Apate dispar Fabricius, 1792, by monotypy.

Antennal club obliquely truncate, type 1 (except A. achaete Smith, Beaver & Cognato, 2020, which is type 2) (Hulcr et al 2007; Smith et al. 2020), club taller than wide (except A. achaete wider than tall); procoxae contiguous or narrowly separated; protibiae slender, obliquely or distinctly triangular, outer margin with 5−8 large socketed denticles on distal half, posterior face flat, unarmed, or with a few small granules; mesonotal mycangial tufts present (except A. achaete, A. carinensis (Eggers, 1923), A. paragogus Smith, Beaver & Cognato, 2020, and A. uniseriatus sp. nov.) along the pronotal base either as a small tuft the length of the scutellum and directly opposite it or extending laterally from the scutellum to striae 3 and with elytral base broadly, shallowly emarginated from the scutellum to striae 3; pronotum anterior margin with a row of serrations, pronotum lateral margins obliquely costate (Smith et al. 2020).

Some Anisandrus species have a median pair of pronotal serrations larger than the remaining serrations and superficially resemble Cnestus Sampson species. The genera are easily separated by the lateral margin of the pronotum which is costate in Anisandrus and carinate in Cnestus.

Holotype : female, Thailand, Chiangmai Province, Chom Thong District, Doi Inthanon National Park, 18°32'03.1"N, 98°29'55.2"E, 1680m, high montane forest, ethanol-baited traps, 01.vi.[20]20, W. Sittichaya (MSUC). Paratypes: same as holotype except: 18°35'10.5"N, 98°29'13.1"E, 2,550 m, 01.iv.[20]19, W. Sittichaya (2, WSTC; 1, THNHM); Chiang Mai, Doi Inthanon NP, Kaew Maepan Trail, 18°33.162'N, 98°28.810'E, 2250 m, Malaise trap, 10–17.xi.2006, Y. Areeluck (1, RABC); as previous except: 18°35.361'N, 98°29.157'E, summit forest, 2500 m, 9–16.viii.2006 (1, RABC); as previous except: 6–13.ix.2006 (1, QSBG).

Figure 1. 

Anisandrus montanus sp. nov. holotype female A dorsal view B postero-lateral view C lateral view D frons E declivital face.

A. eggersi (Beeson, 1930), A. improbus (Sampson, 1913).

2.80–2.92 mm long (mean 2.86, n = 4); 2.33–2.43× as long as wide. This species is similar to A. eggersi, but the body is smaller and more elongate, the declivity is less laterally broadened, and the posterolateral margin is not feebly costate. It is also similar to A. improbus but is distinguished by the following characteristics (A. improbus given first): sparse pubescence vs moderate pubescence, body 3.3–3.4 mm long vs 2.80–2.92 mm long, and body form more elongate, 2.43−2.54× as long as wide, vs stouter, 2.33–2.43× as long as wide.

Female. 2.80–2.92 mm long (mean 2.86, n = 4); 2.33–2.43× as long as wide. Body shiny and black, except appendages yellowish brown; body moderately densely covered with long, yellowish-brown, hair-like setae. Head: epistoma entire, transverse, with a row of hair-like setae; setae moderately long, sparse. Frons feebly convex to upper level of eyes, smooth, moderately shining, rather sparsely punctured, except close to epistoma; punctures bearing long, fine, hair-like setae. Frons with a weakly elevated, glabrous median ridge from epistoma to mid-point; above the eyes slightly domed, less shiny, coriaceous. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum triangular, large, slightly impressed. Antennal scape regularly thick, slightly longer than club (1.1:1). Pedicel as wide as scape, half as long as funicle. Funicle 4-segmented, segment 1 shorter than pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, encircling anterior face, with sharp marginal carina; segment 2 narrow, concave, corneous on anterior face; sutures absent on posterior face. Pronotum: 0.90× as long as wide, in dorsal view rounded, type 1, sides convex, rounded anteriorly; anterior margin with a row of six small serrations; serrations not larger than asperities behind. In lateral view short and tall, type 3, disc as long as anterior slope, summit at midpoint; summit from lateral view weakly raised, disc flat. Anterior slope with moderately spaced, medium-sized, coarse asperities, becoming lower and more strongly transverse towards summit. Disc alutaceous, subshining, sparsely granulate-punctate, with a semi-recumbent, moderately long, fine, forwardly directed, hair-like seta arising from just anterior to each granule. Lateral margins obliquely costate; costa long, slightly elevated. Base slightly, broadly concave; posterior angles angulate. Mycangial tuft present along basal margin; tuft moderately setose, approximately equal to width of scutellum. Elytra: 1.44× as long as wide, 1.63× as long as pronotum. Scutellum moderate in size, flat. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 5/8, then broadly rounded to apex; surface shiny. Disc shiny, moderately convex, without transverse saddle-like depression; striae with broad, shallow punctures separated by 1/2 diameter of a puncture, setose, setae slightly longer than two diameters of a puncture, semi-recumbent, hair-like; interstriae flat, 2–3× as wide as striae, punctate; punctures uniseriate, minute, setose; setae long, erect, hair-like, becoming longer posteriorly; interstriae 2 weakly raised near declivital summit, so that first striae and interstriae appear shallowly sulcate. Declivity occupying approximately 1/3 of elytra, evenly rounded, declivital face narrow, opalescent, weakly bisulcate, moderately impressed between interstriae 1 and 3 in upper part, interstriae 3 weakly inflated near summit, flat below. Declivital striae weakly impressed, strial punctures moderately larger and deeper than those of disc, with setae as described for disc; interstriae impunctate, sparsely minutely granulate; setae 2–3× width of interstriae 2, erect, hair-like; interstriae 2 either as wide as or narrower than interstriae 3 at midpoint of declivity. Declivital summit armed with a small, sharp, backwardly pointed spine on interstriae 2 and 3; spine on interstriae 2 stronger. Posterolateral margin costate to interstriae 5. Legs: procoxae contiguous. Protibiae obliquely triangular, broadest at apical 1/3; posterior face of protibiae punctate, with some punctures near base and inner margin with small, sparse granules; apical ½ of outer margin with six large, socketed denticles, their length longer than basal width. Meso- and metatibiae flattened; outer margins evenly rounded with nine and 10 large socketed denticles, respectively.

Male. Unknown.

Latin adjective montanus, found on mountains. The species is known only from Doi Inthanon, the highest mountain in Thailand, at 1680‒2550 m.

Thailand (Chiangmai Province).

This species prefers montane forest.

Three of the paratypes listed above were previously reported as A. apicalis by Beaver et al. (2014).

Holotype : female, Thailand, Phetchaburi Province, Kaeng Krachan District, Kaeng Krachan National Park, 12°49'43.6"N, 99°21'45.2"E, 900 m, low montane forest, ex Lithocarpus sp., 04.x.22, W. Sittichaya (MSUC). Paratypes: Kanchanaburi Province, Thong Pha Phum District, Thong Pha Phum National Park, 14°41'40.6"N, 98°23'51.9"E, 940 m, low montane forest, ethanol-baited trap, 11.xii.22, W. Sittichaya (1, WSTC; 1, THNHM; 1, RABC).

Figure 2. 

Anisandrus phithakpa sp. nov. holotype female A dorsal view B postero-lateral view C lateral view D frons E antenna.

A. apicalis, A. congruens Smith, Beaver & Cognato, 2020, A. cristatus (Hagedorn, 1908).

2.88–3.02 mm long (mean 2.96 mm, n = 4); 2.06–2.17× as long as wide. Stout species; elytral disc saddle-like, bearing a pair of small spines on interstriae 2. Declivity broad; declivital face with striae 1 and 2 weakly impressed; interstriae 1 and 3 slightly elevated; posterolateral margin rounded, costate only near apex. The species is similar to A. apicalis, A. congruens, and A. cristatus. Anisandrus congruens and A. cristatus are distinguished by the presence on declivital interstriae 3 of a row of 5‒7 regularly spaced, backwardly directed, sharply pointed spines; in A. phithakpa and A. apicalis these are reduced to small granules. Anisandrus phithakpa is distinguished from A. apicalis by the following characters (A. phithakpa given first): declivital interstriae with a pair of minute, pointed granules on interstriae 1, and an equally sized pair on interstriae 2 and 3 vs no granules on interstriae 1, and interstriae 2 with a pair of small, backwardly directed, sharply pointed teeth which are considerably larger than the pointed granules on interstriae 3.

Female. 2.88–3.02 mm long (mean 2.96 mm; n = 4); 2.06–2.17× as long as wide (mean 2.12× as long as wide; n = 4). Body colors: dark brown to black, appendages paler brown. Head: epistoma entire, transverse, with a row of hair-like, moderately long, sparse setae. Frons weakly convex to upper level of eyes, reticulate, subshining, with sparse widely separated, small, shallow, setose punctures each with a long, erect hair-like seta. Medial area inconspicuous, feebly convex (flat in one paratype), glabrous. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum moderate, distinctly triangular, slightly impressed. Antennal scape regularly thick, 1.2× as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented; segment 1 shorter than pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, encircling anterior face; segment 2 narrow, concave, corneous; sutures absent on posterior face. Pronotum: 0.98× as long as wide, in dorsal view rounded, type 1; sides convex, rounded anteriorly; anterior margin with a row of eight small serrations of same size as asperities above; in lateral view, short and tall, type 3; disc as long as anterior slope, summit at midpoint. Anterior slope with densely spaced, very large, coarse asperities, becoming lower and more strongly transverse towards summit. Disc alutaceous, subshiny with moderately dense, large, shallow punctures; punctures with moderate, semi-recumbent, hair-like setae; some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse with posterior angles rounded. Mycangial tuft present along basal margin; tuft moderately setose, approximately the width of scutellum. Elytra: 1.5× as long as wide, 1.5× as long as pronotum. Scutellum broad, large, linguiform, flush with elytra, flat, shiny. Elytral base transverse; edge oblique; humeral angles rounded; elytra parallel-sided in basal ½, then broadly rounded to apex; surface shiny. Disc shiny, with a distinct medial, transverse, saddle-like depression; depressed areas opalescent; striae not impressed; with broad shallow punctures separated by areas less than a diameter of a puncture, setose; setae 2–3× as long as a puncture, recumbent, hair-like; interstriae flat, punctate, with 2 or 3 confused lines of minute punctures, setose; setae long, 1–1.5× width of interstriae 2, erect, hair-like, unarmed by granules. Declivity occupying approximately 1/3 elytra; apex evenly rounded; declivital summit with a pair of minute, pointed granules on interstriae 1, and a slightly larger pair on both interstriae 2 and 3 placed progressively further towards apex; declivital face feebly bisulcate; striae 1 and 2 impressed; interstriae 3 inflated and armed, with 2 or 3 minute granules; strial punctures of similar size and depth to those of disc, bearing setae as described for disc; interstriae impunctate, sparsely, minutely granulate; setae 2× width of interstriae 2, erect, hair-like; interstriae 2 as wide as or narrower than interstriae 3 at midpoint of declivity. Posterolateral margin costate to interstriae 5. Legs: procoxae contiguous; prosternal coxal piece short, inconspicuous. Protibiae obliquely triangular, broadest at apical 1/3; posterior face inflated, punctate, punctures minute; apical 1/2 of outer margin with six socketed denticles, their length 2× their basal as basal width. Meso- and metatibiae flattened; outer margins evenly rounded with seven and eight long slender socketed denticles, respectively.

Male. Unknown.

Thai (พิทักษ์ป่า) “Phithakpa”, forest rangers. The species name indicates our deep appreciation for Thai forest rangers for their hard and selfless work to protect conservation areas in Thailand. Noun in apposition.

Thailand (Kanchanaburi and Phetchaburi provinces).

Recorded from Lithocarpus sp. (Fagaceae).

The paler body colors (brown) of the holotype and some paratypes suggest that they are teneral. One paratype has a consistently dark-brown body with pale appendages.

Holotype : female, Thailand, Phetchaburi Province, Kaeng Krachan District, Kaeng Krachan National Park, 12°49'43.6"N, 99°21'45.2"E, 900 m, low montane forest, ex. Lithocarpus sp., 04.x.22, W. Sittichaya (MSUC). Paratype: Kanchanaburi Province, Thong Pha Phum District, Thong Pha Phum National Park, 14°41'40.6"N, 98°23'51.9"E, 940 m, low montane forest, ethanol-baited trap, 11.xii.22, W. Sittichaya (1 WSTC).

Figure 3. 

Anisandrus tanaosi sp. nov. holotype female A dorsal view B postero-lateral view C lateral view D frons E antenna.

A. auco Smith, Beaver & Cognato, 2020, A. cryphaloides Smith, Beaver & Cognato, 2020, A. triton sp. nov.

2.31–2.70 mm long (mean 2.5; n = 2); 2.20–2.25× as long as wide (mean = 2.23; n = 2). Small and stout species. Pronotal anterior margin slightly angularly projecting; elytral disc convex, without a saddle-like impression, declivital summit armed with a pair of minute, spinulose granules on interstriae 2, declivital face feebly convex, unarmed. Posterolateral margin rounded, costate only near apex. The species is similar to A. auco and A. cryphaloides. It can be distinguished from A. auco by the following characters (A. tanaosi given first): smaller size (2.3‒2.7 mm long vs 2.9‒3.0 mm); anterior margin of pronotum with seven moderately sized asperities vs four large coarse asperities; more steeply sloping elytral declivity occupying 3/8 of elytral length vs more gently sloping occupying 3/5 of length; dark-brown to black body vs light brown to reddish brown. It can be distinguished from A. cryphaloides by the following characters (A. tanaosi given first): pronotum in dorsal view type 0, anterior margin only slightly projecting, with asperities of equal size vs pronotum more strongly conical, type 6, median pair of asperities larger than lateral pairs.

Female. 2.31–2.70 mm long (mean 2.5; n = 2); 2.20–2.25× as long as wide (mean 2.23; n = 2). Body dark brown to black, head and prothorax dark brown, elytra and venter black, appendages yellowish brown. Antennae and legs light brown. Body densely covered with greyish-brown setae. Head: epistoma entire, transverse, with a row of short hair-like setae, setae sparse. Frons feebly convex to upper level of eyes, weakly reticulate, rugulose-punctate, some rugulosities forming longitudinal lines; each puncture with a moderately long, fine, hair-like setae; a weak, impunctate median ridge extends to upper level of eyes. Eyes feebly emarginate just above antennal insertion, upper part slightly smaller than lower part. Submentum triangular, small, slightly impressed. Antennal scape regularly thick, short, as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 as long as pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, encircling anterior face; segment 2 concave, soft and narrow; sutures absent on posterior face. Pronotum: 0.90× as long as wide. In dorsal view, type 0, feebly conical anteriorly, sides convex; anterior margin with a row of seven small, slightly protruding serrations, equal in size to those on anterior slope. In lateral view type 3, short and tall; disc as long as anterior slope, summit at midpoint. Anterior slope with moderately densely spaced, large coarse asperities, becoming lower and more strongly transverse towards summit. Disc alutaceous, subshining with moderately dense, minute granulate punctures, each bearing a short, semi-recumbent, hair-like seta, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles angularly rounded. Mycangial tuft present along basal margin, tuft moderately setose, approximately the width of scutellum. Elytra: 1.22× as long as wide, 1.42× as long as pronotum. Scutellum broad, large, linguiform, flush with elytra, flat, shiny. Elytral base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 2/3, then broadly rounded to apex. Disc subshiny, broadly convex; striae not impressed, with small, shallow, setose punctures separated by 1.5–2× diameters of a puncture, setae 3× as long as diameter of punctures, recumbent, hair-like; interstriae flat; punctures strongly confused, without granules, setose; setae short, as long as strial setae, erect hair-like. Declivity occupying approximately 3/8 elytra; summit with a pair of spinulose granules on interstriae 2; declivital face feebly convex above, flattened below from interstriae 1–3; striae weakly impressed; strial punctures somewhat larger and deeper than those of disc; interstriae sparsely uniseriate punctate, setae 2× width of an interstria, erect, hair-like. Posterolateral margin rounded, unarmed by granules, costate only close to apex. Legs: procoxae contiguous; prosternal coxal piece short, inconspicuous. Protibiae obliquely triangular, broadest at apical 1/3; posterior face minutely granulate; apical 1/3 of outer margin with six small, socketed denticles, their length as long as basal width. Meso- and metatibiae flattened; outer margins evenly rounded with eight large, socketed denticles.

Male. Unknown.

Tanaosi (ตะนาวศรี), Thai name of the Tenasserim mountain range, in reference to the collection locality of the holotype. Noun in apposition.

Thailand (Kanchanaburi and Phetchaburi provinces).

Unknown.

Holotype : female, Thailand, Kanchanaburi Province, Thong Pha Phum District, Thong Pha Phum National Park, 14°41'40.6"N, 98°23'51.9"E, 940m, low montane forest, ethanol-baited trap, 11.xii.22, W. Sittichaya (MSUC).

Figure 4. 

Anisandrus triton sp. nov. holotype female A dorsal view B postero-lateral view C lateral view D frons E antenna.

Anisandrus auco, A. cryphaloides.

4.22 mm long; 1.94× as long as wide. Large, broad, stout species. Elytral disc with a saddle-like, weak impression on middle of disc; declivity longer than disc; interstriae 2 armed with two pairs of spines, backwardly incurved on declivital summit and one additional smaller sized spine on upper portion of declivital face; interstriae 3 armed with a row of 3–5 unequally sized spines and granules; declivital face feebly convex, apex broadly rounded; posterolateral margin rounded, with a short costa near apex. The species is similar to Anisandrus auco, A. cryphaloides, and A. tanaosi. It can be distinguished from them by the following characters (A. triton given first): greater size (4.2 mm long vs 2.1‒3.0 mm) and stouter body (1.94× longer than wide vs 2.2‒2.4×; elytral disc with a weak, saddle-like depression vs elytral disc flat; upper margin of the elytral declivity with a pair of backwardly directed, sharply pointed spines vs a pair of minute, pointed granules.

Female. 4.22 mm long (n = 1); 1.94× as long as wide. Body black except appendages brown; body densely covered with long, erect, yellowish-brown, hair-like setae. Head: epistoma entire, transverse, with a row of short and sparse, hair-like setae, sparser in the middle and on lateral margins below eyes. Frons with a weak median ridge extending to upper margin of eyes, weakly impressed on each side near epistoma, becoming flattened and weakly convex above, reticulate, subshining, with sparse, large, shallow, punctures, each puncture bearing a shorter, finer, erect hair-like seta than those on epistoma; punctures becoming smaller and shallower towards vertex. Eyes large, feebly emarginate just above antennal insertion; upper part of eyes much smaller than lower part. Submentum transversely long, narrowly triangular, slightly impressed. Antennal scape slender, 1.4× as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 as long as pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, encircling anterior face; segment 2 narrow, corneous on anterior face only; sutures absent on posterior face. Pronotum: 0.83× as long as wide. In dorsal view, between type 0 and type 6, sides convex, strongly narrowed anteriorly; anterior margin with a row of seven medium-sized serrations. In lateral view, short and tall, type 3; disc slightly shorter than anterior slope. Anterior slope with moderately dense, large, coarse asperities, becoming lower and more strongly transverse towards summit. Disc convex, moderately shiny with moderately dense, minute, punctures bearing two types of setae: moderately long, erect, hair-like setae and short, semi-recumbent, hair-like setae; some longer, hair-like setae at margins. Base transverse; posterior angles broadly rounded. Mycangial tuft present along basal margin; tuft dense, long, setose, approximately 2× width of scutellum. Elytra: 1.18× as long as wide, 1.63× as long as pronotum. Scutellum small, broad, linguiform, shiny, slightly convex, flush with elytra. Elytral base transverse, edge oblique, humeral angles rounded, parallel-sided in basal ½, then broadly rounded to apex; surface shining. Disc shallowly, transversely impressed; striae not impressed, with medium-sized, shallow punctures separated by the diameter of a puncture; strial setae 1.5× as long as punctures, semi-recumbent, hair-like; discal interstriae 1 and 3 flat, interstriae 4 and 5 weakly convex; near upper margin of declivity, interstriae 1–5 weakly convex; interstriae biseriate punctate, punctures minute, shallow, each bearing an erect hair-like seta; setae on disc as long as interstrial width, some longer setae present on lateral and apical margins of elytra; punctures on lateral margins and near declivital summit replaced by small granules. Declivity occupying approximately 1/2 elytra, evenly rounded, declivital face convex; striae feebly impressed, strial punctures the same size and depth as those of disc; interstriae feebly convex, 1–3 of equal width, biseriate granulate; setae 2× width of an interstria, erect, hair-like; declivital interstria 2 armed with two pairs of spines; the larger on declivital summit backwardly hooked, the smaller on upper portion of declivital face pointed; interstria 3 armed by a row of 5 spinulose granules, the upper two pairs slightly backwardly hooked. Posterolateral margin of declivity rounded, with a short costa near apex, unarmed by granules. Legs: procoxae slightly separated; prosternal coxal piece short, inconspicuous. Protibiae obliquely triangular, broadest at apical 1/3; posterior face inflated, unarmed; apical 1/2 of outer margin with six moderately sized socketed denticles, length approximately equal to basal width. Meso- and metatibiae flattened; outer margins evenly rounded each with eight moderately sized socketed denticles.

Male. Unknown.

Ancient Greek, triton is a fish-tailed sea-god, named after a veteran vehicle used in beetle surveys by the senior author.

Thailand (Kanchanaburi Province).

Unknown.

Holotype , female, Thailand, Nan Province, Pua District, Doi Phu Kha National Park, 19°10'27.4"N, 101°06'19.7"E, 1660 m, montane forest, ethanol-baited traps, 01.viii.19 W. Sittichaya (MSUC). Paratypes: same as holotype except: 30.vi.19 (2) (1, NHMW; 1, RABC); same as holotype except: 10.x.19 (2) (1, THNHM; 1, WSTC).

Figure 5. 

Anisandrus uniseriatus sp. nov. holotype female A dorsal view B lateral view C postero-lateral view D frons E venter and antennae F declivital face.

Anisandrus carinensis.

4.0–4.32 mm long (mean 4.20 mm; n = 5); 2.32–2.60× as long as wide (mean 2.44 mm; n = 5). Species large, robust; body yellowish brown, covered with long erect hair-like setae; pronotal anterior margin broadly rounded and armed with a row of serrations; mesonotal mycangial tuft absent; antennal club type 1 with segment 1 encircling anterior face; elytral disc with a weak transverse saddle-like depression; interstriae uniseriate granulate; declivital interstriae 1 and 3 armed by four or five unequally sized tubercles; elytral apex angularly rounded.

This species is closely related to A. carinensis but is distinguished by the following characteristics (A. uniseriatus given first): discal interstria uniseriate granulate-punctate vs biseriate granulate-punctate; angularly rounded apex vs broadly rounded elytral apex.

Female. 4.0–4.32 mm long (mean 4.20 mm; n = 5); 2.32–2.60× as long as wide (mean 2.44 mm; n = 5). Body yellowish brown to reddish brown, declivity darker. Head, legs, and antennae light brown. Head: epistoma entire, transverse, with a row of golden, long and thick hair-like setae. Frons flat to upper level of eyes, shining, with sparse small granules, each with a fine, long, erect hair-like seta; median line narrowly elevated to upper margin of eyes, glabrous strongly shining. Vertex convex, reticulate, with longitudinal shallow rugae and punctures. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum large, distinctly triangular, slightly impressed. Antennal scape regularly thick, as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 shorter than pedicel. Club wider than long, obliquely truncate, type 1; segment 1 corneous, transverse, occupying basal 2/5, encircling anterior face; segment 2 narrow, concave, corneous; sutures absent on posterior face. Pronotum: 0.89× as long as wide. In dorsal view basic, type 2, sides parallel in basal 1/2, rounded anteriorly; anterior margin without distinct serrations. In lateral view basic, type 0, disc as long as anterior slope, summit at apical 2/5. Anterior slope with densely placed asperities of very variable size, becoming lower and more strongly transverse towards summit. Disc shining, with densely placed, minute asperities and granules, arranged approximately concentrically behind summit; vestiture of long, erect hair-like setae interspersed with shorter, more abundant, semi-recumbent setae directed antero-medially; some longer hair-like setae on convex lateral margins. Base transverse, posterior angles rounded. Mycangial tuft absent. Elytra: 1.55× as long as wide, 1.75× as long as pronotum. Scutellum narrow, moderately sized, linguiform, flush with elytra, flat, shiny. Elytral bases transverse, edge oblique, humeral angles rounded; elytra parallel-sided in basal 2/3, then broadly rounded to apex; surface shiny. Disc with a very slight, transverse, saddle-like depression in middle; only striae 1 impressed, its punctures coarse, shallow, regularly placed, separated by about the diameter of a puncture, and bearing fine, moderately long, hair-like setae; interstriae finely punctate, with some punctures (approximately every second puncture) granulate; interstria 1 uniseriately punctured, but with 2‒3 rows of punctures where widened posteriorly close to declivity; interstria 2 uniseriately punctate along its length; interstriae 3‒5 biseriately punctate close to base, uniseriate posteriorly; granulate punctures on interstriae bearing long, fine, erect hairlike setae, non-granulate punctures with shorter, semi-recumbent setae. Declivity occupying approximately 1/3 of elytra, steeply sloping; declivital face weakly bisulcate between raised interstriae 1 and 3; strial punctures larger and deeper than those of disc; interstriae uniseriate with some punctures replaced by granules or tubercles; interstriae 1 widest at mid-declivity, with 4‒6 larger pointed tubercles, and some smaller granules; interstriae 2 with a few small granules at top of declivity only, narrowed towards apex; interstriae 3 with 5‒7 pointed tubercles, a little smaller than those on interstriae 1; interstrial tubercles and granules bearing long, erect setae 1.5× width of interstriae 2; interstrial punctures with finer, shorter setae. Posterolateral margin of elytra rounded, costate only near apex, unarmed by granules. Legs: procoxae contiguous, prosternal coxal piece tall and pointed. Protibiae distinctly triangular, broadest at apical 4/5, posterior face smooth; apical 1/2 of outer margin with five moderately sized socketed denticles, their length slightly longer than basal width. Meso- and metatibiae flattened, obliquely triangular, their apical 1/2 with 5- or 6-socketed denticles on outer margin.

Male. Unknown.

Latin uniseriatus: uni- meaning one; series meaning row, in reference to a single row of interstrial setae.

Thailand (Nan Province).

Unknown.

3.70–4.26 mm long (mean 4.08; n = 7); 2.27–2.35× as long as wide (mean 2.31×; n = 7). Moderate to large in size, robust form; body yellowish brown, covered with long erect hair-like setae; pronotal anterior margin broadly rounded and armed with a row of serrations on anterior margin of pronotum; mesonotal mycangial tuft absent; antennal club type 1 with segment 1 encircling anterior face; elytral disc with a weak transverse saddle-like depression; interstriae biseriate granulate; declivital interstriae 1 and 3 armed by four or five unequally sized tubercles; elytral apex broadly rounded.

Figure 6. 

Anisandrus carinensis female. A dorsal view B lateral view.

S: Ranong Province, Suk Samran District, Klong Naka Wildlife Sanctuary, 9°27'42.8"N, 98°32'23.3"E, 320 m, tropical lowland forest, ethanol-baited traps, 01.iv.14 (4); Suratthani Province, Ban Ta Khun District, Khao Sok National Park, 8°55'25.6"N, 98°31'19.2"E, 380 m, tropical rain forest, ethanol-baited trap, 01.v.14 (10). W: Kanchanaburi Province, Thong Pha Phum District, Thong Pha Phum National Park, 14°41'40.6"N, 98°23'51.9"E, 940 m, montane forest, ethanol baited trap, 10.ix.22 (3); all W. Sittichaya.

Myanmar (Smith et al. 2020).

Seven specimens were recorded from Chiang Mai province under this name by Beaver et al. (2014). Three of these specimens are now included in the closely similar species, A. montanus sp. nov. (see above). The other four specimens belong to A. cristatus, which was considered a synonym of A. apicalis at the time of Beaver et al.’s publication. It was reinstated as a distinct species by Smith et al. (2020). We know of no specimens of A. apicalis from Thailand, and the species must be removed from the faunal list.