 125 | 2 | 90 | 1 | 77 | 1 |
Oecetis avara (Banks, 1895) | 5654 | 37 | HCIC | 5321 | 269 | 69 | 76 | 2 | 437 | 5144 |
O. cinerascens (Hagen, 1861) | 812 | 89 | HCIC | 199 | 465 | 148 | 494 | 66 | 144 | 8 |
O. disjuncta (Banks, 1920) | 119 | 9 | HCIC | 28 | 90 | 1 | 0 | 1 | 116 | 2 |
O. ditissa Ross, 1966 | 1 | 1 | INHS | 0 | 1 | 0 | 1 | 0 | 0 | 0 |
O. houghtoni Blahnik & Holzenthal, 2014 | 6 | 2 | HCIC | 4 | 2 | 0 | 6 | 0 | 0 | 0 |
O. immobilis (Hagen, 1861) | 28 | 8 | HCIC | 5 | 23 | 0 | 21 | 3 | 3 | 0 |
O. inconspicua (Walker, 1852) | 16220 | 168 | HCIC | 1383 | 12262 | 2550 | 8727 | 2928 | 2184 | 159 |
O. nocturna Ross, 1966 | 2 | 2 | HCIC | 0 | 1 | 1 | 0 | 0 | 2 | 0 |
O. ochracea Curtis, 1825 | 3 | 2 | INHS | 1 | 0 | 2 | 0 | 0 | 1 | 0 |
O. osteni Milne, 1934 | 444 | 55 | HCIC | 71 | 343 | 30 | 333 | 16 | 72 | 17 |
O. persimilis (Banks, 1907) | 1422 | 68 | HCIC | 365 | 987 | 70 | 40 | 222 | 1085 | 72 |
Setodes incertus (Walker, 1852) | 1543 | 23 | HCIC | 905 | 638 | 0 | 3 | 13 | 1371 | 156 |
S. oligius (Ross, 1938) | 308 | 16 | HCIC | 0 | 262 | 46 | 180 | 2 | 79 | 8 |
Triaenodes abus Milne, 1935 | 125 | 14 | HCIC | 4 | 15 | 106 | 8 | 0 | 10 | 2 |
T. baris Ross, 1938 | 57 | 20 | HCIC | 8 | 43 | 6 | 2 | 29 | 18 | 1 |
T. dipsius Ross, 1938 | 98 | 17 | HCIC | 30 | 68 | 0 | 3 | 8 | 87 | 0 |
T. ignitus (Walker, 1852) | 186 | 35 | HCIC | 9 | 131 | 46 | 3 | 13 | 159 | 11 |
T. injustus (Hagen, 1861) | 535 | 56 | HCIC | 152 | 310 | 68 | 224 | 69 | 197 | 4 |
T. marginatus Sibley, 1926 | 334 | 42 | HCIC | 58 | 197 | 83 | 2 | 69 | 188 | 1 |
T. melacus Ross, 1947 | 6 | 3 | HCIC | 0 | 0 | 6 | 4 | 2 | 0 | 0 |
T. nox Ross, 1941 | 107 | 26 | HCIC | 11 | 89 | 7 | 14 | 56 | 34 | 0 |
T. tardus Milne, 1934 | 1015 | 54 | HCIC | 58 | 399 | 557 | 23 | 165 | 288 | 5 |
LIMNEPHILIDAE (49) | ||||||||||
Anabolia bimaculata (Walker, 1852) | 207 | 42 | HCIC | 72 | 114 | 10 | 57 | 21 | 108 | 1 |
A. consocia (Walker, 1852) | 90 | 27 | HCIC | 14 | 64 | 12 | 3 | 12 | 61 | 1 |
A. ozburni Milne, 1935 | 254 | 5 |
|
0 | 23 | 231 | 0 | 0 | 23 | 0 |
A. sordida Hagen, 1861 | 9 | 6 | INHS | 1 | 6 | 2 | 4 | 0 | 4 | 0 |
Asynarchus montanus (Banks, 1907) | 45 | 8 | HCIC | 2 | 15 | 28 | 3 | 1 | 12 | 0 |
A. rossi Leonard & Leonard, 1949 | 15 | 3 |
|
0 | 15 | 0 | 0 | 15 | 0 | 0 |
Frenesia missa (Milne, 1935) | 159 | 13 |
|
1 | 156 | 2 | 1 | 77 | 77 | 0 |
Glyphopsyche irrorata (F., 1781) | 7 | 4 | HCIC | 2 | 5 | 0 | 0 | 5 | 2 | 0 |
Hesperophylax designatus (Walker, 1852) | 154 | 24 | HCIC | 17 | 126 | 11 | 1 | 119 | 37 | 0 |
Hydatophylax argus (Harris, 1869) | 130 | 35 | HCIC | 6 | 119 | 2 | 1 | 71 | 54 | 0 |
Ironoquia lyrata (Ross, 1938) | 4 | 2 | HCIC | 0 | 4 | 0 | 0 | 4 | 0 | 0 |
I. parvula (Banks, 1900) | 4 | 2 | INHS | 4 | 0 | 0 | 2 | 0 | 2 | 0 |
I. punctatissima (Walker, 1852) | 65 | 7 | HCIC | 0 | 26 | 39 | 0 | 25 | 0 | 0 |
Lenarchus crassus (Banks, 1920) | 2 | 1 | HCIC | 2 | 0 | 0 | 0 | 1 | 0 | 0 |
Leptophylax gracilis Banks, 1900 | 11 | 7 |
|
0 | 6 | 5 | 0 | 1 | 0 | 1 |
Limnephilus ademus Ross, 1941 | 1 | 1 | DER | 1 | 0 | 0 | 0 | 0 | 0 | 0 |
L. argenteus Banks, 1914 | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
L. canadensis Banks, 1808 | 7 | 6 |
|
5 | 2 | 0 | 0 | 1 | 5 | 0 |
L. dispar McLachlan, 1875 | 6 | 2 |
|
0 | 0 | 6 | 0 | 0 | 0 | 0 |
L. externus Hagen, 1861 | 3 | 2 |
|
0 | 3 | 0 | 0 | 0 | 3 | 0 |
L. extractus Walker, 1852 | 1 | 1 | INHS | 1 | 0 | 0 | 0 | 0 | 0 | 0 |
L. hyalinus Hagen, 1861 | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
L. indivisus Walker, 1852 | 473 | 46 | HCIC | 2 | 116 | 351 | 12 | 65 | 32 | 6 |
L. infernalis (Banks, 1914) | 15 | 5 |
|
14 | 0 | 0 | 14 | 0 | 0 | 0 |
L. janus Ross, 1938 | 7 | 1 | HCIC | 7 | 0 | 0 | 7 | 0 | 0 | 0 |
L. moestus Banks, 1908 | 186 | 44 | HCIC | 63 | 91 | 32 | 1 | 59 | 70 | 3 |
L. ornatus Banks, 1907 | 97 | 31 | HCIC | 33 | 23 | 41 | 4 | 15 | 30 | 1 |
L. parvulus (Banks, 1905) | 55 | 3 |
|
3 | 0 | 52 | 2 | 0 | 1 | 0 |
L. perpusillus Walker, 1852 | 25 | 2 |
|
0 | 0 | 25 | 0 | 0 | 0 | 0 |
L. rhombicus (L., 1758) | 62 | 13 | HCIC | 14 | 47 | 1 | 2 | 11 | 49 | 0 |
L. sackeni Banks, 1930 | 4 | 4 |
|
2 | 1 | 2 | 1 | 0 | 2 | 1 |
L. samoedus McLachlan, 1880 | 3 | 2 | DER | 0 | 0 | 0 | 2 | 0 | 0 | 0 |
L. secludens Banks, 1914 | 2 | 2 |
|
0 | 0 | 2 | 0 | 0 | 0 | 0 |
L. sericeus (Say, 1824) | 211 | 16 | HCIC | 12 | 195 | 4 | 8 | 25 | 171 | 0 |
L. submonilifer Walker, 1852 | 529 | 34 | HCIC | 19 | 105 | 405 | 0 | 71 | 32 | 0 |
Nemotaulius hostilis (Hagen, 1873) | 45 | 7 | HCIC | 2 | 43 | 0 | 0 | 42 | 2 | 1 |
Onocosmoecus unicolor (Banks, 1897) | 73 | 14 | HCIC | 45 | 28 | 0 | 0 | 14 | 59 | 0 |
Phanocelia canadensis (Banks, 1924) | 4 | 1 |
|
0 | 0 | 4 | 0 | 0 | 0 | 0 |
Platycentropus amicus (Hagen, 1861) | 15 | 4 | HCIC | 4 | 11 | 0 | 0 | 0 | 11 | 4 |
P. radiatus (Say, 1824) | 386 | 68 | HCIC | 61 | 230 | 79 | 64 | 155 | 79 | 0 |
P. indistinctus (Walker, 1852) | 1 | 1 | HCIC | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
Pseudostenophylax sparsus (Banks, 1908) | 20 | 10 | HCIC | 7 | 13 | 0 | 0 | 11 | 8 | 0 |
Psychoglypha subborealis Ross, 1944 | 5 | 2 |
|
0 | 5 | 0 | 0 | 3 | 2 | 0 |
Pycnopsyche antica (Walker, 1852) | 2191 | 12 | HCIC | 0 | 2191 | 0 | 0 | 2165 | 5 | 0 |
P. guttifera (Walker, 1852) | 1387 | 26 | HCIC | 23 | 1348 | 16 | 9 | 1309 | 50 | 0 |
P. indiana (Ross, 1938) | 13 | 2 | HCIC | 0 | 1 | 12 | 0 | 0 | 13 | 0 |
P. lepida (Hagen, 1861) | 236 | 38 | HCIC | 85 | 136 | 11 | 17 | 92 | 100 | 0 |
P. scabripennis (Rambur, 1842) | 4 | 4 | INHS | 0 | 3 | 1 | 0 | 3 | 1 | 0 |
P. subfasciata (Say, 1828) | 62 | 11 | HCIC | 28 | 3 | 30 | 27 | 0 | 0 | 0 |
MOLANNIDAE (5) | ||||||||||
Molanna blenda Sibley, 1926 | 563 | 29 | HCIC | 48 | 513 | 2 | 4 | 513 | 45 | 1 |
M. flavicornis Banks, 1914 | 67 | 12 | HCIC | 11 | 0 | 0 | 10 | 0 | 1 | 0 |
M. tryphena Betten, 1934 | 198 | 42 | HCIC | 111 | 67 | 15 | 2 | 33 | 156 | 0 |
M. ulmerina Navas, 1934 | 22 | 7 | INHS | 2 | 20 | 0 | 17 | 0 | 0 | 5 |
M. uniophila Vorhies, 1909 | 2027 | 65 | HCIC | 105 | 1850 | 68 | 1915 | 3 | 80 | 0 |
ODONTOCERIDAE (1) | ||||||||||
Psilotreta indecisa (Walker, 1852) | 1 | 1 |
|
1 | 0 | 0 | 0 | 1 | 0 | 0 |
PHILOPOTAMIDAE (6) | ||||||||||
Chimarra aterrima Hagen, 1861 | 549 | 44 | HCIC | 136 | 396 | 14 | 45 | 137 | 323 | 16 |
C. feria (Ross, 1941) | 213 | 8 | HCIC | 0 | 209 | 4 | 0 | 210 | 3 | 0 |
C. obscura (Walker, 1852) | 4488 | 62 | HCIC | 69 | 1093 | 3326 | 65 | 136 | 3773 | 510 |
C. socia (Hagen, 1861) | 8744 | 16 | HCIC | 8678 | 68 | 1 | 2 | 0 | 93 | 8646 |
Dolophilodes distinctus (Walker, 1852) | 1343 | 73 | HCIC | 297 | 1043 | 1 | 10 | 750 | 581 | 0 |
Wormaldia moesta (Banks, 1914) | 8 | 3 | HCIC | 8 | 0 | 0 | 0 | 0 | 8 | 0 |
PHRYGANEIDAE (18) | ||||||||||
Agrypnia colorata (Hagen, 1873) | 3 | 3 |
|
1 | 1 | 0 | 1 | 0 | 1 | 0 |
A. improba (Hagen, 1873) | 147 | 22 | HCIC | 130 | 17 | 0 | 46 | 2 | 99 | 0 |
A. macdunnoughi (Milne, 1931) | 6 | 3 | HCIC | 2 | 0 | 0 | 0 | 0 | 2 | 0 |
A. straminea Hagen, 1873 | 18 | 7 | INHS | 18 | 0 | 0 | 18 | 0 | 0 | 0 |
A. vestita (Walker, 1852) | 49 | 13 | HCIC | 2 | 33 | 14 | 5 | 29 | 4 | 0 |
Banksiola crotchi Banks, 1844 | 2219 | 92 | HCIC | 420 | 1644 | 135 | 352 | 1094 | 609 | 19 |
B. dossuaria (Say, 1828) | 108 | 7 | HCIC | 0 | 108 | 0 | 0 | 106 | 1 | 1 |
B. smithi (Banks, 1914) | 73 | 17 | HCIC | 35 | 21 | 16 | 28 | 0 | 27 | 0 |
Beothukus complicatus (Banks, 1924) | 2 | 2 | ? | 2 | 0 | 0 | 2 | 0 | 0 | 0 |
Fabria inornata (Banks, 1907) | 1 | 1 | ? | 0 | 0 | 1 | 0 | 0 | 0 | 0 |
Hagenella canadensis (Banks, 1907) | 50 | 10 | HCIC | 3 | 16 | 31 | 0 | 10 | 9 | 0 |
Oligostomis ocelligera (Walker, 1852) | 10 | 1 |
|
0 | 0 | 0 | 0 | 0 | 0 | 0 |
Phryganea cinerea Walker, 1852 | 213 | 43 | HCIC | 101 | 101 | 2 | 48 | 62 | 89 | 4 |
P. sayi Milne, 1931 | 31 | 10 | HCIC | 0 | 27 | 4 | 1 | 24 | 4 | 0 |
Ptilostomis angustipennis Hagen, 1873 | 44 | 13 | HCIC | 1 | 36 | 7 | 3 | 33 | 1 | 0 |
P. ocellifera (Walker, 1852) | 375 | 59 | HCIC | 79 | 252 | 44 | 42 | 173 | 126 | 3 |
P. postica (Walker, 1852) | 7 | 6 | HCIC | 0 | 3 | 4 | 1 | 1 | 1 | 0 |
P. semifasciata (Say, 1828) | 207 | 41 | HCIC | 131 | 48 | 22 | 14 | 23 | 161 | 0 |
POLYCENTROPODIDAE (28) | ||||||||||
Cernotina spicata Ross, 1938 | 135 | 11 | HCIC | 1 | 70 | 64 | 64 | 3 | 4 | 0 |
Cyrnellus fraternus (Banks, 1905) | 45 | 8 | HCIC | 5 | 2 | 38 | 15 | 0 | 6 | 22 |
Holocentropus flavus Banks, 1908 | 75 | 14 | HCIC | 4 | 18 | 53 | 3 | 11 | 12 | 0 |
H. interruptus Banks, 1914 | 798 | 47 | HCIC | 47 | 246 | 505 | 156 | 95 | 63 | 2 |
H. melanae Ross, 1938 | 45 | 8 | HCIC | 2 | 17 | 26 | 16 | 0 | 3 | 0 |
H. milaca (Etnier, 1968) | 31 | 1 | HCIC | 0 | 31 | 0 | 31 | 0 | 0 | 0 |
H. picicornis (Stephens, 1836) | 20 | 2 | HCIC | 0 | 0 | 20 | 0 | 0 | 0 | 0 |
Neureclipsis bimaculata (L., 1758) | 42 | 5 | HCIC | 36 | 5 | 1 | 4 | 0 | 28 | 9 |
N. crepuscularis (Walker, 1852) | 276 | 59 | HCIC | 69 | 174 | 32 | 17 | 19 | 197 | 39 |
N. piersoni Frazer & Harris, 1991 | 10 | 5 | INHS | 0 | 8 | 2 | 5 | 2 | 3 | 0 |
N. validus (Walker, 1852) | 2 | 1 | HCIC | 2 | 0 | 0 | 2 | 0 | 0 | 0 |
Nyctiophylax affinis (Banks, 1897) | 4982 | 83 | HCIC | 285 | 3496 | 186 | 3027 | 198 | 501 | 236 |
N. moestus Banks, 1911 | 160 | 17 | HCIC | 46 | 84 | 30 | 1 | 2 | 151 | 0 |
N. serratus Lago & Harris, 1985 | 1 | 1 | INHS | 0 | 1 | 0 | 1 | 0 | 0 | 0 |
Plectrocnemia albipuncta Banks, 1930 | 40 | 14 | HCIC | 35 | 5 | 0 | 3 | 2 | 35 | 0 |
P. aureola Banks, 1930 | 862 | 24 | HCIC | 5 | 2 | 816 | 4 | 16 | 26 | 0 |
P. cinerea (Hagen, 1861) | 1147 | 81 | HCIC | 88 | 778 | 276 | 684 | 37 | 117 | 4 |
P. clinei Milne, 1936 | 53 | 17 | HCIC | 12 | 41 | 0 | 10 | 31 | 12 | 0 |
P. crassicornis (Walker, 1852) | 285 | 14 | HCIC | 2 | 20 | 163 | 2 | 2 | 19 | 0 |
P. icula (Ross, 1941) | 10 | 5 | HCIC | 9 | 1 | 0 | 0 | 0 | 10 | 0 |
P. nascotia (Ross, 1941) | 28 | 2 | HCIC | 0 | 0 | 28 | 0 | 0 | 28 | 0 |
P. remota (Banks, 1911) | 230 | 29 | HCIC | 7 | 51 | 171 | 4 | 30 | 25 | 0 |
P. sabulosa (Leonard & Leonard, 1949) | 3 | 1 |
|
0 | 3 | 0 | 0 | 0 | 0 | 0 |
P. weedi (Blickle & Morse, 1955) | 4 | 4 | HCIC | 0 | 2 | 2 | 0 | 1 | 1 | 0 |
Polycentropus centralis Banks, 1914 | 33 | 2 | HCIC | 33 | 0 | 0 | 0 | 0 | 33 | 0 |
P. confusus Hagen, 1861 | 27 | 11 | HCIC | 7 | 16 | 0 | 0 | 0 | 21 | 1 |
P. pentus Ross, 1941 | 678 | 63 | HCIC | 101 | 541 | 36 | 8 | 354 | 296 | 13 |
P. timesis (Denning, 1948) | 15 | 4 | HCIC | 0 | 15 | 0 | 0 | 15 | 0 | 0 |
PSYCHOMYIIDAE (2) | ||||||||||
Lype diversa (Banks, 1914) | 1589 | 94 | HCIC | 391 | 1101 | 97 | 12 | 449 | 1101 | 27 |
Psychomyia flavida Hagen, 1861 | 10574 | 127 | HCIC | 4291 | 6070 | 207 | 248 | 1102 | 6402 | 2644 |
RHYACOPHILIDAE (8) | ||||||||||
Rhyacophila brunnea Banks, 1911 | 78 | 9 | HCIC | 28 | 50 | 0 | 0 | 44 | 19 | 0 |
R. fuscula (Walker, 1852) | 42 | 14 | HCIC | 421 | 1 | 0 | 0 | 0 | 366 | 56 |
R. glaberrima Ulmer, 1907 | 1 | 1 | INHS | 0 | 1 | 0 | 0 | 0 | 1 | 0 |
R. ledra Ross, 1939 | 1 | 1 | HCIC | 0 | 0 | 1 | 0 | 0 | 1 | 0 |
R. lobifera Betten, 1834 | 5 | 1 | HCIC | 0 | 0 | 5 | 0 | 0 | 0 | 5 |
R. mainensis Banks, 1911 | 29 | 6 | HCIC | 3 | 26 | 0 | 0 | 4 | 24 | 1 |
R. manistee Ross, 1939 | 313 | 15 | HCIC | 0 | 313 | 0 | 0 | 32 | 244 | 37 |
R. vibox Milne, 1936 | 246 | 12 | HCIC | 11 | 233 | 0 | 1 | 236 | 7 | 0 |
SERICOSTOMATIDAE (1) | ||||||||||
Agarodes distinctus (Ulmer, 1905) | 125 | 9 | HCIC | 6 | 18 | 101 | 16 | 0 | 5 | 3 |
Thremmatidae (3) | ||||||||||
Neophylax concinnus McLachlan, 1871 | 185 | 32 | HCIC | 86 | 92 | 7 | 11 | 45 | 127 | 1 |
N. fuscus Banks, 1903 | 54 | 4 |
|
0 | 54 | 0 | 0 | 0 | 54 | 0 |
N. oligius Ross, 1938 | 134 | 18 | HCIC | 19 | 80 | 35 | 0 | 62 | 58 | 4 |
Oecetis inconspicua (Walker) (Leptoceridae) was the most widespread species, followed by Psychomyia flavida Hagen (Psychomyiidae) and Helicopsyche borealis (Hagen) (Helicopsychidae) (Table
Nearly all of the species reported here are based on verified adult male specimens. The current location of these specimens is reported in Table
Species from
Species | Explanation |
---|---|
Banksiola selina Betten, 1944 | Junior synonym of B. crotchi ( |
Ceraclea nepha (Ross, 1944) | Reported from “Crawford”. No specimen located |
Ceraclea punctata (Banks, 1894) | Reported from “Crawford”. No specimen located |
Cyrnellus marginalis (Banks, 1930) | Junior synonym of C. fraternus ( |
Dicosmoecus quadrinotatus (Banks, 1908) | Junior synonym of Onocosmoecus unicolor ( |
Hydropsyche alvata Denning, 1949 | Junior synonym of H. incommoda ( |
Hydropsyche bidens Ross, 1938 | Junior synonym of H. incommoda ( |
Hydropsyche bifida Banks, 1905 | Junior synonym of H. morosa ( |
Hydropsyche orris Ross, 1938 | Junior synonym of H. incommoda ( |
Hydropsyche recurvata Banks, 1908 | Junior synonym of H. alternans ( |
Holocentropus glacialis Ross, 1938 | Misidentified. Is Plectrocnemia cinerea (INHS) |
Lepidostoma strophis Ross, 1938 | Junior synonym of L. cinereum ( |
Neophylax autumnus Vorhies, 1909 | Junior synonym of N. concinnus ( |
Nyctiophylax uncus Ross, 1944 | Misidentified. Is N. affinis (INHS) |
Nyctiophylax vestitus (Hagen, 1861) | Nomen dubium ( |
Platycentropus plectrus Ross, 1938 | Junior synonym of P. amicus ( |
Rhyacophila acropedes Banks, 1914 | Junior synonym of R. brunnea ( |
Rhyacophila melita Ross, 1938 | Junior synonym of R. mainensis ( |
Michigan caddisfly species richness appears similar to that of Minnesota (277 total species) and Ohio (272), two adjacent states where surveys of comparable effort have recently occurred (
Michigan and Minnesota also exhibit similar regional patterns, with higher species richness in the Northern Great Lakes and Northern Forested regions than in the Southern Agricultural region (
Although the majority of Michigan caddisflies have also been collected from adjacent states and provinces, and Michigan does not have any known endemic species, there are still some noteworthy Michigan records reported in this study. Polycentropus timesis (Denning) (Polycentropodidae) is known in Michigan from 4 sites in Lake County in the northwestern Lower Peninsula. These sites are separated by >800 km from the other known P. timesis collection sites in Massachusetts and New Hampshire (
Including the current study, 20% of the total caddisfly fauna of Michigan, and almost 40% of the hydroptilid fauna, has been reported during the last 10 years, despite a >80-year collecting history in the state (Table
We appreciate the efforts of all who have collected, sorted, and identified Michigan caddisflies, including Benjamin Albers, Doug Bidlack, Roger Blahnik, Chris Bowyer, Kelsey Brakel, Kiralyn Brakel, the Friends of the Rogue River, Erin Furmaga, Abigail Fusaro, Hannah (Russell) Goble, Bruce McCulloch, Bilyana (Petkova) McLeod, Bridget O’Leary, Joel Parker, Sally Petrella, Logan Shoup, Mary Clare Smith, Erich Steger, Peter Thistleton, Eleanor (Smith) Valle, Jeff Van Zant, Lydia Wassink, and Daniel Wright. We further thank Abigail Fusaro and the Friends of the Rogue River for sharing larval specimens and genetic sequence data for Rhyacophila lobifera. We thank Mark O’Brien (University of Michigan), Ralph Holzenthal (University of Minnesota), Richard Baumann (Brigham Young University), and Boris Kondratieff (Colorado State University) for allowing us access to additional records and collections.
Permit PRD-SU-2011-039 to collect from Michigan State Park habitats was provided by Alicia Selden of the Michigan Department of Natural Resources. Sampling of Sleeping Bear Dunes National Park was conducted under permit SLBE-2014-SCI-0002, facilitated by Kevin Skerl, Chief of Natural Resources for the park. Permission to sample in the Huron Mountains was provided by the Huron Mountains Wildlife Federation. Permission to sample at the Black River Ranch was provided by the Black River Ranch Board of Directors. Permission to sample at Sarah Jane’s Natural Area was provided by John Bagley and Andrew Bacon of the Michigan Nature Association.
Research costs were supported by the Hillsdale College biology department and several LAUREATES grants from Hillsdale College to DCH, AJP, and affiliated students. RED received support from the US Department of Interior (INT RD X-1-R-1) and the National Science Foundation (DEB 09-18805 ARRA). The valuable comments of Ralph Holzenthal, Karl Kjer, and Andy Rasmussen improved earlier versions of the manuscript. This is paper #17 of the G.H. Gordon BioStation Research Series.
Data file for new state records
Data type: speciemens data