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Catalogue of Tenebrionidae (Coleoptera) of North America
expand article infoYves Bousquet, Donald B. Thomas§, Patrice Bouchard, Aaron D. Smith|, Rolf L. Aalbu, M. Andrew Johnston#, Warren E. Steiner Jr.¤
‡ Agriculture and Agri-Food Canada, Ottawa, Canada
§ USDA ARS, Cattle Fever Tick Research Laboratory, Edinburg, United States of America
| Northern Arizona University, Flagstaff, United States of America
¶ California Academy of Sciences, San Francisco, United States of America
# Arizona State University, Tempe, United States of America
¤ mithsonian Institution, Washington, United States of America
† Deceased author
Open Access

Abstract

This catalogue includes all valid family-group (8 subfamilies, 52 tribes, 14 subtribes), genus-group (349 genera, 86 subgenera), and species-group names (2825 species, 215 subspecies) of darkling beetles (Coleoptera: Tenebrionidae) known to occur in North America1 and their available synonyms. Data on extant, subfossil and fossil taxa are given. For each name the author and year and page number of the description are provided, with additional information (e.g., type species for genus-group names, author of synonymies for invalid taxa) depending on the taxon rank.

Several new nomenclatural acts are included. One new genus, Lepidocnemeplatia Bousquet and Bouchard, is described. Spelaebiosis Bousquet and Bouchard [for Ardoinia Özdikmen, 2004], Blapstinus marcuzzii Aalbu [for Blapstinus kulzeri Marcuzzi, 1977], and Hymenorus campbelli Bouchard [for Hymenorus oculatus Doyen and Poinar, 1994] are proposed as new replacement names. Supporting evidence is provided for the conservation of usage of Tarpela micans (Fabricius, 1798) nomen protectum over Tarpela vittata (Olivier, 1793) nomen oblitum. The generic names Psilomera Motschulsky, 1870 [= Stenomorpha Solier, 1836], Steneleodes Blaisdell, 1909 [= Xysta Eschscholtz, 1829], Ooconibius Casey, 1895 and Euconibius Casey, 1895 [= Conibius LeConte, 1851] are new synonyms (valid names in square brackets). The following 127 new synonymies of species-group names, listed in their original combination, are proposed (valid names, in their current combination, placed in square brackets): Bothrasida mucorea Wilke, 1922 [= Pelecyphorus guanajuatensis (Champion, 1884)]; Parasida zacualpanicola Wilke, 1922 [= Pelecyphorus asidoides Solier, 1836]; Stenosides kulzeri Pallister, 1954, Stenosides bisinuatus Pallister, 1954, and Parasida trisinuata Pallister, 1954 [= Pelecyphorus dispar (Champion, 1892)]; Asida favosa Champion, 1884 and Asida similata Champion, 1884 [= Pelecyphorus fallax (Champion, 1884)]; Ologlyptus bicarinatus Champion, 1884 [= Pelecyphorus indutus (Champion, 1884)]; Parasida laciniata Casey, 1912 and Parasida cristata Pallister, 1954 [= Pelecyphorus liratus (LeConte, 1854)]; Parasida esperanzae Wilke, 1922 and Parasida mixtecae Wilke, 1922 [= Pelecyphorus longipennis (Champion, 1884)]; Parasida tolucana Casey, 1912 [= Pelecyphorus scutellaris (Champion, 1884)]; Parasida purpusi Wilke, 1922 [= Pelecyphorus tristis (Champion, 1884)]; Astrotus nosodermoides Champion, 1892 [= Pelecyphorus erosus (Champion, 1892)]; Astrotus seticornis var. humeralis Champion, 1884 [= Pelecyphorus seticornis (Champion, 1884)]; Pactostoma breviuscula Casey, 1912, Pactostoma exoleta Casey, 1912, Pactostoma luteotecta Casey, 1912, Pactostoma monticola Casey, 1912, Pactostoma obtecta Casey, 1912, and Pactostoma sigillata Casey, 1912 [=Pelecyphorus anastomosis (Say, 1824)]; Ologlyptus canus Champion, 1884 and Ologlyptus sinuaticollis Champion, 1884 [= Pelecyphorus graciliformis (Solier, 1836)]; Gonasida elata reducta Casey, 1912, Gonasida elata prolixa Casey, 1912, and Gonasida aucta Casey, 1912 [= Philolithus elatus compar (Casey, 1912)]; Gonasida alaticollis Casey, 1912 [= Philolithus elatus difformis (LeConte, 1854)]; Gonasida gravida Casey, 1912 [= Philolithus elatus elatus (LeConte, 1853)]; Pelecyphorus aegrotus limbatus Casey, 1912 [= Philolithus aegrotus aegrotus (LeConte, 1861)]; Pelecyphorus corporalis Casey, 1912, Pelecyphorus reptans Casey, 1912, Pelecyphorus socer Casey, 1912, Pelecyphorus abscissus Casey, 1912, Pelecyphorus fumosus Casey, 1912, Pelecyphorus parvus Casey, 1912, Pelecyphorus morbillosus pacatus Casey, 1912, Pelecyphorus morbillosus sobrius Casey, 1912, Pelecyphorus piceus Casey, 1912, Pelecyphorus piceus crudelis Casey, 1912, Pelecyphorus snowi Casey, 1912, and Pelecyphorus subtenuis Casey, 1912 [= Philolithus morbillosus (LeConte, 1858)]; Bothrasida sanctae-agnae Wilke, 1922 [= Stenomorpha funesta (Champion, 1884)]; Asida flaccida Horn, 1896 [= Stenomorpha embaphionides (Horn, 1894)]; Asida angustula Casey, 1890, Stethasida stricta Casey, 1912, Stethasida muricatula languida Casey, 1912, Stethasida pertinax Casey, 1912, Stethasida socors Casey, 1912, Stethasida angustula inepta Casey, 1912, Stethasida tenax Casey, 1912, and Stethasida vegrandis Casey, 1912 [= Stenomorpha muricatula (LeConte, 1851)]; Stethasida obsoleta expansa Casey, 1912, Stethasida obsoleta opacella Casey, 1912, Stethasida brevipes Casey, 1912, Stethasida torpida Casey, 1912, Stethasida convergens Casey, 1912, Stethasida discreta Casey, 1912, Stethasida longula Casey, 1912, Stethasida adumbrata Casey, 1912, Stethasida occulta Casey, 1912, Stethasida tarsalis Casey, 1912, Stethasida unica Casey, 1912, and Pelecyphorus laevigatus Papp, 1961 [= Stenomorpha obsoleta (LeConte, 1851)]; Trichiasida eremica Wilke, 1922 [= Stenomorpha difficilis (Champion, 1884)]; Trichiasida lineatopilosa Casey, 1912 [= Stenomorpha hirsuta (LeConte, 1851)]; Trichiasida tenella Casey, 1912 [= Stenomorpha hispidula (LeConte, 1851)]; Trichiasida duplex Casey, 1912 [= Stenomorpha villosa (Champion, 1884)]; Alaudes squamosa Blaisdell, 1919, Alaudes testacea Blaisdell, 1919, and Alaudes fallax Fall, 1928 [= Alaudes singularis Horn, 1870]; Edrotes barrowsi Dajoz, 1999 [=Edrotes ventricosus LeConte, 1851]; Nyctoporis tetrica Casey, 1907 and Nyctoporis maura Casey, 1907 [= Nyctoporis aequicollis Eschscholtz, 1831]; Nyctoporis pullata Casey, 1907 [= Nyctoporis sponsa Casey, 1907]; Eleodes tibialis forma oblonga Blaisdell, 1909 [= Eleodes tibialis Blaisdell, 1909]; Eleodes (manni var.) variolosa Blaisdell, 1917 [= Eleodes constrictus LeConte, 1858]; Eleodes cordata forma sublaevis Blaisdell, 1909, Eleodes cordata forma intermedia Blaisdell, 1909, Eleodes cordata forma oblonga Blaisdell, 1909, Eleodes cordata forma elongata Blaisdell, 1909, and Eleodes (cordata var.) adulterina Blaisdell, 1917 [= Eleodes cordata Eschscholtz, 1829]; Eleodes hornii var. monticula Blaisdell, 1918 and Eleodes manni sierra Blaisdell, 1925 [= Eleodes fuchsii Blaisdell, 1909]; Eleodes parvicollis var. squalida Blaisdell, 1918 [= Eleodes parvicollis Eschscholtz, 1829]; Eleodes reflexicollis Mannerheim, 1843 and Eleodes parvicollis forma farallonicaBlaisdell, 1909 [= Eleodes planata Eschscholtz, 1829]; Eleodes indentata Blaisdell, 1935 [= Eleodes rotundipennis LeConte, 1857]; Eleodes intricata Mannerheim, 1843 [= Eleodes scabrosa Eschscholtz, 1829]; Eleodes horni fenyesi Blaisdell, 1925 [= Eleodes tenebrosa Horn, 1870]; Eleodes cordata var. horrida Blaisdell, 1918 [= Eleodes tuberculata Eschscholtz, 1829]; Eleodes oblonga Blaisdell, 1933 [= Eleodes versatilis Blaisdell, 1921]; Eleodes dentipes marinae Blaisdell, 1921 [= Eleodes dentipes Eschscholtz, 1829]; Eleodes carbonaria forma glabra Blaisdell, 1909 [= Eleodes carbonaria carbonaria (Say, 1824)]; Eleodes granosa forma fortis Blaisdell, 1909 [= Eleodes granosa LeConte, 1866]; Eleodes pilosa forma ordinata Blaisdell, 1909 [= Eleodes pilosa Horn, 1870]; Trogloderus costatus pappi Kulzer, 1960 [= Trogloderus tuberculatus Blaisdell, 1909]; Trogloderus costatus mayhewi Papp, 1961 [= Trogloderus vandykei La Rivers, 1946]; Bolitophagus cristatus Gosse, 1840 [= Bolitotherus cornutus (Fabricius, 1801)]; Eleates explanatus Casey, 1890 [= Eleates depressus (Randall, 1838)]; Blapstinus sonorae Casey, 1890 [= Blapstinus brevicollis LeConte, 1851]; Blapstinus falli Blaisdell, 1929 [= Blapstinus castaneus Casey, 1890]; Blapstinus brunneus Casey, 1890 and Blapstinus coronadensis Blaisdell, 1892 [=Blapstinus histricus Casey, 1890]; Blapstinus hesperius Casey, 1890 [=Blapstinus intermixtus Casey, 1890]; Blapstinus cinerascens Fall, 1929 [= Blapstinus lecontei Mulsant and Rey, 1859]; Blapstinus niger Casey, 1890 and Blapstinus cribricollis Casey, 1890 [= Blapstinus pimalis Casey, 1885]; Blapstinus arenarius Casey, 1890 [= Blapstinus pratensis LeConte, 1859]; Blapstinus gregalis Casey, 1890 [= Blapstinus substriatus Champion, 1885]; Blapstinus hydropicus Casey, 1890 [= Blapstinus sulcatus LeConte, 1851]; Blapstinus hospes Casey, 1890 [= Blapstinus vestitus LeConte, 1859]; Notibius reflexus Horn, 1894 [= Conibius opacus (LeConte, 1866)]; Notibius affinis Champion, 1885 [=Conibius rugipes (Champion, 1885)]; Conibius parallelus LeConte, 1851 [= Conibius seriatus LeConte, 1851]; Nocibiotes rubripes Casey, 1895 [=Nocibiotes caudatus Casey, 1895]; Nocibiotes gracilis Casey, 1895 and Nocibiotes acutus Casey, 1895 [=Nocibiotes granulatus (LeConte, 1851)]; Conibius alternatus Casey, 1890 [= Tonibius sulcatus (LeConte, 1851)]; Pedinus suturalis Say, 1824 [= Alaetrinus minimus (Palisot de Beauvois, 1817)]; Menedrio longipennis Motschulsky, 1872 [= Tenebrio obscurus Fabricius, 1792]; Hymenophorus megops Hatch, 1965 and Telesicles magnus Hatch, 1965 [= Hymenorus sinuatus Fall, 1931]; Andrimus concolor Casey, 1891 and Andrimus convergens Casey, 1891 [= Andrimus murrayi (LeConte, 1866)]; Mycetochara marshalli Campbell, 1978 [= Mycetochara perplexata Marshall, 1970]; Phaleria globosa LeConte, 1857 [= Phaleria picta Mannerheim, 1843]. The following subspecies of Trogloderus costatus LeConte, 1879 are given species rank: Trogloderus nevadus La Rivers, 1943, Trogloderus tuberculatus Blaisdell, 1909, and Trogloderus vandykei La Rivers, 1946. The following taxa, previously thought to be junior synonyms, are considered valid: Amphidora Eschscholtz, 1829; Xysta Eschscholtz, 1829; Helops confluens (Casey, 1924). Two new combinations are proposed: Stenomorpha spinimana (Champion, 1892) and Stenomorpha tenebrosa (Champion, 1892) [from the genus Parasida Casey, 1912]. The type species [placed in square brackets] of the following 12 genus-group taxa are designated for the first time: Lagriola Kirsch, 1874 [Lagriola operosa Kirsch, 1874]; Locrodes Casey, 1907 [Emmenastus piceus Casey, 1890]; Falacer Laporte, 1840 [Acanthopus cupreus Laporte, 1840 (= Helops contractus Palisot de Beauvois, 1812)]; Blapylis Horn, 1870 [Eleodes cordata Eschscholtz, 1829]; Discogenia LeConte, 1866 [Eleodes scabricula LeConte, 1858]; Metablapylis Blaisdell, 1909 [Eleodes nigrina LeConte, 1858]; Steneleodes Blaisdell, 1909 [Eleodes longicollis LeConte, 1851]; Scaptes Champion, 1886 [Scaptes squamulatus Champion, 1886 (= Asida tropica Kirsch, 1866)]; Aspidius Mulsant and Rey, 1859 [Blaps punctata Fabricius, 1792]; Cryptozoon Schaufuss, 1882 [Cryptozoon civile Schaufuss, 1882]; Halophalerus Crotch, 1874 [Phaleria rotundata LeConte, 1851]; Dignamptus LeConte, 1878 [Dignamptus stenochinus LeConte, 1878]. Two species previously known from South America [Nilio lebasi J. Thomson and Platydema erotyloides Chevrolat] are reported for the first time from North America.

Resumen

Este catálogo incluye todos los grupos válidos de nivel famili (8 subfamilias, 52 tribus, 14 subtribus), de nivel género (349 géneros, 86 subgéneros) y de nivel especie (2825 especies, 215 subespecies) de tenebriónidos (Coleoptera: Tenebrionidae) presentes en América del Norte y sus sinónimos disponibles. Se dan datos sobre los táxones existentes, subfósiles y fósiles. Para cada nombre, el autor, el año y el número de página de la descripción se complementan con información adicional dependiendo de la categoría del taxon (por ejemplo, especies tipo para nombres de nivel género, autor de sinonimias para táxones no válidos).

Se incluyen varios nuevos actos nomenclaturales. Un nuevo género, Lepidocnemeplatia Bousquet y Bouchard, se describe. Spelaebiosis Bousquet y Bouchard [para Ardoinia Özdikmen, 2004], Blapstinus marcuzzii Aalbu [para Blapstinuskulzeri Marcuzzi, 1977] e Hymenorus campbelli Bouchard [para Hymenorus oculatus Doyen y Poinar, 1994] se proponen como nuevos nombres de reemplazo. Se proporciona evidencia de uso para la conservación de Tarpela micans (Fabricius, 1798) nomen protectum sobre Tarpela vittata (Olivier, 1793) nomen oblitum. Los nombres genéricos Psilomera Motschulsky, 1870 [= Stenomorpha Solier, 1836], Steneleodes Blaisdell, 1909 [= Xysta Eschscholtz, 1829], Ooconibius Casey, 1895 y Euconibius Casey, 1895 [= Conibius LeConte, 1851] sonnuevos sinónimos (nombres válidos en corchetes). Se proponen las siguientes 127 nuevas sinonimias de nombres de nivel especie (ver lista en el «Resumen»), enumerados en su combinación original (nombres válidos, listados en su combinación actual, colocados entre corchetes). Las siguientes subespecies de Trogloderus costatus LeConte, 1879 reciben categoría de especie: Trogloderus nevadus La Rivers, 1943, Trogloderus tuberculatus Blaisdell, 1909 y Trogloderus vandykei La Rivers, 1946. Los siguientes táxones, que anteriormente se consideraban sinónimos más modernos, se consideran válidos: Amphidora Eschscholtz, 1829, Xysta Eschscholtz, 1829, Helops confluens (Casey, 1924). Se proponen dos nuevas combinaciones: Stenomorpha spinimana (Champion, 1892) y Stenomorpha tenebrosa (Champion, 1892) [del género Parasida Casey, 1912]. Las especies tipo [colocadas entre corchetes] de los siguientes 12 táxones del nivel génerosse designan por primera vez: Lagriola Kirsch, 1874 [Lagriola operosa Kirsch, 1874]; Locrodes Casey, 1907 [Emmenastus piceus Casey, 1890]; Falacer Laporte, 1840 [Acanthopus cupreus Laporte, 1840 (= Helops contractus Palisot de Beauvois, 1812)]; Blapylis Horn, 1870 [Eleodes cordata Eschscholtz, 1829]; Discogenia LeConte, 1866 [Eleodes scabricula LeConte, 1858]; Metablapylis Blaisdell, 1909 [Eleodes nigrina LeConte, 1858]; Steneleodes Blaisdell, 1909 [Eleodes longicollis LeConte, 1851]; Scaptes Champion, 1886 [Scaptes squamulatus Champion, 1886 (= Asida tropica Kirsch, 1866)]; Aspidius Mulsant y Rey, 1859 [Blaps punctata Fabricius, 1792]; Cryptozoon Schaufuss, 1882 [Cryptozoon civile Schaufuss, 1882]; Halophalerus Crotch, 1874 [Phaleria rotundata LeConte, 1851]; Dignamptus LeConte, 1878 [Dignamptus stenochinus LeConte, 1878]. Dos especies previamente conocidas de América del Sur [Nilio lebasi J. Thomson y Platydema erotyloides Chevrolat] se mencionan por primera vez de América del Norte.

Résumé

Ce catalogue liste tous les noms de niveau famille (8 sous-familles, 52 tribus, 14 sous-tribus), genre (349 genres, 86 sous-genres), et espèce (2825 espèces, 215 sous-espèces) de Tenebrionidae (Coleoptera:) recensés en Amérique du Nord ainsi que leurs synonymes disponibles. Le catalogue inclut les taxons actuels, subfossiles et fossiles. L’auteur ainsi que l’année et la pagination de la description sont inclus pour chaque nom scientifique et, pour certains de ces noms, des renseignements additionnels (par exemple, l’espèce-type pour les noms de niveau genre, l’auteur de la synonymie des noms invalides des niveaux genre et espèce).

Plusieurs nouveaux actes nomenclaturaux sont proposés. On décrit un nouveau genre, Lepidocnemeplatia Bousquet et Bouchard. Spelaebiosis Bousquet et Bouchard [pour Ardoinia Özdikmen, 2004], Blapstinus marcuzzii Aalbu [pour Blapstinus kulzeri Marcuzzi, 1977] et Hymenorus campbelli Bouchard [pour Hymenorus oculatus Doyen and Poinar, 1994] sont des nouveaux noms de remplacement. On fournit les évidences nécessaires pour conserver l’usage actuel du nom Tarpela micans (Fabricius, 1798) nomen protectum sur celui de Tarpela vittata (Olivier, 1793) nomen oblitum. Les noms génériques Psilomera Motschulsky, 1870 [= Stenomorpha Solier, 1836], Steneleodes Blaisdell, 1909 [= Xysta Eschscholtz, 1829], Ooconibius Casey, 1895 et Euconibius Casey, 1895 [= Conibius LeConte, 1851] sont des nouveaux synonymes (noms valides placés entre crochets). On propose 127 nouveaux synonymes de noms de niveau espèce (voir la liste dans « Abstract »), listés par leur combinaison originelle (noms valides, listés par leur combinaison actuelle, placés entre crochets). On élève au rang d’espèce les sous-espèces suivantes de Trogloderus costatus LeConte, 1879: Trogloderus nevadus La Rivers, 1943, Trogloderus tuberculatus Blaisdell, 1909, et Trogloderus vandykei La Rivers, 1946. On redonne le statut de taxons valides aux entités suivantes considérées à tort comme invalides: Amphidora Eschscholtz, 1829, Xysta Eschscholtz, 1829, et Helops confluens (Casey, 1924). On propose les combinaisons nouvelles suivantes: Stenomorpha spinimana (Champion, 1892) et Stenomorpha tenebrosa (Champion, 1892) [auparavant placés dans le genre Parasida Casey, 1912]. On désigne pour la première fois l’espèce type [placée entre crochets] des 12 noms suivants de niveau genre: Lagriola Kirsch, 1874 [Lagriola operosa Kirsch, 1874]; Locrodes Casey, 1907 [Emmenastus piceus Casey, 1890]; Falacer Laporte, 1840 [Acanthopus cupreus Laporte, 1840 (= Helops contractus Palisot de Beauvois, 1812)]; Blapylis Horn, 1870 [Eleodes cordata Eschscholtz, 1829]; Discogenia LeConte, 1866 [Eleodes scabricula LeConte, 1858]; Metablapylis Blaisdell, 1909 [Eleodes nigrina LeConte, 1858]; Steneleodes Blaisdell, 1909 [Eleodes longicollis LeConte, 1851]; Scaptes Champion, 1886 [Scaptes squamulatus Champion, 1886 (= Asida tropica Kirsch, 1866)]; Aspidius Mulsant and Rey, 1859 [Blaps punctata Fabricius, 1792]; Cryptozoon Schaufuss, 1882 [Cryptozoon civile Schaufuss, 1882]; Halophalerus Crotch, 1874 [Phaleria rotundata LeConte, 1851]; Dignamptus LeConte, 1878 [Dignamptus stenochinus LeConte, 1878]. Deux espèces, auparavant documentées seulement de l’Amérique du Sud [Nilio lebasi J. Thomson and Platydema erotyloides Chevrolat], sont rapportées pour la première fois d’Amérique du Nord.

Keywords

North America, nomenclature, Lepidocnemeplatia , new genus, new replacement names

Introduction

Darkling beetles (Coleoptera: Tenebrionidae) form a species-rich and morphologically diverse family with approximately 2300 genera and 20000 species worldwide (Matthews et al. 2010), and many more taxa to be described. The first thorough classification of the Tenebrionidae was provided by Lacordaire (1859) and was based entirely on the external morphology of adults. With relatively few exceptions, his classification schema was followed by subsequent workers for approximately 100 years (Watt 1967). The family classification was eventually reviewed using morphological characters of immature stages (Watt 1975) although significant changes did not appear until the first comprehensive investigation of adult internal structures including defense glands, female ovipositor, and female genital tube (Tschinkel and Doyen 1980, Doyen and Tschinkel 1982). The first higher-level phylogeny of the family based on molecular data was published only recently (Kergoat et al. 2014). As a result of these comparative and phylogenetic studies, several taxa previously treated as separate families (e.g., Lagriidae, Alleculidae, Nilionidae) are now included within Tenebrionidae. Additionally, many taxa previously included in Tenebrionidae are now classified in other families (see Table 1 for North American genera, Aalbu (2006) for worldwide taxa).

Table 1.

List of North American genera previously included in Tenebrionidae but currently classified in another family.

Genus Current placement
Boros Herbst, 1797 Boridae
Dacoderus LeConte, 1858 Salpingidae
Megazopherus Casey, 1907 Zopheridae
Meralius Casey, 1907 Zopheridae
Noserodes Casey, 1907 Zopheridae
Noserus LeConte, 1862 Zopheridae
Nosoderma Solier, 1841 Zopheridae
Phellopsis LeConte, 1862 Zopheridae
Phloeodes LeConte, 1862 Zopheridae
Sesaspis Casey, 1907 Zopheridae
Usechus Motschulsky, 1845 Zopheridae
Verodes Casey, 1907 Zopheridae
Zopherinus Casey, 1907 Zopheridae
Zopherodes Casey, 1907 Zopheridae
Zopherus Laporte, 1840 Zopheridae

The aim of this work is to synthesize available taxonomic, nomenclatural, and distributional information for all darkling beetles known from North America.

Methods

Nomenclatural data

All nomenclaturally available family-, genus- and species-group names are included. Extant taxa and subfossils from the Pleistocene (see Doyen and Miller 1980, Doyen and Poinar 1994) are given in the main catalogue. Impression fossils from major North American deposits are listed in Appendix 1, although the taxonomic assignment of these often-fragmentary fossils needs to be confirmed. Fossil species described from amber are listed in Appendix 2. Taxa incorrectly recorded from North America are given in Appendix 3. Subfamilies are listed in a phylogenetic framework but valid tribal, generic, and specific names are given in alphabetic order; listings of all invalid names are chronological.

The author and year and page number of the original description are provided for each scientific name. The type genus for each family-group name and the type species and type fixation for each genus-group name are included. The reference in which a given generic or specific name is first placed in synonymy with the current valid name is listed. Type-species designations in Lucas (1920) were accepted when a single species was listed under a particular genus-group name (see Alonso-Zarazaga and Lyal 2009, Bousquet et al. 2015). For genera with valid subgenera, synonyms are given under the nominotypical subgenus when relevant. Every species-group name are listed in its original combination, as given in the publication even if the agreement in gender with the generic name is incorrect. In cases where a species-group name older than the one currently recognized as valid is available but of doubtful application (e.g., Latridius pubescens Say, 1826), we have retained usage of the younger, accepted name as valid and treated the older name as nomen dubium. The synonymy list for adventive species focuses primarily on names used in the North American literature, other sources (e.g., Löbl et al. 2008a) can be consulted for data on all invalid names.

The classification used follows Bouchard et al. (2011) but also includes corrections and subsequent additions.

The gender of all valid genera listed in the catalogue has been determined following the provision of Article 30 (ICZN 1999) and indicated after the name using the initials M [Masculine], F [Feminine], and N [Neuter]. Therefore, the gender of the generic names Liodema, Platydema, and Scaphidema is herein treated as feminine following Article 30.1.3 (ICZN 1999; see Kerzhner (2003) and Löbl and Smetana (2010: 34) for further comments); the ending is derived from the Greek “demas (body silhouette).” The gender of Alaudes is treated as masculine following Article 30.2.4 (ICZN 1999). The gender of Eleodes, originally treated as feminine by Eschscholtz (1829), was changed to masculine by Somerby and Doyen (1976) and has been followed subsequently in the literature. However, Eleodes is feminine following Article 30.1.4.4 (ICZN 1999) which says that “a compound genus-group name ending in the suffix ... -odes is to be treated as masculine unless its author, when establishing the name, stated that it had another gender or treated it as such by combining it with an adjectival species-group name in another gender form.”

If necessary, the ending of all valid species-group names has been modified according to the gender of the generic name with which the species is currently combined. All specific names that are nouns in apposition need not agree in gender with the generic name and retain their original endings.

The author(s) of every new nomenclatural act proposed in the catalogue is given in square brackets (e.g., “[ADS]”) except for the first typification of genus-group names. One of these new acts needs further development. The genus-group taxon Lepidocnemeplatia was proposed by Kaszab (1938: 80) as a subgenus of Cnemeplatia Costa, 1847 to include two species, Cnemeplatia laticollis Champion, 1885 and C. sericea Horn, 1870. Unfortunately, Kaszab did not designate a type species for his new genus and therefore the name is unavailable from that date (ICZN 1999: Article 13.3). Löbl and Merkl (2003: 245) designated C. sericea Horn, 1870 as type species of Lepidocnemeplatia, the first typification for the taxon, and subsequently Löbl et al. (2008a: 140) credited authorship of the name to Löbl and Merkl (2003). However, since Löbl and Merkl (2003: 245) failed to indicate that they were establishing a new nominal taxon, a mandatory requirement for all new names published after 1999 (ICZN 1999: Article 16.1), the name cannot be attributed to them and is still a nomen nudum. In order to make the name available we here proposed the generic name Lepidocnemeplatia2 Bousquet and Bouchard, new genus; type species (here designated): Cnemeplatia sericea Horn, 1870. The reader is referred to Kaszab (1938: 79–80, couplet 1" of his key) for a description of the character states that differentiate the taxon (ICZN 1999: Article 13.1.2). A summary of all new nomenclatural acts is available in the Abstract.

Distributional data

This catalogue documents all species of Tenebrionidae from Greenland, Alaska, and Canada south to Panama, and also includes islands of the West Indies. Records from the Netherlands Antilles (consisting of several islands in the West Indies), the Venezuelan island Margarita, and Trinidad and Tobago off the northeast coast of Venezuela are not included since the fauna of these islands is more closely affiliated with the South American fauna. When known, the states (for Mexico and continental United States) and the provinces and territories (for Canada) are listed in parentheses for each species. For West Indian records, we give the political unit names for species found in the Lucayan Archipelago and Greater Antilles, occasionally with specific islands in parenthesis; political unit names are not provided for species occurring in the Lesser Antilles (LAN) and Virgin Islands (VIS), though sometimes we include specific islands in parentheses, especially when the species is known from only one or two islands. Further details regarding the West Indies geographical units can be found in Ivie and Hart (2016: Table 1).

Many Mexican state records in this catalogue came from localities listed by Champion in volume IV, parts 1 and 2, of the Biologia Centrali-Americana [1884–1893] and the gazetteer of Selander and Vaurie (1962) was used consistently to associate localities with states (including the federal district).

Amongst the geographical units used (see list below), HIS (for Hispaniola) is used when only the island record is known and LC (for Lower California) when only the peninsula record is known. South America (SA) is placed at the end of a species record list to indicate that the species extends into South America. Distributional records listed in square brackets (e.g., “[NM]”) are considered doubtful. South American species introduced accidentally into North America are indicated with a subscript “i” beside the country record (i.e., “USAi”).

Bibliographic data

References are provided for all scientific names included if a page number (related to the description of the taxon) is provided after the year of publication. Based on evidence previously published (see Bousquet 2016a: 211, 265), the dates of publication of Germar’s Coleopterorum species novae dated 1824 on its title page, and of Hope’s The Coleopterist’s manual, part the third, dated 1840 on its title page, are given as 1823 and 1841 respectively.

As discussed by Bousquet (2016b), Mäklin's “Monographie der Gattung Strongylium” was published in 1867, not 1864 as given by several authors.

List of acronyms used for geographic units3

BAH Bahamas

BEL Belize

BER Bermuda

CAN Canada [AB: Alberta; BC: British Columbia; MB: Manitoba; NB: New Brunswick; NF: Newfoundland and Labrador; NS: Nova Scotia; NT: Northwest Territories; NU: Nunavut; ON: Ontario; PE: Prince Edward Island; QC: Quebec; SK: Saskatchewan; YT: Yukon Territory]

CAY Cayman Islands

CRI Costa Rica

CUB Cuba

DOM Dominican Republic

GRE Greenland

GUA Guatemala

HAI Haiti

HIS Hispaniola

HON Honduras

JAM Jamaica

LAN Lesser Antilles (including among others Anguilla, Antigua and Barbuda, Montserrat, Guadeloupe, Dominica, Martinique, Saint Lucia, Grenada, Barbados)

LC Lower California

MEX Mexico [AG: Aguascalientes; BC: Baja California; BS: Baja California Sur; CA: Campeche; CH: Chihuahua; CI: Chiapas; CL: Colima; CO: Coahuila; DU: Durango; FD: Federal District; GE: Guerrero; GU: Guanajuato; HI: Hidalgo; JA: Jalisco; ME: México; MI: Michoacán; MO: Morelos; NA: Nayarit; NL: Nuevo León; OA: Oaxaca; PU: Puebla; QR: Quintana Roo; QU: Querétaro; SI: Sinaloa; SL: San Luis Potosí; SO: Sonora; TA: Tamaulipas; TB: Tabasco; TL: Tlaxcala; VE: Veracruz; YU: Yucatán; ZA: Zacatecas]

NIC Nicaragua

PAN Panama

PRI Puerto Rico [includes Vieques]

SAL El Salvador

TUR Turks and Caicos Islands

USA United States of America [AK: Alaska; AL: Alabama; AR: Arkansas; AZ: Arizona; CA: California; CO: Colorado; CT: Connecticut; DC: District of Columbia; DE: Delaware; FL: Florida; GA: Georgia; IA: Iowa; ID: Idaho; IL: Illinois; IN: Indiana; KS: Kansas; KY: Kentucky; LA: Louisiana; MA: Massachusetts; MD: Maryland; ME: Maine; MI: Michigan; MN: Minnesota; MO: Missouri; MS: Mississippi; MT: Montana; NC: North Carolina; ND: North Dakota; NE: Nebraska; NH: New Hampshire; NJ: New Jersey; NM: New Mexico; NV: Nevada; NY: New York; OH: Ohio; OK: Oklahoma; OR: Oregon; PA: Pennsylvania; RI: Rhode Island; SC: South Carolina; SD: South Dakota; TN: Tennessee; TX: Texas; UT: Utah; VA: Virginia; VT: Vermont; WA: Washington; WI: Wisconsin; WV: West Virginia; WY: Wyoming]

VIS Virgin Islands [includes US Virgin Islands and British Virgin Islands: Saint Thomas, Saint Croix]

SA South America

Results

Overall diversity

A total of 128 valid and invalid family-group, 612 genus-group, and 4065 species-group taxa (excluding fossils listed in Appendices 1, 2) are listed in this catalogue. The subfamily Pimeliinae is the most diverse with 908 valid species-group taxa, followed by the Tenebrioninae (808), Alleculinae (418), Stenochiinae (361), Diaperinae (260), Lagriinae (256), Phrenapatinae (22), and Nilioninae (6). Thirty-seven species in three subfamilies are adventive (Table 2), several of which are pests of stored grain products. Mexico and the continental United States of America are by far the most diverse political regions with 1224 and 1230 valid species-group taxa respectively, while at the other extreme Greenland and Bermuda only have two each (Table 3).

Table 2.

List of adventive species documented in North America. Data on the origin, date of first detection in North America, and microhabitat associations in nature are given for each species as far as known. * = pest of stored grain products.

Species Origin Date of detection Microhabitat Placement
Alphitobius diaperinus (Panzer, 1797) Africa <1866 *Animal nests, caves, guano Tenebrioninae: Alphitobiini
Alphitobius laevigatus (Fabricius, 1781) Africa <1866 *Animal nests, caves, guano Tenebrioninae: Alphitobiini
Alphitophagus bifasciatus (Say, 1824) Europe, probably <1824 *Animal nests, caves, guano Diaperinae: Diaperini: Adelinina
Anchophthalmops menouxi (Mulsant and Rey, 1853) Africa; probably not established <1870 Leaf litter on sandy soil, probably Tenebrioninae: Pedinini: Platynotina
Blaps (Blaps) lethifera lethifera Marsham, 1802 Europe <1889 Under rocks, wood, in caves Tenebrioninae: Blaptini : Blaptina
Blaps (Blaps) mucronata Latreille, 1804 Europe <1889 Under rocks, wood, in caves Tenebrioninae: Blaptini : Blaptina
Ceropria induta (Wiedemann, 1819) Asia 1998 Polypore fungi, rotten wood Diaperinae: Diaperini: Diaperina
Ellipsodes (Anthrenopsis) ziczac (Motschulsky, 1873) Asia, probably via Antilles 1891 Under leaf litter on sandy soil Diaperinae: Crypticini
Gnatocerus (Echocerus) maxillosus (Fabricius, 1801) Asia, probably via Europe <1866 *Animal nests, caves, under bark Diaperinae: Diaperini: Adelinina
Gnatocerus (Gnatocerus) cornutus (Fabricius, 1798) Asia, probably via Europe <1866 *Animal nests, caves, under bark Diaperinae: Diaperini: Adelinina
Gondwanocrypticus pictus (Gebien, 1928) South America 1954 Under leaf litter near ant nests Diaperinae: Crypticini
Gondwanocrypticus platensis (Fairmaire, 1884) South America 1929 Under leaf litter near ant nests Diaperinae: Crypticini
Gonocephalum (Gonocephalum) sericeum (Baudi di Selve, 1875) Northwest Africa + Arabian Peninsula 1980 Under leaf litter, wood, rocks Tenebrioninae: Opatrini: Opatrina
Latheticus oryzae Waterhouse, 1880 Old World 1908 *Animal nests, caves, under bark Tenebrioninae: Triboliini
Leichenum canaliculatum variegatum (Klug, 1833) Madagascar 1906 Under leaf litter on sandy soil Tenebrioninae: Pedinini: Leichenina
Lyphia tetraphylla (Fairmaire, 1857) Europe <1902 In dead wood, other insect burrows Tenebrioninae: Triboliini
Myrmechixenus lathridioides Crotch, 1873 Europe <1883 Under leaf litter, in soil Diaperina: Myrmechixenini
Opatroides punctulatus Brulle, 1832 Middle East 2003 Under leaf litter, wood, rocks Tenebrioninae: Opatrini: Opatrina
Palorus cerylonoides (Pascoe, 1863) Indo-Malayan, probably 2004 Under bark dry wood, plant debris Tenebrioninae: Palorini
Palorus genalis Blair, 1930 Old World 1937 Under bark dry wood, plant debris Tenebrioninae: Palorini
Palorus ratzeburgii (Wissmann, 1848) North Africa, probably <1897 *Under bark dry wood, plant debris Tenebrioninae: Palorini
Palorus subdepressus (Wollaston, 1864) Africa, probably <1882 *Under bark dry wood, plant debris Tenebrioninae: Palorini
Pentaphyllus testaceus (Hellwig, 1792) Europe 2005 Polypore fungi, under bark Diaperinae: Diaperini: Diaperina
Platydema woldai Triplehorn and Phillips, 1998 Central America, probably 1964 With orchid plants; at lights in forest Diaperinae: Diaperini: Diaperina
Plesiophthalmus spectabilis Harold, 1875 Asia; probably not established 2013 Rotten wood Tenebrioninae: Amarygmini
Poecilocrypticus formicophilus Gebien, 1928 South America 1978 Under leaf litter near ant nests Diaperinae: Crypticini
Strongylium cultellatum Maklin, 1867 Asia 2010 Dead standing wood Stenochiinae: Stenochiini
Tenebrio molitor Linnaeus, 1758 Africa, probably <1837 *Animal nests, caves Tenebrioninae: Tenebrionini
Tenebrio obscurus Fabricius, 1792 Africa, probably <1869 *Animal nests, caves Tenebrioninae: Tenebrionini
Trachyscelis aphodioides Latreille, 1809 Europe <1846 Under plant debris on beach sand Diaperinae: Trachyscelini
Tribolium (Tribolium) castaneum (Herbst, 1797) Africa, probably <1866 *Under bark dry wood, plant debris Tenebrioninae: Triboliini
Tribolium (Tribolium) confusum Jacquelin du Val, 1862 Africa, probably <1893 *Under bark dry wood, plant debris Tenebrioninae: Triboliini
Tribolium (Tribolium) destructor Uyttenboogaart, 1934 Africa <1948 *Under bark dry wood, plant debris Tenebrioninae: Triboliini
Tribolium (Tribolium) madens (Charpentier, 1825) Africa, probably <1866 *Under bark dry wood, plant debris Tenebrioninae: Triboliini
Tyrtaeus dobsoni Hinton, 1947 Unknown; probably via Europe 2002 Under bark and in dead wood Diaperinae: Gnathidiini : Anopidiina
Ulomina carinata Baudi di Selve, 1876 Asia 1952 Under bark dry wood, plant debris Tenebrioninae: Palorini
Ulomoides ocularis (Casey, 1891) Asia <1891 Dry pods of Tamarindus L. Diaperinae: Diaperini: Diaperina
Table 3.

Number of valid species-group taxa by political region. Data excludes impression and amber fossils.

Political region Lagriinae Nilioninae Phrenapatinae Pimeliinae Tenebrioninae Alleculinae Diaperinae Stenochiinae Total
BAH: Bahamas 0 0 0 8 22 14 25 1 70
BEL: Belize 20 0 3 2 20 5 32 28 110
BER: Bermuda 0 0 0 0 1 0 1 0 2
CAN: Canada 7 0 2 8 58 28 24 10 137
CAY: Cayman Islands 1 0 0 1 14 4 16 0 36
CRI: Costa Rica 44 0 6 12 27 23 40 44 196
CUB: Cuba 2 0 0 23 57 25 37 31 175
DOM: Dominican Republic 0 0 0 2 18 13 19 19 71
GRE: Greenland 0 0 0 0 2 0 0 0 2
GUA: Guatemala 60 2 4 20 70 57 66 71 350
HAI: Haiti 2 0 0 3 10 8 13 11 47
HON: Honduras 1 0 0 8 10 5 16 3 43
JAM: Jamaica 2 0 1 3 19 11 17 0 53
LAN: Lesser Antilles 11 0 3 6 50 11 31 18 130
MEX: Mexico 101 1 4 392 348 135 129 115 1225
NIC: Nicaragua 41 1 3 16 49 19 36 62 227
PAN: Panama 70 5 8 13 51 41 60 87 335
PRI: Puerto Rico 3 0 1 2 22 4 19 10 61
SAL: El Salvador 0 0 0 2 4 4 8 0 18
TUR: Turks and Caicos Islands 0 0 0 0 1 2 2 0 5
USA: United States of America 38 0 2 546 362 154 79 46 1227
VIS: Virgin Islands 0 0 0 0 3 2 1 1 7

A significant proportion of new species-group taxa (41%) were described between the years 1880 and 1910 (Fig. 1). A noticeable decrease in the number of new species-group taxa proposed occurred between 1940–1960, with a small resurgence since then (at a rate of approximately 100 new taxa per decade, see Figs 1, 2). Over 3000 North American species-group taxa are currently recognized as valid (Fig. 2), approximately 15% of the world fauna.

Figure 1. 

Number of North American species-group taxa described over time, by decade. Data excludes adventive species as well as impression and amber fossils.

Figure 2. 

Cumulative number of North American species-group taxa described over time, by decade. Data excludes adventive species as well as impression and amber fossils. Line with square marker = total available taxa, line with circular marker = currently valid taxa.

Significant contributions (see Table 4)

The British entomologist George Charles Champion [b. 1851, d. 1927], working on the fauna of Mexico and Central America, proposed the highest number of new tenebrionid taxa found on the continent (83 genus-group, 906 species-group taxa) followed by the American Thomas Lincoln Casey [b. 1857, d. 1925], working mainly on the fauna of the United States of America (80 genus-group, 792 species-group taxa). Frank Ellsworth Blaisdell [b. 1862, d. 1947] (338 species-group taxa), John Lawrence LeConte [b. 1825, d. 1883] (270 species-group taxa), George Henry Horn [b. 1840, d. 1897] (133 species-group taxa), and John Milton Campbell [b. 1935] (103 species-group taxa) also contributed significantly to describing the North American darkling beetle fauna.

Table 4.

Significant contributions to the description of new North American Tenebrionidae genus-group and species-group taxa (list includes the top ten contributors in each category). Data excludes adventive taxa as well as impression and amber fossils. * = given in parentheses is the country where the person produced taxonomic works, when different from the country of origin.

Author Country of origin* New genus-group names New species-group names
Blaisdell, Frank Ellsworth USA 20 338
Campbell, John Milton USA (Canada) 10 103
Casey, Thomas Lincoln USA 80 792
Champion, George Charles United Kingdom 83 906
Dejean, Pierre Francois Marie Auguste France 16 0
Doyen, John Thomas USA 5 79
Fall, Henry Clinton USA 1 66
Horn, George Henry USA 20 133
Laporte, Francois Louis (Comte de Castelnau) United Kingdom (France) 12 40
LeConte, John Lawrence USA 56 277
Maklin, Friedrich [Fredrik] Wilhelm Finland 3 66
Marcuzzi, Giorgio Italy 7 84
Motschulsky, Victor de Russia 12 24
Mulsant, Martial Etienne France 13 18
Pascoe, Francis Polkinghorne United Kingdom 12 4
Solier, Antoine Joseph Jean France 17 35
Triplehorn, Charles Albert USA 4 86

Catalogue of Tenebrionidae (Coleoptera) of North America

Family TENEBRIONIDAE Latreille, 1802

Tenebrionites Latreille, 1802: 165. Type genus: Tenebrio Linnaeus, 1758.

Subfamily LAGRIINAE Latreille, 1825

Lachnaedes Billberg, 1820a: 34. Type genus: Lachna Billberg, 1820 (= Lagria Fabricius, 1775). Nomen oblitum (see Favret and Bouchard 2016).

Lagriariae Latreille, 1825: 381. Type genus: Lagria Fabricius, 1775. Note. Use of younger family-group name conserved over Lachnina Billberg, 1820 (ICZN 1999: Article 40.2) (see Bouchard et al. 2005).

Tribe Belopini Reitter, 1917

Belopinae Reitter, 1917: 59. Type genus: Belopus Gebien, 1911.

Genus Adelonia Laporte, 1840 [F]

Adelonia Laporte, 1840: 221. Type species: Uloma filiformis Laporte, 1840, monotypy.

Merotemnus Horn, 1870: 367. Type species: Merotemnus elongatus Horn, 1870 (= Uloma filiformis Laporte, 1840), monotypy. Synonymy: Spilman (1961a: 49).

Rhacius Champion, 1885: 120. Type species: Rhacius sulcatulus Champion, 1885, subsequent designation (Gebien 1941: 805). Synonymy: Spilman (1961a: 50).

Adelonia filiformis (Laporte, 1840)MEX (BC BS)

Uloma filiformis Laporte, 1840: 221.

Merotemnus elongatus Horn, 1870: 367. Synonymy: Gebien (1908b: 160).

Adelonia insularis Doyen, 1983MEX (NA [Islas Marías])

Adelonia insularis Doyen, 1983: 85.

Adelonia quadricollis (Champion, 1885)GUABELPAN / SA

Rhacius quadricollis Champion, 1885: 121.

Adelonia sulcatula (Champion, 1885)4USA (TX) MEX (GE JA MO OA PU SO VE YU) GUAHONNICCRIPAN / CUBCAYJAM / SA

Rhacius sulcatulus Champion, 1885: 121.

Genus Rhypasma Pascoe, 1862 [N]

Rhypasma Pascoe, 1862: 325. Type species: Rhypasma pusillum Pascoe, 1862, monotypy.

Derosimus Fairmaire, 1904: 62. Type species: Derosimus quadricollis Fairmaire, 1904, monotypy. Synonymy: Blair (1935: 104).

Rhypasma costaricense Marcuzzi, 1976 CRI

Rhypasma costaricense Marcuzzi, 1976: 119.

Rhypasma haitianum Marcuzzi, 1954 CUB HAI

Rhypasma haitianum Marcuzzi, 1954a: 82.

Rhypasma livae Ferrer and Ødegaard, 2005 NIC PAN

Rhypasma livae Ferrer and Ødegaard, 2005: 635.

Tribe Eschatoporiini Blaisdell, 1906

Eschatoporini Blaisdell, 1906: 78. Type genus: Eschatoporis Blaisdell, 1906.

Genus Eschatoporis Blaisdell, 1906 [M]

Eschatoporis Blaisdell, 1906: 76. Type species: Eschatoporis nunenmacheri Blaisdell, 1906, monotypy.

Eschatoporis nunenmacheri Blaisdell, 1906USA (CA)

Eschatoporis nunenmacheri Blaisdell, 1906: 78.

Eschatoporis styx Aalbu, Kanda and Smith, 2017USA (CA)

Eschatoporis styx Aalbu, Kanda and Smith, 2017: 140.

Tribe Goniaderini Lacordaire, 1859

Goniadérides Lacordaire, 1859: 390. Type genus: Goniadera Perty, 1832.

Phobeliina Ardoin, 1961: 33. Type genus: Phobelius Blanchard, 1845.

Genus Anaedus Blanchard, 1842 [M]

Aspisoma Duponchel and Chevrolat, 1841: 210 [junior homonym of Aspisoma Laporte, 1833]. Type species: Aspisoma fulvipenne Duponchel and Chevrolat, 1841, original designation.

Anaedus Blanchard, 1842: pl. 14. Type species: Anaedus punctatissimus Blanchard, 1842, monotypy. Synonymy: Lacordaire (1859: 396).

Anaedes Agassiz, 1846: 36. Unjustified emendation of Anaedus Blanchard, 1842, not in prevailing usage.

Aspidosoma Agassiz, 1846: 36. Unjustified emendation of Aspisoma Duponchel and Chevrolat, 1841, not in prevailing usage.

Anaedus aeneotinctus Champion, 1893MEX (GE)

Anaedus aeneotinctus Champion, 1893a: 543.

Anaedus apicicornis Champion, 1886MEX (JA) PAN

Anaedus apicicornis Champion, 1886: 236.

Anaedus brevicollis Champion, 1886 GUA

Anaedus brevicollis Champion, 1886: 236.

Anaedus brunneus (Ziegler, 1844) [Fig. 3] CAN (ON) USA (AL AR DC FL IN KS KY LA MA MD MO MS NC NJ OH PA RI SC TN VA WI)

Figure 3. 

Anaedus brunneus (Ziegler, 1844). Scale bar = 1 mm.

Pandarus brunneus Ziegler, 1844: 45.

Anaedus impressicollis Pic, 1917 MEX

Anaedus impressicollis Pic, 1917: 19.

Anaedus inangulatus (Pic, 1934) NIC

Aspisoma inangulata Pic, 1934: 35.

Anaedus longicornis Champion, 1886USA (TX) MEX (GU OA) GUA

Anaedus longicornis Champion, 1886: 235.

Anaedus maculatus Champion, 1886 NIC PAN

Anaedus maculatus Champion, 1886: 235.

Anaedus marginatus Champion, 1886 NIC PAN

Anaedus marginatus Champion, 1886: 236.

Anaedus mexicanus Champion, 1886MEX (VE)

Anaedus mexicanus Champion, 1886: 234.

Anaedus nitidissimus Pic, 1917 CRI

Anaedus nitidissimus Pic, 1917: 20.

Anaedus pallidus Schaeffer, 1915USA (TX)

Anaedus pallidus Schaeffer, 1915: 238.

Anaedus punctatissimus Blanchard, 1842MEX (DU JA OA PU SI VE) GUANICCRIPAN/ SA

Anaedus punctatissimus Blanchard, 1842: pl. 14.

Anaedus quadrinotatus Champion, 1896 LAN

Anaedus quadrinotatus Champion, 1896: 26.

Anaedus rotundicollis LeConte, 1851USA (AZ) MEX (BS)

Anoedus rotundicollis LeConte, 1851: 150.

Anaedus setulosus Champion, 1886MEX (TB) NICPAN / SA

Anaedus setulosus Champion, 1886: 237.

Anaedus similis Champion, 1886MEX (VE) GUANIC

Anaedus similis Champion, 1886: 234.

Anaedus texanus Linell, 1899USA (TX)

Anoedus texanus Linell, 1899: 182.

Anaedus villosus Champion, 1893 GUA CRI PAN

Anaedus villosus Champion, 1893a: 543.

Genus Goniadera Perty, 1832 [F]

Goniadera Perty, 1832: 625. Type species: Goniadera crenata Perty, 1832, monotypy.

Goniodera Agassiz, 1846: 165. Unjustified emendation of Goniadera Perty, 1832, not in prevailing usage.

Subgenus Aemymone Bates, 1868

Aemymone Bates, 1868: 314. Type species: Aemymone cariosa Bates, 1868, original designation.

Goniadera championi Ferrer and Delatour, 2007MEX (VE) PAN

Aemymone crenata Champion, 1893a: 542 [junior secondary homonym of Goniadera crenata Perty, 1832].

Goniadera championi Ferrer and Delatour, 2007: 286. Replacement name for Goniadera crenata (Champion, 1893).

Subgenus Goniadera Perty, 1832

Goniadera Perty, 1832: 62. Type species: Goniadera crenata Perty, 1832, monotypy.

Goniadera alternata Champion, 1886MEX (VE) GUABELPAN

Goniadera alternata Champion, 1886: 231.

Goniadera dissipata Kirsch, 1866PAN / JAMLAN / SA

Goniadera dissipata Kirsch, 1866: 197.

Goniadera nicaraguensis Champion, 1886 NIC

Goniadera nicaraguensis Champion, 1886: 230.

Goniadera obscuriceps Pic, 1913NIC / SA

Goniadera obscuriceps Pic, 1913a: 125.

Goniadera oculata oculata Champion, 1886MEX (GE OA VE YU) BELNICCRIPAN

Goniadera oculata Champion, 1886: 230.

Goniadera pilosa Champion, 1886 NIC CRI PAN

Goniadera pilosa Champion, 1886: 230.

Goniadera pseudorepanda Ferrer and Delatour, 2007MEX (GE VE YU) GUANICCRI / SA

Goniadera pseudorepanda Ferrer and Delatour, 2007: 296.

Goniadera repanda (Fabricius, 1801)MEX (JA VE) GUABELNICCRI / SA

Melandrya repanda Fabricius, 1801a: 165.

Subgenus Opatresthes Gebien, 1928

Opatresthes Gebien, 1928b: 192. Type species: Opatresthes binodosus Gebien, 1928, subsequent designation (Gebien 1941: 817).

Goniadera maesi Ferrer and Delatour, 2007 NIC

Goniadera maesi Ferrer and Delatour, 2007: 287.

Goniadera quadrinodosa (Gebien, 1928)6CRI / SA

Opatresthes quadrinodosus Gebien, 1928b: 193.

Genus Paratenetus Spinola, 1844 [M]

Paratenetus Spinola, 1844: 116. Type species: Paratenetus punctatus Spinola, 1844, subsequent designation (Lucas 1920: 483).

Lagriola Kirsch, 1874: 409. Type species: Lagriola operosa Kirsch, 1874, present designation. Synonymy: Matthews and Lawrence (2015: 289).

Storthephora Mäklin, 1875a: 658. Type species: Storthephora denticollis Mäklin, 1875, subsequent designation (Bousquet and Bouchard 2014: 26). Synonymy: Champion (1893c: 47).

Paratenetus brevipennis Champion, 1886 PAN

Paratenetus brevipennis Champion, 1886: 242.

Paratenetus championi Matthews and Lawrence, 2015 PAN

Paratenetus denticulatus Champion, 1886: 243 [secondary homonym of Paratenetus denticulatus (Kirsch, 1874)].

Paratenetus championi Matthews and Lawrence, 2015: 311. Replacement name for Paratenetus denticulatus Champion, 1886.

Paratenetus constrictus Champion, 1893MEX (CI TB VE) GUABELCRIPAN

Paratenetus constrictus Champion, 1893a: 546.

Paratenetus corticarioides Champion, 1886MEX (OA) GUA

Paratenetus corticarioides Champion, 1886: 241.

Paratenetus crenulatus Champion, 1886 PAN

Paratenetus crenulatus Champion, 1886: 242.

Paratenetus exutus Bousquet and Bouchard, 2014CAN (AB MB NB NS ON QC SK) USA (AL AR CT DC FL IA IL IN KS KY LA MD ME MI MN MO MS NC ND NJ NY OH OK PA TN TX VA WI WV)

Paratenetus exutus Bousquet and Bouchard, 2014: 39.

Paratenetus foveithorax Ferrer and Ødegaard, 2005 PAN

Paratenetus foveithorax Ferrer and Ødegaard, 2005: 635.

Paratenetus fuscus LeConte, 1850 [Fig. 4] CAN (AB BC MB NT ON QC SK) USA (CO CT DC IA KS MA MD MI MT ND NE NM NY OH RI SD TN VT WI WY)

Figure 4. 

Paratenetus fuscus LeConte, 1850. Scale bar = 1 mm.

Paratenetus fuscus LeConte, 1850: 223.

Paratenetus crinitus Fall, 1907a: 253. Synonymy: Bousquet and Bouchard (2014: 31).

Paratenetus gibbipennis Motschulsky, 1869CAN (MB ON QC) USA (AL CT GA IL MA ME MI MN MO NC ND NE NH NJ NY OH PA RI SC TN TX VA WI)

Paratenetus gibbipennis Motschulsky, 1869: 193.

Paratenetus cribratus Motschulsky, 1869: 193. Synonymy: Bousquet and Bouchard (2014: 29).

Paratenetus grandicornis Motschulsky, 1869 NIC PAN

Paratenetus grandicornis Motschulsky, 1869: 193.

Paratenetus inermis Champion, 1893 GUA

Paratenetus inermis Champion, 1893a: 545.

Paratenetus koltzei Pic, 1939 MEX

Paratenetus koltzei Pic, 1939: 9.

Paratenetus longicornis Pic, 1925LAN (Guadeloupe)

Paratenetus longicornis Pic, 1925a: 6.

Paratenetus mexicanus Pic, 1925MEX (SI)

Paratenetus mexicanus Pic, 1925a: 6.

Paratenetus nigricornis Champion, 1893MEX (TB VE) GUABELPAN

Paratenetus nigricornis Champion, 1893a: 544.

Paratenetus obovatus Champion, 1886 BEL

Paratenetus obovatus Champion, 1886: 241.

Paratenetus punctatus Spinola, 1844CAN (MB NB ON QC) USA (AR CT DC GA IA IL IN KS KY LA MA MD ME MI MN MO MS NC NH NJ NY OH OK PA RI SC TN TX VA VT WI WV WY)

Latridius pubescens Say, 1826: 265 [nomen dubium, see Bousquet and Bouchard (2014: 33)].

Paratenetus punctatus Spinola, 1844: 118.

Paratenetus punctulatus Champion, 1893MEX (TB VE) BEL

Paratenetus punctulatus Champion, 1893a: 545.

Paratenetus ruficornis Champion, 1886 PAN

Paratenetus ruficornis Champion, 1886: 239.

Paratenetus sexdentatus Champion, 1893 GUA BEL PAN

Paratenetus sexdentatus Champion, 1893a: 546.

Paratenetus testaceus Pic, 1920MEX (PU) CRI

Paratenetus testaceus Pic, 1920: 2.

Paratenetus texanus Bousquet and Bouchard, 2014USA (FL LA TX) MEX (CI NA TA)

Paratenetus texanus Bousquet and Bouchard, 2014: 45.

Paratenetus tibialis Champion, 1886MEX (GE TB VE) GUABEL

Paratenetus tibialis Champion, 1886: 239.

Paratenetus tropicalis Motschulsky, 1869MEX (TB VE) GUABELPAN

Paratenetus tropicalis Motschulsky, 1869: 193.

Paratenetus tuberculatus Champion, 1886 PAN

Paratenetus tuberculatus Champion, 1886: 242.

Paratenetus villosus Champion, 1886MEX (VE) GUAPAN

Paratenetus villosus Champion, 1886: 240.

Genus Phobelius Blanchard, 1842 [M]

Phobelius Blanchard, 1842: pl. 14. Type species: Phobelius crenatus Blanchard, 1842, monotypy.

Phobelius mexicanus Doyen, 1990MEX (JA) GUA

Phobelius mexicanus Doyen, 1990: 217.

Phobelius nevermanni Kulzer, 1961 CRI

Phobelius nevermanni Kulzer, 1961a: 226.

Genus Phymatestes Pascoe, 1867 [M]

Phymatodes Dejean, 1834: 203. Type species: Lagria tuberculata Fabricius, 1792, monotypy. Note. Dejean’s name has been suppressed by the Commission in Opinion 1525 (ICZN 1989).

Phymatestes Pascoe, 1867: 142. Replacement name for Phymatodes Dejean, 1834 [as Phymatodes Blanchard, 1845]. Note. This name has been placed on the Official List of Generic Names in Zoology in Opinion 1525 (ICZN 1989).

Phymatestes agnei Ferrer and Ødegaard, 2005 PAN

Phymatestes agnei Ferrer and Ødegaard, 2005: 634.

Phymatestes charbonnelae Ferrer and Moraguès, 2003LAN (Grenada)

Phymatestes charbonnelae Ferrer and Moraguès, 2003: 161.

Genus Prateus LeConte, 1862 [M]

Prateus LeConte, 1862a: 238. Type species: Prateus fusculus LeConte, 1862, original designation.

Prateus fusculus LeConte, 1862USA (AL AR DC FL MD MS NC NY OH OK SC TN TX VA WV) MEX (TA)

Prateus fusculus LeConte, 1862a: 238.

Genus Xanthicles Champion, 1886 [M]

Xanthicles Champion, 1886: 231. Type species: Xanthicles caraboides Champion, 1886, subsequent designation (Gebien 1941: 815).

Xanthicles caraboides Champion, 1886 CRI

Xanthicles caraboides Champion, 1886: 232.

Xanthicles hirsutus Champion, 1886 CRI

Xanthicles hirsutus Champion, 1886: 232.

Tribe Lagriini Latreille, 1825

Lagriariae Latreille, 1825: 381. Type genus: Lagria Fabricius, 1775.

Subtribe Statirina Blanchard, 1845

Statyrites Blanchard, 1845: 39. Type genus: Statira Lepeletier and Audinet-Serville, 1828.

Genus Arthromacra Kirby, 1837 [F]

Arthromacra Kirby, 1837: 238. Type species: Arthromacra donacioides Kirby, 1837 (= Lagria aenea Say, 1824), monotypy.

Macrarthra Agassiz, 1846: 219. Unjustified emendation of Arthromacra Kirby, 1837, not in prevailing usage.

Arthromacra aenea aenea (Say, 1824)CAN (MB NB NS ON PE QC) USA (CT DC DE MA MD ME MI NC NH NJ NY OH PA TN VA VT WI WV)

Lagria aenea Say, 1824b: 287 [junior primary homonym of Lagria aenea Fabricius, 1775].

Arthromacra donacioides Kirby, 1837: 239. Synonymy: LeConte (1859d: 191).

Lagria viridis Melsheimer, 1845: 311. Synonymy: Parsons (1976: 222).

Arthromacra aenea glabricollis Blatchley, 1910USA (IL IN KY MO OH PA VA WI)

Arthromacra glabricollis Blatchley, 1910: 1285.

Arthromacra aenea lengi Parsons, 1976USA (GA NC PA SC TN WV)

Arthromacra aenea lengi Parsons, 1976: 224.

Arthromacra aenea rugosecollis Leng, 1914USA (GA NC TN)

Arthromacra aenea var. rugosecollis Leng, 1914: 287.

Arthromacra appalachiana Leng, 1917USA (NC SC TN VA)

Arthromacra appalachiana Leng, 1917: 18.

Arthromacra pilosella Leng, 1917USA (KY NC SC TN)

Arthromacra pilosella Leng, 1917: 18.

Arthromacra robinsoni Leng, 1914USA (NC SC VA)

Arthromacra robinsoni Leng, 1914: 286.

Genus Colparthrum Kirsch, 1866 [N]

Colparthrum Kirsch, 1866: 204. Type species: Colparthrum gerstaeckeri Kirsch, 1866, monotypy.

Subgenus Colparthrum Kirsch, 1866

Colparthrum Kirsch, 1866: 204. Type species: Colparthrum gerstaeckeri Kirsch, 1866, monotypy.

Colparthrum aenescens Borchmann, 1936 CRI

Colparthrum aenescens Borchmann, 1936: 444.

Colparthrum decoratum bilunulatum (Pic, 1912) PAN

Statira bilunulata Pic, 1912b: 76.

Colparthrum decoratum decoratum (Mäklin, 1863)MEX (VE) GUANICPAN

Statira decorata Mäklin, 1863: 588.

Colparthrum decoratum maklini Borchmann, 19367 [No region originally mentioned but presumably from Mexico and/or Central America]

Colparthrum decoratum var. mäklini Borchmann, 1936: 447.

Colparthrum foveiceps Champion, 1889 PAN

Colparthrum foveiceps Champion, 1889: 68.

Colparthrum grande Borchmann, 1936 CRI

Colparthrum grandis Borchmann, 1936: 439.

Colparthrum majus Borchmann, 1916 MEX

Colparthrum decoratum var. major Borchmann, 1916: 233.

Subgenus Pseudocolparthrum Borchmann, 1916

Pseudocolparthrum Borchmann, 1916: 236. Type species: Colparthrum calcaratum Champion, 1889, subsequent designation (Borchmann 1936: 452).

Colparthrum calcaratum Champion, 1889 NIC CRI PAN

Colparthrum calcaratum Champion, 1889: 71.

Colparthrum sulcicolle Champion, 1889 NIC PAN

Colparthrum sulcicolle Champion, 1889: 69.

Colparthrum vitticolle Champion, 1889 NIC

Colparthrum vitticolle Champion, 1889: 70.

Genus Disema Mäklin, 1875 [F]

Disema Mäklin, 1875a: 646. Type species: Disema bimaculata Mäklin, 1875, subsequent designation (Lucas 1920: 244).

Disema singularis (Champion, 1889) PAN

Sphragidophorus singularis Champion, 1889: 64.

Genus Epicydes Champion, 1889 [M]

Epicydes Champion, 1889: 60. Type species: Epicydes oculatus Champion, 1889, subsequent designation (Borchmann 1936: 429).

Subgenus Cybostira Borchmann, 1936

Cybostira Borchmann, 1936: 430. Type species: Cybostira caligata Borchmann, 1936, original designation.

Epicydes caligatus (Borchmann, 1936) CRI

Cybostira caligata Borchmann, 1936: 430.

Subgenus Epicydes Champion, 1889

Epicydes Champion, 1889: 60. Type species: Epicydes oculatus Champion, 1889, subsequent designation (Borchmann 1936: 429).

Epicydes oculatus Champion, 1889MEX (OA VE) GUA

Epicydes oculatus Champion, 1889: 61.

Epicydes vicinus Champion, 1889 GUA NIC

Epicydes vicinus Champion, 1889: 61.

Genus Meniscophorus Champion, 1889 [M]

Meniscophorus Champion, 1889: 64. Type species: Meniscophorus amazonicus Champion, 1889, subsequent designation (Lucas 1920: 404).

Meniscophorus costatus Champion, 1889 PAN

Meniscophorus costatus Champion, 1889: 65.

Genus Meropria Borchmann, 1921 [F]

Meropria Borchmann, 1921: 228. Type species: Statira glabrata Mäklin, 1863, original designation.

Meropria chiriquina (Champion, 1889) PAN

Statira chiriquina Champion, 1889: 9.

Meropria denticulata (Champion, 1889)PAN / SA

Statira denticulata Champion, 1889: 7.

Meropria glabrata (Mäklin, 1863)MEX (GE MO OA VE) GUABELCRI

Statira glabrata Mäklin, 1863: 587.

Meropria interrupta (Champion, 1889) GUA NIC PAN

Statira interrupta Champion, 1889: 8.

Meropria unidentata (Champion, 1889)MEX (CI VE) GUABEL

Statira unidentata Champion, 1889: 8.

Genus Nevermanniella Borchmann, 1936 [F]

Nevermanniella Borchmann, 1936: 332. Type species: Statira albolineata Champion, 1889, original designation.

Nevermanniella albolineata (Champion, 1889)MEX (VE) BELNIC

Statira albolineata Champion, 1889: 36.

Genus Othryades Champion, 1889 [M]

Othryades Champion, 1889: 72. Type species: Othryades fragilicornis Champion, 1889, monotypy.

Othryades fragilicornis Champion, 1889 PAN

Othryades fragilicornis Champion, 1889: 72.

Genus Rhaibodera Borchmann, 1921 [F]

Rhaibodera Borchmann, 1921: 219. Type species: Rhaibodera pachycera Borchmann, 1921, original designation.

Rhaibodera crassicornis (Champion, 1889)MEX (TB)

Statira crassicornis Champion, 1889: 18.

Genus Rhosaces Champion, 1889 [M]

Rhosaces Champion, 1889: 73. Type species: Rhosaces clavipes Champion, 1889, monotypy.

Rhosaces clavipes Champion, 1889 PAN

Rhosaces clavipes Champion, 1889: 73.

Genus Sphragidophorus Champion, 1889 [M]

Sphragidophorus Champion, 1889: 61. Type species: Statira cyanipennis Mäklin, 1863, subsequent designation (Lucas 1920: 603).

Sphragidophorus cyanipennis (Mäklin, 1863)MEX (VE) GUAPAN

Statira cyanipennis Mäklin, 1863: 591.

Sphragidophorus ocularis Borchmann, 1936 CRI

Sphragidophorus ocularis Borchmann, 1936: 507.

Sphragidophorus violaceus Champion, 1889 PAN

Sphragidophorus violaceus Champion, 1889: 63.

Genus Statira Lepeletier and Audinet-Serville, 1828 [F]

Statira Lepeletier and Audinet-Serville, 1828: 479. Type species: Statira agroides Lepeletier and Audinet-Serville, 1828, subsequent designation (Blanchard 1844: pl. 53bis).

Subgenus Spinostatira Pic, 1918

Spinostatira Pic, 1918a: 22. Type species: Statira spinipes Pic, 1918, subsequent designation (Borchmann 1936: 247).

Statira costaricensis Champion, 1889CRI / SA

Statira costaricensis Champion, 1889: 36.

Subgenus Statira Lepeletier and Audinet-Serville, 1828

Statira Lepeletier and Audinet-Serville, 1828: 479. Type species: Statira agroides Lepeletier and Audinet-Serville, 1828, subsequent designation (Blanchard 1844: pl. 53bis).

Statira aeneipennis Champion, 1889 GUA

Statira aeneipennis Champion, 1889: 25.

Statira aeneotincta Champion, 1889MEX (VE) GUA

Statira aeneotincta Champion, 1889: 27.

Statira aerata Champion, 1889MEX (VE) GUA

Statira aerata Champion, 1889: 26.

Statira agraeformis Champion, 1889 PAN

Statira agraeformis Champion, 1889: 12.

Statira albofasciata Champion, 1889MEX (VE) GUAPAN

Statira albofasciata Champion, 1889: 44.

Statira alternans Champion, 1889MEX (OA)

Statira alternans Champion, 1889: 30.

Statira amicta Borchmann, 1936 CRI

Statira amicta Borchmann, 1936: 294.

Statira analis Borchmann, 1921 MEX

Statira analis Borchmann, 1921: 273.

Statira angustula Champion, 1889 GUA

Statira angustula Champion, 1889: 15.

Statira antennalis Borchmann, 1936 CRI

Statira antennalis Borchmann, 1936: 313.

Statira asperata Champion, 1889PAN / LAN / SA

Statira asperata Champion, 1889: 49.

Statira antillarum Champion, 1896: 36. Synonymy: Champion (1917: 230).

Statira basalis Horn, 1888USA (AL AR FL GA LA MD MO MS NC SC TX VA)

Statira basalis Horn, 1888: 31.

Statira bicolor Champion, 1889 PAN

Statira bicolor Champion, 1889: 47.

Statira biseriata Borchmann, 1921 MEX

Statira biseriata Borchmann, 1921: 263.

Statira borchmanni Nevermann, 1926 CRI

Statira borchmanni Nevermann, 1926: 113.

Statira brevipilis Champion, 1889MEX (VE)

Statira brevipilis Champion, 1889: 39.

Statira caeruleipennis Champion, 1889MEX (MI)

Statira caeruleipennis Champion, 1889: 14.

Statira ciliata Champion, 1889 GUA

Statira ciliata Champion, 1889: 42.

Statira collaris Champion, 1889MEX (VE)

Statira collaris Champion, 1889: 13.

Statira colorata Fall, 1909MEX (BC BS)

Statira colorata Fall, 1909: 165.

Statira conspicillata conspicillata Mäklin, 1863MEX (CI TB VE) GUABELNICPAN

Statira conspicillata Mäklin, 1863: 589.

Statira conspicillata lateannulata Borchmann, 1936 GUA

Statira conspicillata var. lateannulata Borchmann, 1936: 291.

Statira corrosa Champion, 1889MEXGUA / SA

Statira corrosa Champion, 1889: 37.

Statira cribrata Champion, 1889 GUA

Statira cribrata Champion, 1889: 42.

Statira croceicollis Mäklin, 1863USA (AL FL GA MS)

Statira croceicollis Mäklin, 1863: 594.

Statira cruciata Champion, 1917 NIC

Statira cruciata Champion, 1917: 254.

Statira cupreotincta Champion, 1889 NIC PAN

Statira cupreotincta Champion, 1889: 23.

Statira curticollis Champion, 1889MEX (GE ME MO OA) GUA

Statira curticollis Champion, 1889: 24.

Statira defecta Schaeffer, 1905USA (AZ NM)

Statira defecta Schaeffer, 1905b: 175.

Statira dolera Parsons, 1966USA (FL GA)

Statira dolera Parsons, 1966: 249.

Statira dumalis Parsons, 1973USA (CA)

Statira dumalis Parsons, 1973: 1.

Statira erina Parsons, 1976USA (TX)

Statira erina Parsons, 1976: 219.

Statira evanescens evanescens Champion, 1889MEX (DU GE JA MO OA VE) NIC

Statira evanescens Champion, 1889: 34.

Statira evanescens obscuripennis Borchmann, 1921 MEX

Statira evanescens var. obscuripennis Borchmann, 1921: 260.

Statira flohri Champion, 1893MEX (OA)

Statira flohri Champion, 1893b: 452.

Statira foveicollis Champion, 1889 BEL NIC PAN

Statira foveicollis Champion, 1889: 18.

Statira fulva Fleutiaux and Sallé, 1890 LAN

Statira fulva Fleutiaux and Sallé, 1890: 431.

Statira gagatina Melsheimer, 1845 [Fig. 5] CAN (QC) USA (AL AR CT DC DE IA IL IN KS KY MA MD MI MS NC NJ NY OH PA SC TN VT WI)

Figure 5. 

Statira gagatina Melsheimer, 1845. Scale bar = 1 mm.

Statyra gagatina Melsheimer, 1845: 311.

Statyra resplendens Melsheimer, 1845: 311. Synonymy: Parsons (1976: 220).

Lagria fusca Melsheimer, 1845: 311. Synonymy: Parsons (1966: 252).

Statira glabricollis Borchmann, 1936 CRI

Statira glabricollis Borchmann, 1936: 315.

Statira guatemalensis Champion, 1889 GUA

Statira guatemalensis Champion, 1889: 48.

Statira guttata Borchmann, 1921MEX (“Tenancingo”)

Statira guttata Borchmann, 1921: 250.

Statira haitiensis Champion, 1917 HAI

Statira haitiensis Champion, 1917: 255.

Statira heliconiae Borchmann, 1936 CRI

Statira heliconiae Borchmann, 1936: 292.

Statira heliophila Borchmann, 1936 CRI

Statira heliophila Borchmann, 1936: 276.

Statira hirsuta Champion, 1889USA (TX) MEX (CI GE JA VE) NIC

Statira hirsuta Champion, 1889: 50.

Statira simulans Schaeffer, 1905a: 180. Synonymy: Parsons (1966: 246).

Statira horrida Champion, 1889 GUA

Statira horrida Champion, 1889: 38.

Statira huachucae Schaeffer, 1905USA (AZ NM)

Statira huachucae Schaeffer, 1905b: 176.

Statira ignita Champion, 1889MEX (VE)

Statira ignita Champion, 1889: 23.

Statira inaequicollis Borchmann, 1936 CRI

Statira inaequicollis Borchmann, 1936: 287.

Statira inconstans Champion, 1889 GUA NIC

Statira inconstans Champion, 1889: 16.

Statira ingae Borchmann, 1936 CRI

Statira ingae Borchmann, 1936: 295.

Statira ingens Champion, 1889 NIC CRI PAN

Statira ingens Champion, 1889: 12.

Statira irazuensis Champion, 1889 CRI

Statira irazuensis Champion, 1889: 22.

Statira irregularis Champion, 1889 GUA

Statira irregularis Champion, 1889: 45.

Statira isthmiaca Champion, 1889 PAN

Statira isthmiaca Champion, 1889: 19.

Statira laevicollis Champion, 1889MEX (CL GE)

Statira laevicollis Champion, 1889: 46.

Statira latitator Parsons, 1973USA (CA) MEX (BC)

Statira latitator Parsons, 1973: 3.

Statira leptotracheloides Champion, 1889MEX (DU)

Statira leptotracheloides Champion, 1889: 52.

Statira liebecki Leng, 1924USA (AL FL SC)

Statira liebecki Leng, 1924: 187.

Statira limbata Champion, 1889MEX (TB VE YU) BEL

Statira limbata Champion, 1889: 14.

Statira limonis Borchmann, 1936 CRI

Statira limonis Borchmann, 1936: 315.

Statira marmorata Champion, 1889MEX (VE) GUA

Statira marmorata Champion, 1889: 43.

Statira mediosignata Borchmann, 1921MEX (“Santiago Ixcuintla”)

Statira mediosignata Borchmann, 1921: 278.

Statira melanocephala Mäklin, 1863MEX (VE)

Statira melanocephala Mäklin, 1863: 593.

Statira metallica Champion, 1889 NIC CRI PAN

Statira metallica Champion, 1889: 16.

Statira mexicana Champion, 1889MEX (PU VE)

Statira mexicana Champion, 1889: 26.

Statira microps Champion, 1889MEX (CI TB) GUA

Statira microps Champion, 1889: 44.

Statira minima minima Champion, 1889 NIC PAN

Statira minima Champion, 1889: 47.

Statira minima subatra Borchmann, 1936 “Mittelamerika”

Statira minima var. subatra Borchmann, 1936: 276.

Statira multiformis Champion, 1889MEX (VE) GUANICPAN

Statira multiformis Champion, 1889: 19.

Statira multipunctata Champion, 1889MEX (MO)

Statira multipunctata Champion, 1889: 49.

Statira nevermanni Borchmann, 1936 CRI

Statira nevermanni Borchmann, 1936: 255.

Statira nigripennis affinis Mäklin, 1875 MEX

Statira affinis Mäklin, 1875a: 642.

Statira nigripennis championi Pic, 1912 MEX

Statira nigripennis var. championi Pic, 1912a: 20.

Statira nigripennis humeralis Mäklin, 1875 MEX

Statira humeralis Mäklin, 1875a: 642.

Statira nigripennis nigripennis Mäklin, 1875MEX (JA MO)

Statira nigripennis Mäklin, 1875a: 641.

Statira nigroaenea Champion, 1889MEX (DU)

Statira nigroaenea Champion, 1889: 40.

Statira nigrofasciata Borchmann, 1921MEX (“Navarrete”)

Statira nigrofasciata Borchmann, 1921: 257.

Statira nigromaculata Champion, 1889USA (AZ TX) MEX (OA VE) GUA

Statira nigromaculata Champion, 1889: 33.

Statira nigrosparsa Mäklin, 1863MEX (GE VE) GUANIC

Statira nigrosparsa Mäklin, 1863: 590.

Statira nodulosa Champion, 1889 GUA

Statira nodulosa Champion, 1889: 31.

Statira opaca Borchmann, 1936 CRI

Statira opaca Borchmann, 1936: 283.

Statira opacicollis Horn, 1888USA (AZ)

Statira opacicollis Horn, 1888: 30.

Statira paradoxa Borchmann, 1936 CRI

Statira paradoxa Borchmann, 1936: 306.

Statira patricia Borchmann, 1921 CRI

Statira patricia Borchmann, 1921: 269.

Statira penicillata Champion, 1889MEX (VE)

Statira penicillata Champion, 1889: 30.

Statira perforata Champion, 1917 MEX

Statira perforata Champion, 1917: 262.

Statira pici Blackwelder, 1945 CRI

Statira bimaculata Borchmann, 1936: 291 [junior primary homonym of Statira bimaculata Pic, 1912].

Statira pici Blackwelder, 1945: 500. Replacement name for Statira bimaculata Borchmann, 1936.

Statira picta Champion, 1889 NIC PAN

Statira picta Champion, 1889: 36.

Statira pilifera Champion, 1893MEX (VE)

Statira pilifera Champion, 1893b: 451.

Statira pilipes Champion, 1889MEX (CI)

Statira pilipes Champion, 1889: 43.

Statira pluripunctata Horn, 1888USA (AZ NM TX UT) MEX (GE)

Statira pluripunctata Horn, 1888: 29.

Statira sulcicrus Champion, 1889: 51. Synonymy: Parsons (1966: 245).

Statira pueblensis Champion, 1889MEX (PU)

Statira pueblensis Champion, 1889: 51.

Statira pulchella Mäklin, 1863USA (TX) MEX (SL TA VE) NIC

Statira pulchella Mäklin, 1863: 589.

Statira punctatissima Champion, 1889MEX (CI)

Statira punctatissima Champion, 1889: 38.

Statira punctipennis Champion, 1889 GUA

Statira punctipennis Champion, 1889: 28.

Statira reticulaticollis Borchmann, 1936 NIC CRI

Statira reticulaticollis Borchmann, 1936: 269.

Statira robusta Schaeffer, 1905USA (AZ CO NM TX)

Statira robusta Schaeffer, 1905a: 180.

Statira rugicollis Champion, 1889MEX (VE)

Statira rugicollis Champion, 1889: 48.

Statira rugipes Champion, 1889MEX (DU)

Statira rugipes Champion, 1889: 52.

Statira schmidti Borchmann, 1936 CRI

Statira schmidti Borchmann, 1936: 304.

Statira scitula Champion, 1889MEX (VE) GUA

Statira scitula Champion, 1889: 10.

Statira setigera Champion, 1889MEX (GU)

Statira setigera Champion, 1889: 41.

Statira simplex Borchmann, 1936 CRI

Statira simplex Borchmann, 1936: 284.

Statira sobrina Champion, 1889 MEX

Statira sobrina Champion, 1889: 22.

Statira spiculifera Champion, 1893MEX (VE)

Statira spiculifera Champion, 1893b: 451.

Statira suavis Champion, 1889MEX (DU)

Statira suavis Champion, 1889: 15.

Statira subnitida LeConte, 1866MEX (BC BS)

Statira subnitida LeConte, 1866b: 141.

Statira testacea Champion, 1889MEX (VE)

Statira testacea Champion, 1889: 28.

Statira tolensis Champion, 1889 PAN

Statira tolensis Champion, 1889: 20.

Statira triangulifer Champion, 1889MEX (CI VE) GUABEL

Statira triangulifer Champion, 1889: 34.

Statira tristis Mäklin, 1875 MEX

Statira tristis Mäklin, 1875a: 639.

Statira tropicalis Champion, 1889MEX (VE) GUABELNIC

Statira tropicalis Champion, 1889: 10.

Statira tuberculifera Champion, 1889 GUA

Statira tuberculifera Champion, 1889: 32.

Statira tuberosa Champion, 1889MEX (VE)

Statira tuberosa Champion, 1889: 31.

Statira variabilis inexpecta Borchmann, 1936 PAN

Statira variabilis var. inexpecta Borchmann, 1936: 282.

Statira variabilis variabilis Champion, 1889 GUA PAN

Statira variabilis Champion, 1889: 11.

Statira veraecrucis Champion, 1889MEX (VE)

Statira veraecrucis Champion, 1889: 35.

Statira veraepacis Champion, 1889 GUA

Statira veraepacis Champion, 1889: 24.

Statira vilis Mäklin, 1863MEX (CI TB VE) GUABELPAN

Statira vilis Mäklin, 1863: 592.

Statira villosa Champion, 1889MEX (GE)

Statira villosa Champion, 1889: 39.

Statira viridicollis Champion, 1889 PAN

Statira viridicollis Champion, 1889: 17.

Statira vittata Champion, 1896 LAN

Statira vittata Champion, 1896: 37.

Genus Uroplatopsis Champion, 1889 [F]

Uroplatopsis Champion, 1889: 53. Type species: Uroplatopsis imitator Champion, 1889, subsequent designation (Lucas 1920: 666).

Uroplatopsis appendiculata Champion, 1889 PAN

Uroplatopsis appendiculata Champion, 1889: 59.

Uroplatopsis dilaticornis Champion, 1889 PAN

Uroplatopsis dilaticornis Champion, 1889: 58.

Uroplatopsis excavata Champion, 1889 PAN

Uroplatopsis excavata Champion, 1889: 59.

Uroplatopsis imitator Champion, 1889 NIC

Uroplatopsis imitator Champion, 1889: 54.

Uroplatopsis mimica Champion, 1889 PAN

Uroplatopsis mimica Champion, 1889: 57.

Uroplatopsis planicollis Champion, 1889 PAN

Uroplatopsis planicollis Champion, 1889: 56.

Uroplatopsis reducta Pic, 1931 CRI

Uroplatopsis reducta Pic, 1931: 33.

Uroplatopsis reticulata Champion, 1889 PAN

Uroplatopsis reticulata Champion, 1889: 56.

Uroplatopsis simulans nevermanni Borchmann, 1936 CRI

Uroplatopsis simulans var. nevermanni Borchmann, 1936: 482.

Uroplatopsis simulans simulans Champion, 1889 PAN

Uroplatopsis simulans Champion, 1889: 58.

Uroplatopsis vermiculata Champion, 1889 NIC

Uroplatopsis vermiculata Champion, 1889: 55.

Tribe Lupropini Lesne, 1926

Lypropsini Lesne, 1926: 68. Type genus: Luprops Hope, 1833 [as Lyprops, incorrect subsequent spelling of the type genus name, not in prevailing usage].

Genus Lorelus Sharp, 1876 [M]

Lorelus Sharp, 1876: 76. Type species: Lorelus priscus Sharp, 1876, monotypy.

Lorelopsis Champion, 1896: 15. Type species: Lorelopsis pilosus Champion, 1896, monotypy. Synonymy: Doyen (1993: 295).

Lorelus angustulus Champion, 1913 GUA

Lorelus angustulus Champion, 1913: 165.

Lorelus bicolor Doyen, 1993 PRI

Lorelus bicolor Doyen, 1993: 296.

Lorelus brevicornis Champion, 1896 LAN

Lorelus brevicornis Champion, 1896: 14.

Lorelus breviusculus Champion, 1913 PAN

Lorelus breviusculus Champion, 1913: 165.

Lorelus cribricollis Kaszab, 1940LAN (Guadeloupe)

Lorelus cribricollis Kaszab, 1940: 156.

Lorelus curticollis Champion, 1913MEX (VE) GUAPAN

Lorelus curticollis Champion, 1913: 164.

Lorelus curvipes Champion, 1913 GUA

Lorelus curvipes Champion, 1913: 163.

Lorelus exilis Champion, 1913 GUA

Lorelus exilis Champion, 1913: 166.

Lorelus glabratus Doyen, 1993 PRI

Lorelus glabratus Doyen, 1993: 297.

Lorelus guadeloupensis Kaszab, 1940LAN (Guadeloupe)

Lorelus guadeloupensis Kaszab, 1940: 155.

Lorelus pilosus (Champion, 1896) LAN

Lorelopsis pilosus Champion, 1896: 16.

Lorelus trapeziderus Champion, 1913 GUA PAN

Lorelus trapeziderus Champion, 1913: 167.

Lorelus wolcotti Doyen, 1993 PRI

Lorelus wolcotti Doyen, 1993: 295.

Incertae sedis: Lagriinae

Genus Pseudesarcus Champion, 19138 [M]

Pseudesarcus Champion, 1913: 115. Type species: Pseudesarcus villosus Champion, 1913, original designation.

Pseudesarcus villosus Champion, 1913 PAN

Pseudesarcus villosus Champion, 1913: 116.

Subfamily NILIONINAE Oken, 1843

Nilioniden Oken, 1843: 484. Type genus: Nilio Latreille, 1802.

Genus Nilio Latreille, 1802 [M]

Nilio Latreille, 1802: 179 [as Nilion]9. Type species: Coccinella villosa Fabricius, 1787, monotypy.

Subgenus Nilio Latreille, 1802

Nilio Latreille, 1802: 179 [as Nilion]. Type species: Coccinella villosa Fabricius, 1787, monotypy.

Nilio chiriquensis Champion, 1888 PAN

Nilio chiriquensis Champion, 1888: 471.

Nilio fulvopilosus Champion, 1888 PAN

Nilio fulvo-pilosus Champion, 1888: 471.

Nilio lebasi J. Thomson, 18609PAN / SANew North American record

Nilio lebasei J. Thomson, 1860: 10.

Nilio sallei J. Thomson, 1860MEX (VE) GUA

Nilio sallei J. Thomson, 1860: 10.

Nilio sallaei Champion, 1888: 470. Unjustified emendation of Nilio sallei J. Thomson, 1860, not in prevailing usage.

Nilio thomsoni Champion, 1888 GUA NIC PAN

Nilio thomsoni Champion, 1888: 471.

Nilio villosus (Fabricius, 1787)PAN / SA

Coccinella villosa Fabricius, 1787: 379 [junior primary homonym of Coccinella villosa Geoffroy, 1785].

Subfamily PHRENAPATINAE Solier, 1834

Phrépatides Solier, 1834: 488. Type genus: Phrenapates Gray, 1832.

Tribe Archaeoglenini Watt, 1975

Archaeoglenini Watt, 1975: 412. Type genus: Archaeoglenes Broun, 1893.

Genus Archaeoglenes Broun, 1893 [M]

Archaeoglenes Broun, 1893: 188. Type species: Archaeoglenes costipennis Broun, 1893, monotypy.

Archaeoglenes bollensis Watrous, 1982 PAN

Archaeoglenes bollensis Watrous, 1982: 140.

Archaeoglenes occidentalis Lawrence, 1979MEX (CI) BELPAN / SA

Archaeoglenes occidentalis Lawrence [in Doyen and Lawrence], 1979: 358.

Archaeoglenes pecki Lawrence, 1979 JAM

Archaeoglenes pecki Lawrence [in Doyen and Lawrence], 1979: 358.

Archaeoglenes puntaensis Watrous, 1982 PAN

Archaeoglenes puntaensis Watrous, 1982: 141.

Tribe Penetini Lacordaire, 1859

Pénétides Lacordaire, 1859: 318. Type genus: Peneta Lacordaire, 1859.

Phthorini Boddy, 1965: 144. Type genus: Phtora Mulsant, 1854 [as Phthora, incorrect subsequent spelling, not in prevailing usage].

Genus Clamoris des Gozis, 1886 [F]

Phtora Mulsant, 1854: 228 [junior homonym of Phtora Germar, 1835]. Type species: Phtora crenata Mulsant, 1854, monotypy.

Clamoris des Gozis, 1886: 25. Replacement name for Phtora Mulsant, 1854.

Phthora Champion, 1893a: 531. Unjustified emendation of Phtora Mulsant, 1854, not in prevailing usage.

Clamoris americana (Horn, 1874) [Fig. 6] CAN (BC) USA (CA OR WA)

Figure 6. 

Clamoris americana (Horn, 1874). Scale bar = 1 mm.

Phthora americana Horn, 1874a: 35.

Clamoris armata (Champion, 1893) GUA

Phthora armata Champion, 1893a: 532.

Clamoris elongata (Champion, 1893)MEX (VE) NIC

Phthora elongata Champion, 1893a: 532.

Genus Cleolaus Champion, 1886 [M]

Cleolaus Champion, 1886: 142. Type species: Peneta sommeri Lacordaire, 1859, original designation.

Cleolaus sommeri (Lacordaire, 1859)MEX (OA)

Peneta sommeri Lacordaire, 1859: 320.

Genus Daochus Champion, 1886 [M]

Daochus Champion, 1886: 139. Type species: Daochus mandibularis Champion, 1886, monotypy.

Daochus mandibularis Champion, 1886 GUA BEL

Daochus mandibularis Champion, 1886: 140.

Genus Dioedus LeConte, 1862 [M]

Dioedus LeConte, 1862a: 238. Type species: Dioedus punctatus LeConte, 1862, monotypy.

Arrhabaeus Champion, 1886: 144. Type species: Arrhabaeus convexus Champion, 1886, monotypy. Synonymy: Kaszab (1977b: 314).

Dioedus convexus (Champion, 1886) CRI PAN

Arrhabaeus convexus Champion, 1886: 145.

Dioedus debilis (Champion, 1896) LAN

Arrhabaeus debilis Champion, 1896: 20.

Dioedus guadeloupensis (Fleutiaux and Sallé, 1890) LAN

Arrhabaeus guadeloupensis Fleutiaux and Sallé, 1890: 424.

Dioedus minor (Fleutiaux and Sallé, 1890) LAN

Arrhabaeus guadeloupensis minor Fleutiaux and Sallé, 1890: 425.

Dioedus punctatus LeConte, 1862 [Fig. 7] CAN (ON) USA (AL CT DC FL IA IL IN KS MD MI MO NC NJ NY OH SC VA WI WV) / PRI

Figure 7. 

Dioedus punctatus LeConte, 1862. Scale bar = 1 mm.

Dioedus punctatus LeConte, 1862a: 238.

Genus Peneta Lacordaire, 1859 [F]

Peneta Lacordaire, 1859: 319. Type species: Peneta lebasii Lacordaire, 1859, subsequent designation (Lucas 1920: 492).

Peneta costaricensis Gebien, 1928 CRI

Peneta costaricensis Gebien, 1928a: 147.

Peneta nevermanni Gebien, 1928 CRI

Peneta nevermanni Gebien, 1928a: 148.

Peneta nuchicornis Gebien, 1928 CRI PAN

Peneta nuchicornis Gebien, 1928a: 146.

Peneta obtusicornis Kirsch, 1866PAN / SA

Peneta obtusicornis Kirsch, 1866: 191.

Peneta panamensis Champion, 1886: 142. Synonymy: Champion (1893a: 531).

Genus Telchis Champion, 1886 [M]

Telchis Champion, 1886: 142. Type species: Telchis clavicornis Champion, 1886, monotypy.

Telchis clavicornis Champion, 1886 PAN

Telchis clavicornis Champion, 1886: 143.

Genus Zypoetes Champion, 1893 [M]

Zypoetes Champion, 1893a: 532. Type species: Zypoetes epieroides Champion, 1893, monotypy.

Zypoetes epieroides Champion, 1893MEX (VE) GUABELNIC

Zypoetes epieroides Champion, 1893a: 533.

Tribe Phrenapatini Solier, 1834

Phrépatides Solier, 1834: 488. Type genus: Phrenapates Gray, 1832.

Genus Delognatha Lacordaire, 1859 [F]

Delognatha Lacordaire, 1859: 315. Type species: Delognatha lacordairei Lacordaire, 1859, subsequent designation (Gebien 1940: 756). Note. The name Delognatha Agassiz, 1846 has been suppressed for the purposes of both the Principle of Priority and the Principle of Homonymy in Opinion 2250 (ICZN 2010).

Delognatha persimilis Gebien, 1928 CRI

Delognatha persimilis Gebien, 1928a: 142.

Genus Phrenapates Gray, 1832 [M]

Phrenapates Gray [in Griffith and Pidgeon], 1832: 91. Type species: Phrenapates bennettii Gray, 1832, monotypy.

Phrenapates bennettii Gray, 1832GUANICCRIPAN / SA

Phrenapates bennettii Gray [in Griffith and Pidgeon], 1832: 91.

Subfamily PIMELIINAE Latreille, 1802

Pimeliariae Latreille, 1802: 166. Type genus: Pimelia Fabricius, 1775.

Tribe Anepsiini LeConte, 1862

Anepsiini LeConte, 1862a: 215. Type genus: Anepsius LeConte, 1851.

Batuliini Horn, 1870: 270. Type genus: Batulius LeConte, 1851.

Anchommini Horn, 1878b: 558. Type genus: Anchomma LeConte, 1858.

Genus Anchomma LeConte, 1858 [N]

Anchomma LeConte, 1858b: 63. Type species: Anchomma costatum LeConte, 1858, monotypy.

Anchomma costatum LeConte, 1858USA (CA)

Anchomma costatum LeConte, 1858b: 63.

Genus Anepsius LeConte, 1851 [M]

Anepsius LeConte, 1851: 147. Type species: Anepsius delicatulus LeConte, 1851, monotypy.

Anepsius delicatulus LeConte, 1851USA (AZ CA NV UT) MEX (SO)

Anepsius delicatulus LeConte, 1851: 148.

Anepsius catenulosus Casey, 1907: 505. Synonymy (with A. atratus Casey): Casey (1911: 254).

Anepsius atratus Casey, 1907: 506. Synonymy: Doyen (1987: 351).

Anepsius brunneus Casey, 1907: 506. Synonymy: Doyen (1987: 351).

Anepsius nebulosus Casey, 1907: 507. Synonymy: Doyen (1987: 351).

Anepsius bicolor Casey, 1907: 507. Synonymy: Doyen (1987: 351).

Anepsius deficiens Casey, 1907: 507. Synonymy: Doyen (1987: 351).

Anepsius minutus Doyen, 1987USA (TX) MEX (NL)

Anepsius minutus Doyen, 1987: 352.

Anepsius montanus Casey, 1891 [Fig. 8] CAN (AB) USA (CO ND NE NM WY)

Figure 8. 

Anepsius montanus Casey, 1891. Scale bar = 1 mm.

Anepsius montanus Casey, 1891: 55.

Anepsius valens Casey, 1907USA (AZ)

Anepsius valens Casey, 1907: 504.

Genus Batuliodes Casey, 1907 [M]

Batuliodes Casey, 1907: 499. Type species: Batulius rotundicollis LeConte, 1851, original designation.

Batuliodes confluens (Blaisdell, 1923)MEX (BC BS)

Anepsius confluens Blaisdell, 1923: 243.

Anepsius angulatus Blaisdell, 1923: 244. Synonymy: Doyen (1987: 366).

Batuliodes obesus Doyen, 1987USA (CA)

Batuliodes obesus Doyen, 1987: 369.

Batuliodes rotundicollis (LeConte, 1851)USA (AZ CA NV)

Batulius rotundicollis LeConte, 1851: 148.

Batuliodes spatulatus Doyen, 1987USA (AZ CA UT) MEX (SO)

Batuliodes spatulatus Doyen, 1987: 368.

Batuliodes wasbaueri Doyen, 1987USA (CA) MEX (BC)

Batuliodes wasbaueri Doyen, 1987: 367.

Genus Batuliomorpha Doyen, 1987 [F]

Batuliomorpha Doyen, 1987: 359. Type species: Batuliomorpha comata Doyen, 1987, original designation.

Batuliomorpha comata Doyen, 1987USA (AZ CA)

Batuliomorpha comata Doyen, 1987: 361.

Batuliomorpha imperialis Doyen, 1987USA (CA)

Batuliomorpha imperialis Doyen, 1987: 359.

Batuliomorpha tibiodentata Doyen, 1987MEX (BS)

Batuliomorpha tibiodentata Doyen, 1987: 362.

Genus Batulius LeConte, 1851 [M]

Batulius LeConte, 1851: 148. Type species: Batulius setosus LeConte, 1851, subsequent designation (Casey 1907: 497).

Batulius setosus LeConte, 1851USA (AZ CA) MEX (BC)

Batulius setosus LeConte, 1851: 148.

Tribe Asidini Fleming, 1821

Asidadae Fleming, 1821: 51. Type genus: Asida Latreille, 1802.

Astroti Horn, 1870: 289. Type genus: Astrotus J.L. LeConte, 1858.

Craniotini LeConte and Horn, 1883: 361. Type genus: Craniotus LeConte, 1851.

Genus Ardamimicus Smith, 2013 [M]

Ardamimicus Smith, 2013: 601. Type species: Ardamimicus cognatoi Smith, 2013, original designation.

Ardamimicus cognatoi Smith, 2013USA (TX) MEX (CH DU)

Ardamimicus cognatoi Smith, 2013: 602.

Genus Craniotus LeConte, 1851 [M]

Craniotus LeConte, 1851: 142. Type species: Craniotus pubescens LeConte, 1851, monotypy.

Craniotus mardecortesi Aalbu, Smith and Sánchez Piñero, 2015MEX (BC)

Craniotus mardecortesi Aalbu, Smith and Sánchez Piñero, 2015: 94.

Craniotus pubescens LeConte, 1851USA (AZ CA NV) MEX (BC)

Craniotus pubescens LeConte, 1851: 142.

Craniotus blaisdelli Tanner, 1963: 169. Synonymy: Aalbu et al. (2015: 96).

Craniotus triplehorni Aalbu, Smith and Sánchez Piñero, 2015MEX (BC)

Craniotus triplehorni Aalbu, Smith and Sánchez Piñero, 2015: 95.

Genus Ferveoventer Smith, 2013 [M]

Ferveoventer Smith, 2013: 604. Type species: Ferveoventer browni Smith, 2013, original designation.

Ferveoventer browni Smith, 2013USA (NM TX)

Ferveoventer browni Smith, 2013: 605.

Ferveoventer planatus (Champion, 1884)MEX (MO)

Ologlyptus planatus Champion, 1884: 69.

Genus Heterasida Casey, 1912 [F]

Heterasida Casey, 1912: 76, 165. Type species: Pelecyphorus bifurcus LeConte, 1861, original designation.

Heterasida bifurcus (LeConte, 1861)10MEX (BS)

Pelecyphorus bifurcus LeConte, 1861a: 337.

Heterasida connivens (LeConte, 1866)MEX (BS)

Pelecyphorus connivens LeConte, 1866b: 110.

Heterasida tantilla Casey, 1912: 167. Synonymy: Smith (2013: 607).

Heterasida exilis Casey, 1912: 168. Synonymy: Smith (2013: 607).

Genus Litasida Casey, 1912 [F]

Litasida Casey, 1912: 77, 184. Type species: Litasida townsendi Casey, 1912, original designation.

Litasida townsendi Casey, 1912USA (AZ) MEX (CH)

Litasida townsendi Casey, 1912: 185.

Genus Micrasida Smith, 2013 [F]

Micrasida Smith, 2013: 608. Type species: Micrasida obrienorum Smith, 2013, original designation.

Micrasida obrienorum Smith, 2013MEX (NL)

Micrasida obrienorum Smith, 2013: 608.

Genus Microschatia Solier, 1836 [F]

Microschatia Solier, 1836: 474. Type species: Microschatia punctata Solier, 1836, monotypy.

Pycnonotida Casey, 1912: 75, 89. Type species: Microschatia inaequalis LeConte, 1851, original designation. Synonymy: Brown and Doyen (1992: 546).

Acroschatia Wilke, 1922: 269. Type species: Microschatia robusta Horn, 1893, original designation. Synonymy: Brown and Doyen (1992: 546).

Microschatia cedrosensis Brown and Doyen, 1992MEX (BS)

Microschatia cedrosensis Brown and Doyen, 1992: 568.

Microschatia championi Horn, 1893USA (AZ CA) MEX (BC BS)

Microschatia championi Horn, 1893: 140.

Microschatia costulata Brown and Doyen, 1992USA (CA) MEX (BC)

Microschatia costulata Brown and Doyen, 1992: 570.

Microschatia inaequalis LeConte, 1851USA (CA) MEX (BC)

Microschatia inaequalis LeConte, 1851: 129.

Microschatia puncticollis LeConte, 1851: 129. Synonymy: Horn (1870: 282).

Pycnonotida laxicollis Casey, 1912: 91. Synonymy: Brown and Doyen (1992: 572).

Pycnonotida araneoides Casey, 1912: 92. Synonymy: Brown and Doyen (1992: 572).

Pycnonotida inaequalis diversa Casey, 1912: 92. Synonymy: Brown and Doyen (1992: 572).

Pycnonotida impar Casey, 1912: 93. Synonymy: Brown and Doyen (1992: 572).

Microschatia morata Horn, 1878USA (AZ NM) MEX (CH DU SO)

Microschatia morata Horn, 1878a: 56.

Microschatia planata Doyen and Brown, 1992MEX (BC BS)

Microschatia planata Doyen and Brown [in Brown and Doyen], 1992: 576.

Microschatia polita Horn, 1893USA (AZ)

Microschatia polita Horn, 1893: 141.

Microschatia punctata Solier, 1836MEX (HI QU)

Microschatia punctata Solier, 1836: 475.

Microschatia robusta Horn, 1893USA (TX) MEX (CH CO NL TA)

Microschatia robusta Horn, 1893: 142.

Microschatia rockefelleri Pallister, 1954MEX (CH DU)

Microschatia rockefelleri Pallister, 1954: 15.

Microschatia solieri Brown and Doyen, 1992MEX (HI)

Microschatia solieri Brown and Doyen, 1992: 552.

Microschatia sulcipennis LeConte, 1858USA (TX)

Microschatia sulcipennis LeConte, 1858a: 18.

Genus Pelecyphorus Solier, 1836 [M]

Pelecyphorus Solier, 1836: 467. Type species: Pelecyphorus mexicanus Solier, 1836, subsequent designation (Hope 1841: 110).

Subgenus Astrotus LeConte, 1858

Astrotus LeConte, 1858a: 19. Type species: Microschatia contorta LeConte, 1853, original designation.

Pelecyphorus alveolatus (Casey, 1912)USA (TX)

Astrotus alveolatus Casey, 1912: 83.

Pelecyphorus contortus (LeConte, 1853)USA (TX)

Microschatia contorta LeConte, 1853: 446.

Pelecyphorus fasciculatus (Champion, 1892)MEX (MO)

Asida fasciculata Champion, 1892: 495.

Pelecyphorus guanajuatensis (Champion, 1884)MEX (CH DU GU)

Asida guanajuatensis Champion, 1884: 56.

Bothrasida mucorea Wilke, 1922: 270. New synonymy [ADS].

Pelecyphorus hebes (Champion, 1892)MEX (DU)

Ologlyptus hebes Champion, 1892: 506.

Pelecyphorus regularis (Horn, 1870)USA (TX) MEX (TA)

Astrotus regularis Horn, 1870: 290.

Subgenus Pelecyphorus Solier, 1836

Pelecyphorus Solier, 1836: 467. Type species: Pelecyphorus mexicanus Solier, 1836, subsequent designation (Hope 1841: 110).

Pelecyphorus mexicanus Solier, 1836MEX (PU QU VE)

Pelecyphorus mexicanus Solier, 1836: 469.

Subgenus Pleisiasida Smith, 2013

Parasida Casey, 1912: 76, 126 [junior homonym of Parasida Daday, 1904]. Type species: Parasida laciniata Casey, 1912, original designation.

Pleisiasida Smith, 2013: 610. Replacement name for Parasida Casey, 1912.

Pelecyphorus asidoides Solier, 1836MEX (HI ME)11

Pelecyphorus asidoides Solier, 1836: 471.

Parasida zacualpanicola Wilke, 1922: 272. New synonymy [ADS].

Pelecyphorus bibasalis (Casey, 1912)MEX (DU)

Parasida bibasalis Casey, 1912: 128.

Pelecyphorus dispar (Champion, 1892)MEX (CH DU)

Asida dissimilis Champion, 1884: 59 [junior primary homonym of Asida dissimilis Allard, 1869].

Asida dispar Champion, 1892: 496. Replacement name for Asida dissimilis Champion, 1884.

Stenosides kulzeri Pallister, 1954: 12. New synonymy [ADS].

Stenosides bisinuatus Pallister, 1954: 13. New synonymy [ADS].

Parasida trisinuata Pallister, 1954: 22. New synonymy [ADS].

Pelecyphorus fallax (Champion, 1884)MEX (DU FD GU HI ME PU)

Asida fallax Champion, 1884: 57.

Asida favosa Champion, 1884: 58. New synonymy [ADS].

Asida similata Champion, 1884: 58. New synonymy [ADS].

Pelecyphorus foveolatus Solier, 1836MEX (OA VE)

Pelecyphorus foveolatus Solier, 1836: 472.

Pelecyphorus indutus (Champion, 1884)MEX (OA)

Asida induta Champion, 1884: 56.

Ologlyptus bicarinatus Champion, 1884: 69. New synonymy [ADS].

Pelecyphorus intricatus (Champion, 1892)MEX (CH JA)

Asida intricata Champion, 1892: 493.

Pelecyphorus laticollis (Champion, 1884)MEX (DU GU)

Asida laticollis Champion, 1884: 58.

Pelecyphorus liratus (LeConte, 1854)USA (AZ NM) MEX (CH DU)

Euschides liratus LeConte, 1854c: 223.

Parasida laciniata Casey, 1912: 128. New synonymy [ADS].

Parasida cristata Pallister, 1954: 24. New synonymy [ADS].

Pelecyphorus longipennis (Champion, 1884)MEX (OA PU VE)

Asida longipennis Champion, 1884: 56.

Parasida esperanzae Wilke, 1922: 271. New synonymy [ADS].

Parasida mixtecae Wilke, 1922: 271. New synonymy [ADS].

Pelecyphorus obliviosus (Wilke, 1922)MEX (CH DU)

Parasida obliviosa Wilke, 1922: 270.

Pelecyphorus planatulus (Casey, 1912)MEX (DU)

Parasida planatula Casey, 1912: 129.

Pelecyphorus scutellaris (Champion, 1884)MEX (DU FD ME OA PU VE)

Asida scutellaris Champion, 1884: 56.

Parasida tolucana Casey, 1912: 130. New synonymy [ADS].

Pelecyphorus sexcostatus LeConte, 1861MEX (BS)

Pelecyphorus sexcostatus LeConte, 1861a: 337.

Pelecyphorus tristis (Champion, 1884)MEX (PU VE)

Asida tristis Champion, 1884: 55.

Parasida purpusi Wilke, 1922: 271. New synonymy [ADS].

Subgenus Poliorcetes Champion, 1884

Poliorcetes Champion, 1884: 70. Type species: Poliorcetes platesthoides Champion, 1884, monotypy.

Pelecyphorus platesthoides (Champion, 1884)MEX (OA)

Poliorcetes platesthoides Champion, 1884: 71.

Subgenus Sicharbas Champion, 1884

Sicharbas Champion, 1884: 67. Type species: Sicharbas lobatus Champion, 1884, monotypy.

Pelecyphorus debilis (Champion, 1884)MEX (PU)

Astrotus debilis Champion, 1884: 66.

Pelecyphorus erosus (Champion, 1892)MEX (HI)

Astrotus erosus Champion, 1892: 504.

Astrotus nosodermoides Champion, 1892: 505. New synonymy [ADS].

Pelecyphorus lobatus (Champion, 1884)MEX (GE MO)

Sicharbas lobatus Champion, 1884: 67.

Pelecyphorus seticornis (Champion, 1884)MEX (ME)

Astrotus seticornis Champion, 1884: 67.

Astrotus seticornis var. humeralis Champion, 1884: 67. New synonymy [ADS].

Pelecyphorus undatus (Champion, 1892)MEX (DU)

Astrotus undatus Champion, 1892: 504.

Subgenus Stenosides Solier, 1836

Stenosides Solier, 1836: 484. Type species: Stenosides graciliformis Solier, 1836, monotypy.

Pactostoma LeConte, 1858a: 19. Type species: Asida anastomosis Say, 1824, original designation. Synonymy (with Ologlyptus Lacordaire): LeConte (1862a: 222).

Ologlyptus Lacordaire, 1859: 158. Unnecessary replacement name for Stenosides Solier, 1836.

Pelecyphorus anastomosis (Say, 1824)USA (AR AZ CO KS NM TX) MEX (CH DU)

Asida anastomosis Say, 1824a: 256.

Pactostoma anastomosis salebrosa Casey, 1912: 87. Synonymy: Pallister (1954: 12).

Pactostoma breviuscula Casey, 1912: 87. New synonymy [ADS].

Pactostoma exoleta Casey, 1912: 87. New synonymy [ADS].

Pactostoma luteotecta Casey, 1912: 88. New synonymy [ADS].

Pactostoma monticola Casey, 1912: 88. New synonymy [ADS].

Pactostoma obtecta Casey, 1912: 89. New synonymy [ADS].

Pactostoma sigillata Casey, 1912: 89. New synonymy [ADS].

Pelecyphorus graciliformis (Solier, 1836)MEX (FD HI OA PU SL)

Stenosides graciliformis Solier, 1836: 486.

Ologlyptus canus Champion, 1884: 68. New synonymy [ADS].

Ologlyptus sinuaticollis Champion, 1884: 69. New synonymy [ADS].

Pelecyphorus limosus (Champion, 1884) MEX

Astrotus limosus Champion, 1884: 66.

Pelecyphorus texanus (Wickham, 1903)USA (TX)

Ologlyptus texanus Wickham, 1903: 72.

Subgenus Ucalegon Champion, 1884

Ucalegon Champion, 1884: 65. Type species: Ucalegon pulchellus Champion, 1884, monotypy.

Pelecyphorus pulchellus (Champion, 1884)MEX (GE OA)

Ucalegon pulchellus Champion, 1884: 65.

Subgenus Zaleucus Champion, 1892

Zamolxis Champion, 1884: 70 [junior homonym of Zamolxis Stål, 1865]. Type species: Zamolxis dilatatus Champion, 1884, monotypy.

Zaleucus Champion, 1892: 491. Replacement name for Zamolxis Champion, 1884.

Pelecyphorus dilatatus (Champion, 1884)MEX (PU)

Zamolxis dilatatus Champion, 1884: 70.

Genus Philolithus Lacordaire, 1858 [M]

Philolithus Lacordaire [in LeConte], 1858a: 18. Type species: Pelecyphorus carinatus LeConte, 1851, subsequent designation (Casey 1912: 79).

Subgenus Glyptasida Casey, 1912

Glyptasida Casey, 1912: 75, 95. Type species: Pelecyphorus sordidus LeConte, 1853, original designation.

Philolithus aeger (LeConte, 1858)USA (NM TX)

Pelecyphorus aeger LeConte, 1858a: 19.

Glyptasida sycophanta Casey, 1912: 104. Synonymy: Lockwood and Pollock (2009: 21).

Philolithus rugosissimus (Champion, 1884)USA (AZ NM) MEX (CH CO SL)

Asida rugosissima Champion, 1884: 53.

Philolithus sordidus (LeConte, 1853) [Fig. 9] CAN (AB SK) USA (AZ CO KS MT ND NE NM OK SD TX UT WY) MEX (CH CO DU ZA)

Figure 9. 

Philolithus (Glyptasida) sordidus (LeConte, 1853). Scale bar = 1 mm.

Pelecyphorus sordidus LeConte, 1853: 445.

Pelecyphorus subcostatus LeConte, 1853: 446. Synonymy: Henshaw (1882: 255).

Pelecyphorus irregularis LeConte, 1858a: 19. Synonymy: Champion (1892: 492).

Pelecyphorus costipennis LeConte, 1858a: 20. Synonymy: Champion (1892: 492).

Asida interrupta Champion, 1884: 53. Synonymy: Champion (1892: 492).

Glyptasida parvicollis Casey, 1912: 97. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida sordida porcatula Casey, 1912: 97. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida subpubescens Casey, 1912: 98. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida turgescens Casey, 1912: 98. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida turgescens furtiva Casey, 1912: 99. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida turgescens obesa Casey, 1912: 99. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida procrustes Casey, 1912: 99. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida costipennis fulvisetis Casey, 1912: 100. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida strigipennis Casey, 1912: 100. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida turbulenta Casey, 1912: 101. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida aegra imperfecta Casey, 1912: 102. Synonymy: Lockwood and Pollock (2009: 7).

Glyptasida aegra pigra Casey, 1912: 102. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida aegra plena Casey, 1912: 102. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida heres Casey, 1912: 103. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida crenicollis Casey, 1912: 103. Synonymy: Lockwood and Pollock (2009: 8).

Subgenus Gonasida Casey, 1912

Gonasida Casey, 1912: 75, 117. Type species: Pelecyphorus elatus LeConte, 1853, original designation.

Philolithus elatus compar (Casey, 1912)USA (CO ID KS NE NM NV OR UT WY)

Gonasida compar Casey, 1912: 120.

Gonasida elata reducta Casey, 1912: 121. New synonymy [based on Brown (1971: 262) unpublished thesis].

Gonasida elata prolixa Casey, 1912: 121. New synonymy [based on Brown (1971: 262) unpublished thesis].

Gonasida aucta Casey, 1912: 122. New synonymy [based on Brown (1971: 262) unpublished thesis].

Philolithus elatus difformis (LeConte, 1854)USA (AZ NM UT) MEX (CH)

Pelecyphorus difformis LeConte, 1854c: 223.

Gonasida alaticollis Casey, 1912: 122. New synonymy [based on Brown (1971: 255) unpublished thesis].

Philolithus elatus elatus (LeConte, 1853)USA (AZ NM TX) MEX (CH)

Pelecyphorus elatus LeConte, 1853: 445.

Gonasida gravida Casey, 1912: 119. New synonymy [based on Brown (1971: 251) unpublished thesis].

Philolithus elatus infernus (Casey, 1912)USA (AZ NM UT)

Gonasida inferna Casey, 1912: 119.

Subgenus Herthasida Wilke, 1922

Herthasida Wilke, 1922: 269. Type species: Asida ingens Champion, 1892, monotypy.

Philolithus ingens (Champion, 1892)MEX (CO)

Asida ingens Champion, 1892: 503.

Subgenus Philolithus Lacordaire, 1858

Philolithus Lacordaire [in LeConte], 1858a: 18. Type species: Pelecyphorus carinatus LeConte, 1851, subsequent designation (Casey 1912: 79).

Philolithus actuosus (Horn, 1870)USA (CA)

Asida actuosa Horn, 1870: 284.

Philolithus adversus (Casey, 1912)USA (AZ)

Pelecyphorus adversus Casey, 1912: 114.

Philolithus aegrotus aegrotus (LeConte, 1861)MEX (BC)

Pelecyphorus aegrotus LeConte, 1861a: 337.

Pelecyphorus aegrotus limbatus Casey, 1912: 107. New synonymy [ADS].

Philolithus carinatus (LeConte, 1851)USA (CA)

Pelecyphorus carinatus LeConte, 1851: 128 [junior secondary homonym of Asida carinata Solier, 1836].

Asida carinifera Gebien, 1910a: 128. Replacement name for Asida carinata (LeConte, 1851)12.

Philolithus densicollis (Horn, 1894) [Fig. 10] CAN (BC) USA (OR WA)

Figure 10. 

Philolithus (Philolithus) densicollis (Horn, 1894). Scale bar = 1 mm.

Asida densicollis Horn, 1894b: 417.

Pelecyphorus corrosus Casey, 1912: 117. Synonymy: Boddy (1965: 137).

Philolithus haruspex ellipsipennis (Casey, 1912)USA (UT)

Pelecyphorus haruspex ellipsipennis Casey, 1912: 116.

Philolithus haruspex haruspex (Casey, 1912)USA (AZ ID OR NV UT)

Pelecyphorus haruspex Casey, 1912: 115.

Philolithus jaegeri (Papp, 1961)USA (CA)

Pelecyphorus jaegeri Papp, 1961b: 107.

Philolithus morbillosus (LeConte, 1858)USA (AZ) MEX (SO)

Pelecyphorus morbillosus LeConte, 1858b: 74.

Pelecyphorus corporalis Casey, 1912: 107. New synonymy [ADS].

Pelecyphorus reptans Casey, 1912: 108. New synonymy [ADS].

Pelecyphorus socer Casey, 1912: 108. New synonymy [ADS].

Pelecyphorus abscissus Casey, 1912: 109. New synonymy [ADS].

Pelecyphorus fumosus Casey, 1912: 109. New synonymy [ADS].

Pelecyphorus parvus Casey, 1912: 110. New synonymy [ADS].

Pelecyphorus morbillosus pacatus Casey, 1912: 110. New synonymy [ADS].

Pelecyphorus morbillosus sobrius Casey, 1912: 110. New synonymy [ADS].

Pelecyphorus piceus Casey, 1912: 111. New synonymy [ADS].

Pelecyphorus piceus crudelis Casey, 1912: 111. New synonymy [ADS].

Pelecyphorus snowi Casey, 1912: 111. New synonymy [ADS].

Pelecyphorus subtenuis Casey, 1912: 112. New synonymy [ADS].

Philolithus opimus (Casey, 1912)USA (CA)

Pelecyphorus opimus Casey, 1912: 115.

Philolithus pantex (Casey, 1912)USA (NV UT)

Pelecyphorus pantex Casey, 1912: 116.

Philolithus porcatus (Papp, 1961)USA (CA)

Pelecyphorus porcatus Papp, 1961b: 109.

Philolithus quadripennis (Casey, 1912)MEX (LC)

Pelecyphorus quadripennis Casey, 1912: 113.

Philolithus reflexus (Casey, 1912)USA (CA)

Pelecyphorus reflexus Casey, 1912: 114.

Philolithus rugosus (Papp, 1961)USA (CA)

Pelecyphorus rugosus Papp, 1961c: 157.

Philolithus sophistes (Casey, 1912)USA (CA)

Pelecyphorus sophistes Casey, 1912: 113.

Philolithus uteanus (Casey, 1924)USA (UT)

Pelecyphorus uteanus Casey, 1924: 308.

Subgenus Tisamenes Champion, 1884

Tisamenes Champion, 1884: 64. Type species: Tisamenes truquii Champion, 1884, monotypy.

Philolithus truquii (Champion, 1884)MEX (FD HI)

Tisamenes truquii Champion, 1884: 64.

Genus Stenomorpha Solier, 1836 [F]

Stenomorpha Solier, 1836: 487. Type species: Stenomorpha blapsoides Solier, 1836, subsequent designation (Desmarest 1860: 150).

Euschides LeConte, 1851: 127. Unnecessary replacement name for Stenomorpha Solier, 1836.

Subgenus Asidina Casey, 1912

Asidina Casey, 1912: 169. Type species: Pelecyphorus parallelus LeConte, 1851, original designation.

Stenomorpha confluens (LeConte, 1851)USA (AZ CA) MEX (BC SO)

Pelecyphorus confluens LeConte, 1851: 128.

Stenomorpha parallela (LeConte, 1851)USA (AZ CA) MEX (SO)

Pelecyphorus parallelus LeConte, 1851: 128 [junior secondary homonym of Asida parallela Solier, 1836].

Asida neglecta Gebien, 1910a: 134. Replacement name for Asida parallela (LeConte, 1851)13.

Asidina teres Casey, 1912: 171. Synonymy: Triplehorn and Brown (1971: 84).

Asida parallela terricola Blaisdell, 1923: 254. Synonymy: Triplehorn and Brown (1971: 84).

Stenomorpha rugicollis (Triplehorn and Brown, 1971)USA (AZ)

Asidina rugicollis Triplehorn and Brown, 1971: 76.

Stenomorpha semilaevis (Horn, 1870)USA (AZ CA NV)

Asida semilaevis Horn, 1870: 284.

Subgenus Asidopsis Casey, 1912

Asidopsis Casey, 1912: 77, 185. Type species: Asida opaca Say, 1824, original designation.

Stenomorpha abbreviata (Casey, 1912)USA (NM)

Asidopsis abbreviata Casey, 1912: 198.

Stenomorpha cochisensis (Casey, 1912)USA (AZ NM)

Asidopsis cochisensis Casey, 1912: 189.

Stenomorpha coenosa (Casey, 1912)USA (NM)

Asidopsis coenosa Casey, 1912: 200.

Stenomorpha collaris (Champion, 1892)MEX (AG GU JA)

Asida marginicollis Champion, 1884: 60 [junior primary homonym of Asida marginicollis Rosenhauer, 1856].

Asida collaris Champion, 1892: 499. Replacement name for Asida marginicollis Champion, 1884.

Stenomorpha collega (Casey, 1912)USA (KS)

Asidopsis collega Casey, 1912: 198.

Stenomorpha consentanea (Casey, 1912)USA (CA)

Asidopsis consentanea Casey, 1912: 192.

Stenomorpha divaricata (Blaisdell, 1923)MEX (BS)

Asida divaricata Blaisdell, 1923: 255.

Stenomorpha dolosa (Casey, 1912)USA (MT SD WY)

Asidopsis dolosa Casey, 1912: 194.

Stenomorpha durangoensis (Casey, 1912)MEX (DU)

Asidopsis durangoensis Casey, 1912: 201.

Stenomorpha eximia (Casey, 1912)USA (TX)

Asidopsis eximia Casey, 1912: 188.

Stenomorpha forreri (Champion, 1884)MEX (DU)

Asida forreri Champion, 1884: 55.

Stenomorpha gracilipes (Casey, 1912)USA (AZ)

Asidopsis gracilipes Casey, 1912: 189.

Stenomorpha humeralis (Triplehorn and Flores, 2002)MEX (CH)

Asidopsis humeralis Triplehorn and Flores, 2002: 288.

Stenomorpha immunda (Casey, 1912)USA (NM) MEX (DU)

Asidopsis immunda Casey, 1912: 199.

Stenomorpha macra (Horn, 1883)USA (AZ NM)

Asidopsis macra Horn, 1883: 304.

Stenomorpha mancipata (Horn, 1878)USA (NM) MEX (CH)

Asida mancipata Horn, 1878a: 56.

Asidopsis woodgatei Casey, 1912: 197. Synonymy: Triplehorn and Flores (2002: 285).

Stenomorpha nitidula (Casey, 1912)USA (NM)

Asidopsis nitidula Casey, 1912: 196.

Stenomorpha obsidiana (Casey, 1912)USA (CO)

Asidopsis obsidiana Casey, 1912: 193.

Stenomorpha olsoni (Triplehorn and Flores, 2002)USA (AZ)

Asidopsis olsoni Triplehorn and Flores, 2002: 286.

Stenomorpha opaca (Say, 1824) [Fig. 11] CAN (AB SK) USA (CO KS MT ND NE NM SD TX)

Figure 11. 

Stenomorpha (Asidopsis) opaca (Say, 1824). Scale bar = 1 mm.

Asida opaca Say, 1824a: 254.

Stenomorpha pinalica (Casey, 1912)USA (AZ)

Asidopsis pinalica Casey, 1912: 190.

Stenomorpha planata (Horn, 1894)USA (CA) MEX

Asida planata Horn, 1894b: 415.

Stenomorpha polita futilis (Casey, 1912)USA (KS)

Asidopsis polita futilis Casey, 1912: 194.

Stenomorpha polita polita (Say, 1824)CAN (AB SK) USA (CO ID KS MT NE NM OK SD TX WY)

Asida polita Say, 1824a: 255.

Stenomorpha polita subopaca (Casey, 1912)USA (KS)

Asidopsis polita subopaca Casey, 1912: 193.

Stenomorpha quadricollis (Horn, 1880)USA (AZ NM)

Asidopsis quadricollis Horn, 1880: 151.

Stenomorpha servilis (Casey, 1912)USA (CO)

Asidopsis servilis Casey, 1912: 199.

Stenomorpha suavis (Casey, 1912)USA (AZ)

Asidopsis suavis Casey, 1912: 190.

Stenomorpha tensa (Casey, 1912)USA (CO)

Asidopsis tensa Casey, 1912: 197.

Subgenus Bothrasida Casey, 1912

Bothrasida Casey, 1912: 76, 122. Type species: Asida clathrata Champion, 1884, original designation.

Stenomorpha baroni (Casey, 1912)MEX (GE)

Bothrasida baroni Casey, 1912: 124.

Stenomorpha clathrata (Champion, 1884)MEX (GE ME MO OA PU)

Asida clathrata Champion, 1884: 54.

Stenomorpha funesta (Champion, 1884)MEX (PU)

Asida funesta Champion, 1884: 53.

Bothrasida sanctae-agnae Wilke, 1922: 270. New synonymy [ADS].

Subgenus Megasida Casey, 1912

Megasida Casey, 1912: 77, 202. Type species: Asida obliterata Champion, 1892, original designation.

Stenomorpha foeda (Champion, 1892)MEX (CO DU)

Asida foeda Champion, 1892: 498.

Stenomorpha latissima (Champion, 1892)MEX (DU)

Asida latissima Champion, 1892: 500.

Stenomorpha magnifica (Pallister, 1954)MEX (DU)

Megasida magnifica Pallister, 1954: 30.

Stenomorpha moricoides (Champion, 1892)MEX (CO DU)

Asida moricoides Champion, 1892: 497.

Stenomorpha obliterata (Champion, 1892)USA (NM TX) MEX (CH)

Asida obliterata Champion, 1892: 497.

Stenomorpha rufipes (Champion, 1884)MEX (SL)

Asida rufipes Champion, 1884: 62.

Stenomorpha segregata (Champion, 1892)MEX (CH CO DU)

Asida segregata Champion, 1892: 497.

Stenomorpha tarda (Champion, 1892)MEX (CO)

Asida tarda Champion, 1892: 498.

Stenomorpha tenuicollis (Triplehorn, 1967)USA (AZ NM TX)

Megasida tenuicollis Triplehorn, 1967: 40.

Stenomorpha zacatecensis (Pallister, 1954)MEX (ZA)

Megasida zacatecensis Pallister, 1954: 32.

Subgenus Notiasida Casey, 1912

Notiasida Casey, 1912: 76, 124. Type species: Notiasida abstrusa Casey, 1912, original designation.

Stenomorpha abstrusa (Casey, 1912)MEX (FD)

Notiasida abstrusa Casey, 1912: 125.

Stenomorpha evertissima (Casey, 1912)MEX (CH DU)

Notiasida evertissima Casey, 1912: 126.

Stenomorpha geminata (Champion, 1892)MEX (CH DU)

Asida geminata Champion, 1892: 492.

Stenomorpha lata (Champion, 1884)MEX (SL)

Asida lata Champion, 1884: 60.

Stenomorpha lugubris (Wilke, 1922)MEX (SL)

Pelecyphorus lugubris Wilke, 1922: 269.

Stenomorpha suturalis (Champion, 1884)MEX (FD OA VE)

Asida suturalis Champion, 1884: 55.

Subgenus Platasida Casey, 1912

Platasida Casey, 1912: 77, 182. Type species: Asida embaphionides Horn, 1894, original designation.

Stenomorpha embaphionides (Horn, 1894)MEX (BS)

Asida embaphionides Horn, 1894b: 419.

Asida flaccida Horn, 1896: 379. New synonymy [ADS].

Subgenus Pycnomorpha Motschulsky, 1870

Pycnomorpha Motschulsky, 1870: 398. Type species: Pycnomorpha californica Motschulsky, 1870, monotypy.

Stenomorpha californica (Motschulsky, 1870)MEX (BC)

Pycnomorpha californica Motschulsky, 1870: 399.

Stenomorpha gabbii (Horn, 1880)USA (CA) MEX (BS)

Asida gibbicollis Horn, 1870: 288 [junior primary homonym of Asida gibbicollis Pérez Arcas, 1865].

Asida gabbii Horn, 1880: 152. Replacement name for Asida gibbicollis Horn, 1870.

Stenomorpha tumidicollis Blaisdell, 1943MEX (BC)

Stenomorpha tumidicollis Blaisdell, 1943: 226.

Subgenus Stenomorpha Solier, 1836

Stenomorpha Solier, 1836: 487. Type species: Stenomorpha blapsoides Solier, 1836, subsequent designation (Desmarest 1860: 150).

Psilomera Motschulsky, 1870: 400. Type species: Pelecyphorus angulatus LeConte, 1851, monotypy. New synonymy [YB].

Stenomorpha advena (Casey, 1912)USA (CO)

Euschides advena Casey, 1912: 143.

Stenomorpha amplicollis (Casey, 1912)USA (CA)

Euschides amplicollis Casey, 1912: 153.

Stenomorpha angulata (LeConte, 1851)USA (CA)

Pelecyphorus angulatus LeConte, 1851: 127.

Stenomorpha blanda (Champion, 1884)MEX (AG GU JA)

Asida blanda Champion, 1884: 63.

Stenomorpha blapsoides alutacea Wilke, 1922MEX (ME)

Stenomorpha blapsoides alutacea Wilke, 1922: 272.

Stenomorpha blapsoides blapsoides Solier, 1836MEX (CO FD GU JA ME OA PU SI VE)

Stenomorpha blapsoides Solier, 1836: 491.

Stenomorpha brevimargo (Casey, 1912)USA (AZ)

Euschides brevimargo Casey, 1912: 134.

Stenomorpha caliginosa (Casey, 1912)USA (AZ)

Euschides caliginosus Casey, 1912: 137.

Stenomorpha captiosa (Horn, 1870)USA (CA)

Asida captiosa Horn, 1870: 287.

Stenomorpha clarissae Wilke, 1922MEX (ME)

Stenomorpha clarissae Wilke, 1922: 273.

Stenomorpha compressa (Horn, 1870)USA (CA)

Asida lecontei var. compressa Horn, 1870: 287.

Stenomorpha congruens congruens (Casey, 1912)USA (NM)

Euschides congruens Casey, 1912: 164.

Stenomorpha congruens lubrica (Casey, 1912)USA (AZ)

Euschides congruens lubricus Casey, 1912: 164.

Stenomorpha consobrina (Horn, 1870)USA (ID OR)

Asida consobrina Horn, 1870: 287.

Stenomorpha consors (Casey, 1912)USA (NM)

Euschides consors Casey, 1912: 162.

Stenomorpha consueta (Casey, 1912)USA (AZ)

Euschides consuetus Casey, 1912: 161.

Stenomorpha convexa (LeConte, 1859)USA (CA KS)

Euschides convexa LeConte, 1859a: 14.

Stenomorpha convexicollis (LeConte, 1854)USA (CA CO KS NM TX) MEX (CH DU)

Euschides convexicollis LeConte, 1854c: 224.

Stenomorpha corrugans (Casey, 1912)USA (AZ)

Euschides corrugans Casey, 1912: 143.

Stenomorpha costata Solier, 1836MEX (ME VE)

Stenomorpha costata Solier, 1836: 490.

Stenomorpha crassa (Casey, 1912)USA (CA)

Euschides crassus Casey, 1912: 154.

Stenomorpha cressoni (Blaisdell, 1933)USA (CA)

Euschides cressoni Blaisdell, 1933b: 191.

Stenomorpha cribrata (Casey, 1912)USA (NM)

Euschides cribratus Casey, 1912: 140.

Stenomorpha crinita (Casey, 1912)USA (OR)

Euschides crinitus Casey, 1912: 148.

Stenomorpha deceptor (Casey, 1912)USA (CA)

Euschides deceptor Casey, 1912: 154.

Stenomorpha directa (Casey, 1912)USA (AZ)

Euschides directa Casey, 1912: 141.

Stenomorpha evanescens (Casey, 1912)USA (CA)

Euschides evanescens Casey, 1912: 146.

Stenomorpha facilis (Casey, 1912)USA (KS)

Euschides facilis Casey, 1912: 165.

Stenomorpha fastigiosa (Casey, 1912)USA (NM)

Euschides fastigiosus Casey, 1912: 157.

Stenomorpha globicollis (Casey, 1912)USA (NE)

Euschides globicollis Casey, 1912: 158.

Stenomorpha gracilior (Casey, 1912)USA (KS)

Euschides gracilior Casey, 1912: 160.

Stenomorpha gravidipes (Casey, 1912)USA (CA)

Euschides gravidipes Casey, 1912: 155.

Stenomorpha huachucae (Casey, 1912)USA (AZ)

Euschides huachucae Casey, 1912: 162.

Stenomorpha implicans (Casey, 1912)USA (AZ)

Euschides implicans Casey, 1912: 136.

Stenomorpha inhabilis inhabilis (Casey, 1912)USA (NM)

Euschides inhabilis Casey, 1912: 156.

Stenomorpha inhabilis retusa (Casey, 1912)USA (KS)

Euschides inhabilis retusus Casey, 1912: 157.

Stenomorpha integer (Casey, 1912)USA (CA)

Euschides integer Casey, 1912: 153.

Stenomorpha lecontei lecontei (Horn, 1870)USA (CA)

Pelecyphorus costipennis LeConte, 1859b: 76 [junior primary homonym of Pelecyphorus costipennis LeConte, 1858].

Asida lecontei Horn, 1870: 286. Replacement name for Asida costipennis (LeConte, 1859).

Stenomorpha lecontei gigantea (Blaisdell, 1921)USA (CA)

Euschides lecontei gigantea Blaisdell, 1921b: 209.

Stenomorpha lecontella lecontella (Blaisdell, 1936)USA (CA)

Euschides lecontella Blaisdell, 1936b: 227.

Stenomorpha lecontella tempestalis (Blaisdell, 1936)USA (CA)

Euschides lecontella tempestalis Blaisdell, 1936b: 229.

Stenomorpha luctata (Horn, 1870)USA (CA NV)

Asida luctata Horn, 1870: 286.

Stenomorpha marginata duplicans (Casey, 1912)USA [AZ]

Euschides marginatus duplicans Casey, 1912: 136.

Stenomorpha marginata esuriens (Casey, 1912)USA (CA)

Euschides marginatus esuriens Casey, 1912: 137.

Stenomorpha marginata marginata (LeConte, 1851)USA (AZ CA NM TX) MEX (CH SO)

Pelecyphorus marginatus LeConte, 1851: 128.

Stenomorpha maritima imula (Casey, 1912)USA (CA)

Euschides maritimus imulus Casey, 1912: 151.

Stenomorpha maritima maritima (Casey, 1912)USA (CA)

Euschides maritimus Casey, 1912: 151.

Stenomorpha mckittricki (Pierce, 1954)USA (CA)14

Parasida mckittricki Pierce, 1954a: 43.

Stenomorpha montezuma Wilke, 1922MEX (DU)

Stenomorpha montezuma Wilke, 1922: 272.

Stenomorpha musiva Wilke, 1922MEX (ME)

Stenomorpha musiva Wilke, 1922: 273.

Stenomorpha neutralis (Casey, 1912)USA (CA)

Euschides neutralis Casey, 1912: 146.

Stenomorpha oblonga (Casey, 1924)USA (NM)

Euschides oblongus Casey, 1924: 309.

Stenomorpha obovata gliscans (Casey, 1912)USA (AZ)

Euschides obovatus gliscans Casey, 1912: 161.

Stenomorpha obovata nitidipennis (Casey, 1912)USA (AZ)

Euschides obovatus nitidipennis Casey, 1912: 160.

Stenomorpha obovata obovata (LeConte, 1851)USA (AZ CA TX) MEX (CH)

Euschides obovata LeConte, 1851: 127.

Stenomorpha oregonensis (Casey, 1924)USA (OR)

Euschides oregonensis Casey, 1924: 309.

Stenomorpha orizabae Wilke, 1922MEX (VE)

Stenomorpha orizabae Wilke, 1922: 273.

Stenomorpha papagoana (Casey, 1912)USA (AZ)

Euschides papagoanus Casey, 1912: 163.

Stenomorpha pollens pollens (Casey, 1912)USA (AZ)

Euschides pollens Casey, 1912: 134.

Stenomorpha pollens proxima (Casey, 1912)USA (AZ)

Euschides pollens proximus Casey, 1912: 134.

Stenomorpha procurrens (Casey, 1912)USA (AZ)

Euschides procurrens Casey, 1912: 137.

Stenomorpha puncticollis (LeConte, 1866)USA (OR WA)

Euschides puncticollis LeConte, 1866b: 111 [junior secondary homonym of Asida puncticollis Solier, 1836].

Asida robusta Gebien, 1910a: 135. Replacement name for Asida puncticollis (LeConte, 1866)15.

Stenomorpha rimata rimata (LeConte, 1854)USA (TX)

Pelecyphorus rimatus LeConte, 1854c: 223.

Stenomorpha rimata subplanata (Casey, 1912)USA (AZ)

Euschides rimatus subplanatus Casey, 1912: 139.

Stenomorpha rudis (Casey, 1912)USA (AZ)

Euschides rudis Casey, 1912: 139.

Stenomorpha rugata (Casey, 1912)USA (AZ)

Euschides rugatus Casey, 1912: 138.

Stenomorpha rustica (Casey, 1912)USA (AZ)

Euschides rusticus Casey, 1912: 135.

Stenomorpha satiata (Casey, 1912)USA (AZ)

Euschides satiatus Casey, 1912: 138.

Stenomorpha semirufa (Casey, 1912)USA (AZ)

Euschides semirufus Casey, 1912: 140.

Stenomorpha severa (Casey, 1912)USA (NM)

Euschides severus Casey, 1912: 144.

Stenomorpha socialis (Casey, 1912)USA (NM)

Euschides socialis Casey, 1912: 162.

Stenomorpha speculata (Blaisdell, 1936)USA (CA)

Euschides speculatus Blaisdell, 1936b: 225.

Stenomorpha sphaericollis (Champion, 1884)MEX (AG SL)

Asida sphaericollis Champion, 1884: 64.

Stenomorpha sponsor (Casey, 1912)USA (AZ)

Euschides sponsor Casey, 1912: 135.

Stenomorpha spurcans (Casey, 1912)USA (CA)

Euschides spurcans Casey, 1912: 145.

Stenomorpha strigosula (Casey, 1912)USA (AZ)

Euschides strigosulus Casey, 1912: 163.

Stenomorpha subcruenta (Casey, 1912)USA (NM)

Euschides subcruentus Casey, 1912: 140.

Stenomorpha subcylindrica (Horn, 1870)USA (AZ)

Asida marginata var. subcylindrica Horn, 1870: 288.

Stenomorpha subelegans (Casey, 1912)USA (CA)

Euschides subelegans Casey, 1912: 152.

Stenomorpha tetrica (Casey, 1912)USA (UT)

Euschides tetricus Casey, 1912: 149.

Stenomorpha tularensis (Casey, 1912)USA (CA)

Euschides tularensis Casey, 1912: 152.

Stenomorpha uhdei Wilke, 1922MEX (ME)

Stenomorpha uhdei Wilke, 1922: 273.

Stenomorpha umbrosa (Champion, 1884)MEX (GU)

Asida umbrosa Champion, 1884: 62.

Stenomorpha vigens (Casey, 1912)USA (AZ)

Euschides vigens Casey, 1912: 159.

Subgenus Stethasida Casey, 1912

Stethasida Casey, 1912: 78, 203. Type species: Pelecyphorus muricatulus LeConte, 1851, original designation.

Stenomorpha flohri (Champion, 1892)MEX (JA)

Asida flohri Champion, 1892: 496.

Stenomorpha muricatula (LeConte, 1851)USA (CA)

Pelecyphorus muricatulus LeConte, 1851: 128.

Asida angustula Casey, 1890b: 370. New synonymy [ADS].

Stethasida stricta Casey, 1912: 210. New synonymy [ADS].

Stethasida muricatula languida Casey, 1912: 211. New synonymy [ADS].

Stethasida pertinax Casey, 1912: 211. New synonymy [ADS].

Stethasida socors Casey, 1912: 212. New synonymy [ADS].

Stethasida angustula inepta Casey, 1912: 213. New synonymy [ADS].

Stethasida tenax Casey, 1912: 213. New synonymy [ADS].

Stethasida vegrandis Casey, 1912: 214. New synonymy [ADS].

Stenomorpha obsoleta (LeConte, 1851)USA (CA)

Pelecyphorus obsoletus LeConte, 1851: 128.

Stethasida obsoleta expansa Casey, 1912: 205. New synonymy [ADS].

Stethasida obsoleta opacella Casey, 1912: 205. New synonymy [ADS].

Stethasida brevipes Casey, 1912: 206. New synonymy [ADS].

Stethasida torpida Casey, 1912: 206. New synonymy [ADS].

Stethasida convergens Casey, 1912: 207. New synonymy [ADS].

Stethasida discreta Casey, 1912: 207. New synonymy [ADS].

Stethasida longula Casey, 1912: 207. New synonymy [ADS].

Stethasida adumbrata Casey, 1912: 208. New synonymy [ADS].

Stethasida occulta Casey, 1912: 208. New synonymy [ADS].

Stethasida tarsalis Casey, 1912: 208. New synonymy [ADS].

Stethasida unica Casey, 1912: 209. New synonymy [ADS].

Pelecyphorus laevigatus Papp, 1961c: 159. New synonymy [ADS].

Subgenus Trichiasida Casey, 1912

Trichiasida Casey, 1912: 77, 172. Type species: Pelecyphorus hirsutus LeConte, 1851, original designation.

Stenomorpha acerba (Horn, 1878)USA (AZ NV UT)

Asida acerba Horn, 1878a: 56.

Stenomorpha difficilis (Champion, 1884)MEX (HI ME SL)

Asida difficilis Champion, 1884: 61.

Trichiasida eremica Wilke, 1922: 274. New synonymy [ADS].

Stenomorpha hirsuta (LeConte, 1851)USA (CA)

Pelecyphorus hirsutus LeConte, 1851: 127.

Trichiasida lineatopilosa Casey, 1912: 175. New synonymy [ADS].

Stenomorpha hispidula (LeConte, 1851)USA (AZ CA)

Pelecyphorus hispidulus LeConte, 1851: 127.

Trichiasida tenella Casey, 1912: 177. New synonymy [ADS].

Stenomorpha horrida (Champion, 1892)USA (TX) MEX (TA)

Asida horrida Champion, 1892: 500.

Stenomorpha idahoensis (Boddy, 1957)USA (ID)

Trichiasida idahoensis Boddy, 1957: 187.

Stenomorpha ignava (Casey, 1912)USA (AZ)

Trichiasida ignava Casey, 1912: 180.

Stenomorpha impetrata (Horn, 1894)USA (CA)

Asida impetrata Horn, 1894b: 418.

Stenomorpha impotens (Casey, 1912)USA (AZ)

Trichiasida impotens Casey, 1912: 180.

Stenomorpha lutulenta (Doyen, 1990)MEX (JA)

Trichiasida lutulenta Doyen, 1990: 225.

Stenomorpha palmeri (Champion, 1884)MEX (SL)

Asida palmeri Champion, 1884: 59.

Stenomorpha pubescens (Champion, 1884) MEX

Asida pubescens Champion, 1884: 61.

Stenomorpha subpilosa Solier, 1836MEX (PU)

Stenomorpha subpilosa Solier, 1836: 490.

Stenomorpha thoracica (Champion, 1884) MEX

Asida thoracica Champion, 1884: 62.

Stenomorpha unicostata (Champion, 1892)MEX (GE)

Asida unicostata Champion, 1892: 501.

Stenomorpha villosa (Champion, 1884)MEX (PU)

Asida villosa Champion, 1884: 60.

Trichiasida duplex Casey, 1912: 17816. New synonymy [ADS].

[incertae sedis]

Stenomorpha catalinae (Blaisdell, 1923)MEX (BS)

Asida catalinae Blaisdell, 1923: 256.

Stenomorpha furcata (Champion, 1892)USA (TX) MEX (CO DU)

Asida furcata Champion, 1892: 499.

Stenomorpha granicollis (Blaisdell, 1923)MEX (SO)

Asida granicollis Blaisdell, 1923: 256.

Stenomorpha roosevelti Smith, Miller and Wheeler, 2011MEX (CO)

Stenomorpha roosevelti Smith, Miller and Wheeler, 2011: 30.

Stenomorpha spinimana (Champion, 1892)MEX (DU) New combination [ADS].

Asida spinimanus Champion, 1892: 494.

Stenomorpha subvittata (Horn, 1894)MEX (BS)

Asida subvittata Horn, 1894b: 416.

Stenomorpha tenebrosa (Champion, 1892)MEX (CO) New combination [ADS].

Asida tenebrosa Champion, 1892: 495.

Stenomorpha wickhami (Horn, 1894)USA (AZ CA)

Asida wickhami Horn, 1894b: 420.

Asidina liberta Casey, 1912: 171. Synonymy: Triplehorn and Brown (1971: 84).

Tribe Branchini LeConte, 1862

Branchini LeConte, 1862a: 222. Type genus: Branchus LeConte, 1862

Genus Anectus Horn, 1866 [M]

Anectus Horn, 1866: 399. Type species: Anectus vestitus Horn, 1866, monotypy.

Anectus vestitus Horn, 1866 HON

Anectus vestitus Horn, 1866: 399.

Genus Branchus LeConte, 186217 [M]

Branchus LeConte, 1862a: 222. Type species: Branchus floridanus LeConte, 1862, monotypy.

Branchus floridanus LeConte, 1862USA (FL)

Branchus floridanus LeConte, 1862a: 223.

Branchus geraceorum Steiner, 2006 BAH

Branchus geraceorum Steiner, 2006: 10.

Branchus jamaicensis Marcuzzi, 1977 JAM

Branchus jamaicensis Marcuzzi, 1977: 10.

Branchus obscurus Horn, 1866MEX (GE) GUANIC

Branchus obscurus Horn, 1866: 398.

Branchus opatroides Champion, 1892MEX (JA VE)

Branchus opatroides Champion, 1892: 507.

Branchus saxatilis Steiner, 2005 BAH

Branchus saxatilis Steiner, 2005: 443.

Branchus whiteheadi Steiner, 1991USA (TX)

Branchus whiteheadi Steiner, 1991: 426.

Branchus woodii LeConte, 1866 BAH CUB

Branchus woodii LeConte, 1866b: 111.

Genus Oxinthas Champion, 1884 [M]

Oxinthas Champion, 1884: 72. Type species: Oxinthas praocioides Champion, 1884, monotypy.

Oxinthas nicaraguensis Merkl, 1992 NIC

Oxinthas nicaraguensis Merkl, 1992: 89.

Oxinthas praocioides Champion, 1884MEX (OA)

Oxinthas praocioides Champion, 1884: 72.

Tribe Cnemeplatiini Jacquelin du Val, 1861

Cnéméplatiites Jacquelin du Val, 1861: 286. Type genus: Cnemeplatia Costa, 1847.

Subtribe Cnemeplatiina Jacquelin du Val, 1861

Cnéméplatiites Jacquelin du Val, 1861: 286. Type genus: Cnemeplatia Costa, 1847.

Genus Alaudes Horn, 1870 [M]

Alaudes Horn, 1870: 361. Type species: Alaudes singularis Horn, 1870, monotypy.

Alaudes alternatus Fall, 1928USA (CA)

Alaudes alternata Fall, 1928: 148.

Alaudes setigerus Blaisdell, 1919USA (CA)

Alaudes setigera Blaisdell, 1919a: 310.

Alaudes singularis Horn, 1870USA (CA ID NV OR)

Alaudes singularis Horn, 1870: 362.

Alaudes squamosa Blaisdell, 1919a: 309. New synonymy [RLA].

Alaudes testacea Blaisdell, 1919a: 311. New synonymy [RLA].

Alaudes fallax Fall, 1928: 150. New synonymy [RLA].

Genus Lepidocnemeplatia Bousquet and Bouchard, new genus [F]

Lepidocnemeplatia Bousquet and Bouchard, new genus. Type species: Cnemeplatia sericea Horn, 1870.

Lepidocnemeplatia laticollis (Champion, 1885)MEX (ME) NICCRIPAN / SA

Cnemeplatia laticollis Champion, 1885: 136.

Lepidocnemeplatia sericea (Horn, 1870)USA (AZ CA NV OR TX WA) MEX (BS CH DU MO NL PU SO VE) NIC

Cnemeplatia sericea Horn, 1870: 360.

Tribe Cnemodinini Gebien, 1910

Cnemodini Horn, 1870: 266. Type genus: Cnemodus Horn, 1870 (= Cnemodinus Cockerell, 1906).

Cnemodininae Gebien, 1910a: 4. Type genus: Cnemodinus Cockerell, 1906.

Genus Cnemodinus Cockerell, 1906 [M]

Cnemodus Horn, 1870: 266 [junior homonym of Cnemodus Herrich-Schaeffer, 1850]. Type species: Cnemodus testaceus Horn, 1870, monotypy.

Cnemodinus Cockerell, 1906: 242. Replacement name for Cnemodus Horn, 1870.

Cnemodinus angustus (Casey, 1907)USA (AZ)

Cnemodus angustus Casey, 1907: 284.

Cnemodinus subhyalinus (Casey, 1907)USA (UT)

Cnemodus subhyalinus Casey, 1907: 285.

Cnemodinus testaceus (Horn, 1870)USA (AZ CA)

Cnemodus testaceus Horn, 1870: 266.

Tribe Coniontini G.R. Waterhouse, 1858

Coniontidae G.R. Waterhouse, 1858: 59. Type genus: Coniontis Eschscholtz, 1829.

Coelini Casey, 1907: 500. Type genus: Coelus Eschscholtz, 1829.

Eusatti Doyen, 1984b: 11. Type genus: Eusattus LeConte, 1851.

Genus Coelus Eschscholtz, 1829 [M]

Coelus Eschscholtz, 1829: 5. Type species: Coelus ciliatus Eschscholtz, 1829, monotypy.

Coelomorpha Casey, 1890a: 182. Type species: Coelomorpha maritima Casey, 1890, monotypy. Synonymy: Doyen (1972: 371).

Pseudocoelus Casey, 1908: 152. Type species: Coelus pacificus Fall, 1897, subsequent designation (Doyen 1976: 608). Synonymy: Blaisdell (1919b: 322).

Coelus ciliatus Eschscholtz, 1829 [Fig. 12] CAN (BC) USA (CA OR WA) MEX (BC)

Figure 12. 

Coelus ciliatus Eschscholtz, 1829. Scale bar = 1 mm.

Coelus ciliatus Eschscholtz, 1829: 5.

Coelus arenarius Casey, 1890a: 179. Synonymy: Doyen (1976: 614).

Coelus latus Casey, 1895: 612. Synonymy (with C. arenarius Casey): Fall (1901: 166).

Coelus curtulus Casey, 1895: 612. Synonymy: Fall (1901: 166).

Coelus ciliatus longulus Casey, 1908: 154. Synonymy: Blaisdell (1919b: 333).

Coelus debilis Casey, 1908: 155. Synonymy: Doyen (1976: 614).

Coelus sternalis Casey, 1908: 156. Synonymy: Doyen (1976: 614).

Coelus obscurus Casey, 1908: 156. Synonymy (with C. arenarius Casey): Blaisdell (1919b: 334).

Coelus scolopax Casey, 1908: 157. Synonymy (with C. arenarius Casey): Blaisdell (1919b: 334).

Coelus amplicollis Casey, 1908: 157. Synonymy (with C. latus Casey): Blaisdell (1919b: 334).

Coelus ciliatus var. sparsus Blaisdell, 1919b: 325. Synonymy (with C. ciliatus debilis Casey): Gebien (1938: 409).

Coelus globosus LeConte, 1851USA (CA) MEX (BC)

Coelus globosus LeConte, 1851: 133.

Coelus grossus Casey, 1890a: 178. Synonymy: Doyen (1976: 617).

Coelus solidus Casey, 1908: 153. Synonymy (with C. globosus grossus Casey): Blaisdell (1919b: 333).

Coelus saginatus Casey, 1908: 154. Synonymy: Doyen (1976: 617).

Coelus gracilis Blaisdell, 1939USA (CA)

Coelus gracilis Blaisdell, 1939a: 16.

Coelus maritimus (Casey, 1889)MEX (BC)

Coelomorpha maritima Casey, 1890a: 183.

Coelomorpha pallens Casey, 1908: 160. Synonymy: Doyen (1976: 620).

Coelus pacificus Fall, 1897USA (CA)

Coelus pacificus Fall, 1897: 241.

Coelus remotus Fall, 1897: 241. Synonymy: Doyen (1976: 618).

Genus Coniontis Eschscholtz, 1829 [F]

Coniontis Eschscholtz, 1829: 7. Type species: Coniontis viatica Eschscholtz, 1829, subsequent designation (Casey 1908: 57).

Coelotaxis Horn, 1876a: 200. Type species: Coelotaxis punctulata Horn, 1876, subsequent designation (Gebien 1938: 289). Synonymy: Doyen (1972: 373).

Coniontellus Casey, 1890b: 388. Type species: Coniontis obesa LeConte, 1851, subsequent designation (Casey 1908: 57). Synonymy: Doyen (1972: 373).

Coniontides Casey, 1908: 57, 78. Type species: Coniontis lata LeConte, 1866, original designation. Synonymy: Doyen (1972: 373).

Crypticomorpha Casey, 1908: 81, 140. Type species: Coniontis tenuis Casey, 1908, monotypy. Synonymy: Aalbu et al. (2002: 487).

Brachyontis Casey, 1908: 82, 141. Type species: Coniontis globulina Casey, 1895, monotypy. Synonymy: Aalbu et al. (2002: 487).

Coniontis abdominalis LeConte, 1859USA (CA)

Coniontis abdominalis LeConte, 1859b: 77.

Coniontis strenua Casey, 1908: 84. Synonymy: Doyen (1977: 2).

Coniontis tristis Casey, 1908: 84. Synonymy: Doyen (1977: 2).

Coniontis gravis Casey, 1908: 85. Synonymy: Doyen (1977: 2).

Coniontis rugosa Casey, 1908: 85. Synonymy: Doyen (1977: 2).

Coniontis tenebrosa Casey, 1908: 86. Synonymy: Doyen (1977: 2).

Coniontis abdominalis caseyi Pierce, 1954c: 145. Synonymy: Doyen and Miller (1980: 3).

Coniontis abdominalis labreae Pierce, 1954c: 146. Synonymy: Doyen and Miller (1980: 3).

Coniontis abdominalis fragmans Pierce, 1954c: 148. Synonymy: Doyen and Miller (1980: 3).

Coniontis tristis alpha Pierce, 1954c: 148. Synonymy: Doyen and Miller (1980: 3).

Coniontis tristis asphalti Pierce, 1954c: 149. Synonymy: Doyen and Miller (1980: 3).

Coniontis tristis latigula Pierce, 1954c: 149. Synonymy: Doyen and Miller (1980: 3).

Coniontis blissi Pierce, 1954c: 149. Synonymy: Doyen and Miller (1980: 3).

Coniontis pectoralis paraelliptica Pierce, 1954c: 153. Synonymy: Doyen and Miller (1980: 3).

Coniontis pectoralis interrupta Pierce, 1954c: 154. Synonymy: Doyen and Miller (1980: 3).

Coniontis affinis LeConte, 1851USA (CA OR)

Coniontis affinis LeConte, 1851: 130.

Coniontis expansa Casey, 1908: 120. Synonymy: Doyen (1977: 2).

Coniontis franciscana Casey, 1908: 120. Synonymy: Doyen (1977: 2).

Coniontis truncata Casey, 1908: 120. Synonymy: Doyen (1977: 2).

Coniontis suturalis Casey, 1908: 121. Synonymy: Doyen (1977: 2).

Coniontis audax Casey, 1908: 121. Synonymy: Doyen (1977: 2).

Coniontis symmetrica Casey, 1908: 122. Synonymy: Doyen (1977: 2).

Coniontis convergens Casey, 1908: 122. Synonymy: Doyen (1977: 2).

Coniontis anxia Casey, 1908: 123. Synonymy: Doyen (1977: 2).

Coniontis affinis patruelis Casey, 1908: 123. Synonymy: Doyen (1977: 2).

Coniontis oregona Casey, 1908: 124. Synonymy: Doyen (1977: 2).

Coniontis pagana Casey, 1908: 130. Synonymy: Doyen (1977: 2).

Coniontis atronitens Casey, 1908USA (CA)

Coniontis atronitens Casey, 1908: 110.

Coniontis blaisdelli Casey, 1908USA (CA)

Coniontis blaisdelli Casey, 1908: 97.

Coniontis callida Casey, 1908USA (CA)

Coniontis callida Casey, 1908: 128.

Coniontis shastanica Casey, 1908: 128. Synonymy: Doyen (1977: 2).

Coniontis conferta Casey, 1908: 129. Synonymy: Doyen (1977: 2).

Coniontis agrestis Casey, 1908: 131. Synonymy: Doyen (1977: 2).

Coniontis congesta Casey, 1908: 131. Synonymy: Doyen (1977: 2).

Coniontis costulata Casey, 1908USA (CA)

Coniontis costulata Casey, 1908: 89.

Coniontis elliptica Casey, 1884USA (CA)

Coniontis elliptica Casey, 1884: 46.

Coniontis laevigata Casey, 1908: 88. Synonymy: Doyen (1977: 2).

Coniontis elliptica catalinae Casey, 1908: 88. Synonymy: Doyen (1977: 2).

Coniontis cuneata Casey, 1908: 111. Synonymy: Gebien (1938: 286).

Coniontis elongata Casey, 1890USA (CA)

Coniontis elongata Casey, 1890b: 380.

Coniontis rotundicollis Casey, 1908: 97. Synonymy: Doyen (1977: 3).

Coniontis innocua Casey, 1908: 99. Synonymy: Doyen (1977: 3).

Coniontis elongata limatula Casey, 1908: 99. Synonymy: Doyen (1977: 3).

Coniontis cylindrica Casey, 1908: 100. Synonymy: Doyen (1977: 3).

Coniontis obsidiana Casey, 1908: 100. Synonymy: Blaisdell (1935b: 120).

Coniontis longicollis Casey, 1908: 101. Synonymy: Doyen (1977: 3).

Coniontis eschscholtzii Mannerheim, 1840USA (CA)

Coniontis eschscholtzii Mannerheim, 1840: 138.

Coniontis extricata Casey, 1908USA (CA OR)

Coniontis extricata Casey, 1908: 124.

Coniontis marginata Casey, 1908: 125. Synonymy: Doyen (1977: 3).

Coniontis minuta Casey, 1908: 126. Synonymy: Doyen (1977: 3).

Coniontis parva Casey, 1908: 126. Synonymy: Doyen (1977: 3).

Coniontis perpolita Casey, 1908: 127. Synonymy: Doyen (1977: 3).

Coniontis pudica Casey, 1908: 127. Synonymy: Doyen (1977: 3).

Coniontis nemoralis slevini Blaisdell, 1924a: 86. Synonymy: Doyen (1977: 3).

Coniontis farallonica Casey, 1895USA (CA)

Coniontis farallonica Casey, 1895: 610.

Coniontis genitiva Casey, 1890USA (CA)

Coniontis genitiva Casey, 1890b: 385.

Coniontis verna Casey, 1908: 94. Synonymy: Doyen (1977: 3).

Coniontis opacicollis Casey, 1908: 101. Synonymy: Doyen (1977: 3).

Coniontis globulina Casey, 1895USA (CA)

Coniontis globulina Casey, 1895: 610.

Coniontis histrio Casey, 1908USA (AZ)

Coniontis histrio Casey, 1908: 91.

Coniontis hoppingi Blaisdell, 1918USA (CA)

Coniontis hoppingi Blaisdell, 1918a: 7.

Coniontis inaequalis Casey, 1890USA (CA)

Coniontis inaequalis Casey, 1890b: 375.

Coniontis inornata Casey, 1908USA (CA)

Coniontis inornata Casey, 1908: 130.

Coniontis integer Casey, 1908USA (CA)

Coniontis integer Casey, 1908: 87.

Coniontis keiferi (Blaisdell, 1943)MEX (BC)

Coniontides keiferi Blaisdell, 1943: 189.

Coniontis lamentabilis Blaisdell, 1924USA (CA)

Coniontis lamentabilis Blaisdell, 1924a: 85.

Coniontis lanei Boddy, 1957USA (ORWA)

Coniontis lanei Boddy, 1957: 191.

Coniontis lariversi Blaisdell, 1941USA (NV)

Coniontis lariversi Blaisdell, 1941c: 131.

Coniontis lassenica Casey, 1908USA (CA NV)

Coniontis lassenica Casey, 1908: 95.

Coniontis nevadensis Casey, 1908: 95. Synonymy: Doyen (1977: 3).

Coniontis nevadensis carsonica Casey, 1908: 95. Synonymy: Doyen (1977: 3).

Coniontis lata LeConte, 1866USA (CA)

Coniontis lata LeConte, 1866b: 113.

Coniontis lata var. insularis Casey, 1890b: 377. Synonymy: Doyen (1977: 3).

Coniontides finitimus Casey, 1908: 79. Synonymy: Doyen (1977: 3).

Coniontides clementinus Casey, 1908: 80. Synonymy: Blaisdell (1921b: 212).

Coniontis malkini (Boddy, 1957)USA (CA)

Coniontellus malkini Boddy, 1957: 188.

Coniontis microsticta Casey, 1908USA (CA)

Coniontis microsticta Casey, 1908: 107.

Coniontis inconspicua Casey, 1908: 108. Synonymy: Blaisdell (1924a: 84).

Coniontis muscula Blaisdell, 1918USA (CA)

Coniontis globulina var. muscula Blaisdell, 1918a: 9.

Coniontis nemoralis borealis Boddy, 1957USA (CA OR)

Coniontis nemoralis borealis Boddy, 1957: 192.

Coniontis nemoralis nemoralis Eschscholtz, 1829USA (CA)

Coniontis nemoralis Eschscholtz, 1829: 8.

Coniontis obesa LeConte, 1851USA (CA CO ID MT NV OR WY)

Coniontis obesus LeConte, 1851: 131.

Coniontellus inflatus Casey, 1890b: 389. Synonymy: Doyen (1977: 3).

Coniontellus subglaber Casey, 1890b: 389. Synonymy: Doyen (1977: 3).

Coniontellus hystrix Casey, 1908: 142. Synonymy (with C. inflatus Casey): La Rivers (1947b: 214).

Coniontellus longipennis Casey, 1908: 143. Synonymy (with C. inflatus Casey): La Rivers (1947b: 214).

Coniontellus ampliatus Casey, 1908: 144. Synonymy (with C. inflatus Casey): La Rivers (1947b: 214).

Coniontellus argutus Casey, 1908: 145. Synonymy: Doyen (1977: 3).

Coniontellus micans Casey, 1908: 145. Synonymy: Doyen (1977: 3).

Coniontis oblonga Casey, 1908USA (CA)

Coniontis oblonga Casey, 1908: 92.

Coniontis opaca Horn, 1870USA (CA)

Coniontis opaca Horn, 1870: 296.

Coniontis ancilla Casey, 1908: 91. Synonymy: Doyen (1977: 3).

Coniontis degener Casey, 1908: 93. Synonymy: Doyen (1977: 3).

Coniontis ovalis LeConte, 1851 [Fig. 13] CAN (BC) USA (AK CA CO ID MT NV OR UT WA)

Figure 13. 

Coniontis ovalis LeConte, 1851. Scale bar = 1 mm.

Coniontis ovalis LeConte, 1851: 131.

Coniontis alutacea Casey, 1890b: 383. Synonymy: Doyen (1977: 4).

Coniontis breviuscula Casey, 1908: 133. Synonymy: Boddy (1957: 190).

Coniontis sculptipennis Casey, 1908: 134. Synonymy: Boddy (1957: 190).

Coniontis regularis Casey, 1908: 134. Synonymy: Doyen (1977: 4).

Coniontis punctata Casey, 1908: 135. Synonymy (with C. regularis Casey): Boddy (1965: 141).

Coniontis parilis Casey, 1908: 135. Synonymy: Boddy (1957: 190).

Coniontis vancouveri Casey, 1908: 136. Synonymy: Boddy (1957: 190).

Coniontis uteana Casey, 1908: 136. Synonymy: Doyen (1977: 4).

Coniontis inepta Casey, 1908: 137. Synonymy: Doyen (1977: 4).

Coniontis oblita Casey, 1908: 137. Synonymy: Doyen (1977: 4).

Coniontis arida Casey, 1908: 138. Synonymy: Doyen (1977: 4).

Coniontis weidti Casey, 1908: 138. Synonymy: Doyen (1977: 4).

Coniontis acerba Casey, 1908: 139. Synonymy: Doyen (1977: 4).

Coniontis anita Casey, 1908: 139. Synonymy: Doyen (1977: 4).

Coniontis corvina Casey, 1908: 140. Synonymy: Doyen (1977: 4).

Coniontis ovalis okanagani Boddy, 1957: 190. Synonymy: Doyen (1977: 4).

Coniontis pallidicornis Casey, 1890USA (CA)

Coniontis pallidicornis Casey, 1890b: 385.

Coniontis obsolescens Casey, 1908: 92. Synonymy: Doyen (1977: 4).

Coniontis parallela Casey, 1890USA (CA)

Coniontis parallela Casey, 1890b: 386.

Coniontis parviceps Casey, 1890USA (CA) MEX (BC)

Coniontis parviceps Casey, 1890b: 387.

Coniontis filiola Casey, 1908: 115. Synonymy: Doyen (1977: 4).

Coniontis pectoralis Casey, 1908USA (CA)

Coniontis pectoralis Casey, 1908: 86.

Coniontis levettei Casey, 1908: 87. Synonymy: Doyen (1977: 4).

Coniontis picescens Casey, 1908: 87. Synonymy: Doyen (1977: 4).

Coniontis perspicua Casey, 1908USA (CA)

Coniontis perspicua Casey, 1908: 114.

Coniontis proba Casey, 1908USA (ID OR)

Coniontis proba Casey, 1908: 105.

Coniontis puncticollis LeConte, 1851USA (CA)

Coniontis puncticollis LeConte, 1851: 131.

Coniontis exigua Casey, 1908: 106. Synonymy: Doyen (1977: 4).

Coniontis paupercula Casey, 1908: 106. Synonymy: Doyen (1977: 4).

Coniontis inflexula Casey, 1908: 107. Synonymy (with C. exigua Casey): Casey (1911: 253).

Coniontis picipes Casey, 1908: 107. Synonymy: Doyen (1977: 4).

Coniontis punctipes Casey, 1890USA (CA)

Coniontis punctipes Casey, 1890b: 380.

Coniontis punctulata (Horn, 1876)USA (CA) MEX

Coelotaxis punctulata Horn, 1876a: 201.

Coelotaxis muricata Horn, 1876a: 201. Synonymy: Doyen (1977: 4).

Coelotaxis angustula Casey, 1890a: 177. Synonymy: Doyen (1977: 4).

Coelotaxis densa Casey, 1908: 149. Synonymy: Doyen (1977: 4).

Coelotaxis frontalis Casey, 1908: 149. Synonymy: Doyen (1977: 4).

Coniontis rainieri Boddy, 1957USA (WA)

Coniontis rainieri Boddy, 1957: 191.

Coniontis remnans Pierce, 1954USA (CA)18

Coniontis remnans Pierce, 1954c: 155.

Coniontis robusta Horn, 1870USA (CA)

Coniontis robusta Horn, 1870: 296.

Coniontis luctuosa Casey, 1908: 89. Synonymy: Doyen (1977: 5).

Coniontis sanfordii Blaisdell, 1895USA (CA)

Coniontis sanfordii Blaisdell, 1895: 235.

Coniontis santarosae Blaisdell, 1921USA (CA)

Coniontis santarosae Blaisdell, 1921b: 209.

Coniontis setosa Casey, 1890USA (ID NV OR UT WA)

Coniontis setosus Casey, 1890b: 387.

Coniontis obtusa Casey, 1908: 116. Synonymy: Doyen (1977: 5).

Coniontis wickhami Casey, 1908: 117. Synonymy: Doyen (1977: 5).

Coniontis lanuginosa Casey, 1908: 117. Synonymy: Doyen (1977: 5).

Coniontis pubifera Casey, 1908: 118. Synonymy: Boddy (1965: 140).

Coniontis sparsa Casey, 1908USA (CA)

Coniontis sparsa Casey, 1908: 110.

Coniontis subpubescens LeConte, 1851USA (CA OR)

Coniontis subpubescens LeConte, 1851: 131.

Coniontis montana Casey, 1890b: 384. Synonymy: Doyen (1977: 5).

Coniontis canonica Casey, 1908: 114. Synonymy: Blaisdell (1918a: 13).

Coniontis tenuis Casey, 1908USA (CA)

Coniontis tenuis Casey, 1908: 141.

Coniontis thoracica Casey, 1908USA (CA)

Coniontis thoracica Casey, 1908: 104.

Coniontis timida Casey, 1908USA (CA)

Coniontis timida Casey, 1908: 102.

Coniontis conicicollis Casey, 1908: 102. Synonymy: Doyen (1977: 5).

Coniontis lucidula Casey, 1908: 103. Synonymy: Doyen (1977: 5).

Coniontis protensa Casey, 1908: 104. Synonymy: Doyen (1977: 5).

Coniontis vandykei Blaisdell, 1921USA (CA)

Coniontides vandykei Blaisdell, 1921b: 212.

Coniontis ventura Blaisdell, 1924USA (CA)

Coniontis globulina ventura Blaisdell, 1924a: 83.

Coniontis viatica Eschscholtz, 1829USA (CA)

Coniontis viatica Eschscholtz, 1829: 7.

Coniontis wadei Casey, 1924USA (WA)

Coniontis wadei Casey, 1924: 313.

Genus Conisattus Casey, 1895 [M]

Conisattus Casey, 1895: 614. Type species: Conisattus rectus Casey, 1895, monotypy.

Conisattus rectus Casey, 1895USA (OR WA)

Conisattus rectus Casey, 1895: 614.

Conisattus nelsoni Boddy, 1957: 188. Synonymy: Doyen (1984b: 26).

Genus Eusattus LeConte, 1851 [M]

Eusattus LeConte, 1851: 131. Type species: Eusattus difficilis LeConte, 1851, subsequent designation (Casey 1908: 56).

Discodemus LeConte, 1862a: 223. Type species: Zophosis reticulata Say, 1824, monotypy. Synonymy: LeConte (1866a: 60).

Conipinus LeConte, 1862a: 223. Type species: Eusattus dubius LeConte, 1851, subsequent designation (Gebien 1938: 284). Synonymy: LeConte (1866a: 60).

Nesostes Casey, 1908: 56, 58. Type species: Eusattus robustus LeConte, 1866, original designation. Synonymy: Triplehorn (1968b: 379).

Megasattus Casey, 1908: 56. Type species: Eusattus erosus Horn, 1870, original designation. Synonymy: Triplehorn (1968b: 379).

Eusattodes Casey, 1908: 56. Type species: Eusattus laevis LeConte, 1866, original designation. Synonymy: Triplehorn (1968b: 379).

Sphaeriontis Casey, 1908: 56, 75. Type species: Eusattus muricatus LeConte, 1851, original designation. Synonymy: La Rivers (1949: 180).

Coelosattus Blaisdell, 1927: 166. Type species: Coelosattus fortineri Blaisdell, 1927 (= Eusattus dilatatus LeConte, 1851), monotypy. Synonymy: Doyen (1972: 373).

Eusattus araneosus (Blaisdell, 1923)MEX (BS)

Megasattus araneosus Blaisdell, 1923: 266.

Eusattus arenarius Doyen, 1984MEX (BC BS)

Eusattus arenarius Doyen, 1984b: 75.

Eusattus aridus Doyen, 1984MEX (BC BS)

Eusattus aridus Doyen, 1984b: 43.

Eusattus catalinensis Doyen, 1984MEX (BS)

Eusattus catalinensis Doyen, 1984b: 45.

Eusattus catavinus Doyen, 1984MEX (BC)

Eusattus catavinus Doyen, 1984b: 46.

Eusattus cedrosensis Doyen, 1984MEX (BC)

Eusattus cedrosensis Doyen, 1984b: 47.

Eusattus ceralboensis Doyen, 1984MEX (BS)

Eusattus ceralboensis Doyen, 1984b: 48.

Eusattus cienegus Doyen, 1984MEX (CO)

Eusattus cienegus Doyen, 1984b: 49.

Eusattus ciliatoides Doyen, 1984MEX (BC)

Eusattus ciliatoides Doyen, 1984b: 75.

Eusattus ciliatus Horn, 1894MEX (BC)

Eusattus ciliatus Horn, 1894b: 422.

Eusattus convexus LeConte, 1851USA (AZ CO KS MO NM OK TX UT WY) MEX (CH DU SO)

Eusattus convexus LeConte, 1851: 132.

Eusattus sculptus Champion, 1892: 510. Synonymy: Doyen (1984b: 35).

Eusattus rotundus Casey, 1908: 72. Synonymy: Doyen (1977: 5).

Eusattus turgidus Casey, 1908: 73. Synonymy: Doyen (1977: 5).

Eusattus subnitens Casey, 1908: 73. Synonymy: Doyen (1984b: 35).

Eusattus peropacus Casey, 1908: 74. Synonymy: Doyen (1977: 5).

Eusattus acutus Casey, 1908: 74. Synonymy: Doyen (1977: 5).

Eusattus quadratus Casey, 1924: 311. Synonymy: Doyen (1977: 5).

Eusattus subvelutinus Casey, 1924: 312. Synonymy: Doyen (1977: 5).

Eusattus woodgatei Casey, 1924: 312. Synonymy: Doyen (1977: 5).

Eusattus costatus Horn, 1870USA (CA) MEX (BC BS)

Eusattus costatus Horn, 1870: 293.

Megasattus sternalis Blaisdell, 1923: 268. Synonymy: Doyen (1984b: 50).

Eusattus crypticus Doyen, 1984MEX (BS)

Eusattus crypticus Doyen, 1984b: 90.

Eusattus depressus Champion, 1884MEX (CH NA SI SO)

Eusattus depressus Champion, 1884: 75.

Eusattus puncticeps Blaisdell, 1923: 269. Synonymy: Doyen (1984b: 87).

Eusattus difficilis LeConte, 1851USA (CA NV) MEX (BC SO)

Eusattus difficilis LeConte, 1851: 132.

Eusattus coquilletti Linell, 1899: 180. Synonymy: Doyen (1977: 5).

Eusattus agnatus Casey, 1908: 70. Synonymy: Doyen (1977: 5).

Eusattus compositus Casey, 1908: 71. Synonymy: Doyen (1977: 5).

Eusattus congener Casey, 1908: 71. Synonymy: Doyen (1977: 5).

Eusattus acutangulus Casey, 1908: 72. Synonymy: Doyen (1977: 5).

Eusattus dilatatus LeConte, 1851USA (AZ CA) MEX (SO)

Eusattus dilatatus LeConte, 1851: 132.

Coelosattus fortineri Blaisdell, 1927: 167. Synonymy: Doyen (1977: 5).

Eusattus dubius abditus Doyen, 1984MEX (BC)

Eusattus dubius abditus Doyen, 1984b: 82.

Eusattus dubius arizonensis Doyen, 1984USA (AZ CA NV)

Eusattus dubius arizonensis Doyen, 1984b: 81.

Eusattus dubius dubius LeConte, 1851USA (AZ CA NV UT)

Eusattus dubius LeConte, 1851: 132.

Eusattus nanus Casey, 1895: 613. Synonymy: Doyen (1977: 5).

Eusattus oblongulus Casey, 1908: 67. Synonymy: Doyen (1977: 5).

Conipinus spaldingi Casey, 1924: 313. Synonymy: Doyen (1977: 5).

Eusattus dubius setosus Doyen, 1984MEX (BS)

Eusattus dubius setosus Doyen, 1984b: 82.

Eusattus erosus erosus Horn, 1870MEX (BS)

Eusattus erosus Horn, 1870: 294.

Eusattus erosus laeviventris (Blaisdell, 1923)MEX (BS)

Megasattus laeviventris Blaisdell, 1923: 267.

Eusattus erosus manuelis (Blaisdell, 1923)MEX (BS)

Megasattus erosus manuelis Blaisdell, 1923: 266.

Eusattus franciscanus Doyen, 1984MEX (BS)

Eusattus franciscanus Doyen, 1984b: 54.

Eusattus hirsutus Doyen, 1984USA (NV)

Eusattus hirsutus Doyen, 1984b: 66.

Eusattus laevis LeConte, 1866MEX (BS)

Eusattus laevis LeConte, 1866b: 113.

Eusattus mexicanus Champion, 1892MEX (CO GE JA)

Eusattus mexicanus Champion, 1892: 510.

Eusattus minimus Doyen, 1984MEX (NL)

Eusattus minimus Doyen, 1984b: 38.

Eusattus muricatus diabloensis Doyen, 1984MEX (BC)

Eusattus muricatus diabloensis Doyen, 1984b: 71.

Eusattus muricatus muricatus LeConte, 1851USA (AZ CA CO ID NM NV OR TX UT WA)

Eusattus muricatus LeConte, 1851: 132.

Sphaeriontis acomana Casey, 1908: 76. Synonymy: La Rivers (1949: 180).

Sphaeriontis latissima Casey, 1924: 310. Synonymy: La Rivers (1949: 180).

Sphaeriontis fulvescens Casey, 1924: 310. Synonymy: La Rivers (1949: 180).

Eusattus nitidipennis LeConte, 1851MEX (CO GU NL PU VE ZA)

Eusattus nitidipennis LeConte, 1851: 133.

Eusattus brevis Champion, 1884: 75. Synonymy: Doyen (1977: 6).

Eusattus obliteratus Champion, 1892MEX (CO DU)

Eusattus obliteratus Champion, 1892: 51019.

Eusattus pallidus adustus Doyen, 1984MEX (BS)

Eusattus pallidus adustus Doyen, 1984b: 84.

Eusattus pallidus immaculatus Doyen, 1984MEX (BS)

Eusattus pallidus immaculatus Doyen, 1984b: 85.

Eusattus pallidus pallidus Doyen, 1984MEX (BS)

Eusattus pallidus pallidus Doyen, 1984b: 84.

Eusattus phreatophilus Doyen, 1984USA (CA NV)

Eusattus phreatophilus Doyen, 1984b: 71.

Eusattus planulus Doyen, 1984MEX (BS)

Eusattus planulus Doyen, 1984b: 57.

Eusattus politus cruzensis Doyen, 1984USA (CA)

Eusattus politus cruzensis Doyen, 1984b: 93.

Eusattus politus politus Horn, 1883USA (CA)

Eusattus politus Horn, 1883: 304.

Eusattus vanduzeei Blaisdell, 1921b: 214. Synonymy: Doyen (1984b: 93).

Eusattus pons Triplehorn, 1968USA (TX) MEX (CH CO DU)

Eusattus pons Triplehorn, 1968b: 376.

Eusattus productus LeConte, 1858USA (AZ CA) MEX (BC SO)

Eusattus productus LeConte, 1858a: 20.

Eusattus explanatus Casey, 1908: 68. Synonymy: Doyen (1977: 6).

Eusattus vicinus Casey, 1908: 68. Synonymy: Doyen (1977: 6).

Eusattus lobatus Casey, 1908: 68. Synonymy: Doyen (1977: 6).

Eusattus puberulus LeConte, 1854USA (TX)

Eusattus puberulus LeConte, 1854a: 84.

Eusattus reticulatus (Say, 1824)USA (AZ CO NM OK TX UT) MEX (CH SO)

Tenebrio gibbus DeGeer, 1778: 652 [junior primary homonym of Tenebrio gibbus Linnaeus, 1767].

Tenebrio striatus Retzius, 1783: 134 [junior primary homonym of Tenebrio striatus Müller, 1776]. Replacement name for Tenebrio gibbus DeGeer, 1778.

Zophosis reticulata Say, 1824a: 250. Synonymy: Ferrer and Holston (2011: 252).

Discodemus corrosus Casey, 1908: 61. Synonymy: Doyen (1977: 6).

Discodemus brevipennis Casey, 1908: 61. Synonymy: Doyen (1977: 6).

Discodemus elongatulus Casey, 1908: 61. Synonymy: Doyen (1977: 6).

Discodemus depressulus Casey, 1908: 62. Synonymy: Doyen (1977: 6).

Discodemus subsericeus Casey, 1908: 62. Synonymy: Doyen (1977: 6).

Discodemus knausi Casey, 1908: 62. Synonymy: Doyen (1977: 6).

Eusattus robustus LeConte, 1866USA (CA)

Eusattus robustus LeConte, 1866b: 112.

Eusattus robustus postremus Casey, 1908: 59. Synonymy: Doyen (1977: 6).

Eusattus rudei Doyen, 1984MEX (BS)

Eusattus rudei Doyen, 1984b: 91.

Eusattus secutus Horn, 1894MEX (BS)

Eusattus secutus Horn, 1894b: 421.

Eusattus venosus Champion, 1892MEX (CO JA NA)

Eusattus venosus Champion, 1892: 509.

Eusattus vizcainensis Doyen, 1984MEX (BC BS)

Eusattus vizcainensis Doyen, 1984b: 85.

Tribe Cryptoglossini LeConte, 1862

Centrioptérides Lacordaire, 1859: 134 [nomen oblitum, see Aalbu 2006: 57]. Type genus: Centrioptera Mannerheim, 1843.

Cryptoglossini LeConte, 1862a: 220 [nomen protectum]. Type genus: Cryptoglossa Solier, 1837.

Genus Asbolus LeConte, 1851 [M]

Asbolus LeConte, 1851: 129. Type species: Asbolus verrucosus LeConte, 1851, subsequent designation (Aalbu 2005: 721).

Asbolus laevis LeConte, 1851USA (AZ CA) MEX (BC SO)

Asbolus laevis LeConte, 1851: 130.

Cryptoglossa laevis subsimilis Casey, 1924: 308. Synonymy: Aalbu (2005: 730).

Asbolus mexicanus angularis (Horn, 1894)USA (AZ CA) MEX (BC BS)

Centrioptera angularis Horn, 1894b: 414.

Asbolus mexicanus mexicanus (Champion, 1884)USA (NM TX) MEX (CH CO DU NL)

Cryptoglossa mexicana Champion, 1884: 73.

Cryptoglossa granulifera Champion, 1892: 508. Synonymy: Aalbu (2005: 722).

Asbolus papillosus (Triplehorn, 1964)USA (CA) MEX (SO)

Cryptoglossa laevis papillosa Triplehorn, 1964a: 48.

Asbolus verrucosus LeConte, 1851USA (AZ CA NM NV UT) MEX (BC SO)

Asbolus verrucosus LeConte, 1851: 129.

Cryptoglossa verrucosa carinulatus Blaisdell, 1945: 25. Synonymy: Aalbu (2005: 724).

Genus Cryptoglossa Solier, 1837 [F]

Cryptoglossa Solier, 1837: 680. Type species: Cryptoglossa bicostata Solier, 1837, monotypy.

Centrioptera Mannerheim, 1843: 279. Type species: Centrioptera caraboides Mannerheim, 1843, monotypy. Synonymy: Aalbu et al. (2002: 486).

Oochila LeConte, 1862a: 220. Type species: Asbolus infaustus LeConte, 1854, original designation. Synonymy (with Centrioptera Mannerheim): Horn (1870: 278).

Amblycyphus Motschulsky, 1870: 401. Type species: Amblycyphus asperatus Motschulsky, 1870 (= Centrioptera pectoralis Blaisdell, 1921), monotypy. Synonymy: Aalbu et al. (1995: 483).

Cryptoglossa asperata (Horn, 1870)MEX (BS)

Centrioptera asperata Horn, 1870: 279.

Centrioptera asperata discreta Blaisdell, 1923: 249. Synonymy: Aalbu (2005: 709).

Centrioptera asperata subornata Blaisdell, 1923: 249. Synonymy: Aalbu (2005: 709).

Centrioptera asperata planata Blaisdell, 1923: 250. Synonymy: Aalbu (2005: 709).

Cryptoglossa bicostata Solier, 1837MEX (OA PU)

Cryptoglossa bicostata Solier, 1837: 681.

Cryptoglossa caraboides (Mannerheim, 1843)MEX (GE MO PU)

Centrioptera caraboides Mannerheim, 1843: 280.

Cryptoglossa infausta (LeConte, 1854)USA (TX) MEX (CO DU TA)

Asbolus infaustus LeConte, 1854a: 84.

Centrioptera spiculosa Champion, 1892: 508. Synonymy: Champion (1893a: 572).

Centrioptera texana Blaisdell, 1924b: 88. Synonymy: Aalbu (2005: 705).

Cryptoglossa michelbacheri (Blaisdell, 1943)MEX (BS)

Centrioptera michelbacheri Blaisdell, 1943: 222.

Cryptoglossa muricata (LeConte, 1851)USA (AZ CA NV UT) MEX (BC SO)

Centrioptera muricata LeConte, 1851: 142.

Centrioptera utensis Casey, 1907: 513. Synonymy: Aalbu (2005: 712).

Centrioptera sculptiventris Blaisdell, 1923: 247. Synonymy: Aalbu (2005: 712).

Centrioptera serrata Casey, 1924: 306. Synonymy: Aalbu (2005: 712).

Centrioptera elongata Casey, 1924: 306. Synonymy: Aalbu (2005: 712).

Cryptoglossa seriata cerralvoensis Aalbu, 2005MEX (BS)

Cryptoglossa seriata cerralvoensis Aalbu, 2005: 721.

Cryptoglossa seriata seriata LeConte, 1861USA (AZ CA) MEX (BS)

Cryptoglossa seriata LeConte, 1861a: 337.

Cryptoglossa spiculifera pectoralis (Blaisdell, 1921)USA (CA) MEX (BC BS)

Amblycyphus asperatus Motschulsky, 1870: 404 [junior secondary homonym of Cryptoglossa asperata (Horn, 1870)20].

Centrioptera pectoralis Blaisdell, 1921b: 198. Synonymy: Aalbu et al. (1995: 483).

Centrioptera dulzurae Blaisdell, 1921b: 199. Synonymy: Aalbu et al. (1995: 483).

Centrioptera chamberlini Blaisdell, 1923: 246. Synonymy: Aalbu (2005: 717).

Cryptoglossa spiculifera spiculifera (LeConte, 1861)MEX (BS)

Centrioptera spiculifera LeConte, 1861a: 337.

Cryptoglossa variolosa (Horn, 1870)USA (AZ CA NM) MEX (SI SO)

Centrioptera variolosa Horn, 1870: 280.

Genus Schizillus Horn, 1874 [M]

Schizillus Horn, 1874a: 33. Type species: Schizillus laticeps Horn, 1874, monotypy.

Schizillus laticeps Horn, 1874USA (AZ CA NV UT) MEX (BC)

Schizillus laticeps Horn, 1874a: 33.

Schizillus convexus Blaisdell, 1921b: 203. Synonymy: Aalbu (2005: 732).

Schizillus lomae Blaisdell, 1921b: 206. Synonymy: Aalbu (2005: 732).

Schizillus opacus Casey, 1924: 307. Synonymy: Aalbu (2005: 732).

Schizillus nunenmacheri Blaisdell, 1921USA (AZ CA NV UT)

Schizillus nunenmacheri Blaisdell, 1921b: 204.

Schizillus beali Parker, 1955: 148. Synonymy: Aalbu (2005: 734).

Tribe Edrotini Lacordaire, 1859

Édrotides Lacordaire, 1859: 31. Type genus: Edrotes LeConte, 1851.

Triorophi LeConte and Horn, 1883: 362. Type genus: Triorophus LeConte, 1851.

Auchmobii LeConte and Horn, 1883: 362. Type genus: Auchmobius LeConte, 1851.

Trimytini Casey, 1907: 278. Type genus: Trimytis Leconte, 1851.

Eurymetoponini Casey, 1907: 278. Type genus: Eurymetopon Eschscholtz, 1831.

Trientomini Casey, 1907: 278. Type genus: Trientoma Solier, 1835.

Genus Armalia Casey, 1907 [F]

Armalia Casey, 1907: 289, 330. Type species: Emmenastus texanus LeConte, 1866, original designation.

Armalia alata (Champion, 1884) GUA NIC

Emmenastus alatus Champion, 1884: 13.

Emmenastus salvini Champion, 1884: 13. Synonymy: Champion (1892: 482).

Armalia angularis Casey, 1907USA (TX)

Armalia angularis Casey, 1907: 331.

Armalia belti (Champion, 1884)MEX (YU) GUAHONNIC

Emmenastus belti Champion, 1884: 11.

Emmenastus rotundicollis Champion, 1884: 11. Synonymy: Champion (1892: 480).

Emmenastus intermedius Champion, 1884: 12. Synonymy: Champion (1892: 480).

Armalia brevipennis (Champion, 1884)MEX (NA)

Emmenastus brevipennis Champion, 1884: 10.

Armalia canaliculata (Champion, 1884)MEX (SI)

Emmenastus canaliculatus Champion, 1884: 10.

Armalia chiriquensis (Champion, 1884)PAN / SA

Emmenastus chiriquensis Champion, 1884: 9.

Armalia longicornis (Champion, 1884) GUA

Emmenastus longicornis Champion, 1884: 9.

Armalia solitaria (Champion, 1884)MEX (OA)

Emmenastus solitarius Champion, 1884: 11.

Armalia texana (LeConte, 1866)USA (TX)

Emmenastus texanus LeConte, 1866b: 108.

Armalia variabilis (Champion, 1884)MEX (VE) HON

Emmenastus variabilis Champion, 1884: 10.

Genus Auchmobius LeConte, 1851 [M]

Auchmobius LeConte, 1851: 139. Type species: Auchmobius sublaevis LeConte, 1851, monotypy.

Auchmobius angelicus Blaisdell, 1934USA (CA)

Auchmobius angelicus Blaisdell, 1934b: 249.

Auchmobius parvicollis Blaisdell, 1934USA (CA)

Auchmobius parvicollis Blaisdell, 1934b: 246.

Auchmobius picipes Blaisdell, 1934USA (CA)

Auchmobius picipes Blaisdell, 1934b: 252.

Auchmobius sanfordi Blaisdell, 1934USA (CA)

Auchmobius sanfordi Blaisdell, 1934b: 257.

Auchmobius slevini Blaisdell, 1934USA (CA)

Auchmobius slevini Blaisdell, 1934b: 243.

Auchmobius subboreus Blaisdell, 1934USA (CA NV)

Auchmobius subboreus Blaisdell, 1934b: 254.

Auchmobius sublaevis LeConte, 1851USA (CA)

Auchmobius sublaevis LeConte, 1851: 140.

Auchmobius subovalis Blaisdell, 1934USA (CA)

Auchmobius subovalis Blaisdell, 1934b: 238.

Genus Chilometopon Horn, 1874 [N]

Chilometopon Horn, 1874a: 31. Type species: Trimytis abnormis Horn, 1870, subsequent designation (Casey 1907: 367).

Prometopion Casey, 1907: 366, 370. Type species: Prometopion amplipenne Casey, 1907 (= Chilometopon helopioides Horn, 1874), original designation. Synonymy: MacLachlan and Olson (1990: 72).

Chilometopon abnorme (Horn, 1870)USA (AZ CA NV OR UT) MEX (BC SO)

Trimytis abnormis Horn, 1870: 261.

Chilometopon castaneum Casey, 1907: 373. Synonymy: MacLachlan and Olson (1990: 74).

Chilometopon brevipenne Casey, 1907: 374. Synonymy: MacLachlan and Olson (1990: 74).

Chilometopon ensifer Casey, 1907: 374. Synonymy: MacLachlan and Olson (1990: 74).

Chilometopon brachystomum Doyen, 1983USA (AZ CA NV) MEX (BC)

Chilometopon brachystomum Doyen, 1983: 81.

Chilometopon cribricolle Blaisdell, 1923MEX (BS)

Chilometopon cribricolle Blaisdell, 1923: 230.

Chilometopon helopioides Horn, 1874USA (AZ CA ID NM NV UT) MEX (BC)

Chilometopon helopioides Horn, 1874a: 31.

Prometopion amplipenne Casey, 1907: 372. Synonymy: MacLachlan and Olson (1990: 78).

Chilometopon microps MacLachlan and Olson, 1990USA (CA)

Chilometopon microps MacLachlan and Olson, 1990: 76.

Chilometopon pallidum Casey, 1890USA (AZ CA NM NV TX UT) MEX (BC CH)

Chilometopon pallidum Casey, 1890b: 367.

Chilometopon rugiceps Blaisdell, 1923MEX (BC)

Chilometopon rugiceps Blaisdell, 1923: 229.

Genus Cryptadius LeConte, 1851 [M]

Cryptadius LeConte, 1851: 140. Type species: Cryptadius inflatus LeConte, 1851, monotypy.

Cryptadius inflatus blaisdelli Thomas, 1985MEX (BS)

Cryptadius inflatus blaisdelli Thomas, 1985: 197.

Cryptadius inflatus inflatus LeConte, 1851USA (CA) MEX (BC)

Cryptadius inflatus LeConte, 1851: 140.

Cryptadius oviformis Casey, 1907: 328. Synonymy: Thomas (1985: 196).

Cryptadius punctipennis Casey, 1907: 328. Synonymy: Thomas (1985: 196).

Cryptadius curvipes Casey, 1907: 329. Synonymy: Thomas (1985: 196).

Cryptadius sonorae Berry, 1974MEX (BS SO)

Cryptadius sonorae Berry, 1974: 175.

Cryptadius tarsalis Blaisdell, 1923MEX (BC BS SO)

Cryptadius tarsalis Blaisdell, 1923: 212.

Cryptadius angulatus Blaisdell, 1923: 210. Synonymy: Thomas (1985: 198).

Cryptadius sinuatus Blaisdell, 1923: 211. Synonymy: Thomas (1985: 198).

Cryptadius andrewsi Berry, 1977: 561. Synonymy: Thomas (1985: 198).

Genus Ditaphronotus Casey, 1907 [M]

Ditaphronotus Casey, 1907: 341. Type species: Emmenastus foveicollis Champion, 1884, original designation.

Ditaphronotus championi Casey, 1907 NIC

Ditaphronotus championi Casey, 1907: 342.

Ditaphronotus confusus (Champion, 1884)MEX (CI) GUA

Emmenastus confusus Champion, 1884: 15.

Ditaphronotus foveicollis (Champion, 1884) GUA NIC CRI

Emmenastus foveicollis Champion, 1884: 14.

Ditaphronotus laevicollis (Champion, 1884) PAN

Emmenastus laevicollis Champion, 1884: 15.

Genus Edrotes LeConte, 1851 [M]

Edrotes LeConte, 1851: 140. Type species: Edrotes ventricosus LeConte, 1851, monotypy.

Hedrotes Gemminger [in Gemminger and Harold], 1870: 1816. Unjustified emendation of Edrotes LeConte, 1851, not in prevailing usage.

Subgenus Edrotes LeConte, 1851

Edrotes LeConte, 1851: 140. Type species: Edrotes ventricosus LeConte, 1851, monotypy.

Edrotes fossor Triplehorn, 1972MEX (BS)

Edrotes fossor Triplehorn, 1972: 27.

Edrotes leechi Doyen, 1968USA (AZ CO UT)

Edrotes leechi Doyen, 1968: 218.

Edrotes rotundus (Say, 1824)USA (AZ CO NM TX UT) MEX (BC CH)

Pimelia rotunda Say, 1824a: 251.

Edrotes globosus Casey, 1890a: 175. Synonymy: La Rivers (1947a: 325).

Edrotes inflatus Casey, 1907: 454. Synonymy: La Rivers (1947a: 325).

Edrotes puncticeps Casey, 1907: 454. Synonymy: La Rivers (1947a: 325).

Edrotes intermixtus Casey, 1907: 455. Synonymy: La Rivers (1947a: 325).

Edrotes oblongulus Casey, 1907: 455. Synonymy: La Rivers (1947a: 325).

Edrotes lineatus Casey, 1907: 456. Synonymy: La Rivers (1947a: 325).

Edrotes subaequalis Casey, 1907: 456. Synonymy: La Rivers (1947a: 325).

Edrotes angustulus Casey, 1907: 456. Synonymy: La Rivers (1947a: 325).

Edrotes desertus Blaisdell, 1943: 212. Synonymy: La Rivers (1947a: 325).

Edrotes ventricosus LeConte, 1851USA (AZ CA ID NV OR) MEX (BC BS SO)

Edrotes ventricosus LeConte, 1851: 141.

Edrotes nitidus Casey, 1890a: 175. Synonymy: La Rivers (1947a: 321).

Edrotes orbus Casey, 1907: 452. Synonymy: La Rivers (1947a: 321).

Edrotes angusticollis Casey, 1907: 452. Synonymy: La Rivers (1947a: 321).

Edrotes longipennis Casey, 1907: 453. Synonymy: La Rivers (1947a: 321).

Edrotes mexicanus Blaisdell, 1923: 241. Synonymy: La Rivers (1947a: 321).

Edrotes asperatus Blaisdell, 1923: 241. Synonymy: La Rivers (1947a: 321).

Edrotes laticollis Casey, 1924: 300. Synonymy: La Rivers (1947a: 321).

Edrotes longicornis Casey, 1924: 300. Synonymy: La Rivers (1947a: 321).

Edrotes variipilis Casey, 1924: 301. Synonymy: La Rivers (1947a: 321).

Edrotes barrowsi Dajoz, 1999: 320. New synonymy [RLA].

Subgenus Odrotes La Rivers, 1947

Odrotes La Rivers, 1947a: 320. Type species: Edrotes arens La Rivers, 1947, monotypy.

Edrotes arens La Rivers, 1947USA (AZ CA) MEX (BC)

Edrotes arens La Rivers, 1947a: 320.

Genus Emmenastrichus Horn, 1894 [M]

Emmenastrichus Horn, 1894b: 413. Type species: Emmenastrichus cribratus Horn, 1894, subsequent designation (Casey 1907: 289).

Emmenastrichus cribratus Horn, 1894MEX (BS)

Emmenastrichus cribratus Horn, 1894b: 413.

Emmenastrichus erosus Horn, 1894MEX (BS)

Emmenastrichus erosus Horn, 1894b: 414.

Genus Emmenides Casey, 1907 [M]

Emmenides Casey, 1907: 329. Type species: Emmenastus punctatus LeConte, 1866, original designation.

Emmenides apicalis Blaisdell, 1923MEX (BS)

Emmenides apicalis Blaisdell, 1923: 215.

Emmenides catalinae Blaisdell, 1923MEX (BS)

Emmenides catalinae Blaisdell, 1923: 216.

Emmenides igualensis (Champion, 1892)MEX (GE)

Emmenastus igualensis Champion, 1892: 484.

Emmenides obsoletus Blaisdell, 1923MEX (BS)

Emmenides obsoletus Blaisdell, 1923: 216.

Emmenides punctatus (LeConte, 1866)USA (AZ TX) MEX (BS)

Emmenastus punctatus LeConte, 1866b: 106.

Emmenides subdescalceatus Blaisdell, 1923MEX (BS)

Emmenides subdescalceatus Blaisdell, 1923: 213.

Genus Eremocantor Smith and Wirth, 2016 [M]

Eremocantor Smith and Wirth, 2016: 582. Type species: Eremocantor marioni Smith and Wirth, 2016, original designation.

Eremocantor marioni Smith and Wirth, 2016USA (TX)

Eremocantor marioni Smith and Wirth, 2016: 583.

Genus Eschatomoxys Blaisdell, 1935 [M]

Eschatomoxys Blaisdell, 1935d: 125. Type species: Eschatomoxys wagneri Blaisdell, 1935, original designation.

Eschatomoxys andrewsi Aalbu and Thomas, 2008USA (CA)

Eschatomoxys andrewsi Aalbu and Thomas [in Pape et al.], 2008: 529.

Eschatomoxys paco Aalbu and Thomas, 2008MEX (BC)

Eschatomoxys paco Aalbu and Thomas [in Pape et al.], 2008: 527.

Eschatomoxys pholeter Thomas and Pape, 2008USA (AZ)

Eschatomoxys pholeter Thomas and Pape [in Pape et al.], 2008: 525.

Eschatomoxys rosei Aalbu and Thomas, 2008MEX (BC)

Eschatomoxys rosei Aalbu and Thomas [in Pape et al.], 2008: 530.

Eschatomoxys tanneri Sorenson and Stones, 1959USA (AZ UT)

Eschatomoxys tanneri Sorenson and Stones, 1959: 63.

Eschatomoxys wagneri Blaisdell, 1935USA (AZ CA)

Eschatomoxys wagneri Blaisdell, 1935d: 125.

Genus Eurymetopon Eschscholtz, 1831 [N]

Eurymetopon Eschscholtz, 1831: 5, 8. Type species: Eurymetopon rufipes Eschscholtz, 1831, subsequent designation (Casey 1907: 288).

Eurymetopon ochraceum Eschscholtz, 1831USA (CA)

Eurymetopon ochraceum Eschscholtz, 1831: 8.

Eurymetopon rufipes Eschscholtz, 1831USA (AZ CA) MEX (BS SO)

Eurymetopon rufipes Eschscholtz, 1831: 8.

Genus Garridoa Marcuzzi, 1985 [F]

Garridoa Marcuzzi, 1985: 180. Type species: Garridoa kaszabi Marcuzzi, 1985, monotypy.

Garridoa kaszabi Marcuzzi, 1985 CUB

Garridoa kaszabi Marcuzzi, 1985: 180.

Genus Hylocrinus Casey, 1907 [M]

Hylocrinus Casey, 1907: 289, 331. Type species: Eurymetopon longulum LeConte, 1851, original designation.

Subgenus Hylocrinus Casey, 1907

Hylocrinus Casey, 1907: 289, 331. Type species: Eurymetopon longulum LeConte, 1851, original designation.

Hylocrinus ambiguus (Champion, 1884) PAN

Emmenastus ambiguus Champion, 1884: 13.

Hylocrinus angustus (Casey, 1890)USA (AZ)

Emmenastus angustus Casey, 1890b: 352.

Hylocrinus blaisdelli Casey, 1907USA (CA)

Hylocrinus blaisdelli Casey, 1907: 336.

Hylocrinus breviusculus Casey, 1907USA (TX)

Hylocrinus breviusculus Casey, 1907: 334.

Hylocrinus cunctans Casey, 1907USA (TX)

Hylocrinus cunctans Casey, 1907: 336.

Hylocrinus delicatulus Casey, 1907USA (AZ NV UT)

Hylocrinus delicatulus Casey, 1907: 334.

Hylocrinus depressulus Casey, 1907USA (CA)

Hylocrinus depressulus Casey, 1907: 335.

Hylocrinus filitarsis Casey, 1907USA (CA)

Hylocrinus filitarsis Casey, 1907: 333.

Hylocrinus guatemalensis (Champion, 1884) GUA

Emmenastus guatemalensis Champion, 1884: 14.

Hylocrinus longulus (LeConte, 1851)USA (AZ CA) MEX (BC SO)

Eurymetopon longulum LeConte, 1851: 139.

Hylocrinus magnus Blaisdell, 1923MEX (SO)

Hylocrinus magnus Blaisdell, 1923: 219.

Hylocrinus tenuis Casey, 1907USA (AZ)

Hylocrinus tenuis Casey, 1907: 333.

Subgenus Locrodes Casey, 1907

Locrodes Casey, 1907: 332. Type species: Emmenastus piceus Casey, 1890, present designation.

Hylocrinus brunnescens Casey, 1907USA (UT)

Hylocrinus brunnescens Casey, 1907: 338.

Hylocrinus fraternus Casey, 1907USA (ID UT)

Hylocrinus fraternus Casey, 1907: 338.

Hylocrinus insularis Blaisdell, 1923MEX (BS)

Hylocrinus insularis Blaisdell, 1923: 218.

Hylocrinus laborans Casey, 1907USA (NV UT)

Hylocrinus laborans Casey, 1907: 337.

Hylocrinus mexicanus (Champion, 1892)MEX (FD)

Emmenastus mexicanus Champion, 1892: 481.

Hylocrinus oblongulus Casey, 1907USA (CA) MEX (BC BS)

Hylocrinus oblongulus Casey, 1907: 337.

Hylocrinus parallelus (Champion, 1884)MEX (JA ME MO OA PU SI)

Emmenastus parallelus Champion, 1884: 12.

Hylocrinus piceus (Casey, 1890)USA (CA)

Emmenastus piceus Casey, 1890b: 353.

Hylocrinus seriatus (Champion, 1892)MEX (OA)

Emmenastus seriatus Champion, 1892: 482.

Hylocrinus subapterus (Champion, 1892)MEX (DU)

Emmenastus subapterus Champion, 1892: 481.

Hylocrinus tenebrosus (Champion, 1884)MEX (AG FD GU)

Emmenastus tenebrosus Champion, 1884: 12.

Hylocrinus umbrosus Casey, 1907USA (UT)

Hylocrinus umbrosus Casey, 1907: 338.

Subgenus Paravius Casey, 1907

Paravius Casey, 1907: 332. Type species: Emmenastus marginatus Casey, 1890, monotypy.

Hylocrinus marginatus (Casey, 1890)MEX (BC)

Emmenastus marginatus Casey, 1890b: 351.

Hylocrinus vicinus (Champion, 1884)USA (CA)

Emmenastus vicinus Champion, 1884: 8.

Genus Melanastus Casey, 1907 [M]

Melanastus Casey, 1907: 289. Type species: Eurymetopon atrum LeConte, 1851, original designation.

Melanastus acuminatus Casey, 1907USA (CO)

Melanastus acuminatus Casey, 1907: 362.

Melanastus acutus (Horn, 1870) [Fig. 14] CAN (AB SK) USA (NE)

Figure 14. 

Melanastus acutus (Horn, 1870). Scale bar = 1 mm.

Emmenastus acutus Horn, 1870: 270.

Melanastus aequicollis Casey, 1907USA (CA)

Melanastus aequicollis Casey, 1907: 360.

Melanastus ater (LeConte, 1851)USA (CA ID OR)

Eurymetopon atrum LeConte, 1851: 139.

Melanastus coarcticollis (Casey, 1890)USA (NM)

Emmenastus coarcticollis Casey, 1890b: 364.

Melanastus crassicornis (Casey, 1890)USA (CA)

Emmenastus crassicornis Casey, 1890b: 363.

Melanastus exiguus Casey, 1907USA (CO)

Melanastus exiguus Casey, 1907: 363.

Melanastus exoletus Casey, 1907USA (CA)

Melanastus exoletus Casey, 1907: 357.

Melanastus fallax (Casey, 1890)USA (NM)

Emmenastus fallax Casey, 1890b: 361.

Melanastus finitimus Casey, 1907USA (CO)

Melanastus finitimus Casey, 1907: 359.

Melanastus implicans Casey, 1907USA (CO)

Melanastus implicans Casey, 1907: 358.

Melanastus lucidulus Casey, 1907USA (CA)

Melanastus lucidulus Casey, 1907: 358.

Melanastus ludius Casey, 1907USA (UT)

Melanastus ludius Casey, 1907: 361.

Melanastus moestus Casey, 1907USA (CA)

Melanastus moestus Casey, 1907: 355.

Melanastus nitidus (Casey, 1890)USA (AZ)

Emmenastus nitidus Casey, 1890b: 362.

Melanastus nuperus Casey, 1907USA (AZ)

Melanastus nuperus Casey, 1907: 364.

Melanastus obesus (LeConte, 1851)USA (CA) MEX (BC BS)

Eurymetopon obesum LeConte, 1851: 139.

Emmenastus nanulus Casey, 1884: 45. Synonymy: Casey (1907b: 363).

Melanastus obscurus Blaisdell, 1923 MEX (SO)

Melanastus obscurus Blaisdell, 1923: 226.

Melanastus obtusus (LeConte, 1866)USA (CA)

Emmenastus obtusus LeConte, 1866b: 107.

Melanastus otiosus Casey, 1907USA (CA)

Melanastus otiosus Casey, 1907: 358.

Melanastus parvus Casey, 1907USA (CO)

Melanastus parvus Casey, 1907: 362.

Melanastus sonoricus Casey, 1907MEX (CH MI)

Melanastus sonoricus Casey, 1907: 364.

Melanastus sterilis Casey, 1907USA (CA)

Melanastus sterilis Casey, 1907: 357.

Melanastus texanus Blaisdell, 1926USA (TX)

Melanastus texanus Blaisdell, 1926b: 22.

Melanastus thoracicus (Casey, 1890)USA (CA)

Emmenastus thoracicus Casey, 1890b: 362.

Melanastus vegrandis Casey, 1907USA (CA)

Melanastus vegrandis Casey, 1907: 360.

Genus Mencheres Champion, 1884 [M]

Mencheres Champion, 1884: 5. Type species: Mencheres nicaraguensis Champion, 1884, subsequent designation (Lucas 1920: 403).

Mencheres elongatus Champion, 1884 GUA

Mencheres elongatus Champion, 1884: 6.

Mencheres nicaraguensis Champion, 1884 NIC

Mencheres nicaraguensis Champion, 1884: 5.

Genus Mesabates Champion, 1884 [M]

Mesabates Champion, 1884: 3. Type species: Mesabates latifrons Champion, 1884, monotypy.

Mesabates latifrons Champion, 1884MEX (OA PU)

Mesabates latifrons Champion, 1884: 3.

Mesabates spissicornis Champion, 1892MEX (SI)

Mesabates spissicornis Champion, 1892: 479.

Genus Mesabatodes Casey, 1907 [M]

Mesabatodes Casey, 1907: 517. Type species: Mesabates inaequalis Champion, 1892, original designation.

Mesabatodes inaequalis (Champion, 1892)MEX (AG CH DU)

Mesabates inaequalis Champion, 1892: 480.

Genus Metoponium Casey, 1907 [N]

Metoponium Casey, 1907: 288. Type species: Eurymetopon abnorme LeConte, 1851, original designation.

Subgenus Metoponiopsis Casey, 1907

Metoponiopsis Casey, 1907: 290. Type species: Eurymetopon bicolor Horn, 1870, monotypy.

Metoponium bicolor (Horn, 1870)USA (AZ CA) MEX (BC)

Eurymetopon bicolor Horn, 1870: 268.

Subgenus Metoponium Casey, 1907

Metoponium Casey, 1907: 288. Type species: Eurymetopon abnorme LeConte, 1851, original designation.

Metoponium abnorme abnorme (LeConte, 1851)USA (CA)

Eurymetopon abnorme LeConte, 1851: 138.

Metoponium abnorme faustum Casey, 1907USA (CA)

Metoponium faustum Casey, 1907: 292.

Metoponium abnorme laticolle Casey, 1907USA (AZ) MEX (BC BS)

Metoponium laticolle Casey, 1907: 291.

Metoponium angelicum Blaisdell, 1923MEX (BC)

Metoponium angelicum Blaisdell, 1923: 203.

Metoponium arizonicum Casey, 1907USA (AZ)

Metoponium arizonicum Casey, 1907: 294.

Metoponium candidum Casey, 1907USA (AZ) MEX (SO)

Metoponium candidum Casey, 1907: 292.

Metoponium cognitum Casey, 1907USA (TX)

Metoponium cognitum Casey, 1907: 305.

Metoponium concors Casey, 1907USA (CA)

Metoponium concors Casey, 1907: 305.

Metoponium congener (Casey, 1890)USA (TX)

Eurymetopon congener Casey, 1890b: 333.

Metoponium convexicolle (LeConte, 1851)USA (AZ CA NV) MEX (BC BS)

Eurymetopon convexicolle LeConte, 1851: 139.

Metoponium crassum Casey, 1907USA (AZ)

Metoponium crassum Casey, 1907: 299.

Metoponium cribriceps Casey, 1907USA (NM TX)

Metoponium cribriceps Casey, 1907: 305.

Metoponium cylindricum (Casey, 1890)USA (CA)

Eurymetopon cylindricum Casey, 1890b: 337.

Metoponium dubium (Casey, 1884)USA (AZ)

Eurymetopon dubium Casey, 1884: 44.

Eurymetopon carbonatum Casey, 1884: 43. Synonymy: Horn (1885b: 110).

Metoponium edax Casey, 1907USA (CA)

Metoponium edax Casey, 1907: 309.

Metoponium egregium Casey, 1907USA (CA)

Metoponium egregium Casey, 1907: 300.

Metoponium emarginatum (Casey, 1884)USA (AZ)

Eurymetopon emarginatum Casey, 1884: 41.

Eurymetopon piceum Casey, 1884: 40. Synonymy: Horn (1885b: 110).

Eurymetopon papagonum Casey, 1884: 42. Synonymy: Horn (1885b: 110).

Eurymetopon sculptile Casey, 1884: 44. Synonymy: Horn (1885b: 110).

Metoponium erosum Blaisdell, 1943MEX (BS)

Metoponium erosum Blaisdell, 1943: 175.

Metoponium extensum Casey, 1907USA (AZ)

Metoponium extensum Casey, 1907: 300.

Metoponium fatigans Casey, 1907USA (AZ)

Metoponium fatigans Casey, 1907: 304.

Metoponium fusculum (Casey, 1890)USA (AZ CA)

Eurymetopon fusculum Casey, 1890b: 335.

Metoponium gravidum Casey, 1907USA (CA)

Metoponium gravidum Casey, 1907: 308.

Metoponium gulosum Casey, 1907USA (CA)

Metoponium gulosum Casey, 1907: 307.

Metoponium hebes Casey, 1907USA (AZ)

Metoponium hebes Casey, 1907: 301.

Metoponium insulare Casey, 1907USA (CA)

Metoponium insulare Casey, 1907: 308.

Metoponium integer Casey, 1907USA (CA)

Metoponium integer Casey, 1907: 310.

Metoponium ludificans Casey, 1907USA (TX)

Metoponium ludificans Casey, 1907: 303.

Metoponium molestum Casey, 1907USA (CA)

Metoponium molestum Casey, 1907: 309.

Metoponium nevadense Casey, 1907USA (NV)

Metoponium nevadense Casey, 1907: 307.

Metoponium opacipenne Casey, 1907USA (CA)

Metoponium opacipenne Casey, 1907: 309.

Metoponium pacificum Blaisdell, 1923MEX (BS)

Metoponium pacificum Blaisdell, 1923: 202.

Metoponium pallescens Casey, 1907USA (AZ)

Metoponium pallescens Casey, 1907: 293.

Metoponium parvuliceps Casey, 1907USA (AZ)

Metoponium parvuliceps Casey, 1907: 296.

Metoponium perforatum anceps Casey, 1907USA (NM)

Metoponium anceps Casey, 1907: 294.

Metoponium perforatum congruens Casey, 1907USA (NM)

Metoponium congruens Casey, 1907: 293.

Metoponium perforatum perforatum (Casey, 1890)USA (AZ)

Eurymetopon perforatum Casey, 1890b: 334.

Metoponium phoenicis Casey, 1907USA (AZ)

Metoponium phoenicis Casey, 1907: 301.

Metoponium politum (Casey, 1890)USA (TX)

Eurymetopon politum Casey, 1890b: 338.

Metoponium probatum Casey, 1907USA (CA)

Metoponium probatum Casey, 1907: 310.

Metoponium procerum Casey, 1907USA (AZ)

Metoponium procerum Casey, 1907: 297.

Metoponium prolixum Casey, 1907USA (AZ)

Metoponium prolixum Casey, 1907: 298.

Metoponium rufescens Casey, 1907USA (AZ)

Metoponium rufescens Casey, 1907: 302.

Metoponium rufopiceum Casey, 1907USA (AZ)

Metoponium rufopiceum Casey, 1907: 296.

Metoponium saginatum Casey, 1907USA (TX)

Metoponium saginatum Casey, 1907: 296.

Metoponium socium socium Casey, 1907USA (AZ)

Metoponium socium Casey, 1907: 295.

Metoponium socium subsimile Casey, 1907USA (AZ)

Metoponium subsimile Casey, 1907: 295.

Metoponium subovale Casey, 1907USA (UT)

Metoponium subovale Casey, 1907: 307.

Metoponium tersum Casey, 1907USA (CA)

Metoponium tersum Casey, 1907: 306.

Metoponium testaceum Casey, 1907USA (CA)

Metoponium testaceum Casey, 1907: 303.

Metoponium transversum Blaisdell, 1943MEX (BS)

Metoponium transversum Blaisdell, 1943: 174.

Metoponium truncaticeps Casey, 1907USA (AZ)

Metoponium truncaticeps Casey, 1907: 299.

Genus Micrarmalia Casey, 1907 [F]

Micrarmalia Casey, 1907: 516. Type species: Emmenastus constrictus Champion, 1892, monotypy.

Micrarmalia constricta (Champion, 1892)MEX (GE MO)

Emmenastus constrictus Champion, 1892: 482.

Genus Micromes Casey, 1907 [M]

Micromes Casey, 1907: 432, 441. Type species: Stibia ovipennis Horn, 1874, original designation.

Micromes maritimus (Casey, 1892)USA (CA)

Stibia maritima Casey, 1891: 52.

Micromes ovipennis (Horn, 1874)USA (CA)

Stibia ovipennis Horn, 1874a: 28.

Genus Orthostibia Blaisdell, 1923 [F]

Orthostibia Blaisdell, 1923: 235. Type species: Orthostibia frontalis Blaisdell, 1923, original designation.

Orthostibia fraterna Blaisdell, 1943MEX (BS)

Orthostibia fraterna Blaisdell, 1943: 211.

Orthostibia frontalis Blaisdell, 1923MEX (BS)

Orthostibia frontalis Blaisdell, 1923: 236.

Orthostibia muricata Blaisdell, 1943MEX (BS)

Orthostibia muricata Blaisdell, 1943: 210.

Genus Oxygonodera Casey, 1907 [F]

Oxygonodera Casey, 1907: 433, 444. Type species: Oxygonodera villosa Casey, 1907, original designation.

Oxygonodera grandiceps Casey, 1907USA (UT)

Oxygonodera grandiceps Casey, 1907: 446.

Oxygonodera hispidula (Horn, 1874)USA (ID OR UT WA)

Stibia hispidula Horn, 1874a: 29.

Oxygonodera villosa Casey, 1907USA (UT)

Oxygonodera villosa Casey, 1907: 445.

Genus Pescennius Champion, 1884 [M]

Pescennius Champion, 1884: 3. Type species: Pescennius villosus Champion, 1884, monotypy.

Pescennius villosus Champion, 1884MEX (PU)

Pescennius villosus Champion, 1884: 4.

Genus Pimeliopsis Champion, 1892 [F]

Pimeliopsis Champion, 1892: 477. Type species: Pimeliopsis granulata Champion, 1892, monotypy.

Pimeliopsis granulata Champion, 1892MEX (GE)

Pimeliopsis granulata Champion, 1892: 477.

Genus Posides Champion, 1884 [M]

Posides Champion, 1884: 6. Type species: Posides dissidens Champion, 1884, monotypy.

Posides dissidens Champion, 1884MEX (PU)

Posides dissidens Champion, 1884: 6.

Genus Soemias Champion, 1884 [F]

Soemias Champion, 1884: 4. Type species: Soemias minuta Champion, 1884, monotypy.

Soemias minuta Champion, 1884MEX (VE)

Soemias minuta Champion, 1884: 5.

Genus Steriphanides Casey, 1907 [M]

Steriphanides Casey, 1907: 515. Type species: Emmenastus stolidus Champion, 1892, monotypy.

Steriphanides stolidus (Champion, 1892)MEX (OA)

Emmenastus stolidus Champion, 1892: 483.

Genus Steriphanus Casey, 1907 [M]

Steriphanus Casey, 1907: 289. Type species: Emmenastus conicicollis Casey, 1890, original designation.

Steriphanus aridus Casey, 1907USA (AZ)

Steriphanus aridus Casey, 1907: 347.

Steriphanus conicicollis (Casey, 1890)USA (AZ)

Emmenastus conicicollis Casey, 1890b: 355.

Steriphanus convexus convexus (LeConte, 1866)USA (AZ NM TX)

Emmenastus convexus LeConte, 1866b: 107.

Steriphanus convexus unicolor Casey, 1907USA (NM)

Steriphanus unicolor Casey, 1907: 346.

Steriphanus curtus (Champion, 1884)MEX (DU PU)

Emmenastus curtus Champion, 1884: 16.

Steriphanus discrepans Casey, 1907USA (AZ)

Steriphanus discrepans Casey, 1907: 343.

Steriphanus discretus (Casey, 1890)USA (AZ)

Emmenastus discretus Casey, 1890b: 354.

Steriphanus durus Blaisdell, 1923MEX (BC)

Steriphanus durus Blaisdell, 1923: 224.

Steriphanus ellipticus (Champion, 1884)MEX (“Pensacola”)

Emmenastus ellipticus Champion, 1884: 8.

Steriphanus estebani Blaisdell, 1923MEX (SO)

Steriphanus estebani Blaisdell, 1923: 225.

Steriphanus glabratus (Champion, 1884)MEX (JA OA PU)

Emmenastus glabratus Champion, 1884: 16.

Steriphanus hilaris Casey, 1907USA (AZ UT)

Steriphanus hilaris Casey, 1907: 345.

Steriphanus lentus (Champion, 1884)MEX (CH CO DU)

Emmenastus lentus Champion, 1884: 16.

Steriphanus libertus Casey, 1907USA (AZ)

Steriphanus libertus Casey, 1907: 350.

Steriphanus lubricans Casey, 1907USA (AZ NV)

Steriphanus lubricans Casey, 1907: 345.

Steriphanus lustrans Casey, 1907USA (AZ)

Steriphanus lustrans Casey, 1907: 344.

Steriphanus mancus (Champion, 1884)MEX (GE PU)

Emmenastus mancus Champion, 1884: 15.

Steriphanus nigrans Casey, 1907USA (AZ)

Steriphanus nigrans Casey, 1907: 347.

Steriphanus nitescens Casey, 1907USA (TX)

Steriphanus nitescens Casey, 1907: 344.

Steriphanus perovatus Casey, 1907USA (TX)

Steriphanus perovatus Casey, 1907: 351.

Steriphanus picipes (Champion, 1884)MEX (OA)

Emmenastus picipes Champion, 1884: 17.

Steriphanus placidus Casey, 1907MEX (FD)

Steriphanus placidus Casey, 1907: 347.

Steriphanus proprius Casey, 1907USA (AZ)

Steriphanus proprius Casey, 1907: 347.

Steriphanus pulvinatus (Champion, 1884)MEX (FD HI OA)

Emmenastus pulvinatus Champion, 1884: 17.

Steriphanus rugicollis (Champion, 1884)MEX (SL)

Emmenastus rugicollis Champion, 1884: 17.

Steriphanus rutilans Casey, 1907USA (TX)

Steriphanus rutilans Casey, 1907: 346.

Steriphanus subopacus alutaceus Casey, 1907USA (AZ) MEX (BC SO)

Steriphanus alutaceus Casey, 1907: 347.

Steriphanus subopacus peropacus Casey, 1907USA (AZ)

Steriphanus peropacus Casey, 1907: 349.

Steriphanus subopacus subopacus (Horn, 1870)USA (AZ TX) MEX (BC BS SO)

Emmenastus subopacus Horn, 1870: 269.

Steriphanus torpidus Blaisdell, 1923: 221. Synonymy: Sánchez Piñero and Aalbu (2002: 132).

Steriphanus mucronatus Blaisdell, 1923: 223. Synonymy: Sánchez Piñero and Aalbu (2002: 132).

Steriphanus tardus Blaisdell, 1923MEX (SO)

Steriphanus tardus Blaisdell, 1923: 222.

Genus Stibia Horn, 1870 [F]

Stibia Horn, 1870: 260. Type species: Stibia puncticollis Horn, 1870, monotypy.

Eutriorophus Casey, 1924: 296. Type species: Eutriorophus tuckeri Casey, 1924, original designation. Synonymy: Blaisdell (1933b: 210).

Stibia blairi Blaisdell, 1936USA (AZ CA) MEX (BC)

Stibia blairi Blaisdell, 1936a: 88.

Stibia cribrata Blaisdell, 1923MEX (BS)

Stibia cribrata Blaisdell, 1923: 23921.

Stibia fallaciosa fallaciosa Blaisdell, 1936MEX (BS)

Stibia fallaciosa Blaisdell, 1936a: 70.

Stibia fallaciosa interstitialis Blaisdell, 1936MEX (BS)

Stibia fallaciosa interstitialis Blaisdell, 1936a: 73.

Stibia ferruginea Blaisdell, 1943MEX (BS)

Stibia ferruginea Blaisdell, 1943: 208.

Stibia freyi Kulzer, 1959MEX (SI)

Stibia freyi Kulzer, 1959: 614.

Stibia granulata Blaisdell, 1923MEX (BS)

Stibia granulata Blaisdell, 1923: 238.

Stibia imperialis Blaisdell, 1936USA (AZ CA)

Stibia imperialis Blaisdell, 1936a: 94.

Stibia puncticollis martinensis Blaisdell, 1936MEX (BC)

Stibia puncticollis martinensis Blaisdell, 1936a: 83.

Stibia puncticollis puncticollis Horn, 1870USA (CA) MEX (BC BS SO)

Stibia puncticollis Horn, 1870: 260.

Stibia hannai Blaisdell, 1925b: 329. Synonymy: Blaisdell (1936a: 81).

Stibia sparsa Blaisdell, 1923MEX (BC BS)

Stibia sparsa Blaisdell, 1923: 237.

Stibia tortugensis Blaisdell, 1936a: 100. Synonymy: Sánchez Piñero and Aalbu (2002: 132).

Stibia tanneri Blaisdell, 1936USA (CA)

Stibia tanneri Blaisdell, 1936a: 97.

Stibia tuckeri (Casey, 1924)USA (AZ)

Eutriorophus tuckeri Casey, 1924: 297.

Stibia williamsi Blaisdell, 1925MEX (BC)

Stibia williamsi Blaisdell, 1925b: 328.

Genus Stictodera Casey, 1907 [F]

Stictodera Casey, 1907: 289, 352. Type species: Emmenastus pinguis LeConte, 1866, original designation.

Stictodera pinguis (LeConte, 1866)MEX (BS)

Emmenastus pinguis LeConte, 1866b: 107.

Genus Telabis Casey, 1890 [M]22

Telabis Casey, 1890b: 331. Type species: Eurymetopon longipenne Casey, 1890, subsequent designation (Casey 1907: 288).

Telabis alienus Casey, 1907USA (AZ)

Telabis aliena Casey, 1907: 325.

Telabis amicus Casey, 1907USA (UT)

Telabis amica Casey, 1907: 318.

Telabis asperus Casey, 1907USA (CO)

Telabis aspera Casey, 1907: 322.

Telabis blandus Casey, 1907USA (TX)

Telabis blanda Casey, 1907: 326.

Telabis brevicollis (Champion, 1884)MEX (CO)

Eurymetopon brevicolle Champion, 1884: 7.

Telabis compar Casey, 1907USA (AZ)

Telabis compar Casey, 1907: 321.

Telabis crassulus (Casey, 1890)USA (AZ TX)

Eurymetopon crassulum Casey, 1890b: 344.

Telabis curticollis Casey, 1907USA (AZ)

Telabis curticollis Casey, 1907: 321.

Telabis debilis (Casey, 1890)USA (AZ)

Eurymetopon debile Casey, 1890b: 343.

Telabis discors (Casey, 1890)USA (TX)

Eurymetopon discors Casey, 1890b: 342.

Telabis famelicus Casey, 1907USA (NM)

Telabis famelica Casey, 1907: 323.

Telabis fidelis Casey, 1907USA (CA)

Telabis fidelis Casey, 1907: 320.

Telabis hirtipes Blaisdell, 1923MEX (BC BS)

Telabis hirtipes Blaisdell, 1923: 205.

Telabis histricus (Casey, 1890)USA (AZ)

Eurymetopon histricum Casey, 1890b: 340.

Telabis incisus Casey, 1907USA (CA)

Telabis incisa Casey, 1907: 322.

Telabis inops Casey, 1907USA (AZ)

Telabis inops Casey, 1907: 325.

Telabis latipennis Blaisdell, 1923MEX (BS)

Telabis latipennis Blaisdell, 1923: 207.

Telabis lobifrons Casey, 1907USA (AZ)

Telabis lobifrons Casey, 1907: 318.

Telabis longipennis (Casey, 1890)USA (NM)

Eurymetopon longipenne Casey, 1890b: 339.

Telabis lunulatus Blaisdell, 1923MEX (BC BS)

Telabis lunulata Blaisdell, 1923: 206.

Telabis lustrellus Casey, 1907USA (NM)

Telabis lustrella Casey, 1907: 323.

Telabis mimeticus Casey, 1907USA (TX)

Telabis mimetica Casey, 1907: 319.

Telabis muricatulus (Casey, 1890)USA (AZ TX)

Eurymetopon muricatulum Casey, 1890b: 341.

Telabis nevadensis Blaisdell, 1925USA (NV)

Telabis nevadensis Blaisdell, 1925c: 372.

Telabis obtusus Casey, 1907USA (AZ)

Telabis obtusa Casey, 1907: 317.

Telabis opacellus Casey, 1907USA (CA)

Telabis opacella Casey, 1907: 316.

Telabis ovalis Casey, 1907USA (AZ)

Telabis ovalis Casey, 1907: 324.

Telabis pavidus Casey, 1907USA (NM)

Telabis pavida Casey, 1907: 324.

Telabis prominens Casey, 1907USA (TX)

Telabis prominens Casey, 1907: 314.

Telabis proxima Casey, 1907: 315. Synonymy: Casey (1911: 253).

Telabis punctulatus (LeConte, 1866)MEX (BC BS SO)

Eurymetopon punctulatum LeConte, 1866b: 105.

Telabis rubidus Casey, 1907USA (TX)

Telabis rubida Casey, 1907: 315.

Telabis serratus (LeConte, 1866)USA (AZ CA ID NM NV OR TX) MEX (BC BS SO)

Eurymetopon serratum LeConte, 1866b: 106.

Telabis sodalis (Horn, 1870)USA (AZ CA) MEX (BS)

Eurymetopon sodalis Horn, 1870: 268.

Telabis timidus Casey, 1907USA (AZ)

Telabis timida Casey, 1907: 320.

Telabis uteanus Casey, 1907USA (UT)

Telabis uteana Casey, 1907: 317.

Telabis vafer Casey, 1907USA (AZ)

Telabis vafra Casey, 1907: 315.

Telabis vapidus Casey, 1907USA (TX)

Telabis vapida Casey, 1907: 322.

Genus Telaponium Blaisdell, 1923 [N]

Telaponium Blaisdell, 1923: 209. Type species: Telaponium castaneum Blaisdell, 1923, original designation.

Telaponium castaneum Blaisdell, 1923MEX (BS)

Telaponium castaneum Blaisdell, 1923: 209.

Telaponium pingue Blaisdell, 1943MEX (BS)

Telaponium pingue Blaisdell, 1943: 179.

Genus Texaponium Thomas, 1984 [N]

Texaponium Thomas, 1984: 658. Type species: Cryptadius triplehorni Berry, 1974, original designation.

Texaponium triplehorni (Berry, 1974)USA (TX)

Cryptadius triplehorni Berry, 1974: 172.

Genus Tlascalinus Casey, 1907 [M]

Tlascalinus Casey, 1907: 370. Type species: Trimytis flohri Champion, 1892, monotypy.

Tlascalinus flohri (Champion, 1892)MEX (FD)

Trimytis flohri Champion, 1892: 478.

Genus Trichiotes Casey, 1907 [M]

Trichiotes Casey, 1907: 432, 443. Type species: Trichiotes seriatus Casey, 1907, original designation.

Trichiotes lightfooti Wirth and Smith, 2017MEX (CO)

Trichiotes lightfooti Wirth and Smith, 2017: 535.

Trichiotes seriatus Casey, 1907USA (NM TX) MEX (CO NL)

Trichiotes seriatus Casey, 1907: 444.

Genus Trientoma Solier, 1835 [F]

Trientoma Solier, 1835b: 256. Type species: Trientoma varvasi Solier, 1835, monotypy.

Trientoma cayensis Garrido and Gutiérrez, 1995 CUB

Trientoma cayensis Garrido and Gutiérrez, 1995b: 48.

Trientoma convexipennis Allard, 1883 CUB

Trientoma convexipennis Allard, 1883: 14.

Trientoma garridoi Marcuzzi, 1988 CUB

Trientoma garridoi Marcuzzi, 1988: 69.

Trientoma guadeloupensis Fleutiaux and Sallé, 1890 LAN

Trientoma guadeloupensis Fleutiaux and Sallé, 1890: 421.

Trientoma jilae Steiner, 2006 BAH

Trientoma jilae Steiner, 2006: 3.

Trientoma kaszabi Marcuzzi, 1985 CUB

Trientoma kaszabi Marcuzzi, 1985: 181.

Trientoma kochi Marcuzzi, 1977 CAY

Trientoma kochi Marcuzzi, 1977: 6.

Trientoma laevis Allard, 1883 HAI

Trientoma laevis Allard, 1883: 14.

Trientoma maisiensis Marcuzzi, 1988 CUB

Trientoma maisiensis Marcuzzi, 1988: 67.

Trientoma zayasi Marcuzzi, 1988: 70. Synonymy: Garrido and Gutiérrez (1995b: 48).

Trientoma martinicensis Allard, 1883LAN (Martinique)

Trientoma martinicensis Allard, 1883: 14.

Trientoma puertoricensis Marcuzzi, 1977 PRI

Trientoma puertoricensis Marcuzzi, 1977: 7.

Trientoma rugifrons Champion, 1884MEX / HIS

Trientoma rugifrons Champion, 1884: 2.

Trientoma ryticephala Allard, 1883 HAI

Trientoma ryticephala Allard, 1883: 14.

Trientoma sallei Kraatz, 1865MEX / HAIDOM

Trientoma sallei Kraatz, 1865: 74.

Trientoma mexicana Champion, 1884: 2. Synonymy: Champion (1892: 479).

Trientoma siboneyensis Marcuzzi, 1988 CUB

Trientoma siboneyensis Marcuzzi, 1988: 71.

Trientoma varvasi Solier, 1835 CUB

Trientoma varvasi Solier, 1835b: 257.

Trientoma voegeliorum Steiner, 2006 BAH

Trientoma voegeliorum Steiner, 2006: 8.

Trientoma wickhami Casey, 1907 BAH

Trientoma wickhami Casey, 1907: 377.

Genus Trimytantron Ardoin, 1977 [N]

Trimytantron Ardoin, 1977b: 381. Type species: Trimytantron decui Ardoin, 1977, original designation.

Bielawskia Marcuzzi, 1985: 179. Type species: Bielawskia cubana Marcuzzi, 1985 (= Trimytantron decui Ardoin, 1977), monotypy. Synonymy: Marcuzzi (1998a: 153).

Trimytantron armasi Garrido and Gutiérrez, 1997 CUB

Trimytantron armasi Garrido and Gutiérrez, 1997: 32.

Trimytantron cavernicolous Garrido and Gutiérrez, 1997 CUB

Trimytantron cavernicolous Garrido and Gutiérrez, 1997: 34.

Trimytantron cubanum Ardoin, 1977 CUB

Trimytantron cubanum Ardoin, 1977c: 388.

Trimytantron decui Ardoin, 1977 CUB

Trimytantron decui Ardoin, 1977b: 382.

Bielawskia cubana Marcuzzi, 1985: 179 [junior secondary homonym of Trimytantron cubanum Ardoin, 1977]. Synonymy: Garrido and Gutiérrez (1997: 30).

Trimytantron garridoi Marcuzzi, 1998a: 153. Replacement name for Trimytantron cubanum (Marcuzzi, 1985).

Trimytantron escambrayense Garrido and Gutiérrez, 1997 CUB

Trimytantron escambrayensis Garrido and Gutiérrez, 1997: 33.

Trimytantron litorale Garrido and Gutiérrez, 1997 CUB

Trimytantron litoralis Garrido and Gutiérrez, 1997: 30.

Trimytantron minus 23 Garrido and Gutiérrez, 1997 CUB

Trimytantron minor Garrido and Gutiérrez, 1997: 36.

Trimytantron negreai Ardoin, 1977 CUB

Trimytantron negreai Ardoin, 1977c: 387.

Trimytantron poeyi Ardoin, 1977 CUB

Trimytantron poeyi Ardoin, 1977b: 383.

Trimytantron pumilum Garrido and Gutiérrez, 1997 CUB

Trimytantron pumilus Garrido and Gutiérrez, 1997: 35.

Trimytantron punctulaticeps Garrido and Gutiérrez, 1997 CUB

Trimytantron punctulaticeps Garrido and Gutiérrez, 1997: 34.

Trimytantron sierrae Garrido and Gutiérrez, 1997 CUB

Trimytantron sierrae Garrido and Gutiérrez, 1997: 31.

Trimytantron vinai Ardoin, 1977 CUB

Trimytantron viñai Ardoin, 1977c: 388.

Genus Trimytis LeConte, 1851 [F]

Trimytis LeConte, 1851: 141. Type species: Trimytis pruinosa LeConte, 1851, monotypy.

Pimalius Casey, 1907: 367. Type species: Trimytis pulverea Horn, 1870, original designation. Synonymy: MacLachlan and Olson (1990: 79).

Trimytis ceralboensis Blaisdell, 1943MEX (BS)

Trimytis ceralboensis Blaisdell, 1943: 196.

Trimytis obovata Champion, 1892: 478MEX (CH)

Trimytis obovata Champion, 1892: 478.

Trimytis obtusa Horn, 1894MEX (BS)

Trimytis obtusa Horn, 1894b: 412.

Trimytis pruinosa LeConte, 1851USA (AZ CO KS MT NE NM SD TX WY)

Trimytis pruinosa LeConte, 1851: 141.

Trimytis nympha Casey, 1907: 368. Synonymy: MacLachlan and Olson (1990: 80).

Trimytis tonsa Casey, 1907: 369. Synonymy: MacLachlan and Olson (1990: 80).

Trimytis ignava Casey, 1907: 369. Synonymy: MacLachlan and Olson (1990: 81).

Trimytis trapezifera Casey, 1924: 299. Synonymy: MacLachlan and Olson (1990: 81).

Trimytis pulverea Horn, 1870USA (AZ)

Trimytis pulverea Horn, 1870: 261.

Trimytis subsenilis Blaisdell, 1923MEX (SO)

Trimytis subsenilis Blaisdell, 1923: 227.

Genus Triorophus LeConte, 1851 [M]

Triorophus LeConte, 1851: 141. Type species: Triorophus laevis LeConte, 1851, subsequent designation (Casey 1907: 432).

Triorophus basalis Casey, 1907USA (AZ)

Triorophus basalis Casey, 1907: 437.

Triorophus brevis Casey, 1907USA (TX)

Triorophus brevis Casey, 1907: 439.

Triorophus gracilicornis Casey, 1907USA (CA)

Triorophus gracilicornis Casey, 1907: 437.

Triorophus gravidulus Casey, 1907USA (AZ)

Triorophus gravidulus Casey, 1907: 437.

Triorophus histrio Casey, 1907USA (AZ)

Triorophus histrio Casey, 1907: 437.

Triorophus laevis laevis LeConte, 1851USA (AZ CA NV) MEX (SO)

Triorophus laevis LeConte, 1851: 141.

Triorophus laevis politus Casey, 1907USA (CA NV)

Triorophus politus Casey, 1907: 435.

Triorophus lariversi Blaisdell, 1942USA (NV)

Triorophus lariversi Blaisdell, 1942: 132.

Triorophus laticeps Casey, 1924USA (TX)

Triorophus laticeps Casey, 1924: 297.

Triorophus lecontei Casey, 1890USA (TX) MEX (CH DU)

Triorophus lecontei Casey, 1890b: 327.

Triorophus longicornis Casey, 1907USA (AZ)

Triorophus longicornis Casey, 1907: 438.

Triorophus mixtus Casey, 1907USA (TX)

Triorophus mixtus Casey, 1907: 440.

Triorophus mundulus Casey, 1907USA (AZ)

Triorophus mundulus Casey, 1907: 436.

Triorophus nevadensis Casey, 1924USA (NV)

Triorophus nevadensis Casey, 1924: 298.

Triorophus nodiceps LeConte, 1853USA (TX) MEX (CO)

Triorophus nodiceps LeConte, 1853: 446.

Triorophus puberulus Casey, 1924USA (CA)

Triorophus puberulus Casey, 1924: 298.

Triorophus punctatus LeConte, 1851USA (CA)

Triorophus punctatus LeConte, 1851: 142.

Triorophus rugiceps LeConte, 1851USA (CA ID)

Triorophus rugiceps LeConte, 1851: 142.24

Triorophus simplex Casey, 1907USA (AZ)

Triorophus simplex Casey, 1907: 436.

Triorophus subpubescens Horn, 1870USA (CA)

Triorophus subpubescens Horn, 1870: 259.

Triorophus terebratulus Casey, 1907USA (AZ)

Triorophus terebratulus Casey, 1907: 436.

Triorophus thoracicus Casey, 1924USA (AZ)

Triorophus thoracicus Casey, 1924: 298.

Genus Triphalopsis Blaisdell, 1923 [F]

Triphalopsis Blaisdell, 1923: 232. Type species: Triphalopsis partida Blaisdell, 1923, original designation.

Triphalopsis californica Doyen, 1983USA (CA) MEX (BC)

Triphalopsis californicus Doyen, 1983: 87.

Triphalopsis impressicollis Blaisdell, 1943MEX (BC)

Triphalopsis impressicollis Blaisdell, 1943: 203.

Triphalopsis partida Blaisdell, 1923MEX (BC BS SO)

Triphalopsis partida Blaisdell, 1923: 232.

Triphalopsis minor Blaisdell, 1923: 233. Synonymy: Sánchez Piñero and Aalbu (2002: 132).

Genus Triphalopsoides Doyen, 1990 [M]

Triphalopsoides Doyen, 1990: 222. Type species: Triphalopsoides lasiodorsa Doyen, 1990, monotypy.

Triphalopsoides lasiodorsa Doyen, 1990MEX (JA)

Triphalopsoides lasiodorsa Doyen, 1990: 224.

Genus Triphalus LeConte, 1866 [M]

Triphalus LeConte, 1866b: 104. Type species: Triphalus perforatus LeConte, 1866, monotypy.

Triphalus cribricollis Horn, 1895MEX (BS)

Triphalus cribricollis Horn, 1895: 251.

Triphalus impressifrons Blaisdell, 1943MEX (BS)

Triphalus impressifrons Blaisdell, 1943: 202.

Triphalus perforatus LeConte, 1866MEX (BS)

Triphalus perforatus LeConte, 1866b: 104.

Triphalus subcylindricus Blaisdell, 1923MEX (BS)

Triphalus subcylindricus Blaisdell, 1923: 234.

Genus Troglogeneion Aalbu, 1985 [N]

Troglogeneion Aalbu, 1985: 541. Type species: Troglogeneion zapoteca Aalbu, 1985, monotypy.

Troglogeneion zapoteca Aalbu, 1985MEX (OA)

Troglogeneion zapoteca Aalbu, 1985: 542.

Tribe Epitragini Blanchard, 1845

Lygophila Rafinesque, 1815: 113 [nomen oblitum, see Bouchard et al. (2007: 386)]. Type genus: Lygophilus Rafinesque, 1815 (= Epitragus Latreille, 1802).

Épitragites Blanchard, 1845: 16 [nomen protectum]. Type genus: Epitragus Latreille, 1802.

Genus Bothrotes Casey, 1907 [M]

Bothrotes Casey, 1907: 379, 398. Type species: Epitragus canaliculatus Say, 1824, original designation.

Bothrotes angusticollis (Champion, 1884)MEX (GE JA SI)

Epitragus angusticollis Champion, 1884: 26.

Bothrotes bicarinatus (Champion, 1884)MEX (CL VE)

Epitragus bicarinatus Champion, 1884: 25.

Bothrotes canaliculatus acutus (LeConte, 1866)USA (FL KS NM OK TX) MEX (CO)

Epitragus acutus LeConte, 1866b: 108.

Bothrotes fortis Casey, 1907: 399. Synonymy: Freude (1967: 283).

Bothrotes subrudis Casey, 1907: 400. Synonymy: Freude (1967: 283).

Bothrotes pensus Casey, 1907: 400. Synonymy: Freude (1967: 283).

Bothrotes knausi Casey, 1907: 401. Synonymy: Freude (1967: 283).

Bothrotes canaliculatus arundinis (LeConte, 1866)USA (DE GA MD NC NJ NY SC VA)

Epitragus arundinis LeConte, 1866b: 108.

Bothrotes pinorum Casey, 1924: 304. Synonymy: Freude (1967: 281).

Bothrotes canaliculatus canaliculatus (Say, 1824)USA (AZ CO IL KS MO NM OH SD TX WI) MEX (CH DU SO ZA)

Epitragus canaliculatus Say, 1824b: 281.

Bothrotes canaliculatus mexicanus Freude, 1967MEX (DU NL TA)

Bothrotes canaliculatus mexicanus Freude, 1967: 285.

Bothrotes canus (Champion, 1884)MEX (GE)

Epitragus canus Champion, 1884: 34.

Bothrotes cristatus (Champion, 1892)MEX (CO GE)

Epitragus cristatus Champion, 1892: 485.

Bothrotes foveatus (Champion, 1884)MEX (OA VE)

Epitragus foveatus Champion, 1884: 29.

Bothrotes hoegei (Champion, 1884)MEX (MI VE)

Epitragus högei Champion, 1884: 26.

Bothrotes inaequalis (Champion, 1884)MEX (OA PU VE)

Epitragus inaequalis Champion, 1884: 32.

Bothrotes incisus (Champion, 1884)MEX (NA [Islas Marías])

Epitragus incisus Champion, 1884: 28.

Bothrotes littoralis (Champion, 1884)MEX (GE JA MO NA OA SI)