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Research Article
Taxonomic notes on the collection of the tribe Lamiini (Coleoptera, Cerambycidae) housed in the Natural History Museum, London
expand article infoGuanglin Xie‡§, Maxwell V. L. Barclay‡, Wenkai Wang§
‡ Natural History Museum, London, United Kingdom
§ Yangtze University, Jingzhou, China

Corresponding author: Maxwell V. L. Barclay ( m.barclay@nhm.ac.uk )

Corresponding author: Wenkai Wang ( w_wenkai@hotmail.com )

Academic editor: Francesco Vitali
© 2024 Guanglin Xie, Maxwell V. L. Barclay, Wenkai Wang.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation: Xie G, Barclay MVL, Wang W (2024) Taxonomic notes on the collection of the tribe Lamiini (Coleoptera, Cerambycidae) housed in the Natural History Museum, London. ZooKeys 1205: 239-251. https://doi.org/10.3897/zookeys.1205.127164
ZooBank: urn:lsid:zoobank.org:pub:BFCBF680-68CB-4E0A-A009-23256BE5DDD0
Open Access

Abstract

In the present paper, the genus Mimomyagrus Breuning, 1970 is synonymized with Combe Thomson, 1864, and Mimomyagrus pfanneri Breuning, 1970 is considered a junior synonym of Combe brianus (White, 1858). The female of Arctolamia sinica Bi & Chen, 2022 is described for the first time and this species is reported as new to Myanmar. Type material of Lamia punctator Fabricius, 1776 [= Anoplophora chinensis (Forster, 1771)], Cerambyx galloprovincialis Olivier, 1800 [= Monochamus galloprovincialis (Olivier, 1800)] and Melanauster granulipennis Breuning, 1938 [= Monochamus guerryi Pic, 1903] are confirmed to be preserved in Natural History Museum, London.

Key words

Lamiinae, longhorned beetles, new synonym, taxonomy, type material, zoological collections

Introduction

As one of the largest natural history museums, the Natural History Museum, London [formerly British Museum, Natural History (BMNH), and hereafter NHMUK] houses abundant specimens of longhorn beetles from all over the world.

The first author had an opportunity to visit the NHMUK in 2022, and carried out research related to Cerambycidae for one year under the direction of the second author, Maxwell V. L. Barclay, the Senior Curator in Charge of Coleoptera at the NHMUK; the current paper is part of the research findings, together with four other separately published papers.

On the basis of the study of specimens of the tribe Lamiini preserved in the NHMUK, we propose a genus-level and a species-level new synonymy and confirm that the type material of three species, considered to be ‘lost’ and marked as ‘depositary not known’ (Tavakilian and Chevillotte 2023), are in fact deposited in the NHMUK.

Material and methods

The material examined in this study is deposited in the NHMUK, and has already been identified by earlier specialists, except for the holotypes of Combe fulgurata Thomson, 1864, Mimomyagrus pfanneri Breuning, 1970 and Combe brianus m. rufoantennatus Breuning, 1954 used for comparison in this study, which are deposited in the Muséum national d’Histoire naturelle, Paris, Muséum cantonal des sciences naturelles, Lausanne and Royal Belgian Institute of natural sciences, Brussels, respectively.

The label text, which is reproduced verbatim without corrections or additions, is given in single quotation marks for all studied specimens. Individual labels are separated by a semicolon, and data on different rows by a single slash. Additional and explanatory comments by the authors are given in square brackets.

Photographs were taken using a Canon 7D Mark II DSLR camera with a Canon EFS 100 mm lens and edited using Adobe Photoshop 2020 release. Extended depth of field at magnifications was achieved by combining multiple images from a range of focal planes using Combine ZP or Helicon Focus software.

Results

Based on our study on the tribe Lamiini preserved in the NHMUK, one genus-level synonym and one species-level synonym have been found, the female of one species is newly described with a new distribution record, and the type material of three species previously thought to be lost, has been confirmed to be preserved in the NHMUK. The details are as follows:

Combe Thomson, 1864

Combe Thomson, 1864: 83; Pascoe 1866: 252; Lacordaire 1869: 344; Gemminger and Harold 1873: 3028; Aurivillius 1921: 119; Breuning 1961: 347. Type species:Combe fulgurata Thomson, 1864 [= Combe brianus (White, 1858)].

Mimomyagrus Breuning, 1970: 88. Type species: Mimomyagrus pfanneri Breuning, 1970. Syn. nov.

Combe brianus (White, 1858)

Figs 1, 2, 3

Monohammus brianus White, 1858: 409. Type locality: Nepal.

Combe Fulgurata [sic] Thomson, 1864: 83. Type locality: unspecified.

Combe Brianus [sic]: Pascoe 1866: 244; Lacordaire 1869: 344; Gemminger and Harold 1873: 3028; Aurivillius 1921: 119.

Combe brianus: Breuning 1961: 347; Weigel 2006: 502; Barševskis 2018: 288, 2020: 178.

Combe brianus m. rufoantennatus Breuning, 1954: 7; 1961: 347. Type locality: Sumatra, Indonesia. Unavailable name.

Mimomyagrus pfanneri Breuning, 1970: 88. Type locality: Cameron highlands, Malaysia. Syn. nov.

Type material examined

Holotype of Monohammus brianus White, 1858, female: ‘Type [p, label circular, red frame]; Brianus / n. o [h]; Mon. Brianus White / Nepal [h]; NHMUK013460997 [p]’; holotype of Combe fulgurata Thomson, 1864, female: ‘HOLOTYPE [p, label rectangular, red]; Th. / TYPE [p, black frame]; Ex. Musaeo / JAMES THOMSON [p, black frame]; Ch. J. Gahan / vidit 1895. [p]; Combe / Brianus White / comp. with type / C. J. G. [h]; Brianus / White / Fulguratus / Type Thoms. / Malas. [h, label rectangular, red frame]; Muséum Paris / 1952 / coll. R. Oberthür [p]; HOLOTYPE / Combe / fulgurata Thomson, 1864 [p]; MNHN, Paris / EC23123 [p]’; holotype of Combe brianus m. rufoantennatus Breuning, 1954, female: ‘Holotype [p, label rectangular, red, black frame]; Solok / Sumatra / EX COLL. F. SCHNEIDER [p, black frame]; Coll. R. I. Sc. N. B. / Sumatra [p]; S. Breuning det., 195 [p] 4 [h] / Combe / brianus / rufoantennatus / mihi Typ [h]; L: Bull. Inst. r. sci. Nat. Belg. 1954, 30, 11: F. [h]’; holotype of Mimomyagrus pfanneri Breuning, 1970, female: ‘TYPE [p,label rectangular, red]; Rég. Orientale / Malaisie [h] / Chassot [p] 7. 1968 [h]; GBIFCH / 00338292 [p]’; paratype of Mimomyagrus pfanneri Breuning, 1970, female: ‘Para- / type [p, label circular, yellow frame]; PARATYPE [p, label rectangular, red]; Brit. Mus. / 19 [p] 77–313 [h]; Malaysia [p] / Cameron / h’lds iv/74 [h] / Coll. Pfanner [p]; Mimomyagrus pfanneri Breuning [h]’.

Non-type material examined

Thirteen specimens (5 males, 8 females) identified as ‘Combe brianus’ in NHMUK: Malaysia: 1 male: ‘MALAY PENIN: / Selangor. / Bukit Kutu / 3500 [h] ft. / 14 . 3. [h] 1931 [1 with handwriting] / H. M. Pendlebury. [p]; Ex Coll: / F. M. S. / Museum. [p, reverse side]; Ex F. M. S. Museum. B. M. 1955–354 [p, reverse side]] / NHMUK013461006 [p]’; 1 male: ‘Mal. / P. [h]; Pascoe / Coll. / 93–60 [p]; Combe / brianus / White [h, reverse side]; NHMUK013460999 [p]’; 1 male: ‘Penang [h, label circle]; Bowring. / 63. 47* [p] / Combe / brianus, White [h]; NHMUK013461002’; 1 male: ‘MALAY PENIN. / Kedah Perak. / 1000–2000 [h] ft. / 19 # March, 1928. [p]; Ex F. M. S. / Museum. / B. M. 1955–354 [p]; NHMUK 013461004 [p]’; 1 male: ‘MALAY PENIN [p] / Perak, F. M. S. / Maxwell Hill 3000 / June–July 1916 [h]; 332 [h]; Ex F. M. S. / Museum. / B. M. 1955–354. [p]; NHMUK013387052 [p]’; 1 female: ‘Rantau Panjang / Selangor. / H. C. Robinson. [p] / (12. 16 / V / 04) [h] / 1904–327.[p]; (4 / 1107)[p]; NHMUK013387035 [p]’; 1 female: ‘Malay / Penang [p]; Fry Coll. / 1905. 100. [p]; 17783 [h]; NHMUK013460998 [p]’; 1 female: ‘MALAY PENIN: [p] / Penang F. M. S. / Maxwell Hill 3000 / June–July 1916 [h]; 331 [h]; Ex F. M. S. / Museum. / B. M. 1955–354. [p]; Combe / Brianus, White / 2 [h]; NHMUK013387050 [p]’; 1 female: ‘Malay Penin / Panang, F. M. S. [p] j, / Gap / 2900 / May 1915 [h] 193 [p]2 [h] ; Ex F. M. S. / Museum. / B. M. 1955–354 [p]; NHMUK 013461000 [p]’; 1 female: ‘Malay / Penang [p]; Data unreliable / See Brit. Mus. / 1949–314. [p, label yellow]; Combe / brianus [h]; NHMUK013461003 [p]’; 1 female: ‘Malay / Penang / Batu Feringgi / catchment area [p] / 25 Aug 1963 [h] / H. T. Pagden; Pres by / Com Inst Ent / B. M. 1964-2 [p]; 119 [p]; Combe / brianus White [h] / E. A. J. Duffy det. 1964 [p]; NHMUK013461001 [p]’; 1 female: ‘MALAYA / Kuala Lumpur [p]/ Ampang Village [h] / Feb. 26th[h]19[p]35[h]; Ex F. M. S. / Museum. / B. M. 1955–354 [p]; NHMUK 013387047 [p]’;1 female: ‘MALAY PENIN: / Selangor. / Bukit Kutu / 3300–3500 [h] ft. / 14 . 3. [h] 1931. / H. M. Pendlebury. [p]; Ex Coll: / F. M. S. / Museum. [p, reverse side]; Ex F. M. S. Museum. M. 1955–354 [p, reverse side]; NHMUK013461005 [p]’.

Two specimens (1 male, 1 female) identified as ‘Mimomyagrus pfanneri Breuning, 1970’ in NHMUK: Malaysia: 1 male (Fig. 1a–e): ‘Malaysia [p] / 9. 2. 74 [h] / Coll. Pfanner [p]; NHMUK014596798 [p]’; 1 female (Fig. 1f–j): ‘Malaysia / Cameron Highlands / 1400m 17 iv 73 / via coll. P. Pfanner [h]; Mimomyagrus pfanneri / (Breuning) / Cameron Highlands / (Malaysia 1400 m / 17/4/1973) [p]; Brit. Mus. / 1978.72 [p]; NHMUK014596799 [p]’.

Figure 1. 

Habitus of Combe brianus (White, 1858) a–d male f–i female e, j labels.

Comments

White (1858) described Monohammus brianus (Fig. 2a–d) based on a female specimen without head from Nepal. Thomson (1864) established the genus Combe for Combe fulgurata (Fig. 2e–i) based on a female specimen without indicating locality. Pascoe (1866) transferred Monohammus brianus to the genus.

Figure 2. 

Habitus of Combe brianus (White, 1858) a–c holotype of Monohammus brianus White, 1858, female e–h holotype of Combe fulgurata Thomson, 1864, female j–m holotype of Mimomyagrus pfanneri Breuning, 1970, female d, i, n labels.

Combe and synonymized Combe fulgurata with it based on the male and female specimens from Malacca, Malaysia. Breuning (1954) described a morph, Combe brianus m. rufoantennatus (Fig. 3f–j), based on a specimen of reddish-brown colour from Sumatra. Subsequently, Breuning (1970) described Mimomyagrus pfanneri based on specimens from Cameron Highlands, Malaysia. Through studying these series of type and non-type specimens, we found that, in fact, they belong to the same species. Thus, here we propose the species Mimomyagrus pfanneri as a junior synonym of Combe brianus.

Figure 3. 

Habitus of Combe brianus (White, 1858) a–d paratype of Mimomyagrus pfanneri Breuning, 1970, female f–i holotype of Combe brianus m. rufoantennatus Breuning, 1954, female e, j labels.

Since Mimomyagrus pfanneri is the type species of Mimomyagrus, this genus becomes a junior synonym of the genus Combe.

Breuning (1954) regarded the type specimen of Combe brianus m. rufoantennatus as a male. However, based on the ratio of the antennal length to body length, and the shape of the abdomen, it appears to be a female.

A questionable paratype of Mimomyagrus pfanneri (Fig. 3a–e) is present in the collection of the NHMUK. According to the label, a handwritten collection date indicates that the specimen was collected in April, 1974 (Fig. 3e). The species was described in 1970 and the collection date was clearly recorded as ‘IV–V 1969’ in the original publication by Breuning, although on the label of the holotype a similarly handwritten date is given as ‘July 1968’. In any case, it is impossible for the collection date of the claimed paratype to appear after the publication date. Therefore, this specimen cannot be considered as a paratype.

Arctolamia sinica Bi & Chen, 2022

Figs 4a–j, 5f, i

Arctolamia sinica Bi & Chen, 2022: 199.

Description

Female: similar to male, body length 33.5 mm, humeral width 12.0 mm. Body black, mostly densely clothed with reddish-brown pubescence, each puncture bearing a black or reddish-brown erect hair; glabrous areas showing black integument; elytra provided with five black pubescent patches. Frons, gena and vertex with black hairs, mouthparts with hairs lighter in colour, nearly yellowish-brown. Antennae with scape clothed with reddish-brown pubescence only on lateral margin; antennomeres III–V clothed with reddish-brown pubescence on basal half, antennomeres VI–VIII on basal half and antennomere XI on extreme apex clothed with greyish-yellow pubescence; antennomeres III–IV clothed with black pubescence on apical half, antennomeres V–VIII clothed with pubescence fading to chestnut on apical half; antennomeres IX–X and most of antennomere XI clothed with chestnut pubescence; scape and pedicel clothed with a black long hair on each puncture, denser on inferior margin; antennomeres III–VI fringed with long hairs below, reddish-brown on base and black on apex, antennomeres III–IV tufted with black hairs around apex. Pronotum clothed with black erect hairs on anterior, lateral and posterior margin, more on anterior margin; disc barely clothed with hairs on calli. Scutellum clothed with reddish-brown pubescence, without hairs. Elytra densely clothed with reddish-brown and black hairs, glabrous only on basal granules; each puncture bearing a long erect black or reddish-brown hair, black hairs short and stiff, reddish-brown hairs long and soft, arranged intermixed with each other; elytra with five black pubescent patches: a basal one located around scutellum, subangular; a lateral one on basal quarter after each humerus, smallest, oblique, not reaching the lateral margin; a large oblique one on each side behind the middle, not reaching suture and lateral margin. Underside densely clothed with reddish-brown pubescence, slightly greyish-yellow on mesosternum and mesoepisternum; ventrites furnished with greyish-yellow and black erect hairs. Legs mostly clothed with reddish-brown pubescence, tibiae and tarsi furnished with sparse black bristles.

Head sparsely punctate; frons transverse, slightly convex, with a distinct median sulcus extending to occiput. Eyes coarsely faceted, lower eye lobe transverse, about as long as gena; vertex uneven, with irregular wrinkles. Antennae distinctly shorter than body; antennal insertions conspicuously elevated, broadly separated; scape stout, gradually thickened apically, longest; antennomere III slightly shorter than antennomere IV, antennomere IV about as long as antennomere V, antennomeres V–X gradually shortened in length, antennomere XI about as long as antennomere VII. Pronotum transverse, with a pointed lateral spine on middle of each side; disc convex, coarsely rugose, with developed calli. Scutellum semicircular. Elytra broad, lateral margins gradually expanding outward at basal quarter after humeri, then convergent backward from middle to apices, apices conjointly rounded; about basal fourth provided the sparse, glabrous granules, of which several large granules regularly arranged in a row near the scutellum. Abdomen with first ventrite distinct longer than second and third ventrite, distal ventrite with apical centre slightly depressed, apical margin nearly straight. Legs moderately long and thick, metafemur reaching the middle of fourth ventrite.

Non-type material examined

Myanmar: 1 male: ‘UPPER BURMA: / Nam Tamai Valley / 28. viii. 1938. / R. Kaulback. / B. M. 1938–741. [p]; Alt. 6,000 ft. / Lat. N. 27°42′. / Long. E. 97°54′. [p]; Arctolamia / fruhstorferi / Auriv [h] / DET. –E. F. GILMOUR [p]; NHMUK014596131’; 1 female: ‘UPPER BURMA: / Nam Tamai Valley / 28. viii. 1938. / R. Kaulback. / B. M. 1938–741. [p]; Alt. 6,000 ft. / Lat. N. 27°42′. / Long. E. 97°54′. [p]; NHMUK014596132’.

Comments

Gilmour (1950) misidentified the above pair of specimens as Arctolamia fruhstorferi Aurivillius, 1902 (Fig. 4a–j) and, using this as a comparison, described another species, Arctolamia margaretae Gilmour, 1950 (Fig. 4k–o). In fact, A. margaretae is a junior synonym of A. fruhstorferi, as shown by Pu (1981), whereas the specimens misidentified by Gilmour as A. fruhstorferi represent A. sinica Bi & Chen, 2022.

Figure 4. 

Habitus of Arctolamia spp. a–j Arctolamia sinica Bi & Chen, 2022 k–o holotype of Arctolamia margaretae Gilmour, 1950 a–d male f–i, k–n female e, j, o labels.

Bi and Chen (2022) indicated that A. sinica can be differentiated from A. fruhstorferi by the absence of light-coloured pubescence on the dorsal surface of the scape and numerous large granules on the base of elytra. However, a male specimen of A. sinica from Myanmar shows the scape distinctly clothed with reddish-brown pubescence on dorsal surface (Fig. 4a, b, d) and there is a female specimen in the NHMUK identified as A. fasciata Gestro, 1891 that is actually supposed to be A. fruhstorferi (maybe it is a transitional form), which also has some large granules on the elytral base (Fig. 4a–e). This seems to imply that, given current knowledge, the main feature that distinguishes A. sinica from A. fruhstorferi is the reddish-brown pubescence on its body.

Figure 5. 

Habitus of Arctolamia spp. a–d, g, j Arctolamia fruhstorferi Aurivillius, 1902 f, i Arctolamia sinica Bi & Chen, 2022 h, k holotype of Arctolamia margaretae Gilmour, 1950 e labels a–d, f–k female.

Arctolamia sinica, is also recorded in Myanmar for the first time based on the pair of specimens mentioned above.

Additional discoveries

Type specimens of Lamia punctator Fabricius, 1776 [= Anoplophora chinensis (Forster, 1771)] (Fig. 6a–e) and Cerambyx galloprovincialis Olivier, 1800 [= Monochamus galloprovincialis (Olivier, 1800)] (Fig. 6f–k) and the holotype of Melanauster granulipennis Breuning, 1938 [= Monochamus guerryi Pic, 1903] (Fig. 6l–p) are confirmed to be preserved in the NHMUK; all were considered to be ‘lost’ and marked as ‘depositary not known’ (Tavakilian and Chevillotte 2023). Lamia punctator Fabricius, 1776 and Cerambyx galloprovincialis Olivier, 1800 are marked as having (missing) ‘holotypes’ by Tavakilian and Chevillotte (2023) and are labelled at the NHMUK with standard red framed ‘type’ discs, but there is no evidence of a holotype or of there being only one specimen in their original descriptions, so they are treated as syntypes. Melanauster granulipennis Breuning, 1938 is described from a single female, and this specimen is a holotype.

Figure 6. 

a–d type specimen of Lamia punctator Fabricius, 1776, female f–i type specimen of Cerambyx galloprovincialis Olivier, 1800, female l–o holotype of Melanauster granulipennis Breuning, 1938, female e, j, k, p labels.

Acknowledgments

We wish to express our sincere thanks to Antoine Mantilleri and Christophe Rivier (Muséum national d’Histoire naturelle, Paris) for taking the holotype photographs of Combe fulgurata Thomson, to Alain Drumont and Julien Lalanne (Royal Belgian Institute of Natural Sciences, Brussels) for taking the holotype photographs of Combe brianus m. rufoantennatus Breuning, and to Anne Freitag (Muséum cantonal des sciences naturelles, Lausanne) for taking the holotype photographs of Mimomyagrus pfanneri Breuning, 1970, and especially to Francesco Vitali (Musée national d’histoire naturelle de Luxembourg, Luxembourg), subject editor of ZooKeys, for carefully revising and patiently handling the manuscript. We also wish to express our special thanks to the Coleoptera curatorial team of the NHMUK, past and present, for maintaining and developing the collection and particularly to Keita Matsumoto and Michael Geiser for supporting the first author during his study at the Natural History Museum, London.

Additional information

Conflict of interest

The authors have declared that no competing interests exist.

Ethical statement

No ethical statement was reported.

Funding

This research was supported by the China Scholarship Council (202008420315) and the National Natural Science Foundation of China (No. 31672327).

Author contributions

Conceptualization: GX. Funding acquisition: WW. Resources: MVLB. Writing - original draft: GX. Writing - review and editing: MVLB, WW.

Data availability

All of the data that support the findings of this study are available in the main text.

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