Tortulosa tortuosa A lectotype of Perlisia tweediei NHMUK 1948.10.2.6 from Kaki Bukit, Perlis B holotype of Tortulosa huberi MNHN-IM-2000-34054 from Krabi C holotype of Tortulosa schileykoi MNHN-IM-2000-34055 from Phang Nga D specimen CUMZ 12166 from Tham Suea Temple, Krabi, and E–H operculum of specimen CUMZ 12160 from Tham Kanlayanamit Temple, Nakhon Si Thammarat, showing E outer operculum F inner operculum G side view (inner surface up), and H spring-like inner operculum when extended by force. Photo: M. Caballer, MNHM (B, C).

 
 
  Part of: Jirapatrasilp P, Sutcharit C, Panha S (2022) Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia. ZooKeys 1119: 1-115. https://doi.org/10.3897/zookeys.1119.85400