Shells of Pupina arula species group from mainland Southeast Asia A–G Pupina peguensis A syntype of Pupina blanfordi NHMUK 1888.12.4.100 and specimens B CUMZ 12105 C CUMZ 12050 D CUMZ 12103 E NHMUK ex. Cuming coll. F CUMZ 12094, and G CUMZ 12108 H–J Pupina crosseana H lectotype MNHN-IM-2000-35834 I paralectotype RBINS MT966/10591, and J specimen CUMZ 12049 K, L Pupina siamensis K lectotype SMF 109948 and L specimen CUMZ 12052 M–P Pupina bilabiata sp. nov. M holotype CUMZ 12073/1 and specimens N CUMZ 12079 O CUMZ 12081, and P CUMZ 12086 Q, R Pupina godwinausteni sp. nov. Q holotype CUMZ 12090/1 and R paratype CUMZ 12090/2. Photo: P. Maestrati, MNHN (H), F. Trus, RBINS (I).