Papua New Guinea: Eastern Highlands Province, Aiyura, 06°21.131'S, 145°54.398'E, 1670 m a.s.l.

Holotype : male “3189” [green label], “Papua New Guinea: Eastern Highlands, Aiyura, 1670m, 5.iv.2006, 06.21.131S 145.54.398E, Balke & Sagata (PNG 32)” (ZSM). Paratypes: 2 males, 3 females with the same label as the holotype (NHMW, ZSM). 2 males “Papua New Guinea: Eastern Highlands, Aiyura creek, 1670m, 20.iv.2006, 06.21.131S 145.54.398E, John & Balke (PNG 70)” (ZSM).

Body size and form : Beetle large, oblong-oval to elongate (Fig. 29).

Measurements : TL 6.5–7.6 mm, TL-H 5.9–6.95 mm, MW 2.9–3.6 mm, TL/MW 2.11–2.24; PL 0.9–1.1 mm, PW 2.6–3.05 mm, PL/PW 0.35–0.36; DBE 1.15–1.3 mm, DBE/PW 0.43–0.44. Holotype: TL 7.6 mm, TL-H 6.95 mm, MW 3.6 mm, TL/MW 2.11; PL 1.1 mm, PW 3.05 mm, PL/PW 0.36; DBE 1.3 mm, DBE/PW 0.43.

Colouration : Dorsally piceous, usually with yellowish red head, pronotal sides and at elytral apex between striae (Fig. 29).

Head yellowish red to brown, narrowly darker behind eyes. Pronotum brown to piceous on disc and reddish to yellowish red on sides, especially at anterior angles; in teneral species, gradually paler (to yellowish red) to sides. Elytron brown to piceous, without basal band, sometimes with small, faint, reddish spot at shoulder, but broadly yellow between striae in apical 1/2, sometimes even to shoulder spot laterally, and reddish along suture. Scutellum reddish to brown. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs darker, all legs darker distally. Ventral side brown, yellowish red to reddish on head, prosternum, lower margin of metacoxae and medially and laterally on abdominal ventrites 1–4. Note: the holotype is a sequenced specimen; such specimens are usually darker.

Surface sculpture : Elytron with 11 complete striae, submarginal stria absent: 11+0 (Fig. 29).

Head usually without or with single strioles between eyes, with more or less regular, dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively coarse (usually diameter of punctures equal to diameter of microreticulation cells); microreticulation distinct. Pronotum with numerous strioles, with punctation finer than on head and microreticulation fine. Elytron with 11 complete dorsal striae; submarginal stria absent. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum narrowly rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws long, subequal in length. Median lobe of aedeagus robust, with two lobes of dorsal sclerite subequal in length, with pointed apexes and covered with scale-like structures; left dorsal lobe distinctly concave subapically, with long, narrow apical crest and apex slightly thickened at its tip, also with lateral side concave medially, above this concavity with small convex area covered with spinula-like structures; right dorsal lobe with large, elongate membranous area medially; lobes of ventral sclerite pressed together between lobes of dorsal sclerite, but partly visible in left lateral view; left ventral lobe with relatively broad, elongate sclerotised area, with pointed apex, situated in left part of lobe and mostly visible in left lateral view; membranous right part of left ventral lobe conjoined with sclerotised area and has apical tuft of long setae; right ventral lobe membranous, long and relatively broad. Parameres with dense, long setae occupying slightly more than half of dorsal margin (Fig. 80).

Female : Some females with more numerous strioles on head and pronotum than in males.

There is a variation in body shape, dorsal colouration and striolation described above.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures, e.g., A. centralensis sp. nov. and A. normanbyensis sp. nov. However, it is well-recognisable by its elytral colouration without basal band, 11+0 elytral striation, relatively narrow body shape, and median lobe with left dorsal lobe distinctly concave subapically and having convex area with spinula-like structures medially.

The species is named after Aiyura creek. The name is an adjective in the nominative singular.

Papua New Guinea: Eastern Highlands Province. The species is known only from the type locality (Fig. 107).

The species was collected in rest pools of a temporary stream.

Indonesia: West Papua Province: Manokwari Regency, Kebar, 0°48'27.9"S, 133°03'33.3"E, 584 m a.s.l.

Holotype : male “Indonesia: Papua Barat, Kebar, shaded deep sandy irritation roadside ditches, 584m, 6.xi.2013,”, “-0.80775253 133.05923529, UNIPA Team (BH031)” (MZB). Paratypes: 102 males, 78 females with the same label as the holotype, one male with an additional green text label “6249” (MZB, NHMW, ZSM). 1 male, 1 female “Indonesia: Papua Barat, Kebar Valley, 596m, 6.v.2015, -0,8406 133,2682, UNIPA Team (BH059)” (ZSM). 1 female “IN: West Papua: Manokwari Reg., on road Manokwari-Kebar, near Munbrani vill., ca. 600 m, 8.V., 00°46'21"S 133°22'53"E, roadside ditch (2015-WP36)” (NHMW). 5 males, 4 females “Indonesia: Papua, Manokwari, 140m, 8.ii.2006, 00.55.752S 133.54.448E, UNIPA Team (BH 09)”, one male with an additional green label “4248” (ZSM).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 57).

Measurements : TL 5.7–6.6 mm, TL-H 5.1–5.95 mm, MW 2.7–3.2 mm, TL/MW 2.06–2.12; PL 0.8–0.95 mm, PW 2.3–2.7 mm, PL/PW 0.35–0.36; DBE 0.85–1 mm, DBE/PW 0.37–0.38. Holotype: TL 6.15 mm, TL-H 5.55 mm, MW 2.9 mm, TL/MW 2.12; PL 0.9 mm, PW 2.5 mm, PL/PW 0.36; DBE 0.95 mm, DBE/PW 0.38.

Colouration : With piceous elytra and pronotum, yellowish red head, pronotal sides, and elytral basal band and yellowish between striae laterally (Fig. 57).

Head yellowish red medially, narrowly piceous behind eyes and brownish anteriorly. Pronotum piceous on disc and broadly yellowish red on sides, especially at anterior angles. Elytron piceous, with rather broad yellowish red basal band, usually extending between striae, especially laterally and yellowish between striae laterally and sometimes additionally apically, seldom whole elytron slightly yellowish, with distinct basal band. Scutellum reddish to piceous. Antennae, other head appendages, and pro- and mesolegs yellow, metalegs slightly darker proximally and distinctly distally. Ventral side yellowish red, darker on metaventrite, metacoxal plates and two last abdominal ventrites.

Surface sculpture : Elytron with 11 complete dorsal striae, submarginal stria present, long: 11+1 (Fig. 57).

Head without strioles, with relatively dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (diameter of punctures usually equal to or smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum with numerous, sparse strioles on lateral parts, only middle of disc without strioles, with punctation finer than on head and microreticulation fine. Elytron with 11 complete dorsal striae, striae 1 and 2, seldom 1–3, less strongly impressed dorsally; submarginal stria present, long, often reaching ½ of elytral length. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, strongly convex medially; blade of prosternal process narrow.

Male : Protibia straight, not modified. Proclaws relatively long, simple, subequal in length. Median lobe of aedeagus rather robust, apically thick, with two lobes of dorsal sclerite with broadly pointed apexes, more or less straight and subequal in length; left dorsal lobe with lateral margin not beaded, rather strongly concave at its apical 1/4; lateral side of left dorsal lobe completely covered with tiny spinulae situated in groups on scale-like structures; right dorsal lobe covered with large scale-like structures, with distinct median membranous area; lobes of ventral sclerite partly sclerotised, long, slightly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, with curved apex and slightly sclerotised part distinctly shorter, with dense setae apically; right ventral lobe longer than left one, partly sclerotised, with apex curved left and covered with setae sometimes distinctly sticking out in left lateral view. Parameres with dense, relatively long setae occupying slightly more than half of dorsal margin (Fig. 105).

Female : As males.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe completely covered by scale- and spinula-like structures in left lateral and ventral views. Based on body shape and size and general shape of the median lobe, it is similar and probably closely related to A. epicharis sp. nov. but differs from it in paler elytral colouration, more robust median lobe, and left dorsal lobe of the median lobe with apex thicker and lateral margin more strongly concave apically and therefore ventral lobes more visible in left lateral view. See also under A. yeretuar sp. nov.

The species name is a Greek adjective meaning “nice, pretty” and refers to the nice yellowish red dorsal pattern of the species.

Indonesia: West Papua Province: Manokwari Regency (Fig. 108).

The species was collected in shaded, deep, sandy irritation roadside ditches.

Papua New Guinea: Sandaun Province, Bewani Mts, 03°05.130'S, 141°10.227'E, 200–300 m a.s.l.

Holotype : male “Papua New Guinea: Sandaun, Bewani Stn., stream@base of Bewani Mts., 200–300m, 12.iv.2006, nr. 03.05.130S 141.10.227E, Balke & Sagata (PNG 37)” (MZB). Paratypes: PNG: Sandaun: 4 males, 2 females with the same labels as the holotype (MZB, NHMW, ZSM). 2 females “Papua New Guinea: Sandaun, Bewani Stn., forest puddles @ base of Bewani Mts., 300 m, 12.iv.2006, nr. 03.05.130S 141.10.227E, Balke & Sagata (PNG 38)”, one with an additional green label “3215” (ZSM). IN: Papua: Jayapura Regency: 1 male “Dutch New Guinea: Humbolt Bay [Yos Sudarso Bay] Dist. Pukusam Dist. West of Tami River. Vi.1937.” (BMNH). Sarmi Regency: 1 male, 1 female “Indonesia: Papua, Sarmi, Waaf, N Foja Mts, waterfall in forest, 120m, 23.ix.2014, -2,3317 138,7500, Tim UNCEN: Balke & Menufandu (Pap031)”, male with “6465” [green text] (ZSM). 2 males, 4 females “Indonesia: Papua, Foja Mountains N foot, N Waaf vill., pondok, 150m, 4.-7.vi.2016,”, “-2.06142 138.743949, Sumoked (Pap061)” (NHMW, ZSM). Biak Numfor Regency: 1 male “Indonesia: Papua, Biak, Nimbotong Nimbokramp, 110m 21.iv.2006, Tindige leg.”, “3157” [green label] (ZSM). 1 male “Indonesia: Papua, Biak, Nimbotong Nimbokramp, 110m 21.iv.2006, Tindige”, “1325” [green label] (ZSM). 3 males, 4 females “Indonesia: Papua, Biak, Nimbotong Nimbokramp, 110m 21.iv.2006, Tindige” (NHMW, ZSM).

IN: Papua: Pegunungan Bintang Regency: 1 male “Indonesia: Papua, Dekai, upper Brazza, 273m, 2./3.vi.2015, -4,741084724 139,654211075976, Sumoked (Pap044)” (ZSM). Note: a teneral specimen.

Body size and form : Beetle large, oblong-oval (Fig. 36).

Measurements : TL 6.9–7.8 mm, TL-H 6.35–7.1 mm, MW 3.25–3.6 mm, TL/MW 2.12–2.23; PL 0.95–1.15 mm, PW 2.85–3.2 mm, PL/PW 0.33–0.37; DBE 1.2–1.3 mm, DBE/PW 0.4–0.42. Holotype: TL 7.5 mm, TL-H 6.8 mm, MW 3.4 mm, TL/MW 2.21; PL 1.15 mm, PW 3.1 mm, PL/PW 0.37; DBE 1.3 mm, DBE/PW 0.42.

Colouration : Dorsally piceous, with yellowish red to reddish head, pronotal sides, and elytron with narrow, often faint yellowish red to reddish basal band and yellow apex (Fig. 36).

Head yellowish red to reddish, narrowly darker behind eyes. Pronotum piceous on disc and gradually paler (to yellowish red) to sides. Elytron piceous, with narrow yellowish red to reddish basal band, sometimes broadened between striae making base of elytron yellowish red or sometimes basal band faint, very narrow or even absent; elytral apex broadly or narrowly yellowish red, seldom broadened between striae making elytron yellowish red laterally or even in apical 1/2, striae brown to piceous. Scutellum yellowish red to brown. Antennae, other head appendages, and pro- and mesolegs yellowish red to reddish, metalegs darker, all legs darker distally. Ventral side reddish brown, paler on prosternum.

Surface sculpture : Elytron with 11 dorsal striae, stria 1 shortly reduced basally, submarginal stria present: 11+1 (Fig. 36).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (usually diameter of punctures equal to or smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum with few to several strioles posterolaterally, more numerous in females, with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae: stria 1 shortly reduced basally, seldom striae 1–3 reduced basally; submarginal stria present. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia more or less straight: its ventral margin slightly curved proximally. Proclaws long, subequal in length, anterior claw thicker subapically and more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus robust, with two lobes of dorsal sclerite covered with scale-like structures; left dorsal lobe without distinct median concavity and its whole apical 1/2 broad, more or less evenly tapering to small, curved downwards, slightly truncate apex bearing crest; scale-like structures of left dorsal lobe large in its apical 1/2; right dorsal lobe shorter than left one, broad, with pointed apex and membranous area medially; lobes of ventral sclerite partly sclerotised, pressed together between lobes of dorsal sclerite, not visible in left lateral view due to broad, not concave left dorsal lobe; left ventral lobe consists of two parts: left sclerotised area and less sclerotised right part; sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2; less sclerotised right part long and broad, with its apical part densely covered with long setae; right ventral lobe partly sclerotised, long and broad. Parameres with dense, long setae occupying slightly more than half of dorsal margin (Fig. 86).

Female : Trimorphic: 1) as male but with much more numerous pronotal strioles, 2) matt, with pronotum and elytron very densely covered with fine longitudinal strioles, and 3) less matt with strioles partly absent on pronotal and elytral discs, sometimes completely absent on elytron, but with strong dorsal punctation and microreticulation, especially on elytron.

There is a variation in dorsal colouration and striolation as described above. The male from Dekai (see Additional material) is teneral but most likely belongs to A. bewaniensis sp. nov. It is not A. dekai sp. nov. according to shape of the male proclaws and median lobe though the latter is difficult clearly to interpret due to weak sclerotisation. However, the specimen differs from all other specimens of A. bewaniensis sp. nov. in reduced elytral striation: dorsal stria 1 almost completely reduced, present only as single striole in apical 1/2, striae 3, 5, 7, 9 absent in basal 1/2 and present in apical one as striae or strioles. More material from the area is necessary to clarify specimen affiliation and range of elytral striation variability of the species.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is distinguished by rather dark dorsal colouration and 11+1 elytral striation, and, especially, left dorsal lobe of the median lobe without median concavity and its apical 1/2 covered with large scale-like structures, very broad, evenly tapering to small, curved downwards, slightly truncate apex bearing crest.

The species is named after Bewani Mountains. The name is an adjective in the nominative singular.

Indonesia: Papua Province and Papua New Guinea: Sandaun Province (Fig. 108).

The species was collected in the different forest water bodies: a stream, puddles, and near a waterfall.

Papua New Guinea: Southern Highlands Province, Bosavi Mt, 06°28'S, 142°50'E, 700 m a.s.l.

Holotype : male “Collection Naturhistorisches Museum Basel”, “Papua New Guinea S. Highlands Prov. L.Cizek lgt.”, “Mt Bosavi, 700m 142°50'E 6°28'S 20–27.VI.1999” (NHMB). Paratypes: 18 males, 16 females with the same labels as the holotype (NHMB, NHMW, ZSM).

Body size and form : Beetle large, oblong-oval (Fig. 31).

Measurements : TL 7.25–8.3 mm, TL-H 6.65–7.5 mm, MW 3.5–4.1 mm, TL/MW 2.02–2.07; PL 1.05–1.3 mm, PW 3.05–3.6 mm, PL/PW 0.34–0.36; DBE 1.2–1.4 mm, DBE/PW 0.39. Holotype: TL 8.2 mm, TL-H 7.4 mm, MW 3.95 mm, TL/MW 2.08; PL 1.25 mm, PW 3.45 mm, PL/PW 0.36; DBE 1.35 mm, DBE/PW 0.39.

Colouration : Dorsally piceous, with reddish head, pronotal sides and elytral basal band (Fig. 31).

Head dark yellowish red to reddish, narrowly darker behind eyes. Pronotum dark brown to piceous on disc and sometimes gradually paler (to dark yellowish red) to sides or sides broadly reddish, in contrast to piceous disc. Elytron dark brown to piceous, with narrow, dark yellowish red to reddish basal band differently developed: short, present in shoulder area, faint or distinct at whole base length. Scutellum reddish brown to piceous. Antennae, other head appendages, and pro- and mesolegs reddish, metalegs darker, all legs darker distally. Ventral side reddish brown, darker on posterior margins of abdominal ventrites 1–5 and on abdominal ventrite 6.

Surface sculpture : Elytron with 11 complete striae, submarginal stria present: 11+1 (Fig. 31).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures coarse (usually diameter of punctures larger than diameter of microreticulation cells); microreticulation distinct. Pronotum with strioles posterolaterally, with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae, stria 1 shortly or distinctly reduced basally, sometimes absent in basal 1/2; submarginal stria present. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia slightly modified: its ventral margin slightly curved proximally. Proclaws long and straight, subequal in length, anterior claw medially thicker than posterior one and due to this with a shallow subproximal incision on its inner margin. Median lobe of aedeagus very robust, with two lobes of dorsal sclerite covered with scale-like structures; left dorsal lobe very strongly concave medially, with apical 1/2 very broad and apex slightly rounded and bearing small but rather distinct crest; right dorsal lobe shorter, narrower, more or less evenly tapering to pointed apex, with small membranous area medially; lobes of ventral sclerite mostly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area and less sclerotised right part; sclerotised area with elongate and broad basal 1/2 (visible in left lateral view) and long, thread-like apical 1/2 (not visible); less sclerotised right part long and broad, with its apical part densely covered with long setae (partly visible in left lateral view); right ventral lobe mostly sclerotised, long and broad (partly visible in left lateral view). Parameres with dense, long setae occupying slightly more than half of dorsal margin (Fig. 82).

Female : As male.

There is an insignificant variation in dorsal colouration described above.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is very similar to A. robustus sp. nov. and differs from it in narrower, straighter male anterior claw with inner margin incision situated more proximally and distinctly broader apical 1/2 of the left dorsal lobe of the median lobe. See also under A. robustus sp. nov.

The species is named after Bosavi Mountain. The name is an adjective in the nominative singular.

Papua New Guinea: Southern Highlands Province, Bosavi Mt area (Fig. 107).

Unknown.

Papua New Guinea: Morobe Province, Herzog Mts, Bundun, 06°51.598'S, 146°37.07'E, 700–800 m a.s.l.

Holotype : male “Papua New Guinea: Morobe, Herzog Mts., Bundun, 700–800m, 2.iv.2006, 06.51.598S 146.37.07E, Balke & Sagata (PNG 27)” (ZSM). Paratypes: 16 males, 10 females with the same label as the holotype, three males additionally with green labels “3198”, “3199”, and “3228” (NHMW, ZSM). 1 male “3237” [green label], “Papua New Guinea: Morobe, Huon Pen., rd to Kwapsanek, 250m, 31.iii.2006, 06.30.270S 146.59.581E, Balke & Sagata (PNG 24)” (ZSM). 1 female “3208” [green label], “Papua New Guinea: Morobe, Huon Pen., rd to Kwapsanek, 460m, 31.iii.2006, 06.32.736S 146.59.616E, Balke & Sagata (PNG 26)” (ZSM).

Body size and form : Beetle large, with oblong-oval habitus (Figs 11, 12).

Measurements : TL 7.1–7.9 mm, TL-H 6.5–7.1 mm, MW 3.4–3.7 mm, TL/MW 2.08–2.14; PL 1–1.15 mm, PW 2.9–3.25 mm, PL/PW 0.34–0.35; DBE 1.2–1.3 mm, DBE/PW 0.4–0.41. Holotype: TL 7.7 mm, TL-H 7 mm, MW 3.7 mm, TL/MW 2.08; PL 1.1 mm, PW 3.2 mm, PL/PW 0.34; DBE 1.3 mm, DBE/PW 0.41.

Colouration : With piceous elytra and pronotum and yellowish red head, pronotal sides, and basal band on elytron (Figs 11, 12).

Head yellow to yellowish red, narrowly dark brown behind eyes. Pronotum dark brown to piceous on disc and gradually paler (to yellow) on sides. Elytron dark brown to piceous, with narrow yellow to yellowish red basal band, sometimes with yellowish red apex, seldom yellowish red line along suture too. Scutellum yellowish red to brown. Antennae, other head appendages, and pro- and mesolegs yellow to yellowish red, metalegs darker, all legs darker distally. Ventral side reddish brown.

Surface sculpture : Dorsal elytral striation very variable, submarginal stria usually absent or present as a short stria or few short strioles apically: (0–10)+(0–1) (Figs 11, 12).

Head without strioles, with relatively dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively coarse (diameter of punctures larger than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum with strioles laterally, with punctation and microreticulation slightly finer than on head. Elytron with striation very variable: from without striae, with three puncture lines, with up to ten dorsal striae (as in holotype), of which striae 3, 5–10 usually complete, striae 1 and 2 always strongly reduced basally; submarginal stria usually absent or present as a short stria or few short strioles apically. Elytron with punctation distinctly finer than on pronotum and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins distinctly convergent anteriorly. Base of prosternum narrowly rounded anteriorly, distinctly convex medially; blade of prosternal process relatively broad.

Male : Protibia more or less straight, not modified. Proclaws long, slightly curved, subequal in size and form. Median lobe of aedeagus robust, with two lobes of dorsal sclerite unequal in length and shape, both covered with scale-like structures; left dorsal lobe broader apically and shorter than right dorsal lobe, with a distinct median concavity and its whole apical 1/2 narrowed, its proximal part very large, and its apex more or less straight, with a distinct crest; right dorsal lobe with apex pointed and membranous area medially; lobes of ventral sclerite sclerotised, pressed together between lobes of dorsal sclerite, but partly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area (visible as a triangular in left lateral view) and less sclerotised right part (visible in left lateral view); sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2; less sclerotised right part long and broad, with its apical part densely covered with long setae; right ventral lobe long and broad. Parameres with dense, long setae occupying slightly more than half of dorsal margin (Fig. 64).

Female : Probably dimorphic: all females from the type locality (PNG27) are matt. The only female without this dense striolation is from the locality PNG26, where it was the only specimen collected. It is characterised by the reduced elytral striae (3 more or less complete dorsal striae, with some strioles between them) and presence of numerous strioles on the pronotal sides.

The species is very variable in the elytral striation, as described above, within and among populations. The holotype is with 10+1 elytral striae. Most specimens from the type locality (PNG27) have such striation but some of them have stria reduction, one specimen up to 5 dorsal striae. Only male from the locality PNG24 is without elytral striae. The female from the locality PNG26 is 3 more or less complete dorsal striae, with some strioles between them.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is similar to A. inconstans sp. nov. in size, body form, and dorsal colouration, but distinctly differs from it in shape of the median lobe and simple male proclaws. Also, the species is very similar to A. miltokarenos sp. nov. in median lobe shape but has some differences: apex of left dorsal lobe straighter, with stronger apical crest (in left lateral view) and right dorsal lobe with longer, more curved apex and ventral opening broader (in right lateral view). Additionally, A. bundunensis sp. nov. distinctly differs from A. miltokarenos sp. nov. by presence of a yellowish red basal band on elytron and its smaller, more oval body.

The species is named after its type locality, Bundun. The name is an adjective in the nominative singular.

Papua New Guinea: Morobe Province, Herzog Mts (Fig. 107).

The species was collected in small forest pools.

Papua New Guinea: Central Province, 09°25'47.5"S, 147°32'59.1"E, 755 m a.s.l.

Holotype : male “Papua New Guinea: Central, 755 m 28.x.2009, S9 25 47.5 E147 32 59.1, Sagata (PNG229)” (ZSM). Paratypes: PNG: Central: 10 males, 6 females with the same label as holotype (NHMW, ZSM). 1 male, 4 females “Papua New Guinea: Central, Moroka area, Kailaki Wareaga ridge, 768 m, 27.x.2009, S9 25 42.4 E147 31 06.8, Sagata (PNG227)” (ZSM). 2 males “Papua New Guinea: Central, Moreguina [10°00'57"S, 148°28'27"E] 16.viii.2008 Posman (PNG183) (ZSM). 1 male “Papua New Guinea: Central, Moreguina 18.viii.2008 Posman (PNG184)” (ZSM). 1 male, 1 female “Papua New Guinea: Central, Kokoda Trek, 320m, i.2008, [09°]19.236S 147 31.791E, Posman, (PNG 168)” (ZSM). 2 females “Papua New Guinea: Central, Kokoda Trek, 980m, i.2008, [09°]15.933S 147 36.590E, Posman, (PNG 169)” (ZSM). NCD: 2 males “Papua New Guinea: National Capital District, Varirata NP, 600m, 16.xii.2007, 09.26.13S 147.22.09E, Balke & Sagata (PNG 159)”, one male with an additional green label “2847” (ZSM).

Body size and form : Beetle large, with oblong-oval habitus (Fig. 20).

Measurements : TL 7–8 mm, TL-H 6.3–7.2 mm, MW 3.3–3.8 mm, TL/MW 2.11–2.12; PL 1–1.2 mm, PW 2.9–3.4 mm, PL/PW 0.35–0.36; DBE 1.15–1.35 mm, DBE/PW 0.4. Holotype: TL 7.6 mm, TL-H 6.8 mm, MW 3.6 mm, TL/MW 2.11; PL 1.1 mm, PW 3.1 mm, PL/PW 0.36; DBE 1.25 mm, DBE/PW 0.4.

Colouration : With piceous elytra and pronotum and yellowish red head, pronotal sides, and basal band on elytron (Fig. 20).

Head yellowish red to red, narrowly piceous behind eyes. Pronotum piceous on disc and gradually paler (to yellowish red) on sides, especially at anterior angles. Elytron piceous, with yellowish red to red basal band, which can be distinct or faint, narrower or broader, especially between striae, sometimes due to this elytron reddish laterally; often with yellowish brown apex. Scutellum piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs darker, all legs darker distally. Ventral side reddish brown medially and yellowish red on pro- and metaventrite and abdominal ventrites.

Surface sculpture : Dorsal elytral striation variable, usually with six dorsal striae and stria 1 reduced in basal 1/2, submarginal stria present: (6–10)+1 (Fig. 20).

Head without strioles, with relatively dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively coarse (diameter of punctures larger than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum with strioles laterally (few at posterolateral angles to rather numerous but sparse on sides), with punctation slightly denser and finer than on head and microreticulation fine. Elytron usually with six dorsal striae and some strioles between them, sometimes developed into complete and/or partly complete striae: up to ten dorsal striae; stria 1 always absent in basal 1/2; submarginal stria present, sometimes reduced to short apical strioles. Elytron with very fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins distinctly convergent anteriorly. Base of prosternum broadly rounded anteriorly, distinctly convex medially; blade of prosternal process relatively narrow.

Male : Protibia more or less straight, not modified; its ventral margin can be slightly curved proximally. Proclaws relatively short, subequal in length, anterior claw slightly thicker and more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus robust, with two lobes of dorsal sclerite distinctly narrowed to pointed apexes and covered with scale-like structures; left dorsal lobe longer than right lobe, slightly concave subapically and medially, with apex slightly curved downwards; right dorsal lobe with small membranous area medially; lobes of ventral sclerite sclerotised, pressed together between lobes of dorsal sclerite, but partly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area (not visible in left lateral view) and less sclerotised right part (visible in left lateral view); sclerotised area with elongate and broad basal 1/2 and long, hair-like apical 1/2; less sclerotised right part long and broad, with its apical part with a tuft of long setae; right ventral lobe long and broad. Parameres with dense, long setae occupying approximately half of dorsal margin; more distally situated setae longer than more proximal ones (Fig. 71).

Female : As male.

There is a variation in the elytral striation and colouration described above.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is similar to A. inconstans sp. nov. and A. bundunensis sp. nov. in size, body form, and dorsal colouration, but distinctly differs from them in median lobe with dorsal sclerite lobes distinctly narrowed to pointed apex and elytral striae always present.

The species is named after Central Province. The name is an adjective in the nominative singular.

Papua New Guinea: Central Province (Fig. 107).

Unknown.

Papua New Guinea: Simbu Province, Crater Mts, ca. 06°34'53.5"S, 145°05'26.7"E.

Holotype : male “386” [green label], “PNG: Crater Mts ii.2003, Sagata, MB 386” (ZSM).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 28).

Measurements: Holotype: TL 6.4 mm, TL-H 5.75 mm, MW 3.1 mm, TL/MW 2.07; PL 1 mm, PW 2.8 mm, PL/PW 0.36; DBE 1.2 mm, DBE/PW 0.43.

Colouration : Dorsally yellowish-reddish brown, with darker head and pronotum (Fig. 28).

Head brown, gradually paler (to reddish) anteriorly, narrowly darker behind eyes. Pronotum dark brown on disc and gradually paler (to reddish) to sides. Elytron uneven yellowish-reddish brown due to strongly developed yellowish red colouration between striae. Scutellum brown. Antennae, other head appendages, and pro- and mesolegs reddish, metalegs darker, all legs darker distally. Ventral side brown.

Surface sculpture : Elytron with 11 complete striae, submarginal stria absent: 11+0 (Fig. 28).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures coarse (diameter of punctures larger than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum with strioles on sides, with punctation finer than on head and microreticulation fine. Elytron with 11 complete dorsal striae, stria 1 shortly reduced basally; submarginal stria absent. Elytron with fine, relatively dense punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws long, slightly curved, subequal in length. Median lobe of aedeagus robust, with two lobes of dorsal sclerite unequal in length and shape, both covered with scale-like structures; left dorsal lobe more or less evenly tapering to apex, its apex straight, with small but distinct incision and crest; right dorsal lobe with apex pointed and membranous area medially; lobes of ventral sclerite, pressed together between lobes of dorsal sclerite, but partly visible in left lateral view, with sclerotised areas (since there is only the holotype, we could not dissect the median lobe to study structure of the ventral sclerite lobes in detail). Parameres with dense, long setae occupying distinctly more than half of dorsal margin (Fig. 79).

Female : Unknown.

The species is well distinguishable by its uniform, yellowish-reddish brown elytral colouration, 11+0 elytral striation and smaller body size from species with similar structures of the median lobe: usually, these species have TL > 6.5 mm, e.g., A. inconstans sp. nov.; additionally, the left dorsal lobe of the median lobe of A. craterensis sp. nov. has a very characteristic apex with small but distinct incision and crest.

The species is named after Crater Mountains. The name is an adjective in the nominative singular.

Papua New Guinea: Simbu Province. The species is known only from the type locality (Fig. 107).

Unknown.

Indonesia: Papua Province: Mimika Regency, 04°15'07.3"S, 136°38'36.2"E, 149 m a.s.l.

Holotype : male “7889” [green text], “Indonesia: Papua, Kabupaten Mimika, Timika, 149m, 25–30.v.2017,”, “-4.252020° 136.643384°, B.Sumoked (Pap68-Bob06)” (MZB).

Body size and form : Beetle medium-sized, elongate (Fig. 27).

Measurements: Holotype: TL 5.9 mm, TL-H 5.3 mm, MW 2.72 mm, TL/MW 2.17; PL 0.95 mm, PW 2.4 mm, PL/PW 0.4; DBE 1.05 mm, DBE/PW 0.53.

Colouration : Dorsally dark brown, with paler head, pronotal sides and elytral base and apex (Fig. 27).

Head reddish, narrowly piceous behind eyes and darker posteromedially. Pronotum dark brown on disc and gradually paler (reddish) on sides. Elytron dark brown, reddish basally, laterally and apically, also with faint reddish colouration between striae so that elytron not looking uniformly dark brown. Scutellum reddish brown. Antennae, other head appendages, and legs reddish, darker distally. Ventral side brown.

Surface sculpture : Elytron with 11 complete, relatively strongly impressed striae, submarginal stria present: 11+1 (Fig. 27).

Head without strioles, with distinct, dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively coarse (diameter of punctures slightly larger than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum without strioles, with punctation finer than on head and fine microreticulation. Elytron with 11 complete, relatively strongly impressed striae, submarginal stria present. Elytron with fine punctation and distinct microreticulation. Abdominal ventrite 6 with distinct punctation, sparse medially and forming denser area at each lateral side and with some strioles medially of this area.

Structures : Head large and broad. Pronotum relatively long, its lateral margins rounded, very slightly convergent anteriorly. Base of pronotum slightly narrower than base of elytron, therefore, habitus outline discontinuous. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia slightly modified; its ventral margin curved proximally. Proclaws relatively long, subequal in length, anterior claw slightly thicker and more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite unequal in length, covered with scale-like structures; left dorsal lobe with lateral margin slightly concave medially, with apex truncate and slightly curved downwards, having distinct crest; right dorsal lobe broad, shorter than left lobe, with pointed apex and membranous area medially; lobes of ventral sclerite mostly sclerotised, pressed together between lobes of dorsal sclerite, almost invisible in lateral view: tip of setation of less membranous part of left ventral lobe distinctly sticking out (since there is only the holotype, we could not dissect the median lobe to study structure of the ventral sclerite lobes in detail, see e.g., A. inconstans sp. nov.). Parameres with long and dense setae occupying approximately half of dorsal margin; more distally situated setae longer than more proximal ones (Fig. 78).

Female : Unknown.

The species is well-recognisable by its elongate, slightly discontinuous habitus, almost uniformly reddish brown dorsal colouration, strong elytral striation, and characteristic shape of the median lobe.

The species name is a Latin adjective meaning elegant and refers its elongate, slightly discontinuous habitus, with rounded pronotal sides.

Indonesia: Papua Province: Mimika Regency. The species is known only from the type locality (Fig. 107).

Unknown.

Indonesia: Papua Province: Yahukimo Regency, Dekai, upper Brazza, 04°44'27.9"S, 139°39'15.2"E, 273 m a.s.l.

Holotype : male “Indonesia: Papua, Dekai, upper Brazza, 273m, 2./3.vi.2015, -4,741084724 139,654211075976, Sumoked (Pap044)” (MZB). Paratypes: IN: Papua: Yahukimo Regency: 11 males, 13 females with the same label as the holotype, 2 males and 2 females with additional green text labels “7214”, “7215”, “7216” and “7217”, respectively (MZB, NHMW, ZSM). Pegunungan Bintang Regency: 4 males, 4 females “Indonesia: Papua, S Ok Sibil, tributary of Digul Riv, 359m, 9.vi.2015, -5,05718389526009 140,722535848617, Sumoked (Pap051)” (MZB, NHMW, ZSM). Mimika Regency: 1 male “Indonesia: Papua, Kabupaten Mimika, Timika, 149m, 25–30.v.2017, 04°15.092'S 136°38.597'E, B. Sumoked (Pap68-Bob06)” (ZSM). 1 male, 1 female “Indonesia: Papua, Kabupaten Mimika, Timika, 149m, 25–30.v.2017,”, “-4.252020° 136.643384°, B.Sumoked (Pap68-Bob06)”, with additional green text labels “7887” and “7888”, respectively (ZSM). PNG: SHL: 1 male, 1 female “Collection Naturhistorisches Museum Basel”, “Papua New Guinea S. Highlands Prov. L.Cizek lgt.”, “Mt Bosavi, 700m 142°50'E 6°28'S 20–27.VI.1999” (NHMB).

7 males “New Guinea XII-1942 W.G.Bodenstein” (ZSM). Note: teneral specimens.

Body size and form : Beetle large, oblong-oval (Fig. 32).

Measurements : TL 6.45–7.7 mm, TL-H 5.8–6.95 mm, MW 3.1–3.7 mm, TL/MW 2–2.08; PL 1–1.2 mm, PW 2.8–3.4 mm, PL/PW 0.35–0.36; DBE 1.15–1.35 mm, DBE/PW 0.39–0.41. Holotype: TL 7.2 mm, TL-H 6.5 mm, MW 3.6 mm, TL/MW 2; PL 1.15 mm, PW 3.2 mm, PL/PW 0.36; DBE 1.25 mm, DBE/PW 0.39.

Colouration : Dorsally piceous, with yellowish red to reddish head, pronotal sides and usually elytral basal band (Fig. 32).

Head yellowish red to reddish, narrowly darker behind eyes. Pronotum dark brown to piceous on disc and gradually paler (to yellowish red) to sides. Elytron brown to piceous, without basal band or with narrow, yellowish red to reddish basal band, often short, present in shoulder area, faint. Scutellum brown. Antennae, other head appendages, and pro- and mesolegs yellowish red to reddish, metalegs darker, all legs darker distally. Ventral side reddish brown.

Surface sculpture : Elytron with 10 or 11 differently reduced striae, submarginal stria present or absent: (10–11)+(0–1) (Fig. 32).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (usually diameter of punctures equal to or smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum without strioles, with punctation finer than on head and microreticulation fine. Elytron with 10 or 11 differently reduced dorsal striae: stria 1 usually completely reduced, sometimes more or less complete or present as strioles in apical 1/2, striae 2 and 3 usually reduced basally, striae 5, 7, and 10 usually interrupted in basal 1/2; submarginal stria absent or present, seldom as complete stria, usually as apical strioles. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process broad, weakly convex.

Male : Protibia slightly modified: its ventral margin slightly curved proximally. Proclaws long and straight, subequal in length, anterior claw thicker medially than posterior one. Median lobe of aedeagus robust, with two lobes of dorsal sclerite covered with scale-like structures; left dorsal lobe distinctly broader and longer than right dorsal lobe; left dorsal lobe with a distinct median concavity and its whole apical 1/2 narrowed, its apex straight, without distinct crest, sometimes with a weak crest due to small, very shallow tip concavity, more or less evenly rounded; right dorsal lobe with apex pointed and membranous area medially; lobes of ventral sclerite sclerotised, pressed together between lobes of dorsal sclerite, but visible in left lateral view; left ventral lobe consists of two parts (partly visible in left lateral view): left sclerotised area and less sclerotised right part; sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2; less sclerotised right part long and broad, with its apical part densely covered with long setae; right ventral lobe partly sclerotised, long and broad, curved at apex. Parameres with long and dense setae occupying approximately half of dorsal margin; more distally situated setae longer and denser than more proximal ones (Fig. 83).

Female : Dimorphic: as male, shiny, and matt, with pronotum and elytron very densely covered with fine longitudinal strioles. Matt forms present in two populations, more numerous than shiny ones: e.g., ratio shiny to with strioles is 11:2 in the type locality. Only matt forms are known from the locality “Pap051”.

There is a variation in dorsal colouration and striolation described above.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is similar to A. inconstans sp. nov., A. bundunensis sp. nov. and A. miltokarenos sp. nov. but differs from them in smaller body size, male anterior proclaw long, straight but thicker medially than posterior one and narrower apical 1/2 of the left dorsal lobe of the median lobe.

The species is named after its type locality, Dekai Village. The name is a noun in the nominative singular standing in apposition.

Indonesia: Papua Province and Papua New Guinea: Southern Highlands Province (Fig. 107).

The species was collected side pools of a forest stream.

Indonesia: Papua Province: Nabire Regency, Nabire, Wanggar, Kali Bumi.

Holotype : male “W.-Neuguinea /Paniai Prov./ Wanggar- Kali Bumi / IR 14 30.9 & 1.10.90 leg: Balke & Hendrich” (ZSM). Paratypes: IN: Papua: Nabire Regency: 14 males, 20 females with the same label as the holotype (CLH, ZSM). 3 males, 8 females “IR 90#14: West New Guinea, Nabire→Wanggar, 100m, 30.ix.1990, Balke” (ZSM). 2 males, 8 females “Irian: Nabire – Ikaga [sic! Ilaga], “KM 35”, Seitenstr. nach K. Cemara, 1991, leg. M. Balke” (ZSM). 8 males, 8 females “West New Guinea/Paniai Prov./IR 23 track Nabire-Ilaga km 34 near Topo, 120m, 23.7.1991 leg: Balke & Hendrich” (ZSM). 4 males, 1 female “IRIAN JAYA: Nabire Prov. rd. Nabire – Ilaga, Km 35 Kali Cemara, 100m, 27.9.1997 (IR97#6)” (NHMW). 7 males, 6 females “IRIAN JAYA: Nabire Prov. rd. Nabire – Ilaga, Km 35 Kali Cemara, 100m, 23.10.1997 (IR97#14)” (NHMW). Mimika Regency: 1 female “Indonesia: Papua, Kabupaten Mimika, Timika, 149m, 25–30.v.2017, 04°15.092'S 136°38.597'E, B. Sumoked (Pap68-Bob06)” (ZSM). 9 males, 4 females “Indonesia: Papua, Kabupaten Mimika, 24m, 25–30.v.2017, 04°30.330’”, “E 136°46.53’, B. Sumoked (Pap69-Bob07)” (ZSM). Yahukimo Regency: 2 males, 4 females “Indonesia: Papua, Dekai, upper Brazza, 273m, 2./3.vi.2015, -4,741084724 139,654211075976, Sumoked (Pap044)”, one male with an additional green text label “7221” (ZSM). Aru Islands Regency: 2 males “INDONESIA: Aru Islands, Trangan, 1km E of Ngalgull, 6°48'S 134°4'E, 29.vii.1994, 90m, A.H. Kirk-Spriggs.”, “A.H.Kirk-Spriggs Maluku Tenggara Coll. NMW.Z.1994.061”, “Temporary/Sago pools, Primary mixed forest.” (CLH, ZSM). 1 female “INDONESIA: Aru Islands, Trangan, 1km E of Ngalgull, 6°48'S 134°4'E, 23–28.vii.1994, 90m, A.H. Kirk-Spriggs.”, “A.H.Kirk-Spriggs Maluku Tenggara Coll. NMW.Z.1994.061”, “Temporary/Sago pools, Primary mixed forest.” (CLH, ZSM). 1 male “INDONESIA, SE Moluccas Aru Isls, Wokam Isl. 17 km NE Wakua vill. 1.-7.ii.2022, S. Jákl leg.”, “coll. Jiří Hájek National Museum Prague, Czech Republic” (NMPC). 1 male “INDONESIA, SE Moluccas ARU ISLS, WOKAM I. 17 km NE Wakua vill. 1–7.II.2022, St. Jakl leg.”, “coll. Jiří Hájek National Museum Prague, Czech Republic” (NMPC). West Papua: Kaimana Regency: 1 male, 3 females “INDONESIA W-PAPUA 50km SE Kaimana, Triton bay, vic. Kamaka vill. S3°49'50”/E134°11'27”, 10–50m, 02.-05.II.2011 A. Skale (006)” (ZSM). 1 male “INDONESIA: W-PAPUA vic. Kaimana, road 17 km NE, S3°31'41”/E133°40'51”, 50m, 31.I.2011 A Skale (003)”, “4431” [green label] (ZSM). 47 males, 60 females “IRIAN JAYA: Fak Fak dist. Lake Yamur area, IV.1998 ca. 50 – 100m, Waldtümpel” (NHMW). Raja Ampat Regency: Waigeo: 2 males, 3 females “W-PAPUA Raja Ampat Prov. Waigeo Isl., Lopintol 0°07'54"S, 130°53'45"E 11.i.2004 leg. A.Skale UWP” (CAS). 1 male “760” [green label], “W Papua: Raja Ampat, Waigeo, Lopintol, 11/2005, A. Skale, MB760” (ZSM). 1 female “762” [green label], “W Papua: Raja Ampat, Waigeo, Lopintol, 11/2005, A. Skale, MB 762” (ZSM). Batanta: 1 male “W-PAPUA Raja Ampat Pr. Yensawai Batanta, 9 km W Ross-River 0°49'23"S 130°35'52"E 17.I.2004 leg. A.Weigel UWP KL” (CAS). 1 male, 1 female “Indonesia: Papua, Batanta Selatan, Wailebet, 100m, 17.ii.2006, inland 00.53.957S 130.39.951E, Tindige & Prativi (BH 15)”, with additional green labels “4252” and “4253” (ZSM). 1 male “Indonesia: Papua, Batanta Selatan, Wailebet, 20m, 16.ii.2006, 00.54.003S 130.39.296E (BH 14)” (ZSM). 6 males, 2 females “Indonesia: Papua, Batanta Utara, 20m, 14.ii.2006, 00.50.125S 130.42.856E (BH 12)” (ZSM). 1 male, 1 female “W-PAPUA, Raja Ampat Pr. Waywesar/Batanta 2 km E, 0°45'17"S 130°48'06"E 18.I.2004, leg. A. Weigel” (CAS). 11 males, 21 females “W-PAPUA Raja Ampat Prov. Batanta Isl. bor., 9 km W Yensawai, Ross-River 0°49'23"S, 130°35'52"E 17.I.2004 leg. A. Skale” (CAS, NHMW). 4 males “W-PAPUA Raja Ampat Prov. Batanta Isl. bor., Yensawai, 0°48'05"S, 130°40'36"E 15.-18.I.2004 leg. A.Skale” (CAS, NHMW). 1 female “W-PAPUA Raja Ampat Prov. Batanta Isl. bor., Arefl, 0°47'24,5"S, 130°42'10"E 16.I.2004 leg. A.Skale” (CAS). 7 males, 2 females “W-PAPUA Raja Ampat Prov. Batanta Isl. bor., Waywesar, 0°45'26"S, 130°46'55"E 12.-15.I.2004 leg. A.Skale UWP” (CAS, NHMW). Salawati: 2 males “W-PAPUA Raja Ampat Prov. Salawati Isl. Or., 2–4 km N Kalobo 01°00'56"S, 131°04'58"E 28.I.2004 leg. A.Skale” (CAS). 1 male, 1 female “W-PAPUA Raja Ampat Prov. 1 km E Kalobo, Wajir Island 01°00'S, 131°04'E 26.I.2004 leg. A.Skale UWP/UWS” (CAS). Misool: 23 males, 19 females “Indonesia: West Papua, Raja Ampat, Misool, v.2015, -1.799575° 129.950446, Prativi” (NHMW, ZSM). PNG: SHL: 4 males, 1 female “Collection Naturhistorisches Museum Basel”, “Papua New Guinea S. Highlands Prov. L.Cizek lgt.”, “Mt Bosavi, 700m 142°50'E 6°28'S 20–27.VI.1999” (NHMB, NHMW, ZSM). Central: 6 males, 5 females “Papua New Guinea: Central, Tapini, 870m, 29.x.2007, 08.20.511S 146.59.824E Kinibel (PNG 161)”, one male and one female additionally with green labels “2854” and “2853”, respectively (NHMW, ZSM).

2 males (teneral) “Papua New Guinea: Southern Highlands Province, Tari to Komo, Hides Gas basecamp, 1200m, 13.v.2006, 05.55.223'S 142.46.090'E, Balke (PNG 60)”, one male additionally with a green label “3231” (ZSM).

Body size and form : Beetle small to medium-sized, oblong-oval (Fig. 56).

Measurements : TL 5.1–6.35 mm, TL-H 4.55–5.8 mm, MW 2.45–3.1 mm, TL/MW 2.04–2.8; PL 0.8–0.95 mm, PW 2.1–2.65 mm, PL/PW 0.35–0.38; DBE 0.85–1.1 mm, DBE/PW 0.4–0.42. Holotype: TL 5.6 mm, TL-H 5.1 mm, MW 2.75 mm, TL/MW 2.04; PL 0.85 mm, PW 2.4 mm, PL/PW 0.35; DBE 0.95 mm, DBE/PW 0.4.

Colouration : With piceous elytra and pronotum and yellowish red head, pronotal sides, and elytral basal band and apical spot (Fig. 56).

Head yellowish red medially, narrowly piceous behind eyes and brownish anteriorly. Pronotum piceous on disc and broadly yellowish red on sides, especially at anterior angles. Elytron piceous, with distinct yellowish red basal band, usually not reaching shoulders and suture; elytron usually with yellow spot apically sometimes slightly extending laterally. Scutellum reddish to piceous. Antennae, other head appendages, and pro- and mesolegs yellow, metalegs slightly darker proximally and distinctly distally. Ventral side reddish brown, darker on metaventrite, metacoxal plates and two last abdominal ventrites.

Surface sculpture : Elytron with 11 complete dorsal striae, submarginal stria present, long: 11+1 (Fig. 56).

Head without strioles, with relatively dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (diameter of punctures usually equal to and smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum with several or rather numerous, sparse strioles on lateral parts, sometimes only middle of disc without strioles, with punctation finer than on head and microreticulation fine. Elytron with 11 complete dorsal striae, sometimes strongly impressed, sometimes less, sometimes stria 1 or striae 1 and 2, seldom 1–3, less strongly impressed dorsally; submarginal stria present, long, often reaching ½ of elytral length. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, strongly convex medially; blade of prosternal process narrow.

Male : Protibia straight, not modified. Proclaws relatively long, simple, subequal in length. Median lobe of aedeagus with two lobes of dorsal sclerite with broadly pointed apexes, more or less straight and subequal in length; left dorsal lobe with lateral margin not beaded, slightly concave its apical 1/2; lateral side of left dorsal lobe completely covered with tiny spinulae situated in groups on scale-like structures; right dorsal lobe covered with large scale-like structures, with distinct median membranous area; lobes of ventral sclerite partly sclerotised, long, slightly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, with curved apex and slightly sclerotised part distinctly shorter, with dense setae apically; right ventral lobe longer than left one, partly sclerotised, with apex curved left and covered with setae sometimes distinctly sticking out in left lateral view. Parameres with long and dense setae occupying approximately half of dorsal margin; more distally situated setae longer and denser than more proximal ones (Fig. 106).

Female : As males.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe completely covered by scale- and spinula-like structures in left lateral and ventral views. Based on body shape and size and general shape of the median lobe, it is similar and probably closely related to A. asteios sp. nov. but differs from it in paler elytral colouration, more robust median lobe, and left dorsal lobe of the median lobe with apex thicker and lateral margin more strongly concave apically and therefore ventral lobes more visible in left lateral view. See also under A. yeretuar sp. nov.

Austrelatus epicharis sp. nov. is the second copelatine species described from the Aru Islands. The first one is Copelatus haemorrhoidalis Régimbart, 1883, which was described based on a single female. We have studied habitus photos of the holotype (RMNH.INS.1487394), which is deposited in the Naturalis Biodiversity Center (former Rijksmuseum van Natuurlijke Historie), Leiden, Netherlands (RMNH), and compared them with images and specimens of A. epicharis sp. nov. We believe that they are two different species since 1) thought C. haemorrhoidalis has 11+1 elytral striae, striae 2, 4, 6, 8 are reduced in the apical 1/2, and A. epicharis sp. nov. never demonstrates elytral stria reduction; 2) C. haemorrhoidalis has less prominent yellowish red dorsal colouration and a V-shaped brown spot on head, but head of A. epicharis sp. nov. is always uniformly yellowish red medially (Figs 56, 58). Most likely, C. haemorrhoidalis belongs to the genus Austrelatus but it will be possible to determine with certainty when males of the species are found. Besides these two species, there are two more copelatine species found in the material recently collected from the Aru Islands. They are represented by three females, two of which belong to the A. neoguineensis group and one is most likely a Copelatus species. Thus, Copelatinae fauna of the Aru Islands includes four different species with the certain record of A. epicharis sp. nov. For three other species, more material is necessary to clarify their taxonomic position.

The species name is a Greek adjective meaning “beautiful, graceful” and refers to striation and nice yellowish red dorsal pattern of the species.

The species is widely distributed in Indonesian part of New Guinea including the small islands; it is also known from Central and SHL provinces of PNG (Fig. 108).

All specimens on the track Nabire to Ilaga were collected in shallow (up to 20 cm water depth), shaded or at least partly shaded forest pools and road-side ditches, rich in rotten leaves and twigs (Figs 109, 110). The type specimens were collected in shallow, exposed and flooded meadow puddles and oxbows along Kali (= River) Bumi at Wanggar (Fig. 115). Few specimens were also found in water-filled track hollows. On the Aru Islands, the species was collected in temporary Sago swamp pools, surrounded by primary mixed forest.

Papua New Guinea: East Sepik Province, Lembena, 04°56.859'S, 143°57.379'E, 335 m a.s.l.

Holotype : male “Papua New Guinea: East Sepik, Lembena, 335m, 10.ix.2009, 04 56.859S 143 57.379E, Ibalim & Pius (PNG251)” (ZSM). Paratypes: 3 males, 2 females with the same label as the holotype (NHMW, ZSM). 3 males, 3 females “Papua New Guinea: East Sepik, Lembena, 198m, 3.ix.2009, 04.56.974S 143.56.995E, Ibalim & Pius (PNG241)” (NHMW, ZSM). 3 females “Papua New Guinea: East Sepik, Lembena, 198m, 3.ix.2009, 04 46 [!] 974S 143.56.995E, Ibalim & Pius (PNG243)” (ZSM). 2 males “Papua New Guinea: East Sepik, Lembena, 136m, 3.ix.2009, 04 56.911S 143.56.870E, Ibalim & Pius (PNG244)” (ZSM). 1 male, 6 females “Papua New Guinea: East Sepik, Lembena, 117m, 8.ix.2009, 04.57.513S 14357.296E, Ibalim & Pius (PNG248)”, one male and four females with additional green text labels “6001”, “5999”, “6000”, “6002”, and “6003”, respectively (ZSM). 1 male, 2 females “Papua New Guinea: East Sepik, Lembena, 110m, 10.ix.2009, 04.57.512S 143.57.366E, Ibalim & Pius (PNG249)”, male with a green text label “6033” (ZSM). 2 females “Papua New Guinea: East Sepik, Lembena, 335m, 10.ix.2009, 04.56.859S 143.59.375E, Ibalim & Pius (PNG250)” (ZSM).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 46).

Measurements : TL 5.4–6.1 mm, TL-H 5–5.4 mm, MW 2.65–2.9 mm, TL/MW 2.04–2.1; PL 0.85–0.9 mm, PW 2.3–2.5 mm, PL/PW 0.36–0.37; DBE 0.9–0.95 mm, DBE/PW 0.38–0.39. Holotype: TL 5.5 mm, TL-H 5 mm, MW 2.7 mm, TL/MW 2.04; PL 0.85 mm, PW 2.35 mm, PL/PW 0.36; DBE 0.9 mm, DBE/PW 0.38.

Colouration : With piceous elytra and pronotum and yellowish red head, pronotal sides, and basal band and large apical spot on elytron (Fig. 46).

Head yellowish red, narrowly piceous behind eyes. Pronotum piceous on disc and yellowish red on sides, especially at anterior angles. Elytron piceous, with distinct, rather broad, yellowish red basal band and large, often laterally extending yellow spot on apex. Scutellum reddish to piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs distinctly darker, especially distally. Ventral side reddish brown to brown, paler on pro- and metaventrite and abdominal ventrites 1–3.

Surface sculpture : Elytron with (10–11)+1 striae, sometimes striae differently reduced or interrupted (Fig. 46).

Head without strioles, with rather dense punctation (spaces between punctures 1–3× size of punctures); punctures rather coarse (diameter of punctures usually equal to diameter of microreticulation cells); microreticulation distinct. Pronotum without or with few strioles posterolaterally, with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae, stria 1 differently reduced, often absent in basal 1/2 and present as few strioles in apical 1/2, seldom completely absent, striae 2 and 3 reduced or interrupted basally, striae 5–7, 9 and 10 sometimes shortly reduced or interrupted basally; submarginal stria present, sometimes reduced to few strioles apically. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum broadly rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively long, slightly curved, subequal in length, anterior claw thicker subapically than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, more or less straight, with very broadly pointed apexes, apex of left dorsal lobe slightly curved downwards; left dorsal lobe with lateral side covered with numerous, distinct, strong spinula-like structures; lateral margin apically without surface structures, smooth; right dorsal lobe covered with scale-like structures, with distinct median membranous area; lobes of ventral sclerite partly sclerotised, long, subequal in length, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, concave subapically, with apex pointed and slightly curved to right and membranous right part long, thin, apically covered with setae sometimes visible in left lateral view; right ventral lobe with narrow sclerotised area on right margin. Parameres with long and dense setae occupying approximately half of dorsal margin; more distally situated setae longer and denser than more proximal ones (Fig. 95).

Female : As males but with stronger dorsal punctation and microreticulation and usually more numerous pronotal strioles.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe not completely covered by spinula-like structures, upper lateral margin without spinulae, smooth in left lateral and ventral views. Based on shape of the median lobe, it is similar to A. mianminensis sp. nov. and especially to A. pseudomianminensis sp. nov. (in having lateral margin of left dorsal lobe with edge, below which spinula-like structures situated, more sharp, prominent) but differs from them in more bright dorsal colouration: yellowish red head and lateral sides of pronotum, elytron with broad yellowish red basal band and large apical spot, as well as in less striolated elytron: (10–11)+1 striae, stria 1 strongly reduced basally (visible in traces in apical 1/2), sometimes completely absent, the other striae can be differently reduced. See also under A. maindai sp. nov. and A. kokodensis sp. nov.

The species name is a combination of the Latin words flavus (yellow) and capitatus (heaving a head) and refers to the distinct yellowish head colouration of the species. The species name is an adjective in the nominative singular.

Papua New Guinea: East Sepik Province. The species is known only from Lembena area (Fig. 108).

Unknown.

Papua New Guinea: Central Province, Kokoda, 08°53.481'S, 147°43.648'E, 410 m a.s.l.

Holotype : male “Papua New Guinea: Northern, Kokoda, 410m, i.2008, 53.481S 147.43.648E, Posman, (PNG 174)” (ZSM).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 50).

Measurements: Holotype: TL 6.4 mm, TL-H 5.8 mm, MW 3.1 mm, TL/MW 2.07; PL 0.9 mm, PW 2.7 mm, PL/PW 0.33; DBE 0.95 mm, DBE/PW 0.35.

Colouration : Dorsally almost uniformly piceous (Fig. 50).

Head dark brown, reddish posteriorly between eyes and anteromedially, narrowly piceous behind eyes. Pronotum piceous, dark brown on sides, reddish at anterior angles. Elytron piceous, yellowish red apically, slightly extending laterally. Scutellum piceous. Antennae, other head appendages, and pro- and mesolegs reddish, metalegs distinctly darker, especially distally. Ventral side piceous, with paler prosternum.

Surface sculpture : Elytron with 10+1 striae; submarginal stria present as weak apical strioles (Fig. 50).

Head without strioles, with rather dense punctation (spaces between punctures 1–3× size of punctures); punctures rather coarse (diameter of punctures equal to diameter of microreticulation cells); microreticulation distinct. Pronotum with single strioles at posterior angles, with punctation slightly finer than on head and microreticulation fine. Elytron with ten dorsal striae, striae 1 and 2 absent in basal 1/3, striae 4 and 5 can be interrupted basally, stria 10 reduced and interrupted basally; submarginal stria present only as very weak apical strioles. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively short, subequal in length, anterior claw thicker and more curved subapically than posterior one and due to this with a subapical incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, with broadly pointed apexes; whole lateral side of left dorsal lobe with large but shallow concavity, covered with numerous large spinula-like structures getting longer and denser towards lateral margin and shorter towards dorsal side where they replaced by scale-like structures; lateral margin slightly concave apically so that apex of left dorsal lobe slightly curved upwards; right dorsal lobe covered with large scale-like structures, with small, indistinct median membranous area; lobes of ventral sclerite partly sclerotised, long, subequal in length, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, concave subapically, with apex pointed and slightly curved to right and membranous right part long, thin, apically covered with setae visible in left lateral view; right ventral lobe with narrow sclerotised area on right margin. Parameres with dense, rather short setae occupying more than half of dorsal margin (Fig. 99).

Female : Unknown.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe completely covered by scale- and spinula-like structures in left lateral and ventral views. Based on body size and dark dorsal colouration, it is similar to A. kalibumi sp. nov., A. herzogensis sp. nov., A. mianminensis sp. nov., and A. pseudomianminensis sp. nov. but differs from them in left dorsal lobe of the median lobe with large, shallow concavity and surface of left dorsal lobe completely covered by surface structures. It is also similar to A. sararti sp. nov. but differs from it in larger body size, elytron with 10, not 11, dorsal striae, and left dorsal lobe of the median lobe with concavity and large, long spinula-like structures. See also under A. wasiorensis sp. nov.

The species name is a Latin adjective meaning “dark, dusky” and indicates the dark elytral colouration of the species.

Papua New Guinea: Central Province. The species is known only from the type locality (Fig. 108).

Unknown.

Papua New Guinea: Central Province, Rigo District, ca. 09°46′51.6″S, 147°50′24″E.

Lectotype : male “N. GUINEA MER. RIGO Luglio 1889 L.Loria”, “Neu Guinea Mus. Genua”, “Copelatus Gestroi Reg.” [hw], “Coll. Mus. Vindob.” (NHMW). Paralectotypes: 2 males, 6 females “N. GUINEA MER. RIGO Luglio 1889 L.Loria”, “Neu Guinea Mus. Genua” (NHMW). 1 male “N. GUINEA MER. RIGO Luglio 1889 L.Loria”, “Neu Guinea Mus. Genua”, “Copelatus Gestroi Reg.” [hw] (NHMW). 1 male “N. Guinea Rigo Luglio 1889 L.Loria” [hw], “Neu Guinea Mus. Genua” (NHMW). 2 males “N. GUINEA MER. RIGO Luglio 1889 L.Loria”, “Cotype” [red], “Regimbart det., 1891: COPELATUS (s. str.) Gestroi – Reg.” [partly hw] (ZSM). 2 males “N. GUINEA MER. RIGO Luglio 1889 L.Loria”, “Neu Guinea 7407” [partly hw], “Copelatus Gestroi Rg det. Gschwendt.” [hw], “Staatl. Museum für Tierkunde, Dresden” (MTD). 2 females “N. GUINEA MER. RIGO Luglio 1889 L.Loria”, “Neu Guinea 7407” [partly hw], “Staatl. Museum für Tierkunde, Dresden” (MTD). 3 males, 2 females “N. GUINEA MER. RIGO Luglio 1889 L.Loria”, “Neu Guinea 8947” [partly hw], “Staatl. Museum für Tierkunde, Dresden” (MTD). 1 male “N. GUINEA MER. RIGO Luglio 1889 L.LORIA”, “Regimbart det., 1891: COPELATUS (s. str.) Gestroi – Reg.” [partly hw], “EX TYPUS” (IRSNB). 21 exs. “N. GUINEA MER. RIGO Luglio 1889 L.Loria” (MNHN); these paralectotypes are from the collections of Régimbart (1 ex.), Legros (1 ex.), Guignot (2 ex.), Oberthür (17 exs.), some of them with the hand written labels indicating C. gestroi; median lobes of the specimens were not studied. 1 male “N. GUINEA MER. KELESI Nov. Dic. 1890 L.LORIA”, “MUSEUM PARIS COLL MAURICE REGIMBART 1908”, “MNHN, Paris EC29239 [with QR code]” (MNHN).

1 male “Neu Guinea” [hw by ?Zimmermann], “Samml. A. Zimmermann” (ZSM). 1 male [only aedeagus glued on pined triangular, beetle probably lost] “Penis C. gestroi” [hw by ?Zimmermann], “Samml. A. Zimmermann” (ZSM). PNG: NCD: 1 male “2846”, “Papua New Guinea: National Capital District, Varirata NP, 600m, 16.xii.2007, 09.26.13S 147.22.09E, Balke & Sagata (PNG 159)” (ZSM). 4 males, 5 females “Stn. No. 196”, “Papua: Pt. Moresby Brown R. Rd., 15.III.1965”, “M.E. Bacchus. B.M. 1965-120” (BMNH, NHMW). 1 male, 4 females “Stn. No. 195”, “Papua: Pt. Moresby Brown R. Rd., 15.III.1965”, “M.E.Bacchus, B.M. 1965-120” (BMNH). 2 males, 3 females “Stn. No. 206”, “Papua: Loloki c. 10 m. N. of Pt. Moresby, 19.III.1965.”, “M.E. Bacchus. B.M. 1965-120” (BMNH). 4 males, 10 females “Stn. No. 208”, “Papua: Loloki, c. 10 m. N. of Pt. Moresby 19.III.1965.”, “M.E. Bacchus. B.M. 1965-120” (BMNH, NHMW). 5 males, 3 females “Stn. No. 207a”, “Papua: Moitaka, c. 7 m. N. of Pt. Moresby. 17.III.1965”, “M.E. Bacchus. B.M. 1965-120” (BMNH, NHMW). 1 male, 1 female “4881 New Guinea Port Moresby Dist 24.iii.1956 E.S. Brown” [partly hw], “Pres. By Com.Inst.Ent. B.M. 1958-79” (BMNH). Morobe: 2 males, 1 female “Stn. No. 103”, “New Guinea: Morobe Dist., Finisterre Mts., Mt. Abilala, c.9,000 ft. 19–22.xi.1964.”, “M.E. Bacchus. B.M. 1965-120” (BMNH).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 52).

Measurements : TL 5.8–6.8 mm, TL-H 5.45–6.2 mm, MW 2.9–3.3 mm, TL/MW 2–2.06; PL 0.85–0.95 mm, PW 2.45–2.8 mm, PL/PW 0.34–0.35; DBE 0.9–1 mm, DBE/PW 0.35–0.37. Lectotype: TL 6.4 mm, TL-H 5.8 mm, MW 3.15 mm, TL/MW 2.03; PL 0.9 mm, PW 2.6 mm, PL/PW 0.35; DBE 0.9 mm, DBE/PW 0.35.

Colouration : With piceous elytra and pronotum and yellowish red posterior margin of head, pronotal sides, and elytral basal band and apex (Fig. 52).

Head almost completely brownish or piceous, often paler, to yellowish red posteriorly and anteriorly and darker medially. Pronotum dark brown to piceous on disc and broadly yellowish red on sides, especially at anterior angles. Elytron dark brown to piceous, with yellowish red basal band differently developed: distinct, relatively broad or rather narrow, often notched at posterior margin (expending between striae), seldom reduced near suture and distinct at shoulders; elytron broadly yellow apically, this yellow colouration usually extending between striae, especially laterally (up to elytral 1/2). Scutellum reddish to piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs darker, especially distally. Ventral side reddish brown to brown, darker on coxal plates and abdominal ventrites 4–6.

Surface sculpture : Elytron with (10–11)+1 striae (Fig. 52).

Head without strioles, with rather dense punctation (spaces between punctures 1–3× size of punctures); punctures rather coarse (diameter of punctures usually equal to and larger than diameter of microreticulation cells); microreticulation distinct. Pronotum with few to several strioles at posterior angles or more numerous laterally (in females), with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae, stria 1 usually reduced to strioles in apical 1/2, sometimes completely absent; striae 2 and 3 usually absent in basal 1/4 and additionally interrupted, sometimes uneven dorsal striae interrupted basally, especially striae 5 and 7; submarginal stria present, well-developed. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively long, subequal in length, anterior claw thicker and more curved at its apex than posterior one and due to this with a subapical incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite with broadly pointed apexes, left straight, right slightly curved to left; left dorsal lobe slightly longer than right dorsal lobe, distinctly narrowed to apex in its apical 1/2; lateral side of left dorsal lobe in whole apical 1/2 narrowly and shallowly concave under lateral margin that looks like a bead, covered with large scale-like structures towards dorsal side and smaller scale-like structures towards apex and on lateral margin, with numerous small spinula-like structures in concavity and less numerous large, short spinula-like structures medially; right dorsal lobe covered with large scale-like structures, with small, indistinct median membranous area; lobes of ventral sclerite sclerotised, long, broad, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, with slightly curved apex and broad slightly sclerotised part rounded apically; right ventral lobe longer than left one, slightly sclerotised, with long curved apex visible in left lateral view. Parameres with dense, relatively long setae occupying approximately half of dorsal margin, with single the most proximal setae standing separately (Fig. 101).

Female : As males but with stronger dorsal punctation and microreticulation and more numerous pronotal strioles.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe completely covered by scale- and spinula-like structures in left lateral and ventral views. Based on dorsal colouration and general shape of the median lobe, it is similar to A. asteios sp. nov., A. epicharis sp. nov., A. wasurensis sp. nov., and A. yeretuar sp. nov. but differs from them in larger body size and left dorsal lobe of the median lobe with narrow, shallow, concavity under the lateral margin and different organisation of the surface structures. It is also very similar to A. pseudogestroi sp. nov., see its affinities.

Papua New Guinea: Central (Rigo, Kapakapa, Hula and Kelesi in Régimbart (1892: 991; 1899: 303), Angabanga River as Paumomu River in Régimbart (1899: 303)), Morobe, National Capital District, East New Britain (Kokopo as Herbertshöhe in Zimmermann (1919: 202)) provinces (Fig. 108). The East New Britain record needs to be confirmed.

The species was collected in Varirata NP in small, shallow and shaded forest pools rich in rotten leaves.

Papua New Guinea: Morobe Province, Herzog Mts, backroad to Wagau, 06°46.717'S, 146°38.953'E, 100 m a.s.l.

Holotype : male “Papua New Guinea: Morobe, Herzog Mts., backroad to Wagau, 100m, 3.iv.2006, 6 46.717S 146 38.953E, Balke & Sagata (PNG 29)” (ZSM). Paratypes: PNG: Morobe: 1 male, 7 females with the same label as the holotype (NHMW, ZSM). 1 male “Papua New Guinea: Morobe, Herzog Mts., backroad to Wagau, 100m, 3.iv.2006, Balke & Sagata (PNG 29)”, “3216” [green label] (ZSM). 5 males, 7 females “Stn. No. 158”, “New Guinea: Morobe Distr., Gusap, Markham, Valley c.90 m W. of Lae. 1,000ft. 27–30.i.1965”, “M.E. Bacchus B.M. 1965-120” (BMNH, NHMW). 1 male, 1 female “Stn. No. 162”, “New Guinea: Morobe Distr., Gusap, Markham, Valley c.90 m W. of Lae. 1,000ft. 27–30.i.1965”, “M.E. Bacchus B.M. 1965-120” (BMNH). 1 female “Stn. No. 161”, “New Guinea: Morobe Distr., Gusap, Markham, Valley c.90 m W.of Lae. 1,000ft. 27–30.i.1965”, “M.E. Bacchus B.M. 1965-120” (BMNH). 1 male, 6 females “Stn. No. 114”, “New Guinea: Morobe Distr., Markham R. Valley. Gusap, c.90 m. W. of Lae. 1000ft. 3.xii.1964”, “M.E. Bacchus B.M. 1965-120” (BMNH). Madang: 3 males, 7 females “Papua New Guinea: Madang, middle ramu [Ramu River], Akaraski, 50m, 12.iii.2007, 05.04.787S 144.43.137E, Kinibel (PNG 157)”, two males with additional green labels “2881” and “2882” (NHMW, ZSM). East Sepik: 1 male, 2 females “Papua New Guinea East Sepik Province Amboin Patrol Post Karawari Lodge 7 Jan. 1983, A.C.Messer” (ZSM). IN: West Papua: Yapen Islands Regency: 1 male “Irian Jaya: Japen Isl. W Serui, Panduamin ca. 100m, 19.2.1999 leg. Riedel” (NHMW).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 43).

Measurements : TL 5.65–6.7 mm, TL-H 5.75–6.1 mm, MW 2.8–3.2 mm, TL/MW 2.02–2.09; PL 0.8–0.95 mm, PW 2.3–2.7 mm, PL/PW 0.33–0.37; DBE 0.85–1 mm, DBE/PW 0.37. Holotype: TL 6.3 mm, TL-H 5.75 mm, MW 3.05 mm, TL/MW 2.07; PL 0.95 mm, PW 2.6 mm, PL/PW 0.37; DBE 0.95 mm, DBE/PW 0.37.

Colouration : Dorsally piceous, with yellowish red posterior part of head, pronotal sides, especially anterolaterally, broad, basal band and large, apical spot on elytron (Fig. 43).

Head piceous, yellowish red posteriorly, between eyes; sometimes yellowish red to brown, piceous behind eyes and two spots in middle. Pronotum piceous, with yellowish red sides, often only in anterolateral half. Elytron piceous, with broad, distinct, yellowish red basal band and large, yellow spot apically that usually slightly extending laterally. Scutellum piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs slightly darker, especially distally. Ventral side brown, with paler prosternum and abdominal ventrites.

Surface sculpture : Elytron with (10–11)+0 striae; stria 1 usually strongly reduced, striae 2 and 3 sometimes reduced basally (Fig. 43).

Head without strioles, with rather dense punctation (spaces between punctures 1–3× size of punctures); punctures coarse (diameter of punctures usually larger than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum without strioles or with few strioles posterolaterally or in females, with numerous strioles laterally; with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae, stria 1 often present as strioles in apical 1/2 or sometimes completely absent, striae 2 and 3 usually absent in basal 1/4 or present as strioles, stria 10 reduced at shoulder; submarginal stria absent. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively long, simple, subequal in length. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, more or less straight and evenly tapering to broadly pointed apexes, tip of left dorsal lobe apex with kind of small incision, not completely pointed; left dorsal lobe with lateral side modified: with distinct, median concavity of of triangular shape, covered with numerous, very distinct, strong and long spinula-like structures; lateral margin apically without surface structures, smooth; right dorsal lobe covered with scale-like structures, with median membranous area; lobes of ventral sclerite partly sclerotised, long, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, concave apically, with apex pointed and slightly curved to right and membranous right part with long, thin, upwardly curved apex sometimes sticking out in left lateral view; right ventral lobe with narrow sclerotised area on right margin. Parameres with dense, long setae occupying approximately half of dorsal margin, with single the most proximal setae standing separately (Fig. 92).

Female : As males but usually with stronger elytral punctation and microreticulation and pronotum with numerous strioles occupying sides completely so that only disc without strioles. Interestingly, some females have pronotum with lateral margin slightly to distinctly concave at anterior angles.

There is a variation in dorsal colouration and striolation as described above.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe. Based on body size and shape and shape of the median lobe with the characteristic median, triangle concavity of the left dorsal lobe, it is very similar to A. kalibumi sp. nov. but differs from it in elytron with broad and distinct, yellowish red basal band and slightly differently shaped median lobe, with its apical part distinctly shorter and apex of the left dorsal lobe less pointed. See also under A. sararti sp. nov.

The species is named after Herzog Mts. The name is an adjective in the nominative singular.

According to the studied material, the species has a disjunct areal: all specimens except one are from Papua New Guinea: Madang, Morobe and East Sepik provinces and only one male (its identification is sure) is known from Yapen Island of Indonesia (Fig. 108). Further sampling activities are necessary to show whether the species is indeed distributed in western part of New Guinea or the Yapen specimen is mislabelled.

In the Herzog Mts, the species was collected in shallow and shaded forest pools rich in rotten leaves.

Indonesia: Papua Province: Nabire Regency, road Nabire-Enarotali, 54^th km, 03°29.51'S, 135°43.91'E, 750–800 m a.s.l.

Holotype : male “West New Guinea/Paniai Prov./IR 19 track Nabire-Ilaga km 54 Basecamp, 750–800m, 16.-27.7.1991 leg: Balke & Hendrich” (ZSM). Paratypes: IN: Papua: Nabire Regency: 52 males, 36 females with the same label as the holotype (MZB, NHMW, ZSM). 9 males, 9 females “IR 19-W, New Guinea, Track Nabire-Ilaga KM 54, basecamp, 750–800m, 16.-27.vii.1991 Balke & Hendrich leg.” (NHMW). 41 males, 12 females “W.-Neuguinea/Paniai Prov. Strasse Nabire-Ilaga km 54 700m, 22.-25.9.1990/IR 11 leg: Balke & Hendrich” (NHMW, ZSM). 6 males, 3 females “West New Guinea/Paniai Prov./IR 20 track Nabire-Ilaga KM 59, ca.750m, 18.7.1991 leg: Balke & Hendrich leg.” (ZSM). 17 males, 3 females “IRIAN JAYA: Paniai Prov. road Nabire – Ilaga, km 65, 29.8.1996, 250m (96 # 6)” (NHMW). 1 male “West New Guinea/Paniai Prov./IR 23 track Nabire-Ilaga km 34 near Topo, 120m, 23.7.1991 leg: Balke & Hendrich” (ZSM). 2 males “IRIAN JAYA: Nabire Prov. rd. Nabire – Ilaga, Km 35 Kali Cemara, 100m, 27.9.1997 (IR97#6)” (NHMW). 1 male, 3 females “IRIAN JAYA: Nabire Prov. rd. Nabire – Ilaga, Km 35 Kali Cemara, 100m, 23.10.1997 (IR97#14)” (NHMW). 1 male “IRIAN JAYA: Paniai Prov. road Nabire – Ilaga, km 38 18.9.1996, 150m (96 # 26)” (NHMW). 2 males “West New Guinea/Paniai Prov./IR 18 River n. Nabire, 2m, 15.7.1991 leg: Balke & Hendrich leg.” (ZSM). 54 males, 36 females “West New Guinea/Paniai Prov./IR 22 track Nabire-Ilaga km 62 250m, 24.7.1991, forest pools leg: Balke & Hendrich” (NHMW, ZSM). 1 female “IRIAN JAYA: Kabup. Nabire rd. Nabire – Ilaga, km 62 200m, IX.1998 leg. Konyorah (62)” (NHMW). 23 males, 2 females “West New Guinea/Paniai Prov./IR 21 track Nabire-Ilaga km 65 Kali Utowa, 250m, 18.&19.7.1991 leg: Balke & Hendrich” (NHMW, ZSM). 9 males, 10 females “IR 23-W. New Guinea, track Nabire-Ilaga KM 62, 250m, 24.vii.1991 Balke & Hendrich leg.” (NHMW). 12 males, 13 females “West New Guinea/Paniai Prov./IR 21 track Nabire-Ilaga km 65 Kali Utowa, 250m, 18.&19.7.1991 leg: Balke & Hendrich” (ZSM). 5 males, 3 females “IR 21-W. New Guinea, track Nabire-Ilaga KM 65, Kali Utowa, 250m, 18.-19.vii.1991 Balke & Hendrich leg.” (ZSM). 1 male, 1 female “Indonesia: Papua, Road Nabire-Enarotali KM 55, 774m, 22.x.2011, 03.29.796S 135.43.885E, UNCEN team (PAP09)”, “5127” [green text] and “5126” [green text], respectively (ZSM). 1 male “5136” [green text], “Indonesia: Papua, Road Nabire-Enarotali KM 62, 340m, 22.x.2011, 03.31.684S 135.42.802E, Uncen (PAP11)” (ZSM). 4 males “W.-Neuguinea /Paniai Prov./ Wanggar- Kali Bumi / IR 14 30.9 & 1.10.90 leg: Balke & Hendrich” (ZSM). 1 male, 1 female “IR 90#14: West New Guinea, Nabire→Wanggar, 100m, 30.ix.1990, Balke” (ZSM). Puncak Regency: 2 males, 1 female “Indonesia: Papua, Rouaffer, Iratoi, hill in forest, 164m, 6.ix.2014, -3,2403 137,332 (Pap028)”, one male and female with additional green text labels “6476” and “6475”, respectively (ZSM). West Papua: Kaimana Regency: 1 male “INDONESIA: W-PAPUA vic. Kaimana, road 18 km NE S3°31'11”/E 133°40'15”, 50–80m 21.-25.II.2011 leg. A.Skale (014)” (CAS). Teluk Wondama Regency: 1 male “Indonesia: Irian Jaya, Wandammen, Wasior, DMP, 7.-10.I.2001, Riedel, MB 54”, “54” [green lable] (ZSM). 1 female “Indonesia: West Papua, DMP, Wasior, 7.-10.i.2001, Riedel leg.” (ZSM). 5 males, 2 females “IRIAN JAYA, Wandammen Bay, Wasior, KM 38, Sararti, 100-200 m, 7.-9.I.2001, leg. A. RIEDEL” (SMNS).

Body size and form : Beetle large, with oblong-oval habitus (Figs 13–15).

Measurements : TL 6.7–8.2 mm, TL-H 6–7.3 mm, MW 3.2–3.9 mm, TL/MW 2.09–2.14; PL 0.95–1.25 mm, PW 2.8–3.5 mm, PL/PW 0.34–0.36; DBE 1.1–1.4 mm, DBE/PW 0.39–0.41. Holotype: TL 7.6 mm, TL-H 6.8 mm, MW 3.55 mm, TL/MW 2.14; PL 1.15 mm, PW 3.2 mm, PL/PW 0.36; DBE 1.3 mm, DBE/PW 0.41.

Colouration : With piceous elytra and pronotum and bright yellowish red head, pronotal sides, and basal band on elytron; sometimes yellowish red colouration can be more strongly developed on elytron (Figs 13–15).

Head yellow to yellowish red, narrowly piceous behind eyes. Pronotum dark brown to piceous on disc and gradually paler (to yellow) on sides. Elytron dark brown to piceous, with very distinct yellow to yellowish red basal band, often with yellowish or reddish apex, sometimes also yellow laterally and between striae up to so that whole elytron yellow, with darker disc. Scutellum piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red to yellowish brown, metalegs darker, all legs darker distally. Ventral side reddish brown.

Surface sculpture : Dorsal elytral striation very variable, submarginal stria usually absent or present as a short stria or few short strioles apically: (0–10)+(0–1) (Figs 13–15).

Head without strioles, with relatively dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (diameter of punctures smaller than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum without strioles, with punctation finer than on head and sometimes with longitudinal wrinkles laterally; microreticulation finer than on head. Elytron with striation very variable: from without striae, with three or four puncture lines, which have often some strioles instead of punctures; with up to ten dorsal striae (as in holotype), of which uneven striae usually complete, striae 1 and 2 always reduced basally; submarginal stria usually absent or present as a short stria or few short strioles apically. Elytron with fine punctation and microreticulation. Abdominal ventrite 6 with distinct punctation, sparse medially and forming denser area at each lateral side, with few strioles laterally.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins distinctly convergent anteriorly. Base of prosternum broadly rounded anteriorly, distinctly convex medially; blade of prosternal process broad.

Male : Protibia modified: thinner proximally and broader medially and distally due to its curved ventral margin. Proclaws relatively short, subequal in length, anterior claw thicker subapically and more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus robust, with two lobes of dorsal sclerite unequal in length and shape, both covered with scale-like structures; left dorsal lobe distinctly broader and longer than right dorsal lobe; left dorsal lobe with a distinct median concavity and its whole apical 1/2 narrowed, its basal part very large, and its apex very slightly curved downwards, with a small crest; right dorsal lobe with apex pointed and membranous area medially; lobes of ventral sclerite sclerotised, pressed together between lobes of dorsal sclerite, but partly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area (not visible in left lateral view) and less sclerotised right part (visible in left lateral view); sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2, which apex can be visible at median lobe apex; less sclerotised right part long and broad, with its apical part densely covered with long setae; right ventral lobe long and broad. Parameres with dense, long setae occupying slightly more than half of dorsal margin (Fig. 65).

Female : Dimorphic: as male, shiny, and matt, with pronotum and elytron very densely covered with fine longitudinal strioles. Matt forms present in almost every population, usually less numerous than shiny ones: e.g., ratio shiny to with strioles is 23:13 in the type locality.

The species is very variable in the elytral striation and colouration, as described above, within and among populations. The holotype is characteristic: piceous, with yellowish red head, pronotal sides, and basal band on elytron, and with 10+1 elytral striae. All specimens from the type locality (IR19) have such colouration but most of them have some stria reduction up to six dorsal striae. In the neighbour population (locality IR21), almost all specimens have elytra without striae and approximately half of them have yellow elytral colouration strongly developed. From Papua Province (Kaimana, Wandammen, and Iratoi) only striated and dark specimens are recorded.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is similar to A. bundunensis sp. nov. in size, body form, and dorsal colouration, but distinctly differs from it in shape of the median lobe and modified male proclaws.

The species name is a Latin adjective meaning inconsistent and indicates very variable elytral striation of the species.

Indonesia: West Papua Province and western part of the Papua Province (Fig. 107).

Most specimens were collected in shallow (up to 20 cm water depth), shaded or at least partly shaded forest pools and puddles of different size, rich in rotten leaves and twigs, as, for example, a small forest pool at the 62 km of the Nabire-Ilaga track (Figs 109–113). Few specimens were also found in water-filled track hollows on forest tracks.

Indonesia: Papua Province: Puncak Regency, Iratoi, Rouaffer River, 3°14'25.1"S, 137°19'58.7"E, 164 m a.s.l.

Holotype : male “Indonesia: Papua, Rouaffer, Iratoi, hill in forest, 164m, 6.ix.2014, -3,2403086 137,3329744 (PAP028)” (KSP). Paratypes: 7 males, 9 females with the same label as the holotype, two males with additional labels with green text “6471” and “6472” (MZB, NHMW, ZSM).

Body size and form : Beetle small, elongate (Fig. 25).

Measurements : TL 4.95–5.55 mm, TL-H 4.5–5.1 mm, MW 2.25–2.5 mm, TL/MW 2.2–2.23; PL 0.75–0.85 mm, PW 2–2.3 mm, PL/PW 0.36–0.38; DBE 0.87–0.95 mm, DBE/PW 0.41–0.44. Holotype: TL 5.45 mm, TL-H 4.8 mm, MW 2.45 mm, TL/MW 2.23; PL 0.8 mm, PW 2.22 mm, PL/PW 0.36; DBE 0.95 mm, DBE/PW 0.43.

Colouration : Dorsally piceous, with yellowish red head, pronotal sides and elytral basal band and apex (Fig. 25).

Head yellowish red, narrowly darker behind eyes. Pronotum piceous on disc and gradually paler (to yellowish red) to sides. Elytron piceous, with distinct yellowish red basal band and yellow apically, especially between striae. Scutellum reddish to brown. Antennae, other head appendages, and pro- and mesolegs yellow to yellowish red, metalegs darker, all legs darker distally. Ventral side brown, reddish brown on head, pro- and metaventrite and medially on abdominal ventrites.

Surface sculpture : Elytron with 11 complete striae, submarginal stria present: 11+1 (Fig. 25).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures coarse (diameter of punctures larger than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum with few strioles at posterior margin to several strioles at anterior and posterior margins, with punctation finer than on head and microreticulation fine. Elytron with 11 complete striae, stria 10 shortly reduced basally; submarginal stria present, long. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process narrow.

Male : Protibia more or less straight, not modified; its ventral margin can be slightly curved proximally. Proclaws very long, straight, subequal in length. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, with broadly pointed apexes; left dorsal lobe with lateral margin slightly concave apically and lateral side slightly concave medially, covered laterally with numerous distinct spinula-like structures and dorsally with less numerous scale-like structures, lateral margin without surface structures, smooth; right dorsal lobe covered with scale-like structures; lobes of ventral sclerite partly sclerotised, long, partly visible in left lateral view; left ventral lobe with elongate, narrow sclerotised area, concave apically, with apex pointed and slightly curved; right ventral lobe longer than left lobe, with narrow sclerotised area on right margin, membranous part with long, thin, upwardly curved apex distinctly sticking out in left lateral view. Parameres with dense, long setae occupying distinctly more than half of dorsal margin; on small area proximally shorter and sparser, with several the most proximal setae standing separately (Fig. 76).

Female : Only matt forms present.

The species is similar to A. sumokedi sp. nov. in elongate habitus and dorsal colouration and striation, but differs from it in larger body size, very long, straight male proclaws, and in median lobe structure: slender and more pointed left lobe apex of dorsal sclerite and larger spinula-like structures of left dorsal lobe. In shape and surface structure of the median lobe and long, straight male proclaws, the species is also similar to A. ohu sp. nov. but it is larger than the latter, has more elytral striae and slightly differently shaped left dorsal lobe of the median lobe and its spinula-like structures more numerous.

The species is named after Iriatoi. The name is a noun in the nominative singular standing in apposition.

Indonesia: Papua Province: Puncak Regency. The species is known only from the type locality (Fig. 107).

The species was collected in small, shallow and shaded forest pools rich in rotten leaves.

Indonesia: Papua Province: Nabire Regency, Wanggar, Bumi River.

Holotype : male “W.-Neuguinea /Paniai Prov./ Wanggar- Kali Bumi / IR 14 30.9 & 1.10.90 leg: Balke & Hendrich” (ZSM). Paratypes: 63 males, 29 females with the same label as the holotype (NHMW). 16 males, 8 females “West New Guinea/Paniai Prov./IR 23 track Nabire-Ilaga km 34 near Topo, 120m, 23.7.1991 leg: Balke & Hendrich” (MZB, CLH, ZSM). 11 males, 8 females “Irian: Nabire – Ikaga [sic! Ilaga], “KM 35”, Seitenstr. nach K. Cemara, 1991, leg. M. Balke” (ZSM). 1 female “Indonesia: Papua, Road Nabire-Enarotali KM 35, r. Topo, 130m, 22.x.2011, 03 28.727S 135.38.734E, Uncen (PAP09A)” (ZSM). 1 female “Indonesia: Papua, Road Nabire-Enarotali KM 62, 340m, 22.x.2011, 03.31.684S 135.42.802E, Uncen (PAP11)”, “5144” [green text] (ZSM). 6 males, 2 females “IR 90#14: West New Guinea, Nabire→Wanggar, 100m, 30.ix.1990, Balke” (ZSM). 1 male “IRIAN JAYA: Nabire Prov. rd. Nabire – Ilaga, Km 35 Kali Cemara, 100m, 27.9.1997 (IR97#6)” (NHMW). 1 female “IRIAN JAYA: Nabire Prov. rd. Nabire – Ilaga, Km 35 Kali Cemara, 100m, 27.9.1997 (IR97#5)” (NHMW). 1 female “IRIAN JAYA: Nabire Prov. rd. Nabire – Ilaga, Km 35 Kali Cemara, 100m, 23.10.1997 (IR97#14)” (NHMW). 1 male, 2 females “IRIAN JAYA: Paniai Prov. road Nabire – Ilaga, km 38 18.9.1996, 150m (96 # 26)” (NHMW). 1 female “IRIAN JAYA: Paniai Prov. road Nabire – Ilaga, km 90 1.9.1996, 150m (96 # 11)” (NHMW). 4 females “IRIAN JAYA: Kabup, Nabire 30km S Nabire, Kali Cemara, 150m, 15.8.1998 (CE 1)” (NHMW). 2 males, 2 females “IRIAN JAYA: Kabup, Nabire 30km S Nabire, Kali Cemara, 150m, 15.8.1998 (CE 2)” (NHMW).

1 female “Indonesia: Papua, Road Nabire-Enarotali KM 62, 340m, 22.x.2011, 03.31.684S 135.42.802E, Uncen (PAP11)” (ZSM).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 42).

Measurements : TL 6.1–6.7 mm, TL-H 5.5–6.05 mm, MW 2.85–3.25 mm, TL/MW 2.06–2.14; PL 0.85–1 mm, PW 2.4–2.7 mm, PL/PW 0.35–0.37; DBE 0.9–1 mm, DBE/PW 0.37–0.38. Holotype: TL 6.3 mm, TL-H 5.7 mm, MW 3 mm, TL/MW 2.1; PL 0.95 mm, PW 2.6 mm, PL/PW 0.37; DBE 0.95 mm, DBE/PW 0.37.

Colouration : Dorsally piceous, with yellowish red pronotal sides, especially anterolaterally, and large yellowish red spot on elytral apex (Fig. 42).

Head piceous, paler (sometimes to yellowish red) posteriorly, between eyes, seldom also anteriorly. Pronotum piceous, with yellowish red sides, often only in anterolateral half. Elytron piceous, with large, yellow spot apically that usually slightly extending laterally. Scutellum piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs distinctly darker, especially distally. Ventral side piceous, with slightly paler prosternum.

Surface sculpture : Elytron with (10–11)+0 striae; stria 1 usually strongly reduced, stria 2 reduced basally (Fig. 42).

Head without strioles, with rather dense punctation (spaces between punctures 1–3× size of punctures); punctures fine and weakly impressed (diameter of punctures usually smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum without strioles or in females, with numerous strioles laterally, with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae, stria 1 often completely absent or present as few tiny strioles in apical 1/2, stria 2 usually absent in basal 1/4 or present as strioles, stria 3 sometimes shortly reduced basally, stria 10 reduced at shoulder or interrupted; submarginal stria absent. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively long, simple, subequal in length. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, more or less straight and evenly tapering to broadly pointed apexes; left dorsal lobe with lateral side modified: with distinct, median concavity of triangular shape, covered with numerous, very distinct, strong and long spinula-like structures; lateral margin apically without surface structures, smooth; right dorsal lobe covered with scale-like structures, with median membranous area; lobes of ventral sclerite partly sclerotised, long, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, concave apically, with apex pointed and slightly curved to right and membranous right part with long, thin, upwardly curved apex sometimes sticking out in left lateral view; right ventral lobe longer than left lobe, with narrow sclerotised area on right margin. Parameres with dense, relatively long setae occupying approximately half of dorsal margin (Fig. 91).

Female : As males but with stronger elytral punctation and microreticulation and pronotum with strioles, usually numerous, occupying complete sides so that only disc without them.

There is a variation in dorsal striolation as described above.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe. Based on body size and shape and shape of the median lobe with the characteristic median, triangle concavity of the left dorsal lobe, it is very similar to A. herzogensis sp. nov. but differs from it in dark elytron, without yellow basal band and slightly differently shaped median lobe, with its apical part distinctly longer. See also under A. sararti sp. nov.

The species is named after the Bumi River (= Kali), near Wanggar, Nabire. The name is a noun in the nominative singular standing in apposition.

Indonesia: Papua Province: Nabire and Paniai regencies (Fig. 108).

Specimens at the type locality were collected in shallow, exposed and flooded meadow and oxbows along Kali (=River) Bumi at Wanggar (Fig. 115). All specimens on the track Nabire to Ilaga at km 34 (Topo), were collected in shallow (up to 20 cm water depth), shaded or at least partly shaded forest pools and road side ditches, rich in rotten leaves and twigs (Fig. 109).

Indonesia: West Papua Province: Manokwari Regency, road from Kebar to Manokwari, 0°48'05.0"S, 133°19'20.6"E, 331 m a.s.l.

Holotype : male “Indonesia: Papua Barat, Kebar to Manokwari, 1h from Kebar, limestone creek and roadside pools,”, “331m, 8.xi.2013, -0.8013 133.3223, Unipa team (BH035)” (KSP). Paratypes: 2 males, 2 females with the same label as the holotype, one male additionally with a green text label “6245” (MZB, NHMW, ZSM). 2 males, 3 females “Indonesia: Papua Barat, Sausapor-Fef, 157m, 30.ix.2014, -0,6975004 132,072253 (BH044)”, one male additionally with a green text label “6485” (ZSM). 1 female “Indonesia: Papua Barat, Kebar Valley, 596m, 6.v.2015, -0,8406 133,2682, Unipa team (BH059)” (ZSM). 2 females “Indonesia: Papua Barat, lowland Manokwari, 66m, 8.v.2015, -0,7433 133,3975, Unipa team (BH065)” (ZSM). 1 male, 1 female “Indonesia: Papua Barat, Manokwari to Kebar, forest stream, 302m, 3.xi.2013, -0.8005 133.3321, Unipa team (BH023)”, with two additional green text labels “6222” and “6221”, respectively (ZSM). 2 males, 1 female “Indonesia: Papua, Manokwari, 140m, 8.ii.2006, 00.55.752S 133.54.448E, Tindige & Balke (BH 09)” (ZSM). 1 male, 3 females “Indonesia. West Papua Manokwari distr., Utai. riv. Arfak Mts., 500m 14.xii.2012 J. Horák leg.” (NMPC).

Body size and form : Beetle medium-sized, oblong-oval to elongate (Fig. 17).

Measurements : TL 5.1–5.7 mm, TL-H 4.6–5.2 mm, MW 2.4–2.65 mm, TL/MW 2.13–2.15; PL 0.8–0.9 mm, PW 2.1–2.3 mm, PL/PW 0.37–0.39; DBE 0.87–1 mm, DBE/PW 0.41–0.44. Holotype: TL 5.6 mm, TL-H 5.2 mm, MW 2.6 mm, TL/MW 2.15; PL 0.9 mm, PW 2.3 mm, PL/PW 0.39; DBE 1 mm, DBE/PW 0.44.

Colouration : Dorsally piceous, with yellowish red head, pronotal sides, basal band and apex of elytron (Fig. 17).

Head yellowish red to red, narrowly piceous behind eyes. Pronotum piceous on disc and gradually paler (to yellowish red) to sides. Elytron piceous, with distinct, broad yellowish red basal band and largely yellow apex, especially between striae. Scutellum reddish brown. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs darker, all legs darker distally. Vental side reddish brown.

Surface sculpture : Elytron with six dorsal striae and a submarginal stria: 6+1 (Fig. 17).

Head without strioles, with distinct, relatively sparse punctation (spaces between punctures 1–5× size of punctures); punctures relatively fine (diameter of punctures more or less equal to diameter of microreticulation cells); microreticulation distinct. Pronotum without strioles, with punctation finer than on head and microreticulation distinct. Elytron with six dorsal striae; stria 1 can be shortly reduced basally; striae 5 and sometimes 6 shortly reduced basally or present only in apical 1/2; submarginal stria present. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins slightly convergent anteriorly. Base of prosternum rounded anteriorly, distinctly convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively long, subequal in length, anterior claw thicker subapically and more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, covered with large, broad scale-like structures; left dorsal lobe with lateral margin concave apically and with subapical crest, apex broadly pointed and slightly curved upwards; lobes of ventral sclerite partly sclerotised, subequal in length, long, partly visible in left lateral view; left ventral lobe with elongate, narrow sclerotised area, its apex pointed and curved; right ventral lobe more membranous, slightly longer than left lobe, with apex pointed and slightly curved. Parameres with dense, long setae occupying more than half of dorsal margin; on small area proximally shorter and sparser (Fig. 69).

Female : As male.

There is an insignificant variation in the elytral striation and colouration.

In body size and shape and dorsal colouration, the species is similar to A. yamurensis sp. nov. but distinctly differs from it in the median lobe without submedian knob-like modification of the left dorsal lobe and 6 complete dorsal elytral striae.

The species is named after Kebar Village. The name is an adjective in the nominative singular.

Indonesia: West Papua Province: Manokwari Regency (Fig. 107).

The species was collected in different water bodies: in a limestone creek, roadside pools and a forest stream.

Papua New Guinea: Central Province, Kokoda, 08°53.481'S, 147°43.648'E, 410 m a.s.l.

Holotype : male “Papua New Guinea: Northern, Kokoda, 410m, i.2008, 53.481S 147.43.648E, Posman, (PNG 174)” (ZSM). Paratypes: 1 male, 2 females with the same label as the holotype (NHMW, ZSM).

Body size and form : Beetle medium-sized, rather elongate (Fig. 49).

Measurements : TL 5.6–5.65 mm, TL-H 5.1–5.2 mm, MW 2.6–2.65 mm, TL/MW 2.11–2.17; PL 0.8–0.85 mm, PW 2.25–2.3 mm, PL/PW 0.34–0.37; DBE 0.9–0.95 mm, DBE/PW 0.4–0.41. Holotype: TL 5.6 mm, TL-H 5.2 mm, MW 2.65 mm, TL/MW 2.11; PL 0.85 mm, PW 2.3 mm, PL/PW 0.34; DBE 0.95 mm, DBE/PW 0.41.

Colouration : With piceous elytra and pronotum and yellowish red head, pronotal sides, and basal band and large apical spot on elytron (Fig. 49).

Head yellowish red, narrowly piceous behind eyes. Pronotum piceous on disc and yellowish red on sides, especially at anterior angles. Elytron piceous, with distinct, rather broad, yellowish red basal band and large, yellow spot on apex often extending between striae and laterally. Scutellum reddish brown to brown. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs distinctly darker, especially distally. Ventral side reddish brown to brown, paler on pro- and metaventrite and abdominal ventrites 1–3.

Surface sculpture : Elytron with 11+(0–1) striae (Fig. 49).

Head without strioles, with rather dense punctation (spaces between punctures 1–3× size of punctures); punctures rather coarse (diameter of punctures usually equal to and larger than diameter of microreticulation cells); microreticulation distinct. Pronotum with several strioles or more numerous (in females) laterally, with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae, striae 1 and 10 usually shortly reduced basally, stria 1 additionally can be interrupted; submarginal stria absent or present as few tiny strioles apically. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum broadly rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws relatively long, slightly curved, subequal in length, anterior claw thicker subapically than posterior one and due to this with a shallow median incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, with broadly pointed, more or less straight apexes; lateral side of left dorsal lobe covered with numerous, small spinula-like structures; lateral margin slightly concave apically, without surface structures, smooth; right dorsal lobe covered with large scale-like structures, with large, distinct median membranous area; lobes of ventral sclerite partly sclerotised, long, subequal in length, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, concave apically, with apex pointed and slightly curved and membranous right part with long, thin, upwardly curved apex sometimes sticking out in left lateral view; right ventral lobe with narrow sclerotised area on right margin. Parameres with relatively dense and long setae occupying approximately half of dorsal margin (Fig. 97).

Female : As males but with stronger dorsal punctation and microreticulation and more numerous pronotal strioles.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe not completely covered by spinula-like structures, upper lateral margin without spinulae, smooth in left lateral and ventral views. Based on body size and dorsal colouration, it is similar to A. flavocapitatus sp. nov. but differs from it in more elongate habitus, left dorsal lobe of the median lobe with small spinula-like structures and its apical part slender and more elongate. See also under A. luteomaculatus and A. maindai sp. nov.

The species is named after Kokoda Village. The name is an adjective in the nominative singular.

Papua New Guinea: Central Province. The species is known only from the type locality (Fig. 108).

Unknown.

Papua New Guinea: Sandaun, Bewani Stn., ca 03°05.130'S, 141°10.227'E, 300 m a.s.l.

Holotype : male “Papua New Guinea: Sandaun, Bewani Stn., forest puddles @ base of Bewani Mts., 300 m, 12.iv.2006, nr. 03.05.130S 141.10.227E, Balke & Sagata (PNG 38)”, “3210” [green label] (ZSM). Paratypes: 1 female with the same label as the holotype (ZSM).

Body size and form : Beetle small, distinctly elongate, rather slender (Fig. 1).

Measurements : TL 4.4–4.5 mm, TL-H 4 mm, MW 1.9 mm, TL/MW 2.32–2.37; PL 0.7–0.75 mm, PW 1.8–1.85 mm, PL/PW 0.38–0.42; DBE 0.9 mm, DBE/PW 0.49–0.5. Holotype: TL 4.5 mm, TL-H 4 mm, MW 1.9 mm, TL/MW 2.37; PL 0.75 mm, PW 1.8 mm, PL/PW 0.42; DBE 0.9 mm, DBE/PW 0.5.

Colouration : Dorsally yellowish brown to brown, with distinct yellow basal band on elytron and paler head and pronotal sides; both specimens slightly teneral (Fig. 1).

Head yellowish to yellowish brown, narrowly darker behind eyes. Pronotum brown on disc and paler (yellowish brown to yellow) on sides. Elytron brown, with distinct yellow basal band. Scutellum yellowish to brown. Antennae, other head appendages, and pro- and mesolegs yellowish to yellowish brown, metalegs darker, all legs darker distally. Ventral side yellowish to yellowish brown. Most likely, beetles are darker when more mature.

Surface sculpture : Elytron without striae: 0+0, with three or four indistinct puncture lines (Fig. 1).

Head without strioles, with distinct, uneven punctation (spaces between punctures 1–5× size of punctures); punctures fine (diameter of punctures smaller than or equal to diameter of microreticulation cells); microreticulation distinctly impressed. Pronotum with short, weak strioles along posterior margin but mainly at posterior angles, with punctation finer and sparser than on head and microreticulation distinct. Elytron without striae, with three or four indistinct puncture lines (first line near suture sometimes reduced) and separate large punctures between them. Elytron with very fine, sparse punctation and distinct microreticulation.

Structures : Head large and broad. Pronotum slightly trapezoid, its lateral margins subparallel. Base of prosternum broadly rounded anteriorly, slightly convex medially; blade of prosternal process broad.

Male : Protibia straight, not modified. Proclaws long, slightly curved, subequal in size and form. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, covered with scale-like structures; left dorsal lobe with lateral margin slightly concave apically and with some spinula-like structures medially in left lateral view; apex of left dorsal lobe straight, broadly pointed; left lobe of ventral sclerite with elongate, narrow sclerotised area visible in left lateral view, its apex slightly curved upwards; right lobe of ventral sclerite membranous, with long, thin, upwardly curved apex distinctly sticking out in left lateral view. Parameres with relatively dense and long setae occupying more than half of dorsal margin (Fig. 59).

Female : As male.

There is an insignificant variation in the colouration and dorsal punctation described above.

The species is very distinct due to its small size, elongate, slender habitus, colouration, elytra without striae, and shape of the median lobe.

The species name is a Greek adjective meaning “thin, fine, slender” and indicates small, elongate habitus of the species.

Papua New Guinea: Sandaun Province. The species is known only from the type locality (Fig. 107).

The species was collected in forest puddles.

Papua New Guinea: National Capital District: ca. 16 km N from Port Moresby, Loloki River.

Holotype : male “Stn. No. 208”, “Papua: Loloki c. 10 m. N. of Pt. Moresby, 19.III.1965.”, “M.E. Bacchus. B.M. 1965-120” (BMNH).

Body size and form : Beetle small, with oblong-oval habitus (Fig. 23).

Measurements: Holotype: TL 5 mm, TL-H 4.5 mm, MW 2.4 mm, TL/MW 2.08; PL 0.75 mm, PW 2.1 mm, PL/PW 0.36; DBE 0.8 mm, DBE/PW 0.38.

Colouration : Dorsally dark brown, elytron with faint yellowish shoulder spot and apicolaterally (Fig. 23).

Head dark brown, narrowly piceous behind eyes and paler on clypeus and posteromedially. Pronotum dark brown on disc and reddish on sides. Elytron dark brown, paler basally, with faint yellowish shoulder spot, yellow at apex and narrowly laterally in more than apical 1/2. Scutellum reddish brown. Antennae, other head appendages, and legs yellowish brown, darker distally. Ventral side brown.

Surface sculpture : Elytron with 11 complete, relatively strongly impressed striae, submarginal stria present: 11+1 (Fig. 23).

Head with few strioles between eyes, with distinct, rather dense punctation (spaces between punctures 1–5× size of punctures); punctures relatively coarse (diameter of punctures slightly larger than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum complete covered with sparse but distinct strioles absent at anterior angles, with punctation finer than on head and microreticulation distinct. Elytron with 11 complete, relatively strongly impressed striae, submarginal stria present, long. Elytron with fine punctation and distinct microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws relatively short, slightly curved, subequal in length. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, covered with scale-like structures; left dorsal lobe with lateral margin slightly concave medially, with apex truncate and slightly curved downwards; right dorsal lobe narrowed subapically, with broad, rounded apex; lobes of ventral sclerite elongate, mostly sclerotised, almost invisible in lateral view. Parameres with rather short and sparse setae occupying slightly more than half of dorsal margin (Fig. 74).

Female : Unknown.

The species is well-recognisable by its smaller size, uniform, brown dorsal colouration, strong elytral striation and striolation of head and pronotum, and characteristic shape of the median lobe and paramere setation.

The species is named after Loloki River. The name is a noun in the nominative singular standing in apposition.

Papua New Guinea: National Capital District. The species is known only from the type locality (Fig. 107).

Unknown.

Indonesia: West Papua Province: Raja Ampat Regency, Waigeo Island, Lopintol, Bajon River, 0°07'S, 130°53'E.

Holotype : male “MB759” [green label], “W-PAPUA Raja Ampat Prov. Waigeo Isl., Lopintol, Bajon River, 0°07'S, 130°53'E 11.I.2004 leg. A.,”, “W Papua: Raja Ampat, Waigeo, Lopintol, 11i2004, A. Skale, MB 759” (MZB). Paratype: 1 male “W-PAPUA Raja Ampat Prov. Waigeo Isl., Lopintol 0°07'54"S, 130°53'45"E 11.I.2004 leg. A. Skale UWP” (CAS).

Body size and form : Beetle small, elongate (Fig. 3).

Measurements : TL 4.7–4.85 mm, TL-H 4.3–4.35 mm, MW 2.1–2.2 mm, TL/MW 2.21–2.24; PL 0.7–0.75 mm, PW 1.9–2 mm, PL/PW 0.37–0.38; DBE 0.9–0.95 mm, DBE/PW 0.47–0.48. Holotype: TL 4.7 mm, TL-H 4.3 mm, MW 2.1 mm, TL/MW 2.24; PL 0.7 mm, PW 1.9 mm, PL/PW 0.37; DBE 0.9 mm, DBE/PW 0.47.

Colouration : With dark brown elytra and paler head and pronotal sides; yellow basal band on elytron distinct (Fig. 3).

Head reddish, narrowly darker behind eyes. Pronotum dark brown on disc and gradually paler (to yellowish red) to sides. Elytron dark brown, with distinct yellowish red basal band and pale brown apex. Scutellum brown. Antennae, other head appendages, and pro- and mesolegs yellow, metalegs darker, all legs darker distally. Ventral side yellowish red.

Surface sculpture : Elytron with six dorsal striae and a submarginal stria: 6+1 (Fig. 3).

Head without strioles, with distinct, uneven punctation (spaces between punctures 1–5× size of punctures); punctures relatively fine (diameter of punctures more or less equal to diameter of microreticulation cells); microreticulation distinct. Pronotum without strioles, with punctation sparser and microreticulation finer than on head. Elytron with six dorsal striae, striae 1–4 complete, stria 5 reduced basally, stria 6 reduced apically. Elytron with fine punctation and microreticulation.

Structures : Head large and broad. Pronotum trapezoid, its lateral margins slightly convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws relatively short, subequal in length, anterior claw thicker subapically and more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, covered with scale-like structures; left dorsal lobe with lateral margin thickened apically in kind of bead or crest and concave medially, which gives median lobe calla-like shape; this apical crest of left dorsal lobe with small but distinct numerous spinula-like structures; lobes of ventral sclerite partly sclerotised, long, partly visible in left lateral view; left ventral lobe with elongate, narrow sclerotised area, its apex sharply pointed and curved upwards; right ventral lobe more membranous, longer than left lobe, with pointed apex. Parameres with dense, long setae occupying more than half of dorsal margin, proximally shorter and sparser (Fig. 66).

Female : Unknown.

In body size and shape, elytral striation and dorsal colouration, the species is similar to A. procerus sp. nov. and A. ohu sp. nov. but distinctly differs from them by its calla-like shape of the median lobe and complete, more strongly impressed elytral striae.

The species is named after its type locality, Lopintol. The name is an adjective in the nominative singular.

Indonesia: West Papua Province: Raja Ampat Regency, Waigeo Island (Fig. 107).

The species was collected in a shaded side pool of a larger river.

Papua New Guinea: Madang Province, Astrolabe Bay, Stephansort, 05°26'38.4"S, 145°44'47.8"E.

Holotype : male “N. Guinea Biró 97.”, “Stephansort Astrolabe B”, “Holotypus 1956 ♂ Copelatus luteomaculatus Guignot” [label with red frame, partly hw by Guignot], “Type” [red label], “Dr F. Guignot det., 1955 Copelatus luteomaculatus sp.n. HoloType ♂” [partly hw by Guignot] (HNHM). Paratypes: 1 female “N. Guinea Biró 97.”, “Stephansort Astrolabe Bai // 18.V. [hw on reverse side]”, “♀”, “Paratype” [red label with black frame] (MNHN); 2 males “N. Guinea Biró 1898”, “Simbang Huon Golf // IX.17. [hw on reverse side]”, “♂”, “Paratype” [red label with black frame] (MNHN); 2 females “N. Guinea Biró 1898”, “Simbang Huon Golf // IX.17. [hw on reverse side]”, “Paratypus 1956 Copelatus luteomaculatus Guignot” [label with red frame, partly hw by Guignot] (HNHM).

PNG: Morobe: 2 males, 1 female “Papua New Guinea: Morobe, Patep, 700m, 20.xi.2006, 06.58.267S 146.37.895E, Balke & Kinibel (PNG 105)” (NHMW, ZSM). 1 female “Papua Neuguinea Morobe Provinz Wau Ecology Institute”, “Wau Station 1217m 15.4.1999 073386S 1467083E leg. M.Schaarschmidt/H.Deumer A.Michalczyk” (ZSM). 2 males, 3 females “Papua New Guinea: Morobe, Herzog Mts., Bundun, 700–800m, 2.iv.2006, 06.51.598S 146.37.07E, Balke & Sagata (PNG 27)” (NHMW, ZSM). 2 males “Papua New Guinea: Morobe, Herzog Mts., backroad to Wagau, 100m, 3.iv.2006, Balke & Sagata (PNG 29)” (ZSM). 1 male “3236” [green label], “Papua New Guinea: Morobe, Huon Pen., rd to Kwapsanek, 250m, 31.iii.2006, 06.30.270S 146.59.581E, Balke & Sagata (PNG 24)” (ZSM). 1 female “Papua New Guinea: Morobe, Sattelberg, Iunjaing, 840m, 16.x.2009, 06.27.239S 147.42.531E, Idaho (09) (PNG209)” (ZSM). 25 males, 16 females “Papua New Guinea: Morobe, Menyamya, coffee garten, 1000m, 14.xi.2006, 07.13.714S 146.01.260E, Balke & Kinibel (PNG 99)”, two males with green labels “3225” and “3226” (NHMW, ZSM). 6 males, 8 females “Stn. No. 114”, “New Guinea: Morobe Distr., Markham R. Valley. Gusap, c.90 m. W. of Lae. 1000ft. 3.xii.1964”, “M.E. Bacchus B.M. 1965-120” (BMNH, NHMW). 3 males, 8 females “Stn. No. 158”, “New Guinea: Morobe Distr., Gusap, Markham Valley. c.90 m. W. of Lae. 1,000 ft. 27-30.i.1965”, “M.E. Bacchus B.M. 1965-120” (BMNH, NHMW). 2 males, 1 female “Stn. No. 159”, “New Guinea: Morobe Distr., Gusap, Markham Valley. c.90 m. W of Lae. 1,000 ft. 27-30.I.1965”, “M.E. Bacchus B.M. 1965-120” (BMNH). 1 male, 1 female “Stn. No. 160”, “New Guinea: Morobe Distr., Gusap, Markham Valley. c.90 m. W. of Lae. 1,000 ft. 27-30.i.1965”, “M.E. Bacchus B.M. 1965-120” (BMNH). 1 male, 4 females “Stn. No. 161”, “New Guinea: Morobe Distr., Gusap, Markham Valley. c.90 m. W. of Lae. 1,000 ft. 27-30.i.1965”, “M.E. Bacchus B.M. 1965-120” (BMNH). 1 female “Stn. No. 131”, “New Guinea: Morobe Distr., Lae-Bulolo Rd., 30.xii.1964.”, “M.E. Bacchus B.M. 1965-120” (BMNH). 8 males, 11 females “Stn. No. 127”, “New Guinea: Morobe Distr., Bulolo, 2,070ft. 28.xii.1964.”, “M.E. Bacchus B.M. 1965-120” (BMNH, NHMW). Madand: 2 males “Papua New Guinea: Madang, Mt Tapo, 180m, ii.2008, 5.24.11.00S 145.36.17.16E, BRC leg. (PNG 178)” (ZSM). 1 male, 2 female “Papua New Guinea: Madang, Brahmin, 150m, 26IX2002, 5 44.953S 145 20.844E Balke & Sagata (PNG 024)” (ZSM). Gulf: 2 males “Papua New Guinea: Gulf, 1500m, 13.xi.2006, 07.11.721S 145.54.746E, Balke & Kinibel (PNG 95)”, one male with a green label “4244” (ZSM).

Body size and form : Beetle medium-sized, with oblong-oval habitus (Figs 18, 19).

Measurements : TL 5.55–6.6 mm, TL-H 5.1–6 mm, MW 2.7–3.2 mm, TL/MW 2.06–2.07; PL 0.8–0.95 mm, PW 2.3–2.8 mm, PL/PW 0.34–0.35; DBE 0.85–1.01 mm, DBE/PW 0.36–0.39. Holotype: TL 6.2 mm, TL-H 5.6 mm, MW 3 mm, TL/MW 2.07; PL 0.9 mm, PW 2.6 mm, PL/PW 0.35; DBE 1 mm, DBE/PW 0.39.

Colouration : Dorsally piceous, with yellowish red head, pronotal sides, basal band and apex of elytron (Figs 18, 19).

Head yellow to red, narrowly piceous behind eyes and darker on clypeus. Pronotum piceous, with yellow to red anterior angles and relatively narrowly on sides, darker to posterolateral angles. Elytron piceous, with bright yellow to red basal band and largely yellow apex, especially between striae. Scutellum reddish to piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs darker, all legs darker distally. Ventral side reddish brown, yellowish red on head, pro- and metaventrite. The holotype is a teneral specimen, therefore, distinctly paler.

Surface sculpture : Elytron with six dorsal striae and with or without submarginal stria: 6+(0–1) (Figs 18, 19).

Head without strioles, with distinct, relatively sparse punctation (spaces between punctures 2–5× size of punctures); punctures very fine (diameter of punctures smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum with strioles laterally (few at posterolateral angles to rather numerous but sparse on sides), with punctation slightly denser than on head and microreticulation fine. Elytron with six dorsal striae; striae 1, 5, and 6 reduced basally; striae 1 and 5 sometimes more strongly reduced and present only in apical 1/2; sometimes with single strioles between striae; submarginal stria absent or present, sometimes as fine strioles apically. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum narrowly rounded anteriorly, distinctly convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively short, subequal in length, anterior claw thicker subapically and more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length; left dorsal lobe gradually tapering, covered with very dense spinula-like structures except for thickened lateral margin, with apex rounded and more or less straight; right dorsal lobe covered with large scale-like structures, without membranous area medially; lobes of ventral sclerite partly sclerotised, subequal in length, long, partly visible in left lateral view; left ventral lobe with elongate, narrow sclerotised area, well-visible in left lateral view, its apex pointed and curved; right ventral lobe more membranous, slightly longer than left lobe, with apex pointed and slightly curved. Parameres with dense, long setae occupying approximately half of dorsal margin (Fig. 70).

Female : As male.

There is an insignificant variation in the elytral striation and colouration.

In colouration and in shape and surfaces structure of the median lobe, the species is similar to A. flavocapitatus sp. nov. and A. kokodensis sp. nov. but differs from them in larger size, lesser elytral striation and dorsal sclerite shape of the median lobe. Austrelatus flavocapitatus sp. nov. has 10 dorsal elytral striae and left lobe of the dorsal sclerite with more concave lateral margin and apex gently curved downwards (with more straight lateral margin and apex gently curved upwards in A. luteomaculatus). Austrelatus kokodensis sp. nov. has 11 dorsal elytral striae and left dorsal lobe with more concave lateral margin (with more straight lateral margin in A. luteomaculatus). Two latter species have right lobe of the dorsal sclerite with membranous area medially, which is lacking in A. luteomaculatus.

Papua New Guinea: Madang, Morobe, and Gulf provinces (Fig. 107).

The species was collected in shallow and shaded forest pools.

Papua New Guinea: Madang Province, Wannang, 05°17.235'S, 145°06.160'E, 230 m a.s.l.

Holotype : male “Papua New Guinea: Madang, Wannang, 230m 3.x.2008, 05.17.235S 145.06.160E, Posman (PNG188)” (ZSM). Paratypes: PNG: Madang: 12 males, 12 females with the same labels as the holotype, one of males additionally with a green label “3765” (NHMW, ZSM). 2 males, 1 female “Papua New Guinea: Madang, Wannang, 270m, 31.x.2008, 05.15.458S 145.02.389E, Posman, (PNG187)”, males with additional green labels “4171” and “3792” (ZSM). 1 male “Papua New Guinea: Wanang I, river, 25.ix.2013, Boukal, 44/2013” (NHMW). 1 female “Ibisca Niugini, PNG 2-4.xii.2012 Wanang -5,227670193 145,0797424”, “FIT-WAN-T-8/8-d16 / Plot 20 / P0707 Vial 22371-CODYTI” (ZSM). 1 male, 1 female “Papua New Guinea: Wanang, Kusa, 24.ix.2013, Boukal, 39/2013” (ZSM). 1 male “Papua New Guinea: Wanang - Kusa, Digitam, 24.ix.2013, Boukal, 40/2013” (ZSM). 4 males, 3 females “PNG Madang Province OHU Village, 14.1.2001 leg.: Lukas Cizek Coll. HENDRICH” (CLH). 1 female “Papua New Guinea: Madang, Madang, Ohu Village, 160m, 30.iv.2006, 05.13.923S 145.40.763E, Balke & Manaono (PNG 49) (ZSM). 1 male, 1 female “Papua New Guinea: Madang, Usino, 260m, 15.iii.2007, 05.31.125S 145.25.316E, Kinibel (PNG 158)” (ZSM). 6 males, 7 females “Papua New Guinea: Madang, Akameku-Brahmin, Bismarck Range, 250–500m, 25.xi.2006, nr 05.47.026S 145.24.131E, Balke & Kinibel (PNG 115)” (NHMW, ZSM). 1 female “Ibisca Niugini, PNG 7–9.xi.2012 Mount Wilhelm 200m -5,739897251 145,3297424 MW0200 / P0866 Vial 07444” (ZSM). 1 male “Ibisca Niugini, PNG 1–3.xi.2012 Mount Wilhelm 200m”, “-5,739897251 145,3297424 FIT-MW200-T-4/8-d08 / P0863 Vial 07369-CODYTI” (ZSM). 1 male “Papua New Guinea Madang Prov. 2002 Hulcr”, “coll. Jiří Hájek National Museum Prague, Czech Republic” (NMPC). 3 males, 3 females “Papua New Guinea: Madang, Adelbert Mts., Keki to Sewan, 650m, 7.v.2006, 04.41.802S 145.25.460E, Balke (PNG 54)”, one male with an additional green label “3202” (NHMW, ZSM). 1 male “Papua New Guinea: Madang, Adalbert [sic!] Mts., Keki, 850m, 4.v.2006, nr 04.42.300S 145.25.089E, Balke & Manaono (PNG 52)”, “3206” [green label] (ZSM). 1 male, 1 female “Papua New Guinea: Madang, Keki – Sewan, Adalbert [sic!] Mts., 700m, 30.xi.2006, nr 04.41.802S 145.25.460E, Binatang Boys (PNG 120)” (NHMW, ZSM). 1 male “N. Guinea Biró 97.”, “Stephansort Astrolabe B.”, “♂”, “papuensis Balf.-Br.” [hw, Guignot], “MUSEUM PARIS 1960 Coll. F.GUIGNOT” [pink label], “MNHN, Paris EC29237 [with QR code]” (MNHN). East Sepik: 2 males “Papua New Guinea: East Sepik, Lembena, 198m, 3.ix.2009, 04.56.974S 143.56.995E, Ibalim & Pius (PNG241)” (ZSM). 1 male, 2 females “Papua New Guinea: East Sepik, Lembena, 198m, 3.ix.2009, 04 46 [!] 974S 143.56.995E, Ibalim & Pius (PNG243)” (ZSM). 1 female “Papua New Guinea: East Sepik, Lembena, 127m, 3.ix.2009, 04 56.953S 143 56.614E, Ibalim & Pius (PNG246)” (ZSM). 9 males, 9 females “Papua New Guinea: East Sepik, Lembena, 117m, 8.ix.2009, 04.57.513S 143 57.296E, Ibalim & Pius (PNG248)”, four males and one female with additional green text labels “5989”, “5991”, “5992”, “5993”, and “5990”, respectively (ZSM). 6 males, 2 females “Papua New Guinea: East Sepik, Lembena, 110m, 10.ix.2009, 04.57.512S 143.57.366E, Ibalim & Pius (PNG249)” (ZSM). 2 males, 1 female “Papua New Guinea: East Sepik, Lembena, 335m, 10.ix.2009, 04.56.859S 143.59.375E, Ibalim & Pius (PNG250)” (ZSM). 2 females “Papua New Guinea: East Sepik, Lembena, 335m, 10.ix.2009, 04 56.859S 143 57.379E, Ibalim & Pius (PNG251)” (ZSM). 1 male “Papua New Guinea: East Sepik, Lembena, 335m, 10.ix.2009, 04 56.921S 143 57.478E, Ibalim & Pius (PNG252)” (ZSM). 2 males, 1 female “Papua New Guinea: East Sepik, Prince Alexander Mts., Wewak-Angoram, 300m, 21.iv.2006, 03.42.129S 143., Balke (PNG 46)”, female with a green label “3222” (ZSM). EHL: 1 male “6438”, “Papua New Guinea: Eastern Highlands, Bena – pass to Goroka valley, 1550m, 5.iv.2006, 06.14.567S 145.29.634E, Balke & Sagata (PNG33)” (ZSM).

Body size and form : Beetle large, oblong-oval (Figs 38, 39).

Measurements : TL 6.8–8.15 mm, TL-H 6.2–7.35 mm, MW 3.25–4 mm, TL/MW 2.04–2.09; PL 1–1.2 mm, PW 2.9–3.55 mm, PL/PW 0.34–0.35; DBE 1.15–1.35 mm, DBE/PW 0.37–0.4. Holotype: TL 7.6 mm, TL-H 6.8 mm, MW 3.7 mm, TL/MW 2.08; PL 1.15 mm, PW 3.25 mm, PL/PW 0.35; DBE 1.2 mm, DBE/PW 0.37.

Colouration : Elytron yellowish red to reddish brown between striae, piceous on pronotal disc, with yellowish red head and pronotal sides (Figs 38, 39).

Head yellowish red, narrowly darker behind eyes. Pronotum brown to piceous on disc and gradually paler (to yellowish red) to sides. Elytral colouration varies: usually yellowish red to reddish brown between striae and piceous on striae, sometimes narrowly brown to piceous on disc, seldom elytron brown to piceous, with yellowish red basal band and yellow apex or apical 1/2. Scutellum yellowish red to brown. Antennae, other head appendages, and pro- and mesolegs yellowish red to reddish, metalegs darker, all legs darker distally. Ventral side brown, paler on prosternum and posteriorly and laterally on metacoxal plates and abdominal ventrites.

Surface sculpture : Elytral striation varies: usually elytron with 11 dorsal striae, stria 1 or striae 1–3 reduced in basal 1/2, sometimes stria 1 absent and other reduced or interrupted in basal 1/2, submarginal stria present: (10–11)+1 (Figs 38, 39).

Head without strioles, with rather sparse punctation (spaces between punctures 1–5× size of punctures); punctures rather fine (usually diameter of punctures smaller than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum without or with strioles posterolaterally, more numerous in females, with punctation finer than on head and microreticulation fine. Elytral striation varies: usually elytron with 11 dorsal striae, stria 1 or striae 1–3 absent in basal 1/2, sometimes more striae (e.g., striae 5–7, 9) reduced basally or differently interrupted; maximal reduction with stria 1 absent and almost all other striae absent or interrupted in basal 1/2; submarginal stria present, sometimes as apical strioles. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia more or less straight: its ventral margin slightly curved proximally. Proclaws long, subequal in length, anterior claw thicker apically and more strongly curved downwards than posterior one and due to this with a subapical incision on its inner margin. Median lobe of aedeagus robust, with two lobes of dorsal sclerite covered with scale-like structures; left dorsal lobe slightly concave medially, its whole apical 1/2 broad, more or less evenly tapering to rather large, curved downwards, truncate apex bearing broad crest; right dorsal lobe shorter than left one, broad, with pointed apex and membranous area medially; lobes of ventral sclerite partly sclerotised, pressed together between lobes of dorsal sclerite, slightly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area and less sclerotised right part; sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2; less sclerotised right part long and broad, with its apical part densely covered with long setae (sometimes visible at apex); right ventral lobe partly sclerotised, long and broad. Parameres with dense, long setae occupying approximately half of dorsal margin (Fig. 88).

Female : Dimorphic: as male, but with much more numerous pronotal strioles and distinctly denser punctation and stronger microreticulation on pronotum and elytron, and matt, with pronotum and elytron very densely covered with fine longitudinal strioles. Interestingly, the matt forms are characteristic only for the populations with male specimens having reduced elytral striae, e.g., ones of Adelbert and Alexander Mts or Ohu Village. In latter population, the normal form is also present but less numerous: ratio shiny to with strioles is 1:3.

There is a strong variation in dorsal colouration and striolation as described above. As mentioned above, specimens with reduced elytral striae are observed in three regions: Ohu Village and, especially Adelbert and Alexander Mts. Beetles from the two latter regions also have larger, broader, and more robust habitus and sturdier median lobe of aedeagus.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, in shape of the median lobe, it is similar to A. mamberamo sp. nov. and differs from it in more pale elytral colouration and apex of left dorsal lobe of the median lobe shorter, with more vertical, broader crest. See also under A. mamberamo sp. nov.

The species is named after Madang Province where it is largely distributed. The name is an adjective in the nominative singular.

Papua New Guinea: Madang, Eastern Highlands and East Sepik provinces (Fig. 108).

Unknown.

Indonesia: Papua Province: Sarmi Regency, Waskey, 01°58'17.0"S, 138°50'56.9"E, 70 m a.s.l.

Holotype : male “Indonesia: Papua, Sarmi area, 70m, 25.ix.2014,-1.9713908 138.8491402, Tim UNCEN: Balke & Menufandu (Pap032)” (KSP). Paratypes: 2 males, 1 female with the same label as the holotype, one male with an additional green text label “6469” (MZB, ZSM). 1 male “Indonesia: Papua, Sarmi, Waaf, N Foja Mts, waterfall in forest, 120m, 23.ix.2014, -2.3317793, 138.7500472, Tim UNCEN: Balke & Menufandu (Pap031)”, “6467” [green text] (ZSM). 1 male “Indonesia: Papua, Foja Mountaints N foot, N Waaf vill., pondok, 150m, 4.-7.vi.2016,”, “-2.06142 138.743949, Sumoked (Pap061)” (NHMW). 1 male, 1 female “Indonesia: West Papua, Sarmi Dest, Tor riv. Togonfo vill. 2°15'07.03"S 138°51'25.89"E, 80 m, hollowed log with water, 25.v.2019, leg. Bretschneider Exp.” (ZSM).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 47).

Measurements : TL 5.4–5.7 mm, TL-H 4.95–5.25 mm, MW 2.6–2.7 mm, TL/MW 2.04–2.11; PL 0.8–0.85 mm, PW 2.3–2.4 mm, PL/PW 0.35; DBE 0.9–0.95 mm, DBE/PW 0.38–0.4. Holotype: TL 5.5 mm, TL-H 5.1 mm, MW 2.7 mm, TL/MW 2.04; PL 0.85 mm, PW 2.4 mm, PL/PW 0.35; DBE 0.9 mm, DBE/PW 0.38.

Colouration : With piceous elytra and pronotum and yellowish red head, pronotal sides, and basal band and large apical spot on elytron (Fig. 47).

Head yellowish red, narrowly piceous behind eyes. Pronotum piceous on disc and yellowish red on sides, especially at anterior angles. Elytron piceous, with distinct, rather broad, yellowish red basal band and large, often extending between striae and laterally, yellow spot on apex. Scutellum reddish brown to piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs distinctly darker, especially distally. Ventral side reddish brown to brown, paler on pro- and metaventrite and abdominal ventrites 1–3.

Surface sculpture : Elytron with 11+1 striae (Fig. 47).

Head without strioles, with rather dense punctation (spaces between punctures 1–3× size of punctures); punctures rather coarse (diameter of punctures usually equal to and larger than diameter of microreticulation cells); microreticulation distinct. Pronotum with several strioles posterolaterally or in females, more numerous laterally, with punctation finer than on head and microreticulation fine. Elytron with 11 complete dorsal striae, striae 1 and 10 usually shortly reduced basally; submarginal stria present. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum broadly rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws relatively long, slightly curved, subequal in length, anterior claw thicker subapically than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, with broadly pointed, more or less straight apexes; lateral side of left dorsal lobe with large but shallow, subapical concavity, covered with numerous, small spinula-like structures; lateral margin slightly concave apically, without surface structures, smooth; right dorsal lobe covered with large scale-like structures, with very small, indistinct median membranous area; lobes of ventral sclerite partly sclerotised, long, subequal in length, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, concave subapically, with apex pointed and slightly curved to right and membranous right part with long, thin, upwardly curved apex distinctly projecting in left lateral view; right ventral lobe with narrow sclerotised area on right margin. Parameres with dense, long setae occupying distinctly more than half of dorsal margin; setae on small area proximally shorter and sparser, several most proximal setae standing separately (Fig. 96).

Female : As males but with stronger dorsal punctation and microreticulation and more numerous pronotal strioles.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe not completely covered by spinula-like structures, upper lateral margin without spinulae, smooth in left lateral and ventral views. Based on body size and shape and dorsal colouration, it is similar to A. flavocapitatus sp. nov. and A. kokodensis sp. nov. but differs from them in left dorsal lobe of the median lobe with large lateral concavity.

The species is named after our colleague Tobias Mainda, a great naturalist and next-generation rove beetle expert. The specific epithet is a substantive in the genitive case.

Indonesia: Papua Province: Sarmi Regency (Fig. 108).

In Foja Mts, the species was collected from small puddles near a waterfall in forest. The specimens from Togonfo were sampled from a water-filled hollow in a tree trunk (Fig. 114).

Indonesia: Papua Province: Mamberamo Raya Regency, Rouffaer Mts, Noiadi, ca. 02°46'S, 137°46'E, 150–200 m a.s.l.

Holotype : male “IRAN JAYA: Jayapura Prov. Mamberamo, Rouffaer Mts., Noiadi, 150–200m 17.3.1999, leg. Riedel” (NHMW). Paratypes: 1 male with the same label as the holotype (NHMW).

Body size and form : Beetle large, oblong-oval (Fig. 40).

Measurements : TL 7.5–7.9 mm, TL-H 6.8–7.3 mm, MW 3.6–3.9 mm, TL/MW 2.03–2.08; PL 1.1–1.15 mm, PW 3.3–3.45 mm, PL/PW 0.33; DBE 1.3–1.35 mm, DBE/PW 0.39. Holotype: TL 7.5 mm, TL-H 6.8 mm, MW 3.6 mm, TL/MW 2.08; PL 1.1 mm, PW 3.3 mm, PL/PW 0.33; DBE 1.3 mm, DBE/PW 0.39.

Colouration : Dorsally piceous, with reddish brown head and reddish pronotal sides, elytral basal band and apex (Fig. 40).

Head reddish brown, narrowly darker behind eyes. Pronotum piceous on disc and gradually paler (to reddish) to sides. Elytron piceous, with narrow yellowish red to reddish basal band and yellowish red spot on apex. Scutellum reddish brown to piceous. Antennae, other head appendages, and pro- and mesolegs reddish, metalegs darker, all legs darker distally. Ventral side piceous, paler on prosternum and posteriorly on metacoxal plates and abdominal ventrites.

Surface sculpture : Elytron with 11 dorsal striae, stria 1 reduced to strioles in apical 1/2, some other striae reduced or interrupted basally, submarginal stria present: 11+1 (Fig. 40).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (usually diameter of punctures equal to or smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum without or with few strioles posterolaterally, with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae: stria 1 absent on basal 1/2 and reduced to strioles in apical 1/2, striae 2 and 3 strongly reduced basally, sometimes striae 5–7 shortly reduced basally; submarginal stria present, long. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia modified: thinner proximally and broader medially and distally due to its curved ventral margin. Proclaws long, subequal in length, anterior claw thicker apically and more strongly curved downwards than posterior one and due to this with a subapical incision on its inner margin. Median lobe of aedeagus robust, with two lobes of dorsal sclerite covered with scale-like structures; left dorsal lobe slightly concave medially, its whole apical 1/2 broad, more or less evenly tapering to rather large, elongate, curved downwards, truncate apex bearing broad crest; right dorsal lobe shorter than left one, broad, with pointed apex and membranous area medially; lobes of ventral sclerite partly sclerotised, pressed together between lobes of dorsal sclerite, slightly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area and less sclerotised right part; sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2; less sclerotised right part long and broad, with its apical part densely covered with long setae (sometimes visible at apex); right ventral lobe partly sclerotised, long and broad. Parameres with dense, long setae occupying approximately half of dorsal margin (Fig. 89).

Female : Unknown.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, in shape of the median lobe, it is similar to A. madangensis sp. nov. and differs from it in elytron darker, with narrow, yellowish red basal band and apex of left dorsal lobe of the median lobe more elongate, with more transverse, narrower crest. See also under A. madangensis sp. nov. and A. noiadi sp. nov.

The species is named after the Mamberamo Raya Regency. The name is a noun in the nominative singular standing in apposition.

Indonesia: Papua Province: Mamberamo Raya Regency. The species is known only from the type locality (Fig. 108).

Unknown.

Papua New Guinea: Sandaun Province, Mianmin, 04°53'17.5"S, 141°34'07.1"E, 670 m a.s.l.

Holotype : male “Papua New Guinea: Sandaun, Mianmin, 670m 20.x.2008, 4.53.292S 141.34.118E, Ibalim (PNG 191)” (ZSM). Paratypes: PNG: Sandaun: 22 male, 36 female with the same label as the holotype (NHMW, ZSM). 1 male, 1 female “Papua New Guinea: Sandaun, Mianmin (pool), 990m, 23.x.2008, 4.54.570S 141.35.490E, Ibalim (PNG 193)” (ZSM). 1 male, 1 female “Papua New Guinea: Sandaun, Mianmin (pool), 1080m, 24.x.2008, 04.55.780S 141.38.185E, Ibalim (PNG 196)”, male with an additional green label “3771” (ZSM). 2 males, 2 females “Papua New Guinea: Sandaun, Mianmin (river) 700m, 21.x.2008, 04.52.858S 141.31.706E Ibalim (PNG 197)”, one male with an additional green label “3786” (ZSM). 8 males, 14 females “Papua New Guinea: Sandaun, Mianmin (pool), 700m, 21.x.2008, 04.52.858S 141.31.706E, Ibalim (PNG 198)” (NHMW, ZSM). 7 males, 7 females “Papua New Guinea: Sandaun, Mianmin, 670m, 22.x.2008, 04.53.329S 141.35.263E S. Ibalim PNG189” (NHMW, ZSM). 5 males, 1 female “Papua New Guinea: Sandaun, Mianmin area, >700m, 7.i.2010, Ibalim & Pius (PNG231)”, two males with additional green text labels “6017” and “6018” (ZSM). 1 male “Papua New Guinea: Sandaun, Mianmin, Fak River, 775m, 13.xi.2003, 4.53.53.00S 141.36.39.40E, K. Sagata (WB31)”, “4197” [green label] (ZSM).

PNG: Madang: 1 male, 4 females “Papua New Guinea: Madang, Trans Gogol, 30 m, ii.2008, 5 18.0915S 145 36.4532E, BRC leg. (PNG 179)” [teneral specimens] (ZSM).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 44).

Measurements : TL 5.6–6.4 mm, TL-H 5.2–5.8 mm, MW 2.8–3.1 mm, TL/MW 2–2.07; PL 0.8–0.9 mm, PW 2.3–2.6 mm, PL/PW 0.34–0.35; DBE 0.9–0.95 mm, DBE/PW 0.36–0.39. Holotype: TL 6 mm, TL-H 5.5 mm, MW 2.9 mm, TL/MW 2.07; PL 0.85 mm, PW 2.5 mm, PL/PW 0.34; DBE 0.9 mm, DBE/PW 0.36.

Colouration : Dorsally piceous, with yellowish red to reddish brown head, anterior angles of pronotum, and yellow to yellowish red spot on elytral apex (Fig. 44).

Head yellowish red to reddish brown, narrowly piceous behind eyes. Pronotum piceous, yellowish red to reddish brown in anterior of sides, sometime more, sometimes less extending. Elytron piceous, with yellow to yellowish red apical spot, sometimes rather small and rounded, sometimes larger or even slightly extending laterally. Scutellum piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red to reddish brown, metalegs darker, especially distally. Ventral side piceous, with brown prosternum.

Surface sculpture : Elytron with 11+1 striae; stria 1 often differently reduced (Fig. 44).

Head without strioles, with rather dense punctation (spaces between punctures 1–3× size of punctures); punctures rather coarse (diameter of punctures usually equal to or larger than diameter of microreticulation cells); microreticulation distinct. Pronotum with distinct strioles laterally, with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae, stria 1 usually weakly impressed, often absent in basal 1/2 or present as strioles, maximal reduction to few strioles in apical 1/2, striae 2 and 10 can be shortly reduced basally; submarginal stria present. Elytron with fine but distinct punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively long, slightly curved, subequal in length, anterior claw thicker at its apex than posterior one and due to this with a subapical incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, more or less straight, with very broadly pointed apexes, apex of left dorsal lobe slightly curved downwards; left dorsal lobe with lateral side covered with numerous, distinct, strong spinula-like structures; lateral margin apically without surface structures, smooth; right dorsal lobe covered with scale-like structures, with distinct median membranous area; lobes of ventral sclerite partly sclerotised, long, subequal in length, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, concave subapically, with apex pointed and slightly curved to right and membranous right part long, thin, apically covered with setae sometimes visible in left lateral view; right ventral lobe with narrow sclerotised area on right margin. Parameres with dense, long setae occupying slightly more than 1/2 of dorsal margin (Fig. 93).

Female : As males but with stronger dorsal punctation and microreticulation and pronotum with strioles, usually numerous, occupying sides completely so that only disc without strioles.

There is a variation in dorsal colouration and striolation as described above.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe not completely covered by spinula-like structures, upper lateral margin without spinulae, smooth in left lateral and ventral views. Based on body size and shape and dorsal colouration and striolation, it is similar to A. kalibumi sp. nov. but differs from it in paler head colour and median lobe without concavity of its left dorsal lobe. See also under A. pseudomianminensis sp. nov., A. flavocapitatus sp. nov. and A. sararti sp. nov.

The species is named after Mianmin area. The name is an adjective in the nominative singular.

Papua New Guinea. The species is mainly known from Mianmin area of the Sandaun Province; five teneral specimens, most likely belonged to the species, are recorded from Madang Province (Fig. 108).

In the Mianmin area, the species was collected from a forest pool and a river side pool.

Papua New Guinea: Morobe Province, Garaina, 07°52.516'S, 147°10.427'E, 770 m a.s.l.

Holotype : male “Papua New Guinea: Garaina, 770m, vi.2008 07.52.516S 147.10.427E Ibalim & Sosanika (PNG219)” (ZSM). Paratypes: 2 males, 4 females with the same label as the holotype (NHMW, ZSM). 1 female “Papua New Guinea: Garaina, 770m, 25.vi.2008, 07.50.859S 147.08.614E, Ibalim & Sosanika, (PNG222)” (ZSM)

Body size and form : Beetle large, elongate (Fig. 22).

Measurements : TL 7.3–8.2 mm, TL-H 6.5–7.4 mm, MW 3.3–3.8 mm, TL/MW 2.16–2.21; PL 1.1–1.25 mm, PW 2.95–3.45 mm, PL/PW 0.36–0.37; DBE 1.25–1.4 mm, DBE/PW 0.41–0.42. Holotype: TL 8.2 mm, TL-H 7.4 mm, MW 3.8 mm, TL/MW 2.16; PL 1.25 mm, PW 3.45 mm, PL/PW 0.36; DBE 1.4 mm, DBE/PW 0.41.

Colouration : Dorsally piceous, with yellowish red head and pronotal sides (Fig. 22).

Head yellowish red, narrowly piceous behind eyes. Pronotum reddish brown to piceous on disc and gradually paler on sides, to yellowish red at anterior angles. Elytron piceous, seldom with faint, small reddish spot(s) at shoulder and/or reddish along suture. Scutellum reddish to piceous. Antennae, other head appendages, and legs yellowish red to red, darker distally. Ventral side reddish brown to brown.

Surface sculpture : Dorsal elytral striation variable, usually with ten complete and partly reduced striae, submarginal stria present: (6–11)+1 (Fig. 22).

Head without strioles, with relatively sparse punctation (spaces between punctures 2–5× size of punctures); punctures relatively fine (diameter of punctures smaller than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum with strioles laterally, with punctation finer than on head and microreticulation fine. Elytron with striation variable, usually with ten dorsal, complete or partly reduced, striae, sometimes with six striae with some strioles between them and stria 1 reduced basally (as in holotype), seldom with 11 stria, stria 1 short, present only medially; submarginal stria long, distinct. Elytron with very fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins distinctly convergent anteriorly. Base of prosternum rounded anteriorly, distinctly convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws very long, slightly curved, subequal in size, anterior claw with slight median incision of its inner margin subapically, thinner apically. Median lobe of aedeagus robust, with two lobes of dorsal sclerite subequal in length and shape, both covered with scale-like structures; left dorsal lobe slightly shorter and apically broader than right dorsal lobe, with a distinct median concavity and its whole apical 1/2 narrowed, its proximal part very large, and its apex slightly curved downwards, with a distinct crest; right dorsal lobe with apex pointed and membranous area medially; lobes of ventral sclerite sclerotised, pressed together between lobes of dorsal sclerite, but partly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area (visible as a triangular in left lateral view) and less sclerotised right part (visible in left lateral view); sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2; less sclerotised right part long and broad, with its apical part densely covered with long setae; right ventral lobe long and broad. Parameres with dense, long setae occupying approximately half of dorsal margin (Fig. 73).

Female : As male, except for more numerous pronotal strioles, sometimes absent only on disc.

There is a variation in the elytral striation and colouration described above.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is similar to A. bundunensis sp. nov. in simple male proclaws and shape of the median lobe but differs from it in larger and elongate body, absence of basal elytral band and slightly different structure of median lobe sclerites: see under A. bundunensis sp. nov. Also see under A. xanthocephalus.

The species name is a combination of the Greek words miltos (red lead, red) and karenon (head), meaning red-headed and refers to the distinct yellowish red head colouration of the species. The species name is an adjective in the nominative singular.

Papua New Guinea: Morobe Province (Fig. 107).

Unknown.

Indonesia: Papua Province: Mamberamo Raya Regency, Rouffaer Mts, Noiadi, ca. 02°46'S, 137°46'E, 150–200 m a.s.l.

Holotype : male “IRAN JAYA: Jayapura Prov. Mamberamo, Rouffaer Mts., Noiadi, 150–200m 17.3.1999, leg. Riedel” (NHMW).

Body size and form : Beetle large, oblong-oval (Fig. 37).

Measurements: Holotype: TL 7.9 mm, TL-H 7.1 mm, MW 3.8 mm, TL/MW 2.08; PL 1.1 mm, PW 3.3 mm, PL/PW 0.33; DBE 1.3 mm, DBE/PW 0.39.

Colouration : Elytron piceous in basal 1/2 and yellowish red between striae in apical 1/2 and laterally, with narrow, faint, reddish basal band; head reddish brown, pronotal sides reddish (Fig. 37).

Head reddish brown, narrowly darker behind eyes. Pronotum piceous on disc and gradually paler (to reddish) to sides. Elytron piceous basally on disc and yellowish red between striae in apical 1/2 and laterally, with narrow, faint, reddish basal band, partly conjoined with yellowish red lateral area. Scutellum reddish. Antennae, other head appendages, and pro- and mesolegs reddish, metalegs darker, all legs darker distally. Ventral side piceous, paler on prosternum and posteriorly on abdominal ventrites.

Surface sculpture : Elytron with 11 dorsal striae, stria 1 reduced basally, submarginal stria present: 11+1 (Fig. 37).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (usually diameter of punctures equal to or smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum with strioles posterolaterally, with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae: stria 1 reduced basally, striae 2 and 3 interrupted basally; submarginal stria present. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws relatively long, subequal in length, anterior claw thicker subapically and slightly more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus robust, with two lobes of dorsal sclerite covered with scale-like structures; left dorsal lobe without concavity, its whole apical 1/2 more or less evenly tapering to small, slightly curved downward and truncate apex bearing small crest; right dorsal lobe shorter than left one, broad, with pointed apex and membranous area medially; lobes of ventral sclerite partly sclerotised, pressed together between lobes of dorsal sclerite, slightly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area and less sclerotised right part; sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2; less sclerotised right part long and broad, with its apical part densely covered with long setae (sometimes visible at apex); right ventral lobe partly sclerotised, long and broad. Parameres with dense, long setae occupying approximately half of dorsal margin (Fig. 87).

Female : Unknown.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, in shape of the median lobe, it is similar to A. mamberamo sp. nov. and differs from it in elytron paler apically, male anterior proclaw thicker subapically, with median incision of its inner margin, and apex of left dorsal lobe of the median lobe distinctly shorter, with smaller crest.

The species is named after Noiadi Village. The name is a noun in the nominative singular standing in apposition.

Indonesia: Papua Province: Mamberamo Raya Regency. The species is known only from the type locality (Fig. 108).

Unknown.

Papua New Guinea: Milne Bay Province, Normanby Island, Sewa Bay, Sibonai, 10°02.418'S, 150°58.461'E, 35 m a.s.l.

Holotype : male “Papua New Guinea: Milne Bay, Normanby Isl., Sewa Bay, Sibonai, 35 m”, “30.vi.2017, 10°02.418’S 150°58.461’E, Riedel”, “8722” [green text] (MZB). Paratypes: 3 males with the same lable as holotype and additional labels with green text “8723”, “8724” and “8727” (NHMW, SMNK, ZSM). 6 males, 4 females “Papua New Guinea: Milne Bay Prov., Normanby Isl., Sewa Bay, Sibonai,”, “10°02.418’S 150°58.461’E, 35 m, beaten & hand-collected,”, “30-VI-2017 – position 1, A. Riedel.” (NHMW, SMNK, ZSM). 3 males, 4 females “Papua New Guinea: Milne Bay Prov., Fergusson Isl., Salamo area, Dei Dei,”, “09°39.386’S 150°52.460’E to 09°39.230’S 150°52.586’E, 110 m,”, “beaten & hand-collected, 28-II-2017 – position 1 to 2, A. Riedel.” (NHMW, SMNK, ZSM). 1 male “Papua New Guinea: Milne Bay, Fergusson Isl., Salamo area, Dei Dei,”, “110 m, 28.iii.2017, 09°39.230’S 150°52.586’E, Riedel”, “8729” [green text] (SMNK).

Body size and form : Beetle large, with oblong-oval habitus (Fig. 21).

Measurements : TL 6.5–7 mm, TL-H 5.8–6.3 mm, MW 3.1–3.4 mm, TL/MW 2.06–2.12; PL 1–1.1 mm, PW 2.7–3 mm, PL/PW 0.36–0.37; DBE 1.1–1.2 mm, DBE/PW 0.4–0.41. Holotype: TL 6.9 mm, TL-H 6.1 mm, MW 3.25 mm, TL/MW 2.12; PL 1 mm, PW 2.8 mm, PL/PW 0.36; DBE 1.15 mm, DBE/PW 0.41.

Colouration : With piceous elytra and pronotum and yellowish red head, pronotal sides, and broad basal band on elytron (Fig. 21).

Head yellowish red, narrowly piceous behind eyes. Pronotum piceous, yellowish red on sides, broader at anterior angles. Elytron piceous, with yellowish red broad basal band, often with yellowish on apex, sometimes expanded between striae laterally. Scutellum piceous. Antennae, other head appendages, and legs yellowish red, legs darker distally. Ventral side reddish brown, with paler prosternum.

Surface sculpture : Elytron with six dorsal striae, stria 1 reduced in basal 1/2, submarginal stria present: 6+1 (Fig. 21).

Head without strioles, with relatively dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively coarse (diameter of punctures larger than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum with several strioles at posterolateral angles, with punctation finer than on head and microreticulation fine. Elytron with 6 dorsal striae, stria 1 absent in basal 1/2, submarginal stria present, sometimes only apically. Elytron with very fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins distinctly convergent anteriorly. Base of prosternum broadly rounded anteriorly, distinctly convex medially; blade of prosternal process narrow.

Male : Protibia straight, not modified. Proclaws relatively short, subequal in length, anterior claw slightly thicker subapically and more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus robust, with two lobes of dorsal sclerite subequal in length, with pointed apexes and covered with scale-like structures; left dorsal lobe concave medially, with apex curved downwards, with small apical crest; right dorsal lobe with membranous area medially; lobes of ventral sclerite sclerotised, pressed together between lobes of dorsal sclerite, but partly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area (not visible in left lateral view) and less sclerotised right part (visible in left lateral view); sclerotised area with elongate and broad basal 1/2 and long, hair-like apical 1/2; less sclerotised right part long and broad, with a tuft of long setae; right ventral lobe long and broad. Parameres with dense, long setae occupying approximately half of dorsal margin; more distally situated setae longer than more proximal ones (Fig. 72).

Female : As male.

There is a variation in the elytral striation and colouration described above.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is similar to A. inconstans sp. nov., A. bundunensis sp. nov. and A. centralensis sp. nov. in body form and dorsal colouration, but differs from them in smaller size and median lobe with narrow, distinctly bent downwards apex of left dorsal lobe.

In addition to A. normanbyensis sp. nov., one more Austrelatus species, A. garainensis Shaverdo et al., 2023, is known from Normanby Island (see below). The species belongs to the A. neoguineensis species group, representatives of which have a differently shaped male median lobe without surface sculptures (Shaverdo et al. 2023). The species is distinctly smaller in size than A. normanbyensis sp. nov. and has 11 complete dorsal striae on the elytron.

The species is named after Normanby Island. The name is an adjective in the nominative singular.

Papua New Guinea: Milne Bay Province, Normanby and Fergusson islands (Fig. 107).

Unknown.

Papua New Guinea: Madang Province, 15 km southwest of Madang, Ohu Village.

Holotype : male “PNG Madang Province OHU Village, 14.1.2001 leg.: Lukas Cizek Coll. HENDRICH” (NHMW). Paratypes: 11 males, 21 females with the same label as the holotype (CLH, NHMW, ZSM). 1 male “NEW GUINEA 25 km. N Madang 3 km. W Sempi 27 I 1989 leg. M.&R. Hołyński” (ZSM).

Body size and form : Beetle small, elongate (Fig. 4).

Measurements : TL 4.35–5 mm, TL-H 3.9–4.25 mm, MW 1.9–2.23 mm, TL/MW 2.24–2.29; PL 0.7–0.8 mm, PW 1.73–1.95 mm, PL/PW 0.39–0.42; DBE 0.8–0.95 mm, DBE/PW 0.46–0.5. Holotype: TL 4.8 mm, TL-H 4.25 mm, MW 2.1 mm, TL/MW 2.29; PL 0.75 mm, PW 1.95 mm, PL/PW 0.39; DBE 0.9 mm, DBE/PW 0.46.

Colouration : Dorsally dark brown, with yellowish red head, pronotal sides and basal band on elytron (Fig. 4).

Head yellowish red, narrowly darker behind eyes. Pronotum brown to dark brown on disc and gradually paler (to yellowish red) to sides. Elytron brown to dark brown, with distinct yellowish red basal band and often yellow apex. Scutellum yellowish red to brown. Antennae, other head appendages, and pro- and mesolegs yellowish to yellowish red, metalegs darker, all legs darker distally. Ventral brown. Specimens slightly to strongly teneral.

Surface sculpture : Elytron with six dorsal striae and usually with submarginal stria: 6+(0–1) (Fig. 4).

Head without strioles, with distinct, sparse punctation (spaces between punctures 2–5× size of punctures); punctures relatively fine (diameter of punctures more or less equal to diameter of microreticulation cells); microreticulation distinct. Pronotum without strioles, with punctation as on head and microreticulation distinct. Elytron with six dorsal striae, sometimes striae 1, 3, 5, and 6 shortly reduced basally and/or interrupted, stria 1 can be more strongly reduced; submarginal stria present, seldom absent or represented by tiny strioles. Elytron with fine punctation and microreticulation.

Structures : Head large and broad. Pronotum slightly trapezoid, its lateral margins subparallel. Base of prosternum broadly rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws very long, straight, subequal in length. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, with broadly pointed apexes; left dorsal lobe with lateral margin slightly concave apically and lateral side slightly concave medially, covered laterally with numerous distinct spinula-like structures and dorsally with less numerous scale-like structures, lateral margin without surface structures, smooth; right dorsal lobe covered with scale-like structures; lobes of ventral sclerite partly sclerotised, long, partly visible in left lateral view; left ventral lobe with elongate, narrow sclerotised area, concave apically, with apex pointed and slightly curved; right ventral lobe longer than left lobe, with narrow sclerotised area on right margin, membranous part with long, thin, upwardly curved apex distinctly sticking out in left lateral view. Parameres with relatively dense and long setae occupying distinctly more than half of dorsal margin; setae on small area proximally shorter and sparser, with several most proximal setae standing separately (Fig. 67).

Female : Only matt, strongly striolated form known.

There is an insignificant variation in the elytral striation.

In body size and shape, elytral striation and dorsal colouration, the species is similar to A. procerus sp. nov. and A. lopintolensis sp. nov. but distinctly differs from them by its long, straight male proclaws and shape of the median lobe. See also under A. iriatoi sp. nov.

The species is named after its type locality, Ohu Village. The name is a noun in the nominative singular standing in apposition.

Papua New Guinea: Madang Province (Fig. 107).

Unknown.

Papua New Guinea: Central Province, Kokoda, 08°52'28.4"S, 147°44'16.1"E, ca. 365 m a.s.l. Note: In Balfour-Browne (1939: 87), it is given as “♂ holotype, ♀ allotype, 14 ♂♂ and 10 ♀♀ paratypes, Papua: Kokoda, 1200ft., v.1933 (Miss Chesman) (British Museum, 1933-577).”

Holotype : male “♂” [hw, next to beetle], “Type” [round label with red frame], “PAPUA: Kokoda. 1,200ft. v.1933. L.E.Cheesman. B.M.1933-577.”, “57” [hw on reverse side], “Copelatus papuensis, B-B. ♂ TYPE” [hw by J. Balfour-Browne], “Manuscript name” [printed in red], “Holotype” [red label] (BMNH). Paratypes: 1 female “♀” [hw, next to beetle], “Type” [round label with red frame], “PAPUA: Kokoda. 1,200ft. v.1933. L.E.Cheesman. B.M.1933-577.”, “57” [hw on reverse side], “Copelatus papuensis, B-B. ♀ TYPE” [hw by J. Balfour-Browne] (BMNH). 1 male “♂” [hw, next to genitalia], “Type” [round label with yellow frame], “PAPUA: Kokoda. 1,200ft. v.1933. L.E.Cheesman. B.M.1933-577.”, “57” [hw on reverse side], “Copelatus papuensis, B-B. ♂” [hw by J. Balfour-Browne] (BMNH). 8 males, 7 females “PAPUA: Kokoda. 1,200ft. v.1933. L.E.Cheesman. B.M.1933-577.”, “57” [hw on reverse side], “Type” [round label with yellow frame], “Copelatus papuensis, B-B. ♂ [or ♀]” [hw by J. Balfour-Browne] (BMNH). 1 male “Under stones: river side.”, “PAPUA: Kokoda. 1,200ft. v.1933. L.E.Cheesman. B.M.1933-577.”, “57” [hw on reverse side], “Type” [round label with yellow frame], “Copelatus papuensis, B-B. ♂” [hw by J. Balfour-Browne] (BMNH). 1 female “♀” [hw, next to beetle], “Type” [round label with yellow frame], “PAPUA: Kokoda. 1,200ft. v.1933. L.E.Cheesman. B.M.1933-577.”, “57” [hw on reverse side], “Copelatus papuensis, B-B. ♀” [hw by J. Balfour-Browne] (BMNH). 1 female “♀” [hw, next to beetle], “Type” [round label with yellow frame], “PAPUA: Kokoda. 1,200ft. v.1933. L.E.Cheesman. B.M.1933-577.”, “57” [hw on reverse side], “Copelatus papuensis, B-B. ♀” [hw by J. Balfour-Browne], “measured J. Parkin 73” (BMNH). 1 male “♂” [hw, next to beetle], “Type” [round label with yellow frame], “PAPUA: Kokoda. 1,200ft. v.1933. L.E.Cheesman. B.M.1933-577.”, “57” [hw on reverse side], “Copelatus papuensis, B-B. ♂” [hw by J. Balfour-Browne] (BMNH). 1 male “PAPUA: Kokoda. 1,200ft. v.1933. L.E.Cheesman. B.M.1933-577.”, “57” [hw on reverse side], “Type” [round label with yellow frame], “Copelatus papuensis, B-B. ♂” [hw by J. Balfour-Browne], “measured J. Parkin 72” (BMNH).

PNG: Morobe: 188 males, 98 females “Papua New Guinea: Garaina, 720m, vi.2008 07 51.032S 147 07.007E Ibalim & Sosanika (PNG216)” (NHMW, ZSM). 32 males, 33 females “Papua New Guinea: Garaina 800m, vi.2008 07.53.091S 147.07.915E Ibalim & Sosanika, PNG217” (NHMW, ZSM). 16 males, 9 females “Papua New Guinea: Garaina, 770m, vi.2008 07 52.516S 147.10.427E Ibalim & Sosanika (PNG219)”, one of males additionally with a green label “4074” (NHMW, ZSM). 4 males, 1 female “Papua New Guinea: Garaina, 800m, 27.vi.2009 (PNG220) 7.52.669S 147.07.196E Ibalim & Sosanika”, one of males additionally with a green label “3845” (ZSM). 19 males, 11 females “Papua New Guinea: Garaina, 770m, 25.vi.2008, 07.50.859S 147.08.614E, Ibalim & Sosanika, (PNG222)” (NHMW, ZSM). 1 female “Papua New Guinea: Garaina, 820m, 24.vi.2008 07.52.287S 147.06.297E, Ibalim & Sosanika, (PNG224)” (ZSM). EHL: 1 male, 1 female “Papua New Guinea: Eastern Highlands, Bena Brigde, Unggai Bena, 1393m, 27.viii.2005, K.Sagata, (WB136)” (ZSM).

Body size and form : Beetle large, oblong-oval or elongate (Figs 34, 35).

Measurements : TL 6.9–7.8 mm, TL-H 6.35–7.1 mm, MW 3.25–3.6 mm, TL/MW 2.12–2.23; PL 0.95–1.15 mm, PW 2.85–3.2 mm, PL/PW 0.33–0.37; DBE 1.2–1.3 mm, DBE/PW 0.4–0.42. Holotype: TL 7.5 mm, TL-H 6.8 mm, MW 3.4 mm, TL/MW 2.21; PL 1.15 mm, PW 3.1 mm, PL/PW 0.37; DBE 1.3 mm, DBE/PW 0.42.

Colouration : Elytron yellowish red to reddish brown between striae, narrowly piceous on pronotal and elytral discs, with yellowish red head and pronotal sides (Figs 34, 35).

Head yellowish red, narrowly darker behind eyes. Pronotum piceous on disc and gradually paler (to yellowish red) to sides. Elytron narrowly brown to piceous on disc, with yellowish red basal band broadened at shoulder area, yellow apex, and usually intensive yellowish red colouration between striae, striae brown to piceous. Scutellum yellowish red to brown. Antennae, other head appendages, and pro- and mesolegs yellowish red to reddish, metalegs darker, all legs darker distally. Ventral side reddish brown, paler on prosternum and abdominal ventrites.

Surface sculpture : Elytron with 10–11 dorsal striae, striae 1–3 reduced in basal 1/2, submarginal stria present: (10–11)+1 (Figs 34, 35).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (usually diameter of punctures equal to or smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum with few to several strioles posterolaterally, more numerous in females, with punctation finer than on head and microreticulation fine. Elytron with 10 or 11 dorsal striae: striae 1–3 reduced in basal 1/2, stria 1 strongly to completely reduced, striae 5 and 9 can be interrupted; submarginal stria present, often as apical strioles. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly, seldom rounded so that pronotal base slightly broader than that of elytra. Base of prosternum broadly rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia straight, not modified. Proclaws long, subequal in length, anterior claw thicker subapically and often more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus robust, with two lobes of dorsal sclerite covered with scale-like structures; left dorsal lobe without distinct median concavity and its whole apical 1/2 more or less evenly tapering to elongate, slightly curved, evenly or unevenly narrowly rounded apex bearing small crest; right dorsal lobe shorter, with membranous area medially; lobes of ventral sclerite partly sclerotised, pressed together between lobes of dorsal sclerite, slightly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area and less sclerotised right part; sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2; less sclerotised right part long and broad, with its apical part densely covered with long setae; right ventral lobe partly sclerotised, long and broad. Parameres with dense, long setae occupying approximately half of dorsal margin (Fig. 85).

Female : Dimorphic: as male but with more numerous pronotal strioles and matt, with pronotum and elytron very densely covered with fine longitudinal strioles. Matt forms seldom; in the studied material, known only from the PNG219 population, with ratio shiny to with strioles is 7:3.

There is a variation in dorsal colouration and striolation described above as well as in shape of left dorsal lobe apex of the median lobe: more or less rounded and curved even within one population. Less rounded, curved apex is characteristic of the holotype.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is similar to A. posmani in shape of the median lobe and differs from it in weaker elytral striation, with striae 1–3 reduced in basal 1/2 and stria 1 sometimes absent, male anterior proclaw with median incision of its inner margin, and apex of left dorsal lobe of the median lobe broader, with more distinct crest. See also under A. posmani sp. nov.

Papua New Guinea: Central, Morobe, and Eastern Highlands provinces (Fig. 108). According to Guignot (1956: 52), the species occurs also in Madang Province, Stephansort, Astrolabe Bay; however, this record has not been confirmed. During our study of Guignot’s collection in MNHN, the specimen, on which the records were based, has been found and indentified as A. madangensis sp. nov. (see above).

Unknown.

Papua New Guinea: Central Province, Moreguina, 10°00'57"S, 148°28'27"E.

Holotype : male “Papua New Guinea: Central, Moreguina 16.viii.2008 Posman (PNG183)” (ZSM). Paratypes: 1 male with the same label as the holotype (ZSM).

Body size and form : Beetle large, oblong-oval (Fig. 33).

Measurements : TL 7–7.7 mm, TL-H 6.3–6.9 mm, MW 3.4–3.6 mm, TL/MW 2.06–2.14; PL 1–1.2 mm, PW 2.9–3.15 mm, PL/PW 0.35–0.38; DBE 1.15–1.2 mm, DBE/PW 0.38–0.4. Holotype: TL 7 mm, TL-H 6.3 mm, MW 3.6 mm, TL/MW 2.06; PL 1 mm, PW 2.9 mm, PL/PW 0.35; DBE 1.15 mm, DBE/PW 0.4.

Colouration : With almost completely yellowish red elytron, yellowish red head and pronotal sides (Fig. 33).

Head yellowish red, narrowly darker behind eyes. Pronotum piceous on disc and broadly yellowish red on sides. Elytron yellowish red due to strongly developed basal band and colouration between striae, piceous on small part of disc, on striae and very narrowly on sides. Scutellum yellowish red. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs darker, all legs darker distally. Ventral side reddish brown, paler on prosternum and abdominal ventrites.

Surface sculpture : Elytron with 11 complete, strongly impressed dorsal striae, submarginal stria present: 11+1 (Fig. 33).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (usually diameter of punctures equal to or smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum with strioles laterally, mainly at posterior margin, with punctation slightly finer than on head and microreticulation fine. Elytron with 11 complete, strongly impressed dorsal striae: stria 1 shortly reduced basally; submarginal stria present, long. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly, distinctly rounded so that pronotal base slightly broader than that of elytra. Base of prosternum broadly rounded anteriorly, convex medially; blade of prosternal process broad.

Male : Protibia more or less straight: its ventral margin very slightly curved proximally. Proclaws long, slightly curved, subequal in length, anterior claw thicker at its apex than posterior one and due to this with a subapical incision on its inner margin. Median lobe of aedeagus robust, with two lobes of dorsal sclerite covered with scale-like structures, with their apexes broadly pointed; left dorsal lobe without distinct median concavity and its whole apical 1/2 relatively narrow, more or less evenly tapering to elongate, straight apex bearing small crest; right dorsal lobe shorter, with small membranous area medially; lobes of ventral sclerite partly sclerotised, pressed together between lobes of dorsal sclerite, slightly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area and less sclerotised right part; sclerotised area with elongate and broad basal 1/2 and long, thread-like apical 1/2; less sclerotised right part long and broad, with its apical part densely covered with long setae; right ventral lobe partly sclerotised, long and broad. Parameres with dense, long setae occupying approximately half of dorsal margin (Fig. 84).

Female : Unknown.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is distinguished by yellow colouration and strong striation of its elytra. It is similar to A. papuensis in shape of the median lobe and differs from it in stronger elytral striation, male anterior proclaw thicker at its apex than posterior one and with subapical incision of its inner margin, and apex of left dorsal lobe of the median lobe narrower, with crest weaker. See also under A. papuensis sp. nov.

The species is named after its collector A. Posman. The name is a noun in the genitive case.

Papua New Guinea: Central Province. The species is known only from the type locality (Fig. 108).

Unknown.

Indonesia: West Papua Province: Sorong Regency, road Sorong-Teminabuan, 01°08'08.6"S, 131°54'00.1"E, 130 m a.s.l.

Holotype : male “Indonesia: Papua Barat, Sorong-Teminabuan, 130 m, 2.x.2014, -1,1357267 131,9000149, UNIPA team (BH047)” (KSP). Paratypes: 2 females “Indonesia: Papua Barat, Sorong-Teminabuan, 50 m, 2.x.2014, -1,1092904 131,6125645, B. Sumoked (BH046)”, with two green text labels “6453” and “6454”, respectively (MZB, ZSM).

Body size and form : Beetle small, distinctly elongate (Fig. 2).

Measurements : TL 4.5–4.55 mm, TL-H 4.1 mm, MW 2–2.05 mm, TL/MW 2.22–2.25; PL 0.7–0.75 mm, PW 1.83–1.9 mm, PL/PW 0.38; DBE 0.85–0.9 mm, DBE/PW 0.47. Holotype: TL 4.5 mm, TL-H 4.1 mm, MW 2.05 mm, TL/MW 2.22; PL 0.75 mm, PW 1.9 mm, PL/PW 0.38; DBE 0.9 mm, DBE/PW 0.47.

Colouration : Elytra piceous, head and pronotum paler; yellow basal band on elytron distinct and broad (Fig. 2).

Head yellowish red, narrowly darker behind eyes. Pronotum dark brown on disc and gradually paler (to yellowish red) to sides. Elytron piceous, with broad yellow basal band and yellow apex. Scutellum yellowish red. Antennae, other head appendages, and pro- and mesolegs yellowish to yellowish red, metalegs darker, all leg darker distally. Ventral side yellowish to yellowish red.

Surface sculpture : Elytron without striae or with 6 weakly impressed striae, submarginal stria absent: (0–6)+0 (Fig. 2).

Head without strioles, with distinct, uneven punctation (spaces between punctures 1–5× size of punctures); punctures fine (diameter of punctures smaller than or equal to diameter of microreticulation cells); microreticulation distinct. Pronotum without or with short, weak strioles on sides, with punctation sparser than on head and microreticulation distinct. Elytron without striae, with four puncture lines, or as in holotype, with six weakly impressed striae: striae 2 and 4 complete, other striae strongly reduced, especially basally. Elytron with fine, sparse punctation and distinct microreticulation. Abdominal ventrite 6 with distinct punctation sparse medially and forming denser area at each lateral side, with few strioles laterally.

Structures : Head broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process short and broad.

Male : Protibia straight, not modified. Proclaws long, subequal in length, anterior claw thicker subapically and more strongly curved downwards than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, covered with scale-like structures; left dorsal lobe with lateral margin concave apically, with its side concave subapically and medially, and with some spinula-like structures medially in left lateral view; apex of left dorsal lobe straight, with thickened tip; left lobe of ventral sclerite with elongate, narrow sclerotised area, with its apex visible in left lateral view; right lobe of ventral sclerite membranous. Parameres with relatively long and dense setae occupying distinctly more than half of dorsal margin; more distally situated setae denser and longer than more proximal ones (Fig. 60).

Female : As male, but elytron without striae and pronotum with numerous, weak strioles on its sides.

There is a variation in the elytral striation described above: the holotype has elytron with 6 dorsal striae and both females are without elytral striae.

The species is similar to A. leptos sp. nov. in small size, elongate habitus, and dorsal colouration, but differs from it in sturdier habitus, convergent anteriorly pronotal lateral margins, elytron with striae and broader yellow basal band, and different shape of the median lobe and male proclaws.

The species name is a Latin adjective meaning slender, and indicates the elongate habitus of the species.

Indonesia: West Papua Province: Sorong Regency. The species is known only from the type locality (Fig. 107).

The species was collected in small forest pools rich in rotten leaves.

Papua New Guinea: National Capital District Province, Port Moresby, Brown River Road situated nearby to the suburbs Morata and Waigani.

Holotype : male “Stn. No. 195”, “Papua: Pt. Moresby Brown R. Rd., 15.III.1965”, “M.E.Bacchus, B.M. 1965-120” (BMNH).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 53).

Measurements: Holotype: TL 6.9 mm, TL-H 6.2 mm, MW 3.3 mm, TL/MW 2.09; PL 0.95 mm, PW 2.7 mm, PL/PW 0.35; DBE 1 mm, DBE/PW 0.37.

Colouration : With piceous elytra and pronotum and yellowish red head, pronotal sides, and elytral basal band and apex (Fig. 53).

Head almost completely yellowish red, narrowly piceous behind eyes and with V-shaped, brown spot medially. Pronotum piceous on disc and broadly yellowish red on sides, especially at anterior angles. Elytron piceous, with rather broad yellowish red basal band; elytron broadly yellow apically, usually extending between striae, especially laterally (up to elytral half). Scutellum piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs darker, especially distally. Ventral side piceous, with paler prosternum.

Surface sculpture : Elytron with 11+1 striae, striae 1–3 reduced basally (Fig. 53).

Head without strioles, with relatively dense punctation (spaces between punctures 1–3× size of punctures); punctures relatively fine (diameter of punctures usually equal to and smaller than diameter of microreticulation cells); microreticulation distinct. Pronotum without strioles, with punctation finer than on head and microreticulation fine. Elytron with 11 dorsal striae, stria 1 absent in basal 1/2 and interrupted in apical 1/2; striae 2 and 3 absent in basal ¼ and additionally interrupted basally; submarginal stria present, well-developed. Elytron with fine punctation and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively long; anterior claw shorter, thicker and more curved than posterior one and due to this with a median incision on its inner margin. Median lobe of aedeagus relatively broad, with two lobes of dorsal sclerite with narrowly rounded apexes, left straight, right slightly curved to left; left dorsal lobe slightly longer than right dorsal lobe, distinctly narrowed to apex in its apical 1/3; lateral side of left dorsal lobe covered with large scale-like structures towards dorsal side and smaller scale-like structures towards apex, with numerous small spinula-like structures on slightly beaded lateral margin and numerous large, short spinula-like structures medially; right dorsal lobe covered with large scale-like structures, with small median membranous area; lobes of ventral sclerite sclerotised, long, broad, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, with curved apex and broad slightly sclerotised part rounded apically; right ventral lobe distinctly longer than left one, slightly sclerotised, with long curved apex visible in left lateral view. Parameres with dense, long setae occupying distinctly more than half of dorsal margin; on small area proximally very distinctly shorter and sparser (Fig. 102).

Female : Unknown.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe completely covered by scale- and spinula-like structures in left lateral and ventral views. Based on dorsal colouration and general shape of the median lobe, it is similar to A. asteios sp. nov., A. epicharis sp. nov., A. wasurensis sp. nov., and A. yeretuar sp. nov. but differs from them in larger body size and left dorsal lobe of the median lobe with narrow, shallow, concavity under the lateral margin and different organisation of the surface structures. In body size, shape and colouration, it is very similar to A. gestroi sp. nov. but differs from it in the elytron with a broader yellowish red basal band and different shape of the median lobe: the left dorsal lobe is broader and its apical part is distinctly shorter.

The species is named after its similarity to A. gestroi. The name is a noun in the nominative singular standing in apposition.

Papua New Guinea: National Capital District. The species is known only from the type locality (Fig. 108).

Unknown.

Indonesia: Papua Province: Puncak Regency, Iratoi, Rouaffer River, 3°14'25.1"S, 137°19'58.7"E, 164 m a.s.l.

Holotype : male “Indonesia: Papua, Rouaffer, Iratoi, hill in forest, 164m, 6.ix.2014, -3,2403086 137,3329744, UNCEN team (PAP028)” (KSP). Paratypes: 2 females with the same label as the holotype (MZB, ZSM).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 45).

Measurements : TL 5.3–5.5 mm, TL-H 4.95–5.1 mm, MW 2.6–2.7 mm, TL/MW 2.04–2.06; PL 0.75–0.8 mm, PW 2.3 mm, PL/PW 0.33–0.35; DBE 0.85 mm, DBE/PW 0.37. Holotype: TL 5.5 mm, TL-H 5.05 mm, MW 2.7 mm, TL/MW 2.04; PL 0.8 mm, PW 2.3 mm, PL/PW 0.35; DBE 0.85 mm, DBE/PW 0.37.

Colouration : Dorsally piceous, with yellowish red to reddish brown head, anterior angles of pronotum, and yellow spot on elytral apex (Fig. 45).

Head yellowish red to reddish, narrowly piceous behind eyes. Pronotum piceous, yellowish red to reddish on anterior half of sides. Elytron piceous, with large yellow apical spot, slightly extending laterally. Scutellum reddish to piceous. Antennae, other head appendages, and pro- and mesolegs yellowish red, metalegs distinctly darker, especially distally. Ventral side piceous, with brown prosternum.

Surface sculpture : Elytron with 11+1 complete, rather strongly impressed striae (Fig. 45).

Head without strioles, with rather dense punctation (spaces between punctures 1–3× size of punctures); punctures rather coarse (diameter of punctures usually equal to diameter of microreticulation cells); microreticulation distinct. Pronotum with distinct strioles broadly laterally, only disc without them, with punctation finer than on head and microreticulation fine. Elytron with 11 complete, rather strongly impressed dorsal striae, stria 10 shortly reduced basally; submarginal stria present. Elytron with fine and microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively narrow.

Male : Protibia straight, not modified. Proclaws relatively long, slightly curved, subequal in length, anterior claw thicker at its apex than posterior one and due to this with a subapical incision on its inner margin. Median lobe of aedeagus with two lobes of dorsal sclerite subequal in length, more or less straight, with very broadly pointed apexes, apex of left dorsal lobe slightly curved downwards; left dorsal lobe with lateral side covered with numerous, distinct, strong spinula-like structures; lateral margin apically without surface structures, smooth; right dorsal lobe covered with scale-like structures, with distinct median membranous area; lobes of ventral sclerite partly sclerotised, long, subequal in length, partly visible in left lateral view; left ventral lobe with elongate, strong sclerotised area, concave subapically, with apex pointed and slightly curved to right and membranous right part long, thin, apically covered with setae sometimes visible in left lateral view; right ventral lobe with narrow sclerotised area on right margin. Parameres with dense, long setae occupying approximately half of dorsal margin; more distally situated setae longer than more proximal ones (Fig. 94).

Female : As males but with stronger dorsal punctation.

The species belong to the medium-sized species with median lobe of the aedeagus that has scale- and spinula-like structures on the left dorsal lobe and among them, to the species with surface of left dorsal lobe not completely covered by spinula-like structures, upper lateral margin without spinulae, smooth in left lateral and ventral views. Based on shape of the median lobe, it is very similar to A. mianminensis sp. nov. but differs from it in smaller body size, elytral striae complete and rather strongly impressed, more numerous pronotal strioles, and smaller median lobe, with the lateral margin of left dorsal lobe with edge, below which spinula-like structures situated, more sharp, prominent. See also under A. flavocapitatus sp. nov. and A. sararti sp. nov.

At the beginning, the species was considered to belong to A. mianminensis sp. nov. due to their similarity. The name is a noun in the nominative singular standing in apposition.

Indonesia: Papua Province: Puncak Regency. The species is known only from the type locality (Fig. 108).

The species was collected in small and shaded forest pools rich in rotten leaves.

Papua New Guinea: Madang Province, Highway near Madang, 05°24.405'S, 145°38.213'E, 80 m a.s.l.

Holotype : male “Papua New Guinea: Madang, Highway nr. Madang, ford, 80m, 26.xi./2.-3.xii.1994, 05.24.405S 145.38.213E, Binatang Boys, (PNG 117)” (ZSM). Paratypes: PNG: Madang: 1 male with the same label as the holotype (NHMW). Central: 1 male “Papua New Guinea: Central, Myola, 1110m, i.2008, [09]12.630S 147 31.880E, Posman (PNG 177)”, “4119” [green label] (ZSM).

Body size and form : Beetle large, oblong-oval (Fig. 30).

Measurements : TL 7.45–7.9 mm, TL-H 6.8–7.1 mm, MW 3.75–3.9 mm, TL/MW 1.99–2.03; PL 1.1–1.2 mm, PW 3.35–3.6 mm, PL/PW 0.33–0.33; DBE 1.3–1.45 mm, DBE/PW 0.39–0.4. Holotype: TL 7.45 mm, TL-H 6.8 mm, MW 3.75 mm, TL/MW 1.99; PL 1.1 mm, PW 3.35 mm, PL/PW 0.33; DBE 1.3 mm, DBE/PW 0.39.

Colouration : Dorsally piceous, with reddish head, pronotal sides and elytral base and apex (Fig. 30).

Head red, narrowly darker behind eyes. Pronotum brown on disc and gradually paler (to dark yellowish red) to sides, especially at anterior angles. Elytron dark brown to piceous, with short, faint, dark yellowish red to reddish basal band present on shoulder area and with yellowish red apex, sometimes more developed laterally. Scutellum dark brown. Antennae, other head appendages, and pro- and mesolegs reddish, metalegs darker, all legs darker distally. Ventral side dark brown, sometimes paler medially and on abdominal ventrites.

Surface sculpture : Elytron with 11 complete striae, submarginal stria present: 11+1 (Fig. 30).

Head without strioles, with dense punctation (spaces between punctures 1–3× size of punctures); punctures coarse (usually diameter of punctures larger than diameter of microreticulation cells); microreticulation distinct. Pronotum with few strioles on sides, with punctation finer than on head and microreticulation fine. Elytron with 11 complete striae, striae 1 and 10 usually shortly reduced basally; submarginal stria present. Elytron with fine, sparse punctation and fine microreticulation.

Structures : Head relatively broad. Pronotum trapezoid, its lateral margins convergent anteriorly. Base of prosternum rounded anteriorly, convex medially; blade of prosternal process relatively broad.

Male : Protibia modified: thinner proximally and broader medially and distally due to its curved ventral margin. Proclaws relatively long, subequal in length, anterior claw thicker subapically and more strongly curved downwards than posterior one and due to this with a distinct median incision on its inner margin. Median lobe of aedeagus very robust, with two lobes of dorsal sclerite covered with scale-like structures; left dorsal lobe very strongly concave medially, with apical 1/2 very broad and apex slightly rounded and bearing small but rather distinct crest; right dorsal lobe shorter, narrower, more or less evenly tapering to pointed apex, with small membranous area medially; lobes of ventral sclerite mostly visible in left lateral view; left ventral lobe consists of two parts: left sclerotised area and less sclerotised right part; sclerotised area with elongate and broad basal 1/2 (visible in left lateral view) and long, thread-like apical 1/2 (not visible); less sclerotised right part long and broad, with its apical part densely covered with long setae (partly visible in left lateral view); right ventral lobe mostly sclerotised, long and broad (partly visible in left lateral view). Parameres with dense, long setae occupying slightly more than half of dorsal margin, with proximal setae shorter (Fig. 81).

Female : Unknown.

There is an insignificant variation in dorsal colouration described above.

The species belongs to the large species with robust median lobe, covered mainly with scale-like structures. Among them, it is very similar to A. bosaviensis sp. nov. and differs from it in broader male anterior claw with median inner margin incision and narrower apical 1/2 of the left dorsal lobe of the median lobe. See also under A. bosaviensis sp. nov.

The species name is a Latin adjective meaning “hard and strong like oak” and indicates the large, broad, robust habitus and median lobe of the species.

Papua New Guinea: Madang and Central provinces (Fig. 107).

In Madang, the species was collected at a stream ford.

Indonesia: West Papua: Teluk Wondama Regency, Wasior, Sararti 100–200 m a.s.l.

Holotype : male “Irian Jaya: Wandammen Bay, Wasior, Sararti 100–200 m, 7.-9.I.2001, leg A. Riedel” (SMNS).

Body size and form : Beetle medium-sized, oblong-oval (Fig. 48).

Measurements: Holotype: TL 5.7 mm, TL-H 5.1 mm, MW 2.75 mm, TL/MW 2.07; PL 0.85 mm, PW 2.35 mm, PL/PW 0.36; DBE 0.95 mm, DBE/PW 0.4.