Tree topology obtained by maximum likelihood in IQ-TREE v. 2.0. Numbers at nodes are bootstrap values; other conventions as in Fig. 1. The clade of the three new Luthela species (red font) is nested within Luthela (pink box). Further clades are other genera of Heptathelidae (Heptathela, Ryuthela, Qiongthela, Ganthela, Songthela, and Vinathela are from the bottom up).