Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.).