Female morphology and larval ecology of Phytoliriomyza marchantiae sp. nov. A, B female postabdomen (at Renge-onsen) A oviscape and spermatheca B tergite 10 C–E habitat and mined thalli on Marchantia paleacea paleacea C habitat at type locality D, E mined thalli at Yashajin-toge and Renge-onsen F–H habitat and mined thalli on M. p. diptera F habitat at Kuchisakamoto G, H mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia.