Corresponding author: Jan Růžička (
Academic editor: B. Guéorguiev
Wang C-B, Nishikawa M, Perreau M, Růžička J, Hayashi Y (2016) Revision of the genus
On the island of Taiwan, only three species in the subgenus
In this paper, which is a continuation of our revision of
Specimens were relaxed and softened in a hot saturated solution of potassium hydroxide for 4 minutes (for mounted dry specimens) or 8 minutes (for alcohol-preserved specimens), and then transferred to distilled water to rinse the residual potassium hydroxide off and stop any further bleaching. The softened specimens were moved into glycerine and dissected there to observe morphological details. After examination, the body parts were mounted on a glass slip with Euparal Mounting Medium for future studies. Habitus photographs were taken using a Canon macro photo lens MP-E 65mm on a Canon 550D. Observations, photographs and measurements of morphological details were performed using an Axio Zoom.V16 motorized stereo zoom microscope with an AxioCam MRc 5 in Beijing, or an Olympus BX53 microscope with an Olympus DP73 in Prague. The final deep focus images were created with Helicon Focus 5.3 stacking software in Beijing or Zerene Stacker 1.04 in Prague. The program Adobe Photoshop CS6 was used for post processing. Exact label data are cited for all specimens examined. Authors’ remarks and addenda are placed in square brackets, separate label lines are indicated by a slash (/) and separate labels by a double slash (//). Measurements are mean values based on 5 specimens.
The material examined for this study is deposited in the following collections and museums:
Collection of Cheng-Bin Wang, Chengdu, Sichuan, China
Collection of Jan Růžička, Prague, Czech Republic
Collection of Masaaki Nishikawa, Ebina, Japan
Collection of Michel Perreau, Paris, France
Collection of Yasuhiko Hayashi, Kawanishi, Japan
The following abbreviations are used for the measurements in millimetres (mm):
(antennal length)
(basitarsal width)
(extended body length)
(elytral length)
(elytral width)
(eye width)
(head length)
(head width)
(pronotal length)
(pronotal width)
(tibial width)
Holarctic, north Oriental, north Neotropical.
Palaearctic, north Oriental.
1 | Metathoracic wings absent; aedeagus large and strongly asymmetrical, median lobe turning to right at apex (Figs |
|
– | Metathoracic wings fully developed; aedeagus small and almost symmetrical, median lobe not turning to right at apex; spermatheca J-shaped, distal part simply curved |
|
2 | Elytral apices widely rounded; aedeagus stout and wide; spermatheca not coiled in proximal part |
|
– | Elytral apices narrowly rounded (Figs |
|
3 | Antennomere XI with length/width = 1.9 (Fig. |
|
– | Antennomere XI with length/width = 1.3 (Fig. |
|
Habitus of
Habitus of
Habitus of
Aedeagal apices of
Aedeagi of
Aedeagi of
Distribution map of
3♂♂, Taiwan, 25.V.1977 / Fenchihu [奮起湖, ca.
China (Taiwan) (Fig.
1♀, Mt. YUSHAN [玉山, ca.
This species is exceptional in the genus
In addition, the holotypes of
China (Taiwan) (Fig.
Central Taiwan, Nantou Hsien, Tsuifeng [翠峰, ca.
The specific epithet is dedicated to Cheng-Gong Zheng (1624–1662), a military leader at the end of the Chinese Ming Dynasty, for his feats in 1662 when he defeated the forces of the Dutch East India Company and claimed Taiwan, bringing it under Chinese Han rule.
China (Taiwan) (Fig.
Taiwan, Fushan [福山, ca.
See under
The specific epithet is dedicated to Miss Ting-Ting Song, the first author’s former colleague (
China (Taiwan) (Fig.
We would like to express our sincere thanks to Maxwell V. L. Barclay (BMNH, London, UK), Hideto Hoshina (Fukui