The first part of the name Trigonalyidae refers most likely to protuberance of the second sternite present in females of several spp. (but absent in the species treated in this paper, except Bareogonalos Schulz), which is triangular in lateral view.

Reared from nests of Vespa, Vespula, Dolichovespula and Provespa spp. (Vespinae: Vespidae); the larva of at least one species has a final ectoparasitoid phase (Carmean and Kimsey 1998) after the initial endoparasitoid phase. Collected in August–October.

East Palaearctic, Nearctic (but intruding Central America by reaching SW Mexico), Oriental.

Holotype, ♀ (NWUX) “NW China: Shaanxi, Fengxian County, Baoji, Jialingjiang Riv[er] Source, 1513 m, 34°25'N 106°94'E, 1.ix.2016, JL. Tan & QQ. Tan, NWUX”, “Reared from nest of Vespula structor (Smith)”. Paratypes: 6 ♀ + 8 ♂ (NWUX, RMNH), same label data.

Left mandible with two subapical teeth, ventral tooth shorter than dorsal tooth; maxillary palp with six segments; width of head in dorsal view 1.05–1.10 times maximum width of mesoscutum (Figs 4, 15); anterior half of scutellum flat and at same level as mesoscutum; metanotum with large protuberance and long setae; wing membrane near veins 1-SR and 1-M of fore wing subhyaline as remainder of wing except darkened apical third of wing (Figs 3, 14); hind basitarsus slender (Fig. 12); fourth and fifth tergites largely smooth and shiny; third sternite of ♀ in ventral view distinctly protruding medio-posteriorly (but obscured by second sternite). Differs from the only other known Palaearctic species, B. jezoensis (Uchida) mainly by the colour of the wing membrane (largely subhyaline vs largely yellowish in B. jezoensis; Fig. 23), the median groove of the mesoscutum (absent posteriorly vs present) and the wider head (1.05–1.10 times vs 0.80–0.90 times width of mesoscutum).

Holotype, ♀, length of body 9.2 mm (of fore wing 8.5 mm).

Head. Antenna with 20 segments, segments of apical half 1.1–1.3 times as long as wide (Fig. 13); frons and vertex smooth and strongly shiny, flat except a shallow depression behind each posterior ocellus (Figs 9, 15), with fine and long greyish setae; head gradually narrowed behind eyes and 1.05 times as wide as mesoscutum (Fig. 4); dorsal length of eye 0.9 times length of temple (Fig. 9); temple smooth and shiny; occipital carina thick and moderately lamelliform medio-dorsally and with a short crenula; supra-antennal elevations low, as a thin rim and smooth; clypeus moderately depressed medio-ventrally and flat above depression.

Mesosoma. Length of mesosoma 1.1 times its height (Fig. 5); mesopleuron below transverse mesopleural groove densely moderately reticulate-rugose, narrowly smooth posteriorly and with satin sheen, above groove coarsely reticulate; transverse mesopleural groove wide, deep and coarsely crenulate (Fig. 5); notauli medium-sized and distinctly crenulate; mesoscutum coarsely rugose-reticulate and with satin sheen contrasting with shiny head, antero-medially with parallel-sided flattened area, convex but medially slightly depressed (Fig. 4); scutellar sulcus curved, parallel-sided and coarsely crenulate; scutellum reticulate-rugose, convex and anteriorly above level of mesoscutum; metanotum medially with large and coarsely reticulate trapezoid protuberance (Figs 4, 5), apically shallowly emarginate, in lateral view not or hardly protruding over base of propodeum; propodeum largely finely reticulate-rugose, but anteriorly coarsely crenulate (Fig. 4).

Metasoma. First tergite 0.3 times as long as apically wide (of paratype, obscured by propodeum in holotype), gradually narrowed basally, flat medially and straight apically; second tergite smooth and strongly shiny as following tergites, but with lateral patch of fine punctures; second sternite rather densely finely punctate and shiny, its medio-apical protuberance densely setose and widely truncate medially (Fig. 7), protuberance of third sternite similar but smaller and obscured by second sternite in ventral view (Fig. 7); down curved apex of metasoma nearly up to protuberance of second sternite (Fig. 8).

Legs. Hind coxa and femur smooth and shiny.

Colour. Black; palpi, mandible apically, antenna (but scapus yellow ventrally and basal third of antenna brown), tegulae (but humeral plate brown), first tergite, veins (except yellowish veins 1-SR, 1-M and 1-SR+M of fore wing) and pterostigma dark brown; coxae basally black; femora (except yellow anterior face of fore femur, base and apex of middle and hind femora), and inner side of hind tibia (and partly ventrally) dark brown; 3 apical segments of hind tarsus and infuscate; pronotum dorso-apically, pair of wide patches on mesoscutum anteriorly, axilla, scutellum latero-posteriorly, dorso-apical patch of mesopleuron, pair of large antero-lateral patches on second–sixth tergites (touching each other on fourth and fifth tergites), pair of small subposterior patches on first sternite, large V-shaped patch on second sternite, and remainder of legs yellow; membrane of fore wing subhyaline, but apical 0.4 weakly infuscate (Fig. 3).

Variations. Length of body 8.0–9.3 mm, of fore wing 8.0–8.9 mm; antenna of ♀ with 20(2) or 21(2) segments; width of head in dorsal view 1.05–1.10 times maximum width of mesoscutum; mesoscutum sometimes entirely black, or with additional yellow spot medio-posteriorly; length of vein 1-M of fore wing 2.1–3.1 times as long as vein 1-SR; mandible entirely blackish brown or with some brownish patches subapically; antero-lateral yellow patches of metasoma either all separated from each other, or all touching medially, but sometimes only on second–fourth tergites being separated; yellow patch of second sternite may be separated into large patches; inner side of hind tibia largely dark brown or only apical third.

Male. Length of body 7.2–9.6 mm, of fore wing 6.8–8.0 mm; antenna with 20(4) or 21(1) segments, basal quarter of antenna black, but scapus pale yellow with upper part dark brown and pedicellus brown; mesoscutum, mesopleuron, and metasoma entirely black or mesoscutum with pair of tiny yellow spots anteriorly and second sternite with pair of medium sized yellow spots; veins 1-SR, 1-M and 1-SR+M of fore wing dark brown. Genitalia, see Fig. 22.

The type series was reared from a nest of Vespula structor (Smith, 1870) (Vespidae: Vespinae).

Named after one of the oldest universities in China, the Northwest University in Xi’an, for providing us the facilities to research the biodiversity of Qinling Mts. It also points to the fact that this is the most north-western locality the genus is known from (“xibei” means “northwest” in Chinese).

Figures 1, 2. 

Bareogonalos xibeidai sp. n. 1 female holotype, habitus lateral 2 male paratype, habitus lateral.

Figures 3–13. 

Bareogonalos xibeidai sp. n., female, holotype. 3 fore wing 4 head and mesosoma dorsal 5 mesosoma lateral 6 metasoma dorsal 7 metasoma ventral 8 detail of second and third sternites lateral 9 head dorsal 10 head anterior 11 head lateral 12 hind leg lateral 13 antenna lateral.

Figures 14–22. 

Bareogonalos xibeidai Tan & van Achterberg, sp. n., male, paratype. 14 fore wing 15 head and mesosoma dorsal 16 mesosoma lateral 17 metasoma lateral 18 antenna lateral 19 head anterior 20 head lateral 21 metasoma dorsal 22 genitalia dorsal.

Unknown. Collected in June–November.

China, Japan.

2 ♂ (NWUX, RMNH): “NW China: Shaanxi, Ningqiang, Hanzhong, Tiankeng, Chanjiyan, N32.46° E106.30°, 25.vi-22.vii.2017, b[lack] Mal. trap, alt. 1638 m, Tan Jiangli, NWUX”.

China (*Shaanxi, Sichuan). New for Shaanxi and second record of species.

Holotype, ♀ (NWUX) “NW China: Shaanxi, Upper Changqing Re[ser]v[e], Shanshuping, 1556 m, 33.67N 107.58E, 25.viii.–22.ix.2016, Y[ellow Malaise] T[rap], Zhao Lin-Peng, NWUX”.

Occipital carina very wide medio-dorsally, with pair of curved lamellae (Fig. 35); outer side of supra-antennal elevations subvertical and elevations about 0.6 times as long as scapus (Fig. 35); frons largely coarsely punctate (Fig. 36); supra-antennal elevations largely ivory (Fig. 35); mandible mainly pale brown, except its ivory base and dark teeth (Fig. 36); metasoma dorsally largely smooth, largely black with narrow pale apical bands (Figs 29, 33); first tergite about 0.7 times as long as its apical width (Fig. 33); third sternite about 0.3 times as long as second sternite (Fig. 34). Close to J. satoi (Tsuneki) from Taiwan, from which is differs mainly because of the coarsely and densely punctate frons (sparsely punctulate in J. satoi), the densely rugose and rather matt scutellum (largely smooth and shiny), clypeus with fine acute tooth medio-ventrally (blunt) and the largely ivory supra-antennal elevations (black).

Holotype, ♀, length of body 8.5 mm (of fore wing 7.4 mm).

Head. Antenna with 25 segments; frons coarsely punctate (except anteriorly), interspaces narrow and smooth (Figs 35, 36), with rather long setae; vertex largely smooth behind posterior ocelli and posteriorly, but medially punctate and with some short rugae and antero-laterally with oblique rugulae (Fig. 35); temple largely smooth, punctulate, but near mandible punctate (Fig. 37); head gradually narrowed behind eyes, eye in dorsal view 1.4 times as long as temple (Fig. 35); occipital carina strongly widened and pair of circular lamellae medio-dorsally, without distinct carinae (Figs 31, 35); supra-antennal elevations strongly enlarged (about 0.6 times as long as scapus), outer side subvertical and with distinct rugae; clypeus concave and thick medio-ventrally and area above it convex and acutely protruding (Fig. 36).

Mesosoma. Length of mesosoma 1.5 times its height (Fig. 32); mesopleuron antero-dorsally densely reticulate-rugose, dorso-posteriorly with some rugosity and remainder smooth and shiny, and antero-ventrally curved rugulose (Fig. 32); notauli wide, deep and distinctly crenulated; middle lobe of mesoscutum transversely rugose and with finer interconnected sculpture, lateral lobes mainly rugose except for a smooth shallow groove (Fig. 31); scutellar sulcus very wide and coarsely crenulated; scutellum densely reticulate-rugose, convex laterally and shallowly depressed medially, in lateral view far above level of metanotum (Fig. 29); metanotum medially evenly convex and finely rugose (Fig. 31); propodeum antero-laterally irregularly rugose and interspaces more or less coriaceous, matt, remainder coarsely transversely rugose and shiny but smooth posteriorly (Fig. 31); posterior propodeal carina thick lamelliform (foramen about twice as wide as high medially).

Wings. Fore wing: length of vein 1-M 1.7 times as long as vein 1-SR; third submarginal cell much wider anteriorly than second cell (Fig. 30).

Metasoma. First tergite 0.7 times as long as its apical width, smooth but basal depression anteriorly striate (Fig. 33); second and following tergites smooth and shiny; sternites rather densely finely punctate, with wide smooth interspaces; second sternite weakly curved in lateral view; third sternite about 0.3 times as long as second sternite (Fig. 34); hypopygium triangularly protruding in ventral view (Fig. 34).

Colour. Black; inner orbita vaguely partly orange brown; malar space and supra-antennal elevations largely ivory; minute patch of outer orbita, vertex posteriorly, occipital carina medially dark brown; apex of scapus and pedicellus, and mandible largely pale brown but basally ivory and teeth dark brown (Figs 36, 37); tegulae and pronotal lobe below it pale yellowish; first metasomal tergite apically broadly ivory and narrower laterally, first and second sternites with large ivory patch apico-laterally; with narrow ivory apical band at apex of sternites and of second and following tergites (Figs 29, 33); palpi rather dark brown; remainder of antenna blackish; fore leg (except black coxa and trochanter) yellowish brown; middle and hind legs mainly dark brown but coxae black (except ivory apex of hind coxa) and hind trochanter ivory with some faint infuscation; pterostigma brownish yellow, but anteriorly (except basally) and apically blackish; basal half of marginal cell and third submarginal cell dark brown and remainder of wing membrane subhyaline (Fig. 30).

Male. Unknown.

Unknown.

China (Shaanxi). Collected at 1556 m.

Named after its conspicuously coloured mandible.

Figures 23–28. 

Bareogonalos jezoensis, female, Japan (Kyushu, Kobayashi). 23 wings 24 habitus lateral 25 mesosoma dorsal 26 metasoma lateral 27 metasoma ventral 28 antenna lateral.

Figure 29. 

Jezonogonalos mandibularis Tan & van Achterberg, sp. n., female, holotype, habitus lateral.

Figures 30–38. 

Jezonogonalos mandibularis Tan & van Achterberg, sp. n., female, holotype. 30 fore wing 31 mesosoma dorsal 32 mesosoma lateral 33 metasoma dorsal 34 metasoma ventral 35 head dorsal 36 head anterior 37 head lateral 38 hind leg lateral.

Holotype, ♀ (NWUX) “NW China: Shaanxi, Lower Changqing Re[ser]v[e], Shanshuping Base, 1504 m, 33.67N 107.57E, 23.ix.–10.xi.2016, Y[ellow Malaise] T[rap], Zhao Lin-Peng, NWUX”. Paratypes: 1 ♀ (RMNH), “NW China: Shaanxi, Liping Nat. For. P., MT 1+2, c. 1495 m, 22.vi.–4.ix.2015, 32°47'33"N, 106°39'52"E, JL. Tan & C. v. Achterberg, NWUX”; 1 ♀ (NWUX), “NW. China: Shaanxi, Xunyangba, Ningshan, c. 1300 m, 24.vi. 2011, 33°33'N, 108°32'E, Jiangli Tan, NWUX”.

Occipital carina very wide medio-dorsally, with pair of curved lamellae separated by carina (Fig. 45); outer side of supra-antennal elevations subvertical, largely smooth, and elevations about 0.8 times as long as scapus (Fig. 45); frons moderately punctate (Fig. 46); supra-antennal elevations largely ivory dorsally (Fig. 45); mandible mainly dark brown, except ivory basal patch and pale brown base of teeth (Fig. 46); metasoma dorsally largely smooth and largely black (Figs 39, 43); first tergite about 0.6 times as long as its apical width (Fig. 43); third sternite about 0.4 times as long as second sternite (Fig. 44). Close to J. satoi (Tsuneki) from Taiwan and J. mandibularis from Shaanxi, from which the new species differs because of the largely smooth supra-antennal elevations (with carinae or long grooves and some punctures in J. satoi and J. mandibularis), the densely rugose and rather matt scutellum (largely smooth and shiny), the scutellum somewhat above level of metanotum in lateral view and medially flattened (protruding far above level of metanotum and medially slightly longitudinally depressed), the mandible largely dark brown (largely pale brown and yellowish or ivory); the inner orbita of eye completely ivory (largely dark brown or with incomplete ivory stripe), and the middle lobe of mesoscutum with pair of ivory patches anteriorly (entirely black).

Holotype, ♀, length of body 8.7 mm (of fore wing 7.1 mm).

Head. Antenna with 25 segments; frons moderately punctate with smooth interspaces wider than punctures (Fig. 46), with medium-sized whitish setae; vertex smooth, but remotely punctulate (Fig. 45); temple largely smooth, punctulate, but near mandible punctate (Fig. 47); head hardly narrowed behind eyes, eye in dorsal view 1.2 times as long as temple (Fig. 45); occipital carina strongly widened and pair of circular lamellae medio-dorsally, separated by distinct carina (Figs 41, 45), laterally distinctly crenulated; supra-antennal elevations distinctly enlarged (about 0.8 times as long as scapus), largely smooth (except some punctures) and outer side subvertical; clypeus concave and thick medio-ventrally and area above it convex and obtusely protruding (Fig. 46).

Mesosoma. Length of mesosoma 1.7 times its height (Fig. 42); mesopleuron antero-dorsally spaced rugose, dorso-posteriorly with some fine striae and remainder smooth and shiny, and antero-ventrally curved rugulose; notauli rather narrow, but deep and coarsely crenulate; middle lobe of mesoscutum transversely rugulose and with some punctures in between, lateral lobes mainly punctate except for a smooth line (Fig. 41); scutellar sulcus wide and coarsely crenulate; scutellum remotely coarsely punctate and laterally with rugae, flattened, in lateral view somewhat above level of metanotum (Fig. 39); metanotum medially evenly convex, shiny and largely smooth (Fig. 41); propodeum antero-laterally irregularly rugose or rugulose and interspaces more or less coriaceous, matt, remainder coarsely transversely rugose and shiny medially, and smooth posteriorly (Fig. 41); posterior propodeal carina thick lamelliform (foramen about 4 times as wide as high medially).

Wings. Fore wing: length of vein 1-M 1.5 times as long as vein 1-SR; third submarginal cell much wider anteriorly than petiolate second cell (Fig. 40).

Metasoma. First tergite 0.6 times as long as its apical width, smooth but basal depression anteriorly with some crenulae (Fig. 43); second and following tergites smooth except for punctulation and shiny; sternites rather densely finely punctate, with wide smooth interspaces; second sternite weakly curved in lateral view; third sternite about 0.4 times as long as second sternite (Fig. 44); hypopygium triangularly protruding in ventral view (Fig. 44).

Colour. Black; inner orbita narrowly ivory and connected to ivory malar space; pair of patches on clypeus, basal patch of mandible, large patch on supra-antennal elevations, pair of elongate patches on middle lobe of mesoscutum anteriorly, pair of narrow lines near tegulae, pair of small patches on metanotum medially, epipleura of first tergite, large patch apico-laterally on second tergite and narrow apical bands of sternites ivory; 12^th–22^nd antennal segments largely pale brown ventrally; mandible largely dark brown but teeth light brown basally (Figs 46); tegulae and trochanters mainly dark brown; palpi rather dark brown; legs blackish or dark brown, but fore femur apico-ventrally and tibia ventrally yellowish; pterostigma basally yellow, and remainder dark brown; basal half of marginal cell and to a lesser degree area below it dark brown and remainder of wing membrane subhyaline (Fig. 40).

Variations. Length of body 8.7–10.2 mm, of fore wing 7.1–8.1 mm; antenna of ♀ with 25(2) or 27(1) segments; metanotum black or with pair of ivory spots medially; ivory patches of clypeus and mesoscutum rather small to large; mesoscutum sometimes with minute ivory spot medio-posteriorly; second submarginal cell of fore wing petiolate or sessile anteriorly; length of vein 1-M of fore wing 1.5–1.7 times as long as vein 1-SR; pterostigma largely yellowish brown or largely dark brown; mandible dark brown or ivory subbasally; whitish setae of mesosoma long or medium-sized.

Male. Unknown.

Unknown.

China (Shaanxi). Collected at 1300–1500 m.

Named after the province Shaanxi, where it was collected.

Reared reared as hyperparasitoid of Tachinidae in caterpillars (Carmean and Kimsey 1998; Murphy et al. 2009). Collected in May–August.

Mainly East Palaearctic and Northeast Oriental regions, with few species in East Afrotropical (including Madagascar), Neotropical and Nearctic regions.

Orthogonalys centrimaculata Bischoff, 1951, from N. Vietnam (Sa Pa, Lao Cai) will run in the key to O. robusta, but its vertex has a medio-posterior pale patch (absent in O. robusta), the mesoscutum is very finely and densely transversely rugulose or coriaceous, matt (mainly finely remotely (rugulose-)punctate and with satin sheen), and hind tibia and tarsus are yellowish brown (dark brown). It runs to O. formosana if only the colouration of the vertex is considered, but it has the mesoscutum is very finely and densely transversely rugulose or coriaceous, matt (coarsely punctate and shiny in O. formosana) and hind tibia yellowish brown (apical two-thirds dark brown).

3 ♀ + 9 ♂ (NWUX, RMNH), “NW China: Shaanxi, Lower Changqing Re[ser]v[e], Shanshuping, 1445 m, N33°67' E107°57', 18.vi.–17.vii.2016, Y[ellow Malaise] T[rap], Zhao Lin-Peng, NWUX”; 4 ♀ + 6 ♂ (NWUX, RMNH), “NW China: Shaanxi, Xunyangba, Ningshaan, 1481 m, N33°54' E108°55', 1.vii.–17.viii.2016, Y[ellow] and G[reen Malaise] Trap, Jiangli Tan/Qingqing Tan, NWUX”; 9 ♀ + 13 ♂ (NWUX, RMNH), id., but Green Malaise trap, 20.v.–23.vi.2016; 1 ♀ + 1 ♂ (NWUX), “NW China: Shaanxi, Pingheliang, Ningshaan, N33°47' E108°50', B[lack] Mal[aise] trap, 1.vii.–17.viii.2016, 2188 m, J-L. Tan & Q-Q. Tan, NWUX”; 1 ♂ (NWUX), “NW China: Shaanxi, Ningshaan, from Huangguan to Xunyangba, 1236 m, 33°54'N, 105°36'E, 1.vii.–17.viii.2016, black Mal[aise] trap, J-L Tan & Q-Q Tan, NWUX”; 1 ♀ (NWUX), “NW China: Shaanxi, Ningqiang, Hanzhong, Tiankeng, Chanjiyan, N32.46° E106.30°, 25.vi-22.vii.2017, b[lack] Mal. trap, alt. 1638 m, Tan Jiangli, NWUX”.

China (Guizhou, Ningxia, Shaanxi, Sichuan, Yunnan). Collected at 1445–1650 m.

The series shows a considerable variation in the shape of the second and third submarginal cells, but the third cell remains wider anteriorly than the second cell. The size difference is also considerable, e.g., the length of the fore wing (3 ♀) is 4.6–8.2 mm.

1 ♀ (NWUX), “NW China: Shaanxi, Ningshaan, from Huangguan to Xunyangba, 1236 m, 33°54'N, 105°36'E, 20.v.–20.vi.2016, black Mal[aise] trap, J-L Tan & Q-Q Tan, NWUX”; 1 ♀ (RMNH), “NW China: Shaanxi, Liping Nat. Forest Park, c. 1490 m, 22.vi.2015, 32°48'N, 106°40'E, Jiangli Tan, NWUX”; 1 ♂ (NWUX), “NW China: Shaanxi, Haopingsi to Dadian, Meixian, Taibai Mt., swept, N34.4, E107.46, 16.vii.2017, alt. 1251 m, Tan Jiangli, NWUX”.

China (Henan, *Shaanxi, Sichuan, Tibet); Russia (Far East); Japan (Hokkaido, Honshu).

2 ♀ (NWUX, RMNH), “NW China: Shaanxi, Liping Nat. For. P., MT 1+2, c. 1495 m, 22.vi.–4.ix.2015, 32°47'33"N 106°39'52"E, J-L. Tan & C. v. Achterberg, NWUX”; 2 ♂ (NWUX, RMNH), “NW China: Shaanxi, Lower Changqing Re[ser]v[e], Shanshuping, 1445 m, N33.67 E107.57, 18.vi.–17.vii.2016 & 24.vii–24.viii.2016, Y[ellow Malaise] T[rap], Zhao Lin-Peng, NWUX”; 1 ♂ (NWUX), id., but Shanshuping Base, 1504 m, 25.viii–22.ix.2016; 1 ♀ (NWUX), “NW China: Shaanxi, Ningshaan, from Huangguan to Xunyangba, 1236 m, B[lack] & W[hite Malaise] trap, 17.viii.–5.x.2016, 33°54'N, 105°36'E, J-L. Tan & Q-Q Tan, NWUX”; 1 ♀ (NWUX), “NW China: Shaanxi, Pingheliang, Ningshaan, N33°47' E108°50', B[lack] Mal[aise] trap, 17.viii.–1.x.2016, 2188 m, J-L. Tan & Q-Q. Tan, NWUX”; 1 ♀ + 2 ♂ (NWUX), “NW China: Shaanxi, Ningqiang, Hanzhong, Tiankeng, Chanjiyan, N32.46° E106.30°, 25.vi-22.vii.2017, b[lack] Mal. trap, alt. 1638 m, Tan Jiangli, NWUX”.

China (*Shaanxi, *Sichuan), Japan (Honshu), ?India. Collected at 1445–1495 m in China.

After examination of fresh pale specimens keying out to Orthogonalys elongata Teranishi, it was obvious (e.g. colour pattern of head) that they belong to a separate species for which the name of O. hirasana Teranishi, 1929, is available. In Chen et al. (2014) the figured female from Sichuan belongs also here considering its colour pattern. Here we illustrate the male of this species (Figs 50–59). The female from Pingheliang has the mesopleuron entirely black.

The interpretation of O. albomaculata Bischoff, 1951, from N. India is provisional because only males are known. The propodeum is coarsely reticulate, the pronotal side crenulate medially, the third submarginal cell of fore wing comparatively large and the propodeum has a large ivory patch.

Figure 39. 

Jezonogonalos shaanxiensis Tan & van Achterberg, sp. n., female, holotype, habitus lateral.

Figures 40–48. 

Jezonogonalos shaanxiensis Tan & van Achterberg, sp. n., female, holotype. 40 fore wing 41 mesosoma dorsal 42 mesosoma lateral 43 metasoma dorsal 44 metasoma ventral 45 head dorsal 46 head anterior 47 head lateral 48 hind leg lateral.

Figure 49. 

Orthogonalys hirasana Teranishi, male, Shaanxi (Shanshuping), habitus lateral.

Holotype, ♀ (NWUX), “NW China: Shaanxi, Lower Changqing Re[ser]v[e], Shanshuping, 1445 m, 33.67N 107.57E, 18.vi.–17.vii.2016, Y[ellow Malaise] T[rap], Zhao Lin-Peng, NWUX”. Paratypes: 1 ♂ (NWUX), same data as holotype; 1 ♀ + 8 ♂ (NWUX, RMNH), “NW China: Shaanxi, Xunyangba, Ningshaan, 1481 m, N33°54' E108°55', 1.vii.–17.viii.2016, Y[ellow] and G[reen Malaise] Trap, Jiangli Tan/Qingqing Tan, NWUX”; 2 ♀ + 23 ♂ (NWUX, RMNH), id., but Green Malaise trap (except 1 ♀ from black Malaise trap), 20.v.–23.vi.2016; 3 ♂ (NWUX), “NW China: Shaanxi, Ningshaan, from Huangguan to Xunyangba, 1236 m, 33°54'N, 105°36'E, 1.vii.–17.viii.2016, black Mal[aise] trap, J-L Tan & Q-Q Tan, NWUX”; 4 ♂ (NWUX), id., but 20.v.–20.vi.2016; 2 ♂ (NWUX), “NW China: Shaanxi, Huanghualing, Zhashui, 1408 m, 20.v.–1.vii.2016, 33°80'N, 108°88'E, yellow [Malaise] trap, J-L Tan & Q-Q Tan, NWUX”.

Antenna without subapical ivory band (Figs 60, 61); occipital carina rather narrow and sparsely crenulate dorsally (Fig. 67); frons moderately shiny and largely coarsely punctate (Fig. 68); area behind stemmaticum with some rugae and posteriorly aciculate (Fig. 67); supra-antennal elevations medium-sized and coarsely punctate (Fig. 67); clypeus strongly convex medially, coarsely punctate and medio-ventrally slightly concave (Fig. 68); basal half of mandible ivory; mesoscutum coriaceous-rugulose and anterior pair of smooth stripes inconspicuous (Fig. 64); scutellum coarsely punctate, longitudinally depressed medially and laterally with some longitudinal rugae, and moderately shiny (Fig. 64); mesosoma without pale pattern or spots dorsally (Fig. 64); third submarginal cell 0.4–0.5 times as long as second submarginal cell, but anteriorly much wider than second cell (Figs 61, 62); pterostigma of both sexes dark brown; anterior 0.8–0.9 of first metasomal tergite and anterior half of second tergite black (Fig. 66).

The new species runs in the key to Orthogonalys by Chen et al. (2014) to O. clypeata Chen, van Achterberg, He & Xu, 2014, and differs as follows: mesoscutum coriaceous-rugulose and anterior pair of smooth stripes inconspicuous (densely to remotely punctate and anterior pair of smooth stripes rather conspicuous in O. clypeata), area behind stemmaticum with some fine rugae and posteriorly aciculate (smooth), frons coarsely punctate (at most moderately punctate and often largely smooth), and pterostigma of ♀ dark brown (largely brownish yellow).

Holotype, ♀, length of body 9.2 mm (of fore wing 7.6 mm).

Head. Antenna with 22 segments; frons coarsely punctate; vertex largely smooth laterally, but behind stemmaticum with some fine rugae and posteriorly transversely aciculate (Fig. 67); temple smooth (Fig. 63); head gradually narrowed behind eyes, eye in dorsal view 1.3 times as long as temple (Fig. 67); occipital carina rather narrow and sparsely crenulate dorsally (Fig. 67); supra-antennal elevations medium-sized (about half as long as scapus), outer side subvertical and coarsely densely rugose (Fig. 67); clypeus slightly concave medio-ventrally and strongly convex medially (Fig. 68).

Mesosoma. Length of mesosoma 1.5 times its height; mesopleuron below transverse mesopleural groove with some coarse rugae, above groove largely smooth (Fig. 65); transverse mesopleural groove wide, deep and coarsely crenulate; notauli wide, deep and coarsely crenulate; mesoscutum coriaceous-rugulose and anterior pair of smooth stripes of middle lobe inconspicuous, lateral lobe rugulose and with fine punctures (Fig. 64); scutellar sulcus wide and coarsely crenulate; scutellum coarsely punctate, medially shallowly longitudinally depressed and laterally with some longitudinal rugae (Fig. 64); metanotum medially distinctly convex, smooth and shiny but anteriorly rugulose (Fig. 64); propodeum shiny and irregularly spaced rugose (Fig. 64); posterior propodeal carina thick lamelliform and hardly arched, foramen medially 0.3 times higher than wide basally.

Wings. Fore wing: length of vein 1-M 2.2 times as long as vein 1-SR (Fig. 62); second submarginal cell twice as long as third cell.

Metasoma. First tergite 0.8 times as long as its apical width, smooth and with pair of small depressions medially (Fig. 66); metasoma smooth, but first sternite partly superficially coriaceous (Fig. 69); third sternite about 0.7 times as long as second sternite (Fig. 69); hypopygium triangular (Figs 60, 69).

Colour. Black; palpi and tegulae pale yellow; inner orbita (except dorsally) ivory and connected to broadly ivory malar space; basal half of mandible ivory, apical half pale brown, but teeth dark brown; apical quarter of antenna brownish ventrally; first tergite latero-posteriorly, apical half of second tergite (and medio-anteriorly protruding into black area), pair of large triangular spots on third tergite latero-posteriorly, first sternite laterally, second sternite and fourth sternite laterally and medially yellow (Fig. 69); hind trochanter and trochantellus white; coxae and hind femur black; fore and middle trochanters, base of femora, hind tibia and tarsus dark brown; remainder of legs yellowish brown; pterostigma dark brown; wing membrane subhyaline.

Variations. Length of body 7.1–9.2 mm, of fore wing 5.8–7.6 mm; antenna with 22(1) or 23(1) segments.

Male. Length of body 5.6–10.8 mm, of fore wing 5.9–8.6 mm; antenna with 21(2), 22(15), 23(12), 24(2) segments, apical quarter of antenna brownish ventrally or most of antenna brown; frons densely and coarsely punctate-rugose; clypeus usually entirely black, but sometimes partly or entirely ivory (as in O. clypeata); mesoscutum often less distinctly transversely rugose than in females; metasoma darker than of female, dorsally largely black (only apical margin of tergites brownish) but first sternite laterally, second sternite laterally and medio-posteriorly (or brownish yellow with pair of elongate dark patches) and third sternite partly or entirely brownish yellow, but sometimes entirely dark brown; paramere (Figs 61, 75) smaller than of O. clypeata.

Unknown.

China (Shaanxi).

Named “paraclypeata” because it is similar to O. clypeata and “para” is Greek for “near”.

4 ♀ (NWUX, RMNH), “NW China: Shaanxi, Xunyangba, Ningshaan, 1481 m, N33°54' E108°55', 1.vii.–17.viii.2016, Y[ellow] and G [Malaise] Trap, Jiangli Tan, NWUX”; 1 ♂ (NWUX), “NW China: Shaanxi, Huanghualing, Zhashui, 1408 m, 20.v.–1.vii.2016, 33°80'N, 108°88'E, J-L Tan & Q-Q Tan, NWUX”.

China (Guangxi, Shaanxi). Collected at 1480–1760 m.

Reared as hyperparasitoid of Ichneumonidae in caterpillars and from Diprionidae (Carmean and Kimsey 1998). Collected in April–August.

Palaearctic region, but two species (P. harmandi Schulz, 1907 and P. hahnii (Spinola)) occur in the North Oriental region.

2 ♀ (NWUX, RMNH), “NW. China: Shaanxi, Baolongyu, Mt. Qin, c. 1000 m, 10.vi.2015, 34°03'N, 108°09'E, Jiangli Tan, NWUX”; 1 ♀ (NWUX), “China: Gansu, Xiama, Tianzhu, 16–19.vii.2014, Jiangli Tan”.

China (Beijing, Hebei, Henan, Inner Mongolia, Liaoning, *Shaanxi, Yunnan); Russia; Ukraine; Kazakhstan; Mongolia; Western Europe (Lelej 2003). Collected at 0–1000 m.

Large conspicuous black species with large dark patch near pterostigma of fore wing.

Figures 50–59. 

Orthogonalys hirasana Teranishi, male, Shaanxi (Shanshuping). 50 fore wing 51 mesosoma dorsal 52 mesosoma lateral 53 metasoma dorsal 54 metasoma ventral 55 apical two-thirds of antenna lateral 56 head dorsal 57 head anterior 58 head lateral 59 hind leg lateral.

Figures 60, 61. 

Orthogonalys paraclypeata Tan & van Achterberg, sp. n. 60 female, holotype, habitus lateral 61 male, paratype, habitus lateral.

Figures 62–70. 

Orthogonalys paraclypeata Tan & van Achterberg, sp. n., female, holotype. 62 fore wing 63 head lateral 64 mesosoma dorsal 65 mesosoma lateral 66 metasoma dorsal 67 head dorsal 68 head anterior 69 metasoma ventral 70 hind leg lateral.

Figures 71–79. 

Orthogonalys paraclypeata Tan & van Achterberg, sp. n., male, paratype. 71 mesosoma dorsal 72 mesosoma lateral 73 hind leg lateral 74 metasoma dorsal 75 metasoma ventral 76 head dorsal 77 head anterior 78 head lateral 79 apical two-thirds of antenna lateral.

Reared as hyperparasitoid of parasitoid wasps (Ichneumonidae and Braconidae) and parasitoid flies (Tachinidae) in caterpillars, but some species are primary parasitoids of Pergid sawflies in Australia (Raff 1934; Carne 1969; He and Chen 1986; Weinstein and Austin 1995; Carmean and Kimsey 1998). Collected mainly in April–October, rarely in November or January.

This genus occurs in all major regions, but is unknown from Europe and western Nearctic region. Most of the species occur in the East Palaearctic, Northeast Oriental, and Neotropical regions (Carmean and Kimsey 1998).

Revised part of key to Chinese species of the genus Taeniogonalos Schulz

(for first part, see Chen et al. 2014)

4 ♀ (NWUX), “NW China: Shaanxi, Lower Changqing Re[ser]v[e], Shanshuping, 1445 m, N33.67 E107.57, 18.vi.–17.vii.2016, Y[ellow Malaise] T[rap], Zhao Lin-Peng, NWUX”; 4 ♀ + 8 ♂ (NWUX, RMNH), “NW. China: Shaanxi, Xunyangba, Ningshaan, 1481 m, Y[ellow] and G[reen Malaise] trap, 1.vii.–17.viiii.2016, 33°33'N, 108°32'E, Jiangli Tan/Qingqing Tan, NWUX”; 3 ♂ (NWUX), id., but Green Malaise trap, 20.v.–23.vi.2016; 1 ♀ (NWUX), “NW China: Ningxia, Liupan Mt., Jingyuan, Dongshanpo For[est] Farm, N35°23'26” E106°20'34.27”, 4.viii.2015, c 1800 m, Jiangli Tan, NWUX”; 1 ♀ (NWUX), “NW. China: Shaanxi, Panda valley, Foping, 1411 m, black Mal[aise] trap, 1.vii–18.viii.2016, 33°67'N, 107°97'E, Jiangli Tan, NWUX”; 1 ♂ (NWUX), “NW China: Shaanxi, Huanghualing, Zhashui, 1408 m, 20.v.–1.vii.2016, 33°80'N, 108°88'E, yellow [Malaise] trap, J-L Tan & Q-Q Tan, NWUX”; 1 ♀ (NWUX), “NW China: Shaanxi, Ningshaan, from Huangguan to Xunyangba, 1236 m, 33°54'N, 105°36'E, 1.vii.–17.viii.2016, black Mal[aise] trap, J-L Tan & Q-Q Tan, NWUX”; 1 ♀ (NWUX), “NW China: Shaanxi, Liping Nat. For. P., MT 1+2, c. 1495 m, 22.vi.–4.ix.2015, 32°47'33"N 106°39'52"E, JL. Tan & C. v. Achterberg, NWUX”; 1 ♀ + 2 ♂ (NWUX): “NW China: Shaanxi, Ningqiang, Hanzhong, Tiankeng, Chanjiyan, N32.46° E106.30°, 25.vi-22.vii.2017, b[lack] Mal. trap, alt. 1638 m, Tan Jiangli, NWUX”. First report of female and new for continental China.

Illustrated ♀ from Liupan Mt., length of body 5.6 mm (of fore wing 5.1 mm).

Head. Antenna with 20 segments; frons coarsely punctate, with smooth interspaces narrower than punctures and with some rugae anteriorly (Fig. 88); vertex largely smooth and strongly shiny posteriorly, but spaced moderately punctate anteriorly (Fig. 87); temple smooth except some punctures near eye and mandible (Fig. 89); head gradually narrowed behind eyes, eye in dorsal view 1.1 times as long as temple (Fig. 87); occipital carina narrow, non-lamelliform and smooth dorsally (Fig. 87); supra-antennal elevations medium-sized (about half as long as scapus), outer side subvertical anteriorly and with few rather small punctures and apically with few striae (Fig. 87); clypeus slightly concave medio-ventrally and with blunt tubercle above it.

Mesosoma. Length of mesosoma 1.5 times its height; propleuron largely rugose; mesopleuron below transverse mesopleural groove rugulose, above groove densely rugose but posteriorly largely smooth (Fig. 84); transverse mesopleural groove wide, deep and coarsely crenulated; mesosternum mainly transversely aciculate; notauli rather wide, deep and coarsely crenulated; middle lobe of mesoscutum transversely rugulose, but posteriorly with some rugae and anterior pair of smooth stripes rather conspicuous, lateral lobe with coarse rugae but laterally largely smooth or superficially rugulose (Fig. 83); scutellar sulcus rather wide and coarsely crenulated; scutellum reticulate-rugulose and medially slightly longitudinally depressed laterally (Fig. 83); metanotum rugose and medially flattened, but submedially convex (Fig. 83); propodeum slender, lateral sides nearly straight, its surface shiny, finely rugose anteriorly and superficially rugulose posteriorly (Fig. 83); posterior propodeal carina narrow lamelliform medially and strongly arched, foramen medially 0.4 times higher than wide basally.

Wings. Fore wing: length of vein 1-M 1.7 times as long as vein 1-SR (Fig. 82); second submarginal cell 1.3 times as long as third cell.

Metasoma. First tergite 0.6 times as long as its apical width, smooth and medially largely depressed (Fig. 85); metasoma smooth and strongly shiny, third–fifth tergites superficially punctulate (Fig. 86); second sternite strongly shiny and convex; second–fourth sternites superficially spaced punctulate or finely punctate; third sternite slightly concave and about 0.3 times as long as second sternite (Fig. 86).

Colour. Black; palpi dark brown basally and pale brown apically; antenna pale brown, but scapus and apical 0.4 of antenna darkened; inner orbita slightly brownish near level of antennal sockets; mandible mainly dark brown, but medially pale brown; sixth tergite yellowish ivory; robust hind trochanter and trochantellus mainly white except some slightly brownish small patches; fore and middle tibiae and tarsi brownish yellow, but middle tibia dark brown posteriorly; remainder of legs, tegulae and pterostigma dark brown; wing membrane subhyaline except for infuscated patch near apex of fore wing (Fig. 82).

The holotype of T. alishana fits better with T. alticola than with T. taihorina considering its sculpture. The male holotype has been collected at the same day and locality as the male holotype of T. alticola.

China (*Ningxia, *Shaanxi, Taiwan). Collected at 1236–1800 m.

2 ♂ (NWUX, RMNH), “NW. China: Shaanxi, Xunyangba, Ningshan, c. 1300 m, 2.vi. & 30.ix.2014, 33°33'N, 108°32'E, Jiangli Tan, NWUX”; 4 ♀ + 16 ♂ (NWUX, RMNH), id., but 1481 m, yellow or black Malaise trap, 1.vii.–17.viiii.2016; 9 ♂ (NWUX), id., but 20.v.–23.vi.2016, green Malaise trap, J-L. Tan & Q-Q. Tan; 1 ♀ (NWUX), id., but black trap; 1 ♀ (NWUX), id., but yellow/green Malaise trap, 17.viii.–3.x.2016; 2 ♂ (NWUX), “NW. China: Shaanxi, Qinling Mts, Foping, near Biol. Stat., c. 1400 m, Mal. trap, 29.v.–19.vi.2016, 33°40'N, 107°58'E, J-L. Tan & C. v. Achterberg, NWUX”; 1 ♂ (NWUX), “NW China: Shaanxi, Liping Nat. For. P., MT 1+2, c. 1495 m, 22.vi–4.ix.2015, 32°47'33"N 106°39'52"E, JL. Tan & C. v. Achterberg, NWUX”; 2 ♀ (NWUX, RMNH), “NW China: Shaanxi, Pingheliang, Ningshaan, N33°47' E108°50', B[lack] Mal[aise] trap, 17.viii.–1.x.2016, 2188 m, J-L. Tan & Q-Q. Tan, NWUX”; 4 ♂ (NWUX), “NW China: Shaanxi, Huanghualing, Zhashui, 1408 m, 20.v.–1.vii.2016, 33°80'N, 108°88'E, yellow [Malaise] trap, J-L Tan & Q-Q Tan, NWUX”; 1 ♀ + 1 ♂ (RMNH), “NW China: Shaanxi, Liangfengya, Foping, 1729 m, 19.viii.–13.xi.2016, 33°69'N, 107°90'E, y[ellow] Mal[aise] trap, J-L Tan & Q-Q Tan, NWUX”; 4 ♂ (NWUX), “NW China: Shaanxi, Ningshaan, from Huangguan to Xunyangba, 1236 m, 33°54'N, 105°36'E, 1.vii.–17.viii.2016, black Mal[aise] trap, J-L Tan & Q-Q Tan, NWUX”; 1 ♀ (NWUX), id., but 20.v.–20.vi.2016; 1 ♀ (NWUX), “NW China: Shaanxi, Upper Changqing Re[ser]v[e], Shanshuping, 1556 m, N33.67 E107.58, 24.vii–24.viii.2016, Y[ellow Malaise] T[rap], Zhao Lin-Peng, NWUX”; 2 ♂ (NWUX), “NW. China: Shaanxi, Qinling Mts, Luoyuan, c. 1350 m, 16.iv.–28.v.2016, 34°12'N, 109°50'E, J-L Tan & C. v. Achterberg”; 1 ♀ (NWUX), “NW China: Ningxia, Liupan Mt., Jingyuan, Erlonghe For[est] Farm, N35°23'24.14" E106°20'41.43", 2.viii.2015, c 1800 m, Jiangli Tan, NWUX”; 3 ♀ + 6 ♂ (NWUX): “NW China: Shaanxi, Ningqiang, Hanzhong, Tiankeng, Chanjiyan, N32.46° E106.30°, 25.vi-22.vii.2017, b[lack] Mal. trap, alt. 1638 m, Tan Jiangli, NWUX”.

China (Fujian, Gansu, Henan, Ningxia, Shaanxi, Sichuan, Yunnan, Zhejiang). Collected at 1200–2350 m.

1 ♂ (NWUX), “NW China: Shaanxi, Ningshaan, from Huangguan to Xunyangba, 1236 m, 33°54'N, 105°36'E, 20.v.–20.vi.2016, black Mal[aise] trap, J-L Tan & Q-Q Tan, NWUX”

China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hunan, Jilin, Liaoning, Shaanxi, Taiwan, Zhejiang); Russia (Primorskii krai); Japan; Korea. Collected at 990–1550 m. Also reported from Iran, Malaysia, and Indonesia but these reports need confirmation.

1 ♀ + 4 ♂ (NWUX, RMNH), “NW. China: Shaanxi, Xunyangba, Ningshan, 1481 m, Y[ellow] and G[reen Malaise] trap, 1.vii–17.viiii.2016, 33°33'N, 108°32'E, Jiangli Tan/Qingqing Tan, NWUX”; 3 ♂ (NWUX, RMNH), id., but Green Malaise trap, 20.v.–23.vi.2016; 2 ♂ (NWUX), “NW. China: Shaanxi, Panda valley, Foping, 1411 m, black Mal[aise] trap, 1.vii–18.viii.2016, 33°67'N, 107°97'E, Qingqing Tan, NWUX”; 74 ♂ (NWUX, RMNH), “NW China: Shaanxi, Ningqiang, Hanzhong, Tiankeng, Chanjiyan, N32.46° E106.30°, 25.vi-22.vii.2017, b[lack] Mal. trap, alt. 1638 m, Tan Jiangli, NWUX”.

The specimens from Shaanxi belong to the form intermedia (Chen, 1949) because the hind trochanters (as often fore trochanters) are entirely yellow and with tri-coloured (yellow, black and brown) second tergite and sternite latero-posteriorly. Colour of both mesosoma and head are very variable, e.g. the scutellum is black in most specimens but sometimes it has two yellow patches.

China (Fujian, Guangdong, Guizhou, Henan, Jilin, Ningxia, *Shaanxi, Shanxi, Sichuan, Taiwan, Tibet, Yunnan, Zhejiang); Russia (Far East); Japan. Collected at 800–1638 m.

5 ♀ (NWUX, RMNH), “NW China: Shaanxi, Lower Changqing Re[ser]v[e], Shanshuping, 1445 m, N33.67 E107.57, 18.vi.–17.vii.2016, Y[ellow Malaise] T[rap], Zhao Lin-Peng, NWUX”; 1 ♀ (NWUX), id., but Shanshuping Base, 1504 m, 23.ix.–10.xi.2016; 1 ♀ (NWUX), “NW China: Shaanxi, Liping Nat. For. P., MT 1+2, c. 1495 m, 22.vi.–4.ix.2015, 32°47'33"N 106°39'52"E, JL. Tan & C. v. Achterberg, NWUX”; 6 ♀ (NWUX, RMNH), “NW. China: Shaanxi, Xunyangba, Ningshan, 1481 m, Y[ellow] and G[reen Malaise] trap, 1.vii.–17.viiii.2016, 33°33'N, 108°32'E, Jiangli Tan/Qingqing Tan, NWUX”; 2 ♀ + 4 ♂ (NWUX), id., but 17.viii.–3.x.2016; 2 ♀ + 2 ♂ (NWUX), id., but 20.v.–23.vi.2016, Green trap; 1 ♂ (NWUX), id., but Black Malaise trap; 3 ♀ (NWUX), “NW China: Shaanxi, Ningshaan, from Huangguan to Xunyangba, 1236 m, 33°54'N, 105°36'E, 1.vii.–17.viii.2016, black Mal[aise] trap, J-L Tan & Q-Q Tan, NWUX”; 1 ♀ (NWUX), “NW. China: Shaanxi, Panda valley, Foping, 1411 m, black Mal[aise] trap, 1.vii.–18.viii.2016, 33°67'N, 107°97'E, Jiangli Tan, NWUX”; 1 ♀ + 1 ♂ (NWUX): “NW China: Shaanxi, Ningqiang, Hanzhong, Tiankeng, Chanjiyan, N32.46° E106.30°, 25.vi-22.vii.2017, b[lack] Mal. trap, alt. 1638 m, Tan Jiangli, NWUX”.

In the key to Chinese Taeniogonalos spp. (Chen et al. 2014) T. maga (Teranishi) is retained as valid species because of the more lamelliform occipital carina. The occipital carina of the holotype of T. taihorina (Bischoff) is narrow medio-dorsally (similar to latero-dorsally) and not lamelliform. After examining the series from Shaanxi it is obvious that the width of the occipital carina varies from dorsally as wide as laterally and hardly or not lamelliform to about twice as wide as laterally and distinctly lamelliform. We failed to find additional characters to separate both taxa and, therefore, we synonymize both species.

China (Fujian, Gansu, Guangxi, Heilongjiang, Hubei, Ningxia, *Shaanxi, Sichuan, Taiwan, Tibet, Yunnan, Zhejiang); Russia (Far East); Japan (Hokkaido, Honshu). Collected at 500–2530 m.

2 ♂ (NWUX, RMNH), “NW. China: Shaanxi, Panda valley, Foping, 1411 m, black Mal[aise] trap, 1.vii–18.viii.2016, 33°67'N, 107°97'E, Jiangli Tan, NWUX”; 1 ♂ (NWUX): “NW China: Shaanxi, Ningqiang, Hanzhong, Tiankeng, Chanjiyan, N32.46° E106.30°, 25.vi-22.vii.2017, b[lack] Mal. trap, alt. 1638 m, Tan Jiangli, NWUX”.

China (Henan, Fujian, Guangxi, Guizhou, Hainan, Hubei, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); Korea; Laos; Thailand. Collected at 900–2000 m.

Unknown. Collected in June–September.

China, Japan.

3 ♂ (NWUX, RMNH), “NW China: Ningxia, Liupan Mt., Jingyuan, Erlonghe For[est] Farm, N35°23'24.14” E106°20'41.43”, Mal[aise] tr[ap], 2–5.viii.2015, c 1800 m, Jiangli Tan, NWUX”.

China (Gansu, Ningxia, Sichuan). Collected at 1800–2539 m.

2 ♀ (NWUX, RMNH), “NW China: Shaanxi, Pingheliang, Ningshaan, N33°47' E108°50', B[lack] Mal[aise] trap, 17.viii.–1.x.2016, 2188 m, J-L. Tan & Q-Q. Tan, NWUX”.

China (Gansu, Ningxia, *Shaanxi, Sichuan). Collected at 1400–2188 m.

Figures 80, 81. 

Taeniogonalos alticola (Tsuneki), female, Ningxia (Liupan Mt.). 80 habitus lateral 81 metasomal lateral.

Figures 82–91. 

Taeniogonalos alticola (Tsuneki), female, Ningxia (Liupan Mt.). 82 fore wing 83 mesosoma dorsal 84 mesosoma lateral 85 metasoma dorsal 86 metasoma ventral 87 head dorsal 88 head anterior 89 head lateral 90 antenna lateral 91 hind leg lateral.