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Research Article
The beetle fauna (Insecta, Coleoptera) of the Rawdhat Khorim National Park, Central Saudi Arabia
expand article infoMahmoud S. Abdel-Dayem, Hassan H. Fad§, Ashraf M. El-Torkey|, Ali A. Elgharbawy, Yousif N. Aldryhim, Boris C. Kondratieff#, Amin N. Al Ansi, Hathal M. Aldhafer
‡ King Saud University, Riyadh, Saudi Arabia
§ Ain Shams University, Cairo, Egypt
| Plant Protection Research Institute, Giza, Egypt
¶ Al Azhar University, Nasr City, Egypt
# Colorado State University, Fort Collins, United States of America
Open Access

Abstract

This study was conducted as a part of a comprehensive baseline survey of insect biodiversity of Rawdhat Khorim National Park (RKNP), Central Kingdom of Saudi Arabia (KSA). During this study a total of 262 Coleoptera species belong to 182 genera in 35 families were identified, of which 247 are named at a species level. Fifteen species (6.0%) are apparently endemic to KSA. Thirty-eight species are new to the known beetle fauna of KSA, including 25 species reported from the Arabian Peninsula for the first time. The families Tenebrionidae (45 species), Scarabaeidae (34 species), and Carabidae (27 species) were the most species rich families. About 37% of the beetle abundance was represented by species of Scarabaeidae, especially Aphodius ictericus ghardimaouensis Balthasar. Karumia inaequalis Pic (Dascillidae) was also an abundant species. Approximately 43.5% of beetle species collected during this study are considered very rare taxa in RKNP. The RKNP beetle fauna shows more affinity to Sahro-Arabian (36.4%), Afrotropical-Sahro-Arabian (17.4%) and Palaearctic-Sahro-Arabian (10.5%). Twenty-three species (9.3%) are considered cosmopolitan or subcosmopolitan. The data on month of collection, method of collection, and abundance status within RKNP, together with the distribution within KSA and the general distribution (zoogeography) of each species are presented.

Keywords

Arabian fauna, distribution, endemic species, new records, zoogeography

Introduction

Beetles (Coleoptera) are considered the most taxonomically diverse insect group that comprises major components of ecosystems in terms of biomass, species richness and ecological roles (Stack 2015). About 400,000 species have been described (Hammond 1992), comprising about 25% of the Earth’s total animal diversity (Rosenzweig 1995; Hunt et al. 2007). Beetles play important roles in pollination, herbivory, granivory, predator-prey interactions, decomposition and nutrient cycling, and soil disturbances (Huffaker and Gutierrez 1999).

The foundation of our knowledge of the beetles of the Kingdom of Saudi Arabia (KSA) was presented by various authors in the series “Fauna of Saudi Arabia,” currently published as part of the “Fauna of Arabia” (Wittmer and Buttiker 1979- to date). Additionally, sixteen years ago, Al-Ahmadi and Salem (1999) listed 1,343 beetle species from KSA. Additional data on the beetles in KSA are available in the Catalogue of Palaearctic Coleoptera (Löbl and Smetana 2003–2007).

Rawdhats, naturally occurring moist basins, are one of the major components that considered as biological cores in the hyper-arid ecosystems of Central KSA (Tag El-Din et al. 1994; Al-Farraj et al. 1997; Alfarhan 2001; Al-Qarawi 2011). The flora and fauna of these unique habitats are under constant threat due to increased anthropogenic activities (Al-Nafie 2007; Al-Qarawi 2011). However, relatively little is known about their insect fauna (e.g., Al Dhafer and Platia 2013, Al Dhafer et al. 2016; Sharaf et al. 2013; Alqarni et al. 2015; Abdel-Dayem et al. 2015, 2016) that contribute to the faunal biodiversity of KSA.

Rawdhat Khorim National Park (RKNP) in Riyadh Province is important element in the conservation network of KSA and its landscape is characterized by relative diverse pristine native plant communities. (Tag El-Din et al. 1994; Al-Farraj et al. 1997; Alfarhan 2001; Al-Qarawi 2011). Recent studies of the insects occurring in RKNP have revealed several new taxa, Coleoptera: Dicronychus latifahae (Elateridae) (Al Dhafer and Platia 2013); Reichardtiolus aldhaferi (Histeridae) (Lackner 2014); Boromorphus saudicus (Tenebrionidae) (Schawaller et al. 2013); and Hymenoptera: Tetramorium saudicum (Formicidae) (Sharaf et al. 2013).

The objective of the current study was to provide a comprehensive list of the beetles from one of the more unique habitats in the world, specifically RKNP contributing to the knowledge of the beetle fauna of KSA and the Arabian Peninsula in general.

Materials and methods

Study area

The Rawdhat Khorim National Park is situated in the northeastern Riyadh Province of Central KSA (Fig. 1a). This is part of the Najd Plateau, a sedimentary rectangular plateau of Saudi Arabia. It covers an area of 24 km2 and located about 95 km northeast of the capital of Riyadh (25°23’N, 47°17’E, 560 m.a.s.l.). It is a densely vegetated alluvial basin (Vesey-Fizberald 1957) (Figure 1b) supporting a characteristic floral community in the hyper-arid desert of central Saudi Arabia (Al-Farraj et al. 1997; Alfarhan 2001). This area has low rainfall, but drainage from surrounding foothills of the adjacent highlands provides additional moisture forming temporary water pools and high plant diversity occurs during spring (Al-Farraj et al. 1997).

Figure 1. 

a Map showing the provinces and geographic location of Rawdhat Khorim National Park within the Central Region of Kingdom of Saudi Arabia b Photo of Rawdhat Khorim National Park (Produced by ArcMap 10.4).

Climate

The climate of RKNP is characterized by a hot summer and a mild winter with an average relative humidity of 33%. The average annual temperature is 26°C, June to August is the hottest period of the year (35-37.4 °C) and December to February is the coldest (14.6-15.7 °C). The average annual precipitation is 122 mm, the highest amount of precipitation occurs during March and April (30 and 27 mm, respectively), while no rainfall occurs during June to September (Sharaf et al. 2013).

Flora

The flora of RKNP includes a total of 153 plant species belonging to 32 families (Alfarhan 2001). The vegetation is complex consisting of perennial and annual herbs, shrubs and trees (Al-Farraj et al. 1997). Matricaria aurea (Loefl.) Sch.-Bip. (Compositae)), Plantago boissieri Hausskn. & Bornm., P. ciliata Desf. (Plantaginaceae), and Phalaris minor Retz. (Poaceae) are the dominant annual herbs (Al-Farraj et al. 1997; Alfarhan 2001). Acacia ehrenbergiana Heyne, A. gerrardii Benth. (Fabaceae), Calotropis procera (Aiton) W.T. Aiton (Apocynaceae), Lycium shawii Roem. & Schult. (Solanaceae), Rhazya stricta Decne. (Apocynaceae) and Ziziphus nummularia (Burm. f) Wight & Arn. (Rhamnaceae) are the dominated shrub and trees (Al-Farraj et al. 1997). Plant growth is higher during spring than in other seasons (Al-Farraj et al. 1997).

Beetle collection

The species listed here are based on the survey that was conducted biweekly from October, 2011 to September, 2013 in the RKNP. The material was collected using a variety of collecting methods including pitfall trapping (PT), UV-light trapping (LT), Malaise trapping (MT), net sweeping (SW), beating vegetation (BV), vacuuming (VC), and hand-picking (HP). All collected beetles were sorted, identified and deposited in the King Saud University Museum of Arthropods (KSMA), College of Food and Agricultural Sciences, King Saud University, KSA.

Species format and arrangement

Species identification is based on specialists “see Acknowledgments” and numerous publications not included here. Beetles were not being identified to the species level were included in the study if specimens were morphologically different from other (congeneric) species. The identified species are arranged systematically to subfamily level and alphabetically thereafter. The classification and nomenclature of subfamilies and higher levels follows Bouchard et al. (2011). The valid name followed by the author and date of publication for species were given and these were not cited in the reference section and they can be found in the Catalogue of Palaearctic Coleoptera (Löbl and Smetana 2003–2007). The world distribution for each species is indicated as two capital letters for each country (according ISO 3166: http://www.iso.org/iso/country_codes). The world distribution is based mainly on Catalogue of Palaearctic Coleoptera (Löbl and Smetana 2003–2007), Global Biodiversity Information Facility (GBIF: http://www.gbif.org), and Beetles and Rock Art in Libya (http://jcringenbach.free.fr). The general distribution (zoogeography) of each species is also indicated as a letter code (see “Abbreviations”) corresponding to main zoogeographic regions of the world proposed by Holt et al. (2013) (Fig. 2). The KSA distribution of each species is indicated (abbreviated as two letters for the Saudi provinces) (Fig. 1a). The KSA distributional records are based mainly on published records in the series “Fauna of Saudi Arabia”, being published as part of the “Fauna of Arabia (Wittmer and Buttiker 1979-to date) and other available papers on Saudi beetles. The absence of a geographic entry after a species name indicates that the species was recorded from Arabia or KSA but no locality was specified. For each species the following information is given: local abundance (all values of collected specimens were log transformed and then they classified into five categories using equal interval classification: very rare: <5 individuals, rare: 5-17, frequent: 18-70, common: 71-300, and abundant: > 300); collecting method (see abbreviation in “Beetle collection” above); and months of collection (Roman numerals) within RKNP.

Figure 2. 

The general distributions frequency of the beetle fauna of Rawdhat Khorim National Park, Central Kingdom of Saudi Arabia.

Abbreviations for the zoogeographical regions

AFR Afrotropical

AUS Australian

COS Cosmopolitan

MAD Madagascan

NAR Nearctic

NTR Neotropical

OCN Oceanic

ORR Oriental

PAL Palaearctic

SAR Saharo_Arabian

SCO Subcosmopolitan

SJP Sino_Japanese

Results and discussion

This study represents the first inventory of beetles in RKNP, within arid region of central KSA. In total, 15,658 specimens were collected and identified to 262 species belonging to 182 genera included in 83 subfamilies and 35 families. Forty percent of known faunal diversity is accounted for by three beetle families, Tenebrionidae (45 spp.), Scarabaeidae (34 spp.) and Carabidae (27 spp.) (Table 1). The families with the greatest abundant were Scarabaeidae (36.6%) followed by Tenebrionidae (16.7%) (Table 1). Thirty-eight species have not been previously reported from KSA are listed, including 25 new records for the Arabian Peninsula. Ten families were represented by single species in the RKNP.

Table 1.

The taxonomical and faunistic analysis of the beetle fauna in Rawdhat Khorim National Park, Central Kingdom of Saudi Arabia.

Family Subfamily Genera Species New record Relative abundance
Anthicidae 2 5 7 1 0.27
Bolboceratidae 1 1 1 0 0.01
Bostrichidae 2 3 3 1 1.30
Brentidae 2 2 2 0 1.10
Buprestidae 5 7 10 0 1.30
Carabidae 8 22 27 8 5.36
Cerambycidae 2 2 2 0 0.11
Chrysomelidae 5 8 8 1 3.95
Cleridae 2 4 4 0 0.51
Coccinellidae 1 8 15 0 3.44
Cryptophagidae 2 2 2 1 0.14
Curculionidae 4 15 20 0 1.99
Dascillidae 1 1 1 0 8.02
Dermestidae 4 6 15 5 1.58
Dytiscidae 2 3 3 0 0.56
Elateridae 2 8 13 2 9.30
Glaresidae 0 1 2 0 0.02
Heteroceridae 1 1 2 2 0.08
Histeridae 3 8 14 0 0.54
Hybosoridae 1 1 1 0 3.48
Hydrophilidae 1 1 1 1 0.04
Laemophloeidae 1 1 1 1 0.01
Leiodidae 1 1 1 1 0.01
Meloidae 2 2 4 0 0.17
Melyridae 2 3 4 1 1.23
Mycetophagidae 1 1 1 0 0.02
Nitidulidae 3 3 3 0 0.70
Oedemeridae 1 1 1 0 0.03
Phalacridae 1 1 1 0 0.17
Ptinidae 2 2 3 2 0.06
Scarabaeidae 7 18 34 3 36.62
Scraptiidae 1 1 2 0 0.47
Staphylinidae 5 7 8 4 0.66
Tenebrionidae 4 31 45 4 16.71
Thanerocleridae 1 1 1 0 0.03
Total 35 83 182 262 38

Eleven species (4.2%) are classified as abundant species, from which Aphodius ictericus ghardimaouensis (Scarabaeidae) (1,238 specimens) and Karumia inaequalis(Dascillidae) (1,262 specimens) were the highest abundant species. Thirty-six (13.7%) and 46 (17.6%) species are concidered as common and frequent, respectively. While, 64.5% of the species considered as rare (55 spp.) or very rare (114 spp.).

The RKNP beetle fauna demonstrated variable zoogeographical affinities (Figure 2), 15 species (6.1%) are endemic to KSA. The remainder of the species showed high affinities to SAR (36.4%), AFR_SAR (17.4%), and PAL_SAR (10.5%).

This work has confirmed the occurrence of four species in KSA previously reported as occurring in “Arabia”: Pseudoathyreus flavohirtus (Bolboceratidae) (Paulian, 1980); Saprinus figuratus (Histeridae) (Penati & Vienna, 2006); Sphenoptera gahani (Buprestidae) (Kerremans 1913); and Syntomus lateralis (Carabidae) (Schatzmayr 1936). The holotypes of three species originated from RKNP: Boromorphus saudicus Schawaller, Al Dhafer & Fadl, 2013; Dicronychus latifahae Al Dhafer & Platia, 2013 and Reichardtiolus aldhaferi Lackner, 2014.

In particular, Adelostoma subtile arabicum and A. subtile sabulosum, described from KSA by Kaszab (1981), but the characters for the differentiation of the two subspecies and the nominate A. subtile subtile Reitter that proposed by Kaszab (1981) are not useful for separation, and accordingly, this taxon is treated here as A. subtile.

It is expected that the results of this study of the beetle fauna of RKNP will be used for future studies related to long-term monitoring of the beetle fauna for evaluating impacts of increased use by the growing population of nearby Riyadh and impact of climate change (Bale et al. 2002).

List of species

Suborder: Adephaga

Carabidae

Subfamily: Cicindelinae

Myriochila melancholica (Fabricius, 1798)

World distribution

Africa: AO, BF, BJ, BW, CD, CF, CG, CI, CM, CV, ER, ET, GH, GM, GN, GQ, GW, KE, MG, MS, MW, MZ, NA, NG, SD, SL, SN, SO, TD, TO, TZ, ZA, ZM, ZW. Asia: AE, AF, AZ, BH, CN, CY, EG (Sinai), IL, IN, IQ, IR, JO, KG, KW, KZ, LB, NP, OM, PK, QA, SA, SY, TJ, TM, TR, UZ, YE. Europe: AL, AM, ES, FR, GE, GR, IT, MT, NO (Svalbard), PT. North Africa: DZ, EG, ES (Canary Island), LY, MA, TN.

General distribution

AFR_MAD_ORR_PAL_SAR_SJP.

Local distribution

AS, NJ (Britton 1948; Cassola and Schneider 1997)

Collecting month and method

A very rare species. The adults were collected by LT during V-VI.

Subfamily: Carabinae

Calosoma imbricatum Klug, 1832

World distribution

Africa: DJ, SD, TD (Britton 1948). Asia: AE, QA, SA, YE. North Africa: EG. Widespread all over Africa, the Arabian Peninsula and south Asia (Felix 2009).

General distribution

AFR_ORR_SAR.

Local distribution

AS, MK (Britton 1948), BA (El-Hawagry et al. 2013), EP, JZ, RI (Heinertz 1979).

Collecting month and method

A very rare species collected only by LT through III-IV.

Calosoma olivieri Dejean, 1831

World distribution

Asia: AE (Felix 2009), AZ, IQ, IR, JO, PK, SA, SY, TM, UZ. Europe: MT, PT (Azores). North Africa: DZ, EG, ES (Canary Island), LY, MA, TN.

General distribution

PAL_SAR.

Local distribution

MK (Britton 1948), RI (Heinertz 1979).

Collecting month and method

Very rare species. The beetles were collected by HP, and by PT under canopy of R. stricta in II.

Subfamily: Scaritinae

Distichus planus (Bonelli, 1813)

World distribution

Asia: AZ, IQ, IR, JO, KZ, PK, SA, SY, TJ, TM, UZ, YE. Europe: ES, FR, GE, GR, IT, MT, PT. North Africa: EG, MA, TU.

General distribution

PAL_SAR.

Local distribution

RI (Balkenohl 1994).

Collecting month and method

A frequent species that was collected HP, LT and PT during II, IV-VI, VIII, X, and XII.

Dyschirius beludscha ganglbaueri Znojko, 1927

World distribution

Asia: AE (Felix 2009), AF, EG (Sinai), IL, IQ, IR, KZ, MN, PK, SY, TJ, TM. North Africa: DZ, EG, MA, TU. New to KSA.

General distribution

PAL_SAR.

Collecting month and method

Very rare species that was collected by LT during IV.

Scarites procerus eurytus Fischer von Waldheim, 1828

World distribution

Asia: AF, IQ, IR, KG, KW, KZ, PK, SA, SY, TM, TR, UZ. Europe: ES, GE, GR, IT, PT, RU. North Africa: DZ, EG, LY, MA, TN.

General distribution

PAL_SAR.

Local distribution

EP, QS, RI (Balkenohl 1994), MK (Beccari 1971).

Collecting month and method

Very rare species. The adults were collected by HP and PT during V.

Subfamily: Siagoninae

Siagona europaea Dejean, 1826

World distribution

Asia: AF, CY, EG (Sinai), IL, IN, IQ, IR, JO, KZ, LB, PK, SA, SY, TJ, TM, TR, UZ, YE. Europe: AL, AM, AZ, BG, ES, GE, GR, HR, IT, MK, PT, RU.

General distribution

ORR_PAL_SAR.

Local distribution

MK (Britton 1948), RI (Al Dhafer et al. 2016).

Collecting month and method

A rare species. The beetles were collected by HP, LT and PT through IV-V, VII and XII.

Subfamily: Melaeninae

Cymbionotum pictulum (Bates, 1874)

World distribution

Africa: SD. Asia: Af, IQ, IR, KZ, SA, TM, TR. E: RU.

General distribution

AFR_PAL_SAR.

Local distribution

MK (Britton 1948; Ball and Shpeley 2005), RI (Al Dhafer et al. 2016).

Collecting month and method

Frequent species that was collected by HP and LT during II-V and XII. HP.

Cymbionotum semelederi (Chaudoir, 1864)

World distribution

Africa: MR, NE, SD, SO, TD. Asia: AE (Felix 2009), AF, AZ, CY, IQ, IR, JO, KW, KZ, PK, SA, SY, TM, TR, UZ, YE. Europe: AM, GE, RU. North Africa: DZ, EG, MA, TN.

General distribution

AFR_PAL_SAR.

Local distribution

EP, RI (Basilewsky 1979), MK (Britton 1948).

Collecting month and method

Frequent species. It was collected by LT and PT during II-V and XII.

Subfamily: Trechinae

Bembidion wittmeri (Basilewsky, 1979)

World distribution

Asia: SA.

General distribution

END.

Local distribution

RI (Basilewsky 1979).

Collecting month and method

Common species. Its specimens were collected by HP, LT and PT through I-V and XII.

Tachyura lucasi (Jacquelin du Val, 1852)

World distribution

Africa: CD, CF, CM, ET, GQ, NG, ZA. Asia: CY, IL, IR, LB, SY, TR. Europe: ES, IT, PT. North Africa: DZ, EG, MA, TN. New to Arabian Peninsula.

General distribution

AFR_MAD_PAL_SAR.

Collecting month and method

A rare species. The beetles were collected by LT through IV-V and VII.

Subfamily: Brachininae

Brachinus crepitans (Linné, 1758)

World distribution

Asia: AZ, CY, IQ, IR, KG, KZ, SY, TJ, TM, TR, UZ. Europe: AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IT, LT, LU, LV, MD, NL, NO, PL, PT, RO, RS, RU, SE, SI, SK, UA. New to Arabian Peninsula.

General distribution

PAL_SAR.

Collecting month and method

Rare species. It was collected by HP during IV and XII; and by LT during V.

Brachinus nobilis Dejean, 1831

World distribution

Asia: IL, IQ, IR, SA, SY, TR, YE. North Africa: DZ, MA, TN.

General distribution

SAR.

Local distribution

MK (Britton 1948), RI (Basilewsky 1979).

Collecting month and method

Rare species. The adults were collected by HP under debris and stones around temporary fresh water pool during XII.

Subfamily: Harpalinae

Amara aulica (Panzer, 1796)

World distribution

Asia: AZ, CN, KG, KZ, MN, RU, TJ, TM, TR, UZ. Europe: AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, EE, ES, FI, FO, FR, GB, GE, GR, HR, HU, IE, IT, LI, LT, LV, MD, MK, NL, NO, PL, RO, RS, RU, SK, SE, SI, UA. North Africa: ES (Canary Island). New to Arabian Peninsula.

General distribution

PAL_SAR_SJP.

Collecting month and method

A frequent species that was collected HP and PT through I-IV and XII.

Amara maindroni Bedel, 1907

World distribution

Asia: AE (Felix 2009), AF, IQ, IR, JO, PK, SA, SY. North Africa: DZ, MA, TN.

General distribution

SAR.

Local distribution

RI (Hieke 1993).

Collecting month and method

Very rare species. The adult was very rare and collected by Lt in XI.

Anthia duodecimguttata Bonelli, 1813

World distribution

Asia: AE (Felix 2009), IQ, IR, JO, KW, OM, QA, SA, YE. North Africa: EG.

General distribution

SAR.

Local distribution

AS, JZ, MK, RI (Britton 1948; Beccari 1971; Heinertz 1979).

Collecting month and method

Very rare species. The adult beetles were collected by HP and LT at sandy area during IV.

Calodromius mayeti (Bedel, 1907)

World distribution

Asia: AE (Felix 2009), IR, SA. North Africa: MA, TN.

General distribution

SAR.

Local distribution

MD (Mateu 1986).

Collecting month and method

Very rare species that was collected by HP under stones in IV.

Chlaenius flavipes Menetries, 1832

World distribution

Asia: AF, AZ, IQ, IR, KG, KZ, TJ, TM, TR, UZ. Europe: AL, AM, BA, BG, GE, GR, HR, HU, MD, MK, RO, RS, RU, UA. New to Arabian Peninsula.

General distribution

PAL_SAR.

Collecting month and method

Very rare species that was collected by HP under plant debris during XII.

Harpalus affinis (Schrank, 1781)

World distribution

Asia: AZ, CN, CY, IL, IR, KG, KP, KZ, MN, TR. Australia: AU, NZ. Europe: AD, AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IT, LI, LT, LU, LV, MD, MK, NL, NO, PL, PT, RO, RS, RU, SE, SI, SK, UA. North America: CA, US. New to Arabian Peninsula.

General distribution

SCO.

Collecting month and method

A rare species. The specimens were collected by HP under stones and plant debris during II, IV and XII.

Merizomena buettikeri (Mateu, 1986)

World distribution

Asia: SA.

General distribution

END.

Local distribution

MD (Mateu 1986).

Collecting month and method

Very rare species. It was collected by LT in IV and VI.

Microlestes discoidalis (Fairmaire, 1892)

World distribution

Africa: ER, KE, MR, NE, SD, SO, TD. Asia: AE, AF, IL, IN, IR, SA, TR, YE.

General distribution

AFR_ORR_SAR.

Local distribution

MK (Britton 1948; Mateu 1979).

Collecting month and method

Very rare species that was collected by LT through V.

Orthotrichus cymindoides (Dejean, 1831)

World distribution

Asia: AF, CN, IN, SY. North Africa: EG. New to Arabian Peninsula.

General distribution

ORR_PAL_SAR_SJP.

Collecting month and method

Frequent species. The adults were collected by HP, LT and PT during I-V, IX and XII.

Platytarus faminii faminii (Dejean, 1826)

World distribution

Africa: CV. Asia: AE (Felix 2009), AZ, CY, IL, IQ, KG, KZ, SA, SY, TM, TR, UZ, YE (Socotra). Europe: AM, ES, FR GR, IT, PT. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

AFR_PAL_SAR.

Local distribution

AS, EP, RI (Mateu 1986).

Collecting month and method

Very rare species that was collected by LT in III.

Poecilus wollastoni (Wollaston, 1854)

World distribution

Asia: IQ, KW, SA, YE. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN.

General distribution

SAR.

Local distribution

It was recorded from Arabia without exact locality by Emden (1954) and a recent occurrence in KSA has been confirmed by Al Dhafer et al. (2016).

Collecting month and method

Frequent species, which was collected by HP, LT and PT during I-II, IV-VI and XI-XII.

Stenolophus marginatus Dejean, 1829

World distribution

Asia: AE (Felix 2009), AF, AZ, CY, IL, IQ, IR, JO, KW, KZ, SA, SY, TJ, TM, TR, UZ. Europe: AL, AM, BA, BG, ES, FR, GE, GR, HR, IT, MK, PT, RS, UA. North Africa: DZ, EG, ES (Canary Islands), MA, PT (Madeira Archipelago), TN.

General distribution

PAL_SAR.

Local distribution

EP, RI (Basilewsky 1979).

Collecting month and method

Frequent species. The beetles were collected by LT in IV-V, VII and IX.

Stenolophus pseudoobockianus Felix & Muilwijk, 2009

World distribution

Asia: AE (Felix 2009). New to KSA.

General distribution

SAR.

Collecting month and method

Very rare species that was collected by LT during IV.

Syntomus lateralis (Motschulsky, 1855)

World distribution

Asia: AE (Felix 2009), IL, IQ, IR, SA, SY. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

Local distribution

Schatzmar (1936) mentioned Arabia among the distribution of this species in his work on Carabidae of Egypt, without given any further detailed about the locality.

General distribution

SAR.

Collecting month and method

Common species that was collected by HP, LT and PT in I-XII.

Dytiscidae

Subfamily: Dytiscinae

Eretes sticticus (Linné, 1767)

World distribution

Africa: BW, KE, MG, NA, SD, ZA, ZW. Asia: AE, AZ, BT, CN, CY, EG (Sinai), IN, IQ, IR, JA, á, KW, NP, OM, PK, SA, SY, TM, TR, TW, YE. Europe: AM, BA, BG, ES, FR, GE, GR, HR, HU, IT, PT, RU, SI, UA, RS. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN. North America: MX, PR, US. South America: EC, PE, VE.

General distribution

COS.

Local distribution

AS, EP, MD, RI (Brancucci 19791984).

Collecting month and method

A frequent species. The adult beetles were collected by LT in IV-VI.

Subfamily: Hydroporinae

Hydroglyphus signatellus (Klug, 1834)

World distribution

Africa: ET, KE, SD, SN. Asia: AE (Hájek and Brancucci 2011), AZ, CY, EG (Sinai), IL, IQ, IR, JO, KW, KZ, PK, SA, SY, TJ, TM, TR, UZ, YE. Europe: AM, BA, ES, GE, GR, HR, IT, RS, RU. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_PAL_SAR.

Local distribution

EP, RI (Brancucci 19791984).

Collecting month and method

Very rare species that was collected by LT during V.

Hygrotus inscriptus (Sharp, 1882)

World distribution

Asia: AE, IQ, IR, KW, SA, SY, TM, UZ. North Africa: EG. The distribution is updated from Hájek and Brancucci (2011).

General distribution

SAR.

Local distribution

EP (Brancucci 1984).

Collecting month and method

Frequent species that was collected by LT in VI.

Suborder: Polyphaga

Hydrophilidae

Subfamily: Hydrophilinae

Hydrochara flavipalpis (Boheman, 1851)

World distribution

Asia: OM, YE. New to KSA.

General distribution

SAR.

Collecting month and method

A rare species, which was collected by HP during V and XII.

Histeridae

Subfamily: Abraeinae

Teretrius pulex Fairmaire, 1877

World distribution

Africa: MR, NE, SN. Asia: AE (Kanaar 2007), OM, SA, SY. North Africa: DZ, EG, TN.

General distribution

AFR_SAR.

Local distribution

EP (Mazur 1994; Penati and Vienna 2006).

Collecting month and method

Very rare species that was collected by BV on branches of Z. nummularia during V.

Subfamily: Sapriniae

Pholioxenus sp.1

Collecting month and method

A rare species. The specimens were collected by PT under canopy of A. ehrenbergiana, A. gerrardii and R. stricta during IV-V and X.

Pholioxenus sp.2

Collecting month and method

Frequent species. The beetles were collected by PT under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by LT through V- XI.

Reichardtiolus aldhaferi Lackner, 2014

World distribution

Asia: SA (Lackner 2014).

General distribution

END.

Local distribution

RI (Lackner 2014).

Collecting month and method

A rare species. The beetles were collected by HP and LT during I-II and XII.

Saprinus chalcites (Illiger, 1807)

World distribution

Africa: AO, BF, ET, GM, KE, ML, MR, NA, SD, SN, SO, TZ. Asia: AE, AF, CY, ID, IL, IN, IQ, IR, JO, KW, KZ, MM, MN, OM, PK, SA, TR, YE. Australia: AU. Europe: ES, FR, GR, IT, MT, PT, RU. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN. South America: AR.

General distribution

SCO (Penati and Vienna 2006).

Local distribution

AS, BA, JZ, MK, NJ, RI (Kryzhanovskij 1979; Mazur 1994; Penati and Vienna 2006).

Collecting month and method

Very rare species. The adults were collected by PT under canopies of A. ehrenbergiana, L. shawii and Z. nummularia during IV and VII.

Saprinus confalonierii G. Müller, 1933

World distribution

Africa: MR. Asia: AE (Kanaar 2007), OM, SA. North Africa: DZ, EG, LY, TN.

General distribution

AFR_SAR.

Local distribution

AS, EP, HA, NB, NJ, RI (Mazur 1994; Penati and Vienna 2006).

Collecting month and method

Very rare species that was collected by LT in X.

Saprinus figuratus Marseul, 1855

World distribution

Asia: IL, JO, SA, SY. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

SAR.

Local distribution

Just recorded from Arabia (Penati and Vienna 2006).

Collecting month and method

Very rare species that was collected by HP during II.

Saprinus moyses Marseul, 1862

World distribution

Asia: IR, KW, SA, SY, TR. Europe: GR. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

PAL_SAR.

Local distribution

HA (Mazur 1994; Penati and Vienna 2006).

Collecting month and method

Very rare species. The beetles were collected by BT and LT during V, VII and X.

Saprinus muelleri Mazur, 1997

World distribution

Asia: AE (Kanaar 2007), AF, PK, SA.

General distribution

SAR.

Local distribution

EP, MK, RI (Penati and Vienna 2006).

Collecting month and method

Very rare species that was collected by LT during I-II.

Saprinus sp.

Collecting month and method

A rare species. The adults were collected by LT during V-VI and X.

Xenonychus tridens (Jacquelin Duval, 1853)

World distribution

Africa: CV, MR, NG, TD. Asia: AE (Kanaar 2007), CY, IL, OM, SA, SY, TR.

Europe: ES, FR, GR, IT, PT. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

AFR_PAL_SAR.

Local distribution

EP, MK (Mazur 1994; Penati and Vienna 2006), RI (Al Dhafer et al. 2016).

Collecting month and method

Very rare species that was recorded by PT under canopy of A. ehrenbergiana during IV.

Xenophilothis choumovitchi (Thérond & Hollande, 1965)

World distribution

Asia: AE (Kanaar 2007), OM, SA. North Africa: DZ.

General distribution

SAR.

Local distribution

EP, RI (Penati and Vienna 2006).

Collecting month and method

Very rare species, which was collected by LT in V.

Zorius sp.

Collecting month and method

A rare species. The adults were collected by PT under canopies of A. gerrardii, C. procera, L. shawii and R. stricta during III-IV.

Subfamily: Histerinae

Atholus bimaculatus (Linnaeus, 1758)

World distribution

Africa: BF, CF, CM, DJ, KE, MR, SN, TD. Asia: AE (Kanaar 2007), AF, CN, CY, IL, IN, IQ, IR, JO, JP, KG, KP, KR, KZ, MM, OM, RU, SA, SY, TJ, TM, UZ, YE. Europe: AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IT, LI, LV, NL, NO, PL, PT, RO, RS, RU, SE, SI, SK, UA. North Africa: DZ, ES (Canary Islands), LY, MA, TN. North America: CA, US. South America: AR, BR, CL.

General distribution

SCO (Penati and Vienna 2006).

Local distribution

AS (Kryzhanovskij 1979), RI (Al Dhafer et al. 2016).

Collecting month and method

Very rare species that was collected by PT under canopy of A. ehrenbergiana during IV.

Leiodidae

Subfamily: Leiodinae

Chobautiella anisotomoides (Fairmaire, 1876)

World distribution

Asia: AE (Švec 2010a). North Africa: DZ, ES (Canary Islands), MA, TN (Švec 2010). New to KSA.

General distribution

SAR.

Collecting month and method

Very rare species. The specimens were collected by LT in XII.

Staphylinidae

Subfamily: Pselaphinae

Ctenisomorphus major (Raffray, 1877)

World distribution

Africa: ET. Asia: AE (Besuchet and Cuccodoro 2011), EG (Sinai), IL, IR, JO, SA, TR, YE. North Africa: DZ, EG, TN.

General distribution

AFR_SAR.

Local distribution

AS, BA, EP, MD, RI (Besuchet 1981).

Collecting month and method

Very rare species that was collected by LT in IV-V and VII.

Enoptostomus arabicus Besuchet & Cuccodoro, 2011

World distribution

Asia: AE (Besuchet and Cuccodoro 2011). New to KSA.

General distribution

SAR.

Collecting month and method

Very rare species, which was collected by LT during VII.

Subfamily: Tachyporinae

Sepedophilus sp.

Collecting month and method

Very rare species. It was collected by PT under canopy of Z. nummularia in III.

Subfamily: Aleocharinae

Atheta atramentaria (Gyllenhal, 1810)

World distribution

Africa: MR, ZA. Asia: AZ, CN, CY, IN, IR, JP, KP, KZ, NP, PK, RU, TR. Europe: AL, AT, BE, BG, CH, CZ, DE, DK, EE, ES, FI, FO, FR, GB, GE, GR, HU, IE, IS, IT, LT, LU, NL, NO, PL, PT, RU, SE, SK. North Africa: DZ, EG, ES (Canary Islands), MA, PT (Madeira Archipelago), TN. New to Arabian Peninsula.

General distribution

AFR_PAL_ORR_SAR_SJP.

Collecting month and method

Rare species. The specimens of this species were collected by PT under canopies of A. ehrenbergiana, A. gerrardii, L. shawii, R. stricta, and Z. nummularia in I-II and X-XI; and by SW on branches of C. procera through VII.

Subfamily: Oxytelinae

Bledius niloticus Erichson, 1840

World distribution

Africa: AO, ET, SN, ZA. Asia: CN, IL, IN, JP, KP, KR, LB, LK, SA (Beccari 1971), SY. North Africa: DZ, EG, MA, TN.

General distribution

AFR_ORR_PAL_SAR_SJP.

Local distribution

RI (Beccari 1971).

Collecting month and method

Very rare species that was collected by LT during X.

Carpelimus pusillus (Gravenhorst, 1802)

World distribution

Asia: CY, IR, KZ, RU, TR. Australia: AU. Europe: AL, AM, AT, BA, BE, BG, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IS, IT, LT, LV, NL, NO, PL, PT, RO, RU, SE, SI, SK, UA. North Africa: DZ, EG, ES (Canary Islands), PT (Madeira Archipelago). North America: US. New to Arabian Peninsula.

General distribution

SCO.

Collecting month and method

Very rare species, which was collected by LT through XII.

Carpelimus sp.

Collecting month and method

Very rare species that was collected by LT in VII.

Subfamily: Paederinae

Philonthus sp.

Collecting month and method

Very rare species that was collected by LT through VI.

Glaresidae

Glaresis arabica (Paulian, 1980)

World distribution

Asia: OM, SA.

General distribution

SAR.

Local distribution

EP, RI (Paulian 1980).

Collecting month and method

Very rare species that was collected by LT during V.

Glaresis sp.

Collecting month and method

Very rare species that was collected by LT in IV.

Bolboceratidae

Subfamily: Bolboceratinae

Pseudoathyreus flavohirtus (Wlaker, 1871)

World distribution

Asia: Arabia

General distribution

SAR.

Local distribution

Paulian (1980) recorded this species from Arabia without exact locality.

Collecting month and method

Very rare species that was collected by LT in IV.

Hybosoridae

Subfamily: Hybosorinae

Hybosorus illigeri Reiche, 1853

World distribution

Africa: KM, MG, UG. Asia: AF, AZ, CN, CY, EG (Sinai), IL, IN, IQ, IR, JO, PK, SA, SY, TJ, TM, TR, UZ, YE, VN. Europe: AM, BG, ES, FR, GE, GR, HR, IT, MK, PT, RO, RS, RU, UA. North Africa: DZ, EG, LY, MA, TN. North America: CU, HT, MX, NI, US. South America: VE.

General distribution

SCO.

Local distribution

EP, MD, MK, NJ, RI (Beccari 1971, Kuijten 1980).

Collecting month and method

An abundant species. The adult beetles were collected by HP, LT and PT through II, IV-VIII.

Scarabaeidae

Subfamily Eremazinae

Eremazus giganteus Král, 2015

World distribution

Asia: AE (Král and Batelka 2015). New to KSA.

General distribution

SAR.

Collecting month and method

Very rare species that collected by LT during V.

Eremazus unistriatus Mulsant, 1851

World distribution

Africa: NE, SD. Asia: AE, AF, IL, IN, IQ, IR, KZ, PK, SA, TJ, TM, TR, UZ. Europe: AM, GE. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

AFR_ORR_PAL_SAR.

Local distribution

EP, HA, MD, MK, QS, RI (Pittino 1984).

Collecting month and method

A frequent species. The adults of this species were collected by PT under canopy of A. ehrenbergiana, and by HP and LT during I-II, IV-VII and IX.

Subfamily: Aphodiinae

Aphodius adustus Klug, 1855

World distribution

Africa: CD, CM, DJ (Pittino 1984), GW, KE, MZ, NA, SD (Balthasar 1972), SN, TD, TZ. Asia: SA, YE.

General distribution

AFR_SAR.

Local distribution

The species was reported from Arabia without exact locality (Pittino 1984) and recently its occurrence in KSA: RI has been confirmed by Abdel-Dayem et al. (2016).

Collecting month and method

Very rare species that was collected by LT during IV.

Aphodius arabicus Harold, 1875

World distribution

Asia: EG (Sinai), IL, IR, KW, SA, YE. North Africa: EG

General distribution

SAR.

Local distribution

EP, JZ, MD, MK, RI (Pittino 1984).

Collecting month and method

Common species that was collected by LT through V-VII and IX-X.

Aphodius beluchistanicus Petrovitz, 1962

World distribution

Asia: IQ, IR, SA.

General distribution

SAR.

Local distribution

EP, RI (Pittino 1984).

Collecting month and method

A rare species, which was collected by LT in I and XII.

Aphodius chobauti Clouët, 1896

World distribution

Asia: IL, IQ, IR, JO, SA. North Africa: DZ, EG, LY, MA, TN.

General distribution

SAR.

Local distribution

MD, QS, RI (Pittino 1984).

Collecting month and method

Rare species. The specimens were collected by BV on branches of A. gerrardii, and by HP and LT during IV-V and XI.

Aphodius ictericus ghardimaouensis Balthasar, 1929

World distribution

Asia: CY, IL, IR, JO, LB, SA, SY, TR. Europe: ES, FR, GR, HR, IT, MT, PT. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

PAL_SAR.

Local distribution

EP (Pittino 1984).

Collecting month and method

Abundant species that was collected by LT during I, III and IX.

Aphodius lividus (Olivier, 1789)

World distribution

Africa: MG, NA. Asia: CN, CY, EG (Sinai), IL, IR, KG, KW, KZ, LB, MN, NP, OM, SA, SY, TJ, TM, TR, TW, UZ. Australia: AU, NZ, PG. Europe: AL, AM, AT, BA, BE, BY, CH, CZ, DE, EE, ES, FI, FR, GB, GE, GR, HR, HU, IT, LT, LV, MK, MT, NL, PL, PT, RO, RS, RU, SE, SI, SK, UA. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN. North America: CU, GT, MX, NI, PA, US.

General distribution

SOC.

Local distribution

AS, BA, EP, JZ, MD, MK, QS, RI (Beccari 1971, Pittino 1984; El-Hawagry et al. 2013).

Collecting month and method

Common species. The beetles were collected by LT through IV-VII and IX-XII.

Aphodius luridus (Fabricus, 1775)

World distribution

Asia: CN, CY, IL, IR, KG, KZ, RU, SA, SY, TJ, TM, TR. Europe: AL, AM, AT, BA, BE, BG, BY, CH, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IT, LT, LV, MK, NL, NO, PL, PT, RO, RS, RU, SE, SI, SK, TR, UA. North Africa: MA, TN.

General distribution

PAL_SAR_SJP.

Local distribution

RI (Abdel-Dayem et al. 2016).

Collecting month and method

Very rare species that was collected by LT in I and X.

Aphodius pruinosus Reitter, 1892

World distribution

Africa: MR. Asia: AF, IL, IQ, IR, KG, KW, KZ, LB, OM, PK, SA, TJ, TM, TR, UZ. Europe: RU. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_PAL_SAR.

Local distribution

BA, EP, HA, JZ, MD, MK, QS, RI (Pittino 1984).

Collecting month and method

An abundant species. The adults were collected by PT by PT under canopies of L. shawii and R. stricta; and by LT through II-VI and IX-X.

Aphodius rendallii (Wollaston, 1867)

World distribution

Asia: IL, IQ, JO, SA. North Africa: DZ, EG, LY, MA, TN.

General distribution

SAR.

Local distribution

EP, HA, QS, RI (Pittino 1984) [under the name Aphodius opacior D. Koshantschikov 1894] (Král and Batelka 2015).

Collecting month and method

Common species. The specimens of this species were collected by PT under canopies of A. ehrenbergiana, A. gerrardii, L. shawii, R. stricta and Z. nummularia; and by LT through IV-V and IX-X.

Aphodius translucidus Petrovitz, 1961

World distribution

Asia: AF, IN, IQ, IR, OM, PK, RU, SA, TM. North Africa: EG (Pittino 1984).

General distribution

ORR_SAR.

Local distribution

EP, MD, MK, RI (Pittino 1984).

Collecting month and method

Common species that was collected by LT in IV-V and IX-XI.

Aphodius wollastoni iranicus Balthasar, 1946

World distribution

Asia: EG (Sinai), IN, IQ, IR, JO, OM, PK, SA, SY, YE.

General distribution

ORR_SAR.

Local distribution

EP, HA, MD, QS, RI (Pittino 1984).

Collecting month and method

Abundant species. It was collected by PT under canopies of A. ehrenbergiana and C. procera; and by HP and LT through I-V and XI-XII.

Granulopsammodius plicatulus (Fairmeire, 1892)

World distribution

Africa: ET, SD, SO. Asia: SA, YE. North Africa: DZ, EG, LY, MA.

General distribution

AFR_SAR.

Local distribution

AS, BA, EP, MD, MK, RI (Paulian 1980; Pittino 1984; El-Hawagry et al. 2013).

Collecting month and method

Common species, which was collected by PT under canopy of Z. nummularia; and by LT through IV-VIII.

Leiopsammodius laevicollis (Klug, 1845)

World distribution

Africa: CG, DJ, ER, ET, SD, SO. Asia: IL, SA, SY, YE. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_SAR.

Local distribution

BA, HA, JZ, MD, MK, RI (Pittino 1984; El-Hawagry et al. 2013).

Collecting month and method

A rare species that was collected by PT under canopy of L. shawii and R. stricta; and by LT through IV-VI, IX and XII.

Pararhyssemus coluber (Mayet, 1887)

World distribution

Africa: SO. Asia: AF, EG (Sinai), IR, SA. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_SAR.

Local distribution

AS, BA, HA, MD, MK, RI (Pittino 1984).

Collecting month and method

Very rare species. The beetles were collected by LT in IV.

Pleurophorus arabicus (Pittino & Mariani, 1986)

World distribution

Asia: AZ, IR, JO, SA, TM. Europe: RU.

General distribution

PAL_SAR.

Local distribution

EP, QA, RI (Pittino 1984) [under the name Pleurophorus anatolicus Petrovitz 1961] (Král and Batelka 2015).

Collecting month and method

A common species. The adult beetles were collected by PT under canopies of A. ehrenbergiana, A. gerrardii, C. procera and R. stricta; and by HP and LT through III-V and XII.

Pseudomothon sp.

Collecting month and method

Rare species that was collected by LT during IV.

Rhyssemus brevitarsis Pinttino, 1984

World distribution

Asia: SA.

General distribution

END.

Local distribution

BA, JZ, MD, MK, QS, RI (Pittino 1984; El-Hawagry et al. 2013).

Collecting month and method

Very rare species that was collected by LT during IV.

Rhyssemus granosus (Klug & Erichson, 1842)

World distribution

Africa: CM, CV, ET, KE, ML, MR, NE, NG, SD, SN, SO, TD, TZ. Asia: SA, YE. North Africa: EG.

General distribution

AFR_SAR.

Local distribution

AS, BA, EP, JZ, MD, MK, QS, RI (Beccari 1971, Pittino 1984; El-Hawagry et al. 2013).

Collecting month and method

A rare species. The beetles were collected by LT through IV-V, VII and X.

Rhyssemus saoudi Pittino, 1984

World distribution

Asia: SA.

General distribution

END.

Local distribution

AS, BA, JZ, MK, RI (Pittino 1984; El-Hawagry et al. 2013; Abdel-Dayem et al. 2016).

Collecting month and method

Common species. The adult beetles were collected by PT under canopies of C. procera, L. shawii, R. stricta and Z. nummularia; and by HP and LT through III-V and VII-XI.

Subfamily Scarabaeinae

Metacatharsius inermis (Laporte, 1840)

World distribution

Africa: ER, ET, GM, KE, MR, SD, SN, SO, TD. Asia: IN, IQ, IR, PK, SA. North Africa: EG.

General distribution

AFR_ORR_SAR.

Local distribution

AS, EP, RI (Paulian 1980).

Collecting month and method

Rare species, which was collected during V.

Scarabaeus bannuensis A. Janssens, I940

World distribution

Africa: MR, TD. Asia: IQ, IR, PK, SA. North Africa: DZ, LY, MA, TN.

General distribution

AFR_SAR.

Local distribution

RI (Ziani and Gudenzi 2012)

Collecting month and method

Common species. The specimens were collected by PT under canopies of A. ehrenbergiana, A. gerrardii, L shawii and R. stricta; and by HP, LT, SW during III-VIII.

Scarabaeus cristatus Fabricius, 1775

World distribution

Africa: ER, GN, MR, NE, SD, SN, TD. Asia: AE, AF, IL, IQ, IR, PK, SA. North Africa: EG, LY.

General distribution

AFR_SAR.

Local distribution

AS, EP, MK, RI (Paulian 1980).

Collecting month and method

Very rare species. The specimens were collected by HP and LT during V and VIII.

Subfamily: Dynamopodinae

Orubesa plicifrons (Fairmaire, 1897)

World distribution

Africa: ET, NE, SN. Asia: SA (Paulian 1980). North Africa: MA.

General distribution

AFR_SAR.

Local distribution

RI (Paulian 1980).

Collecting month and method

A common species. The adult beetles were collected by HP and LT in IV.

Subfamily Melolonthinae

Maladera insanabilis (Brenske, 1894)

World distribution

Asia: AE, AF, IL, IN, IQ, IR, JO, KW, NP, OM, PK, SA. North Africa: LY.

General distribution

ORR_SAR.

Local distribution

EP, RI (Ahrens 2000).

Collecting month and method

Very rare species that was collected by LT during V, VII and XI.

Schizonycha buettikeri Sabatinelli & Pontuale, 1998

World distribution

Asia: OM, SA.

General distribution

SAR.

Local distribution

EP, HA, RI (Sabatinelli and Pontuale 1998).

Collecting month and method

A rare species that was collected by LT during IV-V.

Schizonycha flavicornis Brenske, 1898

World distribution

Africa: SD, SO. Asia: SA, YE. North Africa: EG.

General distribution

AFR_SAR.

Local distribution

AS, BA, EP, JZ, MD, MK, RI (Sabatinelli and Pontuale 1998).

Collecting month and method

Rare species, which was collected by LT in IV and X-XI.

Sphaerotrochalus somalicola (Frey, 1960)

World distribution

Africa: ET, SO. Asia: OM, SA, YE.

General distribution

AFR_SAR.

Local distribution

AS, BA, HA, JZ, MK, RI (Ahrens 2000).

Collecting month and method

A common species. The specimens were collected by PT under canopies of A. ehrenbergiana, A. gerrardii and R. stricta; and by LT during I, IV-V and X-XII.

Subfamily Rutelinae

Clipadoretus habibi Král, 2015

World distribution

Asia: AE (Král and Batelka 2015). New to KSA.

General distribution

SAR.

Local distribution

RI (Beccari 1971).

Collecting month and method

A rare species that was recorded by LT in V-VII and IX.

Clipadoretus sp.

Collecting month and method

Moderately common species that was collected by LT during V.

Phaeadoretus syriacus (C. É. Blanchard, 1851)

World distribution

Asia: IQ, IR. New to Arabian Peninsula.

General distribution

SAR.

Collecting month and method

Abundant species. The specimens were collected by PT under canopies of A. ehrenbergiana, A. gerrardii, L. shawii and Z. nummularia; and by HP and LT through I-V.

Subfamily Dynastinae

Pentodon algerinus dispar Baudi, 1870

World distribution

Africa: ER. Asia: CY, IL, IQ, IR, JO, KW, OM, QA, SA, SY, YE. Europe: AM, GR.

General distribution

AFR_PAL_SAR.

Local distribution

EP, MK, RI (Endrödi 1980).

Collecting month and method

Common species. The adults were collected by PT under canopies of R. stricta and Z. nummularia; and by HP and LT through IV-V and VIII-IX.

Podalgus cuniculus arabicus Fairmaire, 1895

World distribution

Asia: AE, EG (Sinai), IL, KW, OM, SA, YE. North Africa: EG.

General distribution

SAR.

Local distribution

EP, MK, RI (Endrödi 1980).

Collecting month and method

A common species that was collected by HP and LT through IV-XI.

Dascillidae

Subfamily: Karumiinae

Karumia inaequalis Pic, 1929

World distribution

Asia: SA.

General distribution

END.

Local distribution

EP, RI (Wittmer 1979).

Collecting month and method

Abundant species that was collected by PT under canopies of A. ehrenbergiana, and by LT during IV-IX.

Buprestidae

Subfamily: Julodinae

Julodis euphratica Laporte & Gory, 1835

World distribution

Asia: AF, EG (Sinai), IQ, IR, JO, OM, SA.

General distribution

SAR.

Local distribution

EP, QS, RI (Bílý 19821985).

Collecting month and method

It is a rare and was collected by BV, HP and VC during I and IV-VI.

Subfamily: Polycestinae

Acmaeoderella arabica Cobos, 1963

World distribution

Asia: IL, IR, OM, SA.

General distribution

SAR.

Local distribution

RI (Cobos 1963).

Collecting month and method

Very rare species, which was collected by BV on branches of Z. nummularia during VI.

Xantheremia pantherina (Bílý, 1979)

World distribution

Asia: IL, IQ, SA. North Africa: EG.

General distribution

SAR.

Local distribution

EP, RI (Bílý 19821985).

Collecting month and method

A rare species. The specimens were collected by BV on branches of L. shawii, HP and by SW of R. stricta during V, VII and VIII.

Subfamily: Chrysochroinae

Sphenoptera gahani Kerremans, 1913

World distribution

Asia: Arabia (Kerremans 1913)

General distribution

END.

Local distribution

This species is described by Kerremans (1913) from Arabia, without any further information about the type locality.

Collecting month and method

Very rare species that was collected only by BV on branches of A. gerrardii during X.

Sphenoptera magna Gory & Laporte, 1839

World distribution

Asia: IL, IQ, IR, JO, SA, SY, TR.

General distribution

PAL_SAR.

Local distribution

EP, QS (Bílý 1980).

Collecting month and method

Very rare species that was collected by HP during V.

Subfamily: Buprestinae

Anthaxia kneuckeri Obenberger, 1920

World distribution

Asia: EG (Sinai), IL, JO, OM, SA.

General distribution

SAR.

Local distribution

AS, MK, RI (Bílý 1980), BA (El-Hawagry et al. 2013).

Collecting month and method

Common species. The adults were collected by BV and VC on branches of A. ehrenbergiana and A. gerrardii, and by HP, LT and PT during IV-XI.

Anthaxia marginifera metallenscens Bílý, 1999

World distribution

Asia: IL, SA.

General distribution

SAR.

Local distribution

RI (Bílý 1979).

Collecting month and method

Very rare species that was collected by BV on branches of A. ehrenbergiana during IV.

Subfamily: Agrilinae

Agrilus desertus (Klug, 1829)

World distribution

Africa: MR, SO, TD. Asia: IL, IR, JO, SA, YE. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_SAR.

Local distribution

AS (Bílý 1982).

Collecting month and method

A rare species. The specimens were collected by BV on branches of A. ehrenbergiana, A. gerrardii and L. shawii, and HP during IV-V, VIII and X.

Agrilus lituratus (Klug, 1829)

World distribution

Africa: MR, SD, SN, TD (Bílý 1982). Asia: IL, IR, JO, SA, SY, YE. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_SAR.

Local distribution

AS, BA, MK, RI (Bílý 1982).

Collecting month and method

Frequent species that was collected by BV, SW and VC on branches of A. ehrenbergiana, A. gerrardii and C. procera, and PT during IV-VII and IX.

Trachys latifrons Kerremans, 1907

World distribution

Africa: ER, ET, SD. Asia: AE, SA, YE.

General distribution

AFR_SAR.

Local distribution

AS (Bílý 1979), RI (Al Dhafer et al. 2016).

Collecting month and method

Very rare species. It was collected by PT under the canopy of Z. nummularia during IV.

Heteroceridae

Subfamily: Heterocerinae

Augyles sericans (Kiesenwetter, 1843)

World distribution

Asia: IL, SY. Europe: AL, AT, BG, CH, CZ, DE, DK, ES, FR, GB, HR, HU, IT, NL, PL, RO, SE, SI, SK, UA. New to Arabian Peninsula.

General distribution

PAL_SAR.

Collecting month and method

Rare species that was collected by Lt during IV-V.

Augyles turanicus (Reitter, 1887)

World distribution

Asia: AE (Mascagni 2009), IL, IQ, IR, KZ, SY, TM, TR, UZ. Europe: GE. North Africa: DZ. New to KSA.

General distribution

PAL_SAR.

Collecting month and method

Very rare species, which was collected by LT in IV.

Elateridae

Subfamily: Agrypninae

Aeoloides grisescens (Germar, 1844)

World distribution

Africa: SD, TD. Asia: AE, AF, AZ, CN, CY, IQ, IR, KG, KZ, MN, OM, PK, QA, SA, SY, TJ, TM, TR, UZ, YE. Europe: AM, GE, GR, RU, UA. North Africa: EG, LY, MA.

General distribution

AFR_PAL_SAR_SJP.

Local distribution

EP, JZ, QS, RI (Platia and Schimmel 1997).

Collecting month and method

Common species. The adults were collected by PT under canopies of A. ehrenbergiana, L. shawii, R. stricta and Z. nummularia; and by HP and LT during I-VII and IX-X.

Aeoloides holzschuhi Platia & Schimmel, 1997

World distribution

Asia: AE (Platia 2007), IR, OM, PA. New to KSA.

General distribution

SAR.

Collecting month and method

Frequent species that was collected by PT under canopies of A. ehrenbergiana, A. gerrardii, L. shawii and R. stricta; and by LT during I-V and XI.

Conoderus productus arabicus (Chassain, 1979)

World distribution

Asia: AE (Platia 2007), IR, OM, QA, SA, YE.

General distribution

SAR.

Local distribution

EP, MK, QS, RI (Chassain 19791983).

Collecting month and method

A frequent species. It was collected by LT through IV-VII.

Heteroderes gallagheri Platia & Schimmel, 1997

World distribution

Asia: AE (Platia 2007), IR, OM, PK. New to KSA.

General distribution

SAR.

Collecting month and method

Frequent species. The adults were collected by LT through IV-VI and X.

Heteroderus ruteri Chassain, 1979

World distribution

Asia: OM, SA.

General distribution

SAR.

Local distribution

MD, MK, RI (Chassain 1979).

Collecting month and method

Very rare species that was collected by PT under canopies of A. gerrardii and C. procera during I-II.

Lacon modestus (Boisduval, 1835)

World distribution

Africa: SN. Asia: AE (Platia 2007), ID, IL, IQ, IR, JP, LA, OM, PK, QA, SA, TW, VN, YE (Socotra). Australia: AU, FJ, MP, NC. North America: GT, US. South America: GF.

General distribution

COS.

Local distribution

EP, MK, RI (Chassain 1983).

Collecting month and method

28.XII.2011

Collecting method

Very rare species, which was collected by HP in XII.

Lanelater buettikeri Chassain, 1983

World distribution

Asia: AE (Platia 2007), OM, SA, YE.

General distribution

SAR.

Local distribution

JZ, MK, NJ, RI (Chassain 1983, Platia and Schimmel 1997).

Collecting month and method

Frequent species. The beetles were collected by LT during VIII-X.

Subfamily: Cardiophorinae

Craspedostethus dilutus (Erichson, 1840)

World distribution

Africa: ER, ET, SO. Asia: AE, EG (Sinai), IR, OM, SA, SY, YE. North Africa: DZ, EG, MA. The distribution was updated from (Platia 2012).

General distribution

AFR_SAR.

Local distribution

BA, EP, MK (Platia and Schimmel 1997), RI (Al Dhafer et al. 2016).

Collecting month and method

Abundant species that was collected by PT under canopies of Z. nummularia and by LT during IV-IX.

Craspedostethus flavescens Chassain, 1979

World distribution

Asia: AE (Platia 2007), OM, SA, YE.

General distribution

SAR.

Local distribution

RI (Chassain 1979).

Collecting month and method

Frequent species, which was collected by LT in IV-V and VII-X.

Dicronychus brancuccii Platia & Schimmel, 1997

World distribution

Asia: AE (Platia 2007), IQ, JO, KW, OM, QA, SA.

General distribution

SAR.

Local distribution

BA, EP, MK (Platia and Schimmel 1997).

Collecting month and method

Frequent species that was collected by LT through IV-VI.

Dicronychus latifae Al Dhafer & Platia, 2013

World distribution

Asia: SA.

General distribution

END.

Local distribution

RI (Al Dhafer and Platia 2013).

Collecting month and method

Abundant species that was collected by LT during IV.

Dicronychus talhouki Platia & Schimmel, 1997

World distribution

Asia: SA, YE.

General distribution

SAR.

Local distribution

AS, BA (Platia and Schimmel 1997), RI (Al Dhafer et al. 2016).

Collecting month and method

Rare species, which was collected by PT under canopy of A. gerrardii, and by HP and LT during II-V.

Drasterius aegypticus Buysson, 1905

World distribution

Asia: SA, YE. North Africa: DZ, EG, LY, MA, TN.

General distribution

SAR.

Local distribution

JZ, RI (Platia and Schimmel 1997).

Collecting month and method

Rare species. It was collected by LT through III and V-VI.

Dermestidae

Subfamily: Dermestinae

Dermestes ater DeGeer, 1774

World distribution

Asia: AE, AF, AZ, CH, CN, CY, EG (Sinai), IL, IN, IQ, IR, JO, JP, KG, KP, KR, KZ, LB, MN, NP, OM, PK, RU, SA, SY, TJ, TM, TR, UZ, YE. Europe: AD, AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, ES, FI, FR, GB, GE, GR, HR, HU, IE, IS, IT, LT, LU, LV, MK, MT, NL, NO, PL, PT, RO, RS, RU, SE, SI, SK, UA. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN.

General distribution

COS (Háva 2013).

Local distribution

EP, RI (Mroczkowski 1979).

Collecting month and method

Very rare species. It was collected by LT during III.

Dermestes maculatus De Geer, 1774

World distribution

Asia: AE, AF, AZ, CN, CY, EG (Sinai), IL, IN, IQ, IR, JO, JP, KG, KP, KR, KZ, LB, MN, NP, OM, PK, RU, SA, SY, TJ, TM, TR, UZ, YE. Europe: AD, AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IS, IT, LT, LU, LV, MK, MT, NL, NO, PL, PT, RO, RS, RU, SE, SI, SK, UA. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN.

General distribution

COS (Háva 2003).

Local distribution

MK (Mroczkowski 1979), RI (Al Dhafer et al. 2016).

Collecting month and method

Very rare species. The adult was collected by LT and PT during III and V.

Subfamily: Thorictinae

Thorictodes heydeni Reitter, 1875

World distribution

Africa: SD. Asia: CN, IL, IN, JP, PK, TR. Australia: AU. Europe: ES, FR, GB, IT, RU. North Africa: DZ, EG, MA. North America: CA, MX, US. New to Arabian Peninsula.

General distribution

SCO.

Collecting month and method

Very rare species. The beetles were collected by PT under the canopies of A. ehrenbergiana, A. gerrardii, C. procera and L. shawii during IV.

Thorictus castaneus Germar, 1834

World distribution

Asia: SY. North Africa: DZ, EG, LY, MA. New to Arabian Peninsula.

General distribution

SAR.

Collecting month and method

Very rare species. It was collected by PT under the canopies of A. gerrardii, C. procera and L. shawii during IV-VI.

Thorictus foreli Wasmann, 1894

World distribution

North Africa: DZ, MA, TN. New to Arabian Peninsula.

General distribution

SAR.

Collecting month and method

Very rare. The species was collected by PT under the canopy of A. gerrardii during XII.

Subfamily: Attageninae

Attagenus fasciolatus (Solsky, 1876) 34

World distribution

Asia: KZ, MN, SA, TJ, TM, UZ.

General distribution

PAL_SAR.

Local distribution

EP (Mroczkowski 1979), RI (Al Dhafer et al. 2016).

Collecting month and method

A frequent species. The adults were collected by BV, SW, VC and PT on branches/under the canopies of A. gerrardii, C. procera, R. stricta and Z. nummularia; and by MT through V-VI and VIII-XI.

Attagenus lobatus Rosenhauer, 1856 1

World distribution

Asia: AE, AF, CN, IN, IQ, IR, KG, KZ, MN, PK, SA, TJ, TM, TR, UZ. Europe: BG, CZ, ES, FR, GR, IT, RO, RU. North Africa: DZ, EG, MA, TN. The distribution is updated from Háva (2013).

General distribution

PAL_ORR_SAR_SJP.

Local distribution

EP, MK (Mroczkowski 1979), RI (Al Dhafer et al. 2016).

Collecting month and method

Very rare species. It is collected by PT under canopy of C. procera during V.

Attagenus posticalis Fairmaire, 1879

World distribution

Africa: MR, NE, SD, SN. Asia: AE, IL, OM, QA, SA, SY, YE. Europe: ES. North Africa: DZ, EG, MA, TN. The distribution is updated from Háva (2013).

General distribution

AFR_SAR.

Local distribution

EP (Háva 2011); RI (Alqarni et al. 2015; Al Dhafer et al. 2016).

Collecting month and method

Frequent species. The adults were collected by BV, SW, VC and PT on branches/under canopies of A. ehrenbergiana, A. gerrardii, C. procera, R. stricta and Z. nummularia through IV-X.

Attagenus reitteri (Mroczkowski, 1968)

World distribution

Europe: ES, PT. North Africa: DZ, MA, TN. New to Arabian Peninsula.

General distribution

SAR.

Collecting month and method

Very rare species that was collected by SW on branches of R. stricta during II.

Attagenus scalaris (Pic, 1893)

World distribution

North Africa: EG, LY. New to Arabian Peninsula.

General distribution

SAR.

Collecting month and method

Very rare species. It was collected by SW during VI.

Subfamily: Megatominae

Anthrenus buettikeri Mruczkowski, 1980

World distribution

Asia: SA.

General distribution

END.

Local distribution

RI (Mroczkowski 1980).

Collecting month and method

Very rare species. The specimens were collected by BV on branches and PT canopies of A. gerrardii, R. stricta and Z. nummularia during IV, VI-VII and X.

Anthrenus flavipes LeConte, 1854

World distribution

Asia: AE (Háva 2013), AF, CN, IL, IR, JP, OM, RU, SA, TJ. Europe: CZ, DE, DK, ES, FI, FR, GB, IT, NL. North Africa: EG, MA, TN.

General distribution

COS (Háva 2013).

Local distribution

MK, RI (Mroczkowski 1979).

Collecting month and method

Very rare species that was collected by BV on branches of Z. nummularia in IX.

Anthrenus malkini Mroczkowski, 1980

World distribution

Asia: AE, OM, QA, SA, YE (Háva 2013).

General distribution

SAR.

Local distribution

AS (Mroczkowski 1980).

Collecting month and method

Very rare species. The beetles were collected by BV and SW on branches of Z. nummularia in IV and IX.

Anthrenus verbasci (Linnaeus, 1767)

World distribution

Asia: AE, AF, AZ, CN, CY, EG (Sinai) IL, IN, IQ, IR, JO, JP, KG, KP, KR, KZ, LB, MN, NP, OM, PK, RU, SA, SY, TJ, TM, TR, UZ, YE. Europe: AD, AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IS, IT, LT, LU, LV, MK, MT, NL, NO, PL, PT, RO, RS, RU, SE, SI, SK, UA. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN. COS.

General distribution

PAL_ORR_SAR_SJP.

Local distribution

RI (Alqarni et al. 2015; Al Dhafer et al. 2016).

Collecting month and method

Common species that was collected by BV, VC, SW and PT on branches/under canopies of A. ehrenbergiana, A. gerrardii, C. procera, R. stricta and Z. nummularia; and by LT through IV-X.

Phradonoma nobile (Reitter, 1881)

World distribution

Africa: ER, NA, SD, TJ, ZA, ZW. Asia: AE, AF, CY, IL, IN, IQ, IR, JO, PK, QA, SA, SY, TJ, TM, UZ. Europe: ES, GB, GR, PT. North Africa: DZ, EG, LY, MA, TN. The distribution is updated from Háva (2013).

General distribution

AFR_ORR_PAL_SAR.

Local distribution

AS, RI (Mroczkowski 1979).

Collecting month and method

A frequent species. The specimens were collected by BV, VC, SW and PT on branches/under canopies of A. ehrenbergiana, A. gerrardii, C. procera, R. stricta and Z. nummularia; and by LT through IV-X.

Bostrichidae

Subfamily: Bostrichinae

Enneadesmus forficula (Fairmaire, 1883)

World distribution

Africa: ER, ET, SO. Asia: AE (Geistharde 2010), IL, IR, JO, OM, PK, SA, YE. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_SAR.

Local distribution

MK, RI (Damoiseau 1979).

Collecting month and method

Common species that was collected by BV and LT during all months except II and VII.

Sinoxylon senegalense Karsch, 1881

World distribution

Africa: SN. Asia: AE (Geistharde 2010), SA, YE. North Africa: DZ, EG, LY, MA.

General distribution

AFR_SAR.

Local distribution

RI (Damoiseau 1979).

Collecting month and method

Frequent species that was collected by BV and LT during all months except II, III and VIII.

Subfamily: Lyctinae

Acantholyctus cornifrons (Lesne, 1898)

World distribution

Africa: DJ, ER, MZ, NA, SN, SO. Asia: AE (Geistharde 2010). North Africa: DZ, EG, MA, TN. New to KSA.

General distribution

AFR_SAR.

Collecting month and method

Rare species and its adults were collected by BV and LT during V, IX and X.

Ptinidae

Subfamily: Xyletininae

Lasioderma baudii Schilsky, 1899

World distribution

Asia: CY, IL, LB, SY. Europe: ES, FR, GR, HR, IT, PT. North Africa: DZ, EG, ES (Canary Islands), LY, TN. New to Arabian Peninsula.

General distribution

PAL_SAR.

Collecting month and method

Very rare species, which was collected by LT during IX-X.

Lasioderma redtenbacheri (Bach, 1852)

World distribution

Asia: AZ, CY, EG (Sinai), IL, IQ, KZ, MN, SY, TM, TR, UZ. Europe: AM, AT, BE, BG, CH, CZ, DE, ES, FR, GE, GR, HR, HU, IT, MK, NL, PL, PT, RO, RU, SK, UA. North Africa: EG, LY, TN. New to Arabian Peninsula.

General distribution

PAL_SAR.

Collecting month and method

A rare species. The adult beetles were collected by LT during IV-V and X.

Subfamily: Dorcatominae

Stagetus montanus Toskina, 1998

World distribution

Asia: OM, SA.

General distribution

SAR.

Local distribution

AS (Toskina 1998).

Collecting month and method

Very rare species. It was collected by LT in V.

Thanerocleridae

Subfamily: Thaneroclerinae

Thaneroclerus buquet (Lefebvre, 1835)

World distribution

Asia: CN, IN, JP, RU, SA, TW. Europe: BE, DE, HU, NL, PL. North Africa: EG.

General distribution

ORR_PAL_SAR_SJP.

Local distribution

BA (El-Hawagry et al. 2013), RI (Al Dhafer et al. 2016).

Collecting month and method

A rare species. It was collected by BV on branches and PT under canopies of A. ehrenbergiana, A. gerrardii and C. procera during V-VII.

Cleridae

Subfamily: Tillinae

Eucymatodera senegalensis (Castelanu, 1832)

World distribution

Africa: SN. Asia: AE, IR, OM, SA, YE. North Africa: DZ, EG, LY, TN. The distribution is updated from Gerstmeier (2010)

General distribution

AFR_SAR.

Local distribution

BA, EP, JZ, RI (Menier 1986).

Collecting month and method

Frequent species that was collected by LT during V and IX-X.

Tillodenops plagiatus (Fairmaire, 1892)

World distribution

Africa: KE, MR, SD, SN, SO, TZ. Asia: AE, IR, OM, SA, YE. The distribution is updated from Gerstmeier (2010)

General distribution

AFR_SAR.

Local distribution

AS, BA, JZ, MD, MK (Menier 1986).

Collecting month and method

Very rare species, which was collected by LT during IV and X.

Wittmeridecus mediozonatus (Fairmaire, 1892)

World distribution

Africa: DJ. Asia: AE, IL, OM, SA, SY, YE. Europe: ES, IT. North Africa: DZ, EG, LY, MA. The distribution is updated from Gerstmeier (2010)

General distribution

AFR_PAL_SAR.

Local distribution

AS, MD, MK, RI (Menier 1986).

Collecting month and method

A frequent species. The adults were collected by VC on branches of A. ehrenbergiana and by LT through IV-X.

Subfamily: Korynetinae

Necrobia rufipes (De Geer, 1775)

World distribution

Asia: AE, AZ, CN, IN, IR, JP, MN, OM, RU, SA, TJ, TR. Europe: AM, AT, BE, BY, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GR, HR, HU, IE, IT, LI, LT, LV, NL, PT, RO, RU, SE, SI, SK. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

COS (Gerstmeier 2010)

Local distribution

BA (El-Hawagry et al. 2013), EP, MK, RI (Menier 1986).

Collecting month and method

Frequent species, which was collected by VC on branches of L. shawii and by LT during IV-VIII and X.

Melyridae
Subfamily: Dasytinae

Danacea sp.

Collecting month and method

Frequent species. The specimens were collected by BV, SW and PT on branches/under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by HP and LT during II-V and XII.

Subfamily: Malachinae

Colotes javeti Du Val, 1852

World distribution

Europe: ES, FR, IT, PT. North Africa: DZ, EG, MA, TN. New to Arabian Peninsula.

General distribution

PAL_SAR.

Collecting month and method

A frequent species. The beetles were collected by BV, SW, VC and PT on branches/under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by HP during II-VI and IX.

Malachius sp.1

Collecting month and method

A common species. The adults were collected by BV, SW, VC and PT on branches/under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by HP and LT through V-IX.

Malachius sp.2

Collecting month and method

A rare species. It was collected by BV on branches of A. gerrardii and Z. nummularia, by PT under canopies of A. ehrenbergiana and C. procera; and by HP in III-IV and IX.

Cryptophagidae
Subfamily: Cryptophaginae

Cryptophagus acutangulus Gyllenhal, 1827

World distribution

Africa: sub-saharan Africa. Asia: AF, AZ, CN, EG (Sinai), IR, JP, KG, KP, KZ, PK, RU, SA, TJ, TM, UZ. Europe: AT, BA, BE, BY, CH, CZ, DE, DK, EE, FI, FR, GB, GE, GR, HU, IE, IS, IT, LT, LV, NL, NO, PL, RO, RS, RU, SE, SI, SK, UA. North Africa: EG, MA. North America: CA, MX, US.

General distribution

COS (Otero 2013).

Local distribution

EP (Johnson 1989), RI (Al Dhafer et al. 2016).

Collecting month and method

Frequent species. The adults were collected by BV on branches and PT under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by LT during I-III, V, VIII and XI-XII.

Subfamily: Atomariinae

Atomaria fuscata (Schönherr, 1808)

World distribution

Asia: AZ, CN, IL, JP, RU. Europe: AM, AT, BA, BE, BY, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IT, LI, LT, LV, NL, NO, PL, RO, RU, SE, SI, SK, UA. North America: CA. New to Arabian Peninsula.

General distribution

NAR_PAL_SAR_SJP.

Collecting month and method

Very rare species that was collected by LT in X.

Phalacridae
Subfamily: Phalacrinae

Olibrosoma testacea Tournier, 1889

World distribution

Asia: AE (Švec 2010b), JO, SA. Europe: ES. North Africa: DZ, EG, MA, TN.

General distribution

SAR.

Local distribution

RI (Al Dhafer et al. 2016).

Collecting month and method

Afrequent species. The adults were collected by PT under canoppy of A. ehrenbergiana, and by LT during IV-X.

Laemophloeidae
Subfamily: Laemophloeinae

Placonotus testaceus (Fabricus, 1787)

World distribution

Asia: AZ, BT, CN, IN, JP, KG, KZ, RU, TW, UZ. Australia: AU. Europe: AM, BE, CH, CZ, DE, DK, ES, FR, GE, IT, PL, PT, SK. North Africa: DZ, TN. New to Arabian Peninsula.

General distribution

SCO.

Collecting month and method

Very rare species that was collected by LT during VIII and XI.

Nitidulidae
Subfamily: Carpophilinae

Carpophilus hemipterus (Linnaeus, 1758)

World distribution

Africa: MR, ZA. Asia: AE, AZ, CN, IL, IN, IQ, IR, JO, JP, LB, SA, TM, TR, TW. Europe: AL, AT, BA, BE, BY, CH, CZ, DE, DK, ES, FI, FR, GB, GR, HR, HU, IE, IS, IT, LV, MD, MT, NL, PL, PT, SE, SK. North Africa: DZ, EG, ES (Canary Islands), LY, PT (Madeira Archipelago), TN. North America: GT, MX, NI, PA, US.

General distribution

COS (Jelinek 1979).

Local distribution

EP, MK, RI (Beccari 1971, Jelinek 19791988).

Collecting month and method

Rare species. The adult beetles were collected by PT under canoppy of R. stricta, and by HP and LT during IV-V and XI.

Subfamily: Nitidulinae

Nitidula eremita Audisio, 1990

World distribution

Asia: AE, IL, IQ, IR, SA. North Africa: DZ, EG, LY, TN.

General distribution

SAR.

Local distribution

EP, RI, (Jelinek 19791988).

Collecting month and method

Frequent species that was collected by PT under canoppy of L. shawii during I, and by LT during III and X-XII.

Subfamily: Cybocephalinae

Cybocephalus rufifrons flaviceps Reitter, 1874

World distribution

Asia: IQ, IR, SA, TR. North Africa: DZ, EG, MA, TN.

General distribution

SAR.

Local distribution

MK (Endrödy-Younga 1979), RI (Beccari 1971).

Collecting month and method

A common species. The specimens were collected by BV, SW and VC on branches of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by MT through II, V-IX and XII.

Coccinellidae
Subfamily: Coccinellinae

Bulaea lividula bocandei Mulsant, 1850

World distribution

Africa: ER. Asia: AE, IL, IQ, IR, JO, PK, SA, SY, YE. North Africa: DZ, EG, MA.

General distribution

AFR_SAR.

Local distribution

EP (Fürsch 1979).

Collecting month and method

Very rare species that was collected by BV on branches of Z. nummularia in VIII.

Coccinella septempunctata Linnaeus, 1758

World distribution

Africa: ZA. Asia: AF, AZ, BT, CN, CY, EG (Sinai), IL, IN, IQ, IR, JO, JP, KG, KP, KR, KW, KZ, LB, MN, NP, PK, RU, SA, SY, TJ, TM, TR, TW, UZ. Europe: AD, AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IT, LI, LT, LU, LV, MD, MK, NL, NO, PL, PT, RO, RS, RU, SE, SI, SK, UA. North Africa: PT (Madeira Archipelago). North America: CA, US.

General distribution

SCO.

Local distribution

RI (Fürsch 1979).

Collecting month and method

Very rare species, which was collected by SW during IV.

Coccinella undecimpunctata menetriesi Mulsant, 1850

World distribution

Asia: AE, AF, CN, EG (Sinai), IL, IN, IQ, IR, JO, KG, KW, KZ, MN, PK, RU, SA, SY, TR. Europe: GR, IT, PT, RU. North Africa: DZ, EG, LY, TN.

General distribution

PAL_ORR_SAR_SJP.

Local distribution

RI (Fürsch 1979).

Collecting month and method

A frequent species. The adult beetles were collected by BV, SW, VC and PT on branches/under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia during I-IV and XI.

Diomus rubidus (Motschulsky, 1837)

World distribution

Asia: IL, IQ, IR, LB, SA, SY, YE. Europe: AM, FR, GR, HR, IT. North Africa: DZ, EG, LY, MA, TN.

General distribution

PAL_SAR.

Local distribution

RI (Fürsch 1979).

Collecting month and method

Frequent species. The specimens were collected by BV and SW on branches and PT under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, and R. stricta through I-VII.

Hippodamia variegata (Goeze, 1777)

World distribution

Africa: SZ, ZA. Asia: AE, AF, AZ, BT, CN, EG (Sinai), IL, IN, IQ, IR, JO, KG, KP, KR, KZ, LB, MN, NP, PK, RU, SA, SY, TJ, TM, TR, UZ, YE. Europe: AD, AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IT, LI, LT, LU, LV, MD, MK, NL, PL, PT, RO, RS, RU, SE, SI, SK, UA. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN. North America: US.

General distribution

COS.

Local distribution

BA (El-Hawagry et al. 2013), JZ (Beccari 1971), RI (Talhouk 1982; Al Dhafer et al. 2016).

Collecting month and method

Frequent species. The specimens were collected by BV, SW, VC and PT on branches/ under canopies of A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by LT and MT in I and III-IV.

Hyperaspis vinciguerrae Capra, 1929

World distribution

Africa: GM, SN. Asia: AE (Raimundo et al. 2007), SA, YE. North Africa: LY.

General distribution

AFR_SAR.

Local distribution

RI (Talhouk 1982)

Collecting month and method

Rare species that was collected by BV, SW and VC on branches of C. procera, L. shawii, R. stricta and Z. nummularia; and MT during V-VIII.

Nephus arcuatus Kapur, 1959

World distribution

Africa: TG. Asia: AE (Raimundo et al. 2007), IR, SA, YE.

General distribution

AFR_SAR.

Local distribution

AS (Fürsch 1979).

Collecting month and method

Very rare species, which was collected by VC on branches of Z. nummularia in VII.

Nephus levaillanti (Mulsant, 1850)

World distribution

Africa: ZA. Asia: AF, CN, IL, IN, IR, JO, JP, LB, PK, RU, SA, TW, YE. Europe: GR, IT. North Africa: EG.

General distribution

AFR_ORR_PAL_SAR_SJP.

Local distribution

RI (Al Dhafer et al. 2016).

Collecting month and method

Very rare species. The specimens were collected by BV on branches of A. gerrardii and Z. nummularia; and by PT under canopies of A. ehrenbergiana and A. gerrardii during IV and VII.

Nephus wittmeri Fürsch, 1979

World distribution

Asia: SA, YE.

General distribution

SAR.

Local distribution

RI (Fürsch 1979).

Collecting month and method

Rare species. The adults were collected by BV on branches of A. ehrenbergianaA. gerrardii and Z. nummularia; by PT under canopy of A. ehrenbergiana; and by VC on branches of R. stricta during II-V and XII.

Parexochomus pubescens (Küster, 1848)

World distribution

Africa: Africa. Asia: AE (Raimundo et al. 2007), AF, IL, IN, IQ, IR, SA, SY, YE. Europe: ES, FR, GR, IT. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_ORR_PAL_SAR

Local distribution

AS, RI (Fürsch 1979).

Collecting month and method

Common beetles that were collected by BV and VC on branches of A. ehrenbergianaA. gerrardii, C. procera and L. shawii throughout the year except in VI and VIII.

Scymnus luxorensis Fürsch, 1989

World distribution

Asia: SA. North Africa: EG.

General distribution

SAR.

Local distribution

RI (Fürsch 1989).

Collecting month and method

Very rare species that was collected by SW on branches of R. stricta during XII.

Scymnus nubilus Mulsant, 1850

World distribution

Africa: KE, SZ, UG. Asia: AE (Raimundo et al. 2007), AF, BD, IL, IN, IQ, IR, JO, JP, KW, LB, NP, OM, PK, SA, SY, TR, TW, YE. Australia: AU. Europe: ES, GR, IT, PT. North Africa: EG, ES (Canary Islands), PT (Madeira Archipelago).

General distribution

SCO.

Local distribution

AS (Fürsch 1979; Raimundo et al 2006); RI (Al Dhafer et al. 2016).

Collecting month and method

A rare species, which was collected by PT under canopy of L. shawii, and by LT and MT during V and VII-VIII.

Scymnus subvillosus (Goeze, 1777)

World distribution

Africa: ZA. Asia: AE, AF, AZ, CY, EG (Sinai), IL, IQ, IR, JO, KG, KW, KZ, LB, PK, QA, SA, SY, TJ, TR, UZ, YE. Europe: AD, AL, AM, AT, BA, BG, CH, CZ, DE, ES, FR, GE, GR, HR, HU, IT, MK, PT, RO, RS, RU, SI, SK, UA. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN.

General distribution

AFR_PAL_SAR.

Local distribution

AS, RI (Fürsch 1979).

Collecting month and method

Common species. The adults were collected by BV, SW, VC and PT on branches/under canopies of A. ehrenbergianaA. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia throughout the year except in X.

Scymnus syriacus (Marseul, 1868)

World distribution

Asia: CY, EG (Sinai), IL, IQ, IR, JO, LB, SA, SY. North Africa: EG.

General distribution

SAR.

Local distribution

BA (El-Hawagry et al. 2013), EP (Fürsch 1989).

Collecting month and method

A common species that was collected by BV, SW and VC on branches of A. ehrenbergianaA. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by HP during II-VII and X-XII.

Scymnus yemenensis (Kapur, 1959)

World distribution

Asia: AE (Raimundo et al. 2007), OM, SA, YE.

General distribution

SAR.

Local distribution

AS, RI (Fürsch 1979).

Collecting month and method

A rare species that was collected by BV on branches of A. ehrenbergiana, by SW and VC on branches of C. procera, and by LT during IV and X.

Mycetophagidae
Subfamily: Mycetophaginae

Typhaea stercorea (Linnaeus, 1758)

World distribution

Africa: GM, MR, ZA. Asia: AE, AF, AZ, BT, CN, CY, EG (Sinai), IL, IQ, IR, JO, JP, KG, KR, KZ, MN, NP, PK, RU, SA, SY, TJ, TM, TR, UZ, YE. Europe: AD, AL, AM, AT, BA, BE, BG, BY, CH, CZ, DE, DK, EE, ES, FI, FO, FR, GB, GE, GR, HR, HU, IE, IS, IT, LI, LT, LU, LV, MD, MK, MT, NL, NO, PL, PT, RO, RS, RU, SE, SI, SK, UA. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN. South America: CL.

General distribution

COS.

Local distribution

BA (El-Hawagry et al. 2013).

Collecting month and method

Very rare species. The adults were collected by LT during V.

Tenebrionidae
Subfamily: Lagriinae

Centorus csikii bagdadensis (Reitter, 1920)

World distribution

Asia: CY, IL, IQ, JO, SA, YE.

General distribution

SAR.

Local distribution

EP, RI (Kaszab 1979).

Collecting month and method

Common beetles that were collected by PT under the canopies A. ehrenbergiana, A. gerrardii and Z. nummularia; and by LT throughout the year except in XI.

Subfamily: Pimeliinae

Adelostoma subtile Reitter, 1900

World distribution

Asia: IL, IR, JO, SA, SY, TR.

General distribution

PAL_SAR.

Local distribution

MD, NJ, RI (Kaszab 19811982).

Collecting month and method

A frequent species. The adults were collected by PT under the canopies A. ehrenbergiana, A. gerrardii, R. stricta and Z. nummularia; and by HP through II, IV-V, and VII-X.

Adesmia cancellata (Klug, 1830)

World distribution

Asia: AE, BH, EG (Sinai), IL, IR, IQ, JO, KW, OM, PK, SA, SY, YE. The distribution is updated from Schawaller (2010).

General distribution

SAR.

Local distribution

AS, BA, EP, JZ, MD, MK, NJ, RI (Kaszab 197919811982; El-Hawagry et al. 2013).

Collecting month and method

Very rare species. It was collected by PT under canopy of A. ehrenbergiana and L. shawii in III and V.

Akis spinosa (Linnaeus, 1764)

World distribution

North Africa: EG. New to Arabian Peninsula,

General distribution

SAR.

Collecting month and method

Very rare species that was collected by PT under canopy of L. shawii during VII.

Akis subtricostata Redtenbacher, 1850

World distribution

Asia: AE (Schawaller 2010), IQ, IR, SY. New to KSA.

General distribution

SAR.

Collecting month and method

Very rare species, which was collected by LT in IX.

Ammogiton sonyae Kaszab, 1979

World distribution

Asia: SA.

General distribution

END.

Local distribution

EP, RI (Kaszab 197919811982).

Collecting month and method

Very rare species that was collected by HP during II.

Apentanodes arabicus (Kirchsberg, 1877)

World distribution

Asia: AE, OM, SA.

General distribution

SAR.

Local distribution

AS, EP, HA, MD, MK, QS, RI, (Kaszab 197919811982).

Collecting month and method

Common beetles that were collected by PT under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by HP and LT through I-VI.

Boromorphus saudicus Schawaller, Al Dhafer & Fadl, 2013

World distribution

Asia: SA (Schawaller et al. 2013).

General distribution

END.

Local distribution

RI (Schawaller et al. 2013).

Collecting month and method

A rare beetle. The adults were collected by PT under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii and Z. nummularia; and by HP during II-IV.

Cyphostethe ferruginea (Marseul, 1867)

World distribution

Asia: AE, IL, SA. North Africa: DZ, LY, TN.

General distribution

SAR.

Local distribution

AS, RI (Kaszab 1981).

Collecting month and method

Very rare species that was collected by PT under canopy of C. procera in X.

Cyphostethe wittmeri Kaszab, 1979

World distribution

Asia: AE, SA.

General distribution

SAR.

Local distribution

EP, HA, MK, RI (Kaszab 197919811982).

Collecting month and method

A rare species. The specimens were collected by LT through V-IX.

Erodius glabratus Solier, 1834

World distribution

Asia: EG (Sinai), SA. North Africa: EG.

General distribution

SAR.

Local distribution

AS, EP, HA, JF, JZ, MD, MK, QS, TB, (Kaszab 197919811982).

Collecting month and method

Very rare species that was collected by LT in IV.

Erodius octocostatus Peyerimhoff, 1907

World distribution

Asia: EG (Sinai), IQ, JO, SA. North Africa: EG.

General distribution

SAR.

Local distribution

ES, NJ (Kaszab 1981).

Collecting month and method

Very rare species. The beetles were collected by HP during II and XII.

Erodius servillei Solier, 1834

World distribution

Asia: AF, IQ, IR, SA, SY.

General distribution

SAR.

Local distribution

MK, QS, RI (Kaszab 19791981)

Collecting month and method

Very rare species that was collected by PT under canopy of R. stricta through III.

Mesostena angustata (Fabricius, 1775)

World distribution

Africa: ER, NG, SD. Asia: EG (Sinai), IL, IQ, IR, JO, LB, SA (Al Dhafer et al. 2016), SY. North Africa: DZ, EG, LY, TN.

General distribution

AFR_SAR.

Local distribution

RI (Al Dhafer et al. 2016).

Collecting month and method

Very rare species. The specimens were collected by PT under canopy of A. gerrardii through V; and by HP in V and XII.

Mesostena puncticollis Solier, 1835

World distribution

Africa: SD, SO. Asia: AE, EG (Sinai), IL, IQ, IR, JO, SA, SY, TM, YE. Europe: AM. North Africa: EG.

General distribution

AFR_PAL_SAR.

Local distribution

AS, EP, HA, MD, MK, NJ, QS, RI, TB (Kaszab 197919811982).

Collecting month and method

An abundant species. The adults of this beetle were collected by HP, LT and PT all over the year except in XI.

Oxycara saudarabica Kaszab, 1979

World distribution

Asia: AE, SA.

General distribution

SAR.

Local distribution

EP, RI (Kaszab 197919811982).

Collecting month and method

Common species that was collected by HP, LT and PT throughout the year.

Paraplatyope arabica arabica (Blair, 1931)

World distribution

Asia: AE, KW, SA.

General distribution

SAR.

Local distribution

EP, MK, RI (Kaszab 19791982).

Collecting month and method

Very rare species that was collected by LT during III.

Pimelia arabica (Klug, 1830)

World distribution

Asia: AE, BH, IL, IQ, JO, KW, OM, PK, SA, SY, YE. North Africa: EG.

General distribution

SAR.

Local distribution

AS, BA, EP, HA, MD, MK, RI (Kaszab 19791982)

Collecting month and method

A rare species. The beetles were collected by HP and PT in X-XII.

Pimelia longula Kwieton, 1981

World distribution

Asia: BH, SA.

General distribution

SAR.

Local distribution

EP, MD, RI, TB (Kaszab 19791982).

Collecting month and method

Very rare species that was collected by HP during I.

Prionotheca coronata ovalis Ancey, 1881

World distribution

Asia: AE, BH, IQ, IR, SA, YE.

General distribution

SAR.

Local distribution

AS, EP, HA, MD, MK, QS, RI (Kaszab 19791982).

Collecting month and method

A rare species. This beetle was collected by HP, LT and PT during II, IV, VI and X.

Thriptera kraatzi Haag-Rutenberg, 1876

World distribution

Asia: AE, EG (Sinai), IN, IR, JO, OM, PK, SA, YE. North Africa: AG.

General distribution

ORR_SAR.

Local distribution

AS, BA, MK, RI (Kaszab 19791982; El-Hawagry et al. 2013).

Collecting month and method

A frequent species. The specimens were collected by HP and PT during III-VI and VIII.

Trachyderma philistina Reiche & Saulcy, 1857

World distribution

Asia: AE, BH, CY, EG (Sinai), IL, IQ, IR, JO, OM, SA, SY, TR, YE. Europe: GR. North Africa: EG. According to the Palaearctic catalogue Trachyderma philistina occurs east of the Arabian Peninsula also in Iran and in the ORR region (Löbl et al. 2008). However, the known distribution of species indicates its absence from Afghanistan, Pakistan, and India. The only record available from western India apparently is by Kaszab (1982). This record is a misidentification and Kaszab (1982) did not specifically mention this species from Afghanistan (W. Schawaller, personal communication, February 29 2016).

General distribution

PAL_SAR.

Local distribution

EP, HA, MK, NJ, QS, RI, TB (Kaszab 19791982)

Collecting month and method

Rare species that was collected by HP and PT through III-VI, X and XII.

Trichosphaena arabica Kaszab, 1961

World distribution

Asia: AE, OM, SA, YE.

General distribution

SAR.

Local distribution

HA, JZ, MK, NJ, QS, RI (Kaszab 197919811982).

Collecting month and method

A frequent species. The adults of this species were collected by BV on branches of A. ehrenbergiana and Z. nummularia, and by HP during IV and VII-IX.

Zophosis punctata Brullé, 1832

World distribution

Asia: AF, AZ, CN, CY, EG (Sinai), IL, IQ, IR, JO, KZ, LB, PK, SA, SY, TJ, TM, TR, UZ. Europe: AL, AM, ES, GR, IT. North Africa: DZ, LY, MA, TN.

General distribution

PAL_SAR_SJP.

Local distribution

MK, QS (Kaszab 19811982), RI (Al Dhafer et al. 2016)

Collecting month and method

Frequent species. The adults were collected by HP and PT through II-IV and VI-X.

Subfamily: Tenebrioninae

Alphitobius laevigatus (Fabricius, 1781)

World distribution

Africa: GH. Asia: AE, AF, BH, BT, CN, CY, IN, IQ, IR, JP, KZ, RU, SA, TW, YE. Europe: AM, BE, CH, DE, ES, GB, GR, HU, IT, LU, MT, NL, PL, PT, RU, SK, UA. North Africa: EG, ES (Canary Islands), LY, PT (Madeira Archipelago), TN. North America: MX, US.

General distribution

COS (Schawaller 2010).

Local distribution

EP, MD, MK, RI (Kaszab 19791982).

Collecting month and method

Very rare species that was collected by LT during X.

Blaps kollari Seidlitz, 1893

World distribution

Asia: AE, IQ, JO, OM, SA, YE. North Africa: EG.

General distribution

SAR.

Local distribution

AS, BA, EP, HA, MD, MK, QS, RI (Kaszab 197919811982; El-Hawagry et al. 2013).

Collecting month and method

A frequent species. The adult beetles were collected by HP, LT and PT during I-II, V-VI and X-XII.

Cheirodes brevicollis (Wollaston, 1864)

World distribution

Africa: SO. Asia: AE, AF, BH, CN, EG (Sinai), IL, IQ, IR, JO, KZ, MN, OM, PK, SA, TM, UZ, YE. Europe: ES, IT, MT. North Africa: DZ, EG, ES (Canary Isalands), LY, MA, TN.

General distribution

AFR_PAL_SAR_SJP.

Local distribution

AS, EP, HA, JZ, MD, MK, RI (Kaszab 19791982). It is reported as Anemia brevicollis (Wollaston 1864) from BA (El-Hawagry et al. 2013).

Collecting month and method

Common species. The specimens were collected by LT and PT through IV-X.

Cheirodes pilosus (Tournier, 1868)

World distribution

Africa: SD, TD. Asia: AE (Schawaller 2010), EG (Sinai), IL, OM, SA, YE. North Africa: DZ, EG, ES (Canary Isalands), LY, MA, TN.

General distribution

AFR_SAR.

Local distribution

EP, HA, MD, MK, QS, RI (Kaszab 19791982).

Collecting month and method

A rare species. The adults were collected by BV and SW on branches of L. shawii and C. procera, respectively during VII; and by LT in V-VI.

Cheirodes sardous (Gené, 1839)

World distribution

Asia: AE, AZ, CY, EG (Sinai), IL, IQ, IR, JO, SA, TR. Australia: AU. Europe: AM, ES, FR, GR, IT, PT. North Africa: EG, ES (Canary Isalands), LY, MA, TN.

General distribution

AUS_PAL_SAR.

Local distribution

AS, EP, HA, MD, MK, NJ, RI, TB (Kaszab 19791982).

Collecting month and method

Very rare species that was collected by LT during VII.

Gonocephalum prolixum (Erichson, 1843)

World distribution

Africa: ZA. Asia: AE, CY, IR, SA, SY. Europe: ES, IT. North Africa: DZ, EG, ES (Canary Isalands), LY, MA, TN.

General distribution

AFR_PAL_SAR.

Local distribution

AS, MD, MK (Kaszab 19791982), RI (Al Dhafer et al. 2016).

Collecting month and method

An abundant species. The beetles were collected by HP, LT and PT throughout the year.

Gonocephalum rusticum (Olivier, 1811)

World distribution

Asia: AF, AZ, CN, EG (Sinai), IL, IQ, IR, KZ, MN, OM, RU, SA, TJ, TM, TR, UZ. Europe: AL, AM, ES, FR, GR, HR, IT, MT, MK, PT, RU, UA. North Africa: EG, ES (Canary Isalands), LY, MA, PT (Madeira Archipelago), TN.

General distribution

PAL_SAR_SJP.

Local distribution

AS, EP, MK, RI (Kaszab 19791982).

Collecting month and method

Common species that was collected by HP, LT and PT all over the year.

Gonocephalum setulosum (Faldermann, 1837)

World distribution

Africa: ZA. Asia: AE (Schawaller 2010), AF, CN, CY, EG (Sinai), IL, IQ, IR, KZ, LB, SA, SY, TJ, TM, TR, UZ, YE. Europe: AM, ES, GR, IT, MT, RU. North Africa: EG, ES (Canary Isalands), LY, MA, TN.

General distribution

AFR_PAL_SAR_SJP.

Local distribution

EP, HA, MD, MK, QS, RI, TB (Kaszab 19791982).

Collecting month and method

Common species that was collected by HP, LT and PT through II, IV-X and XII.

Gonocephalum soricinum (Reiche & Saulcy, 1857)

World distribution

Africa: ET. Asia: AE (Schawaller 2010), AF, EG (Sinai), IL, IR, JO, PK, SA, SY, YE.

General distribution

AFR_SAR.

Local distribution

AS, BA, HA, MK, NJ, RI (Kaszab 19791982).

Collecting month and method

Rare species that was collected by HP, LT and PT in I, V and IX-X.

Opatroides vicinus (Fairmaire, 1896)

World distribution

Asia: AE, AF, BH, IN, IR, KW, NP, OM, PK, SA, YE.

General distribution

ORR_SAR.

Local distribution

RI (Kaszab 19791982).

Collecting month and method

Abundant species. The specimens of this species were collected by BV on branches of A. gerrardii in IV; and by HP, LT and PT throughout the year except in IX and XI.

Palorus ficicola (Wollaston, 1867)

World distribution

Africa: CV, GM. Asia: AE (Schawaller 2010), LK, PK, SA. North Africa: DZ, EG, LY, MA.

General distribution

AFR_ORR_SAR.

Local distribution

MK (Kaszab 19791982).

Collecting month and method

Very rare species, which was collected by LT during VIII.

Praeugena gagatina (Mäklin, 1863)

World distribution

Africa: DJ, ER, ET, SD, SN, SO, TD. Asia: AE (Schawaller 2010), SA, YE.

General distribution

AFR_SAR.

Local distribution

AS, JZ, MD, MK, RI (Kaszab 19791982).

Collecting month and method

Rare species. The adults were collected by PT under canopy of L. shawii during V; and y LT in IV-V.

Prodilamus fausti major Kaszab, 1982

World distribution

Asia: SA.

General distribution

END.

Local distribution

MD, QS (Kaszab 1982).

Collecting month and method

Very rare species that was collected by HP during XII.

Sclerum carinatum Baudi 1875

World distribution

Asia: AF, AZ, CY, IQ, IR, SA, SY, TJ, TM, TR, UZ.

General distribution

PAL_SAR.

Local distribution

RI (Kaszab 1979).

Collecting month and method

Rare species that was collected by HP and LT during II-III, VII and IX.

Sclerum orientale (Fabricius, 1775)

World distribution

Africa: SD. Asia: IL, JO, SA, SI, YE. North Africa: EG.

General distribution

AFR_SAR.

Local distribution

RI (Kaszab 19791982).

Collecting month and method

A common species. The adult beetles were collected by BV on branches of A. ehrenbergiana during X; and by HP and PT all over the year except in VI and IX.

Sclerum sulcatum Baudi, 1876

World distribution

Asia: SA. North Africa: EG.

General distribution

SAR.

Local distribution

HA, MK, RI (Kaszab 19791982; Johnson 1989).

Collecting month and method

A frequent species. The specimens were collected by HP, LT and PT during I-VII, X and XII.

Subfamily: Alluculinae

Cornucistela serrata Campbell, 1980

World distribution

Asia: SA.

General distribution

END.

Local distribution

RI (Campbell 1980).

Collecting month and method

Rare species that was collected by LT during V-VI and VIII-X.

Cteniopus pallidus (Küster, 1850)

World distribution

Asia: CY, IQ, SY, TR. New to Arabian Peninsula.

General distribution

SAR.

Collecting month and method

Very rare species that was collected by SW during IV.

Hymenalia denticulata (Muche, 1982)

World distribution

Asia: AE, OM, SA.

General distribution

SAR.

Local distribution

AS, BA, MD, MK (Muche 1982).

Collecting month and method

Frequent species. The adults were collected by LT during IV-VI, IX and XI.

Mycetocharina bahukalatensis Novak, 2008

World distribution

Asia: IR. New to Arabian Peninsula

General distribution

SAR.

Collecting month and method

A frequent species that was collected by LT in IV-X.

Mycetocharina braaschi Muche, 1982

World distribution

Asia: AE, SA.

General distribution

SAR.

Local distribution

MK, RI (Muche 1982).

Collecting month and method

Common species. The specimens were collected by LT in IV-VI and X.

Oedemeridae
Subfamily: Oedemerinae

Alloxantha talhouki Švihla, 1984

World distribution

Asia: AE, IR, OM, SA.

General distribution

SAR.

Local distribution

EP (Švihla 1984).

Collecting month and method

Very rare species. The specimens were collected by LT during V-VI.

Meloidae
Subfamily: Meloinae

Lydomorphus angusticollis suturellus (Haag-Rutenberg, 1880)

World distribution

Asia: AE, IR, OM, PK, SA, YE.

General distribution

SAR.

Local distribution

EP, MD, MK, RI, (Kaszab 1983, Schneider 1991).

Collecting month and method

Very rare species that was collected by LT through IV.

Lydomorphus brittoni (Kaszab 1953)

World distribution

Asia: AE, IQ, OM, SA.

General distribution

SAR.

Local distribution

EP, MD, MK, QS, RI, (Kaszab 1983, Schneider 1991).

Collecting month and method

Frequent species, which was collected by LT during IV.

Lydomorphus palaestinus (Kirsch, 1871)

World distribution

Asia: EG (Sinai), IL, JO, SA. North Africa: DZ, EG, LY, MA, TN.

General distribution

SAR.

Local distribution

AS, EP, RI (Kaszab 1983, Schneider 1991).

Collecting month and method

Very rare species that was collected by LT during III.

Subfamily: Nemognathinae

Zonitoschema rubricolor Pic, 1924

World distribution

Africa: CD. Asia: AE (Batelka and Geisthardt 2009), IL, SA.

General distribution

AFR_SAR.

Local distribution

BA, RI (Kaszab 1983, Schneider 1991; El-Hawagry et al. 2013).

Collecting month and method

Very rare species. The adults were collected by LT in VI.

Anthicidae
Subfamily: Anthicinae

Anthelephila caeruleipennis (Laferte-Senectere, 1847)

World distribution

Africa: ET, ZA. Asia: AE, EG (Sinai), IL, IQ, IR, JO, LB, OM, PK, SA, SY, YE. Europe: ES, IT. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

AFR_SAR.

Local distribution

AS, BA, JZ, MD, MK, QS (Uhmann 1998; El-Hawagry et al. 2013).

Collecting month and method

Very rare species that was collected by LT during IV and VI.

Anthelephila multiformis Kejval, 2002

World distribution

Asia: AE (Telnov 2007), IR, OM, PK. New to KSA.

General distribution

SAR.

Collecting month and method

Rare species. The specimens of this species were collected by BV, SW, VC and PT on branches/under canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia; and by LT through IV-VI.

Anthicus crinitus Laferte-Senectere, 1849

World distribution

Africa: CF, GM, KE, MR, NA, SL, SN, SZ, TD, ZA. Asia: AE, AF, AZ, CN, CY, EG (Sinai), IL, IN, IQ, IR, JO, JP, KW, NP, OM, PK, SA, SY, TH, TR, TW, UZ, YE. Europe: AL, AM, BG, GR, MT, PT, RU. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN. North America: CU, DO, PR. South America: VE.

General distribution

COS.

Local distribution

AS, MK, QS, (Uhmann 1998), BA (El-Hawagry et al. 2013), RI (Al Dhafer et al. 2016).

Collecting month and method

Rare species. The adults were collected by PT under canopies of A. ehrenbergiana; and by LT during V-VII and IX-X.

Anthicus tristis Schmidt, 1842

World distribution

Asia: AF, AZ, CY, IL, IQ, IR, JO, KZ, LB, SA, SY, TJ, TM, TR, UZ, YE. Europe: AM, BG, ES, FR, GE, GR, HR, IT, MT, RO, RU, SE, SK, UA. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

PAL_SAR.

Local distribution

RI (Uhmann 1992).

Collecting month and method

Very rare species that was collected by LT in IX.

Endomia lefebvrei (LaFerté-Sénectěre, 1849)

World distribution

Africa: GM, TD. Asia: AE, AF, CY, EG (Sinai), IL, IQ, IR, JO, OM, QA, SA, TM, TR, YE. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_PAL_SAR.

Local distribution

BA (El-Hawagry et al. 2013), JZ, MK (Uhmann 1998), RI (Al Dhafer et al. 2016).

Collecting month and method

A rare species. The beetles were collected by PT under canopies of A. gerrardii, C. procera and R. stricta; and by LT during IV-VI.

Omonadus floralis (Linnaeus, 1758)

World distribution

Africa: CM, MR, SN, ZA. Asia: AE (Telnov 2007), AF, AZ, BT, CN, CY, EG (Sinai), IL, IN, IR, JO, JP, LB, MN, NP, OM, PK, RU, SA, SY, TM, TR. Australia: PG. Europe: AL, AR, AT, BA, BE, BG, BY, CH, DE, DK, EE, ES, FI, FR, GB, GE, GR, HR, HU, IE, IT, LI, LT, LV, MD, MK, MT, NL, NO, PL, PT, RO, RS, RU, SE, SK, TR, UA. North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN. North America: CA, US.

General distribution

COS (Uhmann 1998, Telnov 2007).

Local distribution

BA, MK (Uhmann 1998), RI (Alqarni et al. 2015; Al Dhafer et al. 2016).

Collecting month and method

Very rare species that was collected by PT under canopies of Z. nummularia, and by LT during V-VI.

Subfamily: Notoxinae

Mecynotarsus bison (Olivier, 1811)

World distribution

Africa: CG, CI, CV, ET, SD, SO, TD, TZ. Asia: AE, CY, EG (Sinai), IL, IQ, IR, LB, SA, TR, YE. Europe: GR. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

AFR_PAL_SAR.

Local distribution

EP, JZ, MD, MK, NB, RI, TB (Uhmann 1998).

Collecting month and method

Very rare species that was collected by PT under canopy of R. stricta in IV.

Scraptiidae
Subfamily: Anaspidinae

Pentaria arabica Pankow, 1981

World distribution

Asia: SA.

General distribution

END.

Local distribution

AS (Pankow 1981).

Collecting month and method

A frequent species. It was collected by BV and PT on branches/under canopy of A. ehrenbergiana, and by LT and MT through IV-V, VII and X.

Pentaria sp.

Collecting month and method

Rare species. The specimens were collected by BV on branches of Z. nummularia in V, by PT under canopy of L. shawii in V, and by LT during V-VII and IX.

Cerambycidae
Subfamily: Prioninae

Polyarthron philbyi Villiers, 1968

World distribution

Asia: SA.

General distribution

END.

Local distribution

RI (Holzschuh 1993).

Collecting month and method

Very rare species. It was collected by HP during IX.

Subfamily: Lamiinae

Apomecyna lameerei (Pic, 1895)

World distribution

Africa: MR. Asia: AE, IL, IQ, IR, PK, SA. North Africa: EG.

General distribution

AFR_SAR.

Local distribution

MD, MK, RI (Holzschuh and Téocchi 1991).

Collecting month and method

Rare species. The adults were collected by HP and LT during IV-V, X-XI.

Chrysomelidae
Subfamily: Bruchinae

Bruchidius centromaculatus (Allard, 1868)

World distribution

Africa: BF, CD, MR, SD, SN. Asia: IL, SA, YE. North Africa: EG.

General distribution

AFR_SAR.

Local distribution

AS (Anton 1994), RI (Al Dhafer et al. 2016).

Collecting month and method

Common species, which was collected by BV, SW and VC on branches of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia, and by HP, LT and PT throughout the year.

Careydon acaciae (Gyllenhall, 1833)

World distribution

Africa: AO, BF, CD, CM. ET, KE, MR, MZ, NA, NE, NG, RW, SD, SN, SO, SZ, TD, TZ, ZA. Asia: IR, JO, SA, YE. Europe: ES, HU. North Africa: EG.

General distribution

AFR_SAR.

Local distribution

AS, MK (Decelle 1979), RI (Al Dhafer et al. 2016)

Collecting month and method

Common species. The specimens were collected by PT under the canopies of A. ehrenbergiana, A. gerrardii, C. procera and L. shawii; and by SW on branches of C. procera and R. stricta; and by HP, LT and MT throughout the year.

Spermophagus sericeus (Geoffory, 1785)

World distribution

Asia: AF, AZ, CN, CY, IL, IQ, IR, JO, KG, KZ, LB, MN, SA, SY, TJ, TM, TR, UZ. Europe: AL, AM, AT, BA, BE, BG, CH, CZ, DE, DK, ES, FI, FR, GB, GE, GR, HR, HU, IT, LU, MK, MT, NL, PL, PT, RO, RS, RU, SE, SI, SK, UA. North Africa: DZ, MA, TN.

General distribution

PAL_SAR_SJP.

Local distribution

RI (Anton 1994).

Collecting month and method

Very rare that species was collected by SW on branches of C. procera during VIII.

Subfamily: Chrysomelinae

Colaphellus apicalis (Ménétriés, 1849)

World distribution

Asia: AF, IL, IR, KG, KZ, SY, TM, TR, UZ. New to Arabian Peninsula.

General distribution

PAL_SAR.

Collecting month and method

Very rare species. It was collected by PT under the canopy of R. stricta during II.

Subfamily: Galerucinae

Phyllotreta lativittata Kutschera, 1860

World distribution

Asia: AF, AZ, CN, CY, IL, IQ, IR, JO, KG, KZ. LB, OM, RU, SA (Al Dhafer et al. 2016), SY, TJ, TR, TM, UZ. Europe: AM, GR, IT, MT.

General distribution

PAL_SAR_SJP.

Local distribution

RI (Al Dhafer et al. 2016).

Collecting month and method

Common species. The adult beetles were collected by BV, SW, VC and PT on branches/under the canopies of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii. R. stricta and Z. nummularia; and by LT and MT throughout the year.

Psylliodes peyerimhoffi Heikertinger, 1916

World distribution

Asia: AE, EG (Sinai), SA.

General distribution

SAR.

Local distribution

RI (Doguet 1979; Medvedev 1996).

Collecting month and method

Very rare species that was collected by LT during IV.

Subfamily: Cryptocephalinae

Aetheomorpha seminigra pumilio (Lacordaire 1848)

World distribution

Africa: ET, SD. Asia: AF, JO, OM, SA, YE. North Africa: EG.

General distribution

AFR_SAR.

Local distribution

AS, BA, JZ, MK (Medvedev 19931996; El-Hawagry et al. 2013).

Collecting month and method

Very rare species, which was collected by BV on branches of A. gerrardii during V.

Subfamily: Eumolpinae

Macrocoma lefevrei (Baly, 1878)

World distribution

Asia: IR, OM, SA. North Africa: EG.

General distribution

SAR.

Local distribution

MK (Medvedev 1996).

Collecting month and method

Very rare species. It was collected by LT during IV.

Brentidae
Subfamily: Apioninae

Aplemonus arabicus (Wagner, 1909)

World distribution

Africa: MW, ZA. Asia: AE (Magnano et al. 2009), IR, OM, QA (Magnano et al. 2009), SA (Al Dhafer et al. 2016). North Africa: EG.

General distribution

AFR_SAR.

Local distribution

RI (Al Dhafer et al. 2016)

Collecting month and method

Common species. The beetles of this species were collected by BV, SW and VC on branches of A. ehrenbergiana, A. gerrardii, L. shawii and Z. nummularia; by PT under canopies of C. procera; and by LT throughout the year.

Subfamily: Nanophyinae

Allomalia quadrivirgata (A. Costa, 1863)

World distribution

Asia: AZ, CY, IL, KZ, SA (Abdel-Dayem et al. 2015), SY, UZ. Europe: BG, ES, FR, GE, GR, HR, HU, IT, UA. North Africa: DZ, EG, LY, MA, TN.

General distribution

PAL_SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Very rare species that was collected by BV on branches of Z. nummularia in IV.

Curculionidae
Subfamily: Curculioninae

Assuanensius discoidalis (Tournier, 1873)

World distribution

Africa: TD. Asia: IL, SA (Abdel-Dayem et al. 2015). North Africa: DZ, EG, LY, MA.

General distribution

AFR_SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Common species. The specimens were collected by BV on branches of A. ehrenbergiana in III-V.

Assuanensius erectesetosus (Peyerimhoff, 1948)

World distribution

Africa: TD. Asia: IL, SA (Abdel-Dayem et al. 2015). North Africa: EG, LY, MA.

General distribution

AFR_SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Very rare species. The beetles were collected by BV on branches of A. ehrenbergiana, A. gerrardii, C. procera, L. shawii, R. stricta and Z. nummularia during I-VII and IX; by PT under canopies of C. procera and R. stricta in III; and by HP and SW in II and III respectivelly.

Assuanensius peyerimhoffi (Hoffmann, 1963)

World distribution

Africa: TD. Asia: SA (Abdel-Dayem et al. 2015). North Africa: DZ, LY.

General distribution

AFR_SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

A frequent species. The specimens were collected by BV on branches of A. ehrenbergiana, A. gerrardii and Z. nummularia during I-III, V, VIII, and IX-XII.

Mecinus longulus (Desbrochers des Loges, 1893)

World distribution

Asia: IL, SA (Abdel-Dayem et al. 2015). North Africa: DZ, EG, LY, TN.

General distribution

SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Very rare species that was collected by BV on branches of L. shawii and by PT under canopy of C. stricta during III.

Pachytychius cognatus Caldara, 2000

World distribution

Africa: AO, DJ, SD, SN. Asia: SA, YE.

General distribution

AFR_SAR.

Local distribution

JZ (Caldara 2000).

Collecting month and method

Very rare species, which was collected by PT under canopy of C. stricta in XII.

Pseudorchestes letourneuxi (Pic, 1901)

World distribution

Africa: SD. Asia: SA (Abdel-Dayem et al. 2015), TR. North Africa: DZ, EG, LY, MA, TN.

General distribution

AFR_SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Very rare species that was collected by PT under canopy of Z. nummularia during IV.

Sharpia sabulicola Colonnelli, 2009

World distribution

Asia: AE (Magnano et al. 2009), SA (Abdel-Dayem et al. 2015).

General distribution

SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Very rare species and its adult was collected by LT during IV.

Sharpia soluta Faust, 1885

World distribution

Asia: AE (Magnano et al. 2009), AZ, CY, SA (Abdel-Dayem et al. 2015), TM, UZ. Europe: RO.

General distribution

PAL_SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Rare species. It was collected by PT under canopies of A. ehrenbergiana, C. procera, R. stricta, and Z. nummularia during III-IV, X and XII; also collected by LT during V.

Sphincticraerus bruleriei (Desbrochers des Loges, 1873)

World distribution

Asia: IL, SA (Abdel-Dayem et al. 2015). North Africa: EG.

General distribution

SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

A rare species. The adult beetles were collected by BV on branches of Z. nummularia during II and V; and by LT through X.

Tychius banfii Caldara & Fremuth, 1992

World distribution

Asia: IQ, IR, JO, SA, TR. North Africa: EG.

General distribution

PAL_SAR.

Local distribution

EP, RI (Caldara 1993).

Collecting month and method

Rare species. Its adult was collected by BV on branches of A. ehrenbergiana, A. gerrardii and Z. nummularia through XII; by PT under canopy of A. ehrenbergiana in XI; and by LT and MT during V and VII respectivelly.

Tychius mozabitus Pic, 1898

World distribution

Asia: EG (Sinai), IQ, IR, JO, SA. North Africa: DZ, EG, LY, TN.

General distribution

SAR.

Local distribution

MD, RI (Caldara 1993).

Collecting month and method

Very rare species that was collected by PT under canopy of C. procers in II.

Tychius vicinus Roudier, 1954

World distribution

Asia: AE (Magnano et al. 2009), SA, YE. North Africa: DZ, EG.

General distribution

SAR.

Local distribution

AS, RI (Caldara 1993).

Collecting month and method

A frequent species. The adults were collected by BV, SW and VC on branches of A. gerrardii, R. stricta and Z. nummularia during III-V; and by PT under canopy of A. ehrenbergiana during I-VI and XII.

Subfamily: Entiminae

Myllocerus sp.

Collecting month and method

A rare species. It was collected by PT under canopies of R. stricta and Z. nummularia during I-II; and by HP and LT in I, IV and XII.

Tanymecus musculus Fåhraeus, 1840

World distribution

Asia: IL, IQ, SA (Abdel-Dayem et al. 2015), SY. North Africa: EG.

General distribution

SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Very rare species that was collected by LT in I, IV and XII.

Subfamily: Hyperinae

Brachypera isabellina (Boheman, 1834)

World distribution

Asia: AE (Magnano et al. 2009), IL, IQ, IR, JO, KW, QA, SA, SY, TR. Europe: IT (Sicilia). North Africa: DZ, EG, ES (Canary Islands), LY, MA, PT (Madeira Archipelago), TN.

General distribution

SAR.

Local distribution

Heyden (1913) just mentioned Arabia without any further detail about locality. Abdel-Dayem et al. (2015) have been confirmed the occurrence of the species in KSA: RI.

Collecting month and method

A rare species. The beetles were collected by PT under canopies of C. procera, R. stricta and Z. nummularia during II, VIII and XII.

Hypera brunnipennis (Boheman, 1834)

World distribution

Asia: IL, IR, LB, SA (Abdel-Dayem et al. 2015). North Africa: EG. North America: US.

General distribution

NAR_SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Very rare species that was collected by LT during IV.

Subfamily: Lixinae

Hypolixus pica (Fabricius, 1798)

World distribution

Africa: CG, ET, NE, SN, TD, TG. Asia: AE, CY, ID, IQ, IR, JO, PK, SA (Abdel-Dayem et al. 2015), SY, TR, YE (Socotra). Europe: FR. North Africa: EG.

General distribution

AFR_PAL_SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Very rare species that was collected by BV on branches of A. ehrenbergiana in XI.

Larinus elegans Desbrochers des Loges, 1897

World distribution

Asia: SA. North Africa: DZ, EG, LY, MA.

General distribution

SAR.

Local distribution

There is no available information.

Collecting month and method

Very rare species. The adult of this species was collected by BV on branches of Z. nummularia through V.

Lixus ? subfarinosus Desbrochers des Loges, 1893

World distribution

Asia: AE, IQ, SA (Abdel-Dayem et al. 2015), TR, YE. North Africa: EG. TN.

General distribution

SAR.

Local distribution

RI (Abdel-Dayem et al. 2015).

Collecting month and method

Very rare species that was collected by HP in IV.

Pycnodactylopsis tomentosa (Fåhraeus, 1842)

World distribution

Africa: ET, MR, SD, TD. Asia: EG (Sinai), IL, IN, IQ, IR, JO, PK, SA, YE. North Africa: DZ, EG, ES (Canary Islands), LY, MA, TN.

General distribution

AFR_ORR_SAR.

Local distribution

RI (Shalaby 1961, Beccari 1971).

Collecting month and method

Very rare species that was collected by HP in II.

Acknowledgments

We would like to thank the following specialists for their help in identification or confirmation of the identification: David Král (Charles University in Prague, Faculty of Science, Department of Zoology, Czech Republic), Enzo Colonnelli, (Via delle Giunchiglie, 56 00172 Roma, Italy), Giuseppe Platia (Via Molino Vecchio, Gatteo (FC), Italy), Jan Bezděk (Mendel University, Department of Zoology, Zemědělská, Czech Republic); Lev Medvedev (Institute of Animal Morphology and Ecology, Leninsky Prospect, Moscow, Russia), Ron Felix (Naturalis Biodiversity Center Leiden, The Netherland), Tomas Lackner (Munich State Museum, Munich, Germany), and Wolfgang Schawaller (Staatliches Museum für Naturkunde, Stuttgart, Germany). The authors would like to extend their sincere appreciation to the Deanship of Scientific Research at King Saud University for its funding this research group NO (RGP-1437-009).

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