Geophilids with head distinctly elongate; clypeal areas present, variously distinct; labral side-pieces almost touching medially; second maxillary coxosternite medially very short and poorly sclerotized, with statuminia, without anterior inner processes; second maxillary pretarsus claw-like; forcipular tergite distinctly narrower than subsequent tergite, with pleurites exposed dorsally; forcipular coxosternite relatively broad posteriorly, without anterior denticles, with chitin-lines short or absent, with coxopleural sutures subparallel in their anterior half; forcipular trochanteroprefemur distinctly elongate, with distal denticle; forcipular tarsungulum with basal denticle; trunk sternites without “carpophagus” structures; ventral pore-fields usually absent; metasternite of ultimate leg-bearing segment longer than wide; coxopleura usually with sparse pores; legs of the ultimate pair longer than the penultimate legs, often without pretarsus. See Table 2.

Russia: Chukotka autonomous okrug: “Nunamo” (Attems 1909; see Remarks), “Konyam Bay im Senjavin Sund” (Attems 1909) = Penkigney Bay, ca 64°49'N, 172°53'W; USA: Alaska: “Port Clarence” (Attems 1909), ca 65°15'N, 166°51'W.

Syntypes : 7 specimens, including 3 males and 4 females. Deposited in NHMW (Ilie et al. 2009).

A species of Arctogeophilus with first maxillary lappets relatively short; denticles on all forcipular articles; denticles on the forcipular intermediate articles only slightly shorter than those on trochanteroprefemur and tarsungulum; 39 leg-bearing segments, possibly invariably; ventral pore-fields absent; pretarsus of ultimate legs absent.

Far East: Chukotka autonomous okrug (Attems 1909). Outside Asian Russia: Alaska and Canada (e.g., Attems 1909; Chamberlin 1946; Langor and Langor 2022).

The position of the locality “Nunamo” (indicated by Attems 1909) is uncertain: the original material was collected during the Vega expedition, and Nordenskiöld (1882: 565, 567) mentioned a tent-village “Nunamo” located in Chukotka, but he did not indicate the precise position of this place. Instead, Nordenskiöld (1882: 565) provided coordinates for “Konyam Bay” = Penkigney Bay.

Russia: Republic of Tatarstan: Chistopolsky District: “Bliz s. Zmievo” (Folkmanová and Dobroruka 1960) = near Zmievo village, ca 55°23'N, 50°43'E.

Syntypes : 20 specimens. Depository unknown.

A species of Arctogeophilus with first maxillary lappets relatively long; denticles on all forcipular articles, those on intermediate articles smaller than those on trochanteroprefemur and tarsungulum; 35–43 leg-bearing segments; ventral pore-fields on some anterior segments; pretarsus of ultimate legs absent.

Western Siberia: Sverdlovsk, Chelyabinsk, Tomsk, and Kemerovo oblasts, Altai krai, republics of Altai and Khakassia (Byzova and Chadaeva 1965; Zalesskaja et al. 1982; Farzalieva 2008; Nefediev et al. 2017a, 2017c, 2018, 2021; Nefediev 2019). Eastern Siberia: Republic of Tuva and Krasnoyarsk krai (Zalesskaja et al. 1982; Vorobiova 1999; Rybalov 2002; Vorobiova et al. 2002). Far East: Jewish autonomous oblast, Amur oblast, Khabarovsk and Maritime krais, Chukotka autonomous okrug, and Sakhalin oblast (Sakhalin and Kunashir islands) (Zalesskaja et al. 1982; Ganin 1997). Outside Asian Russia: westwards to Transcarpathia (Zalesskaja et al. 1982; Volkova 2016) and Eastern Kazakhstan (Dyachkov and Tuf 2019).

Zalesskaja et al. (1982: 189) suggested this species can be a junior synonym of A. glacialis (Attems, 1909), and this is still in doubt (Nefediev et al. 2017a). Because of this uncertainty, the actual taxonomic status of the populations of Arctogeophilus from the Far East remains to clarify.

The type locality for A. macrocephalus has been sometimes reported erroneously in previous publications (e.g., Bonato and Minelli 2014).

Russia: Sakhalin oblast: “Insel Sachalin” (Verhoeff 1934) = Sakhalin Isl.

Holotype : male. Depository unknown.

A species of Arctogeophilus with first maxillary lappets relatively long; forcipular denticles only on the trochanteroprefemur and tarsungulum, not on the intermediate articles; 39 leg-bearing segments, possibly invariably; ventral pore-fields on some anterior segments; pretarsus of ultimate legs absent.

Far East: Sakhalin oblast (Sakhalin Isl.) (Verhoeff 1934; Takakuwa 1940; Molodova 1973; Kurcheva 1977; Ganin 1997), Maritime krai (Kurcheva 1977; Ganin 1997), and Chukotka autonomous okrug (Kurcheva 1977). Outside Asian Russia: no records.

It has been suggested that A. sachalinus could be a junior synonym of A. glacialis (Attems, 1909) (Nefediev et al. 2017a). More generally, the actual taxonomic status of the populations of Arctogeophilus from the Far East remains to clarify.

Geophilids with head usually only slightly elongate; clypeal areas usually not distinct; labral side-pieces distinctly separated by an intermediate part; second maxillary coxosternite medially long and sclerotized, without both statuminia and anterior inner processes; second maxillary pretarsus claw-like or reduced; forcipular tergite approximately as broad as the subsequent tergite, covering most part of the pleurites; forcipular coxosternite usually wider than long, gradually narrowing posteriorly, without anterior denticles, with chitin-lines, with coxopleural sutures diverging anteriorly also in their anterior half; forcipular trochanteroprefemur only moderately elongate, usually without denticles; forcipular tarsungulum with at most a small basal denticle; trunk sternites often with “carpophagus” pits and often with ventral pore-fields, usually a transverse band on the posterior part of the sternite; metasternite of the ultimate leg-bearing segment usually wider than long; coxopleura with sparse ventral pores, most of them close to metasternite; pretarsus of ultimate leg pair claw-like or reduced. See Table 3.

Russia: Sakhalin oblast: Sakhalin Isl.: “Otako (Chikuka)” (see Remarks) (Takakuwa 1937c).

Syntypes : unknown number of specimens, both sexes. Depository unknown.

A species of Geophilus with head slightly longer than wide, 35–39 leg-bearing segments; “carpophagus” pits present, up to as wide as the metasternites; ventral pore-fields present, an entire posterior diamond on the anterior metasternites, absent on most of the posterior metasternites, two paired posterior groups on the penultimate metasternite; metasternite of ultimate leg-bearing segment wider than long; a few coxal pores on each coxopleuron, all close to the margin of metasternite; pretarsus of ultimate leg pair claw-like; anal pores present.

Far East: Sakhalin oblast (Sakhalin and Kurile Islands) (Takakuwa 1937b; Molodova 1973; Kurcheva 1977). Outside Asian Russia: no records.

The current identity and position of the locality “Otako (Chikuka)” (indicated by Takakuwa 1937c) is uncertain.

Unknown, possibly in France (Bonato and Minelli 2014).

A species of Geophilus with head ~ 1.2× as long as wide; antennal articles ≤ 2× as long as wide; second maxillary pretarsus claw-like, longer than surrounding setae; forcipular trochanteroprefemur ~ 1.5× as long as wide; forcipular trochanteroprefemur, femur, and tibia without denticles; tarsungulum bearing a small basal denticle; 37–61 leg-bearing segments; “carpophagus” pits absent; ventral pore-fields present, an entire transverse posterior band on the anterior metasternites, two paired posterior groups on the posterior metasternites; metasternite of ultimate leg-bearing segment wider than long; up to a dozen coxal pores on each coxopleuron, all close to the margin of metasternite; pretarsus of ultimate leg pair claw-like; anal pores present.

Western Siberia: Tomsk oblast (Nefediev et al. 2017a). Eastern Siberia: Zabaykalsky krai (Gerstfeldt 1859, as Arthronomalus longicornis). Outside Asian Russia: Western Palaearctic (e.g., Bonato et al. 2005; Stoev 2007).

The record from Western Siberia seems to be due to anthropochore introduction (Nefediev et al. 2017a), while the record from Eastern Siberia requires confirmation (see Discussion).

Russia: Khabarovsk krai: “Nikolaevsk na Amure” (Sseliwanoff 1881) = Nikolayevsk-on-Amur city, 53°8'N, 140°42'E.

Holotype : male. Deposited in ZISP.

A species of Geophilus with head slightly longer than wide; forcipular coxosternite with short and wide denticles; all forcipular articles with small denticles, except the tarsungulum; 39 leg-bearing segments; metasternite of ultimate leg-bearing segment longer than wide, only slightly narrowing backwards; numerous coxal pores, on the ventral and lateral sides of coxopleura; anal pores present.

Far East: Khabarovsk krai (Sseliwanoff 1881, 1884). Outside Asian Russia: no records.

The species was originally described under the genus Geophilus, but its taxonomic position is considered uncertain since Attems (1929). The few morphological characters reported in the descriptions and keys by Sseliwanoff (1881, 1884) do not allow it to be assigned confidently to one of the known genera.

Germany: near Regensburg (Bonato and Minelli 2014).

A species of Geophilus with head slightly longer than wide; antennal articles ≤ ~ 1.5× as long as wide; second maxillary pretarsus claw-like, longer than surrounding setae; forcipular trochanteroprefemur slightly longer than wide; forcipular trochanteroprefemur, femur, and tibia without denticles; forcipular tarsungulum bearing a small basal denticle; 45–55 leg-bearing segments; “carpophagus” pits present, up to as wide as the metasternites; ventral pore-fields present, an entire posterior diamond on the anterior metasternites, absent on the posterior metasternites; metasternite of ultimate leg-bearing segment wider than long; up to a dozen coxal pores on each coxopleuron, all close to the margin of metasternite; pretarsus of ultimate leg pair claw-like; anal pores present.

Western Siberia: “Western Siberia” (Sseliwanoff 1881); Sverdlovsk, Chelyabinsk, Tyumen, Omsk, Novosibirsk, Kemerovo, and Tomsk oblasts, Altai krai, Republic of Khakassia (Zalesskaja et al. 1982; Poryadina 1991; Striganova and Poryadina 2005; Farzalieva 2008; Bukhkalo and Sergeeva 2012; Sergeeva 2013, 2014; Bukhkalo et al. 2014; Nefediev et al. 2017a, 2017b, 2021; Nefediev 2019). Outside Asian Russia: Northern Europe (Bonato et al. 2005).

Russia: Sakhalin oblast: “Sachalin (Tomaruoru (= Tomari сity, 47°45'N, 142°3'E), Maoka (= Kholmsk city, 47°02'N, 142°02'E), Sirutori (= Makarov сity, 48°37'N, 142°46'E))” (Takakuwa 1937c).

Syntypes : unknown number of specimens, both sexes. Depository unknown.

A species of Geophilus with head approximately as long as wide; second maxillary pretarsus claw-like and relatively long; forcipular trochanteroprefemur, femur, and tibia without denticles; forcipular tarsungulum bearing a small basal denticle; 43–49 leg-bearing segments; “carpophagus” pits present, up to as wide as the metasternites; ventral pore-fields present, an entire posterior diamond on the anterior metasternites; metasternite of ultimate leg-bearing segment wider than long; a few coxal pores on each coxopleuron, most of them close to the margin of metasternite and one pore located separately; pretarsus of ultimate leg pair claw-like; anal pores present.

Far East: Sakhalin oblast (Sakhalin Isl.) (Takakuwa 1937b, 1937c; Kurcheva 1977). Outside Asian Russia: Japan (Takakuwa 1937c).

Russia: Krasnoyarsk krai: “Krasnojarsk” (Stuxberg 1876b) = Krasnoyarsk city, 56°0'N, 92°53'E.

Syntypes : 3 females. Depository unknown.

A species of Geophilus with head ~ 1.2× as long as wide; tarsungulum without basal denticle; 57–59 leg-bearing segments; more than a dozen coxal pores on each coxopleuron, both on the ventral and lateral sides; pretarsus of ultimate leg pair claw-like; anal pores absent.

Eastern Siberia: Krasnoyarsk krai (Stuxberg 1876b). Outside Asian Russia: no records.

The species was originally described under the genus Geophilus, but its taxonomic position was considered uncertain since Attems (1929). The few morphological characters reported in the descriptions by Stuxberg (1876a, 1876b) do not allow to confidently assign it to one of the known genera.

Japan: Hokkaido: “Sounkyo” (Takakuwa 1937c).

Syntypes : unknown number of specimens, both sexes. Depository unknown.

A species of Geophilus with second maxillary pretarsus claw-like, longer than surrounding setae; forcipular trochanteroprefemur slightly longer than wide; forcipular trochanteroprefemur, femur and tibia without denticles; forcipular tarsungulum bearing a small basal denticle; 55–57 leg-bearing segments; “carpophagus” pits present, up to as wide as the metasternites; ventral pore-fields present, an entire posterior band and other sparse pores on the anterior metasternites, absent on the posterior metasternites; metasternite of ultimate leg-bearing segment wider than long; a few coxal pores on each coxopleuron, most of them close to the margin of metasternite and one pore located separately; anal pores present.

Far East: Maritime krai (Ghilarov and Perel 1973; Kurcheva 1977; Ganin 1997). Outside Asian Russia: Japan (Takakuwa 1937b, 1937c, 1940).

Geophilids with head distinctly elongate; clypeal areas distinct; labral side-pieces distinctly separated by an intermediate part; second maxillary coxosternite medially long and sclerotized, without both statuminia and anterior inner processes; second maxillary pretarsus claw-like; forcipular tergite distinctly narrower than subsequent tergite, with pleurites exposed dorsally; forcipular coxosternite relatively broad posteriorly, with coxopleural sutures subparallel in their anterior half; forcipular trochanteroprefemur distinctly elongate, with distal denticle; chitin-lines present but short; forcipular tarsungulum with basal denticle; trunk sternites without “carpophagus” structures; ventral pore-fields present at least on the anterior part of the trunk, two paired anterior groups and a posterior entire transverse band on each sternite; metasternite of ultimate leg-bearing segment approximately as long as wide or longer than wide; coxopleura with sparse pores; legs of the ultimate pair with claw-like pretarsus. See Table 4.

Germany: Arklee, near Regensburg (Bonato and Minelli 2014).

A species of Pachymerium with two paired clypeal areas; forcipular coxosternite with chitin-lines, which extend for most part of the length of the coxosternite, but do not reach the anterior margin; both forcipular trochanteroprefemur and tarsungulum with relatively small denticles; 41–69 leg-bearing segments; ventral pore-fields present, also on the posterior part of the trunk, where they are reduced to two paired posterior groups on each metasternite; ultimate metasternite trapezoidal, approximately as long as wide, distinctly tapering towards the posterior margin; all coxal pores sparse from the ventral to the dorsal sides of the coxopleura.

Western Siberia: Chelyabinsk, Sverdlovsk, Tyumen, and Tomsk oblasts, Altai krai (Zalesskaja et al. 1982; Farzalieva 2008; Sergeeva 2013; Bukhkalo et al. 2014; Nefediev et al. 2017a). Far East: Amur oblast, Jewish autonomous oblast, and Maritime krai (Zalesskaja et al. 1982; Ganin 1997). Outside Asian Russia: Western Palaearctic.

Russia: Sakhalin oblast: “Sartung, paa Oen Sacolin” (Meinert 1870) = “Sartung” (see Remarks), Sakhalin Isl.

Syntypes : 2 specimens, both sexes. Deposited in NHMD.

A species of Pachymerium with forcipular coxosternite with very short chitin-lines; forcipular tarsungulum without denticle; 45–49 leg-bearing segments; ultimate metasternite narrow, with subparallel lateral edges; all coxal pores sparse from the ventral to the dorsal sides of the coxopleura.

Eastern Siberia: Krasnoyarsk krai, Irkutsk oblast (Stuxberg 1876a, 1876b; Sseliwanoff 1881, 1884; Daday 1889). Far East: Sakhalin oblast (Sakhalin Isl.) (Meinert 1870). Outside Asian Russia: no records.

Sseliwanoff (1884) reported specimens from “near Baikal”, collected by G. Dybovskiy. According to the catalogue of Chilopoda of the ZMMU, these specimens were collected from the Kultuk (urban-type settlement in the Irkutsk oblast, 51°43'N, 103°40'E).

The assignment of this nominal species to the genus Pachymerium is only tentative (Bonato et al. 2016), and is suggested only by the following few characters described by Meinert (1870): head ~ 1.2× as long as wide, forcipular coxosternite very broad and with two anterior denticles, forcipules surpassing the anterior margin of the head, coxopleura elongate and with dense coxal pores not only on the ventral side but also on the lateral and dorsal ones, ultimate legs slightly longer than penultimate legs and bearing a claw.

The current identity and position of the locality “Sartung” (indicated by Meinert 1870) is uncertain.

Geophilids with head slightly wider or as wide as long; clypeal areas absent; labrum without obviously distinct lateral parts; second maxillary coxosternite medially long and sclerotized, without both statuminia and anterior inner processes; second maxillary pretarsus relatively small, claw-like; forcipular tergite approximately as wide as the subsequent tergite, covering pleurites almost completely; forcipular coxosternite distinctly shorter than wide, without both anterior denticles and chitin-lines, with coxopleural sutures distinctly diverging also in their anterior half; forcipular trochanteroprefemur relatively stout, without denticle; forcipular tarsungulum with a relatively large basal denticle; trunk sternites without “carpophagus” structures; ventral pore-fields present, at least two paired ovoid posterior fields on each metasternite; coxal pores only on the ventral side of coxopleura, denser near the metasternite; leg of the ultimate pair usually with claw-like pretarsus. See Table 5.

United Kingdom: “Roborough Down near Plymouth” and “Battersea fields” (Leach 1815).

A species of Strigamia with clypeal setae arranged in an intermediate and two lateral groups; forcipular tarsungula not surpassing the anterior margin of the head; basal denticle of forcipular tarsungulum relatively short and with straight converging margins; 37–43 leg-bearing segments; metasternites of the anterior part of the trunk without a mid-longitudinal sclerotized stripe; ultimate leg-bearing segment with pleuropretergite entire, i.e., without distinct intercalary pleurites, and metasternite approximately as long as wide.

Far East: Amur oblast and Khabarovsk krai (Ganin 1997), Sakhalin oblast (Kuril Islands) (Takakuwa 1933; Kurcheva 1977). Outside Asian Russia: Europe (Barber 2009; Bonato et al. 2012, 2023; Volkova 2016).

All records from Russian Far East need confirmation, as are the records from Japan and the Western part of North America, because of probable confusion with other species including S. chionophila Wood, 1862 (Bonato et al. 2012).

Japan: “Yamanaka, Suruga” and “Bukenji” (Attems 1927).

Syntypes : 2 specimens, both sexes. Deposited in NHMW (Ilie et al. 2009).

A species of Strigamia with forcipular tarsungula not surpassing the anterior margin of the head; basal denticle of forcipular tarsungulum relatively large; internal and external margins of forcipular tarsungulum subparallel in their intermediate part; 51–67 leg-bearing segments; metasternites of the anterior part of the trunk with a mid-longitudinal sclerotized stripe; ultimate leg-bearing segment with distinct intercalary pleurites, and metasternite distinctly wider than long.

Far East: Sakhalin oblast (Sakhalin Isl.) (Kurcheva 1977). Outside Asian Russia: Korean Peninsula and Japan (Bonato et al. 2012).

Japan: “Kanagava”, “Yamanaka (Suruga)”, “Bukengi”, and Negishi” (Attems 1927).

Syntypes : ca 42 specimens, both sexes. Deposited in NHMW (Ilie et al. 2009).

A species of Strigamia with forcipular tarsungula not surpassing the anterior margin of the head; basal denticle of forcipular tarsungulum relatively short; internal and external margins of forcipular tarsungulum gradually converging through the entire length; 39–53 leg-bearing segments; ultimate leg-bearing segment with distinct intercalary pleurites and metasternite longer than wide.

Far East: Maritime krai, Sakhalin oblast (Sakhalin Isl.) (Ghilarov and Perel 1973; Kurcheva 1977; Ganin 1997). Outside Asian Russia: Japan (Attems 1927; Bonato et al. 2012).

The taxonomic distinction of this nominal species from S. sacolinensis is uncertain.

Records from South-Eastern Asia are doubtful (Bonato et al. 2012).

Russia: Moscow oblast: “Zaraysk, Ryazanskoy Gubernii” (Sseliwanoff 1884) = Zaraysk city, 54°45'N, 38°53'E.

Syntypes : 11 specimens, including 4 males and 7 females. Deposited in ZISP.

A species of Strigamia with forcipular tarsungula not surpassing the anterior margin of the head; basal denticle of forcipular tarsungulum relatively short, pointed, and with straight outlines; internal and external margins of forcipular tarsungulum gradually converging through the entire length; 33–39 leg-bearing segments; metasternites of the anterior part of the trunk without a mid-longitudinal sclerotized stripe; ultimate leg-bearing segment with distinct intercalary pleurites and metasternite as long as wide or longer than wide.

Western Siberia: Sverdlovsk oblast (Zalesskaja et al. 1982; Farzalieva 2008), Altai krai (Nefediev et al. 2018; Nefediev 2019), and Republic of Altai (Nefediev et al. 2017c). Eastern Siberia: Republic of Sakha (Yakutia) (Nefediev 2019). Outside Asian Russia: westwards to Central Europe (Zalesskaja et al. 1982; Bonato et al. 2012; Tuf and Tajovsky 2016; Volkova 2016), and southwards to Northern Mongolia (Poloczek et al. 2016; Dyachkov and Farzalieva 2023).

Russia: Sakhalin oblast: “Sartung paa Oen Sacalin” (Meinert 1870) = “Sartung” (see Remarks), Sakhalin Isl.

Holotype : female. Deposited in NHMD.

A species of Strigamia with forcipular tarsungula surpassing the anterior margin of the head; 43–47 leg-bearing segments; metasternites of the anterior part of the trunk without a mid-longitudinal sclerotized stripe.

Far East: Khabarovsk krai (Sseliwanoff 1881, 1884) and Sakhalin oblast (Sakhalin Isl.) (Meinert 1870; Molodova 1973; Kurcheva 1977; Ganin 1997). Outside Asian Russia: no records.

Bonato et al. (2012) suggested the putative projection of the forcipules in front of the anterior margin of the head can be due to some post-mortem displacement of the head with respect to the trunk. Based on the incomplete description provided by Meinert (1870), this nominal species could be a senior synonym of either S. hirsutipes or S. japonica (Verhoeff, 1935).

The current identity and position of the locality “Sartung” (indicated by Meinert 1870) is uncertain.

Russia: Zabaykalsky krai: “Yablonoviy Khrebet” (Sseliwanoff 1881) = Yablonoviy Mt. Range, ca 52°2'N, 113°35'E.

Syntypes : 4 specimens, including 1 male and 3 females. Deposited in ZISP.

A species of Strigamia with forcipular tarsungula not surpassing the anterior margin of the head; 33–35 leg-bearing segments.

Eastern Siberia: Zabaykalsky krai (Sseliwanoff 1881). Outside Asian Russia: no records.

The distinction between this nominal species and S. pusilla is unclear (Bonato et al. 2012).

Russia: Khabarovsk krai: “Nikolaevsk-na-Amure” (Sseliwanoff 1881) = Nikolayevsk-on-Amur city, 53°8'N, 140°42'E.

Syntypes : 3 specimens, including 2 males and 1 female. Deposited in ZISP.

A species of Strigamia with forcipular tarsungula not surpassing the anterior margin of the head; 41–43 leg-bearing segments.

Far East: Khabarovsk krai (Sseliwanoff 1881, 1884; Ganin 1997). Outside Asian Russia: no records.

The distinction between this nominal species and many other congeneric species is unclear (Bonato et al. 2012).

Romania: “Hermannstadt” = Sibiu. Slovenia: “Gottschee” = Kocevje. Austria: “Ostalpen” = Eastern Alps; “Hermagor”; “Arlberg”. Italy: “Schneelagern am Schlüsseljoch beim Brenner” = Colle della Chiave, near Brennero. Slovakia: “Tatra-Höhlenhain” = Tatranska Kotlina. Germany: “Titisee”. Switzerland: “Pilatus Kulm” (Verhoeff 1928).

A species of Strigamia with clypeal setae uniformly spaced in a continuous array, without recognizable gaps between intermediate and lateral groups of setae; forcipular tarsungula not surpassing the anterior margin of the head; basal denticle of forcipular tarsungulum relatively large and with distinctly curved outlines; internal and external margins of forcipular tarsungulum gradually converging through the entire length; 43–57 leg-bearing segments; metasternites of the anterior part of the trunk without a mid-longitudinal sclerotized stripe; ultimate leg-bearing segment with pleuropretergite entire, i.e., without distinct intercalary pleurites, and metasternite approximately as long as wide.

Western Siberia: Altai krai (Nefediev et al. 2018). Far East: Maritime krai, Sakhalin oblast (Sakhalin Isl.) (Kurcheva 1977; Ganin 1997). Outside Asian Russia: westwards to Central Europe (Iorio 2005; Reip and Voigtländer 2009).

Strigamia transsilvanica belongs to a species complex whose taxonomy is only partially resolved (Bonato et al. 2023).

Bonato et al. (2012) suggested that the records from Russian Far East are probably due to misidentification of a different species. Nefediev et al. (2018) suggested the presence of a possible undescribed species similar in morphology to S. transsilvanica from Western Siberia, so that also the presence of S. transsilvanica in Western Siberia is doubtful. Doubtful are also the records from European Russia and Caucasus (Zuev and Evsyukov 2016; Dyachkov et al. 2022; Dyachkov and Zuev 2023), Eastern Kazakhstan (Dyachkov 2018b), Japan, and Taiwan (Bonato et al. 2012).

Family Mecistocephalidae Bollman, 1893

Mecistocephalids with head moderately longer than wide; two clypeal plagulae, separated by a mid-longitudinal areolate stripe and extending to the lateral margins of the clypeus; cephalic pleurites without both spiculum and setae; first maxillary coxosternite medially divided by a sulcus, without antero-lateral corners; second maxillary coxosternite medially undivided, with the grooves from the metameric pores reaching the posterior corners; second maxillary telopodites relatively small, not distinctly overreaching the first maxillary telopodites, without pretarsus; forcipular tergite distinctly wider than long, without a distinct mid-longitudinal sulcus; forcipular trochanteroprefemur with only a distal denticle, tarsungulum with a basal denticle; invariably 41 leg-bearing segments; sternites with non-furcate mid-longitudinal sulcus and without pore-fields; legs of the ultimate pair ending with a short spine. See Table 6.

Russia: Maritime krai: “Sudzuhinsky Zapovednik, Tachingauz” (Titova 1975) = Lazovsky Nature Reserve, Tachingauz bay, ca 43°1'N, 134°8'E.

Holotype : male. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Agnostrup species with body length reaching ≥ 3 cm; clypeal plagulae with an irregular anterior margin and slightly smaller than the areolate part of the clypeus; many setae near the anterior margin of plagulae and on the center of the areolate part of the clypeus; forcipular trochanteroprefemur 1.3× as long as wide, both forcipular femur and tibia with small bulges.

Far East: Maritime krai (Titova 1975; Markelov and Mineeva 1981; Ganin 1997). Outside Asian Russia: no records.

Mecistocephalids with head moderately longer than wide; two clypeal plagulae, separated by a mid-longitudinal areolate stripe and not extending to the lateral margins of the clypeus; cephalic pleurites without both spiculum and setae; first maxillary coxosternite medially undivided, without antero-lateral corners; second maxillary coxosternite medially undivided, with the grooves from the metameric pores reaching the posterior corners; second maxillary telopodites relatively small, not distinctly overreaching the first maxillary telopodites, usually without pretarsus; forcipular tergite distinctly wider than long, without a distinct mid-longitudinal sulcus; forcipular trochanteroprefemur with only a distal denticle, tarsungulum with a basal denticle; invariably 41 leg-bearing segments; sternites with non-furcate mid-longitudinal sulcus and without pore-fields; legs of the ultimate pair ending with a short spine. See Table 6.

Japan: Hokkaido: “Zyôzankei (bei Sapporo)” (Takakuwa 1934b).

Unknown number of specimens, possibly lost (Jonishi and Nakano 2022). Depositary unknown.

An Arrup species with body reaching ≥ 2 cm; second maxillary pretarsi present (see Remarks); forcipular trochanteroprefemur with a large distal denticle, tibia with large denticle, tarsungulum with pointed basal denticle.

Far East: Maritime krai, Sakhalin oblast (Sakhalin and Kuril Islands: Shikotan) (Titova 1975; Ganin 1997). Outside Asian Russia: Japan (Takakuwa 1934b; Shinohara 1972).

The name Prolamnonyx dentatus was validly introduced by Takakuwa (1934a) in a key; specimens were later described in more detail by Takakuwa (1934b, 1934c). Uliana et al. (2007) described the presence of second maxillary pretarsi in this species, according to the original description and Titova (1975), it is absent in this species.

Russia: Maritime krai: “Primorsky Kray, zapovednik Kedrovaya Pad”, Kedrovaya Pad Nature Reserve, ca 43°05'N, 131°30'E, (Titova 1975).

Holotype : female. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Arrup species with body length reaching ≥ 3 cm; second maxillary pretarsi absent; forcipular trochanteroprefemur with a large distal denticle, both femur and tibia with small bulges, tarsungulum with pointed basal denticle.

Far East: Maritime krai (Titova 1975; Ganin 1997). Outside Asian Russia: no records.

The holotype had been previously assigned to Prolamnonyx holstii (Pocock, 1895) by Titova (1969).

Mecistocephalids with head distinctly longer than wide; clypeus with an entire plagula, without mid-longitudinal areolate stripe and extending to the lateral margins of the clypeus; cephalic pleurites without both spiculum and setae; first maxillary coxosternite medially divided by a sulcus, without antero-lateral corners; second maxillary coxosternite medially undivided, with the grooves from the metameric pores reaching the lateral margins; second maxillary telopodites distinctly overreaching the first maxillary telopodites, with claw-like pretarsus; forcipular tergite only slightly wider than long, without a distinct mid-longitudinal sulcus; forcipular trochanteroprefemur with only a distal denticle, tarsungulum without denticle; invariably 43 or 45 leg-bearing segments; sternites with non-furcate mid-longitudinal sulcus and sometimes with pore-fields; legs of the ultimate pair ending with a short spine. See Table 6.

Indonesia: Java: “Buitenzorg”, “Tjibodas” and “Tjompea” (Attems 1929).

A Tygarrup species with body length ≤ 2 cm; no distinct dark patches along the body; second maxillary pretarsus with a long slender point; both forcipular trochanteroprefemur and tibia with denticles; invariably 45 leg-bearing segments; ventral pore-fields absent in females, present in males; metasternite of the ultimate leg-bearing segment slightly wider than long.

Western Siberia: Altai krai (Nefediev 2019). Outside Asian Russia: Southeast Asia and introduced in Europe (e.g., Bonato et al. 2004; Tuf et al. 2018; Damasiewicz and Leśniewska 2020; Gilgado et al. 2022).

The species is regarded as an anthropochore introduction in Asian Russia (Nefediev 2019).

Family Schendylidae Cook, 1896

Schendylids with head slightly longer than wide; antennae gradually tapering; labrum with distinct denticles in the intermediate part; first maxillae with lappets; second maxillary pretarsi fringed by two rows of filaments; forcipular tergite narrower than subsequent tergite; ventral pore-fields absent; coxal pores numerous and scattered; legs of the ultimate pair with two tarsal articles and claw-like pretarsus, swollen in adult males and slender in females; gonopods biarticulated in both sexes. See Table 7.

Russia: Kemerovo oblast: “Prokopyevsky i Novokuznetsky r-ny” (Titova 1973) = Prokopyevsky, ca 53°53'N, 86°43'E, and Novokuznetsky, ca 53°45'N, 87°07'E, districts.

Holotype : female. Paratypes: 10 specimens, including 5 males and 5 females. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Escaryus species with body length reaching ≥ 1.5 cm; clypeus without plagulae; labral arc relatively shallow, with denticles short and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur and femur with small bulges, tibia with a large bulge, tarsungulum with a large basal denticle; 33–35 leg-bearing segments; metasternite of the ultimate leg-bearing segment trapezoid, distinctly wider than long; coxal pores only on the ventral side of coxopleura; anal pores absent.

Western Siberia: Kemerovo oblast, Republic of Altai (Titova 1973; Nefediev et al. 2017c). Eastern Siberia: Krasnoyarsk krai (Rybalov 2002; Vorobiova et al. 2002). Outside Asian Russia: European Russia (Republic of Bashkortostan) (Titova 1973; Volkova 2016), and Northern Mongolia (Poloczek et al. 2016; Dyachkov and Farzalieva 2023).

Japan: Honshu: Saitama Prefecture: “Chichibu (Mt. Kumotori, Kasatori pass, Mt. Kobushi, Kabagoya-ato, Karisaka pass, Jumonji pass, Mt. Shiraiwa, Mt. Mae-Shiraiwa” (Shinohara 1955).

Syntypes : unknown number of specimens, both sexes. Depository unknown.

An Escaryus species with body length reaching ≥ 2 cm; clypeus with small plagulae; labral arc relatively shallow, with denticles short and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur, femur, and tibia with small bulges, tarsungulum without bulge; 35–39 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment trapezoid, approximately as long as wide; coxal pores on both ventral and lateral sides of coxopleura; anal pores present.

Far East: Sakhalin oblast (Kuril Islands: Kunashir and Shikotan) (Titova 1973; Kurcheva 1977). Outside Asian Russia: Japan (Shinohara 1955).

Russia: Maritime krai: “Suputinsky zapovednik” = Ussuriysky Nature Reserve, ca 43°40'N, 132°32'E, and “Kedrovaya Pad” (both Titova 1973) = Kedrovaya Pad Nature Reserve, ca 43°05'N, 131°30'E, (see Remarks).

Holotype : female. Paratypes: 11 specimens, including 6 males and 5 females. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Escaryus species with body length reaching ≥ 2 cm; clypeus without plagulae (polygonal structure poorly visible, but recognizable); labral arc relatively shallow, with denticles short and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur and tarsungulum with large denticles, femur and tibia with small denticles; 37–39 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment trapezoid, slightly longer than wide; coxal pores on both ventral and lateral sides of coxopleura; anal pores present.

Far East: Maritime krai (Titova 1973; Kurcheva 1977; Ganin 1997, 2006). Outside Asian Russia: no records.

Titova (1973) indicated that the type series was from two localities (“Suputinsky zapovednik” and “Kedrovaya Pad”), but she did not state explicitly which is the locality of the holotype.

Russia: Sakhalin oblast: “O-v Sakhalin, Yuzhno-Sakhalinsk, gora Chekhova” (Titova 1973) = Chekhova Mt., near Yuzhno-Sakhalinsk city, ca 47°00'N, 142°50'E.

Holotype: female. Paratypes: 10 specimens, including 5 males and 5 females. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Escaryus species with body length reaching ≥ 2.5 cm; clypeus with large plagulae; labral arc relatively deep, with denticles long and obtuse; first maxillae with one pair of lappets; all forcipular articles with large bulges; 37–39 leg-bearing segments; metasternites with relatively dense setae; metasternite of the ultimate leg-bearing segment almost rectangular, ~ 1.5× as long as wide; coxal pores on both ventral and lateral sides of coxopleura; anal pores present.

Far East: Sakhalin oblast (Sakhalin Isl.) (Titova 1973; Molodova 1973; Kurcheva 1977; Ganin 1997). Outside Asian Russia: no records.

Japan: Hokkaido: “Todohokhe” (Attems 1927).

Syntypes : 2 specimens, including a male and a juvenile. Deposited in NHMW (Ilie et al. 2009).

An Escaryus species with body length reaching ≥ 4.3 cm; clypeus with large plagulae; labral arc relatively shallow, with denticles long and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur with a small distal denticle, all other articles with very small bulges; 43–55 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment rectangular, ~ 2× as long as wide; coxal pores of similar size, on both ventral and lateral sides of coxopleura; anal pores present.

Western Siberia: Sverdlovsk, Chelyabinsk, Tomsk, and Kemerovo oblasts, republics of Altai and Khakassia (Byzova and Chadaeva 1965; Titova 1972, 1973; Farzalieva 2008; Dyachkov 2017; Nefediev et al. 2017a, 2017c; Nefediev 2019). Eastern Siberia: Republic of Buryatia and Krasnoyarsk krai (Alekseeva 1974; Titova 1973; Vorobiova 1999), Magadan oblast (Berman and Leirikh 2019). Far East: Amur oblast, Maritime and Khabarovsk krais, Sakhalin oblast (Sakhalin Isl.) (Titova 1973; Ghilarov and Perel 1973; Molodova 1973; Kurcheva 1977; Markelov and Mineeva 1981; Ganin 1997, 2006). Outside Asian Russia: European Russia (Titova 1973; Farzalieva 2008; Volkova 2016), eastern Kazakhstan (Dyachkov and Tuf 2018), Mongolia (Dyachkov and Farzalieva 2023), Northern China (Takakuwa and Takashima 1949), and Japan (Attems 1927).

The record from the Krasnoyarsk krai by Vorobiova (1999) was questioned by Nefediev et al. (2017a).

North Korea: “Husenzan” (Takakuwa 1937a).

Syntypes : unknown number of specimens, both sexes. Depository unknown.

An Escaryus species with body length reaching ≥ 6.5 cm; clypeus with large plagulae; labral arc relatively shallow, with long denticles, the middle denticles obtuse, the lateral ones pointed; first maxillae with one pair of lappets; forcipular trochanteroprefemur with an small obtuse denticle, femur and tibia without denticles, tarsungulum with a small basal bulge; 43–55 leg-bearing segments; metasternites with relatively dense setae; metasternite of the ultimate leg-bearing segment rectangular, ~ 2× as long as wide; coxal pores of different size, on both ventral and lateral sides of coxopleura, including a pair of much larger pores on each coxopleuron; anal pores present.

Western Siberia: Altai krai, republics of Altai and Khakassia, Kemerovo, Novosibirsk, and Tomsk oblasts (Titova 1972, 1973; Dyachkov 2017; Nefediev et al. 2017a, 2017c, 2018; Nefediev 2019). Eastern Siberia: Krasnoyarsk krai, Irkutskaya oblast (Rybalov 2002; Vorobiova et al. 2002; Nefediev 2019). Far East: Maritime and Khabarovsk krais, Amur Oblast (Titova 1972, 1973; Ghilarov and Perel 1973; Kurcheva 1977; Ganin 1997). Outside Asian Russia: Eastern Kazakhstan (Dyachkov and Tuf 2018), Japan, and North Korea (Takakuwa 1937a, 1940).

Titova (1973: 113) suggested that E. koreanus can be a junior synonym of E. sibiricus Cook, 1899.

Russia: Maritime krai: “Suputinsky zapovednik” = Ussuriysky Nature Reserve, ca 43°40'N, 132°32'E, and “Kangauz” = Anisimovka Village, ca 43°10'N, 132°47'E and “Kedrovaya Pad” = Kedrovaya Pad Nature Reserve, ca 43°05'N, 131°30'E, (all Titova 1973) (see Remarks).

Holotype : male. Paratypes: 10 specimens, including 5 males and 5 females. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Escaryus species with body length reaching ≥ 2 cm; clypeus without plagulae; labral arc relatively deep, with denticles short and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur and tibia with large bulges, femur and tarsungulum with small bulges; 45–49 leg-bearing segments; metasternite of the ultimate leg-bearing segment trapezoid, approximately as long as wide; coxal pores on both ventral and lateral sides of coxopleura, the pair of largest pores close to inner edge of coxopleura; anal pores present.

Far East: Maritime krai (Titova 1973; Kurcheva 1977; Markelov and Mineeva 1981; Ganin 1997, 2006). Outside Asian Russia: no records.

Titova (1973) indicated that the type series was from three localities (“Suputinsky zapovednik”, “Kangauz”, and “Kedrovaya Pad”), but she did not state explicitly which is the locality of the holotype.

Russia: Sakhalin oblast: “O-v Sakhalin, Yuzhno-Sakhalinsk, gora Chekhova” (Titova 1973) = Chekhova Mt., near Yuzhno-Sakhalinsk city, ca 47°0'N, 142°50'E.

Holotype : male. Paratypes: 8 specimens, including 5 males and 3 females. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Escaryus species with body length reaching ≥ 1.4 cm: clypeus with small plagulae; labral arc relatively shallow, with denticles short and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur, femur and tibia with small denticles; forcipular tarsungulum with a large, pointed basal denticle; 35 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment trapezoid, approximately as long as wide; coxal pores only on the ventral side of coxopleura; anal pores present.

Far East: Sakhalin oblast (Sakhalin Isl.) (Titova 1973; Molodova 1973; Kurcheva 1977; Ganin 1997). Outside Asian Russia: no records.

Russia: Maritime krai: “Suputinsky zapovednik” = Ussuriysky Nature Reserve, ca 43°40'N, 132°32'E, and “Rayon r. Sinancha” = near Cheremukhovaya River, inflow of Dzhigitovka River, ca 44°50'N, 136°07'E (both Titova 1973) (see Remarks).

Holotype : female. Paratypes: 6 specimens, including 2 males and 4 females. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Escaryus species with body length reaching ≥ 2 cm; clypeus with small plagulae; labral arc relatively shallow, with denticles short and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur and tibia with small denticles, femur without denticle, tarsungulum with a large basal denticle; 39–43 leg-bearing segments; metasternite of the ultimate leg-bearing segment trapezoid, distinctly wider than long; most of coxal pores on the ventral side of coxopleura, mostly close to metasternite; anal pores present.

Far East: Amur oblast, Maritime and Khabarovsk krais (Titova 1973; Kurcheva 1977; Markelov and Mineeva 1981; Ganin 1997, 2006). Outside Asian Russia: no records.

Titova (1973) indicated that the type series was from two localities (“Suputinsky zapovednik” and “Rayon r. Sinancha”), but she did not state explicitly which is the locality of the holotype.

Russia: Maritime krai: “Suputinsky zapovednik” = Ussuriysky Nature Reserve, ca 43°40'N, 132°32'E, and “Kedrovaya Pad” = Kedrovaya Pad Nature Reserve, ca 43°05'N, 131°30'E, (both Titova 1973).

Holotype : male. Paratypes: 17 specimens, including 9 males and 8 females. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Escaryus species with body length reaching ≥ 2.8 cm; clypeus with small plagulae; labral arc relatively deep, with denticles short and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur, femur, and tibia with small bulges; forcipular tarsungulum without denticle; 39 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment trapezoid, ~ 1.5× as long as wide; coxal pores of different size, on both ventral and lateral sides of coxopleura; anal pores present.

Far East: Maritime krai (Titova 1973; Kurcheva 1977; Markelov and Mineeva 1981; Ganin 1997). Outside Asian Russia: no records.

Titova (1973) indicated that the type series was from two localities (“Suputinsky zapovednik” and “Kedrovaya Pad”), but she did not state explicitly which is the locality of the holotype.

Kyrgyzstan: Issyk-Kul Region: “Przewalsk” (Attems 1904) = Karakol.

Syntypes : 4 specimens, including 2 males and 2 females. Deposited in NHMW (Ilie et al. 2009).

An Escaryus species with body length reaching ≥ 4 cm; clypeus with small plagulae; labral arc relatively deep, with denticles long and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur, femur, and tibia with small denticles, tarsungulum with a small bulge; 49–55 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment trapezoid, approximately as long as wide; coxal pores on both ventral and lateral sides of coxopleura; anal pores absent.

Western Siberia: Altai krai, Republic of Altai, and Kemerovo oblast (Nefediev et al. 2017a, 2017c, 2018; Nefediev 2019). Possibly Far East (Titova 1972, 1973; see Remarks). Outside Asian Russia: westwards to Moldova (Titova 1973; Volkova 2016; Zuev 2016; Zuev and Evsyukov 2016); southwards to Kazakhstan and Kyrgyzstan (Attems 1904; Titova 1972, 1973; Dyachkov and Tuf 2018; Dyachkov 2022).

A total of 19 males and 26 females collected from Trans-Ili Alatau (Almaty Region of Kazakhstan) were indicated by Titova (1973: 110) as lectotypes, but this action is not valid, as the syntypes still exist (Ilie et al. 2009).

Titova (1972: 135; 1973: 116) wrote “... it is possible to distinguish territories from Cisamuria to Kuzbass and Altai, where 4 species are spread: E. koreanus, E. japonicus, E. retusidens, and E. chadaevae”. However, she did not mention material from this area when she listed the studied material of E. retusidens (Titova 1973: 110): “Kazakhstan, Trans-Ili Alatau… Moreover, E. retusidens were studied from the Dzhungarian Alatau, the Greater Caucasus, Crimea, Moldova, Rostov, Voronezh and Voroshilovograd oblasts”.

Russia: Sakhalin oblast: “Sachalin” = Sakhalin Isl. Japan: Hokkaido Isl.: “Sapporo” (both Takakuwa 1935).

Syntypes : 3 specimens. Depository unknown.

An Escaryus species with body length reaching ≥ 3 cm; clypeus without plagulae; labral arc relatively deep, with denticles short and obtuse; first maxillae with lappets; all forcipular articles with small denticles, except tarsungulum; 35–39 leg-bearing segments; metasternite of the ultimate leg-bearing segment rectangular, ~ 1.5× as long as wide; numerous coxal pores on ventral side of coxopleuron; anal pores present.

Far East: Sakhalin oblast (Sakhalin Isl.) (Titova 1973; Kurcheva 1977). Outside Asian Russia: Japan (Takakuwa 1935; Shinohara 1972) and northern China (Shinohara 1972).

Russia: Maritime krai: “Vladivostock” (Cook 1899) = Vladivostok city, 43°7'N, 131°54'E.

Syntypes : 12 specimens, both sexes, including 10 specimens deposited in ZMH (Thofern et al. 2021) and 2 specimens deposited in NHMW (Ilie et al. 2009).

An Escaryus species with body length reaching 6.5 cm; labral arc relatively shallow, with middle denticles obtuse and lateral ones long and pointed; first maxillae with two pairs of lappets; forcipular trochanteroprefemur with a small distal bulge, other forcipular articles without denticles; 49–51 leg-bearing segments; metasternite of the ultimate leg-bearing segment rectangular, ~ 2× as long as wide; coxal pores of different size, on both ventral and lateral sides, including a pair of much larger ventral pores on each coxopleuron; anal pores absent.

Far East: Maritime krai (Cook 1899; Attems 1904) and Amur oblast (Ganin 1997). Outside Asian Russia: no records.

Titova (1969) reported this species from Western Siberia (Kemerovo oblast) but later (Titova 1973) she regarded the same record as E. koreanus, even though she also suggested that the latter species can be a junior synonym of E. sibiricus. The anal pores were indicated as absent by Cook (1899) and Attems (1904) but illustrated as present in specimens identified as E. sibiricus by Thofern et al. (2021), and they are known to be present in E. koreanus.

Russia: Republic of Buryatia: “r. Vitim, Aniboli” (Titova 1973) = ? Anibud river, inflow of Vitim River, ca 53°39'N, 113°53'E.

Holotype : female. Paratypes: 3 specimens, including 1 male and 2 females. Deposited in ZMMU (S. Golovatch and A. Schileyko, pers. comm., 13.II.2023 and XII.2023).

An Escaryus species with body length reaching ≥ 2.7 cm; clypeus with large plagulae; labral arc relatively shallow, with denticles long and obtuse; first maxillae with one pair of lappets; all forcipular articles with small bulges; 37 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment rectangular, ~ 1.5× as long as wide; coxal pores on both ventral and lateral sides of coxopleura, the largest pores near the metasternite; anal pores present.

Eastern Siberia: Republic of Buryatia (Titova 1973; Alekseeva 1974). Outside Asian Russia: no records.