Type material.
Holotype. “Mt. Maruyama / Sapporo-shi / <Hokkaido, JAPAN> / 6. VI. 1988 / M. Maruyama leg. / trail of ants” (LFcfF). Paratypes. 1♂, (ex type series of “Thiasophila rufescens”), “Japan. Lewis.” / “Sharp Coll 1905-313” (no other data) (NHM). Paratypes. 1 unsexed (ex type series of “Thiasophila rufescens”), same data but labelled “rufescens var.?” (NHM). Russia: Primorskyi krai: 1 unsexed, Kamenushka, Ussuryisk (MMLASRUS07), 28. V. 2005, M. Maruyama (LFO); 1 unsexed, same locality (MMLASRUS09), 28. V. 2005, M. Maruyama (LFO) (IBSS); 1 unsexed, 1♂, same locality (MMLASRUS11), 28. V. 2005, M. Maruyama (LFN); 1 unsexed, Bukhta Vityaz, Poluostrov Gamov, Khasanskyi (MMLASRUS14), 29. V. 2005, M. Maruyama (LFO); 2 unsexed, same locality (MMLASRUS17), 31. V. 2005, M. Maruyama (LFO) (IBSS); 2 unsexed, same locality (MMLASRUS19), 31. V. 2005, M. Maruyama (LFFJ); 6 unsexed, 2♀, Okeanskaya, Vladivostok (MMLASRUS23), 1. VI. 2005, M. Maruyama (LFO); 6 unsexed, 1♂, 1♀, Nadezhdinskyi, Khasanskyi, 1. VI. 2005, M. Maruyama (LFFJ) (IBSS); 2 unsexed, 3♂, Arisimovka, 70 km E Vladivostok, [43.11°N, 132.41°E], 5. VI. 1993, L. Zerche (LFO); 2 unsexed, Sibir or. Ussuri Vladivostok, 1919, Dr. Jureček. Korea: 8 unsexed, 1♂, Janggoksa, Chilgab-san, Cheongyang-gun, Chungnam, 23. VI. 2000, Shuhei Nomura (LFS); 70 unsexed, 1♂, Jeonglyeong Chi Samnae Myeon Cheonla-buk Do, 12. VII. 1991, S. Nomura (LFO); 18 unsexed, 3♂, 1♀, same locality, 12. VII. 1991, S. Nomura. Japan: Hokkaido: 23 unsexed, Hyakumatsu-zawa, Sapporo-shi, 8. VI. 1998, M. Maruyama (LFcfF); 50 unsexed, Hakken-zan, Sapporo-shi, 31. V. 2002, M. Maruyama (LFcfF); 54 unsexed, same locality, 31. VI. 2002, M. Maruyama (LFcfF); 17 unsexed, 1♀, same locality, 1. VI. 2002, M. Maruyama (LFO); 92 unsexed, 1♂, 1♀, same locality, 1. VI. 2002, M. Maruyama (LFN); 267 unsexed, 2♂, 2♀, same locality, 1. VI. 2002, M. Maruyama (LFcfF); 2 unsexed, Kannon-zawa, Sapporo-shi, 1. V. 2002, M. Maruyama; 6 unsexed, same locality, 20. V. 2002, M. Maruyama; 27 unsexed, same locality, 31. V. 2002, M. Maruyama (LFcfF); 4 unsexed, 1♂, 5♀, same locality, 1 VI 2002, M. Maruyama (LFcfF); 23 unsexed, Maruyama, Sapporo-shi, 6. VI. 1998, M. Maruyama (LFcfF) (same data as holotype); 3 unsexed, Hitsujigaoka, Sapporo-shi, 18. V. 2000, M. Maruyama (LFS); 2 unsexed, Nopporo, 17. VII. 1990, M. Ohara (HUM); 12 unsexed, Nopporo, Ebetsu-shi, 13. VI. 1999, M. Maruyama (LFcfF); 20 unsexed, 2♀, Ôsawaguchi, Nopporo-shinrin-kôen, Ebetsu-shi, 16–19. VI. 2001, S. Hori; 46 unsexed, same locality, 16–19. VI. 2001, S. Hori (LFO); 25 unsexed, same locality, 2. VI. 1999, M. Maruyama; 2 unsexed, Tomambetsu, Nopporo, Ebetsu-shi, 29. V. 2002, M. Maruyama (LFcfF); 1 unsexed, Nozaki, Bihoro-chô, 23. VI. 2001, Y. Kida; 1 unsexed, Taihei, Maruseppu-chô, 29–31. V. 2000, Y. Kida; 104 unsexed, 1♀, same locality, 16–17. VI. 2000, Y. Kida; 4 unsexed, same locality, 11–12. VIII. 2000, Y. Kida (LFcfF); 36 unsexed, same locality, 19–21. VIII. 2000, Y. Kida (LFcfF); 1 unsexed, 2♂, 1♀, same locality, 25. VIII. 2000, M. Maruyama (LFcfF); 1 unsexed, Makoi, Shari-chô, 27. V. 2000, Y. Kida; 2 unsexed, same locality, 28. V. 2000, Y. Kida; 53 unsexed, 1♂, same locality, 24. VI. 2002, Y. Kida; 4 unsexed, same locality, 13. VI. 2002, Y. Kida; 44 unsexed, Kanayama, same locality, 18–24. VI. 2002, Yasunari Kida; 7 unsexed, Midorioka-kôen, Kitami-shi, 17. VI. 2001, M. Maruyama; 3 unsexed, Hebinuma, Teshio-gawa, Teshio-chô, 9. VII. 1992, S. Hori; 2 unsexed, Kamishunbetu, 20. VII. 1977, Naomi; 2 unsexed, Yûtoku, Ôtaki-mura, 24. VIII. 2001, M. Maruyama; 4 unsexed, Mukôengaru Engaru-chô, 25. V. 2000, Y. Kida (LFcfF); 2 unsexed, Miwa, Koshimizu-chô, 28. VII. 2001, S. Kuwahara; 1 unsexed, Mt. Kariba, Shiribeshi, 13. VI. 1986, S. Nomura; 1 unsexed, Himenuma, Rishiri Is, 4. IX. 1990, T. Kishimoto. Honshû: Aomori-ken: 1 unsexed, Hirosaki, 3. VII. 1960, Y. Murakami; 2 unsexed, same locality, 5. VII. 1960, Y. Murakami. Miyagi-ken: 1 unsexed, Aoba-yama, Sendai-shi, 17. VII. 2004, K. Mizota; 6 unsexed, Naruko-onsen, Naruko-chô, 14–17. VI. 1999, M. Sano (LFO). Akita-ken: 7 unsexed, 1♀, Tamagawa, Tazawako-machi, 12–13. VI. 1999, M. Sano (LFcfF). Fukushima-ken: 6 unsexed, Mizuhiki, Toteiwa-mura, 12. VI. 2004, H. Kamezawa (cKam); 33 unsexed, 2♂, Kashi Spa., Nishigo-mura, 16. VI. 1998, M. Maruyama (LFcfF); 12 unsexed, same locality, 17. VI. 1998, M. Maruyama (LFcfF); 1 unsexed, Nanairi, Hinoemata, 25. VII. 1996, S. Naomi. Ibaraki-ken: 2 unsexed, Inohana Pass, 29. V. 1994, Y. Hagino. Tochigi-ken: 1 unsexed, Yumoto, Nikkô, 29. IV. 1982, S. Naomi; 1 unsexed, Ichikai-machi, Haga-gun, 29. IV.–2. V. 2002, Seidai Nagashima. Gunma-ken: 5 unsexed, Sakurayama, Onishi-chô, 9. V. 1998, Koji Toyoda (LFS); 5 unsexed, same locality, 9. V. 1998, Shiho Arai (LFS); 2 unsexed, same locality, 9. V. 1998, Shiho Arai (LJ); 1 unsexed, same locality, 22. V. 1999, Shiho Arai (LFS); 10 unsexed, Nageishi-tôge, Fujioka-shi, 5. VI. 2001, T. Watanabe; 8 unsexed, Nakanojô-forest-park, Nakanojô-machi, Agatsuma-gun, 8. VI. 2001, T. Watanabe. Saitama-ken: 3 unsexed, Shiro-yama, Kamagata, Ranazan-machi, 17. VI. 2000, K. Toyoda; 1 unsexed, 1♂, Ranzan-keikoku, Kamagata-mura, Ranzan-machi, 14. IV. 2000, K. Toyoda; 1 unsexed, same locality, 25. IV. 2000, K. Toyoda; 1 unsexed, same locality, 21. IV. 1999, K. Toyoda; 4 unsexed, Shioyama, Ranzan-machi, 8. V. 1998, K. Toyoda; 7 unsexed, same locality, 10. V. 1998, K. Toyoda; 11 unsexed, same locality, 10. V. 1998, K. Toyoda (LFcfF); 3 unsexed, 1♂, Toki-gawa, Kamagata-mura, Ranzan-machi, 25. IV. 1999, K. Toyoda; 5 unsexed, Shiroishi-toge, Higashichichibu-mura, 20. VI. 1999, Shiho Arai; 1 unsexed, same locality, 17. VI. 2000, K. Toyoda (LFS); 4 unsexed, Nr. Shiroishi Pass Higashichichibu-mura, 27. VI. 1998, K. Toyoda; 1 unsexed, Chichibu-kôgen, Higashichichibu-mura, 3. VII. 1999, K. Toyoda; 1 unsexed, same locality, 3. VII. 1999, K. Toyoda; 3 unsexed, Hashidake, Chichibu-shi, 17. VII. 1999, Shiho Arai (LJ); 1 unsexed, same locality, 26. IV. 1998, K. Toyoda (LFS); 1 unsexed, Koakazawa, (1,000 m in alt), Iri Kawa, Mt.Hakutai san, Ootaki-chiku, Chichibu-shi, 7. VI. 2002, Koji Toyoda; 1 unsexed, Hashidate Riv., Chichibu-shi, 7. VI. 2002, Koji Toyoda; 3 unsexed, Mt. Jyouminesan Minao-chô, 29. V. 1999, Shiho Arai (LFcfF); 1 unsexed, Nr. Ohno Pass Takigawa-mura, 5. VIII. 1998, K. Toyoda (LFS). Chiba-ken: 2 unsexed, Matuzaki, Inzai-shi, 3. V. 1991, T. Takeda. Tôkyô-to: 1 unsexed, Mt. Takao, 1. V. 1985, S. Kubota; 35 unsexed, 3♂, 2♀, Takao-san (450 m in alt.), Hachiôji-shi, 4. VI. 2001, M. Maruyama (LFS); 4 unsexed, Takao-san, Hachiôji-shi, 1. VII. 1998, M. Maruyama; 2 unsexed, same locality, 4. VII. 1998, M. Maruyama; 1 unsexed, same locality, 4. VII. 1998, M. Maruyama (LFS); 1 unsexed, Oyama, Machida-shi, 19. V. 1991, T. Kishimoto. Kanagawa-ken: 6 unsexed, Kawana, Fujisawa-shi, 14. V. 2001, T. Watanabe (LFS); 4 unsexed, Jimmu-ji, Zushi-shi, 20. VI. 2003, M. Maruyama; 1 unsexed, Toya, Tsukui, 29. IV. 1977, Ryo Kiryu (SCM); 1 unsexed, same locality, 14. V. 1976, Ryo Kiryu (SCM); 1 unsexed, Nanzawa, Tsukui, 14. V. 1977, Ryo Kiryu (SCM); 1 unsexed, Hasuge-san, Aikô, 3. V. 1963, Ryo Kiryu (SCM); 1 unsexed, Minami-Ashigara, 11. II. 1975, Y. Hirano; 2 unsexed, Fudakake, Higashi-Tanzawa, 16. VI. 1984, Y. Hirano; 1 unsexed, Mt. Tanzawa, 26. II. 1973, Y. Hirano; 1 unsexed, Kawasaki, 1. V. 1985, 1. V. 1985; 1 unsexed, Kozuka-yama, Hakozaki-chô, 25–26. V. 1995, Yoshinori Kaneko, Akio Ito, Satoshi Tsuboyama; 6 unsexed, 1♂, Niiharu-shiminno-mori, Yokohama-shi, 21. V. 2001, T. Watanabe (LFO). Niigata-ken: 2 unsexed, Yuzawa-machi, Minami-uonuma-gun, 20–22. VI. 2002, S. Nagashima (LFS); 2 unsexed, Tairai, 3. VII. 1985, S. Nomura. Ishikawa-ken: 2 unsexed, Ichinose pass Shiramine-Village, 25. V–7. VI.2003, Katsuyuki Nakata (fit). Yamanashi-ken: 8 unsexed, Hikawarindô, Daibosatsu-rei, Koushû-shi, 24. VI. 2001, T. Watanabe; 4 unsexed, Ashiyu-onsen, Ashiyu-mura, 23. V. 2001, T. Watanabe (LFcfF); 1 unsexed, same locality, 24. V. 2001, T. Watanabe; 3 unsexed, Shiroidaira Dôshi-mura, 17.VII.2001, T. Watanabe; 4 unsexed, Gozaishi-kôsen (900 m in alt.), Nirasaki-shi, 25. VII. 2002, S. Nomura (LFN); 2 unsexed, Karumizu-rindô, (1400 m in alt.), Narusawa, 29. VI. 2011, T. Watanabe. Nagano-ken: 1 unsexed, Kakuma Valley, Sanada-machi, 17. VI. 2001, T. Watanabe; 1 unsexed, Midarebashi, Honjô-mura, 21. VII. 1999, T. Watanabe; 52 unsexed, Penshionmura, Hara-mura, 11. VII. 1999, Shiho Arai (LFO); 2 unsexed, Otari-onsen (1100 m in alt.), Otari-mura, Kita-azumi-gun, 16. V. 2004, H. Kamezawa (cKam); 2 unsexed, Fujii, Satoyamabe, Matsumoto-shi, 9. VI. 2004, T. Komatsu; 1 unsexed, same locality, 11. VI. 2004, T. Komatsu; 1 unsexed, same locality, 21. VI. 2004, T. Komatsu. Gifu-ken: 6 unsexed, Ogamigo (800 m in alt.) Shôkawa-mura, 5–6. VIII. 1998, M. Maruyama (LFS); 4 unsexed, Mt. Kinkazan, Gifu-shi, 11. VI. 2003, K. Kinomura (LFS); 1 unsexed, Isshiki, Shôkawa-chô, Takayama-shi, 1. VIII. 2004, K. Kinomura (LFN); 51 unsexed, Isshiki, Shôkawa-chô, Takayama-shi, 8. VI. 2013, K. Kinomura (LFM); 29 unsexed, Nabedaira-kôgen, Kansaka, Okuhida-onsenkyô, Takayama-shi, 30. VI. 2013, K. Kinomura (LFN); 20 unsexed, Nabedaira-kôgen, (1305 m in alt.) Kansaka, Okuhida-onsenkyô, Takayama-shi, (1305 m in alt.), 30. VI. 2013, K. Kinomura (LFN). Wakayama-ken: 1 unsexed, Mt. Gomandan, 22–23. VI. 1981, S. Naomi. Tottori-ken: 1♂, Mt. Daisen, 3–5. VI. 1980, S. Naomi; 3 unsexed, same locality, 10. VI. 1986, S. Nomura (LFS); 10 unsexed, 1♂, Urahikimi, Shimane-ken, 6. VI. 1988, S. Nomura (LFS). Okayama-ken: 2 unsexed, Ono Shine, Kawakami, Ohara-chô, Mimasaka-shi, 5. V. 2009, Yoshifumi Fujitani; 1 unsexed, same locality, 24. V. 2009, Yoshifumi Fujitani; 1 unsexed, same locality, 7. VI. 2009, Yoshifumi Fujitani. Hiroshima-ken: 11 unsexed, Nakatsuya, Hatsukaichi-shi, 7. VI. 1987, S. Nomura; 1 unsexed, Mt. Garyu, 27. VI. 1987, S. Nomura (LFcfF). Yamaguchi-ken: 10 unsexed, Nishimagura, (100 m in alt.), Kusunoki, 30. V. – 1. VI. 2000, Toshio Kishimoto; 3 unsexed, Momijidani park, Yokoyama, Iwakuni-shi, 28. V. 2011, M. Shimono (LFS). Shikoku: Kagawa-ken: 5 unsexed, Furodani, Miki-chô, 30. VII. 2000, K. Izawa (LFcfF); 108 unsexed, 3♂, 1♀, Ôtaki-san, Shionoe-chô, 2. VI. 2001, M. Maruyama et al. (LFN); 21 unsexed, Atago-yama, Kotohira-chô, 1. VI. 2001, M. Maruyama et al. (LFS); 22 unsexed, Fujio-jinja, Nishi-ueta-chô, Takamatsu-shi, 31. V. 2001, M. Maruyama et al. (LFN); 2 unsexed, same locality, 1. VI. 2001, M. Maruyama et al. (LFN); 22 unsexed, Daisen-zan, Kotonami-chô, 1. VI. 2001, M. Maruyama et al. (LFM); 13 unsexed, Usa-jinja, Nagaona, Sanuki-shi, 31. V. 2001, M. Maruyama et al. (LFS); 3 unsexed, Shirahige-jinja, Ayakami-chô, 25. IV. 2004, H. Fujimoto (LFM). Ehime-ken: 1♂, Sugitate, 17. VI. 2017, Yu Hisasue; 2 unsexed, Waki-ga-fuchi Park (120–220 m in alt.), Sue-machi, Matsuyama-shi [33.87°N, 132.83°E], 4. V. 2017, Yu Hisasue (Tullgren); 1 unsexed, Mt. Ishizuchi (1400 m in alt.), Saijô-shi, 29. VI. 2014, Yu Hisasue; 1 unsexed, Minamikume-machi, Matsuyama-shi [33.8333°N, 132.8184°E], 25. IV. 2015, Yu Hisasue (Tullgren). Kyûshû: Fukuoka-ken: 1 unsexed, Hakozaki, Fukuoka-shi, 12. V. 1979, K. Yamagishi; 2 unsexed, 1♂, Hikosan, Biol. Lab. KU (750 m in alt.), Soeda-machi, 22. V. 2011, M. Maruyama (LFcfF). Saga-ken: 2 unsexed, Mt. Mifume, 15. V. 2019, S. Nomura; 2 unsexed, Mt. Kurokami-yama (400 m in alt.), Takeo-shi [33.2130°N, 129.9044°E], 9. V. 2021, Yu Hisasue (LFcfF); 4 unsexed, 1♂, Kurokami-yama, Arita-chô, 15. V. 2019, Mitsuyasu Nishida. Nagasaki-ken: 3 unsexed, 1♂, Tanukinoo, Masuragahara-machi, Ômura-shi, 5. V. 2018, Mitsuyasu Nishida; 1 unsexed, Todoroki Valley, 1. VI. 1987, S. Naomi; 1♂, Suwa Shine, Nagasaki-shi, 2. V. 1985, S. Nomura; 1 unsexed, Tomikawa-keikoku, Isahaya-shi, 28. IV.–1. V. 2008, T. Iwai.