Corresponding author: Jan Růžička (
Academic editor: B. Guéorguiev
The conundrum of
Wang C-B, Růžička J, Perreau M, Nishikawa M, Park S-J (2016) Revision of the genus
In the fauna of the Russian Far East, only one species in the subgenus
However, when we examined specimens previously identified as
Specimens were relaxed and softened in a hot saturated solution of potassium hydroxide for 4 minutes (for mounted dry specimens) or 8 minutes (for alcohol-preserved specimens), and then transferred to distilled water to rinse the residual potassium hydroxide off and stop any further bleaching. The softened specimens were moved into glycerin and dissected there to observe morphological details. After examination, the body parts were mounted on a glass slip with Euparal Mounting Medium for future studies. Habitus photographs were taken using a Canon macro photo lens MP-E 65mm on a Canon 550D. Observations, photographs and measurements of morphological details were performed using an Olympus BX53 microscope with an Olympus DP73. The final deep focus images were created with Zerene Stacker 1.04 stacking software. Adobe Photoshop CS6 was used for post processing. Exact label data are cited for specimens examined. Authors’ remarks and addenda are placed in square brackets, separate label lines are indicated by a slash (/), and separate labels by a double slash (//). Measurements are averaged over 5 specimens.
The material examined for this study is deposited in the following collections and museums:
Collection of Cheng-Bin Wang, Chengdu, Sichuan, China
Collection of Jan Růžička, Prague, Czech Republic
Collection of Masaaki Nishikawa, Ebina, Japan
Chungnam National University Insect Collection, Daejeon, Korea
Muséum National d’Histoire Naturelle, France, Paris (T. Deuve, Azadeh Taghavian)
The following abbreviations are used for the measurements in millimetres (mm):
(antennal length)
(basitarsal width)
(extended body length)
(elytral length)
(elytral width)
(eye width)
(head length)
(head width)
(pronotal length)
(pronotal width)
(tibial width)
Holarctic, north Oriental, north Neotropical.
Palaearctic, north Oriental.
Habitus (Fig.
Habitus of
Head transverse,
Pronotum (Fig.
Elytra oval, widest at about basal 1/4,
Prolegs robust, with basal three protarsomeres (Fig.
Abdominal ventrite VIII (Fig.
Aedeagus (Fig.
Furthermore, based on specimens examined,
The specific epithet is dedicated to Mr. Yasuhiko Hayashi (Kawanishi, Japan), a famous independent taxonomist on
Russia (Far East) (Fig.
Distribution map of
Habitus (Figs
Head quite transverse,
Pronotum (Fig.
Elytra oval, widest at about basal 1/3,
Prolegs robust, with basal three protarsomeres (Fig.
Abdominal ventrite VIII (Fig.
Aedeagus (Fig.
See under
Russia (Far East), South Korea (Fig.
We would like to express our sincere thanks to Kee-Jeong Ahn (Chungnam National University, Daejeon, Republic of Korea), Maxwell V. L. Barclay (