Macrotomoderus latipennis Pic, 1901: 741 by monotypy.

Holotype • ♂ NME: “CHINA: Shaanxi/Sichuan Daba Shan, pass 20 km SSE Zhenping ~1750 m leg. Schülke 12.7.2001” [printed]. Paratype • 1♀ DTC: “CHINA: Shaanxi: Dabashan; 20 km SSE Zhenping 1700 m leg. Stary 26.6.2002” [printed].

Holotype, total body length 3.9 mm; head including exposed part of cranial ‘neck’ 0.8 mm long, across eyes 0.8 mm wide, pronotum 1 mm long, maximum width 0.8 mm, minimum width 0.3 mm, elytra 2.1 mm long, combined width 1.4 mm. Paratype ♀ 4 mm long.

Holotype, male. Dorsum and venter uniformly brown, head comparatively slightly darker. Mouthparts, antennae, palps, and legs brownish testaceous. Head transversely ovoid, glossy dorsally and ventrally, with rather small, distinctly ovoid compound eyes which are not protruding beyond lateral or dorsal outline of head. Head rounded in broad arc posterior to eyes. Head dorsal punctures minute and inconspicuous but rather deep. Intervening spaces 4–6 × as wide as diameter of punctures. Head dorsal setae inconspicuous, moderately dense, whitish to yellowish. Antenna reach base of pronotum when directed posteriad. Antennomere 3 subequal in length to antennomere 2, antennomeres 6 and 7 approximately as long as wide, 8–10 transverse, of which 9–10 strongly so. Terminal antennomere strongly asymmetrically triangular with rounded apex, ~ 1.5–1.6 × as long as penultimate antennomere. Terminal maxillary palpomere securiform. Pronotum stout, moderately glossy dorsally and laterally, narrower than head across compound eyes, with broad, medially distinctly notched (in dorsal view) postmedian lateral constriction. Front margin of anterior lobe very broadly rounded, dorsally without modifications and anterior rim. Anterior lobe barely convex in lateral view (Fig. 1B). Lateral constriction barely continues onto pronotal disc in lateral view (Fig. 1B). Lateral pronotal fovea moderately broad at lower (lateroventral) extent of laterally strongly declivous pronotal disc, somewhat widens upwards towards pronotal disc in lateral view, lateral edges of fovea carinate, narrowly separated (all in lateral view), denticle-like in dorsal view. In lateral view anterior and posterior edge of pronotal fovea covered with bristle of short golden setae except at their lower extent. Cavity in lateral wall of pronotum between lateral denticles moderate, deep. In dorsal view, lateral pronotal fovea moderately wide, anterior and posterior denticle poorly visible, glabrous. Pronotal punctures on disc similar to those on head dorsum; lateral constriction dorsally with dense, large, irregularly circular punctures with corrugate backgrounds separated by much less than puncture diameters. Dorsal pronotal setae similar to those on head dorsum, denser on lateral sides of pronotum. Few longer erect tactile setae on lateral sides of anterior lobe. Scutellar shield minute, apically rounded, glabrous, and glossy. Elytra moderately glossy, dorsally elliptical, slightly convex in lateral view, strongly widened laterally around midlength, lateral margins broadly rounded, humerus obsolete (apterous species). Elytral punctures much stronger and larger than those on dorsal forebody, more or less regularly circular, smaller, and less coarse than dorsal punctures on pronotal constriction. Punctures flatter but not much sparser on apical third of elytra. Intervening spaces ~ 1.5–2 × as wide as diameter of punctures. Elytral setae long and sparse, suberect, yellowish. Male tergite and sternite VII broadly rounded at posterior margin. Sternite IX rod-like, strongly sinuous (Fig. 2A). Aedeagus as in Fig. 2B–H, thick and bulbous in basal portion, narrowing towards subtruncate apex. Endophallic armature of numerous minute, peculiarly tack-shaped spines in apical part and, in basal portion, with large, pebble-like sclerites arranged in a kind of garland and intermixed with dense spines filling intervening spaces between sclerites (Fig. 2B–E).

Figure 1. 

Macrotomoderus blinsteini sp. nov. A holotype ♂, habitus, dorsal view B ditto, lateral view C ditto, dorsal forebody D paratype ♀, pronotum, posterodorsal view. Scale bars: 1 mm.

Figure 2. 

Macrotomoderus blinsteini sp. nov. holotype ♂, terminalia and aedeagus A sternite IX, lateral view B aedeagus, ventral view C ditto, lateral view D ditto, endophallic armature in median portion of aedeagus E ditto, endophallic armature in basal portion of aedeagus F ditto, different portion of basale G ditto, lateral view of apical portion with endophallic armature H ditto, ventral view of apex [not to scale].

Sexual dimorphism. Female tergite and sternite VII broadly rounded at posterior margin, pronotum comparatively slenderer and elytra somewhat stronger constricted towards apex than in male.

This species falls in a group of species from continental China including, for example, M. hartmanni Telnov, 2022 and M. korolevi Telnov, 2022 (both from Yunnan) with the lateral constriction area of the pronotum densely and roughly but ordinary punctate and sparsely to moderately densely setose dorsally, lacking the longitudinal carinae, sulci or elongate pores. The aedeagus is differently shaped in M. blinsteini sp. nov., the endophallic armature is smaller, thinner and peculiarly tack-shaped, and the elytral punctures are comparatively coarser and deeper.

Collected at ~ 1700–1750 m a.s.l.

Known from the Daba Mountains in southern Shaanxi Province along boundary with Sichuan, central China.

Patronymic. This species named for Semen Blinstein (Dortmund, Germany; previously Odessa, Ukraine) – a well-known coleopterist who possesses a valuable beetle collection from southern Ukraine and is the author of some Anthicidae species from the region.

Holotype • ♂ NME: “CHINA: Hubei: Dabashan; 13 km NW Muyuping 1900 m leg. Stary 16.7.2002” [printed].

Holotype, total body length 3.8 mm; head including exposed part of cranial ‘neck’ 0.9 mm long, across eyes 0.9 mm wide, pronotum 1.1 mm long, maximum width 0.9 mm, minimum width 0.35 mm, elytra 2.3 mm long, 1.7 mm combined wide.

Holotype, male. Head and pronotum brown, elytra pale brown. Mouthparts, antennae, palps, and legs pale brownish–testaceous. Head ovoid, moderately glossy dorsally and ventrally, with moderate, nearly circular compound eyes which are slightly protruding beyond lateral outline of head. Head rounded in broad arc posterior to eyes. Head dorsal punctures minute but rather deep, denser on occiput and vertex. Intervening spaces 2–4 × as wide as punctures. Head dorsal setae yellowish, inconspicuous on most of head dorsum but long and dense, forming a subconical bristle (in dorsal view) at head base and therefore concealing median part of anterior pronotal margin. Apical portions of these long head–base setae are curled and in part tangled (Fig. 3B, C). Antenna exceeds slightly beyond base of elytra when directed posteriad. Antennomere 3 subequal in length to antennomere 2, antennomeres 6–10 transverse, of which 8–10 strongly so. Terminal antennomere asymmetrically triangular with rounded apex, ~ 1.6 × as long as penultimate antennomere. Terminal maxillary palpomere securiform. Pronotum stout, moderately glossy dorsally and laterally, approximately as wide as head across compound eyes, with broad, medially strongly notched (in dorsal view) postmedian lateral constriction. Anterior lobe much wider than posterior, its front margin broadly rounded, dorsally without modifications and anterior rim. Anterior lobe nearly flat in lateral view (Fig. 3B). Lateral constriction does not continue onto disc in lateral view (Fig. 3B). Lateral pronotal fovea wide at lower (lateroventral) extent of laterally strongly declivous pronotal disc, not widens upwards towards pronotal disc in lateral view, lateral edges of fovea carinate, moderately separated and densely setose (all in lateral view), in dorsal view denticle-like. In lateral view anterior and posterior edge of pronotal fovea completely covered with bristle of short golden setae. Cavity in lateral wall of pronotum moderate. In dorsal view, lateral pronotal fovea wide, anterior and posterior denticle clearly visible, dense brush-like setose (Fig. 3A, C). Pronotal punctures on disc similar to those on head dorsum, intervening spaces 3–7 × as wide as punctures; lateral constriction dorsally with irregularly shaped and variably dense and deep, generally moderately to strongly elongated punctures; median ones particularly large and elongate, groove-like. Dorsal pronotal setae moderately dense, but much denser on lateral sides of pronotum, especially in constriction area where setation effectively conceals shape of lateral pronotal denticles. Several longer erect tactile setae on lateral sides of anterior lobe. Scutellar shield minute, apically rounded, glabrous, and glossy. Elytra moderately glossy, dorsally elongate elliptical, slightly convex in lateral view, widened laterally around midlength, lateral margins broadly rounded, humerus obsolete (apterous species). Elytral punctures stronger and larger than those on dorsal forebody, more or less regularly circular, smaller than dorsal punctures on lateral pronotal constriction. Punctures becoming more flat and somewhat sparser on apical third of elytra. Intervening spaces ~ 3–5 × as wide as diameter of punctures. Elytral setae long and sparse, suberect, yellowish. Male tergite and sternite VII broadly rounded at posterior margin. Sternite IX rod–like, slightly bisinuate (Fig. 4A). Aedeagus as in Fig. 4B–G, elongate, narrows apically, apex somewhat subspatulate, widened and rounded. Endophallic armature of numerous large peculiarly spike/nail-shaped spines (Fig. 4B–F).

Figure 3. 

Macrotomoderus hirsutus sp. nov. holotype ♂ A habitus, dorsal view B ditto, lateral view C dorsal forebody. Scale bars: 1 mm.

Figure 4. 

Macrotomoderus hirsutus sp. nov. holotype ♂, terminalia and aedeagus A sternite IX, lateral view B aedeagus, ventral view C ditto, lateral view D ditto, median portion, endophallic armature E ditto, basal portion, endophallic armature F ditto, endophallic armature, lateral view of basal portion G ditto, ventral view of apex [not to scale].

Sexual dimorphism. Female is unknown.

This species is readily recognized due to the presence of the dense clump of setae on male head base in the combination with the peculiar, spike/nail-like endophallic armature of the aedeagus. The shape of the aedeagus somewhat resembles that of M. dali Telnov, 2022 (Yunnan, China), M. muli Telnov, 2022 (Sichuan, China), and M. wudu Telnov, 2022 (Gansu, China), but all three have entirely different endophallic armatures. The head base with more or less dense setation is present in M. conus Telnov, 2018, M. gracilis Telnov, 2018, M. microscopicus Telnov, 2018, M. monstrificabilis Telnov, 2018, M. perforatus Telnov, 2018, and M. schuelkei Telnov, 2018 (all from Yunnan, China), but other morphological features and the endophallic armature are quite different.

Collected at 1900 m a.s.l.

Known from Daba Mountains in western part of Hubei Province, central China.

From Latin hirsutus – shaggy, hairy, bristly, referring to the bristle of setae on the head base of this species.

Holotype • ♂ IBC: “CH, S Sichuan, S of Xichang, E slope of Mt. ´4282 ´ (NE of Dechang) 3200–2800 m, 6.05.2001 Belousov & Korolev leg.” [printed] // “Macrotomoderus sp.? n. aff. monstrificabilis [handwritten] A. Kovalev det. 20 [printed] 19” [handwritten] [label with black frame]. The holotype will be donated to a local public institution (I. Belousov, pers. comm. viii.2021). Paratype • 1♂ DTC: same labels as holotype.

Holotype, total body length 4.1 mm; head 0.8 mm long, across eyes 0.8 mm wide, pronotum 1.2 mm long, maximum width 1 mm, minimum width 0.25 mm, elytra 2.2 mm long, 1.5 mm combined wide. Paratype ♂ 4 mm long.

Holotype, male. Dorsum and venter brown, posterior lobe of pronotum slightly paler. Mouthparts, antennae, palps, and legs brownish testaceous. Head subtriangular, moderately glossy dorsally and ventrally, with moderate, slightly ovoid compound eyes which are slightly protruding beyond lateral outline of head. Tempus short, constricted towards head base, temporal angle rounded. Head base truncate, declivous. Head dorsal punctures minute and inconspicuous, flat. Intervening spaces much wider than punctures. Head dorsal setae yellowish, short, moderately dense. Head base medially with somewhat longer, apically curled golden setae. Antenna exceeds slightly beyond base of elytra when directed posteriad. Antennomere 3 ~ 2 × as long as antennomere 2, asymmetrical, distal edge obliquely emarginate to accommodate shortened and only slightly longer than wide antennomere 4, antenna appears somewhat bent at area of antennomeres 3–5 (Fig. 5A, B, F). Antennomeres 8–10 transverse, of which antennomere 10 strongly so. Terminal antennomere strongly asymmetrically triangular with rounded apex, ~ 1.8–1.9 × as long as penultimate antennomere. Terminal maxillary palpomere securiform. Pronotum stout, moderately glossy dorsally and laterally, much wider than head across compound eyes, with moderately broad, medially strongly notched (in dorsal view) postmedian lateral constriction. Anterior lobe much wider than posterior, its front margin medially subtruncate, without anterior rim, with moderate mesal impression and transverse median ridge covered with dense golden setae which are directed antero-dorsally (Fig. 5C–E). Pronotum flattened anteroventrally each side of subtruncate median part of its anterior margin (Fig. 5D, E). Anterior lobe convex in lateral view (Fig. 5B). Lateral constriction barely continues onto disc in lateral view (Fig. 5B). Lateral pronotal fovea broad at lower (lateroventral) extent of laterally strongly declivous pronotal disc, not or barely widens upwards towards pronotal disc, lateral edges of fovea carinate, narrowly separated, nearly glabrous (all in lateral view), in dorsal view denticle-like. In lateral view anterior and posterior edge of pronotal fovea covered with a bristle of moderately long whitish setae. In dorsal view, lateral pronotal fovea wide, anterior and posterior denticle clearly visible, in part setose (Fig. 5A, C). There is an inconspicuous trace of short dorsal median longitudinal carina in lateral constriction area. Pronotal punctures on disc small and sparse, intervening spaces 5–8 × as wide as diameter of punctures; lateral constriction dorsally with irregularly ovoid and variably dense, deep punctures. Intervening spaces among punctures in laterally constricted area vary from much narrower than to twice as wide as punctures. Dorsal pronotal setae short and dense, much longer and denser at lateral pronotal fovea, in part concealing its structure in dorsal view. Few longer erect tactile setae on lateral sides of anterior lobe. Scutellar shield minute, apically rounded, moderately glossy. Elytra moderately glossy, dorsally elongate elliptical, moderately convex in lateral view, widened laterally around midlength, lateral margins broadly rounded, humerus broadly rounded. Metathoracic wings fully developed (functional). Elytral punctures stronger and larger than those on dorsal forebody, nearly as large as those on lateral constriction area, more or less regularly circular. Punctures becoming more flat and somewhat sparser on apical third of elytra. Intervening spaces ~ 2.5–4 × as wide as diameter of punctures. Elytral setae long and moderately dense, suberect, yellowish. Male tergite and sternite VII broadly rounded at posterior margin. Aedeagus as in Fig. 6, elongate and narrow, narrows and acutely angulate apically. Endophallic armature of numerous basally multifurcate spines (Fig. 6A–E).

Figure 5. 

Macrotomoderus turpiculus sp. nov. ♂ A holotype, habitus, dorsal view B ditto, lateral view C ditto, dorsal forebody D paratype, forebody, anterodorsal view E ditto, latero-dorsal view F holotype, right antenna. Scale bars: 1 mm.

Figure 6. 

Macrotomoderus turpiculus sp. nov. holotype ♂, aedeagus A ventral view B lateral view C median portion, lateral view D basal portion, lateral view E basal portion, endophallic armature, ventral view F apical portion [not to scale].

Sexual dimorphism. Female is unknown.

This species falls into a group of species from mainland China with strongly widened, apically subtruncate anterior margin of pronotum, long heavy antennae, and slender, dagger-shaped aedeagus: M. imitator Telnov, 2022, M. monstratus Telnov, 2018, and M. monstrificabilis Telnov, 2018 (all from Yunnan). Macrotomoderus turpiculus sp. nov. is peculiar due to the absence of the large elongate median projection on head base (large projection present in M. monstrificabilis, only small projection in M. turpiculus sp. nov.; cf. Fig. 5A, C, E), the truncate and declivous head base (the head base broadly rounded in the similar species, not declivous), the differently modified anterior pronotal margin (see the description) especially the anteroventrally flattened pronotum, the asymmetrical antennomere 3 and the shortened antennomere 4 (both not modified in M. monstratus), the wide head (the head is narrower than the pronotum in M. imitator), and the peculiar endophallic armature not like in other congeners.

Collected between 2800–3200 m a.s.l.

Known only from southern part of Sichuan Province, southwestern China.

From Latin turpiculus – misshaped, ugly, referring to the unusual body shape of this species.