Corresponding author: Maxim Yu. Proshchalykin (
Academic editor: Thorleif Dörfel
The type specimens of the genus
Astafurova YuV, Proshchalykin MYu, Sidorov DA (2022) The bees of the genus
The present paper is the second part of a series of works dealing with the bee taxa described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko deposited in the
More than 140 years ago (1876), the second part of Ferdinand Morawitz’s critical study on the bees collected by Aleksey Fedtschenko 1869–1871 Expeditions in “Turkestan” was published. In the first volume, “
The genus
The nominal taxa of Fedtschenko’s
N | Species name | Sex | Current status | Depositaries of types |
---|---|---|---|---|
1. |
|
♀, ♂ | Valid | |
2. |
|
♀, ♂ | Valid | |
3. |
|
♀ | Valid | |
4. |
|
♂ | Synonym | HT ( |
5. |
|
♀, ♂ | Valid | |
6. |
|
♀ | Valid | HT ( |
7. |
|
♀ | Synonym | |
8. |
|
♀ | Valid | |
9. |
|
♀ | Valid | |
10. |
|
♀, ♂ | Valid | |
11. |
|
♂ | Synonym | |
12. |
|
♀ | Synonym | |
13. |
|
♀, ♂ | Valid | |
14. |
|
♀, ♂ | Valid | |
15. |
|
♀ | Valid | HT ( |
16. |
|
♀, ♂ | Valid | |
17. |
|
♀, ♂ | Valid | |
18. |
|
♀ | Valid | HT ( |
19. |
|
♀, ♂ | Valid | |
20. |
|
♀, ♂ | Valid | |
21. |
|
♀ | Valid | |
22. |
|
♀, ♂ | Valid | |
23. |
|
♂ | Synonym | |
24. |
|
♂ | Valid | |
25. |
|
♀, ♂ | Valid | |
26. |
|
♀ | Valid | |
27. |
|
♂ | Valid | |
28. |
|
♀, ♂ | Valid | |
29. |
|
♂ | Homonym | HT ( |
30. |
|
♀, ♂ | Valid | |
31. |
|
♀, ♂ | Valid | |
32. |
|
♀ | Valid | |
33. |
|
♀ | Valid | |
34. |
|
♀, ♂ | Valid | |
35. |
|
♀ | Valid | HT ( |
36. |
|
♀ | Valid | HT ( |
37. |
|
♀ | Valid | HT ( |
38. |
|
♀ | Synonym | |
39. |
|
♀ | Synonym | |
40. |
|
♀, ♂ | Valid | |
41. |
|
♀ | Valid | |
42. |
|
♀ | Valid | |
43. |
|
♀, ♂ | Valid | |
44. |
|
♂ | Synonym | |
45. |
|
♀, ♂ | Homonym | |
46. |
|
♀, ♂ | Valid | |
47. |
|
♂ | Homonym | |
48. |
|
♂ | Valid | |
49. |
|
♀, ♂ | Synonym | |
50. |
|
♂ | Synonym | HT ( |
51. |
|
♀, ♂ | Valid | |
52. |
|
♀, ♂ | Valid |
Since the 1940s, V. Popov [
Type material of Fedtschenko’s
In 1980 A.Z. Osytshnjuk continued the study of
The present paper is the first complete, critically reviewed, illustrated summary of all species of the genus
All of the material listed below was examined for this study. In the following list, the taxa are treated in alphabetical order of the names used in the original descriptions. Each entry includes the name of the taxon in its original combination, the complete reference to the original description of the species (including the original combination and spelling of the name and the author, year, and page of the description) and a list of type specimens present in the collections of the Zoological Museum of the Moscow State University, Moscow (
Photographs were made using a combination of a stereomicroscope Olympus SZX10 and a digital camera (Olympus OM-D). Illustrations were obtained by montaging from an image series that covers different focal planes into a single in-focus image with the program Helicon Focus 7. The final illustrations were post-processed for contrast and brightness using Adobe Photoshop software.
The classification and current species status mostly follow
Tashkent (Uzbekistan).
Uzbekistan: Tashkent, Urmitan.
♀, Ташкентъ [Uzbekistan, Tashkent,
Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan.
Samarkand (Uzbekistan).
Uzbekistan: Samarkand.
♀, designated by Osychnjuk et al. 2008: 51, 16.[III.1869] // Самаркандъ [Uzbekistan, Samarkand,
1 ♂, 16.[III.1876] // Самаркандъ // Paralectotypus
Uzbekistan.
Chardara (Kazakhstan).
Kazakhstan: Syr Darja River near Chardara.
♀, designated by
Description of male:
Turkmenistan, Uzbekistan, Tajikistan.
Central Asia.
“Turkestan”.
♂, 487. // Туркест.[анский] кр.[ай] [Turkestan] //
Europe, Russia (to East Siberia), Caucasus, Turkey, Israel, Iran, Afghanistan, Central Asia.
Panjakent (Tajikistan).
Uzbekistan: Tashkent, Samarkand, Oalyk Gorge; Tajikistan: Panjakent, Fan [River], Varzaminor [= Ayni], Iori Gorge.
♀, Пянджикентъ [Tajikistan, Panjakent,
There is one female specimen labelled by Warcke as “Lectotype” [31. // Заравш. дол. //
According to
Russia (European part), Caucasus, Iran, Central Asia, India.
Bairkum (Kazakhstan).
Kazakhstan: Bayrakum [= Bairkum].
♀, 4.[V.1871] // Байракумъ [Kazakhstan, Syr-Darya River, Bairkum,
Description of male:
Kazakhstan.
Tashkent (Uzbekistan).
Uzbekistan: Tashkent, Katty-Kurgan, Urmitan [near Katty-Kurgan]; Tajikistan: Panjakent.
♀, 11.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent,
Uzbekistan, Tajikistan.
Samarkand (Uzbekistan).
Uzbekistan: Samarkand.
♀, designated by
Description of male:
Uzbekistan, Tajikistan, Kazakhstan. The record from Russia (Western Siberia) by
Turkistan Province (Kazakhstan).
Kazakhstan: between the Karak Mts and Bairkum.
♀, designated by
Description of male:
Turkmenistan, Uzbekistan, Kazakhstan.
Kattakurgan (Uzbekistan).
Uzbekistan: Tashkent and near Kattakurgan.
♀, 29. [IV.1869] // Верхн.[ий] Заравш.[ан] [Uzbekistan, Upper Zaravshan, near Kattakurgan,
Azerbaijan, Turkey, Syria, Iran, Afghanistan, Uzbekistan, Tajikistan.
Tashkent (Uzbekistan).
Uzbekistan: Tashkent and Samarkand.
♂, 9.[III.1871] // Ташкентъ [Uzbekistan, Tashkent,
Listed as
Russia (European part), Caucasus, Iran, Central Asia, India.
Urmetan (Tajikistan).
Tajikistan: Urmitan.
♀, 2.[V.1869] // Урмитанъ [Tajikistan, Urmetan,
Uzbekistan, Tajikistan, Kazakhstan.
Tashkent (Uzbekistan).
Uzbekistan: Tashkent.
♀, 8.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent,
Uzbekistan, Tajikistan, Kazakhstan.
Chardara (Kazakhstan).
Kazakhstan: Kysyl-Kum [desert] near draw-well Chakany, Karak steppe, Chardara; Uzbekistan: Zeravshan River valley.
♀, Чардара [Kazakhstan, Chardara,
Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan.
Alai Mts, Kavuk Pass (Kyrgyzstan).
Kyrgyzstan: Kavuk Pass.
♀, 24.[VI.1871] // Алай [Kyrgyzstan, Alay Mts, Kavuk Pass,
Description of male:
Iran, Pakistan, Central Asia.
Tashkent (Uzbekistan).
Uzbekistan: Tashkent.
♀, designated by
Uzbekistan.
Tashkent (Uzbekistan).
Uzbekistan: Tashkent.
♂, designated by
The collection date in the original publication is “from 11 February to 11 March”. However, the lectotype and paralectotype specimens designated by A. Osytshnjuk are labelled as collected in April [green label]. Probably “March” was mistakenly mentioned in the Morawitz’ publication instead of April.
Turkmenistan, Uzbekistan, Tajikistan.
Sokh Enclave (Uzbekistan).
Uzbekistan: near Sokh.
♀, 28.[VI.1871] // Сохъ [Uzbekistan, Sokh Enclave,
Description of male:
The lectotype designation by
Iran, Turkmenistan, Uzbekistan, Tajikistan, Pakistan.
Ayni (Tajikistan).
Tajikistan: Varzaminor, near Iskander-Kul Lake; Kyrgyzstan/Tajikistan: Khodzha-Chiburgan River.
♂, designated by by
Afghanistan, Central Asia.
Shardara District (Kazakhstan).
Kazakhstan: “between Keles [River] and Kosaral [Lake]”.
♀, 24.[IV.1871] // Косаралъ [Kazakhstan, “Kosaral Lake” (now Shardara Reservoir), ≈
Male unknown. The single male from the type series belongs to
Uzbekistan, Kazakhstan.
Obburdon (Tajikistan).
Uzbekistan: Gus [near Urgut]; Tajikistan: Pyandzhikent, Obburden [Obburdon].
♀, 4.[VI.1870] // Оббурденъ [Tajikistan, Obburdon,
1 ♀, 24.[V.1869] // Заравш.[анская] дол.[ина] [Zaravshan River Valley] // Paralectotypus
Male unknown (according to
Uzbekistan, Tajikistan.
Shardara District (Kazakhstan).
Kazakhstan: “between Keles [River] and Kosaral [Lake]”, Kyzyl-Kum [desert]; Uzbekistan: Urmitan [near Katty-Kurgan]; Tajikistan: Varzaminor [Ayni], near Obburden [Obburdon].
♀, designated by by
Europe, Russia (to East Siberia), Caucasus, Turkey, Israel, Iran, Afghanistan, Central Asia.
Shardara (Kazakhstan).
Kazakhstan: Syr-Darja River, near Chardara.
♂, 25.[IV.1871] // Чардара [Kazakhstan, Shardara,
Description of female:
Southern Balkans, Cyprus, Turkey, Levant, Caucasus, Ukraine, Russia, Central Asia.
Tashkent (Uzbekistan).
Uzbekistan: Tashkent.
♂, 5.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent,
Female unknown (according to
Uzbekistan.
Samarkand (Uzbekistan).
Uzbekistan: near Samarkand.
♂, designated by
Central Asia.
Bairkum (Kazakhstan).
Kazakhstan: Bairakum, Karak steppe; Uzbekistan: Zeravshan River valley, Dzham Gorge, Oalyk Gorge between Oalyk and Aksay.
♀, designated by
Description of male:
Central Asia.
Ayni (Tajikistan).
Tajikistan: Varzaminor.
♂, designated by
Description of female:
Uzbekistan, Tajikistan, Kyrgyzstan, Kazakhstan, China.
Between Tashkent and Keless River (Uzbekistan).
Uzbekistan: between Tashkent and Keless River, Urmitan [near Katty-Kurgan]; Kazakhstan: “between Keles [River] and Kosaral [Lake]”, Kyzyl Kum [desert].
♀, 23.[IV.1871] // Келесъ [River] [Uzbekistan, between Tashkent and Keless River, ≈
Uzbekistan, Tajikistan, Kazakhstan.
Iskanderkul Lake (Tajikistan).
Tajikistan: Iskanderkul Lake.
♂, 18.[VI.1870] // Искандеръ [Tajikistan, Iskanderkul Lake, Hissar Ridge,
Description of female:
The lectotype designation by
Uzbekistan, Tajikistan, Kyrgyzstan, Kazakhstan.
Kyzylkum Desert (Kazakhstan).
Kazakhstan: Kyzyl-kum Desert; Karak mountains [western part of Kysyl-kum, NW Bairkum].
♀, 29.[IV.1871] // Кизилъкумъ [Kazakhstan, Kyzylkum Desert, NW of Shardara, ≈
1 ♂, 6.[V.1871] // Кизилъкумъ // Paralectotypus
Kazakhstan.
Tashkent (Uzbekistan).
Uzbekistan: Tashkent and Samarkand.
♀, designated by
Uzbekistan, Kazakhstan.
Tashkent (Uzbekistan).
Uzbekistan: near Tashkent.
♀, <golden circle> // Ташкентъ [Uzbekistan, Tashkent,
1 ♀, 5.[IV.1871] // Ташкент //
Description of male:
Europe, Russia (to the Urals), Turkey, Turkmenistan, Uzbekistan, Kazakhstan.
Khodzha-Chiburgan Gorge (Kyrgyzstan).
Kyrgyzstan: Khodzha-Chiburgan Gorge.
♀, <golden circle>, 21.[VI.1871] // Чибурганъ [Kyrgyzstan, Khodzha-Chiburgan Gorge, S Varuch, ≈
Description of male:
Uzbekistan, Kyrgyzstan, Tajikistan, Kazakhstan.
Tashkent (Uzbekistan).
Uzbekistan: near Tashkent.
♀, 23.[III.1871] // Ташкентъ [Uzbekistan, Tashkent,
There is a female specimen labelled by Warcke as a “Lectotypus” [<golden circle> // 5.[IV.1871] // Ташкент //
Uzbekistan, Kazakhstan.
Khodzha-Chiburgan River (Kyrgyzstan).
Kyrgyzstan: “Khodzha-Chiburgan River”.
♀, Чибурганъ [Kyrgyzstan: Khodzha-Chiburgan River/Gorge, S Varuch, ≈
Listed as
Kyrgyzstan.
Iskanderkul Lake (Tajikistan).
Tajikistan: near Iskander-Kul Lake.
♀, 15.[VI.1870] // Искандеръ [Tajikistan, Iskanderkul Lake, Hissar Ridge,
Listed as
Uzbekistan, Tajikistan, Kazakhstan.
Shakhimardan (Uzbekistan).
Uzbekistan: near Shakhimardan.
♀, 7.[VII.1871] // Шагимарданъ [Shakhimardan in the Uzbek enclave in the territory of Kyrgyzstan, Alai Ridge,
Description of male:
Tajikistan, Kazakhstan.
Shardara (Kazakhstan).
Kazakhstan: Chardara.
♀, <golden circle> // 25.[IV.1871] // Чардара [Kazakhstan, Shardara,
1 ♀, 25.[IV.1871] // Чардара //
There is a female specimen (in bad condition) labelled by Warcke as a “Lectotypus” [27.[IV.1871] // Чардара //
Southern Balkans, Cyprus, Turkey, Levant, Caucasus, Ukraine, Russia, Central Asia.
NW of Bairkum (Kazakhstan).
Uzbekistan: Katty-Kurgan; Kazakhstan: Karak steppe.
♀, 5.[V.1871] // Каракск.[ая] степь [Kazakhstan, Karak steppe, NW of Bairkum, ≈
Uzbekistan, Kazakhstan.
Anzob (Tajikistan).
Tajikistan: Anzob.
♂, designated by
♀, 75. // Туркест.[анский] кр.[ай] [Turkestan] // Paralectotypus,
Uzbekistan, Tajikistan, Kazakhstan.
30 km SE of Kozhatogai, Turkistan Province (Kazakhstan).
Kazakhstan: steppe between Tashkent and Syrdarya River.
♀, 20.[V.1871] // Степь м.[ежду] С.[ыр] [-] д.[арьей] и Т.[ашкентом] [Kazakhstan, Turkistan Province, steppe between Syrdarya River and Tashkent, 30 km SE of Kozhatogai,
Description of male:
Uzbekistan, Kazakhstan.
Keles River, Turkistan Province (Kazakhstan).
Kazakhstan: between Keles River and dry Keles; near Kosaral Lake; near Chardara.
♀, designated by
Listed as
Afghanistan, Uzbekistan, Kazakhstan, Mongolia.
Samarkand (Uzbekistan).
Uzbekistan: Samarkand.
♀, Самаркандъ [Uzbekistan, Samarkand,
Listed as
Uzbekistan, Tajikistan.
Samarkand (Uzbekistan).
Uzbekistan: Samarkand.
♂, 11.[IV.1869] // Самаркандъ [Uzbekistan, Samarkand,
There is a male specimen labelled by Warncke as a “Lectotypus” (3.[IV.1869] // Самарканд //
Europe, Russia (to the Urals), Turkey, Turkmenistan, Uzbekistan, Kazakhstan.
Tashkent (Uzbekistan).
Uzbekistan: near Tashkent.
♀, designated by
Uzbekistan.
Samarkand (Uzbekistan).
Uzbekistan: Samarkand.
♀, <golden circle> // 27.[II.1869] // Самаркандъ [Uzbekistan, Samarkand,
Uzbekistan.
Samarkand (Uzbekistan).
Uzbekistan: Samarkand.
♂, designated by
Iran, Turkmenistan, Uzbekistan, Tajikistan, Pakistan.
Tashkent (Uzbekistan).
Uzbekistan: Tashkent.
♂, 10.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent,
Listed as
Uzbekistan, Tajikistan.
Urgut, Zeravshan River valley (Uzbekistan).
Uzbekistan: Zeravshan River valley; steppe between Ulus and Dzham.
♀, Верхн.[ий] Заравш.[ан] [Uzbekistan, Zeravshan River valley, near Urgut,
Listed as “
Europe, North Africa, Russia (Dagestan Rep.), Caucasus, Turkey, Syria, Lebanon, Israel, Jordan, Iran, Afghanistan, Pakistan, Central Asia, India.
Urmetan (Tajikistan).
Tajikistan: near Urmitan [Urmetan].
♂, 2.[V.1869] // Урмитанъ [Tajikistan, Urmetan,
Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan.
Tashkent (Uzbekistan).
Uzbekistan: near Samarkand, near Tashkent, between Tashkent and Keless River.
♀, Ташкентъ [Uzbekistan, Tashkent,
Uzbekistan, Tajikistan.
Tashkent (Uzbekistan).
Uzbekistan: Tashkent, Samarkand.
♀, designated by
Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan.
We are grateful to A.V. Antropov (