Anaka colorata Dworakowska & Viraktamath, 1975 (type locality: India).

Vertex rounded, coronal suture distinct. Face rounded gradually to vertex, flat, lorum broad. Ocelli vestigial. Pronotum ~ 3× longer than head in dorsal view, as broad as head with eyes. Scutum with scutellum distinctly shorter than pronotum. Forewing broad, 3^rd apical cell stalked. Hind wing veins RP and MA confluent in male. Hindwing narrow, membrane smoked, veins dark, area bordered.

Abdominal apodemes well developed, reaching caudal margin of 4^th abdominal sternite. Pygofer side dark, well sclerotized, dorsal lobe the darkest, hind and ventral margins not pigmented, row of microsetae caudad. Subgenital plate has group of few big macrosetae at approximately mid-length, numerous thin short microsetae present at apical 1/3, several small rigid setae near macrosetae, and a row of thin long setae on basal 1/2 of ventral margin. Paramere hooked at apex, strongly attached to subgenital plate. Connective fused with aedeagus (Dworakowska and Viraktamath 1975). Aedeagus tubular, with basal or apical processes. Dorsoatrium well developed. Gonopore apical.

China (Guizhou, Guangdong, Guangxi, Yunnan, Sichuan, Chongqing, Hunan, Zhejiang, Fujian, Taiwan), India, Nepal, Sumatra, Indonesia, Burma.

Holotype , 1♂, China: Guizhou Province, Daozhen. 28.1892°N, 107.4294°E, H, 1700 m, 14.V.2006, collected by Yang Zaihua.

Length : male 4.2 mm. Body (Fig. 1A, B) sandy beige. Crown (Fig. 1C) with two black patches. Face (Fig. 1D, E) yellowish, frontoclypeal area protuberant, anteclypeus broad. Pronotum yellowish brown, wider than crown. Scutellum yellowish with two blackish patches at lateral corner. Forewing (Fig. 1F) infuscate 3^rd apical cell stalked, hind wing (Fig. 1G) transparent.

Figure 1. 

Anaka auricula sp. nov. A male body, dorsal view B male body, lateral view C head and thorax, dorsal view D head and thorax, lateral view E face F forewing G hindwing H abdominal apodeme I male pygofer, lateral view J subgenital plate, ventral view K aedeagus, connective, and paramere, lateral view L aedeagus and connective, ventral view. Scale bars: 0.5 mm (A–E); 0.1 mm (F–L).

Male abdomen (Fig. 1H) well developed and reaching 4^th abdominal sternite. Pygofer side (Fig. 1I) broad, single row of thin setae on central and apical parts. Basal 1/2 of subgenital plate (Fig. 1J) broad, distal 1/2 slender in lateral view, two large macrosetae at approximately mid-length, several small setae near macrosetae, and scattered setae on distal 1/2. Paramere (Fig. 1K) hooked at apex. Aedeagus (Fig. 1K, L) tubular in lateral view, stem curved at middle, apical processes auricle-shaped with small spine at middle. Gonopore apical.

The specific epithet is derived from the Latin word auricula (an ear) referring to the shape of the aedeagal processes.

This species has an aedeagus very similar in form to that of A. blada, but it differs from that species in having elongated apical processes.

Holotype , 1♂, China: Yunnan Province, Pingbian. 22.9101°N, 103.7008°E, H, 2084 m, 22.V.2015, collected by Yan Bin.

Length : male 4.2 mm. Body (Fig. 2A, B) yellowish. Crown (Fig. 2C) obtuse. Coronal suture distinct. Face (Fig. 2D–F) white, frontoclypeal area protuberant, anteclypeus broad. Pronotum yellowish, wider than crown. Scutellum small. Wings (Fig. 2G, H) without patches.

Figure 2. 

Anaka cruciata sp. nov. A male body, dorsal view B male body, lateral view C head and thorax, dorsal view D head and thorax, lateral view E face F head, frontal view G forewing H hindwing I abdominal apodeme J male pygofer, lateral view K subgenital plate, ventral view L aedeagus, connective, and paramere, dorsal view M aedeagus, connective, and paramere, lateral view. Scale bars: 0.5 mm (A–F); 0.1 mm (G–M).

Male abdomen (Fig. 2I) reaching 4^th abdominal sternite. Pygofer side (Fig. 2J) broad, with a small extension and thin setae on central and apical parts. Basal 1/2 of subgenital plate (Fig. 2K) broad, distal 1/2 slender in lateral view, one large macroseta at approximately midlength, several small setae near macrosetae, and scattered setae on distal 1/2. Paramere (Fig. 2L, M) hooked at apex. Connective fused with aedeagus. Aedeagus (Fig. 2L, M) tubular, curved, with two pairs of apical processes, of which each pair are crossed. Gonopore apical.

The specific epithet is derived from the Latin word cruciatus (marked by a cross) referring to the shape formed by the two pairs of aedeagal processes.

This species with two pairs of aedeagal processes differs from all other species of Anaka, and two pairs of processes originate from subapical of stem, but in different positions.

Holotype , 1♂, China: Guangdong Province, Nanling National Natural Reserve, 24.8796°N, 113.0137°E, H, 1340 m. 4.VIII.2006, collected by Zhou Zhonghui. Paratypes, 4♂♂, China: Guangxi Province, Damingshan National Natural Reserve, 23.5049°N, 108.4153°E, H, 1290 m. 15.IV.2012, collected by Long Jiankun; 6♂♂, China: Guangxi Province, Damingshan National Natural Reserve, 23.4898°N, 108.4411°E, H, 1250 m. 14.V.2012, collected by Huang Rong and Yu Xiaofei.

Length : male 4.4–4.5 mm. Body (Fig. 3A, B) brown. Crown (Fig. 3C) with two black patches. Coronal suture distinct. Face (Fig. 3D, E) yellowish brown, frontoclypeal area protuberant, anteclypeus broad. Pronotum brown, wider than crown. Scutellum with a vertical yellow stripe in the median. Forewing (Fig. 3F) infuscate, 3^rd apical cell stalked, hind wing transparent (Fig. 3G).

Figure 3. 

Anaka curvata sp. nov. A male body, dorsal view B male body, lateral view C head and thorax, dorsal view D head and thorax, lateral view E face F forewing G hindwing H abdominal apodeme I male pygofer, lateral view J subgenital plate, ventral view K aedeagus, connective, and paramere, lateral view L aedeagus, connective, ventral view. Scale bars 0.5 mm (A–E); 0.1 mm (F–L).

Male abdomen (Fig. 3H) reaching 4^th abdominal sternite. Pygofer side (Fig. 3I) broad, thin setae on central and apical parts. Basal 1/2 of subgenital plate (Fig. 3G) broad, distal 1/2 slender in lateral view, three large macrosetae at approximately mid-length, several small setae near macrosetae, and scattered setae on distal 1/2. Paramere (Fig. 3K) hooked at apex. Connective fused with aedeagus. Aedeagus (Fig. 3K, L) tubular, curved, with a pair of apical processes, which are curved like a paper clip. Gonopore apical.

The specific epithet is derived from the Latin word curvatus (curved) referring to the shape of the aedeagal processes.

This species is similar to A. blada, but it differs in having the aedeagus processes more strongly curved and less divergent from the stem.

Holotype , 1♂, China: Guizhou Province, Jinsha, 27.4553°N, 106.2667°E, H, 1300 m, 5.VIII.2015, collected by Zhang Yaowen. Paratypes, 3♂9♀, same data as holotype.

Length : male 4.4–4.5 mm. Body (Fig. 4A, B) white with red patches. Crown (Fig. 4C) obtuse, yellowish white. Coronal suture distinct. Face (Fig. 4D, E) red, frontoclypeal area protuberant, anteclypeus broad, yellowish. Pronotum yellowish, with red patches in the central part, wider than crown. Scutellum yellowish. Forewing (Fig. 4F) white with red patches along inside margin, hind wing transparent (Fig. 4G).

Figure 4. 

Anaka rosacea sp. nov. A male body, dorsal view B male body, lateral view C head and thorax, dorsal view D head and thorax, lateral view E face F forewing G hindwing H abdominal apodeme I male pygofer, lateral view J subgenital plate, ventral view K paramere, lateral view L aedeagus and connective, lateral view M aedeagus and connective, dorsal view. Scale bars 0.5 mm (A–E); 0.1 mm (F–M).

Male abdomen (Fig. 4H) weakly developed and reaching 4^th abdominal sternite. Pygofer side (Fig. 4I) broad, apical part elliptical. Basal 1/2 of subgenital plate (Fig. 4J) broad, distal 1/2 slender in lateral view, one large macroseta at approximately midlength. Paramere (Fig. 4K) hooked at apex. Connective fused with aedeagus. Aedeagus (Fig. 4L, M) tubular, stem inflated at apex, with one pair of apical processes, apical processes straight and sculptured, oriented basad. Gonopore apical.

The specific epithet is derived from the Latin word rosaceus (rose-colored) referring to the color of the head.

This species marked with rose-red spots. The aedeagal processes are similar to A. blada and A. spinosa but differs in having the aedeagus with two long apical processes and the processes straight with spiral pattern.

Holotype , 1♂, China: Yunnan Province, Baoshan, 25.1581°N, 99.0814°E, H, 1500 m, 14.V.2016, collected by Li Bin and Ren Guoru. Paratypes, 3♂4♀, same data as holotype.

Length : male 4.4–4.5 mm. Body (Fig. 5A, B) yellow. Crown (Fig. 5C) obtuse. Coronal suture distinct. Face (Fig. 5D–F) white, frontoclypeal area protuberant, anteclypeus broad. Pronotum yellow, wider than crown. Forewing (Fig. 5G) yellow with apical part white, hind wing (Fig. 5H) transparent.

Figure 5. 

Anaka spiralis sp. nov. A male body, dorsal view B male body, lateral view C head and thorax, dorsal view D head and thorax, lateral view E face F head, frontal view G forewing H hindwing I abdominal apodeme J male pygofer, lateral view K male pygofer lobe, dorsal view L subgenital plate, ventral view M aedeagus, connective and paramere, lateral view N aedeagus and connective, dorsal view. Scale bars 0.5 mm (A–F); 0.1 mm (G–N).

Male abdomen (Fig. 5I) well developed and reaching 5^th abdominal sternite. Pygofer side (Fig. 5J, K) broad, with small extension on superior margin, setae along periphery. Basal 1/2 of subgenital plate (Fig. 5L) broad, distal 1/2 slender in lateral view, one large macroseta at approximately mid-length. Paramere (Fig. 5M) hooked at apex. Connective fused with aedeagus. Aedeagus (Fig. 5M, N) tubular, with one pair of basal processes, apical part of processes spiral and not exceeding the stem. Gonopore apical.

The specific epithet is derived from the Latin word spiralis (spiraling) referring to the shape of the aedeagal processes.

In this species the aedeagus has a pair of basal processes like A. burmensis and A. shashidhari, but these basal processes have spiral-shaped top, and do not exceed the stem. These features are also not as long as in A. nepalica.