Body small, usually yellowish brown. Interocellar distance slightly wider than the distance between eyes, narrower than the distance between antennal sockets. The fifth maxillary palp expanded, the third and fourth palpi both longer than the fifth palp. Pronotum subelliptical, broader than long, the disc usually with symmetrical maculae and stripes. Tegmina and hind wings fully developed, mostly both extending beyond the end of abdomen. Tegmina with M and CuA radial, M straight with 4–7 branches. Hind wings with ScP and RA expanded at apex, CuA with 4–6 complete branches. Anteroventral margin of front femur Type B₂ or B₃. Four proximal tarsomeres with pulvilli. The pretarsi with arolium, tarsal claws symmetrical and specialized, with minute denticles on ventral margins. Eighth abdominal tergum usually specialized, with a tuft of setae in the middle near posterior margin. Supra-anal plate usually short and transverse, paraprocts similar and flaky. Cerci long, with setae on the ventral surface. Male subgenital plate symmetrical or slightly asymmetrical. Styli symmetrical and cylindrical, rarely asymmetrical or non-cylindrical. Male genitalia. Left phallomere small, irregularly bone-shaped, mostly with spine-like process. Median phallomere slender, rod-shaped, the apex irregular and variable; accessory sclerite complicated, generally arched. Hook phallomere on right side, apex usually curved inwards.

The genus Margattea is supposedly closely related to Chorisoserrata (Wang et al. 2023) and morphologically similar to Balta, but Margattea could be distinguished from Chorisoserrata and Balta by the following characteristics. The genus Margattea can be distinguished from Balta (Asia, Africa, and parts of Oceania) by the following characteristics: 1) anteroventral margin of front femur Type B₂ or B₃, in contrast to C₂ (but rarely B₃) in Balta; 2) the tarsal claws symmetrical and specialized, but in the latter, the tarsal claws asymmetrical and unspecialized.

The genus Margattea can be distinguished from Chorisoserrata (parts of Asia and Indonesia) by the following characteristics: 1) anteroventral margin of front femur Type B₂ or B₃, in contrast to C₂ (but rarely B₃) in Chorisoserrata; 2) eighth abdominal tergum usually specialized, with a tuft of setae in the middle near posterior margin; while in the latter, abdominal terga unspecialized.

Holotype : China • ♂; Yunnan Province, Dehong Dai and Jingpo Autonomous Prefecture, Yingjiang County, Nabang Town; 282 m; 17 Aug. 2015; Xin-Ran Li, Zhi-Wei Qiu leg; SWU-B-PS000001. Paratypes: China • 1 ♂ & 1 ♀; same data as holotype; SWU-B-PS000002–000003 • 5 ♂ & 2 ♀; Yunnan Province, Dehong Dai and Jingpo Autonomous Prefecture, Yingjiang County, Nabang Town; 282 m; 11 Jul. 2012; Dong Wang leg; SWU-B-PS000004–000010.

(mm). Male (n = 6), pronotum length × width: 2.4–2.7 × 3.4–3.7, tegmina length: 11.8–12.3, body length: 8.8–10.3, overall length: 13.2–13.7. Female (n = 4), pronotum length × width: 2.4–2.9 × 3.2–3.9, tegmina length: 11.2–11.6, body length: 10.9–12.8, overall length: 13.2–14.0.

Male. Coloration. Body pale yellow (Fig. 3A, B). Head yellowish brown. Face pale yellow. Interocular space with a wide brown transverse band. Ocellar spots white, interocellar space with a brown band. Antennal base pale yellowish brown, other segments dark brown (Fig. 3E). Maxillary palpi pale brown (Fig. 3J). Pronotal disc pale yellowish brown with reddish tan stripes, and two lateral borders pale linen-colored and transparent (Fig. 3F). Tegmina pale yellowish brown, hind wings brownish grey (Fig. 3G, H). Legs faint yellow. Abdomen yellowish brown, with black stripes along lateral margins of sterna and reaching the end of abdomen; both sides of each abdominal sternum with one small black spot on the inside of the longitudinal lines (Fig. 3B). Cerci yellowish brown to pale brown (Fig. 3L). Styli yellowish white (Fig. 3N).

Figure 3. 

Margattea pedata Li & Che, sp. nov. A, B, E–O male C, D female A holotype, dorsal view B holotype, ventral view C paratype, dorsal view D paratype, ventral view E pronotum, dorsal view F head, ventral view G tegmen, ventral view H hind wing, ventral view I eighth abdominal tergum, ventral view J maxillary palpi segments 3–5 K front femur, ventral view L supra-anal plate, ventral view M left phallomere, ventral view N subgenital plate and median phallomere, ventral view O hook phallomere, ventral view. Scale bars: 5 mm (A–D, G, H); 1 mm (E, F, I–L, N); 0.5 mm (M, O).

Head. Vertex slightly exposed, interocellar distance slightly wider than the distance between eyes, narrower than the distance between antennal sockets (Fig. 3F). Fifth maxillary palpus expanded, third and fourth maxillary palpi nearly equal in length, both longer than fifth maxillary palpus (Fig. 3J). Pronotum subelliptical, broader than long, anterior and posterior margins nearly straight, and postero-lateral angle blunt and round; disc with symmetrical spots and stripes (Fig. 3E). Tegmina and hind wings. Tegmina and hind wings fully developed, both extending beyond the end of abdomen (Fig. 3A, B). Tegmina with M and CuA radial, M straight with three complete branches and one incomplete branch. Hind wings with ScP and RA expanded at apex, M simple, without branches; CuA with four complete branches (Fig. 3G, H). Legs. Anteroventral margin of front femur Type B₃ (Fig. 3K). Four proximal tarsomeres with pulvilli. The pretarsi with arolium, tarsal claws symmetrical and specialized, with minute denticles on ventral margins.

Abdomen and genitalia. Eighth abdominal tergum specialized, with a heart-shaped transparent area and a tuft of bristles in the middle (Fig. 3I). Supra-anal plate symmetrical, middle posterior margin slightly concave. Paraprocts simple, similar, and flaky (Fig. 3L). Cerci long, with setae on the ventral surface (Fig. 3L). Styli similar, foot-shaped (Fig. 3N). Subgenital plate nearly symmetrical, posterior margin truncate (Fig. 3N). Left phallomere small, irregular bone-shaped, with a long spine (Fig. 3M). Median phallomere slender rod-shaped, with base curved, apex with a row of spines; accessory sclerite I arched, accessory sclerite II complicated with an inverted bell-shaped structure covered with fuzz (Fig. 3N). Hook phallomere on the right side, apex curved inwards with a short spine (Fig. 3O).

Female. Similar to the male (Fig. 3C, D).

This species is similar to M. speciosa Liu & Zhou, 2011 in general appearance, but can be differentiated from the latter by the following characters: 1) styli foot-shaped, while in the latter conical; 2) left phallomere with a long, curved spine, absent in the latter; and 3) accessory sclerite I without a brush-like structure at apex, while in the latter, accessory sclerite I with a brush-like structure at apex.

The specific epithet is derived from the Latin word pedatus, referring to the foot-shaped styli.

China (Yunnan).

Holotype : China • ♂; Guangxi Zhuang Autonomous Region, Laibin City, Jinxiu Yao Autonomous County, Mountain Shengtang; 1182 m; 5 Jun. 2014; Shun-Hua Gui, Xin-Ran Li leg; SWU-B-PS000011. Paratypes: China • 3 ♂ & 1 ♀; same data as holotype; SWU-B-PS000012–000015 • 1 ♂; Guangxi Zhuang Autonomous Region, Laibin City, Jinxiu Yao Autonomous County, Mountain Shengtang; 400 m; 13 Jul. 2015; Lu Qiu, Qi-Kun Bai leg; SWU-B-PS000016.

(mm). Male (n = 6), pronotum length × width: 2.3–2.5 × 3.0–3.4, tegmina length: 11.8–12.5, body length: 9.8–11.6, overall length: 13.8–14.9. Female (n = 2), pronotum length × width: 2.3–2.5 × 3.3, tegmina length: 10.7–11.4, body length: 10.3–11.0, overall length: 13.8–13.9.

Male. Coloration. Body brown (Fig. 4A, B). Head and face yellowish brown. Interocular space with a wide brown transverse band. Ocellar spots yellowish white, interocellar space with a brown band. Antennae blackish brown, antennal space with a brown band (Fig. 4F). Maxillary palpi dark brown (Fig. 4J). Pronotal disc pale brown with dark brown spots and maculae but without stripes, and two lateral borders pale linen-colored and transparent (Fig. 4E). Legs yellowish brown. Tegmina pale yellowish brown, hind wings brownish grey (Fig. 4G, H). Abdomen pale brown. Cerci brown (Fig. 4L). Styli yellowish brown (Fig. 4N).

Figure 4. 

Margattea bisphaerica Li & Che, sp. nov. A, B, E–O male C, D female A holotype, dorsal view B holotype, ventral view C paratype, dorsal view D paratype, ventral view E pronotum, dorsal view F head, ventral view G tegmen, ventral view H hind wing, ventral view I eighth abdominal tergum, ventral view J maxillary palpi segments 3–5 K front femur, ventral view L supra-anal plate, ventral view M left phallomere, ventral view N subgenital plate and median phallomere, ventral view O hook phallomere, ventral view. Scale bars: 5 mm (A–D, G, H); 1 mm (E, F, I–K, N); 0.5 mm (L, M, O).

Head. Vertex slightly exposed, interocellar distance slightly much wider than the distance between eyes, narrower than the distance between antennal sockets (Fig. 4E). Pronotum subelliptical, broader than long, anterior and posterior margins nearly straight, and postero-lateral angle blunt and round; disc with symmetrical but irregular spots and maculae (Fig. 4F). Fifth maxillary palpus expanded, third and fourth maxillary palpi both longer than fifth maxillary palpus (Fig. 4J). Tegmina and hind wings. Tegmina and hind wings fully developed, both extending beyond the end of abdomen (Fig. 4A, B). Tegmina M and CuA radial, M straight with six complete branches. Hind wings with ScP and RA expanded at apex, M simple, without branches; CuA with four complete branches (Fig. 4 G, H). Legs. Anteroventral margin of front femur Type B₂ (Fig. 4K). Four proximal tarsomeres with pulvilli. The pretarsi with arolium, tarsal claws symmetrical and specialized, with minute denticles on ventral margins.

Abdomen and genitalia. Eighth abdominal tergum specialized, with a tuft of bristles in the middle (Fig. 4I). Supra-anal plate symmetrical, anterior margin straight and truncate, the middle of posterior margin slightly concave. Paraprocts simple, similar, and flaky. Cerci long, with setae on the ventral surface (Fig. 4L). Subgenital plate asymmetrical. Styli dissimilar and spherical, the left stylus significantly smaller than the right stylus (Fig. 4N). Left phallomere small, irregular bone-shaped, with a slender curved spine (Fig. 4M). Median phallomere slender rod-shaped, with a curved spine at apex; accessory sclerite I arched, left end expanded, right end with a cluster of thorns; accessory sclerite II brush-shaped; accessory sclerite III slender rod-shaped (Fig. 4N). Hook phallomere on the right side, apex curved hook-shaped (Fig. 4O).

Female. Similar to the male. Subgenital plate symmetrical, middle posterior margin concave inward (Fig. 4C, D).

This species is similar to M. concava Wang, Che & Wang, 2009 in general appearance, but can be differentiated from the latter by the following characters: 1) styli dissimilar and spherical, the left stylus significantly smaller than the right stylus; while in the latter, styli similar and conical; 2) left phallomere with a slender curved spine, absent in the latter.

The specific name is derived from the Latin words, bi and sphaericus, referring to the dissimilar and spherical styli.

China (Guangxi).

Holotype : China • ♂; Chongqing City, Beibei District, Mountain Jinyun; 550 m; 12 Jul. 2016; Lu Qiu, Zhi-Wei Qiu leg; SWU-B-PS000017. Paratypes: China • 10 ♂ & 1 ♀; same data as holotype; SWU-B-PS000018–000028 • 2 ♂ & 1 ♀; Chongqing City, Jiangjin District, Mountain Simian; 425 m; 21 Sep. 2007; Wei-Wei Zhang leg; SWU-B-PS000029–000031 • 1 ♂ & 1 ♀; Chongqing City, Liangping District, Dongshan Forest Park; 2 Oct. 2007; Wei-Wei Zhang leg; SWU-B-PS000032–000033.

(mm). Male (n = 4), pronotum length × width: 2.4–2.9 × 3.6–3.8, tegmina length: 12.8–13.6, body length: 10.4–12.1, overall length: 14.9–16. Female (n = 4), pronotum length × width: 2.3–2.5 × 3.3, tegmina length: 10.7–11.4, body length: 10.3–11.0, overall length: 13.8–13.9.

Male. Coloration. Body, head and face yellowish brown (Fig. 5A, B). Interocular space with a brown transverse band. Ocellar spots small, yellowish white. Antennal base yellowish brown, other segments black-brown. The third and fourth maxillary palpi yellowish brown, the fifth palpus maxillary blackish brown (Fig. 5J). Pronotal disc yellowish brown with reddish tan spots and stripes, and two lateral borders pale linen-colored and transparent (Fig. 5E). Legs yellowish brown, with black spots at the base of the tibial spines. Tegmina yellowish brown, hind wings brownish grey (Fig. 5G, H). Abdomen yellowish brown, both sides of each abdominal sternum with one small round black spot on the inside of the longitudinal lines. Cerci yellowish brown (Fig. 5L). Styli pale yellow (Fig. 5N).

Figure 5. 

Margattea undulata Li & Che, sp. nov. A, B, E–O male C, D female A holotype, dorsal view B holotype, ventral view C paratype, dorsal view D paratype, ventral view E pronotum, dorsal view F head, ventral view G tegmen H hind wing I eighth abdominal tergum, ventral view J maxillary palpi segments 3–5 K front femur, ventral view L supra-anal plate, ventral view M left phallomere, ventral view N subgenital plate and median phallomere, ventral view O hook phallomere, ventral view. Scale bars: 5 mm (A–D, G, H); 1 mm (E, F, I–L, N); 0.5 mm (M, O).

Head. Vertex slightly exposed, interocellar distance slightly wider than the distance between eyes, narrower than the distance between antennal sockets (Fig. 5F). Pronotum subelliptical, broader than long, anterior and posterior margins nearly straight, and postero-lateral margin blunt and round; disc with symmetrical spots and stripes (Fig. 5E). Fifth maxillary palpus expanded, third and fourth maxillary palpi nearly equal in length, both twice as long as fifth maxillary palpus (Fig. 5J). Tegmina and hind wings. Tegmina and hind wings fully developed, both extending beyond the end of abdomen (Fig. 5A, B). Tegmina with M and CuA radial, M straight with seven complete branches. Hind wings with ScP and RA expanded at apex, M simple, without branches; CuA with four complete branches (Fig. 5G, H). Legs. Anteroventral margin of front femur Type B₃ (Fig. 5K). Four proximal tarsomeres with pulvilli. The pretarsi with arolium, tarsal claws symmetrical and specialized, with minute denticles on ventral margins. Abdomen and genitalia. Eighth abdominal tergum specialized, with a tuft of setae near the distinctly concave middle posterior margin (Fig. 5I). Supra-anal plate symmetrical, anterior margin straight and truncate, posterior margin obtusely round. Paraprocts simple, similar and flaky. Cerci long, with setae on the ventral surface (Fig. 5L) Subgenital plate nearly symmetrical, anterior margin distinctly concave in the middle. Styli similar, slender; interstylar margin sinuate, left side with five or six small spines, right side with 5–7 small spines (Fig. 5N). Left phallomere small, irregular bone-shaped, with a small spine (Fig. 5M). Median phallomere slender rod-shaped, with a bristle brush at apex; accessory sclerite I arched, two ends enlarged, right end with a row of spines; accessory sclerite II with three lamellar structures with small spines; accessory sclerite III sickle-shaped (Fig. 5N). Hook phallomere on the right side, apex curved inwards with a short spine (Fig. 5O).

Female. Similar to the male but body and wings somewhat shorter (Fig. 5C, D).

This species is similar to M. flexa Wang et al., 2014 in general appearance, but can be differentiated from the latter by the following characters: 1) interstylar margin sinuate, left side with 4–6 small spines, right side with 4–7 small spines; while in the latter, interstylar margin strongly produced, whose lateral sides upturned and scattered with spines; 2) left phallomere irregular bone-shaped, without a small spine; while in the latter, left phallomere irregular bone-shaped, with two spines; 3) accessory sclerite II with three lamellar structures with small spines; while in the latter, accessory sclerite II with lamellar structure without small spines.

The specific name is derived from the Latin word undulatus, which refers to the sinuate interstylar margin.

China (Chongqing).

Holotype : China • ♂; Guizhou Province, Zunyi City, Suiyang County, Qingbantang Town, Baishao Ditch; 30 Jul. 2013; Xiu-Dan Wang leg; SWU-B-PS000034. Paratype: China • 1 ♂; same data as holotype; SWU-B-PS000035.

(mm). Male (n = 2), pronotum: length × width 2.3–2.5 × 3.0–3.4, tegmina length: 10.7–11.2, body length: 10.5–11.0, overall length: 12.9–13.2.

Male. Coloration. Body, head and face yellowish brown (Fig. 6A, B). Interocular space with a wide brown transverse band. Ocellar spots small, white, with brown spots beside them. Antennal base pale yellowish brown, other segments brown (Fig. 6D). Maxillary palpi dark brown (Fig. 6H). Pronotal disc yellowish brown with reddish tan spots and stripes, and two lateral borders pale linen-colored and transparent (Fig. 6C). Legs yellowish brown, with black spots at the base of the tibial spines. Tegmina pale yellowish brown, hind wings brownish grey (Fig. 6F, G). Abdomen yellowish brown, with black stripes along lateral margins of sterna and reaching the end of abdomen; both sides of each abdominal sternum with one small black spot on the inside of the longitudinal lines (Fig. 6B). Cerci yellowish brown (Fig. 6K). Styli pale yellow (Fig. 6M).

Figure 6. 

Margattea semicircularis Li & Che, sp. nov. A–N male A holotype, dorsal view B holotype, ventral view C pronotum, dorsal view D head, ventral view E eighth abdominal tergum, ventral view F tegmen, ventral view G hind wing, ventral view H maxillary palpi segments 3–5 I front femur, ventral view J Left phallomere, dorsal view K supra-anal plate, ventral view L median phallomere, ventral view M subgenital plate, ventral view N hook phallomere, ventral view. Scale bars: 5 mm (A, B, F, G); 1 mm (C–E, H, I, K–M); 0.5 mm (J, N).

Head. Vertex slightly exposed, interocellar distance wider than the distance between eyes, narrower than the distance between antennal sockets (Fig. 6C, D). Pronotum subelliptical, broader than long, anterior and posterior margins nearly straight, and postero-lateral angle blunt and round; disc with symmetrical spots and stripes (Fig. 6C). Maxillary palpi slender, fifth maxillary palpus expanded, third and fourth maxillary palpi nearly equal in length, both longer than fifth maxillary palpus (Fig. 6H). Tegmina and hind wings. Tegmina and hind wings fully developed, both extending beyond the end of abdomen (Fig. 6A, B). Tegmina with M and CuA radial, M straight with seven complete branches. Hind wings with ScP and RA expanded at apex; M simple, without branches; CuA with five complete branches (Fig. 6F, G). Legs. Anteroventral margin of front femur Type B₃ (Fig. 6I). Four proximal tarsomeres with pulvilli. The pretarsi with arolium, trsal claws symmetrical and specialized, with minute denticles on ventral margins.

Abdomen and genitalia. Eighth abdominal tergum specialized, with a heart-shaped transparent area and a tuft of bristles in the middle (Fig. 6E). Supra-anal plate symmetrical, anterior margin straight and truncate, the middle of posterior margin slightly produced. Paraprocts simple, similar, and flaky. Cerci long, setae on the ventral surface (Fig. 6K). Subgenital plate nearly symmetrical, anterior margin slightly concave in the middle. Styli similar, conical; interstylar margin strongly semicircular produced, both sides with spines (Fig. 6K). Left phallomere small, irregular, bone-shaped, with two small spines (Fig. 6J). Median phallomere curved hook rod-shaped, with a row of spines at apex. Accessory sclerite I arched, two ends enlarged, left end trigonate, right end with a row of spines; accessory sclerite II long transverse (Fig. 6L, M). Hook phallomere on the right side, apex curved inwards with a short spine (Fig. 6N).

This species is similar to M. spinifera Bey-Bienko, 1958a in general appearance, but can be differentiated from the latter by the following characters: 1) left phallomere with two small spines; while the latter, left phallomere with three spine-like processes; 2) accessory sclerite I arched, left end trigonate; while in the latter, accessory sclerite I arched, left end expanded with fuzz; 3) accessory sclerite II long transverse, and with two lamellar structures with a row of spines, while in the latter, without other accessory sclerites.

The scientific name is derived from the Latin word semicircularis, which indicates the interstylar margin has a semicircular protrusion.

China (Guizhou).

Holotype : China • ♂; Hainan Province, Qiongzhong Li and Miao Autonomous County, Mountain Limu; 600 m; 16 May. 2015; Xin-Ran Li, Zhi-Wei Qiu leg; SWU-B-PS000036. Paratypes: China • 2 ♂ & 1 ♀; same data as holotype; SWU-B-PS000037–000039 • 7 ♂ & 3 ♀; Hainan Province, Qiongzhong Li and Miao Autonomous County, Mountain Limu; 600 m; 16 May. 2015; Xin-Ran Li, Zhi-Wei Qiu leg; SWU-B-PS000040–000049.

(mm). Male (n = 7), pronotum length × width: 2.2–2.8 × 3.0–3.6, tegmina length: 11.3–12.2, body length: 10.1–11.1, overall length: 13.0–14.0. Female (n = 5), pronotum length × width: 2.2–2.8 × 3.0–3.6, tegmina length: 10.7–11.4, body length: 9.6–11.7, overall length: 12.9–13.7.

Male. Coloration. Body pale yellowish brown (Fig. 7A, B). Head yellowish brown. Face pale yellow. Interocular space with a brown transverse band. Ocellar spots yellowish white (Fig. 7F). Antennal base pale yellow, other segments yellowish brown. The third and fourth maxillary palpi yellowish brown, the fifth maxillary palpus brown (Fig. 7J). Pronotal disc pale yellowish brown with reddish tan spots but without stripes, and two lateral borders pale linen-colored and transparent (Fig. 7E). Legs faint yellow, with black spots at the base of the tibial spines. Tegmina yellowish brown, hind wings brownish grey (Fig. 7G, H). Abdomen pale yellow, with black stripes along lateral margins of sterna and reaching the end of abdomen. Cerci pale yellow to yellowish brown (Fig. 7L). Styli yellowish brown (Fig. 7N).

Figure 7. 

Margattea parabisignata Li & Che, sp. nov. A, B, E–O male C, D female A holotype, dorsal view B holotype, ventral view C paratype, dorsal view D paratype, ventral view E pronotum, dorsal view F head, ventral view G tegmen, ventral view H hind wing, ventral view I eighth abdominal tergum, ventral view J maxillary palpi segments 3–5 K front femur, ventral view L supra-anal plate, ventral view M left phallomere, ventral view N subgenital plate and median phallomere, ventral view O hook phallomere, ventral view. Scale bars: 5 mm (A–D, G, H); 1 mm (E, F, I–L, N); 0.5 mm (M, O).

Head. Vertex slightly exposed, interocellar distance wider than the distance between eyes, narrower than the distance between antennal sockets (Fig. 7F). Pronotum subelliptical, broader than long, anterior and posterior margins nearly straight, and postero-lateral angle blunt and round; disc with symmetrical spots but without stripes (Fig. 7E). Fifth maxillary palpus expanded, third and fourth maxillary palpi nearly equal in length, both longer than fifth maxillary palpus (Fig. 7J). Tegmina and hind wings. Tegmina and hind wings fully developed, both extending beyond the end of abdomen (Fig. 7A, B). Tegmina with M and CuA radial, M straight with seven complete branches. Hind wings with ScP and RA expanded at apex, M simple, without branches; CuA with four complete branches (Fig. 7G, H). Legs. Anteroventral margin of front femur Type B₂ (Fig. 7K). Four proximal tarsomeres with pulvilli. The pretarsi with arolium, tarsal claws symmetrical and slightly specialized, with minute denticles on ventral margins.

Abdomen and genitalia. Eighth abdominal tergum specialized, with a sparse tuft of bristles in the middle (Fig. 7I). Supra-anal plate symmetrical, the middle of anterior margin slightly concave, posterior margin arcuate produced with setae. Paraprocts simple, similar and flaky, obtuse at apex and each with a spiniform process at the base (Fig. 7L). Subgenital plate nearly symmetrical, anterior margin distinctly concave in the middle, left and right margins both produced in the middle, posterior margin truncate. Styli similar, conical. Cerci long, with setae on the ventral surface (Fig. 7N). Left phallomere large, irregular, bone-shaped, and with spines processes, apex curved upwards with rod-like structure (Fig. 7M). Median phallomere slender rod-shaped, apex with sparse brush-like structure composed of spines of various sizes; accessory sclerite I arched; accessory sclerite II with a lamellar structure with small spines (Fig. 7N). Hook phallomere on the right side, apex curved inwards with a short spine (Fig. 7O).

Female. Similar to the male (Fig. 7C, D).

This species is similar to M. bisignata Bey-Bienko, 1970 in general appearance, but can be differentiated from the latter by the following characters: 1) left phallomere with a short spiny process; the latter with a long spine process; 2) median phallomere apex with sparse brush-like structure composed of spines of varying sizes; while in the latter, median phallomere curved at apex, sheet-like, and with brush-shaped structure.

The species name parabisignata reflects its similarity to M. bisignata Bey-Bienko, 1970.

China (Hainan).

Holotype : China • ♂; Guangdong Province, Zhaoqing City, Fenghuang Town, Jiukeng River, Gold Ditch; 3 Jul. 2015; Zhi-Wei Qiu, Yong-Quan Zhao leg; SWU-B-PS000050. Paratypes: China • 6 ♂ & 1 ♀; same data as holotype; SWU-B-PS000051–000057 • 1 ♂; Guangdong Province, Zhaoqing City, Fenghuang Town, Jiukeng River, Lakeside Villa; 4 Jul. 2015; Zhi-Wei Qiu, Yong-Quan Zhao leg; SWU-B-PS000058.

(mm). Male (n = 4), pronotum length × width: 2.4–2.6 × 3.2–3.4, tegmina length: 10.5–11.5, body length: 10.4–10.8, overall length: 13.1–13.4. Female (n = 2), pronotum length × width: 2.5–2.7 × 3.4–3.6, tegmina length: 11.1–11.5, body length: 10.6–10.7, overall length: 13.4–13.7.

Male. Coloration. Body, head and face yellowish brown (Fig. 8A, B). Interocular space with a wider brown transverse band. Ocellar spots big and white (Fig. 8F). Antennal base pale yellow, other segments yellowish brown to brown. Maxillary palpi yellowish brown (Fig. 8J). Pronotal disc yellowish brown with dark brown spots and maculae, and two lateral borders pale linen-colored and transparent (Fig. 8E). Legs yellowish brown, with black spots at the base of the tibial spines. Tegmina pale yellowish brown, hind wings transparent, brownish grey (Fig. 8G, H). Abdomen yellowish brown, with black stripes along lateral margins of sterna and reaching the end of abdomen; both sides of each abdominal sternum with one small round black spot on the inside of the longitudinal lines. Cerci yellowish brown (Fig. 8L). Styli yellowish white (Fig. 8N).

Figure 8. 

Margattea forcipata Li & Che, sp. nov. A, B, E–O male C, D female A holotype, dorsal view B holotype, ventral view C paratype, dorsal view D paratype, ventral view E pronotum, dorsal view F head, ventral view G tegmen, ventral view H hind wing, ventral view I eighth abdominal tergum, ventral view J maxillary palpi segments 3–5 K front femur, ventral view L supra-anal plate, ventral view M left phallomere, ventral view N subgenital plate and median phallomere, ventral view O hook phallomere, ventral view. Scale bars: 5 mm (A–D, G, H); 1 mm (E, F, I–L, N); 0.5 mm (M, O).

Head. Vertex slightly exposed, interocellar distance wider than the distance between eyes, narrower than the distance between antennal sockets (Fig. 8F). Pronotum subelliptical, broader than long, anterior and posterior margins nearly straight, and postero-lateral angle blunt and round; disc with symmetrical irregular spots and maculae (Fig. 8E). Fifth maxillary palpus expanded, third and fourth maxillary palpi both longer than fifth maxillary palpus (Fig. 8J). Tegmina and hind wings. Tegmina and hind wings fully developed, both extending beyond the end of abdomen (Fig. 8A, B). Tegmina with M and CuA radial, M straight with six complete branches. Hind wings with ScP and RA expanded at apex, M simple, without branches; CuA with four complete branches (Fig. 8G, H). Legs. Anteroventral margin of front femur Type B₂ (Fig. 8K). Four proximal tarsomeres with pulvilli. The pretarsi with arolium, tarsal claws symmetrical and specialized, with minute denticles on ventral margins.

Abdomen and genitalia. Eighth abdominal tergum specialized, with a tuft of bristles in the middle (Fig. 8I). Supra-anal plate symmetrical, anterior margin straight and truncate, posterior margin produced, slightly concave in the middle. Paraprocts simple, similar, and flaky. Cerci long, setae on the ventral surface (Fig. 8L). Subgenital plate nearly symmetrical, anterior margin slightly concave in the middle. Styli similar, slender, conical; interstylar margin irregular produced (Fig. 8N). Left phallomere complex, irregular bone-shaped, with a long spine and three small spines (Fig. 8M). Median phallomere slender rod-shaped, with a forceps-shaped apex. Accessory sclerite I arched; accessory sclerite II with a transverse rod with denticulate (Fig. 8N). Hook phallomere on the right side, apex slightly curved inwards with a short spine (Fig. 8O).

Female. Similar to the male.

This species is similar to M. transversa He & Wang, 2021 in general appearance, but can be differentiated from the latter by the following characters: 1) left phallomere with a long spine; the latter with three long spine-like processes; 2) median phallomere with a forceps-shaped apex; while in the latter, median phallomere apex with a curved spine.

The specific name forcipatus, derived from Latin, refers to the median phallomere with a forceps-shaped apex.

China (Guangdong).